taxonID	type	description	language	source
DD0287CC1038682BFF2CF89DE15C25A8.taxon	materials_examined	Type: — CHINA. Xizang: Cona, Langpo, sandy grassy slope, 27 ° 53 ’ N, 92 ° 0 ’ E, 4020 m, 11 August 2013, FLPH Tibet Exped. 13 - 0894 (holotype PE; isotypes PE). Herbs up to 40 cm tall, perennial, caespitose. Caudex usually branched, apically covered with remains of petioles. Stems usually numerous, simple, whitish, erect, sparsely tomentose. Rosette leaves long petiolate, chartaceous; petiole to 5 cm long; lower leaves lanceolate, 10 – 15 × 1 – 1.5 cm, margin sinuate-dentate to shallowly pinnately lobed, lobe margin usually entire, rarely with 3 – 4 small teeth, adaxially greyish green and white tomentose, abaxially densely white tomentose, base attenuate, apex acuminate. Capitula solitary, terminal on stem. Involucre campanulate, 1.5 – 2.5 cm in diameter. Phyllaries in 4 – 5 rows, purplish brown, pubescent, apex acuminate, usually reflexed; outer phyllaries triangular-lanceolate, 13 – 14 × 3 – 4 mm; middle phyllaries ovate-elliptic, 13 – 15 × 2.5 – 3.5 mm; inner phyllaries linear, 15 – 16 × 1.2 – 1.5 mm. Receptacle bristles subulate, 4 – 8 mm, greyish white. Corolla blue purplish, 1.7 – 1.9 cm long, tube 8 – 9 mm long, limb ca. 7 mm long, lobes ca. 4 mm long. Anther ca. 7 mm long, tails lanate, ca. 1.5 mm long. Achene brown, cylindric, 5 – 5.3 mm long, ribbed, glabrous, apex shortly crowned. Pappus pale brown; outer bristles 4 – 6 mm long, scabrid; inner bristles 1.2 – 1.3 cm long, plumose. from FLPH Tibet Exped. 13 - 0894 (PE).	en	Chen, You-Sheng (2014): Six new species of Saussurea (Asteraceae) from eastern Himalaya. Phytotaxa 177 (4): 191-206, DOI: 10.11646/phytotaxa.177.4.1, URL: http://dx.doi.org/10.11646/phytotaxa.177.4.1
DD0287CC1038682BFF2CF89DE15C25A8.taxon	distribution	Distribution and habitat: — Saussurea austrotibetica is currently known only from Cona, Gyaca and Yadong Counties in southern Xizang, China. It grows on alpine sandy grassy slope at altitudes of 3670 – 4020 m. Phenology: — Flowering and fruiting from July to September.	en	Chen, You-Sheng (2014): Six new species of Saussurea (Asteraceae) from eastern Himalaya. Phytotaxa 177 (4): 191-206, DOI: 10.11646/phytotaxa.177.4.1, URL: http://dx.doi.org/10.11646/phytotaxa.177.4.1
DD0287CC1038682BFF2CF89DE15C25A8.taxon	etymology	Etymology: — The specific epithet is derived from its distribution range in southern Tibet, China. Additional specimens examined (paratypes): — CHINA. Xizang: Gyaca, Longda Shan, 4210 m, 25 July 1975, Qinghai-Xizang Exped. 750695 (PE); Yadong, Kangbo, 27 ° 39 ’ N, 88 ° 57 ’ E, 3670 m, 24 August 2013, FLPH Tibet Exped. 13 - 2040 (PE).	en	Chen, You-Sheng (2014): Six new species of Saussurea (Asteraceae) from eastern Himalaya. Phytotaxa 177 (4): 191-206, DOI: 10.11646/phytotaxa.177.4.1, URL: http://dx.doi.org/10.11646/phytotaxa.177.4.1
