taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
F65487889905FF9AFF33FB797164F891.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8345483/files/figure.png	https://doi.org/10.5281/zenodo.8345483	FIGURE 2. Trematosoma husselae sp. nov. on Setaphyes kielensis. A. Drawing of the holotype with internal developing swarmer ventrally on segment 1 of basibiont (ZMB 12499). B. Paratype in side view ventrally on segment 3 of basibiont (ZMB 12489). Arrow marks apical cleft within lorica. Abbreviations: ca, basal cavity in lorica; lo, lorica; ma, macronucleus; mi, micronucleus; st, stalk with internal striation (information taken from different specimen); sw, swarmer; te, tentacle; zo, zooid. Scale bar 30 µm.	FIGURE 2. Trematosoma husselae sp. nov. on Setaphyes kielensis. A. Drawing of the holotype with internal developing swarmer ventrally on segment 1 of basibiont (ZMB 12499). B. Paratype in side view ventrally on segment 3 of basibiont (ZMB 12489). Arrow marks apical cleft within lorica. Abbreviations: ca, basal cavity in lorica; lo, lorica; ma, macronucleus; mi, micronucleus; st, stalk with internal striation (information taken from different specimen); sw, swarmer; te, tentacle; zo, zooid. Scale bar 30 µm.	2023-09-12	Neuhaus, Birger;Lu, Borong;Yamasaki, Hiroshi;González-Casarrubios, Alberto		Zenodo	biologists	Neuhaus, Birger;Lu, Borong;Yamasaki, Hiroshi;González-Casarrubios, Alberto			
F65487889905FF9AFF33FB797164F891.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8345485/files/figure.png	https://doi.org/10.5281/zenodo.8345485	FIGURE 3. Setaphyes kielensis (A–C, ZMB 12475; D, ZMB 12464; E, F, ZMB 12465; G, ZMB 12462) with specimens of Trematosoma husselae sp. nov., SEM, ventral view. A–C. Specimen with 43 epibionts, segments 1–4 (A), 5–7 (B), and 8–11 (C). D. Segment 8, epibiont surface. E. Segment 6, epibiont surface covered with detritus. F. Segment 8, frontal view on small spherical specimen of Trematosoma husselae sp. nov. G. Segment 10, arrow marks specimen with withdrawn tentacles. Abbreviations: gco, gland cell outlet; lts, lateral terminal spine; s, segment, followed by segment number; se, seta; su, suctorian epibiont; te, tentacle. Scale bar in A–C 40 µm, in D–F 10 µm, and in G 20 µm.	FIGURE 3. Setaphyes kielensis (A–C, ZMB 12475; D, ZMB 12464; E, F, ZMB 12465; G, ZMB 12462) with specimens of Trematosoma husselae sp. nov., SEM, ventral view. A–C. Specimen with 43 epibionts, segments 1–4 (A), 5–7 (B), and 8–11 (C). D. Segment 8, epibiont surface. E. Segment 6, epibiont surface covered with detritus. F. Segment 8, frontal view on small spherical specimen of Trematosoma husselae sp. nov. G. Segment 10, arrow marks specimen with withdrawn tentacles. Abbreviations: gco, gland cell outlet; lts, lateral terminal spine; s, segment, followed by segment number; se, seta; su, suctorian epibiont; te, tentacle. Scale bar in A–C 40 µm, in D–F 10 µm, and in G 20 µm.	2023-09-12	Neuhaus, Birger;Lu, Borong;Yamasaki, Hiroshi;González-Casarrubios, Alberto		Zenodo	biologists	Neuhaus, Birger;Lu, Borong;Yamasaki, Hiroshi;González-Casarrubios, Alberto			
F65487889905FF9AFF33FB797164F891.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8345487/files/figure.png	https://doi.org/10.5281/zenodo.8345487	FIGURE 4. Setaphyes kielensis (A, ZMB 12469; B, ZMB 12489; C, ZMB 12486; D, E, ZMB 12492; F, ZMB 12488) with specimens of Trematosoma husselae sp. nov., SEM (A), and DIC (B–F). A. Segment 1, empty shells of epibionts, ventral view. B. Segments 1–11, optical section. C. Epibionts with capitated tentacles arranged in left and right row. Extended focus image of three images. D. Epibiont with developing swarmer. Arrowheads mark membrane of swarmer. E. Epibiont with swarmer recently freed from its parent. F. Segments 4–6, ventral view. Note thickened lorica in basal two thirds of epibionts.Arrow marks longitudinal striation of stalk of epibiont. Abbreviations: ca, basal cavity in lorica; co, specimen of Cothurnia buetschlii; lo, lorica; lts, lateral terminal spine; ma, macronucleus; mi, micronucleus; s, segment, followed by segment number; su, suctorian epibiont; sw, swarmer; te, tentacle. Scale bar in A, D, E 20 µm, in B 100 µm, in C 30 µm, and in F 50 µm.	FIGURE 4. Setaphyes kielensis (A, ZMB 12469; B, ZMB 12489; C, ZMB 12486; D, E, ZMB 12492; F, ZMB 12488) with specimens of Trematosoma husselae sp. nov., SEM (A), and DIC (B–F). A. Segment 1, empty shells of epibionts, ventral view. B. Segments 1–11, optical section. C. Epibionts with capitated tentacles arranged in left and right row. Extended focus image of three images. D. Epibiont with developing swarmer. Arrowheads mark membrane of swarmer. E. Epibiont with swarmer recently freed from its parent. F. Segments 4–6, ventral view. Note thickened lorica in basal two thirds of epibionts.Arrow marks longitudinal striation of stalk of epibiont. Abbreviations: ca, basal cavity in lorica; co, specimen of Cothurnia buetschlii; lo, lorica; lts, lateral terminal spine; ma, macronucleus; mi, micronucleus; s, segment, followed by segment number; su, suctorian epibiont; sw, swarmer; te, tentacle. Scale bar in A, D, E 20 µm, in B 100 µm, in C 30 µm, and in F 50 µm.	2023-09-12	Neuhaus, Birger;Lu, Borong;Yamasaki, Hiroshi;González-Casarrubios, Alberto		Zenodo	biologists	Neuhaus, Birger;Lu, Borong;Yamasaki, Hiroshi;González-Casarrubios, Alberto			
