identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
ED01879AFFEAFF80FF68FB65FA5D0C59.text	ED01879AFFEAFF80FF68FB65FA5D0C59.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cheliplana asica Marcus 1952	<div><p>Cheliplana asica Marcus, 1952</p><p>Known distribution. Caraguatatuba, San Sebastián and Cananéa, São Paulo, Brazil (Marcus 1952).</p><p>Material and distribution in Cuba. One specimen studied alive and whole mounted (HU IX.4.42), collected in Bahía de Gibara (March 26, 2017), intertidal, upper 20 cm of the fine-grained sand, salinity 35 ‰.</p><p>Remarks. Our specimen is 1.4 mm long, measured on the whole mount, translucent, without eyes. Proboscis hooks curved, 13 µm long; muscular hook supports 9 µm long. A single testis located caudally from the pharynx. Testis connected to two globular seminal vesicles, 27 µm long. Copulatory bulb encloses the prostate vesicle and the unarmed cirrus. Cirrus 170 µm long. Prostate vesicle 38 µm long and 23 µm at the widest point. Ovary rounded and located aside from the copulatory bulb. Globular bursa located at the level of the common gonopore. Bursa connected to the common atrium, and opens externally through a duct into the vagina, at the caudal end of the body.</p></div>	https://treatment.plazi.org/id/ED01879AFFEAFF80FF68FB65FA5D0C59	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Diez, Yander L.;Reygel, Patrick;Artois, Tom	Diez, Yander L., Reygel, Patrick, Artois, Tom (2019): Schizorhynchia (Platyhelminthes, Rhabdocoela) from eastern Cuba, with the description of fifteen new species. Zootaxa 4646 (1): 1-30, DOI: 10.11646/zootaxa.4646.1.1
ED01879AFFEAFF80FF68FC45FDCE0999.text	ED01879AFFEAFF80FF68FC45FDCE0999.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kalyptorhynchia Graff 1905	<div><p>Kalyptorhynchia Graff, 1905</p><p>Schizorhynchia Meixner, 1928</p></div>	https://treatment.plazi.org/id/ED01879AFFEAFF80FF68FC45FDCE0999	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Diez, Yander L.;Reygel, Patrick;Artois, Tom	Diez, Yander L., Reygel, Patrick, Artois, Tom (2019): Schizorhynchia (Platyhelminthes, Rhabdocoela) from eastern Cuba, with the description of fifteen new species. Zootaxa 4646 (1): 1-30, DOI: 10.11646/zootaxa.4646.1.1
ED01879AFFEAFF81FF68F949FB430BC3.text	ED01879AFFEAFF81FF68F949FB430BC3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cheliplana terminalis Brunet 1968	<div><p>Cheliplana terminalis Brunet, 1968</p><p>(Fig. 1)</p><p>Known distribution. Le Brusc, Marseilles, France (Brunet 1968); Djezira, Mogadishu, Somalia (Schockaert 1982); Tudor Creek, Mombasa, Kenya (Jouk &amp; DeVocht 1989).</p><p>Material and distribution in Cuba. Observations on live animals, whole mounted afterwards. Eight whole mounts from Bahía de Santiago de Cuba (January 31, 2017) (HU IX.4.43–IX.4.50), intertidal, superficial finegrained sand with organic matter, salinity 35 ‰. Three whole mounts from Chivirico (March 16, 2017) (HU X.1.01– X.1.03), intertidal up to 0.5 m depth, medium-grained sand with organic matter, surrounded by a seagrass bed of Thalassia testudinum and Syringodium filiforme, with isolated trees of Rhizophora mangle, salinity 35 ‰.</p><p>Remarks. Our animals are 0.5–0.9 mm long (x = 0.7 mm, n = 6), measured on the whole mounts, translucent, pinkish coloured, without eyes. Proboscis hooks curved (Fig. 1A: h), 11–17 µm long (x = 14 µm; n = 5); muscular hook supports (Fig. 1A: ms) 7–8 µm long (x = 7 µm; n = 5). Prostate vesicle (Fig. 1B: pv) elongated, 71–102 µm long (x = 87 µm; n = 2). Seminal vesicles (Fig. 1B: sv) rounded to oval shaped. Cirrus (Fig. 1B: ci) an unarmed and thin sclerotized tube, 200–293 µm long (x = 251 µm; n = 4); it ends in a more or less rectangular sclerotized cap (Fig. 1B: sc), 9–12 µm long (x = 11 µm; n = 4) and 3–5 µm wide (x = 4 µm; n = 4).</p><p>Large bursa located caudally, bipartite, proximally with a well-defined vesicle filled with sperm. It opens externally through a coiled and ribbon shaped duct into the vagina, at the caudal end of the body.</p></div>	https://treatment.plazi.org/id/ED01879AFFEAFF81FF68F949FB430BC3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Diez, Yander L.;Reygel, Patrick;Artois, Tom	Diez, Yander L., Reygel, Patrick, Artois, Tom (2019): Schizorhynchia (Platyhelminthes, Rhabdocoela) from eastern Cuba, with the description of fifteen new species. Zootaxa 4646 (1): 1-30, DOI: 10.11646/zootaxa.4646.1.1
ED01879AFFEBFF81FF68FBD1FE9A0D8E.text	ED01879AFFEBFF81FF68FBD1FE9A0D8E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cheliplana gibarenha Diez & Reygel & Artois 2019	<div><p>Cheliplana gibarenha sp. n.</p><p>(Fig. 2)</p><p>http://zoobank.org/ urn:lsid:zoobank.org:act: 9DDAFE63-D0D3-4B1E-A7B6-B6099BF61011</p><p>Material. One specimen studied alive and whole mounted, designated holotype (FMNH https://id.luomus.fi/ KV.608), collected in Bahía de Gibara (type locality) (March 26, 2017), sublittoral, fine-grained sand, 1 m depth, salinity 35 ‰.</p><p>Etymology. The epithet refers to the name given to the people that live in Gibara, where the type locality is situated.</p><p>Diagnosis. Species of Cheliplana with the proboscis hooks 14 µm long, curved, and the muscular hook supports 5 µm long. A single seminal vesicle located in the caudal end of the body. Seminal vesicle and copulatory bulb directed forward. Cirrus 140 µm long and 4 µm wide distally, with the distal 13-µm-long portion (the cirrus) armed with fine spines about 0.5 µm long.</p><p>Description. Animal 0.8 mm long, measured on the whole mount, translucent, without eyes. Proboscis (Fig. 2A: pr, 2B) with hooks (Fig. 2B: h) curved, 14 µm long; muscular hook supports (Fig. 2B: ms) 5 µm long. Spiny prepharyngeal cavity (Fig. 2A: pph) slightly longer than the pharynx. Mouth (Fig. 2A: m) opens just caudally from the postrostral bulb (Fig. 2A: pb).</p><p>Testis (Fig. 2A: t) located laterally from the pharynx, connected to the single seminal vesicle by a very long vas deferens. Seminal vesicle (Fig. 2A &amp; 2 C–D: sv) located in the caudal body end. Seminal vesicle, including the seminal duct, 184 µm long and 28 µm wide at its widest point, with the seminal duct directed forwards. Copulatory bulb (Fig. 2 C–D) comprising the prostate vesicle and a cirrus, directed forward. Prostate vesicle (Fig. 2A &amp; 2 C–D: pv) long-drawn. Cirrus (Fig. 2A &amp; 2 C–D: ci) 140 µm long and 4 µm wide distally, with the distal 13-µm-long portion armed with small and fine spines (Fig. 2D: sp), about 0.5 µm long. Distal part of the copulatory bulb, surrounding the spiny cirrus, marked by a strong sphincter (Fig. 2D: sph).</p><p>Kidney-shaped ovary (Fig. 2A: ov) located at the level of the connection between the seminal vesicle and the prostate vesicle; it has the oocytes in a row, diminishing in diameter from the most distal to the most proximal end of the ovary. Bursa (Fig. 2A: b) located at the caudal end of the body; it seems to be bipartite and opens caudally into the vagina.</p></div>	https://treatment.plazi.org/id/ED01879AFFEBFF81FF68FBD1FE9A0D8E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Diez, Yander L.;Reygel, Patrick;Artois, Tom	Diez, Yander L., Reygel, Patrick, Artois, Tom (2019): Schizorhynchia (Platyhelminthes, Rhabdocoela) from eastern Cuba, with the description of fifteen new species. Zootaxa 4646 (1): 1-30, DOI: 10.11646/zootaxa.4646.1.1
ED01879AFFECFF87FF68FAE2FA8A0B69.text	ED01879AFFECFF87FF68FAE2FA8A0B69.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cheliplana santiaguera Diez & Reygel & Artois 2019	<div><p>Cheliplana santiaguera sp. n.</p><p>(Fig. 3)</p><p>http://zoobank.org/ urn:lsid:zoobank.org:act: B24F7960-1110-4DB1-8606-48BE1DB8BB87</p><p>Material. Observations on two live animals, whole mounted afterwards, one of which is designated holotype (FMNH https://id.luomus.fi/KV.609), the other in HU (X.1.04), collected in Siboney (type locality) (December 14, 2017), sublittoral, fine-grained sand, 2–3 m depth, salinity 35 ‰.</p><p>Etymology. The epithet refers to the name given to the people that live in Santiago de Cuba, where the type locality is situated.</p><p>Diagnosis. Species of Cheliplana with the proboscis hooks 16 µm long, and the muscular supports 7 µm long. With two seminal vesicles. Copulatory bulb inverted pear shaped, 62 µm long. Ejaculatory duct constricted at its middle by a sphincter. Spiny cirrus 25 µm long, armed with fine spines about 0.5 µm long.</p><p>Description. Animals 0.7–0.8 mm long (n = 2) measured on the whole mounts, pinkish coloured, without eyes. Proboscis (Fig. 3A &amp; 3D: pr, 3B) with curved hooks (Fig. 3B: h), each hook 16 µm long (n = 3); muscular hook supports (Fig. 3B: ms) 6–8 µm long (x = 7 µm; n = 4). Pharynx (Fig. 3A: ph) 1/8 of the body length, 124–126 µm long (n = 2) measured on the whole mounts, connected to the mouth (Fig. 3A: m) by a long prepharyngeal cavity (two times the pharynx length) (Fig. 3A: pph). Wall of the prepharyngial cavity with spines.</p><p>Copulatory bulb (Fig. 3C, 3E) inverted pear shaped, 50–74 µm long (x = 62 µm; n = 2), comprising the prostate vesicle (Fig. 3A, 3C &amp; 3E: pv) and the cirrus. Ejaculatory duct constricted at its middle by a sphincter (Fig. 3C: sph). Cirrus (Fig. 3A, 3C &amp; 3E: ci) 22–28 µm long (x = 25 µm; n = 2), armed with fine, 0.5-µm-long spines.</p><p>The vitellarium (Fig. 3A: vi) extends between the pharynx and the copulatory bulb. Kidney-shaped ovary (Fig. 3A: ov) (with the oocytes in a row, diminishing in diameter from the most distal to the most proximal end of the ovary) and bursa (Fig. 3A: b) located caudally. Bursa connects to the common general atrium and also opens externally through a vagina (Fig. 3A: va), which is situated posteriorly from the common gonopore (Fig. 3A: cg).</p></div>	https://treatment.plazi.org/id/ED01879AFFECFF87FF68FAE2FA8A0B69	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Diez, Yander L.;Reygel, Patrick;Artois, Tom	Diez, Yander L., Reygel, Patrick, Artois, Tom (2019): Schizorhynchia (Platyhelminthes, Rhabdocoela) from eastern Cuba, with the description of fifteen new species. Zootaxa 4646 (1): 1-30, DOI: 10.11646/zootaxa.4646.1.1
