taxonID	type	description	language	source
E33C878A674BFF9FFF0EFCBD98430604.taxon	materials_examined	Type species: Zercon fimbriatus C. L. Koch, 1839 by original designation.	en	Ujvári, Zsolt (2011): Six new species of Prozercon Sellnick, 1943 (Acari, Mesostigmata, Zerconidae) from Greece, with remarks on the genus. Zootaxa 2785: 1-31, DOI: 10.5281/zenodo.201023
E33C878A674BFF9FFF0EFCBD98430604.taxon	materials_examined	Type species. Prozercon lutulentus Halašková, 1963, by original designation.	en	Ujvári, Zsolt (2011): Six new species of Prozercon Sellnick, 1943 (Acari, Mesostigmata, Zerconidae) from Greece, with remarks on the genus. Zootaxa 2785: 1-31, DOI: 10.5281/zenodo.201023
E33C878A674BFF9FFF0EFCBD98430604.taxon	materials_examined	Type species Rafas bisternalis Błaszak, 1979, by original designation.	en	Ujvári, Zsolt (2011): Six new species of Prozercon Sellnick, 1943 (Acari, Mesostigmata, Zerconidae) from Greece, with remarks on the genus. Zootaxa 2785: 1-31, DOI: 10.5281/zenodo.201023
E33C878A674BFF9FFF0EFCBD98430604.taxon	diagnosis	Diagnosis. Podonotal shield carapace-like, expanded anteroventrally and lateroventrally, setae j 1 situated ventrally. Peritrematal shields separated from dorsal shields by a narrow slit of membranous cuticle, their posterolateral tips expanded posteriorly. Peritremes expanded to anterior half of coxae III, straight or slightly bent. Setae r 1 and r 3 shifted ventrally to peritrematal shields, both short, r 3 always smooth and needle-like. Setae z 1 absent. Glands gv 2 absent. Ventrianal shield bearing 19 setae, setae ZV 1 absent. Glands gv 3 situated laterally or anterolaterally to adanal setae. Margin of opisthonotum with six or seven pairs of R-setae. Notes on the genus. Great variety can be observed in the shape of the peritrematal shields and the position of ventral shields in recent Prozercon species. A group of species has freely-ending peritrematal shields in both female and male, and the size of the posterolateral expansion of shields is characteristic for species. However there is precedent for individual variability, even within a population (see P. graecus sp. nov. below). In another group, the peritrematal shields of the female end freely, while they are fused with the ventrianal and dorsal shields in the males (e. g. P. morazae sp. nov.), and in a few species all three shields are fused together in both sexes. The subgenus Prozercon (Plumatozercon) was established by Balan (1992), including two species, P. lutulentus Halašková, 1963 and P. halaskovae Petrova, 1977 b, but Balan failed to designate a type species and the name is therefore not available. The name Plumatozercon was first made available in Urhan & Ayyildiz (1996 c), who gave a short diagnosis accompanied by designation of P. lutulentus as type species of the subgenus. Peritrematal seta r 1 is always short in Prozercon, but may be either pilose or smooth. This is the only character for distinguishing the subgenera Prozercon and Plumatozercon. The distinguishing character of the subgenera Prozercon and Plumatozercon is clearly unstable (as e. g. shape of r 1 varies even within a population of P. graceus sp. nov. from needlelike to pilose), and therefore it does not reflect phylogenetic divergence. Accordingly, I propose that Plumatozercon is a junior synonym of the subgenus Prozercon. The sternal shield of the female and the sternigenital shield of the male are weakly sclerotised in many Prozercon species, which is often expressed in the presence of a weakly sclerotised band between sternal setae st 1 and st 2 (Figs 1 – 4) (e. g. Cälugär, 2004). Rarely it appears that setae st 1 are inserted on small, isolated platelets (Moraza, 1988). A similar phenomenon was described by Błaszak (1979 b) as the sternal shield of the female and sternigenital shield of the male completely divided behind setae st 1, and this served as a major distinguishing feature of the genera Rafas and Prozercon. After reviewing the type specimens of Prozercon bisternalis (Błaszak, 1979 b) (deposited in Collection of Soil Zoology, Hungarian Natural History Museum), I have found that P. bisternalis and many Prozercon species (e. g. some specimens of P. graecus sp. nov.) are identical, and there is no evidence that the anterior sternal region of Rafas develops separately as a jugular shield. Moreover, in the holotype and some paratypes of P. bisternalis, the sternal shield seems to be undivided (Fig. 4), therefore it is a character of individual variability as well. Accordingly, this character should not be used in generic delimitation. Another distinctive feature of Rafas was that there are only seven pairs of marginal setae on the opisthonotum, while the Prozercon species bear eight pairs of setae in the same position (Błaszak, 1979 b). However this is not true inasmuch as numerous species of Prozercon characteristically possess seven pairs of marginal setae (variation in number of opisthomarginal setae is discussed below). I therefore propose that Rafas is a junior synonym of Prozercon, which results in two new combinations as Prozercon bisternalis (Błaszak, 1979 b) comb. nov. and Prozercon blaszaki (Urhan & Ayyildiz, 1996 d) comb. nov. Absence of gland openings gv 2 seems to be a stable apomorphy (however it is shared with many other genera) as it lacks in every Prozercon species I have ever observed. It serves as a basis for distinguishing the genera Aleksozercon Petrova, 1978 (where a single opening of gv 2 present) and Prozercon, while these genera share all other important characters. The single species Aleksozercon zachvatkini Petrova, 1978 is quite similar in general appearance to Prozercon species. It is important to mention that the major distinctive feature in Petrova’s original description was the partial (apparent?) fusion of the opisthonotal and podonotal shields in Aleksozercon. The same phenomenon can be observed in P. semiseparatus Ujvári, 2009, but the latter lacks gv 2. Apparent fusion of the dorsal shields seems to be a rare apomorphy, which may have evolved independently in both groups as a homoplasy, but further observation of Aleksozercon specimens would be necessary for a firm decision. There is a species, Mesozercon changbaiensis (Bei et al., 2002) comb. nov., which was previously described as Prozercon, which possess conspicuous pores gv 2 and ventrianal setae ZV 1. I propose that this species should be transferred to the genus Mesozercon Błaszak, 1976, inasmuch as the posterodorsal muscle scars are unsclerotised and not conspicuous, therefore “ dorsal cavities ” characteristic for most of the Zerconidae genera are lacking, which is a relatively rare apomorphy in the family. Furthermore, other major characters of this species agree with Mesozercon, except the shape of setae r 1, which is plumose in M. changbaiensis and smooth in other Mesozercon species. The number of opisthomarginal setae often varies individually, even within a specimen, and asymmetry of this character is quite common in zerconid mites. In Prozercon, this number varies between five and nine, but specimens with five, six and nine pairs of marginal setae can be considered as aberrant. Observing several specimens of each species it is clear that basically two groups exist, the first one generally bears seven pairs of marginal setae (six R-setae), the second one generally bears eight pairs (seven R-setae). From a zoogeographic point of view it is important to note that most of the species bearing generally seven pairs of opisthomarginal setae are distributed only in the Mediterranean area, with the exception of P. sellnicki Halašková, 1963, which occurs in an area extending to Central Europe as well. Prozercon species with eight pairs of opisthomarginal setae are distributed from the boreal to the Mediterranean zone of Europe and West-Asia.	en	Ujvári, Zsolt (2011): Six new species of Prozercon Sellnick, 1943 (Acari, Mesostigmata, Zerconidae) from Greece, with remarks on the genus. Zootaxa 2785: 1-31, DOI: 10.5281/zenodo.201023
