identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
E84D87D9A002FF94B38A5C460E268154.text	E84D87D9A002FF94B38A5C460E268154.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eutarsopolipus Berlese 1913	<div><p>Genus Eutarsopolipus Berlese, 1913</p><p>Type species: Tarsopolipus lagenaeformis Berlese, 1911, by original designation.</p><p>Diagnosis. Female. Three pairs of well-developed legs. Prodorsal plate present, plates C, D, EF usually present, sometimes divided, rarely absent, plate H usually greatly reduced or absent; setae v1, sc1, sc2, c1, c2, d, f present, setae e absent, setae h present or absent. Pseudolegs usually absent. Respiratory system present or absent. Gnathosoma ovoid, with two pairs of setae. Pretarsi I–III with well-developed ambulacrum, claws present or absent. Tibia II–III without spine-like setae. Male. Three pairs of legs. Genital capsule terminal, elongate or trapezoidal. Prodorsal setae developed to vestigial. Plate C-D-EF with three pairs of setae. Ventral idiosomal setae present. Pretarsi I–III with well-developed ambulacrum, claws present or absent. Femora I–III setation (0-3)-0-0, genua I–III setation (0-2)-(0-1)-(0-1), tibia I with or without seta k. Larva: Plate EF present; plate D sometimes in deep concavity of plate C; plate H present or absent, not on terminal capsule-like segment; ventral idiosomal setae present; pretarsi I–III with well-developed ambulacra, claws present or absent. Larval male: absent. Migrating life stage: probably larva. Host insect family: Carabidae .</p></div>	https://treatment.plazi.org/id/E84D87D9A002FF94B38A5C460E268154	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Seeman, Owen D.	Seeman, Owen D. (2019): Two new species of Australian Eutarsopolipus (Acariformes: Podapolipidae) from Nurus medius (Coleoptera: Carabidae). Zootaxa 4647 (1): 134-153, DOI: 10.11646/zootaxa.4647.1.12
E84D87D9A002FF9DB38A5AA0096D858E.text	E84D87D9A002FF9DB38A5AA0096D858E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eutarsopolipus burwelli Seeman 2019	<div><p>Eutarsopolipus burwelli sp. nov.</p><p>Diagnosis. All life stages: respiratory system absent; tarsus II without solenidion; femur I with three setae; genu I with two small, spine-like setae (l′, l′′ present), genua II–III with one seta (l′ present); claws present on legs I–III. Adult female: gnathosomal length 66–70, width 62–67, cheliceral stylets long, 75–95; idiosomal setae d 10–14, f 8–13, h 5–6; femur I seta l′ 2–4; genu I setae 2–3; tarsi II–III with seta pv′ absent. Larviform adult male: dorsal setae v1, sc1, sc2, c1, c2, d, f developed; leg setation same as female. Larval female: plates C and D separate; seta h2 short, 11–13.</p><p>Type material. 18 females, 5 males, 15 larvae, all ex Nurus medius, as follows. Australia: Queensland. Holotype female, Eungella NP, Mt William, Site 1, 1234 masl, 21.016°S 148.598°E, 14 Nov. 2013 , SC 25844, C. Burwell (QMS 110124). Paratypes : 6 females, 3 larvae, same data as holotype (QMS 110125 -33); 1 female, 1 male, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=148.54251&amp;materialsCitation.latitude=-28.307777" title="Search Plazi for locations around (long 148.54251/lat -28.307777)">Crediton</a> SF, 