taxonID	type	description	language	source
C76D5C1A757B5D059E97E5121FF365C8.taxon	description	Figs 2 A, 7 A, B, 12 A, B, 28	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
C76D5C1A757B5D059E97E5121FF365C8.taxon	diagnosis	Diagnosis. Similar to Ayacucho triarmatus nom. nov. and A. weyrauchi comb. nov. by possessing three rows of spiniform tubercles in male femur IV (Fig. 7 A, B). It differs from A. weyrauchi comb. nov. for having more than four spiniform tubercles in retroventral row of femur IV (Fig. 7 B); higher tubercles in free tergites (Fig. 2 A); areas I-IV with a pair of median tubercles (Fig. 2 A); dorsal process absent in the penis (Fig. 12 A, B). Differs from A. triarmatus nom. nov. by possessing the retroventral row of spiniform tubercles along entire length of femur IV (Fig. 7 B); apical margin of the ventral plate with conspicuous lateral projections (Fig. 12 A, B); dorsal process absent (Fig. 12 A, B).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
C76D5C1A757B5D059E97E5121FF365C8.taxon	distribution	Distribution. (Fig. 28) PERU. Cajamarca. Bambamarca; Cerro Macheipungo.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
C76D5C1A757B5D059E97E5121FF365C8.taxon	materials_examined	Material examined. Type material: Of C. bambamarca: Holotype ♂, ' PERU, Cajamarca, Bambamarca | 2,800 m | 29 / VI / 1956, Weyrauch leg. (SMF RII 11649 / 32) - Paratypes 5 ♂, 3 ♀ ' ditto' (SMF RII 11649 / 32). Of C. triseriata: Holotype ♂ ' PERU, Cajamarca, Cerro Macheipungo | 4 km NE Bambamarca, 3,000 m | 28 / VI / 1956, Weyrauch leg. (SMF RII 11647 / 30) - Paratype ♀ ' ditto' (SMF RII 11647 / 30).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
600BA0C567F5516BBC1AAB18D4488B84.taxon	description	Figs 2 B, 7 C, D, 16 D-F, 22 A, B, 28	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
600BA0C567F5516BBC1AAB18D4488B84.taxon	description	Description. MALE: Measurements (n = 9) DSW: 3.3 - 3.4 (3.3); DSL: 4.6 - 5.2 (4.6); CL: 1.5 - 2.0 (1.5). FIVL: 5.0 - 5.3 (5.0). ChL: 1.7 - 3.0 (2.1). Coloration: (Fig. 22 A): Predominantly orange, with two longitudinal black spots in the lateral portion of areas I-IV. Lateral margins of the dorsal scutum yellowish, with small black spots. Posterior margin of the dorsal scutum, free tergites and legs predominantly black. Chelicerae and pedipalpus orange and reddish. Dorsum: (Fig. 2 B) Alpha-type DSS, with the constrictions weakly marked (when compared to most Ayacucho spp.). Anterior margin of carapace covered with granules in all its width. Ocularium unarmed, smooth or with a few granules in some individuals. Carapace with scattered granules distributed throughout its surface. Areas I-IV with one pair of small median tubercles; densely granulate. Posterior margin of dorsal scutum with a median row of 6 - 8 (6) acuminate tubercles (larger than tubercles of areas I-IV). Free tergites I-III with a row of tubercles similar to those on the posterior margin of dorsal scutum. Chelicerae: (Fig. 2 B) Swollen in some males (not swollen in the holotype). Segment I covered with granules throughout their length. Segment II predominantly smooth; finger with three teeth. Segment III with two teeth. Pedipalpus: With granules sparsely distributed on the dorsal surface of the femur and patella. Trochanter with a distal ventral setiferous tubercle. Femur with a ventrobasal setiferous tubercle and a row of 5 - 6 (5) small ventromedial setiferous tubercles. Tibia: prolateral IIII, retrolateral IIII. Tarsus: prolateral IIi, retrolateral Iii. Venter: Coxa I with scattered small tubercles; coxae II-III, predominantly smooth, with sparse granules; Coxa IV with denser granulation than coxae II-III. Genital area smooth. Free sternites I-IV and anal operculum with granules sparsely distributed. Legs: (Figs 2 B, 7 C, D) Coxa I with a prolateral apophysis; coxa II with a retrolateral and a prolateral apophysis. Coxa IV densely granulate. Trochanters I-IV with few granules, unarmed. Femora I-IV, with granules distributed throughout its length. Tarsal formula: (n = 9) 7 - 8 (7), 11, 7, 8. Penis: (Fig. 16 D-F) VP subrectangular; distal margin with a V-notch, and with conspicuous laterodistal projections; slightly curved in lateral view. MS C 1 - C 7 subdistal long and curved; MS A 1 long and straight; medially placed; MS B 1 sub basal long and straight; MS D 1 subdistal very short, dorsally placed. Lateral sacs long and apically acuminate; with long T 3 - like microsetae. Stylus apically robust and with projections. Dorsal process absent. Promontory convex. - FEMALE: Measurements (n = 10) DSW: 3.5 - 4.0; DSL: 4.5 - 5.2; CL: 1.6 - 1.8. FIVL: 4.7 - 5.6. ChL: 1.7 - 2.1. (Fig. 22 B) Chelicerae not swollen. Strongly similar to small males. Tarsal segmentation: (n = 10) 6, 10 - 11, 7, 8.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
600BA0C567F5516BBC1AAB18D4488B84.taxon	diagnosis	Diagnosis. Similar to Ayacucho pomacocha sp. nov., A. silvae sp. nov., A. titschacki and A. vargasllosai sp. nov. in the following combination of characteristics: dorsal scutum densely granulate; ocularium and areas I-IV of DS unarmed or armed with tiny tubercles, slightly greater than granules; posterior margin of DS and free tergites I-III with median rows of acuminate tubercles (Fig. 2 B); femur IV of males without strong armature (except in ventral surface of femur IV in A. silvae sp. nov., Fig. 7 C, D). It differs from the four previously mentioned species in the following combination of characteristics: DSS with weakly marked constrictions (Fig. 2 B); ocularium unarmed and smooth or with a few granules (Fig. 2 B); male femur IV unarmed (unlike A. silvae sp. nov., Fig. 7 C, D); and penis VP subrectangular with a V-notch on distal margin (Fig. 16 D, F).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
600BA0C567F5516BBC1AAB18D4488B84.taxon	etymology	Derivatio nominis. The specific epithet of masculine gender, in the genitive form, dedicated to the Brazilian filmmaker, actor and writer Glauber de Andrade Rocha (1939 - 1981).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
600BA0C567F5516BBC1AAB18D4488B84.taxon	distribution	Distribution. (Fig. 28) PERU. Huancavellica. Huancavellica, Quebrada Potreros.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
600BA0C567F5516BBC1AAB18D4488B84.taxon	materials_examined	Material examined. Type material: Holotype ♂, ' PERU, Huancavellica, Huancavellica, Quebrada Potreros | 12 ° 46 ′ 10.7 ″ S 75 ° 01 ′ 02.5 ″ W | 28 / IV / 2011, R. Pinto-da-Rocha, A. Benedetti, J. Ochoa & D. Silva leg. (MUBI) - Paratypes 1 ♂, 5 ♀, ' ditto' (MUBI); Paratypes 5 ♂, 11 ♀, ' ditto' (MZSP 36974); Paratypes 2 ♂, 5 ♀, ' ditto' (MUSM).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
1B054189204659C99211985EBCC44301.taxon	description	Figs 2 C, 7 E, F, 12 C, D, 28	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
1B054189204659C99211985EBCC44301.taxon	diagnosis	Diagnosis. Resembles Ayacucho spiniger comb. nov. because two ventral rows of tubercles in femur IV (Fig. 7 F). Differs from A. spiniger comb. nov. because femur IV with spiniform tubercles (Fig. 7 F); ventral plate of penis without lateral projections; VP with short macrosetae; lateral sacs with short microsetae; dorsal process shorter than the stylus (Fig. 12 C, D).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
1B054189204659C99211985EBCC44301.taxon	distribution	Distribution. (Fig. 28) PERU. La Libertad. Huamachuco.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
1B054189204659C99211985EBCC44301.taxon	materials_examined	Material examined. Type material: Holotype ♂, ' PERU, La Libertad, Huamachuco, 3,200 m a. s. l. without date, Weyrauch leg. (SMF RII 11393 / 23) - Paratype ♀, ' ditto' (SMF RII 11393 / 23).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
A257B9E5ECE8555CBEA039657D25FB10.taxon	description	Figs 2 D, 28	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
A257B9E5ECE8555CBEA039657D25FB10.taxon	diagnosis	Diagnosis. It differs from females of Ayacucho uniseriatus comb. nov. and A. weyrauchi comb. nov. because it has one pair of median tubercles in areas I-IV; from A. inermis comb. nov., A. spiniger comb. nov. and A. querococha sp. nov. because it has sparser granulation on carapace, and it is larger than A. inermis comb. nov.; A. bambamarca comb. nov. and A. tapacocha nom. nov. because it has smaller tubercles present in the posterior margin of dorsal scutum and free tergite I-II; and A. roeweri nom. nov. because it does not present a huge median spine in free tergites I-II. (Fig. 7 D).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
A257B9E5ECE8555CBEA039657D25FB10.taxon	distribution	Distribution. (Fig. 28) PERU. La Libertad. Hacienda Llaguen. Rejo Cargaruay forest.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
A257B9E5ECE8555CBEA039657D25FB10.taxon	materials_examined	Material examined. Type material: Holotype ♀, ' PERU, La Libertad. Hacienda Llaguen, Rejo Cargaruay forest, 2,650 m a. s. l., 14 / XII / 1952, Koepcke leg. (SMF RII 9706)	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
A1A323083F4054778BEBA5067216EC3C.taxon	description	Figs 2 E, 7 G, H, 12 E, F, 22 A, 28	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
A1A323083F4054778BEBA5067216EC3C.taxon	description	Description. MALE: Measurements (n = 1) DSW: 3.2; DSL: 3.6; CL: 1.2. FIVL: 5.4. ChL: 1.0. Coloration: (Fig. 22 G) Carapace, chelicerae, pedipalpus, legs and central part of areas of dorsal scutum brown; darker spots behind and laterally to the ocularium. Lateral parts and tubercles of areas of DS and lateral margins of DS with yellowish spots. Spines of area III black. Free tergites I-III yellowish. Dorsum: (Fig. 2 E) Alpha-type DSS, with shallower constrictions (especially constriction II, almost faint). Anterior margin of carapace with median elevation, almost smooth, with very few granules scattered. Ocularium with a median depression little sharped; a pair of spines, with granules in high density in the base. Carapace with few granules concentrated in the lateral regions. Areas I-II and IV with few granules scattered; III with greater quantity of granules than other scutal areas; I-II with a median pair of small tubercles; III totally granulate, with two spines, directed backwards, located at the highest point of the integument; IV with a median pair of small tubercles. Posterior margin of DS smooth with a pair of small median tubercles. Free tergites I-II without granules, with two-three median tubercles. Free tergite III with a pair of large median acuminate tubercles and a pair of smaller lateral acuminate tubercles. Lateral margins of DS with few granules covering its entire length. Chelicerae: (Fig. 2 E) Slightly swollen. Segment I densely covered with granules. Segment II smooth, with four teeth. Segment III with three teeth. Pedipalpus: dorsal region of the femur, tibia and patella granular. Trochanter with two ventroapical setiferous tubercles. Femur with a row of five ventral setiferous tubercles, except in the apical portion. Tibia: retrolateral iIII, prolateral IiII. Tarsus: retrolateral Iiii, prolateral Iiii. Venter: Coxae I-IV densely covered with granules and small tubercles throughout their surface. Genital area, anal operculum and free sternites granulate. Legs: (Figs 2 E, 7 G, H) Coxae I-II each one with a prolateral and a retrolateral apophysis; III with a prolateral apophysis, fused with retrolateral apophysis of coxa II; IV smooth. Trochanters I-IV unarmed and few granulate. Femora I-II unarmed and with small granules; III with a ventral retrobasal row of three tubercles; IV with dense granulation; a retrolateral row of eight-nine acuminate tubercles, growing apically occupying the distal ⅓; a prolateral row of seven acuminate tubercles, smaller than those in retrolateral row, covering the distal ⅓; a prodorsal row of eight small tubercles, extending over the distal ⅓; two ventral rows of nine acuminate tubercles each one on the distal half. Patellae I-III unarmed, with few granules; IV granular, with three tiny dorsoapical acuminate tubercles. Tibiae I-IV unarmed and densely granular. Tarsal segmentation: (n = 1) 7, 13, 10, 11. Penis: (Fig. 12 E, F) VP rectangular; distal margin straight; sinuous in lateral view. MS C 1 - C 4 apical long and slightly curved; MS A 1 - A 2 median long and straight (shorter than MS C); MS B 1 basal (near the lateral sacs) short and straight. Lateral sacs long, robust and apically blunt, with long T 3 - like microsetae. Stylus apically inflated, with slight ventral projection and small projections at the apex. Dorsal process long, cylindrical and apically acuminated. Promontory straight. - FEMALE: Measurements (n = 2) DSW: 3.1; DSL: 3.7 - 4.0; CL: 1.5 - 1.7. FIVL: 5.5 - 5.7. ChL: 1.5 - 1.6. Chelicerae slightly smaller than in male. Femur IV armed, but tubercles smaller than in males. Tarsal segmentation: (n = 2) 7, 12 - 13, 9 - 10, 10 - 11.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
A1A323083F4054778BEBA5067216EC3C.taxon	diagnosis	Diagnosis. It differs from other species of the genus by the set of following characteristics: dorsal scutal area III with a pair of long spines (Fig. 2 E; also present in A. spielbergi sp. nov.); longer legs (ratio between FIVL and DSL greater than 1; 1.5 in A. pasolinii sp. nov.); yellowish spots on the lateral margins of the DS, lateral portions and tubercles of the scutal areas I-IV (Fig. 22 A).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
A1A323083F4054778BEBA5067216EC3C.taxon	etymology	Derivatio nominis. The specific epithet of masculine gender, in the genitive form, dedicated to the Italian writer and filmmaker Pier Paolo Pasolini (1922 - 1975).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
A1A323083F4054778BEBA5067216EC3C.taxon	distribution	Distribution. (Fig. 28) PERU. Pasco. Oxapampa, Parque Nacional Yanachaga-Chemillen.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
A1A323083F4054778BEBA5067216EC3C.taxon	materials_examined	Material examined. Type material: Holotype ♂, ' PERU, Pasco, Oxapampa, Parque Nacional Yanachaga-Chemillen, 10 ° 32 ′ 42.1 ″ S 75 ° 21 ′ 24.4 ″ W, 22 / IV / 2011, R. Pinto-da-Rocha, A. Benedetti, J. Ochoa & D. Silva leg. (MUBI) - Paratypes 2 ♀, ' ditto' (MZSP 36993).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
C92DFA1621E351D9B98FF75832131474.taxon	description	Figs 2 F, 7 I, J, 15 A-C, 22 C, D, 28	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
C92DFA1621E351D9B98FF75832131474.taxon	description	Description. MALE: Measurements (n = 12). DSW: 3.2 - 4.0 (3.2); DSL: 4.7 - 5.3 (4.7); CL: 1.7 - 1.0 (1.7); FIVL: 4.5 - 5.0 (4.5); ChL: 1.2 - 3.5 (2.2). Coloration: (Fig. 22 C) Dorsal scutum ranging from brown to yellowish-brown, with darker regions on the lateral margins of carapace and scutal areas of the DS and others lighter, as bands. Lateral regions of scutal areas darker than the center, or completely brownish. Posterior margin of DS and free tergites brown. Legs dark brown. Chelicerae and pedipalpus yellow. Dorsum: (Fig. 2 F) Alpha-type DSS. Anterior margin of the dorsal scutum completely covered with granules, with median elevation. Ocularium totally covered with granules of equal size (a single specimen [on MZSP], has a pair of larger tubercles). Areas I-IV with a pair of slightly larger median tubercles. Posterior margin of DS and free tergites I-III with rows of tubercles larger than those of areas I-IV. Chelicerae: (Fig. 2 F) Equal to the females (including the holotype), swollen in a single specimen. Segment I covered with granules. Segment II predominantly smooth, with a few hairs on the frontal surface; finger with two teeth. Segment III with two teeth. Pedipalpus: With very small granules sparsely distributed on the dorsal surface of the femur and the patella. Trochanter with a ventrodistal setiferous tubercle. Femur with a ventrobasal setiferous tubercle; a row of five small ventral setiferous tubercles, except at the ends of the article. Some males (n = 3) have (at least in one of pedipalpus) a proapical setiferous tubercle, present in all females, but smaller. Tibia: prolateral IiIi, retrolateral iiIi. Tarsus: prolateral IIi, retrolateral Iii. Venter: Coxae I-II with granules in a row; Coxae III-IV with granules sparsely distributed. Smooth genital area. Free sternites I-IV and anal operculum with rows of small granules. Legs: (Figs 2 F, 7 I, J) Coxae I-II with a retrolateral and a prolateral apophysis; coxa III with a prolateral apophysis; coxa IV with granules distributed throughout its length. Trochanters I-IV with few granules, unarmed. Femora I-III, with granules distributed throughout their length; femur IV with granules densely distributed throughout its length, except for a small ventroapical strip without granules. Patellae I-IV with granules distributed predominantly in the dorsal and lateral regions, scarcer ventrally. Tibiae I-IV with granules throughout their length. Tarsal segmentation: (n = 12) 7, 11 - 12 (11), 7, 7 - 8 (8). Penis: (Fig. 17 A-C) VP hexagonal in dorsal view; subrectangular in ventral view, with distal half larger than basal half; distal margin straight. MS C 1 - C 9 subapical, long and curved; MS A 1 median placed, long and straight (smaller than MS C); MS B 1 sub basal, long and straight (longer than MS A); MS D 1 very short, dorsally placed, near to MS A. Lateral sacs short, apically tapered; with long T 3 - like microsetae. Stylus apically robust, with a large dorsal projection, and several small apical projections. Dorsal process absent. Promontory slightly convex. - FEMALE: Measurements (n = 12) DSW: 3.5 - 4.0; DSL: 4.8 - 5.2; CL: 1.7 - 1.9. FIVL: 4.5 - 5.0. ChL: 1.9 - 2.0. (Fig. 22 D) Chelicerae slightly smaller than those of smaller males. Presence of a proapical setiferous tubercle in femur of pedipalpus (also seen in some males), higher than the tubercles of the ventral femur. Femur IV slightly thinner, with smaller granules. Tarsal segmentation: (n = 12) 6, 11 - 12, 7, 8.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
