identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
FA1487CEFF9D5F36D3B9513B29430E54.text	FA1487CEFF9D5F36D3B9513B29430E54.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euboeus tenebricosus Nabozhenko	<div><p>Euboeus tenebricosus species-group</p><p>This group unites many species from the Balkans in the west to Turkmen-Khorasan mountains in the east. Species representing this group are characterized by matt dorsal side of the body, coarse and dense puncturation of pronotum and strong long setae on the apical part of the abdominal ventrite 5. Usually, representatives of this group are associated with forests, but some species are geobiontic. All Iranian taxa of the group are inhabitants of woodlands, except for probably Euboeus corrugosus .</p></div>	https://treatment.plazi.org/id/FA1487CEFF9D5F36D3B9513B29430E54	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nabozhenko, Maxim V.	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF9E5F37D3B9572F29A10EBC.text	FA1487CEFF9E5F37D3B9572F29A10EBC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euboeus (Pelorinus) corrugosus (Seidlitz 1895)	<div><p>Euboeus (Pelorinus) corrugosus (Seidlitz, 1895)</p><p>(Figs 1, 2)</p><p>Helops corrugosus Seidlitz, 1895: 707, 750 (part);</p><p>Probaticus (Pelorinus) corrugosus: Leo &amp; Ruzzante, 2016: 85 (part);</p><p>= Probaticus (Pelorinus) parthorum G. Medvedev, 1976: 891, syn. nov.</p><p>Type material. Lectotype of Helops corrugosus Seidlitz, 1895 (SNSD), designated here according to Article 61.1 and 74.1 of ICZN (1999), with labels: bronze square, “ subrugosus tut Allard”, “ ♂ astrabad Christoph”, “13916”, “ Helops corrugatus Seidl. ” [handwritten by G. Seidlitz], “ corrugosus Seid. nec corrugatus det.Schuster”. The label “ Helops corrugosus Seidlitz, 1895 des. M. Nabozhenko 2021” was added to the lectotype.</p><p>Holotype of Probaticus parthorum (ZIN), ♂ with the Cyrillic Russian handwritten label “Астрабад, Сев. Иран 9–12.V.1912 Билькевич” (Astrabad, North Iran, leg. Bil’kevich), “ Holotypus Probathicus (Pelorinus) parthorum G. Medvedev, 1977 ” . Paratypes. 1♂ with labels: “Астрабад С. Персия [print] 6.III.904 [handwritten] Филиппович [print italic]” ( Cyrillic, Russian: Astrabad, N Persia, leg. Philippovich) and “ Paratypus Probathicus (Pelorinus) parthorum G. Medvedev, 1977 ” ; 1♂: the same labels, but 18.II.1904; 1♂: the same labels, but 5.V.[19]04; 1♀: the same labels, but 23–24.IV.[1]905; 1♀: the same labels, but 29.V.[19]04 and additional labels “ Probaticus sp. aff. 4-collis Bdi Dr Z. Kaszab det., 1960”, “ Probaticus sp. ” Red labels with “ Holotypus ” and “ Paratypus ” were pinned by a laboratory assistant (A.K. Chistaykova) with erroneous year and the name of the genus.</p><p>Material. 1♂, 1♀ (HNHM): “Persia Astrabad 5.99 Coll. Hauser ” (Iran, Golestan Prov., Gorgan, v.1899); 1♂ (SMNS): Iran, Golestan Prov., Karim Ishan, 70 km NE Gonbad-e-Kavus, 4– 7.04.1999 (leg. J. Rejsek) ; 2♂, 1♀ (SMNS): Iran, Golestan Prov., 10 km S Bandar e Gaz, 27.05– 7.06.2001 (leg. G. Sama) .</p><p>Notes. This species was described from two males from “Klein-Asien” and “Astrabad” (Seidlitz, 1895, p. 707) in the key. Leo and Ruzzante (2016) mentioned that it is a “mysterious” species, described from these two different localities. Sahlberg (1913), Reitter (1922) and Nabozhenko and Löbl (2008) mentioned different species from the Aegean region. Nabozhenko (2020 b) listed E. corrugosus for Greece, Turkey and Iran. One syntype (male) from “ Asia Minor ” was originally mentioned from the collection of G. Kraatz, but it was not found in the Senckenberg Deutsche Entomologische Institut (Müncheberg, Germany). I studied the second Iranian specimen (designated here as the lectotype) (Fig. 1), which is conspecific to Medvedev’s E. parthorum (Fig. 2) also described from Astrabad. Thus, the following new synonymy is proposed: Helops corrugosus Seidlitz, 1895 = Probaticus parthorum G. Medvedev, 1976, syn. nov.</p><p>Distribution. Iran (Golestan Province).</p></div>	https://treatment.plazi.org/id/FA1487CEFF9E5F37D3B9572F29A10EBC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nabozhenko, Maxim V.	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF985F30D3B956D62E200BDE.text	FA1487CEFF985F30D3B956D62E200BDE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euboeus (Pelorinus) gorganicus (G. S. Medvedev 1976)	<div><p>Euboeus (Pelorinus) gorganicus (G.S. Medvedev, 1976)</p><p>(Figs 3, 4)</p><p>Probaticus (Pelorinus) gorganicus G.S. Medvedev, 1976: 892;</p><p>Probaticus (Pelorinus) prometheus: Grimm, 2015: 2010 (misidentification).</p><p>Type material. Holotype, ♂ (ZIN): “Астрабад с. Персия 29.V.905 Филиппович” (Cyrillic, Russian) [N Persia, Astrabad, leg. Filippovich], “ Holotypus Probathicus (Pelorinus) gorganicus G. Medvedev, 1977 ”. The red label “ Holotypus ” was pinned by a laboratory assistant (A.K. Chistaykova) with an erroneous year and the name of the genus.</p><p>Material. 1♀ (HNHM): “ Persia, Astrabad 4.99 Coll. Hauser ” , 1♀ (HNHM), the same label, but 5.[18]99 (Iran, Golestan Prov., Gorgan) ; 2♂ (HNHM): Iran, Ali-abad, E Gorgan, 10.viii.1967 (leg. Heinz) ; 1♀ (HNHM): Iran, Assala, 1200 m, 1970 (leg. Wittmer, v. Bothner); 3♂ (HNHM): Iran Gole [stan], Lovae (= Loveh), 750–1400 m (leg. Wittmer v. Bothner) ; 1♀ (SMNS): Iran, W Mazandaran, Hasankif, 19– 21.04.1999 (leg. J. Rejsek) ; 3♀ (HNHM): Iran, Mazandaran Prov., Ghorogh, 200 m, 23.04.1999 (leg. N. Rahmé) ; 2♂ (NMPC): Iran, Mazandaran Prov., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=51.2&amp;materialsCitation.latitude=36.566666" title="Search Plazi for locations around (long 51.2/lat 36.566666)">10 km S Čalűs</a>, 36°34ʹN, 51°12ʹE, 250–400 m, 21–23.vi.2000 (leg. S. Kadlec) ; 1♂ (HNHM): Iran, Mazandaran Prov., 15 km S of Azad Sar, 24–25.iv.2000 (leg. K. Gaskó) ; 1♂ (HNHM), same locality, 28–29.vi.2000 (leg. G. Rozner); 1♂ (HNHM): Iran, Mazandaran Prov., 12 km S of Azad Sar, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=55.279446&amp;materialsCitation.latitude=37.03" title="Search Plazi for locations around (long 55.279446/lat 37.03)">Sah Mts</a>, 37°01ʹ48ʺN, 55°16ʹ46ʺE, 500 m, 16.v.2000 (leg. K. Székely) ; 1♂ (SMNS): Iran, Mazandaran Prov., 12 km E Kelardasht m., 19.vi.2003 (leg. G. Sama) ; 2♂, 7♀ (PCMN): Iran, Mazandaran Prov. 20 near <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=51.37597&amp;materialsCitation.latitude=36.555798" title="Search Plazi for locations around (long 51.37597/lat 36.555798)">Chalus</a>, 36°33ʹ20.88ʺN, 51°22ʹ33.49ʺE, 25– 27.05.2015 (leg. D.G. Kasatkin) ; 1♂, 1♀ (ZIN): Iran, Mazandaran Prov., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=51.37597&amp;materialsCitation.latitude=36.555798" title="Search Plazi for locations around (long 51.37597/lat 36.555798)">15 km above Chalus</a>, 36°33ʹ20.88ʺN, 51°22ʹ33.49ʺE, 30– 31.05.2017 (leg. D.G. Kasatkin) ; 2♂, 1♀ (PCSB): N Iran, Golestan Prov., 30 km E Minudasht, Gole-Loveh, 750 m, 21–27.iv.1974 (leg. Holzschuh &amp; Ressl) ; 1♀ (NMPC): Iran, Golestan Prov., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=55.988335&amp;materialsCitation.latitude=37.368332" title="Search Plazi for locations around (long 55.988335/lat 37.368332)">2 km E Tunel-e</a> Golestan (streem valley, oak forest, at light), 37°22.1ʹN, 55°59.3ʹE, 835 m, 26–27.v.2006 (leg. J. Hájek, P. Chvojka) ; 3♂ (PCMN): Iran, Golestan Prov., near Gorgan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=54.4688&amp;materialsCitation.latitude=36.672966" title="Search Plazi for locations around (long 54.4688/lat 36.672966)">Ziarat</a>, 36°40ʹ22.67ʺN 54°28ʹ7.68ʺE, 27.05.2016 (leg. D.G. Kasatkin) ; 5♂ (PCMN): Iran, Golestan Prov., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=54.583332&amp;materialsCitation.latitude=36.716667" title="Search Plazi for locations around (long 54.583332/lat 36.716667)">20 km S Gorgan</a> city, 36°43ʹN, 54°35ʹE, 700 m, 9.vi.2009 (leg. A. Klimenko) ; 1♂ (PCMN): Iran, North Khorasan Prov., Golestan <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=55.851543&amp;materialsCitation.latitude=37.379536" title="Search Plazi for locations around (long 55.851543/lat 37.379536)">National</a> park, 37°22ʹ46.33ʺN 55°51ʹ5.56ʺE, ~ 600 m, 24– 25.05.2016 (leg. D.G. Kasatkin) .</p><p>Distribution. Iran: Mazandaran, Golestan and west part of North Khorasan provinces, in forests.</p></div>	https://treatment.plazi.org/id/FA1487CEFF985F30D3B956D62E200BDE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nabozhenko, Maxim V.	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF9A5F33D3B956D62F6B0D74.text	FA1487CEFF9A5F33D3B956D62F6B0D74.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euboeus (Pelorinus) prometheus (Reitter 1902)	<div><p>Euboeus (Pelorinus) prometheus (Reitter, 1902)</p><p>(Fig. 5)</p><p>Helops prometheus Reitter, 1902: 214;</p><p>Probaticus (Pelorinus) prometheus: Reitter, 1922: 43; Abdurakhmanov &amp; Nabozhenko, 2011: 149 (fig. 337), 150 (fig. 346), 169, 307.</p><p>Probaticus prometeus: Kühnelt, 1957: 79</p><p>Full bibliography for this species is published by Abdurakhmanov &amp; Nabozhenko (2011, p. 307).</p><p>Type material. Holotype, ♂ (HNHM): “Talysch.”, “ Coll. Reitter ”, “ H. Prometheus m. 1900 Type.”, “ Holotypus 1901 ♂ Helops prometheus Reitter ”.</p><p>Notes. Reitter (1902) listed “ 1♂ ♀ ” in the original description, but I suggest that it is a misprint, because he described only male characters and only one male is deposited in HNHM. In the same paper, he always indicated whether a character refers to a female or a male for other species and wrote “ 1 ♂ und ♀ ”.</p><p>Distribution. Iran (Talysh). Extensive collecting efforts (more than 190 years) have not produced any records of this species in the territory of the modern Azerbaijani part of Talysh, where only E. quadricollis was collected in alpine meadows. In conclusion, I’m sure that the species occurs Iranian part of the Talysh Mountains and can be excluded from the Azerbaijani faunistic lists. Reitter (1922) also recorded E. prometheus for Astrabad (Now Gorgan, Iran), probably confusing it with E. gorganicus . Kühnelt (1957) recorded E. prometheus from the Caucasus and Central Iran; the second locality is erroneous. Abdurakhamanov &amp; Nabozhenko (2011) partly mixed E. prometheus with females of E. gorganicus and erroneously recorded it for Mazandaran Province.