identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
F46087FB94544C4F908A36D0FCBAFACB.text	F46087FB94544C4F908A36D0FCBAFACB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bucculentum Schweitzer & Feldmann 2009	<div><p>Genus Bucculentum Schweitzer &amp; Feldmann, 2009</p> <p>Diagnosis. After Schweitzer &amp; Feldmann (2009), amended. Carapace roughly rectangular; rostrum tri-lobed, projected well beyond frontal margin of carapace; orbit appearing to be placed under rostrum, augenrest on hepatic region of dorsal carapace, bounded by spines or ridges; hepatic region strongly inflated; groove (named here augenrest groove) leading from mid dorsal margin of augenrest to cervical groove; cervical and branchio-cardiac grooves deep; urogastric region not well defined, depressed below level of metagastric and cardiac regions; cardiac region elevated, its dominant tubercle making highest point of carapace.</p> <p>Type species: Nodoprosopon bucculentum Wehner, 1988.</p> </div>	https://treatment.plazi.org/id/F46087FB94544C4F908A36D0FCBAFACB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Krzemińska, Ewa;Starzyk, Natalia;Fraaije, René H. B.;Schweigert, Günter;Lukeneder, Alexander	Krzemińska, Ewa, Starzyk, Natalia, Fraaije, René H. B., Schweigert, Günter, Lukeneder, Alexander (2021): Jurassic brachyurans of the genus Bucculentum. Zootaxa 5032 (3): 395-410, DOI: 10.11646/zootaxa.5032.3.5
F46087FB94544C4B908A30A4FDC1FD53.text	F46087FB94544C4B908A30A4FDC1FD53.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bucculentum bucculentum (Wehner 1988) bucculentum (Wehner 1988	<div><p>Bucculentum bucculentum (Wehner, 1988)</p> <p>Fig. 1A, B, 2, 3</p> <p>1988 Nodoprosopon bucculentum Wehner, p. 53, pl. 4, figs. 1–2, 6.</p> <p>2009 Bucculentum bucculentum (Wehner, 1988) in: Schweitzer &amp; Feldmann, 2009a: p. 79, fig. 3.3–3.4.</p> <p>Diagnosis. After Schweitzer &amp; Feldmann (2009), amended. Carapace from longer than wide to almost quadrangular, widest across hepatic regions; rostrum tri-lobed, armoured with spines, central lobe being longest and depressed; augenrest positioned perpendicular to body long axis, laterally flanked by short ridge; suborbital spine strong, single, with several sharp terminal spines; cardiac region with three tubercles arranged along body long axis, distalmost tubercle being largest.</p> <p>Age and distribution. From at least Middle Oxfordian (Poland) to Early Kimmeridgian: Platynota-Divisum zones (Germany); the specimen SMNS 66654 described here as „ Bucculentum cf. bucculentum ” would extend the range of this species to the Late Kimmeridgian Pseudomutabilis Zone.</p> <p>Materials examined. Holotype SNS̲-̲SPG 1980 XXX 1255 (Fig. 2A, B), Biburg (Germany), age Early Kimmeridgian: Planula Zone (Schairer 1982; Schweitzer &amp; Feldmann 2009a). Paratype SNS̲-̲SPG 1980 XXX 1254 (Fig. 2C), same locality and age. Specimen SNS̲-̲SPG 1987 I 55 (Fig. 2D, E), Unterwilflingen (Germany), age Early Kimmeridgian: Platynota-Divisum zones (Wehner 1988). Specimens from the Polish Jura Chain described and illustrated by Starzyk et al. (2012) were re-examined for this study; they come from two quarries: Ogrodzieniec (Lower and Middle Oxfordian) and Niegowonice (from Cordatum to Transversarium zones, except discontinuous Mariae Zone; Głowniak 2006).</p> <p>New material: MAB 3572 (Fig. 3 E, F), Geisingen (Germany; N 47 o 93’, E 8 o 67’), age Late Kimmeridgian: Mutabilis/Acanthicum Zone (Van Bakel et al. 2008); MAB 3577, MAB 3579 (Fig. 3G), MAB 3582a (positive, Fig. 3A) and MAB 3582b (negative, Fig. 3B, C, D), Plettenberg quarry n. Balingen, Germany; all coll. R. H. B. Fraaije), age Early Kimmeridgian, Planula Zone (Schweigert &amp; Callomon 1997; Jantschke &amp; Schweigert 2020); I-F/MP/6210/1578/11 (ISEA) and I-F/MP/4648/1534/08 (ISEA), Wysoka n. Łazy (N 50 o 25’44’’, E 19 o 21’13’’), Polish Jura Chain, age Middle Oxfordian (Głowniak 2002).