DD0287CC1038682BFF2CF89DE15C25A8.taxon	discussion	Discussion: — Saussurea austrotibetica belongs to S. subgen. Saussurea sect. Strictae because of its pinnately lobed leaves and solitary capitula. Saussurea austrotibetica is very similar to S. roylei subsp. stenophylla Lipschitz (1968: 225), from which it differs in its leaf margin never entire, receptacle bristles 6 – 8 mm, phyllaries shorter and narrower. Saussurea austrotibetica is also similar to S. lanata Y. L. Chen & S. Yun Liang (Chen et al. 1981: 96) in its long stem, pinnately lobed leaves and reflexed phyllaries. Saussurea lanata was once treated as synonym of S. semilyrata Bureau & Franchet (1891: 76), but the present author thinks it is good species. In fact S. lanata is distinct because of its leaves pinnately lobed, lobes margin coarsely dentate, phyllaries apically not herbaceous, phyllary apex acuminate, uppermost stem leaves not subtending capitula; whereas S. semilyrata differs by its leave blade lyrate, with terminal lobe much larger than lateral lobes, lateral lobes margin entire, apex obtuse to acuminate, phyllaries usually apically herbaceous and reflexed, uppermost stem leaves subtending capitula. Saussurea lanata is different from S. austrotibetica by its leaves adaxially deep green, hispidulous to glabrous, leaf blade usually larger, 9 – 28 × 1.5 – 4.7 cm, lyrate, with terminal lobe much larger than lateral lobes, lobe margin coarsely dentate, capitula solitary or 2 – 3, while S. austrotibetica is characterized by its leaves adaxially greyish green and white tomentose, leaf blade 10 – 15 × 1 – 1.5 cm, margin sinuate-dentate, lobe margin usually entire, capitula always solitary. A comparison of S. austrotibetica, S. lanata and S. roylei subsp. stenophylla is given in Table 1.	en	Chen, You-Sheng (2014): Six new species of Saussurea (Asteraceae) from eastern Himalaya. Phytotaxa 177 (4): 191-206, DOI: 10.11646/phytotaxa.177.4.1, URL: http://dx.doi.org/10.11646/phytotaxa.177.4.1
DD0287CC103A682DFF2CFB4BE73E246C.taxon	materials_examined	Type: — CHINA. Xizang: Cona, Langpo, near Dongzhang waterfall, sandy grassy slopes beside streams, 27 ° 47 ’ N, 91 ° 59 ’ E, 3580 – 3650 m, 11 August 2013, FLPH Tibet Exped. 13 - 910 (holotype PE; isotypes PE). Herbs 20 – 35 cm tall, perennial. Caudex usually sparsely branched, blackish brown. Stems several, erect, simple, lanuginous in upper part. Basal and lower stem leaves petiolate; petiole (2.4 –) 3.5 – 10 (– 14) cm, base sheathed; leaf blade oblong to narrowly ovate-elliptic, base truncate, 4 – 13 × 2.2 – 5 cm, lyrate and runcinate-pinnately lobed; terminal lobe ovate-triangular, apex acute, distinctly larger than other ones; lateral lobes 2 – 4 pairs, triangular, margin sharply toothed, adaxially nearly glabrous or indistinctly scabrid, with multicellular articulate hairs, abaxially white velutinous, rachis nearly wingless; middle and upper stem leaves gradually smaller upward on stem, sessile, usually narrowly ovate-elliptic, undivided but sharply toothed, or with one or two pair of basal lobes, base semiamplexicaul. Uppermost stem leaves sessile, usually narrowly ovate-elliptic, undivided, base sessile and nearly semiamplexicaul, margin sharply toothed. Capitula solitary, terminal on stem. Involucre campanulate, 1.5 – 2.5 cm in diameter. Phyllaries in ca. 5 rows, densely villous, apex acuminate to acute, coriaceous, patent; outer phyllaries triangular-ovate, 9 – 12 × 1.5 – 2 mm; middle phyllaries narrowly elliptic, 12 – 15 × ca. 2 mm; inner phyllaries linear, 20 – 22 × 2.5 – 2.8 mm. Receptacle bristles dense, dirty white, 4 – 5 mm long. Anther dark purple, ca. 8 mm long, tails ca. 1 mm long. Corolla purple, 18 – 19 mm long, tube 1.1 – 1.2 × ca. 0.5 mm, limb 7.5 – 9 mm long, lobes 5 – 6 × 0.2 – 0.3 mm. Achene brown, cylindric, ca. 4 × 1 mm, glabrous. Pappus pale brown; outer bristles 4 – 5 mm long, scabrid; inner bristles ca. 1.4 cm long, plumose.	en	Chen, You-Sheng (2014): Six new species of Saussurea (Asteraceae) from eastern Himalaya. Phytotaxa 177 (4): 191-206, DOI: 10.11646/phytotaxa.177.4.1, URL: http://dx.doi.org/10.11646/phytotaxa.177.4.1