</p></div>	https://treatment.plazi.org/id/FA1487CEFF9A5F33D3B956D62F6B0D74	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nabozhenko, Maxim V.	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF9B5F33D3B954CF28780988.text	FA1487CEFF9B5F33D3B954CF28780988.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euboeus (Pelorinus) vicinus (Allard 1876)	<div><p>Euboeus (Pelorinus) vicinus (Allard, 1876)</p><p>(Fig. 6)</p><p>Helops vicinus Allard, 1876: 14, 34;</p><p>Helops vicinus: Allard, 1877: 26, 91;</p><p>Helops vicinus: Seidlitz, 1895: 706;</p><p>Probaticus (Pelorinus) vicinus: Reitter, 1922: 33, nota 28.;</p><p>Probaticus (Pelorinus) medvedevi Abdurakhmanov et Nabozhenko, 2011, syn. nov.;</p><p>Euboeus (Pelorinus) medvedevi: Nabozhenko et al., 2021b: 441</p><p>Type material. Syntypes of Helops vicinus were not found by curators in the Muséum national d’Histoire naturelle Paris. Holotype, ♀ (ZIN) of Probaticus medvedevi with labels: Cyrilliс “Арм. ССР Алаверди 25.04.86 Мелях С.В.” (Armenian SSR, Alaverdi, 25.iv.1986, leg. S.V. Melyakh), “ Probaticus (Pelorinus) medvedevi Nabozhenko et Abdurakhmanov 2011 ” .</p><p>Material. 1♂, 2♀: “Kaukasus Leder”, “Allard type”, “ Coll. Reitter ”, “ Paratypus 1877 Helops vicinus Allard ” (curator’s label) .</p><p>Notes. This very rare species was described in 1876 based on the specimen(s) from the collection of Reiche (Allard 1876) and redescribed in 1877 using the type specimen(s) and beetles collected by Leder (Allard 1877). Later, Schneider &amp; Leder (1878) again published Allard’s redescription 1877 and added the species’ type locality: “Elisabetthal”. It was a settlement of German colonists, which is now called Asureti (Georgia, 41°35′45″N, 44°40′00″E). Both description and redescription (Allard 1876, 1877) have differences, and the latter is more consistent with specimens from Georgia and northern Armenia, as mentioned by Abdurakhmanov &amp; Nabozhenko (2011). In addition, the Russian authors erroneously interpreted species from South Armenia and Nakhchivan as E. vicinus (described below as E. kalashiani sp. nov.) since Bogachev (1938). As a result, Abdurakhmanov &amp; Nabozhenko described a new species Probaticus medvedevi, from northern Armenia (Figs 6D, F). Allard examined the specimens of Leder and confirmed that they belong to his species Helops vicinus, as he wrote in redescription (Allard 1877). The Leder’s specimens (Figs 6A–C, E, G–K) are not syntypes (as marked on the labels) because the original description was based on other specimens from the Reiche collection. The females from Georgia are conspecific to the female (holotype) of E. medvedevi despite the variability. As a result, I propose the following synonymy: Helops vicinus Allard, 1876 = Probaticus (Pelorinus) medvedevi Abdurakhmanov et Nabozhenko, 2011, syn. nov.</p><p>Distribution. Southern Georgia, northern area of Armenia.</p></div>	https://treatment.plazi.org/id/FA1487CEFF9B5F33D3B954CF28780988	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nabozhenko, Maxim V.	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF955F3ED3B956D628CC0CC8.text	FA1487CEFF955F3ED3B956D628CC0CC8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euboeus (Pelorinus) zoroaster (Seidlitz 1895)	<div><p>Euboeus (Pelorinus) zoroaster (Seidlitz, 1895)</p><p>(Fig. 7)</p><p>Helops zoroaster Seidlitz, 1895: 714, 751;</p><p>Probaticus (Pelorinus) zoroaster: Reitter, 1922: 35; Kühnelt, 1957: 79;</p><p>Probaticus zoroaster: Medvedev &amp; Nepesova, 1985: 149 (fig. 93), 152; Nabozhenko, 2009: 420;</p><p>Helops diabolinus Brancsik, 1899: 101 t. 4, f. 5, synonymized by Nabozhenko (2009);</p><p>Zophohelops diabolinus: Reitter, 1922: 118, nota.</p><p>Full bibliography for this species is published by Nabozhenko (2009).</p><p>Type material. Not studied. The holotype, ♀ (type locality: “ Askabad ”, now Ashgabad, Turkmenistan) originated from the collection of O. Staudinger, which is currently deposited (at least beetles) in SNSD. The holotype was not found by the curator of the SNSD collection Olaf Jäger.</p><p>Material. Turkmenistan. 1♀ (HNHM): “Trans Caspi Eylandt” (Turkmenistan, Transcaspian area); 2♀ (ZIN) : Zakaspiyskaya oblast’ (Turkmenistan, Transcaspian area)), Aydere canyon, 10.v.1890 (leg. Eylandt); 1♂, 1♀ (ZIN) : Turkmenistan, Kopetdag, Aydere, 3.v.1930 (leg. E. Shestopalov); 1♂, 2♀ (ZIN) : Turkmenistan, Aydere canyon, 0–3 km from river mouth, 30.iv.1952 (leg. K.G. Romadina); 2♀ (ZIN) : Turkmenistan, Kara-Kala (= Garrygala), N slope of Syunt Mt., 2.vii.1956 (leg. G.S. Medvedev); 1♂ (ZIN) , 1♀ (HNHM): Turkmenistan, Kara-Kala, a canyon on N slope of Syunt Mt., 6.v.1957 (leg. G.S. Medvedev); 4♀ (ZIN) : Turkmenistan, Kara-Kala, Savudzhe canyon, 8.v.1957, 11.v.1957 (leg. G.S. Medvedev); 1♀ (ZIN) Turkmenistan, N of Kara-Kala, 10.v.1957 (leg. G.S. Medvedev); 1♂, 1♀ (ZIN) : Turkmenistan, W Kopetdag, headwaters of Khozly canyon, 20.04.1975, 23.04.1975 (leg. V. Yanushev); 1♀ (HNHM) : Turkmenistan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=56.766666&amp;materialsCitation.latitude=38.233334" title="Search Plazi for locations around (long 56.766666/lat 38.233334)">Kopet-Dag Mts</a>, 10 km S Aydere, 38°14ʹN, 56°46ʹE, 600–1000 m, 27.vi.1992 (leg. Gy Fábián, B. Herczig, A. Podlussány, Z. Varga) . Iran. 1♂, 1♀ (NMPC): NE Iran, Khorasan, Golestan forest <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=56.0&amp;materialsCitation.latitude=37.333332" title="Search Plazi for locations around (long 56.0/lat 37.333332)">55 km NE Minudast</a>, 37°20ʹN / 56°00ʹE, 840 m, 17.vi.2000 (S. Kadlec); 2♂, 1♀ (HNHM) : Iran, Golestan, Golestan forest, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=55.833332&amp;materialsCitation.latitude=37.383335" title="Search Plazi for locations around (long 55.833332/lat 37.383335)">45 km NE of Minudasht</a>, 584 m, 16.vi.2015 (leg. A. Lasoń); 37°23‘N, 55°50‘E, 9♂, 4♀ (PCMN) : Iran, North Khorassan Prov., Golestan <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=55.851543&amp;materialsCitation.latitude=37.379536" title="Search Plazi for locations around (long 55.851543/lat 37.379536)">National</a> park, 37°22‘46.33“N, 55°51‘5.56“E, ~ 600 m, 24–25.v.2016 (leg. D.G. Kasatkin); 1♂, 2♀ (ZIN) : Iran, North Khorassan Prov., Golestan <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=55.94461&amp;materialsCitation.latitude=37.368332" title="Search Plazi for locations around (long 55.94461/lat 37.368332)">National</a> park, 37°22ʹ6ʺN, 55°56ʹ40.6ʺE, 26–27.v.2017 (leg. D.G. Kasatkin) . Turkmenistan / Iran. 1♀ (HNHM): “Arwas Pers. b.” ( Kopetdag Mts., Arvaz pass between Iran (North Khorasan Prov.) and Turkmenistan) .</p><p>Distribution. South West Turkmenistan (Kopetdag), Iran (North Khorasan Province).</p><p>Note. This is the only species of the genus that occurs in Turkmenistan.</p></div>	https://treatment.plazi.org/id/FA1487CEFF955F3ED3B956D628CC0CC8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nabozhenko, Maxim V.	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF965F3ED3B955932A950B34.text	FA1487CEFF965F3ED3B955932A950B34.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euboeus huedepohli (Kulzer 1964)	<div><p>Euboeus huedepohli species-group</p><p>This group unites alpine and mountain-steppe terrestrial Iranian species with more or less shiny black body in males, fine and sparse puncturation of pronotum (at least in the middle) and strong long setae on the apical part of the abdominal ventrite 5.</p></div>	https://treatment.plazi.org/id/FA1487CEFF965F3ED3B955932A950B34	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nabozhenko, Maxim V.	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF965F38D3B9528F292B0BE8.text	FA1487CEFF965F38D3B9528F292B0BE8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euboeus (Pelorinus) arzanovi Nabozhenko 2022	<div><p>Euboeus (Pelorinus) arzanovi sp. nov.</p><p>(Fig. 8)</p><p>Type material. Holotype, ♂ and paratype, ♂ (HNHM): Iran, Gachsar [Alborz Prov.]— Marzanabad [Mazandaran Prov.], 1700–2500 m, 28.iv.1970 (leg. Wittmer, v. Bothmer).</p><p>Description. Male. Body slender, elongate, black, with dull shine (Fig. 8A). Measurements: Y = 1.47; PH w = 1.44; P w P l = 1.09; E l E w = 1.74; EH w = 1.94; EP w = 1.34; EP l = 2.6–2.61. Body length 12.5–13 mm, width 4.5–4.8 mm.</p><p>Head (Figs 8B, E, F). Anterior margin of epistoma straight (Fig. 8E). Head widest at eye level. Eyes moderately convex, circumocular impression narrow. Lateral margins of genae angularly rounded. Puncturation of head moderately fine and sparse on frons (interpuncture distance near 3 times as long as puncture diameter) and denser on epistoma and genae (interpuncture distance subequal to or near 1.5 times as long as puncture diameter). Ventral side of epicranium coarsely and densely punctured; temples depressed behind eyes in the lower half (Fig. 8B). Antennae comparatively long, with four apical antennomeres, extending beyond base of pronotum, reaching 1/3 of elytral length.</p><p>Prothorax (Figs 8B, E–G). Pronotum weakly transverse, with almost equal length and width, widest at middle (Fig. 8E). Lateral margins of pronotum weakly rounded in middle part, straightly narrowed in apical third and widely emarginated in basal quarter; anterior margin trisinuate; base slightly rounded, almost straight, with emargination at middle. Antero-lateral corners not projected, right, pointed at apex; postero-lateral corners right, narrowly rounded at apex. Lateral margins not beaded; anterior margin only with very short bead marginally; disc completely beaded. Disc of pronotum convex, not flattened along lateral margin, without impressions. Puncturation of disc fine and sparse (puncture diameter 2–4 times as short as interpuncture distance), slightly coarser and denser on lateral sides; punctures round (Figs 8E, F). Prosternum with fine and sparse simple puncturation (Fig. 8B). Outer margin of prothoracic hypomera narrowly flattened only near base. Prohypomera with small confused microwrinkles with poorly distinguished sparse smoothed puncturation against their background (Fig. 8G). Prosternal process smooth and shiny (only rugose near apex), almost without punctures, slightly convex at apex, not beaded (Figs 8B, G).