E33C878A674CFF9EFF0EFDAC998505F0.taxon	materials_examined	Material examined (Fig. 36). E- 2160, Greece, Drama county, Falakro Mts., beech forest beneath the ski centre, 1186 m a. s. l., N 41 ° 17.582 ’ E 24 ° 00.422 ’, leaf litter, 31.03.2007, leg. Dányi, L., Erőss, Z., Fehér, Z., Kontschán, J., Murányi, D. (10 females, 3 males); E- 2171, Greece, Drama county, Falakro Mts., rocky alpine grassland beneath the ski centre, 1350 m a. s. l., N 41 ° 17.373 ’ E 24 ° 01.631 ’, soil of alpine meadow, 30.03.2007, leg. Dányi, L., Erőss, Z., Fehér, Z., Kontschán, J., Murányi, D. (1 female); E- 2201, Greece, Xanthi county, Koula Mts., limestone gorge with beech forest north of Oreo, 820 m a. s. l., N 41 ° 17.485 ’ E 24 ° 50.767 ’, leaf litter and dry rotten tree, 03.04.2007, leg. Dányi, L., Erőss, Z., Fehér, Z., Kontschán, J., Murányi, D. (8 females, 1 m male); E- 2191, Greece, Xanthi county, Koula Mts., Aspro Stream and its gallery beneath Oreo, 550 m a. s. l., N 41 ° 16.369 ’ E 24 ° 51.275 ’, moss on rocks, 03.04.2007, leg. Dányi, L., Erőss, Z., Fehér, Z., Kontschán, J., Murányi, D. (11 females); E- 2158, Greece, Drama county, Orvilos Mts., stream in alder gallery, and limestone rocks above Katafito, 823 m a. s. l., N 41 ° 20.725 ’ E 23 ° 40.463 ’, dry rotten wood, 31.03.2007, leg. Dányi, L., Erőss, Z., Fehér, Z., Kontschán, J., Murányi, D. (3 females); E- 2183, Greece, Serres county, brook in a beech forest along the Serres-Kato Vrondou road, 916 m a. s. l., N 41 ° 14.722 ’ E 23 ° 40.513 ’, leaf litter, 30.03.2007, leg. Dányi, L., Erőss, Z., Fehér, Z., Kontschán, J., Murányi, D. (10 females, 1 male, 1 DN).	en	Ujvári, Zsolt (2011): Six new species of Prozercon Sellnick, 1943 (Acari, Mesostigmata, Zerconidae) from Greece, with remarks on the genus. Zootaxa 2785: 1-31, DOI: 10.5281/zenodo.201023
E33C878A674CFF9EFF0EFDAC998505F0.taxon	discussion	Remarks. This species has been reported from Central Europe (Mašán & Fenďa 2004) and the North Balkan region. This is the first record of the species in Greece.	en	Ujvári, Zsolt (2011): Six new species of Prozercon Sellnick, 1943 (Acari, Mesostigmata, Zerconidae) from Greece, with remarks on the genus. Zootaxa 2785: 1-31, DOI: 10.5281/zenodo.201023
E33C878A674CFF9CFF0EFA37983F07EE.taxon	materials_examined	Material examined (Fig. 39). E- 2556, Greece, Arkadia county, Elliniko, Platanus forest south of the Gortis ruins, 380 m a. s. l., N 37 ° 32.020 ’ E 22 ° 03.191 ’, 06.04.2009, leg. Dányi, L., Kontschán, J., Murányi, D. (3 females, 1 DN).	en	Ujvári, Zsolt (2011): Six new species of Prozercon Sellnick, 1943 (Acari, Mesostigmata, Zerconidae) from Greece, with remarks on the genus. Zootaxa 2785: 1-31, DOI: 10.5281/zenodo.201023
E33C878A674CFF9CFF0EFA37983F07EE.taxon	diagnosis	Diagnosis. Peritrematal shield with posterolateral tips reaching level of R 1. Dorsal podonotal setae pilose except j 5 which is smooth. Marginal R-setae smooth, thorn-like, other opisthonotal setae plumose. Setae S 2 inserted on a line connecting Z 1 and Z 2, equidistantly. Setae S 4 absent. Glands gdS 2 (Po 2) situated medially to a line connecting S 2 and Z 2, gdZ 3 (Po 3) anteromedially to insertions of Z 4. Setae J 3 - J 4 reaching bases of the following setae.	en	Ujvári, Zsolt (2011): Six new species of Prozercon Sellnick, 1943 (Acari, Mesostigmata, Zerconidae) from Greece, with remarks on the genus. Zootaxa 2785: 1-31, DOI: 10.5281/zenodo.201023
E33C878A674CFF9CFF0EFA37983F07EE.taxon	description	Description. Female. Length of idiosoma: 342 – 350 μm (346 μm); width: 235 – 240 μm (238 µm) (n = 3). Dorsal idiosoma (Fig. 5). Podonotum with 20 pairs of setae, j 1 - 6, z 2 - 6, s 1 - 6, r 2 and r 4 - 5 inserted dorsally, setae r 1 and r 3 inserted ventrally, on peritrematal shields. Podonotal setae densely plumose except j 5 short, smooth and needle-like. Setae j 2 - 6, s 1, s 4 - 5, z 2, z 4 - 6 pointed, j 1, z 3, s 2 - 3, s 6, r 2, r 4 - 5 brush-like; s 2, r 2 and r 4 shorter than other brush-like marginal setae. Glands gds 1 (po 1) situated posteromedially to insertions of s 1; gdj 4 (po 2) situated anteromedially to z 4; gds 4 (po 3) on line connecting s 4 and s 5. Podonotal shield covered by reticulate pattern, posterior surface with distinct alveolar pits. Opisthonotum with 20 – 21 pairs of setae, J 1 - 5, Z 1 - 5, S 1 - 5, marginal R-series with five or six pairs of setae, six pairs in majority of specimens. Setae J 1 - 5, Z 1 - 4 and S 2 similar in shape and length, densely pilose, pointed. Setae Z 5, S 3 and S 5 brush-like, plumose, extending beyond margin of opisthonotum. Setae S 1 plumose, twice as long as short, smooth and thorn-like R-setae. Setae S 4 absent. Setae J 1 - 2 not reaching insertions of the following setae, J 3 – 4 reaching insertions of the following setae of the series. Setae J 5 reaching beyond margins of opisthonotum. Setae S 2 inserted on line connecting Z 1 and Z 2, equidistantly. Bases of central opisthonotal setae small to medium-sized. Length of opisthonotal setae and distances between their insertions as in Table 1. Glands gdZ 1 (Po 1) situated anteromedially to insertions of Z 1; gdS 2 (Po 2) medially to line connecting S 2 and Z 2; gdZ 3 (Po 3) anteromedially to insertions of Z 4; gdS 5 (Po 4) medially to S 5. Whole opisthonotal shield covered by relatively large, distinct, irregular pits. Dorsal cavities uniform, weakly developed. Ƥ DN Ƥ DN Ƥ DN J 1 26 17 Z 1 22 17 S 1 15 7 J 1 - J 2 38 33 Z 1 - Z 2 53 43 S 1 - S 2 30 28 J 2 28 19 Z 2 25 21 S 2 23 16 J 2 - J 3 34 27 Z 2 - Z 3 28 24 S 2 - S 3 26 23 J 3 28 21 Z 3 24 18 S 3 30 28 J 3 - J 4 27 24 Z 3 - Z 4 21 18 S 3 - S 5 69 64 J 4 30 20 Z 4 22 18 S 5 26 31 J 4 - J 5 25 20 Z 4 - Z 5 56 46 J 5 24 19 Z 5 25 26 Ventral idiosoma (Fig. 6). Peritrematal shield separated from dorsal shields by a narrow membranous slit, with posterolateral tips reaching level of R 1, covered by fine reticulation. Peritremes straight. Chaetotaxy and poroidotaxy of ventral shields typical for genus Prozercon. Sternal shield well sclerotised, 54 μm long and 43 μm wide at the level of setae st 2, with straight posterior margin and reticulate ornamentation. Ventrianal shield with short, smooth and needle-like preanal and adanal setae, setae ZV 1 absent on anterior margin of the shield. Postanal seta longer than preanal and adanal setae, distally serrate, setae JV 5 similar to R-setae, short, smooth and thorn-like. Anal valves with euanal setae. Glands gv 3 situated anterolaterally to adanal setae. Anterior surface of ventrianal shield covered by squamous pattern to level of JV 3 - ZV 3 - ZV 4. Deutonymph (Fig. 7). Length of idiosoma: 306 μm; width: 215 μm (n = 1). Podonotal setae j 1, z 3, s 3 and s 6 markedly elongate, brush-like, plumose. Marginal setae s 2, r 2 and r 4 short, barbed. Setae j 2, j 6, s 1, z 2, z 4, s 4, s 5 and z 6 similar in shape and length, densely pilose, pointed. Setae j 4 and z 5 finely pilose, j 3 and j 5 smooth and needle-like. Position of gland pores as for the female. Opisthonotal shield covered by weakly-developed, serrulate reticulation. Opisthonotal chaetotaxy and poroidotaxy as for the female, setae S 1 similar in length to marginal R-setae. Length of opisthonotal setae and distances between their insertions as in Table 1. Whole surface of opisthonotum covered by small, irregular pits.	en	Ujvári, Zsolt (2011): Six new species of Prozercon Sellnick, 1943 (Acari, Mesostigmata, Zerconidae) from Greece, with remarks on the genus. Zootaxa 2785: 1-31, DOI: 10.5281/zenodo.201023