28°18′28″S 148°32′33″E, 1 Dec–31 Dec 1993 , IN9787, ANZSES, HR# T110024 (QMS 110134 -35); 7 females, 9 larvae, Eungella NP, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=148.49167&amp;materialsCitation.latitude=-21.136667" title="Search Plazi for locations around (long 148.49167/lat -21.136667)">Dalrymple Heights</a>, 21° 8′12″S 148°29′30″E, IN7864, 6 Jan. 1973 , J. Hammond, HR# T82062 (QMS 110136 -45; 1 female ZMH-A0002396, 1 larva ZMH-A0002398; 1 female ANIC 52-003930, 1 larva ANIC 52-003931); 2 females, 1 male, Eungella NP, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=148.60306&amp;materialsCitation.latitude=-21.028055" title="Search Plazi for locations around (long 148.60306/lat -21.028055)">Upper Cattle Creek</a>, 21°1′41″S 148°36′11″E, 17 Nov. 1992 , IN6186, G. Monteith, G. Thomson, H. Janetzki, HR# T82055 (QMS 110146 -48); 1 female, 1 larva, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=148.59917&amp;materialsCitation.latitude=-21.018055" title="Search Plazi for locations around (long 148.59917/lat -21.018055)">Mount William</a>, 21° 1′ 5″S 148°35′57″E, 21 Dec. 1992 – 10 Jan. 1993 , IN5952, ANZSES, HR# T82066 (QMS 110149 -50); 3 males, 1 larva, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=148.55194&amp;materialsCitation.latitude=-20.8325" title="Search Plazi for locations around (long 148.55194/lat -20.8325)">Mt Macartney</a>, 20°49′57″S 148°33′7″E, 19 Nov. 1992 – 15 Apr. 1993 , IN5936, HR# T82039 (QMS 110151 -52; 1 male ANIC 52-003932; 1 male ZMH-A0002397) .</p><p>Type deposition. Holotype and most paratypes deposited in QM. One female, one male, one larva deposited in ANIC, one female, one male, one larva deposited in ZMH.</p><p>Description. Female (Figs 1–3, n = 18). Gnathosoma . Length 70 (66–70), width 67 (62–67). Palp length 16 (15–16). Cheliceral stylets 90 (75–95), pharynx width 21 (15–21), dorsal gnathosomal setae (ch) 36 (30–37), ventral setae (su) 18 (15–18), distance between ventral setae (su–su) 21 (18–23).</p><p>Idiosoma . Moderately physogastric, ovate. Length 325 (310–450), width 290 (270–370). Respiratory system absent. Prodorsal plate length 88 (83–99), with setae v1 13 (9–13), v2 vestigial, sc1 14 (12–15), sc2 95 (95–120); v2 anteromesad sc1. Distance between setae v1–v1 57 (52–58), sc1–sc1 108 (105–110), v1 –sc1 33 (28–34), v2–v2 68 (65–69), sc2–sc2 155 (125–155), sc1–sc2 42 (35–44). Plate C length 67 (63–70), setae c1 15 (15–17), setae c2 15 (11–16), distance between setae c1–c1 105 (88–110), c1–c2 88 (78–93); often with pair of pore-like structures laterad setae c1. Plate D length 64 (57–64), setae d 13 (10–14), d–d 165 (160–180), cupuli ia anteriad to anterolaterad setae d. Plate EF length 65 (60–65), setae f 13 (8–13), f–f 145 (130–145); cupuli im anteriad to anterolaterad setae f. Plate H length 30 (17–30), width 94 (94–115), setae h 5 (5–6). Venter: coxal setae short, usually simple (anomalous star-like form in holotype specimen), 1a 3 (2–3), 2a 5 (4–6), 3a 7 (6–7), 3b 6 (6–8). Alveoli 1b, 2b apparent. Distance between setae 1a–1a 40 (40–48), 2a–2a 58 (58–66).