taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
DB0F87E19114F11BA7D3FD8A7ED7780C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12189910/files/figure.png	https://doi.org/10.5281/zenodo.12189910	FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.	FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.	2024-06-18	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.		Zenodo	biologists	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.			
DB0F87E19114F11BA7D3FD8A7ED7780C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12189912/files/figure.png	https://doi.org/10.5281/zenodo.12189912	FIGURE3. Maleappendages of Coenagrion spp.ofthe puella-groupin thebroadsense inlateral view:a–c—C.australocaspicum (b—with the appendages expanded); d–k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–q—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; k—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.	FIGURE3. Maleappendages of Coenagrion spp.ofthe puella-groupin thebroadsense inlateral view:a–c—C.australocaspicum (b—with the appendages expanded); d–k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–q—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; k—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.	2024-06-18	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.		Zenodo	biologists	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.			
DB0F87E19114F11BA7D3FD8A7ED7780C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12189914/files/figure.png	https://doi.org/10.5281/zenodo.12189914	FIGURE 4. Male appendages of Coenagrion spp. of the puella-group in the broad sense in posterior view: a–b—C. australocaspicum (b—with the appendages expanded); c–d—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); e-f—C. pulchellum; g–k—C. puella; l—C. persicum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c–d—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; e—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; f—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; g–h—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; i—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, a brook and pond, 25.06.2021; j—Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017. Scale bar 0.5 mm.	FIGURE 4. Male appendages of Coenagrion spp. of the puella-group in the broad sense in posterior view: a–b—C. australocaspicum (b—with the appendages expanded); c–d—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); e-f—C. pulchellum; g–k—C. puella; l—C. persicum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c–d—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; e—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; f—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; g–h—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; i—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, a brook and pond, 25.06.2021; j—Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017. Scale bar 0.5 mm.	2024-06-18	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.		Zenodo	biologists	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.			
DB0F87E19114F11BA7D3FD8A7ED7780C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12189924/files/figure.png	https://doi.org/10.5281/zenodo.12189924	FIGURE 7. Female pronotum of Coenagrion spp. of the puella-group in the broad sense, a–h—in dorsal, with respect to the body main axis (above), and anteriodorsal, perpendicular to the posterior lobe surface (below), views; i–j—in dorsal view only; a—C. australocaspicum; b–g—C. puella; h—C. ponticum; i—C. persicum; j—C. pulchellum; a—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; b—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; c—Sergokala District, 3 km SW of Myurego village, the Inchkheozen’ River upper valley, 19.06.2021; d–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; h—Russia, Chelyabinsk Province, Nagaybakskiy District, small oxbow lakes at the Gumbeyka River left bank at Ostrolenskiy Settlement SE margin, 6.07.2021; i—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; j—Iran, Markazi Ostan, Shazand Shakhrestan, the Garehchay River at Gavigodar village, 34.1078-1111° N 49.3627-3657° E, 1803–1805 m a.s.l., 19.05.2017; k—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.	FIGURE 7. Female pronotum of Coenagrion spp. of the puella-group in the broad sense, a–h—in dorsal, with respect to the body main axis (above), and anteriodorsal, perpendicular to the posterior lobe surface (below), views; i–j—in dorsal view only; a—C. australocaspicum; b–g—C. puella; h—C. ponticum; i—C. persicum; j—C. pulchellum; a—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; b—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; c—Sergokala District, 3 km SW of Myurego village, the Inchkheozen’ River upper valley, 19.06.2021; d–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; h—Russia, Chelyabinsk Province, Nagaybakskiy District, small oxbow lakes at the Gumbeyka River left bank at Ostrolenskiy Settlement SE margin, 6.07.2021; i—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; j—Iran, Markazi Ostan, Shazand Shakhrestan, the Garehchay River at Gavigodar village, 34.1078-1111° N 49.3627-3657° E, 1803–1805 m a.s.l., 19.05.2017; k—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.	2024-06-18	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.		Zenodo	biologists	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.			
DB0F87E19114F11BA7D3FD8A7ED7780C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12189916/files/figure.png	https://doi.org/10.5281/zenodo.12189916	FIGURE 5. Male pronotum of Coenagrion spp. of the puella-group in the broad sense in dorsal view: a—C. persicum; b–c—C. pulchellum; d–e—C. ponticum (males with a more (d) and less (e) expanded black abdominal maculation); f—C. australocaspicum; g–l—C. puella; a—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; b—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; c—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; d–e—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; f—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; g —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; h–i—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021; j—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; k—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; l—Russia, Karachay- Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019. Scale bar 0.5 mm.	FIGURE 5. Male pronotum of Coenagrion spp. of the puella-group in the broad sense in dorsal view: a—C. persicum; b–c—C. pulchellum; d–e—C. ponticum (males with a more (d) and less (e) expanded black abdominal maculation); f—C. australocaspicum; g–l—C. puella; a—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; b—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; c—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; d–e—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; f—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; g —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; h–i—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021; j—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; k—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; l—Russia, Karachay- Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019. Scale bar 0.5 mm.	2024-06-18	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.		Zenodo	biologists	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.			
DB0F87E19117F118A7D3F9B97E6D7D76.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12189924/files/figure.png	https://doi.org/10.5281/zenodo.12189924	FIGURE 7. Female pronotum of Coenagrion spp. of the puella-group in the broad sense, a–h—in dorsal, with respect to the body main axis (above), and anteriodorsal, perpendicular to the posterior lobe surface (below), views; i–j—in dorsal view only; a—C. australocaspicum; b–g—C. puella; h—C. ponticum; i—C. persicum; j—C. pulchellum; a—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; b—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; c—Sergokala District, 3 km SW of Myurego village, the Inchkheozen’ River upper valley, 19.06.2021; d–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; h—Russia, Chelyabinsk Province, Nagaybakskiy District, small oxbow lakes at the Gumbeyka River left bank at Ostrolenskiy Settlement SE margin, 6.07.2021; i—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; j—Iran, Markazi Ostan, Shazand Shakhrestan, the Garehchay River at Gavigodar village, 34.1078-1111° N 49.3627-3657° E, 1803–1805 m a.s.l., 19.05.2017; k—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.	FIGURE 7. Female pronotum of Coenagrion spp. of the puella-group in the broad sense, a–h—in dorsal, with respect to the body main axis (above), and anteriodorsal, perpendicular to the posterior lobe surface (below), views; i–j—in dorsal view only; a—C. australocaspicum; b–g—C. puella; h—C. ponticum; i—C. persicum; j—C. pulchellum; a—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; b—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; c—Sergokala District, 3 km SW of Myurego village, the Inchkheozen’ River upper valley, 19.06.2021; d–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; h—Russia, Chelyabinsk Province, Nagaybakskiy District, small oxbow lakes at the Gumbeyka River left bank at Ostrolenskiy Settlement SE margin, 6.07.2021; i—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; j—Iran, Markazi Ostan, Shazand Shakhrestan, the Garehchay River at Gavigodar village, 34.1078-1111° N 49.3627-3657° E, 1803–1805 m a.s.l., 19.05.2017; k—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.	2024-06-18	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.		Zenodo	biologists	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.			