E33C878A674CFF9CFF0EFA37983F07EE.taxon	discussion	Remarks. This species has been collected in the Aegean region of Turkey (Urhan, 1998 a), and in Crete (Ujvári, 2008). Setae JV 5 are plumose and the peritrematal shields are expanded to the level of setae R 2 on specimens from the Aegean region of Turkey, while JV 5 are smooth and needle-like, peritrematal shields are expanded only to level of S 1 on the ones collected on the Peloponnesos.	en	Ujvári, Zsolt (2011): Six new species of Prozercon Sellnick, 1943 (Acari, Mesostigmata, Zerconidae) from Greece, with remarks on the genus. Zootaxa 2785: 1-31, DOI: 10.5281/zenodo.201023
E33C878A6741FF96FF0EFF579F37016B.taxon	materials_examined	Material examined. Holotype: Female: E- 1979, Greece, Thesprotia county, Petrovitsa, gorge near the village, N 39 ° 33 ’ 28.5 ” E 20 ° 28 ’ 07.8 ”, 314 m a. s. l., moss, 12.05.2006, leg. Dányi, L., Kontschán, J., Murányi, D. Paratypes: Locality and date as for the holotype (1 male); E- 1983, Greece, Ioannina county, east of Metsovo, ” Metzoboy 1987 ” spring, N 39 ° 45 ’ 16.6 ” E 21 ° 08 ’ 56.4 ”, 1027 m a. s. l., leaf litter, 13.05.2006, leg. Dányi, L., Kontschán, J., Murányi, D. (1 male).	en	Ujvári, Zsolt (2011): Six new species of Prozercon Sellnick, 1943 (Acari, Mesostigmata, Zerconidae) from Greece, with remarks on the genus. Zootaxa 2785: 1-31, DOI: 10.5281/zenodo.201023
E33C878A6741FF96FF0EFF579F37016B.taxon	diagnosis	Diagnosis. Peritrematal shield with posterolateral tips reaching level of R 6 - 7. Podonotal central and submarginal setae smooth except j 1. Marginal setae of opisthonotum smooth, thorn-like. Setae Z 5, S 3 - 5, and JV 5 brushlike, plumose. Setae J 1 not reaching insertions of J 2. Setae S 2 smooth, needle-like, half as long as Z 1, situated on line connecting S 1 and Z 1 equidistantly. Glands gdZ 3 (Po 3) situated on line connecting Z 3 and S 4. Surface between J-series covered by relatively large alveolar pits.	en	Ujvári, Zsolt (2011): Six new species of Prozercon Sellnick, 1943 (Acari, Mesostigmata, Zerconidae) from Greece, with remarks on the genus. Zootaxa 2785: 1-31, DOI: 10.5281/zenodo.201023
E33C878A6741FF96FF0EFF579F37016B.taxon	description	Description. Female. Length of idiosoma: 338 μm; width: 257 μm (n = 1). Dorsal idiosoma (Fig. 8). Podonotum with 20 pairs of setae, j 1 - 6, z 2 - 6, s 1 - 6, r 2 and r 4 - 5 inserted dorsally, setae r 1 and r 3 inserted ventrally, on peritrematal shields. Podonotal setae j 2 - 5, z 2, z 4 - 6, s 1 and s 4 - 5 short, smooth and needle-like, s 2 short, finely barbed, j 1, z 3, r 2, r 4 - 5, s 3 and s 6 elongate, feathered. Glands gds 1 (po 1) situated posterolaterally to insertions of s 1; gdj 4 (po 2) situated below line connecting j 4 and z 4; gds 4 (po 3) medially to line connecting s 4 and s 5. Podonotal shield covered by reticulate pattern, posterior surface with distinct alveolar pits. Opisthonotum with 22 pairs of setae (J 1 - 5, Z 1 - 5, S 1 - 5, R 1 - 7). Setae J 1 delicately barbed, reaching approximately half the distance to setae J 2. Setae J 2 - 5 and Z 2 - 4 similar in shape and length, medium-sized, feathered. J 5 not reaching margin of opisthonotum. Z 1 similar in shape and length to J 1. Z 5 elongate, densely pilose. Setae S 2 smooth, pointed, needle-like, situated on line connecting S 1 and Z 1, equidistantly. S 3 – 5 elongate, distally pilose, extending beyond margin of idiosoma. S 1 and R-setae short, smooth and thorn-like. Bases of central opisthonotal setae small to medium-sized. Length of opisthonotal setae and distances between their insertions as in Table 2. Glands gdZ 1 (Po 1) situated anteromedially to insertions of Z 1; gdS 2 (Po 2) situated on line connecting S 2 and Z 2; gdZ 3 (Po 3) on line connecting Z 3 and S 4; gdS 5 (Po 4) anteriorly to S 5. Surface of opisthonotal shield covered by relatively large alveolar pits between J-series, the quadrangle formed by setae J 1 - J 4 - Z 4 - Z 1 with small, distinct spots and fine reticulation, anterolateral surface with tile-like pattern. Dorsal cavities uniform, moderately sclerotised. Ventral idiosoma (Fig. 9). Posterolateral tips of peritrematal shield reaching level of R 6 - 7. Peritremes slightly bent anteriorly. Chaetotaxy and poroidotaxy of ventral shields typical for genus Prozercon. Sternal shield well sclerotised, 55 μm long and 37 μm wide at the level of setae st 2, with straight posterior margin and reticulate ornamentation. Ventrianal shield with short, smooth and needle-like preanal and adanal setae, setae ZV 1 absent on anterior margin of the shield. Postanal seta twice as long as preanal and adanal setae, smooth, setae JV 5 elongate, densely pilose. Anal valves with vestigial euanal setae. Glands gv 3 situated laterally to adanal setae. Anterior surface of ventrianal shield covered by squamous pattern to level of JV 3 - ZV 3 - ZV 4. Male (Figs 10 – 11). Length of idiosoma: 266 – 274 μm (270 μm); width: 220 – 224 μm (222 µm) (n = 2). Chaetotaxy, poroidotaxy and sculptural pattern of dorsal, ventrianal and peritrematal shields similar to those of female. Setae J 1 and Z 1 smooth. Length of opisthonotal setae and distances between their insertions as in Table 2. Dorsal cavities weakly sclerotised. Sternigenital shield well-sclerotised, bearing four pairs of setae. Shape of peritrematal shield similar to that of female, its posterolateral tips reaching level of R 7.	en	Ujvári, Zsolt (2011): Six new species of Prozercon Sellnick, 1943 (Acari, Mesostigmata, Zerconidae) from Greece, with remarks on the genus. Zootaxa 2785: 1-31, DOI: 10.5281/zenodo.201023
E33C878A6741FF96FF0EFF579F37016B.taxon	etymology	Etymology. The species is named after one of the major tribes of ancient Greece, the Achaeans.	en	Ujvári, Zsolt (2011): Six new species of Prozercon Sellnick, 1943 (Acari, Mesostigmata, Zerconidae) from Greece, with remarks on the genus. Zootaxa 2785: 1-31, DOI: 10.5281/zenodo.201023