</p><p>Legs. Setal counts legs I–III, femur-tarsus: 3-2-6(+ φ)-9(+ ω), 0-1-4-7, 0-1-4-7. Leg I: femur I, d minute, l′ 4 (2–4), v′′ 17 (13–17); genu I, l′ 3 (2–3), l′′ 3 (2–3); tibia I, d 82 (70–82), l′ 6 (5–8), l′′ 11 (8–11), k 6 (4–6), v′ 8 (5–8), v′′ 10 (10–14), φ 10 (10); tarsus I, tc′ 17 (14–17), tc′′ 16 (15–17), pl′ 9 (8–11), pl′′ 16 (16–20), pv′ 3 (3), pv′′ 3 (4), s 7 (6–7), ω 3 (3), p′ 2 (2), u′′ 1 (1–2). Leg II: genu II, l′ 4 (3–4); tibia II, d 60 (50–60), l′ 8 (7–8), v′ 14 (12–15), v′′ 25 (23–29); tarsus II, tc′ 6 (5–6), tc′′ 65 (55–65), pl′ 3 (3), pv′′ 3 (3–4), u′ 8 (7–8), u′′ 1 (1–2), p′ 1 (1). Leg III: genu III, l′ 3 (3–4); tibia III, d 47 (45–50), l′ 7 (7–10), v′ 15 (10–15), v′′ 21 (17–26); tarsus III, tc′ 6 (5–6), tc′′ 60 (50–60), pl′ 3 (3–4), pv′′ 3 (3–4), u′ 8 (7–8), u′′ 1 (1), p′ 1 (1). Genu I, l′-l″ blunt, peg-like; tarsus I s spine–like. Tarsi II–III u′ spine-like, bifid, tc′, pv′, pv″ spine-like. Claws on legs I–III (1-2-2) well developed.</p><p>Larviform adult male (Fig. 4; n = 5; square brackets indicate two male specimens with different measurements)</p><p>Gnathosoma . Length 30–33 [37–38], width 29–32 [34–35]. Palp length 10–14. Cheliceral stylets 22–26, pharynx width 6–9, ch 8–11 [13], su 6–9, su–su 13–14.</p><p>Idiosoma . Length 150–175, width 120–145. Prodorsal plate length 52–57 [59–63], width 86–97 [110–115], with setae v1 5–7, v2 vestigial, sc1 4–6, sc2 65–80; v2 anteromesad sc1. Distance between setae v1–v 1 22–26 [27–29], sc1–sc1 45–49 [52–57], v1 –sc 1 15–19 [20–21], v2–v 2 26–29 [30–34], sc2–sc2 53–63 [67], sc1–sc 2 20–22 [25–28]. Plate CD length 45–50 [58–65], width 105–115 [140–145], setae c1 4–7, c2 5–9, d 4–7; distance between setae c1–c1 38–45 [51–56], c1–c 2 28–33 [38–40], d–d 33–36 [44]; cupuli ia anterolaterad setae d. Plate EF bowed, posterior margin concave, length 24–28, width 40–43 [55–56], setae f 2–4, distance f–f 23–27 [35–37]; cupuli im anterolaterad setae f. Genital capsule length 18–26, width 27–28 [33–37], setae h minute. Venter: coxal setae 1a 2–3, 2a 3–4, 3a 3–5, 3b 4–5, alveoli 1b, 2b apparent. Distance between setae 1a–1a 21–24, 2a–2a 23–25 [28–36].</p><p>Legs. Setal counts legs I–III same as female; setal form similar. Leg I: femur I, d minute, l′ 2, v′′ 5–6; genu I, l′, l′′ 1–2; tibia I, d 40–45, l′ 2 [3–4], l′′ 3–4, v′ 2–3, v′′ 7–9, φ 7 [9–10], k 2–3; tarsus I, tc′ 12–14, tc′′ 13–14, pl′ 5–7, pl′′ 13–14, pv′ 1–2, pv′′ 1–2, s 4, p′ 1, u′′ 1, ω 2. Leg II: genu II, l′ 2–3; tibia II, d 25–27, l′ 5–8, v′ 10–14, v′′ 15–17; tarsus II, tc′ 3–5, tc′′ 24–26, pl′ 2–3, pv′′ 1–3, u′ 4–5, u′′ 1. Leg III: genu III, l′ 2; tibia III, d 18–20, l′ 5–6, v′ 8–11, v′′ 14–15; tarsus III, tc′ 3–4, tc′′ 25–27, pl′ 2–3, pv′′ 2–3, u′ 4–6, u′′ 1.</p><p>Larval female (Fig. 5; n = 15). Gnathosoma . Length 55–61, width 52–56. Palp length 15–16. Cheliceral stylets 65–80, pharynx width 12–15, ch 30–38, su 8–12, su–su 15–20.</p><p>Idiosoma . Length 225–280, width 170–245. Prodorsal plate length 76–87, width 135–160, with setae v1 8– 11, v2 vestigial, sc 1 7–10, sc2 130–140; v2 anterolaterad sc1. Distance between setae v1–v1 35 –40, sc1–sc1 72– 78, v1 –sc 1 23–27, v2–v2 44 –47, sc2–sc2 86–98, sc1–sc2 35–45. Plate C length 57–58, setae c 1 12–17, setae c 2 8–12; often with pair of pore-like structures laterad setae c1. Plate D length 48–50, width 105–120, setae d 8–11, distance between setae c1–c1 56–68, c1–c2 47–57, d–d 67–77; cupuli ia anterolateral setae d, distance ia–d 2–4. Plate EF length 40–47, width 85–100, setae f 9–11, distance f–f 54–61; cupuli im anteriad setae f, distance im–f 4–5. Plate H length 29–36, width 21–23, seta h1 long, ca. 180, seta h2 short, 11–13. Venter: coxal setae all short, 1a 3–4, 2a 4–5, 3a 4–6, 3b 6–7. Distance between setae 1a–1a 28–31, 2a–2a 35–39.