identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
EB024F5EFFFFFFB7FF22F8F1FCD2FBF7.text	EB024F5EFFFFFFB7FF22F8F1FCD2FBF7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Miroculis (Miroculis) boldrinii Raimundi & Domínguez & Salles 2021	<div><p>Miroculis (Miroculis) boldrinii sp. nov.</p><p>Figures 1–5; 6</p><p>Diagnosis. Miroculis (Miroculis) boldrinii sp. nov. differs from the other species of Miroculis by the following combination of characters. Male imago: 1) head orange strongly tinged with dark brown; 2) dorsal portion of compound eye on a short stalk, with the inner margin presenting a medial projection (Fig. 1) and dorsal surface with 20 facets on the longest row; 3) fore wing hyaline except for three wide dark brown transversal bands; 4) forceps segment II 0.24× forceps segment I; 5) forceps segment I with subdistomedial protuberance; and 6) forceps brown with inner margin paler on segments II and III.</p><p>Description. Male imago—(in alcohol) Fig. 6.</p><p>MEASURES (mm). Body: 6.0. Fore wing: 4.7; angularity of cubital region 94°; hind wing: 1.4. Fore and middle legs: broken off and missing; hind leg: 3.5. Penis: 0.8. Caudal filament: broken off and missing. RATIOS. Wings: Fore wing width 0.5× fore wing length; hind wing length 0.3× fore wing length; hind wing width 0.6 hind wing length; fork on MA 2 of fore wing 0.5× total length of MA. Legs: segments of hind leg: 0.92: 1.00 (1.41 mm): 0.07: 0.06: 0.06: 0.08. Genitalia: medial length of styliger plate 0.6× maximum width of styliger plate; lateral length of styliger plate 0.7× medial length of styliger plate; lateral length of styliger plate 1.1× lateral length of segment IX. Forceps segment III 0.7× segment II; segment III 0.3× segment I; segment II 0.4× segment I; penis length 1.5× forceps segment I. COLORATION: General coloration: orangish brown. Head: Orangish and strongly tinged with dark brown. Stalk of compound eye black; upper portion with facets separated by dark grooves. Ocellus white surrounded by gray. Antenna brownish. Thorax. Sclerites yellowish brown. Pleura pale; pleural sclerites tinged with dark gray. Ventral surface of cervix with a posteromedial dark gray spot. Pro, meso and metanotum tinged with brown, margins of sclerites darker. Wings (Figs 2–3): Fore wing hyaline marked with two evident wide brownish transversal bands (Fig. 2). Hind wing hyaline, brownish on apical ⅕ and around cross veins (Fig. 3). Legs. Hind leg whitish. Coxa and femur tinged with light brown; femur with dark brown complete band on median and subapical portions. Abdomen. Orangish brown, translucent and tinged with light brown. Tergum IX with posterior gray mark; terga I–VIII with postero-sublateral grayish brown triangular mark; grayish brown marks on posterolateral area of tergum I–VII, and on antero-sublateral areas of terga VI–VII. Sternites orangish brown, translucent; darker on I–VI; sternite X with lateral orangish marks. Spiracles brownish; tracheae gray. Genitalia (Figs 4–5): Styliger plate brown. Forceps brown, inner margin of forceps segment II and III lighter (Fig. 5). Penis dark brown medially, lighter toward base and apex. MORPHOLOGY. Head: posterior margin V-shaped. Upper portion of compound eye on short stalk (wider than long), separated medially by distance of one compound eye; medial projection of dorsal surface present (Fig. 1); dorsal surface elongated with 20 facets on longest row. Thorax: Wings (Figs 2–3). Three bullae visible; 14 cross veins between C and Sc, 10 distal to bulla; fork of MA strongly asymmetric; fork of MP slightly (Fig. 2). Hind wing with apex acute; costal projection poorly developed; Sc ends distally of apex of costal projection; fork of R+MA slightly asymmetric; MP free basaly; CuP present; Anal vein absent (Fig. 3). Genitalia (Figs 4–5): lateral view (Fig. 4); styliger plate with posteromedial portion concave. Forceps segment I with basal third broad, tapering gradually toward apex, and inner margin with subdistomedial portion broader (Fig. 5, arrow). Base of penis broad tapering abruptly toward apex; base of inner margin with row of spines (Fig. 5).