</p><p>Pterothorax (Figs 8A, B). Scutellar shield triangle, moderately coarsely punctured. Elytra strongly elongate, widest slightly behind middle; punctures in striae fine, round, widely spaced; punctures in interstriae fine and sparse, much smaller than strial ones (Fig. 8A). Anterior half of mesoventrite strongly wrinkled, pubescent, with more smooth middle (Fig. 8B). Mesepisterna coarsely and densely punctured by large foveae; puncturation of mesepimera and metepisterna sparser and finer; metaventrite coarsely and moderately sparsely punctured and pubescent with short setae.</p><p>Legs (Figs 8B, G). Trochanters with short dense reddish setae (but not dense brush), protrochanters with two long setae (Fig. 8G), meso- and metatrochanters with one long seta. Basal third of femora with the same dense on flex side (Fig. 8B); another surface of femora with sparse puncturation and short setae. Pro- and metatibiae slightly curved, mesotibiae stright. Pro- and mesotarsomeres 1–4 widened, trasverse, pro- and mesotarsomere 1 as wide as apex of protibia.</p><p>Abdomen (Figs 8B–D, H–J). Abdominal ventrites with moderately dense, fine puncturation and simple, moderate, short setae (shortly pubescent); ventrite 5 with coarser punctures and longer setae at apex. Genitalia (Figs 8C, D, H–J). Apical part of basal piece of aedeagus smooth, without rugosity. Apical piece of aedeagus with wide median longitudinal impression; lateral margins straight at middle and straightly narrowed to narrowly rounded apex.</p><p>Female. Unknown.</p><p>Etymology. The species is named in honour of my first teacher during student times and known specialist on Curculionidae DrSc. Yury Arzanov (Rostov-on-Don).</p><p>Differential diagnosis. The new species is close to E. merkli by the structure of the aedeagus (basal piece without rugosity, apical piece with median longitudinal impression) and the narrow, not beaded pronotum with weakly rounded margins, but differs in following characters: the lower half of temples is impressed behind the eye (weakly rounded in E. merkli); lateral margins of pronotum straight at apical quarter and widely emarginated in basal quarter (regularly rounded in E. merkli); prothoracic hypomera with microrugosity and fine sparse smooth punctures (large, coarse, round foveae and smooth interpuncture space between them in E. merkli); protrochanters with two long setae (one long seta in E. merkli).</p></div>	https://treatment.plazi.org/id/FA1487CEFF965F38D3B9528F292B0BE8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nabozhenko, Maxim V.	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF905F3AD3B952AC2A780A0A.text	FA1487CEFF905F3AD3B952AC2A780A0A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euboeus (Pelorinus) grimmi Nabozhenko 2022	<div><p>Euboeus (Pelorinus) grimmi sp. nov.</p><p>(Fig. 9)</p><p>Type material. Holotype, ♂ and paratype, ♀ (ZIN): Iran, Zanjan Prov., 40 km NE <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=48.731583&amp;materialsCitation.latitude=36.799168" title="Search Plazi for locations around (long 48.731583/lat 36.799168)">Bonab</a>, 36°47ʹ57ʺN, 48°43ʹ53.7ʺE, 2400 m, 9.v.2017 (leg. D.G. Kasatkin) . Paratypes. 1♀ (ZIN): Iran, Zanjan Prov., 36°45ʹ37ʺN, 48°48ʹ13ʺE, 9.v.2017 (leg. A. Zubov); 1♀ (SMS): Iran, Zanjan Prov., Alborz Mts., Sendan Mt., Zanjan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=48.88545&amp;materialsCitation.latitude=36.8077" title="Search Plazi for locations around (long 48.88545/lat 36.8077)">Abbar</a>, 36°48.462ʹN, 48°53.127ʹE, 1480 m, 11–12.v.2008, sigling (leg. T. Hácz, K. Székely, K. Vig) ; 1♀ (HNHM): Iran Zangan [Zanjan] Prov., Zangan [Zanjan], 1800–2000 m, 7.vi.1999 (leg. K. Gaskó) ; 1♀ (HNHM): Iran, Zanjan Prov., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=48.916668&amp;materialsCitation.latitude=36.8" title="Search Plazi for locations around (long 48.916668/lat 36.8)">Sendan Mts.</a>, 20 km SW of Gilvan, 36°48ʹN, 48°55ʹE, 1000 m, 18.v.2001 (leg. B. Benedek, G. Csorba) .</p><p>Description. Male. Body slender, elongate, black, with dull shine. Measurements: Y = 1.47; PH w = 1.6; P w P l = 1.23; E l E w = 1.7; EH w = 1.95; EP w = 1.2; EP l = 2.58. Body length 16 mm, width 5.5 mm.</p><p>Head (Figs 9B, D, E). Anterior margin of epistoma slightly rounded. Head widest at eye level. Eyes moderately convex, circumocular impression narrow. Lateral margins of genae angularly rounded. Puncturation of head fine and sparse, interpuncture distance 1.5–2 times as long as puncture diameter. Ventral side of epicranium coarsely and densely punctured. Antennae comparatively short, with only three apical antennomeres, extending beyond base of pronotum, reaching 1/5 of elytral length.</p><p>Prothorax (Figs 9B, D–F). Pronotum cordate, widest ahead of middle, transverse (Fig. 9D). Lateral margins of pronotum moderately rounded at apical half and slightly widely emarginated at basal half; anterior margin straight at middle and widely emarginated near corners; base slightly rounded. Antero-lateral corners projected, obtuse, widely rounded at apex; postero-lateral corners weakly obtuse, pointed at apex. Lateral margins beaded, interrupted only near antero-lateral corners; bead of anterior margin interrupted in middle. Disc of pronotum evenly weakly convex, not flattened along lateral margin, with oval impression on each lateral side and two small oblique impressions near base. Puncturation of disc fine and sparse, puncture diameter 2–4 times as short as interpuncture distance; punctures round (Figs 9D, E). Prosternum with moderately coarse and sparse raduliform puncturation (Fig. 9B). Prothoracic hypomera not flattened along margin, with small confused microwrinkles with poorly distinguished sparse smoothed puncturation against their background; wrinkles with microgranulation at anterior third (Fig. 9F). Prosternal process smooth and shiny, almost without punctures, slightly convex at apex, beaded only in basal half (Figs 9B, F).</p><p>Pterothorax (Figs 9A, B). Scutellar shield with slightly rounded margins, finely and sparsely punctured. Elytra strongly elongate, widest at middle; punctures in striae fine, slightly elongate, widely spaced; punctures in interstriae fine and sparse, much smaller than strial ones (Fig. 9A). Anterior half of mesoventrite pubescent, with triangle smooth middle and coarsely wrinkled another surface (Fig. 9B). Mesepisterna coarsely and densely punctured by large foveae; puncturation of mesepimera and metepisterna sparser and finer; metaventrite finely and sparsely punctured and covered with short setae.</p><p>Legs. Trochanters with brush of short dark-reddish setae and one long seta (Fig. 9B). Basal third of femora with the same brush on flex side (Fig. 9B); another surface of femora with coarse puncturation and short setae. Pro- and mesotibiae straight, metatibiae weakly bent. Pro- and mesotarsomeres 1–4 strongly widened, trasverse, pro- and mesotarsomere 1 as wide as apex of protibia.</p><p>Abdomen (Figs 9B, G–K). Abdominal ventrites with dense and fine puncturation and simple short setae (not pubescent); ventrite 5 with coarser punctures and long strong setae. Genitalia (Figs 9G–K). Apical part of basal piece of aedeagus smooth, without rugosity. Apical piece of aedeagus with median shiny longitudinal keel; lateral margins slightly emarginated at middle and sharply narrowed at apical quarter.</p><p>Female (Fig. 9C). Body much robuster, elytra matt. Eyes little smaller (see measurements). Antennae short, not reaching base of pronotum. Metatibiae straight. Measurements: Y = 1.4; PH w = 1.7; P w P l = 1.3; E l E w = 1.47; EH w = 2.28; EP w = 1.34; EP l = 2.57. Body length 13.5–17 mm, width 5,5– 6.5 mm.</p><p>Etymology. This species is named in memory of Roland Grimm (1948–2021), my colleague and co-author, a famous specialist on Tenebrionidae of the Old World (Burmeister 2021, Schawaller &amp; Staniczek 2021).</p><p>Differential diagnosis. This new species is different from all Iranian congeners by the presence of shiny keellike longitudinal median elevation on the apical piece of the aedeagus (Fig. 9K); all other species have the apical piece smooth or with the median impression. Euboeus grimmi sp. nov. has the pronotum with beaded lateral margins (Figs 9D, E) and bent metatibiae (Figs 9A, B) as E. hudepohli and E. kasatkini sp. nov. and differs from both species by the absence of longitudinal rugosity in apical part of the basal piece of the aedeagus (Fig. 9K) and the pronotum widest at the apical third (at middle in E. huedepohli and E. kasatkini). In addition, the new species is distinguished from:</p><p>– E. huedepohli by prothoracic hypomera not flattened along margin, covered with small confused microwrinkles with hardly distinguished sparse smoothed puncturation against their background (widely flattened and coarsely longitudinally wrinkled in E. huedepohli) (Fig. 9F); male dorsally is less shiny (lacquered shiny in E. huedepohli);</p><p>– E. kasatkini by prothoracic hypomera not flattened along the margin, covered with small confused microwrinkles with hardly distinguished sparse smoothed puncturation against their background (narrowly fattened and coarsely and densely punctured in E. kasatkini) (Fig. 9F); more bent male metatibiae and less shiny male body.</p></div>	https://treatment.plazi.org/id/FA1487CEFF905F3AD3B952AC2A780A0A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nabozhenko, Maxim V.	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF925F3BD3B953D428100EE4.text	FA1487CEFF925F3BD3B953D428100EE4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euboeus (Pelorinus) huedepohli (Kulzer 1964)	<div><p>Euboeus (Pelorinus) huedepohli (Kulzer, 1964)</p><p>(Figs 10, 11)</p><p>Probaticus huedepohli Kulzer, 1964: 228;</p><p>Probaticus (Probaticus) huedepohli: Grimm, 2015: 310;</p><p>Euboeus (Pelorinus) huedepohli: Nabozhenko et al., 2017 a: 495 (transferred from the nominotypical subgenus to Pelorinus).</p><p>Type material (studied). Holotype, ♂ (NMB): “ Persia Elburs-Gebg. Hüdepohl leg.”, “Lashgarak-Tal 12-4-63 ”, “ Holotypus Probaticus huedepohli n. sp. det H. Kulzer 1963” (Fig. 8C) . Paratype, ♂ (HNHM) with the same geographic label and “ Paratypus 1964 Probaticus huedepohli Külzer ”, “ PARATYPUS Probaticus huedepohli n. sp. det. H. Kulzer 1963”, “ Probaticus huedepohli Kulz. Dr. Kaszab det. 1983” .</p><p>Material. 