taxonID	type	description	language	source
DF3B3510FF878E7FFDA1FE67C6B3ED3E.taxon	description	Description. Male. Body length 1.6 – 1.9 mm, wing length 1.4 – 1.6 mm. Morphological characters as in Cook 1956.	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF878E7FFDA1FE67C6B3ED3E.taxon	materials_examined	Material Examined. 2 ♂, China, Qinghai Province, Qilian [Ēê], Huangzangsi [Wëȕ] [38 ° 25 ’ N, 100 ° 19 ’ E, 2588 m], 2023. VIII. 2, Shang Gao (CAU).	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF878E7FFDA1FE67C6B3ED3E.taxon	distribution	Distribution. China (Qinghai Province); Armenia, Austria, Azerbaijan, Bulgaria, Croatia, Czech Republic, Denmark, Egypt, England, Finland, France, Germany, Greece, Ireland, Italy, Kazakhstan, Malta, Montenegro, Morocco, Norway, Poland, Portugal, Russia, Saudi Arabia, Serbia, Slovakia, Spain (Canary Islands), Spain (Continental), Sweden, Switzerland.	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF878E7FFDA1FE67C6B3ED3E.taxon	discussion	Remarks. According to the literature record, this species has a wide distribution in Eurasia (Ebejer 2021; Haenni 2013; Haenni & Kettani 2011; Hrviniak et al. 2018; Oboňa et al. 2017) and can be separated from other species of this genus by the characters of male terminalia and tergite 7. The two males available were collected by sweeping net in Qinghai Province, belonging to the Palaearctic Region.	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF878E79FDA1FBE5C68CE892.taxon	description	(Figs. 1 A – 1 F)	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF878E79FDA1FBE5C68CE892.taxon	diagnosis	Diagnosis. M fork complete, slightly longer than stem. Male tergite 6 wide and posteriorly with a rounded concavity. Male tergite 7 modified, median process small. A pair of tube-like, pilose structure connected to tergite 7.	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF878E79FDA1FBE5C68CE892.taxon	description	Description. Male (Fig. 1 A). Body length 1.4 – 1.6 mm, wing length 1.5 mm. Head. Dull brown, slightly higher than long; antennae 10 - segmented; flagellum 8 - segmented, a whorl of microtrichia on each flagellomere; last flagellomere elongate, twice as long as preceding one; eyes holoptic; palpus one-segmented, apically acute and pointed at apex. Thorax. Dull brown, stout, and matte; scutum short and dense pilose. Anterior spiracular sclerite small and triangular. Halters slightly darker than scutum, several setae present on stem. Legs concolorous with body; tarsus yellow. Wing (Fig. 1 B). Membrane nearly hyaline; anterior veins sclerotized and light brown, R 1 and R 4 + 5 short, joining costa close to each other, but the distance between the points where R 1 and R 4 + 5 joining the costa larger than usual Swammerdamella species; M fork diverging at the middle of wing and is slightly longer than stem; CuA sigmoid and bent smoothly to posterior margin; a row of macrotrichiae present on posterior margin of wing. Abdomen. Strong, stout and dark brown; segments 1 – 6 visible; sternite 6 (Fig. 1 C) sparsely setose, posterior margin smoothly and widely concave, with a row of setae; tergite 6 (Fig. 1 D) devoid of setae, posterior margin concave. Segment 7 (Fig. 1 F) retracted into segment 6; tergite 7 small, ventral process thin and short, rounded apically, 2 nearly oval lobes connected to tergite 7, with a tube-like, pilose projection; sternite 7 smoothly curved on both margins, with sparse setae on inner margin. Male terminalia (Fig. 1 E). Slightly longer than wide; aedeagus plate oval, sclerotized; aedeagus short; a pair of rod-shaped lobes beside aedeagus plate; another pair of lobes with 2 curves and 3 angles beneath aedeagus plate. Female. Body length 1.6 – 1.8 mm, wing length about 1.5 mm. Similar to male in general morphological characters.	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF878E79FDA1FBE5C68CE892.taxon	materials_examined	Type Material. HOLOTYPE: ♂, China, Gansu Province, Kangxian [DZ 县], Qinghelinchang [ẘñū 场] [105 ° 51 ’ N, 33 ° 12 ’ E, 1456 m], collected by sweep net, 2023. VIII. 26, Liang Wang. (CAU); PARATYPES: 2 ♂ 4 ♀, same data as holotype.	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF878E79FDA1FBE5C68CE892.taxon	distribution	Distribution. China (Gansu Province).	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF878E79FDA1FBE5C68CE892.taxon	etymology	Etymology. This specific name is derived from the Latin adjective “ flexuosus ” which generally means a structure that “ with many curves in it ”, referring to the shape of male terminalia.	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF878E79FDA1FBE5C68CE892.taxon	discussion	Remarks. The M fork of this species is also slightly longer than stem. A pair of unique, tube-like structures are present on male tergite 7, which is different from all other Swammerdamella species. This morphological character is reminiscent of Swammerdamella sessionis Cook, 1971, but the “ trapezoidal sclerites ” of S. sessionis are projected from sternite 7 instead of tergite 7 according to Cook’s description.	