taxonID	type	description	language	source
225423B41B4231E755FE9D671377258D.taxon	diagnosis	Diagnosis. The Graphipterus serrator group is included in the genus Graphipterus based on the following combination of characters: Clypeus concave at anterior margin, posteriorly well separated from front; labrum wide and short, with well-developed microsculpture and six setiferous pores. Mandibles broad at the base, sharp and strongly curved at tip; labial and maxillary palps long and slender, glabrous with exception of distal end of segments which bear a few hairs; last palpal segments slightly thicker than penultimate ones. Pronotum transverse and cordiform, slightly convex, usually ornamented by colored scales at the lateral bead, disc with or without scales. Anterior and posterior angles obtuse. Scutellum triangular, small and short, often hidden by the pronotal base. Flightless. Elytra wide and oval, slightly convex, coalesced along suture, humeri completely rounded; surface covered by dense or sparse scales, white scales creating longitudinal stripes on the radial field and spots on the disc; apex almost truncate. Pygidium not covered by elytra, last visible tergite with colored scales. Legs long, usually black or brown, protibia with clypsetae (antenna cleaner) and dark parallel spurs, as long as 3 / 4 of protarsomere 1. Mesotibia with two long and thin not serrated spurs, metatibia with one long and thin not serrated spur and one shorter, wide and obtuse spur. Claws of all legs long and smooth on median margin.	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
3F331E47F3E3F0B23160994812E2318B.taxon	diagnosis	Diagnosis. Medium-sized species with grayish or yellowish scales usually cover the elytra and sometimes also on the pronotum. Elytra pattern rarely visible with six lateral margin extensions and 18 - 24 isolated white circular to elongated spots occur on elytra.	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
3F331E47F3E3F0B23160994812E2318B.taxon	description	Description. BL male: 13.0 - 17.0 mm, average 15.8 + / - 1.5 mm; BL female: 15.5 - 17.0 mm, average 16.3 + / - 0.6 mm. Grayish with elytral white blurred spots and extensions. Head slender: HW / PW: 0.72; EYL: 1.1 - 1.4 mm; EYL / EL: 0.15. Mentum without teeth (Fig. 3 a). Frontal ridge reduced. In male, apical white frons stripes slenderer than exposed frons (cf. Fig. 4 a). Pronotum wide; PL / PW: 0.63; BPW / BPW / PW: 0.6; posteromedially concave and with white lateral margin, as wide as antennomere I long: white slushy scales cover disc sometimes. Elytra wide, elytron margin almost continuously rounded from humeri to posterolateral angles; EL: 7.1 - 8.8 mm, average 8.2 mm; EW: 6.5 - 7.8 mm, average 7.3 mm; EL / EW: 1.1. Elytra longitudinally flat, usually with grayish scales, disc visible between scales (Fig. 6 c); extensions and lateral margin blurred. Lateral margin nearly as wide as antennomere I long and with six extensions; extension I usually elongated; white posterior margin almost touches suture at apex. Disc with 18 - 24 rounded, usually elongated spots, anterior pair of spots elongate, as wide as extension I; posterior pair of spots rounded, located toward suture; round spots located posterior to third extensions laterally in imaginary lateral line as posterior spots. Apical sinuation slightly developed to straight, apex not protuberant, broadly rounded, especially medially (Fig. 7 c). Suture inconspicuous. Legs medium; MTIL: 4.5 - 6.5 mm, average 5.7 mm; El / MTIL: 1.7. Metatibial secondary spur brown. MTAL: 3.5 - 4.4 mm, average 4 mm; MTAL / MTIL: 0.8. Claws of hind legs brown at base. Median lobe of aedeagus with short unbent tip (Fig. 9 a).	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
3F331E47F3E3F0B23160994812E2318B.taxon	distribution	Distribution. Restricted to north-east Tunisia (Fig. 17).	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
3F331E47F3E3F0B23160994812E2318B.taxon	conservation	Conservation. The restricted distribution range of the endemic species and the decline of the coastal sandy habitat as a result of increasing anthropogenic pressures (e. g., tourism activities, urbanization, etc.) threaten at least the long-term survival of the species.	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
013EE54635BF470E334E0F5716904BE1.taxon	diagnosis	Diagnosis. Large species with 18 - 26 isolated white round spots on elytra, anterior and posterior discal spots larger than other spots; four marginal extensions, anterior extension triangular; median lobe of aedeagus with ventrally bent apex.	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
013EE54635BF470E334E0F5716904BE1.taxon	description	Description. BL male: 17.1 - 20.9 mm, average 18.9 + / - 1.6 mm; BL female: 18 - 20 mm, average 19.4 + / - 1.4 mm. Head slender; HW / PW: 0.71; EYL: 1.5 - 1.9 mm; EYL / EL: 0.16. Mentum with two teeth and concavity between them (Fig. 3 b). Frontal ridge absent. In male, apical white frons stripes slenderer than exposed frons (Fig. 4 a). Pronotum slender; PL / PW: 0.65; BPW / PW: 0.7; posteromedially concave and without white margin; white lateral margin as wide as antennomere I long. Elytra oval, humeri rounded; EL: 10.0 - 12.1 mm, average 10.9 mm; EW: 8.4 - 9.9 mm, average 9.16 mm; EL / EW: 1.2. Lateral cross section convex. Black scales dense, disc not visible between them (Fig. 6 a). White lateral margin almost as wide as antennomere I long and with four extensions; extension I triangular with rounded angels, slightly wider at margin of elytra, slightly elongated, wider and shorter than extension II; the latter one elongated at third quarter of elytra, imaginary line connecting the media ends of the extensions I and II parallel to the suture; white posterior margin forms a gap at suture, wider than lateral margin. Disc usually with 18 - 26 rounded spots; anterior pair of spots rounded, as wide as extension I, usually smaller than posterior spots, larger than spots on mid disc; mid disc spots usually asymmetrically smeared. Posterior pair of spots rounded, one or two small spots located laterally to posterior spots. Apical sinuation strongly developed, apex protruded, almost rectangular, only slightly rounded at most distant tip (Fig. 7 a). Suture inconspicuous. Legs long; MTIL: 5.3 - 7.0 mm, average 6.5 mm; El / MTIL: 1.7. Metatibial secondary spur brown. MTAL: 4.4 - 5.2 mm, average 4.8 mm; MTAL / MTIL: 0.7. Claws of hind legs black at base. Median lobe of aedeagus with apex bent ventrally (Fig. 9 b).	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
013EE54635BF470E334E0F5716904BE1.taxon	distribution	Distribution. Western Lybia (Tripolitania) and western Tunisia (Nefzaoua) (Fig. 18).	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
