taxonID	type	description	language	source
616869CC0FEC5DDAB3D84C8E1BC517E3.taxon	description	Figs 10 A, 11, 37	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
616869CC0FEC5DDAB3D84C8E1BC517E3.taxon	distribution	Distribution. This species is known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
572693BB11185749B4C37128EC9939C6.taxon	description	Figs 10 B, 12, 37	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
572693BB11185749B4C37128EC9939C6.taxon	distribution	Distribution. This species is known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
C03226159ACA5868AB79A36EBACD07A2.taxon	description	Figs 10 C, 13, 14, 37	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
C03226159ACA5868AB79A36EBACD07A2.taxon	diagnosis	Diagnosis. Anauchen species with finely dimpled (pasty) shell surface sculpture (no spiral striae), aperture adnate to last whorl, five strong apertural barriers (parietal, upper palatal, lower palatal, basal, columellar).	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
C03226159ACA5868AB79A36EBACD07A2.taxon	description	Description. Shell conical, consisting of 5.5 – 6 thick-walled, regularly increasing, rounded, convex whorls separated by a moderately deep suture. All shells from the type material are somewhat weathered but colouration is dark reddish brown to pinkish, opaque. Protoconch weathered so the surface sculpture is not clear, not spirally striated, consisting of ~ 1.5 – 1.75 whorls, glossy and slightly darker than the rest of the shell. Teleoconch pasty, very finely radially striated, with no sign of spiral striation. Last whorl adnate to the penultimate and slightly descending near the aperture (~ 10 ° compared to the shell axis) making the aperture profile prosocline to the shell axis. Peristome continuous, leaning on the penultimate whorl at the parietal side in form of a thick and distinct callus. It is slightly paler in colour than the rest of the shell, expanded (particularly on the parietal side where it forms a distinct callus) but not reflected. Aperture equipped with five strong, high, relatively short, and thick barriers (parietal, upper palatal, lower palatal, basal, and columellar). Parietal lamella is the strongest in the aperture, blade-like, slightly oblique, its outer part curved towards the palatal side and only almost reaching the expanding peristome. Upper palatal and lower palatal plicae are similar, the lower one being stronger, more slender, higher, and positioned slightly deeper in the aperture. Basal plica small and very short, knob-like. Columellar lamella thick and horizontal, situated in the middle of the columellar wall. None of these barriers reach the expanding peristome. Surface of all apertural barriers is smooth. Sinulus wide and parabolic (due to the position of angular lamella and upper palatal plica). Umbilicus open and deep but moderately wide, measuring ~ 1 / 6 of the shell width. Only the penultimate whorl is visible through the umbilicus. A very short and shallow groove is present on the last ~ ¼ of the body whorl, running from the peristome towards the umbilicus.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
C03226159ACA5868AB79A36EBACD07A2.taxon	description	Measurements (in mm, n = 5). SH = 4.83 – 5.39; SW = 3.54 – 4.04; AH = 2.15 – 2.24; 2.04 – 2.23.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
C03226159ACA5868AB79A36EBACD07A2.taxon	etymology	Etymology. The specific epithet refers to the thick (Lat. crassus) shell and apertural barriers of this species.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
C03226159ACA5868AB79A36EBACD07A2.taxon	distribution	Distribution. Only known from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
DFE7170FC1AC5C339646558EB32FA710.taxon	description	Figs 10 D, 15, 37	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
DFE7170FC1AC5C339646558EB32FA710.taxon	distribution	Distribution. This species is known only from the imprecise type locality (Shan-Siam boundary).	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
B2B8499F32AE582CAB4D56CB0486E448.taxon	description	Figs 10 E, 16, 17, 37	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
B2B8499F32AE582CAB4D56CB0486E448.taxon	diagnosis	Diagnosis. Anauchen with very slightly keeled last whorl (below the centre of the periphery) which is also detached from the penultimate. Teleoconch finely dimpled, pasty sculptured, with no spiral striae. Five or six apertural barriers present (parietal, infraparietal, upper palatal, lower palatal, basal and columellar), all strongly spiniferous.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
B2B8499F32AE582CAB4D56CB0486E448.taxon	description	Description. Shell conical, brown, consisting of 4.5 – 5 convex whorls separated by a deep suture. Protoconch similar in colour as the rest of the shell, finely pitted, showing no spiralling pattern or spiral striae and consisting of 1.25 – 1.5 whorls. Teleoconch surface sculpture fine, pasty, without spiral striae and with several irregularly spaced whitish radial streaks which are most prominent on the last whorl. Last whorl convex but weakly keeled below the centre of the periphery, slightly detached from the penultimate near the aperture and very slightly ascending (below 5 ° compared to the shell axis). Peristome dirty white, expanded but not reflected. Aperture rounded, equally as wide as high. It is equipped with five or six simple apertural barriers (parietal, infraparietal, upper palatal, lower palatal, basal and columellar). Parietal lamella high and moderately strong, not reaching the expanding peristome. Upper and lower palatal plicae equally strong but the lower palatal is usually slightly higher. Basal plica present as a very weak, dot-like swelling situated halfway between the lower palatal plica and columellar lamella. The latter is sometimes almost as wide as long, provoking a rounded shape. The columellar wall of the aperture is weakly swollen around the columellar lamella. Infraparietal lamella present as a small swelling or completely absent. None of the apertural barriers are reaching the peristome (although the parietal lamella is the closest) and all are roughly spiniferous as well as the apertural surface. Sinulus wide and not strongly isolated from the rest of the aperture due to the relatively weak upper palatal plica. Umbilicus very narrow, measuring ~ 1 / 10 of the shell width. The umbilical groove is present running alongside it. This groove is the broadest near the peristome and regularly tapering towards the umbilicus, ending as a narrow canal at the last ~ ¼ of the body whorl.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
B2B8499F32AE582CAB4D56CB0486E448.taxon	description	Measurements (in mm, n = 5). SW = 2.10 – 2.58; SH = 2.39 – 2.82; AH = 1.10 – 1.22; AW = 1.09 – 1.28.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
B2B8499F32AE582CAB4D56CB0486E448.taxon	etymology	Etymology. The Latin word evanidus means vanishing, which refers to the quarrying of the type locality of this species.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
B2B8499F32AE582CAB4D56CB0486E448.taxon	distribution	Distribution. This species is known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
29EF10771C8F5F4A8D1DC9331002CA1F.taxon	description	Figs 10 F, 18, 37	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
29EF10771C8F5F4A8D1DC9331002CA1F.taxon	diagnosis	Diagnosis. Shell conical-ovoid, teleoconch radially and not spirally striated. Last whorl not detached from the penultimate and slightly descending. Aperture large, with seven weak barriers sitting on the peristome edge. Umbilicus narrow.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
29EF10771C8F5F4A8D1DC9331002CA1F.taxon	description	Description. Shell brown, conical-ovoid with deep suture and bulging whorls. Whorls 3.25, rounded, regularly increasing. Protoconch consists of slightly> 1 whorl, finely pitted, its terminal part with ~ 12 equidistant spiral striae. Teleoconch with inconspicuous, irregular growth lines and finely dimpled (pasty), spiral striation absent. Last whorl rounded, adnate to the penultimate and slightly descending near the aperture (~ 15 ° compared to the shell axis), aperture slightly prosocline to shell axis in lateral view. Aperture conspicuously large to the shell size, parietal part adnate onto penultimate whorl forming a weak callus. Peristome slightly expanded but not reflected. Altogether seven apertural barriers could be found, all being blunt, knob-like, situated on peristome edge, their homologies with traditionally recognised barriers somewhat questionable. Parietal lamella is the largest, but still weak. Suprapalatal and upper palatal plicae situated close to each other, after some distance the next plica is the lower palatal. Lower palatal, basal, subcolumellar, and columellar situated in approximately equal distance from one another, gap between columellar and parietal is approximately as large as between parietal and the suprapalatal. There is a very slight thickening next to lower palatal plica, although it is questionable whether it would develop into a barrier. Surfaces of all apertural barriers are very finely granulated. Sinulus wide but low, not strongly isolated from the rest of the aperture. Umbilicus narrow, elongate, only shows body whorl.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
29EF10771C8F5F4A8D1DC9331002CA1F.taxon	description	Measurements (in mm, n = 1). SH = 1.92; SW = 1.71, AH = 0.96, AW = 0.87.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
29EF10771C8F5F4A8D1DC9331002CA1F.taxon	etymology	Etymology. The specific epithet refers to the relatively large aperture when compared to the shell size.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
29EF10771C8F5F4A8D1DC9331002CA1F.taxon	distribution	Distribution. This species is known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
4805BBACC57E500E8508E7C56A4494C2.taxon	description	Figs 10 G, 19, 37	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
4805BBACC57E500E8508E7C56A4494C2.taxon	distribution	Distribution. This species is known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
E7E18EFE1D36515C9F434D1716E4365F.taxon	description	Figs 10 H, 20, 37	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
E7E18EFE1D36515C9F434D1716E4365F.taxon	distribution	Distribution. This subspecies is known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
EEEA001CBB8D5A5B8BEDCFADE9E22A40.taxon	description	Figs 10 I, 21, 37	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
EEEA001CBB8D5A5B8BEDCFADE9E22A40.taxon	distribution	Distribution. This subspecies is known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
2921D2B84C545541B00129D2DF03FB65.taxon	description	Figs 10 J, 22, 23, 37	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
2921D2B84C545541B00129D2DF03FB65.taxon	diagnosis	Diagnosis. Anauchen species with conical shell, last whorl keeled at its base and slightly ascending near the aperture. Spiral striation absent. Aperture armed with four strong barriers, umbilicus narrow.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
2921D2B84C545541B00129D2DF03FB65.taxon	description	Description. Shell conical, consisting of 5.5 regularly increasing, convex whorls separated by a deep suture. Colour light brown. There is not a clear boundary between the protoconch and the teleoconch due to the similar surface sculpture. Teleoconch pasty, very finely but irregularly radially striated, with no signs of spiral striation. Last whorl adnate to the penultimate and its last quarter slightly ascending near the aperture (~ 10 ° compared to the shell axis), making the aperture profile opisthocline to the shell axis. Last whorl with a blunt keel at its base, making the shell strongly triangular. Peristome continuous and thick, not very strongly expanded on the parietal side and not fully leaning on the penultimate whorl. It is slightly lighter in colour than the rest of the shell, not reflected. Sinulus rounded and well separated from the rest of the aperture. Aperture equipped with four strong but short barriers (parietal, upper palatal, lower palatal and columellar). Parietal lamella is the strongest in the aperture, blade-like. Upper palatal plica ~ 1 / 2 as strong as the parietal lamella and very slightly leaning towards the lower palatal plica. The latter is strong (only slightly weaker than the parietal) and moderately long. There is no basal plica. Columellar lamella thick and relatively low, situated in the middle of the columellar wall. All apertural barriers almost reach the expanding peristome. Surface of all apertural barriers is smooth. Umbilicus rounded, open but narrow, measuring ~ 1 / 11 of the shell width.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
2921D2B84C545541B00129D2DF03FB65.taxon	description	Measurements (in mm, n = 1). SH = 2.48; SW = 2.31; AH = 1.12; AW = 1.04.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
2921D2B84C545541B00129D2DF03FB65.taxon	etymology	Etymology. This species is named after and dedicated to Mór Jókai (1825 – 1904), Hungarian novelist, dramatist, to commemorate the bicentenary of his birth. Jókai was interested in malacology, he had a large collection of (mostly marine) shells, and even published a satirical novel entitled “ Novel of snails ” (A csigák regénye).	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
2921D2B84C545541B00129D2DF03FB65.taxon	distribution	Distribution. Only known from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
54F9A976131F5612ADA27504B83BAF42.taxon	description	Figs 10 K, 24, 37	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
54F9A976131F5612ADA27504B83BAF42.taxon	distribution	Distribution. This species is known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
AE544CE2E66A57289A1D68A7773350CB.taxon	distribution	Distribution. This species has a wide distribution across Vietnam and China. Its northernmost known localities are situated in northern parts of Guangxi province (China) while the southernmost are from Thanh Hoa province (Vietnam).	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
29907147B2E2503C87BEAA83B7395662.taxon	description	Figs 10 M, 26, 27, 37	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
29907147B2E2503C87BEAA83B7395662.taxon	diagnosis	Diagnosis. Anauchen species with conical-ovoid, brown shell, pasty-like surface, last whorl adnate to the penultimate, five apertural barriers (parietal (angulo-parietal), upper palatal, lower palatal, weak basal, and columellar) and a narrow umbilicus.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
29907147B2E2503C87BEAA83B7395662.taxon	description	Description. Shell conical-ovoid, brown, with 4.5 – 5 whorls separated by a deep suture. All whorls convex, bulging, rounded. Protoconch consists of ~ 1.5 whorls, finely pitted and with moderate spiral striae, the same colour as the rest of the shell. Teleoconch fine, pasty, with inconspicuous, irregular growth radial lines which are rarely more pronounced and in form of whitish streaks, most dense right behind the peristome. Spiral striation absent. Last whorl adnate to the penultimate and slightly descending (~ 25 ° compared to the shell axis) making the aperture profile prosocline to the shell axis. Peristome continuous, in form of a short and not much expanded callus leaning on the penultimate whorl at the parietal side. It is white, expanded but not reflected. Aperture equipped with five barriers (parietal (angulo-parietal), upper palatal, lower palatal, basal and columellar). Parietal (angulo-parietal) lamella almost fully reaching the expanded peristome edge. It is the largest in the aperture, sometimes single and sometimes with an additional smaller part, which is probably homologous with the angular lamella in other genera, pointed towards the palatal side. Upper palatal plica is weaker, lower, and less slender than the lower palatal. Lower palatal plica almost as strong as the angulo-parietal lamella. Basal plica in the form of inconspicuous knob, like a very slight swelling in the basal-columellar transition area. Columellar lamella almost reaching the expanded peristome, highest in its middle part and sloping towards the peristome edge where it becomes quite low. It is leaned towards the lower palatal plica. Surface of all apertural barriers is very finely granulated. Sinulus rounded, parabolic and well separated from the rest of the aperture. Umbilicus narrow, ~ 1 / 8 of the shell width, previous whorls not clearly visible through it. Umbilical groove absent.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
29907147B2E2503C87BEAA83B7395662.taxon	description	Measurements (in mm, n = 4). SH = 2.71 – 3; SW = 1.95 – 2.26; AH = 1.1 – 1.27; AW = 1.06 – 1.23.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
29907147B2E2503C87BEAA83B7395662.taxon	etymology	Etymology. This species is named for its globose shell.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
29907147B2E2503C87BEAA83B7395662.taxon	distribution	Distribution. This species is known only from Doi Tung mountain.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
6B2E5B70A63C58D9A55D6C11D0619540.taxon	description	Fig. 37	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
6B2E5B70A63C58D9A55D6C11D0619540.taxon	distribution	Distribution. This species is known only from Yunnan Province, China.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
67988AA56FEB55D088F1104C261F45B0.taxon	description	Figs 10 N, 28, 29, 37	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
67988AA56FEB55D088F1104C261F45B0.taxon	diagnosis	Diagnosis. Anauchen species with distinctly keeled last whorl (slightly below the centre of the periphery) which is also detached near the aperture and descending. Teleoconch spirally striated. Only four weak apertural barriers (parietal, upper palatal, lower palatal and columellar).	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
67988AA56FEB55D088F1104C261F45B0.taxon	description	Description. Shell conical or very slightly conical-ovoid, brown, consisting of 4.5 – 5 whorls separated by a deep suture. Protoconch not much lighter than the rest of the shell, roughly pitted and terminally spirally striated, consisting of 1.25 whorls. Initial teleoconch whorls and the penultimate whorl slightly keeled. Fine spiral striation visible on the teleoconch as well, sometimes stronger near the protoconch (on initial teleoconch whorls), sometimes stronger on the last whorl. There are several irregularly spaced whitish radial streaks crossing the spiral striae. Last whorl enlarged and distinctly keeled slightly below the centre of the periphery. The keel is equally strong along its length but is getting lost right behind the peristome. Last whorl slightly detached from the penultimate near the aperture and slightly descending (~ 15 ° compared to the shell axis), thus making the aperture profile prosocline to the shell axis. Peristome is dirty white, very thick, expanded but not reflected. Aperture rounded, almost as high as wide. There are only four relatively weak apertural barriers (parietal, upper palatal, lower palatal and columellar). Of them, parietal lamella is the strongest and highest and reaching the expanding peristome. Both palatal plicae (upper and lower) are of similar appearance (but lower is usually slightly stronger than the upper), both weak (low and short) and tubercle-like. They are positioned close to each other, probably closer than in any other congener. Columellar lamella is weak and dot-like, usually not present as a distinct barrier but rather as a columellar thickening which is gradually lowering towards the expanding peristome. Surface of all apertural barriers is very finely granulated. Sinulus wide and not strongly isolated from the rest of the aperture due to the relatively weak upper palatal plica. Umbilicus very narrow, elongated and measuring 1 / 8 – 1 / 9 of the shell width. There is a strong umbilical groove running along the umbilicus. This groove is very wide near the peristome and then regularly tapering towards the umbilicus, finally terminating at the ~ ½ of the last whorl.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
67988AA56FEB55D088F1104C261F45B0.taxon	description	Measurements (in mm, n = 5). SH = 2.64 – 2.95; SW = 2.63 – 2.82; AH = 1.23 – 1.38; AW = 1.27 – 1.37.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
67988AA56FEB55D088F1104C261F45B0.taxon	etymology	Etymology. This species looks like an Anauchen with rounded whorls painted in a Pablo Picasso style (i. e., resembling the art style known as Cubism). The specific epithet is to be used as a noun in apposition.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
67988AA56FEB55D088F1104C261F45B0.taxon	distribution	Distribution. This species is known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
5A58BAED0891523EB1CA23BA19BFFE3B.taxon	diagnosis	Diagnosis. This genus is characterised by the combination of characters: i) a single lamella which is formed by a fusion of angular and parietal lamellae (it can appear bifid in some species, see below); ii) last whorl adnate to, or only very slightly detached from the penultimate; iii) shell surface pasty (dimpled) or finely spirally striated (never rough, sandpaper-like surface); iv) surface of apertural barriers smooth.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
C85319C0F7BE53FBBF7E1987BC4A0D43.taxon	description	Figs 10 O, 30, 31, 37, 38	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
C85319C0F7BE53FBBF7E1987BC4A0D43.taxon	distribution	Distribution. This species has a wide distribution extending from the Hunan province, China (type locality of A. angulinus) south to Lang Son province, Vietnam.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
1C6E602308FA51A5A002C3908F2DACFF.taxon	description	Figs 10 P, 32, 33, 34, 37	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
1C6E602308FA51A5A002C3908F2DACFF.taxon	diagnosis	Diagnosis. Shell conical, consisting of 7.5 – 8 whorls. Teleoconch densely spirally striated. Last whorl adnate to the penultimate. Aperture equipped with four main barriers (parietal, upper palatal, lower palatal and columellar), supra-angular and additional two smaller barriers (basal and infraparietal).	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