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF818E79FDA1FE43C763EC72.taxon	description	Description. Male. Body length 1.6 – 1.9 mm, wing length 1.4 – 1.6 mm. Female. Body length 1.7 – 1.8 mm, wing length 1.3 – 1.5 mm. Morphological characters as in Cook 1956, except the description “ fork of M slightly longer than stem ”. The M fork is as long as or slightly shorter than stem both in Cook’s figure and the specimens we have.	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF818E79FDA1FE43C763EC72.taxon	materials_examined	Material Examined. 1 ♂ 1 ♀, China, Beijing, Huayuancun [AE 园ť] [40 ° 32 ’ N, 117 ° 17 ’ E, 672 m], collected by sweep net, 2018. VI. 3, Ding Yang (CAU); 3 ♂ 1 ♀, China, Beijing, Xiaolongmen National Forest Park [小 ẑm 国 ṽ ŭūȓ 园] [39 ° 97 ’ N, 115 ° 43 ’ E, 1330 m], collected by sweep net, 2023. VI. 13, Chen Lin (CAU); 2 ♂ 1 ♀, China, Liaoning Province, Fushun [Kũ], Lijiacun [ṃṽť] [41 ° 60 ’ N, 124 ° 22 ’ E, 540 m], collected by Malaise trap, 2020. VI. 20, Tao Li (CAU); 1 ♂ 1 ♀, China, Hebei Province, Xinglong [Ẋĸ], Longtanpubu [ẑšȃss] [40 ° 23 ’ N, 117 ° 27 ’ E, 1153 m], collected by sweep net, 2018. VI. 2, Ding Yang (CAU); 3 ♂, China, Hebei Province, Xinglong [Ẋĸ], Mingxi [ḛã] [40 ° 37 ’ N, 117 ° 27 ’ E, 1019 m], collected by sweep net, 2018. VI. 2, Ding Yang (CAU); 1 ♀, China, Hebei Province, Xinglong [Ẋĸ], Xicegou [Ʊ 侧ŝ] [40 ° 33 ’ N, 117 ° 29 ’ E, 1112 m], collected by sweep net, 2018. VI. 2, Ding Yang (CAU); 1 ♀, China, Hebei Province, Xinglong [Ẋĸ], Zhonggusi [Ñăȕ] [40 ° 33 ’ N, 117 ° 29 ’ E, 1023 m], collected by sweep net, 2018. VI. 1, Ding Yang (CAU); 5 ♂ 4 ♀, China, Heilongjiang Province, Heihe [Dz ñ], Pingshan Tree Farm [ṮƜū 场] [49 ° 34 ’ N, 126 ° 56 ’ E, 302 m], collected by sweep net, 2024. VII. 6, Yuetian Gao (CAU); 1 ♂, China, Heilongjiang Province, Jagedaqi [加éż奇], Dongfeng Tree Farm [东 ṇū 场] [50 ° 23 ’ N, 124 ° 6 ’ E, 397 m], collected by sweep net, 2024. VII. 14, Yuetian Gao (CAU).	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF818E79FDA1FE43C763EC72.taxon	distribution	Distribution. China (Beijing; Hebei Province; Heilongjiang Province; Liaoning Province); Finland, France, Germany, Norway, Slovakia, Spain, Sweden, Switzerland (Haenni 2013; Haenni & Carles-Tolrá 2020).	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF818E79FDA1FE43C763EC72.taxon	discussion	Remarks. A widely distributed species in Northern China with 26 specimens from four provinces. S. genypodis can be separated from other Swammerdamella species by the spine-like process on male sternite 7 and a pair of tooth-like projections on male terminalia.	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF818E7BFDA1FA23C410EA0E.taxon	description	(Figs. 2 A – 2 F)	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF818E7BFDA1FA23C410EA0E.taxon	diagnosis	Diagnosis. M fork obviously shorter than stem. A rounded posterior lobe on male tergite 6. Male sternite 6 posteriorly truncated with a row of setae. Male tergite 7 modified, with median process elongated and posteriorly pointed. Aedeagus short.	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF818E7BFDA1FA23C410EA0E.taxon	description	Description. Male (Fig. 2 A). Body length 1.6 – 1.8 mm, wing length 1.2 – 1.5 mm. Head. Dull brown, slightly higher than long; antennae 10 - segmented; flagellum 8 - segmented, each flagellomere with a whorl of microtrichia; last flagellomere distinctly elongate, 3 times longer than preceding one; eyes holoptic; palpus one-segmented, brown, apically acute and pointed at apex. Thorax. Dark brown, stout and shining; scutum short brown pilose. Anterior spiracular sclerite obvious, elongated and triangular. Halters concolorous with scutum, several setae present on stem. Legs short, and mostly concolorous with body; each tibia bicolored and tarsus yellow. Wing (Fig. 2 B). Membrane nearly hyaline; anterior veins sclerotized and yellowish brown, R 1 and R 4 + 5 short, joining costa close to each other; M fork complete and highly visible, divergence at apex of wing; CuA sigmoid, bent abruptly towards posterior margin; a row of macrotrichiae present on posterior margin of wing. Abdomen. Strong, stout and dark brown; segments 1 – 6 visible; sternite 6 (Fig. 2 C) sparsely setose, posterior margin truncated and with a row of setae; tergite 6 (Fig. 2 D) sparsely setose, posteriorly with a rounded lobe and a row of setae on margin. Segment 7 (Fig. 2 F) retracted into segment 6; tergite 7 curved, dorsally setose, both arms rounded apically; ventral process long, bent, and posteriorly pointed; sternite 7 rounded in general shape, inner margin smoothly curved and both arms with several setae. Male terminalia (Fig. 2 E). Subequal in length and width; aedeagus plate rectangular and sclerotized; aedeagus short; two apically rounded lobes present on both sides of aedeagus plate, and with another pair of similar lobes beneath. Female. Body length 1.5 – 1.8 mm, wing length 1.1 – 1.5 mm. Similar to male in general morphological characters.	