013EE54635BF470E334E0F5716904BE1.taxon	conservation	Conservation. The restricted distribution range of the endemic species and the decline of the coastal sandy habitat as a result of increasing anthropogenic pressures (e. g., tourism, urbanization etc.) threaten at least the long-term survival of the species.	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
C11D1C1D6CA5A908A582A8EA674E2A15.taxon	diagnosis	Diagnosis. Medium-sized species with 22 - 30 isolated white, usually elongated elytral spots, six very short marginal extensions, and a series of 8 - 12 elongated spots along suture form a broken line. Median lobe of aedeagus with apex slightly bent ventrally.	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
C11D1C1D6CA5A908A582A8EA674E2A15.taxon	description	Description. BL male: 15.0 - 17.5 mm, average 15.8 + / - 1.5 mm; BL female: 15 - 18 mm, average 16.3 + / - 0.6 mm. Head medium; HW / PW: 0.74; EYL: 1 - 1.6 mm; EYL / EL: 0.15. Mentum with two or three teeth. Frontal ridge absent. In male, apical white frons stripes wider than exposed frons, (cf. Fig. 4 b). Pronotum cordiform; PL / PW: 0.64; BPWBPW / PW: 0.61; posteromedially concave and without white margin; white lateral margin as wide as antennomere I long. Elytra oval, humeri rounded; EL: 7.3 - 9.9 mm, average 8.9 mm; EW: 5.8 - 8.4, average 7.7 mm; EL / EW: 1.16. Lateral cross section quite flat suture conspicuous. Scales brown, disc visible between them (Fig. 6 b). White lateral margin narrow, as wide as half antennomere I long and with six extensions, rarely four; extension I usually elongated, sometimes constricted at the base to the lateral margin; extension II elongated, constricted or absent; extension III elongated; white posterior margin almost as wide as the lateral margin, gap at suture smaller than lateral margin or even absent. Disc usually with 22 - 30 mostly elongate small spots; anterior pair of spots slightly elongate, wide as extension I, lateral spots rounded, adjacent or sometimes fused to extension II, posterior pair of spots rounded, slightly larger than others, located toward suture, round spots slightly smaller than all others, located posteriori to third extensions and laterally to posterior spots; a series of 8 - 12 elongated spots along the suture. Apical sinuation developed, apex slightly protruded, strongly rounded (Fig. 7 b). Legs medium; MTIL: 4.5 - 5.9 mm, average 5.4 mm; El / MTIL: 1.7. Metatibial secondary spur brown. MTAL: 3.4 - 4.7 mm, average 4.4 mm; MTAL / MTIL: 0.8. Claws of hind legs brown at base. Median lobe of aedeagus with ventrally bent tip (Fig. 9 c).	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
C11D1C1D6CA5A908A582A8EA674E2A15.taxon	distribution	Distribution. Graphipterus luctuosus presents the widest distribution range of the group: from Laghouat, more than 300 km inland Algeria to the arid and semi-arid regions of north-east Algeria, over most of the Tunisian coast and east up to Sirte on the Libyan coast (Fig. 16).	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
C11D1C1D6CA5A908A582A8EA674E2A15.taxon	conservation	Conservation. The species does not seem to be endangered as it apparently lives in numerous habitats. Consequently, it might not be so strongly affected by human activities.	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
72A7596F66B4213C148E6BD1F35319EF.taxon	diagnosis	Diagnosis. Large species with 20 - 24 white rounded and elongated elytra spots; six white marginal extensions, extension I elongated. Elytra wide, lateral margin strongly and continuous rounded. Aedeagus elongated, thin and with apex slightly bent ventrally (Fig. 9 d).	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
72A7596F66B4213C148E6BD1F35319EF.taxon	description	Description. BL male: 18.3 - 20.1 mm, BL female: unknown. Average 19.4 + / - mm. Head slender; HW / PW: 0.72; EYL: 1.7 mm; EYL / EL: 0.17. Frontal ridge well developed. In male, apical white frons stripes slenderer than exposed frons (cf. Fig. 4 a). Pronotum cordiform; PL / PW: 0.62; BPW / BPW / PW: 0.68; posteromedially concave and without white margin; white lateral margin as wide as antennomere I long. Elytra wide, rounded, rounded-like, humeri strongly narrowed; EL: 9 - 10.7 mm, average 9.7 mm; EW: 8.5 - 9.0 mm, average 8.7 mm; EL / EW: 1.1. Lateral cross section quite flat. Scales black, disc not visible between them (cf. Fig. 6 a). White lateral margin nearly as wide as antennomere I long and with six extensions; extensions I slightly elongated, wider close to the margin; extensions II and III in front of middle. White posterior margin as wide as lateral margin or wider, sutural gap slenderer than lateral margin. Disc usually with 20 (rarely up to 24) spots; anterior pair of spots rounded, wider than extension I, 6 - 8 spots adjacent elongated and parallel to suture, posterior pair of spots rounded, additional 1 - 3 small spots frequently present laterally to posterior ones. Apical sinuation strongly developed, apex protruded, almost rectangular, only slightly rounded at most distant tip (Fig. 7 a). Suture conspicuous. Legs long; MTIL: 6.2 - 6.8, average 6.5 mm; El / MTIL: 1.53. Metatibial secondary spur brown. MTAL: 5.2 mm; MTAL / MTIL: 0.8. Claws of hind legs brown at base. Median lobe of aedeagus long and thin with apex hardly bent ventrally (Fig. 9 d).	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
72A7596F66B4213C148E6BD1F35319EF.taxon	etymology	Etymology. The species name is derived from Latin (magnus) and refers to the large body size.	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
72A7596F66B4213C148E6BD1F35319EF.taxon	conservation	Conservation. Unknown.	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
72A7596F66B4213C148E6BD1F35319EF.taxon	distribution	Distribution. The only known records are from Buq Buq in north-east Egypt (Fig. 19).	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
EC22FA8E268EF64062176E7A448E6E10.taxon	diagnosis	Diagnosis. Medium-sized species with (18 -) 22 white, mostly elongated spots on elytra, anterior and posterior spots larger than other spots; six marginal extensions, extension I usually triangular. Median lobe of aedeagus with short apex unbent ventrally (somewhat similar to that of G. barthelemyi).	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