1C6E602308FA51A5A002C3908F2DACFF.taxon	description	Description. Shell conical to weakly concave-conical, light brown-yellowish in colour, opaque. It is consisting of 7 – 8 regularly increasing whorls separated by a moderately deep suture. All whorls rounded, weakly convex. Protoconch lighter than the rest of the shell, slightly translucent, consisting of ~ 1.25 – 1.5 weakly spirally striated whorls. Teleoconch finely and densely spirally striated, especially on the last ~ 3 whorls. Spiral striae strong, irregularly spaced, curved,> 40 on the last whorl in a standard view. The space between two spiral striae ranges from the width of one to the width of two or even three spiral striae. These striae are occasionally crossed by a few radial whitish streaks, randomly positioned, but mostly present on the last and the penultimate whorl. Last whorl adnate to the penultimate, slightly ascending (~ 5 ° compared to the shell axis) and making the apertural profile slightly opisthocline to the shell axis. Peristome thick, the same colour as the rest of the shell, expanded and not reflected, very finely pitted. It is especially strongly expanded at the parietal side where it leans against the penultimate whorl and forms a thick callus. Aperture equipped with four main barriers (parietal, upper palatal, lower palatal, and columellar) and several smaller ones. Parietal lamella very strong and high, directed towards the lower palatal plica. Upper palatal plica very strong, developed equally as parietal lamella and slightly leaning towards the lower palatal plica. Parietal lamella and upper palatal plica are closely positioned, leaving a narrow spacing between them. Lower palatal plica is strong but weaker than the upper palatal, directed towards the embayment between columellar and parietal lamellae. Columellar lamella slightly oblique, directed towards the lower palatal plica and developed to the same extent. Between these main barriers, smaller number of minor ones are also observed. Supra-angular lamella present, small, and short, leaning towards the palatal wall. Interpalatal plica sometimes present, positioned rather obliquely. Subcolumellar lamella present as weak or absent. Basal plica present, short. A very slight thickening (swelling) like infraparietal lamella is present. This lamella is sometimes much stronger. Surface of all apertural barriers is finely granulated. Sinulus large, distinctly separated from the rest of the aperture due to the strong angulo-parietal lamella and upper palatal plica. Umbilicus narrow, measuring ~ 1 / 8 of the shell width and with almost perpendicular walls, not showing previous whorls.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
1C6E602308FA51A5A002C3908F2DACFF.taxon	description	Measurements (in mm, n = 5). SH = 4.05 – 4.7; SW = 2.72 – 3.43; AH = 1.54 – 1.88; AW = 0.7 – 0.87.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
1C6E602308FA51A5A002C3908F2DACFF.taxon	etymology	Etymology. This species is named after the tower-shaped (turreted) shell.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
1C6E602308FA51A5A002C3908F2DACFF.taxon	distribution	Distribution. This species is known from two localities in Thanh Hoa Province, Vietnam.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
60D512D3B2A758A6B1314737B13FC2FA.taxon	description	Figs 10 Q, 35, 36, 37	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
60D512D3B2A758A6B1314737B13FC2FA.taxon	diagnosis	Diagnosis. This species is most similar to A. huaykhakang and A. banmiensis but they are both spirally striated. Anauchen angthongensis is also spirally striated, has it last whorl regularly rounded and has a narrower umbilicus.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
60D512D3B2A758A6B1314737B13FC2FA.taxon	distribution	Distribution. This species is known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
4A0A2A9498A556FD8FEB6C1E31E2371A.taxon	description	Figs 39 A, 40, 41, 100	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
4A0A2A9498A556FD8FEB6C1E31E2371A.taxon	diagnosis	Diagnosis. Bensonella species with a triangular-conical shell and strongly radially ribbed last whorl, especially near the aperture. Palatal plicae very strong. Umbilicus very narrow and elongated, dot-like.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
4A0A2A9498A556FD8FEB6C1E31E2371A.taxon	description	Description. Shell triangular, conical-ovoid, brown, consisting of 4.5 – 5 convex, rounded whorls separated by a moderately deep suture. Protoconch spirally striated (~ 12 – 15 spiral striae). The protoconch is initially light in colour and then regularly darkening towards the teleoconch. Teleoconch surface very finely spirally striated, striae much weaker than on the protoconch. On the penultimate whorl, relatively coarse, weak, radial growth lines are visible. They become more densely arranged and more prominent on the last whorl and especially near the aperture where they are rib-like (important for species identification!). They are crossed by very delicate and widely spaced spiral striae. Last whorl rounded and adnate to the penultimate. It is slightly ascending near the aperture (~ 20 ° compared to the shell axis), making the aperture profile weakly opisthocline. Peristome lighter than the rest of the shell, not much expanded and not reflected. There is a slight thickening behind the peristome in form of a cervical crest. Aperture equipped with 10 – 12 barriers. Parietal lamella is the strongest in the aperture, long and curved, sinuated in its middle part. Angular lamella long, thin, and continuous, reaching the peristome. There are four main palatal plicae (upper palatal, two interpalatals, and a lower palatal) and usually one smaller suprapalatal plica. Upper palatal plica strong, highest in its middle part and getting lower towards the peristome. Interpalatal plicae similar to each other and the upper palatal. Lower palatal plica is the strongest. There is a strong palatal tubercle sitting on the palatal lip of the peristome, sometimes of almost rectangular shape. Peristome is distinctly sinuated behind this tubercle. Below the palatal tubercle, there are usually one or two more tubercle-like swellings which are of unknown homology. Basal plica very weak and short. Columellar lamella almost horizontal and slightly weaker than upper and lower palatal plicae. Infraparietal lamella similar to the basal plica. Surface of all apertural barriers is finely granulated. Sinulus small, narrow, and distinctly separated from the rest of the aperture. Umbilicus very narrow, dot like and elongated.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
4A0A2A9498A556FD8FEB6C1E31E2371A.taxon	description	Measurements (in mm, n = 5). SH = 1.55 – 1.78; SW = 1.3 – 1.57; AH = 0.65 – 0.80; AW = 0.65 – 0.70.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
4A0A2A9498A556FD8FEB6C1E31E2371A.taxon	etymology	Etymology. The specific epithet refers to the cyclophoroid subfamily Alycaeinae W. T. Blanford, 1864 due to the changing rib density on the last whorl. To be used as a noun in apposition.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
4A0A2A9498A556FD8FEB6C1E31E2371A.taxon	distribution	Distribution. This species is known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
E6A978DAC6675D6DA73A0DD2B0B34AD4.taxon	description	Figs 39 B, 42, 43, 60	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
E6A978DAC6675D6DA73A0DD2B0B34AD4.taxon	distribution	Distribution. This species is described from Java, Indonesia, but is also known from its synonym (B. novemdentata) in Laos. It has also been found in E India.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
1B07E1B30EC657ADB2E3370E1B034C1B.taxon	description	Figs 101 A, 102, 107	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
1B07E1B30EC657ADB2E3370E1B034C1B.taxon	diagnosis	Diagnosis. Bensonella species with obscure, weak spiral striae, asymmetric heart-shaped aperture, an angular lamella, and a palatal tubercle on the peristome and six barriers (upper palatal, lower palatal, basal, columellar, infraparietal, parietal) situated deeper inside the aperture.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
1B07E1B30EC657ADB2E3370E1B034C1B.taxon	description	Description. Shell colourless (white), conical-ovoid with strongly widened last whorl. Whorls 3.5, nearly rounded, slightly shouldered. Protoconch consisting of 1.25 whorls, pitted, showing spiralling pattern, although no clear spiral striae are visible. Teleoconch sculpture roughly pitted. Last whorl adnate to the penultimate, very slightly ascending (~ 10 ° compared to the shell axis), thus making the apertural profile opisthocline to the shell axis. Aperture asymmetric heart-shaped, the parietal part being the shield-like extended peristome. Angular lamella and palatal tubercle situated on the peristome, both are pointed, directed towards each other. Other barriers, altogether 6, are lamella-like, elongated, low. Upper palatal plica sits behind palatal tubercle, lower palatal situated between the basal and upper palatal plica. Columella lamella situated halfway between basal and infraparietal lamella, infraparietal and parietal situated close to each other. Peristome strongly expanded, not reflected. Surface of all apertural barriers is finely granulated. Sinulus relatively narrow and clearly separated from the rest of the aperture due to the strong and closely positioned angular lamella and palatal tubercle. Umbilicus very narrow, shows only last whorl. Last quarter whorl sigmoid.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
1B07E1B30EC657ADB2E3370E1B034C1B.taxon	description	Measurements (in mm, n = 1). SH = 1.12; SW = 1.17; AH = 0.56; AW = 0.58.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
1B07E1B30EC657ADB2E3370E1B034C1B.taxon	etymology	Etymology. Named after the heart-shaped aperture.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
1B07E1B30EC657ADB2E3370E1B034C1B.taxon	distribution	Distribution. This species is known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
6AF9CBF51F8758D7ADBE188AF977CD68.taxon	description	Figs 39 C, 44, 45, 100	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
6AF9CBF51F8758D7ADBE188AF977CD68.taxon	diagnosis	Diagnosis. A small Bensonella species with fine, finely dimpled teleoconch sculpture, rounded last whorl adnate to the penultimate, and a strong, sharp cervical crest right behind the expanding peristome. Palatal plicae long and equally developed; umbilicus very narrow to almost closed.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
6AF9CBF51F8758D7ADBE188AF977CD68.taxon	description	Description. Shell triangular, slightly conical-ovoid, brownish or yellowish, weakly glossy. It is consisting of 4.5 – 5 regularly increasing, convex, rounded whorls separated by a deep suture. Protoconch consisting of ~ 1.5 coarsely spirally striated whorls. There are ~ 13 spiral striae on the protoconch and they are more densely arranged at the bottom of the whorl. Teleoconch surface finely dimpled (pasty) and with not numerous, weak radial growth lines. Spiral striae present initially on the teleoconch but getting lost so that the surface remains finely pasty-like or a very weak spiralling pattern might be visible (only on SEM images). Last whorl rounded, adnate to the penultimate. Right behind the peristome, there is a strong, thick, sharp, white cervical crest which is wider than the peristome and even visible behind it in the apertural view. Peristome thick and expanded but not reflected. Aperture equipped with numerous barriers, four of which are particularly strong (angular, parietal, infraparietal, and columellar), and situated closer to the peristome edge. Parietal lamella is the strongest and highest in the aperture. Angular lamella much longer than the parietal, very slender but continuous. Its inner and outer parts are higher than its middle part. Columellar lamella similar in length to the parietal only slightly weaker. This lamella is almost horizontal but sometimes slightly curved towards the palatal plicae. Infraparietal lamella is roughly half as strong as the parietal. Palatal plicae equidistant, all of equal height and length, altogether four (one upper palatal, two interpalatals, and a lower palatal). They are all high at their inner parts and then regularly lowering and tapering towards their outer parts. Palatal tubercle moderately strong or absent. Basal plica present, the same length as the palatals or slightly shorter. Surface of all apertural barriers is finely granulated. Sinulus small and narrow (resembling a comma sign), distinctly separated from the rest of the aperture. Umbilicus very narrow (measuring ~ 1 / 10 of the shell width) or even dot-like, almost closed.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
6AF9CBF51F8758D7ADBE188AF977CD68.taxon	description	Measurements (in mm, n = 5). SH = 1.77 – 1.86; SW = 1.45 – 1.52; AH = 0.73 – 0.80; AW = 0.63 – 0.76.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
6AF9CBF51F8758D7ADBE188AF977CD68.taxon	etymology	Etymology. This species is named for the strong cervical crest which is the strongest among all the congeners.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
6AF9CBF51F8758D7ADBE188AF977CD68.taxon	distribution	Distribution. This species in known from three localities in Shan State, Myanmar.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
83D19BD115155B0AA694B46165FA96BC.taxon	description	Figs 39 D, 46, 47, 97	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
83D19BD115155B0AA694B46165FA96BC.taxon	diagnosis	Diagnosis. Bensonella species with conical shell which is not spirally striated. Apertural barriers numerous (angular, parietal, five palatals, one basal, one subcolumellar, one columellar, and one infraparietal) and with additional, strong palatal tubercle. Aperture well rounded.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
83D19BD115155B0AA694B46165FA96BC.taxon	description	Description. Shell conical to conical-ovoid, light brownish and weakly glossy, consisting of 4.25 – 5.5 convex rounded whorls separated by a deep suture. Protoconch roughly pitted, sometimes with a spiralling pattern and consisting of ~ 1.5 – 1.75 whorls. Teleoconch with fine pasty structure, not spirally striated but with very fine radial growth lines. Last whorl adnate to the penultimate, aperture profile only slightly prosocline to the shell axis due to the slightly descending last whorl (~ 15 ° compared to the shell axis). Peristome slightly lighter than the rest of the shell, expanded but not reflected. It is particularly expanded on the parietal side where it forms a weak callus which is leaned on the penultimate whorl. Aperture equipped with moderately strong, numerous barriers. Parietal lamella moderately strong and high. Angular lamella longer but overall weaker than the parietal. It is higher in its inner part and regularly tapering towards the peristome, which is not reached. There are two or three very weak barriers inside the sinulus. Palatal wall equipped with five long barriers (suprapalatal, upper palatal, interpalatal, lower palatal, and infrapalatal) and a single strong palatal tubercle sitting on the edge of the palatal lip. Suprapalatal plica weak, not much stronger than the sinulus plicae. Upper palatal plica only slightly stronger than the suprapalatal. Interpalatal moderate. Lower palatal is the strongest among all the palatal plicae. Infrapalatal weak and shorter than others. Basal plica developed to the same extent as the infrapalatal. Subcolumellar lamella weak and short (nearly the same as the basal and infrapalatal). Columellar lamella strong as the lower palatal, almost horizontal. Infraparietal lamella moderate, ~ ½ as strong as the columellar. Surface of all apertural barriers is finely granulated. Sinulus small and not very distinctly separated from the rest of the aperture. Umbilicus very narrow, dot-like.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
83D19BD115155B0AA694B46165FA96BC.taxon	description	Measurements (in mm, n = 13). SH = 1.56 – 2.02; SW = 1.14 – 1.53; AH = 0.64 – 0.78; AW = 0.64 – 0.73.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
83D19BD115155B0AA694B46165FA96BC.taxon	etymology	Etymology. Named after the Burmese word for sword / knife (dha) referring to the blade-like anterior part of the palatal plicae. To be used as a noun in apposition.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
83D19BD115155B0AA694B46165FA96BC.taxon	distribution	Distribution. This species is known from five localities in Shan State, Myanmar.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
EDE3449803FC5A39BA15190A618A5B78.taxon	description	Figs 39 E, 48, 49, 50, 51, 97	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
EDE3449803FC5A39BA15190A618A5B78.taxon	diagnosis	Diagnosis. Bensonella species with triangular, conical-ovoid shell and rounded whorls devoid of spiral striation. There are two palatal tubercles in front of a strong and frontally concave transversal plica. A single, strong basal plica present.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
EDE3449803FC5A39BA15190A618A5B78.taxon	description	Description. Shell triangular, conical-ovoid, consisting of 5 – 5.5 rounded, convex whorls separated by a deep suture. Protoconch consisting of ~ 1.25 – 1.5 roughly pitted whorls. Teleoconch devoid of spiral striation but with some fine radial growth lines, otherwise smooth, pasty. Last whorl adnate to penultimate. Aperture profile slightly opisthocline to the shell axis due to the very slightly ascending last whorl (~ 5 ° compared to the shell axis). In this view, a weak cervical crest could be observed behind the peristome. Peristome thick, expanded (especially on the parietal side where it forms a thick callus) but not reflected. Sinulus small and distinctly separated from the rest of the aperture. Aperture equipped with nine barriers (angular, parietal, upper palatal, transversal, two palatal tubercles, basal, columellar, and infraparietal). Parietal lamella very strong and high, curved and blade-like. Angular lamella much weaker and lower but longer, reaching the expanding peristome. Upper palatal plica moderate, higher in its middle parts. Transversal plica strong and with its frontal surface concave. In front of the transversal plica, there are two strong palatal tubercles. The upper one (usually present in all Bensonella species) is narrower and more acute. The lower palatal tubercle is wider and blunter. Basal plica present, short and ~ ½ as strong as the columellar lamella. The latter is strong, wide, and almost horizontal. Infraparietal lamella as strong as the basal plica. Surface of all apertural barriers is finely granulated. Umbilicus very narrow and slightly elongated, dot-like.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
EDE3449803FC5A39BA15190A618A5B78.taxon	description	Measurements (in mm, n = 6). SH = 1.80 – 2.21; SW = 1.44 – 1.80; AH = 0.64 – 0.96; AW = 0.62 – 0.94.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
EDE3449803FC5A39BA15190A618A5B78.taxon	etymology	Etymology. Two strong palatal tubercles of this species resemble teeth of a vampire. The specific epithet refers to the iconic vampire, Count Dracula. To be used as a noun in apposition.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
EDE3449803FC5A39BA15190A618A5B78.taxon	distribution	Distribution. This species is known from the surroundings of Hopong and Pinlaung in Shan State, as well as Phruno river cave, Kayah State.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
D7497534A48052AD8846965726961082.taxon	description	Figs 39 F, 52, 53, 54, 100	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
D7497534A48052AD8846965726961082.taxon	diagnosis	Diagnosis. Colourless Bensonella species with triangular conical-ovoid and spirally striated whorls. Last whorl slightly ascending near the aperture. Aperture equipped with eight strong barriers (angular, parietal, infraparietal, columellar, lower palatal, interpalatal, upper palatal, and a palatal tubercle). Umbilicus narrow but clearly wider than in majority of congeners.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
D7497534A48052AD8846965726961082.taxon	description	Description. Shell strongly triangular, conical-ovoid, colourless, consisting of 5 – 5.5 regularly growing whorls separated by a moderately deep suture. All whorls rounded, convex. Protoconch-teleoconch boundary not clearly visible due to the similar surface sculpture and colouration. Protoconch with ten coarsely and equidistantly spaced spiral striae. Teleoconch also spirally striated but additionally with coarse radial growth lines which are roughly equidistant. Spacing between two spiral striae irregular, it ranges from the width of two to the width of four spiral striae. Last whorl flat at its base, adnate to the penultimate (but peristome not fully leaning on it), slightly ascending near the aperture (~ 5 ° compared to the shell axis) making the apertural profile weakly opisthocline to the shell axis. Peristome white, expanded but not reflected. Sinulus well-rounded and distinctly separated from the rest of the aperture. Aperture equipped with eight barriers (angular, parietal, infraparietal, columellar, lower palatal, interpalatal, upper palatal, and a palatal tubercle). All barriers relatively strong. Parietal lamella is the strongest in the aperture and wavy. Angular lamella is longer than the parietal and reaching closer to the expanding peristome. It is sinuated in its middle part. Infraparietal lamella ~ 1.5 × weaker than the columellar, short. Columellar lamella strong as the lower palatal or slightly stronger. It is positioned obliquely to the shell axis, directed towards the parietal lamella. Basal plica absent. Palatal plicae all equally strong and relatively short, upper palatal is usually the shortest. Interpalatal situated halfway between the upper and lower palatal. Palatal tubercle prominent, sitting on the palatal lip and roughly in the level with upper palatal plica. Surface of all apertural barriers is finely granulated. Umbilicus narrow (but clearly wider than in the majority of congeners), measuring ~ 1 / 8 – 1 / 9 of the shell width.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
D7497534A48052AD8846965726961082.taxon	description	Measurements (in mm, n = 5). SH = 1.77 – 1.89; SW = 1.58 – 1.71; AH = 0.71 – 0.77; AW = 0.66 – 0.68.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
D7497534A48052AD8846965726961082.taxon	etymology	Etymology. This species is named exploda after its type locality, the “ Big Bang Cave ”.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
D7497534A48052AD8846965726961082.taxon	distribution	Distribution. This species is known only from two localities in Shan State, Pinlaung.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
D8ED8A92D14B5EFAA96C55CF37D5711F.taxon	description	Figs 55, 56	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
D8ED8A92D14B5EFAA96C55CF37D5711F.taxon	distribution	Distribution. This species is known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
E4B317B9D8295265900D725F9B28567D.taxon	description	Figs 101 B, 103, 104, 107	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
E4B317B9D8295265900D725F9B28567D.taxon	diagnosis	Diagnosis. Bensonella species with spirally striated teleoconch and strongly convex whorls separated by a deep suture. Last whorl narrow, slightly detached, rounded. Apertural barriers usually 7. Upper palatal plica bipartite, consisting of anterior and posterior bulge.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