DD0287CC103A682DFF2CFB4BE73E246C.taxon	distribution	Distribution and habitat: — Saussurea langpoensis only occurs in Langpo, Cona County, Xizang, China, very close to the McMahon Line. It grows on sandy or stony mountain slope along the stream bank or mixed Abies and bamboo forests in valley at altitudes of 3580 – 3650 m. Phenology: — Flowering and fruiting from August to September.	en	Chen, You-Sheng (2014): Six new species of Saussurea (Asteraceae) from eastern Himalaya. Phytotaxa 177 (4): 191-206, DOI: 10.11646/phytotaxa.177.4.1, URL: http://dx.doi.org/10.11646/phytotaxa.177.4.1
DD0287CC103A682DFF2CFB4BE73E246C.taxon	etymology	Etymology: — The specific epithet is derived from its type locality, Langpo, a village in Cona County, Xizang.	en	Chen, You-Sheng (2014): Six new species of Saussurea (Asteraceae) from eastern Himalaya. Phytotaxa 177 (4): 191-206, DOI: 10.11646/phytotaxa.177.4.1, URL: http://dx.doi.org/10.11646/phytotaxa.177.4.1
DD0287CC103A682DFF2CFB4BE73E246C.taxon	discussion	Discussion: — Saussurea langpoensis belongs to S. subgen. Saussurea sect. Strictae because of its pinnately lobed leaves and solitary capitula. Saussurea langpoensis is similar to S. semilyrata Bureau & Franchet (1891: 76) in its habit and leaf shape but differs in its stems usually numerous, leaf lobes 2 – 4 pairs, triangular, margin sharply toothed, terminal lobes distinctly larger than lateral ones, rachis nearly wingless, capitula always solitary, phyllary apex acuminate to acute, coriaceous and patent, whereas S. semilyrata is different in its stem usually single, leaf lobes 6 – 10 pairs, ovate-triangular, margin entire, terminal lobes similar to lateral ones, rachis distinctly winged, capitula 1 – 5, phyllary apex obtuse, usually apically herbaceous and reflexed. Saussurea langpoensis occurs in Cona, southern Xizang, whereas S. semilyrata occurs in western Sichuan and northwestern Yunnan. A comparison of S. langpoensis and S. semilyrata is given in Table 2.	en	Chen, You-Sheng (2014): Six new species of Saussurea (Asteraceae) from eastern Himalaya. Phytotaxa 177 (4): 191-206, DOI: 10.11646/phytotaxa.177.4.1, URL: http://dx.doi.org/10.11646/phytotaxa.177.4.1
DD0287CC103C682FFF2CF96CE0A22704.taxon	materials_examined	Type: — CHINA. Xizang: Lhünzê, Sanga Choling, Kala mountain, grassy slopes beside mixed forests in valley, 28 ° 36 ’ N, 92 ° 56 ’ E, 3800 – 4050 m, 9 August 2013, FLPH Tibet Expedition 13 - 0847 (holotype PE; isotypes PE). Perennial herbs 11 – 30 cm tall, usually caespitose. Caudex stout, branched, apically densely covered with blackish brown, sometimes fibrous lacerate remains of petioles. Stems usually several, erect, simple, dark brown glandular pilose. Rosette leaves petiolate; petiole to 5 cm; leaf blade oblong to lanceolate, 7 – 18 × 1 – 2.5 (– 3) cm, pinnately lobed; lateral lobes 7 – 15 pairs, ovate, margin usually entire, or sometimes few denticulate, apex usually mucronate; abaxially greyish white and arachnoid