taxonID	type	description	language	source
F82E87F36228D52F6EA24AA8FD8DFEE2.taxon	materials_examined	Type species: Sciara bruckii Winnertz, 1867 [Winnertz (1867): 38 – 39]. Literature: Menzel & Mohrig (1998): 367 – 369; Menzel & Mohrig (2000): 464 – 480; Mohrig (2003): 34 – 37; Mohrig et al. (2004): 277 – 286. Mohrig et al. (2013): 225 – 231.	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F36228D52F6EA24AA8FD8DFEE2.taxon	materials_examined	Type species: Ostroverkhovana borealis Komarova, 2002 [Komarova (2002): 27 – 29; figs. 1 – 6].	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F36228D52F6EA24AA8FD8DFEE2.taxon	discussion	Comments. Heller (2012) identified the type species of the genus Ostroverkhovana Komarova, 2002 as a species of Pseudolycoriella, similar or identical to Pseudolycoriella nodulosa (Mohrig & Krivosheina, 1985). We did not study the type but we agree with Heller (2012) that the species Ostroverkhovana borealis Komarova belongs to the genus Pseudolycoriella. However, we do not agree with Heller (2012) that the presence of a lobe at the ventral base of the hypopygium is sufficient for the foundation of a new subgenus. Intergonocoxal structures appear in species of Pseudolycoriella as lobes with or without bristles. Diagnostic characters. The genus is characterized as follows: bristles on flagellomeres have mostly large basal pits but sometimes (only in a few species) a net-like surface structure; mouth parts are usually short, sometimes somewhat elongated (1 / 3 of the height of the head); palpi with three segments, basal segment slender, with more than one bristle, without a deepened area of sensillae; scutum and scutellum are mostly densely haired; wings are not reduced, except for Psl. semialata (Edwards, 1913) and with a typical Sciaridae venation; posterior veins without macrotrichia in almost all species; c longer than w; the tibial organ of the fore tibia is variable in shape, often with a semicircular border; claws are usually toothed; gonostylus is variable in shape, always without an apical tooth; in most species with two more subapically inserted spines of different lengths (in some species with one spine only, in others with three or more spines at the apex of the gonostylus) and with one (occasionally with 2 – 3 or none) subapical whiplash hairs, directed towards the base of the hypopygium; whiplash hair (hairs) are as long as or longer than spines and inserted near the apex of the gonostylus (not in or below the middle like in Lycoriella and Mohrigia); the ventral base of the hypopygium (intergonocoxal space) is mostly without a lobe or a distinct bristle patch; tegmen is variable in shape with or without a finger or horseshoe-like inner structure; the aedeagus usually has a large furca.	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F3622BD52E6EA24820FD8AFD86.taxon	description	(Fig. 1 A – C; Plate II, Fig. IV) Locus typicus: USA, Arizona, Cochise County, Huachuca Mts., Ash Cyn. Road, 31.39 ° N, 111.24 ° W.	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F3622BD52E6EA24820FD8AFD86.taxon	materials_examined	Holotype: Male, 24. ix. – 5. x. 1993, oak-pine woodland, leg. N. McFarland, Malaise trap (PWMP). Paratypes: 3 males, same data as holotype (PWMP); 3 males, 5. – 30. viii. 1993, same locality; 2 males, 13. – 22. ix. 1993, same locality (PWMP, 1 male PKHH).	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F3622BD52E6EA24820FD8AFD86.taxon	description	Description. Male. Head: Round; eye bridge 3 facets wide. Antenna brown, scape and pedicel paler; 4 th flagellomere with a l / w index of 1.8 – 2.0, brown, hairs shorter than the flagellomere wide, with a few deepened sensory pits. Palpi 3 - segmented, first segment with 3 – 4 bristles and a flat patch of sensillae. Mouth parts short, 1 / 3 of head height. Thorax: Brownish; scutum with rather long and dark hairs, some lateral and prescutellar bristles longer. Scutellum with 4 robust bristles. Postpronotum bare. Wing brownish; R 