DB0F87E19117F118A7D3F9B97E6D7D76.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12189910/files/figure.png	https://doi.org/10.5281/zenodo.12189910	FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.	FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.	2024-06-18	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.		Zenodo	biologists	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.			
DB0F87E19117F118A7D3F9B97E6D7D76.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12189912/files/figure.png	https://doi.org/10.5281/zenodo.12189912	FIGURE3. Maleappendages of Coenagrion spp.ofthe puella-groupin thebroadsense inlateral view:a–c—C.australocaspicum (b—with the appendages expanded); d–k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–q—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; k—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.	FIGURE3. Maleappendages of Coenagrion spp.ofthe puella-groupin thebroadsense inlateral view:a–c—C.australocaspicum (b—with the appendages expanded); d–k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–q—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; k—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.	2024-06-18	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.		Zenodo	biologists	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.			
DB0F87E19117F118A7D3F9B97E6D7D76.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12189914/files/figure.png	https://doi.org/10.5281/zenodo.12189914	FIGURE 4. Male appendages of Coenagrion spp. of the puella-group in the broad sense in posterior view: a–b—C. australocaspicum (b—with the appendages expanded); c–d—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); e-f—C. pulchellum; g–k—C. puella; l—C. persicum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c–d—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; e—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; f—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; g–h—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; i—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, a brook and pond, 25.06.2021; j—Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017. Scale bar 0.5 mm.	FIGURE 4. Male appendages of Coenagrion spp. of the puella-group in the broad sense in posterior view: a–b—C. australocaspicum (b—with the appendages expanded); c–d—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); e-f—C. pulchellum; g–k—C. puella; l—C. persicum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c–d—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; e—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; f—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; g–h—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; i—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, a brook and pond, 25.06.2021; j—Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017. Scale bar 0.5 mm.	2024-06-18	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.		Zenodo	biologists	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.			
DB0F87E19117F118A7D3F9B97E6D7D76.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12189916/files/figure.png	https://doi.org/10.5281/zenodo.12189916	FIGURE 5. Male pronotum of Coenagrion spp. of the puella-group in the broad sense in dorsal view: a—C. persicum; b–c—C. pulchellum; d–e—C. ponticum (males with a more (d) and less (e) expanded black abdominal maculation); f—C. australocaspicum; g–l—C. puella; a—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; b—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; c—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; d–e—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; f—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; g —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; h–i—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021; j—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; k—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; l—Russia, Karachay- Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019. Scale bar 0.5 mm.	FIGURE 5. Male pronotum of Coenagrion spp. of the puella-group in the broad sense in dorsal view: a—C. persicum; b–c—C. pulchellum; d–e—C. ponticum (males with a more (d) and less (e) expanded black abdominal maculation); f—C. australocaspicum; g–l—C. puella; a—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; b—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; c—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; d–e—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; f—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; g —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; h–i—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021; j—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; k—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; l—Russia, Karachay- Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019. Scale bar 0.5 mm.	2024-06-18	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.		Zenodo	biologists	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.			
DB0F87E19119F10EA7D3FF1F7F207CEF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12189910/files/figure.png	https://doi.org/10.5281/zenodo.12189910	FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.	FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.	2024-06-18	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.		Zenodo	biologists	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.			
DB0F87E19119F10EA7D3FF1F7F207CEF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12189912/files/figure.png	https://doi.org/10.5281/zenodo.12189912	FIGURE3. Maleappendages of Coenagrion spp.ofthe puella-groupin thebroadsense inlateral view:a–c—C.australocaspicum (b—with the appendages expanded); d–k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–q—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; k—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.	FIGURE3. Maleappendages of Coenagrion spp.ofthe puella-groupin thebroadsense inlateral view:a–c—C.australocaspicum (b—with the appendages expanded); d–k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–q—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; k—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.	2024-06-18	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.		Zenodo	biologists	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.			
DB0F87E19119F10EA7D3FF1F7F207CEF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12189914/files/figure.png	https://doi.org/10.5281/zenodo.12189914	FIGURE 4. Male appendages of Coenagrion spp. of the puella-group in the broad sense in posterior view: a–b—C. australocaspicum (b—with the appendages expanded); c–d—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); e-f—C. pulchellum; g–k—C. puella; l—C. persicum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c–d—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; e—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; f—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; g–h—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; i—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, a brook and pond, 25.06.2021; j—Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017. Scale bar 0.5 mm.	FIGURE 4. Male appendages of Coenagrion spp. of the puella-group in the broad sense in posterior view: a–b—C. australocaspicum (b—with the appendages expanded); c–d—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); e-f—C. pulchellum; g–k—C. puella; l—C. persicum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c–d—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; e—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; f—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; g–h—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; i—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, a brook and pond, 25.06.2021; j—Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017. Scale bar 0.5 mm.	2024-06-18	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.		Zenodo	biologists	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.			
DB0F87E19119F10EA7D3FF1F7F207CEF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12189916/files/figure.png	https://doi.org/10.5281/zenodo.12189916	FIGURE 5. Male pronotum of Coenagrion spp. of the puella-group in the broad sense in dorsal view: a—C. persicum; b–c—C. pulchellum; d–e—C. ponticum (males with a more (d) and less (e) expanded black abdominal maculation); f—C. australocaspicum; g–l—C. puella; a—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; b—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; c—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; d–e—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; f—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; g —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; h–i—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021; j—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; k—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; l—Russia, Karachay- Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019. Scale bar 0.5 mm.	FIGURE 5. Male pronotum of Coenagrion spp. of the puella-group in the broad sense in dorsal view: a—C. persicum; b–c—C. pulchellum; d–e—C. ponticum (males with a more (d) and less (e) expanded black abdominal maculation); f—C. australocaspicum; g–l—C. puella; a—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; b—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; c—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; d–e—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; f—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; g —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; h–i—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021; j—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; k—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; l—Russia, Karachay- Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019. Scale bar 0.5 mm.	2024-06-18	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.		Zenodo	biologists	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.			
DB0F87E19119F10EA7D3FF1F7F207CEF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12189920/files/figure.png	https://doi.org/10.5281/zenodo.12189920	FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale.	FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale.	2024-06-18	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.		Zenodo	biologists	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.			
DB0F87E19119F10EA7D3FF1F7F207CEF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12189924/files/figure.png	https://doi.org/10.5281/zenodo.12189924	FIGURE 7. Female pronotum of Coenagrion spp. of the puella-group in the broad sense, a–h—in dorsal, with respect to the body main axis (above), and anteriodorsal, perpendicular to the posterior lobe surface (below), views; i–j—in dorsal view only; a—C. australocaspicum; b–g—C. puella; h—C. ponticum; i—C. persicum; j—C. pulchellum; a—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; b—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; c—Sergokala District, 3 km SW of Myurego village, the Inchkheozen’ River upper valley, 19.06.2021; d–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; h—Russia, Chelyabinsk Province, Nagaybakskiy District, small oxbow lakes at the Gumbeyka River left bank at Ostrolenskiy Settlement SE margin, 6.07.2021; i—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; j—Iran, Markazi Ostan, Shazand Shakhrestan, the Garehchay River at Gavigodar village, 34.1078-1111° N 49.3627-3657° E, 1803–1805 m a.s.l., 19.05.2017; k—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.	FIGURE 7. Female pronotum of Coenagrion spp. of the puella-group in the broad sense, a–h—in dorsal, with respect to the body main axis (above), and anteriodorsal, perpendicular to the posterior lobe surface (below), views; i–j—in dorsal view only; a—C. australocaspicum; b–g—C. puella; h—C. ponticum; i—C. persicum; j—C. pulchellum; a—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; b—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; c—Sergokala District, 3 km SW of Myurego village, the Inchkheozen’ River upper valley, 19.06.2021; d–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; h—Russia, Chelyabinsk Province, Nagaybakskiy District, small oxbow lakes at the Gumbeyka River left bank at Ostrolenskiy Settlement SE margin, 6.07.2021; i—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; j—Iran, Markazi Ostan, Shazand Shakhrestan, the Garehchay River at Gavigodar village, 34.1078-1111° N 49.3627-3657° E, 1803–1805 m a.s.l., 19.05.2017; k—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.	2024-06-18	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.		Zenodo	biologists	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.			