E4B317B9D8295265900D725F9B28567D.taxon	description	Description. Shell conical, light yellowish-brownish, opaque. It is consisting of ~ 5 regularly increasing, strongly convex whorls separated by a deeply impressed suture. Protoconch finely pitted, showing a spiralling pattern (which is stronger terminally) and consisting of ~ 1.5 whorls, coloured as the rest of the shell. Teleoconch with sculpture consisting of fine, raised, and innumerable spiral striae which are occasionally crossed by much weaker radial growth lines (rarely present as strong white streaks). The sculpture of the teleoconch gets more prominent as the whorls are increasing, thus being most noticeable on the last two whorls. Last whorl relatively narrow (low), slightly detached from the penultimate, and slightly ascending near the aperture (~ 5 – 10 ° compared to the shell axis). It is completely rounded and strongly convex. Peristome the same colour as the rest of the shell, strongly expanded with finely pitted surface, not reflected. Aperture equipped with five main barriers (angular, parietal, upper palatal, lower palatal, and columellar). Angular lamella short and thick, directed towards the lower palatal plica. Parietal lamella longer than the angular, leaned towards the upper palatal plica. Upper palatal plica usually bipartite, consisting of anterior and posterior bulge. Anterior bulge is closer to the peristome, smaller and lower. Posterior bulge is longer, higher, and stronger, positioned behind the anterior (deeper in the aperture). Origin of the anterior bulge is not known but it may be homologous with the palatal tubercle usually found in many typical Bensonella species. Columellar lamella developed to the similar extent as the palatal plicae. In front of the columellar lamella, there is sometimes a small swelling on the peristome. Apart from these main barriers, there is usually an additional smaller basal plica and one even smaller knob-like lamella in the columello-parietal embayment. Only the angular lamella and anterior bulge of the upper palatal plica are reaching t peristome. Surface of all apertural barriers is finely granulated. Sinulus small, well separated from the rest of the shell due to the closely situated angular lamella and upper palatal plica. Umbilicus moderately wide (1 / 5 – 1 / 6 of the shell width), showing only the penultimate whorl. Umbilical groove absent.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
E4B317B9D8295265900D725F9B28567D.taxon	description	Measurements (in mm, n = 4). SW 1 = 2.09 – 2.18; SW 2 = 1.31 – 1.48; SH = 1.9 – 2.08; AH = 0.81 – 1.1; AW = 0.89 – 1.23.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
E4B317B9D8295265900D725F9B28567D.taxon	etymology	Etymology. The first examined specimen of this species had a crack on the apical whorls, which gave the specific epithet fracta.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
E4B317B9D8295265900D725F9B28567D.taxon	distribution	Distribution. This species is known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
A2C203FEE1E752A1A91CEDA97DD03617.taxon	description	Figs 39 G, 57, 100	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
A2C203FEE1E752A1A91CEDA97DD03617.taxon	distribution	Distribution. This species is known from different altitudes at the type locality (altitude range: 1650 – 2015 m).	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
7F5AEA59A815501984DEA68197701E66.taxon	description	Figs 39 H, 58	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
7F5AEA59A815501984DEA68197701E66.taxon	distribution	Distribution. SE and SW Himalaya region.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
92F5A80D85FD53E0866D9AF74756E6E0.taxon	description	Figs 39 I, 59, 60	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
92F5A80D85FD53E0866D9AF74756E6E0.taxon	distribution	Distribution. This species is known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
D00007D260145D33AC6E811CD2C945CC.taxon	description	Figs 39 J, 61	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
D00007D260145D33AC6E811CD2C945CC.taxon	distribution	Distribution. This species is well known from Japan as well as Taiwan and China (where it is known from a synonym, B. qingliangfengensis) (Fang et al. 2015; Páll-Gergely and White 2023).	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
975238FFB3ED573D896FDCD05D796E5F.taxon	description	Figs 39 K, 62, 63, 97	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
975238FFB3ED573D896FDCD05D796E5F.taxon	distribution	Distribution. This species is known from three localities in Kalaw.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
DAA9BE7AA09550DDAE4043AC30C7B0BF.taxon	description	Figs 39 L, 64, 65, 95	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
DAA9BE7AA09550DDAE4043AC30C7B0BF.taxon	diagnosis	Diagnosis. Bensonella species with concave-conical, glossy shell that is spirally striated. Apertural barriers (usually eight or nine) and the surface around them prominently spiniferous.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
DAA9BE7AA09550DDAE4043AC30C7B0BF.taxon	description	Description. Shell slightly concave-conical, glossy, light brownish, consisting of 5 – 5.5 convex and rounded whorls separated by a deep suture. Protoconch spirally striated, its boundary with the teleoconch not clear. There are ~ 12 widely spaced spiral striae on the protoconch and they are stronger at the base of the whorl. Teleoconch with relatively coarsely spaced, raised spiral striae (space between two striae equals the width of two or three striae) and with fine radial growth lines. On initial teleoconch whorls, radial lines are more densely arranged (similar to the spacing between the spiral striae) and are crossing the spiral striae forming a reticulated sculpture. Towards the last whorl, radial lines become coarser and the reticulated sculpture becomes weaker. Last whorl rounded, adnate to the penultimate, slightly descending near the aperture (~ 20 ° compared to the shell axis). Peristome whitish, expanded but not reflected. Sinulus parabolic due to the position of the angular lamella and the palatal tubercle. Aperture equipped with numerous barriers which have the typical Bensonella arrangement. These are: angular, parietal, upper palatal, interpalatal, lower palatal, palatal tubercle, basal, columellar, and infraparietal. Parietal lamella strong and moderately high, angular separated into inner and outer parts of almost equal height and width and a strong sinuation between them. Upper and interpalatal plicae positioned close together and almost equally as strong. Lower palatal plica situated some distance from other palatals and stronger than both of them. Palatal tubercle strong and sitting on the edge of the palatal lip. Basal plica weak, low, and short. Columellar lamella nearly as strong as the lower palatal. Infraparietal lamella only very slightly stronger than the basal plica. Surface of all apertural barriers is roughly spiniferous. Umbilicus narrow but relatively deep, measuring ~ 1 / 6 of the shell width.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
DAA9BE7AA09550DDAE4043AC30C7B0BF.taxon	description	Measurements (in mm, n = 4). SH = 1.73 – 2; SW = 1.47 – 1.61; AH = 0.73 – 0.83; AW = 0.63 – 0.75.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
DAA9BE7AA09550DDAE4043AC30C7B0BF.taxon	etymology	Etymology. This species is named for its small apertural barriers.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
DAA9BE7AA09550DDAE4043AC30C7B0BF.taxon	distribution	Distribution. This species is known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
541B399B3B525192928139DDAF9FB6A5.taxon	description	Figs 39 M, 66, 67, 95	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
541B399B3B525192928139DDAF9FB6A5.taxon	diagnosis	Diagnosis. Bensonella species with seven apertural barriers and a prominent hooked upper palatal plica which is pointing laterally (toward the palatal wall).	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
541B399B3B525192928139DDAF9FB6A5.taxon	description	Description. Shell triangular, slightly conical-ovoid, brownish, consisting of 5 – 5.5 whorls separated by a deep suture. All whorls rounded, convex, regularly increasing. Protoconch coarsely spirally striated, ~ 1.5 whorls. Spiral striae much more prominent terminally on the protoconch. Teleoconch with a fine pasty surface sculpture, without spiral striae but with fine radial growth lines. Last whorl rounded, adnate to the penultimate and straight near the aperture. Peristome brownish as the rest of the shell, expanded but not reflected. It is leaning on the penultimate whorl in form of a strong but not much expanded parietal callus. Palatal side of the peristome sinuated in its middle part, next to the palatal tubercle. Behind the expanding peristome, a weak cervical crest is present. Sinulus small and narrow, distinctly separated from the rest of the aperture. Aperture subrectangular, equipped with a smaller number of barriers (angular, parietal, upper palatal, lower palatal, columellar, infraparietal and palatal tubercle) than usual in Bensonella. Parietal lamella long but not reaching the expanding peristome. Angular lamella consisting of two separated parts: inner, which is slightly curved and positioned deeper in the aperture and outer part which is reaching the peristome edge. There is a strong sinuation between the inner and outer parts of the angular lamella. Palatal wall equipped with upper and lower palatal plica. Upper palatal plica of very peculiar appearance: it is very high with a hook positioned laterally (not pointing outside but directly to the palatal wall). Lower palatal plica nearly as strong as the parietal lamella. In front of the upper palatal plica, on the peristome edge, a strong palatal tubercle is observed. Columellar lamella rather weak. Above the columellar lamella, one additional infraparietal is present and it is nearly equally strong as the columellar. Surface of all apertural barriers is finely granulated. Umbilicus oblique (slightly elongated) and narrow, measuring ~ 1 / 6 – 1 / 7 of the shell width.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
541B399B3B525192928139DDAF9FB6A5.taxon	description	Measurements (in mm, n = 5). SH = 2.03 – 2.16; SW = 1.52 – 1.70; AH = 0.80 – 0.88; AW = 0.77 – 0.82	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
541B399B3B525192928139DDAF9FB6A5.taxon	etymology	Etymology. Named for the remarkable upper palatal plica (laterally hooked).	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
541B399B3B525192928139DDAF9FB6A5.taxon	distribution	Distribution. Known from four sampling sites in Kayah state, Hpruso district.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
01F3593007FC5417ACE035B7545C9ABF.taxon	description	Figs 103 C, 105, 107	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
01F3593007FC5417ACE035B7545C9ABF.taxon	diagnosis	Diagnosis. A colourless Bensonella species with shouldered last whorl and spirally striated teleoconch and nearly heart-shaped aperture. Apertural barriers numerous and almost all equally strong, umbilicus moderately wide.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
01F3593007FC5417ACE035B7545C9ABF.taxon	description	Description. Shell conical-ovoid, colourless. Whorls 3.5, initial whorls rounded, last two strongly shouldered, separated by a deep suture. Protoconch consisting of ~ 1 whorl, very finely pitted, with signs of indistinct and widely spaced spiralling pattern. Teleoconch with coarse spiral striae which are crossed by stronger, coarse, not numerous, and almost regularly spaced radial growth lines. Approximately 25 widely spaced spiral striae present on the last whorl in standard apertural view. Last whorl adnate to the penultimate, slightly ascending near the aperture (~ 10 – 15 ° compared to the shell axis). Peristome continuous, strong, and expanded, especially at the parietal side where it leans on the penultimate whorl. Aperture nearly heart-shaped, equipped with a strong angular lamella, which is curved and almost merges with the prominent palatal tubercle, thus nearly enclosing the small sinulus (which makes it strongly separated from the rest of the aperture). Angular lamella reaches the peristome. There are eight to nine other barriers situated deep in the aperture and not reaching the peristome. Additional two lamellae at the parietal side (equally strong parietal and infraparietal), one columellar lamella, one subcolumellar, one basal plica, two palatal plicae (sometimes with additional, weak infrapalatal plica) and a palatal tubercle. Upper and lower palatal plicae strong, the former is situated slightly above the palatal tubercle. Palatal tubercle quite strong and elevated. Basal plica weaker than the palatals but stronger than the subcolumellar lamella. Columellar lamella slightly stronger than them but weaker than upper and lower palatal plica. Surface of all apertural barriers is finely granulated. Umbilicus moderately wide, measuring ~ 1 / 5 of the shell width and showing only the penultimate whorl.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
01F3593007FC5417ACE035B7545C9ABF.taxon	description	Measurements (in mm, n = 5). SH = 1.21 – 1.33; SW = 1.21 – 1.37; AH = 0.57 – 0.62; AW = 0.58 – 0.67.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
01F3593007FC5417ACE035B7545C9ABF.taxon	etymology	Etymology. The appearance of the apertural barriers when the last whorl is broken behind them resembles the morphology of a mitochondria (Fig. 105 F), hence the specific epithet which is to be used as a noun in apposition.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
01F3593007FC5417ACE035B7545C9ABF.taxon	distribution	Distribution. This species in known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
B698D88A4FC859D7B22E92907866E666.taxon	description	Figs 39 N, 68, 95	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
B698D88A4FC859D7B22E92907866E666.taxon	diagnosis	Diagnosis. A Bensonella species with conical, pale yellowish shell. Aperture equipped with numerous barriers, all of which are in form of sharp hooks pointing outside. Parietal lamella with a blunt but long projection in front of the hook, almost reaching the expanding peristome. Umbilicus narrow, dot-like.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
B698D88A4FC859D7B22E92907866E666.taxon	description	Description. Shell pale yellowish, triangular, conical ovoid, whorls 4 – 4.5, convex, rounded, rather regularly increasing. Protoconch consisting of ~ 1.25 whorls, showing spiralling pattern but with no clear spiral striae. Teleoconch with pasty surface, not spirally striated but with coarsely spaced, irregular radial growth lines. Last whorl adnate to the penultimate and slightly ascending near the aperture (~ 15 – 20 ° compared to the shell axis) making the aperture profile opisthocline to the shell axis. Peristome strongly expanded, not reflected. Aperture equipped with numerous barriers, all of which are in form of sharp hooks pointing outside the aperture. All barriers have low, slender projections situated in front of the hooks. A blunt, knob-like palatal tubercle sits anterior to the upper palatal plica on the peristome edge. Parietal lamella in form of a strong hook. A blunt, outer part of the parietal lamella is in form of a low, narrow, and blunt projection (situated in front of the hooked part) almost reaching the peristome edge. Angular lamella strong, consisting of inner and outer part separated by a strong sinuation. Inner part smaller, hooked, pointing towards the outer part which is larger but blunt. There are four weak barriers inside the sinulus. Upper palatal plica ~ 2 × weaker than the parietal and situated slightly above the palatal tubercle. First interpalatal plica as strong as the upper palatal, situated slightly below the palatal tubercle. Second interpalatal plica slightly weaker than the first. Lower palatal plica as strong as the first interpalatal and upper palatal. Infrapalatal plica small, as strong as the basal and subcolumellar. These three plicae are roughly equidistant. Above the subcolumellar, there is a clearly stronger columellar (similar in strength to the lower palatal plica) and one slightly stronger supracolumellar. Infraparietal lamella ~ 1 / 3 weaker than the parietal, with a similar (but shorter) projection in front of the hooked part. There are also some barrier-like swellings, although they never develop into a proper barrier. They can be e. g., between the upper palatal and first interpalatal plica, between the columellar and supracolumellar lamellae, between the supracolumellar and infraparietal lamellae, between the infraparietal and parietal lamellae. Surface of all apertural barriers is smooth. Sinulus parabolic, well separated from the rest of the aperture. Umbilicus very narrow, dot-like.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
B698D88A4FC859D7B22E92907866E666.taxon	description	Measurements (in mm, n = 3). SH = 1.8; SW = 1.4 – 1.6; AH = 0.69 – 0.77; AW = 0.74 – 0.79.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
B698D88A4FC859D7B22E92907866E666.taxon	etymology	Etymology. This species is named after its type locality, the Montawa Cave. The specific epithet is to be used as a noun in apposition.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
B698D88A4FC859D7B22E92907866E666.taxon	distribution	Distribution. This species is only known from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
16D64A5B58B2501DBC9E61C90319576B.taxon	description	Figs 39 O, 69, 70, 71, 72, 100	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
16D64A5B58B2501DBC9E61C90319576B.taxon	diagnosis	Diagnosis. A Bensonella species with high triangular, concave-conical, densely spirally striated shell. Last whorl wide, rounded. Multiple (at least 10) well-developed apertural barriers.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
16D64A5B58B2501DBC9E61C90319576B.taxon	description	Description. Shell light brown, triangular, concave-conical, whorls 5 – 5.25, convex, rounded, rather regularly increasing. Protoconch consisting of ~ 1.25 whorls, its sculpture consists of ~ 10 equidistant spiral striae. Teleoconch finely, very densely spirally striated, and ornamented with a few, inconspicuous radial growth lines. Last whorl adnate to the penultimate and slightly to moderately descending near the aperture (~ range: 15 – 32 ° compared to the shell axis) making the aperture profile prosocline to the shell axis. Aperture subcircular, parietal part strongly extended towards penultimate whorl. Peristome strongly expanded, not reflected. Aperture with numerous and strong barriers. A blunt, knob-like palatal tubercle sits anterior to the upper palatal plica on the peristome edge. Sometimes, this tubercle is very weak or completely absent. All other barriers sit deeper, not reaching the peristome edge. Parietal lamella is the highest among all apertural barriers, long, almost straight, only slightly curved downwards. Angular lamella much lower and more slender than the parietal one. Upper palatal plica short and located slightly above the palatal tubercle. First interpalatal and lower palatal plicae equally strong. There is a smaller and shorter second interpalatal plica between them. Basal part of peristome with two or three low plicae (rarely one), they are situated next to each other, the one situated closer to the columellar lamella may be homologous with the subcolumellar lamella, and the other may be basal plicae. Columellar lamella is weaker than parietal and stronger than the palatal plicae, straight, pointing obliquely upwards. Supracolumellar and infraparietal denticles small, knob-like. Surface of all apertural barriers is finely granulated. Sinulus parabolic, wide, and not strongly separated from the rest of the aperture. Umbilicus wide, covers 1 / 3 – 1 / 4 of shell width.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
16D64A5B58B2501DBC9E61C90319576B.taxon	description	Measurements (in mm, n = 5). SH = 2.40 – 2.51; SW = 1.81 – 1.97; AH = 0.82 – 0.94; AW = 0.85 – 0.95.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
16D64A5B58B2501DBC9E61C90319576B.taxon	etymology	Etymology. Named for the numerous barriers in the aperture.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
16D64A5B58B2501DBC9E61C90319576B.taxon	distribution	Distribution. This species is known from four localities in Chiang Rai province, Thailand.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
748DFECAEA235C35BB5F71A70DC9EA9C.taxon	description	Figs 39 P, 73, 88, 100	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
748DFECAEA235C35BB5F71A70DC9EA9C.taxon	distribution	Distribution. This species is, apart from its type locality, known from one additional site: Doi Ang Khang (1400 m).	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
8AD6F47946E85BFE897FDC3453AF4699.taxon	description	Figs 39 Q, 74, 75, 100	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
8AD6F47946E85BFE897FDC3453AF4699.taxon	diagnosis	Diagnosis. Shell concave-conical, glossy, last whorl strongly enlarged. Teleoconch spirally striated. Last whorl adnate to the penultimate. Aperture equipped with numerous barriers, angular and columellar lamellae reaching the peristome. Umbilicus moderately wide.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
8AD6F47946E85BFE897FDC3453AF4699.taxon	description	Description. Shell concave-conical, brownish, consisting of 5 – 5.5 convex, rounded whorls separated by a deep suture. Last whorl much wider than all preceding ones, resulting in a concave-conical shell shape. Protoconch finely spirally striated initially, more prominently terminally. Its boundary with the teleoconch is not clearly visible due to the similar surface sculpture. Fine spiral striae present across the whole shell surface but quite difficult to observe on initial teleoconch whorls without SEM imaging. They are most clearly visible on the last whorl but still very fine. Spacing between the two spiral striae approximately measures the width of one or two striae. Radial growth lines extremely fine, irregularly spaced and not numerous, most clearly visible on the last and the penultimate whorls. Last whorl rounded, slightly descending (~ 10 ° compared to the shell axis), and adnate to the penultimate near the aperture. Peristome brownish, expanded but not reflected. On the parietal side of the aperture, the peristome leans on the penultimate whorl forming a thin parietal callus. Aperture equipped with numerous barriers (angular, parietal, upper palatal, two interpalatals, lower palatal, palatal tubercle, basal, columellar, and infraparietal). Parietal lamella is the strongest in the aperture, blade like and relatively high. Angular lamella discontinuous, consisting of weaker outer part (which is reaching the expanding peristome) and stronger (higher) inner part. Palatal plicae almost all equidistant. Upper palatal and first interpalatal equally strong, higher in their inner than in outer parts (but upper palatal can be weaker than the interpalatal). There is a strong, white palatal tubercle sitting on the palatal lip of the aperture, in front and between the upper palatal and interpalatal plica. Lower interpalatal plica is the weakest among them all. Lower palatal plica is the strongest among all the palatals. Sometimes, a very weak infrapalatal plica is also present and situated between the lower palatal and basal. All palatal plicae continue as slender projections towards the peristome. Basal plica small and usually the weakest in the aperture. Columellar lamella strong and oblique, leaned towards the parietal lamella. Infraparietal lamella slightly stronger than the basal plica. Surface of all apertural barriers is finely granulated. Sinulus parabolic due to the position of the angular lamella and the palatal tubercle. Umbilicus moderately wide, measuring ~ 1 / 5 of the shell width.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