</p><p>Legs. Setal counts legs I–III same as adult female; setal form similar. Leg I: femur I, d minute, l′ 2–3, v′′ 10–12; genu I, l′ 2, l′′ 2–3; tibia I, d 75–85, l′ 11–13, l′′ 12–15, v′ 4–5, v′′ 12–15, φ 10–12, k 4–5; tarsus I, tc′ 15–17, tc′′ 15–18, pl′ 7–8, pl′′ 19–24, pv′ 1–2, pv′′ 1–2, s 5–6, p′ minute, u′′ 1–2, ω 2–3. Leg II: genu II, l′ 1–3; tibia II, d 40–45, l′ 8–9, v′ 14–15, v′′ 19–21; tarsus II, tc′ 3–5, tc′′ 25–36, pl′ 2–4, pv″ 2–3, u′ 5–6, u′′ 1. Leg III: genu III, l′ 2–3; tibia III, d 45–50, l′ 7–8, v′ 15–17, v′′ 16–20; tarsus III, tc′ 3–5, tc′′ 25–35, pl′ 2–4, pv′′ 1–2, u′ 5–6, u′′ minute–1.</p><p>Etymology. It is with pleasure that I name this species for my colleague Dr Chris Burwell, the collector of several host beetles for Australian mites, including this species.</p><p>Differential diagnosis. Eutarsopolipus burwelli is a member of the ochoai species group (Husband 1995; Constantine and Seeman 2005). Within this group, this species is very similar to E. rutherfordae in having females with a reduced plate H, females and males lacking setae pv′ and tiny, spine-like genual setae l′ and l″. Females and larvae of each species are easily distinguished, but males and larvae are almost identical. Females are most easily separated by the size of the gnathosoma (66–70 × 62–67 in E. burwelli versus 53 × 53 in E. rutherfordae), cheliceral stylets (75–95 in E. burwelli versus 60 in E. rutherfordae), size of setae d and f (8–14 in E. burwelli versus 22–25 in E. rutherfordae), and the size of tibia I seta l′ (5–8 in E. burwelli versus 13 in E. rutherfordae). The larger gnathosoma of E. burwelli also presents in the larva (55–61 × 52–56 in E. burwelli versus 47 × 47 in E. rutherfordae), as do the longer cheliceral stylets (65–80 in E. burwelli versus 48 in E. rutherfordae).</p><p>Remarks. Two males had several measurements somewhat larger than the other specimens. Although these seem different, size variation in male podapolipid mites is a known phenomenon and extreme variation was recorded for Eutarsopolipus pungens Husband and Dastych, 1998 . This variation makes diagnostics based on males difficult. Similar variation in larvae and females has not been reported.</p><p>Constantine and Seeman (2014) described three new species of Eutarsopolipus that they placed in the ochoai species group. This decision was based on the original definition of the ochoai species group (Husband 1995) and the key to species groups in Husband and Husband (2009). Since Husband (1995), the ochoai species group has obtained nine members as defined by Constantine and Seeman (2014), but its definition has become less robust and overlaps with the brettae species group, as proposed by Husband (2002).