DB0F87E19119F10EA7D3FF1F7F207CEF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12189908/files/figure.png	https://doi.org/10.5281/zenodo.12189908	FIGURE 1. Explanation of terminology (see the text) used for morphological details of male terminalia of Coenagrion spp. of the puella-group in the broad sense used in this paper, on examples of a male specimen of C. australocaspicum with an unusual expanded position of the cerci in lateral view (left, the same as Fig. 3a) and a male of C. persicum in dorsal view (right, the same as Fig. 2n).	FIGURE 1. Explanation of terminology (see the text) used for morphological details of male terminalia of Coenagrion spp. of the puella-group in the broad sense used in this paper, on examples of a male specimen of C. australocaspicum with an unusual expanded position of the cerci in lateral view (left, the same as Fig. 3a) and a male of C. persicum in dorsal view (right, the same as Fig. 2n).	2024-06-18	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.		Zenodo	biologists	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.			
DB0F87E19101F10DA7D3FBDE78C47FCF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12189910/files/figure.png	https://doi.org/10.5281/zenodo.12189910	FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.	FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.	2024-06-18	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.		Zenodo	biologists	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.			
DB0F87E19101F10DA7D3FBDE78C47FCF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12189912/files/figure.png	https://doi.org/10.5281/zenodo.12189912	FIGURE3. Maleappendages of Coenagrion spp.ofthe puella-groupin thebroadsense inlateral view:a–c—C.australocaspicum (b—with the appendages expanded); d–k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–q—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; k—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.	FIGURE3. Maleappendages of Coenagrion spp.ofthe puella-groupin thebroadsense inlateral view:a–c—C.australocaspicum (b—with the appendages expanded); d–k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–q—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; k—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.	2024-06-18	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.		Zenodo	biologists	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.			
DB0F87E19101F10DA7D3FBDE78C47FCF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12189914/files/figure.png	https://doi.org/10.5281/zenodo.12189914	FIGURE 4. Male appendages of Coenagrion spp. of the puella-group in the broad sense in posterior view: a–b—C. australocaspicum (b—with the appendages expanded); c–d—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); e-f—C. pulchellum; g–k—C. puella; l—C. persicum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c–d—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; e—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; f—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; g–h—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; i—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, a brook and pond, 25.06.2021; j—Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017. Scale bar 0.5 mm.	FIGURE 4. Male appendages of Coenagrion spp. of the puella-group in the broad sense in posterior view: a–b—C. australocaspicum (b—with the appendages expanded); c–d—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); e-f—C. pulchellum; g–k—C. puella; l—C. persicum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c–d—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; e—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; f—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; g–h—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; i—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, a brook and pond, 25.06.2021; j—Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017. Scale bar 0.5 mm.	2024-06-18	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.		Zenodo	biologists	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.			
DB0F87E19101F10DA7D3FBDE78C47FCF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12189916/files/figure.png	https://doi.org/10.5281/zenodo.12189916	FIGURE 5. Male pronotum of Coenagrion spp. of the puella-group in the broad sense in dorsal view: a—C. persicum; b–c—C. pulchellum; d–e—C. ponticum (males with a more (d) and less (e) expanded black abdominal maculation); f—C. australocaspicum; g–l—C. puella; a—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; b—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; c—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; d–e—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; f—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; g —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; h–i—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021; j—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; k—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; l—Russia, Karachay- Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019. Scale bar 0.5 mm.	FIGURE 5. Male pronotum of Coenagrion spp. of the puella-group in the broad sense in dorsal view: a—C. persicum; b–c—C. pulchellum; d–e—C. ponticum (males with a more (d) and less (e) expanded black abdominal maculation); f—C. australocaspicum; g–l—C. puella; a—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; b—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; c—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; d–e—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; f—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; g —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; h–i—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021; j—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; k—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; l—Russia, Karachay- Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019. Scale bar 0.5 mm.	2024-06-18	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.		Zenodo	biologists	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.			
DB0F87E19101F10DA7D3FBDE78C47FCF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12189924/files/figure.png	https://doi.org/10.5281/zenodo.12189924	FIGURE 7. Female pronotum of Coenagrion spp. of the puella-group in the broad sense, a–h—in dorsal, with respect to the body main axis (above), and anteriodorsal, perpendicular to the posterior lobe surface (below), views; i–j—in dorsal view only; a—C. australocaspicum; b–g—C. puella; h—C. ponticum; i—C. persicum; j—C. pulchellum; a—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; b—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; c—Sergokala District, 3 km SW of Myurego village, the Inchkheozen’ River upper valley, 19.06.2021; d–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; h—Russia, Chelyabinsk Province, Nagaybakskiy District, small oxbow lakes at the Gumbeyka River left bank at Ostrolenskiy Settlement SE margin, 6.07.2021; i—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; j—Iran, Markazi Ostan, Shazand Shakhrestan, the Garehchay River at Gavigodar village, 34.1078-1111° N 49.3627-3657° E, 1803–1805 m a.s.l., 19.05.2017; k—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.	FIGURE 7. Female pronotum of Coenagrion spp. of the puella-group in the broad sense, a–h—in dorsal, with respect to the body main axis (above), and anteriodorsal, perpendicular to the posterior lobe surface (below), views; i–j—in dorsal view only; a—C. australocaspicum; b–g—C. puella; h—C. ponticum; i—C. persicum; j—C. pulchellum; a—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; b—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; c—Sergokala District, 3 km SW of Myurego village, the Inchkheozen’ River upper valley, 19.06.2021; d–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; h—Russia, Chelyabinsk Province, Nagaybakskiy District, small oxbow lakes at the Gumbeyka River left bank at Ostrolenskiy Settlement SE margin, 6.07.2021; i—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; j—Iran, Markazi Ostan, Shazand Shakhrestan, the Garehchay River at Gavigodar village, 34.1078-1111° N 49.3627-3657° E, 1803–1805 m a.s.l., 19.05.2017; k—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.	2024-06-18	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.		Zenodo	biologists	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.			
