taxonID	type	description	language	source
F56787A8FFB6ED0477BE9953FECC5376.taxon	description	Paederus gottschei is the most widespread - and probably also the most common - representative of Harpopaederus in the East Palaearctic region, its distribution ranging from the Russian Far East across Northeast China to North and South Korea. For additional records see LI & ZHOU (2007). So far, all the material of Harpopaederus from this region has proved to belong to P. gottschei, making it seem most unlikely that P. pseudobaudii represents a distinct species. In consequence, P. pseudobaudii is formally placed in synonymy with P. gottschei. The distribution of P. gottschei in China is illustrated in Map 1.	en	Assing, Volker (2015): On the Harpopaederus fauna of China (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 47 (1): 163-190, DOI: 10.5281/zenodo.5414042
F56787A8FFB7ED0277BE994AFED05479.taxon	description	A d d i t i o n a l m a t e r i a l e x a m i n e d China: S h a a n x i: 233, Xunyangba, 1. - 3. VII. 2005, leg. Jeniš (cAss). C o m m e n t: The original description of P. konfuzius is based on thirteen type specimens from three localities in the Qinling Shan, Shaanxi (WILLERS 2001 b). The specimens from one of the localities, at least five paratypes, are not conspecific with the holotype, but belong to P. agnatus. The record from the Taibai Shan is based on three females and consequently somewhat uncertain. PENG et al. (2015) reported additional localities from the Qinling Shan. LI & ZHOU (2007) erroneously reported the species also from Gansu and Sichuan (PENG et al. 2015). Confirmed records are known only from Shaanxi (Map 3).	en	Assing, Volker (2015): On the Harpopaederus fauna of China (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 47 (1): 163-190, DOI: 10.5281/zenodo.5414042
F56787A8FFB0ED0277BE9C3BFC615708.taxon	description	C o m m e n t: The original description is based on a male holotype and a female paratype from Jiuzhaigou in North Sichuan (WILLERS 2001 b). The species was subsequently reported from " Wen County, Qiujia Dam " in Gansu [32 ° 58 ' N, 104 ° 01 ' E] by LI & ZHOU (2007). The currently known distribution is illustrated in Map 2.	en	Assing, Volker (2015): On the Harpopaederus fauna of China (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 47 (1): 163-190, DOI: 10.5281/zenodo.5414042
F56787A8FFB1ED0377BE9918FD0E507D.taxon	description	C o m m e n t: Paederus brevior has been recorded from several localities in the Qinling Shan, Shaanxi (LI et al. 2014; PENG et al. 2015). The distribution apparently strongly overlaps with that of P. konfuzius (Map 3).	en	Assing, Volker (2015): On the Harpopaederus fauna of China (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 47 (1): 163-190, DOI: 10.5281/zenodo.5414042
F56787A8FFB1ED0377BE9838FC5A5314.taxon	description	C o m m e n t: This species was described based on material from several localities in Gansu (LI & ZHOU 2007) and subsequently reported also from two localities in Shaanxi (PENG et al. 2015). If all the records reported in the literature are based on correctly identified material, this species is remarkably widespread in the Qinling Shan and the distribution overlaps with those of several other Harpopaederus species (Map 2).	en	Assing, Volker (2015): On the Harpopaederus fauna of China (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 47 (1): 163-190, DOI: 10.5281/zenodo.5414042
F56787A8FFBCED0E77BE9EBFFD0C51B6.taxon	description	C o m m e n t: The original description is based on six type specimens from " Micang Shan, Daba, 32 ° 40 ' N, 107 ° 02 ' E " and the above specimens (PENG et al. 2015). The currently known distribution is illustrated in Map 3.	en	Assing, Volker (2015): On the Harpopaederus fauna of China (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 47 (1): 163-190, DOI: 10.5281/zenodo.5414042
F56787A8FFBCED0C77BE9862FDDE571B.taxon	description	A d d i t i o n a l m a t e r i a l e x a m i n e d China G a n s u 1 ♀, Qinling Shan, N Chengxian, 34 ° 10 ' N, 105 ° 43 ' E, 1850 m, mixed secondary forest margin, litter sifted, 29. VII. 2012, leg. Wrase (cAss); 13, 1 ♀, Qinling Shan, N Chengxian, 34 ° 10 ' N, 105 ° 42 ' E, 1830 m, stream valley with secondary deciduous forest, moist litter sifted, 29. VII. 2012, leg. Schülke & Wrase (cSch, cAss); 13, 2 ♀♀, Qinling Shan, NW Longnan, Lazikou pass, S-side, Laolong valley, 34 ° 08 ' N, 103 ° 52 ' E, 2300 m, S-slope with pine and spruce forest, litter sifted, 3. VIII. 2012, leg. Assing, Schülke & Wrase (cAss, cSch); 13, Qinling