</p><p>Species in the ochoai group may lack a respiratory system ( E. earnshawi, E. lambkinae, E. rutherfordae; E. scariteus Husband, 2001; E. weatherbyi Husband and Psalmonds, 2004), show only the rudiments of stigmata ( E. ochoai Husband, 1995), or have a well-developed respiratory system ( E. dastychi Husband &amp; Khaustov 2004; E. leytei Husband &amp; Corpuz-Raros, 1995). Species of the brettae species group, which comprises only E. brettae Husband, 2002 and E. obrieni Husband and Husband, 2015, both lack a respiratory system. All these species retain claws on legs I–III, three setae on femur I, and at least one seta on genua I–III.</p><p>The presence/absence of a respiratory system is an important character in Eutarsopolipus . Several species groups are defined by this feature, as shown in the key of Husband and Husband (2009), which has an error in its first couplet (the first couplet needs to be reversed). Therefore, it is unhelpful if this character is not used consistently to define species groups. For instance, E. scariteus —a member of the ochoai species group—will key to the brettae species group in Husband and Husband (2009). Furthermore, the rudimentary stigmata of E. ochoa need to be reassessed as they occur in an atypical position at the anterior margin of the prodorsal shield (and posteromesad the gnathosoma), so these may be another structure, as the stigmata are usually posterolaterad the gnathosoma in Eutarsopolipus .</p><p>A reformation of species groups based on phylogenetic analysis is needed, but in the interim it would seem best to define the ochoai species group in the manner originally intended, so that it includes only species with the respiratory system absent, retaining setae on all genua, and possessing claws on all legs. This definition includes the two species from the brettae species group ( E. brettae, E. obrieni) but excludes E. dastychi and E. leytei . The latter two species could be accommodated in the leytei species group, defined by the retention of a respiratory system, presence of claws on all legs, three femur I setae and presence of setae on all genua. Within the ochoai and leytei species groups, some reductions are apparent. Some species lack setae l″ on genu I ( E. brettae, E. lambkinae, E. obrieni) and males of some species have hypotrichous leg setation ( E. brettae, E. dastychi, E. lambkinae, E. ochoai, E. scariteus). While the former character is homoplasious, the latter is found only in members of the ochoai and leytei species groups, and in the unusual species Eutarsopolipus stammeri Regenfuss, 1968, which has its own species group.</p></div>	https://treatment.plazi.org/id/E84D87D9A002FF9DB38A5AA0096D858E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Seeman, Owen D.	Seeman, Owen D. (2019): Two new species of Australian Eutarsopolipus (Acariformes: Podapolipidae) from Nurus medius (Coleoptera: Carabidae). Zootaxa 4647 (1): 134-153, DOI: 10.11646/zootaxa.4647.1.12
