taxonID	type	description	language	source
FA1487CEFF9D5F36D3B9513B29430E54.taxon	description	This group unites many species from the Balkans in the west to Turkmen-Khorasan mountains in the east. Species representing this group are characterized by matt dorsal side of the body, coarse and dense puncturation of pronotum and strong long setae on the apical part of the abdominal ventrite 5. Usually, representatives of this group are associated with forests, but some species are geobiontic. All Iranian taxa of the group are inhabitants of woodlands, except for probably Euboeus corrugosus.	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF9E5F37D3B9572F29A10EBC.taxon	description	(Figs 1, 2)	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF9E5F37D3B9572F29A10EBC.taxon	materials_examined	Type material. Lectotype of Helops corrugosus Seidlitz, 1895 (SNSD), designated here according to Article 61.1 and 74.1 of ICZN (1999), with labels: bronze square, “ subrugosus tut Allard ”, “ ♂ astrabad Christoph ”, “ 13916 ”, “ Helops corrugatus Seidl. ” [handwritten by G. Seidlitz], “ corrugosus Seid. nec corrugatus det. Schuster ”. The label “ Helops corrugosus Seidlitz, 1895 des. M. Nabozhenko 2021 ” was added to the lectotype. Holotype of Probaticus parthorum (ZIN), ♂ with the Cyrillic Russian handwritten label “ Астрабад, Сев. Иран 9 – 12. V. 1912 Билькевич ” (Astrabad, North Iran, leg. Bil’kevich), “ Holotypus Probathicus (Pelorinus) parthorum G. Medvedev, 1977 ”. Paratypes. 1 ♂ with labels: “ Астрабад С. Персия [print] 6. III. 904 [handwritten] Филиппович [print italic] ” (Cyrillic, Russian: Astrabad, N Persia, leg. Philippovich) and “ Paratypus Probathicus (Pelorinus) parthorum G. Medvedev, 1977 ”; 1 ♂: the same labels, but 18. II. 1904; 1 ♂: the same labels, but 5. V. [19] 04; 1 ♀: the same labels, but 23 – 24. IV. [1] 905; 1 ♀: the same labels, but 29. V. [19] 04 and additional labels “ Probaticus sp. aff. 4 - collis Bdi Dr Z. Kaszab det., 1960 ”, “ Probaticus sp. ” Red labels with “ Holotypus ” and “ Paratypus ” were pinned by a laboratory assistant (A. K. Chistaykova) with erroneous year and the name of the genus. Material. 1 ♂, 1 ♀ (HNHM): “ Persia Astrabad 5.99 Coll. Hauser ” (Iran, Golestan Prov., Gorgan, v. 1899); 1 ♂ (SMNS): Iran, Golestan Prov., Karim Ishan, 70 km NE Gonbad-e-Kavus, 4 – 7.04.1999 (leg. J. Rejsek); 2 ♂, 1 ♀ (SMNS): Iran, Golestan Prov., 10 km S Bandar e Gaz, 27.05 – 7.06.2001 (leg. G. Sama). Notes. This species was described from two males from “ Klein-Asien ” and “ Astrabad ” (Seidlitz, 1895, p. 707) in the key. Leo and Ruzzante (2016) mentioned that it is a “ mysterious ” species, described from these two different localities. Sahlberg (1913), Reitter (1922) and Nabozhenko and Löbl (2008) mentioned different species from the Aegean region. Nabozhenko (2020 b) listed E. corrugosus for Greece, Turkey and Iran. One syntype (male) from “ Asia Minor ” was originally mentioned from the collection of G. Kraatz, but it was not found in the Senckenberg Deutsche Entomologische Institut (Müncheberg, Germany). I studied the second Iranian specimen (designated here as the lectotype) (Fig. 1), which is conspecific to Medvedev’s E. parthorum (Fig. 2) also described from Astrabad. Thus, the following new synonymy is proposed: Helops corrugosus Seidlitz, 1895 = Probaticus parthorum G. Medvedev, 1976, syn. nov.	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF9E5F37D3B9572F29A10EBC.taxon	distribution	Distribution. Iran (Golestan Province).	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF985F30D3B956D62E200BDE.taxon	description	(Figs 3, 4)	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF985F30D3B956D62E200BDE.taxon	materials_examined	Type material. Holotype, ♂ (ZIN): “ Астрабад с. Персия 29. V. 905 Филиппович ” (Cyrillic, Russian) [N Persia, Astrabad, leg. Filippovich], “ Holotypus Probathicus (Pelorinus) gorganicus G. Medvedev, 1977 ”. The red label “ Holotypus ” was pinned by a laboratory assistant (A. K. Chistaykova) with an erroneous year and the name of the genus. Material. 1 ♀ (HNHM): “ Persia, Astrabad 4.99 Coll. Hauser ”, 1 ♀ (HNHM), the same label, but 5. [18] 99 (Iran, Golestan Prov., Gorgan); 2 ♂ (HNHM): Iran, Ali-abad, E Gorgan, 10. viii. 1967 (leg. Heinz); 1 ♀ (HNHM): Iran, Assala, 1200 m, 1970 (leg. Wittmer, v. Bothner); 3 ♂ (HNHM): Iran Gole [stan], Lovae (= Loveh), 750 – 1400 m (leg. Wittmer v. Bothner); 1 ♀ (SMNS): Iran, W Mazandaran, Hasankif, 19 – 21.04.1999 (leg. J. Rejsek); 3 ♀ (HNHM): Iran, Mazandaran Prov., Ghorogh, 200 m, 23.04.1999 (leg. N. Rahmé); 2 ♂ (NMPC): Iran, Mazandaran Prov., 10 km S Čalűs, 36 ° 34 ʹN, 51 ° 12 ʹE, 250 – 400 m, 21 – 23. vi. 2000 (leg. S. Kadlec); 1 ♂ (HNHM): Iran, Mazandaran Prov., 15 km S of Azad Sar, 24 – 25. iv. 2000 (leg. K. Gaskó); 1 ♂ (HNHM), same locality, 28 – 29. vi. 2000 (leg. G. Rozner); 1 ♂ (HNHM): Iran, Mazandaran Prov., 12 km S of Azad Sar, Sah Mts, 37 ° 01 ʹ 48 ʺN, 55 ° 16 ʹ 46 ʺE, 500 m, 16. v. 2000 (leg. K. Székely); 1 ♂ (SMNS): Iran, Mazandaran Prov., 12 km E Kelardasht m., 19. vi. 2003 (leg. G. Sama); 2 ♂, 7 ♀ (PCMN): Iran, Mazandaran Prov. 20 near Chalus, 36 ° 33 ʹ 20.88 ʺN, 51 ° 22 ʹ 33.49 ʺE, 25 – 27.05.2015 (leg. D. G. Kasatkin); 1 ♂, 1 ♀ (ZIN): Iran, Mazandaran Prov., 15 km above Chalus, 36 ° 33 ʹ 20.88 ʺN, 51 ° 22 ʹ 33.49 ʺE, 30 – 31.05.2017 (leg. D. G. Kasatkin); 2 ♂, 1 ♀ (PCSB): N Iran, Golestan Prov., 30 km E Minudasht, Gole-Loveh, 750 m, 21 – 27. iv. 1974 (leg. Holzschuh & Ressl); 1 ♀ (NMPC): Iran, Golestan Prov., 2 km E Tunel-e Golestan (streem valley, oak forest, at light), 37 ° 22.1 ʹN, 55 ° 59.3 ʹE, 835 m, 26 – 27. v. 2006 (leg. J. Hájek, P. Chvojka); 3 ♂ (PCMN): Iran, Golestan Prov., near Gorgan, Ziarat, 36 ° 40 ʹ 22.67 ʺN 54 ° 28 ʹ 7.68 ʺE, 27.05.2016 (leg. D. G. Kasatkin); 5 ♂ (PCMN): Iran, Golestan Prov., 20 km S Gorgan city, 36 ° 43 ʹN, 54 ° 35 ʹE, 700 m, 9. vi. 2009 (leg. A. Klimenko); 1 ♂ (PCMN): Iran, North Khorasan Prov., Golestan National park, 37 ° 22 ʹ 46.33 ʺN 55 ° 51 ʹ 5.56 ʺE, ~ 600 m, 24 – 25.05.2016 (leg. D. G. Kasatkin).	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF985F30D3B956D62E200BDE.taxon	distribution	Distribution. Iran: Mazandaran, Golestan and west part of North Khorasan provinces, in forests.	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF9A5F33D3B956D62F6B0D74.taxon	description	(Fig. 5)	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF9A5F33D3B956D62F6B0D74.taxon	description	Notes. Reitter (1902) listed “ 1 ♂ ♀ ” in the original description, but I suggest that it is a misprint, because he described only male characters and only one male is deposited in HNHM. In the same paper, he always indicated whether a character refers to a female or a male for other species and wrote “ 1 ♂ und ♀ ”.	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF9A5F33D3B956D62F6B0D74.taxon	distribution	Distribution. Iran (Talysh). Extensive collecting efforts (more than 190 years) have not produced any records of this species in the territory of the modern Azerbaijani part of Talysh, where only E. quadricollis was collected in alpine meadows. In conclusion, I’m sure that the species occurs Iranian part of the Talysh Mountains and can be excluded from the Azerbaijani faunistic lists. Reitter (1922) also recorded E. prometheus for Astrabad (Now Gorgan, Iran), probably confusing it with E. gorganicus. Kühnelt (1957) recorded E. prometheus from the Caucasus and Central Iran; the second locality is erroneous. Abdurakhamanov & Nabozhenko (2011) partly mixed E. prometheus with females of E. gorganicus and erroneously recorded it for Mazandaran Province.	