C92DFA1621E351D9B98FF75832131474.taxon	diagnosis	Diagnosis. Similar to Ayacucho glauberrochai sp. nov., A. silvae sp. nov., A. titschacki and A. vargasllosai sp. nov. in the following combination of characteristics: dorsal scutum densely granulate; ocularium and areas I-IV of DS unarmed or armed with tiny tubercles, slightly greater than granules; posterior margin of DS and free tergites I-III with median rows of acuminate tubercles (Fig. 2 F); femur IV of males without strong armature (Fig. 7 I, J; except in ventral surface of femur IV in A. silvae sp. nov.). It differs from the previously mentioned species in the following combination of characteristics: ocularium unarmed and densely granulate (Fig. 2 F); male femur IV unarmed (unlike A. silvae sp. nov.; Fig. 7 I, J); penis VP hexagonal in dorsal view with straight distal margin; 9 pairs of subapical MS C (Fig. 15 A-C).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
C92DFA1621E351D9B98FF75832131474.taxon	etymology	Derivatio nominis. The specific epithet, a noun in apposition, in reference to the type locality, Laguna Pomacocha (Junin, Peru), a beautiful pond surrounded by grass and large rocks that harbor this species.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
C92DFA1621E351D9B98FF75832131474.taxon	distribution	Distribution. (Fig. 28) PERU. Junin. Laguna Pomacocha.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
C92DFA1621E351D9B98FF75832131474.taxon	materials_examined	Material examined. Type material: Holotype ♂, ' PERU, Junin, Laguna Pomacocha, 4,500 m a. s. l., 11 ° 46 ′ 36 ″ S, 75 ° 14 ′ 07 ″ W, 27 / IV / 2011, R. Pinto-da-Rocha, A. Benedetti, J. Ochoa & D. Silva leg. (MUBI) - Paratypes 2 ♂, 3 ♀, ' ditto' (MUBI); Paratypes 3 ♂, 3 ♀, ' ditto' (MUSM); Paratypes 6 ♂, 6 ♀, ' ditto' (MZSP 36970);	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
C3CF922A0FC65156BCE389D286AC0FC6.taxon	description	Figs 2 G, 7 K, L, 15 D-F, 28	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
C3CF922A0FC65156BCE389D286AC0FC6.taxon	description	Description. MALE: Measurements (n = 3) DSW: 3.4 - 4.1 (4.0); DSL: 4.2 - 5.0 (4.8); CL: 1.4 - 1.7 (1.7). FIVL: 3.3 - 4.0 (3.5). ChL: 1.7 - 2.8 (2.4) Coloration (alive): Yellowish with black spots on carapace, lateral part of the scutal areas I-IV, lateral margins of dorsal scutum and legs. Dorsum: (Fig. 2 G) Alpha-type DSS. Anterior margin of DS with median elevation with granules densely distributed. Granular ocularium, with two pairs of small tubercles taller than others. Carapace with granules densely distributed. Areas I-IV densely granulate; each one with a pair of median tubercles. Lateral margins of dorsal scutum with granules distributed throughout their length. Posterior margin of dorsal scutum and free tergites I-III with a row of acuminate tubercles, larger than those tubercles of areas of dorsal scutum, interspersed by small granules. Chelicerae: (Fig. 2 G) Swollen in large males (Swollen in holotype). Slightly larger than females' chelicerae in smaller males. Segment I granulate. Segment II predominantly smooth. Segment III with one tooth. Pedipalpus: Small granules distributed on the dorsal surface. Trochanter with a ventrodistal setiferous tubercle. Femur with a ventrobasal setiferous tubercle; a row of 5 - 6 ventrodistal median setiferous tubercles. Tibia: prolateral II, retrolateral iIii. Tarsus: prolateral Iii, retrolateral Iii. Venter: Coxa I with a median row of 4 - 5 small tubercles. Coxae II-IV and genital area with few scattered granules. Free sternites with a row of small granules. Anal operculum with granules sparsely distributed across its surface. Legs: (Figs 2 G, 7 K, L) Coxae I-II each one with a prolateral and a retrolateral apophyses; III with only prolateral apophysis; coxa II retrolateral and coxa III prolateral apophyses fused; IV with granules distributed throughout its surface, with a proapical spiniform apophysis. Trochanters I-III smooth; IV with 3 - 4 small dorsoapical tubercles and a larger retroapical median acuminate tubercle. Femora I-III with scattered granules; IV granular; with a retroventral row of three small acuminate tubercles and one large apical tubercle; a proventral row of 9 - 10 small tubercles, growing apically, and four large tubercles with curved apex; a prolateral row of 15 - 16 lanceolate tubercles, with the most basal slightly larger than granules, the most apical large, covering the entire length of segment; a retrolateral row of 11 - 12 tubercles. The shape of of retro- and prolateral tubercles are extremely variable and can have a blunt, acuminate or truncated apex. Patellae I-III with sparse granules; IV densely granulate dorsally, with more sparsely distributed granules on the ventral face, with three apical retrodorsal tubercles and a greater dorsoapical acuminate tubercle. Tibiae I-IV granular; IV with a row of 11 - 13 acuminate tubercles, growing apically. Tarsal segmentation: (n = 3) 7, 8 - 10 (10), 6 - 7 (7), 7 - 8 (8). Penis: (Fig. 15 D-F) VP rectangular; distal margin straight, with conspicuous laterodistal projections; slightly curved in lateral view. MS C 1 - C 4 distal long and curved; MS A 1 long and straight, medially placed; MS B 1 sub basal long and straight; MS D 1 very short, laterodorsally placed. Lateral sacs long, robust and with wider apex; with long T 3 - like microsetae. Stylus with triangular apex, with rounded corners; with a ventral projection with setae. Promontory straight. - FEMALE: Measurements (n = 15) DSW: 3.5 - 3.6; DSL: 4.0 - 4.5; CL: 1.4. FIVL: 3.3 - 3.5. ChL: 1.2 - 1.4 Chelicerae similar to that of small males. Pedipalpus femur with a proapical spine. Femur IV unarmed. Tarsal segmentation: (n = 15) 6, 9 - 10, 7, 7.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
C3CF922A0FC65156BCE389D286AC0FC6.taxon	diagnosis	Diagnosis. Similar to Ayacucho tapacocha nom. nov. because the tibia IV is armed (Fig. 7 K, L). Differs from Ayacucho tapacocha nom. nov. because DSS with constriction II most marked (Fig. 2 G); the presence of basal tubercles in a prolateral row in femur IV (totaling 15 - 16; 11 - 12 in A. tapacocha nom. nov.); retroventral row of femur IV with less than 5 tubercles (7 - 9 in A. tapacocha nom. nov.); presence of tubercles on the basal retrolateral row of tibia IV (Fig. 7 K, L); two pairs of small tubercles in ocularium; coxa III retrolateral apophysis absent (Fig. 2 G).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
C3CF922A0FC65156BCE389D286AC0FC6.taxon	etymology	Derivatio nominis. The specific epithet, a noun in apposition, in reference to the type locality, Laguna Querococha, a blue waters lagoon, from glacier of Parque Nacional Huascaran, Department of Ancash, type locality of the species.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
C3CF922A0FC65156BCE389D286AC0FC6.taxon	distribution	Distribution. (Fig. 28) PERU. Ancash. Parque Nacional Huascaran, Laguna Querococha.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
C3CF922A0FC65156BCE389D286AC0FC6.taxon	materials_examined	Material examined. Type material: Holotype ♂, ' PERU, Ancash, Parque Nacional Huascaran, Laguna Querococha, 4,024 m a. s. l., 09 ° 43 ′ 38.8 ″ S 77 ° 19 ′ 47.9 ″ W, 17 / V / 2010, R. Pinto-da-Rocha & D. Silva leg. (MUSM) - Paratypes 7 ♀, ' ditto' (MUSM); Paratypes 2 ♂, 8 ♀, ' ditto' (MZSP 36979).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
4D246E434B5850B880AA7810FE02D661.taxon	description	Figs 2, 3, 7, 8, 12, 13 A-D, 15, 16, 17 A-C, 22, 23; 28	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
4D246E434B5850B880AA7810FE02D661.taxon	distribution	Distribution. PERU. Ancash, Ayacucho, Cajamarca, Huan-cavellica, Junin, La Libertad and Pasco (Fig. 28).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
4D246E434B5850B880AA7810FE02D661.taxon	diagnosis	Diagnosis. Ayacucho can be differentiated from all other Metasarcidae genera by its short leg IV (femur IV length / DS length <1.5). Most of its species can be distinguished from other genera by having, alpha-type DSS; a femur of pedipalpus slightly flattened and males without a proapical spine (present in all other genera), an ocularium high and rounded and a DS densely granulated.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
146030CE3E8C5EF6B050660213E9D58E.taxon	description	Figs 2 H, 28	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
146030CE3E8C5EF6B050660213E9D58E.taxon	diagnosis	Diagnosis. It differs from other species of the genus with known females by having a high median spine in the free tergites I-II (Fig. 2 H).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
146030CE3E8C5EF6B050660213E9D58E.taxon	etymology	Derivatio nominis. The specific epithet of masculine gender, in the genitive form, dedicated to the German arachnologist Carl Friedrich Roewer (1881 - 1963), the author of original name in secondary homonymy.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
146030CE3E8C5EF6B050660213E9D58E.taxon	distribution	Distribution. (Fig. 28) PERU. Ancash. Cajacay.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
146030CE3E8C5EF6B050660213E9D58E.taxon	materials_examined	Material examined. Type material: Hototype ♀, ' PERU. Ancash. Cajacay, Rio Fortaleza, 2,700 m a. s. l., 5 / III / 1956, Weyrauch leg. (SMF RII 11645 / 28).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
A00A0E83A1E1552CB948724920D3FF42.taxon	description	Figs 2 I, 7 M, N, 15 G-I, 22 E, F, 28	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
A00A0E83A1E1552CB948724920D3FF42.taxon	description	Description. MALE: Measurements (n = 10) DSW: 1.7 - 3.0 (2.8); DSL: 4.0 - 4.3 (4.2); CL: 1.3 - 1.5 (1.5). FIVL: 2.6 - 2.8 (2.6). ChL: 1.2 - 2.5 (2.2). Coloration: (Fig. 22 E) Carapace and lateral margins of dorsal scutum yellowish. Scutal areas and posterior margin of dorsal scutum and free tergites brown, with yellowish spots. Legs trochanters yellow, other segments brown. Chelicerae yellow. Pedipalpus brownish. Dorsum: (Fig. 2 I) Alpha-type DSS. Anterior margin of the dorsal scutum completely covered with granules, with median elevation. Ocularium robust, with granules of equal size throughout its length in smaller males, larger males with a pair of small acuminate tubercles. Carapace densely covered with granules. Areas I-IV without larger granules. Posterior margin of the dorsal scutum and free tergites with rows of tubercles, in greater numbers in posterior margin than in free tergites. Chelicerae: (Fig. 2 I) Swollen in larger males (as in the holotype) and at similar size of females in smaller males. Segment I granulate. Segment II predominantly smooth; finger with one tooth. Segment III with one tooth. Pedipalpus: Small granules distributed on the dorsal surface. Trochanter with a ventrodistal setiferous tubercle. Femur with ventrobasal setiferous tubercles, one ventral row of five small setiferous tubercles, distributed throughout the length of segment except at apex. Tibia: prolateral I (i) Ii, retrolateral IIIII. Tarsus: prolateral IIII, retrolateral III. Venter: Coxae I-IV with scattered small granules. Genital area with few granules. Free sternites I-IV with rows of small granules. Anal operculum with granules sparsely distributed throughout its surface. Legs: (Figs 2 I, 7 M, N) Coxae I-II with an anterior and a posterior apophysis. Coxa IV with granules distributed throughout its length. Trochanters I-IV somewhat granulate, unarmed. Femora I-IV with granules densely distributed throughout their dorsal, retrolateral and prolateral surfaces, the ventral surface of femora I-III with two rows of granules, larger than other granules of those segments; femur IV with two ventral rows (in ⅔ distal) of tubercles that increase in size distally, whose size is more pronounced in large males. Patellae I-IV with granules distributed predominantly in the dorsal, retrolateral and prolateral faces, being scarcer ventrally. Tibiae I-IV with granules throughout their length, higher on the ventral side. Tarsal formula: (n = 10) 5 - 6 (6), 7 - 8 (8), 6, 6 - 7 (7). Penis: (Fig. 15 G-I) VP subrectangular, slightly convex, with lateroapical projections. MS C 1 - C 3 subapical, long and straight; MS A 1 median, long and straight (smaller than MS C); MS B 1 - B 2 sub basal, long and straight (MS B 1 longer than MS B 2). Lateral sacs long and apically acuminate, with long T 3 - like microsetae. Stylus with apex inflated, with several apical projections and a small dorsal projection. Dorsal process absent. Promontory convex. - FEMALE: Measurements (n = 12) DSW: 1.8 - 3.1; DSL: 4.0 - 4.2; CL: 1.3. FIVL: 2.8 - 3.0. ChL: 1.2 - 1.4 (Fig. 22 F) Chelicerae slightly smaller than those of small males. Presence of a proapical setiferous tubercle on pedipalpus femur, larger than the tubercles of the ventral surface. Femur IV slightly thinner, with smaller granules and distal two rows of tubercles absent. Tarsal formula: (n = 12) 6 - 7, 8, 6, 5 - 6.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
A00A0E83A1E1552CB948724920D3FF42.taxon	diagnosis	Diagnosis. Similar to Ayacucho glauberrochai sp. nov., A. pomacocha sp. nov., A. titschacki and A. vargasllosai sp. nov. in the following combination of characteristics: dorsal scutum densely granulate; ocularium and areas I-IV of DS unarmed or armed with tiny tubercles, slightly greater than granules; posterior margin of DS and free tergites I-III with median rows of acuminate tubercles (Fig. 2 I). It differs from the previously mentioned species in the following combination of characteristics: ocularium unarmed and densely granulate (Fig. 2 I); male femur IV with ventral surface armed (Fig. 7 N); penis VP subrectangular; three pairs of subapical MS C; two pairs of MS B (Fig. 15 G-I).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
A00A0E83A1E1552CB948724920D3FF42.taxon	etymology	Derivatio nominis. The specific epithet of feminine gender, in the genitive form, dedicated to Dr. Diana Silva D. (MUSM), for her contribution to the knowledge of Peruvian arachnids.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
A00A0E83A1E1552CB948724920D3FF42.taxon	distribution	Distribution. (Fig. 28) PERU. Pasco. Near to Cerro de Pasco.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
A00A0E83A1E1552CB948724920D3FF42.taxon	materials_examined	Material examined. Type material: Holotype ♂, ' PERU, Pasco, Near to Cerro de Pasco, 10 ° 41 ′ 39 ″ S 76 ° 13 ′ 08 ″ W, 23 / IV / 2011, R. Pinto-da-Rocha, A. Benedetti, J. Ochoa & D. Silva leg. (MUBI) - Paratypes 2 ♂, 5 ♀, ' ditto' (MUBI); Paratypes 2 ♂, 5 ♀, ' ditto' (MUSM); Paratypes 5 ♂, 11 ♀, ' ditto' (MZSP 36971).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
40D7F7A9EF4F57D094D22FA682E04E4C.taxon	description	Figs 3 A, 7 O, P, 12 G, H, 22 H, 28	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
40D7F7A9EF4F57D094D22FA682E04E4C.taxon	description	Description. MALE: Measurements (n = 3) DSW: 4.0 - 4.9 (4.9); DSL: 4.6 - 5.0 (5.0); CL: 1.6 - 2.0 (2.0). FIVL: 7.6 - 8.6 (8.6). ChL: 2.1 - 3.9 (3.9). Coloration: (Fig. 22 H) Carapace, lateral margins of DS, spines of scutal area III and legs dark brown. Reddish chelicera. Pedipalpus yellowish with black spots. Posterior margin of DS and free tergites I-III light yellow. Scutal areas light yellow, with series of dark spots. Dorsum: (Fig. 3 A) Gamma-type DSS. Anterior margin of the DS with a median elevation, with few granules sparsely distributed. Ocularium with medial depression; with a pair of spines and few granules near the eyes. Areas I-IV with few granules concentrated in the lateral regions; area III with a pair of spines directed backwards, located at the highest point in elevations of the integument, totally granular; area IV with a pair of small lateral tubercles. Posterior margins of DS smooth, with a pair of medium tubercles. Free tergites I-II with a pair of median tubercles; and a lateral pair of tiny tubercles. Free tergite III with two elongated median tubercles, larger than those of the free tergites I-II. Chelicerae: (Fig. 3 A) Swollen in large males (swollen in holotype); slightly larger than that of the females and small males. Segment I granulate; II with few granules; III with two teeth. Pedipalpus: Trochanter with two ventroapical setiferous tubercles, one large and one smaller. Femur with a row of six ventral setiferous tubercles in the basal ⅔. Patella with a small retroapical tubercle. Tibia: prolateral iiIii, retrolateral IIi. Tarsus: prolateral iiIiIi, retrolateral iIiIi. Venter: Coxa I with a row of 4 - 5 tubercles. Coxae II-IV with granules throughout their length. Genital area, free sternites and anal operculum with few granules. Legs: (Figs 3 A, 7 O, P) Coxae I-II each one with a retrolateral and prolateral apophyses. Coxa III with a prolateral apophysis. Coxa IV with setiferous granules distributed over the entire length. Trochanters I-III unarmed and with few granules. Trochanter IV with few granules and an apical retrolateral blunt tubercle. Femora I-III unarmed and with few granules. Femur IV with sparse granules; a retrolateral row of 15 - 17 acuminate tubercles over the entire length of the segment, except the base; a prolateral row of 10 - 12 acuminate tubercles along the distal ⅔; with four ventrodistal tubercles. Patellae I-III unarmed, with few granules. Patella IV with seven dorsal acuminate tubercles, the two largest apical ones; several granules on the ventral surface. Tibiae I-IV unarmed, with few granules. Tarsal formula: (n = 3) 7 - 8 (8), 12 - 13 (1 3), 10 - 15 (10), 11 - 13 (13). Penis: (Fig. 12 G, H) VP subrectangular, with distal margin straight and lateral projections; straight in lateral view. MS C 1 - C 3 subapical long and straight; MS A 1 median long and straight; MS B 1 sub basal long and straight. Lateral sacs long, apically acuminate, with long T 3 - like microsetae. Stylus with a non-swollen apex and without apical projections. Dorsal process conical, with acuminated apex. Promontory straight. - FEMALE: Measurements (n = 2) DSW: 4.2 - 4.5; DSL: 4.5 - 4.9; CL: 1.5 - 1.7. FIVL: 7.2 - 8.1. ChL: 1.7 - 1.8. Chelicerae not swollen. Femur IV with retrolateral row of 13 - 14 tubercles, smaller than those of the retrolateral row of the males; prolateral row and ventrodistal tubercles absent or in the form of granules. Patella IV with dorsodistal acuminate tubercles, but smaller than in males. Tarsal formula: (n = 2) 7, 11 - 12, 9 - 10, 10 - 12.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