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF818E7BFDA1FA23C410EA0E.taxon	materials_examined	Type Material. HOLOTYPE: ♂, China, Yunnan Province, Yiliang [ḋǝ], Haiziping Station [DZŦ 坪äaeü] [27 ° 53 ’ N, 104 ° 42 ’ E, 1109 m], collected by sweep net, 2024. VI. 8, Yishen Xiao (CAU). PARATYPES: 83 ♂ 70 ♀, same data as holotype.	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF818E7BFDA1FA23C410EA0E.taxon	distribution	Distribution. China (Yunnan Province).	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF818E7BFDA1FA23C410EA0E.taxon	etymology	Etymology. This specific name is derived from the Latin word “ globus ”, which means “ dense mass ” or “ crowd ”, referring to the mating swarm at the time of the first discovery.	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF818E7BFDA1FA23C410EA0E.taxon	discussion	Remarks. This is a distinctive species discovered in Oriental China, which can be easily distinguished from other Swammerdamella species by the rounded posterior lobe on tergite 6 and posteriorly truncated sternite 6. This species was discovered on a privet (Ligustrum sp.) and a pine tree (Taxaceae) as a large mating swarm of about 1000 individuals.	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF838E75FDA1FCF5C192E892.taxon	description	(Figs. 3 A – 3 F)	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF838E75FDA1FCF5C192E892.taxon	diagnosis	Diagnosis. M fork shorter than stem with M 1 and M 2 close to each other. Male sternite 6 unmodified, posteriorly with a row of setae. Posterior margin of male tergite 7 modified and with a triangular projection. Aedeagus plate sclerotised. Aedeagus long and thin.	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF838E75FDA1FCF5C192E892.taxon	description	Description. Male (Fig. 3 A). Body length 1.6 – 2.0 mm, wing length 1.3 – 1.5 mm. Head. Dull brown, slightly higher than long; antennae 10 - segmented and relatively slender; flagellum 8 - segmented, each flagellomere with a whorl of microtrichia; last flagellomere distinctly elongate, twice as long as preceding one; eyes holoptic; palpus one-segmented, brown, and pointed at apex. Thorax. Dull brown, stout and shining; scutum short yellowish brown pilose. Anterior spiracular sclerite obvious and triangular. Halters darker than scutum, with several setae on stem. Legs short, concolorous with body; tarsus and apical part of hind tibia yellowish brown. Wing (Fig. 3 B). Membrane nearly hyaline; anterior veins sclerotised and yellowish brown, R 1 and R 4 + 5 short, joining costa vein very close to each other; M fork complete and narrow, shorter than stem; CuA sigmoid, bent smoothly towards posterior margin; a row of black microtrichia present on posterior margin of wing. Abdomen. Strong and dull brown; segments 1 – 6 visible; sternite 6 (Fig. 3 C) densely setose, posterior margin unmodified and with a row of setae; tergite 6 (Fig. 3 D) densely setose, posterior margin with an obvious, triangular projection with its margin sclerotised. Segment 7 (Fig. 3 F) retracted into segment 6; tergite 7 thick, folded, with both arms densely setose, ventral process long and rectangular; sternite 7 rounded in general shape, both arms setose, with a highly sclerotised median concavity on inner margin. Male terminalia (Fig. 3 E). Longer than wide; aedeagus plate highly sclerotised and circular concave; aedeagus long and slender, swollen apically. Both sides of aedeagus plate with 2 oval, setose projections. Female. Unknown.	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF838E75FDA1FCF5C192E892.taxon	materials_examined	Type Material. HOLOTYPE: ♂, China, Hubei Province, Shennongjia Forestry District [ö ṝĕū 区], Dajiuhu [大九ª] [31 ° 28 ’ N, 110 ° 1 ’ E, 1767 m], collected by sweep net, 2023. VI. 19, Xingnong Xu (CAU). PARATYPES: 1 ♂ 3 ♀, same data as holotype. Other Material. 2 ♂ 2 ♀, China, Heilongjiang Province, Heihe [Dzñ], Pingshan Tree Farm [ṮƜū 场] [49 ° 34 ’ N, 126 ° 56 ’ E, 302 m], collected by sweep net, 2024. VII. 6, Yuetian Gao (CAU); 4 ♂ 3 ♀, China, Heilongjiang Province, Heihe [Dzñ], Pingshan Tree Farm [ṮƜū 场] [49 ° 34 ’ N, 126 ° 56 ’ E, 302 m], collected by sweep net, 2024. VII. 6, Xingnong Xu (CAU).	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF838E75FDA1FCF5C192E892.taxon	distribution	Distribution. China (Heilongjiang Province; Hubei Province).	