</p><p>Etymology. We name this species in honor of Rafael Boldrini for his friendship and important contribution to the knowledge of mayflies.</p><p>Distribution. Brazil: Amazonas State.</p><p>Material examined. HOLOTYPE: male imago: BRAZIL: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-59.94675&amp;materialsCitation.latitude=-2.9887779" title="Search Plazi for locations around (long -59.94675/lat -2.9887779)">Amazonas</a>, Manaus municipality (Reserva Florestal Adolpho Ducke), 02°59’19.6’’S 59°56’48.3’’W, 69 m, 01–03.ii.2012, Vidovix T. A. S. leg., Malaise trap (INPA) . PARATYPE: one male imago: same data as holotype (INPA); one male imago, Manaus municipality, 02º35’50.9’’S, 60º12’54.9’’W, 49 m, 09–12.xi.2008, Neiss. U.G.; Salles; F.F.; Vilela. P.; Laurindo. P., legs. (IBN, Tucumán, Argentina) .</p><p>Discussion. Miroculis (M.) boldrinii sp. nov. shares some characteristics with Miroculis (Miroculis) nebulosus Savage, 1987 such as a medial projection on the inner margin of the compound eye, basal half of penis broad and tapering abruptly toward the apex, and base of the styliger plate broader than apex. However, M. (M.) boldrinii sp. nov. differs from M. (M.) nebulosus by having 20 facets on the longest row of upper portion of compound eye, instead of 11–14 in M. (M.) nebulosus, by the presence of dense pigmentation on the membrane of fore wings and by the size of penis, shorter than the combination of the styliger plate and forceps in the new species. Fore wings of both species are also distinct; in M. (M.) boldrinii sp. nov. three wide and complete brown transversal bands are present, while in M. (M.) nebulosus these bands are narrower and incomplete.</p></div>	https://treatment.plazi.org/id/EB024F5EFFFFFFB7FF22F8F1FCD2FBF7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Raimundi, Erikcsen Augusto;Domínguez, Eduardo;Salles, Frederico Falcão	Raimundi, Erikcsen Augusto, Domínguez, Eduardo, Salles, Frederico Falcão (2021): Description of a new species, nymphs of two known species and a new record of Miroculis Edmunds, 1963 (Ephemeroptera: Leptophlebiidae) from Brazil. Zootaxa 4963 (2): 384-392, DOI: 10.11646/zootaxa.4963.2.9
EB024F5EFFFCFFB2FF22FB26FD1AFBD3.text	EB024F5EFFFCFFB2FF22FB26FD1AFBD3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Miroculis (Miroculis) caparaoensis Salles & Lima 2011	<div><p>Miroculis (Miroculis) caparaoensis Salles &amp; Lima 2011</p><p>Figures 7; 10; 13–14</p><p>Miroculis (Miroculis) caparaoensis Salles &amp; Lima, 2011: 53 (male and female imago); Massariol, Soares &amp; Salles, 2014: 370.</p><p>Diagnosis. The nymph of Miroculis (M.) caparaoensis differs from the remaining species of Miroculis by the following combination of characters: 1) labrum with anterolateral margin rounded (Fig. 7); 2) maxilla with 56–61 pectinate setae along subapical margin; 3) posterolateral spines on abdominal segments V–IX, more developed from VII to IX; and 4) gills narrow, with the lateral lobes present, and trachea unbranched.</p><p>Description. Mature nymph—in alcohol (Fig. 10).