ED01879AFFEDFF85FF68F8A3FA580BA5.text	ED01879AFFEDFF85FF68F8A3FA580BA5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cheliplana spuriaseminalis Diez & Reygel & Artois 2019	<div><p>Cheliplana spuriaseminalis sp. n.</p><p>(Fig. 4)</p><p>http://zoobank.org/ urn:lsid:zoobank.org:act: 0BC1AB04-AAD1-46E4-BB15-ECA04C5AAA77</p><p>Material. Observations on two live animals, whole mounted afterwards, one of which is designated holotype (FMNH https://id.luomus.fi/KV.610), the other in HU (X.1.05), collected in Bueycabón (type locality) (February 6, 2018), fine-grained sand with organic matter, 0.5 m depth, salinity 33 ‰.</p><p>Etymology. The epithet refers to the fact that only one of the seminal vesicles seems to be functional, the second being atrophied and empty.</p><p>Diagnosis. Species of Cheliplana with proboscis hooks 16 µm long and muscular supports 5 µm long. There are two seminal vesicles, one of which is atrophied. Copulatory bulb cylindrical and curved, 94 µm long and 16 µm wide. Cirrus unarmed, 31 µm long, distally widening. Bursa opens externally through a vagina.</p><p>Description. Animals 0.9–1 mm long (n = 2) measured on the whole mounts, pinkish coloured, without eyes. Proboscis (Fig. 4B: pr, 4C) with curved hooks 16 µm long (n = 2); muscular hook supports 5 µm long (n = 3). Phar- ynx (Fig. 4 A–B: ph) connects to the mouth through the prepharyngeal cavity (Fig. 4B: pph); spiny prepharyngeal cavity twice as long as the pharynx.</p><p>Testis located caudally from the pharynx. Atrial organs located in the caudal fourth of the body. Two seminal vesicles open proximally into the copulatory bulb; one seminal vesicle is functional (Fig. 4 D–E: fsv), filled with sperm, 116–119 µm long (x = 118 µm; n = 2), the other one seemingly atrophied (Fig. 4 D–E: asv), empty, 32–40 µm long (x = 36 µm; n = 2). The testis apparently is not connected to the atrophied seminal vesicle. Copulatory bulb (Fig. 4A: cb, 4E, 4D) cylindrical and curved, 83–105 µm long (x = 94 µm; n = 2) and 14–17 µm wide (x = 16 µm; n = 2); it comprises the prostate vesicle (Fig. 4 D–E: pv), (an) accessory gland(s) (Fig. 4 D–E: ag) and the cirrus (Fig. 4 D–E: ci). Cirrus unarmed, 31 µm long (n = 2). It widens distally, where it is surrounded by longitudinal muscles.</p><p>Bursa and ovary caudally from the copulatory bulb. Bursa connects to the common atrium and opens externally through a vagina. The vaginal opening (Fig. 4F: va) is located posteriorly to the common gonopore (Fig. 4F: cg).</p></div>	https://treatment.plazi.org/id/ED01879AFFEDFF85FF68F8A3FA580BA5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Diez, Yander L.;Reygel, Patrick;Artois, Tom	Diez, Yander L., Reygel, Patrick, Artois, Tom (2019): Schizorhynchia (Platyhelminthes, Rhabdocoela) from eastern Cuba, with the description of fifteen new species. Zootaxa 4646 (1): 1-30, DOI: 10.11646/zootaxa.4646.1.1
ED01879AFFE0FF8AFF68FF51FD050F2E.text	ED01879AFFE0FF8AFF68FF51FD050F2E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cheliplana subproximalis Diez & Reygel & Artois 2019	<div><p>Cheliplana subproximalis sp. n.</p><p>(Fig. 5)</p><p>http://zoobank.org/ urn:lsid:zoobank.org:act: 2FEAA7C3-F447-4B35-8FC7-7DD2FBAB3C6D</p><p>Material. Observations on live animals, whole mounted afterwards. Two whole mounts from Bueycabón (type locality) (November 18, 2018), one of which is designated holotype (FMNH https://id.luomus.fi/KV.611), the other one reference material (HU X.1.06), intertidal, upper 10 cm of the fine-grained sand, salinity 31 ‰. One whole mount from Las Sardinas (June 22, 2016), intertidal, upper 10 cm of fine-grained sand, salinity 33 ‰ (HU X.1.07).</p><p>Etymology. The epithet refers to the fact that the seminal duct opens into de copulatory bulb, a little bit distally from the latter’s proximal end.</p><p>Diagnosis. Species of Cheliplana with proboscis hooks 16 µm long and muscular hook supports 7 µm long. Seminal duct opens into the copulatory bulb, subterminally at the latter’s proximal end. At the same level, two separate ducts of unknown origin enter the prostate vesicle. The copulatory bulb ends in a sclerotized cap 16 µm long, which is asymmetrical distally. The spiny cirrus is 48 µm long. It is a narrow tube, widening slightly at the level of the distal cap. The spines are 3 µm long distally and 0.5 µm long in the widened part.</p><p>Description. Animals 0.4–0.7 mm long (x = 0.5 mm; n = 3), measured on the whole mounts, translucent, pinkish coloured, without eyes. Proboscis (Fig. 5A: pr, 5B) with curved hooks 13–18 µm long (x = 16 µm; n = 4); muscular hook supports 5–8 µm long (x = 7 µm; n = 4). Pharynx (Fig. 5A: ph) 1/7–1/8 of the body length in live animals. Pharynx connects to the mouth via a long spiny prepharyngeal cavity (Fig. 5A: pph), twice as long as the pharynx. Mouth (Fig. 5A: m) opens ventrally, subterminally at the anterior body end.</p><p>Testis (Fig. 5A: t) located postero-laterally from the pharynx, connected to two seminal vesicles. Seminal vesicles (Fig. 5A; C–E: sv) and atrial organs located in the last fourth of the body. Seminal vesicles distally fused to form a short seminal duct (Fig. 5C: sd). Seminal duct opens into the copulatory bulb a bit distally from the latter’s proximal end. The copulatory bulb (Fig. 5 C–E) comprises the prostate vesicle and the spiny cirrus. Prostate vesicle (Fig. 5A &amp; 5 C–E: pv) with two proximal ducts (Fig. 5C &amp; 5E: x), which enter the vesicle at the level of the entrance of the seminal duct. It is not clear where exactly these ducts come from (gland necks of the prostate glands?). Distally, the copulatory bulb ends in a sclerotized cap (Fig. 5 C–E: sc). Sclerotized cap distally asymmetric, 10–20 µm long (x = 16 µm; n = 3) and 6–10 µm at its widest point (x = 8 µm; n = 3). Spiny cirrus (Fig. 5A, 5C &amp; 5E: ci) 62–66 µm long (x = 64 µm; n = 2; measured from the most proximal spines to the most distal tip). In the distal portion of the cirrus, the cirrus widens a bit and the spines are 2–3 µm long (x = 3 µm; n = 12). In this part it is surrounded by a sclerotized cap. Proximally from this part, spines are very fine and less distinguishable, about 0.5 µm long.</p><p>The vitellarium (Fig. 5A: vi) extends between the pharynx and the copulatory bulb. Kidney-shaped ovary (Fig. 5A: ov) (with the oocytes in a row, diminishing in diameter from the most distal one to the most proximal one) and bursa located caudally from the common gonopore.</p></div>	https://treatment.plazi.org/id/ED01879AFFE0FF8AFF68FF51FD050F2E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Diez, Yander L.;Reygel, Patrick;Artois, Tom	Diez, Yander L., Reygel, Patrick, Artois, Tom (2019): Schizorhynchia (Platyhelminthes, Rhabdocoela) from eastern Cuba, with the description of fifteen new species. Zootaxa 4646 (1): 1-30, DOI: 10.11646/zootaxa.4646.1.1
ED01879AFFE0FF89FF68FA61FB020D5D.text	ED01879AFFE0FF89FF68FA61FB020D5D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cheliplana verrucosa Diez & Reygel & Artois 2019	<div><p>Cheliplana verrucosa sp. n.</p><p>(Fig. 6)</p><p>http://zoobank.org/ urn:lsid:zoobank.org:act: 0F9C72C8-A3D2-4A73-96CC-2085293D1841</p><p>Material. Observations on one live animal, whole mounted afterwards, designated holotype (FMNH https:// id.luomus.fi/KV.612), collected in Bueycabón (type locality) (February 6, 2018), sublittoral, fine-grained sand with organic matter, 0.5 m depth, salinity 33 ‰ .</p><p>Etymology. The epithet refers to the fact that the epidermal glands resemble small warts.</p><p>Diagnosis. Species of Cheliplana with the epidermis covered by glands that resemble warts, except in the posterior 20% of the body. Proboscis hooks bifurcate at the tip; main hook 15–16 µm long, smallest hook 5–6 µm long, muscular hook supports 16 µm long. Spiny cirrus 28 µm long proximally and 9 µm wide distally, constricted in the middle (4 µm wide); proximal spines 8 µm long, distal ones 4 µm long.</p><p>Description. Live animal about 0.7 mm long, pinkish coloured, without eyes. Epidermis covered by warts-like glands (Fig. 6 A–D: w), except in the caudal 20% of the body. Small rounded rhabdites visible over the entire body surface. Proboscis (Fig. 6B &amp; 6D: pr, 6E) with hooks bifurcate at the tip, forming a second small hook; main hook 15–16 µm long (n = 2), smallest hook 5–6 µm long (n = 2); muscular supports 15–17 µm long (x = 16 µm; n = 2). Measured on the live animal, pharynx (Fig. 6A &amp; 6D: ph) 1/8–1/9 of the body length long, connected to the subterminal mouth (Fig. 6D: m) by a long and spiny prepharyngeal cavity (Fig. 6 A–B &amp; 6D: pph) that is three times longer than the pharynx.