1♂ (ZIN): Iran, Demavend, 24.v.1894 (leg. Glazunov) ; 1♂ (ZIN): Iran, 16 km NE Tehran, foothills of Elburs, 12.v.1955 (leg. D. Shteinberg) ; 2♂ (HNHM): Iran, Mazandaran Prov., Demavend, 3000 m, 3.v.2000 (leg. K. Gaskó) ; 3♂, 3♀ (HNHM): Iran, Tehran / Mazandaran provinces, Polour—Abali, 2100–2800 m, 17.v.1970 (leg. Wittmer. V. Bothmer) ; 2♂, 5♀ (HNHM): Iran, Tehran Prov. [Mazandaran Prov.], Elburz Mts., 10 km W Reine [Rineh], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=52.04&amp;materialsCitation.latitude=35.889446" title="Search Plazi for locations around (long 52.04/lat 35.889446)">Kuh-e Demavend</a>, 35°53ʹ22ʺN, 52°02ʹ24ʺE, 14.v.1998 (Gy. Fábian, K. Székely) ; 1♀, Iran, Mazandaran Prov., Kühha-ye Alborz, Küh-e Damavand, 3500 m, 30.vi.–2.vii.2006 (leg. G. Rozner) ; 1♀ (HNHM): “ Iran, prov. Khorasan, Qudjan (Qućan), 3000 m / Mt. Kopet Dag 1999. V.90 leg.: Gaskó K. ” ; 3♂, 3♀ (SMS): Iran, Mazandaran Prov., Alborz Mts., Damavand Mt., Reyneh [Rineh], 35°52.412ʹ N, 52°06.096ʹ, 2841 m, 14.v.2008, from beneath stone (leg. T. Hácz, K. Székely, K. Vig) ;</p><p>Distribution. Iran (Demavend Mt. from ~ 2800 to 3500 m). I didn’t include Razavi Khorasan Province in the distribution based on the strange locality “Quchan, Kopet Dag, 3000 m ”, because the Iranian part of Kopet Dag mountains does not exceed 2000 m, and only one mount Kuh-e Patu (not Kopet Dag ridge) with the elevation near 3000 m is located 50 km west of Quchan. This mountain belongs to another mountain system (Turkmen-Khorasan mountains) than Demavend and and labelling raises doubts.</p></div>	https://treatment.plazi.org/id/FA1487CEFF925F3BD3B953D428100EE4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nabozhenko, Maxim V.	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF8D5F27D3B956D62E030B58.text	FA1487CEFF8D5F27D3B956D62E030B58.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euboeus (Pelorinus) kalashiani Nabozhenko 2022	<div><p>Euboeus (Pelorinus) kalashiani sp. nov.</p><p>(Fig. 12)</p><p>Type material. Armenia. Holotype, ♂ (ZIN) with the Cyrillic label: “ЛичкваЗ—Нор-Аревик Мегринский р-н, Арм ССР 31.5.1980 А.Л. Лобанов” (Armenia, Lichk, Nor-Arevik, 31.v.1980, leg. A.V. Lobanov) . Paratypes: 1♂ (IZAY): Armenia, Syunik Prov., Tsav, 21.07.1952 (leg. S. Iablokoff-Khnzorian) ; 1♀ (IZAY): Armenia, Syunik Prov., Srashen, 23.vi.1981 (leg. M. Kalashian) ; 1♀ (IZAY): Armenia, Syunik Prov., 4–6 km N Shvanidzor, 20.iv.2001 (leg. K. Aghababyan) ; 1♀ (IZAY): Armenia, Syunik Prov., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=46.2205&amp;materialsCitation.latitude=39.1307" title="Search Plazi for locations around (long 46.2205/lat 39.1307)">Pukhrut</a>, 39.1307°N, 46.2205°E, 2130 m, 25.v.2005 (leg. G. Karagyan) ; 3♂, 1♀ (IZAY): Armenia, Syunik Prov., Meghri, 30–31.v.1923 ( Expedition of Museum of Armenia) ; 1♀ (IZAY): Armenia, Syunik Prov., Meghri, 24.v.1945 (leg. H. Avetian) ; 1♂ (IZAY): Armenia, Syunik Prov., Chakaten, 25.v.1942 (leg. H. Avetian) ; 1♂ (IZAY): Armenia, Syunik Prov., S environs of Lichk, abandoned Nor Arevik vill., 26.iv.1943 (leg. H. Avetian) ; 2♂, 1♀ (IZAY): same locality, 5.v.1954 (leg. S. Iablokoff-Khnzorian); 1♀ (IZAY): Armenia, Syunik Prov., Vahravar, 27.v.1945 (leg. H. Avetian) ; 1♀ (IZAY): Armenia, Syunik Prov., S environs of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=46.1932&amp;materialsCitation.latitude=39.0336" title="Search Plazi for locations around (long 46.1932/lat 39.0336)">Lichk</a>, 39.0336°N, 46.1932°E, 1700 m, 8.v.2016 (leg. M. Kalashian) ; Armenia, Syunik Prov., N <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=46.3746&amp;materialsCitation.latitude=38.9851" title="Search Plazi for locations around (long 46.3746/lat 38.9851)">Shvanidzor</a> vill., 38.9851°N, 46.3746°E, 1320 m 1♂ (IZAY): 9.vi–8.viii.2017 and 1♂, 2♀ (IZAY): 16.vi–26.viii.2018, soil traps (leg. T. Ghrejyan, G. Karagyan); 1♂, 1♀ (IZAY): Armenia, Syunik Prov., E environs of Kapan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=46.424&amp;materialsCitation.latitude=39.2221" title="Search Plazi for locations around (long 46.424/lat 39.2221)">Geghanush</a>, 39.2221°N, 46.4240°E, 1030 m, 8.iv–14.v.2008, soil traps (leg. M. Kalashian) ; 1♂ (IZAY): Armenia, Syunik Prov., N environs of Kapan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=46.388&amp;materialsCitation.latitude=39.2206" title="Search Plazi for locations around (long 46.388/lat 39.2206)">Ghatar</a> locality, 39.2206°N, 46.3880°E, 1120 m, 8.iv–14.v.2008, soil traps (leg. M. Kalashian) ; 1♀ (ZIN): Syunik Prov., fork to Lichk, 7.vi.1993 (leg. M. Kalashian) ; 1♀ (ZIN): Armenia, Syunik Prov., Lichk, 4.vi.1995 (leg. Kalashian) ; 1♀ (ZIN): Armenia, Syunik Prov., Gudemnis, 26.iv.1998 (leg. M. Kalashian) ; 1♀ (ZIN): Armenia, Syunik Prov., Agarak, 14.iv.–20.v.1995 (leg. M. Kalashian) ; Armenia, Vayotsdzor Prov., Areni, 24.v.1988 (leg. M. Kalashian) . Iran. 1♂ (HNHM): Iran, Azerbaijan, Makidi Arasbaran Wildlife Refuge, 1650–1800 m, 4–5.vii.1978 (leg. Martens, Pieper) .</p><p>Description. Male. Body robust, black, shiny (Fig. 12A). Measurements: Y = 1.56; PH w = 1.57; P w P l = 1.18; E l E w = 1.6–1.63; EH w = 2.1; EP w = 1.3; EP l = 2.25. Body length 11.5–16.6 mm, width 4.4–6.4 mm.</p><p>Head (Figs 12C–E). Anterior margin of epistoma slightly rounded. Head widest at eye level. Eyes moderately convex, circumocular impression narrow. Lateral margins of genae evenly rounded. Puncturation of head moderately coarse and sparse, interpuncture distance 1.5 as long as puncture diameter, genae densely and coarsely punctured. Ventral side of epicranium coarsely and densely punctured, gula with coarse transverse wrinkles. Antennae comparatively short, with only three apical antennomeres, extending beyond base of pronotum, reaching 1/5 of elytral length.</p><p>Prothorax (Figs 12C–F). Pronotum subcordate, widest at middle, transverse (Fig. 12D). Lateral margins of pronotum strongly rounded at middle and widely emarginated at basal half; anterior margin widely evenly emarginated; base straight. Antero-lateral corners not projected, obtuse, pointed; postero-lateral corners weakly straight, narrowly rounded at apex. Lateral margins not beaded; anterior margin shortly beaded only near corners; base beaded at middle. Disc of pronotum evenly weakly convex, not flattened along lateral margin, with two small oblique impressions near base. Puncturation of disc fine and sparse at middle and anterior half (puncture diameter 2–3 times as short as interpuncture distance), denser on sides and at basal quarter; punctures round (Figs 12D, E). Prosternum with moderately coarsely and sparse punctured and wrinkled (Fig. 12C). Prothoracic hypomera narrowly flattened along margin, with coarse transverse wrinkles only (Fig. 12F). Prosternal process smooth and shiny, sparsely punctured and pubescent, convex at apex, not beaded (Figs 12C, F).</p><p>Pterothorax (Figs 12A, C). Scutellar shield with triangle, with straight margins, finely and sparsely punctured. Elytra oval, widest at middle; punctures in striae sligly elongate, closely spaced; punctures in interstriae moderately coarse and sparse, little smaller than strial ones (Fig. 12A). Anterior half of mesoventrite pubescent, coarsely wrinkled (Fig. 12C). Mesepisterna, mesepimera and metepisterna coarsely and densely punctured by large foveae; metaventrite finely and sparsely punctured and pubescent with recumbent reddish setae.</p><p>Legs. Trochanters with dense brush of reddish setae and one long seta. Basal half of femora with the same dense brush on flex side (Fig. 12C); another surface of femora with moderately coarse, sparse puncturation and short setae. All tibiae straight. Pro- and mesotarsomeres 1–4 strongly widened, transverse, pro- and mesotarsomere 1 slightly wider than apex of protibia.</p><p>Abdomen (Figs 12C, G–K). Abdominal ventrites with moderately dense and fine puncturation and simple short pubescence; ventrite 5 with coarser punctures and long strong setae. Genitalia (Figs 12G–K). Apical part of basal piece of aedeagus smooth, without rugosity. Apical piece of aedeagus with weak median longitudinal keel; lateral margins almost straight, weakly narrowed to widely rounded apex. Apical third of basal piece smooth, without longitudinal rugosity.</p><p>Female (Figs 12B, C). Body robuster, dull, elytra matt. Eyes slightly smaller (see measurements). Antennae short, not reaching base of pronotum. Metatibiae straight. Measurements: Y = 1.5; PH w = 1.7; P w P l = 1.3; E l E w = 1.47; EH w = 2.28; EP w = 1.34; EP l = 2.57. Body length 13.5–17 mm, width 5.5–6.6 mm.</p><p>Etymology. The species is named in honour of my friend Dr Mark Kalashian (Yerevan, Armenia), a great connoisseur of the insect fauna of the Caucasus and a well-known specialist in Buprestidae .</p><p>Differential diagnosis. The new species is similar to E. quadricollis by the robust body, the form of pronotum with rounded lateral margins in apical 2/3 and widely emarginated in basal third, but differs from it by the following characters: lateral margins of pronotum more rounded and wholly not beaded, the lateral portions of base also not beaded (lateral margins lesser rounded and beaded at least near angles and base completely beaded in E. quadricollis); prothoracic hypomera flattened along outer margin and are with coarse longitudinal wrinkles (the hypomera with irregular fine wrinkles and punctures between them, not flattened along margin in E. quadricollis); the apical piece of the aedeagus is widely rounded at apex, and its dorsal surface is with longitudinal keel (the apical piece is quadrate or emarginated at middle in E. quadricollis and has the smooth surface). The coarsely longitudinally wrinkled prothoracic hypomera are present only in two species, i.e. E. huedepohli and E. kalashiani .</p><p>Notes. This species is interpreted by Russian entomologists as E. vicinus since the paper of Bogatchev (1938), who recorded it for Shikh-Yurdy Mt. (Ordubad District, Nakhichevan Autonomous Republic, Azerbaijan). Unfortunately, the Bogatchev’s specimen from this locality was not found in Russian collections and in the collection of the Institute of Zoology of the Azerbaijan Academy of Sciences. This mount is located just a few kilometers from Armenian localities of E. kalashiani . Iablokoff-Khnzorian (1957) listed both E. quardicollis and E. kalashiani as one species Hedyphanes quardicollis with the wide range from Talysh to Aras valley, but indicated characters for E. kalashiani in the key. Abdurakhmanov &amp; Medvedev (1994) and Abdurakhmanov &amp; Nabozhenko (2011) listed this new species as Probaticus vicinus, and Nabozhenko et al. (2021) indicated it as Euboeus (Pelorinus) sp.