EC22FA8E268EF64062176E7A448E6E10.taxon	description	Description. BL male: 15.1 - 17.5 mm, average 16.6 + / - 1.1 mm. Females were not available. Head medium; HW / PW: 0.76; EYL: 1.3 - 1.5 mm; EYL / EL: 0.16. Mentum with two teeth (cf. Fig. 3 b). Frontal ridge reduced. In male, apical white frons stripes wider than exposed frons (cf. Fig. 4 b). Pronotum cordiform; PL / PW: 0.66; BPW / PW: 0.63; posteriomedially concave and without white margin; white lateral margin as wide as antennomeres I + II long. Elytra relatively elongated oval humeri slightly narrowed; EL: 8.6 - 9.7 mm, average 9.2 mm; EW: 6.8 - 8.0 mm, average 7.4 mm; EL / EW: 1.2. Lateral cross section quite flat. Dense black scales, disc not visible between them (Fig. 6 a). White lateral margin as wide as half of antennomere I long and with six extensions; extension I triangular; extension II shorter than extension III. White posterior margin as wide as lateral margin, sutural gap slenderer than lateral margin. Disc usually with 18 - 22 rounded to elongate spots; anterior spot elongated, as wide as extension I, anterior and posterior spots larger than all other ones, posterior one rounded. Apical sinuation developed, apex slightly protruded, strongly rounded (Fig. 7 b). Suture conspicuous. Legs long; MTIL: 5.3 - 5.9 mm, average 5.7 mm; El / MTIL: 1.61. Metatibial secondary spur brown at base, MTAL: 3.8 - 4.5 mm, average 4.2 mm; MTAL / MTIL: 0.74. Claws of hind legs brown at base. Median lobe of aedeagus with short apex, unbent ventrally (Fig. 9 e).	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
EC22FA8E268EF64062176E7A448E6E10.taxon	etymology	Etymology. The species name is derived from ancient Latin (Mauretania, - ensis).	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
EC22FA8E268EF64062176E7A448E6E10.taxon	distribution	Distribution. As we found in all collections only nine specimens of G. mauretensis sp. n., our knowledge of its distribution range is limited. Known from central coast of Mauritania to more than 400 km inland to Glebat el M'Boza Adrar (Fig. 16).	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
EC22FA8E268EF64062176E7A448E6E10.taxon	conservation	Conservation. Unknown.	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
8DF18127E58A84F2F65213B3D8FA9BE0.taxon	diagnosis	Diagnosis. The two subspecies of G. minutus are distinguished from all other species of the G. serrator group by smaller size, lack of the stridulatory structure, unique pronotum shape (G. serrator group excluding G. minutus: BPW / PW: 0.6 - 0.7, G. minutus: BPW / PW: 0.46) and flat tip of median lobe.	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
8DF18127E58A84F2F65213B3D8FA9BE0.taxon	description	Description. BL male: 10.3 - 13.5 mm, average 12 + / - 1.2 mm; BL female: 10.5 - 15.2 mm, average 13.1 + / - 1.9 mm; Head wide; HW / PW: 0.77; EYL: 1 - 1.8 mm; EYL / EL: 0.15. Frontal ridge slightly developed. Male with two short parallel frontal stripes of white scales usually diverging apically, became wispy, not reach the level of supraorbital setiferous pores. Mentum usually with two pronounced teeth (Fig. 3 e). Pronotum strongly cordiform PL / PW: 0.54; BPW / PW: 0.46; posteromedially flat and without white margin; white lateral margin as twice as antennomere II long. Elytra almost rounded, humeri stringly rounded, lateral margin continuously rounded; EL: 5.3 - 7.5 mm, average 6.6; EW mm: 4.8 - 7.6 mm, average 6.1 mm; EL / EW: 1.16. Suture inconspicuous. Scales black, disc not visible between them (cf. Fig. 6 a). Lateral cross section convex. Apical sinuation almost lacking, apex almost absent, not rounded (Fig. 7 d). White lateral margin usually nearly as wide as antennomere I long and usually with two extensions; extension I elongated from humeri posteriorly; extension II usually absent, sometimes indistinct wider section of lateral margin at its middle. White posterior margin becomes narrower toward the tip, usually disappearing in front of it; gap at suture wider than lateral margin. Disc with 36 - 40 mostly rounded spots; usually 12, sometimes ten or 14 rounded to elongated, not fused spots located parallel to suture; anterior spot as wide as extension I. Stridulatory structure absent. Legs short; MTIL: 2.54 - 4.0 mm, average 3.3 mm; El / MTIL: 1.9 mm. Metatibial secondary spur brown, MTAL length: 2.5 - 3.3 mm, average 2.9 mm; MTAL / MTIL: 0.85. Claws of hind legs brown at base. Median lobe of aedeagus with wide and flat tip (Fig. 9 f).	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
8DF18127E58A84F2F65213B3D8FA9BE0.taxon	distribution	Distribution. Syria, east and south Jordan, north Saudi Arabia, Iraq, and Iran (Fig. 19).	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
8DF18127E58A84F2F65213B3D8FA9BE0.taxon	conservation	Conservation. The species does not seem to be endangered as it has a wide distribution range that is not strongly affected by human activities.	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
FB4892C1E89E4B2BBACBA53B59C0703E.taxon	diagnosis	Diagnosis. Small-sized taxon with 28 - 30 mostly elongated white spots, usually with several spots fused with lateral margin, and with series of usually ten elongated spots, regularly at least several are fused to each other along median suture. Two marginal extensions elongated from humeri posteriorly. Median lobe of aedeagus with wide and flat tip.	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
FB4892C1E89E4B2BBACBA53B59C0703E.taxon	description	Description. BL male: 11.2 - 11.8 mm, average 11.5 + / -. 02 mm; BL female: 11.4 - 13.6 mm, average 12.2 + / - 0.9 mm. Head wide; HW / PW: 0.78; EYL: 1.1 - 1.3 mm; EYL / EL: 0.19. Frontal ridge absent. Male with two short parallel frontal stripes of decumbent white scales usually diverging apically became wispy, not reaching the level of supraorbital setae. Mentum with two pronounced teeth (cf. Fig. 3 e). Pronotum strongly cordiform; PL / PW: 0.57; BPW / PW: 0.5; posteriomedially flat and without white margin; white lateral margin as twice as antennomere II long. Elytra almost rounded, humeri strongly rounded, lateral margin continuously rounded; EL: 5.9 - 7.3 mm, average 6.2 mm; EW: 5.2 - 6.8 mm, average 5.7 mm; EL / EW: 1.1. Lateral cross section convex. Scales black, disc not visible between them (cf. Fig. 6 a). White lateral margin nearly as wide as antennomere I long and with six, rarely fewer, elongated extensions; extensions I elongated from humeri posteriorly; extensions II and III in front and behind the middle of lateral margin, usually much longer than lateral margin wide. White posterior margin becomes narrower toward the tip, usually disappearing in front of it; gap at suture as wide as lateral margin. Disc with 28 - 30 mostly elongated spots, several spots fused with lateral margin resulting in extensions II and III, a series of 10, (rarely 12 - 14), elongated spots fused to each other parallel to suture, anterior spot as wide as extension I. stridulatory structure absent. Apical sinuation almost lacking, apex almost absent, not rounded (Fig. 7 d). Suture inconspicuous. Legs short; MTIL: 2.5 - 3.3 mm, average 3.0 mm; El / MTIL: 1.9. Metatibial secondary spur brown. MTAL: 2.1 - 3.8 mm, average 2.9 mm; MTAL / MTIL: 0.87. Claws of hind legs brown at base. Median lobe of aedeagus with wide and flat tip (Fig. 9 g).	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