E33C878A6741FF96FF0EFF579F37016B.taxon	discussion	Remarks. The new species is closely related to P. carpathicus Balan & Sergienko, 1990 on the basis of the similar position and length of homologous dorsal setae, shape and extension of peritrematal shield, and general appearance of the sculptural pattern. Distingushing characters of the two species are presented in Table 3. However the shape of anterior opisthonotal setae (especially J 1 and Z 1) shows gradual variation in some species — for example Mašán & Fenďa (2004) drew attention to this phenomenon in P. carsticus, supporting the junior synonym status of P. ukrainicus. The other characters listed confirm that specimens from Pindus Mts. are different from P. carpathicus. After examination of P. carpathicus specimens from several locations of the Carpathian basin and the Carpathians it seems that the characters on the basis of which I distinguish the two species are constant and do not show variation, neither at the level of individuals, nor populations. Ƥ 3 Ƥ 3 Ƥ 3 J 1 23 18 Z 1 24 18 S 1 7 8 J 1 - J 2 42 35 Z 1 - Z 2 36 24 S 1 - S 2 23 18 J 2 25 19 Z 2 23 18 S 2 12 11 J 2 - J 3 26 23 Z 2 - Z 3 26 19 S 2 - S 3 25 19 J 3 26 21 Z 3 24 21 S 3 31 26 J 3 - J 4 25 19 Z 3 - Z 4 25 23 S 3 - S 4 42 32 J 4 25 22 Z 4 24 23 S 4 31 26 J 4 - J 5 27 10 Z 4 - Z 5 71 53 S 4 - 5 42 36 J 5 20 17 Z 5 35 28 S 5 33 29 Material examined. Holotype. Female: E- 1426: Greece, Macedonia county, near Thessaloniki Askos, 23.05.1995, leg. Orosz, A. Paratypes. E- 1111: Greece, Pella, excavation site, 04.12.1983, leg. Sin, K. (1 female, 1 male); E- 1427, Greece, Macedonia county, Thessaloniki, Mavrouda, 23.05.1995, leg. Orosz, A. (1 female, 2 ales); E- 1428: Greece, Macedonia county, Halkidiki, Paliokastro, 22.05.1995, leg. Orosz, A. (3 females).	en	Ujvári, Zsolt (2011): Six new species of Prozercon Sellnick, 1943 (Acari, Mesostigmata, Zerconidae) from Greece, with remarks on the genus. Zootaxa 2785: 1-31, DOI: 10.5281/zenodo.201023
E33C878A6741FF96FF0EFF579F37016B.taxon	diagnosis	Diagnosis. Peritrematal shield with posterolateral tips reaching level of R 1 - 2. Dorsal podonotal setae pilose except j 5 smooth. Setae S 1 and marginal R-setae smooth, thorn-like, other opisthonotal setae pilose. Setae S 2 inserted near Z 1, in posterolateral position. Glands gdS 2 (Po 2) situated laterally to line connecting S 2 and Z 2, gdZ 3 (Po 3) laterally to line connecting Z 3 and Z 4. None of J-setae reaching bases of the following setae. J-setae, Z 1 - 4 and S 1 inserted on enlarged setal bases.	en	Ujvári, Zsolt (2011): Six new species of Prozercon Sellnick, 1943 (Acari, Mesostigmata, Zerconidae) from Greece, with remarks on the genus. Zootaxa 2785: 1-31, DOI: 10.5281/zenodo.201023
E33C878A6741FF96FF0EFF579F37016B.taxon	description	Description. Female. Length of idiosoma: 326 – 350 μm (342 μm); width: 241 – 252 μm (246 µm) (n = 6). Holotype: length: 344 µm; width: 248 µm. Dorsal idiosoma (Fig. 12). Podonotum with 20 pairs of setae, j 1 - 6, z 2 - 6, s 1 - 6, r 2 and r 4 - 5 inserted dorsally, r 1 and r 3 inserted ventrally, on peritrematal shields. Podonotal setae densely plumose except j 5 short, smooth and needle-like. Setae j 2 - 6, s 1, s 4 - 5, z 2, z 4 - 6 pointed, j 1, z 3, s 2 - 3, s 6, r 2, r 4 - 5 brush-like. Setae s 2 and r 2 markedly shorter than other brush-like marginal setae. Glands gds 1 (po 1) situated posteriorly to insertions of s 1; gdj 4 (po 2) situated anteromedially to z 4; gds 4 (po 3) on line connecting s 4 and s 5. Podonotal shield covered by reticulate pattern. Opisthonotum with 20 – 22 pairs of setae, J 1 - 5, Z 1 - 5, S 1 - 5, marginal R-series with five to seven pairs of setae, six pairs in majority of specimens. All J-setae, Z 1 - 4 and S 1 situated on enlarged setal bases. J-, Z-setae and S 2 - 5 densely pilose, S 1 and setae of R-series short, smooth and thorn-like. Among pilose setae, J 1 - 5, Z 1 - 4 and S 2 narrow, pointed, Z 5 and S 3 - 5 apically broadening, brush-like. None of J- and Z-setae reaching bases of the following setae in the series. Setae J 5 reaching insertions of Z 5. Setae S 3 - 5 reaching beyond margin of opisthonotum. Setae S 2 situated near Z 1, in posterolateral position. Length of opisthonotal setae and distances between their insertions as in Table 4. Glands gdZ 1 (Po 1) situated anteromedially to insertions of Z 1; gdS 2 (Po 2) laterally to line connecting S 2 and Z 2; gdZ 3 (Po 3) laterally to line connecting Z 3 and Z 4; gdS 5 (Po 4) medially to S 5. Whole opisthonotal shield covered by relatively large, distinct, irregular pits. Dorsal cavities uniform, weakly developed. Ƥ 3 Ƥ 3 Ƥ 3 J 1 20 15 Z 1 20 14 S 1 6 6 J 1 - J 2 33 26 Z 1 - Z 2 35 28 S 1 - S 2 37 27 J 2 21 20 Z 2 20 16 S 2 19 15 J 2 - J 3 34 26 Z 2 - Z 3 31 24 S 2 - S 3 31 27 J 3 23 18 Z 3 22 18 S 3 27 24 J 3 - J 4 31 25 Z 3 - Z 4 34 25 S 3 - S 4 37 33 J 4 22 16 Z 4 17 17 S 4 28 24 J 4 - J 5 29 17 Z 4 - Z 5 62 48 S 4 - 5 44 32 J 5 17 16 Z 5 22 25 S 5 30 25 Ventral idiosoma (Fig. 13). Peritrematal shield separated by a narrow membranous slit from dorsal shields, with posterolateral tips reaching level of R 1 - 2, covered by fine reticulation. Peritremes straight. Chaetotaxy and poroidotaxy of ventral shields typical for genus Prozercon. Sternal shield well sclerotised, 60 μm long and 45 μm wide at the level of setae st 2, with arcuate posterior margin and reticulate ornamentation. Ventrianal shield with short, smooth and needle-like preanal and adanal setae, setae ZV 1 absent on anterior margin of the shield. Postanal seta longer than preanal and adanal setae, distally serrate, setae JV 5 brush-like, plumose. Anal valves with vestigial euanal setae. Glands gv 3 situated laterally to adanal setae. Anterior surface of ventrianal shield covered by squamous pattern to level of JV 3 - ZV 3 - ZV 4. Male (Figs 14 – 15). Length of idiosoma: 280 – 287 μm (285 μm); width: 200 – 205 μm (202 µm) (n = 3). Chaetotaxy, poroidotaxy and sculptural pattern of dorsal, ventrianal and peritrematal shields similar to those of the female. Enlarged setal bases of central opisthonotal setae smaller in proportion than in female. Length of opisthonotal setae and distances between their insertions as in Table 4. Sternigenital shield well-sclerotised, bearing four pairs of setae. Shape of peritrematal shield similar to that of female, its posterolateral tips reaching level of R 2.	en	Ujvári, Zsolt (2011): Six new species of Prozercon Sellnick, 1943 (Acari, Mesostigmata, Zerconidae) from Greece, with remarks on the genus. Zootaxa 2785: 1-31, DOI: 10.5281/zenodo.201023
E33C878A6741FF96FF0EFF579F37016B.taxon	etymology	Etymology. The species name ‘ bulbiferus ’ (= bulbi-bearing) refers to the enlarged, bulblike setal bases observed on opisthonotum.	en	Ujvári, Zsolt (2011): Six new species of Prozercon Sellnick, 1943 (Acari, Mesostigmata, Zerconidae) from Greece, with remarks on the genus. Zootaxa 2785: 1-31, DOI: 10.5281/zenodo.201023
E33C878A6741FF96FF0EFF579F37016B.taxon	discussion	Remarks. The new species is closely related to Prozercon rekae Ujvári, 2008 and Prozercon verruciger Mašán & Fenďa, 2004. These three species bear enlarged opisthonotal setal bases and smooth marginal setae (S 1 and Rsetae) on opisthonotum, and can be distinguished by the characters listed in Table 5.	en	Ujvári, Zsolt (2011): Six new species of Prozercon Sellnick, 1943 (Acari, Mesostigmata, Zerconidae) from Greece, with remarks on the genus. Zootaxa 2785: 1-31, DOI: 10.5281/zenodo.201023