1 = 1 / 2 R; c = 2 / 3 w; y = x, both bare; M-fork as long as M-stem, narrow; CuA-stem somewhat shorter than x; posterior veins distinct, without macrotrichia. Haltere short, brown. Legs yellowish; tibial organ of fore tibia with a small patch of stronger bristles, not bordered. Claws finely toothed. Abdomen: Ventral base of hypopygium without lobe; inner ventral margin of gonocoxites with very long and strong bristles, especially dense basally. Gonostylus apically round, apex densely haired, subapically with 2 long spines and a long whiplash hair. Tegmen as wide as high, apically rounded. Aedeagus short, with large furca. Body length: 2.2 mm.	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F3622BD52E6EA24820FD8AFD86.taxon	discussion	Comments. The species is characterized by long and strong bristles on the ventral inner margin of the gonocoxites and two long subapical spines on the gonostylus. It is similar to Psl. longisetosa sp. n. It differs from it by possessing dense bristles in the basal half of the gonocoxites.	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F3622BD52E6EA24820FD8AFD86.taxon	distribution	Distribution. USA (Arizona).	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F3622AD52E6EA24C1DFDFBFA81.taxon	description	(Fig. 2 A – E; Plate IV, Fig. VII) Locus typicus: USA, Texas, Sam Houston National Forest, Willis, 60 km N of Houston.	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F3622AD52E6EA24C1DFDFBFA81.taxon	materials_examined	Holotype: Male, 3. – 6. xi. 1996, leg. W. Mohrig, yellow pan trap (PWMP). Paratypes: 2 males and 1 female, same data as holotype (PWMP); 1 male, 21. – 23. vii. 1997, USA, Virginia, suburban settlement of Williamsburg, mixed forest, yellow trap, leg. W. Mohrig, (PWMP).	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F3622AD52E6EA24C1DFDFBFA81.taxon	description	Description. Male. Head: Round; eye bridge 2 – 3 facets wide. Antenna uniformly brownish, 4 th flagellomere with a l / w index of 1.4, hairs shorter than the flagellomere width, flagellomere with a few deepened sensorial pits. Palpi short, 3 - segmented, first segment with 1 – 2 bristles and a flat patch of sensillae. Mouth parts short. Thorax: Brownish; scutum with rather short and dark hairs, some lateral and prescutellar bristles longer. Scutellum with 4 more robust bristles. Postpronotum bare. Wing pale, short and wide; R 1 = 1 / 2 R; c = 3 / 4 w; y shorter than x, bare; M-fork as long as M-stem, narrow; CuA-stem shorter than x; posterior veins distinct, without macrotrichia. Haltere short, brown. Legs yellowish; tibial organ of fore tibia with an irregular row of bristles, not bordered. Claws finely toothed. Abdomen: Ventral base of hypopygium without lobe, inner margin of gonocoxites with short sparse hairs. Gonostylus apically rounded, densely haired at the apex, subapically with 2 fine spines not longer than the apical hair and with 1 – 2 short whiplash hairs. Tegmen as wide as high, apically rounded with a horseshoe-like structure. Aedeagus short with a large furca. Body length: 2.5 mm.	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F3622AD52E6EA24C1DFDFBFA81.taxon	discussion	Comments. The species is characterized by a rounded and densely haired apex of the gonostylus with two short subapical spines and 1 – 2 short whiplash hairs.	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F3622AD52E6EA24C1DFDFBFA81.taxon	distribution	Distribution. USA (Texas, Virginia).	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F3622AD52E6EA24B00FD8AF86C.taxon	description	(Fig. 3 E – G) Locus typicus: USA, Arizona, Pima County, Tucson Mountains, 16 km W Tucson, Sonora desert, 32.24 ° N, 111.13 ° W.	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F3622AD52E6EA24B00FD8AF86C.taxon	materials_examined	Holotype: Male, 28. ix. – 7. x. 1995, leg. S. J. Prchal, Malaise trap (PWMP).	