DB0F87E19101F10DA7D3FBDE78C47FCF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12189926/files/figure.png	https://doi.org/10.5281/zenodo.12189926	FIGURE 8. Coenagrion puella (a–d) and C. pulchellum (e–m) in different populations in Dagestan in nature; a, j—copulae; b–c—tandems; d, k–m—males; e–g—females; a—Buynaksk District, Verkhnee Kazanishche village environs, a brook on the Akpeozen’ River valley left slope, 29.05.2023 (178696752); b—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021 (196674839); c—Tabasaran District, the Rubas River valley 2.5 km NE of Sirtych village, Dzharag- Khyan Spring, Sirtych, 28.05.2023 (178695642); d —Gergebil’ District, at Gergebil’ Water Reservoir right bank, 24.05.2023 (167302379); e, h, j—Makhachkala N suburbs, pools at the Krivaya Balka River left bank in its lowermost reaches, 30.05.2023 (178699404, 178699407, 178699406, respectively); f–g, k–l—the same place, 23.05.2023 (166728450, 166734669, 166726653, 166733553, respectively); m—Shuraozen’, 30.05.2023 (181375745); a, c–l—photo by OEK, b, m—by VVO. Not to scale.	FIGURE 8. Coenagrion puella (a–d) and C. pulchellum (e–m) in different populations in Dagestan in nature; a, j—copulae; b–c—tandems; d, k–m—males; e–g—females; a—Buynaksk District, Verkhnee Kazanishche village environs, a brook on the Akpeozen’ River valley left slope, 29.05.2023 (178696752); b—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021 (196674839); c—Tabasaran District, the Rubas River valley 2.5 km NE of Sirtych village, Dzharag- Khyan Spring, Sirtych, 28.05.2023 (178695642); d —Gergebil’ District, at Gergebil’ Water Reservoir right bank, 24.05.2023 (167302379); e, h, j—Makhachkala N suburbs, pools at the Krivaya Balka River left bank in its lowermost reaches, 30.05.2023 (178699404, 178699407, 178699406, respectively); f–g, k–l—the same place, 23.05.2023 (166728450, 166734669, 166726653, 166733553, respectively); m—Shuraozen’, 30.05.2023 (181375745); a, c–l—photo by OEK, b, m—by VVO. Not to scale.	2024-06-18	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.		Zenodo	biologists	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.			
DB0F87E19102F10AA7D3F8F47E277CEF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12189910/files/figure.png	https://doi.org/10.5281/zenodo.12189910	FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.	FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.	2024-06-18	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.		Zenodo	biologists	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.			
DB0F87E19102F10AA7D3F8F47E277CEF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12189912/files/figure.png	https://doi.org/10.5281/zenodo.12189912	FIGURE3. Maleappendages of Coenagrion spp.ofthe puella-groupin thebroadsense inlateral view:a–c—C.australocaspicum (b—with the appendages expanded); d–k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–q—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; k—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.	FIGURE3. Maleappendages of Coenagrion spp.ofthe puella-groupin thebroadsense inlateral view:a–c—C.australocaspicum (b—with the appendages expanded); d–k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–q—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; k—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.	2024-06-18	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.		Zenodo	biologists	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.			
DB0F87E19102F10AA7D3F8F47E277CEF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12189914/files/figure.png	https://doi.org/10.5281/zenodo.12189914	FIGURE 4. Male appendages of Coenagrion spp. of the puella-group in the broad sense in posterior view: a–b—C. australocaspicum (b—with the appendages expanded); c–d—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); e-f—C. pulchellum; g–k—C. puella; l—C. persicum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c–d—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; e—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; f—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; g–h—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; i—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, a brook and pond, 25.06.2021; j—Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017. Scale bar 0.5 mm.	FIGURE 4. Male appendages of Coenagrion spp. of the puella-group in the broad sense in posterior view: a–b—C. australocaspicum (b—with the appendages expanded); c–d—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); e-f—C. pulchellum; g–k—C. puella; l—C. persicum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c–d—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; e—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; f—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; g–h—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; i—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, a brook and pond, 25.06.2021; j—Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017. Scale bar 0.5 mm.	2024-06-18	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.		Zenodo	biologists	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.			
DB0F87E19102F10AA7D3F8F47E277CEF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12189916/files/figure.png	https://doi.org/10.5281/zenodo.12189916	FIGURE 5. Male pronotum of Coenagrion spp. of the puella-group in the broad sense in dorsal view: a—C. persicum; b–c—C. pulchellum; d–e—C. ponticum (males with a more (d) and less (e) expanded black abdominal maculation); f—C. australocaspicum; g–l—C. puella; a—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; b—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; c—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; d–e—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; f—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; g —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; h–i—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021; j—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; k—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; l—Russia, Karachay- Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019. Scale bar 0.5 mm.	FIGURE 5. Male pronotum of Coenagrion spp. of the puella-group in the broad sense in dorsal view: a—C. persicum; b–c—C. pulchellum; d–e—C. ponticum (males with a more (d) and less (e) expanded black abdominal maculation); f—C. australocaspicum; g–l—C. puella; a—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; b—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; c—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; d–e—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; f—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; g —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; h–i—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021; j—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; k—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; l—Russia, Karachay- Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019. Scale bar 0.5 mm.	2024-06-18	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.		Zenodo	biologists	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.			
DB0F87E19102F10AA7D3F8F47E277CEF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12189924/files/figure.png	https://doi.org/10.5281/zenodo.12189924	FIGURE 7. Female pronotum of Coenagrion spp. of the puella-group in the broad sense, a–h—in dorsal, with respect to the body main axis (above), and anteriodorsal, perpendicular to the posterior lobe surface (below), views; i–j—in dorsal view only; a—C. australocaspicum; b–g—C. puella; h—C. ponticum; i—C. persicum; j—C. pulchellum; a—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; b—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; c—Sergokala District, 3 km SW of Myurego village, the Inchkheozen’ River upper valley, 19.06.2021; d–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; h—Russia, Chelyabinsk Province, Nagaybakskiy District, small oxbow lakes at the Gumbeyka River left bank at Ostrolenskiy Settlement SE margin, 6.07.2021; i—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; j—Iran, Markazi Ostan, Shazand Shakhrestan, the Garehchay River at Gavigodar village, 34.1078-1111° N 49.3627-3657° E, 1803–1805 m a.s.l., 19.05.2017; k—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.	FIGURE 7. Female pronotum of Coenagrion spp. of the puella-group in the broad sense, a–h—in dorsal, with respect to the body main axis (above), and anteriodorsal, perpendicular to the posterior lobe surface (below), views; i–j—in dorsal view only; a—C. australocaspicum; b–g—C. puella; h—C. ponticum; i—C. persicum; j—C. pulchellum; a—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; b—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; c—Sergokala District, 3 km SW of Myurego village, the Inchkheozen’ River upper valley, 19.06.2021; d–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; h—Russia, Chelyabinsk Province, Nagaybakskiy District, small oxbow lakes at the Gumbeyka River left bank at Ostrolenskiy Settlement SE margin, 6.07.2021; i—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; j—Iran, Markazi Ostan, Shazand Shakhrestan, the Garehchay River at Gavigodar village, 34.1078-1111° N 49.3627-3657° E, 1803–1805 m a.s.l., 19.05.2017; k—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.	2024-06-18	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.		Zenodo	biologists	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.			