F65487889905FF9AFF33FB797164F891.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8349223/files/figure.png	https://doi.org/10.5281/zenodo.8349223	FIGURE 5. Setaphyes kielensis (A, ZMB 12489; B, ZMB 12500; C, ZMB 12499; D, ZMB 12487; J, ZMB 12486; L, ZMB 12397; M, ZMB 12369) and S. dentatus (E–G, K, ZMB 12503; F, H, I, ZMB 12494) with specimens of Trematosoma husselae sp. nov. (A–G, J–M), Cothurnia buetschlii (J), and a case of fungus or developmental artefact (H, I, ZMB 12494), DIC. A. Segments 1–4, lateral and dorso-ventral view on the calyciform, dorso-ventrally compressed lorica with the zooid, ventrolateral view on basibiont. B. Epibiont with swarmer, stalk, and lorica in optical section. C. Epibiont with two macronuclei not fusing at any focal level. D, E. Young, small, spherical epibionts with few tentacles, not all tentacles in focus. F. Young specimen on cuticle of basibiont with striation pattern on surface (arrows). G. Epibiont with tentacles in empty shell of suctorian epibiont. H, I. Segment 10, dorsally (H) and optical section (I), subcuticular fungus or internal sclerotisation artefact of the cuticle (arrows). J. Assemblage of epibionts on segment 10 of basibiont with C. buetschlii (contracted specimen left, extended specimen right), T. husselae sp. nov., and two kinds of filamentous bacteria. Arrows mark endostyle. K. Suctorian on segment 7 of S. dentatus. L, M. Specimens collected 1998, showing cellular substructures (L) and withdrawn zooid with minimum amount of cellular substructures (M). Extended focus images of six (D), eight (H, L), and three (K) images. Arrowheads mark stalks in B, D, E, G, and J. Abbreviations: ba, bacteria; ca, basal cavity in lorica; co, C. buetschlii; en, endostyle; lo, lorica; ma, macronucleus; s, segment, followed by segment number; sc, spherical cavity; sp, sperm; st, stalk; su, suctorian epibiont; sw, swarmer; te, tentacle; tu, tube on segment 2. Scale bar in A 50 µm, in B 20 µm, valid for B, G–J and M, in C 20 µm, valid for C, K, and L, and in D–F 10 µm.	FIGURE 5. Setaphyes kielensis (A, ZMB 12489; B, ZMB 12500; C, ZMB 12499; D, ZMB 12487; J, ZMB 12486; L, ZMB 12397; M, ZMB 12369) and S. dentatus (E–G, K, ZMB 12503; F, H, I, ZMB 12494) with specimens of Trematosoma husselae sp. nov. (A–G, J–M), Cothurnia buetschlii (J), and a case of fungus or developmental artefact (H, I, ZMB 12494), DIC. A. Segments 1–4, lateral and dorso-ventral view on the calyciform, dorso-ventrally compressed lorica with the zooid, ventrolateral view on basibiont. B. Epibiont with swarmer, stalk, and lorica in optical section. C. Epibiont with two macronuclei not fusing at any focal level. D, E. Young, small, spherical epibionts with few tentacles, not all tentacles in focus. F. Young specimen on cuticle of basibiont with striation pattern on surface (arrows). G. Epibiont with tentacles in empty shell of suctorian epibiont. H, I. Segment 10, dorsally (H) and optical section (I), subcuticular fungus or internal sclerotisation artefact of the cuticle (arrows). J. Assemblage of epibionts on segment 10 of basibiont with C. buetschlii (contracted specimen left, extended specimen right), T. husselae sp. nov., and two kinds of filamentous bacteria. Arrows mark endostyle. K. Suctorian on segment 7 of S. dentatus. L, M. Specimens collected 1998, showing cellular substructures (L) and withdrawn zooid with minimum amount of cellular substructures (M). Extended focus images of six (D), eight (H, L), and three (K) images. Arrowheads mark stalks in B, D, E, G, and J. Abbreviations: ba, bacteria; ca, basal cavity in lorica; co, C. buetschlii; en, endostyle; lo, lorica; ma, macronucleus; s, segment, followed by segment number; sc, spherical cavity; sp, sperm; st, stalk; su, suctorian epibiont; sw, swarmer; te, tentacle; tu, tube on segment 2. Scale bar in A 50 µm, in B 20 µm, valid for B, G–J and M, in C 20 µm, valid for C, K, and L, and in D–F 10 µm.	2023-09-12	Neuhaus, Birger;Lu, Borong;Yamasaki, Hiroshi;González-Casarrubios, Alberto		Zenodo	biologists	Neuhaus, Birger;Lu, Borong;Yamasaki, Hiroshi;González-Casarrubios, Alberto			
F65487889905FF9AFF33FB797164F891.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8345491/files/figure.png	https://doi.org/10.5281/zenodo.8345491	FIGURE 7. Setaphyes kielensis (ZMB 12468) with epibionts, SEM. A. Segments 1–11, ventral-right. B. Segments 4–8, ventral view. Arrows mark laterally recessed anterior margin of lorica of Cothurnia buetschlii. Abbreviations: co, Cothurnia buetschlii; fp, filamentous prokaryotes; lts, lateral terminal spine; s, segment, followed by segment number; se, seta; su, suctorian epibiont; tu, tube. Scale bar in A 100 µm and in B 20 µm.	FIGURE 7. Setaphyes kielensis (ZMB 12468) with epibionts, SEM. A. Segments 1–11, ventral-right. B. Segments 4–8, ventral view. Arrows mark laterally recessed anterior margin of lorica of Cothurnia buetschlii. Abbreviations: co, Cothurnia buetschlii; fp, filamentous prokaryotes; lts, lateral terminal spine; s, segment, followed by segment number; se, seta; su, suctorian epibiont; tu, tube. Scale bar in A 100 µm and in B 20 µm.	2023-09-12	Neuhaus, Birger;Lu, Borong;Yamasaki, Hiroshi;González-Casarrubios, Alberto		Zenodo	biologists	Neuhaus, Birger;Lu, Borong;Yamasaki, Hiroshi;González-Casarrubios, Alberto			