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF9B5F33D3B954CF28780988.taxon	description	(Fig. 6)	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF9B5F33D3B954CF28780988.taxon	materials_examined	Type material. Syntypes of Helops vicinus were not found by curators in the Muséum national d’Histoire naturelle Paris. Holotype, ♀ (ZIN) of Probaticus medvedevi with labels: Cyrilliс “ Арм. ССР Алаверди 25.04.86 Мелях С. В. ” (Armenian SSR, Alaverdi, 25. iv. 1986, leg. S. V. Melyakh), “ Probaticus (Pelorinus) medvedevi Nabozhenko et Abdurakhmanov 2011 ”. Material. 1 ♂, 2 ♀: “ Kaukasus Leder ”, “ Allard type ”, “ Coll. Reitter ”, “ Paratypus 1877 Helops vicinus Allard ” (curator’s label). Notes. This very rare species was described in 1876 based on the specimen (s) from the collection of Reiche (Allard 1876) and redescribed in 1877 using the type specimen (s) and beetles collected by Leder (Allard 1877). Later, Schneider & Leder (1878) again published Allard’s redescription 1877 and added the species’ type locality: “ Elisabetthal ”. It was a settlement of German colonists, which is now called Asureti (Georgia, 41 ° 35 ′ 45 ″ N, 44 ° 40 ′ 00 ″ E). Both description and redescription (Allard 1876, 1877) have differences, and the latter is more consistent with specimens from Georgia and northern Armenia, as mentioned by Abdurakhmanov & Nabozhenko (2011). In addition, the Russian authors erroneously interpreted species from South Armenia and Nakhchivan as E. vicinus (described below as E. kalashiani sp. nov.) since Bogachev (1938). As a result, Abdurakhmanov & Nabozhenko described a new species Probaticus medvedevi, from northern Armenia (Figs 6 D, F). Allard examined the specimens of Leder and confirmed that they belong to his species Helops vicinus, as he wrote in redescription (Allard 1877). The Leder’s specimens (Figs 6 A – C, E, G – K) are not syntypes (as marked on the labels) because the original description was based on other specimens from the Reiche collection. The females from Georgia are conspecific to the female (holotype) of E. medvedevi despite the variability. As a result, I propose the following synonymy: Helops vicinus Allard, 1876 = Probaticus (Pelorinus) medvedevi Abdurakhmanov et Nabozhenko, 2011, syn. nov.	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF9B5F33D3B954CF28780988.taxon	distribution	Distribution. Southern Georgia, northern area of Armenia.	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF955F3ED3B956D628CC0CC8.taxon	description	(Fig. 7)	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF955F3ED3B956D628CC0CC8.taxon	materials_examined	Material. Turkmenistan. 1 ♀ (HNHM): “ Trans Caspi Eylandt ” (Turkmenistan, Transcaspian area); 2 ♀ (ZIN): Zakaspiyskaya oblast’ (Turkmenistan, Transcaspian area )), Aydere canyon, 10. v. 1890 (leg. Eylandt); 1 ♂, 1 ♀ (ZIN): Turkmenistan, Kopetdag, Aydere, 3. v. 1930 (leg. E. Shestopalov); 1 ♂, 2 ♀ (ZIN): Turkmenistan, Aydere canyon, 0 – 3 km from river mouth, 30. iv. 1952 (leg. K. G. Romadina); 2 ♀ (ZIN): Turkmenistan, Kara-Kala (= Garrygala), N slope of Syunt Mt., 2. vii. 1956 (leg. G. S. Medvedev); 1 ♂ (ZIN), 1 ♀ (HNHM): Turkmenistan, Kara-Kala, a canyon on N slope of Syunt Mt., 6. v. 1957 (leg. G. S. Medvedev); 4 ♀ (ZIN): Turkmenistan, Kara-Kala, Savudzhe canyon, 8. v. 1957, 11. v. 1957 (leg. G. S. Medvedev); 1 ♀ (ZIN) Turkmenistan, N of Kara-Kala, 10. v. 1957 (leg. G. S. Medvedev); 1 ♂, 1 ♀ (ZIN): Turkmenistan, W Kopetdag, headwaters of Khozly canyon, 20.04.1975, 23.04.1975 (leg. V. Yanushev); 1 ♀ (HNHM): Turkmenistan, Kopet-Dag Mts, 10 km S Aydere, 38 ° 14 ʹN, 56 ° 46 ʹE, 600 – 1000 m, 27. vi. 1992 (leg. Gy Fábián, B. Herczig, A. Podlussány, Z. Varga). Iran. 1 ♂, 1 ♀ (NMPC): NE Iran, Khorasan, Golestan forest 55 km NE Minudast, 37 ° 20 ʹN / 56 ° 00 ʹE, 840 m, 17. vi. 2000 (S. Kadlec); 2 ♂, 1 ♀ (HNHM): Iran, Golestan, Golestan forest, 45 km NE of Minudasht, 584 m, 16. vi. 2015 (leg. A. Lasoń); 37 ° 23 ‘ N, 55 ° 50 ‘ E, 9 ♂, 4 ♀ (PCMN): Iran, North Khorassan Prov., Golestan National park, 37 ° 22 ‘ 46.33 “ N, 55 ° 51 ‘ 5.56 “ E, ~ 600 m, 24 – 25. v. 2016 (leg. D. G. Kasatkin); 1 ♂, 2 ♀ (ZIN): Iran, North Khorassan Prov., Golestan National park, 37 ° 22 ʹ 6 ʺN, 55 ° 56 ʹ 40.6 ʺE, 26 – 27. v. 2017 (leg. D. G. Kasatkin). Turkmenistan / Iran. 1 ♀ (HNHM): “ Arwas Pers. b. ” (Kopetdag Mts., Arvaz pass between Iran (North Khorasan Prov.) and Turkmenistan).	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF955F3ED3B956D628CC0CC8.taxon	distribution	Distribution. South West Turkmenistan (Kopetdag), Iran (North Khorasan Province). Note. This is the only species of the genus that occurs in Turkmenistan.	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF965F3ED3B955932A950B34.taxon	description	This group unites alpine and mountain-steppe terrestrial Iranian species with more or less shiny black body in males, fine and sparse puncturation of pronotum (at least in the middle) and strong long setae on the apical part of the abdominal ventrite 5.	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF965F38D3B9528F292B0BE8.taxon	description	(Fig. 8)	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF965F38D3B9528F292B0BE8.taxon	materials_examined	Type material. Holotype, ♂ and paratype, ♂ (HNHM): Iran, Gachsar [Alborz Prov.] — Marzanabad [Mazandaran Prov.], 1700 – 2500 m, 28. iv. 1970 (leg. Wittmer, v. Bothmer).	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF965F38D3B9528F292B0BE8.taxon	description	Description. Male. Body slender, elongate, black, with dull shine (Fig. 8 A). Measurements: Y = 1.47; PH w = 1.44; P w P l = 1.09; E l E w = 1.74; EH w = 1.94; EP w = 1.34; EP l = 2.6 – 2.61. Body length 12.5 – 13 mm, width 4.5 – 4.8 mm. Head (Figs 8 B, E, F). Anterior margin of epistoma straight (Fig. 8 E). Head widest at eye level. Eyes moderately convex, circumocular impression narrow. Lateral margins of genae angularly rounded. Puncturation of head moderately fine and sparse on frons (interpuncture distance near 3 times as long as puncture diameter) and denser on epistoma and genae (interpuncture distance subequal to or near 1.5 times as long as puncture diameter). Ventral side of epicranium coarsely and densely punctured; temples depressed behind eyes in the lower half (Fig. 8 B). Antennae comparatively long, with four apical antennomeres, extending beyond base of pronotum, reaching 1 / 3 of elytral length. Prothorax (Figs 8 B, E – G). Pronotum weakly transverse, with almost equal length and width, widest at middle (Fig. 8 E). Lateral margins of pronotum weakly rounded in middle part, straightly narrowed in apical third and widely emarginated in basal quarter; anterior margin trisinuate; base slightly rounded, almost straight, with emargination at middle. Antero-lateral corners not projected, right, pointed at apex; postero-lateral corners right, narrowly rounded at apex. Lateral margins not beaded; anterior margin only with very short bead marginally; disc completely beaded. Disc of pronotum convex, not flattened along lateral margin, without impressions. Puncturation of disc fine and sparse (puncture diameter 2 – 4 times as short as interpuncture distance), slightly coarser and denser on lateral sides; punctures round (Figs 8 E, F). Prosternum with fine and sparse simple puncturation (Fig. 8 B). Outer margin of prothoracic hypomera narrowly flattened only near base. Prohypomera with small confused microwrinkles with poorly distinguished sparse smoothed puncturation against their background (Fig. 8 G). Prosternal process smooth and shiny (only rugose near apex), almost without punctures, slightly convex at apex, not beaded (Figs 8 B, G). Pterothorax (Figs 8 A, B). Scutellar shield triangle, moderately coarsely punctured. Elytra strongly elongate, widest slightly behind middle; punctures in striae