identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
616869CC0FEC5DDAB3D84C8E1BC517E3.text	616869CC0FEC5DDAB3D84C8E1BC517E3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anauchen angthongensis Panha 2002	<div><p>Anauchen angthongensis Panha, 2002</p><p>Figs 10 A, 11, 37</p><p>Anauchen angthongense Panha in Burch &amp; Panha, 2002: 245–246, fig. 4.</p><p>Anauchen angthongense — Panha and Burch 2008: 48, fig. 44; Dumrongrojwattana and Wongkamhaeng 2024: 323, fig. 7.</p><p>Type material examined.</p><p>Thailand • 1 paratype; from the type locality; S. Panha leg.; SMF 3311444 .</p><p>Type locality.</p><p>“ Angthong National Park, Suratthani Province Thailand, 9°38'6"N, 99°41′16"E, 60 meters elevation … ”.</p><p>Differential diagnosis.</p><p>This species is similar to many congeners, mostly based on the appearance of the apertural barriers. For differences, see under the following species: A. crassus sp. nov., A. huaykhakang, A. informis, A. obesus sp. nov., A. utaithaniensis and A. banmiensis .</p><p>Distribution.</p><p>This species is known only from the type locality.</p><p>Remarks.</p><p>Basal plica can be present or absent.</p></div>	https://treatment.plazi.org/id/616869CC0FEC5DDAB3D84C8E1BC517E3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
572693BB11185749B4C37128EC9939C6.text	572693BB11185749B4C37128EC9939C6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anauchen banmiensis Panha & J. B. Burch 2004	<div><p>Anauchen banmiensis Panha &amp; J. B. Burch, 2004</p><p>Figs 10 B, 12, 37</p><p>Anauchen banmiensis Panha &amp; Burch in Panha et al. 2004: 59–60, fig. 2.</p><p>Hypselostoma banmiensis — Panha and Burch 2008: 89–90, fig. 76; Dumrongrojwattana and Wongkamhaeng 2024: 324, fig. 8.</p><p>Type material examined.</p><p>Thailand • 1 paratype; from the type locality; S. Panha leg.; CUMZ ver. 44026 .</p><p>Type locality.</p><p>“ Chang Peuag Cave Mountain, Banmi District, Lopburi Province, 14°54'10"N, 100°30'8"E, 15 meters elevation ” (Thailand).</p><p>Differential diagnosis.</p><p>This species differs from A. angthongensis by its shouldered whorls (not rounded as in the latter) and more enlarged last whorl. For differences from further similar species, see under A. crassus sp. nov., A. huaykhakang, A. informis, and A. utaithaniensis .</p><p>Distribution.</p><p>This species is known only from the type locality.</p><p>Remarks.</p><p>This species was also considered a member of Hypselostoma (Panha and Burch 2008) but given the fact that the shell shape and microsculpture resembles some Anauchen species ( A. huaykhakang, A. utaithaniensis), we assign this species to the latter.</p></div>	https://treatment.plazi.org/id/572693BB11185749B4C37128EC9939C6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
C03226159ACA5868AB79A36EBACD07A2.text	C03226159ACA5868AB79A36EBACD07A2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anauchen crassus Gojsina, Hunyadi & Pall-Gergely 2025	<div><p>Anauchen crassus Gojšina, Hunyadi &amp; Páll-Gergely sp. nov.</p><p>Figs 10 C, 13, 14, 37</p><p>Type material.</p><p>Holotype. Thailand • 1 empty shell (SH: 4.83 mm, SW: 3.54 mm); Ranong Province, south of Kra Buri, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.76555&amp;materialsCitation.latitude=10.324667" title="Search Plazi for locations around (long 98.76555/lat 10.324667)">Tham Phra Kayang</a>, southeastern part of the rock; 10°19.480'N, 98°45.933'E; 10 m a. s. l.; 22 Feb. 2015; A. Hunyadi leg.; CUMZ 14427 . Paratypes. Thailand • 6 shells; same data as for holotype; coll. HA .</p><p>Additional material examined.</p><p>Thailand • 1 shell (damaged, not paratype); same data as for holotype; coll. HA .</p><p>Type locality.</p><p>Thailand, Ranong Province, south of Kra Buri, Tham Phra Kayang, southeastern part of the rock; 10°19.480'N, 98°45.933'E; 10 m a. s. l.</p><p>Diagnosis.</p><p>Anauchen species with finely dimpled (pasty) shell surface sculpture (no spiral striae), aperture adnate to last whorl, five strong apertural barriers (parietal, upper palatal, lower palatal, basal, columellar).</p><p>Description.</p><p>Shell conical, consisting of 5.5–6 thick-walled, regularly increasing, rounded, convex whorls separated by a moderately deep suture. All shells from the type material are somewhat weathered but colouration is dark reddish brown to pinkish, opaque. Protoconch weathered so the surface sculpture is not clear, not spirally striated, consisting of ~ 1.5–1.75 whorls, glossy and slightly darker than the rest of the shell. Teleoconch pasty, very finely radially striated, with no sign of spiral striation. Last whorl adnate to the penultimate and slightly descending near the aperture (~ 10 ° compared to the shell axis) making the aperture profile prosocline to the shell axis. Peristome continuous, leaning on the penultimate whorl at the parietal side in form of a thick and distinct callus. It is slightly paler in colour than the rest of the shell, expanded (particularly on the parietal side where it forms a distinct callus) but not reflected. Aperture equipped with five strong, high, relatively short, and thick barriers (parietal, upper palatal, lower palatal, basal, and columellar). Parietal lamella is the strongest in the aperture, blade-like, slightly oblique, its outer part curved towards the palatal side and only almost reaching the expanding peristome. Upper palatal and lower palatal plicae are similar, the lower one being stronger, more slender, higher, and positioned slightly deeper in the aperture. Basal plica small and very short, knob-like. Columellar lamella thick and horizontal, situated in the middle of the columellar wall. None of these barriers reach the expanding peristome. Surface of all apertural barriers is smooth. Sinulus wide and parabolic (due to the position of angular lamella and upper palatal plica). Umbilicus open and deep but moderately wide, measuring ~ 1 / 6 of the shell width. Only the penultimate whorl is visible through the umbilicus. A very short and shallow groove is present on the last ~ ¼ of the body whorl, running from the peristome towards the umbilicus.</p><p>Differential diagnosis.</p><p>This species is similar to Anauchen angthongensis Panha, 2002 from which it can be distinguished by the lack of spiral striation and much wider umbilicus. Additionally, A. crassus sp. nov. has a shallower suture resulting in less convex whorls than in A. angthongensis . Anauchen informis Vermeulen, Phung &amp; Truong, 2007 is also spirally striated, has narrower umbilicus and its whorls are more convex and less regularly increasing. Anauchen banmiensis is spirally striated and has a shouldered last whorl. For differences from A. obesus sp. nov., see under that species. Boysidia phatangensis Dumrongrojwattana &amp; Assawawattagee, 2018 is more slender and has more whorls which are more convex, with deeper sutures. It usually also has an additional lamella in the columellar-parietal boundary region, which was not (but may be) present in A. crassus sp. nov. Boysidia phatangensis has a bifid lamella (concrescent angular and parietal) on the parietal side while A. crassus sp. nov. has one blade-like lamella. Basal plica in B. phatangensis is seemingly more closely positioned to columellar lamella, while that of A. crassus sp. nov. is somewhere between the lower palatal and columellar or even closer to the lower palatal. Infraparietal lamella is absent in the new species, but present in B. phatangensis . Lastly, the umbilicus of B. phatangensis is slightly more excentrical and wider than in A. crassus sp. nov.</p><p>Measurements</p><p>(in mm, n = 5). SH = 4.83–5.39; SW = 3.54–4.04; AH = 2.15–2.24; 2.04–2.23.</p><p>Etymology.</p><p>The specific epithet refers to the thick (Lat. crassus) shell and apertural barriers of this species.</p><p>Distribution.</p><p>Only known from the type locality.</p></div>	https://treatment.plazi.org/id/C03226159ACA5868AB79A36EBACD07A2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
DFE7170FC1AC5C339646558EB32FA710.text	DFE7170FC1AC5C339646558EB32FA710.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anauchen eotvosi Pall-Gergely 2023	<div><p>Anauchen eotvosi Páll-Gergely, 2023</p><p>Figs 10 D, 15, 37</p><p>Anauchen eotvosi Páll-Gergely, 2023 b: 452–454, fig. 1.</p><p>Anauchen eotvosi — Tongkerd et al. 2024: 165.</p><p>Type material examined.</p><p>Holotype. Thailand-Myanmar • Shan-Siam boundary; Woodthorpe coll.; NHMUK 1903.7. 1.1227. 1 . Paratypes. Thailand-Myanmar •; 6, same data as for holotype; NHMUK 1903.7. 1.1227. 2 – 7 .</p><p>Type locality.</p><p>“ Shan [Shan States, Myanmar] - Siam [Thailand] Boundary ”.</p><p>Differential diagnosis.</p><p>This species differs from its congeners by the presence of weak barriers situated close to the peristome edge. All other congeners similar in shape (e. g., A. angthongensis, A. crassus sp. nov., A. informis), have barriers much stronger and higher. See also under H. iunior sp. nov.</p><p>Distribution.</p><p>This species is known only from the imprecise type locality (Shan-Siam boundary).</p></div>	https://treatment.plazi.org/id/DFE7170FC1AC5C339646558EB32FA710	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
B2B8499F32AE582CAB4D56CB0486E448.text	B2B8499F32AE582CAB4D56CB0486E448.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anauchen evanidus Gojsina & Pall-Gergely 2025	<div><p>Anauchen evanidus Gojšina &amp; Páll-Gergely sp. nov.</p><p>Figs 10 E, 16, 17, 37</p><p>Type material.</p><p>Holotype. Thailand • 1 shell (SH: 2.57 mm, SW: 2.23 mm); Chanthaburi Province, 3.0 km W of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.881&amp;materialsCitation.latitude=12.738" title="Search Plazi for locations around (long 101.881/lat 12.738)">Na Yai Am</a>; 12.738°N, 101.881°E; 23 Apr. 1987; F. G. Thompson leg.; UF 530655 . Paratypes. Thailand • 2 shells; same data as for holotype; CUMZ 14431 • 33 shells; same data as for holotype; UF 591355 .</p><p>Additional material examined.</p><p>Thailand • 2 shells (1 damaged and 1 juvenile, not paratypes); same data as for holotype; UF 583732 .</p><p>Type locality.</p><p>Thailand, Chanthaburi Province, 3.0 km W of Na Yai Am; 12.738°N, 101.881°E.</p><p>Diagnosis.</p><p>Anauchen with very slightly keeled last whorl (below the centre of the periphery) which is also detached from the penultimate. Teleoconch finely dimpled, pasty sculptured, with no spiral striae. Five or six apertural barriers present (parietal, infraparietal, upper palatal, lower palatal, basal and columellar), all strongly spiniferous.</p><p>Description.</p><p>Shell conical, brown, consisting of 4.5–5 convex whorls separated by a deep suture. Protoconch similar in colour as the rest of the shell, finely pitted, showing no spiralling pattern or spiral striae and consisting of 1.25–1.5 whorls. Teleoconch surface sculpture fine, pasty, without spiral striae and with several irregularly spaced whitish radial streaks which are most prominent on the last whorl. Last whorl convex but weakly keeled below the centre of the periphery, slightly detached from the penultimate near the aperture and very slightly ascending (below 5 ° compared to the shell axis). Peristome dirty white, expanded but not reflected. Aperture rounded, equally as wide as high. It is equipped with five or six simple apertural barriers (parietal, infraparietal, upper palatal, lower palatal, basal and columellar). Parietal lamella high and moderately strong, not reaching the expanding peristome. Upper and lower palatal plicae equally strong but the lower palatal is usually slightly higher. Basal plica present as a very weak, dot-like swelling situated halfway between the lower palatal plica and columellar lamella. The latter is sometimes almost as wide as long, provoking a rounded shape. The columellar wall of the aperture is weakly swollen around the columellar lamella. Infraparietal lamella present as a small swelling or completely absent. None of the apertural barriers are reaching the peristome (although the parietal lamella is the closest) and all are roughly spiniferous as well as the apertural surface. Sinulus wide and not strongly isolated from the rest of the aperture due to the relatively weak upper palatal plica. Umbilicus very narrow, measuring ~ 1 / 10 of the shell width. The umbilical groove is present running alongside it. This groove is the broadest near the peristome and regularly tapering towards the umbilicus, ending as a narrow canal at the last ~ ¼ of the body whorl.</p><p>Differential diagnosis.</p><p>This species is the most similar to A. picasso sp. nov. and for differences, see under that species. The new species also bears some resemblance to Aulacospira pluangtong Panha &amp; Burch, 2004, but the latter has less rounded whorls, more elongated umbilicus, smooth apertural barriers (not spiniferous) and it lacks the basal plica.</p><p>Measurements</p><p>(in mm, n = 5). SW = 2.10–2.58; SH = 2.39–2.82; AH = 1.10–1.22; AW = 1.09–1.28.</p><p>Etymology.</p><p>The Latin word evanidus means vanishing, which refers to the quarrying of the type locality of this species.</p><p>Distribution.</p><p>This species is known only from the type locality.</p><p>Remarks.</p><p>Unlike congeners (except for A. kozari), this species has rough, spiniferous surface of the apertural barriers. It is retained in Anauchen because of the similar barrier arrangement to other species (e. g., A. angthongensis, A. crassus sp. nov., A. huaykhakang, A. obesus sp. nov.) and a single lamella on the parietal side.</p></div>	https://treatment.plazi.org/id/B2B8499F32AE582CAB4D56CB0486E448	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
29EF10771C8F5F4A8D1DC9331002CA1F.text	29EF10771C8F5F4A8D1DC9331002CA1F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anauchen grandiportus Gojsina, Grego & Pall-Gergely 2025	<div><p>Anauchen grandiportus Gojšina, Grego &amp; Páll-Gergely sp. nov.</p><p>Figs 10 F, 18, 37</p><p>Type material.</p><p>Holotype. Laos • 1 empty shell (SH: 1.71 mm, SW: 1.92 mm); Khammouane Province, NE foot of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.87168&amp;materialsCitation.latitude=17.5518" title="Search Plazi for locations around (long 104.87168/lat 17.5518)">Mount Pha Soung</a>, caverns among slope boulders; 17°33.108'N, 104°52.301'E; 08 Feb. 2017; J. Grego leg.; CUMZ 14428.</p><p>Type locality.</p><p>Laos, Khammouane Province, NE foot of Mount Pha Soung, caverns among slope boulders; 17°33.108'N, 104°52.301'E.</p><p>Diagnosis.</p><p>Shell conical-ovoid, teleoconch radially and not spirally striated. Last whorl not detached from the penultimate and slightly descending. Aperture large, with seven weak barriers sitting on the peristome edge. Umbilicus narrow.</p><p>Description.</p><p>Shell brown, conical-ovoid with deep suture and bulging whorls. Whorls 3.25, rounded, regularly increasing. Protoconch consists of slightly&gt; 1 whorl, finely pitted, its terminal part with ~ 12 equidistant spiral striae. Teleoconch with inconspicuous, irregular growth lines and finely dimpled (pasty), spiral striation absent. Last whorl rounded, adnate to the penultimate and slightly descending near the aperture (~ 15 ° compared to the shell axis), aperture slightly prosocline to shell axis in lateral view. Aperture conspicuously large to the shell size, parietal part adnate onto penultimate whorl forming a weak callus. Peristome slightly expanded but not reflected. Altogether seven apertural barriers could be found, all being blunt, knob-like, situated on peristome edge, their homologies with traditionally recognised barriers somewhat questionable. Parietal lamella is the largest, but still weak. Suprapalatal and upper palatal plicae situated close to each other, after some distance the next plica is the lower palatal. Lower palatal, basal, subcolumellar, and columellar situated in approximately equal distance from one another, gap between columellar and parietal is approximately as large as between parietal and the suprapalatal. There is a very slight thickening next to lower palatal plica, although it is questionable whether it would develop into a barrier. Surfaces of all apertural barriers are very finely granulated. Sinulus wide but low, not strongly isolated from the rest of the aperture. Umbilicus narrow, elongate, only shows body whorl.</p><p>Differential diagnosis.</p><p>Anauchen grandiportus sp. nov. differs from all other species assigned to Anauchen on the basis of the seven small, marginal denticles and the greatly enlarged aperture.</p><p>Measurements</p><p>(in mm, n = 1). SH = 1.92; SW = 1.71, AH = 0.96, AW = 0.87.</p><p>Etymology.</p><p>The specific epithet refers to the relatively large aperture when compared to the shell size.</p><p>Distribution.</p><p>This species is known only from the type locality.</p></div>	https://treatment.plazi.org/id/29EF10771C8F5F4A8D1DC9331002CA1F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
4805BBACC57E500E8508E7C56A4494C2.text	4805BBACC57E500E8508E7C56A4494C2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anauchen huaykhakang Panha 2002	<div><p>Anauchen huaykhakang Panha, 2002</p><p>Figs 10 G, 19, 37</p><p>Anauchen huaykhakang Panha in Burch &amp; Panha, 2002: 239–244, fig. 2.</p><p>Anauchen huaykhakang — Panha and Burch 2008: 51–52, fig. 47; Dumrongrojwattana and Wongkamhaeng 2024: 323, fig. 7.</p><p>Type material examined.</p><p>Thailand • 1 paratype; from the type locality; 1997; S. Panha leg.; CUMZ ver. 038 .</p><p>Type locality.</p><p>“ Subkao, Huaykhakang Wildlife Sanctuary, Lansak District, Utaithani Province, 15°35'7"N, 99°18'8"E, 310 meters elevation … ” (Thailand).</p><p>Differential diagnosis.</p><p>This species differs from A. angthongensis by the shouldered last whorl and concave-conical shell (due to the strongly enlarged last whorl). Anauchen banmiensis is much more strongly spirally striated, it last whorl is more distinctly shouldered and has a wider umbilicus with a deep groove inside it (absent in A. huaykhakang). For differences from A. utaithaniensis, see under that species.</p><p>Distribution.</p><p>This species is known only from the type locality.</p></div>	https://treatment.plazi.org/id/4805BBACC57E500E8508E7C56A4494C2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
E7E18EFE1D36515C9F434D1716E4365F.text	E7E18EFE1D36515C9F434D1716E4365F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anauchen informis subsp. informis Vermeulen, Phung & Truong 2007	<div><p>Anauchen informis informis Vermeulen, Phung &amp; Truong, 2007</p><p>Figs 10 H, 20, 37</p><p>Anauchen informis informis Vermeulen, Phung &amp; Truong, 2007: 87–89, fig. 6.</p><p>Anauchen informis informis — Schileyko 2011: 4.</p><p>Type material examined.</p><p>Vietnam • holotype; RMNH 108986 .</p><p>Type locality.</p><p>Vietnam, Kien Giang Prov., Kien Luong, Hon Chong hill, doline along East flank.</p><p>Differential diagnosis.</p><p>This species is the most similar to A. angthongensis from which it differs by its more bulging whorls and more ovoid shell. A. banmiensis has a shouldered last whorl and stronger barriers.</p><p>Distribution.</p><p>This subspecies is known only from the type locality.</p></div>	https://treatment.plazi.org/id/E7E18EFE1D36515C9F434D1716E4365F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
EEEA001CBB8D5A5B8BEDCFADE9E22A40.text	EEEA001CBB8D5A5B8BEDCFADE9E22A40.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anauchen informis subsp. parcedentata Vermeulen, Phung & Truong 2007	<div><p>Anauchen informis parcedentata Vermeulen, Phung &amp; Truong, 2007</p><p>Figs 10 I, 21, 37</p><p>Anauchen informis parcedentata Vermeulen, Phung &amp; Truong, 2007: 89, fig. 7.</p><p>Anauchen informis parcedentata — Schileyko 2011: 4.</p><p>Type material examined.</p><p>Vietnam • holotype; RMNH 108987 .</p><p>Type locality.</p><p>“ Vietnam, Kien Giang prov., Kien Luong, Ba Tai hill ”.</p><p>Differential diagnosis.</p><p>Similar to the nominotypical subspecies but with 1–3 barriers in the aperture (parietal always present, both palatals occasionally) and usually more whorls.</p><p>Distribution.</p><p>This subspecies is known only from the type locality.</p></div>	https://treatment.plazi.org/id/EEEA001CBB8D5A5B8BEDCFADE9E22A40	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
2921D2B84C545541B00129D2DF03FB65.text	2921D2B84C545541B00129D2DF03FB65.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anauchen jokaii Gojsina & Pall-Gergely 2025	<div><p>Anauchen jokaii Gojšina &amp; Páll-Gergely sp. nov.</p><p>Figs 10 J, 22, 23, 37</p><p>Type material.</p><p>Holotype. Vietnam • 1 shell (SH: 2.48 mm, SW: 2.31 mm); Nam Pia village, Hung Ba commune, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.08023&amp;materialsCitation.latitude=22.867733" title="Search Plazi for locations around (long 105.08023/lat 22.867733)">Vi Xugen district</a>, Ha Giang Province; 22°52.064'N, 105°4.814'E; 216 m a. s. l.; L. V. Hao, T. Ishibe, Y. Nakahara, K. Ohara, K. Okubo, J. U. Otani, V. P. Sang leg.; IEBR _LS_Anauchen.002 H.</p><p>Type locality.</p><p>Vietnam, Nam Pia village, Hung Ba commune, Vi Xugen district, Ha Giang Province; 22°52.064'N, 105°4.814'E; 216 m a. s. l.</p><p>Diagnosis.</p><p>Anauchen species with conical shell, last whorl keeled at its base and slightly ascending near the aperture. Spiral striation absent. Aperture armed with four strong barriers, umbilicus narrow.</p><p>Description.</p><p>Shell conical, consisting of 5.5 regularly increasing, convex whorls separated by a deep suture. Colour light brown. There is not a clear boundary between the protoconch and the teleoconch due to the similar surface sculpture. Teleoconch pasty, very finely but irregularly radially striated, with no signs of spiral striation. Last whorl adnate to the penultimate and its last quarter slightly ascending near the aperture (~ 10 ° compared to the shell axis), making the aperture profile opisthocline to the shell axis. Last whorl with a blunt keel at its base, making the shell strongly triangular. Peristome continuous and thick, not very strongly expanded on the parietal side and not fully leaning on the penultimate whorl. It is slightly lighter in colour than the rest of the shell, not reflected. Sinulus rounded and well separated from the rest of the aperture. Aperture equipped with four strong but short barriers (parietal, upper palatal, lower palatal and columellar). Parietal lamella is the strongest in the aperture, blade-like. Upper palatal plica ~ 1 / 2 as strong as the parietal lamella and very slightly leaning towards the lower palatal plica. The latter is strong (only slightly weaker than the parietal) and moderately long. There is no basal plica. Columellar lamella thick and relatively low, situated in the middle of the columellar wall. All apertural barriers almost reach the expanding peristome. Surface of all apertural barriers is smooth. Umbilicus rounded, open but narrow, measuring ~ 1 / 11 of the shell width.</p><p>Differential diagnosis.</p><p>See under A. messageri, A. rochebruni, A. pentadens and A. turritus sp. nov.</p><p>Measurements</p><p>(in mm, n = 1). SH = 2.48; SW = 2.31; AH = 1.12; AW = 1.04.</p><p>Etymology.</p><p>This species is named after and dedicated to Mór Jókai (1825–1904), Hungarian novelist, dramatist, to commemorate the bicentenary of his birth. Jókai was interested in malacology, he had a large collection of (mostly marine) shells, and even published a satirical novel entitled “ Novel of snails ” (A csigák regénye).</p><p>Distribution.</p><p>Only known from the type locality.</p></div>	https://treatment.plazi.org/id/2921D2B84C545541B00129D2DF03FB65	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
54F9A976131F5612ADA27504B83BAF42.text	54F9A976131F5612ADA27504B83BAF42.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anauchen kozari Pall-Gergely 2023	<div><p>Anauchen kozari Páll-Gergely, 2023</p><p>Figs 10 K, 24, 37</p><p>Anauchen kozari Páll-Gergely, 2023 a: 64–66, figs 1 D, F, 2, 3.</p><p>Type material examined.</p><p>Laos • holotype; Luang Prabang Province, Phou Xuang Mountain, ca 1.5 km NE of Ban Lak Sip, ca 5 km SE of Luang Prabang, under rocks and logs in old secondary forest under cliff; 19 ° 851.6050 ' N, 10 ° 2811.0810 ' E; 640 m a. s. l.; A. Abdou, I. V Muratov leg.; 24 Nov. 2006; MNHN -IM-2012-27269 .</p><p>Type locality.</p><p>“ Central Laos, Luang Prabang Province, Phou Xuang mountain, ca 1.5 km NE of Ban Lak Sip, ca 5 km SE of Luang Prabang, under rocks and logs in old secondary forest under cliff, 19 ° 851.6050 ' N, 10 ° 2811.0810 ' E, alt. 640 m ”.</p><p>Differential diagnosis.</p><p>See under A. messageri .</p><p>Distribution.</p><p>This species is known only from the type locality.</p><p>Remarks.</p><p>This species is not a typical representative of its genus since it has a palatal tubercle frequently found in Bensonella and a granulated, rough surface of the apertural barriers frequently found in Hypselostoma . It is also the smallest of all Anauchen species. It is however retained in Anauchen for the following reasons: i) the arrangement of apertural barriers is highly reminiscent to those of A. messageri (see Páll-Gergely 2023 a); ii) there is a single lamella on the parietal side; iii) last whorl adnate to the penultimate.</p></div>	https://treatment.plazi.org/id/54F9A976131F5612ADA27504B83BAF42	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
AE544CE2E66A57289A1D68A7773350CB.text	AE544CE2E66A57289A1D68A7773350CB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anauchen messageri (Bavay & Dautzenberg 1904)	<div><p>Anauchen messageri (Bavay &amp; Dautzenberg, 1904)</p><p>Boysidia messageri Bavay &amp; Dautzenberg, 1904: 211, pl. 9, figs 1–4.</p><p>Anauchen messageri — Pilsbry 1917: 189, pl. 33, figs 4–7; Schileyko 2011: 5; Páll-Gergely 2023 a: 67, fig. 1 C.</p><p>Anauchen massageri [sic!] — Burch and Panha 2002: 242, 245, 247.</p><p>Anauchen whitteni Chen, 2023: 90–91, figs 1, 2. syn. nov.</p><p>Type material examined.</p><p>Vietnam • 1 syntype of A. messageri; from the type locality; C. Messager leg.; MNHN -IM-2000-35159 .</p><p>Additional material examined.</p><p>Vietnam • 2 shells; Cao Bang Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.33888&amp;materialsCitation.latitude=22.733017" title="Search Plazi for locations around (long 106.33888/lat 22.733017)">Đeo Ma Phuc - Quang Uyen</a>, left side of the road, rock crevice; 22°43.981'N, 106°20.333'E; 610 m a. s. l.; 14 Nov. 2011; A. Hunyadi leg.; coll. HA • 1 shell; Cao Bang Province, west of Quang Uyen, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.4393&amp;materialsCitation.latitude=22.703133" title="Search Plazi for locations around (long 106.4393/lat 22.703133)">Phi Hao-Đau Tuyen</a>; 22°42.188'N, 106°26.358'E; 500 m a. s. l.; 16. Nov 2011.; A. Hunyadi leg.; coll. HA • 8 shells; Lang Son Province, Huu Lung district, Minh Tien, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.2965&amp;materialsCitation.latitude=21.567066" title="Search Plazi for locations around (long 106.2965/lat 21.567066)">1400 m northeast of Cau Cheo Minh Tien</a>; 21°34.024'N, 106°17.790'E; 30 m a. s. l.; 20 Feb. 2020; A. Hunyadi leg.; coll. HA • 13 shells; Lang Son Province, Huu Lung district, Huu Lien, 33.5 km from junction of roads 1 B and 241 towards <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.38118&amp;materialsCitation.latitude=21.6844" title="Search Plazi for locations around (long 106.38118/lat 21.6844)">Ba Nang</a>, left side of the road; 21°41.064'N, 106°22.871'E; 220 m a. s. l.; 19 Feb. 2020; A. Hunyadi leg; coll. HA • 47 shells; Hoa Binh Province, Kim Boi district, Cao Duong, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.65498&amp;materialsCitation.latitude=20.70985" title="Search Plazi for locations around (long 105.65498/lat 20.70985)">north of Đong Phu, 58 km from Nho Quan towards Hanoi on the Ho Chi Minh road</a>; 20°42.591'N, 105°39.299'E; 10 m a. s. l.; 16 Feb. 2020; A. Hunyadi, H. V. Luong, J. U. Otani &amp; S. V. Pham leg.; coll. HA • 1 shell; Thanh Hoa Province, Xa Ban Cong, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.21187&amp;materialsCitation.latitude=20.381617" title="Search Plazi for locations around (long 105.21187/lat 20.381617)">Xom Sat</a>; 20°22.897'N, 105°12.712'E; 60 m a. s. l.; 18. May 2012 ¸ A. Hunyadi leg.; coll. HA • 15 shells; Thanh Hoa Province, 9.8 km from centre of Ngoc Lac towards Lang Chanh, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.30462&amp;materialsCitation.latitude=20.119867" title="Search Plazi for locations around (long 105.30462/lat 20.119867)">Ngoc Khe</a>, left side of the road, 115 km; 20°7.192'N, 105°18.277'E; 13 Feb. 2020; A. Hunyadi, H. V. Luong, J. U. Otani &amp; S. V. Pham leg.; coll. HA . China • 9 shells; Guangxi, Bose Shi, Leye Xian, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.72075&amp;materialsCitation.latitude=24.657267" title="Search Plazi for locations around (long 106.72075/lat 24.657267)">Buliu He</a>; 24°39.436'N, 106°43.245'E; 540 m a. s. l.; 08 Sept. 2013; A. Hunyadi leg.; coll. HA • 74 shells; Guangxi, Hechi Shi, Huanjiang Maonanzu Zizhixian, south of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.96065&amp;materialsCitation.latitude=25.0995" title="Search Plazi for locations around (long 107.96065/lat 25.0995)">Mulun Guojiaji Ziran Baohuqu</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.96065&amp;materialsCitation.latitude=25.0995" title="Search Plazi for locations around (long 107.96065/lat 25.0995)">Dongning</a>; 25°05.970'N, 107°57.639'E; 530 m a. s. l.; 17 Sept. 2013, A. Hunyadi leg.; coll. HA • 3 shells; Guangxi, Laibin Shi, Xingbin Qu, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=109.071465&amp;materialsCitation.latitude=24.008533" title="Search Plazi for locations around (long 109.071465/lat 24.008533)">Qidong Xiang</a>; 24°00.512'N, 109°04.288'E; 150 m a. s. l.; 20 Sept. 2013; A. Hunyadi leg. • 2 shells; coll. HA • 2 shells; Guangxi, Hechi Shi, Du an <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.0215&amp;materialsCitation.latitude=24.053284" title="Search Plazi for locations around (long 108.0215/lat 24.053284)">Yaozu Zizhixian</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.0215&amp;materialsCitation.latitude=24.053284" title="Search Plazi for locations around (long 108.0215/lat 24.053284)">Gaoling Xiang</a>, 2 km west of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.0215&amp;materialsCitation.latitude=24.053284" title="Search Plazi for locations around (long 108.0215/lat 24.053284)">Dingfucun</a>; 24°03.197'N, 108°01.290'E; 320 m a. s. l.; 08 Oct. 2009; A. Hunyadi leg.; coll. HA • 171 shells; Guangxi, Hechi Shi, Bama Yaozu Zizhixian, southern edge of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.13078&amp;materialsCitation.latitude=24.117416" title="Search Plazi for locations around (long 107.13078/lat 24.117416)">Jiaolecun</a>; 24°07.045'N, 107°07.847'E; 590 m a. s. l.; 10 Sept. 2013; A. Hunyadi leg.; coll. HA • 24 shells; Guangxi, Hechi Shi, Nandan Xian, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.602936&amp;materialsCitation.latitude=25.069384" title="Search Plazi for locations around (long 107.602936/lat 25.069384)">Lihuyaozu Xiang</a>, southeast of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.602936&amp;materialsCitation.latitude=25.069384" title="Search Plazi for locations around (long 107.602936/lat 25.069384)">Encun</a>, rocks above the road; 25°04.163'N, 107°36.176'E; 630 m a. s. l.; 11 Sept. 2013; A. Hunyadi leg.; coll. HA • 10 shells; Guangxi, Guilin Shi, Guilin, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=110.264366&amp;materialsCitation.latitude=25.307766" title="Search Plazi for locations around (long 110.264366/lat 25.307766)">Ludiyan Gongyuan</a>, northeastern side of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=110.264366&amp;materialsCitation.latitude=25.307766" title="Search Plazi for locations around (long 110.264366/lat 25.307766)">Moupanshan</a>; 25°18.466'N, 110°15.862'E 145 m a. s. l.; 22 Sept. 2013; A. Hunyadi leg.; coll. HA • 14 shells; Guangxi, Chongzuo Shi, Longzhou Xian, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.768265&amp;materialsCitation.latitude=22.587317" title="Search Plazi for locations around (long 106.768265/lat 22.587317)">Wude Xiang</a>, vicinity of the junction towards <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.768265&amp;materialsCitation.latitude=22.587317" title="Search Plazi for locations around (long 106.768265/lat 22.587317)">Banxintun</a>; 22°35.239'N, 106°46.096'E; 350 m a. s. l.; 24 Sept. 2013; A. Hunyadi leg.; coll. HA • 257 shells; Guangxi, Hechi Shi, Tiane Xian, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.1945&amp;materialsCitation.latitude=24.852167" title="Search Plazi for locations around (long 107.1945/lat 24.852167)">Qimu Xiang</a>, junction towards <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.1945&amp;materialsCitation.latitude=24.852167" title="Search Plazi for locations around (long 107.1945/lat 24.852167)">Lahaiyan</a>, rock above the tomb; 24°51.130'N, 107°11.670'E; 600 m a. s. l.; 12 Sept. 2013; A. Hunyadi &amp; M. Szekeres leg.; coll. HA • 5 shells; Guangxi, Guilin Shi, Yangshuo Xian, 5 km north of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=110.53095&amp;materialsCitation.latitude=24.955967" title="Search Plazi for locations around (long 110.53095/lat 24.955967)">Xingpingzhen</a>; 24°57.358'N, 110°31.857'E; 150 m a. s. l.; 15 Oct. 2009; A. Hunyadi leg.; coll. HA • 13 shells; Yueliangshan, Gaotianzhen, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=110.472&amp;materialsCitation.latitude=24.724167" title="Search Plazi for locations around (long 110.472/lat 24.724167)">Yangshuoxian</a>, Guangxi <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=110.472&amp;materialsCitation.latitude=24.724167" title="Search Plazi for locations around (long 110.472/lat 24.724167)">Zhuangzu Zizhiqu</a>; 24°43.45'N, 110°28.32'E; K. Ohara, K. Okubo, J. U. Otani leg.; coll. PGB • 17 shells; Daxiaojingfengjingqu, Moyangzhen, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.85722&amp;materialsCitation.latitude=25.562166" title="Search Plazi for locations around (long 106.85722/lat 25.562166)">Luodianxian</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.85722&amp;materialsCitation.latitude=25.562166" title="Search Plazi for locations around (long 106.85722/lat 25.562166)">Guizhousheng</a>; 25°33.73'N, 106°51.4333'E; 450 m a. s. l.; T. Ishibe, K. Okubo, J. U. Otani leg.; coll. PGB • 4 shells; near Hudiequan bridge, Gaotianzhen, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=110.49367&amp;materialsCitation.latitude=24.740833" title="Search Plazi for locations around (long 110.49367/lat 24.740833)">Yangshuoxian</a>, Guangxi <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=110.49367&amp;materialsCitation.latitude=24.740833" title="Search Plazi for locations around (long 110.49367/lat 24.740833)">Zhuangzu Zizhiqu</a>; 24°44.45'N, 110°29.62'E; K. Ohara, K. Okubo, J. U. Otani leg.; coll. PGB .</p><p>Type localities.</p><p>“ Haut-Tonkin ” (Vietnam) ( A. messageri); “ Collected from the rocks, Mt. Nanshe [南蛇山], Yizhou District [宜州区], Hechi City [河池市], Guangxi Zhuang Autonomous Region, China. 24°30'22"N, 108°38'51"E, 270 m … ” ( A. whitteni).</p><p>Differential diagnosis.</p><p>Anauchen jokaii sp. nov. lacks the spiral striation and has its last whorl keeled at the base which separates it from A. messageri . Anauchen kozari is smaller, more depressed and has a palatal tubercle. See also under A. pentadens, A. rochebruni and A. turritus sp. nov.</p><p>Distribution.</p><p>This species has a wide distribution across Vietnam and China. Its northernmost known localities are situated in northern parts of Guangxi province (China) while the southernmost are from Thanh Hoa province (Vietnam).</p><p>Remarks.</p><p>We have observed high morphological variability within this species, particularly in the slenderness of the shell (Fig. 25). The basal plica is also occasionally present or sometimes absent. Anauchen whitteni Z. - Y. Chen, 2023 was described as a distinct species based on the absence of the basal plica. Since the basal plica was found here as a variable character and the species does not differ from A. messageri in any other characters, it is treated here as a junior synonym.</p></div>	https://treatment.plazi.org/id/AE544CE2E66A57289A1D68A7773350CB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