F65487889905FF9AFF33FB797164F891.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8345493/files/figure.png	https://doi.org/10.5281/zenodo.8345493	FIGURE 8. Setaphyes kielensis (A, ZMB 12468; B, ZMB 12474) with epibionts, SEM. A. Segment 6, left side. Arrow marks laterally recessed anterior margin of lorica of Cothurnia buetschlii. B. Segments 8–10, dorsal view.Abbreviations: co, Cothurnia buetschlii; fp, filamentous prokaryotes; s, segment, followed by segment number; su, suctorian epibiont; te, tentacles. Scale bar in A 10 µm and in B 20 µm.	FIGURE 8. Setaphyes kielensis (A, ZMB 12468; B, ZMB 12474) with epibionts, SEM. A. Segment 6, left side. Arrow marks laterally recessed anterior margin of lorica of Cothurnia buetschlii. B. Segments 8–10, dorsal view.Abbreviations: co, Cothurnia buetschlii; fp, filamentous prokaryotes; s, segment, followed by segment number; su, suctorian epibiont; te, tentacles. Scale bar in A 10 µm and in B 20 µm.	2023-09-12	Neuhaus, Birger;Lu, Borong;Yamasaki, Hiroshi;González-Casarrubios, Alberto		Zenodo	biologists	Neuhaus, Birger;Lu, Borong;Yamasaki, Hiroshi;González-Casarrubios, Alberto			
F65487889905FF9AFF33FB797164F891.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8345501/files/figure.png	https://doi.org/10.5281/zenodo.8345501	FIGURE 10. Setaphyes kielensis (A–C, ZMB 12458; D, ZMB 12488) with cuticular marks and filamentous prokaryotes, SEM (A–C) and DIC (D). A. Cuticular marks (arrows) of detached ciliate specimens, laterodorsal view. B. Segment 1 with large mucous? attachment sites to basibiont (arrows), frontal view. C. Segments 7–11, ventral view. D. Segments 8–10, ventral view. Abbreviations: co, Cothurnia buetschlii; fp, filamentous prokaryotes; lts, lateral terminal spine; pl, placid; s, segment, followed by segment number; su, suctorian epibiont. Scale bar in A–C 20 µm and in D 50 µm.	FIGURE 10. Setaphyes kielensis (A–C, ZMB 12458; D, ZMB 12488) with cuticular marks and filamentous prokaryotes, SEM (A–C) and DIC (D). A. Cuticular marks (arrows) of detached ciliate specimens, laterodorsal view. B. Segment 1 with large mucous? attachment sites to basibiont (arrows), frontal view. C. Segments 7–11, ventral view. D. Segments 8–10, ventral view. Abbreviations: co, Cothurnia buetschlii; fp, filamentous prokaryotes; lts, lateral terminal spine; pl, placid; s, segment, followed by segment number; su, suctorian epibiont. Scale bar in A–C 20 µm and in D 50 µm.	2023-09-12	Neuhaus, Birger;Lu, Borong;Yamasaki, Hiroshi;González-Casarrubios, Alberto		Zenodo	biologists	Neuhaus, Birger;Lu, Borong;Yamasaki, Hiroshi;González-Casarrubios, Alberto			
F6548788990FFF82FF33FA8E74FCFC82.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8349223/files/figure.png	https://doi.org/10.5281/zenodo.8349223	FIGURE 5. Setaphyes kielensis (A, ZMB 12489; B, ZMB 12500; C, ZMB 12499; D, ZMB 12487; J, ZMB 12486; L, ZMB 12397; M, ZMB 12369) and S. dentatus (E–G, K, ZMB 12503; F, H, I, ZMB 12494) with specimens of Trematosoma husselae sp. nov. (A–G, J–M), Cothurnia buetschlii (J), and a case of fungus or developmental artefact (H, I, ZMB 12494), DIC. A. Segments 1–4, lateral and dorso-ventral view on the calyciform, dorso-ventrally compressed lorica with the zooid, ventrolateral view on basibiont. B. Epibiont with swarmer, stalk, and lorica in optical section. C. Epibiont with two macronuclei not fusing at any focal level. D, E. Young, small, spherical epibionts with few tentacles, not all tentacles in focus. F. Young specimen on cuticle of basibiont with striation pattern on surface (arrows). G. Epibiont with tentacles in empty shell of suctorian epibiont. H, I. Segment 10, dorsally (H) and optical section (I), subcuticular fungus or internal sclerotisation artefact of the cuticle (arrows). J. Assemblage of epibionts on segment 10 of basibiont with C. buetschlii (contracted specimen left, extended specimen right), T. husselae sp. nov., and two kinds of filamentous bacteria. Arrows mark endostyle. K. Suctorian on segment 7 of S. dentatus. L, M. Specimens collected 1998, showing cellular substructures (L) and withdrawn zooid with minimum amount of cellular substructures (M). Extended focus images of six (D), eight (H, L), and three (K) images. Arrowheads mark stalks in B, D, E, G, and J. Abbreviations: ba, bacteria; ca, basal cavity in lorica; co, C. buetschlii; en, endostyle; lo, lorica; ma, macronucleus; s, segment, followed by segment number; sc, spherical cavity; sp, sperm; st, stalk; su, suctorian epibiont; sw, swarmer; te, tentacle; tu, tube on segment 2. Scale bar in A 50 µm, in B 20 µm, valid for B, G–J and M, in C 20 µm, valid for C, K, and L, and in D–F 10 µm.	FIGURE 5. Setaphyes kielensis (A, ZMB 12489; B, ZMB 12500; C, ZMB 12499; D, ZMB 12487; J, ZMB 12486; L, ZMB 12397; M, ZMB 12369) and S. dentatus (E–G, K, ZMB 12503; F, H, I, ZMB 12494) with specimens of Trematosoma husselae sp. nov. (A–G, J–M), Cothurnia buetschlii (J), and a case of fungus or developmental artefact (H, I, ZMB 12494), DIC. A. Segments 1–4, lateral and dorso-ventral view on the calyciform, dorso-ventrally compressed lorica with the zooid, ventrolateral view on basibiont. B. Epibiont with swarmer, stalk, and lorica in optical section. C. Epibiont with two macronuclei not fusing at any focal level. D, E. Young, small, spherical epibionts with few tentacles, not all tentacles in focus. F. Young specimen on cuticle of basibiont with striation pattern on surface (arrows). G. Epibiont with tentacles in empty shell of suctorian epibiont. H, I. Segment 10, dorsally (H) and optical section (I), subcuticular fungus or internal sclerotisation artefact of the cuticle (arrows). J. Assemblage of epibionts on segment 10 of basibiont with C. buetschlii (contracted specimen left, extended specimen right), T. husselae sp. nov., and two kinds of filamentous bacteria. Arrows mark endostyle. K. Suctorian on segment 7 of S. dentatus. L, M. Specimens collected 1998, showing cellular substructures (L) and withdrawn zooid with minimum amount of cellular substructures (M). Extended focus images of six (D), eight (H, L), and three (K) images. Arrowheads mark stalks in B, D, E, G, and J. Abbreviations: ba, bacteria; ca, basal cavity in lorica; co, C. buetschlii; en, endostyle; lo, lorica; ma, macronucleus; s, segment, followed by segment number; sc, spherical cavity; sp, sperm; st, stalk; su, suctorian epibiont; sw, swarmer; te, tentacle; tu, tube on segment 2. Scale bar in A 50 µm, in B 20 µm, valid for B, G–J and M, in C 20 µm, valid for C, K, and L, and in D–F 10 µm.	2023-09-12	Neuhaus, Birger;Lu, Borong;Yamasaki, Hiroshi;González-Casarrubios, Alberto		Zenodo	biologists	Neuhaus, Birger;Lu, Borong;Yamasaki, Hiroshi;González-Casarrubios, Alberto			