</p><p>Testis (Fig. 6D: t) located ventro-laterally from the pharynx. Testis connected to a single seminal vesicle (Fig. 6 C–D &amp; 6F–G: sv), opening proximally into the copulatory bulb. Prostate glands (Fig. 6D &amp; 6 F–G: prg) enter the copulatory bulb about at the entrance of the seminal vesicle. Copulatory bulb inverted-pear shaped, 49 µm long, comprising the prostate vesicle (Fig. 6 C–D &amp; 6F–G: pv) and the spiny cirrus. Spiny cirrus (Fig. 6 C–D &amp; 6F–G: ci) 28 µm long, 9 µm wide proximally, narrower at about its midpoint (4 µm), subdistally 12 µm wide, and narrowing to a distal triangular-shaped tip; as a whole the cirrus is arrow-like. Spines largest in the proximal end, 7–9 µm long (x = 8 µm; n = 4) and distally 4–5 µm long (x = 4 µm; n = 5).</p><p>Vitellarium (Fig. 6D: vi) extends behind the pharynx to the beginning of the copulatory bulb. Ovary and globular bursa (Fig. 6D: b) located caudally from the copulatory bulb. Ovary (Fig. 6D: ov) kidney shaped, with the oocytes in a row, diminishing in diameter from the most distal to the proximal end of the ovary. Only a connection between the bursa to the common atrium was observed.</p><p>Discussion on Cheliplana . All the above-mentioned species show the typical features of the genus (see de Beauchamp 1927; Schilke 1970a): a proboscis consisting of two muscle bulbs (“hook supports”) with simple, curved hooks, and with soft, not-sclerotized sidepieces, a cylindrical (barrel-shaped) pharynx with a long, often spiny prepharyngeal cavity, and only one ovary.</p><p>The specimen of C. asica collected in Cuba shows the diagnostic features of the species mentioned in the original description by Marcus (1952). Within the genus, an unarmed cirrus ending in a simple papilla as in C. asica is unique. The ejaculatory duct in the specimen from Cuba (170 µm) is shorter than it is in the specimens from Brazil (280 µm). As no other morphological differences between the specimens of both populations were observed, we consider both populations to belong to the same species. Cheliplana terminalis, primarily described as a subspecies of C. asica by Brunet (1968), can be distinguished from C. asica by the long sclerotized ejaculatory which ends in a sclerotized cap. This is clearly present in the specimens from Cuba. The specimens of C. terminalis from Cuba (0.5–0.9 mm long) are noticeably smaller than the specimens recorded from Marseille (1.6 mm long) (Brunet 1968), however, they are similar to the specimens from Somalia (about 1 mm long) (Schockaert 1982). Unfortunately, no measures of the copulatory organ of specimens from Marseilles, Somalia and Kenya are available. However, as there seem to be no morphological differences between the specimens from Cuba and the specimens from the other populations, we prefer to keep them provisionally in the same species.</p><p>Cheliplana verrucosa sp. n. is unique within the genus due it has hook supports that are more or less as long as the hooks themselves. In all other species of Cheliplana, the hook supports are much shorter than the hooks. Considering that only one specimen was examined, it is unclear whether the difference in length between the two muscular hook supports of C. verrucosa sp. n. (see Fig. 6E) is real, or was caused by squeezing. Also the epidermis of the anterior 80% of the body covered with wart-like glands was never recorded from any other species of Cheliplana . The detailed morphology and function of these glands is unknown. Distally bifurcate proboscis hooks as in C. verrucosa sp. n. are only known from C. paradoxa Noldt, 1989 . In C. paradoxa the proboscis hooks are larger (21–25 µm long) than in C. verrucosa sp. n. (17 µm long). Moreover, C. paradoxa has two seminal vesicles (Noldt 1989), whereas in C. verrucosa sp. n. there is only one. Recently, Armonies (2018) erected a new genus, Dactyloplana Armonies, 2018, to include C. paradoxa and Dactyloplana tridigitata Armonies, 2018 . The only feature distinguishing species of Dactyloplana from those of Cheliplana is the distally-split hooks in the former ones. However, considering the lack of other morphological differences and of molecular evidence, we decide here to keep both C. paradoxa and C. verrucosa sp. n. within the genus Cheliplana .</p><p>Also C. gibarenha sp. n. has only one seminal vesicle, a feature shared with only seven other species of Cheliplana: C. verrucosa sp. n., C. evdonini Karling, 1983, C. marcusi (Karling, 1956) Karling, 1983, C. remanei (Meixner, 1928) Karling, 1983, C. sarnensis Gobert, Reygel &amp; Artois, 2017, C. schilkei Noldt, 1989, and C. targa (Marcus, 1952) Karling, 1983 . Cheliplana gibarenha sp. n. is the only one of these species in which the seminal vesicle has a caudal position, and in which the copulatory bulb is oriented forwards, both features unique even within the entire genus. Apart from that it can be distinguished from the other five species with only one seminal vesicle mostly by the detailed structure of the copulatory organ. In C. gibarenha sp. n. the copulatory bulb is very long, with a long cirrus, which only in its most distal, widened part bears small spines. In all other species the cirrus is either short or long, but always completely or for the greater part covered with spines, often showing several species-specific features of its own, such as the presence of an additional stylet in C. targa [for details on the other species: see Karling (1956, 1983); Noldt (1989)]. Of these species, C. marcusi is unique within the genus because its proboscis hooks bear a pair of denticles.</p><p>Cheliplana spuriaseminalis sp. n. is the only species of Cheliplana that has one atrophied and one functional seminal vesicle. Indeed, we could not observe any connection between the atrophied seminal vesicle and the testis. In C. uruguayensis Van Steenkiste, Volonterio, Schockaert &amp; Artois, 2008, there are also two morphologically different seminal vesicles. In this species, however, one show the basic morphology, while the second one is larger and surrounded by a thick muscle coat; both are connected to the single testis (Van Steenkiste et al. 2008) and are filled with sperm, and hence seem to be functional. Members of the related genus Cheliplanilla Meixner, 1938 also have one seminal vesicle that ends blindly. However, in species of this genus both seminal vesicles contain sperm. More- over, species of Cheliplanilla all have denticles on the proboscis hooks, and have heavily-sclerotized rod-shaped proboscis sidepieces (see Meixner 1938; Brunet 1968; Schilke 1970a; Gobert et al. 2017), features not shared by C. spuriaseminalis sp. n. .</p><p>Cheliplana spuriaseminalis sp. n. has a rather short unarmed cirrus. Only few other species of Cheliplana lack hard structures in the copulatory organ, and most of them have a very long (&gt; 100 µm) unarmed cirrus ( C. asica, C. terminalis, C. vaginalis Karling, 1983). Only in C. rubescens Brunet, 1966 and C. hypergyna Boaden, 1965, two species up to now known only from the Mediterranean, is the unarmed cirrus comparable in length to that of C. spuriaseminalis sp. n. . Cheliplana hypergyna can easily be distinguished by the highly muscular vagina that is subdivided into several compartments, which is easily seen even in live animals (Boaden, 1965; own unpublished observations) and by the simple construction of the cirrus, which is no more than a sclerotized tube evenly tapering towards its distal end. According to Boaden, only the most distal part of the cirrus is eversible, and this part “ sometimes displays a number of fine ridges” (Boaden 1965). Such ridges are recorded to be present over the entire length of the cirrus of C. rubescens but are lacking in C. spuriaseminalis sp. n. . The copulatory bulb of C. rubescens is most comparable to that of C. spuriaseminalis sp. n., apparently also showing the curve at about its midpoint (see Brunet 1966: Fig. 8). However, the cirrus of C. spuriaseminalis sp. n. is much less broad, widening abruptly only at the most distal end to a broad space surrounded by longitudinal muscles but without any ridges.</p><p>Cheliplana santiaguera sp. n. and C. subproximalis sp. n. have two functional seminal vesicles and a cirrus armed with spines, without stylet or accessory cirrus (or cirri). This is the most common situation in Cheliplana . Among species with such a combination of features, some species show a long, winding cirrus, only the complete distal part of which is provided with spines ( C. hiemalis Brunet, 1968, C. pacifica Noldt &amp; Hoxhold, 1984, and C. euxeinos Ax, 1959). Other species ( C. setosa Evdonin, 1977, C. evdonini) have a short cirrus, typically consisting of three different regions (for details see Evdonin 1977; Karling 1983), which is not the case in Cheliplana santiaguera sp. n., nor in C. subproximalis sp. n. The remaining species can be divided into two groups, one consisting of species with a cirrus that is at most three times as long as broad ( C. gemmifera Noldt, 1989, C. mamkaevi Evdonin, 1977, C. microcirrus Noldt, 1989, C. piriformis Brunet, 1968, C. pusilla Brunet, 1968), the other group consisting of species with a cirrus that is relatively longer.