tomentose, adaxially green and glandular scabrid, apex obtuse and usually mucronate, base attenuate. Stem leaves 2 – 5, shortly petiolate; leaf blade same in division as basal leaves, but smaller in size. Capitula solitary, terminal on stem. Involucre campanulate, (1 –) 1.5 – 2.5 cm in diam. Phyllaries in 5 – 8 rows, sparsely arachnoid, especially outer ones; outer phyllaries narrowly triangular-subulate, 14 – 17 × 2 – 2.5 mm, apex acute, leafy, green, and reflexed; middle phyllaries subulate-linear, 13 – 17 × 3 – 3.5 mm, apex acute, herbaceous, green, and reflexed; inner phyllaries linear, 17 – 19 × ca. 2 mm, apex blackish, acuminate. Receptacle bristles subulate, 3 – 5 mm long. Corolla light blue, 1.5 – 1.8 cm long, tube 10 – 11 mm long, limb 7 – 8 mm long, lobes 4 – 5 mm long. Anther ca. 7 mm long, tails white lanate, ca. 1 mm long. Achene cylindric, ca. 3 mm long, ribbed, glabrous. Pappus pale brown; outer bristles ca. 2.5 mm long, scabrid; inner bristles 13 – 14 mm long, plumose.	en	Chen, You-Sheng (2014): Six new species of Saussurea (Asteraceae) from eastern Himalaya. Phytotaxa 177 (4): 191-206, DOI: 10.11646/phytotaxa.177.4.1, URL: http://dx.doi.org/10.11646/phytotaxa.177.4.1
DD0287CC103C682FFF2CF96CE0A22704.taxon	distribution	Distribution and habitat: — Saussurea lhunzensis occurs in Lhünzê, Gyaca and Mêdog, southestern Xizang, China. It grows on alpine sandy grassy meadows or thickets beside mixed forests in valleys at altitudes of 3800 – 4450 m. Phenology: — Flowering and fruiting from July to September.	en	Chen, You-Sheng (2014): Six new species of Saussurea (Asteraceae) from eastern Himalaya. Phytotaxa 177 (4): 191-206, DOI: 10.11646/phytotaxa.177.4.1, URL: http://dx.doi.org/10.11646/phytotaxa.177.4.1
DD0287CC103C682FFF2CF96CE0A22704.taxon	etymology	Etymology: — The specific epithet is derived from the type locality, Lhünzê, a county in southern Xizang, China. Additional specimens examined (paratypes): — CHINA. Xizang: Gyaca, Hongqi, Cuogu, 3800 m, 19 August 1972, Xizang Chinese Medic. Exped. 4334 (PE); Gyaca, Hongqi, Cuogu, alpine meadows, 4000 m, 19 August 1972, Xizang Chinese Medic. Exped. 4333 (PE); Lhünzê, Xuesha, Kala mountain, grassy slopes in valley, 28 ° 35 ’ N, 92 ° 29 ’ E, 4340 m, 4 August 2013, FLPH Tibet Expedition 13 - 0260 (PE); Kongbo, [Mêdog], Pero La (at southern Duoxiong La), 29 ° 30 ’ N, 95 ° 00 ’ E, 4115 m, 10 July 1938, F. Ludlow et al. 5203 (BM, E).	en	Chen, You-Sheng (2014): Six new species of Saussurea (Asteraceae) from eastern Himalaya. Phytotaxa 177 (4): 191-206, DOI: 10.11646/phytotaxa.177.4.1, URL: http://dx.doi.org/10.11646/phytotaxa.177.4.1
DD0287CC103C682FFF2CF96CE0A22704.taxon	discussion	Discussion: — Saussurea lhunzensis belongs to S. subgen. Saussurea sect. Strictae because of its pinnately lobed leaves and solitary capitula. Saussurea lhunzensis is similar to S. pachyneura Franchet (1894: 354) in its habit, leaf shape, capitula number and size but differs in its stem covered by glandular pilose hairs, leaf adaxially glandular scabrid, phyllary herbaceous, phyllary apex acute and reflexed, whereas S. pachyneura is different in its stem densely arachnoid, leaf adaxially glabrous, phyllary coriaceous, phyllary apex acuminate and appressed. Saussurea lhunzensis occurs in southerneastern Xizang, whereas S. pachyneura occurs in Gansu, Qinghai, Sichuan, and Xizang. A comparison of S. lhunzensis and S. pachyneura is given in Table 3.	en	Chen, You-Sheng (2014): Six new species of Saussurea (Asteraceae) from eastern Himalaya. Phytotaxa 177 (4): 191-206, DOI: 10.11646/phytotaxa.177.4.1, URL: http://dx.doi.org/10.11646/phytotaxa.177.4.1