29907147B2E2503C87BEAA83B7395662.text	29907147B2E2503C87BEAA83B7395662.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anauchen obesus Gojsina, Hunyadi & Pall-Gergely 2025	<div><p>Anauchen obesus Gojšina, Hunyadi &amp; Páll-Gergely sp. nov.</p><p>Figs 10 M, 26, 27, 37</p><p>Type material.</p><p>Holotype. Thailand • 1 shell (SH: 2.94 mm, SW: 2.26 mm); Chiang Rai Province, Doi Tung, 50 metres before <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.833115&amp;materialsCitation.latitude=20.325666" title="Search Plazi for locations around (long 99.833115/lat 20.325666)">Wat Phra That Doi Tung</a>, vicinity of the parking lot; 20°19.540'N, 99°49.987'E; 1350 m; 12 Feb. 2015; A. Hunyadi leg.; CUMZ 14429 . Paratypes. Thailand • 15 shells; same data as for holotype; coll. HA • 10 shells; Chiang Rai Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.83917&amp;materialsCitation.latitude=20.342222" title="Search Plazi for locations around (long 99.83917/lat 20.342222)">Doi Tung</a>; 20°20′32″N, 99°50′21″E; 1320 m a. s. l.; 08. May 1988; F. G. Thompson leg.; locality code FGT-4417; UF 591335 .</p><p>Additional material examined.</p><p>Thailand • 1 shell (juvenile, not paratype); Chiang Rai Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.83917&amp;materialsCitation.latitude=20.342222" title="Search Plazi for locations around (long 99.83917/lat 20.342222)">Doi Tung</a>; 20°20′32″N, 99°50′21″E; 1320 m a. s. l.; 08. May 1988; F. G. Thompson leg.; locality code FGT-4417; UF 583720 .</p><p>Type locality.</p><p>Thailand, Chiang Rai Province, Doi Tung, 50 metres before Wat Phra That Doi Tung, vicinity of the parking lot; 20°19.540'N, 99°49.987'E; 1350 m.</p><p>Diagnosis.</p><p>Anauchen species with conical-ovoid, brown shell, pasty-like surface, last whorl adnate to the penultimate, five apertural barriers (parietal (angulo-parietal), upper palatal, lower palatal, weak basal, and columellar) and a narrow umbilicus.</p><p>Description.</p><p>Shell conical-ovoid, brown, with 4.5–5 whorls separated by a deep suture. All whorls convex, bulging, rounded. Protoconch consists of ~ 1.5 whorls, finely pitted and with moderate spiral striae, the same colour as the rest of the shell. Teleoconch fine, pasty, with inconspicuous, irregular growth radial lines which are rarely more pronounced and in form of whitish streaks, most dense right behind the peristome. Spiral striation absent. Last whorl adnate to the penultimate and slightly descending (~ 25 ° compared to the shell axis) making the aperture profile prosocline to the shell axis. Peristome continuous, in form of a short and not much expanded callus leaning on the penultimate whorl at the parietal side. It is white, expanded but not reflected. Aperture equipped with five barriers (parietal (angulo-parietal), upper palatal, lower palatal, basal and columellar). Parietal (angulo-parietal) lamella almost fully reaching the expanded peristome edge. It is the largest in the aperture, sometimes single and sometimes with an additional smaller part, which is probably homologous with the angular lamella in other genera, pointed towards the palatal side. Upper palatal plica is weaker, lower, and less slender than the lower palatal. Lower palatal plica almost as strong as the angulo-parietal lamella. Basal plica in the form of inconspicuous knob, like a very slight swelling in the basal-columellar transition area. Columellar lamella almost reaching the expanded peristome, highest in its middle part and sloping towards the peristome edge where it becomes quite low. It is leaned towards the lower palatal plica. Surface of all apertural barriers is very finely granulated. Sinulus rounded, parabolic and well separated from the rest of the aperture. Umbilicus narrow, ~ 1 / 8 of the shell width, previous whorls not clearly visible through it. Umbilical groove absent.</p><p>Differential diagnosis.</p><p>Anauchen obesus sp. nov. is similar in shell shape to A. angthongensis, A. crassus sp. nov., A. eotvosi and A. informis . However, A. obesus sp. nov. can be separated by its smaller size and more globular shell.</p><p>Measurements</p><p>(in mm, n = 4). SH = 2.71–3; SW = 1.95–2.26; AH = 1.1–1.27; AW = 1.06–1.23.</p><p>Etymology.</p><p>This species is named for its globose shell.</p><p>Distribution.</p><p>This species is known only from Doi Tung mountain.</p></div>	https://treatment.plazi.org/id/29907147B2E2503C87BEAA83B7395662	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
6B2E5B70A63C58D9A55D6C11D0619540.text	6B2E5B70A63C58D9A55D6C11D0619540.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anauchen pentadens (D. - N. Chen, M. Wu & G. - Q. Zhang 1999)	<div><p>Anauchen pentadens (D. - N. Chen, M. Wu &amp; G. - Q. Zhang, 1999)</p><p>Fig. 37</p><p>Boysidia (Boysidia) pentadens Chen, Wu &amp; Zhang, 1999: 5 (Chinese description), 11 (English description), fig. 2. — Chen and Zhang 2002: 692.</p><p>Boysidia pentadens — Qian and Zhou 2014: 90–91; Chen et al. 2016: 50; Dumrongrojwattana and Assawawattagee 2018: 216.</p><p>Anauchen pentadens — Chen 2023: 89–90.</p><p>Material examined.</p><p>None.</p><p>Type locality.</p><p>“ collected from Menglun Town (21°5'N, 100°6'E). Mengla County, Yunnan Province ” (China).</p><p>Differential diagnosis.</p><p>This species differs from A. messageri by its less triangular shell, narrower umbilicus, absence of spiral striation, and presence of strong radial ribs on the last whorl (especially near the aperture). Anauchen pentadens differs from A. jokaii sp. nov. by its wider umbilicus, strong ribs on the last whorl, especially near the aperture, and a basal plica. Its last whorl is also regularly rounded, so the is shell less triangular. See also under A. rochebruni .</p><p>Distribution.</p><p>This species is known only from Yunnan Province, China.</p><p>Remarks.</p><p>According to the figures presented in Qian and Zhou (2014) and Chen et al. (2016), Chen (2023) considered it a member of Anauchen rather than Boysidia which we follow herein.</p></div>	https://treatment.plazi.org/id/6B2E5B70A63C58D9A55D6C11D0619540	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
67988AA56FEB55D088F1104C261F45B0.text	67988AA56FEB55D088F1104C261F45B0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anauchen picasso Gojsina & Pall-Gergely 2025	<div><p>Anauchen picasso Gojšina &amp; Páll-Gergely sp. nov.</p><p>Figs 10 N, 28, 29, 37</p><p>Type material.</p><p>Holotype. Thailand • 1 shell (SH: 2.95 mm, SW: 2.79 mm); Prachuap Khiri Khan Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.993&amp;materialsCitation.latitude=12.208" title="Search Plazi for locations around (long 99.993/lat 12.208)">Khao Sam Roi Yot National Park</a>, 3 km S of Park Entrance; 12.208°N, 99.993°E; 31 May 1987; F. G. Thompson leg.; UF 529706 . Paratypes. Thailand • 1 shell; same data as for holotype; CUMZ 14430; 8 shells; same data as for holotype; UF 591339 .</p><p>Additional material examined.</p><p>Thailand • 1 shell (juvenile / broken, not paratype); same data as for holotype; UF 583731 .</p><p>Type locality.</p><p>Thailand, Prachuap Khiri Khan Province, Khao Sam Roi Yot National Park, 3 km S of Park Entrance; 12.208°N, 99.993°E.</p><p>Diagnosis.</p><p>Anauchen species with distinctly keeled last whorl (slightly below the centre of the periphery) which is also detached near the aperture and descending. Teleoconch spirally striated. Only four weak apertural barriers (parietal, upper palatal, lower palatal and columellar).</p><p>Description.</p><p>Shell conical or very slightly conical-ovoid, brown, consisting of 4.5–5 whorls separated by a deep suture. Protoconch not much lighter than the rest of the shell, roughly pitted and terminally spirally striated, consisting of 1.25 whorls. Initial teleoconch whorls and the penultimate whorl slightly keeled. Fine spiral striation visible on the teleoconch as well, sometimes stronger near the protoconch (on initial teleoconch whorls), sometimes stronger on the last whorl. There are several irregularly spaced whitish radial streaks crossing the spiral striae. Last whorl enlarged and distinctly keeled slightly below the centre of the periphery. The keel is equally strong along its length but is getting lost right behind the peristome. Last whorl slightly detached from the penultimate near the aperture and slightly descending (~ 15 ° compared to the shell axis), thus making the aperture profile prosocline to the shell axis. Peristome is dirty white, very thick, expanded but not reflected. Aperture rounded, almost as high as wide. There are only four relatively weak apertural barriers (parietal, upper palatal, lower palatal and columellar). Of them, parietal lamella is the strongest and highest and reaching the expanding peristome. Both palatal plicae (upper and lower) are of similar appearance (but lower is usually slightly stronger than the upper), both weak (low and short) and tubercle-like. They are positioned close to each other, probably closer than in any other congener. Columellar lamella is weak and dot-like, usually not present as a distinct barrier but rather as a columellar thickening which is gradually lowering towards the expanding peristome. Surface of all apertural barriers is very finely granulated. Sinulus wide and not strongly isolated from the rest of the aperture due to the relatively weak upper palatal plica. Umbilicus very narrow, elongated and measuring 1 / 8–1 / 9 of the shell width. There is a strong umbilical groove running along the umbilicus. This groove is very wide near the peristome and then regularly tapering towards the umbilicus, finally terminating at the ~ ½ of the last whorl.</p><p>Differential diagnosis.</p><p>This species is different from other congeners by the strongly keeled last whorl below the centre of the periphery and the presence of four apertural barriers which are quite weak. It is most similar to A. evanidus sp. nov. but A. picasso sp. nov. has a more strongly keeled last whorl, weaker apertural barriers, and is spirally striated.</p><p>Measurements</p><p>(in mm, n = 5). SH = 2.64–2.95; SW = 2.63–2.82; AH = 1.23–1.38; AW = 1.27–1.37.</p><p>Etymology.</p><p>This species looks like an Anauchen with rounded whorls painted in a Pablo Picasso style (i. e., resembling the art style known as Cubism). The specific epithet is to be used as a noun in apposition.</p><p>Distribution.</p><p>This species is known only from the type locality.</p></div>	https://treatment.plazi.org/id/67988AA56FEB55D088F1104C261F45B0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
5A58BAED0891523EB1CA23BA19BFFE3B.text	5A58BAED0891523EB1CA23BA19BFFE3B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anauchen Pilsbry 1917	<div><p>Genus Anauchen Pilsbry, 1917</p><p>Anauchen Pilsbry, 1917: 188 .</p><p>Diagnosis.</p><p>This genus is characterised by the combination of characters: i) a single lamella which is formed by a fusion of angular and parietal lamellae (it can appear bifid in some species, see below); ii) last whorl adnate to, or only very slightly detached from the penultimate; iii) shell surface pasty (dimpled) or finely spirally striated (never rough, sandpaper-like surface); iv) surface of apertural barriers smooth.</p><p>Remarks.</p><p>The following species possess some exceptions from the aforementioned combination of diagnostic characters for the genus: Anauchen kozari and Anauchen evanidus sp. nov. Both of them have rough (granulated) surfaces of the apertural barriers. Furthermore, A. kozari is much smaller (height 1.25 mm, width 1.3 mm) than all other congeners and has a palatal tubercle (typical for many Bensonella species). However, both species are retained in Anauchen for reasons listed under “ Remarks ” section for these species.</p></div>	https://treatment.plazi.org/id/5A58BAED0891523EB1CA23BA19BFFE3B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
C85319C0F7BE53FBBF7E1987BC4A0D43.text	C85319C0F7BE53FBBF7E1987BC4A0D43.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anauchen rochebruni (Mabille 1887) Pall-Gergely 2023	<div><p>Anauchen rochebruni (Mabille, 1887)</p><p>Figs 10 O, 30, 31, 37, 38</p><p>Hypselostoma rochebruni Mabille, 1887 a: 8 .</p><p>Pupa angulina Gredler, 1885: 7 . syn. nov.</p><p>Hypselostoma rochebruni — Mabille 1887 b: 121–122.</p><p>Boysidia gereti Bavay &amp; Dautzenberg, 1904: 212, pl. 9, figs 5–8.</p><p>Hypselostoma rochebrunei [sic] — Fischer and Dautzenberg 1904: 408.</p><p>Anauchen angulinus — Pilsbry 1917: 191, pl. 33, figs 5, 6.</p><p>Anauchen rochebruni — Pilsbry 1917: 190–191; Páll-Gergely 2023 a: 63, 67, figs 1 A, B.</p><p>Anauchen gereti — Pilsbry 1917: 189–190, pl. 33, figs 1–3; Schileyko 1998: 138, fig. 158; Schileyko 2011: 4.</p><p>Hypselostoma rochebruni — Schileyko 2011: 3.</p><p>Boysidia gereti — Páll-Gergely 2023 a: 67, fig. 1 A.</p><p>Type material examined.</p><p>Vietnam • 1 syntype of A. rochebruni; from the type locality; collector unknown; MNHN -IM-2000-35156 • 1 syntype of B. gereti; from the type locality; C. Messager leg.; MNHN -IM-2000-35161 .</p><p>Additional material examined.</p><p>Vietnam • 59 shells; Lang Son Province, Binh Gia, eastern edge of the village, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.39698&amp;materialsCitation.latitude=21.938366" title="Search Plazi for locations around (long 106.39698/lat 21.938366)">Di Chi Keo Leng</a>, rock wall; 21°56.302'N, 106°23.819'E; 375 m a. s. l.; 17 Feb. 2020; A. Hunyadi leg.; coll. HA • 55 shells; Lang Son Province, Bac Son district, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.34907&amp;materialsCitation.latitude=21.890182" title="Search Plazi for locations around (long 106.34907/lat 21.890182)">Quinh Son</a>, northern part of the village; 21°53.411'N, 106°20.944'E; 400 m a. s. l.; 18 Feb. 2020; A. Hunyadi leg.; coll. HA • 31 shells; Lang Son Province, Bac Son district, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.34237&amp;materialsCitation.latitude=21.910267" title="Search Plazi for locations around (long 106.34237/lat 21.910267)">Quinh Son</a>, northeastern part of the village, rock wall; 21°54.616'N, 106°20.542'E; 420 m a. s. l.; 18 Feb. 2020; A. Hunyadi leg.; coll. HA • 25 shells; Lang Son Province, Bac Son district, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.32412&amp;materialsCitation.latitude=21.945467" title="Search Plazi for locations around (long 106.32412/lat 21.945467)">Long Dong</a>, 3.8 km north from junction of roads 1 B and 241; 21°56.728'N, 106°19.447'E; 390 m a. s. l.; 18 Feb. 2020; A. Hunyadi leg.; coll. HA • 4 shells; Lang Son, vicinity of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.74682&amp;materialsCitation.latitude=21.855883" title="Search Plazi for locations around (long 106.74682/lat 21.855883)">Chua Tam Than</a>; 21°51.353'N, 106°44.809'E; 265 m a. s. l.; 21 Feb. 2020; A. Hunyadi leg.; coll. HA • 1 shell; Ha Giang Province, Ha Giang 105.5 km - Đong Van, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.179565&amp;materialsCitation.latitude=23.1514" title="Search Plazi for locations around (long 105.179565/lat 23.1514)">Xa Van Chai</a>, left side of the road; 23°09.084'N, 105°10.774'E; 31. May 2012; A. Hunyadi leg.; coll. HA • 1 shell; Cao Bang Province, Đong Khe 3 km- <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.43228&amp;materialsCitation.latitude=22.403717" title="Search Plazi for locations around (long 106.43228/lat 22.403717)">Đeo Lung Phay</a>, right side of the road; 22°24.223'N, 106°25.937'E; 390 m a. s. l.; 15 Nov. 2011, A. Hunyadi leg.; coll. HA • 1 shell; Cao Ban Province, Tring Khanh district, Canh Tien commune, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.50915&amp;materialsCitation.latitude=22.81568" title="Search Plazi for locations around (long 106.50915/lat 22.81568)">Pac Rao village</a>; 22°48.9408'N, 106°30.549'E; 544 m a. s. l.; L. V. Hao, T. Ishibe, Y. Nakahara, K. Ohara, K. Okubo, J. U. Otani, V. P. Sang leg.; coll. PGB . • 5 shells; Bac Kan Province, Na Ri district, 2 km S of Ban Dem, W slopes of a deep sinkhole covered with forest, in small cavern, in dense rainforest; 06 Apr. 2012; J. Grego, J. Šteffek leg.; coll. JG . China • 2 shells; Hunan; ex. coll. Mollendorff; SMF 42108 • 2 shells; Guangxi, Laibin Shi, Wuxuan Xian, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=109.642235&amp;materialsCitation.latitude=23.497116" title="Search Plazi for locations around (long 109.642235/lat 23.497116)">Wuxuan Tongling Zhen</a> str., north of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=109.642235&amp;materialsCitation.latitude=23.497116" title="Search Plazi for locations around (long 109.642235/lat 23.497116)">Yagangcun</a>, rocks; 23°29.827'N, 109°38.534'E; 100 m a. s. l.; 21 Sept. 2013; A. Hunyadi &amp; M. Szekeres leg. • 17 shells; Guangxi, Hechi Shi, Bama Yaozu Zizhixian, southern edge of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.13078&amp;materialsCitation.latitude=24.117416" title="Search Plazi for locations around (long 107.13078/lat 24.117416)">Jiaolecun</a>; 24°07.045'N, 107°07.847'E; 590 m a. s. l.; 10 Sept. 2013; A. Hunyadi &amp; M. Szekeres leg. • 1 shell; Guangxi, Yulin Shi, Xingye Xian, 2 km southeast frim <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=109.757614&amp;materialsCitation.latitude=22.613567" title="Search Plazi for locations around (long 109.757614/lat 22.613567)">Chenghuang</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=109.757614&amp;materialsCitation.latitude=22.613567" title="Search Plazi for locations around (long 109.757614/lat 22.613567)">Lufengshan Scenic Area</a>; 22°36.814'N, 109°45.457'E; 05 Oct. 2009; A. Hunyadi leg.; coll. HA • 147 shells; Guangxi, Chongzuo Shi, Longzhou Xian, north of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.95595&amp;materialsCitation.latitude=22.486017" title="Search Plazi for locations around (long 106.95595/lat 22.486017)">Lenglei</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.95595&amp;materialsCitation.latitude=22.486017" title="Search Plazi for locations around (long 106.95595/lat 22.486017)">Nonggang Nat. Res.</a>, northern edge; 22°29.161'N, 106°57.357'E; 220 m a. s. l.; 23 Sept. 2013; A. Hunyadi &amp; M. Szekeres leg.; coll. HA • 2 shells; Hunan, coll. Moellendorff; SMF 42108; • 2 shells; Guangxi, Chongzuo Shi, 25 km southeast from the centre of Chongzuo, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.50598&amp;materialsCitation.latitude=22.27155" title="Search Plazi for locations around (long 107.50598/lat 22.27155)">Chongzuo Shengtai Gongyuan</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.50598&amp;materialsCitation.latitude=22.27155" title="Search Plazi for locations around (long 107.50598/lat 22.27155)">Langur Reserve</a>; 22°16.293'N, 107°30.359'E; 220 m a. s. l.; 18 Oct. 2009; A. Hunyadi leg.; coll. HA • 51 shells; Guangxi, Chongzuo Shi, Longzhou Xian, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.768265&amp;materialsCitation.latitude=22.587317" title="Search Plazi for locations around (long 106.768265/lat 22.587317)">Wude Xiang</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.768265&amp;materialsCitation.latitude=22.587317" title="Search Plazi for locations around (long 106.768265/lat 22.587317)">Banxintun</a>; 22°35.239'N, 106°46.096'E; 350 m a. s. l.; 24 Sept. 2013; A. Hunyadi leg.; coll. HA • 7 shells; Tao cun, Tong’anzhen, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=110.9337&amp;materialsCitation.latitude=24.572983" title="Search Plazi for locations around (long 110.9337/lat 24.572983)">Pinglexian</a>, Guangxi <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=110.9337&amp;materialsCitation.latitude=24.572983" title="Search Plazi for locations around (long 110.9337/lat 24.572983)">Zhuangzu Zizhiqu</a>; 24°34.379'N, 110°56.022'E; 163 m a. s. l.; leg. K. Ohara, K. Okubo, J. U. Otani; coll. PGB .</p><p>Type localities.</p><p>“ Tonkin ” (Vietnam) ( A. rochebruni); “ Die nähere Heimat dieses Findelkindes von einer Windelschnecke ist nach obigem vorläufig nimmer wohl festzustellen; doch sicher im Innern des centralen China, in Hunan oder Hupé zu suchen. ” [type locality questionable: Hunan or Hubei provinces, China] ( A. angulinus).</p><p>Differential diagnosis.</p><p>The shell shape is most similar to A. messageri . However, it differs by the much weaker and fewer apertural barriers. Anauchen pentadens is less triangular, has a narrower umbilicus, lacks the spiral striation, has more apertural barriers which are also stronger and has strong ribs on the last whorl (especially near the aperture). Anauchen jokaii sp. nov. lacks the spiral striation and has its last whorl keeled at the base which separates it from A. rochebruni . The apertural barriers are also much weaker in A. rochebruni . See also under A. turritus sp. nov.</p><p>Distribution.</p><p>This species has a wide distribution extending from the Hunan province, China (type locality of A. angulinus) south to Lang Son province, Vietnam.</p><p>Remarks.</p><p>Anauchen angulinus is herein treated a synonym of this species because we observed no significant morphological differences between them.</p><p>There is intraspecific variation in the number of apertural barriers. There are usually three (parietal, palatal, columellar) but there can be as few as one (parietal). The last whorl can be more or less swollen, sometimes resulting in a strongly triangular shell shape.</p></div>	https://treatment.plazi.org/id/C85319C0F7BE53FBBF7E1987BC4A0D43	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
1C6E602308FA51A5A002C3908F2DACFF.text	1C6E602308FA51A5A002C3908F2DACFF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anauchen turritus Gojsina, Hunyadi & Pall-Gergely 2025	<div><p>Anauchen turritus Gojšina, Hunyadi &amp; Páll-Gergely sp. nov.</p><p>Figs 10 P, 32, 33, 34, 37</p><p>Type material.</p><p>Holotype. Vietnam • 1 shell (SH: 4.7 mm, SW: 3.2 mm); Thanh Hoa Province, 23.7 km south from centre of Ngoc Lac, Phuc Thinh, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.369934&amp;materialsCitation.latitude=19.93115" title="Search Plazi for locations around (long 105.369934/lat 19.93115)">Lang Mieng</a>, rock above the village; 19°55.869'N, 105°22.196'E; 65 m a. s. l.; 13 Feb. 2020; A. Hunyadi, H. V. Luong, J. U. Otani &amp; S. V. Pham leg.; IEBR _LS_Anauchen.001 H . Paratypes. Vietnam • 1 shell; same data as for holotype; CUMZ 14432 • 79 shells; same data as for holotype; coll. HA • 1 shell; same data as for holotype; coll. VG • 29 shells; Thanh Hoa Province, Nhur Thanh District, Hai Van, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.57715&amp;materialsCitation.latitude=19.617983" title="Search Plazi for locations around (long 105.57715/lat 19.617983)">Hang Lo Cao Khang Chien</a>, near cave; 19°37.079'N, 105°34.629'E; 22 m a. s. l.; 14 Feb. 2020; A. Hunyadi, H. V. Luong, J. U. Otani &amp; S. V. Pham leg.; coll. HA .</p><p>Additional material examined.</p><p>Vietnam • 13 shells (juveniles, not paratypes); same data as for holotype; coll. HA • 7 shells (juveniles / damaged, not paratypes); Thanh Hoa Province, Nhur Thanh District, Hai Van, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.57715&amp;materialsCitation.latitude=19.617983" title="Search Plazi for locations around (long 105.57715/lat 19.617983)">Hang Lo Cao Khang Chien</a>, near cave; 19°37.079'N, 105°34.629'E; 22 m a. s. l.; 14 Feb. 2020; A. Hunyadi, H. V. Luong, J. U. Otani &amp; S. V. Pham leg.; coll. HA .</p><p>Type locality.</p><p>Vietnam, Thanh Hoa Province, 23.7 km south from centre of Ngoc Lac, Phuc Thinh, Lang Mieng, rock above the village; 19°55.869'N, 105°22.196'E; 65 m a. s. l.</p><p>Diagnosis.</p><p>Shell conical, consisting of 7.5–8 whorls. Teleoconch densely spirally striated. Last whorl adnate to the penultimate. Aperture equipped with four main barriers (parietal, upper palatal, lower palatal and columellar), supra-angular and additional two smaller barriers (basal and infraparietal).</p><p>Description.</p><p>Shell conical to weakly concave-conical, light brown-yellowish in colour, opaque. It is consisting of 7–8 regularly increasing whorls separated by a moderately deep suture. All whorls rounded, weakly convex. Protoconch lighter than the rest of the shell, slightly translucent, consisting of ~ 1.25–1.5 weakly spirally striated whorls. Teleoconch finely and densely spirally striated, especially on the last ~ 3 whorls. Spiral striae strong, irregularly spaced, curved,&gt; 40 on the last whorl in a standard view. The space between two spiral striae ranges from the width of one to the width of two or even three spiral striae. These striae are occasionally crossed by a few radial whitish streaks, randomly positioned, but mostly present on the last and the penultimate whorl. Last whorl adnate to the penultimate, slightly ascending (~ 5 ° compared to the shell axis) and making the apertural profile slightly opisthocline to the shell axis. Peristome thick, the same colour as the rest of the shell, expanded and not reflected, very finely pitted. It is especially strongly expanded at the parietal side where it leans against the penultimate whorl and forms a thick callus. Aperture equipped with four main barriers (parietal, upper palatal, lower palatal, and columellar) and several smaller ones. Parietal lamella very strong and high, directed towards the lower palatal plica. Upper palatal plica very strong, developed equally as parietal lamella and slightly leaning towards the lower palatal plica. Parietal lamella and upper palatal plica are closely positioned, leaving a narrow spacing between them. Lower palatal plica is strong but weaker than the upper palatal, directed towards the embayment between columellar and parietal lamellae. Columellar lamella slightly oblique, directed towards the lower palatal plica and developed to the same extent. Between these main barriers, smaller number of minor ones are also observed. Supra-angular lamella present, small, and short, leaning towards the palatal wall. Interpalatal plica sometimes present, positioned rather obliquely. Subcolumellar lamella present as weak or absent. Basal plica present, short. A very slight thickening (swelling) like infraparietal lamella is present. This lamella is sometimes much stronger. Surface of all apertural barriers is finely granulated. Sinulus large, distinctly separated from the rest of the aperture due to the strong angulo-parietal lamella and upper palatal plica. Umbilicus narrow, measuring ~ 1 / 8 of the shell width and with almost perpendicular walls, not showing previous whorls.</p><p>Differential diagnosis.</p><p>This species is similar to various congeners such as A. rochebruni and A. messageri . However, it can be clearly separated from them by the separate lamellae on the parietal side (angular and parietal) and a generally more elongated shell with slightly more pronounced spiral threads. Anauchen turritus sp. nov. has spiral striation, more apertural barriers and a wider umbilicus than A. jokaii sp. nov. See also under H. annamiticum .</p><p>Measurements</p><p>(in mm, n = 5). SH = 4.05–4.7; SW = 2.72–3.43; AH = 1.54–1.88; AW = 0.7–0.87.</p><p>Etymology.</p><p>This species is named after the tower-shaped (turreted) shell.</p><p>Distribution.</p><p>This species is known from two localities in Thanh Hoa Province, Vietnam.</p><p>Remarks.</p><p>We have observed significant intraspecific variability in this species. Shell can be high conical or conical (even weakly concave-conical). Some specimens (Fig. 32) have only six apertural barriers (supra-angular, parietal, columellar, basal, upper, and lower palatal) while others can have up to nine (including interpalatal, infraparietal and subcolumellar) (Fig. 34). Colouration is usually pale brown but colourless specimens also occur.</p></div>	https://treatment.plazi.org/id/1C6E602308FA51A5A002C3908F2DACFF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