40D7F7A9EF4F57D094D22FA682E04E4C.taxon	diagnosis	Diagnosis. It differs from other species of the genus by the combination of the following characteristics: Gamma-type DSS (Fig. 3 A); area III of DS with a pair of large spines (also present in A. pasolinii sp. nov.); male femur IV armed (Fig. 7 O, P); longer legs (ratio between FIVL and DSL greater than 1; 1.65 in A. spielbergi sp. nov.); and scutal areas light yellow, with series of dark spots (Fig. 22 H).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
40D7F7A9EF4F57D094D22FA682E04E4C.taxon	etymology	Derivatio nominis. The specific epithet of masculine gender, in the genitive form, dedicated to the American filmmaker, producer and screenwriter Steven Allen Spielberg.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
40D7F7A9EF4F57D094D22FA682E04E4C.taxon	distribution	Distribution. (Fig. 28) PERU. Cajamarca. Parque Nacional Cutervo.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
40D7F7A9EF4F57D094D22FA682E04E4C.taxon	materials_examined	Material examined. Type material: Holotype ♂, ' PERU, Cajamarca, Parque Nacional Cutervo, Puente Suro, 6 ° 12 ′ 10 ″ S 78 ° 44 ′ 22 ″ W, 22 / V / 2010, R. Pinto-da-Rocha & D. Silva leg. (MUSM) - Paratypes 2 ♂, 1 ♀, ' ditto' (MZSP 36995); Paratype 1 ♀, ' PERU, Cajamarca, Parque Nacional Cutervo, near to Cueva San Andreas, 22 / V / 2010, R. Pinto-da-Rocha & D. Silva leg. (MZSP 36996). Additional material: 1 ♂, ' PERU, Cajamarca, Parque Nacional Cutervo, San Andreas de Cutervo, 13 / VI / 1996, S. Cordova leg. (MUSM 0501248); 1 ♀, ' ditto' (MUSM 0501247).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
E08A66F2F1B05A1E8BE328BE42094967.taxon	description	Figs 3 B, 7 Q, R, 16 A-C, 23 A, B, 28	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
E08A66F2F1B05A1E8BE328BE42094967.taxon	diagnosis	Diagnosis. Similar to Ayacucho inermis comb. nov. because of two rows of tubercles on the femur IV (pro and retrolateral; Fig. 7 Q, R). Differs from Ayacucho inermis comb. nov. because femur IV with some lanceolate tubercles (Fig. 7 Q, R); ventral plate of penis with conspicuous lateral projections; long MS C on VP; lateral sacs with long T 3 - like microsetae, dorsal process long in relation to stylus (Fig. 16 A-C).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
E08A66F2F1B05A1E8BE328BE42094967.taxon	distribution	Distribution. (Fig. 28) PERU. Junin. Campanillaya. Hacienda Maraynioc. Palca	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
E08A66F2F1B05A1E8BE328BE42094967.taxon	materials_examined	Material examined. Type material: Of P. spiniger: Holotype ♂, ' PERU, Junin, Campanillaya, near to Palca in Rio Tarma, 2,600 m a. s. l., 06 / I / 1955, Weyrauch leg. (SMF RII 11418 / 24). Of P. serrifemur: Holotype ♂, ' PERU, Junin, Hacienda Maraynioc, east Tarma, Chanchamayo bay, 3,500 m a. s. l., 10 / X / 1956, Weyrauch leg. (SMF RII 12764 / 35) - Paratype 1 ♀ ' ditto' (SMF RII 12764 / 35). Additional material: 3 ♂, 3 ♀ ' PERU, Junin, Palca, 11 ° 21 ′ 28.9 ″ S 75 ° 33 ′ 23.9 ″ W, 19 / IV / 2011, R. Pinto-da-Rocha, A. Benedetti, J. Ochoa & D. Silva leg. (MUBI); 3 ♂, 2 ♀, ' ditto' (MUSM); 7 ♂, 5 ♀ ' ditto' (MZSP 36980).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
6EB909EC517954069403CC95046AA667.taxon	description	Figs 3 C, 7 S, T, 16 G-I, 23 C, D, 28	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
6EB909EC517954069403CC95046AA667.taxon	diagnosis	Diagnosis. Similar to Ayacucho querococha sp. nov. because armed tibia IV (Fig. 7 S, T). It differs from Ayacucho querococha sp. nov. by the set of the following characteristics: DSS with constriction II not-so-well marked (almost faint; Fig. 3 C); a pair of spines on ocularium (Fig. 3 C); coxa III with two apophyses (Fig. 3 C; only one apophysis in Ayacucho querococha sp. nov.); femur IV without probasal row of tubercles; tibia IV without basal row of tubercles (Fig. 7 S, T).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
6EB909EC517954069403CC95046AA667.taxon	etymology	Derivatio nominis. The specific epithet, a noun in apposition, in reference to the type locality, Tapacocha, Ancash, Peru, as well as in relation to the name of the genus where the species was originally described by Roewer (1957).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
6EB909EC517954069403CC95046AA667.taxon	distribution	Distribution. (Fig. 28) PERU. Ancash. Huascaran National Park; Tapacocha, Rio Fortaleza.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
6EB909EC517954069403CC95046AA667.taxon	materials_examined	Material examined. Type material: Holotype ♂, ' PERU, Ancash, Tapacocha | Rio Fortaleza, 3,200 m a. s. l., 04 / III / 1956, Weyrauch leg. (SMF RII 11644 / 27). Additional material: 2 ♂, 2 ♀ ' PERU, Ancash, Parque Nacional Huascaran, Chinancoche, 9 ° 04 ′ 18 ″ S 77 ° 38 ′ 38 ″ W | 15 / V / 2011, R. Pinto-da-Rocha & D. Silva leg. (MUSM); 3 ♂, 3 ♀ ' ditto' (MUBI); 5 ♂, 5 ♀ ' ditto' (MZSP 36976). 2 ♂, 4 ♀ ' PERU, Ancash, Parque Nacional Huascaran, Laguna Paron, 8 ° 59 ′ 43 ″ S 77 ° 40 ′ 41 ″ W | 16 / V / 2011, R. Pinto-da-Rocha & D. Silva leg. (MUSM); 2 ♂, 4 ♀ ' ditto' (MUBI); 4 ♂, 8 ♀ ' ditto' (MZSP 36977). 1 ♂, 2 ♀ ' PERU, Ancash, Parque Nacional Huascaran | Llaco, 9 ° 28 ′ 11 ″ S 77 ° 27 ′ 50.8 ″ W, 14 / V / 2011, R. Pinto-da-Rocha & D. Silva leg. (MUSM); 2 ♂, 3 ♀ ' ditto' (MUBI); 3 ♂, 6 ♀ ' ditto' (MZSP 36978).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
A7E8B110FB875A6CBAAC65042B2B7CE5.taxon	description	Figs 3 D, 7 U, V, 12 I, J, 23 E, 28	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
A7E8B110FB875A6CBAAC65042B2B7CE5.taxon	diagnosis	Diagnosis. Similar to Ayacucho glauberrochai sp. nov., A. pomacocha sp. nov., A. silvae sp. nov. and A. vargasllosai sp. nov. in the following combination of characteristics: dorsal scutum densely granulate; ocularium and areas I-IV of DS unarmed or armed with tiny tubercles, slightly greater than granules; posterior margin of DS and free tergites I-III with median rows of acuminate tubercles (Fig. 3 D). It differs from the previously mentioned species in the following combination of characteristics: ocularium with two small acuminate tubercles and densely granulate (Fig. 3 D); male femur IV unarmed (Fig. 7 U, V; unlike A. silvae sp. nov.); penis VP rectangular; five-six pairs of subapical MS C; one pair of MS A and B (Fig. 12 I, J).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
A7E8B110FB875A6CBAAC65042B2B7CE5.taxon	distribution	Distribution. (Fig. 28) PERU. Ayacucho. Ayacucho, Ocollo and Virgem de Cacharras de Cocha.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
A7E8B110FB875A6CBAAC65042B2B7CE5.taxon	materials_examined	Material examined. Type material. Holotype ♀, ' PERU, Ayacucho, Ayacucho, without date and leg. (SMF RII 8589 / 20) - Paratype ♀, ' ditto' (SMF RII 8589 / 20). Additional material. 1 ♂, 2 ♀ ' PERU, Ayacucho, Virgem de Cacharras de Cocha, 13 ° 01 ′ 47 ″ S 73 ° 52 ′ 14 ″ W, without date and leg. (MZSP 36972). 2 ♀ ' PERU, Ayacucho, Near Ocollo, 13 ° 19 ′ 53.0 ″ S 74 ° 30 ′ 17,6 ″ W, 29 / IV / 2011, R. Pinto-da-Rocha, A. Benedetti, J. Ochoa & D. Silva leg. (MUSM); 1 ♂, 2 ♀, ' ditto' (MUBI); 1 ♂, 5 ♀ ' ditto' (MZSP 36973).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
988E8E04A31257CDBF22AC6EDDBA9E3D.taxon	description	Figs 3 E, 7 W, X, 12 K, L, 28	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
988E8E04A31257CDBF22AC6EDDBA9E3D.taxon	diagnosis	Diagnosis. Similar to Ayacucho bambamarca comb. nov. and A. weyrauchi comb. nov. because three rows of spiniform tubercles in femur IV (Fig. 7 W, X). Differs from A. weyrauchi comb. nov. by having nine spiniform tubercles in retrolateral row of femur IV (Fig. 7 W, X); large tubercles in free tergites (Fig. 3 E), distal margin of ventral plate slightly concave (Fig. 12 K). Differs from A. bambamarca comb. nov. by having retrolateral row of tubercles covering only distal half distal of femur IV length (Fig. 7 W, X); areas I-IV unarmed (Fig. 3 E); distal margin of the VP without conspicuous lateral projections; dorsal process present in the penis (Fig. 12 K, L).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
988E8E04A31257CDBF22AC6EDDBA9E3D.taxon	etymology	Derivatio nominis. The specific epithet, an adjective in nominative singular, formed by Latin prefix tri - + Latin armatus, ta, tum (armed), in reference to three rows of tubercles on male femur IV.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
988E8E04A31257CDBF22AC6EDDBA9E3D.taxon	distribution	Distribution. (Fig. 28) PERU. Cajamarca. Cajamarca and near San Juan.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
988E8E04A31257CDBF22AC6EDDBA9E3D.taxon	materials_examined	Material examined. Type material: Holotype ♂, ' PERU, Cajamarca, near San Juan, between Chiclayo and Cajamarca, 1,900 m a. s. l., 06 / VII / 1956, Weyrauch leg. (SMF RII 12763 / 34). Additional material: 1 ♂, ' PERU, Cajamarca, Cajamarca, 3,200 m a. s. l., 6 ° 40 ′ S 78 ° 45 ′ W, 14 / III / 1958, Buschwald leg. (MUSM 0501238).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
DA445EFDF2255C97A4836F908B8D8E1E.taxon	description	Figs 3 H, 8 A, B, 17 A-C, 23 G, H, 28	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
DA445EFDF2255C97A4836F908B8D8E1E.taxon	diagnosis	Diagnosis. It differs from other species of the genus (with males known) by the set of the following characteristics: ocularium with a pair of spines; scutal areas unarmed; free tergites I-III with a pair of tubercles (Fig. 3 H); male femur IV with a retroventral row of long acuminate tubercles (Fig. 8 A, B); penial lateral sacs without T 3 - like microsetae (Fig. 17 A-C).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
DA445EFDF2255C97A4836F908B8D8E1E.taxon	distribution	Distribution. (Fig. 28) PERU. Cajamarca. Cutervo.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
DA445EFDF2255C97A4836F908B8D8E1E.taxon	materials_examined	Material examined. Type material: Holotype ♂, ' PERU, Cajamarca, Cutervo, 15 / VI / 1956, Weyrauch leg. (SMF RII 11648 / 31) - Paratypes 2 ♂, 2 ♀ ' ditto' (SMF RII 11648 / 31). Additional material: 1 ♂, 1 ♀ ' PERU, Cajamarca, near Cutervo, 6 ° 20 ′ 42 " S 78 ° 49 ′ 19 ″ W, 20 / V / 2010, R. Pinto-da-Rocha & D. Silva leg. (MUSM); 2 ♂, 1 ♀ ' ditto' (MZSP 36981).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
F64697E7AAAF5C5A845C7FC2C4942AEC.taxon	description	Figs 3 G, 8 C, D, 13 A, B, 23 F, 28	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
F64697E7AAAF5C5A845C7FC2C4942AEC.taxon	description	Description. MALE: Measurements (n = 1) DSW: 3.3; DSL: 4.6; CL: 1.7. FIVL: 4.7. ChL: 2.5. Coloration: (Fig. 23 F) Chelicerae, pedipalpus, carapace and lateral margin of dorsal scutum and legs orange. Scutal areas, posterior margin of dorsal scutum and free tergites brownish. Dorsum: (Fig. 3 G) Alpha-type DSS, with shallower constrictions. Anterior margin of carapace with median elevation totally covered with granules. Ocularium with tiny acuminate tubercles. Areas I-IV with one pair of small median tubercles (larger than the pair of the ocularium). Posterior margin of dorsal scutum with a transverse row of four tubercles. Free tergites I- - III with a row of 3 - 6 tubercles, larger than tubercles of areas I-IV. Chelicerae: (Fig. 3 G) Swollen in male. Segment I densely covered with granules throughout its length. Segment II predominantly smooth, with some hairs on the front surface; finger with four teeth. Segment III with two teeth. Pedipalpus: With very small granules sparsely distributed on the dorsal surface of the femur-tibia. Trochanter with a distal ventral setiferous tubercle. Femur with a ventrobasal setiferous tubercle and a row of five small ventromedian setiferous tubercles. Tibia: prolateral IIII, IiI; retrolateral iII (II), iIII. Tarsus: prolateral IIi; retrolateral Iii, Ii. Venter: Coxae I-IV and stigmatic area with sparse small granules; coxa I with a longitudinal row of small setiferous tubercles. Genital area smooth. Free sternites I-IV and anal operculum with sparse small granules. Legs: (Figs 3 G, 8 C, D) Coxae I-III each one with a retrolateral and a prolateral apophysis. Coxa IV densely granulate, unarmed. Trochanters I-IV densely granulate, unarmed. Femora I-IV densely granulate throughout their dorsal, retrolateral and prolateral faces. Femora I-III with two rows of granules in the ventral face, larger than other granules of these segments. Femur IV with two ventral rows of granules that increase in size apically, greater than other granules of femur IV. Patellae I-IV densely granulate. Tibiae I-IV granulate (larger granules on ventral face). Tarsal segmentation: (n = 1) 7, 11, 7, 7. Penis: (Fig. 13 A, B) VP subrectangular in dorsal view, with distal half larger than basal half; distal margin straight. MS C 1 - C 7 (C 8) apical, long and curved; MS A 1 median placed, long and straight (smaller than MS C); MS B 1 sub basal, long and straight (smaller than MS A). Lateral sacs long, apically tapered; with long T 3 - like microsetae. Stylus apically inflated, with a conspicuous ventral projection. Dorsal process absent. Promontory straight. - FEMALE: Measurements (n = 3) DSW: 3.0 - 3.1; DSL: 4.4 - 4.5; CL: 1.7. FIVL: 4.5 - 4.7. ChL: 2.2 - 2.3. Areas I-IV with one pair of small median tubercles, smaller than in males. Pedipalpus with ventral tubercles slightly smaller than in males. Tibia: prolateral IiI; retrolateral iIIIi, iIII. Tarsus: retrolateral iIIi, iIii. Genital area with sparse granules. Femur IV with ventral tubercles smaller than in males. Tarsal segmentation (n = 3): 7,11,7,8.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
F64697E7AAAF5C5A845C7FC2C4942AEC.taxon	diagnosis	Diagnosis. Similar to Ayacucho glauberrochai sp. nov. A. pomacocha sp. nov., A. silvae sp. nov., and A. titschacki in the following combination of characteristics: dorsal scutum densely granulate; ocularium and areas I-IV of DS unarmed or armed with tiny tubercles, slightly greater than granules; posterior margin of DS and free tergites I-III with median rows of acuminate tubercles (Fig. 3 G); femur IV of males without strong armature (except in ventral surface of femur IV in A. silvae sp. nov.; Fig. 8 C, D). It differs from the previously mentioned species in the following combination of characteristics: ocularium with a pair of small tubercles; areas I-IV with a pair of tubercles (larger than those in ocularium; Fig. 3 G); male femur IV unarmed (unlike A. silvae sp. nov.; Fig. 8 C, D); and penis VP subrectangular, with 7 - 8 apical MS C (Fig. 13 A, B).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
F64697E7AAAF5C5A845C7FC2C4942AEC.taxon	etymology	Derivatio nominis. The specific epithet of masculine gender, in the genitive form, dedicated to the Peruvian writer, politician, journalist, essayist, filmmaker, college professor and Nobel Prize winner Jorge Mario Pedro Vargas Llosa (born 1936), more commonly known as Mario Vargas Llosa.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
F64697E7AAAF5C5A845C7FC2C4942AEC.taxon	distribution	Distribution. (Fig. 28) PERU. Junin. Cruce Mina.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
F64697E7AAAF5C5A845C7FC2C4942AEC.taxon	materials_examined	Material examined. Type material: Holotype ♂, ' PERU, Junin, Cruce Mina, Cemento Andino, 11 ° 22 ′ 45.4 ″ S 75 ° 52 ′ 43.5 ″ W, | 22 / IV / 2011, R. Pinto-da-Rocha, A. Benedetti, J. Ochoa & D. Silva leg. (MUBI) - Paratype 1 ♀, ' ditto' (MUBI); Paratype 1 ♀, ' ditto' (MZSP 36975); Paratype 1 ♂, 1 ♀, ' ditto' (MZSP 73007).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
968A31B5C99D54DDB93BEA656D95B60E.taxon	description	Figs 3 F, 8 E, F, 13 C, D, 28	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
968A31B5C99D54DDB93BEA656D95B60E.taxon	diagnosis	Diagnosis. Similar to Ayacucho bambamarca comb. nov. and A. triarmatus nom. nov. for presenting three rows of acuminate tubercles in the male femur IV (Fig. 8 E, F). It differs from A. bambamarca comb. nov. and A. triarmatus nom. nov. in having a maximum of four acuminate tubercles in the retrolateral row (Fig. 8 E, F); smaller tubercles in ocularium; free tergites with small tubercles (with wide base); areas I-IV with few granulation (Fig. 3 F). It differs from A. bambamarca comb. nov. in having scutal areas unarmed (Fig. 3 F).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