F6548788990FFF82FF33FA8E74FCFC82.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8345489/files/figure.png	https://doi.org/10.5281/zenodo.8345489	FIGURE 6. A, B. Drawing of Cothurnia buetschlii (specimen 2) ventrally on segment 10 of Setaphyes kielensis (ZMB 12486) as longitudinal optical section (A) and lorica in cross-section (B). Abbreviations: an, annulation on surface of zooid; en, endostyle with striae; inf, withdrawn infundibulum with cilia; lo, lorica; ma, macronucleus; mi, micronucleus; po, pore in cementing mass; sc, spherical cavity; st, stalk with internal striae; zo, zooid. Scale bar 30 µm.	FIGURE 6. A, B. Drawing of Cothurnia buetschlii (specimen 2) ventrally on segment 10 of Setaphyes kielensis (ZMB 12486) as longitudinal optical section (A) and lorica in cross-section (B). Abbreviations: an, annulation on surface of zooid; en, endostyle with striae; inf, withdrawn infundibulum with cilia; lo, lorica; ma, macronucleus; mi, micronucleus; po, pore in cementing mass; sc, spherical cavity; st, stalk with internal striae; zo, zooid. Scale bar 30 µm.	2023-09-12	Neuhaus, Birger;Lu, Borong;Yamasaki, Hiroshi;González-Casarrubios, Alberto		Zenodo	biologists	Neuhaus, Birger;Lu, Borong;Yamasaki, Hiroshi;González-Casarrubios, Alberto			
F6548788990FFF82FF33FA8E74FCFC82.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8345491/files/figure.png	https://doi.org/10.5281/zenodo.8345491	FIGURE 7. Setaphyes kielensis (ZMB 12468) with epibionts, SEM. A. Segments 1–11, ventral-right. B. Segments 4–8, ventral view. Arrows mark laterally recessed anterior margin of lorica of Cothurnia buetschlii. Abbreviations: co, Cothurnia buetschlii; fp, filamentous prokaryotes; lts, lateral terminal spine; s, segment, followed by segment number; se, seta; su, suctorian epibiont; tu, tube. Scale bar in A 100 µm and in B 20 µm.	FIGURE 7. Setaphyes kielensis (ZMB 12468) with epibionts, SEM. A. Segments 1–11, ventral-right. B. Segments 4–8, ventral view. Arrows mark laterally recessed anterior margin of lorica of Cothurnia buetschlii. Abbreviations: co, Cothurnia buetschlii; fp, filamentous prokaryotes; lts, lateral terminal spine; s, segment, followed by segment number; se, seta; su, suctorian epibiont; tu, tube. Scale bar in A 100 µm and in B 20 µm.	2023-09-12	Neuhaus, Birger;Lu, Borong;Yamasaki, Hiroshi;González-Casarrubios, Alberto		Zenodo	biologists	Neuhaus, Birger;Lu, Borong;Yamasaki, Hiroshi;González-Casarrubios, Alberto			
F6548788990FFF82FF33FA8E74FCFC82.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8345493/files/figure.png	https://doi.org/10.5281/zenodo.8345493	FIGURE 8. Setaphyes kielensis (A, ZMB 12468; B, ZMB 12474) with epibionts, SEM. A. Segment 6, left side. Arrow marks laterally recessed anterior margin of lorica of Cothurnia buetschlii. B. Segments 8–10, dorsal view.Abbreviations: co, Cothurnia buetschlii; fp, filamentous prokaryotes; s, segment, followed by segment number; su, suctorian epibiont; te, tentacles. Scale bar in A 10 µm and in B 20 µm.	FIGURE 8. Setaphyes kielensis (A, ZMB 12468; B, ZMB 12474) with epibionts, SEM. A. Segment 6, left side. Arrow marks laterally recessed anterior margin of lorica of Cothurnia buetschlii. B. Segments 8–10, dorsal view.Abbreviations: co, Cothurnia buetschlii; fp, filamentous prokaryotes; s, segment, followed by segment number; su, suctorian epibiont; te, tentacles. Scale bar in A 10 µm and in B 20 µm.	2023-09-12	Neuhaus, Birger;Lu, Borong;Yamasaki, Hiroshi;González-Casarrubios, Alberto		Zenodo	biologists	Neuhaus, Birger;Lu, Borong;Yamasaki, Hiroshi;González-Casarrubios, Alberto			
F6548788990FFF82FF33FA8E74FCFC82.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8345497/files/figure.png	https://doi.org/10.5281/zenodo.8345497	FIGURE 9. Setaphyes kielensis (ZMB 12486), segment 10 with Cothurnia buetschlii (specimens 1–4), ventral view. A. Overview of position of epibionts. Numbers indicate specimens of C. buetschlii (comp. also Table 6). B–G. Specimens 3 (B) and 4 (C–G), images 8 (C), 15 (D), 19 (E), 24 (F) and 28 (G) showing surface annulation of zooid (C, D), internal longitudinal striae of stalk and endostyle (D–F), and attachment site of stalk to the cuticle of the basibiont (G, arrowheads). Note elongate macronucleus and numerous micronuclei (E–G). White arrows (D, E) mark basal outer margin of lorica, black arrows (D, E) point to bottom margin of lorica. Abbreviations: an, annulation on surface of zooid; ci, ciliae; en, endostyle; in, withdrawn infundibulum; lo, lorica; lts, lateral terminal spine; ma, macronucleus; mi, micronucleus; ps, penile spine; sc, spherical cavity; st, stalk; su, suctorian epibiont; zo, zooid. Scale bar in A 50 µm and in D 20 µm, valid for B–G.	