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF838E75FDA1FCF5C192E892.taxon	etymology	Etymology. This specific name is derived from the Latin adjective “ gracilis ”, which means “ slender ” or “ thin ”, referring to the shape of aedeagus.	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF838E75FDA1FCF5C192E892.taxon	discussion	Remarks. S. gracilis has distinctive morphological characters that differ from all known Swammerdamella species, and the general shape of male terminalia is a bit similar to the North American species S. confusa Cook, 1956. Surprisingly, this species has a wide distribution among China and was discovered from 31 ° 27 ’ N to 50 ° 7 ’ N, across the Oriental and the Palaearctic Regions. It can be easily recognized by sclerotized aedeagus plate and a remarkably long aedeagus. Segment 7 of male is also highly sclerotized and densely setose.	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF8D8E75FDA1FE43C6E1EEEE.taxon	description	(Figs. 4 A – 4 F)	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF8D8E75FDA1FE43C6E1EEEE.taxon	diagnosis	Diagnosis. M fork complete and obviously shorter than stem. Male tergite and sternite 6 densely setose. Tergite 6 unmodified, posteriorly with a row of setae. Median process of male tergite 7 obvious and rounded apically. Aedeagus short.	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF8D8E75FDA1FE43C6E1EEEE.taxon	description	Description. Male (Fig. 4 A). Body length 1.6 – 1.8 mm, wing length 1.1 – 1.2 mm. Head. Dull brown, higher than long; antennae 9 - segmented; flagellum 7 - segmented, each flagellomere with a whorl of microtrichia; last flagellomere distinctly elongate, twice as long as the preceding one; eyes holoptic; palpus one-segmented, brown, elongate and rounded apically. Thorax. Dull brown, stout and relatively shining; scutum short and brownish pilose. Anterior spiracular sclerite elongate, triangular. Halters concolourous with scutum, and with a row of setae on stem. Legs concolourous with body, except for yellowish-brown tarsi. Wing (Fig. 4 B). Membrane nearly hyaline; anterior veins sclerotised and brown, R 1 and R 4 + 5 short, joining costa close to each other; M fork highly visible and much shorter than its stem; CuA sigmoid, bent abruptly towards posterior margin; a row of macrotrichiae on posterior margin of wing. Abdomen. Strong and dull brown; segments 1 – 6 visible; sternite 6 (Fig. 4 C) densely setose, with a wide and smooth concavity on posterior margin; tergite 6 (Fig. 4 D) densely setose, rectangular, with a row of setae on posterior margin. Segment 7 (Fig. 4 F) retracted into segment 6; tergite 7 modified, dorsally setose, with both arms short and apically acute; ventral process long, apically swollen and rounded, with a tiny triangular cleft on it; sternite 7 ventrally setose and rounded. Male terminalia (Fig. 4 E). Subequal in length and width; aedeagus plate obvious, dark and elongate; two rectangular projections present on both sides of aedeagus plate with several setae on it; a large, triangular lobe stretched beneath, apically triangularly emarginated; aedeagus short. Female. Body length 1.6 – 1.8 mm, wing length about 1.1 mm. Similar to male in general morphological characters.	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF8D8E75FDA1FE43C6E1EEEE.taxon	materials_examined	Type Material. HOLOTYPE: ♂, China, Jilin Province, Hunchun [NJƌ], Yangpao Tree Farm [ěåū 场] [42 ° 53 ’ N, 130 ° 34 ’ E, 150 m], collected by sweep net, 2023. VI. 30, Yishen Xiao (CAU). PARATYPES: 5 ♂ 5 ♀, same data as holotype. Other Material. 122 ♂ 20 ♀, China, Heilongjiang Province, Heihe [Dzñ], Pingshan Tree Farm [ṮƜū 场] [49 ° 34 ’ N, 126 ° 56 ’ E, 302 m], collected by sweep net, 2024. VII. 6, Yuetian Gao (CAU); 1 ♂ 2 ♀, China, Heilongjiang Province, Jagedaqi [加éż奇], Dongfeng Tree Farm [东 ṇū 场] [50 ° 23 ’ N, 124 ° 6 ’ E, 397 m], collected by sweep net, 2024. VII. 14, Yuetian Gao (CAU); 1 ♂, China, Heilongjiang Province, Tahe [塔ñ], Shibazhan [十Āü] [52 ° 25 ’ N, 124 ° 24 ’ E, 325 m], collected by sweep net, 2024. VII. 11, Yuetian Gao (CAU); 1 ♂ 4 ♀, China, Jilin Province, Wangqing [ĩdz], Dangshuhe Tree Farm [IJŃñū 场] [43 ° 31 ’ N, 129 ° 57 ’ E, 457 m], collected by sweep net, 2023. VII. 4, Yishen Xiao (CAU); 1 ♂, China, Liaoning Province, Fushun [Kũ], Lijiacun [ṃṽť] [41 ° 60 ’ N, 124 ° 22 ’ E, 540 m], collected by Malaise trap, 2020. VI. 20, Tao Li (CAU); 1 ♀, China, Liaoning Province, Fuxin [şǔ], Zhangwu [dz ẋ] [42 ° 37 ’ N, 122 ° 54 ’ E, 83 m], collected by Malaise trap, 2020. VI. 18, Tao Li (CAU).	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF8D8E75FDA1FE43C6E1EEEE.taxon	distribution	Distribution. China (Heilongjiang Province, Jilin Province, Liaoning Province).	