60D512D3B2A758A6B1314737B13FC2FA.text	60D512D3B2A758A6B1314737B13FC2FA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anauchen utaithaniensis Panha 2002	<div><p>Anauchen utaithaniensis Panha, 2002</p><p>Figs 10 Q, 35, 36, 37</p><p>Anauchen utaithaniensis Panha in Burch &amp; Panha, 2002: 244–245, fig. 3.</p><p>Anauchen utaithaniensis — Tongkerd et al. 2004: 143, fig. 2; Panha and Burch 2008: 53–54, fig. 49; Dumrongrojwattana and Wongkamhaeng 2024: 323, fig. 7.</p><p>Type material examined.</p><p>Thailand • holotype; 1997; S. Panha leg.; CUMZ ver. 041 • 2 paratypes; same data as for the holotype; SMF 331454 .</p><p>Type locality.</p><p>“ Patavi Mountain, Utaithani Province, 15°28'20"N, 99°45'17"E, 70 meters elevation … ” (Thailand).</p><p>Diagnosis.</p><p>This species is most similar to A. huaykhakang and A. banmiensis but they are both spirally striated. Anauchen angthongensis is also spirally striated, has it last whorl regularly rounded and has a narrower umbilicus.</p><p>Distribution.</p><p>This species is known only from the type locality.</p><p>Remarks.</p><p>This species shows intra-populational variation of the angulo-parietal lamella. The angulo-parietal lamella of the holotype is bifid, while that of the paratype is almost completely fused.</p></div>	https://treatment.plazi.org/id/60D512D3B2A758A6B1314737B13FC2FA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
4A0A2A9498A556FD8FEB6C1E31E2371A.text	4A0A2A9498A556FD8FEB6C1E31E2371A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bensonella alycaeus Gojsina, Hunyadi & Pall-Gergely 2025	<div><p>Bensonella alycaeus Gojšina, Hunyadi &amp; Páll-Gergely sp. nov.</p><p>Figs 39 A, 40, 41, 100</p><p>Type material.</p><p>Holotype. Thailand • 1 shell (SH: 1.6 mm, SW: 1.3 mm); Chiang Rai Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.839165&amp;materialsCitation.latitude=20.342216" title="Search Plazi for locations around (long 99.839165/lat 20.342216)">Doi Thung</a>; 20°20.533'N, 99°50.350'E; 1320 m a. s. l.; 08 May 1988; F. G. Thompson leg. UF 347144 . Paratypes. Thailand • 1 shell; same data as for holotype; CUMZ 14433 • 7 shells; same data as for holotype; UF 591340 • 1 shell; Chiang Rai Province, Doi Tung, 50 meters before <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.83317&amp;materialsCitation.latitude=20.32605" title="Search Plazi for locations around (long 99.83317/lat 20.32605)">Wat Phra That Doi Tung</a>, vicinity of the parking lot; 20°19.563'N, 99°49.990'E; 1350 m a. s. l.; 12 Feb. 2015; A. Hunyadi leg.; coll. HA .</p><p>Additional material examined.</p><p>Thailand • 1 shell (juvenile, not paratype); same data as for holotype; UF 583721 .</p><p>Type locality.</p><p>Thailand, Chiang Rai Province, Doi Thung; 20°20.533'N, 99°50.350'E; 1320 m a. s. l.</p><p>Diagnosis.</p><p>Bensonella species with a triangular-conical shell and strongly radially ribbed last whorl, especially near the aperture. Palatal plicae very strong. Umbilicus very narrow and elongated, dot-like.</p><p>Description.</p><p>Shell triangular, conical-ovoid, brown, consisting of 4.5–5 convex, rounded whorls separated by a moderately deep suture. Protoconch spirally striated (~ 12–15 spiral striae). The protoconch is initially light in colour and then regularly darkening towards the teleoconch. Teleoconch surface very finely spirally striated, striae much weaker than on the protoconch. On the penultimate whorl, relatively coarse, weak, radial growth lines are visible. They become more densely arranged and more prominent on the last whorl and especially near the aperture where they are rib-like (important for species identification!). They are crossed by very delicate and widely spaced spiral striae. Last whorl rounded and adnate to the penultimate. It is slightly ascending near the aperture (~ 20 ° compared to the shell axis), making the aperture profile weakly opisthocline. Peristome lighter than the rest of the shell, not much expanded and not reflected. There is a slight thickening behind the peristome in form of a cervical crest. Aperture equipped with 10–12 barriers. Parietal lamella is the strongest in the aperture, long and curved, sinuated in its middle part. Angular lamella long, thin, and continuous, reaching the peristome. There are four main palatal plicae (upper palatal, two interpalatals, and a lower palatal) and usually one smaller suprapalatal plica. Upper palatal plica strong, highest in its middle part and getting lower towards the peristome. Interpalatal plicae similar to each other and the upper palatal. Lower palatal plica is the strongest. There is a strong palatal tubercle sitting on the palatal lip of the peristome, sometimes of almost rectangular shape. Peristome is distinctly sinuated behind this tubercle. Below the palatal tubercle, there are usually one or two more tubercle-like swellings which are of unknown homology. Basal plica very weak and short. Columellar lamella almost horizontal and slightly weaker than upper and lower palatal plicae. Infraparietal lamella similar to the basal plica. Surface of all apertural barriers is finely granulated. Sinulus small, narrow, and distinctly separated from the rest of the aperture. Umbilicus very narrow, dot like and elongated.</p><p>Differential diagnosis.</p><p>This species is different from other congeners by the strong, rib-like radial growth lines, especially on the last whorl near the aperture and very strong palatal plicae.</p><p>Measurements</p><p>(in mm, n = 5). SH = 1.55–1.78; SW = 1.3–1.57; AH = 0.65–0.80; AW = 0.65–0.70.</p><p>Etymology.</p><p>The specific epithet refers to the cyclophoroid subfamily Alycaeinae W. T. Blanford, 1864 due to the changing rib density on the last whorl. To be used as a noun in apposition.</p><p>Distribution.</p><p>This species is known only from the type locality.</p></div>	https://treatment.plazi.org/id/4A0A2A9498A556FD8FEB6C1E31E2371A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
E6A978DAC6675D6DA73A0DD2B0B34AD4.text	E6A978DAC6675D6DA73A0DD2B0B34AD4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bensonella boettgeri (Mollendorff 1897)	<div><p>Bensonella boettgeri (Möllendorff, 1897)</p><p>Figs 39 B, 42, 43, 60</p><p>Boysidia boettgeri Möllendorff, 1897: 70.</p><p>Boysidia (Paraboysidia) boettgeri — Pilsbry 1917: 208–209, pl. 34, figs 7, 8.</p><p>Paraboysidia boettgeri — van Benthem Jutting 1950: 38; van Benthem Jutting 1952: 318; Zilch 1984: 164; Maassen 1999: 123.</p><p>Boysidia novemdentata Saurin, 1953: 115–116, fig. 1, pl. 4, fig. 4 a – c. syn. nov.</p><p>Boysidia novemdentata — Inkhavilay et al. 2019: 59, fig. 26 B.</p><p>Bensonella novemdentata — Inkhavilay and Sutcharit 2024: 444, fig. 4.</p><p>Type material examined.</p><p>Indonesia • lectotype of B. boettgeri; W Java; collector unknown; SMF 4616 . Laos • 1 syntype of B. novemdentata; Pah Hia, Laos; collector unknown; MNHN -IM-2000-33881 .</p><p>Additional material examined.</p><p>India • 6 shells; Nagaland, Naga hills; UF 00112290 / 5, UF 179749 / 1 • 3 shells; Eastern Naga Hills; W. Doherty leg.; NHMUK 1903.7. 1.2853.</p><p>Type localities.</p><p>Java, Indonesia ( B. boettgeri); “ environs du village méo de Pah Hia, à 100 kilomètres au Sud de Xieng-Khouang, chef-lieu de la province du Tran Ninh, Laos ” (probably refers to Ban Namthong, Longchaeng District, Xaisomboun Province, Laos, see Páll-Gergely et al. 2016) ( B. novemdentata).</p><p>Differential diagnosis.</p><p>Even though there are no peculiarities regarding the apertural dentition of this species, it can be separated from all its congeners by the far most triangular, elongated shell and relatively shallow suture.</p><p>Distribution.</p><p>This species is described from Java, Indonesia, but is also known from its synonym ( B. novemdentata) in Laos. It has also been found in E India.</p><p>Remarks.</p><p>Boysidia novemdentata was described from northeastern Laos,&gt; 2800 km from the type locality of B. boettgeri . However, these two species are virtually identical in shell morphology. See the discussion section.</p></div>	https://treatment.plazi.org/id/E6A978DAC6675D6DA73A0DD2B0B34AD4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
1B07E1B30EC657ADB2E3370E1B034C1B.text	1B07E1B30EC657ADB2E3370E1B034C1B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bensonella cardiostoma Gojsina, Vermeulen & Pall-Gergely 2025	<div><p>Bensonella cardiostoma Gojšina, Vermeulen &amp; Páll-Gergely sp. nov.</p><p>Figs 101 A, 102, 107</p><p>Type material.</p><p>Holotype. Laos • 1 shell (SH: 1.12 mm; SW: 1.17 mm); Vientiane Province, Vang Vieng District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.37468&amp;materialsCitation.latitude=19.145517" title="Search Plazi for locations around (long 102.37468/lat 19.145517)">Nakay Village</a>, steep forested limestone hill; 19°08.731'N, 102°22.481'E; 323 m a. s. l.; 24 Oct. 2006; A. Schuiteman leg.; locality code JJV 15174; CUMZ 14447.</p><p>Type locality.</p><p>Laos, Vientiane Province, Vang Vieng District, Nakay Village, steep forested limestone hill, 19°08.731'N, 102°22.481'E, 323 m a. s. l.</p><p>Diagnosis.</p><p>Bensonella species with obscure, weak spiral striae, asymmetric heart-shaped aperture, an angular lamella, and a palatal tubercle on the peristome and six barriers (upper palatal, lower palatal, basal, columellar, infraparietal, parietal) situated deeper inside the aperture.</p><p>Description.</p><p>Shell colourless (white), conical-ovoid with strongly widened last whorl. Whorls 3.5, nearly rounded, slightly shouldered. Protoconch consisting of 1.25 whorls, pitted, showing spiralling pattern, although no clear spiral striae are visible. Teleoconch sculpture roughly pitted. Last whorl adnate to the penultimate, very slightly ascending (~ 10 ° compared to the shell axis), thus making the apertural profile opisthocline to the shell axis. Aperture asymmetric heart-shaped, the parietal part being the shield-like extended peristome. Angular lamella and palatal tubercle situated on the peristome, both are pointed, directed towards each other. Other barriers, altogether 6, are lamella-like, elongated, low. Upper palatal plica sits behind palatal tubercle, lower palatal situated between the basal and upper palatal plica. Columella lamella situated halfway between basal and infraparietal lamella, infraparietal and parietal situated close to each other. Peristome strongly expanded, not reflected. Surface of all apertural barriers is finely granulated. Sinulus relatively narrow and clearly separated from the rest of the aperture due to the strong and closely positioned angular lamella and palatal tubercle. Umbilicus very narrow, shows only last whorl. Last quarter whorl sigmoid.</p><p>Differential diagnosis.</p><p>See under B. mitochondria sp. nov. and B. wangviangensis .</p><p>Measurements</p><p>(in mm, n = 1). SH = 1.12; SW = 1.17; AH = 0.56; AW = 0.58.</p><p>Etymology.</p><p>Named after the heart-shaped aperture.</p><p>Distribution.</p><p>This species is known only from the type locality.</p></div>	https://treatment.plazi.org/id/1B07E1B30EC657ADB2E3370E1B034C1B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
6AF9CBF51F8758D7ADBE188AF977CD68.text	6AF9CBF51F8758D7ADBE188AF977CD68.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bensonella cristatissima Gojsina, Hunyadi & Pall-Gergely 2025	<div><p>Bensonella cristatissima Gojšina, Hunyadi &amp; Páll-Gergely sp. nov.</p><p>Figs 39 C, 44, 45, 100</p><p>Type material.</p><p>Holotype. Myanmar • 1 shell (SH: 1.77 mm, SW: 1.51 mm); Shan State, Taunggyi, mountainside above Aye Say Tee, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.0506&amp;materialsCitation.latitude=20.791483" title="Search Plazi for locations around (long 97.0506/lat 20.791483)">Dragon Cave</a>; 20°47.489'N, 97°3.036'E; 1380 m a. s. l.; 08 Oct. 2018; A. Hunyadi, K. Okubo &amp; J. U. Otani leg.; CUMZ 14434 . Paratypes. Myanmar • 95 shells; same data as for holotype; coll. HA • 1 shell; same data as for holotype; coll. VG • 27 shells Shan State, west-southwest from Taunggyi, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.01762&amp;materialsCitation.latitude=20.7547" title="Search Plazi for locations around (long 97.01762/lat 20.7547)">Montawa cave</a>; 20°45.282'N, 97°1.057'E; 1260 m a. s. l.; 05 Oct. 2018; A. Hunyadi, K. Okubo &amp; J. U. Otani leg.; coll. HA • 10 shells; Shan State, Hopong centre ca 22 km towards Namsang, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.33532&amp;materialsCitation.latitude=20.818068" title="Search Plazi for locations around (long 97.33532/lat 20.818068)">Hkoche</a>, near <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.33532&amp;materialsCitation.latitude=20.818068" title="Search Plazi for locations around (long 97.33532/lat 20.818068)">Htem Sann Cave</a>; 20°49.084'N, 97°20.119'E; 1240 m a. s. l.; 06 Oct. 2018; A. Hunyadi, K. Okubo &amp; J. U. Otani leg.; coll. HA .</p><p>Additional material examined.</p><p>Myanmar • 4 shells (3 juveniles and 1 broken, not paratypes); same data as for holotype; coll. HA .</p><p>Type locality.</p><p>Myanmar Shan State, Taunggyi, mountainside above Aye Say Tee, Dragon Cave; 20°47.489'N, 97°3.036'E; 1380 m a. s. l.</p><p>Diagnosis.</p><p>A small Bensonella species with fine, finely dimpled teleoconch sculpture, rounded last whorl adnate to the penultimate, and a strong, sharp cervical crest right behind the expanding peristome. Palatal plicae long and equally developed; umbilicus very narrow to almost closed.</p><p>Description.</p><p>Shell triangular, slightly conical-ovoid, brownish or yellowish, weakly glossy. It is consisting of 4.5–5 regularly increasing, convex, rounded whorls separated by a deep suture. Protoconch consisting of ~ 1.5 coarsely spirally striated whorls. There are ~ 13 spiral striae on the protoconch and they are more densely arranged at the bottom of the whorl. Teleoconch surface finely dimpled (pasty) and with not numerous, weak radial growth lines. Spiral striae present initially on the teleoconch but getting lost so that the surface remains finely pasty-like or a very weak spiralling pattern might be visible (only on SEM images). Last whorl rounded, adnate to the penultimate. Right behind the peristome, there is a strong, thick, sharp, white cervical crest which is wider than the peristome and even visible behind it in the apertural view. Peristome thick and expanded but not reflected. Aperture equipped with numerous barriers, four of which are particularly strong (angular, parietal, infraparietal, and columellar), and situated closer to the peristome edge. Parietal lamella is the strongest and highest in the aperture. Angular lamella much longer than the parietal, very slender but continuous. Its inner and outer parts are higher than its middle part. Columellar lamella similar in length to the parietal only slightly weaker. This lamella is almost horizontal but sometimes slightly curved towards the palatal plicae. Infraparietal lamella is roughly half as strong as the parietal. Palatal plicae equidistant, all of equal height and length, altogether four (one upper palatal, two interpalatals, and a lower palatal). They are all high at their inner parts and then regularly lowering and tapering towards their outer parts. Palatal tubercle moderately strong or absent. Basal plica present, the same length as the palatals or slightly shorter. Surface of all apertural barriers is finely granulated. Sinulus small and narrow (resembling a comma sign), distinctly separated from the rest of the aperture. Umbilicus very narrow (measuring ~ 1 / 10 of the shell width) or even dot-like, almost closed.</p><p>Differential diagnosis.</p><p>See under B. dha sp. nov.</p><p>Measurements</p><p>(in mm, n = 5). SH = 1.77–1.86; SW = 1.45–1.52; AH = 0.73–0.80; AW = 0.63–0.76.</p><p>Etymology.</p><p>This species is named for the strong cervical crest which is the strongest among all the congeners.</p><p>Distribution.</p><p>This species in known from three localities in Shan State, Myanmar.</p></div>	https://treatment.plazi.org/id/6AF9CBF51F8758D7ADBE188AF977CD68	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
83D19BD115155B0AA694B46165FA96BC.text	83D19BD115155B0AA694B46165FA96BC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bensonella dha Gojsina, Hunyadi & Pall-Gergely 2025	<div><p>Bensonella dha Gojšina, Hunyadi &amp; Páll-Gergely sp. nov.</p><p>Figs 39 D, 46, 47, 97</p><p>Type material.</p><p>Holotype. Myanmar • 1 shell (SH: 1.9 mm, SW: 1.3 mm); Shan State, 5.8 km from centre of Hopong towards Namsang, left side of road # 4, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.22448&amp;materialsCitation.latitude=20.817133" title="Search Plazi for locations around (long 97.22448/lat 20.817133)">Hopong Spring Cave</a>; 20°49.028'N, 97°13.469'E; 1110 m a. s. l.; 06 Oct. 2018; A. Hunyadi, K. Okubo &amp; J. U. Otani leg.; CUMZ 14435.</p><p>Paratypes. Myanmar • 34 shells; same data as for holotype; coll. HA • 1 shell; same data as for holotype; coll. VG .</p><p>Type locality.</p><p>Myanmar, Shan State, 5.8 km from centre of Hopong towards Namsang, left side of road # 4, Hopong Spring Cave; 20°49.028'N, 97°13.469'E; 1110 m a. s. l.</p><p>Additional material examined.</p><p>Myanmar • 2 shells (juveniles, not paratypes); same data as for holotype; coll. HA • 4 shells; Shan State, 22 km from centre of Hopong towards Namsang, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.33532&amp;materialsCitation.latitude=20.81806" title="Search Plazi for locations around (long 97.33532/lat 20.81806)">Htem Sann Cave</a>; 20°49.0836'N, 97°20.1192'E; 1240 m a. s. l.; 06 Oct. 2018; A. Hunyadi, K. Okubo &amp; J. U. Otani leg.; coll. HA • 2 shells; Shan State, 13.5 km east-southeast from centre Kalaw, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=96.613235&amp;materialsCitation.latitude=20.590433" title="Search Plazi for locations around (long 96.613235/lat 20.590433)">Myinmati Taung</a>; 20°35.426'N, 96°36.794'E; 1350 m a. s. l.; 03 Oct. 2018; A. Hunyadi, K. Okubo &amp; J. U. Otani leg.; coll. HA • 11 shells; Shan State, 7.4 km from centre of Hopong towards Namsang, 5 km north on road # 4, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.237785&amp;materialsCitation.latitude=20.849384" title="Search Plazi for locations around (long 97.237785/lat 20.849384)">Parpant cave</a>; 20°50.963'N, 97°14.267'E; 1170 m a. s. l.; 06 Oct. 2018; A. Hunyadi, K. Okubo &amp; J. U. Otani leg.; coll. HA • 4 shells; Shan State, Kalaw, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=96.55789&amp;materialsCitation.latitude=20.621027" title="Search Plazi for locations around (long 96.55789/lat 20.621027)">Shwe Oo Min Paya</a>; 20°37.2616'N, 96°33.4735'E; 1340 m a. s. l.; 02 Oct. 2018; A. Hunyadi, K. Okubo &amp; J. U. Otani leg.; coll. HA .</p><p>Diagnosis.</p><p>Bensonella species with conical shell which is not spirally striated. Apertural barriers numerous (angular, parietal, five palatals, one basal, one subcolumellar, one columellar, and one infraparietal) and with additional, strong palatal tubercle. Aperture well rounded.</p><p>Description.</p><p>Shell conical to conical-ovoid, light brownish and weakly glossy, consisting of 4.25–5.5 convex rounded whorls separated by a deep suture. Protoconch roughly pitted, sometimes with a spiralling pattern and consisting of ~ 1.5–1.75 whorls. Teleoconch with fine pasty structure, not spirally striated but with very fine radial growth lines. Last whorl adnate to the penultimate, aperture profile only slightly prosocline to the shell axis due to the slightly descending last whorl (~ 15 ° compared to the shell axis). Peristome slightly lighter than the rest of the shell, expanded but not reflected. It is particularly expanded on the parietal side where it forms a weak callus which is leaned on the penultimate whorl. Aperture equipped with moderately strong, numerous barriers. Parietal lamella moderately strong and high. Angular lamella longer but overall weaker than the parietal. It is higher in its inner part and regularly tapering towards the peristome, which is not reached. There are two or three very weak barriers inside the sinulus. Palatal wall equipped with five long barriers (suprapalatal, upper palatal, interpalatal, lower palatal, and infrapalatal) and a single strong palatal tubercle sitting on the edge of the palatal lip. Suprapalatal plica weak, not much stronger than the sinulus plicae. Upper palatal plica only slightly stronger than the suprapalatal. Interpalatal moderate. Lower palatal is the strongest among all the palatal plicae. Infrapalatal weak and shorter than others. Basal plica developed to the same extent as the infrapalatal. Subcolumellar lamella weak and short (nearly the same as the basal and infrapalatal). Columellar lamella strong as the lower palatal, almost horizontal. Infraparietal lamella moderate, ~ ½ as strong as the columellar. Surface of all apertural barriers is finely granulated. Sinulus small and not very distinctly separated from the rest of the aperture. Umbilicus very narrow, dot-like.</p><p>Differential diagnosis.</p><p>This species differs from B. cristatissima sp. nov. by the thinner peristome, less elongated apertural barriers and by the absence of a strong and sharp cervical crest.</p><p>Measurements</p><p>(in mm, n = 13). SH = 1.56–2.02; SW = 1.14–1.53; AH = 0.64–0.78; AW = 0.64–0.73.</p><p>Etymology.</p><p>Named after the Burmese word for sword / knife (dha) referring to the blade-like anterior part of the palatal plicae. To be used as a noun in apposition.</p><p>Distribution.</p><p>This species is known from five localities in Shan State, Myanmar.</p><p>Remarks.</p><p>We have observed some intraspecific variability between the populations. The population from Shwe Oo Min Paya is the smallest and with the most ovoid shell. While the populations from Myinmati Taung and Parpant cave were the most triangular with slightly shallower sutures.</p></div>	https://treatment.plazi.org/id/83D19BD115155B0AA694B46165FA96BC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
EDE3449803FC5A39BA15190A618A5B78.text	EDE3449803FC5A39BA15190A618A5B78.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bensonella dracula Gojsina, Hunyadi & Pall-Gergely 2025	<div><p>Bensonella dracula Gojšina, Hunyadi &amp; Páll-Gergely sp. nov.</p><p>Figs 39 E, 48, 49, 50, 51, 97</p><p>Type material.</p><p>Holotype. Myanmar • 1 shell (SH: 2 mm, SW: 1.6 mm); Shan State, 22 km from centre of Hopong towards Namsang, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.33532&amp;materialsCitation.latitude=20.818068" title="Search Plazi for locations around (long 97.33532/lat 20.818068)">Hkoche</a>, near <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.33532&amp;materialsCitation.latitude=20.818068" title="Search Plazi for locations around (long 97.33532/lat 20.818068)">Htem Sann Cave</a>; 20°49.084'N, 97°20.119'E; 1240 m a. s. l.; 06 Oct. 2018; A. Hunyadi, K. Okubo &amp; J. U. Otani leg.; CUMZ 14436 . Paratypes. Myanmar • 16 shells; same data as for holotype; coll. HA .</p><p>Type locality.</p><p>Myanmar, Shan State, 22 km from centre of Hopong towards Namsang, Hkoche, near Htem Sann Cave; 20°49.084'N, 97°20.119'E; 1240 m a. s. l.</p><p>Additional material examined.</p><p>Myanmar • 3 shells; Shan State, Pinlaung centre SSW 11 km – Laneli Bridge, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=96.76213&amp;materialsCitation.latitude=20.035233" title="Search Plazi for locations around (long 96.76213/lat 20.035233)">Nam Pam</a>, near “ Upper Spider Cave ”; 20°2.114'N, 96°45.728'E; 1420 m a. s. l.; 04 Oct. 2018; A. Hunyadi, K. Okubo &amp; J. U. Otani leg.; coll. HA • 1 shell; Myanmar, Kayah State, Hpruso district, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.04283&amp;materialsCitation.latitude=19.379066" title="Search Plazi for locations around (long 97.04283/lat 19.379066)">Maw Thi Do Village</a>, entrance of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.04283&amp;materialsCitation.latitude=19.379066" title="Search Plazi for locations around (long 97.04283/lat 19.379066)">Phruno river</a> cave; 19°22.744'N, 97°2.570'E; 12 Feb. 2019; J. Grego leg.; coll. JG .</p><p>Diagnosis.</p><p>Bensonella species with triangular, conical-ovoid shell and rounded whorls devoid of spiral striation. There are two palatal tubercles in front of a strong and frontally concave transversal plica. A single, strong basal plica present.</p><p>Description.</p><p>Shell triangular, conical-ovoid, consisting of 5–5.5 rounded, convex whorls separated by a deep suture. Protoconch consisting of ~ 1.25–1.5 roughly pitted whorls. Teleoconch devoid of spiral striation but with some fine radial growth lines, otherwise smooth, pasty. Last whorl adnate to penultimate. Aperture profile slightly opisthocline to the shell axis due to the very slightly ascending last whorl (~ 5 ° compared to the shell axis). In this view, a weak cervical crest could be observed behind the peristome. Peristome thick, expanded (especially on the parietal side where it forms a thick callus) but not reflected. Sinulus small and distinctly separated from the rest of the aperture. Aperture equipped with nine barriers (angular, parietal, upper palatal, transversal, two palatal tubercles, basal, columellar, and infraparietal). Parietal lamella very strong and high, curved and blade-like. Angular lamella much weaker and lower but longer, reaching the expanding peristome. Upper palatal plica moderate, higher in its middle parts. Transversal plica strong and with its frontal surface concave. In front of the transversal plica, there are two strong palatal tubercles. The upper one (usually present in all Bensonella species) is narrower and more acute. The lower palatal tubercle is wider and blunter. Basal plica present, short and ~ ½ as strong as the columellar lamella. The latter is strong, wide, and almost horizontal. Infraparietal lamella as strong as the basal plica. Surface of all apertural barriers is finely granulated. Umbilicus very narrow and slightly elongated, dot-like.</p><p>Differential diagnosis.</p><p>See under B. nordsiecki, B. obex sp. nov. and B. taiyaiorum .</p><p>Measurements</p><p>(in mm, n = 6). SH = 1.80–2.21; SW = 1.44–1.80; AH = 0.64–0.96; AW = 0.62–0.94.</p><p>Etymology.</p><p>Two strong palatal tubercles of this species resemble teeth of a vampire. The specific epithet refers to the iconic vampire, Count Dracula. To be used as a noun in apposition.</p><p>Distribution.</p><p>This species is known from the surroundings of Hopong and Pinlaung in Shan State, as well as Phruno river cave, Kayah State.</p><p>Remarks.</p><p>Specimens from Pinlaung (Shan) have a shorter transversal plica and have slightly more slender shells (Fig. 50). Specimen from the Phruno river cave in Kayah State has a more ovoid shell (Fig. 51).</p></div>	https://treatment.plazi.org/id/EDE3449803FC5A39BA15190A618A5B78	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
D7497534A48052AD8846965726961082.text	D7497534A48052AD8846965726961082.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bensonella exploda Gojsina, Hunyadi & Pall-Gergely 2025	<div><p>Bensonella exploda Gojšina, Hunyadi &amp; Páll-Gergely sp. nov.</p><p>Figs 39 F, 52, 53, 54, 100</p><p>Type material.</p><p>Holotype. Myanmar • 1 shell (SH: 1.8 mm, SW: 1.8 mm); Shan State, Pinlaung centre N 7.5 km, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=96.7907&amp;materialsCitation.latitude=20.171217" title="Search Plazi for locations around (long 96.7907/lat 20.171217)">Tar Kge</a>, near “ Big Bang Cave ”; 20°10.273'N, 96°47.442'E; 1540 m a. s. l.; 04 Oct. 2018; A. Hunyadi, K. Okubo &amp; J. U. Otani leg.; CUMZ 14446 . Paratypes. Myanmar • 32 shells; same data as for holotype; coll. HA • 1 shell; same data as for holotype; coll. VG .</p><p>Additional material examined.</p><p>Myanmar • 3 shells (juveniles, not paratypes); same data as for holotype; coll. HA • 149 shells; Shan State, 5.7 km south-southwest from centre of Pinlaung, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=96.77053&amp;materialsCitation.latitude=20.0692" title="Search Plazi for locations around (long 96.77053/lat 20.0692)">Wingabar Taung</a>; 20°4.152'N, 96°46.232'E; 1510 m a. s. l.; 04 Oct. 2018; A. Hunyadi, K. Okubo &amp; J. U. Otani leg.; coll. HA .</p><p>Type locality.</p><p>Myanmar, Shan State, Pinlaung centre N 7.5 km, Tar Kge, near “ Big Bang Cave ”; 20°10.273'N, 96°47.442'E; 1540 m a. s. l.</p><p>Diagnosis.</p><p>Colourless Bensonella species with triangular conical-ovoid and spirally striated whorls. Last whorl slightly ascending near the aperture. Aperture equipped with eight strong barriers (angular, parietal, infraparietal, columellar, lower palatal, interpalatal, upper palatal, and a palatal tubercle). Umbilicus narrow but clearly wider than in majority of congeners.</p><p>Description.</p><p>Shell strongly triangular, conical-ovoid, colourless, consisting of 5–5.5 regularly growing whorls separated by a moderately deep suture. All whorls rounded, convex. Protoconch-teleoconch boundary not clearly visible due to the similar surface sculpture and colouration. Protoconch with ten coarsely and equidistantly spaced spiral striae. Teleoconch also spirally striated but additionally with coarse radial growth lines which are roughly equidistant. Spacing between two spiral striae irregular, it ranges from the width of two to the width of four spiral striae. Last whorl flat at its base, adnate to the penultimate (but peristome not fully leaning on it), slightly ascending near the aperture (~ 5 ° compared to the shell axis) making the apertural profile weakly opisthocline to the shell axis. Peristome white, expanded but not reflected. Sinulus well-rounded and distinctly separated from the rest of the aperture. Aperture equipped with eight barriers (angular, parietal, infraparietal, columellar, lower palatal, interpalatal, upper palatal, and a palatal tubercle). All barriers relatively strong. Parietal lamella is the strongest in the aperture and wavy. Angular lamella is longer than the parietal and reaching closer to the expanding peristome. It is sinuated in its middle part. Infraparietal lamella ~ 1.5 × weaker than the columellar, short. Columellar lamella strong as the lower palatal or slightly stronger. It is positioned obliquely to the shell axis, directed towards the parietal lamella. Basal plica absent. Palatal plicae all equally strong and relatively short, upper palatal is usually the shortest. Interpalatal situated halfway between the upper and lower palatal. Palatal tubercle prominent, sitting on the palatal lip and roughly in the level with upper palatal plica. Surface of all apertural barriers is finely granulated. Umbilicus narrow (but clearly wider than in the majority of congeners), measuring ~ 1 / 8–1 / 9 of the shell width.</p><p>Differential diagnosis.</p><p>This species differs from other congeners by the combination of the flat base of the last whorl, spirally striated whorls, and wider umbilicus.</p><p>Measurements</p><p>(in mm, n = 5). SH = 1.77–1.89; SW = 1.58–1.71; AH = 0.71–0.77; AW = 0.66–0.68.</p><p>Etymology.</p><p>This species is named exploda after its type locality, the “ Big Bang Cave ”.</p><p>Distribution.</p><p>This species is known only from two localities in Shan State, Pinlaung.</p><p>Remarks.</p><p>The specimens from Wingabar Taung (Fig. 54) share all the characters with the specimens from Big Bang Cave but have weaker apertural barriers.</p></div>	https://treatment.plazi.org/id/D7497534A48052AD8846965726961082	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