DD0287CC103E682FFF2CFF2AE7552026.taxon	materials_examined	Type: — CHINA. Xizang: Yadong, Pagri, mountain between border of China and Bhutan, scree slope or sandy meadows, 27 ° 37 ’ N, 89 ° 07 ’ E, 4600 – 4870 m, 25 August 2013, FLPH Tibet Exped. 13 - 2141 (holotype PE; isotypes PE). Herbs perennial, usually stemless or shortly stemmed up to 5 cm high, caespitose. Caudex stout, usually numerously branched. Rosette leaves sessile, linear-lanceolate, 2 – 9 × 0.3 – 1 cm, abaxially greyish white and densely tomentosesericeous, adaxially green and sparsely arachnoid, base sheathing, margin pinnately dentate and revolute, apex acute, midvein distinct. Capitula solitary, in center of leaf rosette or terminal on stem, sessile. Involucre campanulate, 1.3 – 2 cm in diameter. Phyllaries in ca. 5 rows, glabrous to sparsely arachnoid, apex obtuse to acute; outer phyllaries narrowly ovate-triangular, 8 – 14 × 3 – 5 mm, basal part green, apical part and margin blackish purple; middle and inner phyllaries narrowly ovate-triangular to narrowly elliptic, 11 – 13 × 2 – 4 mm, basal part pale yellow, apical part blackish purple. Receptacle with dense bristles, straw-color, 4 – 6 mm long. Corolla purplish, 1.5 – 1.7 cm long, tube 7 – 8 mm long, limb 8 – 9 mm long, lobes ca. 4 mm long. Anther ca. 6 mm long, tails lanate, ca. 1 mm long. Achene dark brown, conic, ca. 2.5 mm long, glabrous, ribbed, apex shallowly crowned. Pappus in 2 rows, greyish white; outer bristles 7 – 8 mm long, plumose; inner bristles 11 – 12 mm long, plumose.	en	Chen, You-Sheng (2014): Six new species of Saussurea (Asteraceae) from eastern Himalaya. Phytotaxa 177 (4): 191-206, DOI: 10.11646/phytotaxa.177.4.1, URL: http://dx.doi.org/10.11646/phytotaxa.177.4.1
DD0287CC103E682FFF2CFF2AE7552026.taxon	distribution	Distribution and habitat: — Saussurea pagriensis occurs in northern Bhutan and Yadong, Xizang, China (which is adjacent to Bhutan). It grows on alpine scree slope or sandy meadows at altitudes of 4600 – 4870 m. Phenology: — Flowering and fruiting from August to October.	en	Chen, You-Sheng (2014): Six new species of Saussurea (Asteraceae) from eastern Himalaya. Phytotaxa 177 (4): 191-206, DOI: 10.11646/phytotaxa.177.4.1, URL: http://dx.doi.org/10.11646/phytotaxa.177.4.1
DD0287CC103E682FFF2CFF2AE7552026.taxon	etymology	Etymology: — The specific epithet is derived from the type locality name, Pagri, a frontier town in Yadong, Xizang. Additional specimens examined (paratypes): — BHUTAN. Upper Mo Chu District: [Gasa], south slope of Yale La, 27 ° 47 ’ N, 89 ° 27 ’ E, dry unstable scree, 4780 m, 3 October 1984, I. W. J. Sinclair & D. G. Long 5498 (E, K).	en	Chen, You-Sheng (2014): Six new species of Saussurea (Asteraceae) from eastern Himalaya. Phytotaxa 177 (4): 191-206, DOI: 10.11646/phytotaxa.177.4.1, URL: http://dx.doi.org/10.11646/phytotaxa.177.4.1