</p><p>Cheliplana santiaguera sp. n. belongs to the latter group, with a cirrus that most resembles that of C. barringtonensis Noldt &amp; Hoxhold, 1984, C. californica Karling, 1989, C. canariensis Gobert, Reygel &amp; Artois, 2017, C. elkhornica, and C. orthocirra Ax, 1959 . These species are characterised by a cirrus of moderate length, all between 25 µm ( C. barringtonensis) and 75 µm ( C. elkhornica) long. Cheliplana santiaguera sp. n. can be distinguished from most of these species because the spines are of equal length over the entire surface. Only in C. barringtonensis and C. elkhornica are the spines of equal size over the entire length. However, in C. elkhornica the spines of the cirrus are sturdier and longer than in C. santiaguera sp. n. (4–5 µm vs 0.5 µm), and the cirrus itself is also much longer in C. elkhornica than in C. santiaguera sp. n. (75 µm vs 25 µm). C. barringtonensis has a spiny cirrus of comparable length to that of C. santiaguera sp. n. (20 µm), but also in this species the spines are larger (2–3 µm) than in C. santiaguera sp. n. . Moreover, C. barringtonenis shows a typical narrowing of the distal end of the cirrus, which is unarmed and ends in a 12-µm-long cap that protrudes into the atrium (see Noldt &amp; Hoxhold 1984), a feature that is unique within the group of species mentioned.</p><p>The copulatory organ of C. subproximalis sp. n. is most similar to that of C. hiemalis, as the ejaculatory duct is relatively long and shows a single curve. Both species, however, can easily be distinguished because the spiny cirrus in C. hiemalis is restricted to the most distal part of the copulatory organ, being only 12 µm long and covered with strong spines (see Brunet 1968). In C. subproximalis sp. n., the cirrus consists of two parts, a long proximal part covered with very fine spines, and a distal part of about 16 µm long, which is widened and covered by larger spines. This distal part lies within a sclerotized cap which is lacking in C. hiemalis . Moreover, in C. hiemalis the seminal duct only turns over ±90°, whereas in C. subproximalis sp. n. it shows a 180° turn, and the seminal vesicle is directed forwards. The subterminal entrance of the seminal duct into the prostate vesicle and the two ducts entering it at the same place are two features not recorded for any other species of Cheliplana in literature.</p></div>	https://treatment.plazi.org/id/ED01879AFFE0FF89FF68FA61FB020D5D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Diez, Yander L.;Reygel, Patrick;Artois, Tom	Diez, Yander L., Reygel, Patrick, Artois, Tom (2019): Schizorhynchia (Platyhelminthes, Rhabdocoela) from eastern Cuba, with the description of fifteen new species. Zootaxa 4646 (1): 1-30, DOI: 10.11646/zootaxa.4646.1.1
ED01879AFFE4FF8FFF68FF51FBF90AA8.text	ED01879AFFE4FF8FFF68FF51FBF90AA8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Carcharodorhynchus flavidus Brunet 1967	<div><p>Carcharodorhynchus flavidus Brunet, 1967</p><p>(Fig. 7)</p><p>Known distribution. Several localities in the Mediterranean: France (Brunet 1967); several localities in Sardinia (Italy) (Gobert et al. 2017). Lanzarote (Canary Islands) (Gobert et al. 2017). Possibly North Carolina, United States (Whitson et al. 2011).</p><p>Material and distribution in Cuba. Three specimens studied alive and whole mounted (HU X.1.08– X.1.10), collected in Siboney (September 4, 2016; December 14, 2017), sublittoral, fine-grained sand with organic matter, 0.7–2 m depth, salinity 32–35 ‰.</p><p>Remarks. The live animals from Cuba are about 1.5–2 mm long, yellowish, without eyes. There is a pale belt near the posterior body end, corresponding to the single adhesive girdle. Proboscis (Fig. 7A: pr, 7B–C) symmetric to lightly asymmetric; largest lip 126–130 µm long (n = 2) and the smallest one 118–119 µm long (n = 2). Proboscis with paired fields of shark-tooth-shaped denticles situated along each proboscis lip, separated by a characteristic ‘basal gap’ (see Brunet, 1967; Schilke 1970a; Whitson et al. 2011; Gobert et al. 2017). In the basal half of the toothed belts, the denticles are 1–2 µm long (x = 1 µm; n = 23) and are more or less ordered in three or four rows; in the distal half the denticles appear a bit larger 1–2 µm long (x = 2 µm; n = 12) and are ordered in two rows. In live animals, polygonal glands were observed in the internal surface of the lips (Fig. 7B: prg); the largest ones arranged in two rows on the central longitudinal axis of each lip. These larger glands are accompanied by smaller glands. None of these glands are observable on the whole mounts. Pharynx (Fig. 7A: ph) located at 50%, being about 7% of the body length in live animals.</p><p>Three pairs of testes (Fig. 7A: t) are arranged in two rows in front of the pharynx. Copulatory bulb (Fig. 7D) 63–119 µm long (x = 92 µm; n = 3); it comprises the prostate vesicle and the spiny cirrus. Prostate vesicle (Fig. 7D: pv) 67–87 µm long (x = 77 µm; n = 2). Spiny cirrus (Fig. 7D: ci) 26–35 µm long (x = 32 µm; n = 3), proximally 7–12 µm wide (x = 9 µm; n = 3) and distally 17–19 µm wide (x = 18 µm; n = 3).</p></div>	https://treatment.plazi.org/id/ED01879AFFE4FF8FFF68FF51FBF90AA8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Diez, Yander L.;Reygel, Patrick;Artois, Tom	Diez, Yander L., Reygel, Patrick, Artois, Tom (2019): Schizorhynchia (Platyhelminthes, Rhabdocoela) from eastern Cuba, with the description of fifteen new species. Zootaxa 4646 (1): 1-30, DOI: 10.11646/zootaxa.4646.1.1
ED01879AFFE5FF8CFF68FEC3FB2B09F5.text	ED01879AFFE5FF8CFF68FEC3FB2B09F5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Carcharodorhynchus smilodon Diez & Reygel & Artois 2019	<div><p>Carcharodorhynchus smilodon sp. n.</p><p>(Fig. 8)</p><p>http://zoobank.org/ urn:lsid:zoobank.org:act: 4A 200176 -9D9C-49CF-865D-F56B95ACADD3</p><p>Material. One specimen studied alive and whole mounted, designated holotype (FMNH https://id.luomus.fi/ KV.613), from Siboney (type locality) (December 9, 2017), sublittoral, medium-grained clean sand, 4 m depth, salinity 33 ‰.</p><p>Etymology. The epithet refers to the extinct Saber-tooth tigers Smilodon spp. (Mammalia), as the new species has some large denticles on the proboscis.</p><p>Diagnosis. Species of Carcharodorhynchus with symmetric proboscis, lips 127 µm long. The belts of denticles show a ‘basal gap’; denticles shark-tooth shaped, 2 µm long in the basal half of the belts, ordered in four rows, and 5 µm long in the distal half, ordered in a single row. Spiny cirrus 33 µm long.</p><p>Description. Live animal about 2 mm long, yellowish, caudally dark yellow-brownish, without eyes. There is a pale belt near the posterior body end, corresponding to the single adhesive girdle. Proboscis lips symmetric, 127 µm long. Proboscis (Fig. 8A: pr, 8B–C) with paired fields of shark-tooth-shaped denticles, situated on a belt on each proboscis lip. Denticles at the base of the proboscis separated by a ‘basal gap’. In the basal half of the toothed belts, the denticles are 1–2 µm long (x = 2 µm; n = 17) and are ordered in four to five rows. More distally there is only one row of 4–6-µm-long denticles (x = 5 µm; n = 12). In live animals, polygonal glands are present on the internal surface of the lips. The largest of the glands are arranged into two rows following the central longitudinal axis of each lip. At both sides of the large glands, smaller glands can be observed. These glands are not observable on the whole mounts. Pharynx 133 µm in diameter, located at 60%.</p><p>Testes not observed. A pair of elongated seminal vesicles open proximally into the copulatory bulb. Copulatory bulb (Fig. 8 D–E) 162 µm long, with the same construction as that of C. flavidus; it comprises the very large prostate vesicle (Fig. 8 D–E: pv) and the spiny cirrus. Spiny cirrus (Fig. 8 D–F: ci) 33 µm long, proximally 9 µm wide and distally 16 µm wide, with the same morphology as that of C. flavidus, including the detailed morphology and dimensions of the spines (see Brunet 1967).</p><p>Paired vitellaria extend between the brain (Fig. 8A: br) and the copulatory bulb. Bursa and ovaries (Fig. 8 E–F: ov) are situated just caudally from the copulatory bulb. A single oocyte was observed in each ovary.</p></div>	https://treatment.plazi.org/id/ED01879AFFE5FF8CFF68FEC3FB2B09F5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Diez, Yander L.;Reygel, Patrick;Artois, Tom	Diez, Yander L., Reygel, Patrick, Artois, Tom (2019): Schizorhynchia (Platyhelminthes, Rhabdocoela) from eastern Cuba, with the description of fifteen new species. Zootaxa 4646 (1): 1-30, DOI: 10.11646/zootaxa.4646.1.1
ED01879AFFE6FF8CFF68FC1DFDE10DA9.text	ED01879AFFE6FF8CFF68FC1DFDE10DA9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Carcharodorhynchus papillaris Diez & Reygel & Artois 2019	<div><p>Carcharodorhynchus papillaris sp. n.</p><p>(Fig. 9)</p><p>http://zoobank.org/ urn:lsid:zoobank.org:act: 01734072-BF6B-4DAD-98DD-C4BD54A2B471</p><p>Material. One specimen studied alive and whole mounted, designated holotype (FMNH https://id.luomus.fi/ KV.614), collected in Bueycabón (type locality) (May 25, 2018), sublittoral, fine-grained sand with organic matter, 0.5 m depth, salinity 35 ‰.</p><p>Etymology. The epithet refers to the fact that the copulatory bulb ends in an unarmed papilla.</p><p>Diagnosis. Species of Carcharodorhynchus with symmetric proboscis, lips 145 µm long. The belts of denticles show a ‘basal gap’; denticles shark-tooth shaped, 0.9 µm long in the basal half of the belts and 3 µm long in the distal half. A single seminal vesicle opens into the copulatory bulb. Copulatory bulb 102 µm long and 29 µm at the widest point, without hard structures, distally ending in a muscular penis papilla.