DB0F87E19102F10AA7D3F8F47E277CEF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12189926/files/figure.png	https://doi.org/10.5281/zenodo.12189926	FIGURE 8. Coenagrion puella (a–d) and C. pulchellum (e–m) in different populations in Dagestan in nature; a, j—copulae; b–c—tandems; d, k–m—males; e–g—females; a—Buynaksk District, Verkhnee Kazanishche village environs, a brook on the Akpeozen’ River valley left slope, 29.05.2023 (178696752); b—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021 (196674839); c—Tabasaran District, the Rubas River valley 2.5 km NE of Sirtych village, Dzharag- Khyan Spring, Sirtych, 28.05.2023 (178695642); d —Gergebil’ District, at Gergebil’ Water Reservoir right bank, 24.05.2023 (167302379); e, h, j—Makhachkala N suburbs, pools at the Krivaya Balka River left bank in its lowermost reaches, 30.05.2023 (178699404, 178699407, 178699406, respectively); f–g, k–l—the same place, 23.05.2023 (166728450, 166734669, 166726653, 166733553, respectively); m—Shuraozen’, 30.05.2023 (181375745); a, c–l—photo by OEK, b, m—by VVO. Not to scale.	FIGURE 8. Coenagrion puella (a–d) and C. pulchellum (e–m) in different populations in Dagestan in nature; a, j—copulae; b–c—tandems; d, k–m—males; e–g—females; a—Buynaksk District, Verkhnee Kazanishche village environs, a brook on the Akpeozen’ River valley left slope, 29.05.2023 (178696752); b—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021 (196674839); c—Tabasaran District, the Rubas River valley 2.5 km NE of Sirtych village, Dzharag- Khyan Spring, Sirtych, 28.05.2023 (178695642); d —Gergebil’ District, at Gergebil’ Water Reservoir right bank, 24.05.2023 (167302379); e, h, j—Makhachkala N suburbs, pools at the Krivaya Balka River left bank in its lowermost reaches, 30.05.2023 (178699404, 178699407, 178699406, respectively); f–g, k–l—the same place, 23.05.2023 (166728450, 166734669, 166726653, 166733553, respectively); m—Shuraozen’, 30.05.2023 (181375745); a, c–l—photo by OEK, b, m—by VVO. Not to scale.	2024-06-18	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.		Zenodo	biologists	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.			
DB0F87E19105F108A7D3FBDE78C67F5B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12189910/files/figure.png	https://doi.org/10.5281/zenodo.12189910	FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.	FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.	2024-06-18	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.		Zenodo	biologists	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.			
DB0F87E19105F108A7D3FBDE78C67F5B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12189912/files/figure.png	https://doi.org/10.5281/zenodo.12189912	FIGURE3. Maleappendages of Coenagrion spp.ofthe puella-groupin thebroadsense inlateral view:a–c—C.australocaspicum (b—with the appendages expanded); d–k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–q—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; k—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.	FIGURE3. Maleappendages of Coenagrion spp.ofthe puella-groupin thebroadsense inlateral view:a–c—C.australocaspicum (b—with the appendages expanded); d–k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–q—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; k—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.	2024-06-18	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.		Zenodo	biologists	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.			
DB0F87E19105F108A7D3FBDE78C67F5B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12189914/files/figure.png	https://doi.org/10.5281/zenodo.12189914	FIGURE 4. Male appendages of Coenagrion spp. of the puella-group in the broad sense in posterior view: a–b—C. australocaspicum (b—with the appendages expanded); c–d—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); e-f—C. pulchellum; g–k—C. puella; l—C. persicum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c–d—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; e—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; f—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; g–h—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; i—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, a brook and pond, 25.06.2021; j—Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017. Scale bar 0.5 mm.	FIGURE 4. Male appendages of Coenagrion spp. of the puella-group in the broad sense in posterior view: a–b—C. australocaspicum (b—with the appendages expanded); c–d—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); e-f—C. pulchellum; g–k—C. puella; l—C. persicum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c–d—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; e—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; f—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; g–h—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; i—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, a brook and pond, 25.06.2021; j—Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017. Scale bar 0.5 mm.	2024-06-18	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.		Zenodo	biologists	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.			
DB0F87E19105F108A7D3FBDE78C67F5B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12189916/files/figure.png	https://doi.org/10.5281/zenodo.12189916	FIGURE 5. Male pronotum of Coenagrion spp. of the puella-group in the broad sense in dorsal view: a—C. persicum; b–c—C. pulchellum; d–e—C. ponticum (males with a more (d) and less (e) expanded black abdominal maculation); f—C. australocaspicum; g–l—C. puella; a—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; b—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; c—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; d–e—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; f—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; g —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; h–i—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021; j—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; k—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; l—Russia, Karachay- Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019. Scale bar 0.5 mm.	FIGURE 5. Male pronotum of Coenagrion spp. of the puella-group in the broad sense in dorsal view: a—C. persicum; b–c—C. pulchellum; d–e—C. ponticum (males with a more (d) and less (e) expanded black abdominal maculation); f—C. australocaspicum; g–l—C. puella; a—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; b—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; c—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; d–e—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; f—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; g —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; h–i—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021; j—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; k—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; l—Russia, Karachay- Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019. Scale bar 0.5 mm.	2024-06-18	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.		Zenodo	biologists	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.			
DB0F87E19105F108A7D3FBDE78C67F5B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12189924/files/figure.png	https://doi.org/10.5281/zenodo.12189924	FIGURE 7. Female pronotum of Coenagrion spp. of the puella-group in the broad sense, a–h—in dorsal, with respect to the body main axis (above), and anteriodorsal, perpendicular to the posterior lobe surface (below), views; i–j—in dorsal view only; a—C. australocaspicum; b–g—C. puella; h—C. ponticum; i—C. persicum; j—C. pulchellum; a—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; b—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; c—Sergokala District, 3 km SW of Myurego village, the Inchkheozen’ River upper valley, 19.06.2021; d–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; h—Russia, Chelyabinsk Province, Nagaybakskiy District, small oxbow lakes at the Gumbeyka River left bank at Ostrolenskiy Settlement SE margin, 6.07.2021; i—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; j—Iran, Markazi Ostan, Shazand Shakhrestan, the Garehchay River at Gavigodar village, 34.1078-1111° N 49.3627-3657° E, 1803–1805 m a.s.l., 19.05.2017; k—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.	FIGURE 7. Female pronotum of Coenagrion spp. of the puella-group in the broad sense, a–h—in dorsal, with respect to the body main axis (above), and anteriodorsal, perpendicular to the posterior lobe surface (below), views; i–j—in dorsal view only; a—C. australocaspicum; b–g—C. puella; h—C. ponticum; i—C. persicum; j—C. pulchellum; a—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; b—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; c—Sergokala District, 3 km SW of Myurego village, the Inchkheozen’ River upper valley, 19.06.2021; d–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; h—Russia, Chelyabinsk Province, Nagaybakskiy District, small oxbow lakes at the Gumbeyka River left bank at Ostrolenskiy Settlement SE margin, 6.07.2021; i—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; j—Iran, Markazi Ostan, Shazand Shakhrestan, the Garehchay River at Gavigodar village, 34.1078-1111° N 49.3627-3657° E, 1803–1805 m a.s.l., 19.05.2017; k—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.	2024-06-18	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.		Zenodo	biologists	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.			