Shan, NW Longnan, Lazikou pass, S-side, Zhuli valley, 34 ° 08 ' N, 103 ° 56 ' E, 2260 m, N-slope, mixed forest with oak and pine near stream, litter and dead wood sifted, 3. VIII. 2012, leg. Wrase (cAss); 333, 2 ♀♀, Lazikou valley, 34 ° 10 ' N, 103 ° 48 - 52 ' E, 2120 - 2510 m, 28. VI. 2005, leg. Hájek et al. (cSch, cAss); 13, Lazikou valley, 34 ° 10 ' N, 103 ° 52 ' E, 2120 m, vegetation along meadow, baited pitfall trap (fish, meat, cheese), 28. VI. 2005, leg. Hájek et al. (cSch); 233, Lazikou pass, 34 ° 14 ' N, 103 ° 54 ' E, 3180 m, grassland, 29. VI. 2005, leg. Hájek et al. (cSch, cAss); 233, S Longnan, Min Shan, 33 ° 03 ' N, 104 ° 41 ' E, 2200 m, secondary pine forest with hazelnut, moist litter and roots sifted, 6. VIII. 2012, leg. Assing & Wrase (cAss, cSch). S h a a n x i [see also type material of P. konfuzius]: 233, 1 ♀, Qinling Shan, 105 km SW Xi'an, pass on road Zhouzhi-Foping, 33 ° 46 ' N, 107 ° 58 ' E, 1880 m, 4. VII. 2001, leg. Wrase (cSch); 13, 2 ♀♀, Qinling Shan, 105 km SW Xi'an, pass on road Zhouzhi-Foping, 33 ° 46 ' N, 107 ° 58 ' E, 1700 m, stream valley, mixed deciduous forest, 3. VII. 2001, leg. Wrase (cSch, cAss); 2 ♀♀, Qinling Shan, 105 km SW Xi'an, pass on road Zhouzhi-Foping, 33 ° 44 ' N, 107 ° 59 ' E, 1990 m, 2. - 4. VII. 2001, leg. Schülke & Wrase (cSch, cAss); 1 ♀, Qinling Shan, 52 km SSW Zhouzhi, 33 ° 44 ' N, 107 ° 58 ' E, 1900 m, mixed forest, litter and soil sifted, 25. VII. 2012, leg. Wrase (cSch). C o m m e n t: The original description of P. agnatus is based on two syntypes (" 2 Exemplare ") from " Kan-ssu " (= Gansu) (EPPELSHEIM 1889), that of P. dangchangensis on a male holotype and a male paratype from " Dangchang County, Dahe Dam " (LI & ZHOU 2007). Only one syntype of P. agnatus was found in the Eppelsheim collection at the NHMW. Since it is a female, it is not designated as the lectotype. The second syntype may be in the Potanin collection, whose whereabouts are apparently unknown (HORN et al. 1990), if it still exists. The syntype from the Eppelsheim collection had been examined, but not assigned to a subgenus, by WILLERS (2001 b). Thus, the species is listed as incertae sedis in SMETANA (2004) and SCHÜLKE & SMETANA (in press). Based on the external (coloration of the antennae and legs; shape and coloration of the mandibles; shape of labrum), as well as the female sexual characters (shapes of tergite VIII and sternite VIII), the syntype of P. agnatus is conspecific with P. dangchangensis. Hence the synonymy proposed above. A comparison of the sexual characters of the material from the environs of Chengxian, of the specimens from the regions to the northwest and southwest of Longnan, of material from Shaanxi, and of the aedeagus of the holotype based on photos kindly sent to me by Xiaoyan Li (Beijing) revealed some differences, particularly regarding the shape of the internal structures of the aedeagus (Figs 1 - 12, 17 - 19, 14, 26, 29), the size and number of the denticles on the dorsal plate of the aedeagus, the depth of the posterior excision of the male sternite VIII (Figs 13 - 14, 54), the shape of the posterior margin of the female tergite VIII (Figs 20 - 21), and the shape of the apical projection of the female sternite VIII (Figs 22 - 23). The aedeagi of the holotype and the paratype of P. dangchangensis, as well as those of some males from Shaanxi have smaller denticles on the dorsal plate and an internal structure of slightly different shape. However, no significant differences were found in the external characters and in the general shape of the aedeagus (Figs 1 - 12, 15 - 16, 24 - 27). The question whether or not the observed differences are an expression of inter- or intraspecific variation cannot answered with certainty, but the observation that slight differences may occur even within and between geographically close populations suggests that they are most plausibly interpreted as intraspecific variation. Regardings its male and the female sexual characters, P. agnatus is similar to P. konfuzius, with which is was previously confounded (see type material of P. konfuzius) and from which it is distinguished by the absence of pronounced sexual dimorphisms of the head shape and the mandibles, by the paler tibiae (P. agnatus: usually yellowish, very indistinctly infuscate at most; P. konfuzius: more or less distinctly and more or less extensively infuscate basally), by the smaller aedeagus (P. agnatus: approximately 1.9 mm; P. konfuzius: 2.0 - 2.1 mm), and by the apical internal structure of the aedeagus (P. konfuzius: straight, dagger-shaped, and more strongly sclerotized). For illustrations of P. konfuzius see PENG et al. (2015). The previously unknown female sexual characters of P. agnatus are illustrated in Figs 20 - 23. Paederus agnatus is remarkably widespread in the Qinling Shan, from central southern Gansu to central southern Shaanxi (Map 2).	en	Assing, Volker (2015): On the Harpopaederus fauna of China (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 47 (1): 163-190, DOI: 10.5281/zenodo.5414042