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F3622AD52E6EA24B00FD8AF86C.taxon	description	Description. Male. Head: Round; eye bridge 3 facets wide. Antenna brownish, scape and pedicel somewhat paler; 4 th flagellomere with a l / w index of 1.8 – 2.0, hairs somewhat shorter than the diameter, inserted in rather large pits. Palpi 3 - segmented, first segment with 3 – 4 bristles and a flat patch of sensillae. Mouth parts very short. Thorax: Brown; scutum with dark hairs, some lateral and prescutellar bristles longer. Scutellum with 2 more robust bristles. Postpronotum bare. Wing brownish; R 1 = 1 / 2 R; c somewhat longer than 1 / 2 w; y = x, both bare; Mfork shorter than M-stem, rather wide; CuA-stem = x; posterior veins distinct, without macrotrichia. Haltere short, brown. Legs yellowish; tibial organ with a small patch of stronger bristles, distinctly bordered. Claws finely toothed. Abdomen: Inner margin of gonocoxites with a few longer bristles between short hairs. Gonostylus wide and oval in shape, inner side concave, with dense apical hairs, 3 long spines, and 2 – 3 whiplash hairs that are as long as the spines. Tegmen apically rounded. Aedeagus rather long, with a large furca. Body length: 2.2 mm.	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F3622AD52E6EA24B00FD8AF86C.taxon	discussion	Comments. The species resembles Psl. basisetosa sp. n. but differs from it by having a broader gonostylus with three much longer spines, a few longer bristles on the entire ventral inner margin of the gonocoxites and a distinctly bordered tibial organ apically on the fore tibia.	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F3622AD52E6EA24B00FD8AF86C.taxon	distribution	Distribution. USA (Arizona).	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F3622ED52A6EA24EC6FD80FCE6.taxon	description	(Fig. 3 A – D; Plate II, Fig. III) Locus typicus: USA, Florida, Highlands County, Archibald Biological Station, 13 km SO of Lake Placid.	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F3622ED52A6EA24EC6FD80FCE6.taxon	materials_examined	Holotype: Male, 22. – 29. vii. 1998, Malaise trap leg. D. Wahl & F. Gonsalves (PWMP). Paratypes: 9 males, same data as holotype (PWMP, 1 male in PKHH).	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F3622ED52A6EA24EC6FD80FCE6.taxon	description	Description. Male. Head: Round; eye bridge 3 facets wide. Antenna brownish, scape and pedicel somewhat paler; 4 th flagellomere with a l / w index of 2.0, brownish, hairs not shorter than the diameter, with a few deepened sensorial pits. Palpi 3 - segmented, first segment with 2 bristles and a flat patch of sensillae. Mouth parts very short. Thorax: Brownish, pleural sclerites with yellow spots; scutum rather long and with dark hairs, some lateral and prescutellar bristles longer. Scutellum with 4 stronger bristles, two are long. Posterior pronotum bare. Wing brownish; R 1 = 1 / 2 R; c = 2 / 3 w; y = x, bare; M-fork as long as M-stem, narrow; CuA-stem as long as x; posterior veins distinct and without macrotrichia. Haltere short, brownish. Legs yellowish brown; tibial organ with an undifferentiated patch of strong bristles, not bordered. Claws toothless. Abdomen: Ventral base of hypopygium without lobe and bare; inner margin of gonocoxites with short sparse hairs. Gonostylus apically rounded, with dense hairs apically, subapically with 2 short dark spines, not longer than apical hairs and with a long whiplash hair. Tegmen as wide as high, apically rounded. Aedeagus short and thin. Body length: 2.0 mm.	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F3622ED52A6EA24EC6FD80FCE6.taxon	discussion	Comments. The species is characterized by having two short dark subapical spines as long as the apical hairs, a long whiplash hair, an undifferentiated tibial organ and claws without teeth.	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F3622ED52A6EA24EC6FD80FCE6.taxon	distribution	Distribution. USA (Florida).	