968A31B5C99D54DDB93BEA656D95B60E.taxon	distribution	Distribution. (Fig. 28) PERU. Cajamarca. Cajamarca.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
968A31B5C99D54DDB93BEA656D95B60E.taxon	materials_examined	Material examined. Type material: Of C. weyrauchi: Holotype ♂, ' PERU, Cajamarca, Cajamarca, | 2,750 m a. s. l., without date, Weyrauch leg. (SMF RII 10128 / 22) - Paratypes 2 ♀, ' ditto' (SMF RII 10128 / 22). Of C. affinis: Holotype ♂, ' PERU, Cajamarca, near Cajamarca, 24 km from Cajamarca, road to Celendin, 3,150 m a. s. l., 04 / VIII / 1956, Weyrauch leg. (SMF RII 11646 / 29) - Paratypes 5 ♀, ' ditto' (SMF RII 11646 / 29).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
3E7DF42329B45868B4C3729423361A02.taxon	description	Figs 4 A, 11 A, B, 18 A-C, 24 A, B, 29 A	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
3E7DF42329B45868B4C3729423361A02.taxon	diagnosis	Diagnosis. Huancabamba gen. nov. can be distinguished from all other Metasarcidae genera by the combination of following: Kappa-type DSS; male femur IV at least 1.6 longer than DS length and with low tubercles; ocularium with two low tubercles; area III with spines; coxa IV apex reaching area III; penis with more than 13 MS C; a dry-mark in depression of ocularium, carapace, and lateral region to ocularium.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
3E7DF42329B45868B4C3729423361A02.taxon	description	Description. Kappa-type DSS, with carapace very wide, constriction I weakly marked and coda undefined, coalescing with mid-bulge. Ocularium low, medially depressed. Ocularium with two low tubercles. Areas of dorsal scutum moderately tuberculate. Area I undivided. Area III armed with two high spines. Posterior margin of DS armed with a pair of high tubercles. Coda short, without constriction. Coxa IV reaching area III. Coxa IV unarmed (Fig. 4 A). Femur IV about same size as dorsal scutum length (Figs 11 A, B, 24 A). More than 13 MS C. Penis stylus thin thickness. Penis VP thin thickness (Fig. 18 A-C).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
3E7DF42329B45868B4C3729423361A02.taxon	etymology	Derivatio nominis. The genus name, a noun in the nominative singular, from Quechua, huanca (stone) + bamba (plain). It refers to Huancabamba depression, an interruption of the Andean Mountains, located between southern Ecuador and northern Peru. This depression constitutes a biogeographic barrier between the northern Andes and the central Andes. The name is a reference to this Metasarcidae genus occurred fartherst north. Gender: feminine.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
3E7DF42329B45868B4C3729423361A02.taxon	distribution	Distribution. (Fig. 29 A) PERU. Cajamarca.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
AB4B8BE579B25EDD928A54C11536A1B0.taxon	description	Figs 4 A, 11 A, B, 18 A-C, 24 A, B, 29 A	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
AB4B8BE579B25EDD928A54C11536A1B0.taxon	description	Description. MALE: Measurements (n = 3) DSW: 4.0 - 4.2 (4.2); DSL: 3.6 - 4.3 (4.1); CL: 1.5 - 2.0 (2.0). FIVL: 6.6 - 7.3 (7.3). ChL: 1.2 - 3.6 (3.2). Coloration: (Fig. 24 A) Reddish carapace with blackish sides. Dorsal scutum and free tergites brownish orange. Lateral margins of DS, small tubercles on scutal areas, spines on area III and free tergites yellow. A dry-mark on depression of ocularium, carapace, and lateral region to ocularium. Pedipalpus and chelicerae yellow with small black spots. Legs I-II with coloration similar to dorsal scutum. Legs III-IV reddish brown. Dorsum: (Fig. 4 A) Row of granules on the anterior margin of dorsal scutum and the median elevation. Granules more concentrated on carapace sides and on posterior region to ocularium. Ocularium with a relative strong depr ession; with pair of small tubercles and few granules near the eyes. Dorsal scutum with scutal grooves almost inconspicuous; grooves I and II more visible than others; granules sparsely distributed throughout scutal areas. Areas I-IV with a lateral pair of tubercles; I with a pair of median tubercles, larger than those on the lateral portion of scutal areas; III with a pair of large median blunt spines, base with small tubercles. Lateral margins of DS sparsely covered with granules throughout extension. Posterior margin of DS and free tergites I-III each one with a median pair of tubercles and a row of few granules. Chelicerae: (Fig. 4 A) Swollen in large males, similar to females in the small males. Segment I with three small prolateral tubercles and one large tubercle located medially or retrolaterally. Segment II with small setiferous granules, concentrated more distally; one tooth. Segment III with two teeth. Pedipalpus: Trochanter with a ventroapical setiferous tubercle. Femur with a ventral row of 6 - 7 setiferous tubercles of irregular size, the largest being the most apical; one proapical spine. Patella with a promedian spine. Tibia: retrolateral iiIIIi, prolateral IIII. Tarsus: retrolateral iIiIi, prolateral iIIi. Venter: Coxa I with a median row of 5 - 6 setiferous tubercles, the most median of which is larger; III-IV with small granules sparsely distributed. Rows of tubercles between the coxae II-III and III-IV. Genital area, anal operculum and free sternites with few granules. Legs: (Figs 4 A, 11 A, B) Coxa I-II each one with a prolateral and a retrolateral apophysis. Coxa III unarmed. Coxa IV with setiferous granules distributed throughout its surface, with the largest clustered apically. Trochanters I-IV unarmed and with a few granules. Femora I-III unarmed and with few granules. Femur IV unarmed and with more abundant and larger granulation than those of the other femora. Patellae-tibiae I-IV unarmed with few granules. Tarsal segmentation: (n = 3) 7 - 8 (7), 16 - 17 (17), 15 - 17 (17), 16 - 17 (17). Penis: (18 A-C) VP rectangular, with distal margin straight; VP wide and robust in lateral view. MS C 1 - C 13 (C 14) subapical long and apically curved; MS A 1 median short and straight; MS B 1 - B 2 median-basal long and straight (longer than MS A, shorter than MS C); MS D 1 very short, placed dorsally, near MS C. Lateral sacs very short (twice as long as wide), apically blunt, with short T 3 - like microsetae. Stylus with apex dorsoventrally expanded and with apical projections. Dorsal process absent. Promontory convex. - FEMALE: Measurements (n = 3) DSW: 4.1 - 4.3; DSL: 4.1 - 4.3; CL: 1.6 - 1.8 FIVL: 6.9 - 7.0. ChL: 1.6 - 1.7. (Fig. 24 B) Chelicerae small, not swollen. Femur IV with less dense granulation, with granules smaller than those of males. Tarsal segmentation: (n = 3) 7, 13 - 16, 15 - 17, 15 - 17.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
AB4B8BE579B25EDD928A54C11536A1B0.taxon	diagnosis	Diagnosis. As for the genus.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
AB4B8BE579B25EDD928A54C11536A1B0.taxon	etymology	Derivatio nominis. The specific epithet of masculine gender, in the genitive form, dedicated to the American director, producer and screenwriter Stanley Kubrick (1928 - 1999).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
AB4B8BE579B25EDD928A54C11536A1B0.taxon	distribution	Distribution. (Fig. 29 A) PERU. Cajamarca. Near Cutervo.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
AB4B8BE579B25EDD928A54C11536A1B0.taxon	materials_examined	Material examined. Type material: Holotype ♂, ' PERU, Cajamarca, near Cutervo, 06 ° 20 ′ 42 ″ S 78 ° 49 ′ 19 ″ W, 20 / V / 2011, R. Pinto-da-Rocha & D. Silva leg. (MUSM) - Paratypes 7 ♀, ' ditto' (MUSM); Paratypes 2 ♂, 7 ♀, ' ditto' (MZSP 36989).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
20277AF3E0E556B6A7384331C285FC41.taxon	description	Figs 4 B, 9 A, B, 13 E, F, 30	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
20277AF3E0E556B6A7384331C285FC41.taxon	diagnosis	Diagnosis. It differs from other species of the genus by silver-white patches on carapace, area I and lateral margins of dorsal scutum; femur IV with two rows of acuminate large tubercles, a retroventral one with 35 - 37 tubercles and a proventral one with 31 - 32 tubercles (Fig. 9 A, B).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
20277AF3E0E556B6A7384331C285FC41.taxon	distribution	Distribution. (Fig. 30) PERU. Cusco. La Convencion and Urubamba provinces.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
20277AF3E0E556B6A7384331C285FC41.taxon	materials_examined	Material examined. Type material: Holotype ♂, ' PERU, Cusco, Urubamba province, Ollantaytambo district, Abra de Malaga, Cancayoc, 3,000 m a. s. l., 13 ° 16 ′ S 72 ° 16 ′ W, 27 / VIII / 1995, J. Ochoa leg. (MACN 9549) - Paratype 1 ♀, ' ditto' (MACN 9550). Additional material: 2 ♂, 4 ♀, ' PERU, Cusco, La Convencion province, Carrizales, 3,250 m a. s. l., 13 / IV / 2014, R. Cruz, S. Bejar & M. Serrano leg. (MZSP 76552).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
EAAE10797921525583AEEFA62531F565.taxon	description	Figs 4 C, 9 C, D, 13 G, H, 30	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
EAAE10797921525583AEEFA62531F565.taxon	diagnosis	Diagnosis. It differs from other species of the genus by the set of following characters: DS without silver-white coloration; DS slightly granulate; area I with a pair of lateral tubercles; area III with a pair of spines (Fig. 4 C); male femur IV with a retrolateral and a prolateral rows of tubercles at ⅔ basal (Fig. 9 C, D); penis with short lateral sacs (Fig. 13 G, H).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
EAAE10797921525583AEEFA62531F565.taxon	distribution	Distribution. (Fig. 30) PERU. Cusco. Paucartambo province, Manu National Park.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
EAAE10797921525583AEEFA62531F565.taxon	materials_examined	Material examined. Type material: Holotype ♂, ' PERU, Cusco, Paucartambo province, Manu National Park, road to Paucartambo-Pilcopata, 2,900 m a. s. l., 13 ° 01 ′ 40 ″ S 71 ° 16 ′ 40 ″ W | 19 / II / 1990, A. Cano & D. Silva leg. (MUSM 410) - Paratypes 2 ♀, ' ditto' (MUSM 410); Paratypes 1 ♂, 2 ♀, ' PERU, Cusco, Paucartambo province, Manu National Park, | road to Paucartambo-Pilcopata, 2,900 m, 13 ° 01 ′ 40 ″ S 71 ° 16 ′ 40 ″ W, 14 / II / 1990, A. Cano & D. Silva leg. (MNRJ 5315 ex- MUSM 410).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
F00E3160B87F543380FC151D4683FE95.taxon	description	Figs 4 B-F, 9, 13 E-J, 18 D-I, 30	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
F00E3160B87F543380FC151D4683FE95.taxon	diagnosis	Diagnosis. Incasarcus can be differentiated from other Metasarcidae genera by the combination of following: alpha-type DSS; males with a proapical spine on the pedipalpus femur; area I undivided; male femur IV at least 1.6 longer than dorsal scutum; ocularium with two low tubercles or spines; area III with spines; coxa IV apex reaching area IV or posterior margins of DS; penis with less than 10 MS C and stylus thin thickness.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
F00E3160B87F543380FC151D4683FE95.taxon	distribution	Distribution. (Fig. 30) PERU. Cusco.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
303412CD267E59B7974ED627B3089505.taxon	description	Figs 4 D, 9 J-N, 18 D-F, 30	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
303412CD267E59B7974ED627B3089505.taxon	diagnosis	Diagnosis. It differs from other species of the genus by the set of following characters: DS without silver-white coloration; DS granulate; ocularium, areas I-IV and free tergites I-III unarmed (Fig. 4 D); male femur IV with one ventral row of 23 - 24 acuminate tubercles, a prolateral row of 5 - 7 tubercles and a prodorsal row of 7 - 8 tubercles (virtually inconspicuous in smaller males; Fig. 9 J-N).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
303412CD267E59B7974ED627B3089505.taxon	distribution	Distribution. (Fig. 30) PERU. Cusco. Urubamba province.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
303412CD267E59B7974ED627B3089505.taxon	materials_examined	Material examined. Type material: Holotype ♂, ' PERU, Cusco, Urubamba Province, Huayllabamba district, Yanacocha, Huayocari, Huayoccare, 3,000 - 4,000 m a. s. l., 13 ° 20 ′ S 72 ° 02 ′ W, 14 / XI / 1992, J. C. Chaparro leg. (MACN 9551) - Paratype ♀, ' ditto' (MACN 9552); Paratypes 2 ♀, ' PERU, Cusco, Urubamba Province, Huayllabamba district, Yanacocha, Huayocari, Huayoccare, 3,000 - 4,000 m a. s. l., 13 ° 20 ′ S 72 ° 02 ′ W | 17 / VI / 1995, J. A. Ochoa leg. (MACN 9553); Paratype 2 ♂, ' ditto' (MACN 9554); Paratype 1 ♂, ' ditto' (MNRJ 5401); Paratype 1 ♂, ' ditto' (MNRJ 5402 ♀).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
A8E6C7D4795B522B981AFA465181D2BB.taxon	description	Figs 4 E, 9 E, F, 13 I, J, 30	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
A8E6C7D4795B522B981AFA465181D2BB.taxon	diagnosis	Diagnosis. It differs from other species of the genus by the set of following characters: DS with white coloration on carapace (behind ocularium) and area I; DS granulate; ocularium and areas II and IV unarmed; area I with a pair of tubercles; area III with a pair of spines; free tergites I-III with a row of tubercles (Fig. 4 E); male femur IV with a retroventral row of 28 - 30 acuminate tubercles and an apical proventral row with four tiny tubercles (Fig. 9 E, F).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
A8E6C7D4795B522B981AFA465181D2BB.taxon	distribution	Distribution. (Fig. 30) PERU. Cusco. Winayhuaina.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
A8E6C7D4795B522B981AFA465181D2BB.taxon	materials_examined	Material examined. Type material: Holotype ♂, ' PERU, Cusco, Winayhuaina, Inca trail, 2,700 - 3,100 m a. s. l., 13 ° 09 ′ S 72 ° 31 ′ W, 10 / II / 1990, D. Silva leg. (MUSM 408).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
FC50544FA5EE5F8692A716D33BD72276.taxon	description	Figs 4 F, 9 G-I, 18 G-I, 30	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
FC50544FA5EE5F8692A716D33BD72276.taxon	diagnosis	Diagnosis. It differs from other species of the genus by the set of following characters: DS without white spots on DS; DS granular; ocularium unarmed and with few granules or smooth; areas I, II and IV unarmed; area III with a pair of spines; free tergites I-III with a row of tubercles, unarmed (Fig. 4 F); male femur IV with a retroventral row of 30 - 35 acuminate tubercles (Fig. 9 G-I).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
FC50544FA5EE5F8692A716D33BD72276.taxon	distribution	Distribution. (Fig. 30) PERU. Cusco. Urubamba province.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
FC50544FA5EE5F8692A716D33BD72276.taxon	materials_examined	Material examined. Type material: Holotype ♂, ' PERU, Cusco, Urubamba province, Machu Picchu, 2,200 - 2,500 m a. s. l., 13 ° 07 ′ S 72 ° 34 ′ W, 25 / II / 1994, J. Ochoa & J. Achicahuala leg. (MACN 9547) - Paratype ♀, ' ditto' (MACN 9548); ParatypeS 2 ♀, ' ditto' (MNRJ 5400). Additional material: 2 ♂, 3 ♀ ' PERU, Cusco' (MUBI).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
A37B09607DFD576F9B50FAA9DF224A1B.taxon	description	Figs 6 A, 11 C, D, 17 D-F, 24 C, D, 29 A	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
A37B09607DFD576F9B50FAA9DF224A1B.taxon	description	Description. MALE: Measurements (n = 10) DSW: 4.6 - 5.5 (5.5); DSL: 5.0 - 5.8 (5.6); CL: 1.1 - 1.5 (2.2). FIVL: 10.8 - 12.4 (12.4). ChL: 1.7 - 5.2. Coloration: (Fig. 24 C) Body reddish brown. Dry-marks on the carapace, grooves of dorsal scutum and free tergites. Pedipalpus, chelicerae and legs brown. Dorsum: (Fig. 6 A) Anterior margin of carapace with median elevation, with a few sparsely distributed granules. Ocularium with median saddle-shape depression; a pair of short median spines, and two pairs of tubercles near the eyes; slightly granulate. Carapace with scattered tubercles. Areas I-IV covered with scattered tubercles. Area I divided; with a pair of small median tubercles. Area II invading area I; unarmed. Area III with two long parallel spines, directed posteriorly, located at elevations of integument, totally granulate. Area IV unarmed. Posterior margin of dorsal scutum with two median acuminate tubercles and two small granules. Free tergites I-III with a pair of acuminate tubercles, medially in tergites I-II and laterally in tergite III. Chelicerae: (Fig. 6 A) Conspicuously swollen in large males (as in the holotype), to a lesser extent in smaller males. Segment I granulate. Segment II with few granules; finger with three teeth. Segment III with one tooth. Pedipalpus: Granules on the ventral surface of the femur. Trochanter with a ventroapical setiferous tubercle. Femur with a ventral row of nine large setiferous tubercles (except at the apex), larger in the middle portion; one proapical spine. Patella with a small proapical setiferous tubercle. Tibia: prolateral iIiI, retrolateral IIII. Tarsus: prolateral iIiIi, retrolateral iII. Venter: Coxa I with two rows with 6 - 7 large tubercles. Coxae II-III with two rows of granules. Coxa IV with scattered granules. Row of five small tubercles between the coxae III-IV. Genital area slightly granulate. Free sternites with few granules. Anal operculum granulate. Legs: (Figs 6 A, 11 C, D) Coxae I-II each one with a prolateral and a retrolateral apophyses. Coxa III with a prolateral apophysis Coxa IV with setiferous granules sparsely distributed throughout its surface; unarmed. Trochanters I-III unarmed and with sparse granules. Trochanter IV with few granules; a large retroapical apophysis; two small retrolateral tubercles medially near the retroapical apophysis. Femora I-III with small granules. Femur IV with granules sparsely distributed throughout its length; a retrolateral row of 24 - 26 acuminate tubercles varying in size throughout femur length. Patellae I-IV unarmed, with few granules. Tibiae I-III few granulate; unarmed. Tibia IV with a retrolateral row of 15 - 16 acuminate tubercles, covering the entire length of segment; two dorsal small tubercles. Tarsal segmentation: (n = 5) 9 - 10 (10), 20 - 23 (23), 11 - 12 (12), 11 - 15 (14). Penis: (Fig. 17 D-F) VP rectangular, with distal margin straight; a distal median projection with half the width of VP; Ventral plate strongly thick and straight in lateral view. MS C 1 - C 6 (C 7) subapical long and well curved; MS A 1 - A 2 median to sub basal, long and straight; MS D 1 very short and straight, dorsally placed near MS A. Lateral sacs long, apically tapered; with short T 3 - like microsetae. Stylus elongate and flattened laterally, with lateral supapical filiform projections. Dorsal process absent. Promontory slightly convex. - FEMALE: Measurements (n = 10) DSW: 4.9 - 5.3; DSL: 5.3 - 5.5; CL: 1.0 - 1.2. FIVL: 11.5 - 12.5. ChL: 2.3 - 2.7. (Fig. 24 D) Chelicerae not swollen, smaller or equal to the small males. Areas of the dorsal scutum more granular than that of males. Trochanter IV with retroapical apophysis shorter than the male; without prominent tubercles. Femur IV unarmed. Tarsal segmentation: (n = 10) 8, 19 - 21, 11 - 12, 12 - 13.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