FIGURE 9. Setaphyes kielensis (ZMB 12486), segment 10 with Cothurnia buetschlii (specimens 1–4), ventral view. A. Overview of position of epibionts. Numbers indicate specimens of C. buetschlii (comp. also Table 6). B–G. Specimens 3 (B) and 4 (C–G), images 8 (C), 15 (D), 19 (E), 24 (F) and 28 (G) showing surface annulation of zooid (C, D), internal longitudinal striae of stalk and endostyle (D–F), and attachment site of stalk to the cuticle of the basibiont (G, arrowheads). Note elongate macronucleus and numerous micronuclei (E–G). White arrows (D, E) mark basal outer margin of lorica, black arrows (D, E) point to bottom margin of lorica. Abbreviations: an, annulation on surface of zooid; ci, ciliae; en, endostyle; in, withdrawn infundibulum; lo, lorica; lts, lateral terminal spine; ma, macronucleus; mi, micronucleus; ps, penile spine; sc, spherical cavity; st, stalk; su, suctorian epibiont; zo, zooid. Scale bar in A 50 µm and in D 20 µm, valid for B–G.	2023-09-12	Neuhaus, Birger;Lu, Borong;Yamasaki, Hiroshi;González-Casarrubios, Alberto		Zenodo	biologists	Neuhaus, Birger;Lu, Borong;Yamasaki, Hiroshi;González-Casarrubios, Alberto			
F6548788990FFF82FF33FA8E74FCFC82.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8345481/files/figure.png	https://doi.org/10.5281/zenodo.8345481	FIGURE 1. Map showing the locations of Pycnophyes communis, Setaphyes dentatus, and S. kielensis in European waters and the sampling site of S. dentatus and S. kielensis in the North Sea, island of Sylt (asterisks in inset A).	FIGURE 1. Map showing the locations of Pycnophyes communis, Setaphyes dentatus, and S. kielensis in European waters and the sampling site of S. dentatus and S. kielensis in the North Sea, island of Sylt (asterisks in inset A).	2023-09-12	Neuhaus, Birger;Lu, Borong;Yamasaki, Hiroshi;González-Casarrubios, Alberto		Zenodo	biologists	Neuhaus, Birger;Lu, Borong;Yamasaki, Hiroshi;González-Casarrubios, Alberto			
F65487889912FF86FF33FB4474BAFF6E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8345491/files/figure.png	https://doi.org/10.5281/zenodo.8345491	FIGURE 7. Setaphyes kielensis (ZMB 12468) with epibionts, SEM. A. Segments 1–11, ventral-right. B. Segments 4–8, ventral view. Arrows mark laterally recessed anterior margin of lorica of Cothurnia buetschlii. Abbreviations: co, Cothurnia buetschlii; fp, filamentous prokaryotes; lts, lateral terminal spine; s, segment, followed by segment number; se, seta; su, suctorian epibiont; tu, tube. Scale bar in A 100 µm and in B 20 µm.	FIGURE 7. Setaphyes kielensis (ZMB 12468) with epibionts, SEM. A. Segments 1–11, ventral-right. B. Segments 4–8, ventral view. Arrows mark laterally recessed anterior margin of lorica of Cothurnia buetschlii. Abbreviations: co, Cothurnia buetschlii; fp, filamentous prokaryotes; lts, lateral terminal spine; s, segment, followed by segment number; se, seta; su, suctorian epibiont; tu, tube. Scale bar in A 100 µm and in B 20 µm.	2023-09-12	Neuhaus, Birger;Lu, Borong;Yamasaki, Hiroshi;González-Casarrubios, Alberto		Zenodo	biologists	Neuhaus, Birger;Lu, Borong;Yamasaki, Hiroshi;González-Casarrubios, Alberto			
F65487889912FF86FF33FB4474BAFF6E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8345493/files/figure.png	https://doi.org/10.5281/zenodo.8345493	FIGURE 8. Setaphyes kielensis (A, ZMB 12468; B, ZMB 12474) with epibionts, SEM. A. Segment 6, left side. Arrow marks laterally recessed anterior margin of lorica of Cothurnia buetschlii. B. Segments 8–10, dorsal view.Abbreviations: co, Cothurnia buetschlii; fp, filamentous prokaryotes; s, segment, followed by segment number; su, suctorian epibiont; te, tentacles. Scale bar in A 10 µm and in B 20 µm.	FIGURE 8. Setaphyes kielensis (A, ZMB 12468; B, ZMB 12474) with epibionts, SEM. A. Segment 6, left side. Arrow marks laterally recessed anterior margin of lorica of Cothurnia buetschlii. B. Segments 8–10, dorsal view.Abbreviations: co, Cothurnia buetschlii; fp, filamentous prokaryotes; s, segment, followed by segment number; su, suctorian epibiont; te, tentacles. Scale bar in A 10 µm and in B 20 µm.	2023-09-12	Neuhaus, Birger;Lu, Borong;Yamasaki, Hiroshi;González-Casarrubios, Alberto		Zenodo	biologists	Neuhaus, Birger;Lu, Borong;Yamasaki, Hiroshi;González-Casarrubios, Alberto			
F65487889912FF86FF33FB4474BAFF6E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8345497/files/figure.png	https://doi.org/10.5281/zenodo.8345497	FIGURE 9. Setaphyes kielensis (ZMB 12486), segment 10 with Cothurnia buetschlii (specimens 1–4), ventral view. A. Overview of position of epibionts. Numbers indicate specimens of C. buetschlii (comp. also Table 6). B–G. Specimens 3 (B) and 4 (C–G), images 8 (C), 15 (D), 19 (E), 24 (F) and 28 (G) showing surface annulation of zooid (C, D), internal longitudinal striae of stalk and endostyle (D–F), and attachment site of stalk to the cuticle of the basibiont (G, arrowheads). Note elongate macronucleus and numerous micronuclei (E–G). White arrows (D, E) mark basal outer margin of lorica, black arrows (D, E) point to bottom margin of lorica. Abbreviations: an, annulation on surface of zooid; ci, ciliae; en, endostyle; in, withdrawn infundibulum; lo, lorica; lts, lateral terminal spine; ma, macronucleus; mi, micronucleus; ps, penile spine; sc, spherical cavity; st, stalk; su, suctorian epibiont; zo, zooid. Scale bar in A 50 µm and in D 20 µm, valid for B–G.	FIGURE 9. Setaphyes kielensis (ZMB 12486), segment 10 with Cothurnia buetschlii (specimens 1–4), ventral view. A. Overview of position of epibionts. Numbers indicate specimens of C. buetschlii (comp. also Table 6). B–G. Specimens 3 (B) and 4 (C–G), images 8 (C), 15 (D), 19 (E), 24 (F) and 28 (G) showing surface annulation of zooid (C, D), internal longitudinal striae of stalk and endostyle (D–F), and attachment site of stalk to the cuticle of the basibiont (G, arrowheads). Note elongate macronucleus and numerous micronuclei (E–G). White arrows (D, E) mark basal outer margin of lorica, black arrows (D, E) point to bottom margin of lorica. Abbreviations: an, annulation on surface of zooid; ci, ciliae; en, endostyle; in, withdrawn infundibulum; lo, lorica; lts, lateral terminal spine; ma, macronucleus; mi, micronucleus; ps, penile spine; sc, spherical cavity; st, stalk; su, suctorian epibiont; zo, zooid. Scale bar in A 50 µm and in D 20 µm, valid for B–G.	2023-09-12	Neuhaus, Birger;Lu, Borong;Yamasaki, Hiroshi;González-Casarrubios, Alberto		Zenodo	biologists	Neuhaus, Birger;Lu, Borong;Yamasaki, Hiroshi;González-Casarrubios, Alberto			