F56787A8FFBEED0B77BE9F1EFD6754FA.taxon	description	E t y m o l o g y: The specific epithet (Latin, adjective: without teeth) alludes to the absence of denticles on the dorsal plate of the aedeagus. D e s c r i p t i o n: Body length 10.0 - 11.0 mm; length of forebody 4.7 - 6.0 mm. Coloration: head black; pronotum bright reddish; elytra metallic-blue; scutellum palereddish; abdomen bicoloured with segments III-VI pale-reddish and segments VII-X black; legs dark-yellowish, with the apical halves of the femora blackish; antennae darkyellowish. Head (Fig. 30) transverse, 1.13 - 1.19 times as broad as long; punctation rather fine and sparse in median and anterior dorsal portions, coarser and denser in lateral and posterior dorsal portions; interstices without microsculpture. Eyes noticeably shorter than postocular region from posterior margin of eye to posterior constriction of head. Antenna 2.9 - 3.0 mm long; antennomere IV approximately twice as long as broad; antennomere X approximately 1.5 times as long as broad. Labrum with rather deeply and broadly excavate anterior margin (Figs 31 - 32). Pronotum (Fig. 30) 1.06 - 1.07 times as long as broad and 1.02 - 1.06 times as broad as head, strongly convex in cross-section; dorsal series each composed of approximately 10 punctures. Elytra (Fig. 30) approximately 0.68 - 0.75 times as long as pronotum and of trapezoid shape; punctation moderately coarse and of somewhat variable density; interstices without microsculpture. Hind wings completely reduced. Protarsomeres I-IV with pronounced sexual dimorphism. Metatarsomere I approximately as long as the combined length of II and III. Abdomen approximately 1.1 times as broad as elytra; punctation moderately fine and rather dense; pubescence long and blackish; interstices with fine and transverse microsculpture; posterior margin of tergite VII without palisade fringe; tergite VIII sexually dimorphic. 3: protarsomeres I-IV strongly dilated; tergite VIII strongly convex posteriorly; posterior margin of sternite VII weakly concave in the middle; sternite VIII (Fig. 33) weakly transverse, posterior excision nearly half as deep as length of sternite; tergites IX- X as in Fig. 34; aedeagus (Figs 35 - 42) 2.0 mm long and nearly symmetric; ventral process very weakly sclerotized; dorsal plate long, basally slender, apically acute and not curved in lateral view, extending far beyond apices of parameres, without denticles; parameres apically very weakly curved; internal sac with a clip-shaped weakly sclerotized basal structure, but without sclerotized spines. ♀: protarsomeres I-IV moderately dilated, much less so than in male; tergite VIII (Fig. 43) strongly tapering posteriad, posterior margin blackish, weakly convex in the middle; sternite VIII (Fig. 44) posteriorly with median process of triangular shape, postero-lateral angles acute and projecting posteriad; tergites IX and X as in Fig. 45. C o m p a r a t i v e n o t e s: The new species differs from the geographically close and externally highly similar P. agnatus, as well as from other micropterous Harpopaederus species recorded from the western Qinling Shan by the morphology of the aedeagus (dorsal plate rather slender basally, without denticles, apically not curved in lateral view; apices of parameres only very weakly curved), and additionally as follows: from P. agnatus by the shapes of the female tergite VIII and sternite VIII (see Figs 9 - 10); from P. antennocinctus, with which it shares the absence of denticles on the dorsal plate of the aedeagus, by uniformly yellowish antennae (P. antennocinctus: antennomeres III- XI apically infuscate), the yellowish tibiae and basal halves of the femora, the differently shaped female tergite VIII (P. antennocinctus: apically strongly convex), and the much shorter and less slender median projection of the posterior margin of the female sternite VIII (see description and figures in WILLERS 2001 b); from P. gracilacutus, with which it shares a similar shape of the labrum, by the acutely angled and distinctly projecting postero-lateral angles of the female sternite VIII; for illustrations of P. gracilacutus see PENG et al. (2015). D i s t r i b u t i o n a n d n a t u r a l h i s t o r y: Thespecieswasdiscoveredin two localities situated to the southeast of Longnan, southern Gansu. The specimens were collected by sifting litter and moss in a mixed pine and birch forest and in a shrub habitat on a slope with scree at altitudes of 2060 and 2080 m.	en	Assing, Volker (2015): On the Harpopaederus fauna of China (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 47 (1): 163-190, DOI: 10.5281/zenodo.5414042