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF8D8E75FDA1FE43C6E1EEEE.taxon	etymology	Etymology. This specific name is derived from the Latin adjective “ latus ” and noun “ lobus ”, which means “ wide ” and “ hull ”, referring to the large, concave lobe of male terminalia.	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF8D8E75FDA1FE43C6E1EEEE.taxon	discussion	Remarks. This is a widely distributed species in Northeast China with about 150 specimen records. The antennae of S. latilobata are 9 - segmented, the same as some other Palaearctic species like S. acuta, S. adercotris and S. brevicornis, but the terminalia of S. latilobata are distinct from them.	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF8C8E76FDA1F8A6C647EAF6.taxon	description	(Figs. 5 A – 5 F)	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF8C8E76FDA1F8A6C647EAF6.taxon	diagnosis	Diagnosis. M fork complete and shorter than stem. The posterior margin of male tergite and sternite 6 modified, both truncated. Median process of male tergite 7 obvious and posteriorly pointed. Aedeagus large and sclerotized.	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF8C8E76FDA1F8A6C647EAF6.taxon	description	Description. Male (Fig. 5 A). Body length 1.5 – 1.9 mm, wing length 1.4 – 1.7 mm. Head. Dark brown, slightly higher than long; antennae 10 - segmented; flagellum 8 - segmented, each flagellomere with a whorl of microtrichia; last flagellomere distinctly elongate, twice as long as preceding one; eyes holoptic; palpus one-segmented, brown, apically rounded and pointed at apex. Thorax. Dark brown and shining; scutum short, brown pilose. Anterior spiracular sclerite elongated and triangular. Halters concolorous with scutum, stem with several setae. Legs short, and concolorous with body. Wing (Fig. 5 B). Membrane nearly hyaline; anterior veins sclerotized and brown, R 1 and R 4 + 5 short, joining the costa close to each other; M fork complete but hardly visible, divergence near the apex of wing; CuA sigmoid and bent abruptly to posterior margin; a sparse row of macrotrichiae on posterior margin of wing. Abdomen. Strong, stout and dark brown; segments 1 – 6 visible; sternite 6 (Fig. 5 C) sparsely setose, posterior margin truncated and devoid of setae; tergite 6 (Fig. 5 D) sparsely setose, posterior margin with a rectangular projection and devoid of setae. Segment 7 (Fig. 5 F) retracted into segment 6; tergite 7 highly modified, with a long, triangular, and posteriorly pointed projection; both arms short and apically acute; sternite 7 rounded, with a shallow, rectangular median concavity on inner margin. Male terminalia (Fig. 5 E). Subequal in length and width; aedeagus plate oval and dark; aedeagus long, sclerotized and truncate apically; a pair of elongate, acute lobes beside aedeagus plate. Female. Body length 1.6 – 1.8 mm, wing length about 1.5 mm. Similar to male in general morphological characters.	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF8C8E76FDA1F8A6C647EAF6.taxon	materials_examined	Type Material. HOLOTYPE: ♂, China, Yunnan Province, Kunming [ḆOi], Changchongshan [ẗƦƜ] [27 ° 10 ’ N, 102 ° 44 ’ E, 2005 m], collected by sweep net, 2024. VI. 10, Yishen Xiao (CAU). PARATYPES: 3 ♂ 3 ♀, same data as holotype.	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF8C8E76FDA1F8A6C647EAF6.taxon	distribution	Distribution. China (Yunnan Province).	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF8C8E76FDA1F8A6C647EAF6.taxon	etymology	Etymology. This specific name is derived from the Latin adjective “ papillatus ”, which means “ teat-like ”, referring to the shape of male tergite 6.	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF8C8E76FDA1F8A6C647EAF6.taxon	discussion	Remarks. This species can be easily recognized by the elongate and posteriorly pointed ventral process of male tergite 7, and a triangular truncated projection on posterior margin of tergite 6. This species shares the same posteriorly pointed median process with S. globus. S. papillata is another species discovered in the Oriental Region, collected from a dry mountain in Kunming, Yunnan Province.	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF8E8E76FDA1FBADC11DEF71.taxon	description	Description. Male. Body length 1.5 – 1.6 mm, wing length 1.1 – 1.2 mm. Female. Body length 1.6 – 1.7 mm, wing length 1.3 mm. Morphological characters as in Cook 1956.	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF8E8E76FDA1FBADC11DEF71.taxon	materials_examined	Material Examined. 2 ♂ 1 ♀, China, Heilongjiang Province, Heihe [Dzñ], Pingshan Tree Farm [ṮƜū 场] [49 ° 34 ’ N, 126 ° 56 ’ E, 302 m], collected by sweep net, 2024. VII. 5, Yuetian Gao (CAU); 15 ♂ 10 ♀, China, Liaoning Province, Fushun [Kũ], Lijiacun [ṃṽť] [41 ° 60 ’ N, 124 ° 22 ’ E, 540 m], collected by Malaise trap, 2020. VI. 20, Tao Li (CAU).	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF8E8E76FDA1FBADC11DEF71.taxon	distribution	Distribution. China (Heilongjiang Province, Liaoning Province); Germany, Hungary, Russia, Slovakia, Sweden, Switzerland.	