D8ED8A92D14B5EFAA96C55CF37D5711F.text	D8ED8A92D14B5EFAA96C55CF37D5711F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bensonella fengxianensis (D. - N. Chen, Y. - H. Liu & W. - X. Wu 1995)	<div><p>Bensonella fengxianensis (D. - N. Chen, Y. - H. Liu &amp; W. - X. Wu, 1995)</p><p>Figs 55, 56</p><p>Boysidia fengxianensis Chen, Liu &amp; Wu, 1995: 275–276 (Chinese description), 276–277 (English description), fig. 2.</p><p>Bensonella fengxianensis — Schileyko 1998: 140; Páll-Gergely and White 2023: 2016.</p><p>Boysidia (Boysidia) fengxianensis — Chen and Zhang 2002: 692.</p><p>Boysidia (Bensonella) fengxianensis — Zhang et al. 2014: 572.</p><p>Material examined.</p><p>China • 7 shells; Shaanxi province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.764465&amp;materialsCitation.latitude=33.961815" title="Search Plazi for locations around (long 108.764465/lat 33.961815)">Chang’an district</a>, subtropical broadleaf forest, 1.2 km SE Xiaoqian Goukou village; 33°57.709'N, 108°45.868'E; 780 m a. s. l.; 08 July 2010; D. M. Palatov leg.; coll. JG • 4 shells; Hubei, Enshi Tujiazu Miaozu Zizhizhou, Enshi Shi, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.57335&amp;materialsCitation.latitude=30.577717" title="Search Plazi for locations around (long 108.57335/lat 30.577717)">Taiyanghe Xian</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.57335&amp;materialsCitation.latitude=30.577717" title="Search Plazi for locations around (long 108.57335/lat 30.577717)">Suobuya Shilin Jingqu</a>, split rock; 30°34.663'N, 108°34.401'E; 975 m a. s. l.; 06 Nov. 2010; A. Hunyadi leg.; coll. HA • 1 shell; Hubei, Enshi Tujiazu Miaozu Zizhizhou, Badong Xian, E <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=110.4214&amp;materialsCitation.latitude=31.024534" title="Search Plazi for locations around (long 110.4214/lat 31.024534)">Badong</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=110.4214&amp;materialsCitation.latitude=31.024534" title="Search Plazi for locations around (long 110.4214/lat 31.024534)">Bashan Senlin Gongyuan</a> (near <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=110.4214&amp;materialsCitation.latitude=31.024534" title="Search Plazi for locations around (long 110.4214/lat 31.024534)">Xinglingzhen</a>); 31°01.472'N, 110°25.284'E; 225 m; 03 Nov. 2011; A. Hunyadi leg.; coll. HA • 1 shell; Longfengyan, Hengjiangqiaoxiang, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=109.5775&amp;materialsCitation.latitude=26.456367" title="Search Plazi for locations around (long 109.5775/lat 26.456367)">Jingzhoumiaozudongzuzizhixian</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=109.5775&amp;materialsCitation.latitude=26.456367" title="Search Plazi for locations around (long 109.5775/lat 26.456367)">Hunansheng</a>; 26°27.382'N, 109°34.650'E; 357 m a. s. l.; K. Ohara, K. Okubo, J. U. Otani leg.; coll. PGB • 5 shells; Sanjiaoyan. Zhilouzangzuxiang, Wenxian, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.4112&amp;materialsCitation.latitude=32.886333" title="Search Plazi for locations around (long 104.4112/lat 32.886333)">Gansusheng Wenxian</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.4112&amp;materialsCitation.latitude=32.886333" title="Search Plazi for locations around (long 104.4112/lat 32.886333)">Gansasheng</a>; 32°53.180′N,, 104°24.672'E; 1646 m a. s. l.; Y. Nakahara, K. Okubo, J. U. Otani leg.; coll. PGB • 1 shell; along walking path below Cuying Lake (翠映湖), Qingchenghoushan (青城后山), <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.47914&amp;materialsCitation.latitude=30.93717" title="Search Plazi for locations around (long 103.47914/lat 30.93717)">Qingchengshanzhen</a> (青城山镇), <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.47914&amp;materialsCitation.latitude=30.93717" title="Search Plazi for locations around (long 103.47914/lat 30.93717)">Dujiangyanshì</a> (都江堰市), <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.47914&amp;materialsCitation.latitude=30.93717" title="Search Plazi for locations around (long 103.47914/lat 30.93717)">Chengdushì</a> (成都市), <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.47914&amp;materialsCitation.latitude=30.93717" title="Search Plazi for locations around (long 103.47914/lat 30.93717)">Sìchuansheng</a> (四川省); 30°56.2302'N, 103°28.7484'E; 1310 m a. s. l.; A. Hunyadi, T. Ishibe, K. Okubo, J. U. Otani, M. Szekeres leg.; coll. HA .</p><p>Type locality.</p><p>“ Fengxian County (33°08'N, 106°05'E), Shaanxi Province ” (China).</p><p>Distribution.</p><p>This species is known only from the type locality.</p><p>Remarks.</p><p>We could not locate the type specimens of this species. The original illustration of the holotype is inadequate. Based on the presence of the palatal tubercle, arrangement of apertural barriers, and shell shape, this species must belong to Bensonella (as already suggested by Páll-Gergely and White 2023). The specimens examined by us, collected in Shaanxi province (China) where the type locality of B. fengxianensis is located, may belong to this species, but this cannot be confirmed due to the absence of the type specimen. The same can be said for other samples in the nearby provinces (see Fig. 56).</p></div>	https://treatment.plazi.org/id/D8ED8A92D14B5EFAA96C55CF37D5711F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
E4B317B9D8295265900D725F9B28567D.text	E4B317B9D8295265900D725F9B28567D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bensonella fracta Gojsina, Hunyadi & Pall-Gergely 2025	<div><p>Bensonella fracta Gojšina, Hunyadi &amp; Páll-Gergely sp. nov.</p><p>Figs 101 B, 103, 104, 107</p><p>Type material.</p><p>Holotype. Thailand • 1 shell (SH: 2.08 mm; SW: 2.09 mm); Loei Province, Nong Hin district, 20.3 km southwest from centre of Nong Hin towards <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.727585&amp;materialsCitation.latitude=17.041183" title="Search Plazi for locations around (long 101.727585/lat 17.041183)">Pha Wai</a>, left side of road no. 3029; 17°2.471'N, 101°43.655'E; 705 m a. s. l.; 28 Feb. 2023; A. Hunyadi &amp; J. U. Otani leg.; CUMZ 14448.</p><p>Paratypes. Thailand • 5 shells; same data as for holotype; coll. HA .</p><p>Type locality.</p><p>Thailand, Loei Province, Nong Hin district, 20.3 km southwest from centre of Nong Hin, Pha Wai; left side of road no. 3029; 17°2.471'N, 101°43.655'E; 705 m a. s. l.</p><p>Diagnosis.</p><p>Bensonella species with spirally striated teleoconch and strongly convex whorls separated by a deep suture. Last whorl narrow, slightly detached, rounded. Apertural barriers usually 7. Upper palatal plica bipartite, consisting of anterior and posterior bulge.</p><p>Description.</p><p>Shell conical, light yellowish-brownish, opaque. It is consisting of ~ 5 regularly increasing, strongly convex whorls separated by a deeply impressed suture. Protoconch finely pitted, showing a spiralling pattern (which is stronger terminally) and consisting of ~ 1.5 whorls, coloured as the rest of the shell. Teleoconch with sculpture consisting of fine, raised, and innumerable spiral striae which are occasionally crossed by much weaker radial growth lines (rarely present as strong white streaks). The sculpture of the teleoconch gets more prominent as the whorls are increasing, thus being most noticeable on the last two whorls. Last whorl relatively narrow (low), slightly detached from the penultimate, and slightly ascending near the aperture (~ 5–10 ° compared to the shell axis). It is completely rounded and strongly convex. Peristome the same colour as the rest of the shell, strongly expanded with finely pitted surface, not reflected. Aperture equipped with five main barriers (angular, parietal, upper palatal, lower palatal, and columellar). Angular lamella short and thick, directed towards the lower palatal plica. Parietal lamella longer than the angular, leaned towards the upper palatal plica. Upper palatal plica usually bipartite, consisting of anterior and posterior bulge. Anterior bulge is closer to the peristome, smaller and lower. Posterior bulge is longer, higher, and stronger, positioned behind the anterior (deeper in the aperture). Origin of the anterior bulge is not known but it may be homologous with the palatal tubercle usually found in many typical Bensonella species. Columellar lamella developed to the similar extent as the palatal plicae. In front of the columellar lamella, there is sometimes a small swelling on the peristome. Apart from these main barriers, there is usually an additional smaller basal plica and one even smaller knob-like lamella in the columello-parietal embayment. Only the angular lamella and anterior bulge of the upper palatal plica are reaching t peristome. Surface of all apertural barriers is finely granulated. Sinulus small, well separated from the rest of the shell due to the closely situated angular lamella and upper palatal plica. Umbilicus moderately wide (1 / 5–1 / 6 of the shell width), showing only the penultimate whorl. Umbilical groove absent.</p><p>Differential diagnosis.</p><p>Bensonella fracta sp. nov. is much larger than all other representatives of Bensonella wangviangensis group. See also under B. wangviangensis .</p><p>Measurements</p><p>(in mm, n = 4). SW 1 = 2.09–2.18; SW 2 = 1.31–1.48; SH = 1.9–2.08; AH = 0.81–1.1; AW = 0.89–1.23.</p><p>Etymology.</p><p>The first examined specimen of this species had a crack on the apical whorls, which gave the specific epithet fracta.</p><p>Distribution.</p><p>This species is known only from the type locality.</p><p>Remarks.</p><p>This species is not a typical representative of this group since the angular lamella is quite small and not almost enclosing the sinulus. However, it is placed here because of the high morphological similarity with mostly B. wangviangensis . A tubercle-like swelling in the columellar-parietal embayment was found to be missing in one specimen. Otherwise, this species does not seem to be much variable regarding apertural barriers.</p></div>	https://treatment.plazi.org/id/E4B317B9D8295265900D725F9B28567D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
A2C203FEE1E752A1A91CEDA97DD03617.text	A2C203FEE1E752A1A91CEDA97DD03617.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bensonella geminounca Jirapatrasilp & Tongkerd 2024	<div><p>Bensonella geminounca Jirapatrasilp &amp; Tongkerd, 2024</p><p>Figs 39 G, 57, 100</p><p>Bensonella geminounca Jirapatrasilp &amp; Tongkerd in Jirapatrasilp et al. 2024: 90–91, fig. 2.</p><p>Type material examined.</p><p>Thailand • holotype; 27. May 1997, S. Panha, J. B. Burch, P. Dumrongrojwattana leg.; CUMZ 14366 .</p><p>Type locality.</p><p>“ Doi Chiang Dao, Chiang Dao Wildlife Sanctuary, Chiang Dao District, Chiang Mai Province, Thailand (19°22′046.3″N, 098°51′08.1″E, 2,015 m a. m. s. l.) ”.</p><p>Differential diagnosis.</p><p>This species has a unique appearance of the infraparietal lamella which readily distinguishes it from all other congeners: it is consisting of two hooked parts pointing toward each other, outer (pointing inwards) and inner (pointing outwards).</p><p>Distribution.</p><p>This species is known from different altitudes at the type locality (altitude range: 1650–2015 m).</p></div>	https://treatment.plazi.org/id/A2C203FEE1E752A1A91CEDA97DD03617	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
7F5AEA59A815501984DEA68197701E66.text	7F5AEA59A815501984DEA68197701E66.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bensonella hooki Pall-Gergely 2023	<div><p>Bensonella hooki Páll-Gergely, 2023</p><p>Figs 39 H, 58</p><p>Bensonella hooki Páll-Gergely in Páll-Gergely and White 2023: 2023–2025, figs 6, 9 c, d.</p><p>Type material examined.</p><p>India • holotype; Godwin-Austen coll.; NHMUK 20191114 .</p><p>Type locality.</p><p>“ … Cherrapunji … ” (India).</p><p>Differential diagnosis.</p><p>See under B. lakainguta, B. montawa sp. nov., B. multihami, B. plicidens and B. spinosa sp. nov.</p><p>Distribution.</p><p>SE and SW Himalaya region.</p><p>Remarks.</p><p>After long-lasting taxonomic confusion, it has been shown that this species (hooked apertural barriers) is distinct from B. plicidens (blunt barriers) even though both “ forms ” were found mixed together in the same specimen lot (for details see Páll-Gergely and White 2023).</p></div>	https://treatment.plazi.org/id/7F5AEA59A815501984DEA68197701E66	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
92F5A80D85FD53E0866D9AF74756E6E0.text	92F5A80D85FD53E0866D9AF74756E6E0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bensonella karoensis Maassen 1999	<div><p>Bensonella karoensis Maassen, 1999</p><p>Figs 39 I, 59, 60</p><p>Bensonella karoensis Maassen, 1999: 123–126, figs 8, 9.</p><p>Type material examined.</p><p>Indonesia • holotype; Jul 1996; W. Maassen leg.; RMNH.Moll.84438 .</p><p>Type locality.</p><p>“ N. Sumatra, Karo Highlands, Kuta Buluh, 40 km N of Brastagi, near the entrance of the cave Liangdehar in leaf litter at the foot of limestone rocks ”.</p><p>Differential diagnosis.</p><p>See under B. plicidens .</p><p>Distribution.</p><p>This species is known only from the type locality.</p><p>Remarks.</p><p>This species is known only by a holotype (Maassen 1999).</p></div>	https://treatment.plazi.org/id/92F5A80D85FD53E0866D9AF74756E6E0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
D00007D260145D33AC6E811CD2C945CC.text	D00007D260145D33AC6E811CD2C945CC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bensonella lakainguta C. - C. Hwang 2014	<div><p>Bensonella lakainguta C. - C. Hwang, 2014</p><p>Figs 39 J, 61</p><p>Bensonella plicidens lakainguta Hwang, 2014: 18–19, figs 2–4.</p><p>Boysidia (Bensonella) qingliangfengensis Fang, Wang &amp; Chen, 2015: 692, fig. 1.</p><p>Bensonella lakainguta — Páll-Gergely and White 2023: 2025–2026, figs 7, 8, 9 e – h.</p><p>Bensonella qingliangfengensis — Páll-Gergely and White 2023: 2020, fig. 7 d – f.</p><p>Type material examined.</p><p>Taiwan • holotype; 27 Aug. 1998, C-C. Hwang leg.; NMNS - 7244-00 .</p><p>Additional material examined.</p><p>Japan • 4 shells; limestone outcrop along forest rd., upstream Ugui-gawa, Aiga, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=136.36975&amp;materialsCitation.latitude=34.96518" title="Search Plazi for locations around (long 136.36975/lat 34.96518)">Tsuchiyama-cho</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=136.36975&amp;materialsCitation.latitude=34.96518" title="Search Plazi for locations around (long 136.36975/lat 34.96518)">Koka</a>, Shiga; 34°57.9108'N, 136°22.185'E; 461 m a. s. l.; Kawase, J. U. Otani leg.; coll. PGB .</p><p>Type locality.</p><p>“ TAIWAN: Mt. Zhibenzhushan, Wutai, Pingdung, 22°43'32.6"N, 120°52'50.2"E, alt. 2100 m ”.</p><p>Differential diagnosis.</p><p>This species differs from B. hooki by its more triangular shell, more pointed apex, and parietal callus adnate to the penultimate whorl (not so in B. hooki, see Páll-Gergely and White 2023). For differences from B. montawa sp. nov., B. multihami, B. plicidens, and B. spinosa sp. nov. see under those species.</p><p>Distribution.</p><p>This species is well known from Japan as well as Taiwan and China (where it is known from a synonym, B. qingliangfengensis) (Fang et al. 2015; Páll-Gergely and White 2023).</p><p>Remarks.</p><p>This species was originally described as a subspecies of B. plicidens . Páll-Gergely and White (2023) have shown that it is distinct as it has hooked and not blunt apertural barriers, so. B. plicidens does not inhabit Japan and Taiwan where only B. lakainguta occurs.</p></div>	https://treatment.plazi.org/id/D00007D260145D33AC6E811CD2C945CC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
975238FFB3ED573D896FDCD05D796E5F.text	975238FFB3ED573D896FDCD05D796E5F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bensonella lophiodera Tongkerd & Panha 2024	<div><p>Bensonella lophiodera Tongkerd &amp; Panha, 2024</p><p>Figs 39 K, 62, 63, 97</p><p>Bensonella lophiodera Tongkerd &amp; Panha in Tongkerd et al. 2024: 176–178, figs 7 C, 9, 13 I.</p><p>Type material examined.</p><p>Myanmar • holotype; collector unknown; CUMZ 14378 .</p><p>Additional material examined.</p><p>Myanmar • 158 shells; Shan State, NNE of Kalaw, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=96.582115&amp;materialsCitation.latitude=20.655333" title="Search Plazi for locations around (long 96.582115/lat 20.655333)">Osei Mountain Pagoda NW 150 m</a>; 20°39.320'N, 96°34.927'E; 1565 m a. s. l.; 03 Oct. 2018; A. Hunyadi, K. Okubo &amp; J. U. Otani leg.; coll. HA • 181 shells; Shan State, Kalaw ESE 13.5 km, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=96.61323&amp;materialsCitation.latitude=20.59044" title="Search Plazi for locations around (long 96.61323/lat 20.59044)">Myinmati Taung</a>; 20°35.4264'N, 96°36.7938'E; 1350 m a. s. l.; 03 Oct. 2018; A. Hunyadi, K. Okubo &amp; J. U. Otani leg; coll. HA .</p><p>Type locality.</p><p>“ Myinmati Cave, Kalaw Township, Taunggyi District, Shan State, Myanmar (20°35'24.6"N, 96°36'42.1"E; 1312 m a. s. l.) ”.</p><p>Differential diagnosis.</p><p>The shell of this species is relatively large when compared to other geographically approximate congeners and the transversal palatal plica is rather weak and small. See under B. taiyaiorum .</p><p>Distribution.</p><p>This species is known from three localities in Kalaw.</p><p>Remarks.</p><p>This species lacks the palatal tubercle but is still clearly a member of Bensonella because of the triangular-conical shell shape and three barriers on the parietal side. In our specimens collected from Osei Mountain Pagoda (see above under “ Material examined ”), there is a transversal plica which is usually only partly merged, i. e., interpalatal and lower palatal plicae are clearly discernible (Fig. 63). Even though in the original description, this barrier is listed as a lower palatal, we believe that it is actually a transversal plica but just much weaker than in other congeners. Furthermore, Tongkerd et al. (2024: fig. 9 C) shows a lower palatal plica that is wider than normal and positioned transversally indicating the fusion of the interpalatal and lower palatal plicae. Otherwise, our examined specimens fit perfectly in the description of B. lophiodera by all shell characters (size, colour, shape, surface sculpture, and apertural dentition).</p></div>	https://treatment.plazi.org/id/975238FFB3ED573D896FDCD05D796E5F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
DAA9BE7AA09550DDAE4043AC30C7B0BF.text	DAA9BE7AA09550DDAE4043AC30C7B0BF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bensonella microdentata Gojsina & Pall-Gergely 2025	<div><p>Bensonella microdentata Gojšina &amp; Páll-Gergely sp. nov.</p><p>Figs 39 L, 64, 65, 95</p><p>Type material.</p><p>Holotype. Thailand • 1 shell (SH: 2 mm; SW: 1.5 mm); Lampang Province limestone dome at pass 11 km SSW of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.80639&amp;materialsCitation.latitude=18.466112" title="Search Plazi for locations around (long 99.80639/lat 18.466112)">Ban Pang La</a>; 18°27.9667'N, 99°48.3833'E; 610 m a. s. l.; 14. May 1988; F. G. Thompson leg.; UF 346912 . Paratypes. Thailand • 1 shell; same data as for holotype; CUMZ 14437 • 6 shells; same data as for holotype; UF 591333 .</p><p>Additional material examined.</p><p>Thailand • 3 shells (juveniles, not paratypes); same data as the holotype; UF 583726 .</p><p>Type locality.</p><p>Thailand, Lampang Province limestone dome at pass 11 km SSW of Ban Pang La; 18°27.9667'N, 99°48.3833'E; 610 m a. s. l.</p><p>Diagnosis.</p><p>Bensonella species with concave-conical, glossy shell that is spirally striated. Apertural barriers (usually eight or nine) and the surface around them prominently spiniferous.</p><p>Description.</p><p>Shell slightly concave-conical, glossy, light brownish, consisting of 5–5.5 convex and rounded whorls separated by a deep suture. Protoconch spirally striated, its boundary with the teleoconch not clear. There are ~ 12 widely spaced spiral striae on the protoconch and they are stronger at the base of the whorl. Teleoconch with relatively coarsely spaced, raised spiral striae (space between two striae equals the width of two or three striae) and with fine radial growth lines. On initial teleoconch whorls, radial lines are more densely arranged (similar to the spacing between the spiral striae) and are crossing the spiral striae forming a reticulated sculpture. Towards the last whorl, radial lines become coarser and the reticulated sculpture becomes weaker. Last whorl rounded, adnate to the penultimate, slightly descending near the aperture (~ 20 ° compared to the shell axis). Peristome whitish, expanded but not reflected. Sinulus parabolic due to the position of the angular lamella and the palatal tubercle. Aperture equipped with numerous barriers which have the typical Bensonella arrangement. These are: angular, parietal, upper palatal, interpalatal, lower palatal, palatal tubercle, basal, columellar, and infraparietal. Parietal lamella strong and moderately high, angular separated into inner and outer parts of almost equal height and width and a strong sinuation between them. Upper and interpalatal plicae positioned close together and almost equally as strong. Lower palatal plica situated some distance from other palatals and stronger than both of them. Palatal tubercle strong and sitting on the edge of the palatal lip. Basal plica weak, low, and short. Columellar lamella nearly as strong as the lower palatal. Infraparietal lamella only very slightly stronger than the basal plica. Surface of all apertural barriers is roughly spiniferous. Umbilicus narrow but relatively deep, measuring ~ 1 / 6 of the shell width.</p><p>Differential diagnosis.</p><p>This species is most similar to B. nitens (see under that species for differences). See also under B. paviei and B. tamphathai .</p><p>Measurements</p><p>(in mm, n = 4). SH = 1.73–2; SW = 1.47–1.61; AH = 0.73–0.83; AW = 0.63–0.75.</p><p>Etymology.</p><p>This species is named for its small apertural barriers.</p><p>Distribution.</p><p>This species is known only from the type locality.</p></div>	https://treatment.plazi.org/id/DAA9BE7AA09550DDAE4043AC30C7B0BF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
541B399B3B525192928139DDAF9FB6A5.text	541B399B3B525192928139DDAF9FB6A5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bensonella mirabilis Gojsina, Grego & Pall-Gergely 2025	<div><p>Bensonella mirabilis Gojšina, Grego &amp; Páll-Gergely sp. nov.</p><p>Figs 39 M, 66, 67, 95</p><p>Type material.</p><p>Holotype. Myanmar • 1 shell (SH: 2.05 mm, SW: 1.65 mm); Kayah State, Hpruso district, Maw Thi Do, road towards <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.03588&amp;materialsCitation.latitude=19.382767" title="Search Plazi for locations around (long 97.03588/lat 19.382767)">Han Li village</a>, under bridge over <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.03588&amp;materialsCitation.latitude=19.382767" title="Search Plazi for locations around (long 97.03588/lat 19.382767)">Phruno river</a>; 19°22.966'N, 97°2.153'E; 12 Feb. 2019; J. Grego leg.; CUMZ 14438 . Paratypes. Myanmar • 9 shells; same data as for holotype; coll. JG • 13 shells; Kayah state, Hpruso district, Maw Thi Do Village, entrance of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.04283&amp;materialsCitation.latitude=19.379066" title="Search Plazi for locations around (long 97.04283/lat 19.379066)">Phruno river</a> cave; 19°22.744'N, 97°2.570'E; 12 Feb. 2019; J. Grego leg.; coll. JG • 1 shell; same data as previous; coll. HA • 1 shell; same data as previous; coll. VG • 2 shells; Kayah State, Hpruso distr., Hoyar village, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=96.93993&amp;materialsCitation.latitude=19.3016" title="Search Plazi for locations around (long 96.93993/lat 19.3016)">Kwar Yin cave</a>; 19°18.096'N, 96°56.396'E; 13 Feb. 2019; J. Grego leg.; coll. JG • 1 shell; Kayah state, Hpruso district, Maw Thi Do, road towards <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.03513&amp;materialsCitation.latitude=19.383516" title="Search Plazi for locations around (long 97.03513/lat 19.383516)">Han Li village</a>, rocks above bridge over <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.03513&amp;materialsCitation.latitude=19.383516" title="Search Plazi for locations around (long 97.03513/lat 19.383516)">Phruno river</a>; 19°23.011'N, 97°2.108'E; 12 Feb. 2019; J. Grego leg.; coll. JG .</p><p>Additional material examined.</p><p>Myanmar • 5 shells (juveniles, not paratypes); same data as for holotype; coll. JG .</p><p>Type locality.</p><p>Myanmar, Kayah State, Hpruso district, Maw Thi Do, road towards Han Li village, under bridge over Phruno river; 19°22.966'N, 97°2.153'E.</p><p>Diagnosis.</p><p>Bensonella species with seven apertural barriers and a prominent hooked upper palatal plica which is pointing laterally (toward the palatal wall).</p><p>Description.</p><p>Shell triangular, slightly conical-ovoid, brownish, consisting of 5–5.5 whorls separated by a deep suture. All whorls rounded, convex, regularly increasing. Protoconch coarsely spirally striated, ~ 1.5 whorls. Spiral striae much more prominent terminally on the protoconch. Teleoconch with a fine pasty surface sculpture, without spiral striae but with fine radial growth lines. Last whorl rounded, adnate to the penultimate and straight near the aperture. Peristome brownish as the rest of the shell, expanded but not reflected. It is leaning on the penultimate whorl in form of a strong but not much expanded parietal callus. Palatal side of the peristome sinuated in its middle part, next to the palatal tubercle. Behind the expanding peristome, a weak cervical crest is present. Sinulus small and narrow, distinctly separated from the rest of the aperture. Aperture subrectangular, equipped with a smaller number of barriers (angular, parietal, upper palatal, lower palatal, columellar, infraparietal and palatal tubercle) than usual in Bensonella . Parietal lamella long but not reaching the expanding peristome. Angular lamella consisting of two separated parts: inner, which is slightly curved and positioned deeper in the aperture and outer part which is reaching the peristome edge. There is a strong sinuation between the inner and outer parts of the angular lamella. Palatal wall equipped with upper and lower palatal plica. Upper palatal plica of very peculiar appearance: it is very high with a hook positioned laterally (not pointing outside but directly to the palatal wall). Lower palatal plica nearly as strong as the parietal lamella. In front of the upper palatal plica, on the peristome edge, a strong palatal tubercle is observed. Columellar lamella rather weak. Above the columellar lamella, one additional infraparietal is present and it is nearly equally strong as the columellar. Surface of all apertural barriers is finely granulated. Umbilicus oblique (slightly elongated) and narrow, measuring ~ 1 / 6–1 / 7 of the shell width.</p><p>Differential diagnosis.</p><p>This species is easily distinguished from all other congeners by the presence of a laterally hooked upper palatal plica.</p><p>Measurements</p><p>(in mm, n = 5). SH = 2.03–2.16; SW = 1.52–1.70; AH = 0.80–0.88; AW = 0.77–0.82</p><p>Etymology.</p><p>Named for the remarkable upper palatal plica (laterally hooked).</p><p>Distribution.</p><p>Known from four sampling sites in Kayah state, Hpruso district.</p></div>	https://treatment.plazi.org/id/541B399B3B525192928139DDAF9FB6A5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
01F3593007FC5417ACE035B7545C9ABF.text	01F3593007FC5417ACE035B7545C9ABF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bensonella mitochondria Gojsina, Vermeulen & Pall-Gergely 2025	<div><p>Bensonella mitochondria Gojšina, Vermeulen &amp; Páll-Gergely sp. nov.</p><p>Figs 103 C, 105, 107</p><p>Type material.</p><p>Holotype. Laos • 1 shell (SH: 1.21 mm; SW: 1.27 mm); Khammouane Province, Thakhek Distr., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.13133&amp;materialsCitation.latitude=17.433216" title="Search Plazi for locations around (long 105.13133/lat 17.433216)">Ban Nakok</a>, base of forested limestone hill; 17°25.993'N, 105°07.880'E; 222 m a. s. l.; 14 Nov. 2006; A. Schuiteman leg.; locality code JJV 15157; CUMZ 14449 . Paratypes. Laos • 9 shells; same data as for holotype; coll. JJV .</p><p>Additional material examined.</p><p>Laos • 2 shells (juveniles, not paratypes); same data as for holotype; coll. JJV .</p><p>Type locality.</p><p>Laos, Khammouane Province, Thakhek Distr., Ban Nakok, base of forested limestone hill; 17°25.993'N, 105°07.880'E; 222 m a. s. l.</p><p>Diagnosis.</p><p>A colourless Bensonella species with shouldered last whorl and spirally striated teleoconch and nearly heart-shaped aperture. Apertural barriers numerous and almost all equally strong, umbilicus moderately wide.</p><p>Description.</p><p>Shell conical-ovoid, colourless. Whorls 3.5, initial whorls rounded, last two strongly shouldered, separated by a deep suture. Protoconch consisting of ~ 1 whorl, very finely pitted, with signs of indistinct and widely spaced spiralling pattern. Teleoconch with coarse spiral striae which are crossed by stronger, coarse, not numerous, and almost regularly spaced radial growth lines. Approximately 25 widely spaced spiral striae present on the last whorl in standard apertural view. Last whorl adnate to the penultimate, slightly ascending near the aperture (~ 10–15 ° compared to the shell axis). Peristome continuous, strong, and expanded, especially at the parietal side where it leans on the penultimate whorl. Aperture nearly heart-shaped, equipped with a strong angular lamella, which is curved and almost merges with the prominent palatal tubercle, thus nearly enclosing the small sinulus (which makes it strongly separated from the rest of the aperture). Angular lamella reaches the peristome. There are eight to nine other barriers situated deep in the aperture and not reaching the peristome. Additional two lamellae at the parietal side (equally strong parietal and infraparietal), one columellar lamella, one subcolumellar, one basal plica, two palatal plicae (sometimes with additional, weak infrapalatal plica) and a palatal tubercle. Upper and lower palatal plicae strong, the former is situated slightly above the palatal tubercle. Palatal tubercle quite strong and elevated. Basal plica weaker than the palatals but stronger than the subcolumellar lamella. Columellar lamella slightly stronger than them but weaker than upper and lower palatal plica. Surface of all apertural barriers is finely granulated. Umbilicus moderately wide, measuring ~ 1 / 5 of the shell width and showing only the penultimate whorl.</p><p>Differential diagnosis.</p><p>This species differs from B. cardiostoma sp. nov. most clearly by the spirally striated shell and a wider umbilicus.</p><p>Measurements</p><p>(in mm, n = 5). SH = 1.21–1.33; SW = 1.21–1.37; AH = 0.57–0.62; AW = 0.58–0.67.</p><p>Etymology.</p><p>The appearance of the apertural barriers when the last whorl is broken behind them resembles the morphology of a mitochondria (Fig. 105 F), hence the specific epithet which is to be used as a noun in apposition.</p><p>Distribution.</p><p>This species in known only from the type locality.</p></div>	https://treatment.plazi.org/id/01F3593007FC5417ACE035B7545C9ABF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