fine, round, widely spaced; punctures in interstriae fine and sparse, much smaller than strial ones (Fig. 8 A). Anterior half of mesoventrite strongly wrinkled, pubescent, with more smooth middle (Fig. 8 B). Mesepisterna coarsely and densely punctured by large foveae; puncturation of mesepimera and metepisterna sparser and finer; metaventrite coarsely and moderately sparsely punctured and pubescent with short setae. Legs (Figs 8 B, G). Trochanters with short dense reddish setae (but not dense brush), protrochanters with two long setae (Fig. 8 G), meso- and metatrochanters with one long seta. Basal third of femora with the same dense on flex side (Fig. 8 B); another surface of femora with sparse puncturation and short setae. Pro- and metatibiae slightly curved, mesotibiae stright. Pro- and mesotarsomeres 1 – 4 widened, trasverse, pro- and mesotarsomere 1 as wide as apex of protibia. Abdomen (Figs 8 B – D, H – J). Abdominal ventrites with moderately dense, fine puncturation and simple, moderate, short setae (shortly pubescent); ventrite 5 with coarser punctures and longer setae at apex. Genitalia (Figs 8 C, D, H – J). Apical part of basal piece of aedeagus smooth, without rugosity. Apical piece of aedeagus with wide median longitudinal impression; lateral margins straight at middle and straightly narrowed to narrowly rounded apex. Female. Unknown.	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF965F38D3B9528F292B0BE8.taxon	etymology	Etymology. The species is named in honour of my first teacher during student times and known specialist on Curculionidae DrSc. Yury Arzanov (Rostov-on-Don).	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF965F38D3B9528F292B0BE8.taxon	diagnosis	Differential diagnosis. The new species is close to E. merkli by the structure of the aedeagus (basal piece without rugosity, apical piece with median longitudinal impression) and the narrow, not beaded pronotum with weakly rounded margins, but differs in following characters: the lower half of temples is impressed behind the eye (weakly rounded in E. merkli); lateral margins of pronotum straight at apical quarter and widely emarginated in basal quarter (regularly rounded in E. merkli); prothoracic hypomera with microrugosity and fine sparse smooth punctures (large, coarse, round foveae and smooth interpuncture space between them in E. merkli); protrochanters with two long setae (one long seta in E. merkli).	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF905F3AD3B952AC2A780A0A.taxon	description	(Fig. 9)	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF905F3AD3B952AC2A780A0A.taxon	materials_examined	Type material. Holotype, ♂ and paratype, ♀ (ZIN): Iran, Zanjan Prov., 40 km NE Bonab, 36 ° 47 ʹ 57 ʺN, 48 ° 43 ʹ 53.7 ʺE, 2400 m, 9. v. 2017 (leg. D. G. Kasatkin). Paratypes. 1 ♀ (ZIN): Iran, Zanjan Prov., 36 ° 45 ʹ 37 ʺN, 48 ° 48 ʹ 13 ʺE, 9. v. 2017 (leg. A. Zubov); 1 ♀ (SMS): Iran, Zanjan Prov., Alborz Mts., Sendan Mt., Zanjan, Abbar, 36 ° 48.462 ʹN, 48 ° 53.127 ʹE, 1480 m, 11 – 12. v. 2008, sigling (leg. T. Hácz, K. Székely, K. Vig); 1 ♀ (HNHM): Iran Zangan [Zanjan] Prov., Zangan [Zanjan], 1800 – 2000 m, 7. vi. 1999 (leg. K. Gaskó); 1 ♀ (HNHM): Iran, Zanjan Prov., Sendan Mts., 20 km SW of Gilvan, 36 ° 48 ʹN, 48 ° 55 ʹE, 1000 m, 18. v. 2001 (leg. B. Benedek, G. Csorba).	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF905F3AD3B952AC2A780A0A.taxon	description	Description. Male. Body slender, elongate, black, with dull shine. Measurements: Y = 1.47; PH w = 1.6; P w P l = 1.23; E l E w = 1.7; EH w = 1.95; EP w = 1.2; EP l = 2.58. Body length 16 mm, width 5.5 mm. Head (Figs 9 B, D, E). Anterior margin of epistoma slightly rounded. Head widest at eye level. Eyes moderately convex, circumocular impression narrow. Lateral margins of genae angularly rounded. Puncturation of head fine and sparse, interpuncture distance 1.5 – 2 times as long as puncture diameter. Ventral side of epicranium coarsely and densely punctured. Antennae comparatively short, with only three apical antennomeres, extending beyond base of pronotum, reaching 1 / 5 of elytral length. Prothorax (Figs 9 B, D – F). Pronotum cordate, widest ahead of middle, transverse (Fig. 9 D). Lateral margins of pronotum moderately rounded at apical half and slightly widely emarginated at basal half; anterior margin straight at middle and widely emarginated near corners; base slightly rounded. Antero-lateral corners projected, obtuse, widely rounded at apex; postero-lateral corners weakly obtuse, pointed at apex. Lateral margins beaded, interrupted only near antero-lateral corners; bead of anterior margin interrupted in middle. Disc of pronotum evenly weakly convex, not flattened along lateral margin, with oval impression on each lateral side and two small oblique impressions near base. Puncturation of disc fine and sparse, puncture diameter 2 – 4 times as short as interpuncture distance; punctures round (Figs 9 D, E). Prosternum with moderately coarse and sparse raduliform puncturation (Fig. 9 B). Prothoracic hypomera not flattened along margin, with small confused microwrinkles with poorly distinguished sparse smoothed puncturation against their background; wrinkles with microgranulation at anterior third (Fig. 9 F). Prosternal process smooth and shiny, almost without punctures, slightly convex at apex, beaded only in basal half (Figs 9 B, F). Pterothorax (Figs 9 A, B). Scutellar shield with slightly rounded margins, finely and sparsely punctured. Elytra strongly elongate, widest at middle; punctures in striae fine, slightly elongate, widely spaced; punctures in interstriae fine and sparse, much smaller than strial ones (Fig. 9 A). Anterior half of mesoventrite pubescent, with triangle smooth middle and coarsely wrinkled another surface (Fig. 9 B). Mesepisterna coarsely and densely punctured by large foveae; puncturation of mesepimera and metepisterna sparser and finer; metaventrite finely and sparsely punctured and covered with short setae. Legs. Trochanters with brush of short dark-reddish setae and one long seta (Fig. 9 B). Basal third of femora with the same brush on flex side (Fig. 9 B); another surface of femora with coarse puncturation and short setae. Pro- and mesotibiae straight, metatibiae weakly bent. Pro- and mesotarsomeres 1 – 4 strongly widened, trasverse, pro- and mesotarsomere 1 as wide as apex of protibia. Abdomen (Figs 9 B, G – K). Abdominal ventrites with dense and fine puncturation and simple short setae (not pubescent); ventrite 5 with coarser punctures and long strong setae. Genitalia (Figs 9 G – K). Apical part of basal piece of aedeagus smooth, without rugosity. Apical piece of aedeagus with median shiny longitudinal keel; lateral margins slightly emarginated at middle and sharply narrowed at apical quarter. Female (Fig. 9 C). Body much robuster, elytra matt. Eyes little smaller (see measurements). Antennae short, not reaching base of pronotum. Metatibiae straight. Measurements: Y = 1.4; PH w = 1.7; P w P l = 1.3; E l E w = 1.47; EH w = 2.28; EP w = 1.34; EP l = 2.57. Body length 13.5 – 17 mm, width 5,5 – 6.5 mm.	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF905F3AD3B952AC2A780A0A.taxon	etymology	Etymology. This species is named in memory of Roland Grimm (1948 – 2021), my colleague and co-author, a famous specialist on Tenebrionidae of the Old World (Burmeister 2021, Schawaller & Staniczek 2021).	