taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
8BF8C77C59735206B8A3A8C72DCE898F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349087	https://doi.org/10.3897/phytokeys.134.37979.figure5	Figure 5. Virola allenii A leaves B fruit C partial inflorescences. Drawn by Pedro Juarez based on P. H. Allen 6727 (A), R. Aguilar 2224 (B) and R. Aguilar (C) from a photo, physical specimen not seen.	Figure 5. Virola allenii A leaves B fruit C partial inflorescences. Drawn by Pedro Juarez based on P. H. Allen 6727 (A), R. Aguilar 2224 (B) and R. Aguilar (C) from a photo, physical specimen not seen.	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
8BF8C77C59735206B8A3A8C72DCE898F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349088	https://doi.org/10.3897/phytokeys.134.37979.figure6	Figure 6. Virola allenii A treetop B branching C lower trunk and buttress D branch with leaves, showing the adaxial surface E leaf blades on abaxial surface F exudate of the trunk G leaf blade surfaces, adaxial (above) and abaxial (below) H petiole and leaf base I mature fruit, inset left and arrow showing the galls on leaves and branches, respectively J fruits K fruit close-up. Photos by Reinaldo Aguilar.	Figure 6. Virola allenii A treetop B branching C lower trunk and buttress D branch with leaves, showing the adaxial surface E leaf blades on abaxial surface F exudate of the trunk G leaf blade surfaces, adaxial (above) and abaxial (below) H petiole and leaf base I mature fruit, inset left and arrow showing the galls on leaves and branches, respectively J fruits K fruit close-up. Photos by Reinaldo Aguilar.	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
8BF8C77C59735206B8A3A8C72DCE898F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349089	https://doi.org/10.3897/phytokeys.134.37979.figure7	Figure 7. Comparisons of Virola macrocarpa with similar species in Mesoamerica A V. allenii (P. H. Allen 5763, F; inset fruits, from P. H. Allen 6727, GH) B V. amistadensis (G. McPherson 9717, MO; inset fruits, from G. McPherson 9715, MO) C V. macrocarpa (A. E. Lawrance 675, MO) D V. otobifolia (J. P. Folsom 1440, MO). Image courtesy of Field Museum (A).	Figure 7. Comparisons of Virola macrocarpa with similar species in Mesoamerica A V. allenii (P. H. Allen 5763, F; inset fruits, from P. H. Allen 6727, GH) B V. amistadensis (G. McPherson 9717, MO; inset fruits, from G. McPherson 9715, MO) C V. macrocarpa (A. E. Lawrance 675, MO) D V. otobifolia (J. P. Folsom 1440, MO). Image courtesy of Field Museum (A).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
8BF8C77C59735206B8A3A8C72DCE898F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349090	https://doi.org/10.3897/phytokeys.134.37979.figure8	Figure 8. Pattern of the lateral veins in Mesoamerican Virola A Virola allenii (K. Thomsen 1284, NO) B V. amistadensis (G. McPherson 8703, MO) C V. chrysocarpa (B. Hammel et al. 16864, MO) D V. elongata (M. Correa & R. L. Dressler 1078, MO) E V. fosteri (G. de Nevers 7226, MO) F V. guatemalensis (G. Ibarra 957, MO) G V. koschnyi (J. Miller & J. C. Sandino 1110, MO) H V. laevigata (N. Zamora & T. D. Pennington 1583, MO) I V. megacarpa (G. de Nevers 5184, MO) J V. montana (E. Lepiz & J. F. Morales 284, MO) K V. multiflora (J. Manzanares 3561, MO) L V. nobilis from Barro Colorado (R. J. Schmalzel 320, MO) M V. otobifolia (G. de Nevers et al. 7530, MO) N V. sebifera (T. B. Croat 5959, MO) O V. nobilis from Osa Peninsula (R. Aguilar 11186, MO).	Figure 8. Pattern of the lateral veins in Mesoamerican Virola A Virola allenii (K. Thomsen 1284, NO) B V. amistadensis (G. McPherson 8703, MO) C V. chrysocarpa (B. Hammel et al. 16864, MO) D V. elongata (M. Correa & R. L. Dressler 1078, MO) E V. fosteri (G. de Nevers 7226, MO) F V. guatemalensis (G. Ibarra 957, MO) G V. koschnyi (J. Miller & J. C. Sandino 1110, MO) H V. laevigata (N. Zamora & T. D. Pennington 1583, MO) I V. megacarpa (G. de Nevers 5184, MO) J V. montana (E. Lepiz & J. F. Morales 284, MO) K V. multiflora (J. Manzanares 3561, MO) L V. nobilis from Barro Colorado (R. J. Schmalzel 320, MO) M V. otobifolia (G. de Nevers et al. 7530, MO) N V. sebifera (T. B. Croat 5959, MO) O V. nobilis from Osa Peninsula (R. Aguilar 11186, MO).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
8BF8C77C59735206B8A3A8C72DCE898F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349085	https://doi.org/10.3897/phytokeys.134.37979.figure3	Figure 3. Trichomes in Mesoamerican Virola A Virola allenii (R. Aguilar 2224, MO) B V. amistadensis (G. McPherson 8703, MO) C V. chrysocarpa (O. Valverde 970, MO) D V. elongata (G. McPherson 13674, MO) E V. fosteri (G. McPherson 20913, MO) F V. guatemalensis (R. Cedillo 3349, MO) G V. koschnyi (R. E. Gereau et al. 3455, MO) H V. laevigata (R. Aguilar 2004, MO) I V. megacarpa (J. A. Duke 15261, MO) J V. montana (E. Bello 855, MO) K V. multiflora (R. Rueda et al. 8196, MO; inset, J. C. Sandino 3302, MO) L V. nobilis from Barro Colorado (R. J. Schmalzel 320, MO) M V. otobifolia (G. de Nevers & H. Herrera 7056, MO) N V. sebifera (F. Almeda et al. 5553, MO) O V. nobilis from Osa Peninsula (R. Aguilar 11186, MO).	Figure 3. Trichomes in Mesoamerican Virola A Virola allenii (R. Aguilar 2224, MO) B V. amistadensis (G. McPherson 8703, MO) C V. chrysocarpa (O. Valverde 970, MO) D V. elongata (G. McPherson 13674, MO) E V. fosteri (G. McPherson 20913, MO) F V. guatemalensis (R. Cedillo 3349, MO) G V. koschnyi (R. E. Gereau et al. 3455, MO) H V. laevigata (R. Aguilar 2004, MO) I V. megacarpa (J. A. Duke 15261, MO) J V. montana (E. Bello 855, MO) K V. multiflora (R. Rueda et al. 8196, MO; inset, J. C. Sandino 3302, MO) L V. nobilis from Barro Colorado (R. J. Schmalzel 320, MO) M V. otobifolia (G. de Nevers & H. Herrera 7056, MO) N V. sebifera (F. Almeda et al. 5553, MO) O V. nobilis from Osa Peninsula (R. Aguilar 11186, MO).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
8BF8C77C59735206B8A3A8C72DCE898F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349086	https://doi.org/10.3897/phytokeys.134.37979.figure4	Figure 4. Fruits of Mesoameican Virola, organised in groups of morphologically similar species A V. allenii (P. H. Allen 6727, GH) B V. amistadensis (G. McPherson 9715, MO) C V. otobifolia (A. Gentry & S. Mori 14199, MO) D V. macrocarpa (A. E. Lawrance 675, MO) E V. chrysocarpa (B. Hammel et al. 16864, MO) F V. koschnyi (R. Robles 1587, MO) G V. guatemalensis (S. Sinaca 1542, MO) H V. montana (J. Utley & K. Utley 5054, MO) I V. fosteri (S. Mori & J. Kallunki 4891, MO) J V. multiflora (R. Rueda et a. l 9960, MO) K V. sebifera (T. B. Croat 5959, MO) L V. elongata (J. D. Dwyer & M. Correa 8420, MO) M V. laevigata (R. Aguilar 1585, MO) N V. nobilis from Barro Colorado (T. B. Croat 8090, MO) O V. nobilis from the Osa Peninsula (G. Herrera 4222, MO) P V. megacarpa (G. de Nevers 5184, MO). Scale bars: 1 cm.	Figure 4. Fruits of Mesoameican Virola, organised in groups of morphologically similar species A V. allenii (P. H. Allen 6727, GH) B V. amistadensis (G. McPherson 9715, MO) C V. otobifolia (A. Gentry & S. Mori 14199, MO) D V. macrocarpa (A. E. Lawrance 675, MO) E V. chrysocarpa (B. Hammel et al. 16864, MO) F V. koschnyi (R. Robles 1587, MO) G V. guatemalensis (S. Sinaca 1542, MO) H V. montana (J. Utley & K. Utley 5054, MO) I V. fosteri (S. Mori & J. Kallunki 4891, MO) J V. multiflora (R. Rueda et a. l 9960, MO) K V. sebifera (T. B. Croat 5959, MO) L V. elongata (J. D. Dwyer & M. Correa 8420, MO) M V. laevigata (R. Aguilar 1585, MO) N V. nobilis from Barro Colorado (T. B. Croat 8090, MO) O V. nobilis from the Osa Peninsula (G. Herrera 4222, MO) P V. megacarpa (G. de Nevers 5184, MO). Scale bars: 1 cm.	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
8BF8C77C59735206B8A3A8C72DCE898F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349091	https://doi.org/10.3897/phytokeys.134.37979.figure9	Figure 9. Geographic distribution of Virola allenii (A), V. amistadensis (B), V. chrysocarpa (C), V. elongata (D), V. fosteri (E) and V. guatemalensis (F).	Figure 9. Geographic distribution of Virola allenii (A), V. amistadensis (B), V. chrysocarpa (C), V. elongata (D), V. fosteri (E) and V. guatemalensis (F).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
8BF8C77C59735206B8A3A8C72DCE898F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349092	https://doi.org/10.3897/phytokeys.134.37979.figure10	Figure 10. Virola calophylla A branch with leaves, showing both sides B branch with inflorescences. Virola elongata C leafy branch with inflorescences, inset fruit D inflorescences. Virola guatemalensis E, F leaf blades showing adaxial (E) and abaxial (F) surface. G young twigs and petioles. Photos by Robin Foster (A, B), from https: // plantidtools. fieldmuseum. org / en / nlp; Steven Paton (C, D), Rolando Perez (C inset), from https: // stricollections. org / portal / index. php and Angela Rojas (E-G).	Figure 10. Virola calophylla A branch with leaves, showing both sides B branch with inflorescences. Virola elongata C leafy branch with inflorescences, inset fruit D inflorescences. Virola guatemalensis E, F leaf blades showing adaxial (E) and abaxial (F) surface. G young twigs and petioles. Photos by Robin Foster (A, B), from https: // plantidtools. fieldmuseum. org / en / nlp; Steven Paton (C, D), Rolando Perez (C inset), from https: // stricollections. org / portal / index. php and Angela Rojas (E-G).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
71D7988BA64F5D009A972802DD6B6058.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349089	https://doi.org/10.3897/phytokeys.134.37979.figure7	Figure 7. Comparisons of Virola macrocarpa with similar species in Mesoamerica A V. allenii (P. H. Allen 5763, F; inset fruits, from P. H. Allen 6727, GH) B V. amistadensis (G. McPherson 9717, MO; inset fruits, from G. McPherson 9715, MO) C V. macrocarpa (A. E. Lawrance 675, MO) D V. otobifolia (J. P. Folsom 1440, MO). Image courtesy of Field Museum (A).	Figure 7. Comparisons of Virola macrocarpa with similar species in Mesoamerica A V. allenii (P. H. Allen 5763, F; inset fruits, from P. H. Allen 6727, GH) B V. amistadensis (G. McPherson 9717, MO; inset fruits, from G. McPherson 9715, MO) C V. macrocarpa (A. E. Lawrance 675, MO) D V. otobifolia (J. P. Folsom 1440, MO). Image courtesy of Field Museum (A).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
71D7988BA64F5D009A972802DD6B6058.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349093	https://doi.org/10.3897/phytokeys.134.37979.figure11	Figure 11. Virola amistadensis A leaf with staminate inflorescences B leaf C fruits D pericarp E seed. Drawn by Pedro Juarez, based on G. McPherson 9717 (A) and G. McPherson 9715 (B-E).	Figure 11. Virola amistadensis A leaf with staminate inflorescences B leaf C fruits D pericarp E seed. Drawn by Pedro Juarez, based on G. McPherson 9717 (A) and G. McPherson 9715 (B-E).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
71D7988BA64F5D009A972802DD6B6058.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349090	https://doi.org/10.3897/phytokeys.134.37979.figure8	Figure 8. Pattern of the lateral veins in Mesoamerican Virola A Virola allenii (K. Thomsen 1284, NO) B V. amistadensis (G. McPherson 8703, MO) C V. chrysocarpa (B. Hammel et al. 16864, MO) D V. elongata (M. Correa & R. L. Dressler 1078, MO) E V. fosteri (G. de Nevers 7226, MO) F V. guatemalensis (G. Ibarra 957, MO) G V. koschnyi (J. Miller & J. C. Sandino 1110, MO) H V. laevigata (N. Zamora & T. D. Pennington 1583, MO) I V. megacarpa (G. de Nevers 5184, MO) J V. montana (E. Lepiz & J. F. Morales 284, MO) K V. multiflora (J. Manzanares 3561, MO) L V. nobilis from Barro Colorado (R. J. Schmalzel 320, MO) M V. otobifolia (G. de Nevers et al. 7530, MO) N V. sebifera (T. B. Croat 5959, MO) O V. nobilis from Osa Peninsula (R. Aguilar 11186, MO).	Figure 8. Pattern of the lateral veins in Mesoamerican Virola A Virola allenii (K. Thomsen 1284, NO) B V. amistadensis (G. McPherson 8703, MO) C V. chrysocarpa (B. Hammel et al. 16864, MO) D V. elongata (M. Correa & R. L. Dressler 1078, MO) E V. fosteri (G. de Nevers 7226, MO) F V. guatemalensis (G. Ibarra 957, MO) G V. koschnyi (J. Miller & J. C. Sandino 1110, MO) H V. laevigata (N. Zamora & T. D. Pennington 1583, MO) I V. megacarpa (G. de Nevers 5184, MO) J V. montana (E. Lepiz & J. F. Morales 284, MO) K V. multiflora (J. Manzanares 3561, MO) L V. nobilis from Barro Colorado (R. J. Schmalzel 320, MO) M V. otobifolia (G. de Nevers et al. 7530, MO) N V. sebifera (T. B. Croat 5959, MO) O V. nobilis from Osa Peninsula (R. Aguilar 11186, MO).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