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF8E8E76FDA1FBADC11DEF71.taxon	discussion	Remarks. The cup-shaped process on male tergite 7 and a highly sclerotized aedeagus can easily distinguish this species. S. pediculata has been recorded in several European countries (Haenni 2013) since the original description and the specimens examined in this paper were collected from Northeast China.	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF8E8E70FDA1F92FC658EAD2.taxon	description	(Figs. 6 A – 6 F)	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF8E8E70FDA1F92FC658EAD2.taxon	diagnosis	Diagnosis. M fork complete, slightly longer than stem. Posterior margin of male tergite 6 unmodified. Male tergite 7 with a small ventral process and a pair of lobes. Aedeagus obvious, flattened and strongly sclerotized.	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF8E8E70FDA1F92FC658EAD2.taxon	description	Description. Male (Fig. 6 A). Body length 1.8 – 2.2 mm, wing length 2.0 – 2.1 mm. Head. Dull brown, slightly higher than long; antennal flagellum 8 - segmented, with a whorl of microtrichia on each flagellomere; last flagellomere elongate, approximately twice as long as preceding one; eyes holoptic; palpus one-segmented, brown, apically acute and pointed at apex. Thorax. Dull brown, stout, and matte; scutum densely short pilose. Anterior spiracular sclerite relatively small, short and triangular. Halters concolourous with scutum, stem with a row of setae. Legs short, and mostly concolourous with body, except for yellowish brown hind tarsus and distal hind tibia. Wing (Fig. 6 B). Membrane nearly hyaline; anterior veins sclerotized and light brown, R 1 and R 4 + 5 short, joining costa close to each other; M fork divergent at middle of wing and slightly longer than stem; CuA sigmoid and gently bent to posterior margin; only few macrotrichiae present on posterior margin of wing. Abdomen. Strong, stout and dark brown; segments 1 – 6 visible; sternite 6 (Fig. 6 C) sparsely setose, posterior margin smoothly concave; tergite 6 (Fig. 6 D) sparsely setose, posterior margin flat and unmodified. Segment 7 (Fig. 6 F) retracted into segment 6; tergite 7 modified, ventral process small, rounded apically, with a pair of oval lobes on both sides; sternite 7 rounded. Male terminalia (Fig. 6 E). Slightly longer than wide; aedeagus oval, dark and flattened, basally with 2 symmetric processes; a large and apically twisted projection present beneath aedeagus. Female. Unknown.	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF8E8E70FDA1F92FC658EAD2.taxon	materials_examined	Type Material. HOLOTYPE: ♂, China, Gansu Province, Tanchang [âē], Zedangcuo [āīDz] [104 ° 22 ’ N, 33 ° 52 ’ E, 2649 m], collected by sweep net, 2024. VIII. 1, Yishen Xiao (CAU). PARATYPES: 2 ♂, same data as holotype.	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF8E8E70FDA1F92FC658EAD2.taxon	distribution	Distribution. China (Gansu Province).	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF8E8E70FDA1F92FC658EAD2.taxon	etymology	Etymology. This specific name is derived from the Latin adjective “ planus ”, which means “ flat ”, referring to the flattened aedeagus.	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF8E8E70FDA1F92FC658EAD2.taxon	discussion	Remarks. This species can be readily distinguished from most species of this genus by the length of M fork, which is slightly longer than its stem. S. planus is confirmed to belong to Swammerdamella because of the typically concealed, modified tergite 7 and adjacent R 1 and R 4 + 5. The male tergite 7 of this species is also highly recognizable by the short and small ventral process and a pair of oval lobes.	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF888E72FDA1FB89C02CE8CA.taxon	description	(Figs. 7 A – 7 E)	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF888E72FDA1FB89C02CE8CA.taxon	diagnosis	Diagnosis. M fork complete and shorter than stem. Sternite 6 of male unmodified, with a row of setae on posterior margin. Male tergite 7 devoid of median process and with 3 concavities. Aedeagus long, sclerotised and thin.	