FB4892C1E89E4B2BBACBA53B59C0703E.taxon	distribution	Distribution. Saudi Arabia and Iraq (Fig. 19). There are old records from Iran (Perse), but without indication of exact locality.	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
FB4892C1E89E4B2BBACBA53B59C0703E.taxon	conservation	Conservation. The species does not seem to be endangered as it has a wide distribution range in desert regions that are not strongly affected by human activities.	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
22129DDD1DF909F221AA7DC3FC7B6970.taxon	diagnosis	Diagnosis. Small species with 16 - 20 white, mostly elongated spots on elytra, only posterior discal spots rounded; 4 - 6 marginal extensions, extension I oriented slightly posteriorly. Median lobe of aedeagus with ventrally short, unbent tip.	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
22129DDD1DF909F221AA7DC3FC7B6970.taxon	description	Description. BL male: 10.0 - 15.0 mm, average 13.0 + / - 1.3 mm; BL female: 11.5 - 16.0 mm, average 14.0 + / - 1.2 mm. Head wide; HW / PW: 0.76; EYL: 1.0 - 1.6 mm; EYL / EL: 0.17. Frontal ridge slightly developed. In male, apical white frons stripes wider than exposed frons (Fig. 4 a); stripes elongate, reaching the level of supraorbital setae (populations east of the Dead Sea-Rift Valley), or being shorter (populations west of the Dead Sea-Rift Valley). Mentum with 2 - 3 teeth. Pronotum cordiform; PL / PW: 0.66; BPW / PW: 0.64; posteromedially concave and without white margin; white lateral margin as wide antonomer 1 long. Elytra oval, humeri rounded; EL: 4.5 - 9.1 mm, average 7.7 mm; EW: 4.1 - 8.0 mm, average 6.4 mm; EL / EW: 1.2. Lateral cross section quite flat. Elytra with Dense black scales, disc not visible between them (cf. Fig. 6 a). White lateral margin nearly as wide as half of antennomere I long and with 6, sometimes four extensions; extension I medium long, shorter than anterior spot, but longer than extension II and shorter than extension III, which is wider than lateral margin; extension II sometimes constricted, rarely absent or fused with lateral disc spot. White posterior margin as wide as lateral margin, gap at suture smaller than lateral margin. Disc usually with 16 sometimes 18 rounded to elongate spots; anterior spot slightly elongate, longer than extension I; lateral spots rounded, adjacent, or sometimes fused to extension II, six spots forming an arch pattern anteriorly and laterally to posterior rounded larger spots. Apical sinuation slightly developed to straight, apex not protuberant, broadly rounded, especially on the medial side (Fig. 7 c). Suture conspicuous. Legs long; MTIL: 3.7 - 5.5 mm, average 4.7 mm; El / MTIL: 1.6. Metatibial secondary spur brown at base, MTAL: 3 - 4.5 mm, average 3.7 mm; MTAL / MTIL: 0.8. Claws of hind legs black at base. Median lobe of aedeagus with short, unbent tip (Fig. 9 h).	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
22129DDD1DF909F221AA7DC3FC7B6970.taxon	distribution	Distribution. Egypt, Israel, and Jordan (Fig. 19). The only Jordanian population of which we are aware lives between Aqaba to Ma'an, and inhabits a flat sand dune area without or only slightly developed crust. In the same habitat Anthia (Thermophilum) duodecimguttata (Bonelli, 1813) and Amara maindromi Bedel, 1907 occur.	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
22129DDD1DF909F221AA7DC3FC7B6970.taxon	conservation	Conservation. The species does not seem to be endangered as it has a wide distribution range and it prefers habitats that are not strongly affected by human activities. However, in Israel, in the Dead Sea region and the Arava Valley, G. multiguttatus has been collected in the past, but no longer exists there. The latest records from these regions are Ein Gedi, 1976; Ein Husub, 1956 (leg. unknown, both specimens preserved in KCE); Sedom road, 1953; Ein-Radian, 1958 (leg. Ch. Lewinsohn, both specimens preserved in SMNHTAU). Habitats for G. multiguttatus on the Israeli side of the Arava valley may have disappeared. Anthia (Thermophilum) duodecimguttata (Bonelli, 1813), one of the co-occurring ground beetle species of the Jordanian population of G. multiguttatus, was last found in 2003 in Israel (coll. U. Shanas, V. Chikatunov, SMNHTAU; pers. obs.).	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
22129DDD1DF909F221AA7DC3FC7B6970.taxon	biology_ecology	Biology. Seasonality and daily activity time are in the same as in G. serrator (see there), but the species seems to spend more time under shrubs. Graphipterus multiguttatus prefers stabilized and semi-stabilized sand with high vegetation. The population densities in the sympatric areas of the distribution ranges are lower than those for G. serrator. The beetles dig burrows between the hard crust layer and the soft sand, sometimes close to the dwarf-shrubs. Frequently, the openings do not collapse or become covered by sand. The beetles sometimes close the openings with sand from inside. Diet, intraspecific behavior including copulation and the chirping sounds produced by the stridulatory structure, are same as in G. serrator.	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
147945E8C7AD6A4F87B4CC5F518CDC26.taxon	diagnosis	Diagnosis. Predominantly dark, medium-sized species with 18 - 24 small, mostly rounded white spots on elytra, four usually short marginal extensions. Median lobe of aedeagus with ventral, short, unbent tip.	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