8AD6F47946E85BFE897FDC3453AF4699.taxon	description	Measurements (in mm, n = 5). SH = 1.91 – 2.00; SW = 1.57 – 1.87; AH = 0.86 – 0.96; AW = 0.86 – 0.94.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
8AD6F47946E85BFE897FDC3453AF4699.taxon	etymology	Etymology. The specific epithet refers to the shiny shell surface.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
8AD6F47946E85BFE897FDC3453AF4699.taxon	distribution	Distribution. This species is known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
A705D508F16E53AF899F5E6715BBC51A.taxon	description	Figs 39 R, 76, 100	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
A705D508F16E53AF899F5E6715BBC51A.taxon	distribution	Distribution. This species is known only from the type locality, with an altitude range from 605 m to 2015 m.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
C2E17F1CDC7F5F8CA4222653DDAB8757.taxon	description	Figs 39 S, 77, 78, 97	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
C2E17F1CDC7F5F8CA4222653DDAB8757.taxon	diagnosis	Diagnosis. Bensonella species with a pitted protoconch and conical-ovoid shell. Last whorl slightly ascending near the aperture. Peristome usually wrinkled due to several tubercles. Transversal plica strong and very weakly concave.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
C2E17F1CDC7F5F8CA4222653DDAB8757.taxon	description	Description. Shell conical-ovoid, consisting of 4.5 – 5 rounded, convex, regularly increasing whorls separated by a deep suture. Shell colour slightly weathered in the holotype but probably light brownish. Protoconch consisting of 1.75 whorls, finely pitted or sometimes spirally striated, especially terminally. Teleoconch finely pitted, with a pasty sculpture and without visible spiral striae but with coarsely and irregularly spaced radial growth lines. Last whorl enlarged, adnate to the penultimate and slightly ascending near the aperture (~ 20 – 25 ° compared to the shell axis). Aperture profile opisthocline to the shell axis. Peristome white and expanded (at columellar but especially at the parietal side where it forms a distinct callus) but not reflected. Aperture equipped with numerous barriers. Parietal lamella strong and high, almost reaching the expanding peristome. It sometimes has a heart-like appearance due to the distinct sinuation in the middle. Angular lamella weakly wavy, slightly weaker than the parietal but longer, reaching the expanding peristome. It also has a sinuation as in the parietal lamella but weaker. Upper palatal plica very short and moderately strong. Transversal plica strong, slightly concave on the front surface. A strong palatal tubercle is sitting in front and between the upper palatal and transversal plica. Below this tubercle, there are several more tubercle-like swellings which give the peristome a wrinkled appearance. Infrapalatal plica elongated and low. Basal plicae number ranging from one to four, similarly shaped as the infrapalatal plica. Subcolumellar lamella (if present) shares the same appearance as the former two. Columellar lamella moderate, ~ ½ as strong as the parietal. Supracolumellar lamella present but can be strong or quite weak. Infraparietal lamella slightly weaker than the columellar or even half as strong. Surface of all apertural barriers is finely granulated. Sinulus relatively narrow, small, and distinctly separated from the rest of the aperture. Umbilicus very narrow, measuring ~ 1 / 10 of the shell width, slightly elongated.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
C2E17F1CDC7F5F8CA4222653DDAB8757.taxon	description	Measurements (in mm, n = 10). SH = 1.42 – 1.8; SW = 1.36 – 1.5; AH = 0.61 – 0.81; AW = 0.63 – 0.70.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
C2E17F1CDC7F5F8CA4222653DDAB8757.taxon	etymology	Etymology. The specific epithet comes from a Latin word for a barricade (obex), which refers to the transversal palatal plica in the form of a barrier / obstacle. To be used as a noun in apposition.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
C2E17F1CDC7F5F8CA4222653DDAB8757.taxon	distribution	Distribution. This species is known from three localities around Pinlaung, Shan State.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
1AD8DA2A599D566DB664956EEFFC3F2B.taxon	description	Figs 39 T, 79, 100	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
1AD8DA2A599D566DB664956EEFFC3F2B.taxon	distribution	Distribution. This species in known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
C408234488FD516CB093F4F379DCA9C0.taxon	description	Figs 39 U, 80, 100	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
C408234488FD516CB093F4F379DCA9C0.taxon	distribution	Distribution. This species is known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
601594F9B8475CC9882C68E6D3EFCFE1.taxon	description	Figs 39 V, 81, 100	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
601594F9B8475CC9882C68E6D3EFCFE1.taxon	distribution	Distribution. This species is known from several sampling sites, all located in Mae Hong Son province where the type locality is also situated.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
3A2619A2810D57C7A30768F2911CAA88.taxon	description	Figs 39 W, 82, 83, 100	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
3A2619A2810D57C7A30768F2911CAA88.taxon	distribution	Distribution. This species is known from several localities in Laos (Xieng Khouang, Udomxai and Luang Prabang provinces), Vietnam (Son La province), Myanmar (Shan State, see Tongkerd et al. 2024, Mandalay) as well as Thailand (Chiang Mai, Mae Hong Son and Chaiyapumi provinces).	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
8E04333ECE0E542BAEA5ACF60204DC97.taxon	description	Figs 39 X, 84, 85, 95	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
8E04333ECE0E542BAEA5ACF60204DC97.taxon	diagnosis	Diagnosis. Bensonella species with a conical, brownish shell that is not spirally striated. Apertural barriers numerous: five plicae (4 palatals and one basal), all weak and low. Palatal tubercle strong.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
8E04333ECE0E542BAEA5ACF60204DC97.taxon	description	Description. Shell conical, pale brown, consisting of 5 – 5.5 regularly increasing, rounded whorls. Protoconch consisting of 1.75 whorls, same colour as the rest of the shell, finely pitted and spirally striated. Teleoconch with fine pasty like surface sculpture and coarse radial growth lines which are irregularly spaced. Rarely, these radial lines appear as stronger, whitish streaks. A spiralling pattern is very slightly visible on the last whorl (only under SEM) although there are no spiral striae. Last whorl rounded, adnate to the penultimate and slightly descending near the aperture (~ 5 – 10 ° compared to the shell axis). Apertural profile slightly prosocline to the shell axis. Peristome of same colour as the rest of the shell, expanded but not reflected. There is a weak cervical crest just behind the peristome. Aperture equipped with numerous barriers. Parietal lamella strong and blade-like, not reaching the peristome. Angular lamella closer to the peristome but appears discontinuous as its outer and inner parts are separated by a distinct sinuation. These two parts are roughly equally strong. There are one or two barriers inside the sinulus. Four palatal plicae (upper palatal, interpalatal, lower palatal and infrapalatal). Upper, inter and lower palatal are moderately long and high, equally developed and equally distanced. Sometimes, upper palatal is weak and lower palatal is stronger than others. Infrapalatal plica is much weaker (lower and shorter). Palatal tubercle is strong, sitting on the palatal lip of the peristome just below the line of upper palatal plica. Basal plica situated close to the infrapalatal and equally developed or sometimes weaker. Columellar lamella stronger than in most of the congeners (quite broad and high), almost horizontal, sometimes appears slightly wavy. Infraparietal lamella moderate, weaker than other lamellae but stronger than all the plicae. Surface of all apertural barriers is finely granulated. Additionally, plicae in the palatal and basal areas as well as the infraparietal and angular lamellae are equipped with isolated but strong spines which are positioned medially. Sinulus small and distinctly separated from the rest of the aperture. Umbilicus narrow, slightly elongated, measuring ~ 1 / 5 – 1 / 6 of the shell width.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
8E04333ECE0E542BAEA5ACF60204DC97.taxon	description	Measurements (in mm, n = 5). SH = 2.91 – 3.09; SW = 2.08 – 2.23; AH = 1.15 – 1.30; AW = 1.2 – 1.37.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
8E04333ECE0E542BAEA5ACF60204DC97.taxon	etymology	Etymology. This species is named for the exceptionally fine surface sculpture.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
8E04333ECE0E542BAEA5ACF60204DC97.taxon	distribution	Distribution. This species is known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
BB0DFD4224905C5380E3F1D9D7472CA7.taxon	description	Figs 39 Y, 86	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
BB0DFD4224905C5380E3F1D9D7472CA7.taxon	distribution	Distribution. SW and SE Himalaya.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
2950672BE1265E3B98FFBFAE09F1D367.taxon	description	Figs 39 Z, 87, 88, 100	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
2950672BE1265E3B98FFBFAE09F1D367.taxon	diagnosis	Diagnosis. A Bensonella species with conical to conical-ovoid shell, very dense spiral striation, and a small number of apertural barriers, only four of which are strong (parietal, upper palatal, lower palatal and columellar).	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
2950672BE1265E3B98FFBFAE09F1D367.taxon	description	Description. Shell brown, conical to conical-ovoid, whorls 4.5 – 5, convex, regularly increasing. Protoconch consisting of ~ 1.5 whorls, its sculpture consists of ~ 10 roughly equidistant spiral striae. Teleoconch finely, very densely spirally striated, and ornamented with a few, inconspicuous radial growth lines. Last whorl adnate to the penultimate and slightly descending near the aperture (~ 15 ° compared to the shell axis) making the aperture profile prosocline to the shell axis. Peristome slightly expanded, not reflected. A blunt, weak, knob-like palatal tubercle sits anterior and slightly below the upper palatal plica on the peristome edge. Parietal lamella is the strongest in the aperture and slightly leaned towards the palatal wall. Angular lamella weak, its outer part reduced, inner part sits deep in the aperture. Upper palatal plica in form of a very weak swelling, situated slightly above the palatal tubercle. Interpalatal plica strong, located slightly below the palatal tubercle. Lower palatal plica nearly as strong as the parietal, stronger than the interpalatal. There can be two additional swellings: one between the upper and interpalatal and one between the interpalatal and lower palatal. Basal plica absent. Columellar lamella strong and slightly oblique. Infraparietal lamella in form of indistinct swelling located close to the parietal lamella. Surface of all apertural barriers is smooth. Sinulus wide and relatively well separated from the rest of the aperture due to the strong parietal lamella. Umbilicus moderately wide, measuring 1 / 5 of shell width.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
2950672BE1265E3B98FFBFAE09F1D367.taxon	description	Measurements (in mm, n = 4). SH = 2.26 – 2.39; SW = 1.69 – 1.73; AH = 0.94 – 1.01; AW = 0.87 – 0.94.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
2950672BE1265E3B98FFBFAE09F1D367.taxon	etymology	Etymology. The specific epithet refers to the silky surface of the shell (as if “ covered in silk ”).	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
2950672BE1265E3B98FFBFAE09F1D367.taxon	distribution	Distribution. This species is known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
F1476E8059305973AD61C74A6F4A36EE.taxon	diagnosis	Diagnosis. Bensonella species with pasty shell surface, last whorl adnate to the penultimate and palatal plicae with serrated, elevated inner and very low outer parts.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
F1476E8059305973AD61C74A6F4A36EE.taxon	description	Description. Shell triangular, slightly conical-ovoid, brown, not very glossy, consisting of 5 – 5.5 regularly increasing whorls separated by a deep suture. All whorls convex, rounded. Protoconch consisting of 1.5 whorls, finely pitted, showing spiralling pattern even though without clear spiral striae. The boundary between the protoconch and the teleoconch is visible due to the presence of radial growth lines on the teleoconch. Teleoconch surface finely dimpled, pasty and with less numerous, weak radial growth lines, spiral striae absent. Last whorl also rounded, adnate to the penultimate near the aperture. Last whorl slightly descending immediately behind the aperture (~ 10 ° compared to the shell axis) making the apertural profile slightly prosocline to the shell axis. Peristome weakly expanded but not reflected, its edges are thickened. Aperture equipped with numerous barriers, two of them relatively strong (parietal, columellar), others weak. Parietal lamella is the strongest and highest in the aperture. Angular lamella is the longest in the aperture, much longer than the parietal, very slender and low but continuous. Its inner and outer parts are higher than its middle part. Columellar lamella similar in length to the parietal but much weaker. There is one weak infraparietal lamella. On the palatal side, there are four main palatal plicae which are all almost equally strong and long (upper palatal, two interpalatals, and a lower palatal). They are all elevated at their inner ends and sloping towards the outer end, ending very low. These low parts are equipped with numerous small spines (serrated), thus resembling a saw. Palatal tubercle, sitting on the palatal lip, is weak but clearly present. There are several suprapalatal plicae (with similar morphology as in the main palatals) all weak and located inside the sinulus (the length and number (usually two to four) of these plicae is variable and not useful in species identification). There are usually two basal plicae (although one of them could be homologous with the infrapalatal plica), which are long but can occasionally be shorter. Surface of all barriers is finely granulated. Sinulus not strongly isolated from the rest of the aperture due to the low angular lamella and weak palatal tubercle. Umbilicus is narrow, measuring 1 / 8 of the shell width.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
F1476E8059305973AD61C74A6F4A36EE.taxon	description	Measurements (in mm, n = 5). SH = 2.52 – 2.86; SW = 1.80 – 2.06; AH = 0.94 – 1.03; AW = 0.83 – 0.94.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
F1476E8059305973AD61C74A6F4A36EE.taxon	etymology	Etymology. This species is named for the serrated (saw-like) palatal plicae.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
F1476E8059305973AD61C74A6F4A36EE.taxon	distribution	Distribution. This species is known from two localities situated ca 18 km apart (straight-line distance).	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
85FB2666144451FFA1AD40E199238617.taxon	diagnosis	Diagnosis. Bensonella species with 7 – 9 apertural barriers, most peculiar of these is the transversal plica (strong, high but with a concave frontal surface resembling a cave entrance).	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
85FB2666144451FFA1AD40E199238617.taxon	description	Description. Shell triangular, conical-ovoid, brownish, consisting of 5 – 5.5 convex, regularly increasing, rounded whorls separated by a deep suture. Boundary between the protoconch and the teleoconch is only clearly visible after SEM imaging because of the delicate sculpture and somehow weathered type series. Protoconch is spirally striated, probably ~ 1.75 whorls). Teleoconch of fine, pasty sculpture with dense but fine radial growth lines and without signs of spiral striation. Last whorl rounded, adnate to the penultimate. Aperture profile sinuated in its upper portion (concave) but convex in its middle and lower sections. Peristome expanded, especially at the parietal side where it leans on the penultimate whorl in form of a strong callus but not reflected. There is a moderate cervical crest located a short distance behind the peristome. Aperture equipped with relatively small number of barriers, seven to nine. Angular, parietal, upper palatal, transversal, palatal tubercle, columellar and infraparietal are always present, while basal and supracolumellar only sometimes. Parietal lamella heart-shaped in profile due to the sinuation located in its middle part (this makes the middle part of the lamella lower than outer and inner parts). Angular lamella also appears curvy, and it is curved to the palatal side. Upper palatal plica relatively weak and low. Transversal plica large (the largest in the aperture), high but with a strongly concave frontal surface (this gives the appearance of the transversal plica as an entrance to a cave). Basal very short and low when present. Columellar lamella very strong and thick (roughly the same as the parietal lamella). Supracolumellar lamella weak, dot like (sometimes absent). Infraparietal lamella present but much weaker than both the parietal and the columellar. Surface of all apertural barriers is granulated to almost spiniferous. Sinulus small and distinctly separated from the rest of the aperture due to the strong and closely positioned angular lamella and palatal tubercle. Umbilicus initially narrow and the widening at the last whorl, elongated, measuring ~ 1 / 6 – 1 / 7 of the shell width.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
85FB2666144451FFA1AD40E199238617.taxon	description	Measurements (in mm, n = 5). SH = 2.48 – 3.38; SW = 1.98 – 2.4; AH = 1.08 – 1.30; AW = 0.95 – 1.12.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
85FB2666144451FFA1AD40E199238617.taxon	etymology	Etymology. Named after the cave-like transversal plica.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
85FB2666144451FFA1AD40E199238617.taxon	distribution	Distribution. This species is known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
86E9288D36235017999C4A2A38DBE0DA.taxon	diagnosis	Diagnosis. Bensonella with triangular-conical, reddish-brown shell. Teleoconch fine, pasty, not spirally striated. All barriers and aperture surface around them very roughly spiniferous.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
86E9288D36235017999C4A2A38DBE0DA.taxon	description	Description. Shell triangular, slightly conical-ovoid, reddish brown, consisting of 5 – 5.5 convex, rounded whorls separated by a deep suture. Protoconch of ~ 1.5 whorls, finely pitted, showing very weak spiralling pattern, lighter than the rest of the shell but with no clearly visible boundary with the teleoconch. Teleoconch finely dimpled (pasty) and finely radially striated but without spiral striae. Last whorl adnate to the penultimate and very slightly ascending near the aperture (~ 5 – 10 ° compared to the shell axis), making the aperture profile weakly opisthocline to the shell axis. Peristome of the same colour as the rest of the shell, expanded and not reflected. Aperture equipped with numerous barriers. Parietal lamella is the strongest in the aperture, high but not curved, directed towards the palatal wall. Angular lamella slightly lower than the parietal but otherwise very similar, continuous. There are three main palatal plicae (upper palatal, interpalatal and lower palatal) and one additional small interpalatal plica situated below the stronger interpalatal and a lower palatal. Upper palatal as strong as the main interpalatal. Lower palatal plica is the strongest. Infrapalatal plica weak. Basal plica 2 × stronger than the infrapalatal. All palatal plicae are higher at their inner parts and sloping towards their outer parts. There is a strong palatal tubercle sitting on the palatal lip of the peristome, in front of the upper palatal plica. Peristome is distinctly sinuated behind this tubercle. Columellar lamella ~ 2 × stronger than the basal and positioned obliquely, directed towards the parietal lamella. Subcolumellar similar to the infrapalatal. Supracolumellar lamella similar to the subcolumellar. Infraparietal lamella strong as the columellar. All apertural barriers are very strongly spiniferous as well as the surface of the aperture around them. Sinulus small and well separated from the rest of the aperture due to the strong angular lamella. Umbilicus very narrow, dot like.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
86E9288D36235017999C4A2A38DBE0DA.taxon	description	Measurements (in mm, n = 5). SH = 1.81 – 2.14; SW = 1.37 – 1.5; AH = 0.66 – 0.79; AW = 0.63 – 0.75.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
86E9288D36235017999C4A2A38DBE0DA.taxon	etymology	Etymology. The specific epithet is due to the numerous spines in the aperture.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
86E9288D36235017999C4A2A38DBE0DA.taxon	distribution	Distribution. This species is known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
E5B7E7D8D8AE5096BD83A826BFED1EC7.taxon	distribution	Distribution. This species is known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
13D456061CD557038CD69312C6349E9E.taxon	distribution	Distribution. This species is known from Lampang Province, Thailand.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
4F4D5B05735E58788058202718A703A4.taxon	description	Fig. 108	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
CBE9A7CC1E8358C7BB3297846454363A.taxon	description	Figs 103 D, 106, 107	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
CBE9A7CC1E8358C7BB3297846454363A.taxon	distribution	Distribution. This species is known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