A37B09607DFD576F9B50FAA9DF224A1B.taxon	diagnosis	Diagnosis. Similar to Lumieria woodyalleni gen. et sp. nov. because ocularium is armed with a pair of spines, area I divided, area III with a great pair of spines, free tergites with large tubercles (Fig. 6 A) and femur-tibia IV with a retrolateral row of acuminate tubercles (Fig. 11 C, D). Differs from Lumieria woodyalleni gen. et sp. nov. because the area II invading the area I; increased granulation on carapace and areas of dorsal scutum (Fig. 6 A); male femur IV with less than 20 retrolateral acuminate tubercles (Fig. 11 C, D); VP with distal margin straight and with a distal median projection (Fig. 17 D-F).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
A37B09607DFD576F9B50FAA9DF224A1B.taxon	etymology	Derivatio nominis. The specific epithet of masculine gender, in the genitive form, dedicated to the Italian filmmaker, editor, screenwriter, painter and writer Michelangelo Antonioni (1912 - 2007).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
A37B09607DFD576F9B50FAA9DF224A1B.taxon	distribution	Distribution. (Fig. 29 A) PERU. Junin. Near Comas.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
A37B09607DFD576F9B50FAA9DF224A1B.taxon	materials_examined	Material examined. Type material: Holotype ♂, ' PERU, Junin, Centro Turistico Ilpa, near Comas, 11 ° 42 ′ 37.1 ″ S 75 ° 04 ′ 20.2 ″ W, 27 / IV / 2011, R. Pinto-da-Rocha, A. Benedetti, J. Ochoa & D. Silva leg. (MUBI) - Paratypes 5 ♂, 5 ♀, ' ditto' (MUSM); Paratypes 11 ♂, 11 ♀, ' ditto' (MZSP 36982); Paratypes 4 ♂, 6 ♀, ' ditto' (MUBI).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
393C453EA15457D4BEC083540A2EA2E5.taxon	description	Figs 6 A, B, 11 C-F, 14 A, B, 17 D-F, 24 C-F, 29 A, B	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
393C453EA15457D4BEC083540A2EA2E5.taxon	diagnosis	Diagnosis. Lumieria gen. nov. can be distinguished from all other Metasarcidae genera by the combination of following: Kappa-type DSS; area I divided; ocularium medially depressed, with two high tubercles, area III with two spines; posterior margin and free tergites with one or two higher tubercles; femur IV much longer than dorsal scutum length; penis VP and stylus robust and thick in lateral view; conspicuous dry-marks on the carapace, grooves of DS and free tergites.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
393C453EA15457D4BEC083540A2EA2E5.taxon	description	Description. Kappa-type DSS, with constriction I well marked and constriction II absent. Coda undefined, coalescing with mid-bulge. Ocularium low, medially depressed. Ocularium unarmed, small tuberculate or smooth. Areas of dorsal scutum small to moderately tuberculate. Area I divided in two halves. Area III armed with two high spines. Posterior margin armed with one or a pair of high tubercles. Coda short, without constriction. Coxa IV reaching area IV or posterior margin. Coxa IV unarmed (Figs 6 A, B, 24 C, E). Femur IV much longer than dorsal scutum length (Figs 11 C-F, 24 C, E). Less than 10 macrosetae C. Penis stylus elongate, robust, very thick and flattened latterally (with a almost subrectangular or claviform appearance in the lateral view), with apical or subapical projections. Penis VP thick in lateral view, with dorsal side with a robust curvature resulting in a laterally convex appearance (Figs 14 A, B, 17 D-F).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
393C453EA15457D4BEC083540A2EA2E5.taxon	etymology	Derivatio nominis. The genus name, a noun in the nominative singular, is derived from Auguste Marie Louis Nicholas Lumiere (1862 - 1954) and Louis Jean Lumiere (1864 - 1948), the Lumiere brothers, who were the inventors of cinematograph, being frequently referred like the parents of the " Cinema ". Gender feminine.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
393C453EA15457D4BEC083540A2EA2E5.taxon	distribution	Distribution. (Fig. 29) PERU. Junin.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
723040C3E1B4504996E34788507261CA.taxon	description	Figs 6 B, 11 E, F, 14 A, B, 24 E, F, 29 B	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
723040C3E1B4504996E34788507261CA.taxon	description	Description. MALE: Measurements (n = 1) DSW: 5.4; DSL: 5.8; CL: 1.3. FIVL: 11,8. ChL: 3.5. Coloration: (Fig. 24 E) Pedipalpus and chelicerae brown-orange, dark in tone. DS, free tergites and legs reddish brown; dry-marks around the scutal grooves and on the carapace. Distal parts of legs and lateral parts of free tergites dark brown. Dorsum: (Fig. 6 B) Anterior margin of carapace with median elevation, covered with a few scattered granules. Ocularium with pronounced median depression; a pair of spines, the right one bifurcated. Area I divided (although the division is barely conspicuous in the region close to the longitudinal groove that divides the area); with a lateral and a median pair of small tubercles. Area II with two lateral tubercles. Area III with two long spines, directed posteriorly, located at elevations of integument, totally granulate. Area IV with five sparse tubercles. Lateral margins of DS with granules, except on carapace. Posterior margin of the DS and free tergite I smooth, each one with an acuminate tubercle. Free tergites II-III with two lateral acuminate tubercles. Chelicerae: (Fig. 6 B) Segment I granular. Segment II with few granules; finger with four teeth. Segment III with three teeth. Pedipalpus: Coxa with 1 - 2 retrodorsal tubercles. Trochanter with a ventroapical setiferous tubercle. Femur with a ventrobasal setiferous tubercle; a ventral row of 3 - 4 small granules; a small proapical tubercle. Patella with a small proapical tubercle. Tibia: prolateral II, retrolateral iIii. Tarsus: prolateral Iii, retrolateral Iii. Venter: Coxa I with a median row of 4 - 5 setiferous tubercles. Coxae II-IV with granules throughout their surface. Rows of small tubercles between the coxae II-III and III-IV. Genital area slightly granulate. Free sternites with rows of small granules. Anal operculum granulate. Legs: (Figs 6 B, 11 E, F) Coxae I-II each one with a prolateral and a retrolateral apophysis. Coxa III with a prolateral apophysis. Coxa IV with few sparse granules. Trochanters I-III smooth. Trochanter IV with 1 - 2 retroapical tubercles. Femur I-III unarmed, with small granules. Femur IV with sparse granules; a retrolateral row of 24 tubercles, these of varying size, most with acuminate apex. Patellae I-IV unarmed, with few granules. Tibiae I-III unarmed and few granular. Tibia IV with a retrolateral row of 16 acuminate tubercles along the basal 3 / 4. Tarsal segmentation: (n = 1) 8, 18, 10, 11. Penis: (Fig. 14 A, B) VP rectangular, with distal margin slightly concave; VP strongly thick and straight in lateral view. MS C 1 - C 5 subapical long and straight (some with curved apex); MS A 1 basal, short and straight. Lateral sacs long, with acuminate apex; with short T 3 - like microsetae. Stylus extended and wide, with swollen apical region; with many small apical projections. Dorsal process absent. Promontory slightly convex. - FEMALE: Measurements (n = 1) DSW: 5.5; DSL: 6.4; CL: 1.5. FIVL: 11.5. ChL: 1.5. (Fig. 24 F) Chelicerae not swollen. Femur-tibia IV unarmed. Tarsal segmentation (n = 1) 8, 16, 9, 11.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
723040C3E1B4504996E34788507261CA.taxon	diagnosis	Diagnosis. Similar to Lumieria antonionii gen. et. sp. nov. by combination of following characters: ocularium with a pair of spines, area I divided, a long pair of spines in area III, free tergites with large tubercles (Fig. 6 B) and femur-tibia IV with a retrolateral row of tubercles (Fig. 11 E, F). Differs from Lumieria antonionii gen. et. sp. nov. because the division of the area I is less conspicuous; area II not invading the area I; carapace and scutal areas with very few granules (Fig. 6 B); male femur IV with more than 20 retrolateral acuminate tubercles (Fig. 11 E, F); VP with distal margin slightly concave (Fig. 14 A, B).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
723040C3E1B4504996E34788507261CA.taxon	etymology	Derivatio nominis. The specific epithet of masculine gender, in the genitive form, dedicated to the American filmmaker, actor, musician and writer Heywood Allen (born Allan Stewart Koenigsberg in 1935), known as Woody Allen.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
723040C3E1B4504996E34788507261CA.taxon	distribution	Distribution. (Fig. 29 B) PERU. Junin. Near Comas.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
723040C3E1B4504996E34788507261CA.taxon	materials_examined	Material examined. Type material: Holotype ♂, ' PERU, Junin, Centro Turistico Ilpa, near Comas, 1 ° 42 ′ 37.1 ″ S 75 ° 04 ′ 20.2 ″ W, 27 / IV / 2011, R. Pinto-da-Rocha, A. Benedetti, J. Ochoa & D. Silva leg. (MUBI) - Paratype ♀, ' ditto' (MZSP 51851).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
CC91D7ABA56E59DEA00AF50EDEA0A505.taxon	description	Figs 5 A, 10 A, B, 14 C, D, 31	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
CC91D7ABA56E59DEA00AF50EDEA0A505.taxon	description	Description. MALE: Measurements (n = 2) DSW: 5.0 - 5.1 (5.1); DSL: 5.0 - 5.2 (5.2); CL: 1.2 - 1.3 (1.3). FIVL: 16.0 - 17.0 (17). ChL: 3.3 - 3.4 (3.4). Coloration (in ethanol): Yellow. Dorsum: (Fig. 5 A) Gamma-P-type DSS. Anterior margin of carapace with median elevation, with few granules. Carapace with granules concentrated in the lateral regions to ocularium and sparse behind ocularium. Ocularium with pronounced median depression; granulate and unarmed. Areas I-II granulate; III-IV with sparse granulation. Area I divided by longitudinal groove; with one pair of median tubercles. Areas II and IV unarmed. Area III with two long spines, directed posteriorly. Posterior margin of dorsal scutum and free tergites with few granules distributed. Lateral margins of dorsal scutum with granules along the posterior ⅔. Chelicerae: (Fig. 5 A) Swollen. Segment I granular. Segment II with few granules; finger with seven teeth. Segment III with four teeth. Pedipalpus: Femur-tibia with small tubercles on dorsal surface. Trochanter with a ventroapical setiferous tubercle. Femur a ventral row of 5 - 6 large spines; a large retroapical spine. Patella with a large retroapical and smaller proapical spines. Tibia: prolateral IIII, retrolateral IIII. Tarsus: prolateral IIi, retrolateral iIi. Venter: Coxa I with a row of 5 - 6 tubercles. Coxae II-IV with sparse granules. Rows of four tiny tubercles between coxae II-III and six tubercles between coxae III-IV. Genital area, free sternites and anal operculum few granulate. Legs: (Figs 5 A, 10 A, B) Coxae I-II each with a prolateral and a retrolateral apophysis. Coxa III unarmed. Coxa IV with few sparse granules. Trochanters I-III unarmed and smooth. Trochanter IV unarmed and granular. Femora I-III unarmed and with small granules. Femur IV granular; a distal retrodorsal small apophysis; a distal retroventral row of 11 - 12 small tubercles. Patella-tibiae I-IV unarmed, with few granules. Tarsal segmentation (n = 2) 7, 16 - 17 (17), 10, 11 - 12 (12). Penis: (Fig. 14 C, D) VP rectangular; distal margin slightly straight; straight in lateral view, with apex slightly inclined. MS C 1 - C 4 (C 5) subapical long and straight (some with curved apex); MS A 1 - A 2 (A 3) median long and straight, shorter than MS C. Lateral sacs long, with acuminate apex; with long T 3 - like microsetae. Stylus with apical region cylindrical, not swollen; with many apical small projections. Dorsal process conical, with acuminate apex, smaller than stylus. Promontory convex. - FEMALE: Measurements (n = 3) DSW: 5.2 - 5.5; DSL: 4.5 - 4.7; CL: 1.7 - 1.8. FIVL: 11.8 - 12.4. ChL: 2.4 - 2.5. Chelicerae not swollen. Femur IV unarmed. Tarsal segmentation: (n = 3) 7, 13, 10, 11.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
CC91D7ABA56E59DEA00AF50EDEA0A505.taxon	diagnosis	Diagnosis. Similar to Metasarcus kurosawai sp. nov. because ocularium unarmed, area I divided, area III with a pair of long spines (Fig. 5 A) and femur IV of males slender and without rows of tubercles (Fig. 10 A, B). It differs from M. kurosawai sp. nov. by gamma-P-type DSS; coxa III unarmed (Fig. 5 A); male femur IV with a distal retrodorsal small apophysis (Fig. 10 A, B) and penis with dorsal process and long lateral sacs (Fig. 14 C, D).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
CC91D7ABA56E59DEA00AF50EDEA0A505.taxon	etymology	Derivatio nominis. The specific epithet, a noun in apposition, in reference to Beni Department (" El Beni "), Bolivia, Department of the type locality of the species.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
CC91D7ABA56E59DEA00AF50EDEA0A505.taxon	distribution	Distribution. (Fig. 31) BOLIVIA. Beni. Southwest of Yucumo.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
CC91D7ABA56E59DEA00AF50EDEA0A505.taxon	materials_examined	Material examined. Type material: Holotype ♂, ' BOLIVIA, Beni, southwest of Yucumo | 15 ° 23 ′ S 66 ° 59 ′ W | 15 - 19 / XI / 1989, Coddington, Griswold, Silva, Larcher & Penaranda leg. (USNM) - Paratypes 1 ♂, 3 ♀, ' ditto' (USNM).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
775C1A1E890053C5A50DD784E840DDB6.taxon	description	Figs 5 B, 10 C, D, 19 A-C, 25 A, B, 31	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
775C1A1E890053C5A50DD784E840DDB6.taxon	description	Description. MALE: Measurements (n = 5) DSW: 5.2 - 6.0 (5.2); DSL: 5.5 - 6.8 (5.5); CL: 2.1 - 2.5 (2.1). FIVL: 12.5 - 13.0 (12.9). ChL: 2.0 - 2.1 (2.1). Coloration (Fig. 25 A): Chelicerae, pedipalpus and trochanters I-III yellowish green. Ocularium, anterior and lateral margin of DS, anterior part of the carapace, coxa and trochanter IV reddish. Posterior part of the carapace and areas I-IV orange, in lighter tone. Spines of area III, apophysis of coxa IV and anterior angle on area I most blackened. Granules of posterior margin of DS and free tergites I-III orange. Grooves of dorsal scutum white. Dorsum: (Fig. 5 B) Gamma-P-type DSS, with concave posterior margin of DS. Anterior margin of carapace with median elevation, covered with granules. Ocularium with median depression granular, and 4 - 5 small tubercles near the eyes. Carapace with sparse granules. Areas I-IV covered by few granules; Area I divided by a longitudinal groove and invaded by area II; with a pair of median small tubercles. Area II and IV unarmed. Area III with a pair of median spines, parallel, directed posteriorly. Lateral margins of DS granular. Posterior margin of DS and free tergites with a row of small granules. Free tergite III with a spiniform apophysis slightly longer than the length of this tergite. Chelicerae: (Fig. 5 B) Not swollen, similar to females. Segment I covered with small granules. Segment II covered with granules larger than those of segment I; finger with four teeth. Segment III with three teeth. Pedipalpus: Trochanter with a ventroapical setiferous tubercle. Femur with a ventral row of four spines and a proapical spine. Patella with a proventral small setiferous tubercle. Tibia: prolateral IiII / iIiII, retrolateral IiIIi. Tarsus: prolateral iIiIi, retrolateral iIiIi / iIiIii. Venter: Coxa I with a median row of 5 - 6 setiferous tubercles and small sparse granules. Coxae II-III with setiferous tubercles and sparse granules. Coxa IV with granules smaller than those on other coxae, sparsely distributed. Smooth genital area. Free sternites with few granules. Legs: (Figs 5 B, 10 C, D) Coxae I-II each with a prolateral and a retrolateral apophysis. Coxa III unarmed. Coxa IV with a large retrolateral apophysis, curved posteriorly and laterally. Trochanters I-IV with small granules. Trochanter IV with prolateral base expanded and a retrolateral tubercle. Femora I-III straight and unarmed, with few small granules. Femur IV straight, with few sparsely scattered small granules and a small retrobasal blunt tubercle. Tarsal segmentation: (n = 5) 9, 17 - 19 (18), 11 - 12 (12), 12 - 14 (14). Penis: (Fig. 19 A-C) VP rectangular, distal margin straight and with small lateral projections; with distal laterodorsal depressions; slightly concave in lateral view. MS C 1 - C 3 subapical long and straight; MS A 1 sub basal long and straight; MS B 1 basal short and straight. Lateral sacs long and robust, with large base and blunt apex; with long tapered T 3 - like microsetae near the base and short and flattened near the apex. Stylus long and cylindrical (apex cannot be viewed in SEM). Dorsal process absent. Promontory conspicuous and convex. - FEMALE: Measurements (n = 3) DSW: 5.0 - 5.2; DSL: 5.2 - 5.3; CL: 2.0. FIVL: 12.1 - 11.3. ChL: 2.0 - 2.2. (Fig. 25 B) Granulation denser than in male, mainly in the areas of the dorsal scutum. Spine of area III with multiple granules at base. Retrolateral apophysis in the coxa IV short, much shorter than in male. Trochanter IV without expansion of the prolateral base. Apophysis of free tergite III longer than in male. Tarsal segmentation: (n = 3) 8 - 9, 16 - 18, 10, 11 - 12.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