</p><p>MEASUREMENTS (mm). Body: 4.7–5.1 Thorax: fore leg: 3.5–3.7; mid leg: 3.2–3.4; hind leg: 3.5–3.6. RATIOS. Mouthparts: segment III of maxillary palp 0.5× segment II; segment II 1.0–1.2× segment I; segment III of labial palp 0.4–0.5× segment II; segment III 0.4–0.6× segment I; segment II 1.2–1.4× segment I. Legs: width of fore femur 0.3× length; width of mid-femur 0.4–0.5× length. Abdomen: width of lamella of gill in its wider part 0.13× length of gill; length of medial filament of gill 0.9–1.0× length of lamella. COLORATION. Body. Light brown. Head: yellowish brown, pale yellow marks along epicranial and frontal sutures, and between compound eye and lateral ocellus; grayish mark among lateral and medial ocelli and antenna; antenna brown with scape and flagellum lighter. Labrum brown basally, lighter toward apex. Mandible yellowish brown; maxilla, hypopharynx, and labium washed with grayish brown. Thorax: pro- and mesonotum yellowish brown, except for grayish brown marks on sublateral and anteromedial margins of pronotum. Sterna yellowish. Prosternum light yellowish with oblique gray mark sublaterally. Fore and hind wing pads pigmented. Legs: pale yellow. Femora with subapical gray mark on dorsal and ventral surfaces; tibiae with incomplete brown band subbasally, and complete brown band apically; tarsi washed with light brown. Abdomen: terga yellowish brown; terga I–X with longitudinal grayish brown stripe on anterosubmedial margin, and with grayish brown mark along posterior margin; terga I–VII with antero-sublateral brown band. Sterna light yellow; grayish brown marks sublaterally on sterna I–VII and submedial on sterna I–VIII. Gills with membrane grayish and trachea dark gray (Fig. 11). MORPHOLOGY. Head: clypeus anteriorly concave; anterolateral margin of labrum rounded (Fig. 7); maxilla with 56–61 pectinate setae along subapical margin. Abdomen: posterolateral spines present on segment IV or V–IX, more developed on VII–IX. Gills with trachea unbranched; lengths of inner and outer margin of lamella subequal (Fig. 11).</p><p>Male and female imagos (Figs 13–14). Properly described by Salles &amp; Lima (2011); additional information as follows. Male imago: MEASUREMENT (mm): fore wings: 5.9–7.3; hind wing: 1.6–2.3. RATIOS: lateral length of styliger plate: 0.5–0.7 lateral margin; penis length 0.7–0.8 forceps segment I. MORPHOLOGY: inner and outer margins of forceps with different types of rugosity (Fig. 13). In the inner margin they are short, high, with crenulated edges, and in the outer, they are long, low and with smooth edges (Fig. 14).</p><p>Distribution. BRAZIL: Espírito Santo and Minas Gerais.</p><p>Material examined. BRAZIL: Espírito Santo: 1 nymph (n), <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-40.85522&amp;materialsCitation.latitude=-20.549694" title="Search Plazi for locations around (long -40.85522/lat -20.549694)">Alfredo Chaves</a>, municipality, 20°32’58.9’’S, 40°51’18.8’’W, 566m, 31.i.2012, D-net sampler ; 13n, Iúna (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-41.83433&amp;materialsCitation.latitude=-20.41075" title="Search Plazi for locations around (long -41.83433/lat -20.41075)">Parque Nacional do Caparaó</a> — PNC), 20°24’38.7’’S, 41°50’3.6’’W, 1976m, 13.x.2011, Raimundi, E. R.; Massariol, F.C., D-net sampler; 32n, Ibitirama (PNC), 20°23’48.1’’S, 41°44’8.1’’W, 1076m, 15.x.2011, Raimundi, E. R.; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-41.722916&amp;materialsCitation.latitude=-20.469112" title="Search Plazi for locations around (long -41.722916/lat -20.469112)">Massariol</a>, F.C., D-net sampler; 9n, Ibitirama (PNC), 20°28’8.8’’S, 41°43’22.5’’W, 