71D7988BA64F5D009A972802DD6B6058.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349085	https://doi.org/10.3897/phytokeys.134.37979.figure3	Figure 3. Trichomes in Mesoamerican Virola A Virola allenii (R. Aguilar 2224, MO) B V. amistadensis (G. McPherson 8703, MO) C V. chrysocarpa (O. Valverde 970, MO) D V. elongata (G. McPherson 13674, MO) E V. fosteri (G. McPherson 20913, MO) F V. guatemalensis (R. Cedillo 3349, MO) G V. koschnyi (R. E. Gereau et al. 3455, MO) H V. laevigata (R. Aguilar 2004, MO) I V. megacarpa (J. A. Duke 15261, MO) J V. montana (E. Bello 855, MO) K V. multiflora (R. Rueda et al. 8196, MO; inset, J. C. Sandino 3302, MO) L V. nobilis from Barro Colorado (R. J. Schmalzel 320, MO) M V. otobifolia (G. de Nevers & H. Herrera 7056, MO) N V. sebifera (F. Almeda et al. 5553, MO) O V. nobilis from Osa Peninsula (R. Aguilar 11186, MO).	Figure 3. Trichomes in Mesoamerican Virola A Virola allenii (R. Aguilar 2224, MO) B V. amistadensis (G. McPherson 8703, MO) C V. chrysocarpa (O. Valverde 970, MO) D V. elongata (G. McPherson 13674, MO) E V. fosteri (G. McPherson 20913, MO) F V. guatemalensis (R. Cedillo 3349, MO) G V. koschnyi (R. E. Gereau et al. 3455, MO) H V. laevigata (R. Aguilar 2004, MO) I V. megacarpa (J. A. Duke 15261, MO) J V. montana (E. Bello 855, MO) K V. multiflora (R. Rueda et al. 8196, MO; inset, J. C. Sandino 3302, MO) L V. nobilis from Barro Colorado (R. J. Schmalzel 320, MO) M V. otobifolia (G. de Nevers & H. Herrera 7056, MO) N V. sebifera (F. Almeda et al. 5553, MO) O V. nobilis from Osa Peninsula (R. Aguilar 11186, MO).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
71D7988BA64F5D009A972802DD6B6058.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349086	https://doi.org/10.3897/phytokeys.134.37979.figure4	Figure 4. Fruits of Mesoameican Virola, organised in groups of morphologically similar species A V. allenii (P. H. Allen 6727, GH) B V. amistadensis (G. McPherson 9715, MO) C V. otobifolia (A. Gentry & S. Mori 14199, MO) D V. macrocarpa (A. E. Lawrance 675, MO) E V. chrysocarpa (B. Hammel et al. 16864, MO) F V. koschnyi (R. Robles 1587, MO) G V. guatemalensis (S. Sinaca 1542, MO) H V. montana (J. Utley & K. Utley 5054, MO) I V. fosteri (S. Mori & J. Kallunki 4891, MO) J V. multiflora (R. Rueda et a. l 9960, MO) K V. sebifera (T. B. Croat 5959, MO) L V. elongata (J. D. Dwyer & M. Correa 8420, MO) M V. laevigata (R. Aguilar 1585, MO) N V. nobilis from Barro Colorado (T. B. Croat 8090, MO) O V. nobilis from the Osa Peninsula (G. Herrera 4222, MO) P V. megacarpa (G. de Nevers 5184, MO). Scale bars: 1 cm.	Figure 4. Fruits of Mesoameican Virola, organised in groups of morphologically similar species A V. allenii (P. H. Allen 6727, GH) B V. amistadensis (G. McPherson 9715, MO) C V. otobifolia (A. Gentry & S. Mori 14199, MO) D V. macrocarpa (A. E. Lawrance 675, MO) E V. chrysocarpa (B. Hammel et al. 16864, MO) F V. koschnyi (R. Robles 1587, MO) G V. guatemalensis (S. Sinaca 1542, MO) H V. montana (J. Utley & K. Utley 5054, MO) I V. fosteri (S. Mori & J. Kallunki 4891, MO) J V. multiflora (R. Rueda et a. l 9960, MO) K V. sebifera (T. B. Croat 5959, MO) L V. elongata (J. D. Dwyer & M. Correa 8420, MO) M V. laevigata (R. Aguilar 1585, MO) N V. nobilis from Barro Colorado (T. B. Croat 8090, MO) O V. nobilis from the Osa Peninsula (G. Herrera 4222, MO) P V. megacarpa (G. de Nevers 5184, MO). Scale bars: 1 cm.	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
71D7988BA64F5D009A972802DD6B6058.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349091	https://doi.org/10.3897/phytokeys.134.37979.figure9	Figure 9. Geographic distribution of Virola allenii (A), V. amistadensis (B), V. chrysocarpa (C), V. elongata (D), V. fosteri (E) and V. guatemalensis (F).	Figure 9. Geographic distribution of Virola allenii (A), V. amistadensis (B), V. chrysocarpa (C), V. elongata (D), V. fosteri (E) and V. guatemalensis (F).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
71D7988BA64F5D009A972802DD6B6058.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349092	https://doi.org/10.3897/phytokeys.134.37979.figure10	Figure 10. Virola calophylla A branch with leaves, showing both sides B branch with inflorescences. Virola elongata C leafy branch with inflorescences, inset fruit D inflorescences. Virola guatemalensis E, F leaf blades showing adaxial (E) and abaxial (F) surface. G young twigs and petioles. Photos by Robin Foster (A, B), from https: // plantidtools. fieldmuseum. org / en / nlp; Steven Paton (C, D), Rolando Perez (C inset), from https: // stricollections. org / portal / index. php and Angela Rojas (E-G).	Figure 10. Virola calophylla A branch with leaves, showing both sides B branch with inflorescences. Virola elongata C leafy branch with inflorescences, inset fruit D inflorescences. Virola guatemalensis E, F leaf blades showing adaxial (E) and abaxial (F) surface. G young twigs and petioles. Photos by Robin Foster (A, B), from https: // plantidtools. fieldmuseum. org / en / nlp; Steven Paton (C, D), Rolando Perez (C inset), from https: // stricollections. org / portal / index. php and Angela Rojas (E-G).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
71D7988BA64F5D009A972802DD6B6058.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349088	https://doi.org/10.3897/phytokeys.134.37979.figure6	Figure 6. Virola allenii A treetop B branching C lower trunk and buttress D branch with leaves, showing the adaxial surface E leaf blades on abaxial surface F exudate of the trunk G leaf blade surfaces, adaxial (above) and abaxial (below) H petiole and leaf base I mature fruit, inset left and arrow showing the galls on leaves and branches, respectively J fruits K fruit close-up. Photos by Reinaldo Aguilar.	Figure 6. Virola allenii A treetop B branching C lower trunk and buttress D branch with leaves, showing the adaxial surface E leaf blades on abaxial surface F exudate of the trunk G leaf blade surfaces, adaxial (above) and abaxial (below) H petiole and leaf base I mature fruit, inset left and arrow showing the galls on leaves and branches, respectively J fruits K fruit close-up. Photos by Reinaldo Aguilar.	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
71D7988BA64F5D009A972802DD6B6058.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349105	https://doi.org/10.3897/phytokeys.134.37979.figure23	Figure 23. Virola otobifolia A branch with leaf blades showing both surfaces and inflorescences B leaf blade, venation and base on abaxial surface C infructescence D fruits E detail of fruit, showing an aril of an immature fruit. Virola surinamensis. F Branch with inflorescences G, H leaf blade on adaxial (G) and abaxial surface (H). I Infructescence J detail of fruits, including a longitudinal section of an immature fruit. Photos by Rolando Perez (A, B from https: // stricollections. org / portal / index. php); Jerry Harrison (C), Alwyn H. Gentry (D, E from http: // www. tropicos. org) and John P. Janovec (F-J from http: // atrium. andesamazon. org /).	Figure 23. Virola otobifolia A branch with leaf blades showing both surfaces and inflorescences B leaf blade, venation and base on abaxial surface C infructescence D fruits E detail of fruit, showing an aril of an immature fruit. Virola surinamensis. F Branch with inflorescences G, H leaf blade on adaxial (G) and abaxial surface (H). I Infructescence J detail of fruits, including a longitudinal section of an immature fruit. Photos by Rolando Perez (A, B from https: // stricollections. org / portal / index. php); Jerry Harrison (C), Alwyn H. Gentry (D, E from http: // www. tropicos. org) and John P. Janovec (F-J from http: // atrium. andesamazon. org /).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
A0D0CB899ED955D1ABE63768352D7FDA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349094	https://doi.org/10.3897/phytokeys.134.37979.figure12	Figure 12. Virola chrysocarpa A branch with leaves and fruits B base of young leaf C fruit D partial staminate inflorescences E staminate flower. Drawn by Pedro Juarez, based on R. Aguilar 11190 (A-C) and R. Aguilar 11569 (D, E).	Figure 12. Virola chrysocarpa A branch with leaves and fruits B base of young leaf C fruit D partial staminate inflorescences E staminate flower. Drawn by Pedro Juarez, based on R. Aguilar 11190 (A-C) and R. Aguilar 11569 (D, E).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
A0D0CB899ED955D1ABE63768352D7FDA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349095	https://doi.org/10.3897/phytokeys.134.37979.figure13	Figure 13. Vegetative characteristics of Virola chrysocarpa A tree showing deciduous nature of the species B trunk C buttress D branch showing new leaves E leaf blades on abaxial surface F leaf blades on abaxial surface, showing margin detail and venation G leaf apex H new branch and leaf base. Photos by Reinaldo Aguilar.	Figure 13. Vegetative characteristics of Virola chrysocarpa A tree showing deciduous nature of the species B trunk C buttress D branch showing new leaves E leaf blades on abaxial surface F leaf blades on abaxial surface, showing margin detail and venation G leaf apex H new branch and leaf base. Photos by Reinaldo Aguilar.	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
A0D0CB899ED955D1ABE63768352D7FDA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349096	https://doi.org/10.3897/phytokeys.134.37979.figure14	Figure 14. Fertile characteristics of Virola chrysocarpa A branch with staminate inflorescences B staminate inflorescences C staminate flower buds and open flower D Infructescence E open fruit F seeds. Photos by Reinaldo Aguilar.	Figure 14. Fertile characteristics of Virola chrysocarpa A branch with staminate inflorescences B staminate inflorescences C staminate flower buds and open flower D Infructescence E open fruit F seeds. Photos by Reinaldo Aguilar.	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
A0D0CB899ED955D1ABE63768352D7FDA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349085	https://doi.org/10.3897/phytokeys.134.37979.figure3	Figure 3. Trichomes in Mesoamerican Virola A Virola allenii (R. Aguilar 2224, MO) B V. amistadensis (G. McPherson 8703, MO) C V. chrysocarpa (O. Valverde 970, MO) D V. elongata (G. McPherson 13674, MO) E V. fosteri (G. McPherson 20913, MO) F V. guatemalensis (R. Cedillo 3349, MO) G V. koschnyi (R. E. Gereau et al. 3455, MO) H V. laevigata (R. Aguilar 2004, MO) I V. megacarpa (J. A. Duke 15261, MO) J V. montana (E. Bello 855, MO) K V. multiflora (R. Rueda et al. 8196, MO; inset, J. C. Sandino 3302, MO) L V. nobilis from Barro Colorado (R. J. Schmalzel 320, MO) M V. otobifolia (G. de Nevers & H. Herrera 7056, MO) N V. sebifera (F. Almeda et al. 5553, MO) O V. nobilis from Osa Peninsula (R. Aguilar 11186, MO).	Figure 3. Trichomes in Mesoamerican Virola A Virola allenii (R. Aguilar 2224, MO) B V. amistadensis (G. McPherson 8703, MO) C V. chrysocarpa (O. Valverde 970, MO) D V. elongata (G. McPherson 13674, MO) E V. fosteri (G. McPherson 20913, MO) F V. guatemalensis (R. Cedillo 3349, MO) G V. koschnyi (R. E. Gereau et al. 3455, MO) H V. laevigata (R. Aguilar 2004, MO) I V. megacarpa (J. A. Duke 15261, MO) J V. montana (E. Bello 855, MO) K V. multiflora (R. Rueda et al. 8196, MO; inset, J. C. Sandino 3302, MO) L V. nobilis from Barro Colorado (R. J. Schmalzel 320, MO) M V. otobifolia (G. de Nevers & H. Herrera 7056, MO) N V. sebifera (F. Almeda et al. 5553, MO) O V. nobilis from Osa Peninsula (R. Aguilar 11186, MO).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
A0D0CB899ED955D1ABE63768352D7FDA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349090	https://doi.org/10.3897/phytokeys.134.37979.figure8	Figure 8. Pattern of the lateral veins in Mesoamerican Virola A Virola allenii (K. Thomsen 1284, NO) B V. amistadensis (G. McPherson 8703, MO) C V. chrysocarpa (B. Hammel et al. 16864, MO) D V. elongata (M. Correa & R. L. Dressler 1078, MO) E V. fosteri (G. de Nevers 7226, MO) F V. guatemalensis (G. Ibarra 957, MO) G V. koschnyi (J. Miller & J. C. Sandino 1110, MO) H V. laevigata (N. Zamora & T. D. Pennington 1583, MO) I V. megacarpa (G. de Nevers 5184, MO) J V. montana (E. Lepiz & J. F. Morales 284, MO) K V. multiflora (J. Manzanares 3561, MO) L V. nobilis from Barro Colorado (R. J. Schmalzel 320, MO) M V. otobifolia (G. de Nevers et al. 7530, MO) N V. sebifera (T. B. Croat 5959, MO) O V. nobilis from Osa Peninsula (R. Aguilar 11186, MO).	Figure 8. Pattern of the lateral veins in Mesoamerican Virola A Virola allenii (K. Thomsen 1284, NO) B V. amistadensis (G. McPherson 8703, MO) C V. chrysocarpa (B. Hammel et al. 16864, MO) D V. elongata (M. Correa & R. L. Dressler 1078, MO) E V. fosteri (G. de Nevers 7226, MO) F V. guatemalensis (G. Ibarra 957, MO) G V. koschnyi (J. Miller & J. C. Sandino 1110, MO) H V. laevigata (N. Zamora & T. D. Pennington 1583, MO) I V. megacarpa (G. de Nevers 5184, MO) J V. montana (E. Lepiz & J. F. Morales 284, MO) K V. multiflora (J. Manzanares 3561, MO) L V. nobilis from Barro Colorado (R. J. Schmalzel 320, MO) M V. otobifolia (G. de Nevers et al. 7530, MO) N V. sebifera (T. B. Croat 5959, MO) O V. nobilis from Osa Peninsula (R. Aguilar 11186, MO).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
A0D0CB899ED955D1ABE63768352D7FDA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349091	https://doi.org/10.3897/phytokeys.134.37979.figure9	Figure 9. Geographic distribution of Virola allenii (A), V. amistadensis (B), V. chrysocarpa (C), V. elongata (D), V. fosteri (E) and V. guatemalensis (F).	Figure 9. Geographic distribution of Virola allenii (A), V. amistadensis (B), V. chrysocarpa (C), V. elongata (D), V. fosteri (E) and V. guatemalensis (F).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