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF905F3AD3B952AC2A780A0A.taxon	diagnosis	Differential diagnosis. This new species is different from all Iranian congeners by the presence of shiny keellike longitudinal median elevation on the apical piece of the aedeagus (Fig. 9 K); all other species have the apical piece smooth or with the median impression. Euboeus grimmi sp. nov. has the pronotum with beaded lateral margins (Figs 9 D, E) and bent metatibiae (Figs 9 A, B) as E. hudepohli and E. kasatkini sp. nov. and differs from both species by the absence of longitudinal rugosity in apical part of the basal piece of the aedeagus (Fig. 9 K) and the pronotum widest at the apical third (at middle in E. huedepohli and E. kasatkini). In addition, the new species is distinguished from: – E. huedepohli by prothoracic hypomera not flattened along margin, covered with small confused microwrinkles with hardly distinguished sparse smoothed puncturation against their background (widely flattened and coarsely longitudinally wrinkled in E. huedepohli) (Fig. 9 F); male dorsally is less shiny (lacquered shiny in E. huedepohli); – E. kasatkini by prothoracic hypomera not flattened along the margin, covered with small confused microwrinkles with hardly distinguished sparse smoothed puncturation against their background (narrowly fattened and coarsely and densely punctured in E. kasatkini) (Fig. 9 F); more bent male metatibiae and less shiny male body.	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF925F3BD3B953D428100EE4.taxon	description	(Figs 10, 11)	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF925F3BD3B953D428100EE4.taxon	materials_examined	Type material (studied). Holotype, ♂ (NMB): “ Persia Elburs-Gebg. Hüdepohl leg. ”, “ Lashgarak-Tal 12 - 4 - 63 ”, “ Holotypus Probaticus huedepohli n. sp. det H. Kulzer 1963 ” (Fig. 8 C). Paratype, ♂ (HNHM) with the same geographic label and “ Paratypus 1964 Probaticus huedepohli Külzer ”, “ PARATYPUS Probaticus huedepohli n. sp. det. H. Kulzer 1963 ”, “ Probaticus huedepohli Kulz. Dr. Kaszab det. 1983 ”. Material. 1 ♂ (ZIN): Iran, Demavend, 24. v. 1894 (leg. Glazunov); 1 ♂ (ZIN): Iran, 16 km NE Tehran, foothills of Elburs, 12. v. 1955 (leg. D. Shteinberg); 2 ♂ (HNHM): Iran, Mazandaran Prov., Demavend, 3000 m, 3. v. 2000 (leg. K. Gaskó); 3 ♂, 3 ♀ (HNHM): Iran, Tehran / Mazandaran provinces, Polour — Abali, 2100 – 2800 m, 17. v. 1970 (leg. Wittmer. V. Bothmer); 2 ♂, 5 ♀ (HNHM): Iran, Tehran Prov. [Mazandaran Prov.], Elburz Mts., 10 km W Reine [Rineh], Kuh-e Demavend, 35 ° 53 ʹ 22 ʺN, 52 ° 02 ʹ 24 ʺE, 14. v. 1998 (Gy. Fábian, K. Székely); 1 ♀, Iran, Mazandaran Prov., Kühha-ye Alborz, Küh-e Damavand, 3500 m, 30. vi. – 2. vii. 2006 (leg. G. Rozner); 1 ♀ (HNHM): “ Iran, prov. Khorasan, Qudjan (Qućan), 3000 m / Mt. Kopet Dag 1999. V. 90 leg.: Gaskó K. ”; 3 ♂, 3 ♀ (SMS): Iran, Mazandaran Prov., Alborz Mts., Damavand Mt., Reyneh [Rineh], 35 ° 52.412 ʹ N, 52 ° 06.096 ʹ, 2841 m, 14. v. 2008, from beneath stone (leg. T. Hácz, K. Székely, K. Vig);	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF925F3BD3B953D428100EE4.taxon	distribution	Distribution. Iran (Demavend Mt. from ~ 2800 to 3500 m). I didn’t include Razavi Khorasan Province in the distribution based on the strange locality “ Quchan, Kopet Dag, 3000 m ”, because the Iranian part of Kopet Dag mountains does not exceed 2000 m, and only one mount Kuh-e Patu (not Kopet Dag ridge) with the elevation near 3000 m is located 50 km west of Quchan. This mountain belongs to another mountain system (Turkmen-Khorasan mountains) than Demavend and and labelling raises doubts.	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF8D5F27D3B956D62E030B58.taxon	description	(Fig. 12)	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF8D5F27D3B956D62E030B58.taxon	materials_examined	Type material. Armenia. Holotype, ♂ (ZIN) with the Cyrillic label: “ ЛичкваЗ — Нор-Аревик Мегринский р-н, Арм ССР 31.5.1980 А. Л. Лобанов ” (Armenia, Lichk, Nor-Arevik, 31. v. 1980, leg. A. V. Lobanov). Paratypes: 1 ♂ (IZAY): Armenia, Syunik Prov., Tsav, 21.07.1952 (leg. S. Iablokoff-Khnzorian); 1 ♀ (IZAY): Armenia, Syunik Prov., Srashen, 23. vi. 1981 (leg. M. Kalashian); 1 ♀ (IZAY): Armenia, Syunik Prov., 4 – 6 km N Shvanidzor, 20. iv. 2001 (leg. K. Aghababyan); 1 ♀ (IZAY): Armenia, Syunik Prov., Pukhrut, 39.1307 ° N, 46.2205 ° E, 2130 m, 25. v. 2005 (leg. G. Karagyan); 3 ♂, 1 ♀ (IZAY): Armenia, Syunik Prov., Meghri, 30 – 31. v. 1923 (Expedition of Museum of Armenia); 1 ♀ (IZAY): Armenia, Syunik Prov., Meghri, 24. v. 1945 (leg. H. Avetian); 1 ♂ (IZAY): Armenia, Syunik Prov., Chakaten, 25. v. 1942 (leg. H. Avetian); 1 ♂ (IZAY): Armenia, Syunik Prov., S environs of Lichk, abandoned Nor Arevik vill., 26. iv. 1943 (leg. H. Avetian); 2 ♂, 1 ♀ (IZAY): same locality, 5. v. 1954 (leg. S. Iablokoff-Khnzorian); 1 ♀ (IZAY): Armenia, Syunik Prov., Vahravar, 27. v. 1945 (leg. H. Avetian); 1 ♀ (IZAY): Armenia, Syunik Prov., S environs of Lichk, 39.0336 ° N, 46.1932 ° E, 1700 m, 8. v. 2016 (leg. M. Kalashian); Armenia, Syunik Prov., N Shvanidzor vill., 38.9851 ° N, 46.3746 ° E, 1320 m 1 ♂ (IZAY): 9. vi – 8. viii. 2017 and 1 ♂, 2 ♀ (IZAY): 16. vi – 26. viii. 2018, soil traps (leg. T. Ghrejyan, G. Karagyan); 1 ♂, 1 ♀ (IZAY): Armenia, Syunik Prov., E environs of Kapan, Geghanush, 39.2221 ° N, 46.4240 ° E, 1030 m, 8. iv – 14. v. 2008, soil traps (leg. M. Kalashian); 1 ♂ (IZAY): Armenia, Syunik Prov., N environs of Kapan, Ghatar locality, 39.2206 ° N, 46.3880 ° E, 1120 m, 8. iv – 14. v. 2008, soil traps (leg. M. Kalashian); 1 ♀ (ZIN): Syunik Prov., fork to Lichk, 7. vi. 1993 (leg. M. Kalashian); 1 ♀ (ZIN): Armenia, Syunik Prov., Lichk, 4. vi. 1995 (leg. Kalashian); 1 ♀ (ZIN): Armenia, Syunik Prov., Gudemnis, 26. iv. 1998 (leg. M. Kalashian); 1 ♀ (ZIN): Armenia, Syunik Prov., Agarak, 14. iv. – 20. v. 1995 (leg. M. Kalashian); Armenia, Vayotsdzor Prov., Areni, 24. v. 1988 (leg. M. Kalashian). Iran. 1 ♂ (HNHM): Iran, Azerbaijan, Makidi Arasbaran Wildlife Refuge, 1650 – 1800 m, 4 – 5. vii. 1978 (leg. Martens, Pieper).	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF8D5F27D3B956D62E030B58.taxon	description	Description. Male. Body robust, black, shiny (Fig. 12 A). Measurements: Y = 1.56; PH w = 1.57; P w P l = 1.18; E l E w = 1.6 – 1.63; EH w = 2.1; EP w = 1.3; EP l = 2.25. Body length 11.5 – 16.6 mm, width 4.4 – 6.4 mm. Head (Figs 12 C – E). Anterior margin of epistoma slightly rounded. Head widest at eye level. Eyes moderately convex, circumocular impression narrow. Lateral margins of genae evenly rounded. Puncturation of head moderately coarse and sparse, interpuncture distance 1.5 as long as puncture diameter, genae densely and coarsely punctured. Ventral side of epicranium coarsely and densely punctured, gula with coarse transverse wrinkles. Antennae comparatively short, with only three apical antennomeres, extending beyond base of pronotum, reaching 1 / 5 of elytral length. Prothorax (Figs 12 C – F). Pronotum subcordate, widest at middle, transverse (Fig. 12 D). Lateral margins of pronotum strongly rounded at middle and widely emarginated at basal half; anterior margin widely evenly emarginated; base straight. Antero-lateral corners not projected, obtuse, pointed; postero-lateral corners weakly straight, narrowly rounded at apex. Lateral margins not beaded; anterior margin shortly beaded only near corners; base beaded at middle. Disc of pronotum evenly weakly convex, not flattened along lateral margin, with two small oblique impressions near base. Puncturation of disc fine and sparse at middle and anterior half (puncture diameter 2 – 3 times as short as interpuncture distance), denser on sides and at basal quarter; punctures round (Figs 12 D, E). Prosternum with moderately coarsely and sparse punctured and wrinkled (Fig. 12 C). Prothoracic hypomera narrowly flattened along margin, with coarse transverse wrinkles only (Fig. 12 F). Prosternal process smooth and shiny, sparsely punctured and pubescent, convex at apex, not beaded (Figs 12 C, F). Pterothorax (Figs 12 A, C). Scutellar shield with triangle, with straight margins, finely and sparsely punctured. Elytra oval, widest at middle; punctures in striae sligly elongate, closely spaced; punctures in interstriae moderately coarse and sparse, little smaller than strial ones (Fig. 12 A). Anterior half of mesoventrite pubescent, coarsely wrinkled (Fig. 12 C). Mesepisterna, mesepimera and metepisterna coarsely and densely punctured by large foveae; metaventrite finely and sparsely punctured and pubescent with recumbent reddish setae. Legs. Trochanters with dense brush of reddish setae and one long seta. Basal half of femora with the same dense brush on flex side (Fig. 12 C); another surface of femora with moderately coarse, sparse puncturation and short setae. All tibiae straight. Pro- and mesotarsomeres 1 – 4 strongly widened, transverse, pro- and mesotarsomere 1 slightly wider than apex of protibia. Abdomen (Figs 12 C, G – K). Abdominal ventrites with moderately dense and fine puncturation and simple short pubescence; ventrite 5 with coarser punctures and long strong setae. Genitalia (Figs 12 G – K). Apical part of basal piece of aedeagus smooth, without rugosity. Apical piece of aedeagus with weak median longitudinal keel; lateral margins almost straight, weakly narrowed to widely rounded apex. Apical third of basal piece smooth, without longitudinal rugosity. Female (Figs 12 B, C). Body robuster, dull, elytra matt. Eyes slightly smaller (see measurements). Antennae short, not reaching base of pronotum. Metatibiae straight. Measurements: Y = 1.5; PH w = 1.7; P w P l = 1.3; E l E w = 1.47; EH w = 2.28; EP w = 1.34; EP l = 2.57. Body length 13.5 – 17 mm, width 5.5 – 6.6 mm.	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF8D5F27D3B956D62E030B58.taxon	etymology	Etymology. The species is named in honour of my friend Dr Mark Kalashian (Yerevan, Armenia), a great connoisseur of the insect fauna of the Caucasus and a well-known specialist in Buprestidae.	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF8D5F27D3B956D62E030B58.taxon	diagnosis	Differential diagnosis. The new species is similar to E. quadricollis by the robust body, the form of pronotum with rounded lateral margins in apical 2 / 3 and widely emarginated in basal third, but differs from it by the following characters: lateral margins of pronotum more rounded and wholly not beaded, the lateral portions of base also not beaded (lateral margins lesser rounded and beaded at least near angles and base completely beaded in E. quadricollis); prothoracic hypomera flattened along outer margin and are with coarse longitudinal wrinkles (the hypomera with irregular fine wrinkles and punctures between them, not flattened along margin in E. quadricollis); the apical piece of the aedeagus is widely rounded at apex, and its dorsal surface is with longitudinal keel (the apical piece is quadrate or emarginated at middle in E. quadricollis and has the smooth surface). The coarsely longitudinally wrinkled prothoracic hypomera are present only in two species, i. e. E. huedepohli and E. kalashiani. Notes. This species is interpreted by Russian entomologists as E. vicinus since the paper of Bogatchev (1938), who recorded it for Shikh-Yurdy Mt. (Ordubad District, Nakhichevan Autonomous Republic, Azerbaijan). Unfortunately, the Bogatchev’s specimen from this locality was not found in Russian collections and in the collection of the Institute of Zoology of the Azerbaijan Academy of Sciences. This mount is located just a few kilometers from Armenian localities of E. kalashiani. Iablokoff-Khnzorian (1957) listed both E. quardicollis and E. kalashiani as one species Hedyphanes quardicollis with the wide range from Talysh to Aras valley, but indicated characters for E. kalashiani in the key. Abdurakhmanov & Medvedev (1994) and Abdurakhmanov & Nabozhenko (2011) listed this new species as Probaticus vicinus, and Nabozhenko et al. (2021) indicated it as Euboeus (Pelorinus) sp.	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF8F5F21D3B9522329610B34.taxon	description	(Fig. 13)	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF8F5F21D3B9522329610B34.taxon	materials_examined	Type material. Holotype, ♂ and two paratypes, ♂ and ♀ (ZIN): Iran, Alborz Prov., Central Elburs Mts., Gachsar env., H = 2600 – 2720 m, 36 ° 05 ʹ 51.88 ʺN, 51 ° 19 ʹ 46.53 ʺE, 30. v. 2016 (leg. D. G. Kasatkin). Paratypes. 1 ♂, 1 ♀ (HNHM): N Iran, 8 km W Gachsar, 6 – 8. vii. 1977, “ Loc. No. 396 Exped. Nat. Mus. Praha ”, “ Probaticus huedepohli Kulzer, 1964 det. Z. Kaszab, 1983 ”; 1 ♀ (HNHM): Iran, Tehran Prov, Elburs-Gebirge, N Tehran, Shenshak, 2650 – 2900 m, 17 – 25. vi. 1978 (leg. Martens and Pleper); 2 ♀ (PCMN): Iran, Tehran Prov., Alborz Mts., Shemshak vill., road to Dizin, 3000 m, 1. vi. 2014 (D. Murastyi);	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF8F5F21D3B9522329610B34.taxon	description	Description. Male. Body slender, black, pronotum and head dorsally shiny, elytra lesser shiny (Fig. 13 A). Measurements: Y = 1.44; PH w = 1.5; P w P l = 1.2; E l E w = 1.54; EH w = 1.86; EP w = 1.24; EP l = 2.38. Body length 12 mm, width 4.4 mm. Head (Figs 13 C, E, F). Anterior margin of epistoma slightly rounded and emarginated near angles. Head widest at eye level. Eyes moderately convex, circumocular impression narrow. Lateral margins of genae angularly rounded. Puncturation of head fine and sparse, interpuncture distance 1.5 – 2 times as long as puncture diameter. Ventral side of epicranium coarsely and densely punctured. Antennae comparatively short, with only three apical antennomeres, extending beyond base of pronotum, reaching less than 1 / 4 of elytral length. Prothorax (Figs 13 C, E – G). Pronotum widest at middle, transverse (Fig. 13 E). Lateral margins of pronotum weakly evenly rounded; anterior margin weakly rounded at middle and slightly widely emarginated near corners; base weakly rounded at middle, slightly bisinuate. Antero-lateral corners not projected, obtuse, narrowly rounded at apex; postero-lateral corners weakly obtuse, pointed at apex. Lateral margins beaded; anterior margin beaded only laterally (Figs 13 E, F). Disc of pronotum evenly weakly convex, not flattened along lateral margin, with weak oval impression on each lateral side and two small impressions at base. Puncturation of disc fine and sparse, puncture diameter 3 – 5 times as short as interpuncture distance; punctures round; punctures in impressions slightly larger (Figs 13 E, F). Prosternum with moderate and very sparse simple puncturation (Fig. 13 C). Prothoracic hypomera very narrowly weakly flattened along margin, with coarse and dense punctures among wrinkles; surface without microgranulation (Fig. 13 G). Prosternal process smooth and shiny, with two terminal lines of punctures, slightly convex at apex, not beaded (Figs 13 C, G). Pterothorax (Figs 13 A, C). Scutellar shield with straight margins, coarsely sparsely punctured. Elytra moderately elongate, widest at middle; punctures in striae fine, slightly elongate, widely spaced; punctures in interstriae fine and sparse, much smaller than strial ones (Fig. 13 A). Anterior half of mesoventrite pubescent, coarsely and densely wrinkled. Mesepisterna, mesepimera and metepisterna with coarse and dense puncturation of large foveae; metaventrite and intercoxal part of mesoventrite finely and sparsely punctured and covered with short setae (Fig. 13 C). Legs. Trochanters with brush of short dark-reddish setae and one long seta (Fig. 13 C). Basal third of femora with the same brush on flex side (Fig. 13 C); another surface of femora with coarse puncturation and short setae. Pro- and mesotibiae straight, metatibiae very weakly bent (Fig. 13 B). Pro- and mesotarsomeres 1 – 4 strongly widened, trasverse, pro- and mesotarsomere 1 wider than apex of protibia. Abdomen (Figs 13 C, H – M). Abdominal ventrites with fine and sparse puncturation and short setation, ventrite 5 with pubescence of long strong setae. Genitalia (Figs 13 H – M). Apical part of the basal piece of aedeagus with longitudinal rugosity (Fig. 13 M). Lateral sides of apical piece of aedeagus straight in apical two thirds (Fig. 13 M). Female (Fig. 13 D). Body much more robust, elytra duller in specimen from the type locality and the same shiny as male in specimens from Shemshak). Antennae short, not reaching base of pronotum. Pronotum widest at middle or sometimes behind middle. Metatibiae straight. Measurements: Y = 1.44; PH w = 1.65; P w P l = 1.24 – 26; E l E w = 1.46; EH w = 2.13; EP w = 1.29; EP l = 2.41. Body length 11 – 16 mm, width 4 – 6.5 mm.	