</p></div>	https://treatment.plazi.org/id/FA1487CEFF8D5F27D3B956D62E030B58	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nabozhenko, Maxim V.	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF8F5F21D3B9522329610B34.text	FA1487CEFF8F5F21D3B9522329610B34.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euboeus (Pelorinus) kasatkini Nabozhenko 2022	<div><p>Euboeus (Pelorinus) kasatkini sp. nov.</p><p>(Fig. 13)</p><p>Type material. Holotype, ♂ and two paratypes, ♂ and ♀ (ZIN): Iran, Alborz Prov., Central Elburs Mts., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=51.32959&amp;materialsCitation.latitude=36.097744" title="Search Plazi for locations around (long 51.32959/lat 36.097744)">Gachsar</a> env., H= 2600–2720 m, 36°05ʹ51.88ʺN, 51°19ʹ46.53ʺE, 30.v.2016 (leg. D.G. Kasatkin) . Paratypes. 1♂, 1♀ (HNHM): N Iran, 8 km W Gachsar, 6–8.vii.1977, “Loc. No. 396 Exped. Nat. Mus. Praha”, “ Probaticus huedepohli Kulzer, 1964 det. Z. Kaszab, 1983” ; 1♀ (HNHM): Iran, Tehran Prov, Elburs-Gebirge, N Tehran, Shenshak, 2650–2900 m, 17–25.vi.1978 (leg. Martens and Pleper) ; 2♀ (PCMN): Iran, Tehran Prov., Alborz Mts., Shemshak vill., road to Dizin, 3000 m, 1.vi.2014 (D. Murastyi) ;</p><p>Description. Male. Body slender, black, pronotum and head dorsally shiny, elytra lesser shiny (Fig. 13A). Measurements: Y = 1.44; PH w = 1.5; P w P l = 1.2; E l E w = 1.54; EH w = 1.86; EP w = 1.24; EP l = 2.38. Body length 12 mm, width 4.4 mm.</p><p>Head (Figs 13C, E, F). Anterior margin of epistoma slightly rounded and emarginated near angles. Head widest at eye level. Eyes moderately convex, circumocular impression narrow. Lateral margins of genae angularly rounded. Puncturation of head fine and sparse, interpuncture distance 1.5–2 times as long as puncture diameter. Ventral side of epicranium coarsely and densely punctured. Antennae comparatively short, with only three apical antennomeres, extending beyond base of pronotum, reaching less than 1/4 of elytral length.</p><p>Prothorax (Figs 13C, E–G). Pronotum widest at middle, transverse (Fig. 13E). Lateral margins of pronotum weakly evenly rounded; anterior margin weakly rounded at middle and slightly widely emarginated near corners; base weakly rounded at middle, slightly bisinuate. Antero-lateral corners not projected, obtuse, narrowly rounded at apex; postero-lateral corners weakly obtuse, pointed at apex. Lateral margins beaded; anterior margin beaded only laterally (Figs 13E, F). Disc of pronotum evenly weakly convex, not flattened along lateral margin, with weak oval impression on each lateral side and two small impressions at base. Puncturation of disc fine and sparse, puncture diameter 3–5 times as short as interpuncture distance; punctures round; punctures in impressions slightly larger (Figs 13E, F). Prosternum with moderate and very sparse simple puncturation (Fig. 13C). Prothoracic hypomera very narrowly weakly flattened along margin, with coarse and dense punctures among wrinkles; surface without microgranulation (Fig. 13G). Prosternal process smooth and shiny, with two terminal lines of punctures, slightly convex at apex, not beaded (Figs 13C, G).</p><p>Pterothorax (Figs 13A, C). Scutellar shield with straight margins, coarsely sparsely punctured. Elytra moderately elongate, widest at middle; punctures in striae fine, slightly elongate, widely spaced; punctures in interstriae fine and sparse, much smaller than strial ones (Fig. 13A). Anterior half of mesoventrite pubescent, coarsely and densely wrinkled. Mesepisterna, mesepimera and metepisterna with coarse and dense puncturation of large foveae; metaventrite and intercoxal part of mesoventrite finely and sparsely punctured and covered with short setae (Fig. 13C).</p><p>Legs. Trochanters with brush of short dark-reddish setae and one long seta (Fig. 13C). Basal third of femora with the same brush on flex side (Fig. 13C); another surface of femora with coarse puncturation and short setae. Pro- and mesotibiae straight, metatibiae very weakly bent (Fig. 13B). Pro- and mesotarsomeres 1–4 strongly widened, trasverse, pro- and mesotarsomere 1 wider than apex of protibia.</p><p>Abdomen (Figs 13C, H–M). Abdominal ventrites with fine and sparse puncturation and short setation, ventrite 5 with pubescence of long strong setae. Genitalia (Figs 13H–M). Apical part of the basal piece of aedeagus with longitudinal rugosity (Fig. 13M). Lateral sides of apical piece of aedeagus straight in apical two thirds (Fig. 13M).</p><p>Female (Fig. 13D). Body much more robust, elytra duller in specimen from the type locality and the same shiny as male in specimens from Shemshak). Antennae short, not reaching base of pronotum. Pronotum widest at middle or sometimes behind middle. Metatibiae straight. Measurements: Y = 1.44; PH w = 1.65; P w P l = 1.24–26; E l E w = 1.46; EH w = 2.13; EP w = 1.29; EP l = 2.41. Body length 11–16 mm, width 4–6.5 mm.</p><p>Etymology. The new species is named in honour of my friend, the collector of many specimens of Euboeus and other Tenebrionidae from Iran, Dr Denis Kasatkin (Rostov Branch of the Federal State Budgetary Institution All-Russian Plant Quarantine Centre, Rostov-on-Don, Russia).</p><p>Differential diagnosis. This species is very close to E. huedepohli from Demavend by the beaded lateral margins of pronotum (Figs 11E, F, 13E, F) and longitudinal rugosity in apical part of the basal piece of the aedeagus (Figs 11M, 13M), but differs by the following characters: body dorsally lesser shiny, pronotum without lacquer gloss; prothoracic hypomera with coarse puncturation among smooth wrinkles (Fig. 13G) (coarse longitudinal wrinkles in E. huedepohli (Fig. 11G)); lateral margin of prothoracic hypomera very narrowly weakly flattened (Fig. 13G) (widely clearly flattened in E. huedepohli (Fig. 11G)); metatibiae very weakly bent (Fig. 13B) (strongly bent in E. huedepohli (Fig. 11C)); lateral margins of the apical piece of the aedeagus straight in apical half (Fig. 13M) (weakly rounded in E. huedepohli (Fig. 11M)).</p></div>	https://treatment.plazi.org/id/FA1487CEFF8F5F21D3B9522329610B34	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nabozhenko, Maxim V.	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF895F23D3B9528F2A0F091A.text	FA1487CEFF895F23D3B9528F2A0F091A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euboeus (Pelorinus) krivokhatskyi Nabozhenko 2022	<div><p>Euboeus (Pelorinus) krivokhatskyi sp. nov.</p><p>(Fig 14)</p><p>Type material. Holotype, ♂ (HNHM): Iran, Gilan Prov.: “ Kelishum [ Klishom] 2000 m südl. Langerud [Langarud] 4.viii.1967 ”, “ Gilan (Iran) Heinz leg.”, “ Probaticus sp. det. Kaszab ” . Paratype, ♀ (SMNS): Iran, Gilan Prov., 20 km SE Deilaman, 1900 m, 22.04.2003 (leg. G. Sama) ; 1♂ (NMPC): Iran, Koedestan [Kurdestan] Prov., Pass 26 km SE Saqqez, 17.iv.2002 (leg. S. Kadlec) ; 1♀ (NME): Iran, Luristan [Lorestan] Prov., pass 2300 m W Nour-Abad (SE Nahavand), oberh. Grotte v. Gamasiab [upper grotto of Gamasiab], 10.iv.2014 (leg. Heinz) .</p><p>Description. Male. Body slender, black, shiny (Fig. 14A), but weakly shiny in the Lorestan Population. Measurements: Y = 1.59; PH w = 1.55; P w P l = 1.3; E l E w = 1.6; EH w = 1.83; EP w = 1.17; EP l = 2.48. Body length 12.5 mm, width 4.5 mm.</p><p>Head (Fig. 14B, D, E). Anterior margin of epistoma slightly rounded and emarginated near angles. Head widest at eye level. Eyes strongly convex, angulate in dorsal view, circumocular impression deep, moderately wide (Figs 14B, D, E). Lateral margins of genae angularly rounded. Puncturation of head fine and sparse, interpuncture distance 1.5–2 times as long as puncture diameter; punctures coarser on genae. Ventral side of epicranium coarsely and densely punctured around submentum and coarse and sparse on temples. Antennae comparatively short, with only three apical antennomeres, extending beyond base of pronotum, reaching less than 1/4 of elytral length.</p><p>Prothorax (Figs 14D–F). Pronotum widest ahead of middle, transverse (Fig. 14D). Lateral margins of pronotum moderately rounded, slightly emarginated near base; anterior margin weakly widely emarginated; base straight. Antero-lateral corners not projected, obtuse, narrowly rounded at apex; postero-lateral corners weakly obtuse, narrowly rounded at apex. Lateral margins not beaded; anterior margin beaded only laterally (Figs 14D, E). Disc of pronotum evenly weakly convex, not flattened along lateral margin, without impressions. Puncturation of disc fine and sparse, punctures small, but surface around punctures impressed, puncture diameter 3 times as short as interpuncture distance; punctures round. Prosternum with coarse and sparse simple puncturation (Figs 14B, F). Prothoracic hypomera not flattened flattened along margin, with coarse, sparse, large, round punctures on smooth surface at outer half and dense and coarse merged punctures on inner half; surface without microgranulation (Fig. 14F). Prosternal process shiny, sparsely punctured, slightly convex at apex, not beaded (Figs 14B, F).</p><p>Pterothorax (Figs 14A, B). Scutellar shield with very weakly rounded margins, coarsely sparsely punctured. Elytra shiny, moderately elongate, widest at middle; punctures in striae fine, round, widely spaced; punctures in interstriae fine and sparse, much smaller than strial ones (Fig. 14A). Anterior half of mesoventrite pubescent, coarsely and densely wrinkled. Mesepisterna, mesepimera and metepisterna with coarse and dense puncturation of large foveae; metaventrite and intercoxal part of mesoventrite with moderately coarse and moderately dense punctures; metaventrite covered with very short setae (Fig. 14B).</p><p>Legs. Trochanters with brush of short dark-reddish setae and one long seta (Fig. 14B). Basal third of meso- and metafemora with the same brush on flex side, the same part of profemora without this brush (Fig. 14B). Protibiae weakly S-shaped, mesotibiae weakly curved, metatibiae weakly bent inward. Pro- and mesotarsomeres 1–4 strongly widened, trasverse, but pro- and mesotarsomere 1 not wider than apex of protibia.