</p> <p>Additional description. The rostrum is tri-lobed, the central lobe being the longest and directed downwards while the two flanking lobes are raised; together, they form a kind of a scoop armoured with terminal rows of sharp spines. The entire rostrum, or almost so, is preserved only in the specimens MAB 3572 from Plettenberg and I-F/ MP/6210/1578/11 from Poland (Fig. 3E, H, respectively), but it is fragmentarily present in several other specimens, including the holotype (Fig. 2A, B) and specimen SNSB-BSPG 1987 I 55 (Fig. 2D, E). The augenrest is convex and directed frontally, i.e, at an angle c. 90 o to the body long axis. The suborbital spine is single, massive, and oval in cross section. The distal portion of this spine is armoured with few teeth (traces of four teeth are visible in two specimens, Fig. 2B and 3G). The epibranchial region of the holotype (Fig. 2A) and MAB 3572 has six ridges ending in rows of tubercles, fan-like spreading just over the cardiac region; in other specimens this feature is weakly expressed or absent. Along the lateral sides there are strong spines (Fig. 3D, F, G).</p> <p>Remarks. Noteworthy is the specimen MAB 3852, preserved in positive (convex) and negative (concave) parts (Fig. 3A, B, resp.), which supplement each other with details; the concave mould of the rostrum with white depressions left by the warts is completely preserved in the negative part (Fig. 3C); this part retained also the traces of lateral spines along the metabranchial region (Fig. 3D), and almost complete cover of white cuticle, visible from the inner side. The specimen MAB 3572 (Fig. 3F) shows that the pattern of tubercles is preserved both on the cuticle and on the surface of the mould beneath.</p></div> 	https://treatment.plazi.org/id/F46087FB94544C4B908A30A4FDC1FD53	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Krzemińska, Ewa;Starzyk, Natalia;Fraaije, René H. B.;Schweigert, Günter;Lukeneder, Alexander	Krzemińska, Ewa, Starzyk, Natalia, Fraaije, René H. B., Schweigert, Günter, Lukeneder, Alexander (2021): Jurassic brachyurans of the genus Bucculentum. Zootaxa 5032 (3): 395-410, DOI: 10.11646/zootaxa.5032.3.5
F46087FB94504C48908A37DCFCFBFE9F.text	F46087FB94504C48908A37DCFCFBFE9F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bucculentum Schweitzer & Feldmann 2009	<div><p>Bucculentum cf. B. bucculentum</p> <p>Fig. 4</p> <p>Material examined. Specimen SMNS 66654: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=9.983334&amp;materialsCitation.latitude=48.633335" title="Search Plazi for locations around (long 9.983334/lat 48.633335)">Gundershofen-Springen</a>, source of Schmiech river, southern Germany (N 48 o 38’, E 9 o 59’); age: Late Kimmeridgian, Pseudomutabilis Zone (according to Ziegler 1959).</p> <p>Description. The specimen is characterized by two unique features: the shape of the suborbital spine and mesobranchial regions. The suborbital spine (Fig. 4A, B) is very broad, flat, and curved. The mesobranchial regions are very small and of semicircular outline, while in other specimens of the genus they are larger and triangular. Other features are concordant with those of B. bucculentum.</p> <p>Remarks. The specimen is symmetrical and well preserved, therefore, the carapace seems to present genuine characters. The suborbital spine such as in this specimen is not encountered in any other species of Bucculentum. However, it cannot be excluded that its shape results from taphonomic conditions; the second spine is not preserved, therefore it is not possible to compare their symmetry. Since we are not sure of the genuine shape, we refrain from description of a new species. On the other hand, if the specimen represents really B. bucculentum, the range of this species would be extended to the Late Kimmeridgian (Fig. 11).</p> </div>	https://treatment.plazi.org/id/F46087FB94504C48908A37DCFCFBFE9F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Krzemińska, Ewa;Starzyk, Natalia;Fraaije, René H. B.;Schweigert, Günter;Lukeneder, Alexander	Krzemińska, Ewa, Starzyk, Natalia, Fraaije, René H. B., Schweigert, Günter, Lukeneder, Alexander (2021): Jurassic brachyurans of the genus Bucculentum. Zootaxa 5032 (3): 395-410, DOI: 10.11646/zootaxa.5032.3.5
F46087FB94534C49908A3490FB08FD37.text	F46087FB94534C49908A3490FB08FD37.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bucculentum bachmayeri Schweitzer & Feldmann 2009	<div><p>Bucculentum bachmayeri Schweitzer &amp; Feldmann, 2009</p> <p>Fig. 1C, D, 5</p> <p>2009a Bucculentum bachmayeri in: Schweitzer &amp; Feldmann, Ann. Naturhist. Mus. Wien 110 A: 81, fig. 1.4, 3.1, 3.2. Diagnosis. Carapace widest across hepatic regions which are more expanded laterally than in Bucculentum bucculentum, but flattened (in frontal view); augenrest positioned perpendicular to body long axis, bounded by small tubercles on outer-orbital edge; suborbital spine bifid, its two prongs being thin and long; cardiac region small, ovate, with one dominant, sharp tubercle.