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF8F5F21D3B9522329610B34.taxon	etymology	Etymology. The new species is named in honour of my friend, the collector of many specimens of Euboeus and other Tenebrionidae from Iran, Dr Denis Kasatkin (Rostov Branch of the Federal State Budgetary Institution All-Russian Plant Quarantine Centre, Rostov-on-Don, Russia).	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF8F5F21D3B9522329610B34.taxon	diagnosis	Differential diagnosis. This species is very close to E. huedepohli from Demavend by the beaded lateral margins of pronotum (Figs 11 E, F, 13 E, F) and longitudinal rugosity in apical part of the basal piece of the aedeagus (Figs 11 M, 13 M), but differs by the following characters: body dorsally lesser shiny, pronotum without lacquer gloss; prothoracic hypomera with coarse puncturation among smooth wrinkles (Fig. 13 G) (coarse longitudinal wrinkles in E. huedepohli (Fig. 11 G )); lateral margin of prothoracic hypomera very narrowly weakly flattened (Fig. 13 G) (widely clearly flattened in E. huedepohli (Fig. 11 G )); metatibiae very weakly bent (Fig. 13 B) (strongly bent in E. huedepohli (Fig. 11 C )); lateral margins of the apical piece of the aedeagus straight in apical half (Fig. 13 M) (weakly rounded in E. huedepohli (Fig. 11 M )).	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF895F23D3B9528F2A0F091A.taxon	description	(Fig 14)	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF895F23D3B9528F2A0F091A.taxon	materials_examined	Type material. Holotype, ♂ (HNHM): Iran, Gilan Prov.: “ Kelishum [Klishom] 2000 m südl. Langerud [Langarud] 4. viii. 1967 ”, “ Gilan (Iran) Heinz leg. ”, “ Probaticus sp. det. Kaszab ”. Paratype, ♀ (SMNS): Iran, Gilan Prov., 20 km SE Deilaman, 1900 m, 22.04.2003 (leg. G. Sama); 1 ♂ (NMPC): Iran, Koedestan [Kurdestan] Prov., Pass 26 km SE Saqqez, 17. iv. 2002 (leg. S. Kadlec); 1 ♀ (NME): Iran, Luristan [Lorestan] Prov., pass 2300 m W Nour-Abad (SE Nahavand), oberh. Grotte v. Gamasiab [upper grotto of Gamasiab], 10. iv. 2014 (leg. Heinz).	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF895F23D3B9528F2A0F091A.taxon	description	Description. Male. Body slender, black, shiny (Fig. 14 A), but weakly shiny in the Lorestan Population. Measurements: Y = 1.59; PH w = 1.55; P w P l = 1.3; E l E w = 1.6; EH w = 1.83; EP w = 1.17; EP l = 2.48. Body length 12.5 mm, width 4.5 mm. Head (Fig. 14 B, D, E). Anterior margin of epistoma slightly rounded and emarginated near angles. Head widest at eye level. Eyes strongly convex, angulate in dorsal view, circumocular impression deep, moderately wide (Figs 14 B, D, E). Lateral margins of genae angularly rounded. Puncturation of head fine and sparse, interpuncture distance 1.5 – 2 times as long as puncture diameter; punctures coarser on genae. Ventral side of epicranium coarsely and densely punctured around submentum and coarse and sparse on temples. Antennae comparatively short, with only three apical antennomeres, extending beyond base of pronotum, reaching less than 1 / 4 of elytral length. Prothorax (Figs 14 D – F). Pronotum widest ahead of middle, transverse (Fig. 14 D). Lateral margins of pronotum moderately rounded, slightly emarginated near base; anterior margin weakly widely emarginated; base straight. Antero-lateral corners not projected, obtuse, narrowly rounded at apex; postero-lateral corners weakly obtuse, narrowly rounded at apex. Lateral margins not beaded; anterior margin beaded only laterally (Figs 14 D, E). Disc of pronotum evenly weakly convex, not flattened along lateral margin, without impressions. Puncturation of disc fine and sparse, punctures small, but surface around punctures impressed, puncture diameter 3 times as short as interpuncture distance; punctures round. Prosternum with coarse and sparse simple puncturation (Figs 14 B, F). Prothoracic hypomera not flattened flattened along margin, with coarse, sparse, large, round punctures on smooth surface at outer half and dense and coarse merged punctures on inner half; surface without microgranulation (Fig. 14 F). Prosternal process shiny, sparsely punctured, slightly convex at apex, not beaded (Figs 14 B, F). Pterothorax (Figs 14 A, B). Scutellar shield with very weakly rounded margins, coarsely sparsely punctured. Elytra shiny, moderately elongate, widest at middle; punctures in striae fine, round, widely spaced; punctures in interstriae fine and sparse, much smaller than strial ones (Fig. 14 A). Anterior half of mesoventrite pubescent, coarsely and densely wrinkled. Mesepisterna, mesepimera and metepisterna with coarse and dense puncturation of large foveae; metaventrite and intercoxal part of mesoventrite with moderately coarse and moderately dense punctures; metaventrite covered with very short setae (Fig. 14 B). Legs. Trochanters with brush of short dark-reddish setae and one long seta (Fig. 14 B). Basal third of meso- and metafemora with the same brush on flex side, the same part of profemora without this brush (Fig. 14 B). Protibiae weakly S-shaped, mesotibiae weakly curved, metatibiae weakly bent inward. Pro- and mesotarsomeres 1 – 4 strongly widened, trasverse, but pro- and mesotarsomere 1 not wider than apex of protibia. Abdomen (Figs 14 B, G – K). Abdominal ventrites with the same puncturation as on metaventrite and very short setation (not pubescent) (Fig. 14 B); Ventrite 5 with several sparse long setae at apex. Genitalia (Figs 14 G – K). Apical part of the basal piece of aedeagus with longitudinal rugosity (Fig. 14 K). Apical piece subtriangle, narrowly rounded at apex, dorsal surface with longitudinal small median groove (Fig. 14 K). Female (Fig. 14 C). Body much robuster and dull. Eyes smaller (Y = 1.36 vs. 1.59 in male). Antennae short, not reaching base of pronotum. Lateral margins of pronotum beaded at basal half. Postero-lateral corners of pronotum right. Metatibiae straight. Measurements: Y = 1.36; PH w = 1.67; P w P l = 1.29; E l E w = 1.37; EH w = 2.2; EP w = 1.32; EP l = 2.34. Body length 17 mm, width 7 mm.	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF895F23D3B9528F2A0F091A.taxon	etymology	Etymology. The species is named in memory of my colleague Viktor Anatolievich Krivokhatsky (1954 – 2021), the world specialist on Neuroptera, Myrmeleontidae (Volkovitsh et al. 2021, Ovtshinnikova et al. 2021).	