B698D88A4FC859D7B22E92907866E666.text	B698D88A4FC859D7B22E92907866E666.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bensonella montawa Gojsina, Hunyadi & Pall-Gergely 2025	<div><p>Bensonella montawa Gojšina, Hunyadi &amp; Páll-Gergely sp. nov.</p><p>Figs 39 N, 68, 95</p><p>Type material.</p><p>Holotype. Myanmar • 1 shell (SH: 1.8 mm, SW: 1.5 mm); Shan State, west-southwest from Taunggyi, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.01762&amp;materialsCitation.latitude=20.7547" title="Search Plazi for locations around (long 97.01762/lat 20.7547)">Montawa cave</a>; 20°45.282'N, 97°1.057'E; 1260 m a. s. l.; 05 Oct. 2018; A. Hunyadi, K. Okubo &amp; J. U. Otani leg.; CUMZ 14471 . Paratypes. Myanmar • 30 shells; same data as for holotype; coll. HA .</p><p>Type locality.</p><p>Myanmar, Shan State, west-southwest from Taunggyi, Montawa cave; 20°45.282'N, 97°1.057'E; 1260 m a. s. l.</p><p>Diagnosis.</p><p>A Bensonella species with conical, pale yellowish shell. Aperture equipped with numerous barriers, all of which are in form of sharp hooks pointing outside. Parietal lamella with a blunt but long projection in front of the hook, almost reaching the expanding peristome. Umbilicus narrow, dot-like.</p><p>Description.</p><p>Shell pale yellowish, triangular, conical ovoid, whorls 4–4.5, convex, rounded, rather regularly increasing. Protoconch consisting of ~ 1.25 whorls, showing spiralling pattern but with no clear spiral striae. Teleoconch with pasty surface, not spirally striated but with coarsely spaced, irregular radial growth lines. Last whorl adnate to the penultimate and slightly ascending near the aperture (~ 15–20 ° compared to the shell axis) making the aperture profile opisthocline to the shell axis. Peristome strongly expanded, not reflected. Aperture equipped with numerous barriers, all of which are in form of sharp hooks pointing outside the aperture. All barriers have low, slender projections situated in front of the hooks. A blunt, knob-like palatal tubercle sits anterior to the upper palatal plica on the peristome edge. Parietal lamella in form of a strong hook. A blunt, outer part of the parietal lamella is in form of a low, narrow, and blunt projection (situated in front of the hooked part) almost reaching the peristome edge. Angular lamella strong, consisting of inner and outer part separated by a strong sinuation. Inner part smaller, hooked, pointing towards the outer part which is larger but blunt. There are four weak barriers inside the sinulus. Upper palatal plica ~ 2 × weaker than the parietal and situated slightly above the palatal tubercle. First interpalatal plica as strong as the upper palatal, situated slightly below the palatal tubercle. Second interpalatal plica slightly weaker than the first. Lower palatal plica as strong as the first interpalatal and upper palatal. Infrapalatal plica small, as strong as the basal and subcolumellar. These three plicae are roughly equidistant. Above the subcolumellar, there is a clearly stronger columellar (similar in strength to the lower palatal plica) and one slightly stronger supracolumellar. Infraparietal lamella ~ 1 / 3 weaker than the parietal, with a similar (but shorter) projection in front of the hooked part. There are also some barrier-like swellings, although they never develop into a proper barrier. They can be e. g., between the upper palatal and first interpalatal plica, between the columellar and supracolumellar lamellae, between the supracolumellar and infraparietal lamellae, between the infraparietal and parietal lamellae. Surface of all apertural barriers is smooth. Sinulus parabolic, well separated from the rest of the aperture. Umbilicus very narrow, dot-like.</p><p>Differential diagnosis.</p><p>This species is very similar to B. hooki and B. lakainguta . Bensonella hooki is however more ovoid, reddish-brown, and does not have a slender projection in front of the parietal lamella. The latter also distinguishes the new species from B. lakainguta, which is also more triangular (i. e., pointed). See also under B. multihami .</p><p>Measurements</p><p>(in mm, n = 3). SH = 1.8; SW = 1.4–1.6; AH = 0.69–0.77; AW = 0.74–0.79.</p><p>Etymology.</p><p>This species is named after its type locality, the Montawa Cave. The specific epithet is to be used as a noun in apposition.</p><p>Distribution.</p><p>This species is only known from the type locality.</p></div>	https://treatment.plazi.org/id/B698D88A4FC859D7B22E92907866E666	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
16D64A5B58B2501DBC9E61C90319576B.text	16D64A5B58B2501DBC9E61C90319576B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bensonella multidentata Gojsina, A. Reischutz & Pall-Gergely 2025	<div><p>Bensonella multidentata Gojšina, A. Reischütz &amp; Páll-Gergely sp. nov.</p><p>Figs 39 O, 69, 70, 71, 72, 100</p><p>Type material.</p><p>Holotype. Thailand • 1 shell (SH: 2.40 mm, SW: 1.95 mm); Chiang Rai Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.77679&amp;materialsCitation.latitude=19.91285" title="Search Plazi for locations around (long 99.77679/lat 19.91285)">Wat Phra That Tham Doi Khong Khao Meditation Centre</a>, cave clay at the entrance of the cave, ca 6 km W of Chiang Rai; 19°54.771'N, 99°46.607'E; ca 420 m a. s. l.; Sept. 2017; A. Reischütz leg.; NHMW - MO-113729 . Paratypes. Thailand • 1 shell; same data as for holotype; CUMZ 14439 • 29 shells; same data as for holotype; coll. REI • 1 shell; same data as for holotype; coll. VG • 41 shells; Chiang Rai Province, 5 km west from centre of Chiang Rai, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.77737&amp;materialsCitation.latitude=19.913666" title="Search Plazi for locations around (long 99.77737/lat 19.913666)">vicinity of Wat Phra That Tham Doi Kong Khao</a>; 19°54.820'N, 99°46.642'E; 420 m a. s. l.; 11 Feb. 2015; A. Hunyadi leg.; coll. HA .</p><p>Additional material examined.</p><p>Thailand • 3 shells (damaged / juveniles, not paratypes); same data as for holotype; coll. REI • 2 shells; Chiang Rai Province, south-southwest from Mae Sai, vicinity of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.86362&amp;materialsCitation.latitude=20.328716" title="Search Plazi for locations around (long 99.86362/lat 20.328716)">Wat Tham Pla</a>; 20°19.723'N, 99°51.817'E; 400 m a. s. l.; 12 Feb. 2015; A. Hunyadi leg.; coll. HA • 22 shells; Chiang Rai Province, limestone Mtn. 6 km by road W of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.444725&amp;materialsCitation.latitude=19.648611" title="Search Plazi for locations around (long 99.444725/lat 19.648611)">Ban Mae Suai</a>; 19°38′55″N, 99°26′41″E; 650 m a. s. l.; 07. May 1988; F. G. Thompson leg.; locality code FGT-4415; UF 347105 .</p><p>Type locality.</p><p>Thailand, Chiang Rai Province, Wat Phra That Tham Doi Khong Khao Meditation Centre, cave clay at the entrance of the cave, ca 6 km NW of Chiang Rai, 19°54.771'N, 99°46.607'E, ca 420 m a. s. l.</p><p>Diagnosis.</p><p>A Bensonella species with high triangular, concave-conical, densely spirally striated shell. Last whorl wide, rounded. Multiple (at least 10) well-developed apertural barriers.</p><p>Description.</p><p>Shell light brown, triangular, concave-conical, whorls 5–5.25, convex, rounded, rather regularly increasing. Protoconch consisting of ~ 1.25 whorls, its sculpture consists of ~ 10 equidistant spiral striae. Teleoconch finely, very densely spirally striated, and ornamented with a few, inconspicuous radial growth lines. Last whorl adnate to the penultimate and slightly to moderately descending near the aperture (~ range: 15–32 ° compared to the shell axis) making the aperture profile prosocline to the shell axis. Aperture subcircular, parietal part strongly extended towards penultimate whorl. Peristome strongly expanded, not reflected. Aperture with numerous and strong barriers. A blunt, knob-like palatal tubercle sits anterior to the upper palatal plica on the peristome edge. Sometimes, this tubercle is very weak or completely absent. All other barriers sit deeper, not reaching the peristome edge. Parietal lamella is the highest among all apertural barriers, long, almost straight, only slightly curved downwards. Angular lamella much lower and more slender than the parietal one. Upper palatal plica short and located slightly above the palatal tubercle. First interpalatal and lower palatal plicae equally strong. There is a smaller and shorter second interpalatal plica between them. Basal part of peristome with two or three low plicae (rarely one), they are situated next to each other, the one situated closer to the columellar lamella may be homologous with the subcolumellar lamella, and the other may be basal plicae. Columellar lamella is weaker than parietal and stronger than the palatal plicae, straight, pointing obliquely upwards. Supracolumellar and infraparietal denticles small, knob-like. Surface of all apertural barriers is finely granulated. Sinulus parabolic, wide, and not strongly separated from the rest of the aperture. Umbilicus wide, covers 1 / 3–1 / 4 of shell width.</p><p>Differential diagnosis.</p><p>See under B. nitens sp. nov., B. paviei, B. sericata sp. nov. and B. tamphathai .</p><p>Measurements</p><p>(in mm, n = 5). SH = 2.40–2.51; SW = 1.81–1.97; AH = 0.82–0.94; AW = 0.85–0.95.</p><p>Etymology.</p><p>Named for the numerous barriers in the aperture.</p><p>Distribution.</p><p>This species is known from four localities in Chiang Rai province, Thailand.</p><p>Remarks.</p><p>The population found near Ban Mae Suai is characterised by shells that lack the palatal tubercle (or have a very slight swelling indicating its presence) and smaller number of apertural barriers (only one basal and one interpalatal). However, we do not intend to regard this population as distinct due to the lack of extensive material and the fact that main shell characters are shared with B. multidentata sp. nov. (shell shape, surface sculpture, umbilicus width and appearance of other barriers).</p></div>	https://treatment.plazi.org/id/16D64A5B58B2501DBC9E61C90319576B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
748DFECAEA235C35BB5F71A70DC9EA9C.text	748DFECAEA235C35BB5F71A70DC9EA9C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bensonella multihami Jirapatrasilp & Tongkerd 2024	<div><p>Bensonella multihami Jirapatrasilp &amp; Tongkerd, 2024</p><p>Figs 39 P, 73, 88, 100</p><p>Bensonella multihami Jirapatrasilp &amp; Tongkerd in Jirapatrasilp et al. 2024: 95–96, fig. 5.</p><p>Type material examined.</p><p>Thailand • holotype; 27. May 1997, S. Panha, J. B. Burch, P. Dumrongrojwattana leg.; CUMZ 14366 .</p><p>Type locality.</p><p>“ Doi Chiang Dao, Chiang Dao Wildlife Sanctuary, Chiang Dao District, Chiang Mai province, Thailand (19°22′046.3″N, 098°51′08.1″E, 2,015 m. a. m. s. l.) ”</p><p>Differential diagnosis.</p><p>This species is strikingly similar to B. hooki, B. lakainguta, and B. montawa sp. nov. from which it differs in its blunt parietal lamella (hooked in former three)</p><p>Distribution.</p><p>This species is, apart from its type locality, known from one additional site: Doi Ang Khang (1400 m).</p></div>	https://treatment.plazi.org/id/748DFECAEA235C35BB5F71A70DC9EA9C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
8AD6F47946E85BFE897FDC3453AF4699.text	8AD6F47946E85BFE897FDC3453AF4699.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bensonella nitens Gojsina & Pall-Gergely 2025	<div><p>Bensonella nitens Gojšina &amp; Páll-Gergely sp. nov.</p><p>Figs 39 Q, 74, 75, 100</p><p>Type material.</p><p>Holotype. Thailand • 1 shell (SH: 1.91 mm; SW: 1.77 mm); Phrae Province, 11 km W of Phrae, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.07056&amp;materialsCitation.latitude=18.178055" title="Search Plazi for locations around (long 100.07056/lat 18.178055)">on Road 1023 limestone knoll</a>, leaf litter; 18°10′41″N, 100°4′14″E; 290 m a. s. l.; 16. May 1988; F. G. Thompson leg.; locality code FGT-4447, UF 346975 . Paratypes. Thailand • 2 shells; same data as for holotype; CUMZ 14440 • 45 shells; same data as for holotype; UF 591334 .</p><p>Additional material examined.</p><p>Thailand • 4 shells (juveniles, not paratypes); same data as for holotype; UF 583730 .</p><p>Type locality.</p><p>Thailand, Phrae Province, 11 km W of Phrae, on Road 1023 limestone knoll, leaf litter; 18°10′41″N, 100°4′14″E; 290 m a. s. l.</p><p>Diagnosis.</p><p>Shell concave-conical, glossy, last whorl strongly enlarged. Teleoconch spirally striated. Last whorl adnate to the penultimate. Aperture equipped with numerous barriers, angular and columellar lamellae reaching the peristome. Umbilicus moderately wide.</p><p>Description.</p><p>Shell concave-conical, brownish, consisting of 5–5.5 convex, rounded whorls separated by a deep suture. Last whorl much wider than all preceding ones, resulting in a concave-conical shell shape. Protoconch finely spirally striated initially, more prominently terminally. Its boundary with the teleoconch is not clearly visible due to the similar surface sculpture. Fine spiral striae present across the whole shell surface but quite difficult to observe on initial teleoconch whorls without SEM imaging. They are most clearly visible on the last whorl but still very fine. Spacing between the two spiral striae approximately measures the width of one or two striae. Radial growth lines extremely fine, irregularly spaced and not numerous, most clearly visible on the last and the penultimate whorls. Last whorl rounded, slightly descending (~ 10 ° compared to the shell axis), and adnate to the penultimate near the aperture. Peristome brownish, expanded but not reflected. On the parietal side of the aperture, the peristome leans on the penultimate whorl forming a thin parietal callus. Aperture equipped with numerous barriers (angular, parietal, upper palatal, two interpalatals, lower palatal, palatal tubercle, basal, columellar, and infraparietal). Parietal lamella is the strongest in the aperture, blade like and relatively high. Angular lamella discontinuous, consisting of weaker outer part (which is reaching the expanding peristome) and stronger (higher) inner part. Palatal plicae almost all equidistant. Upper palatal and first interpalatal equally strong, higher in their inner than in outer parts (but upper palatal can be weaker than the interpalatal). There is a strong, white palatal tubercle sitting on the palatal lip of the aperture, in front and between the upper palatal and interpalatal plica. Lower interpalatal plica is the weakest among them all. Lower palatal plica is the strongest among all the palatals. Sometimes, a very weak infrapalatal plica is also present and situated between the lower palatal and basal. All palatal plicae continue as slender projections towards the peristome. Basal plica small and usually the weakest in the aperture. Columellar lamella strong and oblique, leaned towards the parietal lamella. Infraparietal lamella slightly stronger than the basal plica. Surface of all apertural barriers is finely granulated. Sinulus parabolic due to the position of the angular lamella and the palatal tubercle. Umbilicus moderately wide, measuring ~ 1 / 5 of the shell width.</p><p>Differential diagnosis.</p><p>This species is similar to Bensonella microdentata sp. nov., but the barriers are more numerous and not spiniferous, the umbilicus is wider and the last whorl is more enlarged. Bensonella nitens sp. nov. is smaller, glossier and has a weaker surface sculpture than B. multidentata sp. nov. The apertural barrier arrangement is strikingly similar in these two species but B. multidentata sp. nov. has stronger barriers and palatal plicae without slender projections towards the peristome. These projections are always present in B. nitens sp. nov. See also under B. sericata sp. nov. and B. tamphathai .</p><p>Measurements</p><p>(in mm, n = 5). SH = 1.91–2.00; SW = 1.57–1.87; AH = 0.86–0.96; AW = 0.86–0.94.</p><p>Etymology.</p><p>The specific epithet refers to the shiny shell surface.</p><p>Distribution.</p><p>This species is known only from the type locality.</p></div>	https://treatment.plazi.org/id/8AD6F47946E85BFE897FDC3453AF4699	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
A705D508F16E53AF899F5E6715BBC51A.text	A705D508F16E53AF899F5E6715BBC51A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bensonella nordsiecki Jirapatrasilp & Tongkerd 2024	<div><p>Bensonella nordsiecki Jirapatrasilp &amp; Tongkerd, 2024</p><p>Figs 39 R, 76, 100</p><p>Bensonella nordsiecki Jirapatrasilp &amp; Tongkerd in Jirapatrasilp et al. 2024: 91–93, fig. 3.</p><p>Type material examined.</p><p>Thailand • holotype; 27. May 1997, S. Panha, J. B. Burch, P. Dumrongrojwattana leg.; CUMZ 14372 .</p><p>Type locality.</p><p>“ Doi Chiang Dao, Chiang Dao Wildlife Sanctuary, Chiang Dao District, Chiang Mai province, Thailand (19°22′046.3″N, 098°51′08.1″E, 2,015 m. a. m. s. l.) ”.</p><p>Differential diagnosis.</p><p>This species is much more ovoid (less slender and triangular) than Bensonella dracula sp. nov. which also has a strong second palatal tubercle (very weak or absent in B. nordsiecki). The umbilicus is wider in B. dracula sp. nov. See also under B. obex sp. nov.</p><p>Distribution.</p><p>This species is known only from the type locality, with an altitude range from 605 m to 2015 m.</p></div>	https://treatment.plazi.org/id/A705D508F16E53AF899F5E6715BBC51A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
C2E17F1CDC7F5F8CA4222653DDAB8757.text	C2E17F1CDC7F5F8CA4222653DDAB8757.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bensonella obex Gojsina, Hunyadi & Pall-Gergely 2025	<div><p>Bensonella obex Gojšina, Hunyadi &amp; Páll-Gergely sp. nov.</p><p>Figs 39 S, 77, 78, 97</p><p>Type material.</p><p>Holotype. Myanmar • 1 shell (SH: 1.8 mm, SW: 1.5 mm); Shan State, 5.7 km south-southwest from Pinlaung, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=96.77053&amp;materialsCitation.latitude=20.0692" title="Search Plazi for locations around (long 96.77053/lat 20.0692)">Wingabar Taung</a>; 20°4.152'N, 96°46.232'E; 1510 m a. s. l.; 04 Oct. 2018; A. Hunyadi, K. Okubo &amp; J. U. Otani leg.; CUMZ 14441 . Paratypes. Myanmar • 32 shells; same data as for holotype; coll. HA .</p><p>Additional material examined.</p><p>Myanmar • 12 shells; Shan State, Pinlaung centre N 7.5 km, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=96.7907&amp;materialsCitation.latitude=20.171217" title="Search Plazi for locations around (long 96.7907/lat 20.171217)">Tar Kge</a>, near ” Big Bang Cave ”; 20°10.273'N, 96°47.442'E; 1540 m a. s. l.; A. Hunyadi, K. Okubo &amp; J. U. Otani leg.; coll. HA • 4 shells; Shan State, Pinlaung centre SSW 11 km – Laneli Bridge, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=96.76213&amp;materialsCitation.latitude=20.035233" title="Search Plazi for locations around (long 96.76213/lat 20.035233)">Nam Pam</a>, near “ Upper Spider Cave ”; 20°2.114'N, 96°45.728'E; 1420 m a. s. l.; 04 Oct. 2018; A. Hunyadi, K. Okubo &amp; J. U. Otani leg.; coll. HA .</p><p>Type locality.</p><p>Myanmar, Shan State, 5.7 km south-southwest from Pinlaung, Wingabar Taung; 20°4.152'N, 96°46.232'E; 1510 m a. s. l.</p><p>Diagnosis.</p><p>Bensonella species with a pitted protoconch and conical-ovoid shell. Last whorl slightly ascending near the aperture. Peristome usually wrinkled due to several tubercles. Transversal plica strong and very weakly concave.</p><p>Description.</p><p>Shell conical-ovoid, consisting of 4.5–5 rounded, convex, regularly increasing whorls separated by a deep suture. Shell colour slightly weathered in the holotype but probably light brownish. Protoconch consisting of 1.75 whorls, finely pitted or sometimes spirally striated, especially terminally. Teleoconch finely pitted, with a pasty sculpture and without visible spiral striae but with coarsely and irregularly spaced radial growth lines. Last whorl enlarged, adnate to the penultimate and slightly ascending near the aperture (~ 20–25 ° compared to the shell axis). Aperture profile opisthocline to the shell axis. Peristome white and expanded (at columellar but especially at the parietal side where it forms a distinct callus) but not reflected. Aperture equipped with numerous barriers. Parietal lamella strong and high, almost reaching the expanding peristome. It sometimes has a heart-like appearance due to the distinct sinuation in the middle. Angular lamella weakly wavy, slightly weaker than the parietal but longer, reaching the expanding peristome. It also has a sinuation as in the parietal lamella but weaker. Upper palatal plica very short and moderately strong. Transversal plica strong, slightly concave on the front surface. A strong palatal tubercle is sitting in front and between the upper palatal and transversal plica. Below this tubercle, there are several more tubercle-like swellings which give the peristome a wrinkled appearance. Infrapalatal plica elongated and low. Basal plicae number ranging from one to four, similarly shaped as the infrapalatal plica. Subcolumellar lamella (if present) shares the same appearance as the former two. Columellar lamella moderate, ~ ½ as strong as the parietal. Supracolumellar lamella present but can be strong or quite weak. Infraparietal lamella slightly weaker than the columellar or even half as strong. Surface of all apertural barriers is finely granulated. Sinulus relatively narrow, small, and distinctly separated from the rest of the aperture. Umbilicus very narrow, measuring ~ 1 / 10 of the shell width, slightly elongated.</p><p>Differential diagnosis.</p><p>Bensonella nordsiecki has a smooth protoconch and more ovoid shell. The transversal plica in B. nordsiecki is without a concave frontal surface. Bensonella dracula sp. nov. is less ovoid, does not have a wrinkled peristome (but two palatal tubercles) and has a wider umbilicus.</p><p>Measurements</p><p>(in mm, n = 10). SH = 1.42–1.8; SW = 1.36–1.5; AH = 0.61–0.81; AW = 0.63–0.70.</p><p>Etymology.</p><p>The specific epithet comes from a Latin word for a barricade (obex), which refers to the transversal palatal plica in the form of a barrier / obstacle. To be used as a noun in apposition.</p><p>Distribution.</p><p>This species is known from three localities around Pinlaung, Shan State.</p></div>	https://treatment.plazi.org/id/C2E17F1CDC7F5F8CA4222653DDAB8757	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
1AD8DA2A599D566DB664956EEFFC3F2B.text	1AD8DA2A599D566DB664956EEFFC3F2B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bensonella pahpetensis (Saurin 1953) Gojšina & Hunyadi & Sutcharit & Tongkerd & Auffenberg & Grego & Vermeulen & Reischütz & Páll-Gergely 2025	<div><p>Bensonella pahpetensis (Saurin, 1953) comb. nov.</p><p>Figs 39 T, 79, 100</p><p>Boysidia pahpetensis Saurin, 1953: 116, fig. 2, pl. 4, fig. 5 a – c.</p><p>Boysidia pahpetensis — Inkhavilay et al. 2019: 59, fig. 26 C; Inkhavilay and Sutcharit 2024: 442, fig. 2 A.</p><p>Type material examined.</p><p>Laos • 1 syntype; Pah Hia, Laos; collector unknown; MNHN -IM-2000-33880 .</p><p>Type locality.</p><p>“ environs du village méo de Pah Hia, à 100 kilomètres au Sud de Xieng-Khouang, chef-lieu de la province du Tran Ninh, Laos ” (probably refers to Ban Namthong, Longchaeng District, Xaisomboun Province, Laos, see Páll-Gergely et al. 2016).</p><p>Differential diagnosis.</p><p>This species is unique due to the presence of one concrescent barrier on the parietal side in combination with the palatal tubercle typical for the genus Bensonella .</p><p>Distribution.</p><p>This species in known only from the type locality.</p><p>Remarks.</p><p>Bensonella pahpetensis is not a typical representative of its genus due to the concrescent angular and parietal lamellae. However, we have placed it in this genus because of the presence of the palatal tubercle as well as the triangular-conical shell shape.</p></div>	https://treatment.plazi.org/id/1AD8DA2A599D566DB664956EEFFC3F2B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
C408234488FD516CB093F4F379DCA9C0.text	C408234488FD516CB093F4F379DCA9C0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bensonella palatotridens Jirapatrasilp & Tongkerd 2024	<div><p>Bensonella palatotridens Jirapatrasilp &amp; Tongkerd, 2024</p><p>Figs 39 U, 80, 100</p><p>Bensonella palatotridens Jirapatrasilp &amp; Tongkerd in Jirapatrasilp et al. 2024: 93, fig. 4.</p><p>Type material examined.</p><p>Thailand • holotype; 27. May 1997, S. Panha, J. B. Burch, P. Dumrongrojwattana leg.; CUMZ 14370 .</p><p>Type locality.</p><p>“ Doi Chiang Dao, Chiang Dao Wildlife Sanctuary, Chiang Dao District, Chiang Mai province, Thailand (19°22′046.3″N, 098°51′08.1″E, 2,015 m. a. m. s. l.) ”.</p><p>Differential diagnosis.</p><p>This species is characterised by three plicae on the palatal side of the aperture. See under B. paviei and B. tamphathai .</p><p>Distribution.</p><p>This species is known only from the type locality.</p></div>	https://treatment.plazi.org/id/C408234488FD516CB093F4F379DCA9C0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
601594F9B8475CC9882C68E6D3EFCFE1.text	601594F9B8475CC9882C68E6D3EFCFE1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bensonella pangmapaensis (Panha & J. B. Burch 2002)	<div><p>Bensonella pangmapaensis (Panha &amp; J. B. Burch, 2002)</p><p>Figs 39 V, 81, 100</p><p>Paraboysidia pangmapaensis Panha &amp; Burch, 2002 c: 87, fig. 5.</p><p>Paraboysidia pangmapaensis — Panha and Burch 2008: 114, fig. 97; Dumrongrojwattana and Wongkamhaeng 2024: 324.</p><p>Material examined.</p><p>Thailand • 6 shells; Mae Hong Son Province, Mae Hong Son, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.96667&amp;materialsCitation.latitude=19.416666" title="Search Plazi for locations around (long 97.96667/lat 19.416666)">Road 1095</a>; 19°25'N, 97°58'E; 390 m a. s. l.; 22 Mar. 1988; K. Auffenberg leg.; locality code KA-0596; UF 345697 • 5 shells; Mae Hong Son Province, Mae Hong Son, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.96667&amp;materialsCitation.latitude=19.416666" title="Search Plazi for locations around (long 97.96667/lat 19.416666)">Road 1095</a>; 19°25'N, 97°58'E; 390 m a. s. l.; 22 Mar. 1988; K. Auffenberg leg.; locality code KA-0595 B; UF 345721 • 7 shells; Mae Hong Son Province, 21 km N Mae Hong Son, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.96667&amp;materialsCitation.latitude=19.433332" title="Search Plazi for locations around (long 97.96667/lat 19.433332)">Fish Cave</a>; 19°26'N, 97°58'E; 380 m a. s. l.; 22 Mar. 1988; K. Auffenberg leg.; locality code KA-0593; UF 345671 • 20 shells; Mae Hong Son Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.96667&amp;materialsCitation.latitude=19.483334" title="Search Plazi for locations around (long 97.96667/lat 19.483334)">35.5 km NNE of Mae Hong Son on road to Ban Huai Phung off Road 1095</a>; 19°29'N, 97°58'E; 450 m a. s. l.; 21 Mar. 1988; K. Auffenberg leg.; locality code KA-0591; UF 345665 • 34 shells; Mae Hong Son Province, 35.1 km S Mae Hong Son, 1.7 km E of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.96667&amp;materialsCitation.latitude=19.083334" title="Search Plazi for locations around (long 97.96667/lat 19.083334)">Route 108</a>; 19°5'N, 97°58'E; 850 m a. s. l.; 23 Mar. 1988; K. Auffenberg leg.; locality code KA-0579; UF 345727 .</p><p>Type locality.</p><p>“ Lod cave, Pang Ma Pa District, Mae Hong Son Province, 19°29'36"N, 98°17'18"E and 19°34′03″N, 98°16′41″E, 800 meters elevation … ” (Thailand).</p><p>Differential diagnosis.</p><p>This species is clearly distinguishable by its rough sculpture showing the spiralling pattern in combination with hooked basal and palatal plicae, as well as the inner part of the angular lamella.</p><p>Distribution.</p><p>This species is known from several sampling sites, all located in Mae Hong Son province where the type locality is also situated.</p></div>	https://treatment.plazi.org/id/601594F9B8475CC9882C68E6D3EFCFE1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
3A2619A2810D57C7A30768F2911CAA88.text	3A2619A2810D57C7A30768F2911CAA88.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bensonella paviei (Bavay & Dautzenberg 1912)	<div><p>Bensonella paviei (Bavay &amp; Dautzenberg, 1912)</p><p>Figs 39 W, 82, 83, 100</p><p>Boysidia paviei Bavay &amp; Dautzenberg, 1912: 20, pl. 3, figs 4–6.</p><p>Boysidia (Paraboysidia) paviei — Pilsbry 1917: 203–204, pl. 35, figs 7–8; Schileyko 1998: 138, fig. 156; Chen et al. 1999: 1; Schileyko 2011: 2.</p><p>Paraboysidia paviei — van Benthem Jutting 1950: 39.</p><p>Paraboysidia nabhitabhatai Panha &amp; Burch, 2002 c: 81–84, fig. 3. syn. nov.</p><p>Paraboysidia nabhitabhatai — Panha and Burch 2008: 113–114, fig. 96.</p><p>Paraboysidia anguloobtusus Inkhavilay &amp; Panha in Inkhavilay et al. 2016: 215–217, figs 1, 2 D – F, 4 B. syn. nov.</p><p>Paraboysidia anguloobtusa — Inkhavilay et al. 2019: 61, fig. 26 F.</p><p>Bensonella paviei — Páll-Gergely and White 2023: 2015, fig. 3 f – k.</p><p>Bensonella anguloobtusa — Tongkerd et al. 2024: 171–174, figs 6 B, 7 A, 13 G.</p><p>Bensonella nabhitabhatai — Dumrongrojwattana and Wongkamhaeng 2024: 324.</p><p>Type material examined.</p><p>Vietnam • 1 syntype; Pac-Kha: Long-ping; C. Messager leg.; MNHN -IM-2000-35158 . Thailand • holotype of P. nabhitabhatai; 1998; S. Panha, P. Dumrongrojwattana, C. Sutcharit, S. Tumpeesuwan leg.; CUMZ ver. 064 .</p><p>Additional material examined.</p><p>Vietnam • 4 shells; Son La Province, 27 km east-southeast from centre of Phu Yen, Muong Do, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.785484&amp;materialsCitation.latitude=21.195517" title="Search Plazi for locations around (long 104.785484/lat 21.195517)">Ban Han Mot, southern part of the village</a>; 21°11.731'N, 104°47.129'E; 800 m a. s. l.; 06 Feb. 2020; A. Hunyadi, J. U. Otani &amp; S. V. Pham leg.; coll. HA • 23 shells; Son La Province, Moc Chau district, Van Ho, northeastern edge of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.75338&amp;materialsCitation.latitude=20.766684" title="Search Plazi for locations around (long 104.75338/lat 20.766684)">Pa Cop</a> towards <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.75338&amp;materialsCitation.latitude=20.766684" title="Search Plazi for locations around (long 104.75338/lat 20.766684)">Bo Nhang</a>; 20°46.001'N, 104°45.203'E; 980 m a. s. l.; 10 Feb. 2020; A. Hunyadi, H. V. Luong, J. U. Otani &amp; S. V. Pham leg.; coll. HA • 31 shells; Son La Province, Yen Chau district, Xa Chieng On, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.187836&amp;materialsCitation.latitude=20.991383" title="Search Plazi for locations around (long 104.187836/lat 20.991383)">Ban</a> Tram Hoc, Hang Nha Nhung, vicinity of the cave; 20°59.483'N, 104°11.270'E; 970 m a. s. l.; 09 Feb. 2020; A. Hunyadi, H. V. Luong, J. U. Otani &amp; S. V. Pham leg.; coll. HA • 110 shells; Son La Province, Quynh Nhai district, 20 km east from cross towards Thuan Chau, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.681816&amp;materialsCitation.latitude=21.55735" title="Search Plazi for locations around (long 103.681816/lat 21.55735)">Chieng Khoang</a>, cave above the village; 21°33.441'N, 103°40.909'E; 315 m a. s. l.; 07 Feb. 2020; A. Hunyadi, H. V. Luong, J. U. Otani &amp; S. V. Pham leg.; coll. HA • 10 shells; Son La Province, Moc Chau district, Long Luong, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.84586&amp;materialsCitation.latitude=20.780582" title="Search Plazi for locations around (long 104.84586/lat 20.780582)">Ban</a> Tan Lap, northern edge of the village; 20°46.835'N, 104°50.752'E; 1165 m a. s. l.; 05 Feb. 2020; A. Hunyadi, H. V. Luong, J. U. Otani &amp; S. V. Pham leg.; coll. HA • 4 shells; Sơn La Province, Thu ận Châu district, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.52793&amp;materialsCitation.latitude=21.3531" title="Search Plazi for locations around (long 103.52793/lat 21.3531)">900 m from centre of Co M ạ</a>, rock wall above the road; 21°21.186'N, 103°31.676'E; 1300 m a. s. l.; 08 Feb. 2020; A. Hunyadi leg. ; Laos • 37 shells; Udomxai Province, 6.5 km southeast from centre of Na Mor towards Udomxai, Ban Nathong, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.78302&amp;materialsCitation.latitude=20.872816" title="Search Plazi for locations around (long 101.78302/lat 20.872816)">Tham Nathong</a>, above cave spring; 20°52.369'N, 101°46.981'E; 635 m a. s. l.; 07 Oct. 2019; A. Hunyadi leg.; coll. HA • 1 shell; Luang Prabang Province, 28.5 km south from Pak Mong towards Luang Prabang, northern bank of Nam Nga, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.33463&amp;materialsCitation.latitude=20.370333" title="Search Plazi for locations around (long 102.33463/lat 20.370333)">Tham Nam Nga</a>; 20°22.220'N, 102°20.078'E; 350 m a. s. l.; 09 Oct. 2019; A. Hunyadi leg.; coll. HA • 45 shells; Luang Prabang Province, 10.3 km from centre of Luang Prabang towards Xiang Ngeum, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.185&amp;materialsCitation.latitude=19.858767" title="Search Plazi for locations around (long 102.185/lat 19.858767)">east, 1800 m on a path, Phou Xuang</a>, eastern mountain slope; 19°51.526'N, 102°11.100'E; 730 m a. s. l.; 10 Oct. 2019; A. Hunyadi leg.; coll. HA • 26 shells; Luang Prabang Province, 12 km from centre of Nong Khiaw towards Pak Xeng, northeast from <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.68602&amp;materialsCitation.latitude=20.543083" title="Search Plazi for locations around (long 102.68602/lat 20.543083)">Ban</a> Huai Lek, left side of the road; 20°32.585'N, 102°41.161'E; 350 m a. s. l.; 05 Oct. 2019; A. Hunyadi leg.; coll. HA • 103 shells; Luang Prabang Province, 2.8 km northeast from <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.438965&amp;materialsCitation.latitude=19.44645" title="Search Plazi for locations around (long 102.438965/lat 19.44645)">Phou Khoun</a>, rock wall above the cave; 19°26.787'N, 102°26.338'E; 1200 m a. s. l.; 11 Oct. 2019; A. Hunyadi leg.; coll. HA • 113 shells; Udomxai Province, 10 km southeast from centre of Na Mor, 3.8 km east-northeast from <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.81605&amp;materialsCitation.latitude=20.889517" title="Search Plazi for locations around (long 101.81605/lat 20.889517)">Na Xay</a>, northern rock wall; 20°53.371'N, 101°48.963'E; 660 m a. s. l.; 07 Oct. 