A0D0CB899ED955D1ABE63768352D7FDA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349086	https://doi.org/10.3897/phytokeys.134.37979.figure4	Figure 4. Fruits of Mesoameican Virola, organised in groups of morphologically similar species A V. allenii (P. H. Allen 6727, GH) B V. amistadensis (G. McPherson 9715, MO) C V. otobifolia (A. Gentry & S. Mori 14199, MO) D V. macrocarpa (A. E. Lawrance 675, MO) E V. chrysocarpa (B. Hammel et al. 16864, MO) F V. koschnyi (R. Robles 1587, MO) G V. guatemalensis (S. Sinaca 1542, MO) H V. montana (J. Utley & K. Utley 5054, MO) I V. fosteri (S. Mori & J. Kallunki 4891, MO) J V. multiflora (R. Rueda et a. l 9960, MO) K V. sebifera (T. B. Croat 5959, MO) L V. elongata (J. D. Dwyer & M. Correa 8420, MO) M V. laevigata (R. Aguilar 1585, MO) N V. nobilis from Barro Colorado (T. B. Croat 8090, MO) O V. nobilis from the Osa Peninsula (G. Herrera 4222, MO) P V. megacarpa (G. de Nevers 5184, MO). Scale bars: 1 cm.	Figure 4. Fruits of Mesoameican Virola, organised in groups of morphologically similar species A V. allenii (P. H. Allen 6727, GH) B V. amistadensis (G. McPherson 9715, MO) C V. otobifolia (A. Gentry & S. Mori 14199, MO) D V. macrocarpa (A. E. Lawrance 675, MO) E V. chrysocarpa (B. Hammel et al. 16864, MO) F V. koschnyi (R. Robles 1587, MO) G V. guatemalensis (S. Sinaca 1542, MO) H V. montana (J. Utley & K. Utley 5054, MO) I V. fosteri (S. Mori & J. Kallunki 4891, MO) J V. multiflora (R. Rueda et a. l 9960, MO) K V. sebifera (T. B. Croat 5959, MO) L V. elongata (J. D. Dwyer & M. Correa 8420, MO) M V. laevigata (R. Aguilar 1585, MO) N V. nobilis from Barro Colorado (T. B. Croat 8090, MO) O V. nobilis from the Osa Peninsula (G. Herrera 4222, MO) P V. megacarpa (G. de Nevers 5184, MO). Scale bars: 1 cm.	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
A0D0CB899ED955D1ABE63768352D7FDA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349099	https://doi.org/10.3897/phytokeys.134.37979.figure17	Figure 17. Virola koschnyi A lower trunk and buttresses B trunk C branching D leaf blades on abaxial surface E leaf blades on adaxial surface F twig, petiole and leaf base G leaf venation on adaxial surface (left) and abaxial surface (right) H branch with fruits, showing the interior part of the fruit I seed J mature fruit. All photos by Reinaldo Aguilar from https: // sura. ots. ac. cr / florula 4 / index. php, except G by D. Santamaria-Aguilar.	Figure 17. Virola koschnyi A lower trunk and buttresses B trunk C branching D leaf blades on abaxial surface E leaf blades on adaxial surface F twig, petiole and leaf base G leaf venation on adaxial surface (left) and abaxial surface (right) H branch with fruits, showing the interior part of the fruit I seed J mature fruit. All photos by Reinaldo Aguilar from https: // sura. ots. ac. cr / florula 4 / index. php, except G by D. Santamaria-Aguilar.	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
051616A20EFF5C8280C7375A645056AA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349092	https://doi.org/10.3897/phytokeys.134.37979.figure10	Figure 10. Virola calophylla A branch with leaves, showing both sides B branch with inflorescences. Virola elongata C leafy branch with inflorescences, inset fruit D inflorescences. Virola guatemalensis E, F leaf blades showing adaxial (E) and abaxial (F) surface. G young twigs and petioles. Photos by Robin Foster (A, B), from https: // plantidtools. fieldmuseum. org / en / nlp; Steven Paton (C, D), Rolando Perez (C inset), from https: // stricollections. org / portal / index. php and Angela Rojas (E-G).	Figure 10. Virola calophylla A branch with leaves, showing both sides B branch with inflorescences. Virola elongata C leafy branch with inflorescences, inset fruit D inflorescences. Virola guatemalensis E, F leaf blades showing adaxial (E) and abaxial (F) surface. G young twigs and petioles. Photos by Robin Foster (A, B), from https: // plantidtools. fieldmuseum. org / en / nlp; Steven Paton (C, D), Rolando Perez (C inset), from https: // stricollections. org / portal / index. php and Angela Rojas (E-G).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
051616A20EFF5C8280C7375A645056AA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349085	https://doi.org/10.3897/phytokeys.134.37979.figure3	Figure 3. Trichomes in Mesoamerican Virola A Virola allenii (R. Aguilar 2224, MO) B V. amistadensis (G. McPherson 8703, MO) C V. chrysocarpa (O. Valverde 970, MO) D V. elongata (G. McPherson 13674, MO) E V. fosteri (G. McPherson 20913, MO) F V. guatemalensis (R. Cedillo 3349, MO) G V. koschnyi (R. E. Gereau et al. 3455, MO) H V. laevigata (R. Aguilar 2004, MO) I V. megacarpa (J. A. Duke 15261, MO) J V. montana (E. Bello 855, MO) K V. multiflora (R. Rueda et al. 8196, MO; inset, J. C. Sandino 3302, MO) L V. nobilis from Barro Colorado (R. J. Schmalzel 320, MO) M V. otobifolia (G. de Nevers & H. Herrera 7056, MO) N V. sebifera (F. Almeda et al. 5553, MO) O V. nobilis from Osa Peninsula (R. Aguilar 11186, MO).	Figure 3. Trichomes in Mesoamerican Virola A Virola allenii (R. Aguilar 2224, MO) B V. amistadensis (G. McPherson 8703, MO) C V. chrysocarpa (O. Valverde 970, MO) D V. elongata (G. McPherson 13674, MO) E V. fosteri (G. McPherson 20913, MO) F V. guatemalensis (R. Cedillo 3349, MO) G V. koschnyi (R. E. Gereau et al. 3455, MO) H V. laevigata (R. Aguilar 2004, MO) I V. megacarpa (J. A. Duke 15261, MO) J V. montana (E. Bello 855, MO) K V. multiflora (R. Rueda et al. 8196, MO; inset, J. C. Sandino 3302, MO) L V. nobilis from Barro Colorado (R. J. Schmalzel 320, MO) M V. otobifolia (G. de Nevers & H. Herrera 7056, MO) N V. sebifera (F. Almeda et al. 5553, MO) O V. nobilis from Osa Peninsula (R. Aguilar 11186, MO).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
051616A20EFF5C8280C7375A645056AA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349090	https://doi.org/10.3897/phytokeys.134.37979.figure8	Figure 8. Pattern of the lateral veins in Mesoamerican Virola A Virola allenii (K. Thomsen 1284, NO) B V. amistadensis (G. McPherson 8703, MO) C V. chrysocarpa (B. Hammel et al. 16864, MO) D V. elongata (M. Correa & R. L. Dressler 1078, MO) E V. fosteri (G. de Nevers 7226, MO) F V. guatemalensis (G. Ibarra 957, MO) G V. koschnyi (J. Miller & J. C. Sandino 1110, MO) H V. laevigata (N. Zamora & T. D. Pennington 1583, MO) I V. megacarpa (G. de Nevers 5184, MO) J V. montana (E. Lepiz & J. F. Morales 284, MO) K V. multiflora (J. Manzanares 3561, MO) L V. nobilis from Barro Colorado (R. J. Schmalzel 320, MO) M V. otobifolia (G. de Nevers et al. 7530, MO) N V. sebifera (T. B. Croat 5959, MO) O V. nobilis from Osa Peninsula (R. Aguilar 11186, MO).	Figure 8. Pattern of the lateral veins in Mesoamerican Virola A Virola allenii (K. Thomsen 1284, NO) B V. amistadensis (G. McPherson 8703, MO) C V. chrysocarpa (B. Hammel et al. 16864, MO) D V. elongata (M. Correa & R. L. Dressler 1078, MO) E V. fosteri (G. de Nevers 7226, MO) F V. guatemalensis (G. Ibarra 957, MO) G V. koschnyi (J. Miller & J. C. Sandino 1110, MO) H V. laevigata (N. Zamora & T. D. Pennington 1583, MO) I V. megacarpa (G. de Nevers 5184, MO) J V. montana (E. Lepiz & J. F. Morales 284, MO) K V. multiflora (J. Manzanares 3561, MO) L V. nobilis from Barro Colorado (R. J. Schmalzel 320, MO) M V. otobifolia (G. de Nevers et al. 7530, MO) N V. sebifera (T. B. Croat 5959, MO) O V. nobilis from Osa Peninsula (R. Aguilar 11186, MO).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
051616A20EFF5C8280C7375A645056AA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349086	https://doi.org/10.3897/phytokeys.134.37979.figure4	Figure 4. Fruits of Mesoameican Virola, organised in groups of morphologically similar species A V. allenii (P. H. Allen 6727, GH) B V. amistadensis (G. McPherson 9715, MO) C V. otobifolia (A. Gentry & S. Mori 14199, MO) D V. macrocarpa (A. E. Lawrance 675, MO) E V. chrysocarpa (B. Hammel et al. 16864, MO) F V. koschnyi (R. Robles 1587, MO) G V. guatemalensis (S. Sinaca 1542, MO) H V. montana (J. Utley & K. Utley 5054, MO) I V. fosteri (S. Mori & J. Kallunki 4891, MO) J V. multiflora (R. Rueda et a. l 9960, MO) K V. sebifera (T. B. Croat 5959, MO) L V. elongata (J. D. Dwyer & M. Correa 8420, MO) M V. laevigata (R. Aguilar 1585, MO) N V. nobilis from Barro Colorado (T. B. Croat 8090, MO) O V. nobilis from the Osa Peninsula (G. Herrera 4222, MO) P V. megacarpa (G. de Nevers 5184, MO). Scale bars: 1 cm.	Figure 4. Fruits of Mesoameican Virola, organised in groups of morphologically similar species A V. allenii (P. H. Allen 6727, GH) B V. amistadensis (G. McPherson 9715, MO) C V. otobifolia (A. Gentry & S. Mori 14199, MO) D V. macrocarpa (A. E. Lawrance 675, MO) E V. chrysocarpa (B. Hammel et al. 16864, MO) F V. koschnyi (R. Robles 1587, MO) G V. guatemalensis (S. Sinaca 1542, MO) H V. montana (J. Utley & K. Utley 5054, MO) I V. fosteri (S. Mori & J. Kallunki 4891, MO) J V. multiflora (R. Rueda et a. l 9960, MO) K V. sebifera (T. B. Croat 5959, MO) L V. elongata (J. D. Dwyer & M. Correa 8420, MO) M V. laevigata (R. Aguilar 1585, MO) N V. nobilis from Barro Colorado (T. B. Croat 8090, MO) O V. nobilis from the Osa Peninsula (G. Herrera 4222, MO) P V. megacarpa (G. de Nevers 5184, MO). Scale bars: 1 cm.	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
051616A20EFF5C8280C7375A645056AA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349091	https://doi.org/10.3897/phytokeys.134.37979.figure9	Figure 9. Geographic distribution of Virola allenii (A), V. amistadensis (B), V. chrysocarpa (C), V. elongata (D), V. fosteri (E) and V. guatemalensis (F).	Figure 9. Geographic distribution of Virola allenii (A), V. amistadensis (B), V. chrysocarpa (C), V. elongata (D), V. fosteri (E) and V. guatemalensis (F).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
5D1C8010D8B4501F81CAC3DE1AF0229A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349097	https://doi.org/10.3897/phytokeys.134.37979.figure15	Figure 15. Virola fosteri A branch with leaves and inflorescences B partial inflorescences C fruit D seed with aril E seed. Drawn by Pedro Juarez, based on C. Galdames et al. 6422 (A-E).	Figure 15. Virola fosteri A branch with leaves and inflorescences B partial inflorescences C fruit D seed with aril E seed. Drawn by Pedro Juarez, based on C. Galdames et al. 6422 (A-E).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
5D1C8010D8B4501F81CAC3DE1AF0229A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349098	https://doi.org/10.3897/phytokeys.134.37979.figure16	Figure 16. Virola fosteri A lower trunk and buttresses B branch with staminate inflorescences C leaf blades showing adaxial (left) and abaxial (right) surface; also demonstrating the revolute base D branch with staminate inflorescences and leaf blades on abaxial surface E close up of staminate inflorescences F fruit G aril covering the seed H seed Photos by Rolando Perez (A), Carmen Galdames (B, D, E), Steven Paton (C, F, G, H); all photos from https: // stricollections. org / portal / index. php.	Figure 16. Virola fosteri A lower trunk and buttresses B branch with staminate inflorescences C leaf blades showing adaxial (left) and abaxial (right) surface; also demonstrating the revolute base D branch with staminate inflorescences and leaf blades on abaxial surface E close up of staminate inflorescences F fruit G aril covering the seed H seed Photos by Rolando Perez (A), Carmen Galdames (B, D, E), Steven Paton (C, F, G, H); all photos from https: // stricollections. org / portal / index. php.	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
5D1C8010D8B4501F81CAC3DE1AF0229A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349086	https://doi.org/10.3897/phytokeys.134.37979.figure4	Figure 4. Fruits of Mesoameican Virola, organised in groups of morphologically similar species A V. allenii (P. H. Allen 6727, GH) B V. amistadensis (G. McPherson 9715, MO) C V. otobifolia (A. Gentry & S. Mori 14199, MO) D V. macrocarpa (A. E. Lawrance 675, MO) E V. chrysocarpa (B. Hammel et al. 16864, MO) F V. koschnyi (R. Robles 1587, MO) G V. guatemalensis (S. Sinaca 1542, MO) H V. montana (J. Utley & K. Utley 5054, MO) I V. fosteri (S. Mori & J. Kallunki 4891, MO) J V. multiflora (R. Rueda et a. l 9960, MO) K V. sebifera (T. B. Croat 5959, MO) L V. elongata (J. D. Dwyer & M. Correa 8420, MO) M V. laevigata (R. Aguilar 1585, MO) N V. nobilis from Barro Colorado (T. B. Croat 8090, MO) O V. nobilis from the Osa Peninsula (G. Herrera 4222, MO) P V. megacarpa (G. de Nevers 5184, MO). Scale bars: 1 cm.	Figure 4. Fruits of Mesoameican Virola, organised in groups of morphologically similar species A V. allenii (P. H. Allen 6727, GH) B V. amistadensis (G. McPherson 9715, MO) C V. otobifolia (A. Gentry & S. Mori 14199, MO) D V. macrocarpa (A. E. Lawrance 675, MO) E V. chrysocarpa (B. Hammel et al. 16864, MO) F V. koschnyi (R. Robles 1587, MO) G V. guatemalensis (S. Sinaca 1542, MO) H V. montana (J. Utley & K. Utley 5054, MO) I V. fosteri (S. Mori & J. Kallunki 4891, MO) J V. multiflora (R. Rueda et a. l 9960, MO) K V. sebifera (T. B. Croat 5959, MO) L V. elongata (J. D. Dwyer & M. Correa 8420, MO) M V. laevigata (R. Aguilar 1585, MO) N V. nobilis from Barro Colorado (T. B. Croat 8090, MO) O V. nobilis from the Osa Peninsula (G. Herrera 4222, MO) P V. megacarpa (G. de Nevers 5184, MO). Scale bars: 1 cm.	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