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF895F23D3B9528F2A0F091A.taxon	diagnosis	Differential diagnosis. The new species is close to E. merkli by having not beaded lateral margins of pronotum (Figs 14 E, 15 E) and smooth punctured sculpture of the prothoracic hypomera (Figs 14 F, 15 F), but is different from other congeners of the huedepohli species-group by strongly convex (lateral margin angulate in dorsal view) eyes (Fig. 14 D) and smooth prothoracic hypomera with sparse coarse punctures (except for E. merkli, which has the similar sculpture of hypomera). Euboeus krivokhatskyi has a similar shape of the pronotum as E. grimmi, not beaded male pronotum as is in E. merkli and the apical part of the basal piece of the aedeagus with longitudinal rugosity as in E. huedepohli and E. kasatkini. The new species additionally differs from: – E. huedepohli and E. kasatkini by not beaded laterally male pronotum and partly beaded female pronotum (Figs 14 D, E) (lateral margins of the pronotum are beaded in both comparable species (Figs 11 E, F, 13 E, F )), and the absence of the dense brush of setae at the base of the flex side of profemora (Fig. 14 B), the shape of the apical piece of aedeagus; – E. grimmi by shiny body dorsally, at least in the Elburs population (Fig. 14 A) (weakly shiny in E. grimmi (Fig. 9 A )), not beaded laterally male pronotum (Figs 14 D, E) (lateral margins of the pronotum are beaded in E. grimmi (Fig. 9 D, E )), the absence of the dense brush of setae at the base of the flex side of profemora (Fig. 14 B), the apical part of the basal piece of the aedeagus with longitudinal rugosity (Fig. 14 K) (smooth in E. grimmi (Fig. 9 K )), the apical piece of the aedeagus with longitudinal median groove (Fig. 14 K) (median keel in E. grimmi (Fig. 9 K )); – E. merkli by the shiny male body at least in the Elburs population (14 A) (weakly shiny in E. merkli (15 A )), wider and the widest ahead of the middle pronotum with rounded lateral margins (Fig. 14 D) (E. merkli has narrower, widest at middle pronotum, with very weakly rounded lateral margins (Fig. 15 D )), the presence of dense brush of setae at the base of the flex side of meso- and metafemora (Fig. 14 B) (absence of the femoral brush in E. merkli (Fig. 15 B )) and almost bare abdominal ventrites (Fig. 14 B) (pubescent in E. merkli (Fig. 15 B )).	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF8B5F2DD3B950E22A760918.taxon	description	(Fig. 15)	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF8B5F2DD3B950E22A760918.taxon	materials_examined	Type material. Holotype, ♂ (HNHM) and paratypes, 3 ♂ (two in SMS, one in HNHM), 1 ♀ (SMS): Iran, Mazandaran Prov., Alborz Mts., Damavand Mt., Reyneh [Rineh], 35 ° 52.412 ʹ N, 52 ° 06.096 ʹ, 2841 m, 14. v. 2008, from beneath stone (leg. T. Hácz, K. Székely, K. Vig). Paratypes: 1 ♂ (HNHM): Iran, Abgarm (Damavand), 2100 – 2400 m, 5. v. 1970 (Wittmer, v. Bothmer), “ zoroaster Sdl det. Kaszab ”; 1 ♀ (HNHM): Iran, Tehran Prov. [Mazandaran Prov.], Elburz Mts., 10 km W Reine [Rineh], Kuh-e Demavend, 35 ° 53 ʹ 22 ʺN, 52 ° 02 ʹ 24 ʺE, 5. v. 1998 (Gy. Fábian, K. Székely); 1 ♂ (HNHM): Iran, Mazandaran Prov., Demavend Mt., 3000 m, 4. vi. 1999 (leg. T. Hácz, G. Kőszegi, K. Gaskó); 2 ♀ (HNHM): Iran, Mazandaran Prov., Kühha-ye Alborz, Küh-e Damavand, 3500 m, 30. vi. – 2. vii. 2006 (leg. G. Rozner); 1 ♂, 2 ♀ (HNHM): Iran, Mazandaran Prov., Alborz Mts., 15 km E of Lar-e Polur, 35 ° 52 ʹ 28 ʺN, 52 ° 06 ʹ 10 ʺE, 2580 m, 12 – 13. v. 2001 (leg. Gy. Fábian, K. Vig).	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF8B5F2DD3B950E22A760918.taxon	description	Description. Male. Body narrow, slender, black, moderately shiny (Fig. 15 A). Measurements: Y = 1.55; PH w = 1.42; P w P l = 1.08; E l E w = 1.57; EH w = 2; EP w = 1.4; EP l = 2.46. Body length 12 – 14 mm, width 4 – 5 mm. Head (Figs 15 B, D, E). Anterior margin of epistoma slightly rounded. Head widest at eye level. Eyes moderately convex, large in dorsal view, circumocular impression narrow (Fig. 12 D). Lateral margins of genae evenly rounded. Puncturation of head fine and sparse, interpuncture distance 2 – 3 times as long as puncture diameter. Ventral side of epicranium coarsely and densely punctured by merged punctures. Antennae comparatively short, with only three apical antennomeres, extending beyond base of pronotum, reaching 1 / 5 of elytral length. Prothorax (Figs 15 D – F). Pronotum subquadrate, widest at middle, very weakly transverse, almost with subequal width and length (Fig. 15 D). Lateral margins of pronotum very weakly evenly rounded; anterior margin slightly rounded, almost straight; base weakly rounded. Antero-lateral corners not projected, obtuse, narrowly rounded at apex; postero-lateral corners weakly obtuse, pointed at apex. Lateral margins not beaded; anterior margin unclearly beaded only near corners (Fig. 15 E). Disc of pronotum evenly strongly convex (only area near postero-lateral corners slightly flattened), not flattened along lateral margin, with basal impressions: oval transverse at middle and two obliques laterally. Puncturation of disc fine and sparse, puncture diameter 4 – 5 times as short as interpuncture distance in middle and 2 – 3 times on lateral sides; punctures round (Fig. 15 D). Prosternum with coarse and very sparse simple puncturation (Fig. 15 B). Prothoracic hypomera not flattened along margin, with coarse, sparse, large, round punctures along margin and near procoxae and dense and coarse merged punctures at middle; surface without microgranulation (Fig. 15 F). Prosternal process shiny, moderately punctured, slightly convex at apex, not beaded, pubescent by recumbent setae (Figs 15 B, F). Pterothorax (Figs 15 A, B). Scutellar shield with straight margins, finely and densely punctured. Elytra slightly shiny than pronotum, elongate, widest at middle; punctures in striae fine, round, widely spaced; punctures in interstriae fine and sparse, much smaller than strial ones (Fig. 15 A). Anterior half of mesoventrite densely pubescent, coarsely and densely wrinkled. Mesepisterna, mesepimera and metepisterna with coarse and dense puncturation of moderate in size punctures and short setae; metaventrite with moderately coarse and moderately dense punctures; intercoxal part of mesoventrite finely and sparsely punctured and sparsely pubescent; metaventrite pubescent with recumbent comparatively long setae (Fig. 15 B). Legs. Trochanters with brush of short dark-reddish setae and one long seta (Fig. 15 B). Basal third of meso- and metafemora without the same brush on flex side, but with simple recumbent setation (Fig. 15 B). Tibiae straight. Pro- and mesotarsomeres 1 – 4 strongly widened, trasverse, subequal to width of apex of protibia. Abdomen (Figs 15 B, G – K). Abdominal ventrites 1 – 3 with the same puncturation as on metaventrite, ventrites 4 – 5 much sparser and finer punctured. All ventrites pubescent with long recumbent setae (Fig. 15 B); ventrite 5 with several sparse, longer and stronger setae at apex. Genitalia (Figs 15 G – K). Apical part of the basal piece of aedeagus smooth, without longitudinal rugosity (Fig. 15 K). Apical piece narrowly rounded at apex, dorsal surface with longitudinal small median groove (Fig. 15 K). Female (Fig. 15 C). Body robuster, elytra dull. Eyes smaller (see mesuarements). Antennae short, not reaching base of pronotum. Lateral margins of pronotum little more narrowed to base. Measurements: Y = 1.43; PH w = 1.42; P w P l = 1.1 – 1.12; E l E w = 1.58; EH w = 2; EP w = 1.4; EP l = 2.64 – 2.65. Body length 12 – 16.5 mm, width 4.5 – 6 mm.	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF8B5F2DD3B950E22A760918.taxon	etymology	Etymology. The new species is named in memory of Ottó Merkl (1957 – 2021), my friend, colleague and coauthor of several papers, the famous Hungarian entomologist (Schawaller 2021, Szél et al. 2021).	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF8B5F2DD3B950E22A760918.taxon	diagnosis	Differential diagnosis. This new species is similar to E. krivokhatskyi by the surface of prothoracic hypomera (Figs 11 F, 12 F) and not beaded lateral margins of the pronotum (Figs 11 E, 12 E). See differences in the diagnosis of E. krivokhatskyi.	