</p><p>Abdomen (Figs 14B, G–K). Abdominal ventrites with the same puncturation as on metaventrite and very short setation (not pubescent) (Fig. 14B); Ventrite 5 with several sparse long setae at apex. Genitalia (Figs14G–K). Apical part of the basal piece of aedeagus with longitudinal rugosity (Fig. 14K). Apical piece subtriangle, narrowly rounded at apex, dorsal surface with longitudinal small median groove (Fig. 14K).</p><p>Female (Fig. 14C). Body much robuster and dull. Eyes smaller (Y=1.36 vs. 1.59 in male). Antennae short, not reaching base of pronotum. Lateral margins of pronotum beaded at basal half. Postero-lateral corners of pronotum right. Metatibiae straight. Measurements: Y = 1.36; PH w = 1.67; P w P l = 1.29; E l E w = 1.37; EH w = 2.2; EP w = 1.32; EP l = 2.34. Body length 17 mm, width 7 mm.</p><p>Etymology. The species is named in memory of my colleague Viktor Anatolievich Krivokhatsky (1954–2021), the world specialist on Neuroptera, Myrmeleontidae (Volkovitsh et al. 2021, Ovtshinnikova et al. 2021).</p><p>Differential diagnosis. The new species is close to E. merkli by having not beaded lateral margins of pronotum (Figs 14E, 15E) and smooth punctured sculpture of the prothoracic hypomera (Figs 14F, 15F), but is different from other congeners of the huedepohli species-group by strongly convex (lateral margin angulate in dorsal view) eyes (Fig. 14D) and smooth prothoracic hypomera with sparse coarse punctures (except for E. merkli, which has the similar sculpture of hypomera). Euboeus krivokhatskyi has a similar shape of the pronotum as E. grimmi, not beaded male pronotum as is in E. merkli and the apical part of the basal piece of the aedeagus with longitudinal rugosity as in E. huedepohli and E. kasatkini . The new species additionally differs from:</p><p>– E. huedepohli and E. kasatkini by not beaded laterally male pronotum and partly beaded female pronotum (Figs 14D, E) (lateral margins of the pronotum are beaded in both comparable species (Figs 11E, F, 13E, F)), and the absence of the dense brush of setae at the base of the flex side of profemora (Fig. 14B), the shape of the apical piece of aedeagus;</p><p>– E. grimmi by shiny body dorsally, at least in the Elburs population (Fig. 14A) (weakly shiny in E. grimmi (Fig. 9A)), not beaded laterally male pronotum (Figs 14D, E) (lateral margins of the pronotum are beaded in E. grimmi (Fig. 9D, E)), the absence of the dense brush of setae at the base of the flex side of profemora (Fig. 14B), the apical part of the basal piece of the aedeagus with longitudinal rugosity (Fig. 14K) (smooth in E. grimmi (Fig. 9K)), the apical piece of the aedeagus with longitudinal median groove (Fig. 14K) (median keel in E. grimmi (Fig. 9K));</p><p>– E. merkli by the shiny male body at least in the Elburs population (14A) (weakly shiny in E. merkli (15A)), wider and the widest ahead of the middle pronotum with rounded lateral margins (Fig. 14D) ( E. merkli has narrower, widest at middle pronotum, with very weakly rounded lateral margins (Fig. 15D)), the presence of dense brush of setae at the base of the flex side of meso- and metafemora (Fig. 14B) (absence of the femoral brush in E. merkli (Fig. 15B)) and almost bare abdominal ventrites (Fig. 14B) (pubescent in E. merkli (Fig. 15B)).</p></div>	https://treatment.plazi.org/id/FA1487CEFF895F23D3B9528F2A0F091A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nabozhenko, Maxim V.	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF8B5F2DD3B950E22A760918.text	FA1487CEFF8B5F2DD3B950E22A760918.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euboeus (Pelorinus) merkli Nabozhenko 2022	<div><p>Euboeus (Pelorinus) merkli sp. nov.</p><p>(Fig. 15)</p><p>Type material. Holotype, ♂ (HNHM) and paratypes, 3♂ (two in SMS, one in HNHM), 1♀ (SMS): Iran, Mazandaran Prov., Alborz Mts., Damavand Mt., Reyneh [Rineh], 35°52.412ʹ N, 52°06.096ʹ, 2841 m, 14.v.2008, from beneath stone (leg. T. Hácz, K. Székely, K. Vig) . Paratypes: 1♂ (HNHM): Iran, Abgarm ( Damavand), 2100–2400 m, 5.v.1970 (Wittmer, v. Bothmer) , “ zoroaster Sdl det. Kaszab”; 1♀ (HNHM): Iran, Tehran Prov. [Mazandaran Prov.], Elburz Mts., 10 km W Reine [Rineh], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=52.04&amp;materialsCitation.latitude=35.889446" title="Search Plazi for locations around (long 52.04/lat 35.889446)">Kuh-e Demavend</a>, 35°53ʹ22ʺN, 52°02ʹ24ʺE, 5.v.1998 (Gy. Fábian, K. Székely) ; 1♂ (HNHM): Iran, Mazandaran Prov., Demavend Mt., 3000 m, 4.vi.1999 (leg. T. Hácz, G. Kőszegi, K. Gaskó) ; 2♀ (HNHM): Iran, Mazandaran Prov., Kühha-ye Alborz, Küh-e Damavand, 3500 m, 30.vi.–2.vii.2006 (leg. G. Rozner) ; 1♂, 2♀ (HNHM): Iran, Mazandaran Prov., Alborz <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=52.102776&amp;materialsCitation.latitude=35.874443" title="Search Plazi for locations around (long 52.102776/lat 35.874443)">Mts.</a>, 15 km E of Lar-e Polur, 35°52ʹ28ʺN, 52°06ʹ10ʺE, 2580 m, 12–13.v.2001 (leg. Gy. Fábian, K. Vig) .</p><p>Description. Male. Body narrow, slender, black, moderately shiny (Fig. 15A). Measurements: Y = 1.55; PH w = 1.42; P w P l = 1.08; E l E w = 1.57; EH w = 2; EP w = 1.4; EP l = 2.46. Body length 12–14 mm, width 4–5 mm.</p><p>Head (Figs 15B, D, E). Anterior margin of epistoma slightly rounded. Head widest at eye level. Eyes moderately convex, large in dorsal view, circumocular impression narrow (Fig. 12D). Lateral margins of genae evenly rounded. Puncturation of head fine and sparse, interpuncture distance 2–3 times as long as puncture diameter. Ventral side of epicranium coarsely and densely punctured by merged punctures. Antennae comparatively short, with only three apical antennomeres, extending beyond base of pronotum, reaching 1/5 of elytral length.</p><p>Prothorax (Figs 15D–F). Pronotum subquadrate, widest at middle, very weakly transverse, almost with subequal width and length (Fig. 15D). Lateral margins of pronotum very weakly evenly rounded; anterior margin slightly rounded, almost straight; base weakly rounded. Antero-lateral corners not projected, obtuse, narrowly rounded at apex; postero-lateral corners weakly obtuse, pointed at apex. Lateral margins not beaded; anterior margin unclearly beaded only near corners (Fig. 15E). Disc of pronotum evenly strongly convex (only area near postero-lateral corners slightly flattened), not flattened along lateral margin, with basal impressions: oval transverse at middle and two obliques laterally. Puncturation of disc fine and sparse, puncture diameter 4–5 times as short as interpuncture distance in middle and 2–3 times on lateral sides; punctures round (Fig. 15D). Prosternum with coarse and very sparse simple puncturation (Fig. 15B). Prothoracic hypomera not flattened along margin, with coarse, sparse, large, round punctures along margin and near procoxae and dense and coarse merged punctures at middle; surface without microgranulation (Fig. 15F). Prosternal process shiny, moderately punctured, slightly convex at apex, not beaded, pubescent by recumbent setae (Figs 15B, F).</p><p>Pterothorax (Figs 15A, B). Scutellar shield with straight margins, finely and densely punctured. Elytra slightly shiny than pronotum, elongate, widest at middle; punctures in striae fine, round, widely spaced; punctures in interstriae fine and sparse, much smaller than strial ones (Fig. 15A). Anterior half of mesoventrite densely pubescent, coarsely and densely wrinkled. Mesepisterna, mesepimera and metepisterna with coarse and dense puncturation of moderate in size punctures and short setae; metaventrite with moderately coarse and moderately dense punctures; intercoxal part of mesoventrite finely and sparsely punctured and sparsely pubescent; metaventrite pubescent with recumbent comparatively long setae (Fig. 15B).</p><p>Legs. Trochanters with brush of short dark-reddish setae and one long seta (Fig. 15B). Basal third of meso- and metafemora without the same brush on flex side, but with simple recumbent setation (Fig. 15B). Tibiae straight. Pro- and mesotarsomeres 1–4 strongly widened, trasverse, subequal to width of apex of protibia.</p><p>Abdomen (Figs 15B, G–K). Abdominal ventrites 1–3 with the same puncturation as on metaventrite, ventrites 4–5 much sparser and finer punctured. All ventrites pubescent with long recumbent setae (Fig. 15B); ventrite 5 with several sparse, longer and stronger setae at apex. Genitalia (Figs 15G–K). Apical part of the basal piece of aedeagus smooth, without longitudinal rugosity (Fig. 15K). Apical piece narrowly rounded at apex, dorsal surface with longitudinal small median groove (Fig. 15K).</p><p>Female (Fig. 15C). Body robuster, elytra dull. Eyes smaller (see mesuarements). Antennae short, not reaching base of pronotum. Lateral margins of pronotum little more narrowed to base. Measurements: Y = 1.43; PH w = 1.42; P w P l = 1.1–1.12; E l E w = 1.58; EH w = 2; EP w = 1.4; EP l = 2.64–2.65. Body length 12–16.5 mm, width 4.5–6 mm.</p><p>Etymology. The new species is named in memory of Ottó Merkl (1957–2021), my friend, colleague and coauthor of several papers, the famous Hungarian entomologist (Schawaller 2021, Szél et al. 2021).</p><p>Differential diagnosis. This new species is similar to E. krivokhatskyi by the surface of prothoracic hypomera (Figs 11F, 12F) and not beaded lateral margins of the pronotum (Figs 11E, 12E). See differences in the diagnosis of E. krivokhatskyi .</p></div>	https://treatment.plazi.org/id/FA1487CEFF8B5F2DD3B950E22A760918	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nabozhenko, Maxim V.	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF855F2ED3B950E329770EC0.text	FA1487CEFF855F2ED3B950E329770EC0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euboeus (Pelorinus) quadricollis (Menetries 1832)	<div><p>Euboeus (Pelorinus) quadricollis (Ménétriés, 1832)</p><p>(Figs 16, 17)</p><p>Hedyphanes quadraticollis Ménétriés, 1832: 196 .</p><p>Type material. Holotype, ♂ (ZIN): “Mont. Taliish”, “ quadraticollis Menet. Zouvant ”, bronze square, “ Holotypus Helops quadraticollis Ménétriés, 1832 ” (Nabozhenko’s label) (Fig. 13B).</p><p>Material. Azerbaijan. 1♂, 4♀ (ZIN): Azerbaijan, 25 km below Yardýmlý, meadows, 2–3.vi.2008 (leg. D. Kasatkin) . Iran. 1♂ (SMNH): Iran, Ardabil Prov., Nir env., 27.iv.2006 (leg. M. Machálik) ; 1♀ (SMNH); 1♂ (NMPC): Iran, Azerbaijan-e Sharqi Prov., Sefide Khan, 25 km S Tabriz, 7.v.2002 (S. Kadlec).