F56787A8FFB9ED0877BE9CBEFDB75799.taxon	description	E t y m o l o g y: The specific epithet is an adjective derived from the name of the mountain range where the species was discovered. D e s c r i p t i o n: Body length 12.0 mm; length of forebody 6.0 mm. Coloration: head black; pronotum dark-reddish; elytra metallic-blue; scutellum dark-reddish; abdomen bicoloured with segments III-VI pale-reddish and segments VII-X black; legs with the tibiae and tarsi yellowish-brown, and the femora blackish with the bases narrowly and diffusely yellowish-brown; antennae and maxillary palpi dark-yellowish; labial palpi brown with the terminal joint yellowish-brown. Head (Fig. 46) transverse, 1.2 times as broad as long; punctation rather fine and sparse in median and anterior dorsal portions, denser and with interspersed coarser punctures in lateral and posterior dorsal portions; interstices without microsculpture. Eyes much shorter than postocular region from posterior margin of eye to posterior constriction of head. Antenna 3.3 mm long; antennomere IV more than twice as long as broad; antennomere X slightly more than 1.5 times as long as broad. Anterior margin of labrum broadly excavate and with a distinct tooth on either side of middle (Fig. 47). Pronotum (Fig. 46) 1.09 times as long as broad and 1.02 times as broad as head, strongly convex in cross-section; dorsal series each composed of approximately 10 punctures; lateral portions with sparse and fine punctation. Elytra (Fig. 46) 0.7 times as long as pronotum and of trapezoid shape; punctation moderately coarse and dense; interstices without microsculpture. Hind wings completely reduced. Metatarsomere I approximately as long as the combined length of II and III. Abdomen approximately 1.1 times as broad as elytra; punctation moderately fine and rather dense, somewhat sparser on tergites VII and VIII than on anterior tergites; pubescence rather short and blackish; interstices with fine and transverse microsculpture; posterior margin of tergite VII without palisade fringe. 3: protarsomeres I-IV distinctly dilated (Fig. 46); tergite VIII strongly convex posteriorly; posterior margin of sternite VII not distinctly concave in the middle; sternite VIII (Fig. 48) about as broad as long, posterior excision 0.55 times as deep as length of sternite; aedeagus (Figs 49 - 53) conspicuously long (2.7 mm) and slender; ventral process very weakly sclerotized; dorsal plate long, basally slender, apically acute and distinctly curved in lateral view, extending beyond apices of parameres, dorsally with two rows of 3 - 4 denticles at some distance from the slightly hooked apex; parameres apically hooked; internal sac with a clip-shaped weakly sclerotized basal structure, apical structure not sclerotized. ♀: unknown. C o m m e n t: The dorsal plate of the aedeagus of the holotype has a conspicuous flagshaped dorsal process subapically. Most likely, this structure is a teratotology and not a constant character. C o m p a r a t i v e n o t e s: Paederus minicus differs from other geographically close Harpopaederus species particularly by the conspicuously long and slender aedeagus, the absence of a distinct apical internal structure of the aedeagus, and the presence of two series of denticles on the dorsal plate of the aedeagus situated at some distance from the apex. From most species recorded from the Qinling Shan and adjacent mountain ranges, it is additionally distinguished by the dark-reddish pronotum, the dark femora, and the deep posterior incision of the male sternite VIII. D i s t r i b u t i o n a n d n a t u r a l h i s t o r y: The type locality is situated in the Min Shan in southern Gansu (Map 2) at an altitude between 2300 and 3300 m. Additional data are not available.	en	Assing, Volker (2015): On the Harpopaederus fauna of China (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 47 (1): 163-190, DOI: 10.5281/zenodo.5414042