5D1C8010D8B4501F81CAC3DE1AF0229A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349085	https://doi.org/10.3897/phytokeys.134.37979.figure3	Figure 3. Trichomes in Mesoamerican Virola A Virola allenii (R. Aguilar 2224, MO) B V. amistadensis (G. McPherson 8703, MO) C V. chrysocarpa (O. Valverde 970, MO) D V. elongata (G. McPherson 13674, MO) E V. fosteri (G. McPherson 20913, MO) F V. guatemalensis (R. Cedillo 3349, MO) G V. koschnyi (R. E. Gereau et al. 3455, MO) H V. laevigata (R. Aguilar 2004, MO) I V. megacarpa (J. A. Duke 15261, MO) J V. montana (E. Bello 855, MO) K V. multiflora (R. Rueda et al. 8196, MO; inset, J. C. Sandino 3302, MO) L V. nobilis from Barro Colorado (R. J. Schmalzel 320, MO) M V. otobifolia (G. de Nevers & H. Herrera 7056, MO) N V. sebifera (F. Almeda et al. 5553, MO) O V. nobilis from Osa Peninsula (R. Aguilar 11186, MO).	Figure 3. Trichomes in Mesoamerican Virola A Virola allenii (R. Aguilar 2224, MO) B V. amistadensis (G. McPherson 8703, MO) C V. chrysocarpa (O. Valverde 970, MO) D V. elongata (G. McPherson 13674, MO) E V. fosteri (G. McPherson 20913, MO) F V. guatemalensis (R. Cedillo 3349, MO) G V. koschnyi (R. E. Gereau et al. 3455, MO) H V. laevigata (R. Aguilar 2004, MO) I V. megacarpa (J. A. Duke 15261, MO) J V. montana (E. Bello 855, MO) K V. multiflora (R. Rueda et al. 8196, MO; inset, J. C. Sandino 3302, MO) L V. nobilis from Barro Colorado (R. J. Schmalzel 320, MO) M V. otobifolia (G. de Nevers & H. Herrera 7056, MO) N V. sebifera (F. Almeda et al. 5553, MO) O V. nobilis from Osa Peninsula (R. Aguilar 11186, MO).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
5D1C8010D8B4501F81CAC3DE1AF0229A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349091	https://doi.org/10.3897/phytokeys.134.37979.figure9	Figure 9. Geographic distribution of Virola allenii (A), V. amistadensis (B), V. chrysocarpa (C), V. elongata (D), V. fosteri (E) and V. guatemalensis (F).	Figure 9. Geographic distribution of Virola allenii (A), V. amistadensis (B), V. chrysocarpa (C), V. elongata (D), V. fosteri (E) and V. guatemalensis (F).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
55D4AA719DD15A3EBD311FDFAEFE9F58.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349092	https://doi.org/10.3897/phytokeys.134.37979.figure10	Figure 10. Virola calophylla A branch with leaves, showing both sides B branch with inflorescences. Virola elongata C leafy branch with inflorescences, inset fruit D inflorescences. Virola guatemalensis E, F leaf blades showing adaxial (E) and abaxial (F) surface. G young twigs and petioles. Photos by Robin Foster (A, B), from https: // plantidtools. fieldmuseum. org / en / nlp; Steven Paton (C, D), Rolando Perez (C inset), from https: // stricollections. org / portal / index. php and Angela Rojas (E-G).	Figure 10. Virola calophylla A branch with leaves, showing both sides B branch with inflorescences. Virola elongata C leafy branch with inflorescences, inset fruit D inflorescences. Virola guatemalensis E, F leaf blades showing adaxial (E) and abaxial (F) surface. G young twigs and petioles. Photos by Robin Foster (A, B), from https: // plantidtools. fieldmuseum. org / en / nlp; Steven Paton (C, D), Rolando Perez (C inset), from https: // stricollections. org / portal / index. php and Angela Rojas (E-G).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
55D4AA719DD15A3EBD311FDFAEFE9F58.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349085	https://doi.org/10.3897/phytokeys.134.37979.figure3	Figure 3. Trichomes in Mesoamerican Virola A Virola allenii (R. Aguilar 2224, MO) B V. amistadensis (G. McPherson 8703, MO) C V. chrysocarpa (O. Valverde 970, MO) D V. elongata (G. McPherson 13674, MO) E V. fosteri (G. McPherson 20913, MO) F V. guatemalensis (R. Cedillo 3349, MO) G V. koschnyi (R. E. Gereau et al. 3455, MO) H V. laevigata (R. Aguilar 2004, MO) I V. megacarpa (J. A. Duke 15261, MO) J V. montana (E. Bello 855, MO) K V. multiflora (R. Rueda et al. 8196, MO; inset, J. C. Sandino 3302, MO) L V. nobilis from Barro Colorado (R. J. Schmalzel 320, MO) M V. otobifolia (G. de Nevers & H. Herrera 7056, MO) N V. sebifera (F. Almeda et al. 5553, MO) O V. nobilis from Osa Peninsula (R. Aguilar 11186, MO).	Figure 3. Trichomes in Mesoamerican Virola A Virola allenii (R. Aguilar 2224, MO) B V. amistadensis (G. McPherson 8703, MO) C V. chrysocarpa (O. Valverde 970, MO) D V. elongata (G. McPherson 13674, MO) E V. fosteri (G. McPherson 20913, MO) F V. guatemalensis (R. Cedillo 3349, MO) G V. koschnyi (R. E. Gereau et al. 3455, MO) H V. laevigata (R. Aguilar 2004, MO) I V. megacarpa (J. A. Duke 15261, MO) J V. montana (E. Bello 855, MO) K V. multiflora (R. Rueda et al. 8196, MO; inset, J. C. Sandino 3302, MO) L V. nobilis from Barro Colorado (R. J. Schmalzel 320, MO) M V. otobifolia (G. de Nevers & H. Herrera 7056, MO) N V. sebifera (F. Almeda et al. 5553, MO) O V. nobilis from Osa Peninsula (R. Aguilar 11186, MO).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
55D4AA719DD15A3EBD311FDFAEFE9F58.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349090	https://doi.org/10.3897/phytokeys.134.37979.figure8	Figure 8. Pattern of the lateral veins in Mesoamerican Virola A Virola allenii (K. Thomsen 1284, NO) B V. amistadensis (G. McPherson 8703, MO) C V. chrysocarpa (B. Hammel et al. 16864, MO) D V. elongata (M. Correa & R. L. Dressler 1078, MO) E V. fosteri (G. de Nevers 7226, MO) F V. guatemalensis (G. Ibarra 957, MO) G V. koschnyi (J. Miller & J. C. Sandino 1110, MO) H V. laevigata (N. Zamora & T. D. Pennington 1583, MO) I V. megacarpa (G. de Nevers 5184, MO) J V. montana (E. Lepiz & J. F. Morales 284, MO) K V. multiflora (J. Manzanares 3561, MO) L V. nobilis from Barro Colorado (R. J. Schmalzel 320, MO) M V. otobifolia (G. de Nevers et al. 7530, MO) N V. sebifera (T. B. Croat 5959, MO) O V. nobilis from Osa Peninsula (R. Aguilar 11186, MO).	Figure 8. Pattern of the lateral veins in Mesoamerican Virola A Virola allenii (K. Thomsen 1284, NO) B V. amistadensis (G. McPherson 8703, MO) C V. chrysocarpa (B. Hammel et al. 16864, MO) D V. elongata (M. Correa & R. L. Dressler 1078, MO) E V. fosteri (G. de Nevers 7226, MO) F V. guatemalensis (G. Ibarra 957, MO) G V. koschnyi (J. Miller & J. C. Sandino 1110, MO) H V. laevigata (N. Zamora & T. D. Pennington 1583, MO) I V. megacarpa (G. de Nevers 5184, MO) J V. montana (E. Lepiz & J. F. Morales 284, MO) K V. multiflora (J. Manzanares 3561, MO) L V. nobilis from Barro Colorado (R. J. Schmalzel 320, MO) M V. otobifolia (G. de Nevers et al. 7530, MO) N V. sebifera (T. B. Croat 5959, MO) O V. nobilis from Osa Peninsula (R. Aguilar 11186, MO).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
55D4AA719DD15A3EBD311FDFAEFE9F58.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349086	https://doi.org/10.3897/phytokeys.134.37979.figure4	Figure 4. Fruits of Mesoameican Virola, organised in groups of morphologically similar species A V. allenii (P. H. Allen 6727, GH) B V. amistadensis (G. McPherson 9715, MO) C V. otobifolia (A. Gentry & S. Mori 14199, MO) D V. macrocarpa (A. E. Lawrance 675, MO) E V. chrysocarpa (B. Hammel et al. 16864, MO) F V. koschnyi (R. Robles 1587, MO) G V. guatemalensis (S. Sinaca 1542, MO) H V. montana (J. Utley & K. Utley 5054, MO) I V. fosteri (S. Mori & J. Kallunki 4891, MO) J V. multiflora (R. Rueda et a. l 9960, MO) K V. sebifera (T. B. Croat 5959, MO) L V. elongata (J. D. Dwyer & M. Correa 8420, MO) M V. laevigata (R. Aguilar 1585, MO) N V. nobilis from Barro Colorado (T. B. Croat 8090, MO) O V. nobilis from the Osa Peninsula (G. Herrera 4222, MO) P V. megacarpa (G. de Nevers 5184, MO). Scale bars: 1 cm.	Figure 4. Fruits of Mesoameican Virola, organised in groups of morphologically similar species A V. allenii (P. H. Allen 6727, GH) B V. amistadensis (G. McPherson 9715, MO) C V. otobifolia (A. Gentry & S. Mori 14199, MO) D V. macrocarpa (A. E. Lawrance 675, MO) E V. chrysocarpa (B. Hammel et al. 16864, MO) F V. koschnyi (R. Robles 1587, MO) G V. guatemalensis (S. Sinaca 1542, MO) H V. montana (J. Utley & K. Utley 5054, MO) I V. fosteri (S. Mori & J. Kallunki 4891, MO) J V. multiflora (R. Rueda et a. l 9960, MO) K V. sebifera (T. B. Croat 5959, MO) L V. elongata (J. D. Dwyer & M. Correa 8420, MO) M V. laevigata (R. Aguilar 1585, MO) N V. nobilis from Barro Colorado (T. B. Croat 8090, MO) O V. nobilis from the Osa Peninsula (G. Herrera 4222, MO) P V. megacarpa (G. de Nevers 5184, MO). Scale bars: 1 cm.	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
55D4AA719DD15A3EBD311FDFAEFE9F58.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349091	https://doi.org/10.3897/phytokeys.134.37979.figure9	Figure 9. Geographic distribution of Virola allenii (A), V. amistadensis (B), V. chrysocarpa (C), V. elongata (D), V. fosteri (E) and V. guatemalensis (F).	Figure 9. Geographic distribution of Virola allenii (A), V. amistadensis (B), V. chrysocarpa (C), V. elongata (D), V. fosteri (E) and V. guatemalensis (F).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
A5F614C7BCBE54FE9642232C771993C1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349099	https://doi.org/10.3897/phytokeys.134.37979.figure17	Figure 17. Virola koschnyi A lower trunk and buttresses B trunk C branching D leaf blades on abaxial surface E leaf blades on adaxial surface F twig, petiole and leaf base G leaf venation on adaxial surface (left) and abaxial surface (right) H branch with fruits, showing the interior part of the fruit I seed J mature fruit. All photos by Reinaldo Aguilar from https: // sura. ots. ac. cr / florula 4 / index. php, except G by D. Santamaria-Aguilar.	Figure 17. Virola koschnyi A lower trunk and buttresses B trunk C branching D leaf blades on abaxial surface E leaf blades on adaxial surface F twig, petiole and leaf base G leaf venation on adaxial surface (left) and abaxial surface (right) H branch with fruits, showing the interior part of the fruit I seed J mature fruit. All photos by Reinaldo Aguilar from https: // sura. ots. ac. cr / florula 4 / index. php, except G by D. Santamaria-Aguilar.	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
A5F614C7BCBE54FE9642232C771993C1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349085	https://doi.org/10.3897/phytokeys.134.37979.figure3	Figure 3. Trichomes in Mesoamerican Virola A Virola allenii (R. Aguilar 2224, MO) B V. amistadensis (G. McPherson 8703, MO) C V. chrysocarpa (O. Valverde 970, MO) D V. elongata (G. McPherson 13674, MO) E V. fosteri (G. McPherson 20913, MO) F V. guatemalensis (R. Cedillo 3349, MO) G V. koschnyi (R. E. Gereau et al. 3455, MO) H V. laevigata (R. Aguilar 2004, MO) I V. megacarpa (J. A. Duke 15261, MO) J V. montana (E. Bello 855, MO) K V. multiflora (R. Rueda et al. 8196, MO; inset, J. C. Sandino 3302, MO) L V. nobilis from Barro Colorado (R. J. Schmalzel 320, MO) M V. otobifolia (G. de Nevers & H. Herrera 7056, MO) N V. sebifera (F. Almeda et al. 5553, MO) O V. nobilis from Osa Peninsula (R. Aguilar 11186, MO).	Figure 3. Trichomes in Mesoamerican Virola A Virola allenii (R. Aguilar 2224, MO) B V. amistadensis (G. McPherson 8703, MO) C V. chrysocarpa (O. Valverde 970, MO) D V. elongata (G. McPherson 13674, MO) E V. fosteri (G. McPherson 20913, MO) F V. guatemalensis (R. Cedillo 3349, MO) G V. koschnyi (R. E. Gereau et al. 3455, MO) H V. laevigata (R. Aguilar 2004, MO) I V. megacarpa (J. A. Duke 15261, MO) J V. montana (E. Bello 855, MO) K V. multiflora (R. Rueda et al. 8196, MO; inset, J. C. Sandino 3302, MO) L V. nobilis from Barro Colorado (R. J. Schmalzel 320, MO) M V. otobifolia (G. de Nevers & H. Herrera 7056, MO) N V. sebifera (F. Almeda et al. 5553, MO) O V. nobilis from Osa Peninsula (R. Aguilar 11186, MO).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
A5F614C7BCBE54FE9642232C771993C1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349090	https://doi.org/10.3897/phytokeys.134.37979.figure8	Figure 8. Pattern of the lateral veins in Mesoamerican Virola A Virola allenii (K. Thomsen 1284, NO) B V. amistadensis (G. McPherson 8703, MO) C V. chrysocarpa (B. Hammel et al. 16864, MO) D V. elongata (M. Correa & R. L. Dressler 1078, MO) E V. fosteri (G. de Nevers 7226, MO) F V. guatemalensis (G. Ibarra 957, MO) G V. koschnyi (J. Miller & J. C. Sandino 1110, MO) H V. laevigata (N. Zamora & T. D. Pennington 1583, MO) I V. megacarpa (G. de Nevers 5184, MO) J V. montana (E. Lepiz & J. F. Morales 284, MO) K V. multiflora (J. Manzanares 3561, MO) L V. nobilis from Barro Colorado (R. J. Schmalzel 320, MO) M V. otobifolia (G. de Nevers et al. 7530, MO) N V. sebifera (T. B. Croat 5959, MO) O V. nobilis from Osa Peninsula (R. Aguilar 11186, MO).	Figure 8. Pattern of the lateral veins in Mesoamerican Virola A Virola allenii (K. Thomsen 1284, NO) B V. amistadensis (G. McPherson 8703, MO) C V. chrysocarpa (B. Hammel et al. 16864, MO) D V. elongata (M. Correa & R. L. Dressler 1078, MO) E V. fosteri (G. de Nevers 7226, MO) F V. guatemalensis (G. Ibarra 957, MO) G V. koschnyi (J. Miller & J. C. Sandino 1110, MO) H V. laevigata (N. Zamora & T. D. Pennington 1583, MO) I V. megacarpa (G. de Nevers 5184, MO) J V. montana (E. Lepiz & J. F. Morales 284, MO) K V. multiflora (J. Manzanares 3561, MO) L V. nobilis from Barro Colorado (R. J. Schmalzel 320, MO) M V. otobifolia (G. de Nevers et al. 7530, MO) N V. sebifera (T. B. Croat 5959, MO) O V. nobilis from Osa Peninsula (R. Aguilar 11186, MO).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