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F3622ED52A6EA24DE5FBABFA82.taxon	description	(Fig. 4 A – C; Plate I, Fig. I)	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F3622ED52A6EA24DE5FBABFA82.taxon	discussion	Comment. The species is characterized by having an intergonocoxal lobe on the ventral base of the hypopygium, densely covered with long, lanceolate hyaline bristles. The apex of the gonostylus is broadly rounded, with a few fine hyaline spines in between hairs. It resembles Psl. subjucunda sp. n.	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F3622ED52A6EA24DE5FBABFA82.taxon	distribution	Distribution. USA (Arizona, New York, Rhode Island, Virginia, Washington).	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F3622ED52A6EA24B09FC49F976.taxon	description	(Fig. 5 A – D; Plate III, Fig. V)	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F3622ED52A6EA24B09FC49F976.taxon	discussion	Comments. The species is characterized by having a basal lobe on the ventral side of the hypopygium, densely covered with macrotrichia. The gonostylus is strongly curved with two long spines and a whiplash hair shorter than spines. It seems to be closely related to the Palaearctic species Pseudolycoriella nodulosa (Mohrig & Krivosheina, 1985), but it does not have a hyaline spine in the middle of the gonostylus, instead it has just a patch of short fine bristles.	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F3622ED52A6EA24B09FC49F976.taxon	distribution	Distribution. USA (Arizona, California); Canada (British Columbia).	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F36220D5276EA24BABFD8AFEB1.taxon	description	(Fig. 1 D – E) Locus typicus: USA, Arizona, Cochise County, Huachuca Mountains, Ash Canyon Road, 31.39 ° N, 111.24 ° W.	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F36220D5276EA24BABFD8AFEB1.taxon	materials_examined	Holotype: Male, 22. – 31. x. 1993, oak-pine woodland, Malaise trap, leg. N. McFarland, (PWMP).	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F36220D5276EA24BABFD8AFEB1.taxon	description	Description. Male. Head: Round; eye bridge 3 facets wide. Antenna brown; 4 th flagellomere with a l / w index of 2.2, brownish, with hairs as long as the diameter. Palpi 3 - segmented, long; first segment with 5 – 6 bristles and a flat patch of short sensillae. Mouth parts short. Thorax: Brown, scutum with short sparse hairs, some lateral and prescutellar bristles longer. Scutellum with 4 stronger bristles. Postpronotum bare. Wing brownish; R 1 = 2 / 3 R; c longer than 1 / 2 w; y = x, both bare; M-fork as long as M-stem, narrow; CuA-stem shorter than x; posterior veins distinct and without macrotrichia. Haltere short, brownish. Legs yellowish brown; tibial organ small, with row like bristles, weakly bordered. Claws toothed. Abdomen: Ventral base of hypopygium without lobe and bare; inner margin of gonocoxites with short sparse hairs in the proximal half and densely covered with long strong bristles in the distal half. Gonostylus broadly ovoid, densely haired apically, subapically with 3 fine hyaline spines and a long whiplash hair. Body length: 3.5 mm.	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F36220D5276EA24BABFD8AFEB1.taxon	discussion	Comments. The species is characterized by having a large ovoid gonostylus with three fine hyaline spines, a longer whiplash hair and unusually long strong dense bristles in the distal half of the gonocoxites. It is similar to Psl. chlorothoracica sp. n., but differs from it by possessing long and strong bristles in the apical half of the gonocoxites, shorter spines on the gonostylus and uniformly brown antennae.	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F36220D5276EA24BABFD8AFEB1.taxon	distribution	Distribution. USA (Arizona).	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F36223D5276EA24FD0FDA1FBDA.taxon	description	(Fig. 6 A – B) Locus typicus: Canada, Alberta, Munn Creek, 53.30 ° N, 118.10 ° W.	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F36223D5276EA24FD0FDA1FBDA.taxon	materials_examined	Holotype: Male, 11. vi. – 24. vii. 1994, spruce forest, Malaise trap, leg. E. Fuller, (PWMP).	