F56787A8FFBAED1777BE9F9CFEF35056.taxon	description	E t y m o l o g y: The specific epithet is an adjective derived from the Latin noun cultellus (small knife) and alludes to the shape of the apical internal structure of the aedeagus. D e s c r i p t i o n: Body length 10.0 - 11.5 mm; length of forebody 5.0 - 5.2 mm. Coloration: head black; pronotum bright to dark reddish; elytra metallic-blue; scutellum reddish; abdomen bicoloured with segments III-VI pale-reddish and segments VII-X black; legs dark-yellowish, with the apical halves of the femora blackish; antennae and maxillary palpi dark-yellowish; labial palpi uniformly yellowish or with the basal palpomeres brown. Head (Fig. 55) transverse, approximately 1.15 times as broad as long; punctation rather fine and sparse in median and anterior dorsal portions, slightly less sparse and with interspersed coarser punctures in lateral and posterior dorsal portions; interstices without microsculpture. Eyes slightly more than half as long as postocular region from posterior margin of eye to posterior constriction of head. Antenna 3.1 - 3.2 mm long; antennomere IV approximately twice as long as broad; antennomere X approximately 1.5 times as long as broad. Anterior margin of labrum in the middle with broadly U-shaped excision, on either side of this incision with a more or less distinct tooth (Fig. 56). Pronotum (Fig. 55) approximately 1.1 times as long as broad and as broad as head, strongly convex in cross-section; dorsal series each composed of approximately 10 punctures; lateral portions with sparse and fine punctation. Elytra (Fig. 55) approximately 0.7 times as long as pronotum; punctation moderately coarse and dense; interstices without microsculpture. Hind wings completely reduced. Metatarsomere I approximately as long as the combined length of II and III. Abdomen approximately 1.1 times as broad as elytra; punctation moderately fine and moderately dense; pubescence rather short and brownish; interstices with fine and transverse microsculpture; posterior margin of tergite VII without palisade fringe. 3: protarsomeres I-IV strongly dilated; tergite VIII strongly convex posteriorly; posterior margin of sternite VII not distinctly concave in the middle; sternite VIII (Fig. 57) about as broad as long, posterior excision nearly 0.6 times as deep as length of sternite, posterior margin on either side of this incision strongly oblique; aedeagus (Figs 58 - 62) approximately 2.4 mm long; ventral process very weakly sclerotized; dorsal plate long, basally rather broad, gradually tapering apicad, and with long and slender apical portion, apically weakly curved dorsad and reaching far beyond apices of parameres, dorsally with two rows of 2 - 6 denticles; internal sac with a basal clip-shaped structure and a long (ca. 0.85 mm) dagger-shaped, apically acute, and moderately sclerotized apical structure (Fig. 62). ♀: unknown. C o m p a r a t i v e n o t e s: Based on the similar morphology of the aedeagus, P. cultellatus is closely allied to P. agnatus and particularly to P. konfuzius, suggesting that the unknown female secondary sexual characters are similar, too. It differs from both species by the longer aedeagus with a longer narrow apical portion of the dorsal plate, and additionally as follows: from P. konfuzius (aedeagus 2.0 - 2.1 mm long) by the yellowish tibiae, on average slightly smaller eyes, the apex of the dorsal plate of the aedeagus reaching much farther beyond the apices of the parameres, and by the larger and longer apical internal structure of the aedeagus (P. konfuzius: internal structure 0.65 - 0.70 mm long and more slender); from P. agnatus (aedeagus approximately 1.9 mm long) by the deeper posterior incision of the male sternite VIII, the apex of the dorsal plate extending slightly farther beyond the apices of the parameres, and particularly by the much more strongly sclerotized and differently shaped apical internal structure of the aedeagus. The new species is distinguished from the rather widespread P. gracilacutus (aedeagus of similar length) by slightly smaller eyes, by the deeper posterior incision of the male sternite VIII, by the less slender and apically less acute dorsal plate (P. gracilacutus: apex of dorsal plate extending even farther beyond the apices of the parameres), and by the much more strongly sclerotized and differently shaped apical internal structure of the aedeagus. For illustrations of P. agnatus, P. konfuzius, and P. gracilacutus see Figs 1 - 29, 54 and PENG et al. (2015), respectively. D i s t r i b u t i o n a n d n a t u r a l h i s t o r y: The type locality is situated in the border region between the provinces Shaanxi, Hubei, and Chongqing (Map 3). The specimens were sifted from leaf litter and moss in a young mixed forest at an altitude of 1700 - 1800 m.	en	Assing, Volker (2015): On the Harpopaederus fauna of China (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 47 (1): 163-190, DOI: 10.5281/zenodo.5414042