A5F614C7BCBE54FE9642232C771993C1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349086	https://doi.org/10.3897/phytokeys.134.37979.figure4	Figure 4. Fruits of Mesoameican Virola, organised in groups of morphologically similar species A V. allenii (P. H. Allen 6727, GH) B V. amistadensis (G. McPherson 9715, MO) C V. otobifolia (A. Gentry & S. Mori 14199, MO) D V. macrocarpa (A. E. Lawrance 675, MO) E V. chrysocarpa (B. Hammel et al. 16864, MO) F V. koschnyi (R. Robles 1587, MO) G V. guatemalensis (S. Sinaca 1542, MO) H V. montana (J. Utley & K. Utley 5054, MO) I V. fosteri (S. Mori & J. Kallunki 4891, MO) J V. multiflora (R. Rueda et a. l 9960, MO) K V. sebifera (T. B. Croat 5959, MO) L V. elongata (J. D. Dwyer & M. Correa 8420, MO) M V. laevigata (R. Aguilar 1585, MO) N V. nobilis from Barro Colorado (T. B. Croat 8090, MO) O V. nobilis from the Osa Peninsula (G. Herrera 4222, MO) P V. megacarpa (G. de Nevers 5184, MO). Scale bars: 1 cm.	Figure 4. Fruits of Mesoameican Virola, organised in groups of morphologically similar species A V. allenii (P. H. Allen 6727, GH) B V. amistadensis (G. McPherson 9715, MO) C V. otobifolia (A. Gentry & S. Mori 14199, MO) D V. macrocarpa (A. E. Lawrance 675, MO) E V. chrysocarpa (B. Hammel et al. 16864, MO) F V. koschnyi (R. Robles 1587, MO) G V. guatemalensis (S. Sinaca 1542, MO) H V. montana (J. Utley & K. Utley 5054, MO) I V. fosteri (S. Mori & J. Kallunki 4891, MO) J V. multiflora (R. Rueda et a. l 9960, MO) K V. sebifera (T. B. Croat 5959, MO) L V. elongata (J. D. Dwyer & M. Correa 8420, MO) M V. laevigata (R. Aguilar 1585, MO) N V. nobilis from Barro Colorado (T. B. Croat 8090, MO) O V. nobilis from the Osa Peninsula (G. Herrera 4222, MO) P V. megacarpa (G. de Nevers 5184, MO). Scale bars: 1 cm.	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
A5F614C7BCBE54FE9642232C771993C1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349100	https://doi.org/10.3897/phytokeys.134.37979.figure18	Figure 18. Geographic distribution of Virola koschnyi (A), V. laevigata (B), V. megacarpa (C), V. montana (D), V. multiflora (E) and V. nobilis (F).	Figure 18. Geographic distribution of Virola koschnyi (A), V. laevigata (B), V. megacarpa (C), V. montana (D), V. multiflora (E) and V. nobilis (F).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
CA381C53D596526F974179D43FC36DB2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349101	https://doi.org/10.3897/phytokeys.134.37979.figure19	Figure 19. Virola laevigata A branch with staminate inflorescences B leaf blades on abaxial surface, inset showing the glabrous midvein C twig, petiole and leaf on adaxial surface D close up of staminate inflorescences E leaf base F leaf apex, inset showing leaf punctations G branch with immature fruits H close-up of immature fruit I open fruit, showing the aril. Photos by Reinaldo Aguilar.	Figure 19. Virola laevigata A branch with staminate inflorescences B leaf blades on abaxial surface, inset showing the glabrous midvein C twig, petiole and leaf on adaxial surface D close up of staminate inflorescences E leaf base F leaf apex, inset showing leaf punctations G branch with immature fruits H close-up of immature fruit I open fruit, showing the aril. Photos by Reinaldo Aguilar.	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
CA381C53D596526F974179D43FC36DB2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349085	https://doi.org/10.3897/phytokeys.134.37979.figure3	Figure 3. Trichomes in Mesoamerican Virola A Virola allenii (R. Aguilar 2224, MO) B V. amistadensis (G. McPherson 8703, MO) C V. chrysocarpa (O. Valverde 970, MO) D V. elongata (G. McPherson 13674, MO) E V. fosteri (G. McPherson 20913, MO) F V. guatemalensis (R. Cedillo 3349, MO) G V. koschnyi (R. E. Gereau et al. 3455, MO) H V. laevigata (R. Aguilar 2004, MO) I V. megacarpa (J. A. Duke 15261, MO) J V. montana (E. Bello 855, MO) K V. multiflora (R. Rueda et al. 8196, MO; inset, J. C. Sandino 3302, MO) L V. nobilis from Barro Colorado (R. J. Schmalzel 320, MO) M V. otobifolia (G. de Nevers & H. Herrera 7056, MO) N V. sebifera (F. Almeda et al. 5553, MO) O V. nobilis from Osa Peninsula (R. Aguilar 11186, MO).	Figure 3. Trichomes in Mesoamerican Virola A Virola allenii (R. Aguilar 2224, MO) B V. amistadensis (G. McPherson 8703, MO) C V. chrysocarpa (O. Valverde 970, MO) D V. elongata (G. McPherson 13674, MO) E V. fosteri (G. McPherson 20913, MO) F V. guatemalensis (R. Cedillo 3349, MO) G V. koschnyi (R. E. Gereau et al. 3455, MO) H V. laevigata (R. Aguilar 2004, MO) I V. megacarpa (J. A. Duke 15261, MO) J V. montana (E. Bello 855, MO) K V. multiflora (R. Rueda et al. 8196, MO; inset, J. C. Sandino 3302, MO) L V. nobilis from Barro Colorado (R. J. Schmalzel 320, MO) M V. otobifolia (G. de Nevers & H. Herrera 7056, MO) N V. sebifera (F. Almeda et al. 5553, MO) O V. nobilis from Osa Peninsula (R. Aguilar 11186, MO).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
CA381C53D596526F974179D43FC36DB2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349086	https://doi.org/10.3897/phytokeys.134.37979.figure4	Figure 4. Fruits of Mesoameican Virola, organised in groups of morphologically similar species A V. allenii (P. H. Allen 6727, GH) B V. amistadensis (G. McPherson 9715, MO) C V. otobifolia (A. Gentry & S. Mori 14199, MO) D V. macrocarpa (A. E. Lawrance 675, MO) E V. chrysocarpa (B. Hammel et al. 16864, MO) F V. koschnyi (R. Robles 1587, MO) G V. guatemalensis (S. Sinaca 1542, MO) H V. montana (J. Utley & K. Utley 5054, MO) I V. fosteri (S. Mori & J. Kallunki 4891, MO) J V. multiflora (R. Rueda et a. l 9960, MO) K V. sebifera (T. B. Croat 5959, MO) L V. elongata (J. D. Dwyer & M. Correa 8420, MO) M V. laevigata (R. Aguilar 1585, MO) N V. nobilis from Barro Colorado (T. B. Croat 8090, MO) O V. nobilis from the Osa Peninsula (G. Herrera 4222, MO) P V. megacarpa (G. de Nevers 5184, MO). Scale bars: 1 cm.	Figure 4. Fruits of Mesoameican Virola, organised in groups of morphologically similar species A V. allenii (P. H. Allen 6727, GH) B V. amistadensis (G. McPherson 9715, MO) C V. otobifolia (A. Gentry & S. Mori 14199, MO) D V. macrocarpa (A. E. Lawrance 675, MO) E V. chrysocarpa (B. Hammel et al. 16864, MO) F V. koschnyi (R. Robles 1587, MO) G V. guatemalensis (S. Sinaca 1542, MO) H V. montana (J. Utley & K. Utley 5054, MO) I V. fosteri (S. Mori & J. Kallunki 4891, MO) J V. multiflora (R. Rueda et a. l 9960, MO) K V. sebifera (T. B. Croat 5959, MO) L V. elongata (J. D. Dwyer & M. Correa 8420, MO) M V. laevigata (R. Aguilar 1585, MO) N V. nobilis from Barro Colorado (T. B. Croat 8090, MO) O V. nobilis from the Osa Peninsula (G. Herrera 4222, MO) P V. megacarpa (G. de Nevers 5184, MO). Scale bars: 1 cm.	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
CA381C53D596526F974179D43FC36DB2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349100	https://doi.org/10.3897/phytokeys.134.37979.figure18	Figure 18. Geographic distribution of Virola koschnyi (A), V. laevigata (B), V. megacarpa (C), V. montana (D), V. multiflora (E) and V. nobilis (F).	Figure 18. Geographic distribution of Virola koschnyi (A), V. laevigata (B), V. megacarpa (C), V. montana (D), V. multiflora (E) and V. nobilis (F).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
5FE769BF14B05418B4D5ED9739AE500C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349102	https://doi.org/10.3897/phytokeys.134.37979.figure20	Figure 20. A. Virola megacarpa A branch with fruits, both leaf blade surfaces shown B fruit. Virola nobilis from Barro Colorado C lower trunk and buttress D branch with fruits, leaf blades on both surfaces E pistillate flowers F staminate flower and bud G fruits. Photos by Carmen Galdames (A, B), Rolando Perez (C, D, G), Steven Paton (E, F); all photos from https: // stricollections. org / portal / index. php.	Figure 20. A. Virola megacarpa A branch with fruits, both leaf blade surfaces shown B fruit. Virola nobilis from Barro Colorado C lower trunk and buttress D branch with fruits, leaf blades on both surfaces E pistillate flowers F staminate flower and bud G fruits. Photos by Carmen Galdames (A, B), Rolando Perez (C, D, G), Steven Paton (E, F); all photos from https: // stricollections. org / portal / index. php.	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
5FE769BF14B05418B4D5ED9739AE500C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349085	https://doi.org/10.3897/phytokeys.134.37979.figure3	Figure 3. Trichomes in Mesoamerican Virola A Virola allenii (R. Aguilar 2224, MO) B V. amistadensis (G. McPherson 8703, MO) C V. chrysocarpa (O. Valverde 970, MO) D V. elongata (G. McPherson 13674, MO) E V. fosteri (G. McPherson 20913, MO) F V. guatemalensis (R. Cedillo 3349, MO) G V. koschnyi (R. E. Gereau et al. 3455, MO) H V. laevigata (R. Aguilar 2004, MO) I V. megacarpa (J. A. Duke 15261, MO) J V. montana (E. Bello 855, MO) K V. multiflora (R. Rueda et al. 8196, MO; inset, J. C. Sandino 3302, MO) L V. nobilis from Barro Colorado (R. J. Schmalzel 320, MO) M V. otobifolia (G. de Nevers & H. Herrera 7056, MO) N V. sebifera (F. Almeda et al. 5553, MO) O V. nobilis from Osa Peninsula (R. Aguilar 11186, MO).	Figure 3. Trichomes in Mesoamerican Virola A Virola allenii (R. Aguilar 2224, MO) B V. amistadensis (G. McPherson 8703, MO) C V. chrysocarpa (O. Valverde 970, MO) D V. elongata (G. McPherson 13674, MO) E V. fosteri (G. McPherson 20913, MO) F V. guatemalensis (R. Cedillo 3349, MO) G V. koschnyi (R. E. Gereau et al. 3455, MO) H V. laevigata (R. Aguilar 2004, MO) I V. megacarpa (J. A. Duke 15261, MO) J V. montana (E. Bello 855, MO) K V. multiflora (R. Rueda et al. 8196, MO; inset, J. C. Sandino 3302, MO) L V. nobilis from Barro Colorado (R. J. Schmalzel 320, MO) M V. otobifolia (G. de Nevers & H. Herrera 7056, MO) N V. sebifera (F. Almeda et al. 5553, MO) O V. nobilis from Osa Peninsula (R. Aguilar 11186, MO).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
5FE769BF14B05418B4D5ED9739AE500C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349086	https://doi.org/10.3897/phytokeys.134.37979.figure4	Figure 4. Fruits of Mesoameican Virola, organised in groups of morphologically similar species A V. allenii (P. H. Allen 6727, GH) B V. amistadensis (G. McPherson 9715, MO) C V. otobifolia (A. Gentry & S. Mori 14199, MO) D V. macrocarpa (A. E. Lawrance 675, MO) E V. chrysocarpa (B. Hammel et al. 16864, MO) F V. koschnyi (R. Robles 1587, MO) G V. guatemalensis (S. Sinaca 1542, MO) H V. montana (J. Utley & K. Utley 5054, MO) I V. fosteri (S. Mori & J. Kallunki 4891, MO) J V. multiflora (R. Rueda et a. l 9960, MO) K V. sebifera (T. B. Croat 5959, MO) L V. elongata (J. D. Dwyer & M. Correa 8420, MO) M V. laevigata (R. Aguilar 1585, MO) N V. nobilis from Barro Colorado (T. B. Croat 8090, MO) O V. nobilis from the Osa Peninsula (G. Herrera 4222, MO) P V. megacarpa (G. de Nevers 5184, MO). Scale bars: 1 cm.	Figure 4. Fruits of Mesoameican Virola, organised in groups of morphologically similar species A V. allenii (P. H. Allen 6727, GH) B V. amistadensis (G. McPherson 9715, MO) C V. otobifolia (A. Gentry & S. Mori 14199, MO) D V. macrocarpa (A. E. Lawrance 675, MO) E V. chrysocarpa (B. Hammel et al. 16864, MO) F V. koschnyi (R. Robles 1587, MO) G V. guatemalensis (S. Sinaca 1542, MO) H V. montana (J. Utley & K. Utley 5054, MO) I V. fosteri (S. Mori & J. Kallunki 4891, MO) J V. multiflora (R. Rueda et a. l 9960, MO) K V. sebifera (T. B. Croat 5959, MO) L V. elongata (J. D. Dwyer & M. Correa 8420, MO) M V. laevigata (R. Aguilar 1585, MO) N V. nobilis from Barro Colorado (T. B. Croat 8090, MO) O V. nobilis from the Osa Peninsula (G. Herrera 4222, MO) P V. megacarpa (G. de Nevers 5184, MO). Scale bars: 1 cm.	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
5FE769BF14B05418B4D5ED9739AE500C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349100	https://doi.org/10.3897/phytokeys.134.37979.figure18	Figure 18. Geographic distribution of Virola koschnyi (A), V. laevigata (B), V. megacarpa (C), V. montana (D), V. multiflora (E) and V. nobilis (F).	Figure 18. Geographic distribution of Virola koschnyi (A), V. laevigata (B), V. megacarpa (C), V. montana (D), V. multiflora (E) and V. nobilis (F).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
5FE769BF14B05418B4D5ED9739AE500C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349090	https://doi.org/10.3897/phytokeys.134.37979.figure8	Figure 8. Pattern of the lateral veins in Mesoamerican Virola A Virola allenii (K. Thomsen 1284, NO) B V. amistadensis (G. McPherson 8703, MO) C V. chrysocarpa (B. Hammel et al. 16864, MO) D V. elongata (M. Correa & R. L. Dressler 1078, MO) E V. fosteri (G. de Nevers 7226, MO) F V. guatemalensis (G. Ibarra 957, MO) G V. koschnyi (J. Miller & J. C. Sandino 1110, MO) H V. laevigata (N. Zamora & T. D. Pennington 1583, MO) I V. megacarpa (G. de Nevers 5184, MO) J V. montana (E. Lepiz & J. F. Morales 284, MO) K V. multiflora (J. Manzanares 3561, MO) L V. nobilis from Barro Colorado (R. J. Schmalzel 320, MO) M V. otobifolia (G. de Nevers et al. 7530, MO) N V. sebifera (T. B. Croat 5959, MO) O V. nobilis from Osa Peninsula (R. Aguilar 11186, MO).	Figure 8. Pattern of the lateral veins in Mesoamerican Virola A Virola allenii (K. Thomsen 1284, NO) B V. amistadensis (G. McPherson 8703, MO) C V. chrysocarpa (B. Hammel et al. 16864, MO) D V. elongata (M. Correa & R. L. Dressler 1078, MO) E V. fosteri (G. de Nevers 7226, MO) F V. guatemalensis (G. Ibarra 957, MO) G V. koschnyi (J. Miller & J. C. Sandino 1110, MO) H V. laevigata (N. Zamora & T. D. Pennington 1583, MO) I V. megacarpa (G. de Nevers 5184, MO) J V. montana (E. Lepiz & J. F. Morales 284, MO) K V. multiflora (J. Manzanares 3561, MO) L V. nobilis from Barro Colorado (R. J. Schmalzel 320, MO) M V. otobifolia (G. de Nevers et al. 7530, MO) N V. sebifera (T. B. Croat 5959, MO) O V. nobilis from Osa Peninsula (R. Aguilar 11186, MO).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