147945E8C7AD6A4F87B4CC5F518CDC26.taxon	description	Description. BL male: 13.9 - 14.8 mm, average 14.3 + / - 0.4 mm; BL female: 11.5 - 16.1 mm, average 13.6 + / - 1.8 mm. Head medium; HW / PW: 0.76; EYL: 1 - 1.5 mm; EYL / EL: 0.17. Mentum with usually three teeth (Fig. 3 d). Frontal ridge absent. In male, apical white frons stripes slenderer than exposed frons (cf. Fig. 4 a). Pronotum strongly cordiform, PL / PW: 0.63; BPW / PW: 0.6; posteromedially concave and without white margin; white lateral margin as wide as antennomere I long. Elytra oval, humeri rounded; EL: 7.4 - 9.1 mm, average 8; EW: 5.7 - 7.9 mm, average 7.0 mm; EL / EW: 1.15. Elytra with brown scales, disc of elytra visible between scales (cf. Fig. 6 b). White lateral margin wide as half antennomere I long and with four extensions; extensions often constricted; extension I elongated, shorter than extension II; sutural gap of white posterior margin wider than lateral margin. Disc usually with 18 - 24 mostly rounded spots; anterior pair of spots slightly elongate, as wide as extension I, lateral spots rounded, adjacent or sometimes fused to extension II, posterior pair of spots rounded, slightly larger than others, located toward suture. Lateral cross section quite flat. Apical sinuation developed, apex slightly protruded, strongly rounded (cf. Fig. 7 b). Suture inconspicuous. Legs long; MTIL: 3.8 - 5.3 mm, average 4.8 mm; El / MTIL: 1.7. Metatibial secondary spur brown. MTAL: 2.8 - 4.0 mm, average 3.5 mm; MTAL / MTIL: 0.7 (all other species of the G. serrator group El / MTIL: 0.8). Claws of hind legs brown at base. Median lobe of aedeagus with ventrally short, not bent tip (Fig. 9 i).	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
147945E8C7AD6A4F87B4CC5F518CDC26.taxon	distribution	Distribution. North-west Algeria and north Morocco (Fig. 17).	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
147945E8C7AD6A4F87B4CC5F518CDC26.taxon	conservation	Conservation. The species does not seem to be threatened as it has a wide distribution range that appears to be mostly not strongly affected by human activities.	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
EBD5D3B65FFE2B0B0BE63BEDD81C75AC.taxon	diagnosis	Diagnosis. Medium-sized species with usually 24 white large rounded and elongated spots on elytra; posterior discal spots slightly larger than other spots; six marginal extensions (Fig. 25 a). Median lobe of aedeagus with slightly bent tip.	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
EBD5D3B65FFE2B0B0BE63BEDD81C75AC.taxon	description	Description. BL male: 15.5 - 19.8 mm, average 17.5 + / - 2.1 mm; BL female: 17 - 17.9 mm, average 17.5 + / - 0.3 mm. Head medium; HW / PW: 0.76; EYL: 1.5 - 1.8 mm; EYL / EL: 0.17. Mentum with two or three teeth. Frontal ridge absent. In male, apical white frons stripes wider than exposed frons (cf. Fig. 4 b). Pronotum cordiform; PL / PW: 0.63; BPW / PW: 0.66; posteromedially concave and without white margin; white lateral margin as wide antennomere I long. Elytra oval, humeri rounded, but slightly protruding; EL: 8.2 - 11.0 mm, average 9.4 mm; EW: 6.5 - 9.2 mm, average 8.1 mm; EL / EW: 1.2. Lateral cross section quite flat. Suture conspicuous. Black scales dense, disc not visible between scales (cf. Fig. 6 a). White lateral margin nearly as wide as 11 / 2 antennomere I long and with six extensions; extension I triangular, slightly elongated and posteriori oriented; extensions II and III frequently constricted at base, usually wider than lateral margin. White posterior margin as wide as lateral margin or wider, not becoming narrower towards the suture; gap at suture smaller than lateral margin. Disc usually with 24 rounded to elongate, moderate large spots; anterior spot elongated, as wide as extension I, posterior discal spots slightly larger than other spots, series of six elongated spots along suture, sometimes fused to each other; posterior discal spots larger than other spots. Apical sinuation strongly developed, apex protruded, almost rectangular, only slightly rounded at most distant tip (cf. Fig. 7 a). Legs long; MTIL: 5.3 - 6.6 mm, average 6.1 mm; El / MTIL: 1.54. Metatibial secondary spur brown at base, MTAL: 4 - 5.7.0 mm, average 4.9 mm; MTAL / MTIL: 0.8. Claws of hind legs brown at base. Median lobe of aedeagus with slightly bent tip (Fig. 9 j).	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
EBD5D3B65FFE2B0B0BE63BEDD81C75AC.taxon	etymology	Etymology. The species is dedicated to Pinchas (Pini) Amitai, an inspiring entomologist and mentor who wrote the first Hebrew photographed insect guide.	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
EBD5D3B65FFE2B0B0BE63BEDD81C75AC.taxon	distribution	Distribution. Restricted to Central Tunisia, from the vicinity of Kebili to Gabes (Fig. 18).	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
EBD5D3B65FFE2B0B0BE63BEDD81C75AC.taxon	conservation	Conservation. The species does not seem to be endangered, as the preferred habitat is not strongly affected by human activities.	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
F091445356C4F7E2BF3122F396406632.taxon	diagnosis	Diagnosis. Large species with 20 - 24 isolated white round spots on elytra, six marginal extensions, extension II short, almost triangular. Humeri very narrowed, maximum width of elytra at interior rear third. The discal elytra pattern comprises a group of 8 - 12 elongated spots in an order parallel to the suture. Median lobe of aedeagus with ventrally bent tip.	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
F091445356C4F7E2BF3122F396406632.taxon	description	Description. BL male: 17 - 18, average 17.6 + / - 0.4 mm; BL female: 17.4 - 21.4, average 19.3 + / - 2 mm. Head medium; HW / PW: 0.76; EYL: 1.6 - 1.8 mm; EYL / EL: 0.17. Mentum with three teeth (cf. Fig. 3 d). Frontal ridge absent. In male, apical white frons stripes wider than exposed frons (Fig. 4 b). Pronotum wide; PL / PW: 0.72; BPW / PW: 0.63; posteromedially concave and without white margin; white lateral margin as wide as antennomere I long. Elytra with strongly narrowed humeri; EL: 9.4 - 10.3 mm, average 9.75 mm; EW: 8.0 - 8.5, average 8.3 mm; EL / EW: 1.2. Lateral cross section convex. Suture conspicuous. Scales brown, disc visible between them (Fig. 6 b). White lateral margin nearly as wide as antennomere I long and with six extensions; extension I triangular with rounded tip, slightly more elongated than in G. serrator, wider and shorter than extensionII; the latter one elongated, at third quarter of elytra. White posterior margin commonly slightly wider than lateral margin, gap at suture smaller than lateral margin, usually with a small, indistinct tip anteriorly. Disc usually with 20 - 24 (rarely 18), mainly rounded spots; anterior pair of spots rounded, wide as extension I, usually smaller than posterior spots, but larger than spots on central disc; central disc spots usually asymmetrically smeared; posterior pair of spots rounded; one or two small additional spots adjacent laterally to the posterior ones. Apical sinuation strongly developed, apex protruded, almost rectangular, only slightly rounded at most distant tip (Fig. 7 a). Legs long; MTIL: 5.8 - 6.1 mm, average 5.9 mm; El / MTIL: 1.6. Metatibial secondary spur brown. MTAL length: 4.2 - 5, average 4.7 mm; MTAL / MTIL: 0.8. Claws of hind legs black at base. Median lobe of aedeagus with ventrally bent tip (Fig. 9 k).	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