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F36223D5276EA24FD0FDA1FBDA.taxon	description	Description. Male. Head: Round; eye bridge 4 – 5 facets wide. Antenna brown; 2 nd flagellomere with a l / w index of 2.0, hairs pale, as long as the diameter, no deepened sensorial pits. Palpi 3 - segmented, long, brown; first segment with 5 – 6 bristles and a flat patch of sensillae. Mouth parts rather short. Thorax: Dark brown, scutum long and densely haired. Scutellum with 4 stronger bristles. Postpronotum bare; mediotergite with a few bristles. Wing brownish; R 1 = 2 / 3 R; c = 1 / 2 w; y longer than x, x bare and y with 3 – 4 macrotrichia; M-fork shorter than M-stem, rather wide; CuA-stem longer than x; posterior veins distinct and without macrotrichia. Haltere brown and rather long. Legs brown; tibial organ indistinct with a few short bristles, nearly comb-like, not bordered. Claws toothless. Abdomen: Ventral base of hypopygium without lobe, inner margin of gonocoxites with rather strong but not dense hairs. Gonostylus in profile (ventro-lateral aspect) rather narrow, apex densely haired, inner side broad and somewhat spoon-like; subapically with a few (3 – 4) fine, hyaline spines within the hairs and a long whiplash hair, inserted subapically. Body length: 4.0 mm.	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F36223D5276EA24FD0FDA1FBDA.taxon	discussion	Comments. The species is characterized by having a uniformly dark brown body colour, macrotrichia on the apical part of y, a gonostylus that is broad on the inner side (spoon-like), a few bristles on the mediotergite (unusual for Pseudolycoriella), toothless claws and a weak, somewhat comb-like tibial organ on the apex of the fore tibia. It resembles Psl. planiforceps (Steffan), but differs by having a darker body colour, a long whiplash hair and a weakly structured tibial organ.	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F36223D5276EA24FD0FDA1FBDA.taxon	distribution	Distribution. Canada (Alberta).	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F36223D5276EA24AF9FAD1FA40.taxon	description	(Fig. 7 B; Plate III, Fig. VI)	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F36223D5276EA24AF9FAD1FA40.taxon	discussion	Comments. The species is characterized by having an elongate gonostylus, narrowed to the apex, with two long robust apical spines and a somewhat longer whiplash hair. Distribution. Costa Rica; Cuba; Dominican Republic; Puerto Rica; USA (Alabama, Florida, Kansas).	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F36223D5276EA24843FE84F830.taxon	description	(Fig. 8 A – D; Plate IV, Fig. VIII)	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F36223D5276EA24843FE84F830.taxon	discussion	Comments. The species shows characters that are typical for the genus Pseudolycoriella like the shape of the tegmen, a semicircular-bordered tibial organ on the apex of the fore tibia and flagellomeres with large cuticular pits at the bases of bristles. The species is unique since it does not have a whiplash hair and typical apical spines on the gonostylus. Instead the gonostylus is wide and spoon-like in shape, having many fine spines in between short spine-like bristles along the inner side. The shape of the gonostylus is similar to Psl. nocturna.	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F36223D5276EA24843FE84F830.taxon	distribution	Distribution. USA (Arizona, Texas). Also known from Ecuador (Galapagos Islands) and Mexico (Baja California).	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F36224D5236EA248C1FD8AFF7F.taxon	description	(Fig. 7 A)	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F36224D5236EA248C1FD8AFF7F.taxon	materials_examined	Further material: (not mentioned in Mohrig et al. (2013): 1 male, 13. – 22. ix. 1993, USA, Arizona, Cochise County, Huachuca Mts., Ash Cyn. Rd. 31.39 ° N, 1.24 ° W, oak-pine woodland, Malaise trap, leg. N. McFarland, (PWMP).	