A1E94AB74FA55164BE92C204B2AF880E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349103	https://doi.org/10.3897/phytokeys.134.37979.figure21	Figure 21. Virola montana A branch with leaves and inflorescences B fruit C flowers. Drawn by Pedro Juarez based on G. Herrera et al. 514 (A-C), and R. Lent 3899 (B).	Figure 21. Virola montana A branch with leaves and inflorescences B fruit C flowers. Drawn by Pedro Juarez based on G. Herrera et al. 514 (A-C), and R. Lent 3899 (B).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
A1E94AB74FA55164BE92C204B2AF880E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349104	https://doi.org/10.3897/phytokeys.134.37979.figure22	Figure 22. Virola montana A juvenile tree showing branching pattern B leaf blades on adaxial side C leaf blades on abaxial side D venation. Virola multiflora. E Branch with staminate inflorescences, inset showing twig and flowers F twig, leaf blade on abaxial surface and inflorescences, inset showing immature fruit G lateral view of a perianth H close-up of staminate flowers. Photos by J. Esteban Jimenez (A-D); B. Hammel (E-H), Indiana Coronado (F, inset).	Figure 22. Virola montana A juvenile tree showing branching pattern B leaf blades on adaxial side C leaf blades on abaxial side D venation. Virola multiflora. E Branch with staminate inflorescences, inset showing twig and flowers F twig, leaf blade on abaxial surface and inflorescences, inset showing immature fruit G lateral view of a perianth H close-up of staminate flowers. Photos by J. Esteban Jimenez (A-D); B. Hammel (E-H), Indiana Coronado (F, inset).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
A1E94AB74FA55164BE92C204B2AF880E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349085	https://doi.org/10.3897/phytokeys.134.37979.figure3	Figure 3. Trichomes in Mesoamerican Virola A Virola allenii (R. Aguilar 2224, MO) B V. amistadensis (G. McPherson 8703, MO) C V. chrysocarpa (O. Valverde 970, MO) D V. elongata (G. McPherson 13674, MO) E V. fosteri (G. McPherson 20913, MO) F V. guatemalensis (R. Cedillo 3349, MO) G V. koschnyi (R. E. Gereau et al. 3455, MO) H V. laevigata (R. Aguilar 2004, MO) I V. megacarpa (J. A. Duke 15261, MO) J V. montana (E. Bello 855, MO) K V. multiflora (R. Rueda et al. 8196, MO; inset, J. C. Sandino 3302, MO) L V. nobilis from Barro Colorado (R. J. Schmalzel 320, MO) M V. otobifolia (G. de Nevers & H. Herrera 7056, MO) N V. sebifera (F. Almeda et al. 5553, MO) O V. nobilis from Osa Peninsula (R. Aguilar 11186, MO).	Figure 3. Trichomes in Mesoamerican Virola A Virola allenii (R. Aguilar 2224, MO) B V. amistadensis (G. McPherson 8703, MO) C V. chrysocarpa (O. Valverde 970, MO) D V. elongata (G. McPherson 13674, MO) E V. fosteri (G. McPherson 20913, MO) F V. guatemalensis (R. Cedillo 3349, MO) G V. koschnyi (R. E. Gereau et al. 3455, MO) H V. laevigata (R. Aguilar 2004, MO) I V. megacarpa (J. A. Duke 15261, MO) J V. montana (E. Bello 855, MO) K V. multiflora (R. Rueda et al. 8196, MO; inset, J. C. Sandino 3302, MO) L V. nobilis from Barro Colorado (R. J. Schmalzel 320, MO) M V. otobifolia (G. de Nevers & H. Herrera 7056, MO) N V. sebifera (F. Almeda et al. 5553, MO) O V. nobilis from Osa Peninsula (R. Aguilar 11186, MO).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
A1E94AB74FA55164BE92C204B2AF880E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349086	https://doi.org/10.3897/phytokeys.134.37979.figure4	Figure 4. Fruits of Mesoameican Virola, organised in groups of morphologically similar species A V. allenii (P. H. Allen 6727, GH) B V. amistadensis (G. McPherson 9715, MO) C V. otobifolia (A. Gentry & S. Mori 14199, MO) D V. macrocarpa (A. E. Lawrance 675, MO) E V. chrysocarpa (B. Hammel et al. 16864, MO) F V. koschnyi (R. Robles 1587, MO) G V. guatemalensis (S. Sinaca 1542, MO) H V. montana (J. Utley & K. Utley 5054, MO) I V. fosteri (S. Mori & J. Kallunki 4891, MO) J V. multiflora (R. Rueda et a. l 9960, MO) K V. sebifera (T. B. Croat 5959, MO) L V. elongata (J. D. Dwyer & M. Correa 8420, MO) M V. laevigata (R. Aguilar 1585, MO) N V. nobilis from Barro Colorado (T. B. Croat 8090, MO) O V. nobilis from the Osa Peninsula (G. Herrera 4222, MO) P V. megacarpa (G. de Nevers 5184, MO). Scale bars: 1 cm.	Figure 4. Fruits of Mesoameican Virola, organised in groups of morphologically similar species A V. allenii (P. H. Allen 6727, GH) B V. amistadensis (G. McPherson 9715, MO) C V. otobifolia (A. Gentry & S. Mori 14199, MO) D V. macrocarpa (A. E. Lawrance 675, MO) E V. chrysocarpa (B. Hammel et al. 16864, MO) F V. koschnyi (R. Robles 1587, MO) G V. guatemalensis (S. Sinaca 1542, MO) H V. montana (J. Utley & K. Utley 5054, MO) I V. fosteri (S. Mori & J. Kallunki 4891, MO) J V. multiflora (R. Rueda et a. l 9960, MO) K V. sebifera (T. B. Croat 5959, MO) L V. elongata (J. D. Dwyer & M. Correa 8420, MO) M V. laevigata (R. Aguilar 1585, MO) N V. nobilis from Barro Colorado (T. B. Croat 8090, MO) O V. nobilis from the Osa Peninsula (G. Herrera 4222, MO) P V. megacarpa (G. de Nevers 5184, MO). Scale bars: 1 cm.	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
A1E94AB74FA55164BE92C204B2AF880E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349100	https://doi.org/10.3897/phytokeys.134.37979.figure18	Figure 18. Geographic distribution of Virola koschnyi (A), V. laevigata (B), V. megacarpa (C), V. montana (D), V. multiflora (E) and V. nobilis (F).	Figure 18. Geographic distribution of Virola koschnyi (A), V. laevigata (B), V. megacarpa (C), V. montana (D), V. multiflora (E) and V. nobilis (F).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
A1E94AB74FA55164BE92C204B2AF880E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349090	https://doi.org/10.3897/phytokeys.134.37979.figure8	Figure 8. Pattern of the lateral veins in Mesoamerican Virola A Virola allenii (K. Thomsen 1284, NO) B V. amistadensis (G. McPherson 8703, MO) C V. chrysocarpa (B. Hammel et al. 16864, MO) D V. elongata (M. Correa & R. L. Dressler 1078, MO) E V. fosteri (G. de Nevers 7226, MO) F V. guatemalensis (G. Ibarra 957, MO) G V. koschnyi (J. Miller & J. C. Sandino 1110, MO) H V. laevigata (N. Zamora & T. D. Pennington 1583, MO) I V. megacarpa (G. de Nevers 5184, MO) J V. montana (E. Lepiz & J. F. Morales 284, MO) K V. multiflora (J. Manzanares 3561, MO) L V. nobilis from Barro Colorado (R. J. Schmalzel 320, MO) M V. otobifolia (G. de Nevers et al. 7530, MO) N V. sebifera (T. B. Croat 5959, MO) O V. nobilis from Osa Peninsula (R. Aguilar 11186, MO).	Figure 8. Pattern of the lateral veins in Mesoamerican Virola A Virola allenii (K. Thomsen 1284, NO) B V. amistadensis (G. McPherson 8703, MO) C V. chrysocarpa (B. Hammel et al. 16864, MO) D V. elongata (M. Correa & R. L. Dressler 1078, MO) E V. fosteri (G. de Nevers 7226, MO) F V. guatemalensis (G. Ibarra 957, MO) G V. koschnyi (J. Miller & J. C. Sandino 1110, MO) H V. laevigata (N. Zamora & T. D. Pennington 1583, MO) I V. megacarpa (G. de Nevers 5184, MO) J V. montana (E. Lepiz & J. F. Morales 284, MO) K V. multiflora (J. Manzanares 3561, MO) L V. nobilis from Barro Colorado (R. J. Schmalzel 320, MO) M V. otobifolia (G. de Nevers et al. 7530, MO) N V. sebifera (T. B. Croat 5959, MO) O V. nobilis from Osa Peninsula (R. Aguilar 11186, MO).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
A1E94AB74FA55164BE92C204B2AF880E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349092	https://doi.org/10.3897/phytokeys.134.37979.figure10	Figure 10. Virola calophylla A branch with leaves, showing both sides B branch with inflorescences. Virola elongata C leafy branch with inflorescences, inset fruit D inflorescences. Virola guatemalensis E, F leaf blades showing adaxial (E) and abaxial (F) surface. G young twigs and petioles. Photos by Robin Foster (A, B), from https: // plantidtools. fieldmuseum. org / en / nlp; Steven Paton (C, D), Rolando Perez (C inset), from https: // stricollections. org / portal / index. php and Angela Rojas (E-G).	Figure 10. Virola calophylla A branch with leaves, showing both sides B branch with inflorescences. Virola elongata C leafy branch with inflorescences, inset fruit D inflorescences. Virola guatemalensis E, F leaf blades showing adaxial (E) and abaxial (F) surface. G young twigs and petioles. Photos by Robin Foster (A, B), from https: // plantidtools. fieldmuseum. org / en / nlp; Steven Paton (C, D), Rolando Perez (C inset), from https: // stricollections. org / portal / index. php and Angela Rojas (E-G).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
04AE7760CAC258C993F6C8A0A4F5374C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349104	https://doi.org/10.3897/phytokeys.134.37979.figure22	Figure 22. Virola montana A juvenile tree showing branching pattern B leaf blades on adaxial side C leaf blades on abaxial side D venation. Virola multiflora. E Branch with staminate inflorescences, inset showing twig and flowers F twig, leaf blade on abaxial surface and inflorescences, inset showing immature fruit G lateral view of a perianth H close-up of staminate flowers. Photos by J. Esteban Jimenez (A-D); B. Hammel (E-H), Indiana Coronado (F, inset).	Figure 22. Virola montana A juvenile tree showing branching pattern B leaf blades on adaxial side C leaf blades on abaxial side D venation. Virola multiflora. E Branch with staminate inflorescences, inset showing twig and flowers F twig, leaf blade on abaxial surface and inflorescences, inset showing immature fruit G lateral view of a perianth H close-up of staminate flowers. Photos by J. Esteban Jimenez (A-D); B. Hammel (E-H), Indiana Coronado (F, inset).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
04AE7760CAC258C993F6C8A0A4F5374C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349085	https://doi.org/10.3897/phytokeys.134.37979.figure3	Figure 3. Trichomes in Mesoamerican Virola A Virola allenii (R. Aguilar 2224, MO) B V. amistadensis (G. McPherson 8703, MO) C V. chrysocarpa (O. Valverde 970, MO) D V. elongata (G. McPherson 13674, MO) E V. fosteri (G. McPherson 20913, MO) F V. guatemalensis (R. Cedillo 3349, MO) G V. koschnyi (R. E. Gereau et al. 3455, MO) H V. laevigata (R. Aguilar 2004, MO) I V. megacarpa (J. A. Duke 15261, MO) J V. montana (E. Bello 855, MO) K V. multiflora (R. Rueda et al. 8196, MO; inset, J. C. Sandino 3302, MO) L V. nobilis from Barro Colorado (R. J. Schmalzel 320, MO) M V. otobifolia (G. de Nevers & H. Herrera 7056, MO) N V. sebifera (F. Almeda et al. 5553, MO) O V. nobilis from Osa Peninsula (R. Aguilar 11186, MO).	Figure 3. Trichomes in Mesoamerican Virola A Virola allenii (R. Aguilar 2224, MO) B V. amistadensis (G. McPherson 8703, MO) C V. chrysocarpa (O. Valverde 970, MO) D V. elongata (G. McPherson 13674, MO) E V. fosteri (G. McPherson 20913, MO) F V. guatemalensis (R. Cedillo 3349, MO) G V. koschnyi (R. E. Gereau et al. 3455, MO) H V. laevigata (R. Aguilar 2004, MO) I V. megacarpa (J. A. Duke 15261, MO) J V. montana (E. Bello 855, MO) K V. multiflora (R. Rueda et al. 8196, MO; inset, J. C. Sandino 3302, MO) L V. nobilis from Barro Colorado (R. J. Schmalzel 320, MO) M V. otobifolia (G. de Nevers & H. Herrera 7056, MO) N V. sebifera (F. Almeda et al. 5553, MO) O V. nobilis from Osa Peninsula (R. Aguilar 11186, MO).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
04AE7760CAC258C993F6C8A0A4F5374C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349090	https://doi.org/10.3897/phytokeys.134.37979.figure8	Figure 8. Pattern of the lateral veins in Mesoamerican Virola A Virola allenii (K. Thomsen 1284, NO) B V. amistadensis (G. McPherson 8703, MO) C V. chrysocarpa (B. Hammel et al. 16864, MO) D V. elongata (M. Correa & R. L. Dressler 1078, MO) E V. fosteri (G. de Nevers 7226, MO) F V. guatemalensis (G. Ibarra 957, MO) G V. koschnyi (J. Miller & J. C. Sandino 1110, MO) H V. laevigata (N. Zamora & T. D. Pennington 1583, MO) I V. megacarpa (G. de Nevers 5184, MO) J V. montana (E. Lepiz & J. F. Morales 284, MO) K V. multiflora (J. Manzanares 3561, MO) L V. nobilis from Barro Colorado (R. J. Schmalzel 320, MO) M V. otobifolia (G. de Nevers et al. 7530, MO) N V. sebifera (T. B. Croat 5959, MO) O V. nobilis from Osa Peninsula (R. Aguilar 11186, MO).	Figure 8. Pattern of the lateral veins in Mesoamerican Virola A Virola allenii (K. Thomsen 1284, NO) B V. amistadensis (G. McPherson 8703, MO) C V. chrysocarpa (B. Hammel et al. 16864, MO) D V. elongata (M. Correa & R. L. Dressler 1078, MO) E V. fosteri (G. de Nevers 7226, MO) F V. guatemalensis (G. Ibarra 957, MO) G V. koschnyi (J. Miller & J. C. Sandino 1110, MO) H V. laevigata (N. Zamora & T. D. Pennington 1583, MO) I V. megacarpa (G. de Nevers 5184, MO) J V. montana (E. Lepiz & J. F. Morales 284, MO) K V. multiflora (J. Manzanares 3561, MO) L V. nobilis from Barro Colorado (R. J. Schmalzel 320, MO) M V. otobifolia (G. de Nevers et al. 7530, MO) N V. sebifera (T. B. Croat 5959, MO) O V. nobilis from Osa Peninsula (R. Aguilar 11186, MO).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