F091445356C4F7E2BF3122F396406632.taxon	distribution	Distribution. Morocco (Fig. 16).	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
F091445356C4F7E2BF3122F396406632.taxon	conservation	Conservation. Unknown.	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
D8615B1867B16CDD64AB9B9D1DCE011E.taxon	diagnosis	Diagnosis. Small species with large distribution range, high variation in size (15 - 19 mm) and variation in elytra pattern (4 - 6 extensions, 16 - 22 spots). posterior discal spots larger than other spots; six spots usually forming an arc pattern anterior and lateral to posterior spots; Median lobe of aedeagus with short, slightly bent tip.	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
D8615B1867B16CDD64AB9B9D1DCE011E.taxon	description	Description. BL male: 15.0 - 19.0 mm, average 17.4 + / - 1.5 mm; BL female: 15.4 - 17.1 mm, average 16.1 + / - 1.3 mm. Head slender; HW / PW: 0.72; EYL: 1.4 - 1.7 mm; EYL / EL: 0.16. Mentum with 2 - 3 teeth. Frontal ridge slightly developed. In male, apical white frons stripes wider than exposed frons (Fig. 4 b). Pronotum cordiform; PL / PW: 0.65; BPW / PW: 0.69; posteromedially concave and without white margin; white lateral margin as wide antonomer 1 long. Elytra oval, humeri rounded; EL: 8.9 - 11.0 mm, average 9.7 mm; EW: 7.0 - 8.7 mm, average 7.8 mm; EL / EW: 1.25. Lateral cross section quite flat. Dense black scales, disc not visible between scales (Fig. 6 a). White lateral margin nearly as wide as half antennomere I long and with six, sometimes fouor extensions; extension I triangular to slightly elongated; extension II absent or only weakly developed, rarely fused with lateral disc spot. White posterior margin becomes narrower towards the suture, gap at suture smaller than lateral margin. Disc usually with 18, sometimes 16 or 22 rounded to weakly elongate spots; anterior spot slightly elongated, wide as extension I, six spots usually forming an arc pattern anterior and lateral to posterior rounded, larger spots. Apical sinuation strongly developed, apex protruded, almost rectangular, only slightly rounded at most distant tip (Fig. 7 a). Suture conspicuous. Legs long; MTIL: 4.3 - 5.2 mm, average 4.7 mm; El / MTIL: 1.63. Metatibial secondary spur dark at base, MTAL: 5.4 - 6.9 mm, average 6 mm; MTAL / MTIL: 0.8. Claws of hind legs brown at base. Median lobe of aedeagus with slightly bent tip (Fig. 9 l).	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
D8615B1867B16CDD64AB9B9D1DCE011E.taxon	distribution	Distribution. Algeria, Tunisia, and the coastal region of west Libya (Fig. 18).	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
D8615B1867B16CDD64AB9B9D1DCE011E.taxon	conservation	Conservation. The species does not seem to be endangered as it has a wide distribution range which is not strongly affected by human activities.	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
D8615B1867B16CDD64AB9B9D1DCE011E.taxon	biology_ecology	Biology. The three larval stages develop during the summer inside ant nests. The first larval instar is nearly 4 mm long and creeps into nests of large ant species, digs there a chamber, preys on the ant's brood and pupates within the nest. When the first larval instar tries to enter nests of small ants, it is attacked by the ants (Paarmann 1985; Dinter et al. 2002). The larval instars have a mandibular suctorial tube to suck hemolymph from their prey (Brandmayr 1994 a, 1994 b). Four specimens from the species studied by Wilfried Paarmann, Pietro Brandmayr, and their co-workers were examined; the material belongs to G. rotundatus and not to G. serrator as noted in their publications.	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
5509E63269C3FF275AC39F4B7F99BA28.taxon	diagnosis	Diagnosis. Large species with 10 - 12 isolated white round spots on elytra: anterior and posterior discal spots larger than other spots, six smaller spots near suture form circular pattern on disc; four white marginal extensions present, extension I triangular. Median lobe of aedeagus with ventrally bent tip.	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
5509E63269C3FF275AC39F4B7F99BA28.taxon	description	Description. BL male: 17 - 18 mm, average 17.6 + / - 0.4 mm; BL female: 17.4 - 21.4 mm, average 19.3 + / - 2 mm. Head medium; HW / PW: 0.76; EYL: 1.6 - 1.8.0 mm; EYL / EL: 0.16. Mentum with three teeth, mid tooth shallow (Fig. 3 f). Frontal ridge absent. In male, Apical white frons stripes slenderer than exposed frons (Fig. 4 b). Pronotum wide; PL / PW: 0.58; BPW / PW: 0.65; posteromedially concave and without white margins; white lateral margin as wide as antennomere I long. Elytra oval, humeri rounded; EL: 9.3 - 11.3 mm, average 10.3 mm; EW: 7.0 - 9.8 mm, average 8.4 mm; EL / EW: 1.2. Lateral cross section convex. Elytra with dense black scales, disc of elytra not visible between scales (Fig. 6 a). White lateral margin nearly as wide as antennomere I long and with four extensions; extension I triangular with rounded angels, margin of elytra wider and shorter than extension II; the latter one elongated; at third quarter of elytra, imaginary line connecting the medial ends of the extension I and I parallel to the suture. White posterior margin forming gap at suture which is wider than lateral margin. Disc usually with 10, sometimes 12 round spots; anterior pair of spots circular to slightly elongate, narrower than extension I, larger than the six central spots forming a circular pattern; anterior and posterior pair of spots circular rounded, larger than other spots; small additional spots frequently present laterally to the posterior spots. Apical sinuation strongly developed, apex protruded, almost rectangular, only slightly rounded at most distant tip (Fig. 7 a). Suture inconspicuous. Legs long; MTIL: 4.8 - 7.4 mm, average 6.1 mm; El / MTIL: 1.7. Metatibial secondary spur dark. MTAL: 4.0 - 5.3 mm, average 4.6 mm; MTAL / MTIL: 0.8. Claws of hind legs black at base. Median lobe of aedeagus with ventrally bent tip (Fig. 9 m).	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
5509E63269C3FF275AC39F4B7F99BA28.taxon	distribution	Distribution. North-east Egypt (incl. Sinai) and Israel. In Israel it is restricted to the western Negev sand dunes (Fig. 19).	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
5509E63269C3FF275AC39F4B7F99BA28.taxon	conservation	Conservation. The sand dunes in the western Negev suffer from two major threats: agricultural development that has caused a significant loss of the sands' range (Ben David and Avni 2013), and a stabilizing process of the shifting sand resulting from a bio-crust (Kidron and Abeliovich 2009). In the Sinai Peninsula, a lack of shrubs as a result of overgrazing threats the population.	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