</p><p>Note. Full bibliography for this species covering complicated historical and nomenclatural issues with application of two names quadraticollis / quadricollis for this species, discussion about the author of the taxon, as well as short redescription and figures were published by Abdurakhmanov &amp; Nabozhenko (2011).</p><p>Distribution. Azerbaijan (Talysh Mts.: Yardýmlý and Lerik districts), North Iran (Ardabil and Azerbaijan-e Sharqi provinces). New record for Iran.</p></div>	https://treatment.plazi.org/id/FA1487CEFF855F2ED3B950E329770EC0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nabozhenko, Maxim V.	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF805F28D3B956D6291F084F.text	FA1487CEFF805F28D3B956D6291F084F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euboeus (Pelorinus) zubovi Nabozhenko 2022	<div><p>Euboeus (Pelorinus) zubovi sp. nov.</p><p>(Fig. 18)</p><p>Type material. Holotype, ♂ (ZIN): Iran, Kurdistan Prov., between <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=48.80361&amp;materialsCitation.latitude=36.776943" title="Search Plazi for locations around (long 48.80361/lat 36.776943)">Saqqez</a> and Baneh, 36°46ʹ37ʺ N, 48°48ʹ13ʺE (leg. A. Zubov).</p><p>Description. Male. Body slender, black, moderately shiny. Measurements: Y = 1.56; PH w = 1.35; P w P l = 1.15; E l E w = 1.7; EH w = 1.8; EP w = 1.34; EP l = 2.7. Body length 11.5 mm, width 3.9 mm.</p><p>Head (Figs 18A). Anterior margin of epistoma straght. Head widest at eye level. Eyes moderately convex, large in dorsal view, circumocular impression wide (Fig. 18D). Lateral margins of genae angularly rounded. Puncturation of head moderately coarse and sparse, interpuncture distance 1.5–2 times as long as puncture diameter. Head pubescent with sparse, reddish, moderately long, recumbent setae (Figs 18D, E). Ventral side of epicranium coarsely and densely punctured by raduliform punctures, gula almost smooth (Fig. 18B). Antennae comparatively short, with only two apical antennomeres, extending beyond base of pronotum, reaching 1/6 of elytral length.</p><p>Prothorax (Figs 18B, D–F). Pronotum weakly cordate, widest at middle, weakly transverse (Fig. 18D). Lateral margins of pronotum moderately rounded, slightly emarginated near antero-lateral corners and stronger emarginated near postero-lateral corners; anterior margin slightly rounded at middle and widely emarginated laterally; base weakly bisinuate. Antero-lateral corners projected, acute and pointed at apex; postero-lateral corners straight, pointed at apex. Lateral margins beaded (Figs 18D, E); anterior margin with interrupted bead at middle. Disc of pronotum evenly strongly convex, not flattened along lateral margin, only lateral sides more gentle. Puncturation of disc fine and sparse, puncture diameter near 3–4 times as short as interpuncture distance in middle and 3 times on lateral sides; punctures round (Fig. 18D). Prosternum with coarse, sparse, short transverse wrinkles and pubescent with long, recumbent, sparse setae (Fig. 18B). Prothoracic hypomera not flattened along margin, with smooth, weak, sparse wrinkles and distinct sparse granules bearing long seta (Fig. 18F). Prosternal process shiny, moderately punctured on sides, densely pubescent with subrecumbent reddish setae slightly convex at apex, beaded (Figs 18B, F).</p><p>Pterothorax (Figs 18A, B). Scutellar shield triangle, with straight margins, coarsely and densely punctured. Elytra elongate-oval, widest at middle; punctures in striae fine, slightly elongate, widely spaced; punctures in interstriae fine and sparse, with almost the same size than strial ones, but deeper (Fig. 18A). Anterior half of mesoventrite densely pubescent, coarsely and densely wrinkled. Mesepisterna, mesepimera and metepisterna with coarse and dense puncturation and long recumbent setae; intercoxal part of mesoventrite finely and sparsely punctured and sparsely pubescent; metaventrite with moderately coarse and moderately dense punctures, pubescent with long recumbent setae (Fig.18B).</p><p>Legs. Trochanters with brush of short dark-reddish setae and one long seta (Fig. 18B). Basal third of meso- and metafemora without the same brush on flex side (Fig. 18B). Tibiae straight. Pro- and mesotarsomeres 1–4 strongly widened, trasverse, subequal to width of apex of protibia.</p><p>Abdomen (Figs 18B, C, G–J). Abdominal ventrites with moderately sparse, fine raduliform puncturation and pubescence of long, recumbent, reddish setae. Genitalia (Figs 18C, G–J). Apical part of the basal piece of aedeagus almost smooth, with only pair small areas of fine longitudinal rugosity (Fig. 18J). Apical piece strongly narrower from base to apex, widely rounded at apex, dorsal surface without median groove or keel (Fig. 18J).</p><p>Female unknown.</p><p>Etymology. The new species is named in honour of Andrey Zubov (Moscow, Russia), my colleague and excellent insect collector.</p><p>Differential diagnosis. The new species differs from all representatives of the huedepohli species-group by the following characters: head pubescent with recumbent setae; pronotum with projected, acute, pointed at apex anterolateral corners; prothoracic hypomera with sparse distinct granules; the apical piece of the aedeagus sub-triangle, strongly narrowed from base to apex.</p></div>	https://treatment.plazi.org/id/FA1487CEFF805F28D3B956D6291F084F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nabozhenko, Maxim V.	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF835F2BD3B956D62FE10DB1.text	FA1487CEFF835F2BD3B956D62FE10DB1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euboeus (Pelorinus) chorasanicus (G. Medvedev 1976)	<div><p>Euboeus (Pelorinus) chorasanicus (G. Medvedev, 1976)</p><p>(Figs 19, 20)</p><p>Probaticus (Pelorinus) chorasanicus Medvedev, 1976: 893;</p><p>Euboeus (Pelorinus) chorasanicus: Nabozhenko, 2019 b: 1033, fig. 21.</p><p>Type material. Holotype, ♂ (ZIN): “В. Персiя Имам-Гуляръ—Кучанъ 16.III.98 Зарудный” (Cyrillic, old Russian) [E Persia, Imam-Gulyar—Quchan, leg. Zarudny], “ Holotypus Probathicus (Pelorinus) chorasanicus G. Medvedev, 1977 ” (Fig. 16D) . Paratype, ♀ (ZIN): “ Хорасан М. Кучан—Имам Гуляр 16.III.98 Н. Зарудный” (Cyrillic, Russian) [Khorasan, Kuchan—Imam-Gulyar, leg. N. Zarudny] (Fig. 15D). Red labels with “ Holotypus ” and “ Paratypus ” were pinned by a laboratory assistant (A.K. Chistaykova) with erroneous year and the name of the genus .</p><p>Material. 1♂, 1♀ (SMNS): Iran, North Khorasan Prov., Asadij (20 km SE Bojnurd), 1700 m, 4.04.2001 (leg. G. Sama) ; 1♂ (ZIN): Iran, North Khorassan Prov., Bojnurd area, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=57.216667&amp;materialsCitation.latitude=37.283333" title="Search Plazi for locations around (long 57.216667/lat 37.283333)">Gerivan</a>, 37°17ʹN, 57°13ʹE, 1800 m, 12.v.2009 (leg. A. Klimenko) ; 1♀ (ZIN): the same label, but 8.vi.2009; 1♀ (NMPC): NE Iran, Razavi Khorasan Province, Kuh-e- Binalud, 20 km NE Nishapur, 13–15.vi.1977, 2500 m, Loc. No 366, (leg. Exped. Nat. Mus. Praha) .</p><p>Distribution. Iran (North Khorasan and Razavi Khorasan provinces). This species is terrestrial and inhabits xerophytic areas.</p><p>Variability. The female from Binalud Mt. differs by the weakly shiny body and dull elytra.</p></div>	https://treatment.plazi.org/id/FA1487CEFF835F2BD3B956D62FE10DB1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nabozhenko, Maxim V.	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFF835F14D3B951CF2EB9088E.text	FA1487CEFF835F14D3B951CF2EB9088E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euboeus Boieldieu 1865	<div><p>Key to species of the genus Euboeus of the Caucasus, Iran and Turkmenistan, on males</p><p>1 Pronotum and head dark-bluish or dark-violet, elytra black (Figs 19A, 20A, B)........................ E. chorasanicus</p><p>- Body dorsally completely black or sometimes black-brown................................................... 2</p><p>2 Puncturation of head and pronotum coarse and dense, punctures subequal or larger than interpuncture distance (sometimes punctures coarse but slightly sparser at middle of pronotum) (Figs 1A, 2E, F, 4F, G, 5D, E, 6A, 7F, G). Male body dull dorsally (Figs 1A, 2A, 3A, 4A, 5A, 7A) or rarely shiny (Fig. 7A)...................................................... 3</p><p>- Puncturation of head and pronotum fine and sparse, puncture diameter much lesser than interpuncture distance, at least at middle and anterior third (Figs 9D, E, 10A, B, 11E, F, 12D, E, 13E, F, 14D, E, 15D, E, 17D, E). Male body more or less shiny dorsally (8A, 9A, 10A, 11A, 12A, 13A, 14A, 15A, 17A, 18A)................................................. 7</p><p>3 Pronotum with very coarse and dense puncturation of longitudinal elongated punctures, especially on lateral sides of disc; Median smooth line of pronotum elevated near base and separated from other surface by impression with very dense merged punctures (Figs 1A, 2E, F).................................................................. E. corrugosus</p><p>- Pronotum with coarse and dense puncturation of simple round, not merged punctures; median line not elevated near base or absent (Figs 4F, G, 5D, E, 6A, 7F, G)..................................................................... 4</p><p>4 Head densely pubescent with recumbent reddish setae (Figs 6A–D). Pro- and mesotarsi slightly widened, much narrower than apex of protibia (Fig. 6A)....................................................................... E. vicinus</p><p>- Head bare, only with very short simple seta in each puncture (Figs 4G, 5E, 7G). Pro- and mesotarsi stronger widened, equal or wider than width of apex of protibia (Figs 4A, 5A, 7A)....................................................... 5</p><p>5 Metaventrite and abdominal ventrites pubescent with long recumbent hairs (Fig. 4C, D). Pronotum very narrow P w P l = 1.13 (Fig. 4F)................................................................................. E. gorganicus</p><p>- Metaventrite and abdominal ventrites bare, covered with simple very short setae (Figs 5B, 7C, D). Pronotum wider, P w P l = 1.16–1.28 (Figs 5D, 7F)............................................................................... 