2019; A. Hunyadi leg.; coll. HA • 5 shells; Xieng Khouang Province, ca 15 km SW of Ban Ko Kieng, ca 19 km SW of Vieng Thong, on the left side of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.26272&amp;materialsCitation.latitude=19.937733" title="Search Plazi for locations around (long 103.26272/lat 19.937733)">Nam Khan</a>, under rocks and logs in old forest; 19°56.264'N, 103°15.763'E; 570 m a. s. l.; 27 Oct. 2006; A. Abdou, I. V. Muratov leg.; MNHN -IM-2012-27287 • 3 shells; Luang Prabang Province, ca <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.25322&amp;materialsCitation.latitude=19.744833" title="Search Plazi for locations around (long 102.25322/lat 19.744833)">6 km E of Muang Xiang Ngeun</a>, under rocks in old secondary forest under cliff; 19°44.690'N, 102°15.193'E; 390 m a. s. l.; 21 Nov. 2006; A. Abdou, I. V. Muratov leg.; MNHN -IM-2012-27288 • 2 shells; Luang Prabang Province, Phou Xuang mountain, ca 1.5 km NE of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.184685&amp;materialsCitation.latitude=19.860083" title="Search Plazi for locations around (long 102.184685/lat 19.860083)">Ban Lak Sip</a>, ca 5 km SE of Luang Prabang, under rocks and logs in old secondary forest under cliff; 19°51.605'N, 102°11.081'E; 640 m a. s. l.; 24 Nov. 2006; A. Abdou, I. V. Muratov leg.; MNHN -IM-2012-27270 • 1 shell; Luang Prabang Province, ca 18 km SE of Muang Xiang Ngeun, on the left side of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.32905&amp;materialsCitation.latitude=19.682184" title="Search Plazi for locations around (long 102.32905/lat 19.682184)">Nam Khan</a>, under rocks and logs in old forest; 19°40.931'N, 102°19.743'E; 455 m a. s. l.; 30 Oct. 2006; A. Abdou, I. V. Muratov leg.; MNHN -IM-2012-27285 • 9 shells; Xieng Khouang Province, ca 30 km WNW of Phonsavan, ca 47 km ENE of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.86501&amp;materialsCitation.latitude=19.53365" title="Search Plazi for locations around (long 102.86501/lat 19.53365)">Phou Khoun</a>, limestone, black soil in limestone pockets, clay, under rocks in old secondary forest at base of cliff near cave; 19°32.019'N, 102°51.901'E; 1131 m a. s. l.; 15 Nov. 2006; A. Abdou, I. V. Muratov leg.; MNHN -IM-2012-27272 • 6 shells; Luang Prabang Province, just <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.43816&amp;materialsCitation.latitude=19.4464" title="Search Plazi for locations around (long 102.43816/lat 19.4464)">NE of Phou Khoun</a>, under rocks in old secondary forest above large cave; 19°26.784'N, 102°26.290'E; 1177 m a. s. l.; 15 Nov. 2006; A. Abdou, I. V. Muratov leg.; MNHN -IM-2012-27289 • 12 shells; Xieng Khouang Province, Plane of Jars, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.1536&amp;materialsCitation.latitude=19.430317" title="Search Plazi for locations around (long 103.1536/lat 19.430317)">Site</a> 1, ca 6 km S of Phonsavan, limestone, clay, under rocks near small trees under cliff surrounded by grassland; 19°25.819'N, 103°09.216'E; 1097 m a. s. l.; 14 Nov. 2006; I. V. Muratov leg.; MNHN -IM-2012-27286 • 12 shells; Luang Prabang Province, ca 17 km SE of Muang Xiang Ngeun, along small stream on the left side (ca 2 km S) of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.30897&amp;materialsCitation.latitude=19.6698" title="Search Plazi for locations around (long 102.30897/lat 19.6698)">Nam Khan</a>, on and under rocks and logs in old forest with banana; 19°40.188'N, 102°18.538'E; 525 m a. s. l.; 31 Oct. 2006; A. Abdou, I. V. Muratov leg.; MNHN -IM-2012-27273 • 22 shells; Luang Prabang Province, ca <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.40784&amp;materialsCitation.latitude=19.494217" title="Search Plazi for locations around (long 102.40784/lat 19.494217)">6 km N of Phou Khoun</a>, under rocks in dry secondary forest under and above cliff; 19°29.653'N, 102°24.470'E; 1244 m a. s. l.; 16 Nov. 2006; A. Abdou, I. V. Muratov leg.; MNHN -IM-2012-27271 • 28 shells; Xieng Khouang Province, ca 18 km SW of Ban Ko Kieng, ca 22 km SW of Vieng Thong, on the right side of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.23987&amp;materialsCitation.latitude=19.914034" title="Search Plazi for locations around (long 103.23987/lat 19.914034)">Nam Khan</a>, under rocks and logs at base of cliff in secondary forest; 19°54.842'N, 103°14.392'E; 560 m a. s. l.; 28 Oct. 2006; A. Abdou, I. V. Muratov leg.; MNHN -IM-2012-27284 . Myanmar • 1 shell; Mandalay, Anesakhan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=96.38527&amp;materialsCitation.latitude=21.979334" title="Search Plazi for locations around (long 96.38527/lat 21.979334)">Dat Taw Gyaint Waterfall</a>; 21°58.760'N, 96°23.116'E; 610 m a. s. l.; 18 Oct. 2018; Hunyadi leg; coll. HA . Thailand • 51 shells; Chiang Mai Province, northeastern side of Doi Chiang Dao, 2 km northwest from <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.91133&amp;materialsCitation.latitude=19.400267" title="Search Plazi for locations around (long 98.91133/lat 19.400267)">Wat Tham Chiang Dao</a>; 19°24.016'N, 98°54.680'E; 835 m a. s. l.; 07 Feb. 2015, A. Hunyadi leg.; coll. HA • 3 shells; Mae Hong Song Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.05&amp;materialsCitation.latitude=19.55" title="Search Plazi for locations around (long 98.05/lat 19.55)">10.2 km WNW of Soppong</a>, road 1095; 19°33'N, 98°3'E; 820 m a. s. l.; 23 Mar. 1998; K. Auffenberg leg.; locality code KA-0586; UF 345620 .</p><p>Type localities.</p><p>“ Pac-Kha ”, Vietnam ( B. paviei); “ Limestone wall outside of Kao Rao. Cave, Vieng Phouka District, Luang Namtha Province, Laos (20°43'30.1"N, 101°9'4.3"E), 732 m amsl ” ( Paraboysidia anguloobtusus); “ Phadevada, Phukhieo Wildlife Sanctuary, Chaiyapumi Province, 16°3'20"N, 101°34'14"E, 110 meters elevation … ” ( P. nabhitabhatai).</p><p>Differential diagnosis.</p><p>This species is smaller, with less numerous apertural barriers and a narrower umbilicus than Bensonella multidentata sp. nov. B. palatotridens is not spirally striated. This species differs from B. microdentata sp. nov. in the less glossy shell surface, stronger apertural barriers and non-spiniferous aperture surface (including the barriers). See also under B. tamphathai . This species is also similar to B. nitens but can be separated by the much less glossy shell surface and stronger and fewer apertural barriers.</p><p>Distribution.</p><p>This species is known from several localities in Laos (Xieng Khouang, Udomxai and Luang Prabang provinces), Vietnam (Son La province), Myanmar (Shan State, see Tongkerd et al. 2024, Mandalay) as well as Thailand (Chiang Mai, Mae Hong Son and Chaiyapumi provinces).</p><p>Remarks.</p><p>Paraboysidia anguloobtusa described from a Kao Rao Cave, Vieng Phouka District, Luang Namtha Province, Laos (20°43'30.1"N, 101°9'4.3"E) and Paraboysidia nabhitabhatai described from Phadevada, Phukhieo Wildlife Sanctuary, Chaiyapumi Province, Thailand (16°3'20"N, 101°34'14"E) are both herein considered junior synonyms of B. paviei since no morphological differences were observed. Inkhavilay et al. (2016) stated that P. anguloobtusa has a different shell shape than B. paviei and that it has a blunt angular lamella. However, the shell shape of both species is the same (conical to slightly concave-conical) and both species have a blunt angular lamella. Other differences mentioned for apertural barriers (infraparietal situated closer to the parietal and angular lamellae, suprapalatal absent, lamellae more twisted) are considered minor intraspecific variation. The umbilicus in B. paviei is not widely opened as stated by Inkhavilay et al. (2016) but rather narrow and of the same appearance as that in Paraboysidia anguloobtusa . Sometimes, the last whorl of B. paviei is more or less enlarged, but the apertural barrier arrangement is largely consistent. Spiral striation can vary in strength but is always present.</p></div>	https://treatment.plazi.org/id/3A2619A2810D57C7A30768F2911CAA88	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
8E04333ECE0E542BAEA5ACF60204DC97.text	8E04333ECE0E542BAEA5ACF60204DC97.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bensonella perfecta Gojsina & Pall-Gergely 2025	<div><p>Bensonella perfecta Gojšina &amp; Páll-Gergely sp. nov.</p><p>Figs 39 X, 84, 85, 95</p><p>Type material.</p><p>Holotype. Laos • 1 shell (SH: 2.9 mm; SW: 2.1 mm); Central Laos, Xieng Khouang Province, ca 30 km WNW of Phonsavan, ca 47 km ENE of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.86501&amp;materialsCitation.latitude=19.53365" title="Search Plazi for locations around (long 102.86501/lat 19.53365)">Phou Khoun</a>, limestone, black soil in limestone pockets, clay, under rocks in old secondary forest at base of cliff; 19°32.019'N, 102°51.901'E; 1131 m a. s. l.; 15 Nov. 2006; A. Abdou, I. V. Muratov leg.; MNHN -IM-2000-39829.</p><p>Paratypes. Laos • 2 shells; same data as for holotype; CUMZ 14442; 174 shells; same data as for holotype; MNHN -IM-2000-39830 .</p><p>Additional material examined.</p><p>Laos • 94 shells (damaged and juveniles, not paratypes); same data as for holotype; MNHN -IM-2012-25390 .</p><p>Type locality.</p><p>Central Laos, Xieng Khouang Province, ca 30 km WNW of Phonsavan, ca 47 km ENE of Phou Khoun, limestone, black soil in limestone pockets, clay, under rocks in old secondary forest at base of cliff; 19°32.019'N, 102°51.901'E; 1131 m a. s. l.</p><p>Diagnosis.</p><p>Bensonella species with a conical, brownish shell that is not spirally striated. Apertural barriers numerous: five plicae (4 palatals and one basal), all weak and low. Palatal tubercle strong.</p><p>Description.</p><p>Shell conical, pale brown, consisting of 5–5.5 regularly increasing, rounded whorls. Protoconch consisting of 1.75 whorls, same colour as the rest of the shell, finely pitted and spirally striated. Teleoconch with fine pasty like surface sculpture and coarse radial growth lines which are irregularly spaced. Rarely, these radial lines appear as stronger, whitish streaks. A spiralling pattern is very slightly visible on the last whorl (only under SEM) although there are no spiral striae. Last whorl rounded, adnate to the penultimate and slightly descending near the aperture (~ 5–10 ° compared to the shell axis). Apertural profile slightly prosocline to the shell axis. Peristome of same colour as the rest of the shell, expanded but not reflected. There is a weak cervical crest just behind the peristome. Aperture equipped with numerous barriers. Parietal lamella strong and blade-like, not reaching the peristome. Angular lamella closer to the peristome but appears discontinuous as its outer and inner parts are separated by a distinct sinuation. These two parts are roughly equally strong. There are one or two barriers inside the sinulus. Four palatal plicae (upper palatal, interpalatal, lower palatal and infrapalatal). Upper, inter and lower palatal are moderately long and high, equally developed and equally distanced. Sometimes, upper palatal is weak and lower palatal is stronger than others. Infrapalatal plica is much weaker (lower and shorter). Palatal tubercle is strong, sitting on the palatal lip of the peristome just below the line of upper palatal plica. Basal plica situated close to the infrapalatal and equally developed or sometimes weaker. Columellar lamella stronger than in most of the congeners (quite broad and high), almost horizontal, sometimes appears slightly wavy. Infraparietal lamella moderate, weaker than other lamellae but stronger than all the plicae. Surface of all apertural barriers is finely granulated. Additionally, plicae in the palatal and basal areas as well as the infraparietal and angular lamellae are equipped with isolated but strong spines which are positioned medially. Sinulus small and distinctly separated from the rest of the aperture. Umbilicus narrow, slightly elongated, measuring ~ 1 / 5–1 / 6 of the shell width.</p><p>Differential diagnosis.</p><p>See under H. frequens .</p><p>Measurements</p><p>(in mm, n = 5). SH = 2.91–3.09; SW = 2.08–2.23; AH = 1.15–1.30; AW = 1.2–1.37.</p><p>Etymology.</p><p>This species is named for the exceptionally fine surface sculpture.</p><p>Distribution.</p><p>This species is known only from the type locality.</p></div>	https://treatment.plazi.org/id/8E04333ECE0E542BAEA5ACF60204DC97	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
674B841E5AD9520CAA49283BE0141E54.text	674B841E5AD9520CAA49283BE0141E54.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bensonella Pilsbry & Vanatta 1900	<div><p>Genus Bensonella Pilsbry &amp; Vanatta, 1900</p><p>Bifidaria (Bensonella) Pilsbry &amp; Vanatta, 1900: 591.</p><p>Boysidia (Bensonella) — Pilsbry 1917: 198.</p><p>Boysidia (Paraboysidia) Pilsbry, 1917: 174, 201.</p><p>Remarks.</p><p>We have divided the genus Bensonella in two species groups:</p><p>Group 1: Bensonella plicidens group which includes the majority of species. This group can be recognised by the combination of the following traits: i) a distinct palatal tubercle on the palatal lip (Fig. 9); ii) three barriers on the parietal side (angular, parietal, and infraparietal) (Fig. 9). Several species within the Bensonella plicidens group are not typical because they either lack a palatal tubercle or some of the three barriers on the parietal side (see remarks sections under those species).</p><p>Group 2: Bensonella wangviangensis species group includes only four species ( B. wangviangensis, B. cardiostoma sp. nov., B. fracta sp. nov., and B. mitochondria sp. nov.). This group is recognised by the combination of the following traits: i) triangular-conical shell shape; ii) angular lamella as strong as the parietal or stronger, fully reaching the peristome; iii) palatal tubercle not typical but clearly represents a slightly discontinued part of the upper palatal plica (thus, it is of more lamella-like than tubercle-like). This part of the upper palatal plica is most probably homologous to the palatal tubercle found in other Bensonella species; iv) angular lamella and palatal tubercle are situated close together leaving a relatively narrow canal to the sinulus.</p><p>1. Bensonella plicidens group</p><p>Diagnosis. This species group is characterised by a distinct palatal tubercle on the peristome. as well as three barriers on the parietal side (angular, parietal, and infraparietal).</p><p>Remarks. The group includes 32 species distributed from eastern India all the way to northern Vietnam but also present in Japan, China, Taiwan, and south to Indonesia.</p></div>	https://treatment.plazi.org/id/674B841E5AD9520CAA49283BE0141E54	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
BB0DFD4224905C5380E3F1D9D7472CA7.text	BB0DFD4224905C5380E3F1D9D7472CA7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bensonella plicidens (Benson 1849)	<div><p>Bensonella plicidens (Benson, 1849)</p><p>Figs 39 Y, 86</p><p>Pupa plicidens Benson, 1849: 126 .</p><p>Pupa plicidens — Küster 1852: 136, pl. 17, figs 23, 24.</p><p>Boysidia plicidens — Pfeiffer 1853: 553; Hanley and Theobald 1874: pl. 100, fig. 8; Sowerby 1876: pl. 16, fig. 151; Ancey 1881: 373; Gude 1914: 294.</p><p>Vertigo (Odontocyclas) plicidens — Adams and Adams 1858: 173.</p><p>Pupa (Scopelophila) plicidens — Albers and Martens 1860: 296.</p><p>Pupa (Odontocyclas) plicidens — Pfeiffer 1879: 350.</p><p>Bifidaria (Bensonella) plicidens — Pilsbry and Vanatta 1900: 591.</p><p>Bifidaria (Bensonella) landurensis [sic] Pilsbry, 1915: 73.</p><p>Boysidia landurensis — Pilsbry 1917: 204, pl. 35, fig. 9.</p><p>Paraboysidia landourensis — van Benthem Jutting 1950: 39.</p><p>Boysidia (Bensonella) plicidens — Chen and Zhang 2002: 693; Zhang et al. 2014: 572.</p><p>Bensonella plicidens — Hwang 2014: 17–20 (partim: fig. 5 shows B. plicidens); Páll-Gergely and White 2023: 2018–2023, figs 3 a – e, 4, 5, 9 a, b.</p><p>Bensonella landourensis and plicidens — Budha and Backeljau 2017. (partim: figs 2 B, 3, 5 A, F show B. plicidens).</p><p>Type material examined.</p><p>India • lectotype; R. McAndrew coll. (ex. W. H. Benson coll.); UMZC I. 103325 .</p><p>Type locality.</p><p>“ ad Landour et Moussoorie, montibus Himalayanis ”.</p><p>Differential diagnosis.</p><p>This species is similar to B. lakainguta, B. multihami, and B. hooki, but differs from all by the presence of spiral striation on the shell and absence of hooked apertural barriers. Bensonella karoensis is also not spirally striated, is more ovoid with a blunter apex and shallower suture. See also under B. spinosa sp. nov.</p><p>Distribution.</p><p>SW and SE Himalaya.</p></div>	https://treatment.plazi.org/id/BB0DFD4224905C5380E3F1D9D7472CA7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
2950672BE1265E3B98FFBFAE09F1D367.text	2950672BE1265E3B98FFBFAE09F1D367.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bensonella sericata Gojsina & Pall-Gergely 2025	<div><p>Bensonella sericata Gojšina &amp; Páll-Gergely sp. nov.</p><p>Figs 39 Z, 87, 88, 100</p><p>Type material.</p><p>Holotype. Thailand • 1 shell (SH: 2.38 mm, SW: 1.69 mm); Chiang Rai Province, limestone knoll, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.55167&amp;materialsCitation.latitude=20.193611" title="Search Plazi for locations around (long 99.55167/lat 20.193611)">Ban Mae Song Nai</a>, 4.0 km N, 6.0 km NW Mae Chan, limestone domes, shaded ledges; 20°11′37″N, 99°33′6″E; 520 m a. s. l.; 10. May 1988; F. G. Thompson leg.; locality code FGT-4425; UF 380270 . Paratypes. Thailand • 4 shells; same data as for holotype; UF 591360 .</p><p>Additional material examined.</p><p>Thailand • 5 shells (juveniles, not paratypes); same data as for holotype; UF 591368 .</p><p>Type locality.</p><p>Thailand, Chiang Rai Province, limestone knoll, Ban Mae Song Nai, 4.0 km N, 6.0 km NW Mae Chan, limestone domes, shaded ledges; 20°11′37″N, 99°33′6″E; 520 m a. s. l.</p><p>Diagnosis.</p><p>A Bensonella species with conical to conical-ovoid shell, very dense spiral striation, and a small number of apertural barriers, only four of which are strong (parietal, upper palatal, lower palatal and columellar).</p><p>Description.</p><p>Shell brown, conical to conical-ovoid, whorls 4.5–5, convex, regularly increasing. Protoconch consisting of ~ 1.5 whorls, its sculpture consists of ~ 10 roughly equidistant spiral striae. Teleoconch finely, very densely spirally striated, and ornamented with a few, inconspicuous radial growth lines. Last whorl adnate to the penultimate and slightly descending near the aperture (~ 15 ° compared to the shell axis) making the aperture profile prosocline to the shell axis. Peristome slightly expanded, not reflected. A blunt, weak, knob-like palatal tubercle sits anterior and slightly below the upper palatal plica on the peristome edge. Parietal lamella is the strongest in the aperture and slightly leaned towards the palatal wall. Angular lamella weak, its outer part reduced, inner part sits deep in the aperture. Upper palatal plica in form of a very weak swelling, situated slightly above the palatal tubercle. Interpalatal plica strong, located slightly below the palatal tubercle. Lower palatal plica nearly as strong as the parietal, stronger than the interpalatal. There can be two additional swellings: one between the upper and interpalatal and one between the interpalatal and lower palatal. Basal plica absent. Columellar lamella strong and slightly oblique. Infraparietal lamella in form of indistinct swelling located close to the parietal lamella. Surface of all apertural barriers is smooth. Sinulus wide and relatively well separated from the rest of the aperture due to the strong parietal lamella. Umbilicus moderately wide, measuring 1 / 5 of shell width.</p><p>Differential diagnosis.</p><p>This species differs from B. multidentata sp. nov. by the less numerous apertural barriers (including the lack of the basal plica) and not concave-conical shell. The shell in B. multidentata sp. nov. is also more pointed. Bensonella nitens sp. nov. is glossier due to the very coarse spiral striae (very dense in B. sericata sp. nov.), concave conical and has slender and sharp projections proximally and distally from the palatal plicae. See also under B. tamphathai .</p><p>Measurements</p><p>(in mm, n = 4). SH = 2.26–2.39; SW = 1.69–1.73; AH = 0.94–1.01; AW = 0.87–0.94.</p><p>Etymology.</p><p>The specific epithet refers to the silky surface of the shell (as if “ covered in silk ”).</p><p>Distribution.</p><p>This species is known only from the type locality.</p><p>Remarks.</p><p>Coordinates of the sample point in Myanmar just next to the Thai border.</p></div>	https://treatment.plazi.org/id/2950672BE1265E3B98FFBFAE09F1D367	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
F1476E8059305973AD61C74A6F4A36EE.text	F1476E8059305973AD61C74A6F4A36EE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bensonella serrata Gojsina, Hunyadi & Pall-Gergely 2025	<div><p>Bensonella serrata Gojšina, Hunyadi &amp; Páll-Gergely sp. nov.</p><p>Type material.</p><p>Holotype. Myanmar • 1 shell (SH: 2.71 mm, SW: 1.92 mm); Shan State, 16 km from centre of Taunggyi towards Hopong, 1.5 km along road # 4, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.137314&amp;materialsCitation.latitude=20.787716" title="Search Plazi for locations around (long 97.137314/lat 20.787716)">Shwe Pyi Aunchonda</a> monastery; 20°47.263'N, 97°8.239'E; 1110 m a. s. l.; 08 Oct. 2018; A. Hunyadi, K. Okubo &amp; J. U. Otani leg.; CUMZ 14443 . Paratypes. Myanmar • 15 shells; same data as for holotype; coll. HA • 13 shells; Shan State, Hopong, Sam Phu, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.269966&amp;materialsCitation.latitude=20.685051" title="Search Plazi for locations around (long 97.269966/lat 20.685051)">Cave</a> Ae- 5 at ridge above village <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.269966&amp;materialsCitation.latitude=20.685051" title="Search Plazi for locations around (long 97.269966/lat 20.685051)">Ho Hwe</a>; 20°41.103'N, 97°16.198'E; 02 Feb. 2019; J. Grego leg., coll. JG .</p><p>Additional material examined.</p><p>Myanmar • 3 shells (damaged, not paratypes); same data as for holotype; coll. HA • 5 shells (damaged / juveniles, not paratypes); Shan State, Hopong, Sam Phu, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.269966&amp;materialsCitation.latitude=20.685051" title="Search Plazi for locations around (long 97.269966/lat 20.685051)">Cave</a> Ae- 5 at ridge above village <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.269966&amp;materialsCitation.latitude=20.685051" title="Search Plazi for locations around (long 97.269966/lat 20.685051)">Ho Hwe</a>; 20°41.103'N, 97°16.198'E; 02 Feb. 2019; J. Grego leg., coll. JG .</p><p>Type locality.</p><p>Myanmar, Shan State, 16 km from centre of Taunggyi towards Hopong, 1.5 km along road # 4, Shwe Pyi Aunchonda monastery; 20°47.263'N, 97°8.239'E; 1110 m a. s. l.</p><p>Diagnosis.</p><p>Bensonella species with pasty shell surface, last whorl adnate to the penultimate and palatal plicae with serrated, elevated inner and very low outer parts.</p><p>Description.</p><p>Shell triangular, slightly conical-ovoid, brown, not very glossy, consisting of 5–5.5 regularly increasing whorls separated by a deep suture. All whorls convex, rounded. Protoconch consisting of 1.5 whorls, finely pitted, showing spiralling pattern even though without clear spiral striae. The boundary between the protoconch and the teleoconch is visible due to the presence of radial growth lines on the teleoconch. Teleoconch surface finely dimpled, pasty and with less numerous, weak radial growth lines, spiral striae absent. Last whorl also rounded, adnate to the penultimate near the aperture. Last whorl slightly descending immediately behind the aperture (~ 10 ° compared to the shell axis) making the apertural profile slightly prosocline to the shell axis. Peristome weakly expanded but not reflected, its edges are thickened. Aperture equipped with numerous barriers, two of them relatively strong (parietal, columellar), others weak. Parietal lamella is the strongest and highest in the aperture. Angular lamella is the longest in the aperture, much longer than the parietal, very slender and low but continuous. Its inner and outer parts are higher than its middle part. Columellar lamella similar in length to the parietal but much weaker. There is one weak infraparietal lamella. On the palatal side, there are four main palatal plicae which are all almost equally strong and long (upper palatal, two interpalatals, and a lower palatal). They are all elevated at their inner ends and sloping towards the outer end, ending very low. These low parts are equipped with numerous small spines (serrated), thus resembling a saw. Palatal tubercle, sitting on the palatal lip, is weak but clearly present. There are several suprapalatal plicae (with similar morphology as in the main palatals) all weak and located inside the sinulus (the length and number (usually two to four) of these plicae is variable and not useful in species identification). There are usually two basal plicae (although one of them could be homologous with the infrapalatal plica), which are long but can occasionally be shorter. Surface of all barriers is finely granulated. Sinulus not strongly isolated from the rest of the aperture due to the low angular lamella and weak palatal tubercle. Umbilicus is narrow, measuring 1 / 8 of the shell width.</p><p>Differential diagnosis.</p><p>This species can be separated from its congeners by the presence of equally strong palatal plicae with elevated inner and very low outer portions equipped with numerous spines.</p><p>Measurements</p><p>(in mm, n = 5). SH = 2.52–2.86; SW = 1.80–2.06; AH = 0.94–1.03; AW = 0.83–0.94.</p><p>Etymology.</p><p>This species is named for the serrated (saw-like) palatal plicae.</p><p>Distribution.</p><p>This species is known from two localities situated ca 18 km apart (straight-line distance).</p></div>	https://treatment.plazi.org/id/F1476E8059305973AD61C74A6F4A36EE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
85FB2666144451FFA1AD40E199238617.text	85FB2666144451FFA1AD40E199238617.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bensonella spelaea Gojsina, Grego & Pall-Gergely 2025	<div><p>Bensonella spelaea Gojšina, Grego &amp; Páll-Gergely sp. nov.</p><p>Type material.</p><p>Holotype. Myanmar • 1 shell (SH: 3.3 mm, SW: 2.4 mm); Kayah State, Hpruso district, Maw Thi Do Village, entrance of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.04283&amp;materialsCitation.latitude=19.379066" title="Search Plazi for locations around (long 97.04283/lat 19.379066)">Phruno river</a> cave; 19°22.744'N, 97°2.570'E; 12 Feb. 2019; J. Grego leg.; CUMZ 14444 . Paratypes. Myanmar • 7 shells; same data as for holotype; coll. JG • 1 shell; same data as for holotype; coll. HA .</p><p>Type locality.</p><p>Myanmar, Kayah State, Hpruso district, Maw Thi Do Village, entrance of Phruno river cave; 19°22.744'N, 97°2.570'E.</p><p>Diagnosis.</p><p>Bensonella species with 7–9 apertural barriers, most peculiar of these is the transversal plica (strong, high but with a concave frontal surface resembling a cave entrance).</p><p>Description.</p><p>Shell triangular, conical-ovoid, brownish, consisting of 5–5.5 convex, regularly increasing, rounded whorls separated by a deep suture. Boundary between the protoconch and the teleoconch is only clearly visible after SEM imaging because of the delicate sculpture and somehow weathered type series. Protoconch is spirally striated, probably ~ 1.75 whorls). Teleoconch of fine, pasty sculpture with dense but fine radial growth lines and without signs of spiral striation. Last whorl rounded, adnate to the penultimate. Aperture profile sinuated in its upper portion (concave) but convex in its middle and lower sections. Peristome expanded, especially at the parietal side where it leans on the penultimate whorl in form of a strong callus but not reflected. There is a moderate cervical crest located a short distance behind the peristome. Aperture equipped with relatively small number of barriers, seven to nine. Angular, parietal, upper palatal, transversal, palatal tubercle, columellar and infraparietal are always present, while basal and supracolumellar only sometimes. Parietal lamella heart-shaped in profile due to the sinuation located in its middle part (this makes the middle part of the lamella lower than outer and inner parts). Angular lamella also appears curvy, and it is curved to the palatal side. Upper palatal plica relatively weak and low. Transversal plica large (the largest in the aperture), high but with a strongly concave frontal surface (this gives the appearance of the transversal plica as an entrance to a cave). Basal very short and low when present. Columellar lamella very strong and thick (roughly the same as the parietal lamella). Supracolumellar lamella weak, dot like (sometimes absent). Infraparietal lamella present but much weaker than both the parietal and the columellar. Surface of all apertural barriers is granulated to almost spiniferous. Sinulus small and distinctly separated from the rest of the aperture due to the strong and closely positioned angular lamella and palatal tubercle. Umbilicus initially narrow and the widening at the last whorl, elongated, measuring ~ 1 / 6–1 / 7 of the shell width.</p><p>Differential diagnosis.</p><p>This species is the largest one with a transversal plica. This plica shows the peculiar appearance with the strongly concave frontal surface resembling a cave entrance.</p><p>Measurements</p><p>(in mm, n = 5). SH = 2.48–3.38; SW = 1.98–2.4; AH = 1.08–1.30; AW = 0.95–1.12.</p><p>Etymology.</p><p>Named after the cave-like transversal plica.</p><p>Distribution.</p><p>This species is known only from the type locality.</p></div>	https://treatment.plazi.org/id/85FB2666144451FFA1AD40E199238617	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
86E9288D36235017999C4A2A38DBE0DA.text	86E9288D36235017999C4A2A38DBE0DA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bensonella spinosa Gojsina, Hunyadi & Pall-Gergely 2025	<div><p>Bensonella spinosa Gojšina, Hunyadi &amp; Páll-Gergely sp. nov.</p><p>Type material.</p><p>Holotype. Myanmar • 1 shell (SH: 2 mm, SW: 1.5 mm); Shan State, 5.7 km south-southwest from centre of Pinlaung, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=96.77053&amp;materialsCitation.latitude=20.0692" title="Search Plazi for locations around (long 96.77053/lat 20.0692)">Wingabar Taung</a>; 20°4.152'N, 96°46.232'E; 1510 m a. s. l.; 04 Oct. 2018; A. Hunyadi, K. Okubo &amp; J. U. Otani leg.; CUMZ 14445 . Paratypes. Myanmar • 14 shells; same data as for holotype; coll. HA .</p><p>Type locality.</p><p>Myanmar, Shan State, 5.7 km south-southwest from centre of Pinlaung, Wingabar Taung; 20°4.152'N, 96°46.232'E; 1510 m a. s. l.</p><p>Diagnosis.</p><p>Bensonella with triangular-conical, reddish-brown shell. Teleoconch fine, pasty, not spirally striated. All barriers and aperture surface around them very roughly spiniferous.</p><p>Description.</p><p>Shell triangular, slightly conical-ovoid, reddish brown, consisting of 5–5.5 convex, rounded whorls separated by a deep suture. Protoconch of ~ 1.5 whorls, finely pitted, showing very weak spiralling pattern, lighter than the rest of the shell but with no clearly visible boundary with the teleoconch. Teleoconch finely dimpled (pasty) and finely radially striated but without spiral striae. Last whorl adnate to the penultimate and very slightly ascending near the aperture (~ 5–10 ° compared to the shell axis), making the aperture profile weakly opisthocline to the shell axis. Peristome of the same colour as the rest of the shell, expanded and not reflected. Aperture equipped with numerous barriers. Parietal lamella is the strongest in the aperture, high but not curved, directed towards the palatal wall. Angular lamella slightly lower than the parietal but otherwise very similar, continuous. There are three main palatal plicae (upper palatal, interpalatal and lower palatal) and one additional small interpalatal plica situated below the stronger interpalatal and a lower palatal. Upper palatal as strong as the main interpalatal. Lower palatal plica is the strongest. Infrapalatal plica weak. Basal plica 2 × stronger than the infrapalatal. All palatal plicae are higher at their inner parts and sloping towards their outer parts. There is a strong palatal tubercle sitting on the palatal lip of the peristome, in front of the upper palatal plica. Peristome is distinctly sinuated behind this tubercle. Columellar lamella ~ 2 × stronger than the basal and positioned obliquely, directed towards the parietal lamella. Subcolumellar similar to the infrapalatal. Supracolumellar lamella similar to the subcolumellar. Infraparietal lamella strong as the columellar. All apertural barriers are very strongly spiniferous as well as the surface of the aperture around them. Sinulus small and well separated from the rest of the aperture due to the strong angular lamella. Umbilicus very narrow, dot like.</p><p>Differential diagnosis.</p><p>This species is most similar to B. plicidens from which it differs by the absence of spiral striation as well as spiniferous barriers and aperture surface (both smooth in B. plicidens). Bensonella lakainguta, B. multihami and B. hooki all have more numerous and hooked apertural barriers and they are not spiniferous.</p><p>Measurements</p><p>(in mm, n = 5). SH = 1.81–2.14; SW = 1.37–1.5; AH = 0.66–0.79; AW = 0.63–0.75.</p><p>Etymology.</p><p>The specific epithet is due to the numerous spines in the aperture.</p><p>Distribution.</p><p>This species is known only from the type locality.</p></div>	https://treatment.plazi.org/id/86E9288D36235017999C4A2A38DBE0DA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