</p><p>Description. Live animal about 2 mm long, yellowish, without eyes. Proximal end of the body uncoloured. There is a pale belt near the posterior body end, corresponding to the single adhesive girdle. Proboscis symmetric, 145 µm long. Proboscis (Fig. 9 A–B: pr, 9C) with paired fields of shark-tooth-shaped denticles on each proboscis lip. Toothed belts from both lips separated by a ‘basal gap’. In the basal half of the toothed belts, the denticles are 0.6–1 µm long (x = 0.9 µm; n = 20) and are ordered in five to six rows. In the distal half the teeth are 3–4 µm long (x = 3 µm; n = 26) and are ordered in two to three rows. In live animals, polygonal glands are present on the internal surface of the lips. The largest of the glands are arranged into two rows following the central longitudinal axis of each lip. At both sides of the large glands, smaller glands can be observed. These glands are not observable on the whole mounts. Pharynx (Fig. 9A: ph) 95 µm in diameter, located at 75%.</p><p>A pair of testes is located rostrally from the pharynx. The vasa deferentia (Fig. 9F: vad) open separately into a single seminal vesicle (Fig. 9E: sv), which itself opens proximally into the copulatory bulb. Copulatory bulb (Fig. 9A: cb, 9D–F) 102 µm long and 29 µm at the widest point, without hard structures, ending in a muscular papilla. The prostate vesicle is short and rounded. Two types of prostate secretions were observed in the prostate vesicle of the live animal (Fig. 9 D–E: pv): in the proximal third a fine-grained secretion and in the middle third a coarsegrained secretion. Distal third of the copulatory bulb muscular, forming a penis papilla (Fig. 9 D–F: pp).</p><p>Paired vitellaria extend between the brain (Fig. 9A: br) and the copulatory bulb. Ovaries and bursa located caudally from the common gonopore.</p></div>	https://treatment.plazi.org/id/ED01879AFFE6FF8CFF68FC1DFDE10DA9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Diez, Yander L.;Reygel, Patrick;Artois, Tom	Diez, Yander L., Reygel, Patrick, Artois, Tom (2019): Schizorhynchia (Platyhelminthes, Rhabdocoela) from eastern Cuba, with the description of fifteen new species. Zootaxa 4646 (1): 1-30, DOI: 10.11646/zootaxa.4646.1.1
ED01879AFFE7FF93FF68F88EFC6C09D1.text	ED01879AFFE7FF93FF68F88EFC6C09D1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Carcharodorhynchus spiniferus Diez & Reygel & Artois 2019	<div><p>Carcharodorhynchus spiniferus sp. n.</p><p>(Fig. 10)</p><p>http://zoobank.org/ urn:lsid:zoobank.org:act: D32E2CF9-4285-4535-B79C-8C1DD6ACA8EB</p><p>Material. Observations on live animals, whole mounted afterwards. Three whole mounts from Siboney (type local- ity) (February 7 &amp; May 15, 2016; April 4, 2017), one of which is designated holotype (FMNH https://id.luomus. fi/KV.615), the others in HU (X.1.11– X.1.12), one specimen collected in fine-grained sand, 0.2 m depth; the second one in the intertidal, upper 20 cm of coarse-grained sand; and the third one on Ulva lactuca, 0.3 m depth, salinity 34–35 ‰. One whole mount from Macabí (April 23, 2017) (HU X.1.13), intertidal, upper 10 cm of fine-grained sand, salinity 34 ‰.</p><p>Etymology. The epithet refers to the fact that the cirrus is armed with long spines.</p><p>Diagnosis. Species of Carcharodorhynchus with proboscis asymmetric; largest lip 87 µm long and the smallest 70 µm. Inner side of both lips covered with small denticles. Largest lip with about 4–6 rows of denticles, smallest one with 2–3 rows of denticles. Belts of denticles continuous between both lips. The denticles on the largest lip are about 0.3 µm long in the proximal third, and 0.6 µm in the middle and distal thirds. On the smallest lip the ±0.6-µmlong denticles are located in the proximal and the middle third. Cirrus 50 µm long, armed with long spines, 23 µm long in the proximal two-thirds of its length and 5 µm long in the distal third. The largest spines bear a proximal small tooth in their middle.</p><p>Description. Living animals slender, 1–1.5 mm long, 0.6–1.5 mm long in the whole mounts (x = 1 mm; n = 4), yellow, without eyes. Epidermis opaque, thick and hampering the study of the internal morphology on live animals. Proboscis (Fig. 10A: pr, 10B–C) with asymmetric lips; largest lip 82–90 µm long (x = 87 µm; n = 3) and the smallest 68–73 µm long (x = 70 µm; n = 3). The inner side of the largest lip is covered by 4–6 rows of denticles from its most basal part towards the distal tip, leaving only the distal 20% without denticles. These denticles are ± 0.3 µm long in the basal third and ± 0.6 µm long over the distal two-thirds. The inner side of the smallest lip is covered by 2–3 rows of denticles from its most basal part towards its middle third, leaving the distal ±30% without denticles; these denticles are equal in size, ± 0.6 µm long. The belts of denticles are continuous between the lips. In live animals, spherical to oval-shaped glands (Fig. 10B: prg) were observed on the inner surface of the lips. Pharynx (Fig. 10A &amp; 10D: ph) located at 70%, 94–129 µm in diameter (x = 107 µm; n = 3).</p><p>Testes were observed only in one specimen (Macabí). There is one pair, which is located rostrally from the pharynx. Two large seminal vesicles (Fig. 10D: sv) open separately into the copulatory bulb. Copulatory bulb (Fig. 10A: cb, 10D–F) situated just caudally from the pharynx, 80–94 µm long, comprising the prostate vesicle (Fig. 10D: pv) and the armed cirrus. Cirrus (Fig. 10D: ci, 10E–F) 48–53 µm long (x = 50 µm; n = 4), armed with 21–25-µm-long spines in its proximal two-thirds (x = 23 µm; n = 6), 3–6-µm-long in its distal third (x = 5 µm; n = 8). The largest spines show a small tooth more or less in their middle (Fig. 10 E–F: to).</p><p>The paired vitellaria extend at both sides of the body from just behind the brain to the distal body third. Globular bursa located caudally from the copulatory bulb. Ovaries not observed.</p></div>	https://treatment.plazi.org/id/ED01879AFFE7FF93FF68F88EFC6C09D1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Diez, Yander L.;Reygel, Patrick;Artois, Tom	Diez, Yander L., Reygel, Patrick, Artois, Tom (2019): Schizorhynchia (Platyhelminthes, Rhabdocoela) from eastern Cuba, with the description of fifteen new species. Zootaxa 4646 (1): 1-30, DOI: 10.11646/zootaxa.4646.1.1
ED01879AFFF9FF96FF68FC26FD3A0B69.text	ED01879AFFF9FF96FF68FC26FD3A0B69.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Carcharodorhynchus nativus Diez & Reygel & Artois 2019	<div><p>Carcharodorhynchus nativus sp. n.</p><p>(Fig. 11)</p><p>http://zoobank.org/ urn:lsid:zoobank.org:act: D3B6D290-80A3-44F8-99CA-F7CC6CC66BF8</p><p>Material. Observations on live animals, whole mounted afterwards. One whole mount from Bueycabón (type locality) (November 18, 2018), designated holotype (FMNH https://id.luomus.fi/KV.616), intertidal, upper 10 cm of fine-grained sand, salinity 31 ‰. Five whole mounts from El Masío (June 30, 2016) (HU X.1.14– X.1.18), intertidal, upper 10 cm of medium-grained sand, salinity 33 ‰.</p><p>Etymology. Species named in tribute to the autochthonous populations (named “nativos” in Spanish) who dwelled around the distribution area of the new species before, and few years after the discovery of Cuba by Christopher Columbus.</p><p>Diagnosis. Species of Carcharodorhynchus with asymmetric proboscis; largest lip 119 µm long and smallest 95 µm long. In the smallest lip, the denticles are 1–2 µm long, ordered in four rows. In the largest lip, the denticles are 0.3–0.5 µm long in the basal half, ordered in six to seven rows, and 1–2 µm long in the distal half, ordered in four to six rows. Spiny cirrus 45 μm long; densely covered with small spines, which are 3 µm long proximally, 6 µm in the midpart, and 9 µm distally.</p><p>Description. Live animals slender, 1–1.5 mm long, light-brown to orange-brown, without eyes. There is a pale belt near the posterior body end, corresponding to the single adhesive girdle. Epidermis opaque, thick, hampering the study of the internal morphology on live animals. Proboscis (Fig. 11A: pr, 11B–C) asymmetric: largest lip 105–146 µm long (x = 119 µm; n = 3), smallest 85–112 µm long (x = 95 µm; n = 3). In the smallest lip, denticles are 1–2 µm long, ordered in four rows. In the largest lip, denticles are 0.3–0.5 µm long in the basal half, ordered in six to seven rows, and 1–2 µm long in the distal half, ordered in four to six rows. The toothed belts are continuous between both lips. In live animals, spherical to oval-shaped glands (Fig. 11B: prg) were observed along the internal surface of the lips. Pharynx (Fig. 11A: ph) 64–142 µm in diameter (x = 100 µm; n = 3), located at 60%. Several diatoms were observed in the gut.</p><p>Testes were observed only in one specimen (Bueycabón). There is one pair, which is located rostrally from the pharynx. Copulatory bulb (Fig. 11A: cb, 11D–E) located caudally from the pharynx. Two elongated seminal vesicles (Fig. 11 D–E: sv) narrow towards the copulatory bulb, which they enter separately. Elongated copulatory bulb 86–101 μm long (x = 92 µm; n = 3), comprising the prostate vesicle and the spiny cirrus. Prostate vesicle (Fig. 11 D–E: pv) spherical, 31–37 μm long (x = 35 µm; n = 3). At the place where the seminal vesicles enter the prostate vesicle, two small vesicles (Fig. 11E: ev) can be observed, which are the necks of prostate glands. Spiny cirrus (Fig. 11 D–E: ci) 46–48 μm long (x = 45 µm; n = 3) and 9–10 μm wide proximally (n = 2). Spines densely packed together. Proximally the spines are 2–3 µm long (x = 3 µm; n = 10), in the middle part 5–7 µm (x = 6 µm; n = 10), and distally 8–10 µm (x = 9 µm; n = 10). Distally from the cirrus, the copulatory bulb continues as a long and nar- row male atrium (Fig. 11E: ma) towards the common genital atrium.