DD0287CC103E682FFF2CFF2AE7552026.taxon	discussion	Discussion: — Saussurea pagriensis belongs to S. subgen. Saussurea sect. Strictae because of its pinnately dentate leaves and solitary capitula. This species is distinct in its outer pappus plumose. Saussurea pagriensis is somewhat similar to S. purpurascens Y. L. Chen & S. Yun Liang (Chen et al. 1981: 105) and S. taraxacifolia (Lindley ex Royle 1835: 251) Wallich ex Candolle (1838: 532) in its leaf shape. However S. pagriensis is distinct in its usually stemless habit, caudex numerous branched, leaves sessile and linear-lanceolate, margin denate, capitula usually smaller, outer pappus plumose. A comparison of S. pagriensis, S. purpurascens and S. taraxacifolia is given in Table 4.	en	Chen, You-Sheng (2014): Six new species of Saussurea (Asteraceae) from eastern Himalaya. Phytotaxa 177 (4): 191-206, DOI: 10.11646/phytotaxa.177.4.1, URL: http://dx.doi.org/10.11646/phytotaxa.177.4.1
DD0287CC10306821FF2CFDD3E0CB23F3.taxon	materials_examined	Type: — CHINA. Xizang: Yadong, Pagri, mountains between border of China and Bhutan, 27 ° 37 ’ N, 89 ° 07 ’ E, sandy meadows, 4870 m, 25 August 2013, FLPH Tibet Exped. 13 - 2131 (holotype PE; isotypes PE). Herbs 1 – 4 cm tall, perennial, stemless, caespitose. Caudex 2 – 3 cm in diam., usually much branched. Rosette leaves sessile, linear, 10 – 25 × 1 – 1.5 mm, abaxially greyish white and densely tomentose-sericeous, adaxially green, shiny, and glabrous, base enlarged, sheathing, and white villous, margin entire and revolute, apex acute. Uppermost leaves merging into phyllaries, supporting capitula. Capitula solitary, in center of leaf rosette or terminal on stem, sessile, concealed by villous hairs. Involucre campanulate, 1 – 1.6 cm in diam. Phyllaries in ca. 6 rows, apex acuminate to caudate; outer phyllaries narrowly ovate-triangular, 14 – 16 × 3 – 4 mm, basal part dark green, apical part greenish, sparsely villous, and reflexed; middle and inner phyllaries narrowly ovate-triangular to narrowly elliptic-linear, 11 – 13 × ca. 2 mm, basal part pale yellow, apical part blackish purple, villous, and reflexed. Receptacle bristles very few, 1 – 2 mm long. Corolla purplish, 1.3 – 1.5 cm long, tube 7 – 8 mm long, limb 6.5 – 8 mm long, lobes 3.5 – 4 mm long. Achene dark brown, conic, ca. 3 mm long, glabrous. Pappus in 2 rows; outer bristles white, 2 – 3 mm long, scabrid; inner bristles pale brown, 9 – 10 mm long, plumose.	en	Chen, You-Sheng (2014): Six new species of Saussurea (Asteraceae) from eastern Himalaya. Phytotaxa 177 (4): 191-206, DOI: 10.11646/phytotaxa.177.4.1, URL: http://dx.doi.org/10.11646/phytotaxa.177.4.1
DD0287CC10306821FF2CFDD3E0CB23F3.taxon	distribution	Distribution and habitat: — Saussurea bhutanensis occurs in northern Bhutan and adjacent border of Yadong, Xizang, China. It grows on alpine sandy meadows (including Saussurea, Aster, Primula, Arenaria, Gentiana, Swertia, Cyananthus, Saxifraga and Delphinium species), alpine scree or limestones at altitudes of 4500 – 4900 m. Phenology: — Flowering and fruiting from August to October.	en	Chen, You-Sheng (2014): Six new species of Saussurea (Asteraceae) from eastern Himalaya. Phytotaxa 177 (4): 191-206, DOI: 10.11646/phytotaxa.177.4.1, URL: http://dx.doi.org/10.11646/phytotaxa.177.4.1