DB0F87E19105F108A7D3FBDE78C67F5B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12189928/files/figure.png	https://doi.org/10.5281/zenodo.12189928	FIGURE 9. Males (a–b) and female (c) of Coenagrion ponticum in nature by a pond in the Fanagoriyskoe village NW environs, Goryachiy Klyuch Municipality, Krasnodarskiy Kray, Russia, 27.04.2019 (108924871, 37680876, 37681071, respectively); a—photo by OEK, b, c—by VVO. Not to scale.	FIGURE 9. Males (a–b) and female (c) of Coenagrion ponticum in nature by a pond in the Fanagoriyskoe village NW environs, Goryachiy Klyuch Municipality, Krasnodarskiy Kray, Russia, 27.04.2019 (108924871, 37680876, 37681071, respectively); a—photo by OEK, b, c—by VVO. Not to scale.	2024-06-18	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.		Zenodo	biologists	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.			
DB0F87E19107F108A7D3F84A7F557999.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12189930/files/figure.png	https://doi.org/10.5281/zenodo.12189930	FIGURE 10. Male appendages of Coenagrion ornatum specimens from Dagestan in dorsal view: a–b—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021; c–d—Tabasaran District, the Rubas River valley 2.5 km NE of Sirtych village, a swampy Dzharag-Khyan Spring, 28.05.2023. Scale bar 0.5 mm.	FIGURE 10. Male appendages of Coenagrion ornatum specimens from Dagestan in dorsal view: a–b—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021; c–d—Tabasaran District, the Rubas River valley 2.5 km NE of Sirtych village, a swampy Dzharag-Khyan Spring, 28.05.2023. Scale bar 0.5 mm.	2024-06-18	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.		Zenodo	biologists	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.			
DB0F87E19107F108A7D3F84A7F557999.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12189932/files/figure.png	https://doi.org/10.5281/zenodo.12189932	FIGURE 11. Coenagrion ornatum (a–d) and C. scitulum (e–k) in different populations in the Russian part of the Caucasus in nature; a–c, e–f—males; d—copula; g–j—females; k—tandems; a—Buynaksk District, Verkhnee Kazanishche village environs, a brook on the Akpeozen’ River valley left slope, 29.05.2023 (180159191); b—Tabasaran District, the Rubas River valley 2.5 km NE of Sirtych village, Dzharag-Khyan Spring, Sirtych, 28.05.2023 (178695635); c—Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021 (178693662); d—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021 (98025103); e, j—Krasnodarskiy Kray, Anapa Municipality, E suburbs of Supsekh settlement, Lake Zmeinoe env., 24.06.2019 (108802758, 108804470, respectively); f—Karachay-Cherkes Rpeublic, Malokarachaevsk District, Uchkeken village E environs, 9.07.2019 (109974039, Fig. 11f); g, k—Dagestan, Makhachkala N suburbs, pools at the Krivaya Balka River left bank in its lowermost reaches, 30.05.2023 (176987626, 182772933, respectively); h—Derbent District, 3.5 km NE of Belidzhi village, thickets at a swamp, 25.05.2023 (186590343); i—Dagestan, Buynaksk District, Nizhnee Kazanishche village environs, a water reservoir, 29.05.2023 (176987387); a, g–i, k—photo by VVO, b–f, j—by OEK. Not to scale.	FIGURE 11. Coenagrion ornatum (a–d) and C. scitulum (e–k) in different populations in the Russian part of the Caucasus in nature; a–c, e–f—males; d—copula; g–j—females; k—tandems; a—Buynaksk District, Verkhnee Kazanishche village environs, a brook on the Akpeozen’ River valley left slope, 29.05.2023 (180159191); b—Tabasaran District, the Rubas River valley 2.5 km NE of Sirtych village, Dzharag-Khyan Spring, Sirtych, 28.05.2023 (178695635); c—Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021 (178693662); d—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021 (98025103); e, j—Krasnodarskiy Kray, Anapa Municipality, E suburbs of Supsekh settlement, Lake Zmeinoe env., 24.06.2019 (108802758, 108804470, respectively); f—Karachay-Cherkes Rpeublic, Malokarachaevsk District, Uchkeken village E environs, 9.07.2019 (109974039, Fig. 11f); g, k—Dagestan, Makhachkala N suburbs, pools at the Krivaya Balka River left bank in its lowermost reaches, 30.05.2023 (176987626, 182772933, respectively); h—Derbent District, 3.5 km NE of Belidzhi village, thickets at a swamp, 25.05.2023 (186590343); i—Dagestan, Buynaksk District, Nizhnee Kazanishche village environs, a water reservoir, 29.05.2023 (176987387); a, g–i, k—photo by VVO, b–f, j—by OEK. Not to scale.	2024-06-18	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.		Zenodo	biologists	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.			
DB0F87E19106F109A7D3F9B97A857FB6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12189920/files/figure.png	https://doi.org/10.5281/zenodo.12189920	FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale.	FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale.	2024-06-18	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.		Zenodo	biologists	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.			
DB0F87E19106F109A7D3FB1978B779B7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12189932/files/figure.png	https://doi.org/10.5281/zenodo.12189932	FIGURE 11. Coenagrion ornatum (a–d) and C. scitulum (e–k) in different populations in the Russian part of the Caucasus in nature; a–c, e–f—males; d—copula; g–j—females; k—tandems; a—Buynaksk District, Verkhnee Kazanishche village environs, a brook on the Akpeozen’ River valley left slope, 29.05.2023 (180159191); b—Tabasaran District, the Rubas River valley 2.5 km NE of Sirtych village, Dzharag-Khyan Spring, Sirtych, 28.05.2023 (178695635); c—Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021 (178693662); d—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021 (98025103); e, j—Krasnodarskiy Kray, Anapa Municipality, E suburbs of Supsekh settlement, Lake Zmeinoe env., 24.06.2019 (108802758, 108804470, respectively); f—Karachay-Cherkes Rpeublic, Malokarachaevsk District, Uchkeken village E environs, 9.07.2019 (109974039, Fig. 11f); g, k—Dagestan, Makhachkala N suburbs, pools at the Krivaya Balka River left bank in its lowermost reaches, 30.05.2023 (176987626, 182772933, respectively); h—Derbent District, 3.5 km NE of Belidzhi village, thickets at a swamp, 25.05.2023 (186590343); i—Dagestan, Buynaksk District, Nizhnee Kazanishche village environs, a water reservoir, 29.05.2023 (176987387); a, g–i, k—photo by VVO, b–f, j—by OEK. Not to scale.	FIGURE 11. Coenagrion ornatum (a–d) and C. scitulum (e–k) in different populations in the Russian part of the Caucasus in nature; a–c, e–f—males; d—copula; g–j—females; k—tandems; a—Buynaksk District, Verkhnee Kazanishche village environs, a brook on the Akpeozen’ River valley left slope, 29.05.2023 (180159191); b—Tabasaran District, the Rubas River valley 2.5 km NE of Sirtych village, Dzharag-Khyan Spring, Sirtych, 28.05.2023 (178695635); c—Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021 (178693662); d—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021 (98025103); e, j—Krasnodarskiy Kray, Anapa Municipality, E suburbs of Supsekh settlement, Lake Zmeinoe env., 24.06.2019 (108802758, 108804470, respectively); f—Karachay-Cherkes Rpeublic, Malokarachaevsk District, Uchkeken village E environs, 9.07.2019 (109974039, Fig. 11f); g, k—Dagestan, Makhachkala N suburbs, pools at the Krivaya Balka River left bank in its lowermost reaches, 30.05.2023 (176987626, 182772933, respectively); h—Derbent District, 3.5 km NE of Belidzhi village, thickets at a swamp, 25.05.2023 (186590343); i—Dagestan, Buynaksk District, Nizhnee Kazanishche village environs, a water reservoir, 29.05.2023 (176987387); a, g–i, k—photo by VVO, b–f, j—by OEK. Not to scale.	2024-06-18	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.		Zenodo	biologists	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.			