E1CB45B827A65B21BF4D022023DB33F5.taxon	diagnosis	Diagnosis. The genus Boysidia remains unrevised. This is urgently needed because many species are misunderstood due to poor original descriptions / drawings or the uncertain status of type specimens. The genus currently includes the following species: Boysidia chiangmaiensis Panha & J. B. Burch, 2002, B. conspicua (Möllendorff, 1885), B. dilamellaris D. - N. Chen, Y. - H. Liu & W. - X. Wu, 1995, B. dorsata (Ancey, 1881), B. gracilis F. Haas, 1937, B. guiyangensis T. - C. Luo, D. - N. Chen, G. - Q. Zhang, La. Zhang, Li Zhang & T. - W. Li, 1998, B. hangchowensis (Pilsbry & Y. Hirase, 1908), B. houaphanica Inkhavilay & Sutcharit, 2024, B. huangguoshuensis T. - C. Luo, D. - N. Chen & G. - Q. Zhang, 2000, B. hunana (Gredler, 1881), B. jinpingensis M. Tian, B. Fao & Y. - X. Chen, 2015, B. nanjiangensis Z. - L. Zhang, W. - H. Zhang & T. - C. Luo, 2011, B. orientalis B. Rensch, 1935, B. phatangensis Dumrongrojwattana & Assawawattagee, 2018, B. ringens van Benthem Jutting, 1950; B. robusta Bavay & Dautzenberg, 1912, B. shilinensis D. - N. Chen, M. Wu & G. - Q. Zhang, 1999, B. strophostoma (Möllendorff, 1885), B. taibaiensis D. - N. Chen, M. Wu & G. - Q. Zhang, 1999, B. tamtouriana Pokryszko & Auffenberg, 2009, B. tholos Panha & J. B. Burch, 2002, B. tongguanensis D. - N. Chen & W. - H. Zhang, 2002, B. xianfengensis W. - H. Zhang, D. - N. Chen & W. - C. Zhou, 2014, B. xiaoguanensis W. - H. Zhang, D. - N. Chen & W. - C. Zhou, 2014, B. xishanensis D. - N. Chen, M. Wu & G. - Q. Zhang, 1999, B. xiuwenensis W. - H. Zhang, T. - C. Luo & W. - C. Zhou, 2010. The species listed below are excluded from the genus Bensonella and transferred to Boysidia due to the following traits (not shared among all these species): i) apertural dentition more similar to Boysidia (columellar lamella very strong and long, absence of three barriers on the parietal side); ii) surface sculpture similar to Boysidia (radial growth lines, no spiral striae); iii) larger shell sizes (frequently over 4 mm), which is more common in Boysidia representatives.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
F896FB6E8C4E59BA880CF22CD4EB712C.taxon	description	Figs 109	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
78541F9F33E659D68EAC2028999B26AD.taxon	description	Fig. 110	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
78541F9F33E659D68EAC2028999B26AD.taxon	distribution	Distribution. This species is known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
3057154D05485D0A908072205C62B448.taxon	description	Figs 111 A, 112, 113, 154	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
3057154D05485D0A908072205C62B448.taxon	distribution	Distribution. This species in known from several localities in Thailand and one in Peninsular Malaysia. Our record from Malaysia is the first record of this species in the country.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
A7F1DE4ED3615EDBA8AA9BE749373388.taxon	description	Figs 111 B, 114, 115, 158	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
A7F1DE4ED3615EDBA8AA9BE749373388.taxon	diagnosis	Diagnosis. Hypselostoma species with conical-ovoid shell, spirally striated whorls, and the last whorl with a keel below the centre of the periphery. Apertural barriers three but the parietal lamella is the only tooth-like barrier. Palatal plica and columellar lamella are present as swellings on the peristome.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
A7F1DE4ED3615EDBA8AA9BE749373388.taxon	description	Description. Shell conical-ovoid, pale yellowish, consisting of nearly 4.5 whorls separated by a deep suture. All whorls, except the last one, regularly rounded and convex. Last whorl with a blunt keel below the centre of the periphery and weakly concave to almost flat whorl outline both above and below the keel. Protoconch of same colour as the rest of the shell, finely pitted showing spiralling pattern which continues as more distinct spiral striae terminally. It consists of ~ 1.75 whorls. Teleoconch coarsely sculptured with irregularly spaced radial growth lines but also with dense spiral striae. Spacing between the two spiral striae equals the width of two or three striae. Last whorl adnate to the penultimate and very slightly ascending (~ 7 ° compared to the shell axis), making the apertural profile weakly opisthocline to the shell axis. Peristome white, thick, and expanded (especially on the parietal side) but not reflected. Apertural barriers largely reduced and of very unusual appearance for any hypselostomatid. Parietal lamella is the only „ tooth-like “ barrier in the aperture but it is very low and short. Palatal plica and columellar lamella not present in their typical forms but rather as very weak and indistinct swellings on the peristome, visible only in the frontal view. These swellings are probably homologous to the two mentioned barriers. There are no other barriers in the aperture. Surface of all apertural barriers is very finely granulated (almost smooth). Sinulus wide due to the weak parietal lamella and palatal plica. Umbilicus extremely narrow, dot like and slightly elongated. Umbilical groove absent.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
A7F1DE4ED3615EDBA8AA9BE749373388.taxon	description	Measurements (in mm, n = 1): SH = 2.1; SW = 1.7; AH = 0.97; AW = 0.93.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
A7F1DE4ED3615EDBA8AA9BE749373388.taxon	etymology	Etymology. The specific epithet aenigma comes from a Latin word for a riddle (which refers to the unique appearance of the shell and unclear relationship with other congeners) and is to be used as a noun in apposition.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
A7F1DE4ED3615EDBA8AA9BE749373388.taxon	distribution	Distribution. This species is known only from its type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
9BB9CA10F75D525994361730BFCC2306.taxon	description	Figs 111 C, 116, 154	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
9BB9CA10F75D525994361730BFCC2306.taxon	distribution	Distribution. This species is known only from Annam, Vietnam.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
F07A17348C1257F0B396DF7E9AAE416F.taxon	description	Figs 111 D, 117, 118, 177	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
F07A17348C1257F0B396DF7E9AAE416F.taxon	diagnosis	Diagnosis. A Hypselostoma species with a slender, concave-conical shell, bluntly and strongly keeled last whorl (and a deep groove above it), raised spiral striation, hooked apertural barriers (upper palatal, and lower palatal) and a wide umbilicus which is initially narrow.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
F07A17348C1257F0B396DF7E9AAE416F.taxon	description	Description. Shell concave-conical, light-brown, consisting of 4.5 – 5 whorls separated by a deep suture. Protoconch nearly white, consisting of ~ 1.25 coarsely spirally striated whorls. There are ~ 12 weak spiral striae on the protoconch. Protoconch and initial teleoconch whorls rounded, penultimate weakly convex. Last whorl with strong but blunt keel positioned at the centre of the periphery. Above the keel there is a deep groove on the last 0.75 – 1 whorl. All whorls densely sculptured with strong, thick spiral striae crossed by less dense radial growth lines and occasionally by a few thin, whitish radial streaks. Spacing between the spiral striae irregular and ranges from the width of two to the width of three spiral striae. Last whorl moderately detached from the penultimate and weakly ascending (≤ 10 ° compared to the shell axis). Peristome expanded, not reflected, its surface finely pitted. Aperture equipped with four strong, main barriers (angulo-parietal, upper palatal, lower palatal and columellar) and several smaller ones. Angulo-parietal lamella strong and leaned towards the palatal side, almost reaching the expanding peristome. It is consisting of main, parietal part, and a smaller, pointed angular part which are separated by a weak sinuation. Upper and lower palatal plicae roughly the same size, both divided and the inner parts are hooked (with the tip of hooks pointing outside), slender and high, while the outer parts are low and slender. A weak swelling is also observed at the palatal side (in front of the upper palatal plicae) which is probably homologous to the palatal tubercle in the majority of Bensonella species. Columellar lamella almost horizontal, approximately as high as the palatal plicae. There are four smaller barriers, two interpalatal plicae, one basal plica and one infraparietal lamella. Surface of all apertural barriers is smooth or with very weak granules. Sinulus rounded, distinctly separated from the rest of the aperture. Umbilicus initially narrow and then abruptly widening, measuring between 1 / 3 and 1 / 4 of the shell width. There is a periumbilical keel situated on the last ~ 0.75 whorl near the centre of the last whorl, but rather slightly towards the umbilicus. Along this keel there are shallow grooves on both sides.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
F07A17348C1257F0B396DF7E9AAE416F.taxon	description	Measurements (in mm, n = 5). SH = 1.91 – 2.04; SW 1 = 2.16 – 2.39; SW 2 = 1.35 – 1.42; AH = 0.80 – 0.86; AW = 0.69 – 0.82.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
F07A17348C1257F0B396DF7E9AAE416F.taxon	etymology	Etymology. This species is named for the apertural barriers resembling claws of an eagle (aquila = eagle in Latin). To be used as a noun in apposition.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
F07A17348C1257F0B396DF7E9AAE416F.taxon	distribution	Distribution. This species is known from two localities in Steung Treng province.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
4CB784AA75FB58F1ADD3E08328FCC5AE.taxon	description	Figs 186 A, 187, 223	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
4CB784AA75FB58F1ADD3E08328FCC5AE.taxon	distribution	Distribution. This species is known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
C8B98D2512815848A1F273EC40401E94.taxon	description	Figs 111 E, 119	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
C8B98D2512815848A1F273EC40401E94.taxon	distribution	Distribution. This species in known from several localities in N Australia as well as under several synonyms (Iredale 1937; Solem 1981).	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
85DACDD1EDF45505908AF4380E1483E5.taxon	distribution	Distribution. This subspecies is known from several localities in Napier Range, western Australia.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
3A1138DB104A5264AEB0BC20F3A8A63D.taxon	description	Figs 111 F, 120, 182	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
3A1138DB104A5264AEB0BC20F3A8A63D.taxon	distribution	Distribution. This species is known from Kampot province, Cambodia.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
3B18896022AF5A64BE96BC58CF98526A.taxon	description	Figs 111 G, 121, 158	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
3B18896022AF5A64BE96BC58CF98526A.taxon	distribution	Distribution. This species is known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
22793DE134D1595F82D6D1F19C6CE6D9.taxon	description	Figs 186 B, 188, 189, 223	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
22793DE134D1595F82D6D1F19C6CE6D9.taxon	diagnosis	Diagnosis. Shell depressed, concave-conical. Teleoconch finely radially and spirally striated (localised, just below and above the keel on the last whorl). Last whorl detached from the penultimate and strongly ascending. Aperture wide, equipped with five main barriers (angular, parietal, upper palatal, lower palatal and columellar) and two smaller barriers (basal and infraparietal). Umbilicus wide, measuring 50 % of the shell width.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
22793DE134D1595F82D6D1F19C6CE6D9.taxon	description	Description. Shell depressed, concave-conical, chestnut brown, very weakly glossy and not translucent. Whorls 3.5 – 4, initially weakly convex, separated by a deep suture, regularly increasing until the last whorl which is strongly enlarged and flat. Protoconch lighter than the rest of the shell, finely pitted without spiralling pattern, consisting of ~ 1.25 whorls. Teleoconch very finely granulated (sandpaper-like surface) and radially striated with additional whitish radial streaks which are weak, widely, and irregularly spaced. Spiral striae present but localised, clearly visible only on the last whorl above and below the keel, elsewhere absent. Sometimes, spiral striae are very difficult to observe on the whole shell. Last whorl strongly enlarged in comparison to previous ones, bluntly keeled very slightly above the centre of the periphery (or sometimes at the centre of the periphery) and almost flat above and weakly convex below the keel. Keel is strongest near the aperture. Last whorl moderately to strongly ascending near the aperture (~ 50 – 65 ° compared to the shell axis) and moderately detached from the penultimate whorl. Peristome very pale brownish, strongly expanded but not reflected. Aperture equipped with five main barriers (angular, parietal, upper palatal, lower palatal, and columellar). Angular lamella short and small, directed towards the upper palatal plica. Parietal lamella strong and high, not much longer than the angular and only reaching its profile. Upper palatal plica slightly weaker than the parietal lamella and bent towards the lower palatal plica. Lower palatal plica straight, equally strong as the upper palatal but not bent. Columellar lamella is the broadest in the aperture but relatively short. Between these main barriers, additional small basal and infraparietal plicae visible. Surface of all apertural barriers is very finely spiniferous medially and very finely granulated laterally. Sinulus small and distinctly separated from the rest of the aperture. Umbilicus very wide, measuring half of the shell width and showing all previous whorls. A deep and wide groove is running from the columellar side of the peristome alongside the umbilicus.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
22793DE134D1595F82D6D1F19C6CE6D9.taxon	description	Measurements (in mm, n = 5). SH = 2 – 2.23; SW 1 = 3.40 – 4.13; SW 2 = 1.95 – 2.41; AH = 1.44 – 1.78; AW = 1.33 – 1.61.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
22793DE134D1595F82D6D1F19C6CE6D9.taxon	etymology	Etymology. The type locality of this species, Gua Kerbau, means “ Buffalo cave ” in Malay. The specific epithet bubalus is also a scientific name for a buffalo, which is to be used as a noun in apposition.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
22793DE134D1595F82D6D1F19C6CE6D9.taxon	distribution	Distribution. This species is known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
30801580F371587FB602B63E31FA6065.taxon	description	Figs 242 A, 243, 259	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
30801580F371587FB602B63E31FA6065.taxon	distribution	Distribution. This species is known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
48CA0F81C7CF5488867777A1045A9067.taxon	description	Figs 111 H, 122, 182	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
48CA0F81C7CF5488867777A1045A9067.taxon	distribution	Distribution. This species in known from Kampot province (Cambodia) and Kien Giang province (Vietnam) (Vermeulen et al. 2019).	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
D8BFAE571B8E536B961463946671F915.taxon	description	Figs 186 C, 190, 223	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
D8BFAE571B8E536B961463946671F915.taxon	distribution	Distribution. Apart from the type locality, this species is also known from one more nearby locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
481B339CBD1D5132BEE354F346A4FDCB.taxon	description	Figs 111 I, 123, 182	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
481B339CBD1D5132BEE354F346A4FDCB.taxon	distribution	Distribution. This species is known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
F0E65A96447B5B67A47F6595A1846C14.taxon	description	Figs 111 J, 124, 154	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
F0E65A96447B5B67A47F6595A1846C14.taxon	distribution	Distribution. This species is known from Nakhon Sawan and Kampangpet provinces.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
44A493DFBA625C4FB85B476735F7175E.taxon	description	Figs 111 K, 125, 126, 154	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
44A493DFBA625C4FB85B476735F7175E.taxon	diagnosis	Diagnosis. Hypselostoma species with very deep sutures and spirally striated whorls. Penultimate and the last whorl are keeled. Last whorl not detached from the penultimate. Aperture equipped with six barriers (angulo-parietal, upper palatal, lower palatal, basal, columellar and a small infraparietal). Umbilicus initially narrow, abruptly widening at the last whorl.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
44A493DFBA625C4FB85B476735F7175E.taxon	description	Description. Shell conical-ovoid, consisting of 4 – 4.5 whorls separated by a deep suture. Protoconch pitted, showing some spiralling pattern, the same colour as the rest of the shell, consisting of ~ 1.5 whorls. Teleoconch surface sculptured with coarsely spaced radial lines crossed by much stronger and dense spiral striae. Spacing between two spiral striae approximates the width of one or two striae. First teleoconch whorl rounded. All other teleoconch whorls strongly keeled, last one the most prominent. Surface above the keel on the penultimate whorl is almost flat, (like a platform) but is much more steep (sloping) on the last whorl. Last whorl adnate to the penultimate and slightly ascending near the aperture (~ 5 – 10 ° compared to the shell axis). Peristome not very strong but expanded, not reflected. It is forming a weak callus on the parietal side. Aperture equipped with five main barriers (angulo-parietal, upper palatal, lower palatal, basal and columellar) and one small infraparietal lamella which is knob-like (although it can be absent as well). Angulo-parietal lamella is the strongest in the aperture, consisting of larger parietal part and indistinct angular part so that it appears slightly bilobed. Upper palatal plica short, weaker than the lower palatal and slightly bent towards it. Lower palatal plica roughly as strong as the columellar, situated halfway between upper palatal and basal plica. Basal plica small, slightly weaker than the upper palatal. Columellar lamella strong, directed towards the upper palatal plica and almost reaching the expanding peristome. Peristome is weakly swollen in front of the columellar lamella. Surface of all apertural barriers is finely granulated to almost smooth. Sinulus wide and not distinctly separated from the rest of the aperture. Umbilicus initially very narrow and then abruptly widening at the last whorl, measuring ~ 1 / 5 – 1 / 6 of the shell width. A weak groove is visible running alongside the umbilicus but it is getting lost near the middle part of the last whorl.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
44A493DFBA625C4FB85B476735F7175E.taxon	description	Measurements (in mm, n = 4). SH = 1.93 – 2.46; SW = 2.09 – 2.24; AH = 1.02 – 1.16; AW = 1.05 – 1.18.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
44A493DFBA625C4FB85B476735F7175E.taxon	etymology	Etymology. The two largest whorls (last and the penultimate) are keeled which gives the initial impression that the whole shell is keeled, hence the specific epithet.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
44A493DFBA625C4FB85B476735F7175E.taxon	distribution	Distribution. This species is known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
6625028B828659898D58D5F30D7FF4F1.taxon	description	Figs 186 D, 191, 192, 223	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
6625028B828659898D58D5F30D7FF4F1.taxon	diagnosis	Diagnosis. Shell depressed-conical, yellowish, and glossy. Last whorl bluntly keeled. Aperture detached from the penultimate whorl and slightly ascending upwards. Teleoconch finely granulated and radially striated. Spiral striation absent. Aperture equipped with five main strong barriers (parietal, upper and lower palatal, basal, and columellar) and additional smaller one of variable number.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
6625028B828659898D58D5F30D7FF4F1.taxon	description	Description. The yellowish, glossy, and weakly translucent, depressed-conical shell is consisting of 4 – 4.5 whorls separated by a deep suture. The finely pitted protoconch is showing a weak spiralling pattern, consisting of ~ 1.5 whorls without a clear boundary with teleoconch. Teleoconch finely granulated, sandpaper-like (surface microsculpture similar to H. hungerfordianum) and inconspicuously regularly radially striated. Last whorl bluntly keeled, slightly detached from the penultimate and slightly to moderately ascending upwards (~ 25 – 35 ° compared to the shell axis). Peristome expanded and not reflected, with sharp edges. Aperture equipped with five main barriers (parietal, angular, upper palatal, lower palatal, and columellar) and variable number of smaller ones. Angular lamella is the weakest among the main five, the rest of the main barriers are developed roughly to the same level. They are all high and straight. Parietal and angular lamellae are directed towards the palatal wall. In front of the upper palatal plica, a small swelling is present which is probably homologous with the palatal tubercle present in other genera (e. g., Bensonella, see Páll-Gergely and White 2023). Between the angular lamella and upper palatal plica, one or two additional plicae are present. There are also usually two interpalatal plicae, two between lower palatal and columellar (including basal plica) and two between the columellar and parietal lamellae. None of these smaller barriers reach the profiles of the main ones, and none of the apertural barriers reach the peristome. Surface of all apertural barriers is very finely spiniferous medially and very finely granulated laterally. Sinulus parabolic and not very strongly separated from the rest of the aperture. Umbilicus open, initially narrow, and then abruptly widening at the last whorl, measuring ~ ¼ of the shell width and showing the penultimate whorl. A strong groove can be observed running from the peristome towards the inner side of the umbilicus.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
6625028B828659898D58D5F30D7FF4F1.taxon	description	Measurements. SH = 2.05 – 2.32; SW 1 = 3.12 – 3.37; SW 2 = 1.85 – 2.09; AH = 1.33 – 1.58; AW = 1.28 – 1.40.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
6625028B828659898D58D5F30D7FF4F1.taxon	etymology	Etymology. The specific epithet is due to its “ leathery ” surface sculpture.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
6625028B828659898D58D5F30D7FF4F1.taxon	distribution	Distribution. Known only from Tanjung Rhu, Langkawi Island, Malaysia.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
9CCDDBD20882511E8F199CB8F5C96193.taxon	description	Figs 242 B, 244, 245, 246, 247, 248, 249, 250, 251	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
9CCDDBD20882511E8F199CB8F5C96193.taxon	distribution	Distribution. This species is known from a wide area covering China, N Vietnam, Myanmar, and Laos (from where it is known as G. plesiolopa).	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
A15C56E7CB1B581C8B04FC2C5095E548.taxon	description	Figs 111 L, 128, 154	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
A15C56E7CB1B581C8B04FC2C5095E548.taxon	distribution	Distribution. This species is known from the type locality and its immediate surroundings.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
4F79B0374FEE57BFA78B9D8FC032481A.taxon	description	Figs 111 M, 127, 154	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
4F79B0374FEE57BFA78B9D8FC032481A.taxon	distribution	Distribution. This species is, apart from the type locality in Loei province, known from two more localities: one in Lampang and one in Chonburi province.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