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF855F2ED3B950E329770EC0.taxon	description	(Figs 16, 17)	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF855F2ED3B950E329770EC0.taxon	materials_examined	Type material. Holotype, ♂ (ZIN): “ Mont. Taliish ”, “ quadraticollis Menet. Zouvant ”, bronze square, “ Holotypus Helops quadraticollis Ménétriés, 1832 ” (Nabozhenko’s label) (Fig. 13 B). Material. Azerbaijan. 1 ♂, 4 ♀ (ZIN): Azerbaijan, 25 km below Yardýmlý, meadows, 2 – 3. vi. 2008 (leg. D. Kasatkin). Iran. 1 ♂ (SMNH): Iran, Ardabil Prov., Nir env., 27. iv. 2006 (leg. M. Machálik); 1 ♀ (SMNH); 1 ♂ (NMPC): Iran, Azerbaijan-e Sharqi Prov., Sefide Khan, 25 km S Tabriz, 7. v. 2002 (S. Kadlec). Note. Full bibliography for this species covering complicated historical and nomenclatural issues with application of two names quadraticollis / quadricollis for this species, discussion about the author of the taxon, as well as short redescription and figures were published by Abdurakhmanov & Nabozhenko (2011).	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF855F2ED3B950E329770EC0.taxon	distribution	Distribution. Azerbaijan (Talysh Mts.: Yardýmlý and Lerik districts), North Iran (Ardabil and Azerbaijan-e Sharqi provinces). New record for Iran.	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF805F28D3B956D6291F084F.taxon	description	(Fig. 18)	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF805F28D3B956D6291F084F.taxon	materials_examined	Type material. Holotype, ♂ (ZIN): Iran, Kurdistan Prov., between Saqqez and Baneh, 36 ° 46 ʹ 37 ʺ N, 48 ° 48 ʹ 13 ʺE (leg. A. Zubov).	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF805F28D3B956D6291F084F.taxon	description	Description. Male. Body slender, black, moderately shiny. Measurements: Y = 1.56; PH w = 1.35; P w P l = 1.15; E l E w = 1.7; EH w = 1.8; EP w = 1.34; EP l = 2.7. Body length 11.5 mm, width 3.9 mm. Head (Figs 18 A). Anterior margin of epistoma straght. Head widest at eye level. Eyes moderately convex, large in dorsal view, circumocular impression wide (Fig. 18 D). Lateral margins of genae angularly rounded. Puncturation of head moderately coarse and sparse, interpuncture distance 1.5 – 2 times as long as puncture diameter. Head pubescent with sparse, reddish, moderately long, recumbent setae (Figs 18 D, E). Ventral side of epicranium coarsely and densely punctured by raduliform punctures, gula almost smooth (Fig. 18 B). Antennae comparatively short, with only two apical antennomeres, extending beyond base of pronotum, reaching 1 / 6 of elytral length. Prothorax (Figs 18 B, D – F). Pronotum weakly cordate, widest at middle, weakly transverse (Fig. 18 D). Lateral margins of pronotum moderately rounded, slightly emarginated near antero-lateral corners and stronger emarginated near postero-lateral corners; anterior margin slightly rounded at middle and widely emarginated laterally; base weakly bisinuate. Antero-lateral corners projected, acute and pointed at apex; postero-lateral corners straight, pointed at apex. Lateral margins beaded (Figs 18 D, E); anterior margin with interrupted bead at middle. Disc of pronotum evenly strongly convex, not flattened along lateral margin, only lateral sides more gentle. Puncturation of disc fine and sparse, puncture diameter near 3 – 4 times as short as interpuncture distance in middle and 3 times on lateral sides; punctures round (Fig. 18 D). Prosternum with coarse, sparse, short transverse wrinkles and pubescent with long, recumbent, sparse setae (Fig. 18 B). Prothoracic hypomera not flattened along margin, with smooth, weak, sparse wrinkles and distinct sparse granules bearing long seta (Fig. 18 F). Prosternal process shiny, moderately punctured on sides, densely pubescent with subrecumbent reddish setae slightly convex at apex, beaded (Figs 18 B, F). Pterothorax (Figs 18 A, B). Scutellar shield triangle, with straight margins, coarsely and densely punctured. Elytra elongate-oval, widest at middle; punctures in striae fine, slightly elongate, widely spaced; punctures in interstriae fine and sparse, with almost the same size than strial ones, but deeper (Fig. 18 A). Anterior half of mesoventrite densely pubescent, coarsely and densely wrinkled. Mesepisterna, mesepimera and metepisterna with coarse and dense puncturation and long recumbent setae; intercoxal part of mesoventrite finely and sparsely punctured and sparsely pubescent; metaventrite with moderately coarse and moderately dense punctures, pubescent with long recumbent setae (Fig. 18 B). Legs. Trochanters with brush of short dark-reddish setae and one long seta (Fig. 18 B). Basal third of meso- and metafemora without the same brush on flex side (Fig. 18 B). Tibiae straight. Pro- and mesotarsomeres 1 – 4 strongly widened, trasverse, subequal to width of apex of protibia. Abdomen (Figs 18 B, C, G – J). Abdominal ventrites with moderately sparse, fine raduliform puncturation and pubescence of long, recumbent, reddish setae. Genitalia (Figs 18 C, G – J). Apical part of the basal piece of aedeagus almost smooth, with only pair small areas of fine longitudinal rugosity (Fig. 18 J). Apical piece strongly narrower from base to apex, widely rounded at apex, dorsal surface without median groove or keel (Fig. 18 J). Female unknown.	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF805F28D3B956D6291F084F.taxon	etymology	Etymology. The new species is named in honour of Andrey Zubov (Moscow, Russia), my colleague and excellent insect collector.	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF805F28D3B956D6291F084F.taxon	diagnosis	Differential diagnosis. The new species differs from all representatives of the huedepohli species-group by the following characters: head pubescent with recumbent setae; pronotum with projected, acute, pointed at apex anterolateral corners; prothoracic hypomera with sparse distinct granules; the apical piece of the aedeagus sub-triangle, strongly narrowed from base to apex.	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF835F2BD3B956D62FE10DB1.taxon	description	(Figs 19, 20)	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF835F2BD3B956D62FE10DB1.taxon	materials_examined	Type material. Holotype, ♂ (ZIN): “ В. Персiя Имам-Гуляръ — Кучанъ 16. III. 98 Зарудный ” (Cyrillic, old Russian) [E Persia, Imam-Gulyar — Quchan, leg. Zarudny], “ Holotypus Probathicus (Pelorinus) chorasanicus G. Medvedev, 1977 ” (Fig. 16 D). Paratype, ♀ (ZIN): “ Хорасан М. Кучан — Имам Гуляр 16. III. 98 Н. Зарудный ” (Cyrillic, Russian) [Khorasan, Kuchan — Imam-Gulyar, leg. N. Zarudny] (Fig. 15 D). Red labels with “ Holotypus ” and “ Paratypus ” were pinned by a laboratory assistant (A. K. Chistaykova) with erroneous year and the name of the genus. Material. 1 ♂, 1 ♀ (SMNS): Iran, North Khorasan Prov., Asadij (20 km SE Bojnurd), 1700 m, 4.04.2001 (leg. G. Sama); 1 ♂ (ZIN): Iran, North Khorassan Prov., Bojnurd area, Gerivan, 37 ° 17 ʹN, 57 ° 13 ʹE, 1800 m, 12. v. 2009 (leg. A. Klimenko); 1 ♀ (ZIN): the same label, but 8. vi. 2009; 1 ♀ (NMPC): NE Iran, Razavi Khorasan Province, Kuh-e- Binalud, 20 km NE Nishapur, 13 – 15. vi. 1977, 2500 m, Loc. No 366, (leg. Exped. Nat. Mus. Praha).	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF835F2BD3B956D62FE10DB1.taxon	distribution	Distribution. Iran (North Khorasan and Razavi Khorasan provinces). This species is terrestrial and inhabits xerophytic areas. Variability. The female from Binalud Mt. differs by the weakly shiny body and dull elytra.	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFFBD5F15D3B956D62EB80BB3.taxon	description	(without E. prometheus, E. arzanovi and E. zubovi whith unknown females)	en	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