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF888E72FDA1FB89C02CE8CA.taxon	description	Description. Male (Fig. 7 A). Body length about 1.3 mm, wing length 1.1 mm. Head. Dull brown, higher than long; antennae 10 - segmented; flagellum 8 - segmented, each flagellomere with a whorl of microtrichia; last flagellomere distinctly elongate, twice as long as preceding one, with about 3 whorls of microtrichia; eyes holoptic; palpus one-segmented, light brown, and elongate. Thorax. Dull brown, stout and matte; scutum densely pilose. Anterior spiracular sclerite triangular. Halters concolorous with scutum, stem with a row of setae. Legs short, mostly concolorous with body except for the yellowish-brown tarsi. Wing (Fig. 7 B). Membrane nearly hyaline; anterior veins sclerotized and light brown, R 1 and R 4 + 5 joining costa close to each other; M fork complete but hardly visible, shorter than stem of M; CuA sigmoid, smoothly bent; a sparse row of macrotrichiae on whole posterior margin of wing. Abdomen. Strong, stout and dull brown; segments 1 – 6 visible; sternite 6 (Fig. 7 C) wide, sparsely setose, posterior margin unmodified; tergite 6 missing during the dissection, but obviously with a relatively unmodified posterior margin (Fig. 7 A). Segment 7 (Fig. 7 E) retracted into segment 6; tergite 7 without setae and ventral process, lateral arms wide and rounded apically; sternite 7 rounded, inner margin setose, with 2 symmetric angles and a shallow median concavity. Male terminalia (Fig. 7 D). Longer than wide; aedeagus thin, long, and darker than other parts, rounded apically and basally with 2 sharp angles. Two lobes dorsally pointed, sclerotized, and bifurcated apically, forming a pair of twig-like projections. Female. Unknown.	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF888E72FDA1FB89C02CE8CA.taxon	materials_examined	Type Material. HOLOTYPE: ♂, China, Guangxi Province, Leye [乐业], Yachang Nature Reserve [ǎ ẗnẍ 保ae区] [24 ° 50 ’ N, 106 ° 12 ’ E, 612 m], collected by sweep net, 2024. VIII. 8, Wei Zeng (CAU).	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF888E72FDA1FB89C02CE8CA.taxon	distribution	Distribution. China (Guangxi Province).	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF888E72FDA1FB89C02CE8CA.taxon	etymology	Etymology. This specific name is derived from the Latin noun “ ramulus ”, which means “ twig ” or “ little branch ”, referring to the twig-like projections of male terminalia.	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF888E72FDA1FB89C02CE8CA.taxon	discussion	Remarks. S. ramulosa is another new species from the Oriental Region and can be distinguished from other Swammerdamella species by the characteristics of tergite 7. The shape of tergite 7 is somewhat twisted and concave, with a large concavity in the middle and other 2 smaller ones on both sides, and without ventral process.	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF8A8E72FDA1FDB1C7C2EE1E.taxon	description	(Figs. 8 A – 8 F)	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF8A8E72FDA1FDB1C7C2EE1E.taxon	diagnosis	Diagnosis. Sternite 6 of male triangularly concave. Male tergite 6 unmodified and posteriorly with a row of setae. Arms of male tergite 7 long and thin, apically acute. Three spines on the position of median process. Aedeagus long and spiral.	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF8A8E72FDA1FDB1C7C2EE1E.taxon	description	Description. Male (Fig. 8 A). Body length 1.4 – 1.5 mm, wing length 1.2 – 1.3 mm. Head. Dull brown, higher than long; antennae 10 - segmented; flagellum 8 - segmented, each flagellomere with a whorl of microtrichia; last flagellomere distinctly elongate, 3 times longer than preceding one; eyes holoptic; palpus one-segmented, yellowish brown, elongate and pointed at apex. Thorax. Dull brown, stout and matte; scutum brownish pilose. Anterior spiracular sclerite triangular. Halters concolorous with scutum, with a row of setae on stem. Legs short, and mostly concolorous with body except for yellowish-brown tarsi. Wing (Fig. 8 B). Membrane nearly hyaline; anterior veins sclerotized and light brown, R 1 and R 4 + 5 joining costa vein very close to each other; M fork complete and visible, obviously shorter than stem of M; CuA sigmoid, bent abruptly; a row of macrotrichiae on the posterior margin. Abdomen. Strong, stout, and dull brown; segments 1 – 6 visible; sternite 6 (Fig. 8 C) wide, sparsely setose, with posterior margin triangularly concave; tergite 6 (Fig. 8 D) wide, sparsely setose, with posterior margin smoothly curved and with a row of setae. Segment 7 (Fig. 8 F) retracted into segment 6; tergite 7 devoid of setae, both arms thin and narrow, apically acute, 3 spines in the middle of tergite 7 as ventral process; sternite 7 rounded and both arms setose apically, with 2 symmetrical sclerotised, dark-colored areas on inner margin. Male terminalia (Fig. 8 E). Slightly longer than wide; aedeagus long, spiral, and darker than other parts; a large lobe folded posteriorly on both sides, forming a semi-enclosed structure around the base of aedeagus; another large, densely pilose projection posteriorly stretched with a deep concavity on it. Female. Unknown.	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF8A8E72FDA1FDB1C7C2EE1E.taxon	materials_examined	Type Material. HOLOTYPE: ♂, China, Jilin Province, Helong [和 ẑ], Wolong Station [卧 ẑäaeü] [42 ° 46 ’ N, 128 ° 45 ’ E, 737 m], collected by sweep net, 2023. VII. 6, Yishen Xiao (CAU). PARATYPE: ♂, same data as holotype.	