1006C0BD8D015D499CF0FC8C8121A780.taxon	description	Figs 242 C, 252, 259	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
1006C0BD8D015D499CF0FC8C8121A780.taxon	distribution	Distribution. This species is known from Peninsular Malaysia.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
5708339FAFB45FC3B90AB009FD182773.taxon	description	Figs 111 N, 129, 182	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
5708339FAFB45FC3B90AB009FD182773.taxon	distribution	Distribution. This species is known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
9D3638C42F885881A13CF3B751B4DEE6.taxon	description	Figs 111 O, 130, 131, 154	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
9D3638C42F885881A13CF3B751B4DEE6.taxon	distribution	Distribution. Apart from the type locality, this species is known from one other sampling site near the type locality as well as from the Rayong, Sa Kaeo, Prachinburi, and Songkhla provinces (Dumrongrojwattana and Wongkamhaeng 2024).	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
B839F2EC620B5114979CA2D3A83BB218.taxon	description	Figs 111 P, 132, 182	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
B839F2EC620B5114979CA2D3A83BB218.taxon	distribution	Distribution. This species is known from the Vietnam part of the Mekong Delta limestone.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
DE7A437EA5D75E6890195C1AB86ED722.taxon	description	Figs 111 Q, 133, 182	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
DE7A437EA5D75E6890195C1AB86ED722.taxon	distribution	Distribution. This species in known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
1F3670D619EB5212998F3831EF24233F.taxon	description	Figs 111 R, 134, 154	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
1F3670D619EB5212998F3831EF24233F.taxon	distribution	Distribution. This species in known from Lampang and Chiang Rai provinces in Thailand.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
90B0F1AA1BE2571ABB7A656C2E813548.taxon	description	Figs 224 A, 225, 241	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
90B0F1AA1BE2571ABB7A656C2E813548.taxon	distribution	Distribution. This species in known from Bukit Tenggek and Gunung Sagu (IUCN, 2015).	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
956C9AEC0FC25B8AABB2932BFBB74568.taxon	description	Figs 224 B, 226, 241	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
956C9AEC0FC25B8AABB2932BFBB74568.taxon	distribution	Distribution. This species is known from Bukit Chuping (type locality) and Bukit Keteri (this study), both located in Perlis, N Peninsular Malaysia.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
988ED4596B5B5B2588653E486C6D3832.taxon	description	Figs 111 S, 135, 154	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
988ED4596B5B5B2588653E486C6D3832.taxon	distribution	Distribution. This species is known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
7A5D2E449F7D5849B4C7E64BDBD5A197.taxon	description	Figs 111 T, 136, 137, 140	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
7A5D2E449F7D5849B4C7E64BDBD5A197.taxon	distribution	Distribution. This species is known from Timor island, Kalao island, Sulawesi, Java, Bali, Lesser Sunda islands and Tanimbar islands (latter is the type locality of H. dohertyi).	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
F491D1DE91CC5D1A9CD30E7865390DA9.taxon	description	Figs 186 E, 193, 194, 195, 223	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
F491D1DE91CC5D1A9CD30E7865390DA9.taxon	diagnosis	Diagnosis. Hypselostoma with depressed-conical shell, teleoconch with finely sandpaper-like surface and radially but not spirally striated. Aperture equipped with only four relatively weak barriers (angulo-parietal, upper palatal, lower palatal and columellar).	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
F491D1DE91CC5D1A9CD30E7865390DA9.taxon	description	Description. Shell depressed-conical, chestnut brown, glossy, not translucent. It is consisting of 3.5 – 4 regularly increasing, convex whorls separated by a deep suture. Protoconch of the same colour as the teleoconch, finely pitted, not showing spiralling pattern and consisting of ~ 1.25 whorls. Teleoconch sculpture finely granulated (sandpaper-like) with additional radial growth lines which are more or less regularly spaced. Last whorl bluntly keeled at the centre of the periphery, slightly to moderately detached from the penultimate whorl and strongly ascending near the aperture (usually ranging from 65 to almost 90 ° compared to the shell axis). The keel is the most prominent on the last whorl behind the peristome. Peristome brown, weakly expanded, not reflected. Aperture equipped with four main barriers (angulo-parietal, upper palatal, lower palatal and columellar), all of which are relatively weak. Angulo-parietal lamella is the strongest in the aperture, consisting of a high parietal part and weak, tubercle-like angular part which is sometimes weakly sinuated in the middle. The angular part is sometimes also more distinctly separated from the parietal. Upper palatal plica relatively weak and straight. Lower palatal plica only slightly stronger (broader and higher) than the upper palatal. Columellar lamella horizontal, developed to the same extent as the lower palatal plica. Weak basal plica and infraparietal lamella sometimes present. All barriers weaker than in the majority of the congeners, all finely spiniferous medially but indistinctly granulated to almost smoot laterally. Sinulus wide and not strongly separated from the rest of the aperture. Umbilicus moderately wide to wide, measuring ~ 1 / 3 – 1 / 5 of the shell width and clearly showing the penultimate whorl. A relatively shallow groove runs towards the inner side of the umbilicus, terminating at the penultimate whorl.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
F491D1DE91CC5D1A9CD30E7865390DA9.taxon	description	Measurements (in mm, n = 5). SH = 1.46 – 1.58; SW 1 = 2.67 – 2.90; SW 2 = 1.74 – 1.92; AH = 1.06 – 1.26; AW = 1.05 – 1.23.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
F491D1DE91CC5D1A9CD30E7865390DA9.taxon	etymology	Etymology. The shape of the aperture, between the barriers, resembles a four-leaf clover which is a common symbol of luck, hence the specific epithet which is derived from the Latin word for lucky (fortunatus).	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
F491D1DE91CC5D1A9CD30E7865390DA9.taxon	distribution	Distribution. This species is known from Phattalung and Songkhla provinces (S Thailand).	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
7ABF7A5A9DF350338C9846886C710393.taxon	description	Figs 224 C, 227, 241	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
7ABF7A5A9DF350338C9846886C710393.taxon	distribution	Distribution. This species is known from Pahang, Kedah, Perak, and Selangor provinces of Peninsular Malaysia. Possibly more widely spread.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
CB7BF46F98FE5A1089D4556B076B7402.taxon	description	Figs 111 U, 138, 139, 140	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
CB7BF46F98FE5A1089D4556B076B7402.taxon	distribution	Distribution. This species in known from Lesser Sunda islands, Java and Sulawesi islands, Indonesia.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
F3E4D81E73B55F01BAD8C70631BE810A.taxon	description	Figs 111 V, 141, 142, 177	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
F3E4D81E73B55F01BAD8C70631BE810A.taxon	diagnosis	Diagnosis. Hypselostoma species with a large last whorl (previous whorls sunken into it) and spirally striated teleoconch. Last whorl strongly detached from the penultimate and descending at almost 80 ° compared to the shell axis. Aperture equipped with four main barriers (angulo-parietal, upper palatal, lower palatal and columellar) and some smaller ones. Basal furrow present. Umbilicus extremely wide, almost as wide as the whole shell.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
F3E4D81E73B55F01BAD8C70631BE810A.taxon	description	Description. Shell depressed-conical, brown, and not glossy, consisting of 3.5 – 4 whorls separated by a deep suture. Protoconch rounded, roughly pitted and consisting of one whorl. Initial teleoconch whorls bluntly keeled and sunken into the last whorl (but sometimes they appear more elevated). Last whorl contributing to almost 100 % of the shell height. It is extending obliquely around the previous whorls and is strongly shouldered both on its upper and lower sides, which gives the shell a unique appearance. Surface of the teleoconch is irregularly, very finely and densely spirally striated. Whitish radial streaks are numerous and present on all teleoconch whorls but unevenly spaced (rather randomly positioned). Last whorl is strongly detached from the penultimate and strongly descending (~ 80 ° compared to the shell axis). This makes the aperture profile almost perpendicular to the shell axis. The so called “ trumpet ” is quite broad (similar to e. g., H. chaunosalpinx). Peristome is dirty white (or light brown), thick and strongly expanded, not reflected. Aperture is equipped with four main barriers (angulo-parietal, upper palatal, lower palatal and columellar) and additional smaller ones. The parietal part of the angulo-parietal lamella is strong and very high (blade-like), but the angular is very short and pointed towards the palatal side. Upper palatal plica is directed towards the lower palatal plica but is weaker. Lower palatal plica is long and low in its inner part and much higher in its outer part (closer to the peristome). This plica is similar to the columellar lamella but more slender. Basal (?) plica is situated closer to the lower palatal. It is not clear whether this plica actually represents the infrapalatal or the basal. In the region where usually the basal plica is present, there is a deep basal furrow which is also rarely encountered in this group (similar structure is also present in H. torta sp. nov.). This furrow is stretching all the way to the expanding peristome. Columellar lamella almost horizontal, thick, and long. A distinct infraparietal lamella is present (sometimes absent) between the columellar and parietal lamellae. Of all these barriers, only the angulo-parietal and columellar lamellae are reaching the expanding peristome. Surface of all apertural barriers is finely granulated. Sinulus narrow, well separated from the rest of the aperture. Umbilicus extremely wide (almost as wide as the whole shell). The lower edge of the last whorl (towards the umbilicus) is very strongly ridged (almost forming a cliff-like structure), leaving the walls of the umbilicus very steep (like a playground slide). The walls of the umbilicus are especially densely sculptured with white radial streaks.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
F3E4D81E73B55F01BAD8C70631BE810A.taxon	description	Measurements (in mm, n = 5). SH 1 = 2.83 – 2.97; SH 2 = 1.62 – 1.85; SW = 3.09 – 3.85; AH = 1.82 – 2; AW = 1.77 – 2.05.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
F3E4D81E73B55F01BAD8C70631BE810A.taxon	etymology	Etymology. This species’ shape resembles a mushroom, which gave its specific epithet (Lat. fungus). The detached last whorl resembles a stalk while the rest of the shell resembles a cap. To be used as a noun in apposition.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
F3E4D81E73B55F01BAD8C70631BE810A.taxon	distribution	Distribution. This species is known only from its type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
FD90B3CA36CF5905B48509C7BCA2100D.taxon	description	Figs 111 W, 143, 144, 154	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
FD90B3CA36CF5905B48509C7BCA2100D.taxon	diagnosis	Diagnosis. Shell concave-conical, last whorl shouldered, previous whorls weakly convex. Teleoconch with raised spiral striae and strong whitish streaks. Last whorl detached from the penultimate, slightly turned downwards. Aperture equipped with five barriers (angulo-parietal, upper palatal, lower palatal, basal, and columellar). Umbilicus initially narrow but suddenly expanding at the last whorl.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
FD90B3CA36CF5905B48509C7BCA2100D.taxon	description	Description. Shell concave-conical (due to the strongly expanded last whorl), light brown, weakly glossy, opaque. It is consisting of 4.5 – 5 step-like, regularly increasing whorls separated by a very deep suture. All whorls except for the last one are rounded, convex. Protoconch slightly lighter than the rest of the shell, spirally striated (~ 25 densely arranged spiral striae) and consisting of ~ 1.25 whorls. Boundary between protoconch and teleoconch not particularly clear but visible as stronger spiral striation and darker surface. Teleoconch densely and strongly spirally striated, especially on the last whorl. Spiral striae slightly almost regularly spaced, leaving the space between the two striae up to the width of three or rarely two striae. Spiral striae get more densely arranged at the lower part of the last whorl than those at the centre of the periphery. There are ~ 33 spiral striae on the last whorl in standard view. Strong and wide radial white streaks cross the spiral striae. These streaks vary in number and are mostly present on the last three whorls. Last whorl shouldered, slightly detached from the penultimate and slightly descending near the aperture (~ 5 – 10 ° compared to the shell axis). Peristome of the same colour as the rest of the shell, or very slightly lighter. It is expanded and not reflected. Aperture equipped with four main barriers (angulo-parietal, upper palatal, lower palatal and columellar). Angular part of the angulo-parietal lamella reaching the peristome, very small and pointed, leaned towards the upper palatal plica. Parietal part of the angulo-parietal lamella is much stronger and high, blade shaped. Upper palatal plica moderately developed, slightly curved towards the lower palatal and similarly strong. Lower palatal plica high and narrow, making it particularly slender. Columellar lamella strong as the palatal plicae, almost horizontal and very slightly leaned towards the lower palatal plica. Between these main barriers, only a small and weak basal one is additionally observed. Surface of all apertural barriers is smooth. Sinulus wide, not strongly separated from the rest of the aperture. Umbilicus initially very narrow, suddenly expanded at the last whorl, measuring between 1 / 3 and ¼ of the shell width. A deep groove is running from the columellar side of the peristome towards the inner walls of the umbilicus.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
FD90B3CA36CF5905B48509C7BCA2100D.taxon	description	Measurements (in mm, n = 5). SW 1 = 2.48 – 2.74; SW 2 = 1.68 – 1.75; SH = 2.62 – 2.92; AH = 1.09 – 1.21; AW = 1.12 – 1.25.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
FD90B3CA36CF5905B48509C7BCA2100D.taxon	etymology	Etymology. The specific epithet is due to the fact that there were many geckos at its habitat.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
FD90B3CA36CF5905B48509C7BCA2100D.taxon	distribution	Distribution. This species is known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
BBEC67F6B35455BFB0DBB1AD4A6CD73C.taxon	description	Figs 111 X, 145, 154	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
BBEC67F6B35455BFB0DBB1AD4A6CD73C.taxon	distribution	Distribution. This species is known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
FEDD6A30FC4F5729BA22BE5005C13FA0.taxon	description	Figs 186 F, 196, 197, 198, 199, 200, 201	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
FEDD6A30FC4F5729BA22BE5005C13FA0.taxon	distribution	Distribution. This species is widely spread in Malaysia and adjacent regions of Thailand. Also known from Myanmar where the northernmost locality of the species is known (Tongkerd et al. 2024). The southernmost localities are positioned in and near the city of Kuala Lumpur (Malaysia).	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
54099BE0415F52958A15FEBE8D8920BB.taxon	description	Figs 111 Y, 146, 147	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
54099BE0415F52958A15FEBE8D8920BB.taxon	distribution	Distribution. This species is known from the type locality and Kenting Tropical Botanical Garden, Kenting, Hengchun, Pingdung, Taiwan (21 ° 58 ' 04.5 " N, 120 ° 48 ' 55.3 " E). The latter is the type locality of H. kentingensis.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
AF42A8DDD59656FDA54909F2206CF774.taxon	description	Figs 111 Z, 148, 149, 154	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
AF42A8DDD59656FDA54909F2206CF774.taxon	diagnosis	Diagnosis. Hypselostoma with very dense spiral striation on the teleoconch. Apertural barriers very weak and four (parietal, upper palatal, lower palatal and columellar). Umbilicus narrow, widening at the last whorl.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
AF42A8DDD59656FDA54909F2206CF774.taxon	description	Description. Shell conical, light yellow to brown, consisting of 5 – 5.5 convex, rounded whorls separated by a deep suture. Protoconch slightly lighter than the rest of the shell, consisting of ~ 1.5 whorls, densely spirally striated (striae clearer terminally). Teleoconch very finely, densely spirally striated crossed by stronger and much coarser, irregularly spaced radial growth lines. Last whorl rounded to slightly shouldered, adnate to the penultimate and slightly to moderately descending (~ 25 – 35 ° compared to the shell axis) making the aperture profile prosocline to the shell axis. Peristome white but not much expanded, not reflected. It is only particularly expanded on the columellar side where it forms a strong margin which is, however, not covering the umbilicus. The peristome is discontinuous on the parietal side where it is completely merged with the penultimate whorl. Aperture almost as wide as high. Apertural barriers few and much weaker than in the majority of congeners. There are four barriers: parietal, upper palatal, lower palatal and columellar. Parietal lamella is the strongest in the aperture but still somehow weak, moderately high and curved towards the palatal side. Two palatal plicae (upper and lower palatal) are very weak, like small thickenings. Columellar lamella also weak and shares the same appearance as the palatal plicae but situated slightly closer to the peristome edge. None of the apertural barriers are reaching the thin peristome. Surface of all apertural barriers is almost smooth but very finely granulated. Sinulus wide and not strongly isolated from the rest of the aperture. Umbilicus moderately wide, measuring 1 / 5 of the shell width, initially narrow and suddenly widening at the last whorl (which gives it a slightly elongated appearance).	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
AF42A8DDD59656FDA54909F2206CF774.taxon	description	Measurements (in mm, n = 6). SH = 2.27 – 2.48; SW = 1.67 – 1.72; AH = 0.81 – 0.92; AW = 0.84 – 0.96.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
AF42A8DDD59656FDA54909F2206CF774.taxon	etymology	Etymology. This species is named iunior for the very weak teeth and weakly expanded peristome which is common in subadult hypselostomatid specimens.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
AF42A8DDD59656FDA54909F2206CF774.taxon	distribution	Distribution. This species is known only from its type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
67FBD2C26B6E5F399C99D0BC2039C6A9.taxon	description	Figs 224 D, 228, 229, 230, 241	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
67FBD2C26B6E5F399C99D0BC2039C6A9.taxon	distribution	Distribution. This species is known from Kelantan, Pahang, and Perak provinces in Peninsular Malaysia.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
61815C3DF1905356966A2C30D2E9D12C.taxon	distribution	Distribution. This species is, apart from the type locality, known also from Sichang Island (type locality of H. srirachaensis).	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
9D27F6D2C6B2533BBAFB726332D252B5.taxon	distribution	Distribution. This species is known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
5FE53108D50858959AEB2A12B43BEA2A.taxon	distribution	Distribution. Clearly a widespread species: its distribution stretches from Chiang Rai province in N Thailand to the southernmost known localities in Ratchaburi and Chonburi provinces. Westwards, this species stretches to Taunggyi in Myanmar and the easternmost locality is known from Luang Prabang province in Laos.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
A0D2A6E194C752C9A7AC44D38F0BCDD3.taxon	description	Fig. 185	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
A0D2A6E194C752C9A7AC44D38F0BCDD3.taxon	distribution	Distribution. This species is known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
68759A089ECD5694B83ABEF1F1366DBD.taxon	description	Figs 186 G, 202, 223	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
68759A089ECD5694B83ABEF1F1366DBD.taxon	distribution	Distribution. This species is known from the type locality in Cambodia and is now reported for the first time in Thailand (Sa Kaeo province).	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
651CF96A72E658D6A9D44462F99BCC7C.taxon	distribution	Distribution. This species is known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
75F14CD8BADC5C099EDD1807B489DA0F.taxon	description	Figs 186 H, 203, 223	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
75F14CD8BADC5C099EDD1807B489DA0F.taxon	distribution	Distribution. Known from Loei, Phitsanulok, Khon Kaen and Nong Bua Lamphu provinces in Thailand.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
63EBE0B33D4D577BA5158FC59B135BDD.taxon	description	Figs 186 I, 204, 205, 223	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
63EBE0B33D4D577BA5158FC59B135BDD.taxon	distribution	Distribution. This species is known from Peninsular Malaysia and Yala Province in Thailand.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
E46B9263724853E2947D2A28B51E5DDC.taxon	description	Figs 224 E, 231, 241	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
E46B9263724853E2947D2A28B51E5DDC.taxon	distribution	Distribution. This species is known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
0E054234EDE15B4F921E1FBBCFC7B938.taxon	description	Figs 242 D, 253, 259	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
0E054234EDE15B4F921E1FBBCFC7B938.taxon	distribution	Distribution. This species is known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
94691362CCF75175A3DC2CE31A574849.taxon	description	Figs 186 J, 206, 207, 223	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
94691362CCF75175A3DC2CE31A574849.taxon	diagnosis	Diagnosis. A Hypselostoma species with sandpaper-like shell surface and a bluntly keeled last whorl which is detached from the penultimate and slightly ascending upwards. Five main barriers in the aperture (parietal, angular, upper palatal, lower palatal and columellar). Parietal lamella very long and wavy. Umbilicus moderately wide.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