E5B7E7D8D8AE5096BD83A826BFED1EC7.text	E5B7E7D8D8AE5096BD83A826BFED1EC7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bensonella taiyaiorum Tongkerd & Panha 2024	<div><p>Bensonella taiyaiorum Tongkerd &amp; Panha, 2024</p><p>Bensonella taiyaiorum Tongkerd &amp; Panha in Tongkerd et al. 2024: 174–176, figs 7 B, 8, 13 H.</p><p>Type material examined.</p><p>Myanmar • holotype; collector unknown; CUMZ 14380 .</p><p>Type locality.</p><p>“ Dragon Rock, Pindaya Township, Taunggyi District, Shan State, Myanmar (20°55'31.5"N, 96°39'01.2"E; 1300 m a. s. l.) ”.</p><p>Differential diagnosis.</p><p>This species differs from B. lophiodera by the stronger apertural barriers, narrower umbilicus, and the lack of a sharp cervical crest. Bensonella dracula sp. nov. is smaller, has a second palatal tubercle and lacks the suprapalatal plica.</p><p>Distribution.</p><p>This species is known only from the type locality.</p></div>	https://treatment.plazi.org/id/E5B7E7D8D8AE5096BD83A826BFED1EC7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
13D456061CD557038CD69312C6349E9E.text	13D456061CD557038CD69312C6349E9E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bensonella tamphathai (Panha & J. B. Burch 2000)	<div><p>Bensonella tamphathai (Panha &amp; J. B. Burch, 2000)</p><p>Paraboysidia tamphathai Panha &amp; Burch, 2000: 107, fig. 2.</p><p>Paraboysidia tamphathai — Panha and Burch 2008: 115, fig. 98.</p><p>Bensonella tamphathai — Dumrongrojwattana and Wongkamhaeng 2024: 324.</p><p>Material examined.</p><p>Thailand • 4 shells; Lampang Province, limestone ridge, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.842224&amp;materialsCitation.latitude=18.585278" title="Search Plazi for locations around (long 99.842224/lat 18.585278)">Ban Pang La</a>; 18°35′7″N, 99°50′32″E; 400 m a. s. l.; 14. May 1988; F. G. Thompson leg.; locality code FGT-4438; UF 346895 • 11 shells; Lampang Province, limestone knoll <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.86972&amp;materialsCitation.latitude=18.554167" title="Search Plazi for locations around (long 99.86972/lat 18.554167)">1 km NE of Ban Pang La</a>; 18°33′15″N, 99°52′11″E; 400 m a. s. l.; 14. May 1988; F. G. Thompson leg.; locality code FGT-4436; UF 346878 • 14 shells; same data as previous; locality code FGT-4435; UF 380388 • 3 shells; Lampang Province, limestone mtn. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.71361&amp;materialsCitation.latitude=18.234167" title="Search Plazi for locations around (long 99.71361/lat 18.234167)">3 km SE of Ban Mae Moa</a> ledge on NW slope of mtn., leaf litter; 18°14′3″N, 99°42′49″E; 500 m a. s. l.; 15. May 1988; F. G. Thompson leg.; locality code FGT-4443; UF 346921 • 2 shells; Lampang Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.851944&amp;materialsCitation.latitude=17.964724" title="Search Plazi for locations around (long 98.851944/lat 17.964724)">6 km SW Mae Pak</a>, limestone hill, 17°57′53″N, 98°51′7″E (provided coordinates point in Lamphun province); 600 m a. s. l.; 13 June 1987; F. G. Thompson leg.; locality code FGT-4328; UF 346579 .</p><p>Type locality.</p><p>“ Tam Pha Thai National Park, Lampang Province, 18°36'20"N, 99°53'49"E, 490 meters elevation ” (Thailand).</p><p>Differential diagnosis.</p><p>The most similar species is B. paviei which is however smaller and has less pronounced spiral sculpture. Apertural barriers are also more numerous than in B. paviei . Bensonella palatotridens is not spirally striated. Bensonella tamphathai has a blunter conical shell (i. e., more barrel-shaped), narrower umbilicus and fewer basal plicae than B. multidentata sp. nov. Bensonella multidentata sp. nov. is also much more densely spirally striated which makes it shell surface less shiny than in B. tamphathai . Bensonella nitens sp. nov. has a wider umbilicus, is more concave-conical and has a thin, sharp, and long projections of palatal plicae (proximally and distally). Also, the last whorl is B. nitens sp. nov. is much narrower than in B. tamphathai (i. e., it contributes less to the total shell height than in B. tamphathai). Bensonella sericata sp. nov. is much more densely spirally striated, lacks the basal plica and has weaker upper palatal plica. Bensonella microdentata sp. nov. has weaker apertural barriers which are roughly spiniferous.</p><p>Distribution.</p><p>This species is known from Lampang Province, Thailand.</p></div>	https://treatment.plazi.org/id/13D456061CD557038CD69312C6349E9E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
25F2B0EE465256B4A40293225CF307F8.text	25F2B0EE465256B4A40293225CF307F8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bensonella undefined-1	<div><p>Bensonella sp. 1</p><p>Material examined.</p><p>Thailand • 1 shell; Phangnga Province, 32 km NE Khok Kloi, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.45&amp;materialsCitation.latitude=8.45" title="Search Plazi for locations around (long 98.45/lat 8.45)">Wat Tham Suwan Khuha</a>; 8°27'N, 98°27'E; 27 Mar. 1988; 20 m a. s. l.; K. Auffenberg leg.; locality code KA-0601; UF 345752 .</p><p>Remarks.</p><p>Only a single, weathered shell was in this sample which made the precise idenitifcation impossible. Based on the morphology of apertural barriers, they were possibly hooked.</p></div>	https://treatment.plazi.org/id/25F2B0EE465256B4A40293225CF307F8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
2011362CF1845D21BCE9062C3F325157.text	2011362CF1845D21BCE9062C3F325157.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bensonella undefined-2	<div><p>Bensonella sp. 2</p><p>Material examined.</p><p>Thailand • 1 shell; Chiang Rai prov., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.85778&amp;materialsCitation.latitude=20.369446" title="Search Plazi for locations around (long 99.85778/lat 20.369446)">Tham Tumluang cave</a>, 7 km S, 2 k W of Mae Sai. Around caves, leaf litter; 20°22.1667'N, 99°51.4667'E; 08. May 1988; F. G. Thompson leg.; locality code FGT-4419; UF 380223 .</p><p>Remarks.</p><p>A single, damaged specimen was in this sample which made its identification impossible.</p></div>	https://treatment.plazi.org/id/2011362CF1845D21BCE9062C3F325157	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
4F4D5B05735E58788058202718A703A4.text	4F4D5B05735E58788058202718A703A4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bensonella undefined-3	<div><p>Bensonella sp. 3</p><p>Fig. 108</p><p>Material examined.</p><p>Laos • Luang Namtha Province, Luang Namtha ca 43.8 km southwest – Vieng Phou Kha, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.18502&amp;materialsCitation.latitude=20.74213" title="Search Plazi for locations around (long 101.18502/lat 20.74213)">Phou Lan, right side of the road 100 m</a>; 20°44.5278'N, 101°11.1011'E; 770 m a. s. l.; 08 Oct. 2019; A. Hunyadi, K. Okubo, J. U. Otani leg., coll. HA .</p><p>Remarks.</p><p>Only a single, rather weathered, and partly damaged specimen of Bensonella which is the most similar to B. lakainguta and related species ( B. hooki, B. montawa sp. nov., B. multihami) could be examined from Laos. Most of the aperture is obstructed by the dirt that could not be cleaned which is why majority of the barriers were invisible. The identity of this species remains unresolved until more specimens are collected.</p></div>	https://treatment.plazi.org/id/4F4D5B05735E58788058202718A703A4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
81B09EC7FFAF5287BFE1EC2B8F089434.text	81B09EC7FFAF5287BFE1EC2B8F089434.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bensonella wangviangensis (Panha & Tongkerd 2003)	<div><p>2. Bensonella wangviangensis group</p><p>Diagnosis.</p><p>This species group is characterised by angular lamella as strong as the parietal or even stronger, fully reaching the peristome edge, palatal tubercle not in its typical form but clearly represents a slightly discontinued part of the upper palatal plica (thus, it is of more lamella-like than tubercle-like appearance) and closely situated angular lamella and palatal tubercle which clearly separated the sinulus from the rest of the aperture.</p><p>Remarks.</p><p>Four species are assigned to this group and they inhabit northern Thailand (Loei Province) as well as adjacent territories in Laos (Vientiane and Khammouane provinces). Due to the same distribution and very similar overall shell morphology, these species are probably closely related.</p></div>	https://treatment.plazi.org/id/81B09EC7FFAF5287BFE1EC2B8F089434	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
CBE9A7CC1E8358C7BB3297846454363A.text	CBE9A7CC1E8358C7BB3297846454363A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bensonella wangviangensis (Panha & Tongkerd 2003)	<div><p>Bensonella wangviangensis (Panha &amp; Tongkerd, 2003)</p><p>Figs 103 D, 106, 107</p><p>Paraboysidia wangviangensis Panha &amp; Tongkerd in Panha et al. 2003: 123–128, figs 2–3.</p><p>Paraboysidia wangviangensis — Inkhavilay et al. 2019: 62.</p><p>Type material examined.</p><p>Laos • 1 paratype; from the type locality; 1998; S. Panha leg.; CUMZ ver. 089 .</p><p>Type locality.</p><p>“ Tam Chang Cave, Wangviang, Laos, 18°54′47″N, 102°26′33″E, 140 meters elevation … ” (Laos).</p><p>Differential diagnosis.</p><p>This species differs from B. cardiostoma sp. nov. and B. mitochondria sp. nov. by the more elongated shell, larger size as well as depressed and more elongated aperture. Additionally, B. cardiostoma sp. nov. is not spirally striated. Apart from the clearly smaller size, this species can further be separated from B. fracta sp. nov. by the shouldered last whorl, which is rounded in B. fracta sp. nov.</p><p>Distribution.</p><p>This species is known only from the type locality.</p><p>Remarks.</p><p>We examined the types of this species and could add several notes to the original description. This species is originally described as having six apertural barriers (angular, parietal, infraparietal, two upper palatals and a lower palatal). From the Fig. 106, it can be clearly seen that there are eight barriers, the ones already described and additionally one interpalatal and a columellar lamella. The “ second upper palatal plica ” from the original description is in the position of usually mentioned palatal tubercle (typical for Bensonella), although it does not look like a tubercle but more like a proper lamella. Nevertheless, we treat them homologous.</p></div>	https://treatment.plazi.org/id/CBE9A7CC1E8358C7BB3297846454363A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
E1CB45B827A65B21BF4D022023DB33F5.text	E1CB45B827A65B21BF4D022023DB33F5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Boysidia Ancey 1881	<div><p>Genus Boysidia Ancey, 1881</p><p>Pupa (Boysidia) Ancey, 1881: 373 .</p><p>Diagnosis.</p><p>The genus Boysidia remains unrevised. This is urgently needed because many species are misunderstood due to poor original descriptions / drawings or the uncertain status of type specimens. The genus currently includes the following species: Boysidia chiangmaiensis Panha &amp; J. B. Burch, 2002, B. conspicua (Möllendorff, 1885), B. dilamellaris D. - N. Chen, Y. - H. Liu &amp; W. - X. Wu, 1995, B. dorsata (Ancey, 1881), B. gracilis F. Haas, 1937, B. guiyangensis T. - C. Luo, D. - N. Chen, G. - Q. Zhang, La. Zhang, Li Zhang &amp; T. - W. Li, 1998, B. hangchowensis (Pilsbry &amp; Y. Hirase, 1908), B. houaphanica Inkhavilay &amp; Sutcharit, 2024, B. huangguoshuensis T. - C. Luo, D. - N. Chen &amp; G. - Q. Zhang, 2000, B. hunana (Gredler, 1881), B. jinpingensis M. Tian, B. Fao &amp; Y. - X. Chen, 2015, B. nanjiangensis Z. - L. Zhang, W. - H. Zhang &amp; T. - C. Luo, 2011, B. orientalis B. Rensch, 1935, B. phatangensis Dumrongrojwattana &amp; Assawawattagee, 2018, B. ringens van Benthem Jutting, 1950; B. robusta Bavay &amp; Dautzenberg, 1912, B. shilinensis D. - N. Chen, M. Wu &amp; G. - Q. Zhang, 1999, B. strophostoma (Möllendorff, 1885), B. taibaiensis D. - N. Chen, M. Wu &amp; G. - Q. Zhang, 1999, B. tamtouriana Pokryszko &amp; Auffenberg, 2009, B. tholos Panha &amp; J. B. Burch, 2002, B. tongguanensis D. - N. Chen &amp; W. - H. Zhang, 2002, B. xianfengensis W. - H. Zhang, D. - N. Chen &amp; W. - C. Zhou, 2014, B. xiaoguanensis W. - H. Zhang, D. - N. Chen &amp; W. - C. Zhou, 2014, B. xishanensis D. - N. Chen, M. Wu &amp; G. - Q. Zhang, 1999, B. xiuwenensis W. - H. Zhang, T. - C. Luo &amp; W. - C. Zhou, 2010 .</p><p>The species listed below are excluded from the genus Bensonella and transferred to Boysidia due to the following traits (not shared among all these species): i) apertural dentition more similar to Boysidia (columellar lamella very strong and long, absence of three barriers on the parietal side); ii) surface sculpture similar to Boysidia (radial growth lines, no spiral striae); iii) larger shell sizes (frequently over 4 mm), which is more common in Boysidia representatives.</p></div>	https://treatment.plazi.org/id/E1CB45B827A65B21BF4D022023DB33F5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
ACA175368885544E829597B068E1C844.text	ACA175368885544E829597B068E1C844.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Boysidia gittenbergeri (Maassen 2008) Gojšina & Hunyadi & Sutcharit & Tongkerd & Auffenberg & Grego & Vermeulen & Reischütz & Páll-Gergely 2025	<div><p>Boysidia gittenbergeri (Maassen, 2008) comb. nov.</p><p>Paraboysidia gittenbergeri Maassen, 2008: 237–239, figs 5, 6.</p><p>Paraboysidia gittenbergeri — Inkhavilay et al. 2016: 215–216, figs 1, 2 A – C, 4 A, tables 1, 2; Inkhavilay et al. 2019: 61, fig. 27 A.</p><p>Material examined.</p><p>None.</p><p>Type locality.</p><p>“ NW Laos, province Luang Namtha, district Vieng Phouka, Oung (= Khama word for cave) Pra Ngiene, 500 m SE of Ban (= village) Phou Lek, at the entrance of the cave ”.</p><p>Remarks.</p><p>We have excluded this species from Bensonella and classified it in the genus Boysidia due to the combination of the following traits: i) large shell size (nearly 6 mm); ii) absence of the palatal tubercle typical for Bensonella; iii) shell shape more similar to some Boysidia species (e. g., B. chiangmaiensis, B. tholos, B. lamothei).</p></div>	https://treatment.plazi.org/id/ACA175368885544E829597B068E1C844	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
F896FB6E8C4E59BA880CF22CD4EB712C.text	F896FB6E8C4E59BA880CF22CD4EB712C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Boysidia hupeana (Gredler 1901)	<div><p>Boysidia hupeana (Gredler, 1901)</p><p>Figs 109</p><p>Hypselostoma hupeanum Gredler, 1901: 151.</p><p>Boysidia (Paraboysidia) hupeana — Pilsbry 1917: 206; Haas 1937: 8–9.</p><p>Paraboysidia hupeana — van Benthem Jutting 1950: 38; Zilch 1984: 165.</p><p>Paraboysidia kitteli Maassen, 1999: 123, figs 5–7. syn. nov.</p><p>Boysidia (Paraboysidia) hubeina [sic] — Chen and Zhang 2002: 692.</p><p>Boysidia (Boysidia) gongyaoshanensis Yang, Zhang &amp; Chen, 2012: 550 (English description), 550–551 (Chinese description), figs 1–3. syn. nov.</p><p>Type material examined.</p><p>Indonesia • 1 paratypes of P. kitteli; from the type locality; 17 Aug. 1993; ex. coll. Hemmen, Wiesbaden; coll. PGB • 1 paratype of P. kitteli; from the type locality; 16 June 1990; ex. coll. Hemmen, Wiesbaden; coll. PGB • 4 paratypes of P. kitteli; from the type locality; Aug. 1993; ex. coll. W. J. M. Maassen; coll. PGB .</p><p>Additional material examined.</p><p>China • 1 shell; Badung, Hubei, coll. Moellendorff; SMF 10775 .</p><p>Type localities.</p><p>“ Südwest Hupé ”, China ( B. hupeana); “ Gongyao Hill, Xingyi County (25°N, 104°08'E), Guizhou province, China ” ( B. gongyaoshanensis); “ N. Sumatra, Karo Highlands, Kuta Buluh, 40 km N of Brastagi, near the entrance of the cave Liangdehar in leaf litter at the foot of limestone rocks ” ( B. kitteli).</p><p>Remarks.</p><p>Boysidia (Boysidia) gongyaoshanensis, described from Gongyao Hill, Xingyi County (25°N, 104°08'E), Guizhou province, China, is herein treated a junior synonym of B. hupeana since no morphological differences were noticed. Also, no differences in shell morphology were noticed between B. kitteli and B. hupeana even though they are described from localities which are ca 3200 km from each other. Even though the distance is enormous, they are treated conspecific due to the lack of conchological differences. Both new synonyms were not compared to B. hupeana in their respective original descriptions (Maassen 1999; Yang et al. 2012).</p></div>	https://treatment.plazi.org/id/F896FB6E8C4E59BA880CF22CD4EB712C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
63963AC6652A566AA731262CF2E57043.text	63963AC6652A566AA731262CF2E57043.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Boysidia lamothei Bavay & Dautzenberg 1912	<div><p>Boysidia lamothei Bavay &amp; Dautzenberg, 1912</p><p>Boysidia lamothei Bavay &amp; Dautzenberg, 1912: 21, pl. 3, figs 7–9.</p><p>Boysidia (Paraboysidia) lamothei — Pilsbry 1917: 202–203, pl. 35, figs 3–6; Schileyko 2011: 2.</p><p>Paraboysidia lamothei — van Benthem Jutting 1950: 39.</p><p>Type material examined.</p><p>Vietnam • syntype; MNHN -IM-2000-35160 .</p><p>Type locality.</p><p>“ Ban-Lao ”, Vietnam.</p><p>Remarks.</p><p>This species is treated as a member of Boysidia due to the shell shape being more similar to some other congeners such as B. robusta and B. jinpingensis and a strong and oblique columellar lamella which extends to the expanding peristome.</p></div>	https://treatment.plazi.org/id/63963AC6652A566AA731262CF2E57043	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
26135924C7615D7EA8060B6583719D01.text	26135924C7615D7EA8060B6583719D01.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Boysidia paralella (Inkhavilay & Panha 2016) Gojšina & Hunyadi & Sutcharit & Tongkerd & Auffenberg & Grego & Vermeulen & Reischütz & Páll-Gergely 2025	<div><p>Boysidia paralella (Inkhavilay &amp; Panha, 2016) comb. nov.</p><p>Paraboysidia paralella Inkhavilay &amp; Panha in Inkhavilay et al. 2016: 220, figs 1, 3 A – C, 4 C.</p><p>Paraboysidia paralella — Inkhavilay et al. 2019: 62, fig. 27 B.</p><p>Material examined.</p><p>Laos • holotype; CUMZ 7059 .</p><p>Type locality.</p><p>“ Limestone wall near the entrance of Kao Rao Cave, Vieng Phouka District, Luang Namtha Province, Laos (20°43'30.1"N, 101°9'4.3"E), 732 m amsl ”.</p><p>Remarks.</p><p>This species is assigned to Boysidia because of similar shell shape, surface sculpture and apertural barrier arrangement (especially strong columellar lamella) to some Boysidia species such as B. gittenbergeri, and B. hupeana .</p></div>	https://treatment.plazi.org/id/26135924C7615D7EA8060B6583719D01	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
78541F9F33E659D68EAC2028999B26AD.text	78541F9F33E659D68EAC2028999B26AD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Boysidia ringens van Benthem Jutting 1950	<div><p>Boysidia ringens van Benthem Jutting, 1950</p><p>Fig. 110</p><p>Boysidia ringens van Benthem Jutting, 1950: 11–12, fig. 3.</p><p>Type material examined.</p><p>Malaysia • 23 paratypes; from the type locality; M. W. F. Tweedie leg.; RMNH Moll.137127 .</p><p>Additional material examined.</p><p>Malaysia • 7 shells; Kedah, Gunung Keriang (N of Alor Setar), Oct. 1998, Hemmen, leg. Wiesbaden ex. coll.; coll. PGB • 1 shell; Kedah, Gunung Keriang, 7.5 km NW of Alor Setar; ex. coll. Maassen; coll. PGB .</p><p>Type locality.</p><p>“ Mount Keriang, Kedah ”, Malaysia.</p><p>Cross-diagnosis.</p><p>This species is not similar to any other congener.</p><p>Distribution.</p><p>This species is known only from the type locality.</p></div>	https://treatment.plazi.org/id/78541F9F33E659D68EAC2028999B26AD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
E7FD78A318A85E2FAE11487531096348.text	E7FD78A318A85E2FAE11487531096348.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Boysidia salwiniana (Theobald 1870)	<div><p>Boysidia salwiniana (Theobald, 1870)</p><p>Pupa salwiniana Theobald, 1870: 400 .</p><p>Pupa salwiniana — Hanley and Theobald 1874: pl. 100, fig. 9; Sowerby 1876: pl. 16, fig. 150; Nevill 1878: 23.</p><p>Pupa salwineana [sic] — Godwin-Austen 1888: 244.</p><p>Boysidia (Paraboysidia) salwiniana — Pilsbry 1917: 206–208, pl. 33, fig. 11.</p><p>Paraboysidia salwiniana — van Benthem Jutting 1950: 40.</p><p>Bensonella salwiniana — Tongkerd et al. 2024: 167–171, figs 4, 5, 6 A, 13 F.</p><p>Material examined.</p><p>Myanmar • 1 shell; Burma NHMUK 1912.4. 16.66.</p><p>Type locality.</p><p>“ Shan States ”, Myanmar.</p><p>Remarks.</p><p>Tongkerd et al. (2024) provided the first description of the genitalia and radula of this species. They have also pointed out that the type specimens could not be located.</p></div>	https://treatment.plazi.org/id/E7FD78A318A85E2FAE11487531096348	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
4007E7BA430456AE915078EC0F36563A.text	4007E7BA430456AE915078EC0F36563A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Boysidia shuitianbaensis T. - C. Luo, D. - N. Chen, G. - Q. Zhang, La. Zhang, Li Zhang & T. - W. Li 1998	<div><p>Boysidia shuitianbaensis T. - C. Luo, D. - N. Chen, G. - Q. Zhang, La. Zhang, Li Zhang &amp; T. - W. Li, 1998</p><p>Boysidia (Bensonella) shuitianbaensis Luo et al., 1998: 2 (Chinese description), 3–4 (English description), fig. 2.</p><p>Material examined.</p><p>None.</p><p>Type locality.</p><p>“ Collected from Shuitianba town (26°7'N, 106°8'E), Guiyang City, Guizhou Province, China ”.</p><p>Remarks.</p><p>According to the original description, this species should be assigned to Boysidia due to the presence of only four barriers and a merged angulo-parietal lamella.</p></div>	https://treatment.plazi.org/id/4007E7BA430456AE915078EC0F36563A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
3057154D05485D0A908072205C62B448.text	3057154D05485D0A908072205C62B448.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma adela (F. G. Thompson & Upatham 1997) Gojšina & Hunyadi & Sutcharit & Tongkerd & Auffenberg & Grego & Vermeulen & Reischütz & Páll-Gergely 2025	<div><p>Hypselostoma adela (F. G. Thompson &amp; Upatham, 1997) comb. nov.</p><p>Figs 111 A, 112, 113, 154</p><p>Gyliotrachela adela Thompson &amp; Upatham, 1997: 234–235, figs 44–47.</p><p>Gyliotrachela adela — Tongkerd et al. 2004: 143, fig. 2; Panha and Burch 2008: 65–66, fig. 57; Dumrongrojwattana and Wongkamhaeng 2024: 323, fig. 8.</p><p>Type material examined.</p><p>Thailand • holotype; 03 June 1987; F. G. Thompson leg.; UF 00113508; 4 paratypes; from the type locality; 03 June 1987; F. G. Thompson leg.; UF 00113509 .</p><p>Additional material examined.</p><p>Thailand-South • 16 shells; Suratthani Province, 28.4 km E Surat Thani, 19 km S <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.6&amp;materialsCitation.latitude=9.15" title="Search Plazi for locations around (long 99.6/lat 9.15)">Hwy.</a> 401, dry, scrubby evergreen forest, leaf litter, rocks, boulders; 9.15°N, 99.6°E; 40 m a. s. l.; 19 Apr. 1988; K. Auffenberg leg.; locality code KA-0675; UF 591322 • 15 shells; Suratthani Province, limestone outcrop along Hwy. 401, 2.7 km W junc <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.666664&amp;materialsCitation.latitude=9.166667" title="Search Plazi for locations around (long 99.666664/lat 9.166667)">Hwys.</a> 4142 &amp; 401; 9°10'N, 99°40'E; 90 m a. s. l.; 19 Apr. 1988; K. Auffenberg leg.; locality code KA-0673; UF 591323 • 2 shells; Suratthani Province, 15.3 km NE Ban Na San off road to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.433334&amp;materialsCitation.latitude=8.883333" title="Search Plazi for locations around (long 99.433334/lat 8.883333)">Tha Rom Yen National Park</a>; 8°53'N, 99°26'E; 40 m a. s. l.; 20 Apr. 1988; K. Auffenberg leg.; locality code KA-0678; UF 345389 • 5 shells; Suratthani Province, 13.8 km SE Don Sak, 5 km SW <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.71667&amp;materialsCitation.latitude=9.3" title="Search Plazi for locations around (long 99.71667/lat 9.3)">Hwy.</a> 4142; 9°18'N, 99°43'E; 50 m a. s. l.; 14 Apr. 1988; K. Auffenberg leg.; locality code KA-0670; UF 345284 • 1 shell; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.382&amp;materialsCitation.latitude=8.812" title="Search Plazi for locations around (long 99.382/lat 8.812)">Suratthani Province</a>, 1 km NE of Na San, 100 m limestone mountain; 8.812°N, 99.382°E; 20 June 1987; F. G. Thompson leg.; locality code FGT-4309; UF 529617 • 35 shells; Suratthani Province, Hwy. 401, 2.7 km W junction <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.666664&amp;materialsCitation.latitude=9.166667" title="Search Plazi for locations around (long 99.666664/lat 9.166667)">Hwy</a>. 4142 and 401, evergreen forest on rocky hillside, below cliff, base of cliff; 9°10'N, 99°40'E; 90 m a. s. l.; 19 Apr. 1988; K. Auffenberg leg.; locality code KA-0674; UF 345331 • 3 shells; Suratthani Province, 4.5 km S Don Sak, E side of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.7&amp;materialsCitation.latitude=9.3" title="Search Plazi for locations around (long 99.7/lat 9.3)">Hwy</a>. 4142, evergreen forest, limestone outcrop, base of cliff; 9°18'N, 99°42'E; 40 m a. s. l.; 18 Apr. 1988; K. Auffenberg leg.; locality code KA-0672; UF 345302 • 5 shells; Suratthani Province, 15.3 km NE Ban Na San off road to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.433334&amp;materialsCitation.latitude=8.883333" title="Search Plazi for locations around (long 99.433334/lat 8.883333)">Thi Rom Yen National Park</a>; 8°53'N, 99°26'E; 40 m a. s. l.; 20 Apr. 1988; K. Auffenberg leg.; locality code KA-0678; UF 591320 • 12 shells; Suratthani Province, 13.8 km SE Don Sak, 5 km SW <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.71667&amp;materialsCitation.latitude=9.3" title="Search Plazi for locations around (long 99.71667/lat 9.3)">Hwy.</a> 4142; 9°18'N, 99°43'E; 50 m a. s. l.; 17 Apr. 1988; K. Auffenberg leg.; locality code KA-0669; UF 345277 • 9 shells; Suratthani Province, 15.3 km NE Ban Na San off road to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.433334&amp;materialsCitation.latitude=8.883333" title="Search Plazi for locations around (long 99.433334/lat 8.883333)">Tha Rom Yen National Park</a>; 8°53'N, 99°26'E; 40 m a. s. l.; 20 Apr. 1988; K. Auffenberg leg.; locality code KA-0679; UF 345397 • 16 shells; Suratthani Province, 28.4 km E Surat Thani, 1.9 km S <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.6&amp;materialsCitation.latitude=9.15" title="Search Plazi for locations around (long 99.6/lat 9.15)">Hwy.</a> 401, dry, scrubby evergreen forest, leaf litter, rocks, boulders; 9.15°N, 99.6°E; 40 m a. s. l.; 19 Apr. 1988; K. Auffenberg leg.; locality code KA-0675; UF 436269 • 2 shells; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.37815&amp;materialsCitation.latitude=8.693167" title="Search Plazi for locations around (long 99.37815/lat 8.693167)">Suratthani Province</a>, limestone hill (with monastery) left side off rd. # 4009 (~ 11 km N of Waing Sa); 08°41.590'N, 099°22.689'E; ex. coll. Hemmen, Wiesbaden; coll. PGB • 8 shells; Nakhon Si Thammarat Province, 15.6 km SE Don Sak, 1 km SW <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.76667&amp;materialsCitation.latitude=9.283334" title="Search Plazi for locations around (long 99.76667/lat 9.283334)">Hwy.</a> 4142; 9°17'N, 99°46'E; 50 m a. s. l.; 15 Apr. 1988; K. Auffenberg leg.; locality code KA-0667; UF 345253 • 16 shells; Nakhon Sri Thammarat Province, 15.6 km SE Don Sak, 1 km SW <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.76667&amp;materialsCitation.latitude=9.283334" title="Search Plazi for locations around (long 99.76667/lat 9.283334)">Hwy.</a> 4142; 9°17'N, 99°46'E; 50 m a. s. l.; 17 Apr. 1988; K. Auffenberg leg.; locality code KA-0668; UF 345258 • 2 shells; Nakhon Sri Thammarat Province, limestone hill right off road # 4142 ca km 38.2 (road Khanom-Don Sak); Mar. 2000; ex. coll. Hemmen, Wiesbaden; coll. PGB • 4 shells; Krabi Province, Phi Phi islands, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.765785&amp;materialsCitation.latitude=7.7343836" title="Search Plazi for locations around (long 98.765785/lat 7.7343836)">Phi Phi Don island</a>, climbing rock above the western end of the beach in <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.765785&amp;materialsCitation.latitude=7.7343836" title="Search Plazi for locations around (long 98.765785/lat 7.7343836)">Ton Sai Bay</a>, at the base of limestone rocks in the forest; 7°44.063'N, 98°45.947'E; 50 m a. s. l.; Mar. 2010; A. Reischütz leg; coll. REI • 1 shell; Phattalung Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.08884&amp;materialsCitation.latitude=7.6251" title="Search Plazi for locations around (long 100.08884/lat 7.6251)">Khao Ok Thalu</a>, rock wall; 07°37.506'N, 100°05.330'E; 19 Feb. 2015; A. Hunyadi leg.; coll. HA • 1 shell; Phatthalung Province, Khao Pu-Khao Ya Nat. Parl, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.84703&amp;materialsCitation.latitude=7.6789665" title="Search Plazi for locations around (long 99.84703/lat 7.6789665)">Si Banphot district</a>, ca 6 km NNW of Taphaen, 7°40.738'N, 99°50.822'E; 24 Feb. 2023; A. Hunyadi, K. Okubo &amp; J. U. Otani leg.; coll. HA • 1 shell; Songkhla Province, 31.3 km NW <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.26667&amp;materialsCitation.latitude=7.1666665" title="Search Plazi for locations around (long 100.26667/lat 7.1666665)">Hat Yai</a>, 1.2 km W of Hwy. 43; 7°10'N, 100°16'E; 80 m a. s. l.; 08 Apr. 1988; K. Auffenberg leg.; locality code KA-0636 B; UF 344945. Malaysia • 3 shells; Pahang, Bandar Pusat Jengka, west 15 km, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.433&amp;materialsCitation.latitude=3.6977" title="Search Plazi for locations around (long 102.433/lat 3.6977)">Hutan Lipur Gunung Senyum</a>; 03°41.862'N, 102°25.980'E; 85 m a. s. l.; 23 Jan. 2013; A. Hunyadi leg.; coll. HA .</p><p>Type locality.</p><p>“ Thailand, Surat Thani Province, box canyon in limestone range 6 km south of Na San; 100 m altitude (08°39.6'N, 99°23.9'E) ”.</p><p>Differential diagnosis.</p><p>This species is not similar to any other congener and is unique due to the combination of the following characters: small size, yellowish shell, spirally striated teleoconch with descending detached last whorl and aperture equipped with several equally developed barriers (without smaller ones). See under H. cucumense .</p><p>Distribution.</p><p>This species in known from several localities in Thailand and one in Peninsular Malaysia. Our record from Malaysia is the first record of this species in the country.</p><p>Remarks.</p><p>Apertural barriers were constant in number in all the specimens examined. We have examined the material from northern Thailand hosted in UF with the following data: 1 shell; Mae Hong Song Province, Tham Lod Cave, 1.5 km SE Ban Tham Lod Village, 6 km N Sappong on Road 1095; 19°34'N, 98°9'E; 720 m a. s. l.; 19 Mar. 1988; K. Auffenberg leg.; locality code KA-0576; UF 591321; 7 shells; Mae Hong Song Province, Tham Lod Cave, 1.5 km SE Ban Tham Lod Village, 6 km N Sappong on Road 1095; 19°34'N, 98°9'E; 720 m a. s. l.; 19 Mar. 1988; K. Auffenberg leg.; locality code KA-0674; UF 345523. Since these localities are quite far from the actual distribution area of this species, we believe that these samples were mislabelled and that this species actually does not occur in Mae Hong Son Province.</p></div>	https://treatment.plazi.org/id/3057154D05485D0A908072205C62B448	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