5509E63269C3FF275AC39F4B7F99BA28.taxon	biology_ecology	Biology. Adults emerge immediately after the first significant rainfall and inhabit sandy dunes or sand and loess plains and edges of salt lakes. In the spring following an average rainy winter, the species can densely populate the dunes (one observer can locate up to 40 individuals within one hour). Their diet is based mainly on ants and occasionally on other small insects, as well as on dead insects and dead reptiles. Activity is limited by temperature: it begins at a soil temperature of approximetly 18 ° C, and ceases at a soil temperature of approximetly 39 ° C. By moving between sun-exposed microhabitats and the shadow of dwarf-shrubs can prolong the activity period. Strong wind halts activity due to the beetle's sensitivity to dehydration. Some activity also occurs in the afternoon, but it is significantly lower than in the morning peak hours. Prior to commencing inactivity, the beetle digs a short burrow with a narrow elliptic cross-section into the dune's slope. The digging is performed mainly with the hind legs and secondarily with the middle legs. The well-developed, spoon-shaped metatibial spurs (see fig. 4 a in Assmann et al. 2015) seem to function as a shovel. The burrow's opening usually collapses behind the beetle or is covered by shifting sand. In the burrow, a few centimeters below the sand surface, the beetle is relatively protected from predation and can probably still detect the outside temperature and light conditions. In enclosure experiments with individual markings and variation in population density, one of us found that even during the peak activity season, most of the specimens spend most of the days without displaying epigeic activity. An encounter between two individuals of any gender immediately develops into a short, hasty, bite battle and the escape of the loser. In some regions, shade is a limited resource and the battle occurs mainly under bushes and dwarf-shrubs. An encounter between male and female starts with an aggressive fight. The persistent male will then mount the back of the female. His forelegs grasp the female between the basal part of the pronotum and the elytral humeri, while the female tries to grab the male with her hind legs. The copulation lasts for approximately 30 minutes and occurs mostly beneath perennial vegetation. During the fight, the beetles stridulate. This sound is produced when the beetles are threatened by other individuals or by potential predators (Renan unpublished data, based on field observations and arena experiments).	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
C15937F8E4B6AA23B6904D783092F4A5.taxon	diagnosis	Diagnosis. Medium-sized species with 12 - 18 white elytral spots; the anterior and central ones usually elongated, the posterior ones rounded; six marginal extensions, extension II triangular. Median lobe of aedeagus with bent tip.	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
C15937F8E4B6AA23B6904D783092F4A5.taxon	description	Description. BL male: 15.0 - 18.0 mm, average 16.5 + / - 0.8 mm; BL female: 16.0 - 193 mm., average 17.6 + / - 0.8 mm; Head medium; HW / PW: 0.74 mm; EYL: 1.2 - 1.7 mm; EYL / EL: 0.15. Mentum with two teeth and shallow depression between (cf. Fig. 3 b). Frontal ridge slightly developed. In male, apical white frons stripes wider than exposed frons (Fig. 4 b). Pronotum cordiform; PL / PW: 0.66; BPW / PW: 0.66; posteromedially concave and without white margin; white lateral margin as wide as antennomere I long. Elytra oval, humeri rounded; EL: 8.1 - 10.3 mm, average 9.2 mm; EW: 6.2 - 8.8 mm, average 7.8 mm, (EL / EW: 1.3). Lateral cross section flat. Dense black scales, disc not visible between scales (Fig. 6 a). White lateral margin as wide as half antennomere I long and with six extensions; extension I triangular with rounded angels, as wide as or wider than elytra margin, wider and shorter than extension III; extension II smaller and usually shorter than two other ones; extension III often constricted at base. White posterior margin almost continuously rounded, only slightly becoming narrower, gap at suture smaller than lateral margin. Disc with 14, sometimes 12 or 18 spots; most anterior pair of spots slightly elongate to rounded, usually wide as extension I, second anterior pair of spots strongly elongate, nearly two times as long as wide; the two lateral pairs of spots rounded, adjacent or sometimes fused to extension II; the tow to four posterior pairs of spots rounded; the medial, most posterior pair of spots larger than all other spots; the outer most posterior pair of spots much smaller than the latter one. Apical sinuation slightly developed to straight, apex not protuberant, broadly rounded, especially on the medial side (Fig. 7 c). Suture conspicuous. Legs long; MTIL: 4.9 - 6.1 mm, average 5.6 mm; El / MTIL: 1.7. Metatibial secondary spur brown. MTAL: 4.0 - 5.0 mm, average: 4.5 mm; MTAL / MTIL: 0.8. Claws of hind legs black at base. Median lobe of aedeagus with bent tip (Fig. 9 n).	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
C15937F8E4B6AA23B6904D783092F4A5.taxon	etymology	Etymology. The species is dedicated to Sharon Renan, biologist, conservationist, and the first author's wife.	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
C15937F8E4B6AA23B6904D783092F4A5.taxon	distribution	Distribution. Endemic to the Mediterranean coastal plains from north-east Sinai (El Arish) to central Israel south of Nahal Alexander (Fig. 19).	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
C15937F8E4B6AA23B6904D783092F4A5.taxon	conservation	Conservation. The coastal plain sand dunes of Israel form the largest part of the entire distribution range of G. sharonae sp n. As a result of land use changes and urbanization, less than 25 % of the Israel sandy habitats remain and a further decline can be expected. In addition, the remaining dune habitats are under extreme anthropogenic disturbance and highly fragmented (Achiron-Frumkin et al. 2003). The following records are examples of sites that were populated by G. sharonae sp. n. in the past, but where their populations no longer exist: Kefar Bilu, Rehovot, Nes Ziyyona, Bat Yam, Holon, Tel Aviv, Ramat Gan, Bene Beraq, Ra'ananna, Yafo (based on SMNHTAU collection and the authors' experience). Despite having no precise data, the habitats in the Gaza Strip and north-eastern Egypt seem also to have declined as these areas feature a strong increase in human population density. In a faunistic survey of the ground beetles of the Sinai Peninsula, Abdel-Dayem et al. (2004) did not record Graphipterus from El-Arish, where it had been present nearly a century ago (records in London, cf. Schatzmayr 1936). El Surtasi et al. (2012) demonstrated the negative effect of urbanization on G. serrator population in Egypt. Both the restricted distribution range of the endemic species G. sharonae sp. n. and the decline in coastal sandy habitats threaten the long-term survival of the species.	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