</p><p>Paired vitellaria lie at both body sides, between the brain and the copulatory bulb. Globular bursa located caudally from the copulatory bulb. Globular ovaries located caudally from the bursa.</p><p>Discussion on Carcharodorhynchus . The new species from Cuba can easily be attributed to Carcharodorhynchus as they show all diagnostic features of the genus. Species of that genus all have a proboscis consisting of two sturdy lips that bear belts of small teeth (= denticles) at their inner surface, and have two ovaries, whereas most other schizorhynchs only have one (Meixner 1938; Ax 1951; Schilke 1970a; Reygel et al. 2014). All species have a very similar habitus and internal organisation, and can mainly be distinguished from each other based on details in the structure of the proboscis (symmetric or asymmetric), the shape and organisation of the proboscis denticles, and the morphology of the copulatory organ proper.</p><p>Seventeen species of Carcharodorhynchus have been described, only one of which ( C. flavidus) was described with a proboscis with separated belts of teeth; i.e. the rows of teeth of the ventral and dorsal lip are separated from each other by an area lacking teeth at the place where the two lips join each other (see Brunet, 1967). This characteristic “basal gap” is also seen in two of the new species: C. smilodon sp. n. and C. papillaris sp. n. . Our examination of the specimen of C. polyorchis L’Hardy, 1963 deposited in the SMNH (Nr. 54059) clearly shows that also in this species this ‘basal gap’ is present, a fact never mentioned in literature. As such, the four species mentioned are clearly different from all other species of Carcharodorhynchus . The subtle differences distinguishing C. polyorchis and C. flavidus pertain to the presence/absence of an adhesive belt, the presence/absence of pigment grains in the epidermis and details in the male genital system, and are discussed by Brunet (1967). The length of the cirrus in the specimens of C. flavidus collected in Cuba (26–35 µm) clearly fits the range provided by Brunet (1967) (15–39 µm), as did that of the populations recorded from North Carolina (19–22 μm; Whitson et al. 2011), Lanzarote (26–67 μm; Gobert et al. 2017) and from Sardinia (30–34 μm; Gobert et al. 2017). Whether the different populations represent a complex of ‘cryptic’ species or whether C. flavidus is a very wide-spread species remains enigmatic.</p><p>Carcharodorhynchus smilodon sp. n. and C. papillaris sp. n. resemble C. flavidus most as they both lack pigment granules, possess an adhesive belt near the caudal end, and have long-drawn seminal vesicles (in C. smilodon sp. n.). Carcharodorhynchus smilodon sp. n. is extremely similar to C. flavidus, and can mainly be distinguished from that species by the dimensions of the denticles on the proboscis belts. Carcharodorhynchus smilodon sp. n. has denticles of 2 µm long at the base of the lips, whereas the most distal ones are ± 5 µm long (up to 6 µm long for the largest ones). In C. flavidus, denticles are at most 3 µm long, and are more or less of the same length over the entire length of the proboscis lips, without large size differences.</p><p>Carcharodorhynchus papillaris sp. n. can easily be distinguished from C. flavidus, as well as from all other species of Carcharodorhynchus as it is the only representative of Carcharodorhynchus with a single seminal vesicle and a copulatory bulb without any hard structures. All other species show a stylet, a spiny cirrus, or a combination of both. Carcharodorhynchus spiniferus sp. n. and C. nativus sp. n. both have a cirrus only. Carcharodorhynchus spiniferus sp. n. differs from all the other species in that the cirrus is armed with large, up to 23-µm-long spines proximally, and much shorter spines distally. In all other species with an armed cirrus the spines are more or less equally long over the entire length of the cirrus (except for C. nativus sp. n.; see further), and never as long as 25 µm. Moreover, the tooth that each of the larger spines in the cirrus of C. spiniferus sp. n. shows is a unique feature within the genus.</p><p>Carcharodorhynchus nativus sp. n. is characterized by the cirrus spines very densely packed together: proximally 2–3 µm long, 5–7 µm long in the middle part, and 8–10 µm long distally. Such an arrangement is unique within the genus. As to proboscis dentition, the distribution pattern as seen in C. nativus sp. n. is similar to that in C. spiniferus sp. n., with two regions of denticles of different lengths, which is a quite an uncommon feature within Carcharodorhynchus . In C. worsaae Reygel, Janssens &amp; Artois, 2014, the denticles also show differences in length and morphology. On the longer proboscis lip of this species, each field is proximally armed with denticles ±1 μm wide and more distally denticles of ±2.5 μm wide. The most distal part shows denticles up to 6.5 μm wide and 2.5 μm high with a pointed end bending towards the base of the proboscis. The smaller proboscis lip bears denticles of the same size and with the same arrangement as the distal zone of the larger lip (see Reygel et al. 2014). However, this situation differs from that in the two species from Cuba. In C. spiniferus sp. n. the largest lip bears triangular denticles ± 0.3 µm long in the proximal third, and ± 0.6 µm in the middle and distal thirds; in the smallest lip the denticles are equal in size; ±0.6-µm-long. In C. nativus sp. n. the largest lip bears denticles 0.3–0.5 µm long in the basal half and 1–2 µm long in the distal half; in the smallest lip the triangular denticles are 1–2 µm. The proboscis of C. isolatus Schilke, 1970 also bears two kind of teeth: proximally in the largest lip the denticles are triangular and 2.4 µm long, and distally there are shark-teeth shaped and 4.8 µm long. In the smallest lip the teeth are similar to those in the distal part of the largest lip. Also C. longicirrus Noldt &amp; Hoxhold, 1984 and C. tabulaeferus Noldt &amp; Hoxhold, 1984 bear denticles of two different sizes on the proboscis: the smallest lip and the proximal part of the largest lip bears 0.5-µm-long teeth, and in the largest one there are 1.2 µm long ( C. tabulaeferus) or 0.9–1.2 µm long ( C. longicirrus). Furthermore, the cirrus of C. nativus sp. n. (45 µm long) and C. spiniferus sp. n. (50 µm long) greatly differs from that of C. isolatus (14 µm long and armed with small spines), C. tabulaeferus (25 µm long and armed with 9-µm-long spines), and C. longicirrus (90 µm long and bears three cuticular strips and 10-µm-long spines) (see Schilke 1970a; Noldt &amp; Hoxhold 1984).</p></div>	https://treatment.plazi.org/id/ED01879AFFF9FF96FF68FC26FD3A0B69	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Diez, Yander L.;Reygel, Patrick;Artois, Tom	Diez, Yander L., Reygel, Patrick, Artois, Tom (2019): Schizorhynchia (Platyhelminthes, Rhabdocoela) from eastern Cuba, with the description of fifteen new species. Zootaxa 4646 (1): 1-30, DOI: 10.11646/zootaxa.4646.1.1
ED01879AFFFCFF97FF68FE61FBEE09D1.text	ED01879AFFFCFF97FF68FE61FBEE09D1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Schizochilus espinosai Diez & Reygel & Artois 2019	<div><p>Schizochilus espinosai sp. n.</p><p>(Fig. 12 A–B, 15E)</p><p>http://zoobank.org/ urn:lsid:zoobank.org:act: 2CF67117-9665-4779-96D4-2B0A4272D392</p><p>Material. One specimen studied alive and whole mounted, designated holotype (FMNH https://id.luomus.fi/ KV.617), collected in Bueycabón (type locality) (February 21, 2018), sublittoral, fine-grained sand with organic matter, 0.5 m depth, salinity 33 ‰.</p><p>Etymology. Species dedicated to Prof. Em. Dr. José Espinosa Sáez (Instituto de Ciencias del Mar, La Habana, Cuba), specialist in ecology and taxonomy of molluscs, and awarded with the National Award of Marine Sciences of Cuba (2018).</p><p>Diagnosis. Species of Schizochilus with a spiny cirrus 71 μm long; spines talon-shaped, about 3 μm long. A 52-μm-long, needle-shaped stylet runs centrally through the cirrus. Copulatory bulb distally forming a 12-μm-long cap.</p><p>Description. Animal 0.9 mm long, measured on the whole mount, translucent, without eyes. Proboscis lips (Fig. 12A: pr) 77–92 μm long (x = 85 μm; n = 2), with a pair of caudal glands. Pharynx (Fig. 12A: ph) 72 μm in diameter, located at 70%.</p><p>Four testes (Fig. 12A: t) lie in a row rostrally from the pharynx, fused two by two into two pairs. Copulatory bulb (Fig. 12A: cb, 15E) 77 μm long, comprising the prostate vesicle and the spiny cirrus. The cirrus is 71 μm long and 6 μm wide, measured over its midlength. The spines are talon-shaped and about 2–3 μm long (x = 3 μm; n = 2). A 52-µm-long stylet (Fig. 12B) runs centrally through the cirrus. It is 6 μm wide proximally, needle shaped, and tapers to a distal sharp point. The copulatory bulb forms a distal sclerotized cap of 12 μm long and 9 μm wide at the widest point. As such, it surrounds the cirrus for 23% of the latter’s length. The common gonopore (Fig. 12A: cg) opens at 90%.</p><p>A pair of vitellaria (Fig. 12A: vi) extend between the brain (Fig. 12A: br) and the copulatory bulb, one vitellarium at each side of the body. A bursa (Fig. 12A: b) and ovary (Fig. 12A: ov) are located in the caudal body end. The uterus (Fig. 12A: ut) is directed forwards, situated beside the copulatory bulb.</p></div>	https://treatment.plazi.org/id/ED01879AFFFCFF97FF68FE61FBEE09D1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Diez, Yander L.;Reygel, Patrick;Artois, Tom	Diez, Yander L., Reygel, Patrick, Artois, Tom (2019): Schizorhynchia (Platyhelminthes, Rhabdocoela) from eastern Cuba, with the description of fifteen new species. Zootaxa 4646 (1): 1-30, DOI: 10.11646/zootaxa.4646.1.1