F56787A8FFA5ED1277BE9843FE23578F.taxon	description	E t y m o l o g y: The specific epithet is the present participle of the Latin verb deplecti (to claw) and alludes to the claw-like denticles on the dorsal plate of the aedeagus. D e s c r i p t i o n: Body length 11.0 - 12.5 mm; length of forebody 5.1 - 5.3 mm. Coloration: head black; pronotum reddish; elytra metallic-blue; scutellum reddish; abdomen bicoloured with segments III-VI pale-reddish and segments VII-X black; legs blackish, with the apices of the mesotibiae more or less distinctly paler and the tarsi brown to dark-brown; antennae and palpi yellowish. Head (Fig. 63) transverse, approximately 1.15 times as broad as long; punctation rather fine and sparse in median and anterior dorsal portions, less sparse and with interspersed coarser punctures in lateral and posterior dorsal portions; interstices without microsculpture. Eyes 0.6 - 0.7 times as long as postocular region from posterior margin of eye to posterior constriction of head. Antenna 3.3 - 3.4 mm long; antennomere IV more than twice as long as broad; antennomere X approximately 1.5 times as long as broad. Anterior margin of labrum in the middle with broadly U-shaped excision, on either side of this incision with a more or less distinct tooth (Fig. 64). Pronotum (Fig. 63) approximately 1.1 times as long as broad and as broad as head, strongly convex in cross-section; dorsal series each composed of approximately 10 punctures; lateral portions with sparse and moderately fine punctation. Elytra (Fig. 63) approximately 0.7 times as long as pronotum; punctation moderately coarse and dense; interstices without microsculpture. Hind wings completely reduced. Protarsomeres I-IV with pronounced sexual dimorphism. Metatarsomere I approximately as long as the combined length of II and III, or nearly so. Abdomen 1.05 - 1.10 times as broad as elytra; punctation moderately fine and moderately dense; pubescence moderately short and brownish; interstices with fine and transverse microsculpture; posterior margin of tergite VII without palisade fringe; tergite VIII with pronounced sexual dimorphism. 3: protarsomeres I-IV strongly dilated (Fig. 63); tergite VIII strongly tapering posteriad and very strongly convex posteriorly; posterior margin of sternite VII weakly concave in the middle; sternite VIII (Fig. 70) weakly transverse, posterior incision 0.53 times as deep as length of sternite; aedeagus (Figs 65 - 69) 2.4 mm long; ventral process very weakly sclerotized; dorsal plate apically far from reaching apices of parameres, with one large apical and two pairs of large subapical denticles; internal sac with a basal clipshaped structure and a broad and somewhat asymmetric apical internal structure (Fig. 69). ♀: protarsomeres I-IV moderately dilated; tergite VIII (Fig. 71) triangularly projecting posteriorly, middle of posterior margin obtusely pointed; sternite VIII shaped as in Fig. 72. C o m p a r a t i v e n o t e s: Based on the male and female sexual characters, particularly the large claw-shaped denticles on the dorsal plate of the aedeagus, P. deplectens is closely allied to P. chinensis, from which it differs by slightly smaller size, the slightly paler average coloration of the pronotum, the smaller aedeagus (P. chinensis: approximately 2.7 mm long), the presence of five (P. chinensis: two) claw-like denticles of the dorsal plate, and the apically slightly narrower female tergite VIII. For illustrations of P. chinensis see Figs 73 - 79. D i s t r i b u t i o n a n d n a t u r a l h i s t o r y: The known distribution is confined to two localities in northwestern Sichuan (Map 2). The examined type specimens were collected in a coniferous forest and on a stream bank. The altitudes range from 1500 to 2500 m.	en	Assing, Volker (2015): On the Harpopaederus fauna of China (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 47 (1): 163-190, DOI: 10.5281/zenodo.5414042