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F36224D5236EA248C1FD8AFF7F.taxon	discussion	Comments. The species is characterized by having dense short bristles on the inner ventral bases of the gonocoxites, an elongate gonostylus densely covered with short hairs at the apex, with one spine and one long whiplash hair. It is similar to Pseudolycoriella curviseta, Mohrig from Costa Rica (Mohrig 2003). Distribution. USA (Arizona).	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F36227D5226EA24951FD58FCCA.taxon	description	(Fig. 9 A – D; Plate I, Fig. II) Locus typicus: USA, Pennsylvania, Spring Brook.	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F36227D5226EA24951FD58FCCA.taxon	materials_examined	Holotype: Male, 1945, leg. F. R. Shaw (PWMP). Paratypes: 2 males, same data as holotype (PWMP); 1 male, 30. iii. 2002, USA, Alabama, Baldwin County, Byrnes Lake, 30.47 ° N, 87.54 ° W, Malaise trap, leg. J. W. McCreadie, (PWMP).	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F36227D5226EA24951FD58FCCA.taxon	description	Description. Male. Head: Round; eye bridge 3 facets wide. Antenna brownish; 4 th flagellomere with a l / w index of 2.0 and pale hairs as long as the diameter of the basal node. Palpi 3 - segmented, rather short; first segment with 4 – 5 bristles and a flat patch of sensillae. Mouth parts short. Thorax: Brown, scutum with long dense hairs. Scutellum with 5 – 6 robust bristles. Postpronotum bare; mediotergite sometimes with a few short bristles. Wing brownish; R 1 nearly as long as R; c = 3 / 4 w; y = x, x bare and y without or with a few macrotrichia; M-fork as long as M-stem, narrow; CuA-stem shorter than x; posterior veins distinct and without macrotrichia. Haltere short, brownish. Legs brownish; tibial organ of the fore tibia not visible. Claws toothless. Abdomen: Tergites rather long and densely haired. Ventral base of hypopygium with a large intergonocoxal lobe, covered with long dark bristles. Gonostylus short, broadly oval, apex densely haired; subapically with 2 short dark spines, not longer than the apical hair and with a longer whiplash hair. Tegmen somewhat broader than high, apically rounded. Aedeagus short and robust. Body length: 5 mm.	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F36227D5226EA24951FD58FCCA.taxon	discussion	Comments. The species is characterized by having a large intergonocoxal lobe covered with long dark bristles. It is similar to Psl. jucunda regarding this character. It differs from it by having shorter apical spines on the gonostylus, long fine bristles on the basal lobe of the hypopygium, a differently shaped tegmen and bristles only in the distal half of the ventral side of the gonocoxites. The specimen from Alabama differs from the two specimens from Pennsylvania regarding the setosity of the basal lobe of the hypopygium. In this specimen nearly the whole surface is covered, whereas in the other specimens bristles occur at the margins only and the mediotergite is without short bristles. The small number of specimens currently known to us does not yet allow us to establish another species.	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F36227D5226EA24951FD58FCCA.taxon	distribution	Distribution. USA (Alabama, Pennsylvania).	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F36226D5226EA24BB5FB4AF84F.taxon	materials_examined	Type species: Sciara halterata Lengersdorf, 1926: 250, fig. 12. Selected literature: Tuomikoski (1960): 103 – 110; Mohrig & Menzel (1994): 167 – 210; Hippa & Vilkamaa (1991): 113 – 155; Vilkamaa (1996): 1 – 57; Menzel & Mohrig (2000): 429 – 451.	