E84D87D9A00BFF9BB38A5EDE0EF085AA.text	E84D87D9A00BFF9BB38A5EDE0EF085AA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eutarsopolipus echinatus Seeman 2019	<div><p>Eutarsopolipus echinatus sp. nov.</p><p>Diagnosis. All life stages: respiratory system absent; tarsus II without solenidion; femur I with two setae (v″ absent); genu I–III lacking setae; claws absent on legs I–III; setae ch, v1, sc1, c1, c2, d, f short, thorn-like. Adult female: gnathosomal length 44–46, width 43–46, cheliceral stylets long, 44–50; seta h absent; tarsi II–III with setae u′ and p′ in proximal position, seta pv′ absent. Larviform adult male: setae v1, sc1, c1, c2, f short, length 2–3; leg setation same as female. Larval female: plates C and D separate; seta h2 long, 33–44.</p><p>Type material: 4 females, 1 male, 5 larvae, all ex Nurus medius, as follows. Australia: Queensland. Holotype female, Eungella NP, Mt William, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=148.598&amp;materialsCitation.latitude=-21.016" title="Search Plazi for locations around (long 148.598/lat -21.016)">Site</a> 1, 1234 masl, 21.016°S 148.598°E, 14 Nov. 2013, SC 25844, C. Burwell (QMS 110153, on same slide as paratype larva of E. burwelli QMS 110133). Paratypes : 1 female, 1 male, 1 larva, same data as holotype (QMS 110154 -56); 1 female, Broken River, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=148.50833&amp;materialsCitation.latitude=-21.168055" title="Search Plazi for locations around (long 148.50833/lat -21.168055)">Eungella</a> NP, 21° 10′ 5″S 148°30′30″E, 17 Nov. 1992 – 15 Apr. 1993, IN7677, G. Monteith, D. Cook, HR# T82045 (ANIC 52-003933) ; 1 female, Eungella NP, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=148.49167&amp;materialsCitation.latitude=-21.136667" title="Search Plazi for locations around (long 148.49167/lat -21.136667)">Dalrymple Heights</a>, 21° 8′12″S 148°29′30″E, IN7864, 6 Jan. 1973, J. Hammond, HR# T82062 (QMS 110157) ; 1 female, 4 larvae, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=148.55194&amp;materialsCitation.latitude=-20.8325" title="Search Plazi for locations around (long 148.55194/lat -20.8325)">Mt Macartney</a>, 20°49′57″S 148°33′7″E, 19 Nov. 1992 – 15 Apr. 1993, IN5936, HR#T82035 (1 female, 1 larva ZMH-A0002399, ZMH-A0002400, 3 larvae QMS 110158 -60); 1 larva, Mt Macartney, same data except, HR# T82039 (ANIC 52-003934) .</p><p>Type deposition. Holotype and most paratypes deposited in QM. One female, one larva deposited in ANIC, one female, one larva deposited in ZMH.</p><p>Description. Female (Figs 6–8, n = 4). Gnathosoma . Length 45 (44–46), width 46 (43–46). Palp length 14 (13–15). Cheliceral stylets 45 (44–50), pharynx width 13 (11–13), dorsal gnathosomal setae (ch) thorn-like, 2 (2–4), ventral setae (su) 7 (7–10), distance between ventral setae (su–su) 17 (17–19).</p><p>Idiosoma . Body not to slightly physogastric, ovate. Length 285 (215–305), width 210 (185–275). Respiratory system absent. Dorsal setae v1, sc1, c1, c2, d, f thorn-like. Prodorsal plate length 71 (65–75), with setae v1 6 (6–7), v2 vestigial, sc1 7 (7–8), sc2 32 (31–35); v2 anteromesad sc1. Distance between setae v1–v1 30 (28–35), sc1–sc1 79 (79–86), v1 –sc1 37 (37–40), v2–v2 40 (39–44), sc2–sc2 89 (89–111), sc1–sc2 10 (10–12). Plates C, D, EF, H variously eroded medially, making widths variable. Plate C length 43 (38–43), setae c1 6 (5–6), setae c2 4 (4–5), distance between setae c1–c1 73 (73–94), c1–c2 54 (52–62). Plate D length 24 (24–40), setae d 4 (4–6), d–d 91 (84–93), cupuli ia anteromesad setae d. Plate EF length 30 (26–33), setae f 3 (2–3), f–f 66 (66–85); cupuli im anterolaterad setae f. Plate H length 11 (11–15), width 61 (55–62), setae h absent. Venter: small, thorn-like, 1a minute (minute–1), 2a 3 (2–3), 3a 3 (2–3), 3b 3 (2–3). Alveoli 1b, 2b apparent. Distance between setae 1a–1a 29 (24–28), 2a–2a 41 (30–41).