F56787A8FFA0ED1077BE9E6AFE8C5479.taxon	description	NW Chengdu, Qingcheng Hou Shan [approx. 30 ° 56 ' N, 103 ° 28 ' E], 1400 m, 2. - 4. V. 2006, leg. Murzin & Shokhin (cSch, cAss); 333, 6 ♀♀ [identified by Zhong Peng], Dujiangyan, Qingcfheng Shan, 30 ° 58 ' N, 103 ° 29 ' E, 1950 m, 30. VII. 2012, leg. Peng (SNUC). C o m m e n t: The original description of P. chinensis is based on an unspecified number of syntypes from " Szetschwan: Tatsienlu [= Kangding; 30 ° 03 ' N, 102 ° 01 ' E] " deposited in " der Sammlung des Dresdener Staatsmuseums und in meiner eigenen " (BERNHAUER 1931). One syntype, unfortunately a female, was located in the collections of the SNSD. WILLERS (2001 b) studied a male syntype from the Bernhauer collection, which he erroneously referred to as the holotype, and recorded the species from Guangkou [31 ° 00 ' N, 103 ° 37 ' E] and, based exclusively on females, from the environs of Wolong [30 ° 58 ' N, 103 ° 11 ' E]. He concluded from the structure of the aedeagus that P. chinensis did not belong to Harpopaederus. Paederus chinensis is the type species and sole representative of the subgenus Oedopaederus SCHEERPELTZ, 1957. SCHEERPELTZ (1957), however, had misinterpreted this species. The material from the Tianmu Shan in Zhejiang, which he had identified as P. chinensis and on which he based his description of Oedopaederus, was described as P. describendus by WILLERS (2001 b), who stated that " Oedopaederus Scheerp. muss jedoch (obwohl inhaltsleer geworden und umzuinterpretieren) mit der Art. [sic] P. chinensis als Typus subgeneris verbunden bleiben " [sic]. Based on the male and female sexual characters (Figs 73 - 79), the above material is conspecific with the syntype studied by WILLERS (2001 b). However, I have been unable to find any characters suggesting that P. chinensis should not belong to Harpopaederus. As in P. deplectens nov. sp., which P. chinensis is evidently closely related to, the dorsal plate of the aedeagus has conspicuously large apical and subapical denticles. In other respects, the aedeagus is similar also to the aedeagi of other Harpopaederus species. Consequently, P. chinensis is assigned to Harpopaederus and Oedopaederus is synonymized with Harpopaederus. Based on the illustrations and description provided by WILLERS (2001 b), P. describendus clearly does not belong to Harpopaederus. The revised distribution of P. chinensis is confined to three localities in West Sichuan (Map 2). The female-based record from Wolong (WILLERS 2001 b) most likely refers to P. deplectens.	en	Assing, Volker (2015): On the Harpopaederus fauna of China (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 47 (1): 163-190, DOI: 10.5281/zenodo.5414042
F56787A8FFA2ED1077BE9C3BFD5C56AA.taxon	description	C o m m e n t: The original description is based on a male holotype and twelve paratypes from Da Shennongjia in Hubei (WILLERS 2001 a). The species was subsequently reported from additional localities in Shennongjia County by LI & ZHOU (2007) and PENG et al. (2015). The record from Shaanxi in SMETANA (2004) is based on misidentified material. The currently known distribution is illustrated in Map 3. This species is characterized particularly by the pronounced dimorphism of the head (larger, more transverse, and with more distinctly marked posterior angles in males than in females) and of the mandibles (longer, curved, and with the molar teeth distinctly separated in males), by the coloration (legs blackish; antennomeres IV-XI apically more or less distinctly infuscate; preapical palpomere of the maxillary palpi apically infuscate), the morphology of the aedeagus, the broadly semi-transparent and strongly convex posterior margin of the female tergite VIII, and by the shape of the female sternite VIII. For illustrations of P. apfelsinicus see WILLERS (2001 a).	en	Assing, Volker (2015): On the Harpopaederus fauna of China (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 47 (1): 163-190, DOI: 10.5281/zenodo.5414042
F56787A8FFA2ED1F77BE994FFD7D5116.taxon	description	C o m m e n t: The original description is based on 16 type specimens from " Sichuan Province, Wu Mountain, Liziping " (LI & ZHOU 2007). As can be inferred from the map provided by LI & ZHOU (2007), the type locality is not Liziping in Sichuan [[ 28 ° 59 ' N, 102 ° 17 ' E], but Liziping wood farm [31 ° 12 ' N, 110 ° 00 ' E] in what has been Chonqing since 1997, near the border with Hubei. Since the illustrations of some body parts and the original description may be misleading, a redescription and new illustrations are provided. R e d e s c r i p t i o n: Body length 10.0 - 12.5 mm; length of forebody 4.8 - 5.6 mm. Coloration: head black; pronotum bright reddish; elytra metallic-blue; scutellum reddish to brown; abdomen bicoloured with segments III-VI pale-reddish and