94691362CCF75175A3DC2CE31A574849.taxon	description	Description. Shell shape concave-conical (due to the strongly enlarged last whorl), consisting of 4.5 – 5 convex whorls separated by a deep suture. Colouration dark brown, weakly glossy and opaque. Protoconch finely pitted, showing very weak spiralling pattern, consisting of ~ 1.25 – 1.5 whorls, coloured the same as the teleoconch. Teleoconch very finely granulated (sandpaper-like) with additional radial growth lines. Also, some irregularly spaced radial white streaks are occasionally visible. Last whorl bluntly keeled, moderately detached from the penultimate and slightly ascending upwards (~ 20 ° compared to the shell axis). In lateral view, two strong grooves are visible near the aperture, one above and one below the central keel. Peristome expanded and not reflected, slightly lighter than the rest of the shell. Aperture equipped with five strong and thick barriers (parietal, angular, upper palatal, lower palatal, and columellar) and several smaller ones. Parietal lamella very strong, wavy, and very long. Angular lamella short and sometimes separated in inner and outer part. Inner part is weaker. Both palatal plicae very strong, nearly as strong as the parietal lamella or even stronger but clearly shorter. In front of the upper palatal plica, a small swelling is present which is probably homologous with the palatal tubercle present in other genera (e. g., Bensonella, see Páll-Gergely and White 2023) Columellar lamella nearly horizontal, stronger only than the angular among the main barriers. Additionally, smaller barriers are present between the main ones. There is usually one in the interpalatal region, one basal and one between the columellar and parietal. None of these barriers reach the peristome. Surface of all barriers is very finely spiniferous medially and very finely granulated laterally. Sinulus rounded and visibly separated from the rest of the aperture. Umbilicus open and moderately wide, measuring 1 / 3 of the shell width and showing the penultimate whorl. A strong groove running from the peristome towards the umbilicus is visible. This groove is getting lost near the penultimate whorl.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
94691362CCF75175A3DC2CE31A574849.taxon	description	Measurements (in mm, n = 4). SH = 1.76 – 2.1; SW 1 = 2.38 – 2.6; SW 2 = 1.48 – 1.54; AH = 0.88 – 0.95; AW = 0.85 – 0.89.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
94691362CCF75175A3DC2CE31A574849.taxon	etymology	Etymology. The specific epithet is derived from the Greek work ophis (snake) which is in special reference to the curved and long parietal lamella (which is also one of the longest in all hitherto described Hypselostoma species). To be used as a noun in apposition.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
94691362CCF75175A3DC2CE31A574849.taxon	distribution	Distribution. This species is known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
5676833304895855ADBD69D06A8CB2DC.taxon	description	Figs 224 F, 232, 239	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
5676833304895855ADBD69D06A8CB2DC.taxon	distribution	Distribution. This species is known only from Sarawak, Borneo.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
4D3E43294DE85AF594AC5CA1C0254D1E.taxon	description	Fig. 185	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
4D3E43294DE85AF594AC5CA1C0254D1E.taxon	distribution	Distribution. This species is known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
4338866150C656CF8C5239786E1CA04A.taxon	description	Figs 242 E, 254, 259	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
4338866150C656CF8C5239786E1CA04A.taxon	distribution	Distribution. This species is known from Phattalung and Songkhla provinces, Thailand.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
70BC59E38C9A575A9A6FEC2897365365.taxon	description	Figs 242 F, 255, 259	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
70BC59E38C9A575A9A6FEC2897365365.taxon	distribution	Distribution. This species is known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
A03D76F2657950ABBD54A01D5609C6CF.taxon	description	Figs 242 G, 256, 259	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
A03D76F2657950ABBD54A01D5609C6CF.taxon	distribution	Distribution. This species is known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
6BC13778D39F59229375461E690CAAB2.taxon	description	Figs 186 K, 208, 223	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
6BC13778D39F59229375461E690CAAB2.taxon	distribution	Distribution. This species is known only from the type locality and its surroundings.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
DF28882580A65621A850B69ACDD03977.taxon	diagnosis	Diagnosis. Hypselostoma species with bluntly keeled penultimate (at the centre of the periphery) and shouldered last whorl. Teleoconch densely spirally striated. Last whorl very slightly detached from the penultimate and with a sharp shoulder. Apertural barriers few (angulo-parietal (parietal), upper palatal, lower palatal and columellar) and relatively weak. Umbilicus initially narrow and then abruptly widening at the last whorl.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
DF28882580A65621A850B69ACDD03977.taxon	description	Description. Shell conical or slightly concave-conical, dark brown, consisting of 4 – 4.5 whorls separated by a deep suture. Protoconch rounded, slightly glossy, roughly pitted, consisting of ~ 1.5 whorls. Initial teleoconch whorls rounded, penultimate whorl bluntly keeled. Teleoconch sculptured with strong, raised spiral striae which are crossed by weaker radial growth lines usually in form of whitish radial streaks. There are ~ 30 spiral striae on the last whorl in standard view. The spacing between two striae ranges to ~ 2 – 3 × the width of one stria. Basis of the last whorl flat. Last whorl with a sharp shoulder (almost like a keel), adnate or very slightly detached from the penultimate, very slightly descending near the aperture (less than 5 ° compared to the shell axis), almost straight. Peristome brown, not very strong but still expanded and not reflected. Aperture equipped with four main barriers (parietal (angulo-parietal), upper palatal, lower palatal and columellar), none of which reach the expanding peristome. Angulo-parietal lamella is the strongest, longest and the highest in the aperture. There is usually no sign of the angular part. Upper and lower palatal plicae both low and relatively weak, almost equally strong. Basal plica clearly present and moderately strong or absent (with only a slight swelling in its place). Columellar lamella developed to a similar extent as both palatal plicae. There is one additional swelling in the columello-parietal embayment which corresponds to the infraparietal lamella. Surface of all apertural barriers is finely granulated. Sinulus wide and not distinctly separated from the rest of the aperture. Umbilicus initially narrow, but abruptly widening at the last whorl, becoming very wide, and measuring slightly> ½ of the shell width. There is a deep groove running from the peristome edge towards the umbilicus along the inner side of the last whorl. This groove is terminating at the transition from the last whorl to penultimate.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
DF28882580A65621A850B69ACDD03977.taxon	description	Measurements (in mm, n = 5). SH = 2.36 – 2.75; SW = 2.56 – 2.85; AH = 1.20 – 1.32; AW = 1.25 – 1.47.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
DF28882580A65621A850B69ACDD03977.taxon	etymology	Etymology. This species is named for the flat base of the shell.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
DF28882580A65621A850B69ACDD03977.taxon	distribution	Distribution. This species is only known from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
8506BD4320BD50BDB23729928BE771F5.taxon	diagnosis	Diagnosis. Shell depressed, concave-conical, last whorl strongly detached from the penultimate and ascending at nearly 60 – 70 ° angle. Teleoconch strongly spirally striated. Aperture equipped with four blunt barriers (angulo-parietal, upper palatal, lower palatal, and columellar). Umbilicus very wide.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
8506BD4320BD50BDB23729928BE771F5.taxon	description	Description. Shell depressed, concave-conical (due to the strongly enlarged last whorl), consisting of 3.75 – 4.25 whorls separated by a very deep suture. Colouration yellowish or very pale brownish, opaque. Protoconch initially almost smooth and then finely pitted, showing spiralling pattern and consisting of 1.25 – 1.5 whorls. Boundary between the protoconch and teleoconch clearly visible due to the change of microsculpture. Teleoconch strongly and regularly spirally striated, spiral striae thread-like raised. Space between two spiral striae measuring approximately the width of two to four spiral striae. There are ~ 30 spiral striae on the last whorl in standard view. Weaker, irregularly spaced radial growth lines are also visible crossing the spiral striae. Some irregularly spaced radial white streaks are occasionally present. Last whorl shouldered, strongly detached from the penultimate and ascending strongly near the aperture (~ 60 – 70 ° compared to the shell axis). Peristome lighter than the rest of the shell, fragile, expanded and not reflected. Aperture equipped with four moderately strong barriers (angulo-parietal, upper palatal, lower palatal, and columellar). Angulo-parietal lamella is the strongest in the aperture, consisting of two parts corresponding to angular and parietal lamellae in the former Gyliotrachela. The angular part of the angulo-parietal lamella is very small and pointy, leaning towards the palatal wall. The parietal part of the angulo-parietal lamella is much stronger and positioned deeper in the aperture, leaned towards the columellar lamella. Both palatal plicae and a columellar lamella are developed to similar level, relatively short and blunt. Upper palatal plica nearly fully rectangular in shape, lower palatal plica and columellar lamella more rounded and positioned even deeper in the aperture than the upper palatal plica. The apertural barriers are not very variable. None of these barriers reach the peristome. Surface of all apertural barriers is finely granulated. Sinulus rounded and well separated from the rest of the aperture. Umbilicus open, very wide, measuring ~ 1 / 3 of the shell width and showing all previous whorls.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
8506BD4320BD50BDB23729928BE771F5.taxon	description	Measurements (in mm, n = 5). SH = 1.33 – 1.41; SW = 2.13 – 2.22; AH = 0.79 – 0.86; AW = 0.7 – 0.87.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
8506BD4320BD50BDB23729928BE771F5.taxon	etymology	Etymology. Named after the fact that many sampling sites of this species are popular touristic destinations.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
8506BD4320BD50BDB23729928BE771F5.taxon	distribution	Distribution. This species is known from Krabi, Chumphon and Ranong provinces in Thailand.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
AF8B9E6C07B65A5A9511FBBE4CECAEAF.taxon	description	Figs 224 G, 233, 239	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
AF8B9E6C07B65A5A9511FBBE4CECAEAF.taxon	distribution	Distribution. This species is known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
DC1C4FFC49A95C5F84B051C6EBCE57D1.taxon	distribution	Distribution. This species in known from several localities in Kien Giang province (Vietnam). We have also found this species in Kampot province, Cambodia (see under Material examined).	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
8DB4A8D817915734988534CE3BDB6270.taxon	distribution	Distribution. This species is known only from the limited area (Raub) in Malaysia and Suratthani, Phattalung and Songkhla provinces in Thailand.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
FBA32FF5A19C5B32A91D752F4B11A18A.taxon	distribution	Distribution. This species is known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
3570D9ADBB175A1DB6E8A3D70AE6E5B5.taxon	diagnosis	Diagnosis. Hypselostoma species with all teleoconch whorls keeled, detached last whorl and strongly spirally striated shell surface. There are five strong barriers in the aperture (angulo-parietal, upper palatal, lower palatal, basal and columellar). Umbilicus initially narrow, suddenly widening at the last whorl.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
3570D9ADBB175A1DB6E8A3D70AE6E5B5.taxon	description	Description. Shell conical-ovoid, brownish, not glossy. It is consisting of 4 – 4.5 whorls separated by a deep suture. Protoconch rounded, of ~ 1.75 finely pitted whorls of the same colour as the rest of the shell. Teleoconch sculptured with strong, raised spiral striae. Spacing between two striae equals the width of one stria or is twice as wide as a stria. Spiral striae are more densely arranged on the initial teleoconch whorls, and more coarsely on the last whorl. The number of striae on the last whorl is ~ 25 in standard view. There are several white, unevenly positioned, radial streaks crossing the spiral striae. All teleoconch whorls are keeled, and the strength of the keel is increasing towards the last whorl which is especially strongly keeled. Surfaces of the whorls above and below the centrally positioned keel are flat, to very slightly concave (above the keel on the last whorl). Last whorl moderately detached from the penultimate, distanced from the shell axis. From the lateral view (Fig. 168 B), it can be seen that the last whorl is initially slightly ascending and then straightening right near the aperture so that the aperture profile is almost parallel to the shell axis (leaving the aperture only slightly ascending at ~ 5 – 10 ° compared to the shell axis). Peristome light brownish, expanded and not reflected. Aperture equipped with five barriers (angulo-parietal, upper palatal, lower palatal, basal, and columellar). Angulo-parietal strong and high, lamella usually appears as a single barrier (parietal) but has a very short swollen part in front of it which most probably represents the remains of the angular part. Upper palatal plica moderately high, slightly curved towards the lower palatal plica (somehow similar to the upper palatal plica in H. serpa). Lower palatal plica slightly higher, longer, and more slender than the upper palatal but similar in width. It is pointing towards the embayment between the columellar and parietal lamellae. Basal plica more closely positioned to the lower palatal (not in its usual position between the lower palatal and columellar), ~ 2 × lower than the latter and short. Columellar lamella thick and broad, more closely resembling a very large swelling than a distinct barrier. There is also a small swelling between the columellar and parietal lamellae. Surface of all apertural barriers is finely granulated. Sinulus narrow, distinctly separated from the rest of the aperture. Umbilicus initially very narrow and suddenly widening at the last whorl. It is measuring between ½ and ¹ ⁄ 3 of the shell width. There is a deep groove running from the peristome edge towards the umbilicus along the inner side of the last whorl. This groove is terminating at the transition from the last whorl to penultimate.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
3570D9ADBB175A1DB6E8A3D70AE6E5B5.taxon	description	Measurements (in mm, n = 5). SH = 2.35 – 2.70; SW 1 = 3.52 – 3.80; SW 2 = 2.08 – 2.18; AH = 1.32 – 1.62; AW = 1.40 – 1.79.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
3570D9ADBB175A1DB6E8A3D70AE6E5B5.taxon	etymology	Etymology. This species is named for the strong surface sculpture of its shell.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
3570D9ADBB175A1DB6E8A3D70AE6E5B5.taxon	distribution	Distribution. This species is only known from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
7ED1CB55E4BB592083FE443726198F80.taxon	description	Figs 224 H, 234, 241	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
7ED1CB55E4BB592083FE443726198F80.taxon	distribution	Distribution. This species is known from Kedah, Malaysia (the type locality) as well as from Krabi Province and Ko Tarutao Island, Satun province, Thailand (the type locality of G. tarutao).	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
95DC87B3CABE5743B611F5C3C4BD9EF8.taxon	description	Figs 186 L, 209, 210, 223	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
95DC87B3CABE5743B611F5C3C4BD9EF8.taxon	distribution	Distribution. This species is, apart from the type locality, known from one more sampling site in Sa Kaeo Province.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
7F69B785C94B5EDF9CC905798722236A.taxon	description	Figs 242 H, 257, 258, 259	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
7F69B785C94B5EDF9CC905798722236A.taxon	diagnosis	Diagnosis. Shell triangular conical. Teleoconch almost smooth, finely radially but not spirally striated. Last whorl detached from the penultimate and ascending, keeled below the centre of the periphery. Aperture equipped with weak barriers, five main (angular, parietal, upper palatal, lower palatal and columellar) and several smaller ones. Umbilicus narrow, measuring 1 / 7 – 1 / 8 of the shell width.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
7F69B785C94B5EDF9CC905798722236A.taxon	description	Description. Shell conical, light brownish, opaque, consisting of 3.75 – 4.25 weakly convex and regularly increasing whorls separated by a relatively shallow suture. Protoconch finely pitted, without spiral striae, consisting of ~ 1.5 whorls, same colour as the rest of the shell. Teleoconch sculpture delicate, very finely dimpled (pasty) and crossed by relatively weak radial growth lines. Spiral striae absent. Occasionally, a few stronger whitish radial streaks can be observed at the last whorl near the aperture. Last whorl weakly detached from the penultimate and slightly ascending near the aperture (~ 15 – 20 ° compared to the shell axis). It is bluntly but prominently keeled below the periphery, making the last whorl oblique to the shell axis and not convex but with a sloping, flat outline. There is a slight furrow above the keel. Peristome thick, whitish, expanded and not reflected. All apertural barriers are weak. Aperture equipped with five main barriers (angular, parietal, upper palatal, lower palatal, and columellar) and several smaller ones of variable number and appearance. There are usually no plicae in the sinulus, usually one plica in the interpalatal region, two or three plicae in the region of the basal plica and usually one or two lamellae in the columello-parietal region. Angular lamella tubercle-like, almost reaching the peristome. Behind it, more deeply situated, there is one additional part which also belongs to the angular lamella. Parietal lamella stronger and higher when compared to the others (except lower palatal plica), not reaching the profile of the angular tubercle. Lower palatal plica developed to the same extent as the parietal lamella and slightly higher and stronger than the upper palatal plica. Columellar lamella horizontal, developed equally as the upper palatal plica. Anterior to the upper palatal plica, there is one more tubercle-like part, possibly homologous with the palatal tubercle found in the genus Bensonella (see Páll-Gergely and White 2023) or it even might represent a part of the upper palatal plica. Weaker barriers are all positioned deeper in the aperture and never reaching the profiles of the main ones. A tubercle-like swelling is present in the columellar-parietal transition embayment. All barriers are strongly and coarsely spiniferous. Sinulus small, rounded and distinctly separated from the rest of the aperture. Umbilicus narrow, with almost perpendicular walls, showing only a part of the penultimate whorl and measuring ~ 1 / 7 – 1 / 8 of the shell width.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
7F69B785C94B5EDF9CC905798722236A.taxon	etymology	Etymology. Named after the fact that this species is in shell shape very similar to some Hypselostoma species inhabiting the Philippines.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
7F69B785C94B5EDF9CC905798722236A.taxon	description	Measurements (in mm, n = 3). SH = 1.82 – 1.91; SW 1 = 2.34 – 2.51; SW 2 = 1.45 – 1.51; AH = 0.98 – 1.01; AW = 0.9 – 0.98.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
7F69B785C94B5EDF9CC905798722236A.taxon	distribution	Distribution. This species is known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
968874338F085E538FEF1899AD1850DA.taxon	distribution	Distribution. This species is known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
F0E153FF8E5E5689ADB24DF76E7EEE3F.taxon	diagnosis	Diagnosis. A Hypselostoma species with depressed, concave-conical shell, bluntly and strongly keeled last whorl, raised spiral striation, hooked apertural barriers (upper palatal, interpalatal, lower palatal and basal) and very wide umbilicus. Angular and parietal lamellae fused. Shell width reaches 4 mm.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
F0E153FF8E5E5689ADB24DF76E7EEE3F.taxon	description	Description. Shell depressed, concave-conical, brown to yellowish-brown, consisting of 4.75 – 5.5 whorls separated by a deep suture. Protoconch slightly lighter than the rest of the shell, finely pitted, showing spiralling pattern initially and clearer spiral striae terminally. It is consisting of ~ 1.25 – 1.5 whorls. Protoconch and initial teleoconch whorls rounded, penultimate weakly convex. Last whorl with strong but blunt keel positioned at the centre of the periphery. Above the keel there is a deep or rarely shallow groove. All whorls densely sculptured with strong, raised spiral striae crossed by less dense radial growth lines and occasionally by a few thick, whitish radial streaks. Spacing between the spiral striae irregular, but usually around the width of two to four spiral striae. Last whorl moderately detached from the penultimate and moderately ascending (~ 35 ° compared to the shell axis). Peristome expanded, not reflected, its surface finely pitted. Aperture equipped with four main barriers (angulo-parietal, upper palatal, lower palatal, and columellar) and several smaller ones. Angulo-parietal lamella strong, its angular part is leaned towards the palatal side while the parietal part is leaned towards the columellar side. Angular part is pointed and smaller than the parietal. It is the only barrier reaching the expanding peristome. Sometimes, there is a deep constriction between the angular and parietal part so that these lamellae appear almost separated (as in former Gyliotrachela). Upper and lower palatal plicae are hooked, pointing outside and roughly the same size. Columellar lamella thick and almost horizontal. Of smaller barriers, there is usually one hooked interpalatal, one infrapalatal, one basal plica (hooked or blunt) and one blunt lamella in the columello-parietal transition embayment. Of them, the strongest one is the basal, which is almost as strong as the palatals. A distinct swelling is also observed at the palatal side (in front of the upper palatal and interpalatal plicae) which is probably homologous to the palatal tubercle in the majority of Bensonella species. Surface of all apertural barriers is finely granulated. Sinulus rounded, distinctly separated from the rest of the aperture. Umbilicus very wide, measuring ~ ½ of the shell width. There is a periumbilical keel situated on the last ~ 0.75 whorl right above the umbilicus. Along this keel there is a deep groove on the umbilical side.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