6</p><p>6 Pronotum much wider (P w P l = 1.28), with lateral strongly emarginated at basal quarter (Fig. 5D). Abdominal ventrite 5 with double puncturation: large sparse punctures among fine dense puncturation (Fig. 5B). Pro- and mesotarsi more elongate, pro- and mesotarsomere 1 longitudinal, pro- and mesotarsomere 2 with subequal length and width (Fig. 5A)..... E. prometheus</p><p>- Pronotum narrower (P w P l = 1.16), with lateral margins straight or weakly rounded at base (Fig. 7D). Abdominal ventrite 5 with simple fine and sparse puncturation (Fig 7B). Pro- and mesotarsi wider, pro- and mesotarsomeres 1 and 2 transverse (Fig. 7A).......................................................................................... E. zoroaster</p><p>7 Head pubescent with long recumbent setae (Fig. 18E). Pronotum with projected, acute, pointed at apex antero-lateral corners (Fig. 18D). Prothoracic hypomera with sparse distinct granules (Fig. 18F)................................. E. zubovi</p><p>- Head bare (Figs 8F, 9E, 11F, 12E, 13F, 14E, 15E, 17E). Antero-lateral angles of pronotum not projected, right or obtuse (Figs 8E, 9D, 11E, 12D, 13E, 14D, 15D, 17D). Prothoracic hypomera with wrinkles and punctures, separately or together, but without sparse granules (Figs 8G, 9F, 11G, 12F, 13G, 14F, 15F, 17F)................................................ 8</p><p>8 Lateral margins of pronotum not completely beaded, bead widely interrupted at middle (Figs 17D, E). Apical piece of aedeagus subquadrate at apex or apex strongly emarginated at middle (Fig. 17K)............................... E. quadricollis</p><p>- Lateral margins of pronotum completely beaded (Figs 9E, F, 11E, F, 13E, F) or completely not beaded (Figs 14D, E, 15D, E). Apical piece of aedeagus round at apex (Figs 8D, 9K, 11M, 12K, 13M, 14K, 15K)................................. 9</p><p>9 Lateral margins of pronotum completely beaded (Figs 9E, F, 11E, F, 13E, F)..................................... 10</p><p>- Lateral margins of pronotum not beaded (Figs 8E, F, 12D, E, 14D, E, 15D, E).................................... 12</p><p>10 Pronotum widest ahead of middle (Fig. 9D). Prothoracic hypomera not flattened along margin, with small confused microwrinkles with poorly distinguished sparse smoothed puncturation against their background; wrinkles with microgranulation at anterior third (Fig. 9F). Apical piece of aedeagus with longitudinal median keel (Fig. 9K). Apical part of the basal piece of aedeagus smooth, without longitudinal rugosity (Fig. 9K).............................................. E. grimmi</p><p>- Pronotum widest at middle (Figs 11E, 13E). Prothoracic hypomera narrowly (Fig. 13G) or widely (Fig. 11G) flattened along margin, coarsely and densely punctured (Fig. 13G) or coarsely longitudinally wrinkled (Fig. 11G), without microgranulation. Apical piece of aedeagus with longitudinal median groove or wide impression (Figs 11M, 13M)..................... 11</p><p>11 ody dorsally strongly shiny, especially pronotum with lacquer gloss (Figs 10B, 11E, F). Prothoracic hypomera with coarse longitudinal wrinkles; lateral margin of prothoracic hypomera clearly widely flattened (Fig. 11G). Metatibiae strongly bent (Fig. 11C). Lateral margins of apical piece of aedeagus weakly rounded (Fig. 11M)........................... E. huedepohli</p><p>- Body dorsally lesser shiny, pronotum without lacquer gloss (Figs 13E, F). Prothoracic hypomera with coarse puncturation and smooth wrinkles at middle; lateral margin of prothoracic hypomera very narrowly weakly flattened (Fig. 13G). Metatibiae very weakly bent (Fig. 13B). Lateral margins of apical piece of aedeagus straight in apical half (Fig. 13M)......... E. kasatkini</p><p>12 Prothoracic hypomera with large and coarse longitudinal smooth wrinkles (Fig. 12F). Pronotum with strongly rounded lateral margins at middle (Fig. 12D)................................................................. E. kalashiani</p><p>- Prothoracic hypomera punctured or with punctation and microwrinkles (Figs 8G, 14F, 15F). Lateral margins of pronotum weakly rounded..................................................................................... 13</p><p>13 Prothoracic hypomera with dense uneven small uneven wrinkles and small sparse punctures on their background (Fig. 8G)............................................................................................ E. arzanovi</p><p>- Prothoracic hypomera with large coarse foveolate punctures, without microwrinkles (Figs 14F, 15F).................. 14</p><p>14 Eyes strongly convex (lateral margin angulate in dorsal view) (Figs 14B, D). Pronotum wider (P w P l = 1.3), widest ahead of the middle, with moderately rounded lateral margins (Fig. 14D). Base of flex side of meso- and metafemora with dense brush of setae (Fig. 14B). Abdominal ventrites bare, with only simple very short setae (Fig. 14B)................ E. krivokhatskyi</p><p>- Eyes moderately convex (lateral margin rounded in dorsal view) (Figs 15B, D). Pronotum narrow, almost with subequal width and length (P w P l = 1.08), widest at middle, with very weakly rounded lateral margins (Fig. 15D). Base of flex side of meso- and metafemora with simple recumbent setae, without separate dense brush (Fig. 15B).......................... E. merkli</p></div>	https://treatment.plazi.org/id/FA1487CEFF835F14D3B951CF2EB9088E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nabozhenko, Maxim V.	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
FA1487CEFFBD5F15D3B956D62EB80BB3.text	FA1487CEFFBD5F15D3B956D62EB80BB3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euboeus Boieldieu 1865	<div><p>Key to species of the genus Euboeus of the Caucasus, Iran and Turkmenistan on females</p><p>(without E. prometheus, E. arzanovi and E. zubovi whith unknown females)</p><p>1 Pronotum and head dorsally strongly shiny, dark-bluish or dark-violet, elytra black (Fig. 19B)............ E. chorasanicus</p><p>- Body dorsally dull, completely black (Figs 2D, 4E, 7E, 9C, 11D, 12B, 13D, 14C, 15C, 17C)......................... 2</p><p>2 Puncturation of head and pronotum coarse and dense, punctures subequal or larger than interpuncture distance; sometimes punctures coarse and sparse at middle of pronotum (Figs 1A, 2E, F, 4F, G, 5D, E, 6B–D, 7F, G)...................... 3</p><p>- Puncturation of head and pronotum fine and sparse, puncture diameter much lesser than interpuncture distance (Figs 9D, E, 10A, B, 11E, F, 13E, F, 14D, E, 15D, E, 17D, E), at least in anterior half (Fig. 12B)................................ 6</p><p>3 Pronotum with very coarse and dense puncturation of longitudinally elongated punctures, especially on lateral sides of disc; median smooth line of pronotum elevated near base and separated from another surface by impression with very dense merged punctures (Fig. 2D, E, F).................................................................... E. corrugosus</p><p>- Pronotum with coarse and dense puncturation of simple round, not merged punctures; median line not elevated near base or absent (Figs 4E, 6B–D, 7E)............................................................................. 4</p><p>4 Head densely pubescent by reddish, recumbent, long setae (Figs 6B–D)................................... E. vicinus</p><p>- Head bare (Figs 4E, 7E)............................................................................... 5</p><p>5 Metaventrite and abdominal ventrites pubescent with long recumbent hairs (Fig. 4C, D).................. E. gorganicus</p><p>- Metaventrite and abdominal ventrites bare, covered with simple very short setae (Fig. 7C, D)............... E. zoroaster</p><p>6 Lateral margins of pronotum not completely beaded, bead widely interrupted at middle (Figs 14C, 17C)................ 7</p><p>- Lateral margins of pronotum completely beaded (Figs 9C, 11D, 13D) or completely not beaded (Fig. 12B, 15C)......... 8</p><p>7 Prothoracic hypomera wrinkled (Fig. 17F)...................................................... E. quadricollis</p><p>- Prothoracic hypomera coarsely punctured (Fig. 14F)............................................. E. krivokhatskyi</p><p>8 Lateral margins of pronotum not beaded (Figs 12B, 15C)..................................................... 9</p><p>- Lateral margins of pronotum completely beaded (Figs. 7C, D, E, 9D, E, F, 10D, E, F).............................. 10</p><p>9 Prothoracic hypomera with coarse large longitudinal wrinkles (Fig. 12F). Lateral margins of pronotum strongly rounded at middle and widely emarginated at base (Fig. 12B)................................................. E. kalashiani</p><p>- Prothoracic hypomera with dense, large, foveolate punctures (Fig. 15F). Lateral margins of pronotum weakly evenly rounded (Fig. 15C)................................................................................... E. merkli</p><p>10 Pronotum widest ahead of middle (Fig. 9C). Prothoracic hypomera not flattened along margin, with small confused microwrinkles with poorly distinguished sparse smoothed puncturation against their background; wrinkles with microgranulation at anterior third (Fig. 9F).......................................................................... E. grimmi</p><p>- Pronotum widest at middle or behind middle (Figs 11D, 13D). Prothoracic hypomera narrowly (Fig. 13G) or widely (Fig. 11G) flattened along margin, coarsely and densely punctured (Fig. 13G) or coarsely longitudinally wrinkled (Fig. 11G), without microgranulation.................................................................................... 11</p><p>11 Prothoracic hypomera with coarse longitudinal wrinkles; lateral margin of prothoracic hypomera widely flattened (Fig. 11G)......................................................................................... E. huedepohli</p><p>- Prothoracic hypomera with coarse puncturation and smooth wrinkles at middle; lateral margin of prothoracic hypomera very narrowly weakly flattened (Fig. 13G)............................................................ E. kasatkini</p></div>	https://treatment.plazi.org/id/FA1487CEFFBD5F15D3B956D62EB80BB3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nabozhenko, Maxim V.	Nabozhenko, Maxim V. (2022): A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan. Zootaxa 5159 (4): 451-486, DOI: 10.11646/zootaxa.5159.4.1