F65487889912FF86FF33FB4474BAFF6E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8349223/files/figure.png	https://doi.org/10.5281/zenodo.8349223	FIGURE 5. Setaphyes kielensis (A, ZMB 12489; B, ZMB 12500; C, ZMB 12499; D, ZMB 12487; J, ZMB 12486; L, ZMB 12397; M, ZMB 12369) and S. dentatus (E–G, K, ZMB 12503; F, H, I, ZMB 12494) with specimens of Trematosoma husselae sp. nov. (A–G, J–M), Cothurnia buetschlii (J), and a case of fungus or developmental artefact (H, I, ZMB 12494), DIC. A. Segments 1–4, lateral and dorso-ventral view on the calyciform, dorso-ventrally compressed lorica with the zooid, ventrolateral view on basibiont. B. Epibiont with swarmer, stalk, and lorica in optical section. C. Epibiont with two macronuclei not fusing at any focal level. D, E. Young, small, spherical epibionts with few tentacles, not all tentacles in focus. F. Young specimen on cuticle of basibiont with striation pattern on surface (arrows). G. Epibiont with tentacles in empty shell of suctorian epibiont. H, I. Segment 10, dorsally (H) and optical section (I), subcuticular fungus or internal sclerotisation artefact of the cuticle (arrows). J. Assemblage of epibionts on segment 10 of basibiont with C. buetschlii (contracted specimen left, extended specimen right), T. husselae sp. nov., and two kinds of filamentous bacteria. Arrows mark endostyle. K. Suctorian on segment 7 of S. dentatus. L, M. Specimens collected 1998, showing cellular substructures (L) and withdrawn zooid with minimum amount of cellular substructures (M). Extended focus images of six (D), eight (H, L), and three (K) images. Arrowheads mark stalks in B, D, E, G, and J. Abbreviations: ba, bacteria; ca, basal cavity in lorica; co, C. buetschlii; en, endostyle; lo, lorica; ma, macronucleus; s, segment, followed by segment number; sc, spherical cavity; sp, sperm; st, stalk; su, suctorian epibiont; sw, swarmer; te, tentacle; tu, tube on segment 2. Scale bar in A 50 µm, in B 20 µm, valid for B, G–J and M, in C 20 µm, valid for C, K, and L, and in D–F 10 µm.	FIGURE 5. Setaphyes kielensis (A, ZMB 12489; B, ZMB 12500; C, ZMB 12499; D, ZMB 12487; J, ZMB 12486; L, ZMB 12397; M, ZMB 12369) and S. dentatus (E–G, K, ZMB 12503; F, H, I, ZMB 12494) with specimens of Trematosoma husselae sp. nov. (A–G, J–M), Cothurnia buetschlii (J), and a case of fungus or developmental artefact (H, I, ZMB 12494), DIC. A. Segments 1–4, lateral and dorso-ventral view on the calyciform, dorso-ventrally compressed lorica with the zooid, ventrolateral view on basibiont. B. Epibiont with swarmer, stalk, and lorica in optical section. C. Epibiont with two macronuclei not fusing at any focal level. D, E. Young, small, spherical epibionts with few tentacles, not all tentacles in focus. F. Young specimen on cuticle of basibiont with striation pattern on surface (arrows). G. Epibiont with tentacles in empty shell of suctorian epibiont. H, I. Segment 10, dorsally (H) and optical section (I), subcuticular fungus or internal sclerotisation artefact of the cuticle (arrows). J. Assemblage of epibionts on segment 10 of basibiont with C. buetschlii (contracted specimen left, extended specimen right), T. husselae sp. nov., and two kinds of filamentous bacteria. Arrows mark endostyle. K. Suctorian on segment 7 of S. dentatus. L, M. Specimens collected 1998, showing cellular substructures (L) and withdrawn zooid with minimum amount of cellular substructures (M). Extended focus images of six (D), eight (H, L), and three (K) images. Arrowheads mark stalks in B, D, E, G, and J. Abbreviations: ba, bacteria; ca, basal cavity in lorica; co, C. buetschlii; en, endostyle; lo, lorica; ma, macronucleus; s, segment, followed by segment number; sc, spherical cavity; sp, sperm; st, stalk; su, suctorian epibiont; sw, swarmer; te, tentacle; tu, tube on segment 2. Scale bar in A 50 µm, in B 20 µm, valid for B, G–J and M, in C 20 µm, valid for C, K, and L, and in D–F 10 µm.	2023-09-12	Neuhaus, Birger;Lu, Borong;Yamasaki, Hiroshi;González-Casarrubios, Alberto		Zenodo	biologists	Neuhaus, Birger;Lu, Borong;Yamasaki, Hiroshi;González-Casarrubios, Alberto			
F65487889916FF8AFF33F9937438FDAA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8345483/files/figure.png	https://doi.org/10.5281/zenodo.8345483	FIGURE 2. Trematosoma husselae sp. nov. on Setaphyes kielensis. A. Drawing of the holotype with internal developing swarmer ventrally on segment 1 of basibiont (ZMB 12499). B. Paratype in side view ventrally on segment 3 of basibiont (ZMB 12489). Arrow marks apical cleft within lorica. Abbreviations: ca, basal cavity in lorica; lo, lorica; ma, macronucleus; mi, micronucleus; st, stalk with internal striation (information taken from different specimen); sw, swarmer; te, tentacle; zo, zooid. Scale bar 30 µm.	FIGURE 2. Trematosoma husselae sp. nov. on Setaphyes kielensis. A. Drawing of the holotype with internal developing swarmer ventrally on segment 1 of basibiont (ZMB 12499). B. Paratype in side view ventrally on segment 3 of basibiont (ZMB 12489). Arrow marks apical cleft within lorica. Abbreviations: ca, basal cavity in lorica; lo, lorica; ma, macronucleus; mi, micronucleus; st, stalk with internal striation (information taken from different specimen); sw, swarmer; te, tentacle; zo, zooid. Scale bar 30 µm.	2023-09-12	Neuhaus, Birger;Lu, Borong;Yamasaki, Hiroshi;González-Casarrubios, Alberto		Zenodo	biologists	Neuhaus, Birger;Lu, Borong;Yamasaki, Hiroshi;González-Casarrubios, Alberto			