F0E153FF8E5E5689ADB24DF76E7EEE3F.taxon	description	Measurements (in mm, n = 15). SH = 1.62 – 2.78; SW 1 = 2.44 – 4.01; SW 2 = 1.57 – 2.56; AH = 0.85 – 1.71; AW = 0.81 – 1.46.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
F0E153FF8E5E5689ADB24DF76E7EEE3F.taxon	etymology	Etymology. The name of this species is derived from two Latin words, soror meaning sister and major meaning greater, larger, or elder, which serves as a comparison with similar but smaller H. sororminor sp. nov.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
F0E153FF8E5E5689ADB24DF76E7EEE3F.taxon	distribution	Distribution. This species is known from four closely situated localities in Steung Treng Province.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
B902884D17DF528C95464B957F1870A0.taxon	diagnosis	Diagnosis. A Hypselostoma species with depressed, concave-conical shell, bluntly and strongly keeled last whorl, raised spiral striation, blunt apertural barriers (altogether 6 or 7) and very wide umbilicus. Angular and parietal lamellae fused.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
B902884D17DF528C95464B957F1870A0.taxon	description	Description. Shell depressed, concave-conical, brown, consisting of 5 – 5.5 whorls separated by a deep suture. Protoconch lighter than the rest of the shell, coarsely spirally striated (~ 12 striae) and consisting of 1.25 – 1.5 whorls. Protoconch and initial teleoconch whorls rounded, penultimate weakly convex. Last whorl with strong but blunt keel positioned at the centre of the periphery. Above the keel there is a deep groove. All whorls densely sculptured with strong, raised spiral striae crossed by less dense radial growth lines and occasionally by a few thin, whitish radial streaks. Spacing between the spiral striae irregular, but usually around the width of two, three, or four spiral striae. Last whorl slightly to moderately detached from the penultimate and slightly to moderately ascending near the aperture (~ 25 – 35 ° compared to the shell axis). Peristome expanded, not reflected, its surface finely pitted. Aperture equipped with four strong, main barriers (angulo-parietal, upper palatal, lower palatal, and columellar) and several smaller ones. Angulo-parietal lamella strong, its angular part is leaned towards the palatal side while its parietal part is leaned towards the columellar side. Angular part is pointed and smaller than the parietal. It is the only barrier reaching the expanding peristome. Upper and lower palatal plicae approximately the same size, the lower palatal being longer and more slender. Upper palatal plica slightly bent towards the lower palatal. Columellar lamella very thick and almost horizontal. Of smaller barriers, a small and weak interpalatal plica is present, stronger (but still weak) basal and a moderate knob-like plica in the columello-parietal transition embayment. A weak swelling is also observed at the palatal side (in front of the upper palatal plicae) which is probably homologous to the palatal tubercle in the majority of Bensonella species. Surface of all apertural barriers is densely granulated. Sinulus rounded, distinctly separated from the rest of the aperture. Umbilicus very wide, measuring between 1 / 3 and ½ of the shell width. There is a periumbilical keel situated on the last ~ 0.75 whorl right above the umbilicus. Along this keel there is a deep groove on the umbilical side.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
B902884D17DF528C95464B957F1870A0.taxon	description	Measurements (in mm, n = 5). SH = 2.37 – 2.57; SW 1 = 3.20 – 3.61; SW 2 = 2.05 – 2.40; AH = 1.25 – 1.42; AW = 1.22 – 1.26.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
B902884D17DF528C95464B957F1870A0.taxon	etymology	Etymology. The name of this species is derived from two Latin words, soror meaning sister and minor meaning smaller which serves as a comparison with similar but usually larger H. sorormajor sp. nov.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
B902884D17DF528C95464B957F1870A0.taxon	distribution	Distribution. This species is only known from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
A887FB65FABB5BFDACD4C14C7083EBAF.taxon	description	Figs 186 M, 211, 223	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
A887FB65FABB5BFDACD4C14C7083EBAF.taxon	distribution	Distribution. This species is known from four sampling sites, two in Sa Kaeo province (Thailand) and two in Banteay Meanchey province (Cambodia) (Sutcharit et al. 2023).	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
C1EECA34E5E15C94A4A28728002CE654.taxon	description	Figs 186 N, 212, 223	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
C1EECA34E5E15C94A4A28728002CE654.taxon	distribution	Distribution. This species is known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
4C91D7CE8F855BCC907456E6CD0C0B92.taxon	distribution	Distribution. This species is known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
DE00A42C28DD5827AF5C620726E1EE1F.taxon	description	Figs 224 I, 235, 236, 237, 241	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
DE00A42C28DD5827AF5C620726E1EE1F.taxon	distribution	Distribution. This species is known from Pahang and Selangor provinces in Peninsular Malaysia.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
4133167011B95CA49DC7D3CF8F5B35DB.taxon	description	Figs 224 J, 238, 239	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
4133167011B95CA49DC7D3CF8F5B35DB.taxon	etymology	Etymology. The specific epithet is made of tertius meaning third and frater meaning brother, given the fact that this is one of three Hypselostoma species described from Borneo.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
4133167011B95CA49DC7D3CF8F5B35DB.taxon	distribution	Distribution. This species in known from Sarawak and Sabah, Borneo.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
EDCFC5D2D0C25AF6B17E4596E8649E84.taxon	diagnosis	Diagnosis. A Hypselostoma with all teleoconch whorls shouldered and very weakly spirally striated. Spiral striae not raised. Last whorl detached from the penultimate and slightly descending. Four weak barriers in the aperture. Columellar lamella is like a strong swelling, forming a basal furrow.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
EDCFC5D2D0C25AF6B17E4596E8649E84.taxon	description	Description. Shell concave-conical (due to the strongly enlarged last whorl), brownish, consisting of 5 – 5.5 whorls separated by a very deep suture. Protoconch finely pitted, not showing spiralling pattern, rounded, same colour as the initial teleoconch whorls but slightly lighter than the last whorl. First two teleoconch whorls weakly shouldered, penultimate strongly and the last whorl is very strongly shouldered. The outline of the last and the penultimate whorl above the shoulder is concave (forming a flat platform). This surface is on initial teleoconch whorls more sloping. Whole teleoconch surface is weakly spirally striated (these spiral striae are not thread-like raised as in many congeners, but very blunt and indistinct) and occasionally crossed by radial growth lines and several thicker, whitish radial streaks which are rather unevenly positioned. These whitish streaks seem to be the densest on the penultimate whorl. Last whorl convex at its base, towards the umbilicus. It is slightly to moderately detached from the penultimate whorl and slightly descending near the aperture (~ 15 – 25 ° compared to the shell axis). Peristome very strong, dirty white, expanded but not reflected. Aperture equipped with three or four barriers (parietal (angulo-parietal?), upper palatal, lower palatal (sometimes absent), and columellar). There are no smaller barriers between the main ones. Parietal lamella is strong, high, and triangular when observed in perpendicular view. There is a very small tubercle-like swelling in front of the parietal lamella which may be homologous with the angular part of the usually present angulo-parietal (or just angular) in other congeners. Upper palatal plica moderately strong, almost twice as weak as the parietal lamella. Lower palatal plica very weak and low, dot-like, sometimes absent. Columellar lamella not present in its usual form (its borders are not clearly delimited) but as a rather strong, swelled part of the aperture which forms a prominent basal furrow below it (similar to the one present in H. fungus sp. nov.). The shape of the columellar side of the aperture is also reminiscent of the playground slide. Surface of all apertural barriers is smooth. Sinulus wide, distinctly separated from the rest of the aperture. Umbilicus wide, measuring ~ 1 / 3 of the shell width, expanding at the last whorl. Inside the umbilicus, there is a strong groove present, stretching across the whole last whorl.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
EDCFC5D2D0C25AF6B17E4596E8649E84.taxon	description	Measurements (in mm, n = 5). SH = 3.45 – 3.8; SW = 2.83 – 3.62; AH = 1.62 – 1.82; AW = 1.38 – 1.84.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
EDCFC5D2D0C25AF6B17E4596E8649E84.taxon	etymology	Etymology. Since all whorls of this species are shouldered, the shell has an appearance of the wedding cake. The specific epithet torta comes from a Serbian word torta meaning a cake. To be used as a noun in apposition.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
EDCFC5D2D0C25AF6B17E4596E8649E84.taxon	distribution	Distribution. This species is known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
B2D28E0DA1275D35B9CDD2BA3574FFE6.taxon	distribution	Distribution. This species is known only from the Battambang province, Cambodia.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
D376B9833A165298A454CD46DC6DD0E3.taxon	description	Figs 186 O, 213, 214, 223	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
D376B9833A165298A454CD46DC6DD0E3.taxon	distribution	Distribution. This species in known from Chumphon and Suratthani provinces in Thailand.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
3F54769239FE575F881C38DB805CFDE1.taxon	distribution	Distribution. This species is, apart from the type locality, also known from Phnom Prampi Cave in Battambang Province which is a first record from Cambodia.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
64C84E2FAE2A5EC2AA14A4B042FB6EA4.taxon	description	Figs 224 K, 240, 241	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
64C84E2FAE2A5EC2AA14A4B042FB6EA4.taxon	distribution	Distribution. This species is known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
F475DBCA40045DADA0BA4AF63CEA4EF7.taxon	description	Figs 242 I, 260, 261	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
F475DBCA40045DADA0BA4AF63CEA4EF7.taxon	distribution	Distribution. This species is, apart from the type locality, known from several localities in Mandalay, Myanmar.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
E032DFB56C205AAC9BADEEFA5BC7BC80.taxon	distribution	Distribution. This species is known from two provinces in Thailand, Supanburi (type locality) and Kanchanaburi (this study).	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
A02A9D5E1C08555BA9BECCC1A11E5401.taxon	description	Figs 186 P, 215, 216, 223	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
A02A9D5E1C08555BA9BECCC1A11E5401.taxon	distribution	Distribution. This species is known only from three localities in Peninsular Malaysia (Gunong Pondok, Kota Tongkat and Gua Kota Gelanggi).	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
3E3B47F9AE5F59EA86FC7143B4614F37.taxon	description	Figs 186 Q, 217, 218, 223	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
3E3B47F9AE5F59EA86FC7143B4614F37.taxon	diagnosis	Diagnosis. Shell concave-conical, with roughly granulated (sandpaper-like) surface sculpture. Last two whorls keeled below the periphery, keel especially strong on the last whorl. Last whorl detached from the penultimate, directed upwards. Aperture equipped with five main barriers and usually five smaller ones. All barriers are very broad and thick, very strongly spiniferous. Umbilicus very narrow, dot-like.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
3E3B47F9AE5F59EA86FC7143B4614F37.taxon	description	Description. Shell concave-conical (due to the strongly enlarged last whorl), pinkish-brown in colour, corneous. It is consisting of 4.5 – 5 regularly increasing whorls separated by a deeply impressed suture. Protoconch finely pitted, not showing spiralling pattern, consisting of 1.25 – 1.5 whorls and slightly darker than the rest of the shell. Protoconch- teleoconch boundary clearly visible due to the change in shell surface sculpture. Teleoconch roughly granulated, sandpaper-like. On the last whorl near the keel, granules can in some specimens be arranged in a way to provoke a spiralling pattern although no regular spiral striae are developed. Radial lines are strong, widely and irregularly spaced, ~ 17 radial lines on the last whorl in standard view. They are more densely arranged on the initial teleoconch whorls. Protoconch and initial teleoconch whorls rounded and convex, last two whorls are keeled below the centre of the periphery. Last whorl especially strongly keeled, with a deep groove above the keel. It is also moderately detached from the penultimate near the aperture, and slightly ascending (~ 15 – 25 ° compared to the shell axis). Peristome whitish, strongly expanded but not reflected. Aperture large, equipped with five main barriers (parietal, angular, upper palatal, lower palatal and columellar). Angular lamella is the weakest and the shortest among the main five. All other main barriers are relatively thick and broad, almost equally developed. Parietal lamella is the longest in the aperture and surpassing the profile of the angular. Between the main barriers, several smaller ones could be observed. There are usually two interpalatal plicae, two between the lower palatal and the columellar and one knob-like infraparietal lamella. All barriers, as well as the aperture surface, are very strongly spiniferous. These spines are widely spaced (especially on the barriers) and much stronger than in other congeners. Sinulus not very distinctly separated from the rest of the aperture. Umbilicus is very narrow, dot-like, not showing previous whorls and measuring slightly> 1 / 10 of the shell width. A very shallow groove is visible running alongside the umbilicus.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
3E3B47F9AE5F59EA86FC7143B4614F37.taxon	description	Measurements (in mm, n = 5). SH = 2.10 – 2.35; SW 1 = 2.53 – 3.12; SW 2 = 1.49 – 1.77; AH = 1.10 – 1.23; AW = 1.05 – 1.25.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
3E3B47F9AE5F59EA86FC7143B4614F37.taxon	etymology	Etymology. Named after Dr. Nikola Vesović, a prominent Serbian entomologist and a friend of the first author.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
3E3B47F9AE5F59EA86FC7143B4614F37.taxon	distribution	Distribution. Known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
80BF9D4EEC8F5050810CAE16FBD94350.taxon	description	Figs 186 R, 219, 220, 223	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
80BF9D4EEC8F5050810CAE16FBD94350.taxon	diagnosis	Diagnosis. A Hypselostoma species with roughly granulated (not spirally striated) teleoconch, and keeled, detached, and ascending last whorl. Apertural barriers few (five main ones and usually three smaller ones).	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
80BF9D4EEC8F5050810CAE16FBD94350.taxon	description	Description. Shell depressed, concave-conical (due to the strongly enlarged last whorl), chestnut brown and weakly glossy, consisting of 3.75 – 4.25 convex whorls separated by a deep suture. Protoconch is of the same colouration as the teleoconch, finely pitted and showing no spiralling pattern. Boundary between the protoconch and the teleoconch not clear under microscope because of the very similar surface sculpture of the two regions, but the former is ~ 1.5 whorls. Teleoconch roughly granulated (sandpaper-like), weakly sculptured with radial growth lines but devoid of spiral striation (these radial lines are much harder to observe than the prominent granulation). Occasionally, thicker whitish radial streaks are unevenly placed across the surface of the shell. Last whorl with a very blunt keel (which is getting stronger near the aperture) positioned at the centre of the periphery. Shell surface is immediately below and above the keel concave, and then again convex towards the suture and the umbilicus. Last whorl is also very slightly detached from the penultimate and moderately ascending upwards (~ 40 – 50 ° compared to the shell axis). Peristome dirty white, expanded but not reflected, its surface very finely pitted. There are altogether five main apertural barriers (angular, parietal, upper palatal, lower palatal and columellar) and several smaller ones of variable number. Parietal lamella is the strongest and highest in the aperture, straight but directed towards the palatal side. Angular lamella ~ 2 × lower and shorter than the parietal, but extends closer to the expanding peristome, almost reaching it. Upper and lower palatal plicae almost the same in height and length, the former may be slightly weaker than the latter. Columellar lamella horizontal, almost identical to the lower palatal plica. Among the smaller barriers, there is usually one interpalatal plica which is very small, dot-like. Slightly larger basal plica is present nearly halfway between the lower palatal and columellar. One dot-like swelling is present in the columello-parietal transition embayment. Surface of all apertural barriers is very finely spiniferous medially and very finely granulated laterally. Sinulus wide, not distinctly separated from the rest of the aperture. Umbilicus wide, measuring slightly> 1 / 4 of the shell width. A shallow groove runs alongside the umbilicus but is visible only around the last quarter of the last whorl.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
80BF9D4EEC8F5050810CAE16FBD94350.taxon	description	Measurements (in mm, n = 5). SH = 1.73 – 2.06; SW 1 = 2.49 – 2.85; SW 2 = 1.63 – 1.79; AH = 0.93 – 1.19; AW = 0.94 – 1.10.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
80BF9D4EEC8F5050810CAE16FBD94350.taxon	etymology	Etymology. The specific epithet vicinum comes from the Latin word for neighbour. This name is provided because the most similar species to the new one, H. troglodytes, lives in the neighbouring Peninsular Malaysia.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
80BF9D4EEC8F5050810CAE16FBD94350.taxon	distribution	Distribution. This species is known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
BC7FB5AE88D6531187EFB079DAE51A20.taxon	description	Figs 186 S, 221, 222, 223	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
BC7FB5AE88D6531187EFB079DAE51A20.taxon	diagnosis	Diagnosis. A Hypselostoma species with 5.75 – 6 roughly granulated whorls and a very deep suture. Last whorl somehow canted and adnate to the penultimate near the aperture. Palatal plicae and basal plica hooked. Other barriers (angulo-parietal, columellar and infraparietal) blunt. Umbilicus narrow.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
BC7FB5AE88D6531187EFB079DAE51A20.taxon	description	Description. Shell high, concave-conical, shiny brown or even colourless, consisting of 5 – 5.5 whorls separated by a very deep suture. Protoconch almost smooth, with very weak spiralling pattern, consisting of ~ 1.5 whorls but its boundary with the teleoconch is not clearly visible due to the similar surface sculpture. Teleoconch with weak and coarse radial growth lines, which are on the last two whorls visible as white streaks, and rough granulation (sandpaper-like) but no spiral striation. Initial teleoconch whorls rounded, penultimate only very slightly less rounded. Last whorl adnate to slightly detached from the penultimate, with a very weak keel situated near the centre of the periphery. It is slightly descending near the aperture (~ 5 – 10 ° compared to the shell axis). Peristome dirty white, expanded but not reflected. Aperture rounded (almost as wide as high), equipped with six main barriers (angulo-parietal, upper palatal, interpalatal, lower palatal, basal and columellar) and one weak infraparietal swelling-like lamella. Angulo-parietal lamella is the strongest in the aperture, with clearly distinguishable angular and parietal part and a prominent sinuation between them. Parietal part stronger and higher than the angular, both curved towards the palatal wall. All palatal plicae hooked, pointing outside. Upper palatal and interpalatal plica positioned close together, almost equally as strong. Lower palatal plica stronger than others and situated some distance from the interpalatal. Basal plica also hooked but less clearly than the palatals, strong as the interpalatal and upper palatal but situated closer to the columellar lamella than to the lower palatal plica. Columellar lamella not hooked, long and almost horizontal. A small swelling-like infraparietal lamella is present. Angulo-parietal lamella is reaching the expanding peristome, columellar lamella almost reaching it. Surface of all apertural barriers is finely granulated. Sinulus wide, not distinctly separated from the rest of the aperture. Umbilicus always open but of variable width, measuring ~ 1 / 6 or ≤ 1 / 3 of the shell width. Umbilical groove is strong in specimens with wider and weak in specimens with narrower umbilicus.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
BC7FB5AE88D6531187EFB079DAE51A20.taxon	description	Measurements (in mm, n = 4). SH = 2.36 – 2.62; SW = 1.78 – 1.93; AH = 0.92 – 0.96; AW = 0.85 – 0.93.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
BC7FB5AE88D6531187EFB079DAE51A20.taxon	etymology	Etymology. Named after Mihailo Vujić, a prominent Serbian entomologist and a friend of the first author.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
BC7FB5AE88D6531187EFB079DAE51A20.taxon	distribution	Distribution. This species is known only from the type locality.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
4B7B972D41A757539C4BD3938597E898.taxon	diagnosis	Diagnosis. This genus is subdivided into four species groups: i) H. bensonianum group ii) H. hungerfordianum group; iii) H. terae group and iv) H. tubiferum group. This subdivision is based on the shell surface sculpture (coupled with apertural barrier morphology and geographic distribution in H. terae group) which is raised spirally striated in H. bensonianum group, granulated (sandpaper-like) in H. hungerfordianum group and fine, pasty in H. terae and H. tubiferum group. Most representatives share the depressed-conical or conical shell shape with last whorl adnate to or detached from the penultimate.	en	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