775C1A1E890053C5A50DD784E840DDB6.taxon	diagnosis	Diagnosis. It differs from other species of the genus by having a large retrolateral apophysis on coxa IV, curved toward externally, and a short spiniform apophysis in free tergite III (Fig. 5 B).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
775C1A1E890053C5A50DD784E840DDB6.taxon	etymology	Derivatio nominis. The specific epithet of masculine gender, in the genitive form, dedicated to the Swedish filmmaker, director, producer and writer Ernst Ingmar Bergman (1918 - 2007).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
775C1A1E890053C5A50DD784E840DDB6.taxon	distribution	Distribution. (Fig. 31) BOLIVIA. Cochabamba. Near Corani.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
775C1A1E890053C5A50DD784E840DDB6.taxon	materials_examined	Material examined. Type material: Holotype ♂, ' BOLIVIA, Cochabamba, road next to Corani, | 17 ° 11 ′ 18.9 ″ S 65 ° 53 ′ 49.2 ″ W, 01 / XII / 2010, R. Pinto-da-Rocha, A. Benedetti, J. Ochoa & A. Saravia leg. (CBF) - Paratypes 1 ♂, 1 ♀, ' ditto' (CBF); Paratypes 3 ♂, 3 ♀, ' ditto' (MZSP 36983).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
A36AB90A04FB54DCA7842D21AA4A8089.taxon	description	Figs 5 C	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
A36AB90A04FB54DCA7842D21AA4A8089.taxon	diagnosis	Diagnosis. Very similar to females of Metasarcus fellinii sp. nov. because a long spiniform apophysis in tergite III (Fig. 5 C). Differs from M. fellinii sp. nov. because longer DS (DSL greater than 5); a sparser granulation in dorsal scutum (Fig. 5 C); longer legs (FIVL greater than 11) and coxa III unarmed (Fig. 5 C).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
A36AB90A04FB54DCA7842D21AA4A8089.taxon	distribution	Distribution. BOLIVIA. Chaco.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
A36AB90A04FB54DCA7842D21AA4A8089.taxon	materials_examined	Material examined. Type material: Of M. bolivianus: Holotype ♀, ' BOLIVIA, Gran Chaco, whitout date and leg. (SMF 873). Of M. armatipalpus: Holotype ♀, ' BOLIVIA, Chaco | whitout date and leg. (SMF 1003 / 2).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
9116DAEFC376544EBEC3CF0CDEAAD952.taxon	description	Figs 5 D, 10 E, F, 19 D-F, 25 C, D, 31	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
9116DAEFC376544EBEC3CF0CDEAAD952.taxon	diagnosis	Diagnosis. Differs from other species of the genus by having femur IV swollen and with a long retrobasal apophysis; trochanter IV with long retroapical spiniform apophysis (Fig. 10 E, F).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
9116DAEFC376544EBEC3CF0CDEAAD952.taxon	distribution	Distribution. (Fig. 31) BOLIVIA. La Paz. Coroico. La Paz. Yanacachi.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
9116DAEFC376544EBEC3CF0CDEAAD952.taxon	materials_examined	Material examined. Type material: Lectotype ♂, ' BOLIVIA, La Paz. Without date and collector data. ' (SMF 1006 / SMF 1467 / 11); paralectotype ♂, ' Without date and collector data. ' (MNHN; male not examined). Additional material: 1 ♂, 2 ♀ ' BOLIVIA, La Paz, Road La Paz-Coroico, 16 ° 12 ′ 57 ″ S 67 ° 49 ′ 24.7 ″ W, 30 / XI / 2010, R. Pinto-da-Rocha, A. Benedetti, J. Ochoa & A. Saravia leg. (CBF); 2 ♂, 2 ♀, ' ditto' (MZSP 36985). 1 ♂, 1 ♀ ' BOLIVIA, La Paz, Yanacachi, 16 ° 23 ′ 54.5 ″ S 67 ° 44 ′ 11.6 ″ W, 12 / XII / 2010, R. Pinto-da-Rocha, A. Benedetti & A. Saravia leg. (CBF); 1 ♂, 2 ♀, ' ditto' (MZSP 36986).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
40CCEEBC6EA55C65B8143BC4A9673AD0.taxon	description	Figs 5 E, 10 G, H, 19 G-I, 25 E, F, 31	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
40CCEEBC6EA55C65B8143BC4A9673AD0.taxon	description	Description. MALE: Measurements (n = 5) DSW: 4.0 - 4.7 (4.7); DSL: 3.8 - 4.3 (4.3); CL: 1.5 - 1.7 (1.7). FIVL: 7.4 - 9.0 (9.0). ChL: 1.5 - 1.6 (1.6). Coloration: (Fig. 25 E) Most body brown. Apophysis of tergite III dark, almost black. Lateral margins of DS dark brown. Trochanters yellowish brown. Area III armature and granules of DS and free tergites I-III orange. Dorsum: (Fig. 5 E) Gamma-type DSS, with concave posterior margin of DS. Anterior margin of carapace with a median elevation, covered with irregular row of granules. Ocularium slightly medially depressed, unarmed, covered with granules. Carapace densely covered with granules of different sizes along its entire length. Areas I-IV densely covered with granules of different sizes. Area I divided by a longitudinal groove; with a median pair of acuminate tubercles. Areas II and IV unarmed. Area III armed with a pair of median spines. Lateral margins of DS densely granular, the granules being slightly more robust to those of carapace and areas. Posterior margin of DS and free tergites I-II with a row of granules similar to those on lateral margins of DS. Free tergite III with a large arched apophysis, apically forked, directed posteriorly, covered with small granules at base. Chelicerae: (Fig. 5 E) Slightly larger than those of females. Segment I covered with small granules. Segment II with 3 - 4 small granules; finger with three teeth. Segment III with four teeth. Pedipalpus: With granules irregularly distributed over the entire length, except tibia and tarsus. Trochanter with a ventroapical spine. Femur with a small ventroapical setiferous tubercle; a ventral row of four spines along its length and a proapical spine. Patella with a proapical spine. Tibia retrolateral IiII, prolateral iIIIi / iiIi. Tarsus: retrolateral Iii / iIiIi / iiII, prolateral II. Venter: Coxae I-IV densely covered of granules. Coxa I with a median row of 3 - 4 small setiferous tubercles. Genital area with a few sparse granules. Free sternites I-IV with a row of small tubercles. Legs: (Figs 5 E, 10 G, H) Coxae I-II each with a prolateral and a retrolateral apophysis. Coxa III with a prolateral apophysis. Coxa IV with a dorsoapical row of seven small tubercles; two small retrolateral tubercles and granules of varied sizes throughout its extension. Trochanters I-II smooth. Trochanters III-IV with a dorsoapical wide base blunt tubercle. Femora I-III straight and unarmed, with small granules. Femur IV slightly curved and unarmed, granular. Tarsal segmentation: (n = 5) 6 - 8 (8), 9 - 11 (11), 7 - 8 (8), 8 - 9 (8). Penis: (Fig. 19 G-I) VP subrectangular, with parallel sides; distal margin straight; straight in lateral view; VP with two distal lateroventral depressions. MS C 1 - C 4 apical long and straight; MS A 1 median long and straight (about half the length of the MS C); MS B 1 - B 2 sub basal short and straight (MS B 2 longer than MS B 1); MS D 1 short and straight, dorsally placed, near MS C 4. Lateral sacs short, with short T 3 - like microsetae. Stylus with projections at the apex and an armed projection directed ventrally. Dorsal process short and blunt. Promontory truncated. - FEMALE: Measurements (n = 10) DSW: 3.5 - 4.0; DSL: 3.5 - 4.1; CL: 1.2 - 1.5. FIVL: 6.6 - 7.2. ChL: 1.2 - 1.8. (Fig. 25 F) Chelicerae slightly smaller. Granules in dorsal scutum denser concentrated than in male. Apophysis of free tergite III short, spiniform. Tarsal segmentation: (n = 10) 6, 7 - 11, 7 - 9, 8 - 9	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
40CCEEBC6EA55C65B8143BC4A9673AD0.taxon	diagnosis	Diagnosis. It differs from other species of the genus because males present an extremely long apophysis in free tergite III with forked apex (Fig. 5 E). The females have short and simple apophysis, and are similar to Metasarcus bolivianus (whose male is unknown). It differs from M. bolivianus to present a greater density of granules in the dorsal scutum (Fig. 25 F) and being smaller.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
40CCEEBC6EA55C65B8143BC4A9673AD0.taxon	distribution	Distribution. (Fig. 31) BOLIVIA. Tarija. Near Entre Rios.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
40CCEEBC6EA55C65B8143BC4A9673AD0.taxon	materials_examined	Material examined. Type material: Holotype ♂, ' BOLIVIA, Tarija, near Entre Rios, Paraiso del Tordo, 21 ° 35 ′ 54.7 ″ S 64 ° 08 ′ 58.4 ″ W, 05 / XII / 2010, R. Pinto-da-Rocha, A. Benedetti, J. Ochoa & A. Saravia leg. (CBF) - Paratypes 1 ♂, 5 ♀, ' ditto' (CBF); Paratypes 3 ♂, 5 ♀, ' ditto' (MZSP 36987).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
E46DDAA163A75B47939A6252EF6D01CA.taxon	description	Figs 5 F, 10 I, J, 14 E, F, 25 G, H, 31	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
E46DDAA163A75B47939A6252EF6D01CA.taxon	description	Description. MALE: Measurements (n = 2) DSW: 4.0; DSL: 4.0 - 4.5 (4.5); CL: 1.5 FIVL: 9.2 - 10.0 (9.2). ChL: 3.3 - 3.8 (3.8). Coloration: (Fig. 25 G) Chelicerae, trochanter IV and basal portion of femur IV orange. Pedipalpus, legs I-III, remaining parts of leg IV and dorsal scutum black. Dorsum: (Fig. 5 F) Kappa- type DSS, with straight posterior margin of DS; constriction I shallow, constriction II absent, mid-bulge slightly larger than carapace and coda undefined, coalescing with mid-bulge. Sparse row of granules on the anterior margin of DS, including the median elevation. Clusters of granules concentrated on the lateral of the carapace and the posterior region to ocularium. Ocularium with depression well marked, unarmed and with granules concentrated near the eyes. Areas I-IV with a few scattered granules. Area I divided by the scutal grooves I and II; with a pair of median tubercles. Area II invading the area I; unarmed. Area III with a median pair of large spines, whose base elevation is irregularly covered by small tubercles. Area IV unarmed. Lateral margins of DS covered with granules. Posterior margin of DS and free tergites I-III with a row of granules. Chelicerae: (Fig. 5 F) Swollen. Segment I with few granules. Segment II with small setiferous granules; finger with twelve small teeth. Segment III with five teeth. Pedipalpus: Trochanter with a long ventroapical acuminate setiferous tubercle. Femur with a ventral row of six setiferous tubercles of irregular size, most long and two short. Patella with a proapical small tubercle, whose base is wider than tall. Tibia: retrolateral iIiIii, prolateral IiIi. Tarsus: retrolateral iIiIi, prolateral IiIii. Venter: Coxa I with a median row of five setiferous tubercles. Coxa II with a row of 5 - 6 setiferous tubercles in the basal portion. Coxae III-IV with small sparse granules. Genital area, free sternites and anal operculum with scattered granules. Legs: (Figs 5 F, 10 I, J) Coxae I-III each one with a prolateral and a retrolateral apophysis. Coxa IV covered with setiferous granules. Trochanters I-IV unarmed and granular. Femora-patellae I-IV unarmed and with few small granules. Tibiae I-IV unarmed, with sparse granules. Tarsal segmentation: (n = 2) 8, 15 - 17 (17), 10, 10 - 12 (12). Penis: (Fig. 14 E, F) VP rectangular, with distal margin straight; straight in lateral view. MS C 1 - C 3 (sub) apical long and straight; MS A 1 median long and straight (smaller than MS C and MS B); MS B 1 basal long and straight; MS D 1 very short, near MS C 3, placed more ventrally; MS E 1 very short, near MS C 1, placed more ventrally. Lateral sacs short and robust, with short T 3 - like microsetae. Stylus long with truncated apex and small apical projections. Dorsal process absent. Promontory truncated / straight. - FEMALE: Measurements (n = 2) DSW: 4.0; DSL: 4.2 - 4.3; CL: 1.4; FIVL: 8.6 - 9.0; ChL: 1.0 - 1.7 (Fig. 25 H). Chelicerae small, not swollen. Tarsal segmentation: (n = 2) 7, 15, 10, 11.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
E46DDAA163A75B47939A6252EF6D01CA.taxon	diagnosis	Diagnosis. Similar to Metasarcus beni sp. nov. because the ocularium is unarmed, area I divided, a pair of spines in the area III (Fig. 5 F) and male femur IV slender and without major rows of tubercles (Fig. 10 I, J). It differs from M. beni sp. nov. because kappa type DSS, coxa III with two apophyses (Fig. 5 F), femur IV without a distal retrodorsal small apophysis (Fig. 10 I, J) and by having chelicerae orange colored (Fig. 25 G).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
E46DDAA163A75B47939A6252EF6D01CA.taxon	etymology	Derivatio nominis. The specific epithet of masculine gender, in the genitive form, dedicated to the Japanese filmmaker, producer and screenwriter Akira Kurosawa (黒澤明; 1910 - 1998).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
E46DDAA163A75B47939A6252EF6D01CA.taxon	distribution	Distribution. (Fig. 31) BOLIVIA. La Paz. Zongo.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
E46DDAA163A75B47939A6252EF6D01CA.taxon	materials_examined	Material examined. Type material: Holotype ♂, ' BOLIVIA, La Paz, Zongo, 16 ° 10 ′ 32.4 ″ S 68 ° 08 ′ 11.9 ″ W, 10 / XII / 2010, R. Pinto-da-Rocha, A. Benedetti & A. Saravia leg. (CBF) - Paratype ♀, ' ditto' (CBF). Paratype ♀, ' ditto' (MZSP 76551). Paratype 1 ♂ ' BOLIVIA, La Paz, Road La Paz-Coroico, 16 ° 12 ′ 57 ″ S 67 ° 49 ′ 24.7 ″ W, 30 / XI / 2010, R. Pinto-da-Rocha, A. Benedetti, J. Ochoa & A. Saravia leg. (MZSP).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
8D8A109AA8895DA1BA59C34BA7990A55.taxon	description	Figs 5 G, 10 K, L, 14 G, H, 26 A, B, 31	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
8D8A109AA8895DA1BA59C34BA7990A55.taxon	description	Description. MALE: Measurements (n = 1) DSW: 6.1; DSL: 6.8; CL: 1.4. FIVL: 14.2. ChL: 2.4. Coloration: (Fig. 26 A) Yellow background with black spots covering virtually the entire DS; scutal area III, pedipalpus, chelicerae and legs I-III brown. Leg IV black. Dorsum: (Fig. 5 G) Gamma-P-type DSS, with strongly concave posterior margin of DS. Median elevation of anterior margin of DS with some scattered granules. Ocularium with median depression well marked with large and small granules around the eyes, and in lower concentrations in depression. Carapace with sparse granules. Areas I-IV with small-scattered granules. Area I undivided; with a pair of median tubercles. Areas II unarmed. Area III with a median pair of spines. Area IV with a pair of median tubercles. Lateral margins of DS with a denser concentration of granules. Posterior margin of DS and free tergites I-III with a row of granules. Free tergite III with a long spiniform apophysis, curved laterally and with a ventral short and sharp projection (This curvature is probably an individual anomaly. Furthermore, the first author has already had the opportunity to examine a photograph taken by Dr. Arthur Anker of a live male specimen whose apophysis was straight). Chelicerae: (Fig. 5 G) Not swollen. Segment I smooth. Segment II with granules concentrated near the fixed finger; finger with three teeth. Segment III with four teeth. Pedipalpus: Trochanter with a ventroapical setiferous tubercle. Femur with two small ventrobasal spines; a ventral row of large spines and a proapical spine. Patella with a proapical spine. Tibia retrolateral iiiIi, prolateral IiIi. Tarsus: retrolateral iIiI, prolateral IiIi / iIiIi. Venter: Coxa I with a median row of 5 - 6 small setiferous tubercles near the apex. Coxae II-IV with sparse granules. Coxa IV with the lower granules. Genital area with few granules. Free sternites with rows of larger granules. Anal operculum with granules, concentrated in the apical part. Legs: (Figs 5 G, 10 K, L) Coxa I-II each one with a retrolateral and a prolateral apophysis. Coxa III with dorsal small tubercles. Coxa IV with sparse granules; with 4 - 5 dorsoapical tubercles; a retrolateral short apophysis, with a tubercle at the base of apophysis, and a retolateral tubercle located more ventrally than apophysis. Trochanters I-III with sparse granules. Trochanter IV with a small probasal spiniform apophysis; a retrobasal and large retroapical spiniform apophyses. Femora I-III unarmed, granular. Femur IV slightly sigmoid, with a large dorsoapical tubercle; granular. Patellae I-IV unarmed. Tarsal segmentation: (n = 1) 8, 16, 10, 11. Penis: (Fig. 14 G, H) VP rectangular, with distal margin straight; straight in lateral view. MS C 1 - C 3 apical long and straight; MS A 1 median long and slightly curved; MS B 1 sub basal long and slightly curved. Lateral sacs short, apically tapered, with short T 3 - like microsetae. Stylus with inflated apex, with an elongated ventral projection and tiny projections in the apical portion. Dorsal process long. Promontory convex. - FEMALE: Measurements (n = 2) DSW: 5.6 - 5.7; DSL: 5.3 - 5.7; CL: 1.1 - 1.2. FIVL: 11.2 - 14.0. ChL: 2.0 - 2.3. (Fig. 26 B) Areas I-II with three pairs of small tubercles. Area III with a paramedian pair of small spines. Trochanter IV unarmed. Apophysis of free tergite III shorter and without ventral projection. Coloration with less brown spots, highlighting the most yellow body, resulting in a spotty pattern. Area IV with brown pigmentation more obvious. Tarsal segmentation: (n = 2) 7 - 8, 13 - 15, 9 - 11, 7 - 10.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