DB0F87E1910FF101A7D3FC71795D7E93.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12189930/files/figure.png	https://doi.org/10.5281/zenodo.12189930	FIGURE 10. Male appendages of Coenagrion ornatum specimens from Dagestan in dorsal view: a–b—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021; c–d—Tabasaran District, the Rubas River valley 2.5 km NE of Sirtych village, a swampy Dzharag-Khyan Spring, 28.05.2023. Scale bar 0.5 mm.	FIGURE 10. Male appendages of Coenagrion ornatum specimens from Dagestan in dorsal view: a–b—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021; c–d—Tabasaran District, the Rubas River valley 2.5 km NE of Sirtych village, a swampy Dzharag-Khyan Spring, 28.05.2023. Scale bar 0.5 mm.	2024-06-18	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.		Zenodo	biologists	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.			
DB0F87E1910EF101A7D3F80979D87817.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12189914/files/figure.png	https://doi.org/10.5281/zenodo.12189914	FIGURE 4. Male appendages of Coenagrion spp. of the puella-group in the broad sense in posterior view: a–b—C. australocaspicum (b—with the appendages expanded); c–d—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); e-f—C. pulchellum; g–k—C. puella; l—C. persicum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c–d—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; e—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; f—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; g–h—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; i—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, a brook and pond, 25.06.2021; j—Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017. Scale bar 0.5 mm.	FIGURE 4. Male appendages of Coenagrion spp. of the puella-group in the broad sense in posterior view: a–b—C. australocaspicum (b—with the appendages expanded); c–d—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); e-f—C. pulchellum; g–k—C. puella; l—C. persicum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c–d—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; e—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; f—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; g–h—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; i—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, a brook and pond, 25.06.2021; j—Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017. Scale bar 0.5 mm.	2024-06-18	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.		Zenodo	biologists	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.			
DB0F87E1910EF101A7D3F80979D87817.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12189942/files/figure.png	https://doi.org/10.5281/zenodo.12189942	FIGURE 16. Morphological details of male specimens of Coenagrion syriacum kept in the RMNH collection, courtesy by Charlotte Hartong: a–c—“Turkey, Adana / River Ceyhan, Ca. 20 km NW of / Osmaniye (37SBB42) River and / pools. / 11 May 2002. / Leg.: K. Goudsmits & V.J. Kalkman” (RMNH.INS.1468579); d–f—“Turkey, Adana / Aydinlar, 15 km W of Kadirli / (37SBB33) Well vegetated, slowly flowing / channel. / 10 May 2002. / Leg.: K. Goudsmits & V.J. Kalkman” (RMNH. INS.1468580); a, d—end of abbdomen in lateral view; b, e—ditto, dorsal view; a, d—ditto, caudal view.	FIGURE 16. Morphological details of male specimens of Coenagrion syriacum kept in the RMNH collection, courtesy by Charlotte Hartong: a–c—“Turkey, Adana / River Ceyhan, Ca. 20 km NW of / Osmaniye (37SBB42) River and / pools. / 11 May 2002. / Leg.: K. Goudsmits & V.J. Kalkman” (RMNH.INS.1468579); d–f—“Turkey, Adana / Aydinlar, 15 km W of Kadirli / (37SBB33) Well vegetated, slowly flowing / channel. / 10 May 2002. / Leg.: K. Goudsmits & V.J. Kalkman” (RMNH. INS.1468580); a, d—end of abbdomen in lateral view; b, e—ditto, dorsal view; a, d—ditto, caudal view.	2024-06-18	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.		Zenodo	biologists	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.			
DB0F87E1910EF101A7D3F80979D87817.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12189910/files/figure.png	https://doi.org/10.5281/zenodo.12189910	FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.	FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.	2024-06-18	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.		Zenodo	biologists	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.			
DB0F87E1910EF101A7D3F80979D87817.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12189912/files/figure.png	https://doi.org/10.5281/zenodo.12189912	FIGURE3. Maleappendages of Coenagrion spp.ofthe puella-groupin thebroadsense inlateral view:a–c—C.australocaspicum (b—with the appendages expanded); d–k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–q—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; k—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.	FIGURE3. Maleappendages of Coenagrion spp.ofthe puella-groupin thebroadsense inlateral view:a–c—C.australocaspicum (b—with the appendages expanded); d–k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–q—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; k—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.	2024-06-18	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.		Zenodo	biologists	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.			
DB0F87E1910EF101A7D3F80979D87817.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12189916/files/figure.png	https://doi.org/10.5281/zenodo.12189916	FIGURE 5. Male pronotum of Coenagrion spp. of the puella-group in the broad sense in dorsal view: a—C. persicum; b–c—C. pulchellum; d–e—C. ponticum (males with a more (d) and less (e) expanded black abdominal maculation); f—C. australocaspicum; g–l—C. puella; a—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; b—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; c—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; d–e—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; f—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; g —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; h–i—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021; j—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; k—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; l—Russia, Karachay- Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019. Scale bar 0.5 mm.	FIGURE 5. Male pronotum of Coenagrion spp. of the puella-group in the broad sense in dorsal view: a—C. persicum; b–c—C. pulchellum; d–e—C. ponticum (males with a more (d) and less (e) expanded black abdominal maculation); f—C. australocaspicum; g–l—C. puella; a—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; b—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; c—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; d–e—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; f—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; g —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; h–i—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021; j—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; k—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; l—Russia, Karachay- Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019. Scale bar 0.5 mm.	2024-06-18	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.		Zenodo	biologists	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.			
DB0F87E1910EF101A7D3F80979D87817.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12189920/files/figure.png	https://doi.org/10.5281/zenodo.12189920	FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale.	FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale.	2024-06-18	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.		Zenodo	biologists	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.			
DB0F87E1910EF101A7D3FF8D788579FF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12189910/files/figure.png	https://doi.org/10.5281/zenodo.12189910	FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.	FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.	2024-06-18	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.		Zenodo	biologists	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.			
DB0F87E1910EF101A7D3FF8D788579FF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12189912/files/figure.png	https://doi.org/10.5281/zenodo.12189912	FIGURE3. Maleappendages of Coenagrion spp.ofthe puella-groupin thebroadsense inlateral view:a–c—C.australocaspicum (b—with the appendages expanded); d–k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–q—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; k—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.	FIGURE3. Maleappendages of Coenagrion spp.ofthe puella-groupin thebroadsense inlateral view:a–c—C.australocaspicum (b—with the appendages expanded); d–k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–q—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; k—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.	2024-06-18	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.		Zenodo	biologists	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.			