ED01879AFFFDFF97FF68FC26FBFA0D56.text	ED01879AFFFDFF97FF68FC26FBFA0D56.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Schizochilus atlanticus Diez & Reygel & Artois 2019	<div><p>Schizochilus atlanticus sp. n.</p><p>(Fig. 12 C–D, 15A–B)</p><p>http://zoobank.org/ urn:lsid:zoobank.org:act: 7C16446E-7CE1-40F9-8786- 171035 B7C507</p><p>Material. Observations on live animals, whole mounted afterwards. Four whole mounts from Macabí (type locality) (April 23, 2017), one of which is designated holotype (FMNH https://id.luomus.fi/KV.618), the other material in HU (X.1.19– X.1.21), intertidal, upper 10 cm of fine-grained sand, salinity 34 ‰. Two whole mounts from Siboney (April 4, 2017) (HU X.1.22– X.1.23), intertidal, upper 10 cm of fine-grained sand, salinity 34 ‰.</p><p>Etymology. The epithet refers to the fact that the species was found at the Atlantic coast of Cuba.</p><p>Diagnosis. Species of Schizochilus with a spiny cirrus; fine spines about 1.5 μm long. A straight, tubular, ±39- µm-long stylet is present in the cirrus. Copulatory bulb with a distal cap of ±19 μm long.</p><p>Description. Live animals about 1–1.5 mm long, translucent, without eyes. Habitus and internal organisation as in S. espinosai sp. n. . Proboscis lips 162–184 μm long (x = 176 μm; n = 12), with a pair of gland sacs that enter the proboscis at its proximal end. Pharynx 141–172 μm in diameter (x = 162 μm; n = 5), located at 70%.</p><p>Four (in the specimens from Macabí) to five or six (in the specimens from Siboney) oval- to kidney-shaped testes lie in a single row rostrally from the pharynx. Two vitellaria extend between the brain and the copulatory bulb, one at each side of the body. The rest of the genital organs are situated in the caudal fourth of the body, with the ovary and the bursa near to the caudal body end. Copulatory bulb 81–131 μm long (x = 107 μm; n = 6), com- prising the prostate vesicle and a spiny cirrus. Cirrus (Fig. 12 C–D, 15A–B) ornamented with 0.5–1.8-μm-long fine spines (x = 1.2 μm; n = 30). A tubular stylet (Fig. 12 C–D, 15A–B) lies centrally in the cirrus. Cirrus and stylet are 34–43 μm long (x = 39 μm; n = 6) and proximally 5–7 μm wide (x = 6 μm; n = 6). The distal walls of the copula- tory bulb are sclerotized, forming a 13–24-μm-long cap (x = 19 μm; n = 6) that is 9–14 μm at the widest point (x = 11 μm; n = 6). As such it surrounds the cirrus for 38–56% of the latter’s length.</p></div>	https://treatment.plazi.org/id/ED01879AFFFDFF97FF68FC26FBFA0D56	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Diez, Yander L.;Reygel, Patrick;Artois, Tom	Diez, Yander L., Reygel, Patrick, Artois, Tom (2019): Schizorhynchia (Platyhelminthes, Rhabdocoela) from eastern Cuba, with the description of fifteen new species. Zootaxa 4646 (1): 1-30, DOI: 10.11646/zootaxa.4646.1.1
ED01879AFFFDFF94FF68F8BAFB720885.text	ED01879AFFFDFF94FF68F8BAFB720885.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Schizochilus banesensis Diez & Reygel & Artois 2019	<div><p>Schizochilus banesensis sp. n.</p><p>(Fig. 12G, 15C)</p><p>http://zoobank.org/ urn:lsid:zoobank.org:act: 8B63964A-DB9B-4BE3-A0EC-07176AD0F423</p><p>Material. Observations on one live animal, whole mounted afterwards, designated holotype (FMNH https:// id.luomus.fi/KV.619), collected in Playa Morales (type locality) (January 4, 2017), intertidal, upper 10 cm of finegrained sand, salinity 35 ‰.</p><p>Etymology. The epithet refers to its occurrence in Banes, area of the type locality.</p><p>Diagnosis. Species of Schizochilus with a spiny cirrus, covered with fine spines of ±2 μm long.A needle-shaped, 70-μm-long stylet runs centrally through the cirrus. Copulatory bulb with a 32-µm-long sclerotized distal cap.</p><p>Description. Live animal about 1.5 mm long, translucent, without eyes. Habitus and internal organisation as in S. espinosai sp. n. . Proboscis lips 279 μm long (n = 2). Pharynx 226 μm in diameter, located at 75%.</p><p>Two vitellaria extend between the brain and the copulatory bulb, one at each side of the body. The rest of the genital organs are situated in the caudal fourth of the body, with the ovary and the bursa near to the caudal body end. Two long testes are located laterally between the brain and the pharynx, one on each body side. Seminal vesicles open proximally into the copulatory bulb. Copulatory bulb 144 μm long, comprising the prostate vesicle and the cir- rus. Cirrus (Fig. 12G, 15C) 70 μm long and ±8 μm wide, armed with fine spines that are 1.7–2.3 μm long (x = 2 μm; n = 9). A stylet (Fig. 12G, 15C) lies centrally in the cirrus. It is slightly curved in the live animal; more curved in the whole mount (probably because of the squeezing). It is 70 μm long and 8 μm wide proximally, needle shaped, taper- ing to a distal sharp tip. The distal walls of the copulatory bulb are sclerotized, forming a 32-μm-long cap, which is 13 μm wide at the widest point. As such, it surrounds the cirrus for 46% of the latter’s length.</p></div>	https://treatment.plazi.org/id/ED01879AFFFDFF94FF68F8BAFB720885	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Diez, Yander L.;Reygel, Patrick;Artois, Tom	Diez, Yander L., Reygel, Patrick, Artois, Tom (2019): Schizorhynchia (Platyhelminthes, Rhabdocoela) from eastern Cuba, with the description of fifteen new species. Zootaxa 4646 (1): 1-30, DOI: 10.11646/zootaxa.4646.1.1
ED01879AFFFEFF94FF68FC8DFF060C59.text	ED01879AFFFEFF94FF68FC8DFF060C59.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Schizochilus bueycabonensis Diez & Reygel & Artois 2019	<div><p>Schizochilus bueycabonensis sp. n.</p><p>(Fig. 12 E–F, 14C, 15D) http://zoobank.org/ urn:lsid:zoobank.org:act: 2362B7AC-F4D5-45CA-8C70-29AADD5E3DB4</p><p>Material. Observations on live animals, whole mounted afterwards. Four whole mounts from Bueycabón (type locality) (November 18, 2017), one of which designated holotype (FMNH https://id.luomus.fi/KV.620), other materials in HU (X.1.24– X.1.26), intertidal, upper 10 cm of fine-grained sand, salinity 31 ‰.</p><p>Etymology. The epithet refers to its occurrence in Bueycabón, the type locality of the new species.</p><p>Diagnosis. Species of Schizochilus with a spiny cirrus 37 μm long, armed with fine spines ±0.5 μm long. Tubular stylet lying inside the cirrus, ±31 μm long, with an oblique distal end. Copulatory bulb without a distal sclerotized cap.</p><p>Description. Animals 1–1.5 mm (x = 1.3 mm; n = 3), measured on whole mounts, translucent, without eyes. Habitus and internal organisation as in S. espinosai sp. n. . Proboscis lips 140–189 μm long (x = 162 μm; n = 8), with a pair of caudal glands. Pharynx 125–201 μm in diameter (x = 154 μm; n = 4), located at 70%.</p><p>Copulatory bulb (Fig. 14C, 15D) 84–104 μm long (x = 97 μm; n = 4), comprising the prostate vesicle and the spiny cirrus. The cirrus is 32–41 μm long (x = 37 μm; n = 4) and 4–7 μm wide proximally (x = 5 μm; n = 4), mea- sured over its midlength. The spines are very fine and ±0.5 μm long. A tubular stylet (Fig. 12 E–F, 14C, 15D) runs centrally through the cirrus. It is 27–33 μm long (x = 31 μm; n = 4) and 5–7 μm wide proximally (x = 6 μm; n = 4). It has an oblique distal end. In two specimens, including the holotype, the stylet is constricted just distally from the proximal funnel-shaped end of the stylet. The distal wall of the copulatory bulb is not sclerotized, and does not form a cap.</p></div>	https://treatment.plazi.org/id/ED01879AFFFEFF94FF68FC8DFF060C59	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Diez, Yander L.;Reygel, Patrick;Artois, Tom	Diez, Yander L., Reygel, Patrick, Artois, Tom (2019): Schizorhynchia (Platyhelminthes, Rhabdocoela) from eastern Cuba, with the description of fifteen new species. Zootaxa 4646 (1): 1-30, DOI: 10.11646/zootaxa.4646.1.1
ED01879AFFFEFF9BFF68F949FD9D0968.text	ED01879AFFFEFF9BFF68F949FD9D0968.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Schizochilus favus Diez & Reygel & Artois 2019	<div><p>Schizochilus favus sp. n.</p><p>(Fig. 13, 14 A–B)</p><p>http://zoobank.org/ urn:lsid:zoobank.org:act: 23613044-29FD-45CF-A694-B984B0164D12</p><p>Material. Observations on live animals, whole mounted afterwards. Six whole mounts from Siboney (type locality) (February 7, 2016), one of which is designated holotype (FMNH https://id.luomus.fi/KV.621), the others in HU (X.1.27– X.1.31), intertidal, upper 20 cm of the fine-grained sand, salinity 35 ‰.</p><p>Etymology. The epithet reflects the fact that the bases of the cirrus spines together have the appearance of a honeycomb. Lat. favus: honeycomb.</p><p>Diagnosis. Species of Schizochilus with a single large testis rostrally from the pharynx. Spiny cirrus 37 μm long; spines about 2 μm long in the proximal half of the cirrus and 7 μm long in the distal half. The bases of the spines are arranged in such a way that in the live animal they have the appearance of a honeycomb. Inside the cirrus a ±36-μm-long stylet is present. Copulatory organ without a distal cap.</p><p>Description. Live animals about 1–1.5 mm long, 0.6–1.4 mm long in the whole mounts (x = 1 mm; n = 5), translucent, without eyes. Habitus and internal organisation as in S. espinosai sp. n. . Proboscis (Fig. 13A: pr) 84– 88 μm long (x = 86 μm; n = 6), with a pair of gland sacs (Fig. 13A: pg) that enter the proboscis at its proximal end. Rostrally from the proboscis, in the proximal tip of the body, a pair of brownish-pink glands (Fig. 13A: gl) can be found, one on each body side. The brain (Fig. 13A: br) is located caudally from the proboscis. Pharynx (Fig. 13 A–B: ph) has a diameter of 93–98 μm (x = 96 μm; n = 4), and is located at 70%.