</p> <p>Material examined. Holotype, NHMW 1990 /0041/3376, near Dörfles, Vienna, Ernstbrunn Limestone, Tithonian. Specimen SMNS 70606: Bad Überkingen-Oberböhringen near Geislingen an der Steige, southern Germany (N 48 o 61’, E 9 o 79’); Upper Kimmeridgian, Pseudomutabilis Zone (leg. H. Kuschel).</p> <p>Additional description. The frontal view of the holotype exposes a very characteristic feature of this species: the hepatic regions are broad and flattened when compared to B. bucculentum, so that the distance between the upper margin of the augenrest and the surface of the hepatic region over it is very short (marked with an arrow in Fig. 5A); the same character is visible in the specimen SMNS 70606 (Fig. 5C, arrow). The area of the augenrest is flat and smaller than in B. bucculentum; the plane of the augenrest is perpendicular to the long body axis, similar to B. bucculentum. The suborbital spine in the holotype is bifid, and its two prongs are thin (Fig. 5B); in the specimen SMNS 70606 these spines are not preserved. The lateral sides of carapace are endowed with strong, short spurs partially preserved in SMNS 70606 (Fig. 5E, F).</p> <p>The known specimens delimit the range of this species to Late Kimmeridgian-Tithonian.</p></div> 	https://treatment.plazi.org/id/F46087FB94534C49908A3490FB08FD37	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Krzemińska, Ewa;Starzyk, Natalia;Fraaije, René H. B.;Schweigert, Günter;Lukeneder, Alexander	Krzemińska, Ewa, Starzyk, Natalia, Fraaije, René H. B., Schweigert, Günter, Lukeneder, Alexander (2021): Jurassic brachyurans of the genus Bucculentum. Zootaxa 5032 (3): 395-410, DOI: 10.11646/zootaxa.5032.3.5
F46087FB94524C49908A3638FE45F93A.text	F46087FB94524C49908A3638FE45F93A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bucculentum horstkuscheli Krzemińska & Starzyk & Fraaije & Schweigert & Lukeneder 2021	<div><p>Bucculentum horstkuscheli, n. sp.</p> <p>Fig. 1E, F, 6</p> <p>Diagnosis. Hepatic regions large, much wider than metabranchial regions; metabranchial region not expanded laterally; augenrest directed laterad at c. 60 o to body long axis and aligned with proximal portions of hepatic region; suborbital spine wide, directed outwards and downwards, and ending in two short, triangular teeth; cardiac region with one distalmost dominant tubercle accompanied anteriorly by row of small tubercles.</p> <p>Etymology. Trivial name is dedicated to Horst Kuschel (Faurndau), who found and donated the holotype of this species.</p> <p>Material examined. Holotype: # SMNS 70607 (leg. H. Kuschel), Germany: Bad Überkingen-Oberböhringen, near Geislingen a. d. Steige. Age: Late Kimmeridgian: Acanthicum Zone (Fraaije et al. 2017).</p> <p>Description. The holotype represents an internal mould, with only the traces of the white cuticle in the grooves. It is excellently preserved and symmetrical, lacking only the rostrum. The hepatic regions are very wide, c. 1.4x wider than the mesobranchial region and 1.2x wider than the metabranchial region. The frontal portions containing the augenrests and anterior portions of hepatic regions are aligned, obliquely set to the long axis at an angle of 57 o, symmetrical on both sides. Augenrests are almost perpendicular to the plane of carapace, only very slightly shifted dorsally (Fig. 6B), at an angle of c. 80 o to the upper surface of the carapace; they are laterally flanked by single, broad, triangular tubercles. The suborbital spine is very broad, supporting most of the lower margin of the augenrest, directed down- and outwards, and terminating in two short, triangular teeth (Fig. 6B, C, arrows). The epibranchial region is narrow; the cervical groove is almost straight. The cardiac region is equipped at its distal end with a large tubercle (which is the highest point of the carapace) and a median row of small tubercles towards the mesobranchial region. The metabranchial regions are narrow, almost rectangular; the distal end is broad, stretched over the entire distal portion of the metabranchial region and upturned.</p> <p>Remarks. The new species resembles B. bachmayeri in having a large hepatic region, a divided suborbital spine, and a cardiac region with one large tubercle, but differs in having the augenrests directed outwards and positioned obliquely to body long axis. The suborbital spine is divided into two short, triangular lobes, unlike the long, thin prongs in B. bachmayeri.</p> </div>	https://treatment.plazi.org/id/F46087FB94524C49908A3638FE45F93A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Krzemińska, Ewa;Starzyk, Natalia;Fraaije, René H. B.;Schweigert, Günter;Lukeneder, Alexander	Krzemińska, Ewa, Starzyk, Natalia, Fraaije, René H. B., Schweigert, Günter, Lukeneder, Alexander (2021): Jurassic brachyurans of the genus Bucculentum. Zootaxa 5032 (3): 395-410, DOI: 10.11646/zootaxa.5032.3.5