8D8A109AA8895DA1BA59C34BA7990A55.taxon	diagnosis	Diagnosis. It differs from other species of the genus because by having area I undivided; long spiniform apophysis on free tergite III (Fig. 5 G); trochanter IV with a probasal, a retrobasal and a retroapical apophyses (Fig. 10 K, L); DS with yellow background with black spots (Fig. 26 A).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
8D8A109AA8895DA1BA59C34BA7990A55.taxon	etymology	Derivatio nominis. The specific epithet of masculine gender, in the genitive form, dedicated Miguel Limachi, from Coleccion Boliviana de Fauna (CBF), who provided invaluable help with all formalities for collecting in Bolivia.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
8D8A109AA8895DA1BA59C34BA7990A55.taxon	distribution	Distribution. (Fig. 31) BOLIVIA. La Paz. La Paz-Coroico; Yanacachi.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
8D8A109AA8895DA1BA59C34BA7990A55.taxon	materials_examined	Material examined. Type material: Holotype ♂, ' BOLIVIA, La Paz, Yanacachi, 16 ° 23 ′ 54.5 ″ S 67 ° 44 ′ 11.6 ″ W, 12 / XII / 2010, R. Pinto-da-Rocha, A. Benedetti, J. Ochoa & A. Saravia leg. (CBF) - Paratype ♀, ' ditto' (MZSP 36991). Paratype ♀, ' BOLIVIA, La Paz, road La Paz-Coroico, 16 ° 13 ′ 45 ″ S 67 ° 49 ′ 17.1 ″ W, 30 / XI / 2010, R. Pinto-da-Rocha, A. Benedetti, J. Ochoa & A. Saravia leg. (MZSP 36992).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
F527066760BD5643AE59B52817A99A28.taxon	description	Figs 5, 10, 14 C-H, 19, 20, 25, 26, 31	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
F527066760BD5643AE59B52817A99A28.taxon	diagnosis	Diagnosis. Metasarcus can be diagnosed from other Metasarcidae genera with long legs (except Ayacucho) by the combination of the following features: gamma-type; gamma-P-type or kappa-type DSS; ocularium unarmed; area III with two spines. Half of Metasarcus species present a spine, long projection or trifid projection in free tergite III, present in no other Metasarcidae species; coxa III with two apophyses; coxa IV reaching posterior margin of dorsal scutum or surpassing it; femur IV smooth or slightly granulate; penis stylus and VP thin thickness, with less than 10 MS C	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
F527066760BD5643AE59B52817A99A28.taxon	distribution	Distribution. (Fig. 31) BOLIVIA. Beni, Cochabamba, La Paz and Tarija.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
AACBD9DCD9F357B9897853CF2951C824.taxon	description	Figs 5 H, 10 M, N, 20 A-C, 26 C, D, 31	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
AACBD9DCD9F357B9897853CF2951C824.taxon	description	Description. MALE: Measurements (n = 5) DSW: 6.5 - 7.7 (7.6); DSL: 6.4 - 7.0 (7.0); CL: 2.3 - 2.8 (2.6). FIVL: 10 - 12 (11.5). ChL: 2.5 - 3.0 (3.0). Coloration: (Fig. 26 C) Red, with elongated spots of dark pigmentation throughout the carapace, DS and coxa IV. Posterior margin of DS, free tergites and coxa IV apex black. Apophysis of the free tergite III black. Chelicerae, pedipalpus and legs (except the coxa IV) black. Arthrodial membrane between the posterior margin of DS and free tergites I-III green. Dorsum: (Fig. 5 H) Gamma-P-type DSS, with strongly concave posterior margin of DS. Anterior margin of the carapace with median elevation partially covered with sparse granules. Ocularium unarmed, with few small granules in the depression and a higher concentration of larger granules close to the eyes. Carapace with granules sparsely distributed throughout its length. DS with slightly conspicuous grooves, not being possible to notice the existence of a fourth area in some specimens. Area I undivided or with an extremely shallow longitudinal groove, practically inconspicuous. Areas I-III with small sparse granules. Area III with a median pair of setiferous tubercles. Lateral margins of DS with granules distributed throughout their length. Posterior margin of DS and free tergites I-II with rows of small granules. Free tergite III with a large trifurcated and acuminate apophysis, robust, smooth, with a broad base, median branch larger than lateral ones. Chelicerae: (Fig. 5 H) Not swollen. Segment I covered by small granules. Finger of segment II with two teeth. Segment III with six teeth. Pedipalpus: Trochanter with a ventrodistal setiferous tubercle. Femur with a ventrobasal row of three small spines; a retroventral row of three large spines; and a prodistal spine. Patella with a prodistal spine. Tibia: prolateral (i) iiIi, retrolateral IiIi. Tarsus: prolateral iiIii / iiIiii / iIiIi / iIiI, retrolateral iIiIi / iIiiIi. Venter: Coxae I-IV with small sparse granules. Coxa I with a median row of 5 - 7 small setiferous tubercles. Smooth genital area. Free sternites with few granules. Legs: (Figs 5 H, 10 M, N) Coxae I-II each one with a prolateral and a retrolateral apophysis. Coxa III with a prolateral apophysis. Coxa IV with small sparse granules. Trochanters I-IV smooth. Femora I-II straight. Femur III slightly curved at the apex and base. Femur IV slightly sigmoid; with few granules and a retrodistal spiniform apophysis. Patella-tibiae I-IV granular. Tarsal formula: (n = 5) 7 - 8 (8) / 13 - 16 (16) / 9 - 10 (9) / 10 - 12 (10). Penis: (Fig. 20 A-C) VP subrectangular, with diverging lateral margins at the apex, resulting in a narrower base than the apex; distal margin straight; straight in lateral view. MS C 1 - C 2 subdistal long and curved; MS A 1 long and straight; medially placed; MS A 2 sub basal long and straight (longer than MS A 1 and MS C); MS B 1 basal very short (near lateral sacs); MS D 1 long and straight (half the length of MS C), dorsally placed, near MS C 2. Lateral sacs long and apically blunt; with short T 3 - like microsetae. Stylus with a serrated inflated apex and a ventral projection. Dorsal process rounded, bag-shaped. Promontory convex. - FEMALE: Measurements (n = 6) DSW: 5.5 - 6.0; DSL: 5.5 - 6.5; CL: 1.8 - 2.6. FIVL: 9.0 - 9.5. ChL: 2.0 - 2.6. (Fig. 26 D) Denser granulation in ocular depression and in the anterior part of the carapace. Femora III-IV straight. Femur IV without retrodistal apophysis. Apophysis of the free tergite III shorter and unbranched, smaller and less wide than the males. Tarsal segmentation: (n = 6) 7 - 8, 12 - 14, 9 - 10, 10 - 11.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
AACBD9DCD9F357B9897853CF2951C824.taxon	diagnosis	Diagnosis. It differs from other species of the genus by a trifurcated and acuminate apophysis in free tergite III (Fig. 5 H); male femur IV with a retrodistal spiniform apophysis (Fig. 10 M, N); scutal grooves almost inconspicuous (Fig. 5 H); red (Fig. 26 C).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
AACBD9DCD9F357B9897853CF2951C824.taxon	etymology	Derivatio nominis. The specific epithet, an adjective in nominative singular, formed by Latin prefix tri - + Latin spīnōsus, a, um (thorny), in reference to the trifurcated apophysis of free tergite III.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
AACBD9DCD9F357B9897853CF2951C824.taxon	distribution	Distribution. (Fig. 31) BOLIVIA. Cochabamba. Road to Corani.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
AACBD9DCD9F357B9897853CF2951C824.taxon	materials_examined	Material examined. Type material: Holotype ♂, ' BOLIVIA, Cochabamba, road to Corani, 17 ° 11 ′ 18.9 ″ S 65 ° 53 ′ 49.2 ″ W, 01 / XII / 2010, R. Pinto-da-Rocha, A. Benedetti, J. Ochoa & A. Saravia leg. (CBF) - Paratypes 1 ♂, 3 ♀, ' ditto' (CBF); Paratypes 3 ♂, 3 ♀, ' ditto' (MZSP 36997).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
77ECF8DB232D54A889CC709E7E0F7A37.taxon	description	Figs 5 I, 10 O, P, 20 D-F, 26 E, F, 31	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
77ECF8DB232D54A889CC709E7E0F7A37.taxon	description	Description. MALE: Measurements (n = 4) DSW: 4.2 - 4.9 (4.7); DSL: 5.1 - 5.4 (5.4); CL: 2.1 - 2.5 (2.5). FIVL: 13.0 - 14.2 (14.2). ChL: 2.9 - 4.2 (3.6). Coloration: (Fig. 26 E) Orange brown. Brown spots on the carapace and on the lateral margins of DS. Free tergites I-III and anal operculum dark brown. Reddish brown legs, except for the coxae and trochanters, in a lighter shade of orange brown. Dorsum: (Fig. 5 I) Kappa-type DSS, with straight posterior margin of DS; constriction II present, coda long. Anterior margin of the carapace with median elevation, with few granules. Ocularium with very subtle median depression, almost inconspicuous. Carapace with sparse granules. DS with four areas covered by few granules, unarmed. Area I undivided. Lateral margins of DS with rows of granules in greater density than in the dorsal areas of the DS and carapace. Posterior margin of DS and free tergites I-III unarmed and with few granules. Chelicerae: (Fig. 5 I) Swollen in large males (as in the holotype), similar to females in the small males. Segment I with sparse small granules. Segment II smooth, swollen in some males, finger with one tooth. Segment III with two teeth. Pedipalpus: Sparse granules throughout the appendage, less numerous on the ventral surfaces. Coxa with a small retrobasal tubercle. Trochanter with two ventroapical setiferous tubercles, the apical one being smaller. Femur with a row of five ventral setiferous tubercles, two more basal and three median and a proapical spine. Patella smooth. Tibia: prolateral iIiIIi, retrolateral IIii. Tarsus: prolateral iiIiIii, retrolateral iIiIi. Venter: Coxa I with a median row of five setiferous tubercles, a parallel row of small tubercles and scattered granules. Coxa II covered with sparse granules; with an apical row of two-three setiferous tubercles. Coxa III with a proventral row of five tubercles and a retroventral row of eight. Coxa IV with sparse granules. Smooth genital area. Free sternites and anal operculum with few granules. Legs: (Figs 5 I, 10 O, P) Coxa I with a retrolateral apophysis. Coxa II with one prolateral and two retrolateral apophyses. Coxa III unarmed. Coxa IV with sparse small granules. Trochanters with sparse granules. Femora I-IV straight, unarmed and with granules. Tarsal formula: (n = 4) 8, 17 - 18 (18), 10 - 11 (10), 12 - 13 (13). Penis: (Fig. 20 D-F) VP subrectangular, long, with apex narrower than the base; convex distal margin; curved in lateral view. MS C 1 - C 3 subdistal long and straight (or slightly curved); MS A 1 sub basal long and straight (slightly shorter than MS C); MS B 1 basal (near lateral sacs) long and straight (or apically curved); MS D 1 short and straight, medially placed; MS E 1 - E 2 very short, ventrally placed (MS E 1 between MS C 2 and MS C 3; MS E 2 near MS D 1). Lateral sacs long and tapered; with long T 3 - like microsetae. Stylus with wide apex and projections. Dorsal process long and apically tapered. Promontory convex. - FEMALE: Measurements (n = 5) DSW: 4.1 - 4.5; DSL: 5.0 - 5.7; CL: 2.1 - 2.3. FIVL: 12.7 - 14.6. ChL: 2.3 - 2.6. (Fig. 26 F) Chelicerae smaller than in males. Tarsal segmentation: (n = 5) 7 - 8, 14 - 16, 10 - 11, 11 - 13.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
77ECF8DB232D54A889CC709E7E0F7A37.taxon	diagnosis	Diagnosis. It differs from other species of the genus by the following set of characteristics: alpha type DSS; scutal area I undivided; areas I-IV unarmed (Fig. 5 I); male femur IV unarmed (Fig. 10 O, P); body orange brown (Fig. 26 E); VP subrectangular and long; dorsal process present (Fig. 20 D-F).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
77ECF8DB232D54A889CC709E7E0F7A37.taxon	etymology	Derivatio nominis. The specific epithet of feminine gender, in the genitive form, in honor to Maria Rene Vacaflores, from Coleccion Boliviana de Fauna (CBF), who provided invaluable help with all formalities for collecting in Bolivia.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
77ECF8DB232D54A889CC709E7E0F7A37.taxon	distribution	Distribution. (Fig. 31) BOLIVIA. La Paz. Zongo.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
77ECF8DB232D54A889CC709E7E0F7A37.taxon	materials_examined	Material examined. Type material: Holotype ♂, ' BOLIVIA, La Paz, Zongo, 16 ° 10 ′ 32.4 ″ S 68 ° 08 ′ 11.9 ″ W, 10 / XII / 2010, R. Pinto-da-Rocha, A. Benedetti, J. Ochoa & A. Saravia leg. (CBF) - Paratypes 2 ♂, 5 ♀, ' ditto' (CBF); Paratypes 4 ♂, 6 ♀, ' ditto' (MZSP 36998).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
DA7962F9568D543EBFD28BA3F07254F1.taxon	diagnosis	Diagnosis. Metasarcidae can be easily diagnosed by other Gonyleptoidea by only one feature, the penis with lateral finger-like sacs. Only one genus (Metalibitia, Cosmetidae) of Gonyleptoidea possess lateral sacs on ventral plate but its shape and position is different from it and not homologous. It differs from Stygnidae by having ocularium undivided; by Gonyleptidae by pedipalpal femur with long spines; by Cosmetidae by pedipalpus somewhat cylindrical and with spines; by Agoristenidae by having tarsal process; by Cranaidae by pedipalpal femur smooth or small-tuberculate.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
DA7962F9568D543EBFD28BA3F07254F1.taxon	distribution	Distribution. The family Metasarcidae occurs in Andean Mountains of Bolivia and Peru, the southern limit being the border with Argentina and the northern limit the Huancabamba depression, situated in northern Peru (Figs 28 - 31). Most species are found in moderate to high altitudes (circa 4,000 m above sea level). The only exception to is Metasarcus beni sp. nov. (about 170 m above sea level) which occurs in the mountain foothills. A second species, the type-species of the genus Metasarcus, M. bolivianus, has been attributed to the Bolivian lowland region, the Chaco Province, without mention of a more precise locality. Most species (28 spp.) are known only from their type-locality, and those known from a few records of distribution (6 spp.) are endemic to small areas, where the maximum distance between two records is 150 km. A few localities possess sympatric species, such as: Parque Nacional Yanachaga-Chemillen / Peru (T. chaplini sp. nov., T. joseochoai sp. nov., A. pasolinii sp. nov., T. scorsesei sp. nov.), Centro Turistico Ilpa / Peru (L. woodyalleni gen. et sp. nov., L. antonionii gen. et sp. nov.), Zongo / Bolivia (M. kurosawai sp. nov., M. vacafloresae sp. nov.), Cutervo / Peru (A. uniseriatus comb. nov., H. kubricki gen. et sp. nov., A. spielbergi sp. nov.). Ayacucho titschacki, which occurs in the Peruvian Central Andes, (near to Ocollo, Virgem de Cacharras de Cocha) and Ayacucho tapacocha nom. nov., which occurs in northern Peru, are the species with the largest distributions recorded in the Ayacucho and Ancash areas, respectively. This high level of endemism is comparable to the eastern coast of South America, where most species occupy small areas of endemism (see Da-Silva et al. 2017). However, the harvestmen fauna from Peru and Bolivia is poorly sampled, which prevents a more detailed comparison with other regions. All Bolivian species of Metasarcidae belong to the type genus, Metasarcus, and occur in the eastern Andes from La Paz to Tarija Province (M. fellinii sp. nov. is the southernmost species of the family), the Altiplano being the northern distribution limit. Its sister genus, Incasarcus is present only in the Peruvian Cusco Department, in Montane tall grass vegetation (Puna) and scrub and montane Rain Forest. Both genera are separated by Puna Seca and Titicaca lake, which means the Altiplano. Ayacucho is widespread in most of the Peruvian Andean region, from Cajamarca to Ayacucho departments, the Rio Apurimac being the southern limit of its distribution. The only metasarcid species recorded from the western Andean foothills is A. roeweri nom. nov., from Rio Fortaleza (2700 m above sea level, Ancash, Cajacay, Peru), where the riparian forest El Bosque de Fortaleza is found. Most species can be found in two types of vegetation, the Mountain short grass and Andean wastes (Quechua) and Mountain tall grass and scrub (Puna). One species, A. pasolinii sp. nov. was recorded from the Mountain Rain Forest (Parque Nacional Yanachaga-Chemillen, Oxapampa, Peru). The monotypic genus Huancabamba gen. nov. is recorded only in Cutervo (Cajamarca Department), in Mountain Rain Forest. Lumieria gen. nov. has only two species, sympatrically distributed at Centro Turistico Ilpa (Junin Department - Bolivia). This locality is covered by Mountain tall grass and scrub. Tschaidicancha has four species recorded in only three regions, two of which are very close to each other. T. scorsesei sp. nov., T. joseochoai sp. nov., and T. chaplini sp. nov. occur in Mountain Rain Forest, and T. weyrauchi in areas with scrubs of Mountain Tall Grass and Scrub.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
64BC536009F75377927848379F23E6AF.taxon	description	Figs 6 C, 11 G, H, 21 A-C, 27 C, D, 29 A	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