1063m, 21.iv.2008, D-net sampler; 13n, Ibitirama (PNC), 20°28’8.8’’S, 41°43’22.5’’W, 959m, 21.iv.2008, D-net sampler; 3n, Santa Teresa (border of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-40.529804&amp;materialsCitation.latitude=-19.875473" title="Search Plazi for locations around (long -40.529804/lat -19.875473)">Reserva Biológica Augusto Ruschi</a>), 19°52’31.7’’S, 40°31’47.3’’W, 705m, 16.xi.2008, D-net sampler ; 5n, Santa Teresa (border of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-41.836555&amp;materialsCitation.latitude=-20.419888" title="Search Plazi for locations around (long -41.836555/lat -20.419888)">Reserva Biológica Augusto Ruschi</a>), 19°52’32.6’’S, 40°31’49.8’’W, 721m, 19.i.2008, D-net sampler. Minas Gerais: 10n, Alto Caparaó (PNC), 20°25’11.6’’S, 41°50’11.6’’W, 1309m, 12.x.2011, Raimundi, E. R.; Massariol, F.C., D-net sampler; 32n, Espera Feliz (PNC), 20°37’30.3’’S, 41°49’27.1’’W, 884m, 15.x.2011, Raimundi, E. R.; Massariol, F.C., D-net sampler; 44n, Alto Caparaó (PNC), 20°28’19.5’’S, 41°49’41.7’’W, 1972m, 16.x.2011, Raimundi, E. R .; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-41.830666&amp;materialsCitation.latitude=-20.482529" title="Search Plazi for locations around (long -41.830666/lat -20.482529)">Massariol</a>, F.C., D-net sampler; 1n, Alto Caparaó (PNC), 20°28’57.1’’S, 41°49’50.4’’W, 1850m, 22.iv.2008, D-net sampler ; 5n, Alto Caparaó (PNC), 20°25’11.6’’S, 41°50’11.6’’W, 1309m, 23.iii.2009, Raimundi, E. R.; Massariol, F.C., D-net sampler; 3n, Alto Caparaó (PNC), 20°27’21.4’’S, 41°48’’32.1’’W, 2250m, 23.iv.2008, Raimundi, E. R.; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-41.829056&amp;materialsCitation.latitude=-20.481277" title="Search Plazi for locations around (long -41.829056/lat -20.481277)">Massariol</a>, F.C., D-net sampler; 2n, Alto Caparaó (PNC), 20°28’52.6’’S, 41°49’44.6’’W, 1854m, 23.iv.2008, D-net sampler; 1n, Alto Caparaó (PNC), 20°28’57’’S, 41°49’50.2’’W, 1855m, 25.iii.2009, D-net sampler; 6♀, 4♂, Espera Feliz (PNC), 20°37’30.3”S, 41°49’27.1”W, 884m, 14.x.2011, Raimundi, EA; Massariol, FC, Insect Net. All the specimens are housed at UFVB .</p><p>Biological aspects. Miroculis (M.) caparaoensis is distributed in the Atlantic Forest biome, mainly associated with protected areas (such as the Parque Nacional do Caparaó) and adjacencies, including on the border of Reserva Biológica Augusto Ruschi, in altitudes from 566 to 2250 m.a.s.l. Nymphs occur in a range of small to large streams, but they always are associated with leaf litter, among rocky substrates, in weak to moderate current. A flight of M. (M.) caparaoensis was recorded between 5 and 6 PM, approximately 3m above ground, with an abundance of about 50 specimens, both male and female.</p></div>	https://treatment.plazi.org/id/EB024F5EFFFCFFB2FF22FB26FD1AFBD3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Raimundi, Erikcsen Augusto;Domínguez, Eduardo;Salles, Frederico Falcão	Raimundi, Erikcsen Augusto, Domínguez, Eduardo, Salles, Frederico Falcão (2021): Description of a new species, nymphs of two known species and a new record of Miroculis Edmunds, 1963 (Ephemeroptera: Leptophlebiidae) from Brazil. Zootaxa 4963 (2): 384-392, DOI: 10.11646/zootaxa.4963.2.9