04AE7760CAC258C993F6C8A0A4F5374C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349086	https://doi.org/10.3897/phytokeys.134.37979.figure4	Figure 4. Fruits of Mesoameican Virola, organised in groups of morphologically similar species A V. allenii (P. H. Allen 6727, GH) B V. amistadensis (G. McPherson 9715, MO) C V. otobifolia (A. Gentry & S. Mori 14199, MO) D V. macrocarpa (A. E. Lawrance 675, MO) E V. chrysocarpa (B. Hammel et al. 16864, MO) F V. koschnyi (R. Robles 1587, MO) G V. guatemalensis (S. Sinaca 1542, MO) H V. montana (J. Utley & K. Utley 5054, MO) I V. fosteri (S. Mori & J. Kallunki 4891, MO) J V. multiflora (R. Rueda et a. l 9960, MO) K V. sebifera (T. B. Croat 5959, MO) L V. elongata (J. D. Dwyer & M. Correa 8420, MO) M V. laevigata (R. Aguilar 1585, MO) N V. nobilis from Barro Colorado (T. B. Croat 8090, MO) O V. nobilis from the Osa Peninsula (G. Herrera 4222, MO) P V. megacarpa (G. de Nevers 5184, MO). Scale bars: 1 cm.	Figure 4. Fruits of Mesoameican Virola, organised in groups of morphologically similar species A V. allenii (P. H. Allen 6727, GH) B V. amistadensis (G. McPherson 9715, MO) C V. otobifolia (A. Gentry & S. Mori 14199, MO) D V. macrocarpa (A. E. Lawrance 675, MO) E V. chrysocarpa (B. Hammel et al. 16864, MO) F V. koschnyi (R. Robles 1587, MO) G V. guatemalensis (S. Sinaca 1542, MO) H V. montana (J. Utley & K. Utley 5054, MO) I V. fosteri (S. Mori & J. Kallunki 4891, MO) J V. multiflora (R. Rueda et a. l 9960, MO) K V. sebifera (T. B. Croat 5959, MO) L V. elongata (J. D. Dwyer & M. Correa 8420, MO) M V. laevigata (R. Aguilar 1585, MO) N V. nobilis from Barro Colorado (T. B. Croat 8090, MO) O V. nobilis from the Osa Peninsula (G. Herrera 4222, MO) P V. megacarpa (G. de Nevers 5184, MO). Scale bars: 1 cm.	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
04AE7760CAC258C993F6C8A0A4F5374C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349100	https://doi.org/10.3897/phytokeys.134.37979.figure18	Figure 18. Geographic distribution of Virola koschnyi (A), V. laevigata (B), V. megacarpa (C), V. montana (D), V. multiflora (E) and V. nobilis (F).	Figure 18. Geographic distribution of Virola koschnyi (A), V. laevigata (B), V. megacarpa (C), V. montana (D), V. multiflora (E) and V. nobilis (F).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
7C429EDB2AC852BDBB9EDF297C84E0CF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349102	https://doi.org/10.3897/phytokeys.134.37979.figure20	Figure 20. A. Virola megacarpa A branch with fruits, both leaf blade surfaces shown B fruit. Virola nobilis from Barro Colorado C lower trunk and buttress D branch with fruits, leaf blades on both surfaces E pistillate flowers F staminate flower and bud G fruits. Photos by Carmen Galdames (A, B), Rolando Perez (C, D, G), Steven Paton (E, F); all photos from https: // stricollections. org / portal / index. php.	Figure 20. A. Virola megacarpa A branch with fruits, both leaf blade surfaces shown B fruit. Virola nobilis from Barro Colorado C lower trunk and buttress D branch with fruits, leaf blades on both surfaces E pistillate flowers F staminate flower and bud G fruits. Photos by Carmen Galdames (A, B), Rolando Perez (C, D, G), Steven Paton (E, F); all photos from https: // stricollections. org / portal / index. php.	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
7C429EDB2AC852BDBB9EDF297C84E0CF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349106	https://doi.org/10.3897/phytokeys.134.37979.figure24	Figure 24. Comparisons of Virola surinamensis with collections of V. nobilis A V. surinamensis (S. Mori & R. Cardoso 17467, NY; inset showing fruits from W. E. Broadway 5264 a, MO) B V. nobilis (T. B. Croat 7488, MO) C V. nobilis from Costa Rica (B. Hammel 18017, MO). A, image courtesy of New York Botanical Garden.	Figure 24. Comparisons of Virola surinamensis with collections of V. nobilis A V. surinamensis (S. Mori & R. Cardoso 17467, NY; inset showing fruits from W. E. Broadway 5264 a, MO) B V. nobilis (T. B. Croat 7488, MO) C V. nobilis from Costa Rica (B. Hammel 18017, MO). A, image courtesy of New York Botanical Garden.	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
7C429EDB2AC852BDBB9EDF297C84E0CF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349085	https://doi.org/10.3897/phytokeys.134.37979.figure3	Figure 3. Trichomes in Mesoamerican Virola A Virola allenii (R. Aguilar 2224, MO) B V. amistadensis (G. McPherson 8703, MO) C V. chrysocarpa (O. Valverde 970, MO) D V. elongata (G. McPherson 13674, MO) E V. fosteri (G. McPherson 20913, MO) F V. guatemalensis (R. Cedillo 3349, MO) G V. koschnyi (R. E. Gereau et al. 3455, MO) H V. laevigata (R. Aguilar 2004, MO) I V. megacarpa (J. A. Duke 15261, MO) J V. montana (E. Bello 855, MO) K V. multiflora (R. Rueda et al. 8196, MO; inset, J. C. Sandino 3302, MO) L V. nobilis from Barro Colorado (R. J. Schmalzel 320, MO) M V. otobifolia (G. de Nevers & H. Herrera 7056, MO) N V. sebifera (F. Almeda et al. 5553, MO) O V. nobilis from Osa Peninsula (R. Aguilar 11186, MO).	Figure 3. Trichomes in Mesoamerican Virola A Virola allenii (R. Aguilar 2224, MO) B V. amistadensis (G. McPherson 8703, MO) C V. chrysocarpa (O. Valverde 970, MO) D V. elongata (G. McPherson 13674, MO) E V. fosteri (G. McPherson 20913, MO) F V. guatemalensis (R. Cedillo 3349, MO) G V. koschnyi (R. E. Gereau et al. 3455, MO) H V. laevigata (R. Aguilar 2004, MO) I V. megacarpa (J. A. Duke 15261, MO) J V. montana (E. Bello 855, MO) K V. multiflora (R. Rueda et al. 8196, MO; inset, J. C. Sandino 3302, MO) L V. nobilis from Barro Colorado (R. J. Schmalzel 320, MO) M V. otobifolia (G. de Nevers & H. Herrera 7056, MO) N V. sebifera (F. Almeda et al. 5553, MO) O V. nobilis from Osa Peninsula (R. Aguilar 11186, MO).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
7C429EDB2AC852BDBB9EDF297C84E0CF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349086	https://doi.org/10.3897/phytokeys.134.37979.figure4	Figure 4. Fruits of Mesoameican Virola, organised in groups of morphologically similar species A V. allenii (P. H. Allen 6727, GH) B V. amistadensis (G. McPherson 9715, MO) C V. otobifolia (A. Gentry & S. Mori 14199, MO) D V. macrocarpa (A. E. Lawrance 675, MO) E V. chrysocarpa (B. Hammel et al. 16864, MO) F V. koschnyi (R. Robles 1587, MO) G V. guatemalensis (S. Sinaca 1542, MO) H V. montana (J. Utley & K. Utley 5054, MO) I V. fosteri (S. Mori & J. Kallunki 4891, MO) J V. multiflora (R. Rueda et a. l 9960, MO) K V. sebifera (T. B. Croat 5959, MO) L V. elongata (J. D. Dwyer & M. Correa 8420, MO) M V. laevigata (R. Aguilar 1585, MO) N V. nobilis from Barro Colorado (T. B. Croat 8090, MO) O V. nobilis from the Osa Peninsula (G. Herrera 4222, MO) P V. megacarpa (G. de Nevers 5184, MO). Scale bars: 1 cm.	Figure 4. Fruits of Mesoameican Virola, organised in groups of morphologically similar species A V. allenii (P. H. Allen 6727, GH) B V. amistadensis (G. McPherson 9715, MO) C V. otobifolia (A. Gentry & S. Mori 14199, MO) D V. macrocarpa (A. E. Lawrance 675, MO) E V. chrysocarpa (B. Hammel et al. 16864, MO) F V. koschnyi (R. Robles 1587, MO) G V. guatemalensis (S. Sinaca 1542, MO) H V. montana (J. Utley & K. Utley 5054, MO) I V. fosteri (S. Mori & J. Kallunki 4891, MO) J V. multiflora (R. Rueda et a. l 9960, MO) K V. sebifera (T. B. Croat 5959, MO) L V. elongata (J. D. Dwyer & M. Correa 8420, MO) M V. laevigata (R. Aguilar 1585, MO) N V. nobilis from Barro Colorado (T. B. Croat 8090, MO) O V. nobilis from the Osa Peninsula (G. Herrera 4222, MO) P V. megacarpa (G. de Nevers 5184, MO). Scale bars: 1 cm.	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
7C429EDB2AC852BDBB9EDF297C84E0CF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349100	https://doi.org/10.3897/phytokeys.134.37979.figure18	Figure 18. Geographic distribution of Virola koschnyi (A), V. laevigata (B), V. megacarpa (C), V. montana (D), V. multiflora (E) and V. nobilis (F).	Figure 18. Geographic distribution of Virola koschnyi (A), V. laevigata (B), V. megacarpa (C), V. montana (D), V. multiflora (E) and V. nobilis (F).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
7C429EDB2AC852BDBB9EDF297C84E0CF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349105	https://doi.org/10.3897/phytokeys.134.37979.figure23	Figure 23. Virola otobifolia A branch with leaf blades showing both surfaces and inflorescences B leaf blade, venation and base on abaxial surface C infructescence D fruits E detail of fruit, showing an aril of an immature fruit. Virola surinamensis. F Branch with inflorescences G, H leaf blade on adaxial (G) and abaxial surface (H). I Infructescence J detail of fruits, including a longitudinal section of an immature fruit. Photos by Rolando Perez (A, B from https: // stricollections. org / portal / index. php); Jerry Harrison (C), Alwyn H. Gentry (D, E from http: // www. tropicos. org) and John P. Janovec (F-J from http: // atrium. andesamazon. org /).	Figure 23. Virola otobifolia A branch with leaf blades showing both surfaces and inflorescences B leaf blade, venation and base on abaxial surface C infructescence D fruits E detail of fruit, showing an aril of an immature fruit. Virola surinamensis. F Branch with inflorescences G, H leaf blade on adaxial (G) and abaxial surface (H). I Infructescence J detail of fruits, including a longitudinal section of an immature fruit. Photos by Rolando Perez (A, B from https: // stricollections. org / portal / index. php); Jerry Harrison (C), Alwyn H. Gentry (D, E from http: // www. tropicos. org) and John P. Janovec (F-J from http: // atrium. andesamazon. org /).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
7C429EDB2AC852BDBB9EDF297C84E0CF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349107	https://doi.org/10.3897/phytokeys.134.37979.figure25	Figure 25. Virola cf. nobilis from the Osa Peninsula, Costa Rica A trunk B branch with fruits C leaf blades on adaxial surface D twig, petiole and leaf base on adaxial surface E exudate on a twig F, G leaf blades on abaxial surface and trichomes (G). H Leaf margin I leaf apex J staminate inflorescences K detail of staminate inflorescences L detail of staminate flower M branch with fruits N fruits O seed. Photos by Reinaldo Aguilar.	Figure 25. Virola cf. nobilis from the Osa Peninsula, Costa Rica A trunk B branch with fruits C leaf blades on adaxial surface D twig, petiole and leaf base on adaxial surface E exudate on a twig F, G leaf blades on abaxial surface and trichomes (G). H Leaf margin I leaf apex J staminate inflorescences K detail of staminate inflorescences L detail of staminate flower M branch with fruits N fruits O seed. Photos by Reinaldo Aguilar.	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
8269F333FCF1547F96290F179013CC7F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349089	https://doi.org/10.3897/phytokeys.134.37979.figure7	Figure 7. Comparisons of Virola macrocarpa with similar species in Mesoamerica A V. allenii (P. H. Allen 5763, F; inset fruits, from P. H. Allen 6727, GH) B V. amistadensis (G. McPherson 9717, MO; inset fruits, from G. McPherson 9715, MO) C V. macrocarpa (A. E. Lawrance 675, MO) D V. otobifolia (J. P. Folsom 1440, MO). Image courtesy of Field Museum (A).	Figure 7. Comparisons of Virola macrocarpa with similar species in Mesoamerica A V. allenii (P. H. Allen 5763, F; inset fruits, from P. H. Allen 6727, GH) B V. amistadensis (G. McPherson 9717, MO; inset fruits, from G. McPherson 9715, MO) C V. macrocarpa (A. E. Lawrance 675, MO) D V. otobifolia (J. P. Folsom 1440, MO). Image courtesy of Field Museum (A).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
8269F333FCF1547F96290F179013CC7F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349105	https://doi.org/10.3897/phytokeys.134.37979.figure23	Figure 23. Virola otobifolia A branch with leaf blades showing both surfaces and inflorescences B leaf blade, venation and base on abaxial surface C infructescence D fruits E detail of fruit, showing an aril of an immature fruit. Virola surinamensis. F Branch with inflorescences G, H leaf blade on adaxial (G) and abaxial surface (H). I Infructescence J detail of fruits, including a longitudinal section of an immature fruit. Photos by Rolando Perez (A, B from https: // stricollections. org / portal / index. php); Jerry Harrison (C), Alwyn H. Gentry (D, E from http: // www. tropicos. org) and John P. Janovec (F-J from http: // atrium. andesamazon. org /).	Figure 23. Virola otobifolia A branch with leaf blades showing both surfaces and inflorescences B leaf blade, venation and base on abaxial surface C infructescence D fruits E detail of fruit, showing an aril of an immature fruit. Virola surinamensis. F Branch with inflorescences G, H leaf blade on adaxial (G) and abaxial surface (H). I Infructescence J detail of fruits, including a longitudinal section of an immature fruit. Photos by Rolando Perez (A, B from https: // stricollections. org / portal / index. php); Jerry Harrison (C), Alwyn H. Gentry (D, E from http: // www. tropicos. org) and John P. Janovec (F-J from http: // atrium. andesamazon. org /).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