E33C878A6744FF94FF0EFE549B8203BA.taxon	materials_examined	Material examined. Holotype. Female: E- 2158, Greece, Drama county, Orvilos Mts., stream in alder gallery, and limestone rocks above Katafito, 823 m a. s. l., N 41 ° 20.725 ’ E 23 ° 40.463 ’, dry rotten wood, 31.03.2007, leg. Dányi, L., Erőss, Z., Fehér, Z., Kontschán, J., Murányi, D. Paratypes. Locality and date as for the holotype (17 females, 8 m males); E- 2165, Drama county, Dit-Rodopi Mts., stream in beech forest north of Kalikarpos, 869 m a. s. l., N 41 ° 27.959 ’ E 24 ° 14.389 ’, moss and fern on rock, 01.04.2007, leg. Dányi, L., Erőss, Z., Fehér, Z., Kontschán, J., Murányi, D. (3 males); E- 2160, Greece, Drama county, Falakro Mts., beech forest beneath the ski centre, 1186 m a. s. l., N 41 ° 17.582 ’ E 24 ° 00.422 ’, leaf litter, 31.03.2007, leg. Dányi, L., Erőss, Z., Fehér, Z., Kontschán, J., Murányi, D. (10 females, 2 males).	en	Ujvári, Zsolt (2011): Six new species of Prozercon Sellnick, 1943 (Acari, Mesostigmata, Zerconidae) from Greece, with remarks on the genus. Zootaxa 2785: 1-31, DOI: 10.5281/zenodo.201023
E33C878A6744FF94FF0EFE549B8203BA.taxon	diagnosis	Diagnosis. Peritrematal shield with posterolateral tips reaching level of R 5. Podonotal central and submarginal setae smooth except j 1. Setae S 1 pilose, other marginal R-setae smooth, thorn-like. Setae J 1, Z 1 and S 2 smooth and needle-like, other opisthonotal setae variably pilose. Setae S 2 inserted on a line connecting S 1 and Z 1, equidistantly. Glands gdS 2 (Po 2) situated on a line connecting S 2 and Z 2, gdZ 3 (Po 3) medial to line connecting Z 3 and Z 4. No opisthonotal setae reaching bases of the following setae. Opisthonotum covered by small, distinct alveolar pits, anterior surface reticulate. Outer pair of dorsal cavities twice as big as inner pair.	en	Ujvári, Zsolt (2011): Six new species of Prozercon Sellnick, 1943 (Acari, Mesostigmata, Zerconidae) from Greece, with remarks on the genus. Zootaxa 2785: 1-31, DOI: 10.5281/zenodo.201023
E33C878A6744FF94FF0EFE549B8203BA.taxon	description	Description. Female. Length of idiosoma: 332 – 350 μm (342 μm); width: 262 – 274 μm (267 µm) (n = 10). Holotype: length: 338 µm; width: 268 µm. Dorsal idiosoma (Fig. 16). Podonotum with 20 pairs of setae, j 1 - 6, z 2 - 6, s 1 - 6, r 2 and r 4 - 5 inserted dorsally, r 1 and r 3 inserted ventrally, on peritrematal shields. Podonotal setae j 1, z 3, s 2 - 3, r 2, r 4 - 5 and s 6 densely plumose, brush-like, other setae on the shield short, smooth and needle-like. Glands gds 1 (po 1) situated anteromedial to insertions of s 1; gdj 4 (po 2) situated on line connecting j 4 and z 4; gds 4 (po 3) lateral to line connecting s 4 and s 5. Podonotal shield covered by reticulate pattern, with small pits on its central and posterior surface. Opisthonotum with 21 – 23 pairs of setae, J 1 - 5, Z 1 - 5, S 1 - 5, marginal R-series with six to eight pairs of setae, seven pairs in majority of specimens. Setae J 1, Z 1 and S 2 similar in shape and length, short, smooth and needlelike. Setae J 2 - 5 and Z 2 - 4 short, finely pilose and pointed. Setae S 1 similar to marginal setae of podonotum, densely pilose. Setae S 3 - 5 and Z 5 twice as long as central opisthonotal setae, brush-like and plumose. Setae S 3 – 5 reaching beyond margin of opisthonotum. Setae S 2 situated on line connecting S 1 and Z 1, equidistantly. Marginal R-setae short, smooth and thorn-like. No opisthonotal setae reaching bases of the following setae of the series. Setae S 2 situated near Z 1, in posterolateral position. Length of opisthonotal setae and distances between their insertions as in Table 6. Glands gdZ 1 (Po 1) situated anterolateral to insertions of Z 1; gdS 2 (Po 2) situated on line connecting S 2 and Z 2, closer to S 2; gdZ 3 (Po 3) medial to line connecting Z 3 and Z 4; gdS 5 (Po 4) near S 5, in medial position. Whole opisthonotal shield covered by small, alveolar pits, anterior surface with reticulate pattern to level of J 2 - Z 2. Dorsal cavities well-developed, lateral pair strongly sclerotised, twice as large as less-sclerotised medial pair. Ventral idiosoma (Fig. 17). Slit between peritrematal shield and dorsal shields inconspicuous. Peritrematal shield with posterolateral tips reaching level of R 5, covered by fine reticulation. Peritremes slightly bent anteriorly. Chaetotaxy and poroidotaxy of ventral shields typical for genus Prozercon. Sternal shield well sclerotised, 50 μm long and 40 μm wide at the level of setae st 2, with straight posterior margin and reticulate ornamentation. Ventrianal shield with short, smooth and needle-like preanal and adanal setae, setae ZV 1 absent. Postanal seta longer than preanal and adanal setae, smooth or finely serrate distally, setae JV 5 brush-like, plumose. Anal valves with vestigial euanal setae. Glands gv 3 situated anterolateral to adanal setae. Anterior surface of ventrianal shield covered by squamous pattern to level of JV 3 - ZV 4 - JV 4. Male (Figs 18 – 19). Length of idiosoma: 257 – 268 μm (263 μm); width: 200 – 208 μm (203 µm) (n = 10). Chaetotaxy, poroidotaxy and sculptural pattern of dorsal, ventrianal and peritrematal shields similar to those of female, except setae j 2 often delicately barbed. Length of opisthonotal setae and distances between their insertions as in Table 6. Sternigenital shield well-sclerotised, bearing four pairs of setae. Peritrematal shield laterally fused to ventrianal shield, the membranous slit between dorsal and ventral shields lacking.	en	Ujvári, Zsolt (2011): Six new species of Prozercon Sellnick, 1943 (Acari, Mesostigmata, Zerconidae) from Greece, with remarks on the genus. Zootaxa 2785: 1-31, DOI: 10.5281/zenodo.201023
E33C878A6744FF94FF0EFE549B8203BA.taxon	etymology	Etymology. The species name ‘ dramaensis ’ refers to Drama county (Greece), where the specimens were collected.	en	Ujvári, Zsolt (2011): Six new species of Prozercon Sellnick, 1943 (Acari, Mesostigmata, Zerconidae) from Greece, with remarks on the genus. Zootaxa 2785: 1-31, DOI: 10.5281/zenodo.201023
E33C878A6744FF94FF0EFE549B8203BA.taxon	discussion	Remarks. The new species is closely related to Prozercon buraki Urhan, 2008 on the basis of the smooth central podonotal setae and unaequal dorsal cavities. The two species can be distinguished by the characters listed in Table 7.	en	Ujvári, Zsolt (2011): Six new species of Prozercon Sellnick, 1943 (Acari, Mesostigmata, Zerconidae) from Greece, with remarks on the genus. Zootaxa 2785: 1-31, DOI: 10.5281/zenodo.201023