F65487889916FF8AFF33F9937438FDAA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8345487/files/figure.png	https://doi.org/10.5281/zenodo.8345487	FIGURE 4. Setaphyes kielensis (A, ZMB 12469; B, ZMB 12489; C, ZMB 12486; D, E, ZMB 12492; F, ZMB 12488) with specimens of Trematosoma husselae sp. nov., SEM (A), and DIC (B–F). A. Segment 1, empty shells of epibionts, ventral view. B. Segments 1–11, optical section. C. Epibionts with capitated tentacles arranged in left and right row. Extended focus image of three images. D. Epibiont with developing swarmer. Arrowheads mark membrane of swarmer. E. Epibiont with swarmer recently freed from its parent. F. Segments 4–6, ventral view. Note thickened lorica in basal two thirds of epibionts.Arrow marks longitudinal striation of stalk of epibiont. Abbreviations: ca, basal cavity in lorica; co, specimen of Cothurnia buetschlii; lo, lorica; lts, lateral terminal spine; ma, macronucleus; mi, micronucleus; s, segment, followed by segment number; su, suctorian epibiont; sw, swarmer; te, tentacle. Scale bar in A, D, E 20 µm, in B 100 µm, in C 30 µm, and in F 50 µm.	FIGURE 4. Setaphyes kielensis (A, ZMB 12469; B, ZMB 12489; C, ZMB 12486; D, E, ZMB 12492; F, ZMB 12488) with specimens of Trematosoma husselae sp. nov., SEM (A), and DIC (B–F). A. Segment 1, empty shells of epibionts, ventral view. B. Segments 1–11, optical section. C. Epibionts with capitated tentacles arranged in left and right row. Extended focus image of three images. D. Epibiont with developing swarmer. Arrowheads mark membrane of swarmer. E. Epibiont with swarmer recently freed from its parent. F. Segments 4–6, ventral view. Note thickened lorica in basal two thirds of epibionts.Arrow marks longitudinal striation of stalk of epibiont. Abbreviations: ca, basal cavity in lorica; co, specimen of Cothurnia buetschlii; lo, lorica; lts, lateral terminal spine; ma, macronucleus; mi, micronucleus; s, segment, followed by segment number; su, suctorian epibiont; sw, swarmer; te, tentacle. Scale bar in A, D, E 20 µm, in B 100 µm, in C 30 µm, and in F 50 µm.	2023-09-12	Neuhaus, Birger;Lu, Borong;Yamasaki, Hiroshi;González-Casarrubios, Alberto		Zenodo	biologists	Neuhaus, Birger;Lu, Borong;Yamasaki, Hiroshi;González-Casarrubios, Alberto			
F65487889916FF8AFF33F9937438FDAA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8349223/files/figure.png	https://doi.org/10.5281/zenodo.8349223	FIGURE 5. Setaphyes kielensis (A, ZMB 12489; B, ZMB 12500; C, ZMB 12499; D, ZMB 12487; J, ZMB 12486; L, ZMB 12397; M, ZMB 12369) and S. dentatus (E–G, K, ZMB 12503; F, H, I, ZMB 12494) with specimens of Trematosoma husselae sp. nov. (A–G, J–M), Cothurnia buetschlii (J), and a case of fungus or developmental artefact (H, I, ZMB 12494), DIC. A. Segments 1–4, lateral and dorso-ventral view on the calyciform, dorso-ventrally compressed lorica with the zooid, ventrolateral view on basibiont. B. Epibiont with swarmer, stalk, and lorica in optical section. C. Epibiont with two macronuclei not fusing at any focal level. D, E. Young, small, spherical epibionts with few tentacles, not all tentacles in focus. F. Young specimen on cuticle of basibiont with striation pattern on surface (arrows). G. Epibiont with tentacles in empty shell of suctorian epibiont. H, I. Segment 10, dorsally (H) and optical section (I), subcuticular fungus or internal sclerotisation artefact of the cuticle (arrows). J. Assemblage of epibionts on segment 10 of basibiont with C. buetschlii (contracted specimen left, extended specimen right), T. husselae sp. nov., and two kinds of filamentous bacteria. Arrows mark endostyle. K. Suctorian on segment 7 of S. dentatus. L, M. Specimens collected 1998, showing cellular substructures (L) and withdrawn zooid with minimum amount of cellular substructures (M). Extended focus images of six (D), eight (H, L), and three (K) images. Arrowheads mark stalks in B, D, E, G, and J. Abbreviations: ba, bacteria; ca, basal cavity in lorica; co, C. buetschlii; en, endostyle; lo, lorica; ma, macronucleus; s, segment, followed by segment number; sc, spherical cavity; sp, sperm; st, stalk; su, suctorian epibiont; sw, swarmer; te, tentacle; tu, tube on segment 2. Scale bar in A 50 µm, in B 20 µm, valid for B, G–J and M, in C 20 µm, valid for C, K, and L, and in D–F 10 µm.	