DB0F87E1910EF101A7D3FF8D788579FF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12189924/files/figure.png	https://doi.org/10.5281/zenodo.12189924	FIGURE 7. Female pronotum of Coenagrion spp. of the puella-group in the broad sense, a–h—in dorsal, with respect to the body main axis (above), and anteriodorsal, perpendicular to the posterior lobe surface (below), views; i–j—in dorsal view only; a—C. australocaspicum; b–g—C. puella; h—C. ponticum; i—C. persicum; j—C. pulchellum; a—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; b—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; c—Sergokala District, 3 km SW of Myurego village, the Inchkheozen’ River upper valley, 19.06.2021; d–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; h—Russia, Chelyabinsk Province, Nagaybakskiy District, small oxbow lakes at the Gumbeyka River left bank at Ostrolenskiy Settlement SE margin, 6.07.2021; i—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; j—Iran, Markazi Ostan, Shazand Shakhrestan, the Garehchay River at Gavigodar village, 34.1078-1111° N 49.3627-3657° E, 1803–1805 m a.s.l., 19.05.2017; k—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.	FIGURE 7. Female pronotum of Coenagrion spp. of the puella-group in the broad sense, a–h—in dorsal, with respect to the body main axis (above), and anteriodorsal, perpendicular to the posterior lobe surface (below), views; i–j—in dorsal view only; a—C. australocaspicum; b–g—C. puella; h—C. ponticum; i—C. persicum; j—C. pulchellum; a—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; b—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; c—Sergokala District, 3 km SW of Myurego village, the Inchkheozen’ River upper valley, 19.06.2021; d–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; h—Russia, Chelyabinsk Province, Nagaybakskiy District, small oxbow lakes at the Gumbeyka River left bank at Ostrolenskiy Settlement SE margin, 6.07.2021; i—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; j—Iran, Markazi Ostan, Shazand Shakhrestan, the Garehchay River at Gavigodar village, 34.1078-1111° N 49.3627-3657° E, 1803–1805 m a.s.l., 19.05.2017; k—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.	2024-06-18	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.		Zenodo	biologists	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.			
DB0F87E19131F13DA7D3FC9079B07C7B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12189924/files/figure.png	https://doi.org/10.5281/zenodo.12189924	FIGURE 7. Female pronotum of Coenagrion spp. of the puella-group in the broad sense, a–h—in dorsal, with respect to the body main axis (above), and anteriodorsal, perpendicular to the posterior lobe surface (below), views; i–j—in dorsal view only; a—C. australocaspicum; b–g—C. puella; h—C. ponticum; i—C. persicum; j—C. pulchellum; a—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; b—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; c—Sergokala District, 3 km SW of Myurego village, the Inchkheozen’ River upper valley, 19.06.2021; d–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; h—Russia, Chelyabinsk Province, Nagaybakskiy District, small oxbow lakes at the Gumbeyka River left bank at Ostrolenskiy Settlement SE margin, 6.07.2021; i—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; j—Iran, Markazi Ostan, Shazand Shakhrestan, the Garehchay River at Gavigodar village, 34.1078-1111° N 49.3627-3657° E, 1803–1805 m a.s.l., 19.05.2017; k—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.	FIGURE 7. Female pronotum of Coenagrion spp. of the puella-group in the broad sense, a–h—in dorsal, with respect to the body main axis (above), and anteriodorsal, perpendicular to the posterior lobe surface (below), views; i–j—in dorsal view only; a—C. australocaspicum; b–g—C. puella; h—C. ponticum; i—C. persicum; j—C. pulchellum; a—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; b—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; c—Sergokala District, 3 km SW of Myurego village, the Inchkheozen’ River upper valley, 19.06.2021; d–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; h—Russia, Chelyabinsk Province, Nagaybakskiy District, small oxbow lakes at the Gumbeyka River left bank at Ostrolenskiy Settlement SE margin, 6.07.2021; i—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; j—Iran, Markazi Ostan, Shazand Shakhrestan, the Garehchay River at Gavigodar village, 34.1078-1111° N 49.3627-3657° E, 1803–1805 m a.s.l., 19.05.2017; k—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.	2024-06-18	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.		Zenodo	biologists	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.			
DB0F87E19131F13DA7D3FC9079B07C7B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12189910/files/figure.png	https://doi.org/10.5281/zenodo.12189910	FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.	FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.	2024-06-18	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.		Zenodo	biologists	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.			
DB0F87E19131F13DA7D3FC9079B07C7B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12189912/files/figure.png	https://doi.org/10.5281/zenodo.12189912	FIGURE3. Maleappendages of Coenagrion spp.ofthe puella-groupin thebroadsense inlateral view:a–c—C.australocaspicum (b—with the appendages expanded); d–k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–q—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; k—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.	FIGURE3. Maleappendages of Coenagrion spp.ofthe puella-groupin thebroadsense inlateral view:a–c—C.australocaspicum (b—with the appendages expanded); d–k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–q—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; k—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.	2024-06-18	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.		Zenodo	biologists	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.			
DB0F87E19131F13DA7D3FC9079B07C7B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12189914/files/figure.png	https://doi.org/10.5281/zenodo.12189914	FIGURE 4. Male appendages of Coenagrion spp. of the puella-group in the broad sense in posterior view: a–b—C. australocaspicum (b—with the appendages expanded); c–d—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); e-f—C. pulchellum; g–k—C. puella; l—C. persicum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c–d—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; e—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; f—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; g–h—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; i—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, a brook and pond, 25.06.2021; j—Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017. Scale bar 0.5 mm.	FIGURE 4. Male appendages of Coenagrion spp. of the puella-group in the broad sense in posterior view: a–b—C. australocaspicum (b—with the appendages expanded); c–d—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); e-f—C. pulchellum; g–k—C. puella; l—C. persicum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c–d—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; e—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; f—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; g–h—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; i—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, a brook and pond, 25.06.2021; j—Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017. Scale bar 0.5 mm.	2024-06-18	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.		Zenodo	biologists	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.			
DB0F87E19131F13DA7D3FC9079B07C7B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12189916/files/figure.png	https://doi.org/10.5281/zenodo.12189916	FIGURE 5. Male pronotum of Coenagrion spp. of the puella-group in the broad sense in dorsal view: a—C. persicum; b–c—C. pulchellum; d–e—C. ponticum (males with a more (d) and less (e) expanded black abdominal maculation); f—C. australocaspicum; g–l—C. puella; a—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; b—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; c—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; d–e—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; f—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; g —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; h–i—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021; j—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; k—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; l—Russia, Karachay- Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019. Scale bar 0.5 mm.	FIGURE 5. Male pronotum of Coenagrion spp. of the puella-group in the broad sense in dorsal view: a—C. persicum; b–c—C. pulchellum; d–e—C. ponticum (males with a more (d) and less (e) expanded black abdominal maculation); f—C. australocaspicum; g–l—C. puella; a—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; b—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; c—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; d–e—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; f—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; g —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; h–i—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021; j—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; k—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; l—Russia, Karachay- Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019. Scale bar 0.5 mm.	2024-06-18	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.		Zenodo	biologists	Kosterin, Oleg E.;Onishko, Vladimir V.;Ilyina, Elena V.;Chepurnov, Grigory Yu.;Blinov, Alexander G.			