E33C878A6746FF8FFF0EF9AC9A010553.taxon	materials_examined	Material examined. Holotype. Female: E- 2580, Greece, Larisa county, Ossa Mts, Karitsa, oak forest south of the village, 520 m a. s. l., N 39 ° 49.615 ’ E 22 ° 46.174 ’, 09.04.2009, leg. Dányi, L., Kontschán, J., Murányi, D. Paratypes. Locality and date as for the holotype (2 females); E- 2165, Drama county, Dit-Rodopi Mts., stream in beech forest north of Kalikarpos, 869 m a. s. l., N 41 ° 27.959 ’ E 24 ° 14.389 ’, moss and fern on rock, 01.04.2007, leg. Dányi, L., Erőss, Z., Fehér, Z., Kontschán, J., Murányi, D. (2 females); E- 2160, Greece, Drama county, Falakro Mts., beech forest beneath the ski centre, 1186 m a. s. l., N 41 ° 17.582 ’ E 24 ° 00.422 ’, leaf litter, 31.03.2007, leg. Dányi, L., Erőss, Z., Fehér, Z., Kontschán, J., Murányi, D. (1 female); E- 2560, Greece, Larisa county, Ossa Mts., Stomio, mixed forest (deciduous and conifer trees) south of the village, 85 m a. s. l., N 39 ° 51.524 ’ E 22 ° 44.613 ’, 09.04.2009, leg. Dányi, L., Kontschán, J., Murányi, D. (4 females); E- 2209, Greece, Rodopi county, Papikio Mts., brook in a secondary forest, 5 km north of Sostis, 442 m a. s. l., N 41 ° 09.859 ’ E 25 ° 16.939 ’, leaf litter, 04.04.2007, leg. Dányi, L., Erőss, Z., Fehér, Z., Kontschán, J., Murányi, D. (11 females, 1 PN); E- 2191, Greece, Xanthi county, Koula Mts., Aspro Stream and its gallery beneath Oreo, 550 m a. s. l., N 41 ° 16.369 ’ E 24 ° 51.275 ’, moss on rocks, 03.04.2007, leg. Dányi, L., Erőss, Z., Fehér, Z., Kontschán, J., Murányi, D. (3 females); E- 2174, Greece, Drama county, Dit- Rodopi Mts., gallery forest stream south of Mikromilia, 430 m a. s. l., N 41 ° 23.326 ’ E 24 ° 10.078 ’, leaf litter, 31.03.2007, leg. Dányi, L., Erőss, Z., Fehér, Z., Kontschán, J., Murányi, D. (1 female); E- 2167, Greece, Serres county, Orvilos Mts., rocky forest 8 km east of Angistrou, 661 m a. s. l., N 41 ° 23.936 ’ E 23 ° 30.321 ’, soil, 30.03.2007, leg. Dányi, L., Erőss, Z., Fehér, Z., Kontschán, J., Murányi, D. (1 female); E- 2158: Greece, Drama county, Orvilos Mts., stream in alder gallery, and limestone rocks above Katafito, 823 m a. s. l., N 41 ° 20.725 ’ E 23 ° 40.463 ’, dry rotten wood, 31.03.2007, leg. Dányi, L., Erőss, Z., Fehér, Z., Kontschán, J., Murányi, D. (10 females, 2 males); E- 2188, Greece, Serres county, Strimonas River, grassland and limestone rocks east of Neo Petrisi, 77 m a. s. l., N 41 ° 17.000 ’ E 23 ° 19.994 ’, termite nest, 30.03.2007, leg. Dányi, L., Erőss, Z., Fehér, Z., Kontschán, J., Murányi, D. (1 female); E- 2183, Greece, Serres county, brook in a beech forest along the Serres- Kato Vrondou road, 916 m a. s. l., N 41 ° 14.722 ’ E 23 ° 40.513 ’, leaf litter, 30.03.2007, leg. Dányi, L., Erőss, Z., Fehér, Z., Kontschán, J., Murányi, D. (1 female, 1 male).	en	Ujvári, Zsolt (2011): Six new species of Prozercon Sellnick, 1943 (Acari, Mesostigmata, Zerconidae) from Greece, with remarks on the genus. Zootaxa 2785: 1-31, DOI: 10.5281/zenodo.201023
E33C878A6746FF8FFF0EF9AC9A010553.taxon	diagnosis	Diagnosis. All dorsal podonotal setae densely pilose in female. Setae S 1 and marginal R-setae smooth, thornlike, other opisthonotal setae densely pilose. Setae S 2 inserted near Z 1, in lateral position. Glands gdS 2 (Po 2) situated slightly lateral to line connecting S 2 and Z 2, gdZ 3 (Po 3) lateral to line connecting Z 3 and Z 4. Setae J 3 - 4 reaching bases of the following setae.	en	Ujvári, Zsolt (2011): Six new species of Prozercon Sellnick, 1943 (Acari, Mesostigmata, Zerconidae) from Greece, with remarks on the genus. Zootaxa 2785: 1-31, DOI: 10.5281/zenodo.201023
E33C878A6746FF8FFF0EF9AC9A010553.taxon	description	Description. Female. Length of idiosoma: 303 – 340 μm (318 μm); width: 215 – 234 μm (220 µm) (n = 10). Holotype: length: 320 µm; width: 215 µm. Dorsal idosoma (Fig. 20). Podonotum with 20 pairs of setae, j 1 - 6, z 2 - 6, s 1 - 6, r 2 and r 4 - 5 inserted dorsally, setae r 1 and r 3 inserted ventrally, on peritrematal shields. All podonotal setae densely plumose. Setae j 2 - 6, s 1, s 4 - 5, z 2, z 4 - 6 pointed, j 1, z 3, s 2 - 3, s 6, r 2, r 4 - 5 brush-like. Setae s 2 and r 2 markedly shorter than other brush-like marginal setae. Glands gds 1 (po 1) situated posterior to insertions of s 1; gdj 4 (po 2) situated on a line connecting j 4 and z 4; gds 4 (po 3) medial to a line connecting s 4 and s 5. Podonotal shield covered by reticulate pattern, with irregular pits on its posterior and posterolateral surface. Opisthonotum with 20 – 21 pairs of setae, J 1 - 5, Z 1 - 5, S 1 - 5, marginal R-series with five or six pairs of setae, six pairs in majority of specimens. Setae J 1 - 5, Z 1 - 5 and S 2 - 5 densely plumose, S 1 and marginal R-setae short, smooth and thorn-like. Plumose opisthonotal setae Z 4 - 5 and S 3 - 5 brush-like, others pointed. Setae J 1 - 2 not reaching insertions of the following setae of the series, J 3 - 4 reaching insertions of the following setae of the series. Setae J 5 and S 3 - 5 reaching beyond margins of opisthonotum. Setae S 2 situated on line connecting Z 1 and S 1, near Z 1. Bases of central opisthonotal setae medium-sized. Length of opisthonotal setae and distances between their insertions as in Table 8. Glands gdZ 1 (Po 1) situated anterior or anterolateral to insertions of Z 1; gdS 2 (Po 2) slightly lateral to a line connecting S 2 and Z 2; gdZ 3 (Po 3) lateral to line connecting Z 3 and Z 4; gdS 5 (Po 4) posteromedial to S 5. Whole opisthonotal shield covered by relatively large, distinct, irregular pits. Dorsal cavities uniform, of normal size and appearance. Ƥ 3 PN Ƥ 3 PN Ƥ 3 PN J 1 26 20 6 Z 1 17 17 6 S 1 6 5 - J 1 - J 2 31 23 23 Z 1 - Z 2 32 26 23 S 1 - S 2 29 25 - J 2 25 20 7 Z 2 23 18 7 S 2 22 19 18 J 2 - J 3 29 21 24 Z 2 - Z 3 28 19 21 S 2 - S 3 30 20 21 J 3 28 23 7 Z 3 25 17 9 S 3 32 21 28 J 3 - J 4 22 20 19 Z 3 - Z 4 29 21 21 S 3 - S 4 33 26 30 J 4 21 15 6 Z 4 23 16 12 S 4 31 21 28 J 4 - J 5 18 15 14 Z 4 - Z 5 55 40 33 S 4 - 5 34 26 28 J 5 20 14 6 Z 5 33 23 26 S 5 33 20 31 Ventral idiosoma (Fig. 21). Slit between peritrematal shield and dorsal shields inconspicuous. Peritrematal shield with posterolateral tips reaching level of R 5 - 6, covered by fine reticulation. Peritremes straight. Chaetotaxy and poroidotaxy of ventral shields typical for genus Prozercon, peritrematal setae r 1 smooth or barbed. Sternal shield 51 μm long and 42 μm wide at the level of setae st 2, with arcuate posterior margin and reticulate ornamentation. A weakly sclerotised slit between level of setae st 1 and st 2 can often be observed. Ventrianal shield with short, smooth and needle-like preanal and adanal setae, setae ZV 1 absent. Postanal seta longer than preanal and adanal setae, distally serrate, setae JV 5 brush-like, plumose. Anal valves with euanal setae. Glands gv 3 situated anterolateral to adanal setae. Anterior surface of ventrianal shield covered by squamous pattern to level of JV 3 - ZV 4 - JV 4. Male (Figs 22 – 23). Length of idiosoma: 245 – 252 μm (247 μm); width: 188 – 193 μm (190 µm) (n = 3). Chaetotaxy, poroidotaxy and sculptural pattern of dorsal, ventrianal and peritrematal shields similar to those of female, except setae j 5 smooth and needle-like. Peritrematal setae r 1 often barbed. Length of opisthonotal setae and distances between their insertions as in Table 8. Sternigenital shield bearing four pairs of setae, a weakly sclerotised slit between level of setae st 1 and st 2 can often be observed. Shape of peritrematal shield similar to that of female, its posterolateral tips reaching level of R 5. Protonymph (Fig. 24). Length of idiosoma: 237 μm; width: 158 μm (n = 1). Podonotal setae j 1, j 3, z 4, r 2 and s 4 elongate, brush-like, plumose, others short. Short setae densely plumose except j 5 and z 5 barbed. Podonotal ornamentation weakly developed, gland openings not conspicuous. On opisthonotum, Z 5 and S 3 - 5 markedly elongate, brush-like, plumose. S 2 half as long as S 3, plumose. Setae J 1 - 5 and Z 1 - 4 shorter, J 1 and Z 1 - 4 pilose, J 2 - 5 delicately barbed. Length of opisthonotal setae and distances between their insertions as in Table 8. Opisthonotum covered by large, irregular pits. Dorsal cavities weakly developed.	en	Ujvári, Zsolt (2011): Six new species of Prozercon Sellnick, 1943 (Acari, Mesostigmata, Zerconidae) from Greece, with remarks on the genus. Zootaxa 2785: 1-31, DOI: 10.5281/zenodo.201023