FIGURE 5. Setaphyes kielensis (A, ZMB 12489; B, ZMB 12500; C, ZMB 12499; D, ZMB 12487; J, ZMB 12486; L, ZMB 12397; M, ZMB 12369) and S. dentatus (E–G, K, ZMB 12503; F, H, I, ZMB 12494) with specimens of Trematosoma husselae sp. nov. (A–G, J–M), Cothurnia buetschlii (J), and a case of fungus or developmental artefact (H, I, ZMB 12494), DIC. A. Segments 1–4, lateral and dorso-ventral view on the calyciform, dorso-ventrally compressed lorica with the zooid, ventrolateral view on basibiont. B. Epibiont with swarmer, stalk, and lorica in optical section. C. Epibiont with two macronuclei not fusing at any focal level. D, E. Young, small, spherical epibionts with few tentacles, not all tentacles in focus. F. Young specimen on cuticle of basibiont with striation pattern on surface (arrows). G. Epibiont with tentacles in empty shell of suctorian epibiont. H, I. Segment 10, dorsally (H) and optical section (I), subcuticular fungus or internal sclerotisation artefact of the cuticle (arrows). J. Assemblage of epibionts on segment 10 of basibiont with C. buetschlii (contracted specimen left, extended specimen right), T. husselae sp. nov., and two kinds of filamentous bacteria. Arrows mark endostyle. K. Suctorian on segment 7 of S. dentatus. L, M. Specimens collected 1998, showing cellular substructures (L) and withdrawn zooid with minimum amount of cellular substructures (M). Extended focus images of six (D), eight (H, L), and three (K) images. Arrowheads mark stalks in B, D, E, G, and J. Abbreviations: ba, bacteria; ca, basal cavity in lorica; co, C. buetschlii; en, endostyle; lo, lorica; ma, macronucleus; s, segment, followed by segment number; sc, spherical cavity; sp, sperm; st, stalk; su, suctorian epibiont; sw, swarmer; te, tentacle; tu, tube on segment 2. Scale bar in A 50 µm, in B 20 µm, valid for B, G–J and M, in C 20 µm, valid for C, K, and L, and in D–F 10 µm.	2023-09-12	Neuhaus, Birger;Lu, Borong;Yamasaki, Hiroshi;González-Casarrubios, Alberto		Zenodo	biologists	Neuhaus, Birger;Lu, Borong;Yamasaki, Hiroshi;González-Casarrubios, Alberto			
F65487889918FF88FF63FED87634FC82.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8349223/files/figure.png	https://doi.org/10.5281/zenodo.8349223	FIGURE 5. Setaphyes kielensis (A, ZMB 12489; B, ZMB 12500; C, ZMB 12499; D, ZMB 12487; J, ZMB 12486; L, ZMB 12397; M, ZMB 12369) and S. dentatus (E–G, K, ZMB 12503; F, H, I, ZMB 12494) with specimens of Trematosoma husselae sp. nov. (A–G, J–M), Cothurnia buetschlii (J), and a case of fungus or developmental artefact (H, I, ZMB 12494), DIC. A. Segments 1–4, lateral and dorso-ventral view on the calyciform, dorso-ventrally compressed lorica with the zooid, ventrolateral view on basibiont. B. Epibiont with swarmer, stalk, and lorica in optical section. C. Epibiont with two macronuclei not fusing at any focal level. D, E. Young, small, spherical epibionts with few tentacles, not all tentacles in focus. F. Young specimen on cuticle of basibiont with striation pattern on surface (arrows). G. Epibiont with tentacles in empty shell of suctorian epibiont. H, I. Segment 10, dorsally (H) and optical section (I), subcuticular fungus or internal sclerotisation artefact of the cuticle (arrows). J. Assemblage of epibionts on segment 10 of basibiont with C. buetschlii (contracted specimen left, extended specimen right), T. husselae sp. nov., and two kinds of filamentous bacteria. Arrows mark endostyle. K. Suctorian on segment 7 of S. dentatus. L, M. Specimens collected 1998, showing cellular substructures (L) and withdrawn zooid with minimum amount of cellular substructures (M). Extended focus images of six (D), eight (H, L), and three (K) images. Arrowheads mark stalks in B, D, E, G, and J. Abbreviations: ba, bacteria; ca, basal cavity in lorica; co, C. buetschlii; en, endostyle; lo, lorica; ma, macronucleus; s, segment, followed by segment number; sc, spherical cavity; sp, sperm; st, stalk; su, suctorian epibiont; sw, swarmer; te, tentacle; tu, tube on segment 2. Scale bar in A 50 µm, in B 20 µm, valid for B, G–J and M, in C 20 µm, valid for C, K, and L, and in D–F 10 µm.	FIGURE 5. Setaphyes kielensis (A, ZMB 12489; B, ZMB 12500; C, ZMB 12499; D, ZMB 12487; J, ZMB 12486; L, ZMB 12397; M, ZMB 12369) and S. dentatus (E–G, K, ZMB 12503; F, H, I, ZMB 12494) with specimens of Trematosoma husselae sp. nov. (A–G, J–M), Cothurnia buetschlii (J), and a case of fungus or developmental artefact (H, I, ZMB 12494), DIC. A. Segments 1–4, lateral and dorso-ventral view on the calyciform, dorso-ventrally compressed lorica with the zooid, ventrolateral view on basibiont. B. Epibiont with swarmer, stalk, and lorica in optical section. C. Epibiont with two macronuclei not fusing at any focal level. D, E. Young, small, spherical epibionts with few tentacles, not all tentacles in focus. F. Young specimen on cuticle of basibiont with striation pattern on surface (arrows). G. Epibiont with tentacles in empty shell of suctorian epibiont. H, I. Segment 10, dorsally (H) and optical section (I), subcuticular fungus or internal sclerotisation artefact of the cuticle (arrows). J. Assemblage of epibionts on segment 10 of basibiont with C. buetschlii (contracted specimen left, extended specimen right), T. husselae sp. nov., and two kinds of filamentous bacteria. Arrows mark endostyle. K. Suctorian on segment 7 of S. dentatus. L, M. Specimens collected 1998, showing cellular substructures (L) and withdrawn zooid with minimum amount of cellular substructures (M). Extended focus images of six (D), eight (H, L), and three (K) images. Arrowheads mark stalks in B, D, E, G, and J. Abbreviations: ba, bacteria; ca, basal cavity in lorica; co, C. buetschlii; en, endostyle; lo, lorica; ma, macronucleus; s, segment, followed by segment number; sc, spherical cavity; sp, sperm; st, stalk; su, suctorian epibiont; sw, swarmer; te, tentacle; tu, tube on segment 2. Scale bar in A 50 µm, in B 20 µm, valid for B, G–J and M, in C 20 µm, valid for C, K, and L, and in D–F 10 µm.	2023-09-12	Neuhaus, Birger;Lu, Borong;Yamasaki, Hiroshi;González-Casarrubios, Alberto		Zenodo	biologists	Neuhaus, Birger;Lu, Borong;Yamasaki, Hiroshi;González-Casarrubios, Alberto			