segments VII-X black; legs with blackish femora and metatibiae, and with dark-brown to blackish-brown pro- and mesotibiae and tarsi; antennae yellowish, with antennomeres V-XI often more or less distinctly infuscate in apical half, pro- and mesocoxae reddish, metacoxae blackish; maxillary palpi yellowish, with preapical palpomere more or less distinctly and more or less extensively infuscate; labial palpi dark-brown to blackish, with the terminal palpomere yellowish to reddish. Head (Figs 80 - 82) transverse, 1.1 - 1.2 times as broad as long, with moderately pronounced sexual dimorphism, on average larger and more transverse in males than in females; posterior angles weakly to moderately marked; punctation rather fine and sparse in median and anterior dorsal portions, coarser and denser in lateral and posterior dorsal portions; interstices without microsculpture. Eyes distinctly shorter than postocular region from posterior margin of eye to posterior constriction of head. Antenna 2.8 - 3.0 mm long; antennomere IV approximately twice as long as broad; antennomere X more than 1.5 times as long as broad. Labrum of somewhat variable shape, but anterior margin with more or less pronounced U-shaped median incision. Mandibles with distinct sexual dimorphism: longer and more slender in males than in females; molar teeth often separated in males, basally fused in females (Figs 81 - 82). Pronotum (Fig. 80) 1.04 - 1.06 times as long as broad and 1.00 - 1.08 times as broad as head, strongly convex in cross-section; dorsal series each composed of at least 10 punctures. Elytra (Fig. 80) 0.65 times as long as pronotum and of trapezoid shape; punctation rather fine and dense; interstices without microsculpture. Hind wings completely reduced. Protarsomeres I-IV with pronounced sexual dimorphism. Metatarsomere I approximately as long as the combined length of II and III. Abdomen approximately 1.05 times as broad as elytra; punctation moderately fine and rather dense; pubescence long and blackish; interstices with fine and transverse microsculpture; posterior margin of tergite VII without palisade fringe; tergite VIII sexually dimorphic. 3: protarsomeres I-IV strongly dilated (Fig. 80); tergite VIII posteriorly obtusely pointed; sternite VIII (Fig. 89) weakly transverse, posterior excision approximately 0.55 times as deep as length of sternite; aedeagus (Figs 83 - 88) approximately 2.9 mm long and nearly symmetric; ventral process very weakly sclerotized; dorsal plate long, basally slender, gradually tapering apicad, apically acute and curved ventrad in lateral view, extending distinctly beyond apices of parameres, with two series of rather small and numerous denticles; parameres apically very weakly curved; internal sac with a clipshaped weakly sclerotized basal structure and with an additional, moderately sclerotized, and apically needle-shaped structure. ♀: protarsomeres I-IV moderately dilated, much less so than in male; tergite VIII (Fig. 90) strongly tapering posteriad, posterior margin not distinctly semi-transparent, weakly convex in the middle; sternite VIII (Fig. 91) posteriorly with long and apically very acute median process, postero-lateral angles acute and projecting posteriad. C o m p a r a t i v e n o t e s: Based on the morphology of the aedeagus (dorsal plate long and slender, dorsally with two rows of numerous small denticles, shape of internal structure), as well as on the sexual dimorphism of the shapes of the head and of the mandibles, P. lineodenticulatus is closely allied to P. apfelsinicus, from which it differs by the less dark pro- and mesotibiae, the less distinctly infuscate antennomeres V-XI, the more strongly pointed male tergite VIII, the longer aedeagus (P. apfelsinicus (2.4 - 2.7 mm) with an apically more slender dorsal plate, the more slender female tergite VIII with a less convex and not distinctly semitransparent apical margin, and by the longer and more acute postero-median process of the female sternite VIII. D i s t r i b u t i o n a n d n a t u r a l h i s t o r y: The currently known distribution is confined to the western Daba Shan from the environs of Zhenping in southern Shaanxi to the Wu Shan in northern Chongqing (Map 3). Previous records from Sichuan are incorrect. The material listed above was sifted from leaf litter in various forest habitats at altitudes between 1680 and 2600 m.	en	Assing, Volker (2015): On the Harpopaederus fauna of China (Coleoptera: Staphylinidae: Paederinae). Linzer biologische Beiträge 47 (1): 163-190, DOI: 10.5281/zenodo.5414042