8269F333FCF1547F96290F179013CC7F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349108	https://doi.org/10.3897/phytokeys.134.37979.figure26	Figure 26. Virola otobifolia A branch with leaves and inflorescences B fruits. Drawn by Pedro Juarez, based on S. Mori & J. Kallunki 5542 (A) and J. P. Folsom 1440 (B).	Figure 26. Virola otobifolia A branch with leaves and inflorescences B fruits. Drawn by Pedro Juarez, based on S. Mori & J. Kallunki 5542 (A) and J. P. Folsom 1440 (B).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
8269F333FCF1547F96290F179013CC7F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349090	https://doi.org/10.3897/phytokeys.134.37979.figure8	Figure 8. Pattern of the lateral veins in Mesoamerican Virola A Virola allenii (K. Thomsen 1284, NO) B V. amistadensis (G. McPherson 8703, MO) C V. chrysocarpa (B. Hammel et al. 16864, MO) D V. elongata (M. Correa & R. L. Dressler 1078, MO) E V. fosteri (G. de Nevers 7226, MO) F V. guatemalensis (G. Ibarra 957, MO) G V. koschnyi (J. Miller & J. C. Sandino 1110, MO) H V. laevigata (N. Zamora & T. D. Pennington 1583, MO) I V. megacarpa (G. de Nevers 5184, MO) J V. montana (E. Lepiz & J. F. Morales 284, MO) K V. multiflora (J. Manzanares 3561, MO) L V. nobilis from Barro Colorado (R. J. Schmalzel 320, MO) M V. otobifolia (G. de Nevers et al. 7530, MO) N V. sebifera (T. B. Croat 5959, MO) O V. nobilis from Osa Peninsula (R. Aguilar 11186, MO).	Figure 8. Pattern of the lateral veins in Mesoamerican Virola A Virola allenii (K. Thomsen 1284, NO) B V. amistadensis (G. McPherson 8703, MO) C V. chrysocarpa (B. Hammel et al. 16864, MO) D V. elongata (M. Correa & R. L. Dressler 1078, MO) E V. fosteri (G. de Nevers 7226, MO) F V. guatemalensis (G. Ibarra 957, MO) G V. koschnyi (J. Miller & J. C. Sandino 1110, MO) H V. laevigata (N. Zamora & T. D. Pennington 1583, MO) I V. megacarpa (G. de Nevers 5184, MO) J V. montana (E. Lepiz & J. F. Morales 284, MO) K V. multiflora (J. Manzanares 3561, MO) L V. nobilis from Barro Colorado (R. J. Schmalzel 320, MO) M V. otobifolia (G. de Nevers et al. 7530, MO) N V. sebifera (T. B. Croat 5959, MO) O V. nobilis from Osa Peninsula (R. Aguilar 11186, MO).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
8269F333FCF1547F96290F179013CC7F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349085	https://doi.org/10.3897/phytokeys.134.37979.figure3	Figure 3. Trichomes in Mesoamerican Virola A Virola allenii (R. Aguilar 2224, MO) B V. amistadensis (G. McPherson 8703, MO) C V. chrysocarpa (O. Valverde 970, MO) D V. elongata (G. McPherson 13674, MO) E V. fosteri (G. McPherson 20913, MO) F V. guatemalensis (R. Cedillo 3349, MO) G V. koschnyi (R. E. Gereau et al. 3455, MO) H V. laevigata (R. Aguilar 2004, MO) I V. megacarpa (J. A. Duke 15261, MO) J V. montana (E. Bello 855, MO) K V. multiflora (R. Rueda et al. 8196, MO; inset, J. C. Sandino 3302, MO) L V. nobilis from Barro Colorado (R. J. Schmalzel 320, MO) M V. otobifolia (G. de Nevers & H. Herrera 7056, MO) N V. sebifera (F. Almeda et al. 5553, MO) O V. nobilis from Osa Peninsula (R. Aguilar 11186, MO).	Figure 3. Trichomes in Mesoamerican Virola A Virola allenii (R. Aguilar 2224, MO) B V. amistadensis (G. McPherson 8703, MO) C V. chrysocarpa (O. Valverde 970, MO) D V. elongata (G. McPherson 13674, MO) E V. fosteri (G. McPherson 20913, MO) F V. guatemalensis (R. Cedillo 3349, MO) G V. koschnyi (R. E. Gereau et al. 3455, MO) H V. laevigata (R. Aguilar 2004, MO) I V. megacarpa (J. A. Duke 15261, MO) J V. montana (E. Bello 855, MO) K V. multiflora (R. Rueda et al. 8196, MO; inset, J. C. Sandino 3302, MO) L V. nobilis from Barro Colorado (R. J. Schmalzel 320, MO) M V. otobifolia (G. de Nevers & H. Herrera 7056, MO) N V. sebifera (F. Almeda et al. 5553, MO) O V. nobilis from Osa Peninsula (R. Aguilar 11186, MO).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
8269F333FCF1547F96290F179013CC7F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349086	https://doi.org/10.3897/phytokeys.134.37979.figure4	Figure 4. Fruits of Mesoameican Virola, organised in groups of morphologically similar species A V. allenii (P. H. Allen 6727, GH) B V. amistadensis (G. McPherson 9715, MO) C V. otobifolia (A. Gentry & S. Mori 14199, MO) D V. macrocarpa (A. E. Lawrance 675, MO) E V. chrysocarpa (B. Hammel et al. 16864, MO) F V. koschnyi (R. Robles 1587, MO) G V. guatemalensis (S. Sinaca 1542, MO) H V. montana (J. Utley & K. Utley 5054, MO) I V. fosteri (S. Mori & J. Kallunki 4891, MO) J V. multiflora (R. Rueda et a. l 9960, MO) K V. sebifera (T. B. Croat 5959, MO) L V. elongata (J. D. Dwyer & M. Correa 8420, MO) M V. laevigata (R. Aguilar 1585, MO) N V. nobilis from Barro Colorado (T. B. Croat 8090, MO) O V. nobilis from the Osa Peninsula (G. Herrera 4222, MO) P V. megacarpa (G. de Nevers 5184, MO). Scale bars: 1 cm.	Figure 4. Fruits of Mesoameican Virola, organised in groups of morphologically similar species A V. allenii (P. H. Allen 6727, GH) B V. amistadensis (G. McPherson 9715, MO) C V. otobifolia (A. Gentry & S. Mori 14199, MO) D V. macrocarpa (A. E. Lawrance 675, MO) E V. chrysocarpa (B. Hammel et al. 16864, MO) F V. koschnyi (R. Robles 1587, MO) G V. guatemalensis (S. Sinaca 1542, MO) H V. montana (J. Utley & K. Utley 5054, MO) I V. fosteri (S. Mori & J. Kallunki 4891, MO) J V. multiflora (R. Rueda et a. l 9960, MO) K V. sebifera (T. B. Croat 5959, MO) L V. elongata (J. D. Dwyer & M. Correa 8420, MO) M V. laevigata (R. Aguilar 1585, MO) N V. nobilis from Barro Colorado (T. B. Croat 8090, MO) O V. nobilis from the Osa Peninsula (G. Herrera 4222, MO) P V. megacarpa (G. de Nevers 5184, MO). Scale bars: 1 cm.	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
8269F333FCF1547F96290F179013CC7F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349109	https://doi.org/10.3897/phytokeys.134.37979.figure27	Figure 27. Geographic distribution of Virola otobifolia (A) and V. sebifera (B).	Figure 27. Geographic distribution of Virola otobifolia (A) and V. sebifera (B).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
8269F333FCF1547F96290F179013CC7F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349088	https://doi.org/10.3897/phytokeys.134.37979.figure6	Figure 6. Virola allenii A treetop B branching C lower trunk and buttress D branch with leaves, showing the adaxial surface E leaf blades on abaxial surface F exudate of the trunk G leaf blade surfaces, adaxial (above) and abaxial (below) H petiole and leaf base I mature fruit, inset left and arrow showing the galls on leaves and branches, respectively J fruits K fruit close-up. Photos by Reinaldo Aguilar.	Figure 6. Virola allenii A treetop B branching C lower trunk and buttress D branch with leaves, showing the adaxial surface E leaf blades on abaxial surface F exudate of the trunk G leaf blade surfaces, adaxial (above) and abaxial (below) H petiole and leaf base I mature fruit, inset left and arrow showing the galls on leaves and branches, respectively J fruits K fruit close-up. Photos by Reinaldo Aguilar.	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
6543C7FD082F51418D924FD226374777.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349110	https://doi.org/10.3897/phytokeys.134.37979.figure28	Figure 28. Virola sebifera A leaf blade on adaxial surface B leaf blade on abaxial surface C, E leaf base on both sides D flowering branch G staminate flowers H infructescence I fruits and aril. Photos by Reinaldo Aguilar.	Figure 28. Virola sebifera A leaf blade on adaxial surface B leaf blade on abaxial surface C, E leaf base on both sides D flowering branch G staminate flowers H infructescence I fruits and aril. Photos by Reinaldo Aguilar.	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
6543C7FD082F51418D924FD226374777.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349085	https://doi.org/10.3897/phytokeys.134.37979.figure3	Figure 3. Trichomes in Mesoamerican Virola A Virola allenii (R. Aguilar 2224, MO) B V. amistadensis (G. McPherson 8703, MO) C V. chrysocarpa (O. Valverde 970, MO) D V. elongata (G. McPherson 13674, MO) E V. fosteri (G. McPherson 20913, MO) F V. guatemalensis (R. Cedillo 3349, MO) G V. koschnyi (R. E. Gereau et al. 3455, MO) H V. laevigata (R. Aguilar 2004, MO) I V. megacarpa (J. A. Duke 15261, MO) J V. montana (E. Bello 855, MO) K V. multiflora (R. Rueda et al. 8196, MO; inset, J. C. Sandino 3302, MO) L V. nobilis from Barro Colorado (R. J. Schmalzel 320, MO) M V. otobifolia (G. de Nevers & H. Herrera 7056, MO) N V. sebifera (F. Almeda et al. 5553, MO) O V. nobilis from Osa Peninsula (R. Aguilar 11186, MO).	Figure 3. Trichomes in Mesoamerican Virola A Virola allenii (R. Aguilar 2224, MO) B V. amistadensis (G. McPherson 8703, MO) C V. chrysocarpa (O. Valverde 970, MO) D V. elongata (G. McPherson 13674, MO) E V. fosteri (G. McPherson 20913, MO) F V. guatemalensis (R. Cedillo 3349, MO) G V. koschnyi (R. E. Gereau et al. 3455, MO) H V. laevigata (R. Aguilar 2004, MO) I V. megacarpa (J. A. Duke 15261, MO) J V. montana (E. Bello 855, MO) K V. multiflora (R. Rueda et al. 8196, MO; inset, J. C. Sandino 3302, MO) L V. nobilis from Barro Colorado (R. J. Schmalzel 320, MO) M V. otobifolia (G. de Nevers & H. Herrera 7056, MO) N V. sebifera (F. Almeda et al. 5553, MO) O V. nobilis from Osa Peninsula (R. Aguilar 11186, MO).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
6543C7FD082F51418D924FD226374777.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349090	https://doi.org/10.3897/phytokeys.134.37979.figure8	Figure 8. Pattern of the lateral veins in Mesoamerican Virola A Virola allenii (K. Thomsen 1284, NO) B V. amistadensis (G. McPherson 8703, MO) C V. chrysocarpa (B. Hammel et al. 16864, MO) D V. elongata (M. Correa & R. L. Dressler 1078, MO) E V. fosteri (G. de Nevers 7226, MO) F V. guatemalensis (G. Ibarra 957, MO) G V. koschnyi (J. Miller & J. C. Sandino 1110, MO) H V. laevigata (N. Zamora & T. D. Pennington 1583, MO) I V. megacarpa (G. de Nevers 5184, MO) J V. montana (E. Lepiz & J. F. Morales 284, MO) K V. multiflora (J. Manzanares 3561, MO) L V. nobilis from Barro Colorado (R. J. Schmalzel 320, MO) M V. otobifolia (G. de Nevers et al. 7530, MO) N V. sebifera (T. B. Croat 5959, MO) O V. nobilis from Osa Peninsula (R. Aguilar 11186, MO).	Figure 8. Pattern of the lateral veins in Mesoamerican Virola A Virola allenii (K. Thomsen 1284, NO) B V. amistadensis (G. McPherson 8703, MO) C V. chrysocarpa (B. Hammel et al. 16864, MO) D V. elongata (M. Correa & R. L. Dressler 1078, MO) E V. fosteri (G. de Nevers 7226, MO) F V. guatemalensis (G. Ibarra 957, MO) G V. koschnyi (J. Miller & J. C. Sandino 1110, MO) H V. laevigata (N. Zamora & T. D. Pennington 1583, MO) I V. megacarpa (G. de Nevers 5184, MO) J V. montana (E. Lepiz & J. F. Morales 284, MO) K V. multiflora (J. Manzanares 3561, MO) L V. nobilis from Barro Colorado (R. J. Schmalzel 320, MO) M V. otobifolia (G. de Nevers et al. 7530, MO) N V. sebifera (T. B. Croat 5959, MO) O V. nobilis from Osa Peninsula (R. Aguilar 11186, MO).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
6543C7FD082F51418D924FD226374777.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349086	https://doi.org/10.3897/phytokeys.134.37979.figure4	Figure 4. Fruits of Mesoameican Virola, organised in groups of morphologically similar species A V. allenii (P. H. Allen 6727, GH) B V. amistadensis (G. McPherson 9715, MO) C V. otobifolia (A. Gentry & S. Mori 14199, MO) D V. macrocarpa (A. E. Lawrance 675, MO) E V. chrysocarpa (B. Hammel et al. 16864, MO) F V. koschnyi (R. Robles 1587, MO) G V. guatemalensis (S. Sinaca 1542, MO) H V. montana (J. Utley & K. Utley 5054, MO) I V. fosteri (S. Mori & J. Kallunki 4891, MO) J V. multiflora (R. Rueda et a. l 9960, MO) K V. sebifera (T. B. Croat 5959, MO) L V. elongata (J. D. Dwyer & M. Correa 8420, MO) M V. laevigata (R. Aguilar 1585, MO) N V. nobilis from Barro Colorado (T. B. Croat 8090, MO) O V. nobilis from the Osa Peninsula (G. Herrera 4222, MO) P V. megacarpa (G. de Nevers 5184, MO). Scale bars: 1 cm.	Figure 4. Fruits of Mesoameican Virola, organised in groups of morphologically similar species A V. allenii (P. H. Allen 6727, GH) B V. amistadensis (G. McPherson 9715, MO) C V. otobifolia (A. Gentry & S. Mori 14199, MO) D V. macrocarpa (A. E. Lawrance 675, MO) E V. chrysocarpa (B. Hammel et al. 16864, MO) F V. koschnyi (R. Robles 1587, MO) G V. guatemalensis (S. Sinaca 1542, MO) H V. montana (J. Utley & K. Utley 5054, MO) I V. fosteri (S. Mori & J. Kallunki 4891, MO) J V. multiflora (R. Rueda et a. l 9960, MO) K V. sebifera (T. B. Croat 5959, MO) L V. elongata (J. D. Dwyer & M. Correa 8420, MO) M V. laevigata (R. Aguilar 1585, MO) N V. nobilis from Barro Colorado (T. B. Croat 8090, MO) O V. nobilis from the Osa Peninsula (G. Herrera 4222, MO) P V. megacarpa (G. de Nevers 5184, MO). Scale bars: 1 cm.	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
6543C7FD082F51418D924FD226374777.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/349109	https://doi.org/10.3897/phytokeys.134.37979.figure27	Figure 27. Geographic distribution of Virola otobifolia (A) and V. sebifera (B).	Figure 27. Geographic distribution of Virola otobifolia (A) and V. sebifera (B).	2019-12-31	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.		Zenodo	biologists	Santamaria-Aguilar, Daniel;Aguilar, Reinaldo;Lagomarsino, Laura P.			