</p><p>Legs. Setal counts legs I–III, femur-tarsus: 2-0-5(+ φ)-9(+ ω), 0-0-4-7, 0-0-4-7. Leg I: femur I, d minute, l′ 3 (2–3); tibia I, d 28 (25–28), l′ 5 (4–5), l′′ 1 (1–2), v′ 5 (4–5), v′′ 4 (4–5), φ 7 (7–9); tarsus I, tc′ 10 (9–10), tc′′ 9 (9–10), pl′ 7 (7–10), pl′′ 12 (10–12), pv′ 3 (3–4), pv′′ 3 (3–4), s 6 (6–7), ω 2 (2), p′ 1 (1), u′′ 2 (2–3). Leg II: tibia II, d 8 (8–10), l′ 4 (4–5), v′ 4 (4–5), v′′ 8 (8–12); tarsus II, tc′ 5 (5), tc′′ 25 (23–26), pl′ 17 (17–19), pv′′ 3 (3), u′ 5 (5), u′′ 1 (1–2), p′ 1 (1–2). Leg III: tibia III, d 7 (7–10), l′ 4 (3–4), v′ 4 (4–5), v′′ 7 (7–10); tarsus III, tc′ 5 (5), tc′′ 32 (26–35), pl′ 17 (17–21), pv′′ 2 (2–3), u′ 5 (5), u′′ 1 (1–2), p′ 3 (2–3). Tibia I seta v′ spine-like; tarsus I setae pv′, pv″ s spine–like. Tibia II–III, seta l′ spine-like; tarsi II–III setae u′ spine-like, setae p′, pv″ spine-like. Claws on legs I-III absent.</p><p>Larviform adult male (Fig. 7; n = 1). Gnathosoma . Length 33, width 35. Palp length 10. Cheliceral stylets 25, pharynx width 9, ch 2, su 5, su–su 14.</p><p>Idiosoma . Length 180, width 155. Dorsal setae v1, sc1, c1, c2, d, f small, thickened to thorn-like. Prodorsal plate length 61, width 130, with setae v1 2, v2 vestigial, sc1 2, sc2 16; v2 anteromesad sc1. Distance between setae v1–v1 16, sc1–sc1 65, v1 –sc 1 28, v2–v2 23, sc2–sc2 69, sc1–sc2 12. Plate CD length 54, width 140, setae c1 3, c2 3, d 2; distance between setae c1–c1 52, c1–c2 36, d–d 39; cupuli ia anteriad setae d. Plate EF divided, each plate length 17, width 37, setae f 2, distance f–f 23; cupuli im anterolaterad setae f. Genital capsule length 23, width 37, setae h minute. Venter: coxal setae minute, alveoli 1b, 2b apparent. Distance between setae 1a–1a 22, 2a–2a 24.</p><p>Legs. Setal counts for legs I–III same as female; setal form similar. Leg I: femur I, d minute, l′ 2; tibia I, d 25, l′ 2, l′′ 2, v′ 3, v′′ 5, φ 5, tarsus I, tc′ 11, tc′′ 11, pl′ 5, pl′′ 9, pv′ 3, pv′′ 2, s 3, p′ minute, u′′ 1, ω 2, left tarsus I with additional thorn-like seta between (pv). Leg II: tibia II, d 5, l′ 4, v′ 5, v′′ 9; tarsus II, tc′ 5, tc′′ 16 (possibly broken), pl′ 16, pv′′ 2, u′ 4, p′ 2, u′′ minute. Leg III: tibia III, d 4, l′ 3, v′ 5, v′′ 10; tarsus III, tc′ 5, tc′′ 24, pl′ 16, pv′′ 2, u′ 5, p′ 2, u′′ minute.</p><p>Larval female (Fig. 10; n = 5)</p><p>Gnathosoma . Length 38–41, width 39–51. Palp length 11–14. Cheliceral stylets 41–46, pharynx width 10–12, ch 1–2, su 4–5, su–su 14–17.</p><p>Idiosoma . Length 175–215, width 135–240. Prodorsal plate length 64–69, width 115–135, with setae v1 4–5, v2 vestigial, sc1 4–5, sc2 70–85; v2 anteromesad sc1. Distance between setae v1–v 1 23–26, sc1–sc1 60– 69, v1 –sc 1 31–36, v2–v 2 25–30, sc2–sc2 73–81, sc1–sc 2 15–19. Plate C length 36–43, setae c1 4–5, setae c2 4–5. Plate D length 37–42, width 65–70, setae d 3–4, distance between setae c1–c1 51–56, c1–c2 49–57, d–d 37–41; cupuli ia anteriad setae d, distance ia–d 4–6. Plate EF length 32–40, width 55–65, setae f 2–3, distance f–f 38–41; cupuli im anteriad setae f, distance im–f 5–7. Plate H length 25–35, width 34–40, seta h1 long, ca. 120, seta h2 long, 33–44. Venter: coxal setae minute–1, distance between setae 1a–1a 25–29, 2a–2a 25–30.