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F36226D5226EA24BB5FB4AF84F.taxon	discussion	Comments. Phytosciara s. str. is not species-rich. It is known from the Palaearctic region as well as one species from Papua New Guinea (Mohrig 1999). Dolichosciara is more species-rich and distributed in the Holarctic region, reaching the Neotropical region (Mohrig 2003) and Australasia / Oceania (Mohrig 1999; Steffan 1969; Mohrig et al. 2017). The subgenus Prosciara is species-rich in the Eastern Palaearctic, in the Oriental region, particularly in the Indomalayan subregion (Vilkamaa & Hippa 1996), and five species are known from the western Palaearctic as well (Hippa & Vilkamaa 1991; Menzel & Mohrig 2000). Mohrig et al. (2013) reported two species, Ph. multisetifera (Pettey) and Ph. ornata (Winnertz) of the subgenus Dolichosciara, and one species, Ph. prosciaroides (Tuomikoski), of the subgenus Prosciara from North America. Phytosciara greylockensis was reported by Eiseman et al. (2018) as the first North American representative of Phytosciara s. str.	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F36226D5226EA24DC3FBDFFA19.taxon	description	(Fig. 6 C – D)	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F36226D5226EA24DC3FBDFFA19.taxon	materials_examined	Further material: (not mentioned in Mohrig et al. 2013). 1 male, 5. ix. 2000, USA, Alabama, Baldwin Country, Byrnes lake, 30.79 ° N, 87.91 ° W, leg. J. W. McCreadie, pitfall trap (SDEI).	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F36226D5226EA24DC3FBDFFA19.taxon	discussion	Comments. The species is characterized by having a lightly sclerotized, broad ventral base of the hypopygium, tending to form an intergonocoxal lobe. The gonostylus is elongate and bulbous on the inner side, with two long subapical spines of different lengths, a whiplash hair somewhat longer than the spines. 1 st and 2 nd antennal flagellomeres are pale. Flagellomeres are sparsely haired and with rather long dark bristles but also densely covered with fine hyaline sensory-like hairs inserted in rather large basal pits. The tibial organ on the fore tibia is nearly comblike and not bordered. It is nearly identical to Psl. villaborensis Heller, 2012 from Spain, except that European specimens possess a more distinct intergonocoxal lobe on the ventral base of the hypopygium.	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F36226D5226EA24DC3FBDFFA19.taxon	distribution	Distribution. Holarctic. Canada (New Brunswick); USA (Alabama, New York).	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F36239D5396EA24EC6FD01F85D.taxon	description	(Fig. 10 A)	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F36239D5396EA24EC6FD01F85D.taxon	materials_examined	Material: 1 male, 11. vi. – 24. vii. 1994, Canada, Berland River at Hwy 40, 53.42 ° N, 118.20 ° W, pine forest, Malaise trap, leg. E. Fuller (PWMP).	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F36239D5396EA24EC6FD01F85D.taxon	discussion	Comments. The species is characterized by having 5 apical spines on the gonostylus, a distinct intergonocoxal lobe on the ventral base of the hypopygium and bare posterior wing veins. It is similar to Ph. quadriangulata Mohrig & Krivosheina, 1985 from the Russian Far East but differs by having five instead of four apical spines.	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F36239D5396EA24EC6FD01F85D.taxon	distribution	Distribution. Canada (Alberta). New for North America. PLATE I. Figure I. Pseudolycoriella jucunda (Johannsen). Hypopygium ventral view. Figure II. Pseudolycoriella subjucunda sp. n. Hypopygium ventral view.	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
F82E87F36239D5396EA24EC6FD01F85D.taxon	description	PLATE II. Figure III. Pseudolycoriella flavipila sp. n. Hypopygium ventral view. Figure IV. Pseudolycoriella basisetosa sp. n. Hypopygium ventral view. PLATE III. Figure V. Pseudolycoriella lobosa (Pettey). Hypopygium ventral view. Figure VI. Pseudolycoriella parilis (Pettey). Hypopygium ventral view. PLATE IV. Figure VII. Pseudolycoriella brevialata sp. n. Hypopygium ventral view. Figure VIII. Pseudolycoriella planiforceps (Steffan). Hypopygium ventral view.	en	Mohrig, Werner, Kauschke, Ellen (2019): New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part V. Genera Pseudolycoriella Menzel & Mohrig and Phytosciara Frey. Zootaxa 4543 (2): 261-283, DOI: 10.11646/zootaxa.4543.2.5