</p><p>A large testis, up to 272 μm long in one whole mount (Fig. 13 A–B: t), is located rostrally from the pharynx. Two vitellaria (Fig. 13 A–B: vi) extend between the brain and the copulatory bulb, one at each side of the body. The rest of the genital organs are situated in the caudal fourth of the body, with the ovary and the bursa near to the caudal body end. Copulatory bulb (Fig. 13A: cb, 14A–B) comprising the prostate vesicle and a spiny cirrus. Spiny cirrus (Fig. 13B: ci, 13C–F, 14A–B) 34–38 μm long (x = 37 μm; n = 5) and 9–11 μm wide (x = 10 μm; n = 5). In the proximal half of the cirrus, the spines are about 2 μm long (n = 18); in the distal half they are 5–12 μm long (x = 7 μm; n = 12). The spines are arranged in such a way, that their bases together form a honeycomb-like struc- ture (see Fig. 13C), mainly in the distal half of the cirrus. Inside the cirrus lies a tubular stylet, 35–37 μm long (x = 36 μm; n = 2) and 8–12 μm wide proximally (x = 10 μm; n = 4). This stylet bears longitudinal ridges (see Fig. 13D, 14A). In general the stylet is not well visible because the spines hamper observation. Often only the proximal part is visible, as shown in Fig. 14A. The copulatory bulb does not form a sclerotized cap surrounding its distal tip. The common gonopore opens at 90%.</p></div>	https://treatment.plazi.org/id/ED01879AFFFEFF9BFF68F949FD9D0968	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Diez, Yander L.;Reygel, Patrick;Artois, Tom	Diez, Yander L., Reygel, Patrick, Artois, Tom (2019): Schizorhynchia (Platyhelminthes, Rhabdocoela) from eastern Cuba, with the description of fifteen new species. Zootaxa 4646 (1): 1-30, DOI: 10.11646/zootaxa.4646.1.1
ED01879AFFF1FF99FF68FCA7FDB20E9D.text	ED01879AFFF1FF99FF68FCA7FDB20E9D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Schizochilus maximus Diez & Reygel & Artois 2019	<div><p>Schizochilus maximus sp. n.</p><p>(Fig. 12H, 15F)</p><p>http://zoobank.org/ urn:lsid:zoobank.org:act: 471E20E5-1C96-4EC0-A164-DEE5E3FAE86C</p><p>Material. One specimen studied alive and whole mounted, designated holotype (FMNH https://id.luomus.fi/ KV.622), collected in Siboney (type locality) (February 7, 2016), fine-grained sand with small stones and silt, 0.5 m depth, salinity 35 ‰.</p><p>Etymology. The epithet refers to the fact that the new species has the largest cirrus and stylet among all the known species of the genus.</p><p>Diagnosis. Species of Schizochilus with a cirrus 129 μm long, unarmed in the proximal fourth of its length, ornamented with fine spines in the rest of its length; spines 3 μm long. Needle-shaped stylet 120 μm long, running centrally through the cirrus. Copulatory bulb with a distal cap, 19 μm long.</p><p>Description. Live animal about 2 mm long, translucent, without eyes. Habitus and internal organisation as in S. espinosai sp. n. . Proboscis lips 70 μm long, representing 3,5% of the body length in the live animal, with a pair of glands caudally. Pharynx located at the 75%, impossible to measure in the whole mount.</p><p>Copulatory bulb (Figs 12H, 15F) located caudally from the pharynx; it comprises the prostate vesicle and a spiny cirrus. The cirrus is 129 μm long, 7 μm wide proximally and 6 μm wide distally, unarmed in the proximal fourth of its length, the rest of the length is ornamented with fine spines. These spines are 2–4 μm long (x = 3 μm; n = 9). A 120-μm-long stylet lies centrally in the cirrus. It is needle-shaped, 5 μm wide proximally, 2 μm wide at the base of the proximal funnel, gradually tapering towards its sharp distal tip. Distal wall of the copulatory bulb sclerotized, forming a 19-μm-long cap. As such, it surrounds the cirrus for 15% of the latter’s length. Testes and female system not observed.</p><p>Discussion on Schizochilus . All new species described show the diagnostic features of the genus Schizochilus (see Boaden 1963; Gobert et al. 2017): symmetrical proboscis without any hard structures; pharynx situated in the posterior body half; a single ovary; several testes before the pharynx in a single row, often merged into a single or a few larger lobed testes; a copulatory organ proper consisting of a spiny cirrus, sometimes supplemented with a central stylet. Up to now, eleven species of Schizochilus have been described (Tyler et al. 2006–2018 b). Of these eleven species, eight have a copulatory organ proper consisting of a spiny cirrus surrounding a central stylet, as is the case in all six new species described in this contribution: S. bruneti Noldt &amp; Hoxhold, 1984, S. caecus L’Hardy, 1963, S. choriurus Boaden, 1963, S. hoxholdi Karling, 1989, S. lanzarotensis Gobert, Reygel &amp; Artois, 2017, S. parvulus Brunet, 1970, S. santacruzensis Noldt &amp; Hoxhold, 1984, and S. tubulatus Brunet, 1970 . An overview of the distinguishing characters among these known species is provided by Gobert et al. (2017).</p><p>Schizochilus maximus sp. n. can easily be recognised by its largest stylet (120 μm long) of all species of Schizochilus . Moreover, a cirrus unarmed in its proximal fourth is also a unique feature within the genus. In other species with a partly-unarmed cirrus, either the distal end is unarmed ( S. santacruzensis) (see Noldt &amp; Hoxhold 1984) or the cirrus bears longitudinal ridges and the spines are restricted to the distal part ( S. caecus and S. choriurus) (see Boaden 1963; L’Hardy 1963).</p><p>The five other species described in this contribution have a cirrus that is completely covered with spines, i.e. over the entire length of the stylet. This situation is also found in three described species: S. lanzarotensis, S. hoxholdi and S. parvulus . An important distinguishing feature between these species is the fact whether or not the copulatory bulb forms a distal, sclerotized cap, which surrounds the distal part of the cirrus/stylet. In the three species mentioned, such a cap is only lacking in S. lanzarotensis . This is also the case in two of the newly-described species from Cuba: S. favus sp. n. and S. bueycabonensis sp. n. .</p><p>Schizochilus bueycabonensis sp. n. can easily be recognized from S. lanzarotensis by the detailed morphology of the stylet and the cirrus. In S. bueycabonensis sp. n. the stylet is tubular, with a proximal funnel, 31 μm long, and has an oblique distal end. The cirrus of this species is armed over its entire length with fine, 0.5-µm-long spines. By studying the holotype of S. lanzarotensis for sake of detailed comparison, we noticed that the measurements for this species in Gobert et al. (2017) were erroneous, most probably because of a scale conversion mistake, and hence we provide new measurements for this species. In S. lanzarotensis, the stylet (Fig. 14D) is 80 μm long. It lacks the proximal funnel, and tapers to a distal sharp tip, and is therefore clearly distinguishable from this of S. bueycabonensis sp. n. The cirrus of the latter species is ornamented with two groups of shark-tooth-shaped spines: the smaller spines (3–5 μm long) proximally, and the larger ones (8–10 μm long) distally.</p><p>Schizochilus favus sp. n. can be distinguished from S. lanzarotensis, as well as from all other representatives of Schizochilus because it is the only species in which the spines are arranged in a way that the bases form a honeycomb structure in the right focal plane. Also the stylet is typical of this is species, lacking a proximal funnel and with longitudinal ridges, the latter also being a unique character for this species.</p><p>The general morphology of the cirrus and the stylet of S. atlanticus sp. n. is similar to that of S. bueycabonensis sp. n. . However, both species can easily be distinguished by the presence of a sclerotized cap in S. atlanticus sp. n. and its stylet ends in a sharp point distally, while it ends obliquely in S. bueycabonensis sp. n. .</p><p>The general morphology of the cirrus and the stylet of S. espinosai sp. n. also resembles that of S. lanzarotensis . However, both species can easily be distinguished by the presence of a sclerotized cap in S. espinosai sp. n., which is lacking in S. lanzarotensis . Moreover, the stylet of S. espinosai sp. n. is needle shaped, with a proximal funnel, and is 52 μm long, while in S. lanzarotensis it is tubular, lacks the proximal funnel, and is 80 μm long. Both species have a cirrus armed over its entire length with shark-tooth-shaped spines, but in S. lanzarotensis these are distributed into two size groups (see above), while all spines are more or less 3 µm long in S. espinosai sp. n. .</p><p>The general morphology of the copulatory organ of S. banesensis sp. n. is somewhat comparable to that of S. espinosai sp. n. . However, in the former species the cirrus is covered by much more, and much finer spines, and the sclerotized distal cap is much larger compared to the length of the stylet than it is in S. espinosai sp. n. . These features make both easily distinguishable.</p></div>	https://treatment.plazi.org/id/ED01879AFFF1FF99FF68FCA7FDB20E9D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Diez, Yander L.;Reygel, Patrick;Artois, Tom	Diez, Yander L., Reygel, Patrick, Artois, Tom (2019): Schizorhynchia (Platyhelminthes, Rhabdocoela) from eastern Cuba, with the description of fifteen new species. Zootaxa 4646 (1): 1-30, DOI: 10.11646/zootaxa.4646.1.1