C15937F8E4B6AA23B6904D783092F4A5.taxon	biology_ecology	Biology. Seasonal and daily activity time, as well as diet, intraspecific behavior, including copulation and the chirping sounds produced by the stridulatory structure, are as in G. serrator. Graphipterus sharonae sp. n. prefers stabilized sands with high vegetation cover, and its population density is higher than that of G. serrator.	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
FDC2AE3C9B69567D0B58B4197B399078.taxon	diagnosis	Diagnosis. Large species with 16 white rounded to elongated spots on elytra, anterior and posterior pair of spots larger than others; six marginal extensions, extension I triangular, extension I and II elongated. Elytra widest at interior rear third, drop-like shape. Median lobe of aedeagus with slightly bent tip.	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
FDC2AE3C9B69567D0B58B4197B399078.taxon	description	Description. BL male: 17.2 - 20.1 mm, average 18.8 + / - 1 mm; BL female: 18.4 - 19.8 mm, average 18.9 + / - 0.6 mm. Head slender; HW / PW: 0.7; EYL 1.1 - 1.7 mm; EYL / EL: 0.16. Mentum with two teeth (Fig. 3 b). Frontal ridge slightly developed. Male, apical white frons stripes wider than exposed frons (Fig. 4 b). Pronotum strongly cordiform; PL / PW: 0.66. BPW / PW: 0.64; posteromedially concave and without white margin; white lateral margin as wide as antennomere I long. Elytra droplet-like, humeri strongly narrowed; EL: 9.1 - 11.1 mm, average 8.4 mm; EW: 7.5 - 9.0 mm, average 8.4 mm; EL / EW: 1.1 - 1.5. Lateral cross section convex. Dense black scales, disc not visible between them (Fig. 6 a). White lateral margin nearly as half wide as antennomere I long and with six extensions; extension I triangular with rounded angels, as wide as lateral margin, posteriorly oriented; extension II small, often constricted at base, as wide as lateral margin; extension III large, elongated, posteriorly oriented. White posterior margin becomes narrower towards suture; gap at suture wider than lateral margin. Disc usually with 16 spots; anterior pair of spots elongate, as wide as extension I; posterior pair of spots rounded and larger than other ones; six spots forming arch pattern anterior and lateral to posterior rounded larger spots. Apical sinuation slightly developed to straight, apex not protuberant, broadly rounded, especially on the median side (Fig. 7 c). Suture inconspicuous. Legs long; MTIL: 6.0 - 7.0 mm, average 6.5 mm; El / MTIL: 1.6. Metatibial secondary spur black. MTAL: 4.7 - 5.3 mm, average 4.9 mm; MTAL / MTIL: 0.8. Claws of hind legs black at base. Median lobe of aedeagus with bent tip (Fig. 9 o).	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
FDC2AE3C9B69567D0B58B4197B399078.taxon	etymology	Etymology. The name is derived from ancient Greek (σταγών, oψις) and means " drop-like " which refers to the shape of the elytra.	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
FDC2AE3C9B69567D0B58B4197B399078.taxon	distribution	Distribution. Central and west Algeria (Fig. 18).	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
FDC2AE3C9B69567D0B58B4197B399078.taxon	conservation	Conservation. Unknown.	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
0EB3AD64F86514D39EC5E5B67B2DF35C.taxon	diagnosis	Diagnosis. Large species with 10 - 16 white round spots on elytra; anterior and posterior discal spots larger than other spots; four white marginal extensions, oval elytra, extension I triangular. Median lobe of aedeagus with bent tip.	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
0EB3AD64F86514D39EC5E5B67B2DF35C.taxon	description	Description. BL male: 14.8 - 19.0 mm, average 17.1 + / - 1.7 mm; BL female: 18.6 - 20.5 mm, average 18.6 + / - 1.9 mm. Head slender; HW / PW: 0.71; EYL: 1.4 - 1.8 mm; EYL / EL: 0.15. Mentum with mentum with two teeth as margin between them slightly convex in middle (Fig. 3 c). Frontal ridge slightly developed. In male, apical white frons stripes slenderer than exposed frons (Fig. 4 a). Pronotum cordiform; PL / PW: 0.64; BPW / PW: 0.7; posteromedially concave and without white margin; white lateral margin as wide as antennomere I long. Elytra oval, relatively elongated, humeri rounded; EL: 8.1 - 12 mm, average 10.6 mm; EW: 6.5 - 9.1 mm, average 8.0 mm; EL / EW: 1.3. Lateral cross section convex. Dense black scales, disc not seen between scales (Fig. 6 a). White lateral margin as wide as half antennomere I long and with four extensions; extension I triangular with rounded angels, much wider than lateral margin, but shorter than extensionII; the latter one elongated, positioned at second third of elytra. Apical sinuation strongly sinuated, apex strongly protruded, forming almost a rectangular. White posterior margin evenly rounded, not becoming narrower towards the suture; gap at suture wider than lateral margin. Disc with (10 -) 14 (- 16) spots; anterior pair of spots rounded to slightly elongate, much smaller than extension I; anterior and posterior pair of spots round, larger than other spots; small additional spots located medially to extension II and laterally to posterior spots. Apical sinuation strongly developed, apex protruded, almost rectangular, only slightly rounded at most distant tip (Fig. 7 a). Suture inconspicuous. Legs long; MTIL: 4.9 - 7.0 mm, average 6.3 mm; El / MTIL: 1.7. Metatibial secondary spur dark. MTAL: 5.4 - 6.9 mm, average 6.0 mm; MTAL / MTIL: 0.72. Claws of hind legs brown at base. Median lobe of aedeagus with bent tip (Fig. 9 p).	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
0EB3AD64F86514D39EC5E5B67B2DF35C.taxon	distribution	Distribution. The arid and semi-arid regions of north-east Algeria from Ghardaia, to Bou-Saada and Tebessa (Fig. 17).	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
0EB3AD64F86514D39EC5E5B67B2DF35C.taxon	conservation	Conservation. Unknown.	en	Renan, Ittai, Assmann, Thorsten, Freidberg, Amnon (2018): Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species. ZooKeys 753: 23-82, DOI: http://dx.doi.org/10.3897/zookeys.753.22366, URL: http://dx.doi.org/10.3897/zookeys.753.22366