64BC536009F75377927848379F23E6AF.taxon	description	Description. MALE: Measurements (n = 5) DSW: 4.4 - 4.7 (4.6); DSL: 4.6 - 5.5 (5.5); CL: 1.9 - 2.4 (2.2). FIVL: 14.1 - 14.7 (14.7). ChL: 2.9 - 4.5 (4.5). Coloration: (Figs 27 C) Reddish brown. Chelicerae, pedipalpus and trochanters I-IV pale brown. White spots on the lateral margins of the DS. Dorsum: (Fig. 6 C) Kappa-type DSS. Anterior margin of carapace with median elevation, with granules sparsely distributed. Ocularium with one pair of parallel spines. Carapace with few sparse granules. Areas I-IV with sparse granules (lower density in area III). Area I undivided longitudinally. Area III with two parallel spines, directed posteriorly, located at elevations of integument, totally granulate. Posterior margin of DS with a row of few granules. Lateral margins of DS with sparse granules. Free tergites I-II with a row of granules. Free tergite III with two median spiniform tubercles, longer than the length of this tergite. Chelicerae: (Fig. 6 C) Swollen. Segment I with three granules. Segment II with few granules; finger with two teeth. Segment III with three teeth. Pedipalpus: With very few granules on the dorsal surface of the femur, tibia and patella. Trochanter with a ventroapical setiferous tubercle. Femur with a ventral row of eight setiferous tubercles, larger in the median portion, distributed throughout the length of femur except the apex; one large proapical spine. Patella with a proapical tubercle. Tibia: retrolateral iiIi, prolateral IiIi. Tarsus: prolateral iIiIi, retrolateral iIiIi. Venter: Coxae I-II with a median row of 4 - 6 tubercles. Coxae III-IV with granules and setae scattered. Genital area slightly granulate. Free sternites with few granules. Anal operculum few granulate. Legs: (Figs 6 C, 11 G, H) Coxae I-II each one with a prolateral and a retrolateral apophysis. Coxa III with a prolateral apophysis. Coxa IV with sparse setiferous granules. Trochanters I-IV granulate and unarmed. Femora I-III unarmed, with small granules. Femur IV with granules sparsely distributed in higher density on the ventral side; a row of retrolateral small 36 - 39 acuminate tubercles. Patellae I-III with few granules. Patella IV with three tiny retroapical tubercles. Tibiae I-III unarmed and few granulate. Tibia IV with a retrolateral row of 16 - 19 small acuminate tubercles along the basal 1 / 2. Tarsal segmentation: (n = 3) 8 - 9 (9), 18, 10 - 12 (12), 10 - 14 (14). Penis: (Fig. 21 A-C) VP subrectangular, long; distal margin slightly convex, with distal-lateral projections; slightly curved in lateral view. MS C 1 - C 3 (4) subdistal long and slightly curved; MS A 1 long and straight (slightly shorter than MS C and MS B), medially placed; MS B 1 sub basal long and straight; MS D 1 short and straight, placed near MS C 3. Lateral sacs long and apically blunt; with long T 3 - like microsetae. Stylus laterally flattened, apex slightly inflated, with several apical projections. Dorsal process laterally flattened, with slender apex. Promontory convex. - FEMALE: Measurements (n = 3) DSW: 4.5 - 4.7; DSL: 4.7 - 5.3; CL: 1.8 - 2.0. FIVL: 13.8 - 14.2. ChL: 1.5 - 1.7. (Fig. 27 D) Chelicerae smaller than in smaller males. Femur IV unarmed. Tarsal segmentation: (n = 3) 8, 16 - 17, 11 - 12, 12 - 14.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
64BC536009F75377927848379F23E6AF.taxon	diagnosis	Diagnosis. Similar to Tschaidicancha scorsesei sp. nov. by the set of the following characteristics: reddish-brown coloration (Fig. 27 C); femur IV with a retrolateral row with more than 30 small tubercles; tibia IV with a retrolateral row with more than ten small tubercles (Fig. 11 G, H). Differs from T. scorsesei sp. nov. by the set of the following characteristics: the area I undivided longitudinally; free tergite III with a pair of spiniform tubercles (Fig. 6 C); tibia IV with more than 15 retrolateral tubercles (Fig. 11 G, H); VP with distal-lateral projections (Fig. 21 A-C).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
64BC536009F75377927848379F23E6AF.taxon	etymology	Derivatio nominis. The specific epithet of masculine gender, in the genitive form, dedicated to the English actor, composer, director and producer Sir Charles Spencer Chaplin (1889 - 1977), a worldwide icon in the era of silent film through his screen persona " The Tramp ".	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
64BC536009F75377927848379F23E6AF.taxon	distribution	Distribution. (Fig. 29 A) PERU. Pasco. Oxapampa. Parque Nacional Yanachaga-Chemillen.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
64BC536009F75377927848379F23E6AF.taxon	materials_examined	Material examined. Type material: Holotype ♂, ' PERU, Pasco, Oxapampa, Parque Nacional Yanachaga-Chemillen, 10 ° 32 ′ 42.1 ″ S 75 ° 21 ′ 24.4 ″ W, 22 / IV / 2011, R. Pinto-da-Rocha, A. Benedetti, J. Ochoa & D. Silva leg. (MUBI) - Paratype 1 ♀, ' ditto' (MUBI); Paratypes 1 ♂, 1 ♀, ' ditto' (MUSM); Paratypes 3 ♂, 3 ♀, ' ditto' (MZSP 36984).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
A1610A2DB89353C2957F92D8AB247D0F.taxon	description	Figs 6 D, 11 I, J, 14 I, J, 27 A, B, 29 B	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
A1610A2DB89353C2957F92D8AB247D0F.taxon	description	Description. MALE: Measurements (n = 1) DSW: 4,4; DSL: 4,2; CL: 1,7; FIVL: 7,8; ChL: 3,0. Coloration: (Fig. 27 A) Predominantly orange brown, with two white spots on the carapace, laterally posterior to the ocularium. Dark brown legs. Chelicerae orange brown (in a shade darker than body). Tubercles of the DS areas, free tergites and lateral margins whitish. Dorsum: (Fig. 6 D) Gamma-P-type DSS, with concave posterior margin of DS. Anterior margin of the carapace with medium elevation, practically smooth, with few sparse granules. Ocularium with mild median depression; a pair of acuminate spines. Carapace with few granules, distributed mainly in the lateral region. Areas I-IV without granules, except elevation on area III. Area I undivided longitudinally; with a median pair of small tubercles. Area II with a median pair of tubercles and a right lateral tubercle. Area III with two spines, directed posteriorly located in elevations of the integument, very granular. Area IV with a median pair of tubercles. Posterior margin of DS smooth; with a left lateral tubercle. Free tergites I-III smooth. Free tergite I unarmed. Free tergite II with a pair of lateral tubercles. Free tergite III with three tubercles. Lateral margins of DS smooth; seven-nine tubercles in the posterior half, close to areas II-IV and posterior margin of DS. Chelicerae: (Fig. 6 D) Swollen. Segments I-II with few granules. Segment II with finger with one tooth. Segment III with 5 - 6 tiny teeth. Pedipalpus: With sparse granules dorsally in the femur. Trochanter with a ventroapical setiferous tubercle. Femur with a ventrobasal tubercle; a ventral row of four setiferous tubercles, the basal-most being smallest and the remaining three of equal size, distributed throughout the article, except at the apex; a proapical spine. Patella with a proapical tubercle. Tibia: prolateral iiIii, retrolateral IiIi. Tarsus: prolateral iIiii, retrolateral IiIi. Venter: Coxae I-II with rows of four and six tubercles, respectively, those of the coxa I larger. Coxae III-IV with granules. Genital area with few granules. Free sternites with row of granules. Anal operculum granular. Legs: (Figs 6 D, 11 I, J) Coxae I-II each one with a prolateral and a retrolateral apophysis. Coxa III with a prolateral apophysis. Coxa IV with 7 - 10 tubercles, distributed on the prodorsal side. Trochanters I-IV unarmed and with few granules. Femora I-III unarmed and with small granules. Femur IV with dense dorsal granulation; a prolateral row of 29 acuminate tubercles along apcal ⅔; a retrolateral row of 34 acuminate tubercles; a prodorsal row of five acuminate tubercles and three blunt tubercles along basal ⅓, smaller than the other tubercles present in the article; a ventral row of granules. Patellae I-III unarmed, with few granules. Patella IV granular; with three dorsoapical spiniform tubercles. Tibiae I-IV unarmed and densely granular. Tarsal segmentation: (n = 1) 9 / 16 / 14 / 17. Penis: (Fig. 14 I, J) VP subrectangular, long; distal margin slightly convex, with distal-lateral projections; straight in lateral view. MS C 1 - C 3 (4) subdistal long (MS C 1 - C 2 shorter than the others) and slightly curved; MS A 1 short and straight, medially placed; MS B 1 sub basal long and straight. Lateral sacs long and apically blunt; with long T 3 - like microsetae. Stylus expanded in the shape of a wedge and flattened laterally, with serrated edges; with large projection in the dorsal direction and with several tiny apical projections. Dorsal process subrectangular. - FEMALE: Measurements (n = 1) DSW: 4.2; DSL: 4.1; CL: 1.5. FIVL: 8.7. ChL: 1.7. (Fig. 27 B) Chelicerae smaller than the male. Femur IV unarmed. Patella IV with acuminate tubercles as in the male, but less robust and smaller. Tarsal segmentation: (n = 1) 8, 15, 14, 17.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
A1610A2DB89353C2957F92D8AB247D0F.taxon	diagnosis	Diagnosis. It differs from other species of the genus by the following set of characteristics: gamma-P-type DSS; ocularium with a pair of acuminate tubercles; DS and scutal areas smooth; scutal area I undivided; free tergites I-III unarmed (Fig. 6 D); male femur IV with a prolateral row of 29 acuminate tubercles and a retrolateral row of 34 acuminate tubercles (Fig. 11 I, J); body predominantly orange-brown with two white spots on the carapace; tubercles of the DS areas, free tergites and lateral margins whitish (Fig. 27 A).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
A1610A2DB89353C2957F92D8AB247D0F.taxon	etymology	Derivatio nominis. The specific epithet of masculine gender, in the genitive form, dedicated Dr. Jose Ochoa C. (MUBI), for his great help in fieldwork in Bolivia and Peru.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
A1610A2DB89353C2957F92D8AB247D0F.taxon	distribution	Distribution. (Fig. 29 B) PERU. Pasco. Oxapampa. Parque Nacional Yanachaga-Chemillen.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
A1610A2DB89353C2957F92D8AB247D0F.taxon	materials_examined	Material examined. Holotype ♂, ' PERU, Pasco, Oxapampa, Parque Nacional Yanachaga-Chemillen 10 ° 32 ′ 42.1 ″ S 75 ° 21 ′ 24.4 ″ W, 22 / IV / 2011, R. Pinto-da-Rocha, A. Benedetti, J. Ochoa & D. Silva leg. (MUBI) - Paratype 1 ♀, ' ditto' (MZSP 36988). Paratype 1 ♂, ' PERU, Pasco, Oxapampa, Parque Nacional Yanachaga-Chemillen 10 ° 32 ′ 42.5 ″ S 75 ° 21 ′ 29.8 ″ W, 9 - 11 / IV / 2011, S. Cardonel & J. Grados leg. (MUSM); Paratypes 3 ♂, 3 ♀, ' ditto' (MZSP 36984).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
EDF422241D4A5171AF58CC04A22D7425.taxon	description	Figs 6 C-F, 11 G-N, 14 I-L, 21, 27, 29 A, B	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
EDF422241D4A5171AF58CC04A22D7425.taxon	diagnosis	Diagnosis. Tschaidicancha can be distinguished from all other Metasarcidae genera by the combination of following: Kappa-type or gamma-P-type DSS; ocularium low, with two pairs of high spines; area III with two pairs of high spines; long leg IV (femur IV length / DS length> 1.6); male coxa IV apex reaching posterior margin; penis VP subrectangular, with distal-lateral projections (in most species), thin thickness; with less than five MS C; stylus laterally flattened, with apex inflated; dorsal process present.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
EDF422241D4A5171AF58CC04A22D7425.taxon	distribution	Distribution. (Fig. 29) PERU. Huanuco and Pasco.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
942B04D6850E510C815449E7BE9AA7EA.taxon	description	Figs 6 E, 11 K, L, 21 D-F, 29 A	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
942B04D6850E510C815449E7BE9AA7EA.taxon	description	Description. MALE: Measurements (n = 10) DSW: 4.1 - 5.5 (5.5); DSL: 4.6 - 5.6 (5.6); CL: 1.6 - 2.2 (2.2). FIVL: 11.2 - 15.0 (12.4). ChL: 2.8 - 5.2 (5.2). Coloration: Brownish red DS. Chelicerae, pedipalpus and legs brown. Dorsum: (Fig. 6 E) Kappa-type DSS. Anterior margin of the carapace with median elevation; with sparse granules. Ocularium with pronounced median depression; with a pair of parallel spines. Carapace with few sparse granules. Areas I-IV with few sparse granules. Area I divided; with a medium pair of acuminate tubercles. Area III with a pair of spines, directed posteriorly, located in tegument elevations, very granular. Area IV short, with a pair of medium tubercles (smaller than those on Area I). Posterior margin of DS and free tergites I-III with a row of granules. Lateral margins of DS smooth. Chelicerae: (Fig. 6 E) Swollen on large males (as in the holotype). Slightly larger than female chelicerae in smaller males. Segment I slightly granular. Segment II with few granules; with five teeth. Segment III with two teeth. Pedipalpus: Trochanter with a ventroapical setiferous tubercle. Femur with a ventral row of 8 - 9 large tubercles (the middle ones largest) along the basal ⅔; one large prosubapical spine. Patella with a proapical tubercle. Tibia: retrolateral iIiIi, prolateral IiIi. Tarsus: retrolateral iiIiIi, prolateral iIIIi. Venter: Coxa I with a row of 4 - 5 tubercles. Coxae II-IV with granules throughout their surface. Genital area with few granules. Free sternites smooth. Anal operculum with few granules. Legs: (Figs 6 E, 11 K, L) Coxae I-II each one with an anterior and a posterior apophysis. Coxa III unarmed. Coxa IV with sparse setiferous granules. Trochanters I-III unarmed and slightly granular. Trochanter IV with few granules and a retroapical small tubercle. Femora I-III unarmed and with small granules. Femur IV with sparse granules; a retrolateral row of 33 - 35 small acuminate tubercles, except at the apex. Patellae I-III unarmed, with few granules. Patella IV with three retroapical small tubercles. Tibiae I-III unarmed and with few granules. Tibia IV with a retrolateral row of 11 - 12 small acuminate tubercles along the basal 1 / 2. Tarsal segmentation: (n = 10) 9 - 10 (10), 16 - 23 (23), 11 - 13 (12), 13 - 14 (14). Penis: (Fig. 21 D-F) VP subrectangular, long; distal margin straight; slightly curved in lateral view. MS C 1 - C 2 (3) subdistal long and slightly curved; MS A 1 long and straight (slightly shorter than MS C and MS B), medially placed; MS B 1 sub basal long and straight; MS D 1 short and straight, placed below MS C. Lateral sacs long and apically blunt; with long T 3 - like microsetae. Stylus with several apical projections. Dorsal process present. Promontory convex. - FEMALE: Measurements (n = 5) DSW: 3.8 - 5.0; DSL: 4.3 - 5.6; CL: 1.5 - 2.2. FIVL: 10.5 - 15.3. ChL: 2.0 - 2.7. Chelicerae slightly smaller than that of small males and not swollen as in large males. Femur IV unarmed. Tarsal segmentation: (n = 5) 7 - 9 / 15 - 17 / 11 - 14 / 12 - 14.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
942B04D6850E510C815449E7BE9AA7EA.taxon	diagnosis	Diagnosis. Similar to Tschaidicancha chaplini sp. nov. by the set of the following characteristics: reddish brown coloration; femur IV with a retrolateral row with more than 30 small tubercles; tibia IV with a retrolateral row with more than ten small tubercles (Fig. 11 K, L). Differs from T. chaplini sp. nov. by the set of the following characteristics: the area I divided longitudinally; free tergite III unarmed (Fig. 6 E); tibia IV with less than 15 retrolateral tubercles (Fig. 11 K, L); VP without distal-lateral projections (Fig. 21 D-F).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
942B04D6850E510C815449E7BE9AA7EA.taxon	etymology	Derivatio nominis. The specific epithet of masculine gender, in the genitive form, dedicated to the American director, producer, screenwriter and cinema historian Martin Charles Scorsese (1942 -), an exponent of New Hollywood (American New Wave).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
942B04D6850E510C815449E7BE9AA7EA.taxon	distribution	Distribution. (Fig. 29 A) PERU. Pasco. Oxapampa. Parque Nacional Yanachaga-Chemillen.	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
942B04D6850E510C815449E7BE9AA7EA.taxon	materials_examined	Material examined. Type material: Holotype ♂, ' PERU, Pasco, Oxapampa, Parque Nacional Yanachaga-Chemillen, 10 ° 32 ′ 42.1 ″ S 75 ° 21 ′ 24.4 ″ W, 22 / IV / 2011, R. Pinto-da-Rocha, A. Benedetti, J. Ochoa & D. Silva leg. (MUBI) - Paratypes 1 ♂, 1 ♀, ' ditto' (MZSP 36994). Additional material: 16 ♂, 4 ♀, ' PERU, Pasco, Oxapampa, Parque Nacional Yanachaga-Chemillen 10 ° 32 ′ 42.1 ″ S 75 ° 21 ′ 24.4 ″ W | 9 - 11 / IV / 2011, S. Cardonel & J. Grados leg. (MUSM 0500505).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
2C03425E591E58E09B054C64830F1E0A.taxon	description	Figs 6 F, 11 M, N, 14 K, L, 27 E, F, 29 A	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
2C03425E591E58E09B054C64830F1E0A.taxon	diagnosis	Diagnosis. It differs from other species of the genus by the following set of characteristics: kappa-type DSS; ocularium with a pair of long spines; scutal area I undivided; free tergite III with a pair of large spiniform tubercles (Fig. 6 F); male femur IV with a dorsoapical row of acuminate tubercles; male patella IV with three long acuminate tubercles (Fig. 11 M, N); chelicerae orange and body predominantly dark reddish brown (Fig. 27 E). VP with long lateroapical projections (Fig. 14 K, L).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
2C03425E591E58E09B054C64830F1E0A.taxon	distribution	Distribution. (Fig. 29 A) PERU. Huanuco. Tschaidicancha (near Huanuco); Bosque Carpish (SW Tingo Maria).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
2C03425E591E58E09B054C64830F1E0A.taxon	materials_examined	Material examined. Type material: Holotype ♂, ' PERU, Huanuco, Tschaidicancha, near Huanuco, 2,800 m a. s. l., 2 / VIII / 1955, Weyrauch leg. (SMF RII 11417 / 20). Additional material: 1 ♂, 1 ♀ ' PERU, Huanuco, Bosque Carpish, southwest Tingo Maria, 2,590 m a. s. l., 09 ° 42 ′ 46.6 ″ S 76 ° 05 ′ 26.7 ″ W, 23 / IV / 2011, A. Benedetti, J. Ochoa, R. Pinto-da-Rocha & D. Silva leg. (MZSP 76550).	en	Benedetti, Alipio Rezende, Pinto-da-Rocha, Ricardo (2022): Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species. Arthropod Systematics & amp; Phylogeny 80: 309-388, DOI: http://dx.doi.org/10.3897/asp.80.e73829, URL: http://dx.doi.org/10.3897/asp.80.e73829