E33C878A6746FF8FFF0EF9AC9A010553.taxon	etymology	Etymology. The species name ‘ graecus ’ refers to Greece, where the specimens were collected.	en	Ujvári, Zsolt (2011): Six new species of Prozercon Sellnick, 1943 (Acari, Mesostigmata, Zerconidae) from Greece, with remarks on the genus. Zootaxa 2785: 1-31, DOI: 10.5281/zenodo.201023
E33C878A6746FF8FFF0EF9AC9A010553.taxon	discussion	Remarks. The new species belongs to the group characterised by the following features: most podonotal setae densely pilose; central and submarginal setae of opisthonotum densely pilose, S 1 and R-setae short, smooth and thorn-like; setae S 3 - 5 elongate, brush-like, extending beyond margin of opisthonotum; glands gdZ 3 (Po 3) situated lateral to a line connecting Z 3 - 4. Within this group, Prozercon graecus sp. nov. is most similar to Prozercon plumosus Cälugär, 2004, P. tragardhi (Halbert, 1923) and P. blaszaki (Urhan & Ayyildiz, 1996 d) comb. nov., by the similar shape and position of setae S 2 (elongate, inserted near Z 1, in posterolateral position), position of gdS 2 (Po 2) (on line connecting S 2 and Z 2). The four species can be distinguished by the characters listed in Table 9. The species seems to show a large degree of variation in characters which previously served as basis for specific and even generic determination. Both divided and entire sternal shield (Figs 1, 3), smooth and pilose r 1 setae occur even within a population (Figs 28 – 29), and the posterolateral expansion of peritrematal shield varies within a wide range (Figs 25 – 27).	en	Ujvári, Zsolt (2011): Six new species of Prozercon Sellnick, 1943 (Acari, Mesostigmata, Zerconidae) from Greece, with remarks on the genus. Zootaxa 2785: 1-31, DOI: 10.5281/zenodo.201023
E33C878A675CFF8CFF0EFF5799BB02D0.taxon	materials_examined	Material examined. Holotype. Female: E- 2551, Greece, Arkadia county, Aroania Mts., Planitero, Planitero (Landon) Springs, platanus gallery and limestone rocks in the village, 640 m a. s. l., N 37 ° 56.022 ’ E 22 ° 09.971 ’, 07.04.2009, leg. Dányi, L., Kontschán, J., Murányi, D. Paratypes. Locality and date as for the holotype (3 females); E- 2550, Greece, Arkadia county, Tetrazi Mts., Ag. Theodora, rocky maple forest, 490 m a. s. l., N 37 ° 21.269 ’ E 21 ° 58.782 ’, 05.04.2009, leg. Dányi, L., Kontschán, J., Murányi, D. (2 females, 1 male).	en	Ujvári, Zsolt (2011): Six new species of Prozercon Sellnick, 1943 (Acari, Mesostigmata, Zerconidae) from Greece, with remarks on the genus. Zootaxa 2785: 1-31, DOI: 10.5281/zenodo.201023
E33C878A675CFF8CFF0EFF5799BB02D0.taxon	diagnosis	Diagnosis. Peritrematal shield with posterolateral tips reaching level of R 1. Dorsal podonotal setae pilose, except j 5 smooth. Marginal R-setae smooth, thorn-like, other opisthonotal setae plumose. Setae S 2 inserted slightly lateral to a line connecting Z 1 and Z 2, equidistantly. Setae S 3 and S 4 pointed, not extending beyond margins of opisthonotum. Glands gdS 2 (Po 2) situated on a line connecting S 2 and Z 2, gdZ 3 (Po 3) anterior to insertions of Z 4. Setae J 3 - 4 reaching bases of the following setae.	en	Ujvári, Zsolt (2011): Six new species of Prozercon Sellnick, 1943 (Acari, Mesostigmata, Zerconidae) from Greece, with remarks on the genus. Zootaxa 2785: 1-31, DOI: 10.5281/zenodo.201023
E33C878A675CFF8CFF0EFF5799BB02D0.taxon	description	Description. Female. Length of idiosoma: 330 – 347 μm (338 μm); width: 228 – 242 μm (235 µm) (n = 6). Holotype: length: 340 µm; width: 238 µm. Dorsal idiosoma (Fig. 30). Podonotum with 20 pairs of setae, j 1 - 6, z 2 - 6, s 1 - 6, r 2 and r 4 - 5 inserted dorsally, setae r 1 and r 3 inserted ventrally, on peritrematal shields. Podonotal setae pilose except j 5 short, smooth and needle-like. Setae j 2 - 6, s 1, s 4 - 5, z 2, z 4 - 6 pointed, j 1, z 3, s 2 - 3, s 6, r 2, r 4 - 5 brush-like. Setae s 2, r 2 and r 4 shorter than other brush-like marginal setae. Glands gds 1 (po 1) situated posteromedial to insertions of s 1; gdj 4 (po 2) situated on line connecting j 4 and z 4; gds 4 (po 3) on line connecting s 4 and s 5. Podonotal shield covered by reticulate pattern. Opisthonotum with 19 – 21 pairs of setae, J 1 - 5, Z 1 - 5, S 1 - 5, marginal R-series with four to six pairs of setae, six pairs in majority of specimens. Setae J 1 - 5, Z 1 - 4 and S 2 - 4 similar in shape and length, densely pilose, pointed. Setae Z 5 and S 5 brush-like, plumose, extending beyond margin of opisthonotum. Setae S 1 plumose, twice as long as short, smooth and thorn-like R-setae. No J-, Z- and S-setae reaching insertions of the following setae of the series. Setae J 5 extending beyond margins of opisthonotum. Setae S 2 inserted slightly lateral to line connecting Z 1 and Z 2, equidistantly. Bases of central opisthonotal setae medium-sized. Length of opisthonotal setae and distances between their insertions as in Table 10. Glands gdZ 1 (Po 1) situated anteromedial to insertions of Z 1; gdS 2 (Po 2) on line connecting S 2 and Z 2; gdZ 3 (Po 3) anterior to insertions of Z 4; gdS 5 (Po 4) near JV 5, in anterolateral position. Whole opisthonotal shield covered by relatively large, distinct, irregular pits. Dorsal cavities uniform, weakly developed. Ƥ 3 Ƥ 3 Ƥ 3 J 1 25 23 Z 1 25 20 S 1 14 10 J 1 - J 2 28 31 Z 1 - Z 2 44 37 S 1 - S 2 33 25 J 2 27 23 Z 2 24 20 S 2 21 19 J 2 - J 3 32 19 Z 2 - Z 3 30 18 S 2 - S 3 25 17 J 3 28 23 Z 3 22 20 S 3 21 21 J 3 - J 4 31 18 Z 3 - Z 4 27 25 S 3 - S 4 35 30 J 4 25 23 Z 4 17 17 S 4 20 18 J 4 - J 5 31 21 Z 4 - Z 5 44 36 S 4 - 5 28 18 J 5 19 21 Z 5 24 19 S 5 28 18 Ventral idiosoma (Fig. 31). Peritrematal shield separated from dorsal shields by a narrow membranous slit, with posterolateral tips reaching level of R 1, covered by fine reticulation. Peritremes slightly bent. Chaetotaxy and poroidotaxy of ventral shields typical for genus Prozercon. Sternal shield well sclerotised, 45 μm long and 54 μm wide at the level of setae st 2, with straight posterior margin and reticulate ornamentation. Ventrianal shield with short, smooth and needle-like preanal and adanal setae, setae ZV 1 absent. Postanal seta slightly broadening apically, longer than preanal and adanal setae, setae JV 5 similar in shape and length to setae S 1, plumose. Anal valves with vestigial euanal setae. Glands gv 3 situated lateral to adanal setae. Anterior surface of ventrianal shield covered by squamous pattern to level of JV 3 - ZV 3 - ZV 4. Male (Figs 32 – 33). Length of idiosoma: 258 μm; width: 194 μm (n = 1). Chaetotaxy, poroidotaxy and sculptural pattern of dorsal, ventrianal and peritrematal shields similar to those of the female, except that setae j 4 barely pilose in male and setae S 5 of male similar in shape to Z 1 - 4 and S 2 - 4, pointed. Length of opisthonotal setae and distances between their insertions as in Table 10. Sternigenital shield bearing four pairs of setae. Peritrematal shield laterally fused to ventrianal shield, the membranous slit between dorsal and ventral shields lacking.	en	Ujvári, Zsolt (2011): Six new species of Prozercon Sellnick, 1943 (Acari, Mesostigmata, Zerconidae) from Greece, with remarks on the genus. Zootaxa 2785: 1-31, DOI: 10.5281/zenodo.201023