DD0287CC10306821FF2CFDD3E0CB23F3.taxon	etymology	Etymology: — The specific epithet is derived from its main distribution area, Bhutan. Additional specimens examined (paratypes): — BHUTAN. [Bumthang district]: Tolegang, Tsampa, steep open hillside, 4720 m, 2 October 1949, F. Ludlow, G. Sherriff & J. H. Hicks 19784 (BM, E); [Wangdi district]: Wangdue-Phodrang, Thampe La, limestone, 27 ° 43 ’ N, 90 ° 18 ’ E, 4500 – 4600 m, 28 September 2000, G. & S. Miehe 00 - 421 - 01 (E).	en	Chen, You-Sheng (2014): Six new species of Saussurea (Asteraceae) from eastern Himalaya. Phytotaxa 177 (4): 191-206, DOI: 10.11646/phytotaxa.177.4.1, URL: http://dx.doi.org/10.11646/phytotaxa.177.4.1
DD0287CC10306821FF2CFDD3E0CB23F3.taxon	discussion	Discussion: — Saussurea bhutanensis belongs to S. subgen. Saussurea sect. Lagurostemon on account of its leaf blade being linear, entire, grasslike, and the capitula solitary, 1 – 1.6 cm in diameter. Grierson & Springate (2001) reported one specimen from Bhutan (F. Ludlow, G. Sherriff & J. H. Hicks 19784) that is related to Saussurea columnaris Handel-Mazzetti (1937: 652), but they noted this specimen is different from the typical S. columnaris in its receptacle naked, achene glandular-stipitate above and habit usually stemless. This kind of specimen was also found from the Tibetean border with Bhutan (FLPH Tibet Exped. 13 - 2131) by the present author in 2013. But the present author found its receptacle is not entirely naked but with few short bristles 1 – 2 mm long, achenes glabrous, leaves 1 – 2.5 cm long, phyllaries usually narrower and apex acuminate, capitula concealed by dense villous hairs. Typical specimens of S. columnaris from Yunnan are different in the receptacle bristles ca. 7 mm long and plants usually stemmed, leaves 2 – 7 cm long, phyllaries usually broader and apex acute, capitula not concealed by villous hairs. Saussurea bhutanensis is also similar to S. durgae Jeffrey & Srivastava (1997: 235) in its caespitose habit and linear grassy leaves, but S. bhutanensis differs in its leaves adaxially green, shiny, and glabrous, capitula concealed by villous hairs, and corolla purplish, whereas S. durgae differs in its leaves densely clothed with greyish white wooly hairs on both surfaces, capitula not concealed by villous hairs, and corolla pink. A comparison of S. bhutanensis, S. columnaris and S. durgae is given in Table 5.	en	Chen, You-Sheng (2014): Six new species of Saussurea (Asteraceae) from eastern Himalaya. Phytotaxa 177 (4): 191-206, DOI: 10.11646/phytotaxa.177.4.1, URL: http://dx.doi.org/10.11646/phytotaxa.177.4.1
DD0287CC10316827FF2CFBBAE6A227B0.taxon	materials_examined	Type: — CHINA. Xizang: Lhozhag, Lakang, mountains behind Kajiu monastery, scree slope or sandy meadows, 28 ° 5 ’ N, 91 ° 8 ’ E, 4330 – 4570 m, 18 August 2013, FLPH Tibet Exped. 13 - 1587 (holotype PE; isotypes PE). Herbs 2 – 5 cm tall, perennial, polycarpic, stemless or shortly stemmed. Roots elongate, thickened. Rhizomes usually branched. Flowering stems erect, few, simple, 2 – 5 cm tall, base with persistent withered leaves. Lower leaves narrowly oblanceolate to narrowly oblong, 2 – 5 × 0.4 – 1 cm, margin with 2 – 4 pairs of teeth, apex acute or obtuse, midrib conspicuous, abaxially greyish white and covered with dense white wool-like multicellular hairs, adaxially light green, nearly glabrous, sparsely pilose along midrib. Upper leaves surrounding capitula, ovate-triangular, with dense woollike hairs. Capitula 5 – 25, ca. 5 mm in diameter, densely clustered on