EB024F5EFFF9FFB3FF22FB42FE5BF9BC.text	EB024F5EFFF9FFB3FF22FB42FE5BF9BC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Miroculis (Ommaethus) misionensis Dominguez 2007	<div><p>Miroculis (Ommaethus) misionensis Domínguez 2007</p><p>Figures 8; 12</p><p>Miroculis (Ommaethus) misionensis Domínguez, 2007: 99 (male and female imago).</p><p>Diagnosis. The nymph of Miroculis (O.) misionensis differs from the remaining species of Miroculis by the following combination of characters: 1) labrum with anterolateral margin rounded; 2) maxilla with 53 pectinate setae along subapical margin; 3) posterolateral spines on abdominal segments V–IX, more developed from VII to IX; and 4) gills broad, with the lateral lobes present and trachea unbranched.</p><p>Description. Mature nymph (exuviae)—in alcohol.</p><p>MEASURES (mm). Body: 5.9. Thorax: fore leg: 3.0; middle leg: 2.8; hind leg: 3.1. RATIOS. Mouthparts: segment III of maxillary palp 0.3× segment II; segment II 1.2× segment I; segment III of labial palp 0.5× segment I; (segment I of labial palp damaged). Legs: width of fore femur 0.4× length; width of hind-femur 0.4× length. Abdomen: width of lamella of gill in its wider part 0.17× length of gill (Fig. 12); length of medial filament of gill 0.8× length of gill body. COLORATION. Body. Thorax: pattern of marks as in male imago, with posterior margin of all segments dark, terga II–VII with U-shaped lateral marks on each segment and tergum VII with submedial longitudinal lines. Legs: pattern of marks as in male imago, except for sub-basal band on tibiae; femora with no subapical marks on dorsal and ventral surfaces. Abdomen: pattern of marks as in male imago (Domínguez, 2007). Gills with inner margin of membrane darker than outer margin. MORPHOLOGY. Head: clypeus anteriorly concave; anterolateral margin of labrum rounded (Fig. 8); maxilla with 53 pectinate setae along subapical margin. Abdomen: posterolateral spines present on segments V–IX, more developed on VII–IX. Gills broad, with lateral lobes present and length of inner and outer margin of lamella subequal; with trachea unbranched (Fig. 12).</p><p>Male imago. Properly described by Domínguez (2007) with the additional information: RATIOS. Forceps with segment III 0.9 × segment II; forceps with segment II 0.4 × segment I; styliger plate with lateral length 0.5 × medial width; penis length 0.8 × length of forceps segment I. MORPHOLOGY. Styliger plate with anterior margin straight; forceps segment II darker than segment I; 1/4 basal of forceps segment I broad narrowing abruptly toward apex.</p><p>Material examined. BRAZIL: Santa Catarina: 1 male imago and nymphal exuviae, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-51.322556&amp;materialsCitation.latitude=-27.331696" title="Search Plazi for locations around (long -51.322556/lat -27.331696)">Erval Velho</a> municipality, 27°19’54.1’’S, 51°19’21.2’’W, 760m, 18.ix.2011, Pes, A.; Boldrini, R.; Cruz, P.; Hamada, N. col, reared .</p><p>Distribution. ARGENTINA: Misiones. BRAZIL: Santa Catarina.</p><p>Biological aspects. Originally collected in a protected area (Parque Provincial de Urugua-í, Misiones, Argentina; Domínguez 2007; Manzo et al. 2014), and in low altitudes (from 271 to 410 m.a.s.l.), the nymph collected in Brazil was found in a medium order river surrounded by agricultural areas with relatively higher altitude (760 m. a.s.l.). The substrate was rocky and with particulate organic matter. The species, so far, is distributed in the Atlantic Forest biome.</p></div>	https://treatment.plazi.org/id/EB024F5EFFF9FFB3FF22FB42FE5BF9BC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Raimundi, Erikcsen Augusto;Domínguez, Eduardo;Salles, Frederico Falcão	Raimundi, Erikcsen Augusto, Domínguez, Eduardo, Salles, Frederico Falcão (2021): Description of a new species, nymphs of two known species and a new record of Miroculis Edmunds, 1963 (Ephemeroptera: Leptophlebiidae) from Brazil. Zootaxa 4963 (2): 384-392, DOI: 10.11646/zootaxa.4963.2.9