E33C878A675CFF8CFF0EFF5799BB02D0.taxon	etymology	Etymology. The new species is named in honor of the prominent acarologist, Dr. María Lourdes Moraza (Departamento de Zoología y Ecología, Facultad de Ciencias, Universidad de Navarra).	en	Ujvári, Zsolt (2011): Six new species of Prozercon Sellnick, 1943 (Acari, Mesostigmata, Zerconidae) from Greece, with remarks on the genus. Zootaxa 2785: 1-31, DOI: 10.5281/zenodo.201023
E33C878A675CFF8CFF0EFF5799BB02D0.taxon	discussion	Remarks. The new species belongs to the group characterised by the following features: podonotal setae plumose with the exception of j 5; setae S 1 plumose; S-series complete; setae S 3 similar in shape and length to S 2, not reaching beyond margin of opisthonotum. Distingushing characters of the four species of the group (Prozercon morazae sp. nov., Prozercon escalai Moraza, 1988, Prozercon orhani Urhan & Ayyildiz, 1996 c and Prozercon umidicola Urhan, 2002) are presented in Table 11.	en	Ujvári, Zsolt (2011): Six new species of Prozercon Sellnick, 1943 (Acari, Mesostigmata, Zerconidae) from Greece, with remarks on the genus. Zootaxa 2785: 1-31, DOI: 10.5281/zenodo.201023
E33C878A675EFF80FF0EFB2A9EEE0553.taxon	materials_examined	Material examined. Holotype. Female: E- 1991, Greece, Thesprotia, oak wood in the vicinity of Vrosina, N 39 ° 37 ’ 14.9 ” E 20 ° 31 ’ 21.3 ”, 360 m a. s. l., soil and moss on tree bark, 12.05.2006, leg. Dányi, L., Kontschán, J., Murányi, D. Paratypes. Locality and date as for the holotype (2 females); E- 2549, Greece, Phocis county, Parnassos Mts., Eptolofos, spruce forest, southeast of the village, 1115 m a. s. l., N 38 ° 35.245 ’ E 22 ° 30.267 ’, 08.04.2009, leg. Dányi, L., Kontschán, J., Murányi, D. (1 female).	en	Ujvári, Zsolt (2011): Six new species of Prozercon Sellnick, 1943 (Acari, Mesostigmata, Zerconidae) from Greece, with remarks on the genus. Zootaxa 2785: 1-31, DOI: 10.5281/zenodo.201023
E33C878A675EFF80FF0EFB2A9EEE0553.taxon	diagnosis	Diagnosis. Peritrematal shield with posterolateral tips reaching level of R 1. Dorsal podonotal setae pilose, except j 5 smooth. Marginal R-setae smooth, thorn-like, other opisthonotal setae plumose. Setae S 2 inserted slightly lateral to line connecting Z 1 and Z 2, equidistantly. Setae S 3 - 5 brush-like, plumose, extending beyond margin of opisthonotum. Glands gdS 2 (Po 2) situated on a line connecting S 2 and Z 2, gdZ 3 (Po 3) anteromedial to insertions of Z 4. Setae J 3 - 4 reaching bases of the following setae.	en	Ujvári, Zsolt (2011): Six new species of Prozercon Sellnick, 1943 (Acari, Mesostigmata, Zerconidae) from Greece, with remarks on the genus. Zootaxa 2785: 1-31, DOI: 10.5281/zenodo.201023
E33C878A675EFF80FF0EFB2A9EEE0553.taxon	description	Description. Female. Length of idiosoma: 322 – 341 μm (332 μm); width: 248 – 257 μm (252 µm) (n = 4). Holotype: length: 333 µm; width: 254 µm. Dorsal idiosoma (Fig. 34). Podonotum with 20 pairs of setae, j 1 - 6, z 2 - 6, s 1 - 6, r 2 and r 4 - 5 inserted dorsally, setae r 1 and r 3 inserted ventrally, on peritrematal shields. Setae j 5 short, smooth and needle-like, other podonotal setae pilose. Setae j 2 - 6, s 1, s 4 - 5, z 2, z 4 - 6 pointed, j 1, z 3, s 2 - 3, s 6, r 2, r 4 - 5 brush-like. Among marginal setae, s 2, shorter than submarginal setae, r 2 and r 4 - 5 approximately as long as submarginal setae, j 1, z 3, s 3 and s 6 longer than other podonotal setae. Glands gds 1 (po 1) situated posteromedial to insertions of s 1; gdj 4 (po 2) situated anteromedial to z 4; gds 4 (po 3) on a line connecting s 4 and s 5. Podonotal shield covered by reticulate pattern, posterior surface with small, alveolar pits. Opisthonotum with 21 pairs of setae, J 1 - 5, Z 1 - 5, S 1 - 5, marginal R-series with six pairs of setae. Setae J 1 - 5, Z 1 - 4 and S 2 similar in shape and length, densely pilose, pointed. Setae Z 5, and S 3 - 5 brush-like, plumose, extending beyond margin of opisthonotum. Setae S 1 plumose, three times as long as short, smooth and thorn-like R-setae. Setae J 1 - 2 not reaching insertions of the following setae of the series, J 3 - 4 reaching insertions of the following setae of the series. Setae J 5 reaching beyond margins of opisthonotum. Setae S 2 inserted lateral to line connecting Z 1 and Z 2, equidistantly. Bases of central opisthonotal setae medium-sized. Length of opisthonotal setae and distances between their insertions as in Table 12. Glands gdZ 1 (Po 1) situated in the anterior area of insertions of Z 1; gdS 2 (Po 2) on line connecting S 2 and Z 2; gdZ 3 (Po 3) anteromedial to insertions of Z 4; gdS 5 (Po 4) medial to S 5. Whole opisthonotal shield covered by relatively small, distinct, irregular pits. Dorsal cavities uniform, weakly developed. Ventral idiosoma (Fig. 35). Peritrematal shield separated from dorsal shields by a narrow membranous slit, with posterolateral tips reaching level of R 1, covered by fine reticulation. Peritremes very slightly bent. Chaetotaxy and poroidotaxy of ventral shields typical for genus Prozercon. Sternal shield well sclerotised, 59 μm long and 43 μm wide at the level of setae st 2, with straight posterior margin and reticulate ornamentation. Ventrianal shield with short, smooth and needle-like preanal and adanal setae, setae ZV 1 absent. Postanal seta distally serrate, longer than preanal and adanal setae, setae JV 5 similar in shape to setae S 1, plumose. Anal valves with euanal setae. Glands gv 3 situated anterolaterally to adanal setae. Anterior surface of ventrianal shield covered by squamous pattern to level of JV 3 - ZV 3 - JV 4.	en	Ujvári, Zsolt (2011): Six new species of Prozercon Sellnick, 1943 (Acari, Mesostigmata, Zerconidae) from Greece, with remarks on the genus. Zootaxa 2785: 1-31, DOI: 10.5281/zenodo.201023
E33C878A675EFF80FF0EFB2A9EEE0553.taxon	etymology	Etymology. The new species is dedicated in honor of my dear friend, Nóra Cieleszky.	en	Ujvári, Zsolt (2011): Six new species of Prozercon Sellnick, 1943 (Acari, Mesostigmata, Zerconidae) from Greece, with remarks on the genus. Zootaxa 2785: 1-31, DOI: 10.5281/zenodo.201023
E33C878A675EFF80FF0EFB2A9EEE0553.taxon	discussion	Remarks. The new species belongs to the group characterised by the following features: majority of podonotal setae pilose; setae S 1 plumose; S-series complete; setae S 3 - 5 brush-like, extending beyond margin of opisthonotum; glands gdZ 3 (Po 3) situated medial to line of Z-series. Prozercon halaskovae Petrova, 1977 b matches the first four criteria, however the situation of pores gdZ 3 is currently unknown. Distingushing characters of the three species of the group (Prozercon norae sp. nov., Prozercon dominiaki Błaszak, 1979 a and Prozercon kamili Urhan & Ayyildiz, 1996 c) and the morphologically similar Prozercon halaskovae are presented in Table 13. Ƥ Ƥ Ƥ J 1 29 Z 1 23 S 1 16 J 1 - J 2 30 Z 1 - Z 2 43 S 1 - S 2 40 J 2 27 Z 2 25 S 2 23 J 2 - J 3 32 Z 2 - Z 3 24 S 2 - S 3 30 J 3 33 Z 3 27 S 3 27 J 3 - J 4 22 Z 3 - Z 4 22 S 3 - S 4 43 J 4 34 Z 4 26 S 4 28 J 4 - J 5 32 Z 4 - Z 5 57 S 4 - 5 39 J 5 29 Z 5 29 S 5 29 P. norae sp. nov. P. dominiaki P. halaskovae P. kamili	en	Ujvári, Zsolt (2011): Six new species of Prozercon Sellnick, 1943 (Acari, Mesostigmata, Zerconidae) from Greece, with remarks on the genus. Zootaxa 2785: 1-31, DOI: 10.5281/zenodo.201023