flat top of stem in a synflorescence 2.5 – 6 cm in diameter, hidden by dense arachnoid hairs at anthesis. Involucre cylindric, ca. 5 mm in diameter. Phyllaries in 3 or 4 rows, membranous; outer phyllaries spatulate to narrowly obovate, 8 – 11 × 2 – 3.5 mm, apex acute and arachnoid; middle and inner phyllaries slightly smaller, 7 – 8 × 2 – 3.5 mm, apex acute, sparsely arachnoid to glabrous. Receptacle naked. Corolla white, ca. 8.5 mm long, tube ca. 4 long, limb ca. 4.5 mm long, lobes ca. 1 mm long. Anther (including tails) 4.5 – 5 mm long, tails lanate, ca. 1.5 mm long. Achene blackish, conic, ca. 1.5 mm long, glabrous, ribbed. Pappus dark brown; outer bristles 3 – 3.5 mm long, scabrid; inner bristles ca. 8 mm long, plumose.	en	Chen, You-Sheng (2014): Six new species of Saussurea (Asteraceae) from eastern Himalaya. Phytotaxa 177 (4): 191-206, DOI: 10.11646/phytotaxa.177.4.1, URL: http://dx.doi.org/10.11646/phytotaxa.177.4.1
DD0287CC10316827FF2CFBBAE6A227B0.taxon	distribution	Distribution and habitat: — Saussurea lhozhagensis occurs in Lhozhag County, Xizang, China, adjacent to Bhutan. It grows on scree slopes or sandy meadows at altitudes of 4330 – 4570 m. Phenology: — Flowering and fruiting from August to September.	en	Chen, You-Sheng (2014): Six new species of Saussurea (Asteraceae) from eastern Himalaya. Phytotaxa 177 (4): 191-206, DOI: 10.11646/phytotaxa.177.4.1, URL: http://dx.doi.org/10.11646/phytotaxa.177.4.1
DD0287CC10316827FF2CFBBAE6A227B0.taxon	etymology	Etymology: — The specific epithet is derived from its type locality, Lhozhag.	en	Chen, You-Sheng (2014): Six new species of Saussurea (Asteraceae) from eastern Himalaya. Phytotaxa 177 (4): 191-206, DOI: 10.11646/phytotaxa.177.4.1, URL: http://dx.doi.org/10.11646/phytotaxa.177.4.1
DD0287CC10316827FF2CFBBAE6A227B0.taxon	discussion	Discussion: — Saussurea lhozhagensis obviously belongs to S. subgen. Eriocoryne because its capitula are clustered in densely congested hemispheric synflorescences surrounded by densely lanate leaves. Saussurea lhozhagensis is similar to S. conaensis (Liu 1984: 69) Fujikawa & Ohba (2010: 73) in its leaf shape and naked receptacle, but S. lhozhagensis is different in its usually polycarpic habit, stem 2 – 5 cm tall, leaf blade narrowly oblanceolate to narrowly oblong, 2 – 5 × 0.4 – 1 cm, capitula 5 – 25, whereas S. conaensis is different in its usually monocarpic habit, stem 12 – 20 (– 30) cm tall, leaf blade linear to narrowly oblanceolate to narrowly oblong, margin sparsely denticulate, 5 – 17 × 0.5 – 2 (– 3) cm, capitula solitary. Saussurea lhozhagensis is also similar to S. nishiokae Kitamura (1969: 6) in its leaf shape and mumerous capitula but differs in its usually polycarpic habit, stem 2 – 5 cm tall, leaf blade narrowly oblanceolate to narrowly oblong, margin with 2 – 4 pairs of teeth, receptacle naked, whereas S. nishiokae is different in its monocarpic habit, stem 9 – 30 cm tall, leaf blade narrowly elliptic to oblong, margin entire, receptacles densely papillose, papillae ca. 0.5 mm long. A comparison of S. conaensis, S. lhozhagensis and S. nishiokae is given in Table 6.	en	Chen, You-Sheng (2014): Six new species of Saussurea (Asteraceae) from eastern Himalaya. Phytotaxa 177 (4): 191-206, DOI: 10.11646/phytotaxa.177.4.1, URL: http://dx.doi.org/10.11646/phytotaxa.177.4.1