F46087FB94524C46908A3228FED5F84A.text	F46087FB94524C46908A3228FED5F84A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bucculentum plettenbergense Krzemińska & Starzyk & Fraaije & Schweigert & Lukeneder 2021	<div><p>Bucculentum plettenbergense, n. sp.</p> <p>Fig. 1G, 7</p> <p>Diagnosis. Metabranchial region large, more than half as long as carapace, and a little wider than hepatic region; augenrest round, slightly concave, shifted more dorsally than in other species of genus, and flanked laterally by long, crescent ridge; suborbital spine undivided, curved upwards; epibranchial region very narrow.</p> <p>Etymology. The new species is named after its type locality.</p> <p>Material examined. Holotype MAB 3576, Germany: Plettenberg quarry n. Balingen (N 48 o 21’, E 8 o 80’), coll. RHB Fraaije, Late Oxfordian: Planula Zone (Schweigert &amp; Callomon 1997; Jantschke &amp; Schweigert 2020).</p> <p>Description. The carapace is 8.43 mm long. The rostrum and centro-anterior portion have been split off. Left augenrest complete, large, round, shifted to dorsal side at an angle c. 50 o to the surface of carapace and delimited laterally by a narrow, crescent-like ridge. The suborbital spine is directed upwards (Fig. 7A, B, arrows). The epibranchial region is very narrow (less than 10% of the total lateral length of carapace), with two symmetric swellings over the cardiac region. The cardiac region has three tubercles arranged axially, the distalmost one being the largest, and also the highest point of the carapace. The metabranchial region is characteristically large, a little bit wider than the hepatic region, and more than half as long as the carapace.</p> <p>Remarks. Bucculentum plettenbergense, n. sp., is distinguished because of having the largest metabranchial region amongst all bucculentids, combined with the narrowest epibranchial region. Such proportions are not met in any other congener.</p> </div>	https://treatment.plazi.org/id/F46087FB94524C46908A3228FED5F84A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Krzemińska, Ewa;Starzyk, Natalia;Fraaije, René H. B.;Schweigert, Günter;Lukeneder, Alexander	Krzemińska, Ewa, Starzyk, Natalia, Fraaije, René H. B., Schweigert, Günter, Lukeneder, Alexander (2021): Jurassic brachyurans of the genus Bucculentum. Zootaxa 5032 (3): 395-410, DOI: 10.11646/zootaxa.5032.3.5
F46087FB945E4C45908A35B1FA3DFE26.text	F46087FB945E4C45908A35B1FA3DFE26.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bucculentum Schweitzer & Feldmann 2009	<div><p>Genus Bucculentum, key to the species</p> <p>1. Metabranchial region nearly as wide as hepatic region; epibranchial region only c. 1/3 as wide as hepatic region........................................................................................ B. plettenbergense, n. sp.</p> <p>- metabranchial region narrower than hepatic region......................................................... 2.</p> <p>2. Augenrest vertical to long body axis or almost so (dorsal view)............................................... 3.</p> <p>- augenrest oblique, aligned with proximal portions of hepatic regions.......................... B. horstkuscheli, n. sp.</p> <p>3. Suborbital spur divided; in frontal view augenrest is close to dorsal side of carapace (Fig. 1D) and flanked with several small tubercles................................................................................ B. bachmayeri.</p> <p>- suborbital spur undivided; in frontal view augenrest is close to ventral side of carapace (Fig. 1B) and flanked laterally with one elongated protuberance.................................................................... B. bucculentum.</p> </div>	https://treatment.plazi.org/id/F46087FB945E4C45908A35B1FA3DFE26	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Krzemińska, Ewa;Starzyk, Natalia;Fraaije, René H. B.;Schweigert, Günter;Lukeneder, Alexander	Krzemińska, Ewa, Starzyk, Natalia, Fraaije, René H. B., Schweigert, Günter, Lukeneder, Alexander (2021): Jurassic brachyurans of the genus Bucculentum. Zootaxa 5032 (3): 395-410, DOI: 10.11646/zootaxa.5032.3.5