</p><p>Legs. Setal counts for legs I–III same as female; setal form similar. Leg I: femur I, d minute, l′ 2; tibia I, d 40–47, l′ 5–6, l′′ 2–3, v′ 3, v′′ 8–10, φ 6–7; tarsus I, tc′ 10–11, tc′′ 11–12, pl′ 8–10, pl′′ 15–16, pv′ 3, pv′′ 2, s 4–5, p′ 2, u′′ 2, ω 3. Leg II: tibia II, d 7–9, l′ 3–4, v′ 6–8, v′′ 10–15; tarsus II, tc′ 4–5, tc′′ 31–41, pl′ 17–20, pv″ 2, p′ 3, u′ 4, u′′ 2. Leg III: tibia III, d 7–8, l′ 3–4, v′ 6–7, v′′ 10–13; tarsus III, tc′ 5, tc′′ 26–34, pl′ 16–20, pv′′ 2–3, p′ 3, u′ 4–5, u′′ 2.</p><p>Etymology. The name echinatus (Latin: prickly, spiny; gender masculine) refers to the spine-like setae on the dorsum.</p><p>Differential diagnosis. The short, thorn-like setae are unlike any other species of Eutarsopolipus . This species fits the concept of the pterostichi species group, which is a heterogeneous collection of eight species that requires revision. Members of this group are generally regarded as species that lack both a respiratory system and setae on genu II–III. However, it includes species with or without ambulacral claws and variable femur I and genu I setation, suggesting either homoplasious morphological reductions within this group or that it is polyphyletic. In lacking ambulacral claws on all legs and all genual setae, E. echinatus sp. nov. is similar to E. inermis Regenfuss, 1974 and E. osunaharae Husband and Kurosa, 2012, which are the only other species of the pterostichi group that share these character states. These species also share the loss of seta v″ on femur I and seta k on tibia I. In addition to the absence of spine-like setae, E. inermis and E. osunaharae also differ in retaining a solenidion on tarsus II and having larvae with fused plates C and D.</p><p>Remarks. The species was found cohabiting with E. burwelli on the type host beetle and beetles T82062 and T82039.</p><p>The notations for the p′ setae on tarsus II–III are tentative. Species of Eutarsopolipus may present a minute to small, distal seta on the anterior side of the tarsus. When this seta is proximal to the insertion of the spur-like seta u″, I have interpreted it as the retention of seta pv′, as in E. earnshawi (Constantine &amp; Seeman 2013) . When this seta is distal to the insertion of u″, I have designated the seta as p′. In E. echinatus sp. nov., seta u′ is in an unusual proximal position, so according to my interpretation, I have designated the small spine-like anteroventral seta as p′, which is assumed to have migrated ventrally with u′.</p></div>	https://treatment.plazi.org/id/E84D87D9A00BFF9BB38A5EDE0EF085AA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Seeman, Owen D.	Seeman, Owen D. (2019): Two new species of Australian Eutarsopolipus (Acariformes: Podapolipidae) from Nurus medius (Coleoptera: Carabidae). Zootaxa 4647 (1): 134-153, DOI: 10.11646/zootaxa.4647.1.12