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF8A8E72FDA1FDB1C7C2EE1E.taxon	distribution	Distribution. China (Jilin Province).	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF8A8E72FDA1FDB1C7C2EE1E.taxon	etymology	Etymology. This specific name is derived from the Latin adjective “ tricuspis ”, meaning “ which has three points ”, referring to the shape of the ventral process on male tergite 7.	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF8A8E72FDA1FDB1C7C2EE1E.taxon	discussion	Remarks. This species shows some common Swammerdamella characteristics on wing and body, but the shape of male tergite 7 and terminalia are relatively unique and highly recognizable. The terminalia is also morphologically different from all other Swammerdamella species. The structure of male terminalia is simple, only with a long, spiral aedeagus while the aedeagus plate seems missing.	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF948E6FFDA1FF04C45DEDBA.taxon	description	(Figs. 9 A – 9 F)	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF948E6FFDA1FF04C45DEDBA.taxon	diagnosis	Diagnosis. M fork complete, slightly longer than stem. Posterior margin of male sternite 6 smoothly concave. Male tergite 6 setose, with its posterior margin truncated. A wide and trapezoidal median process on male tergite 7. Aedeagus oval and obvious.	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF948E6FFDA1FF04C45DEDBA.taxon	description	Description. Male (Fig. 9 A). Body length about 1.6 mm, wing length 1.9 mm. Head. Yellowish brown, slightly higher than long; antennae 10 - segmented; flagellum 8 - segmented, each flagellomere with a whorl of microtrichia; last flagellomere distinctly elongate, 3 times longer than preceding one; eyes holoptic, meeting over antennae; palpus one-segmented, light brown, elongate and pointed at apex. Thorax. Yellowish brown, stout and shining; scutum darker colored, with sparse and brownish setae. Anterior spiracular sclerite relatively small, triangular. Halters concolorous with scutum, with a row of setae on stem. Legs short, concolorous with body, distal part of tibia slightly lighter than its basal part. Wing (Fig. 9 B). Membrane nearly hyaline; anterior veins sclerotized and light brown, R 1 and R 4 + 5 joining costa close to each other; M fork complete, highly visible, slightly longer than stem; CuA sigmoid, smoothly angled; a row of macrotrichiae on posterior margin. Abdomen. Strong and stout, tergites and sternites dull brown; segments 1 – 6 visible; sternite 6 (Fig. 9 C) wide, setose and smoothly concave on its posterior margin; tergite 6 (Fig. 9 D) relatively rectangular, setose, truncated posteriorly. Segment 7 (Fig. 9 F) retracted into segment 6; tergite 7 with thin lateral arms and apically acute, ventral process wide and trapezoidal; sternite 7 rounded, inner margin smoothly rounded and sparsely setose, ending laterally with 2 apically truncated arms. Male terminalia (Fig. 9 E). Slightly longer than wide; aedeagus plate oval, obviously darker than other parts, with 2 oval lobes on each side; a pair of thin, slender lobes beneath the aedeagus, elongate, dorsally pointed, and gently twisted apically like a hook. Female. Unknown.	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF948E6FFDA1FF04C45DEDBA.taxon	materials_examined	Type Material. HOLOTYPE: ♂, China, Hubei Province, Shennongjia Forestry District [ö ṝĕū 区], Shennongding [ö ṝȗ] [31 ° 26 ’ N, 110 ° 18 ’ E, 2394 m], collected by Malaise trap, 2022. VIII. 26, Siqi Wang & Bintao Du (CAU).	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF948E6FFDA1FF04C45DEDBA.taxon	distribution	Distribution. China (Hubei Province).	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF948E6FFDA1FF04C45DEDBA.taxon	etymology	Etymology. This specific name is derived from the Latin adjective “ venustus ”, which means “ charming ”, referring to the unusual morphological character of wings and the elegant shape of male terminalia.	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
DF3B3510FF948E6FFDA1FF04C45DEDBA.taxon	discussion	Remarks. S. venusta is another species from the Oriental Region. This species shares the unique elongated M fork with S. planus sp. nov. and S. flexuosa sp. nov. mentioned above. S. venusta can be easily distinguished from other species in this genus by the delicately curved male terminalia and the trapezoidal, truncated ventral process of male tergite 7.	en	Xiao, Yishen, Hong, Dawei, Li, Zhu, Yang, Ding (2025): First report of the genus Swammerdamella Enderlein, 1912 (Diptera: Scatopsidae) in China, with descriptions of nine new species. Zootaxa 5723 (1): 49-70, DOI: 10.11646/zootaxa.5723.1.2, URL: https://doi.org/10.11646/zootaxa.5723.1.2