A7F1DE4ED3615EDBA8AA9BE749373388.text	A7F1DE4ED3615EDBA8AA9BE749373388.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma aenigma Gojsina, Grego & Pall-Gergely 2025	<div><p>Hypselostoma aenigma Gojšina, Grego &amp; Páll-Gergely sp. nov.</p><p>Figs 111 B, 114, 115, 158</p><p>Type material.</p><p>Holotype. Myanmar • 1 shell (SH: 2.1 mm; SW: 1.7 mm); Kayin State, Kamarmaung, Phon Tho Village, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.72474&amp;materialsCitation.latitude=17.322624" title="Search Plazi for locations around (long 97.72474/lat 17.322624)">Pan Do Mi Mountain</a> spring cave at W foot of the mountain; 17°19.3575'N, 97°43.4842'E 19. May 2019; J. Grego leg; CUMZ 14452.</p><p>Type locality.</p><p>Myanmar, Kayin State, Kamarmaung, Phon Tho Village, Pan Do Mi Mountain spring cave at W foot of the mountain; 17°19.3575'N, 97°43.4842'E.</p><p>Diagnosis.</p><p>Hypselostoma species with conical-ovoid shell, spirally striated whorls, and the last whorl with a keel below the centre of the periphery. Apertural barriers three but the parietal lamella is the only tooth-like barrier. Palatal plica and columellar lamella are present as swellings on the peristome.</p><p>Description.</p><p>Shell conical-ovoid, pale yellowish, consisting of nearly 4.5 whorls separated by a deep suture. All whorls, except the last one, regularly rounded and convex. Last whorl with a blunt keel below the centre of the periphery and weakly concave to almost flat whorl outline both above and below the keel. Protoconch of same colour as the rest of the shell, finely pitted showing spiralling pattern which continues as more distinct spiral striae terminally. It consists of ~ 1.75 whorls. Teleoconch coarsely sculptured with irregularly spaced radial growth lines but also with dense spiral striae. Spacing between the two spiral striae equals the width of two or three striae. Last whorl adnate to the penultimate and very slightly ascending (~ 7 ° compared to the shell axis), making the apertural profile weakly opisthocline to the shell axis. Peristome white, thick, and expanded (especially on the parietal side) but not reflected. Apertural barriers largely reduced and of very unusual appearance for any hypselostomatid. Parietal lamella is the only „ tooth-like “ barrier in the aperture but it is very low and short. Palatal plica and columellar lamella not present in their typical forms but rather as very weak and indistinct swellings on the peristome, visible only in the frontal view. These swellings are probably homologous to the two mentioned barriers. There are no other barriers in the aperture. Surface of all apertural barriers is very finely granulated (almost smooth). Sinulus wide due to the weak parietal lamella and palatal plica. Umbilicus extremely narrow, dot like and slightly elongated. Umbilical groove absent.</p><p>Differential diagnosis.</p><p>This species is not similar to any other congener or species within other genera due to the unique appearance of apertural barriers.</p><p>Measurements</p><p>(in mm, n = 1): SH = 2.1; SW = 1.7; AH = 0.97; AW = 0.93.</p><p>Etymology.</p><p>The specific epithet aenigma comes from a Latin word for a riddle (which refers to the unique appearance of the shell and unclear relationship with other congeners) and is to be used as a noun in apposition.</p><p>Distribution.</p><p>This species is known only from its type locality.</p></div>	https://treatment.plazi.org/id/A7F1DE4ED3615EDBA8AA9BE749373388	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
9BB9CA10F75D525994361730BFCC2306.text	9BB9CA10F75D525994361730BFCC2306.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma annamiticum Mollendorff 1900	<div><p>Hypselostoma annamiticum Möllendorff, 1900</p><p>Figs 111 C, 116, 154</p><p>Hypselostoma annamiticum Möllendorff, 1900: 133.</p><p>Hypselostoma annamiticum — Möllendorff 1901 a: 50; Fischer and Dautzenberg 1904: 408; Pilsbry 1917: 180–181; Zilch 1984: 161; van Benthem Jutting 1950: 41.</p><p>Hypselostoma annamiticum altius Pilsbry, 1917: 181, pl. 31, figs 11–13. syn. nov.</p><p>Hypselostoma annamiticum altius — van Benthem Jutting 1950: 41; Schileyko 2011: 3.</p><p>Hypselostoma annamiticum annamiticum — Schileyko 2011: 3.</p><p>Type material examined.</p><p>Vietnam • holotype of H. annamiticum annamiticum; “ Phuc-Son, Touranne, Annam ”; ex. coll. Möllendorff; SMF 4871 • 11 paratypes of H. annamiticum annamiticum; same data as for the holotype; ex. coll. Möllendorff; SMF 4872 • 1 paralectotype of H. annamiticum altius; from the type locality; ANSP 45167 .</p><p>Type localities.</p><p>“ Phuc-son ”, Vietnam ( H. annamiticum annamiticum), “ Annam (Fruhstorfer) ”, Vietnam ( H. annamiticum altius).</p><p>Differential diagnosis.</p><p>This species is superficially similar and geographically approximate to A. turritus sp. nov., but the suture of this species is shallower, the shell is more slender, the umbilicus is wider and the apertural dentition is also different (no concrescent barriers on the parietal side in the new species vs angulo-parietal lamella in H. annamiticum).</p><p>Distribution.</p><p>This species is known only from Annam, Vietnam.</p><p>Remarks.</p><p>Holotype of H. annamiticum annamiticum has a vestigial lamella between the columellar lamella and angulo-parietal lamella. Some paratypes do not have this lamella or it is further less developed, sometimes also located much deeper inside the aperture. In some specimens, upper palatal plica has an additional palatal tubercle which is sometimes connected with the former. Hypselostoma annamiticum altius is synonymised with the nominotypical subspecies since we could observe no morphological differences. The fact that it is higher than the nominotypical subspecies (Pilsbry 1917) does not seem to justify its distinctness.</p></div>	https://treatment.plazi.org/id/9BB9CA10F75D525994361730BFCC2306	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
F07A17348C1257F0B396DF7E9AAE416F.text	F07A17348C1257F0B396DF7E9AAE416F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma aquila Gojsina, Hunyadi & Pall-Gergely 2025	<div><p>Hypselostoma aquila Gojšina, Hunyadi &amp; Páll-Gergely sp. nov.</p><p>Figs 111 D, 117, 118, 177</p><p>Type material.</p><p>Holotype. Cambodia • 1 shell (SH: 1.98 mm; SW 1: 2.39 mm); Steung Treng Province, 55.7 km northwest + 2 km north from Stung Treng Mekong Bridge, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.60342&amp;materialsCitation.latitude=13.797017" title="Search Plazi for locations around (long 105.60342/lat 13.797017)">Chap Phleung Mt.</a> (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.60342&amp;materialsCitation.latitude=13.797017" title="Search Plazi for locations around (long 105.60342/lat 13.797017)">Neak Khiev Mt.</a>); 13°47.821'N, 105°36.205'E; 135 m a. s. l.; 26 Oct. 2023, A. Hunyadi &amp; J. U. Otani leg.; CUMZ 14461 . Paratypes. Cambodia • 107 shells; same data as for holotype; coll. HA • 1 shell; same data as for holotype; coll. VG • 229 shells; Steung Treng Province, Stung Treng Mekong Bridge, 36.3 km NWN + 5 km N, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.74796&amp;materialsCitation.latitude=13.7762165" title="Search Plazi for locations around (long 105.74796/lat 13.7762165)">Phnom Chhnok</a>; 13°46.573'N, 105°44.878'E; 120 m a. s. l.; 25 Oct. 2023; A. Hunyadi &amp; J. U. Otani leg.; coll. HA .</p><p>Additional material examined.</p><p>Cambodia • 4 shells (juveniles / damaged, not paratypes); Steung Treng Province, Stung Treng Mekong Bridge, 36.3 km NWN + 5 km N, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.74796&amp;materialsCitation.latitude=13.7762165" title="Search Plazi for locations around (long 105.74796/lat 13.7762165)">Phnom Chhnok</a>; 13°46.573'N, 105°44.878'E; 120 m a. s. l.; 25 Oct. 2023; A. Hunyadi &amp; J. U. Otani leg.; coll. HA .</p><p>Type locality.</p><p>Cambodia, Steung Treng Province, 55.7 km northwest + 2 km north from Stung Treng Mekong Bridge, Chap Phleung Mt. (Neak Khiev Mt.); 13°47.821'N, 105°36.205'E; 135 m a. s. l.</p><p>Diagnosis.</p><p>A Hypselostoma species with a slender, concave-conical shell, bluntly and strongly keeled last whorl (and a deep groove above it), raised spiral striation, hooked apertural barriers (upper palatal, and lower palatal) and a wide umbilicus which is initially narrow.</p><p>Description.</p><p>Shell concave-conical, light-brown, consisting of 4.5–5 whorls separated by a deep suture. Protoconch nearly white, consisting of ~ 1.25 coarsely spirally striated whorls. There are ~ 12 weak spiral striae on the protoconch. Protoconch and initial teleoconch whorls rounded, penultimate weakly convex. Last whorl with strong but blunt keel positioned at the centre of the periphery. Above the keel there is a deep groove on the last 0.75–1 whorl. All whorls densely sculptured with strong, thick spiral striae crossed by less dense radial growth lines and occasionally by a few thin, whitish radial streaks. Spacing between the spiral striae irregular and ranges from the width of two to the width of three spiral striae. Last whorl moderately detached from the penultimate and weakly ascending (≤ 10 ° compared to the shell axis). Peristome expanded, not reflected, its surface finely pitted. Aperture equipped with four strong, main barriers (angulo-parietal, upper palatal, lower palatal and columellar) and several smaller ones. Angulo-parietal lamella strong and leaned towards the palatal side, almost reaching the expanding peristome. It is consisting of main, parietal part, and a smaller, pointed angular part which are separated by a weak sinuation. Upper and lower palatal plicae roughly the same size, both divided and the inner parts are hooked (with the tip of hooks pointing outside), slender and high, while the outer parts are low and slender. A weak swelling is also observed at the palatal side (in front of the upper palatal plicae) which is probably homologous to the palatal tubercle in the majority of Bensonella species. Columellar lamella almost horizontal, approximately as high as the palatal plicae. There are four smaller barriers, two interpalatal plicae, one basal plica and one infraparietal lamella. Surface of all apertural barriers is smooth or with very weak granules. Sinulus rounded, distinctly separated from the rest of the aperture. Umbilicus initially narrow and then abruptly widening, measuring between 1 / 3 and 1 / 4 of the shell width. There is a periumbilical keel situated on the last ~ 0.75 whorl near the centre of the last whorl, but rather slightly towards the umbilicus. Along this keel there are shallow grooves on both sides.</p><p>Differential diagnosis.</p><p>See under H. benetuitum, H. sorormajor sp. nov., and H. sororminor sp. nov.</p><p>Measurements</p><p>(in mm, n = 5). SH = 1.91–2.04; SW 1 = 2.16–2.39; SW 2 = 1.35–1.42; AH = 0.80–0.86; AW = 0.69–0.82.</p><p>Etymology.</p><p>This species is named for the apertural barriers resembling claws of an eagle (aquila = eagle in Latin). To be used as a noun in apposition.</p><p>Distribution.</p><p>This species is known from two localities in Steung Treng province.</p><p>Remarks.</p><p>The largest, and still unexplored limestone hill in Steung Treng Province (13°47.941'N, 105°43.7671'E) was inaccessible during the collecting efforts in 2023 due to the works of a cement factory. Due to the proximity of this hill to the type locality of H. aquila sp. nov., it is possible that this species can be found here as well and be threatened by quarrying.</p></div>	https://treatment.plazi.org/id/F07A17348C1257F0B396DF7E9AAE416F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
4CB784AA75FB58F1ADD3E08328FCC5AE.text	4CB784AA75FB58F1ADD3E08328FCC5AE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma aunglini (Tongkerd & Panha 2024) Gojšina & Hunyadi & Sutcharit & Tongkerd & Auffenberg & Grego & Vermeulen & Reischütz & Páll-Gergely 2025	<div><p>Hypselostoma aunglini (Tongkerd &amp; Panha, 2024) comb. nov.</p><p>Figs 186 A, 187, 223</p><p>Gyliotrachela aunglini Tongkerd &amp; Panha in Tongkerd et al. 2024: 184–187, figs 12, 13 P.</p><p>Type material examined.</p><p>Myanmar • holotype; collector unknown; CUMZ 14383 .</p><p>Type locality.</p><p>“ Kaw Gon Cave, Hpa-An, Kayin State, Myanmar (…; 16°49'22.2"N, 97°35'08.9"E) ”.</p><p>Differential diagnosis.</p><p>The most similar congener from Myanmar is H. bensonianum which is however spirally striated (granulated in H. aunglini) and has its last whorl keeled (shouldered in H. aunglini).</p><p>Distribution.</p><p>This species is known only from the type locality.</p></div>	https://treatment.plazi.org/id/4CB784AA75FB58F1ADD3E08328FCC5AE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
C8B98D2512815848A1F273EC40401E94.text	C8B98D2512815848A1F273EC40401E94.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma australe subsp. australe Odhner 1917	<div><p>Hypselostoma australe australe Odhner, 1917</p><p>Figs 111 E, 119</p><p>Hypselostoma australis Odhner, 1917: 98, pl. 3, figs 107–109.</p><p>Gyliotrachela australis — Iredale 1937: 305; van Benthem Jutting 1950: 44; Pokryszko 1996: 1117, figs 31, 33, 34.</p><p>Gyliotrachela ningbingia Solem, 1981: 91, figs 3, 4, 10, 14–16, 18, 19.</p><p>Gyliotrachela catherina Solem, 1981: 91–92, figs 5, 6, 11, 17; Pokryszko 1996: 1117.</p><p>Gyliotrachela ninbingia [sic] — Pokryszko 1996: 1117.</p><p>Type material examined.</p><p>Australia • syntype of H. australis australis; SMNH 1667 .</p><p>Type locality.</p><p>“ caves at Chillagoe ” (North Queensland, Australia).</p><p>Differential diagnosis.</p><p>This species is similar in shape to several Southeast Asian representatives, but it can be separated clearly by its different shell surface sculpture which consists of equally strong, rib-like radial growth lines and spiral striae. See also under H. australe napierana .</p><p>Distribution.</p><p>This species in known from several localities in N Australia as well as under several synonyms (Iredale 1937; Solem 1981).</p><p>Remarks.</p><p>Pokryszko (1996) synonymised G. catherina and G. ningbingia with H. australe, mentioning that the former two show only biometrical differences from H. australe which does not justify their distinctness.</p></div>	https://treatment.plazi.org/id/C8B98D2512815848A1F273EC40401E94	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
85DACDD1EDF45505908AF4380E1483E5.text	85DACDD1EDF45505908AF4380E1483E5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma australe subsp. napieranum (Solem 1981)	<div><p>Hypselostoma australe napieranum (Solem, 1981) comb. nov.</p><p>Gyliotrachela napierana Solem, 1981: 90–91, figs 1, 2, 9, 13.</p><p>Gyliotrachela australis napierana — Pokryszko 1996: 1117–1118, fig. 32.</p><p>Material examined.</p><p>None.</p><p>Type locality.</p><p>“ Station WA-325, 0.6 km south-west of road along north side of Napier Range, 5.9 km north-west of Yammera Gap, Western Australia ”.</p><p>Differential diagnosis.</p><p>This subspecies is different from the nominotypical by the more numerous and stronger apertural barriers.</p><p>Distribution.</p><p>This subspecies is known from several localities in Napier Range, western Australia.</p></div>	https://treatment.plazi.org/id/85DACDD1EDF45505908AF4380E1483E5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
3A1138DB104A5264AEB0BC20F3A8A63D.text	3A1138DB104A5264AEB0BC20F3A8A63D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma benetuitum Vermeulen, Luu, Theary & Anker 2019	<div><p>Hypselostoma benetuitum Vermeulen, Luu, Theary &amp; Anker, 2019</p><p>Figs 111 F, 120, 182</p><p>Hypselostoma benetuitum Vermeulen et al., 2019: 38–39, figs 64–68.</p><p>Hypselostoma benetuitum — Sutcharit et al. 2020: 18, fig. 7 E, F; Sutcharit et al. 2023: 1290.</p><p>Type material examined.</p><p>Cambodia • holotype; J. Vermeulen leg.; NHMUK 20180575 .</p><p>Type locality.</p><p>“ Phnom Kampong Trach ”, Cambodia.</p><p>Differential diagnosis.</p><p>This species is similar in shell shape to other Cambodian representatives ( H. discobasis, H. sorormajor sp. nov., H. sororminor sp. nov., H. aquila sp. nov.) but differs from all of them by the presence of a keel with a concave surface below it but flat above it (vice versa in other species). Furthermore, this species has hooked apertural barriers in contrast to H. discobasis and H. sororminor sp. nov. See also under H. cambodjense .</p><p>Distribution.</p><p>This species is known from Kampot province, Cambodia.</p></div>	https://treatment.plazi.org/id/3A1138DB104A5264AEB0BC20F3A8A63D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
3B18896022AF5A64BE96BC58CF98526A.text	3B18896022AF5A64BE96BC58CF98526A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma bensonianum W. T. Blanford 1863	<div><p>Hypselostoma bensonianum W. T. Blanford, 1863</p><p>Figs 111 G, 121, 158</p><p>Hypselostoma bensonianum Blanford, 1863: 326–327.</p><p>Hypselostoma bensonianum — Hanley and Theobald 1870: 4, pl. 8 fig. 2; Pfeiffer 1875: 488; Möllendorff 1891: 338; Gude 1914: 299.</p><p>Pupa (Hypselostoma) bensoni — Nevill 1878: 193.</p><p>Gyliauchen bensonianus — Pilsbry 1917: 211; pl. 37 figs 11, 12.</p><p>Gyliotrachela bensoniana — van Benthem Jutting 1950: 44.</p><p>Gyliotrachela bensonianum — Gojšina et al. 2022: 136–137, figs 4–5; Tongkerd et al. 2024: 179, fig. 13 L.</p><p>Type material examined.</p><p>Myanmar • 5 syntypes; NHMUK 20191141 .</p><p>Type locality.</p><p>“ Mya Leit Doung, Ava ” (nowadays Melai Taung 21.8°N, 96.2°E).</p><p>Differential diagnosis.</p><p>This species differs from all other congeners by its keeled last whorl and spirally striated teleoconch in combination with a small number of relatively strong apertural barriers (usually up to seven). See also under H. aunglini .</p><p>Distribution.</p><p>This species is known only from the type locality.</p></div>	https://treatment.plazi.org/id/3B18896022AF5A64BE96BC58CF98526A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
22793DE134D1595F82D6D1F19C6CE6D9.text	22793DE134D1595F82D6D1F19C6CE6D9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma bubalus Gojsina, Hunyadi & Pall-Gergely 2025	<div><p>Hypselostoma bubalus Gojšina, Hunyadi &amp; Páll-Gergely sp. nov.</p><p>Figs 186 B, 188, 189, 223</p><p>Type material.</p><p>Holotype. Malaysia • 1 shell (SH: 2 mm; SW: 3.7 mm); Kedah, 1.5 km east from Kodiang, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.31506&amp;materialsCitation.latitude=6.3905168" title="Search Plazi for locations around (long 100.31506/lat 6.3905168)">Gua Kerbau</a>; 06°23.431'N, 100°18.904'E, 14 Jan. 2013; A. Hunyadi leg.; CUMZ 14466.</p><p>Paratypes. Malaysia • 216 shells; same data as for holotype; coll. HA • 1 shell; same data as for holotype; coll. VG .</p><p>Additional material examined.</p><p>Malaysia • 25 shells (damaged, not paratypes); same data as for holotype; coll. HA .</p><p>Type locality.</p><p>Malaysia, Kedah, 1.5 km east from Kodiang, Gua Kerbau; 06°23.431'N, 100°18.904'E.</p><p>Diagnosis.</p><p>Shell depressed, concave-conical. Teleoconch finely radially and spirally striated (localised, just below and above the keel on the last whorl). Last whorl detached from the penultimate and strongly ascending. Aperture wide, equipped with five main barriers (angular, parietal, upper palatal, lower palatal and columellar) and two smaller barriers (basal and infraparietal). Umbilicus wide, measuring 50 % of the shell width.</p><p>Description.</p><p>Shell depressed, concave-conical, chestnut brown, very weakly glossy and not translucent. Whorls 3.5–4, initially weakly convex, separated by a deep suture, regularly increasing until the last whorl which is strongly enlarged and flat. Protoconch lighter than the rest of the shell, finely pitted without spiralling pattern, consisting of ~ 1.25 whorls. Teleoconch very finely granulated (sandpaper-like surface) and radially striated with additional whitish radial streaks which are weak, widely, and irregularly spaced. Spiral striae present but localised, clearly visible only on the last whorl above and below the keel, elsewhere absent. Sometimes, spiral striae are very difficult to observe on the whole shell. Last whorl strongly enlarged in comparison to previous ones, bluntly keeled very slightly above the centre of the periphery (or sometimes at the centre of the periphery) and almost flat above and weakly convex below the keel. Keel is strongest near the aperture. Last whorl moderately to strongly ascending near the aperture (~ 50–65 ° compared to the shell axis) and moderately detached from the penultimate whorl. Peristome very pale brownish, strongly expanded but not reflected. Aperture equipped with five main barriers (angular, parietal, upper palatal, lower palatal, and columellar). Angular lamella short and small, directed towards the upper palatal plica. Parietal lamella strong and high, not much longer than the angular and only reaching its profile. Upper palatal plica slightly weaker than the parietal lamella and bent towards the lower palatal plica. Lower palatal plica straight, equally strong as the upper palatal but not bent. Columellar lamella is the broadest in the aperture but relatively short. Between these main barriers, additional small basal and infraparietal plicae visible. Surface of all apertural barriers is very finely spiniferous medially and very finely granulated laterally. Sinulus small and distinctly separated from the rest of the aperture. Umbilicus very wide, measuring half of the shell width and showing all previous whorls. A deep and wide groove is running from the columellar side of the peristome alongside the umbilicus.</p><p>Differential diagnosis.</p><p>See under H. luctans .</p><p>Measurements</p><p>(in mm, n = 5). SH = 2–2.23; SW 1 = 3.40–4.13; SW 2 = 1.95–2.41; AH = 1.44–1.78; AW = 1.33–1.61.</p><p>Etymology.</p><p>The type locality of this species, Gua Kerbau, means “ Buffalo cave ” in Malay. The specific epithet bubalus is also a scientific name for a buffalo, which is to be used as a noun in apposition.</p><p>Distribution.</p><p>This species is known only from the type locality.</p><p>Remarks.</p><p>Although this species shows some spiral striation, it is placed into H. hungerfordianum group because of the predominant sandpaper-like surface on the teleoconch.</p></div>	https://treatment.plazi.org/id/22793DE134D1595F82D6D1F19C6CE6D9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
30801580F371587FB602B63E31FA6065.text	30801580F371587FB602B63E31FA6065.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma burchi (Panha 1998) Gojšina & Hunyadi & Sutcharit & Tongkerd & Auffenberg & Grego & Vermeulen & Reischütz & Páll-Gergely 2025	<div><p>Hypselostoma burchi (Panha, 1998) comb. nov.</p><p>Figs 242 A, 243, 259</p><p>Gyliotrachela burchi Panha, 1998 c: 123–124, fig. 2.</p><p>Gyliotrachela burchi — Hemmen and Hemmen 2001: 41, fig. 9; Panha and Burch 2008: 66–67, fig. 58; Dumrongrojwattana and Wongkamhaeng 2024: 323, fig. 8.</p><p>Type material examined.</p><p>Thailand • holotype; 1996; S. Panha leg.; CUMZ ver. 015 .</p><p>Type locality.</p><p>“ Pattalung province, 7°40′17″N, 100°1′11″E, 70 meters elevation ”. (Thailand)</p><p>Differential diagnosis.</p><p>This species is similar in shell shape to H. taehwani but the latter is spirally striated and has concrescent angular and parietal lamellae. Also, H. taehwani has a stronger keel than H. burchi and deeper grooves both above and below the keel. It also resembles H. modestum but the last whorl in H. burchi is keeled and more strongly detached and descending. The umbilicus is also wider in H. burchi .</p><p>Distribution.</p><p>This species is known only from the type locality.</p></div>	https://treatment.plazi.org/id/30801580F371587FB602B63E31FA6065	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
48CA0F81C7CF5488867777A1045A9067.text	48CA0F81C7CF5488867777A1045A9067.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma cambodjense van Benthem Jutting 1962	<div><p>Hypselostoma cambodjense van Benthem Jutting, 1962</p><p>Figs 111 H, 122, 182</p><p>Hypselostoma cambodjense van Benthem Jutting, 1962: 3–5, fig. 1.</p><p>Hypselostoma cambodjense — Vermeulen et al. 2019: 39; Sutcharit et al. 2020: 18, figs 3 H, 8 A – C; Sutcharit et al. 2023: 1290.</p><p>Type material examined.</p><p>Cambodia • 4 paratypes; from the type locality; 1960; E. Saurin leg.; RMNH.Moll.137169 .</p><p>Type locality.</p><p>“ Phnom Can Long, à 6 km au Sud de Tuk Méas, Cambodge ”.</p><p>Differential diagnosis.</p><p>This species differs from H. benetuitum by the absence of hooked apertural barriers. Also, the keel is slightly sharper in H. benetuitum with a more concave surface below it.</p><p>Distribution.</p><p>This species in known from Kampot province (Cambodia) and Kien Giang province (Vietnam) (Vermeulen et al. 2019).</p><p>Remarks.</p><p>Of four paratypes examined, one of them had the last whorl fully adnate to the penultimate.</p></div>	https://treatment.plazi.org/id/48CA0F81C7CF5488867777A1045A9067	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
D8BFAE571B8E536B961463946671F915.text	D8BFAE571B8E536B961463946671F915.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma chatnareeae (Panha & J. B. Burch 2003) Gojšina & Hunyadi & Sutcharit & Tongkerd & Auffenberg & Grego & Vermeulen & Reischütz & Páll-Gergely 2025	<div><p>Hypselostoma chatnareeae (Panha &amp; J. B. Burch, 2003) comb. nov.</p><p>Figs 186 C, 190, 223</p><p>Anauchen chatnaeerae Panha &amp; Burch in Burch et al. 2003: 132–138, figs 2, 3.</p><p>Anauchen chatnaeerae — Tongkerd et al. 2004: 143, fig. 2; Panha and Burch 2008: 49–50, fig. 45; Dumrongrojwattana and Wongkamhaeng 2024: 323, fig. 7.</p><p>Type material examined.</p><p>Thailand • 3 paratypes; from the type locality; SMF 331447 .</p><p>Additional material examined.</p><p>Thailand • 1 shell; Tam Sua Hill, Utong, Suphanburi; 21 Dec. 2002; K. Okubo leg.; coll. PGB .</p><p>Type locality.</p><p>“ Tam Sua, on the limestone hill on the west side of Tam Sua Temple, Utong District, Suphanburi Province, 14°17'31"N, 100°7'31"E, 20 meters elevation, Thailand ”.</p><p>Differential diagnosis.</p><p>This species is not similar to any of its congeners. The shell shape (conical) and apertural dentition (one barrier on the parietal side) strongly resembles some Anauchen species (e. g., A. angthongensis) but the surface is granulated as in H. hungerfordianum and all whorls are keeled. See also under H. khmerianum .</p><p>Distribution.</p><p>Apart from the type locality, this species is also known from one more nearby locality.</p><p>Remarks.</p><p>Even though this species has high morphological similarities to some Anauchen species (e. g., A. angthongensis), the shell surface sculpture is characteristically granulated (as in H. hungerfordianum) which is the reason it is transferred to this genus.</p></div>	https://treatment.plazi.org/id/D8BFAE571B8E536B961463946671F915	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
481B339CBD1D5132BEE354F346A4FDCB.text	481B339CBD1D5132BEE354F346A4FDCB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma chaunosalpinx (Vermeulen, Luu, Theary & Anker 2019) Gojšina & Hunyadi & Sutcharit & Tongkerd & Auffenberg & Grego & Vermeulen & Reischütz & Páll-Gergely 2025	<div><p>Hypselostoma chaunosalpinx (Vermeulen, Luu, Theary &amp; Anker, 2019) comb. nov.</p><p>Figs 111 I, 123, 182</p><p>Anauchen chaunosalpinx Vermeulen et al., 2019: 35, figs 57–59.</p><p>Anauchen chaunosalpinx — Sutcharit et al. 2023: 1290.</p><p>Type material examined.</p><p>Cambodia • holotype; NHMUK 20180573 .</p><p>Type locality.</p><p>“ Cambodia: Kampot Province, Kampong Trach area: Phnom Kampong Trach ”.</p><p>Differential diagnosis.</p><p>This species is not similar to any other congener due to the combination of conical shell with wide last whorl which is strongly descending and has several hooked apertural barriers. See under H. depressum .</p><p>Distribution.</p><p>This species is known only from the type locality.</p></div>	https://treatment.plazi.org/id/481B339CBD1D5132BEE354F346A4FDCB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
F0E65A96447B5B67A47F6595A1846C14.text	F0E65A96447B5B67A47F6595A1846C14.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma chedi Panha 1998	<div><p>Hypselostoma chedi Panha, 1998</p><p>Figs 111 J, 124, 154</p><p>Hypselostoma chedi Panha, 1998 b: 61–63, fig. 2.</p><p>Anauchen chedi — Burch and Panha 2002: 246, fig. 4; Tongkerd et al. 2004: 143, fig. 2; Panha and Burch 2008: 50–51, fig. 46; Dumrongrojwattana and Wongkamhaeng 2024: 323, fig. 7.</p><p>Hypselostoma chedi — Hemmen and Hemmen 2001: 41.</p><p>Type material examined.</p><p>Thailand • paratype; S. Panha leg.; CUMZ ver. 004; 2 paratypes; from the type locality; SMF 331470 .</p><p>Additional material examined.</p><p>Thailand • 36 shells; Nakhon Sawan Province, 19.5 km NE intersection <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.26667&amp;materialsCitation.latitude=15.233334" title="Search Plazi for locations around (long 100.26667/lat 15.233334)">Hwys</a> 11 and 32; 15°14'N, 100°16'E; 50 m a. s. l.; 02. May 1988; K. Auffenberg leg.; locality code KA-0696; UF 346127 / 2, UF 346138 / 34 .</p><p>Type locality.</p><p>“ Thailand, Tebpratan Nature Reserve area, Kampangpet Province, 15°54′52″N, 99 ° 52 ′ 63 ″ E, 140 meters elevation ”.</p><p>Differential diagnosis.</p><p>see under H. smokon and H. khaochakan .</p><p>Distribution.</p><p>This species is known from Nakhon Sawan and Kampangpet provinces.</p><p>Remarks.</p><p>Coordinates provided in the original description are not correct.</p></div>	https://treatment.plazi.org/id/F0E65A96447B5B67A47F6595A1846C14	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
44A493DFBA625C4FB85B476735F7175E.text	44A493DFBA625C4FB85B476735F7175E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma circumcarinatum Gojsina, Auffenberg & Pall-Gergely 2025	<div><p>Hypselostoma circumcarinatum Gojšina, Auffenberg &amp; Páll-Gergely sp. nov.</p><p>Figs 111 K, 125, 126, 154</p><p>Type material.</p><p>Holotype. Thailand • 1 shell (SH: 2.3 mm; SW: 2.2 mm); Kanchanaburi Province, 4.3 km SW Kanchanaburi, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.51667&amp;materialsCitation.latitude=13.983334" title="Search Plazi for locations around (long 99.51667/lat 13.983334)">Taoist Buddhist temple</a>; 13°59'N, 99°31'E; 40 m a. s. l.; 13. May 1988, K. Auffenberg leg.; UF 345933 . Paratypes. Thailand • 1 shell; same data as for holotype; CUMZ 14456 • 3 shells (2 whole and 1 with complete aperture); same data as for holotype; UF 591336 .</p><p>Additional material examined.</p><p>Thailand • 4 shells (damaged, without last whorl, not paratypes); same data as for holotype; UF 583722 .</p><p>Type locality.</p><p>Kanchanaburi Province, 4.3 km SW Kanchanaburi, Taoist Buddhist temple; 13°59'N, 99°31'E; 40 m a. s. l.</p><p>Diagnosis.</p><p>Hypselostoma species with very deep sutures and spirally striated whorls. Penultimate and the last whorl are keeled. Last whorl not detached from the penultimate. Aperture equipped with six barriers (angulo-parietal, upper palatal, lower palatal, basal, columellar and a small infraparietal). Umbilicus initially narrow, abruptly widening at the last whorl.</p><p>Description.</p><p>Shell conical-ovoid, consisting of 4–4.5 whorls separated by a deep suture. Protoconch pitted, showing some spiralling pattern, the same colour as the rest of the shell, consisting of ~ 1.5 whorls. Teleoconch surface sculptured with coarsely spaced radial lines crossed by much stronger and dense spiral striae. Spacing between two spiral striae approximates the width of one or two striae. First teleoconch whorl rounded. All other teleoconch whorls strongly keeled, last one the most prominent. Surface above the keel on the penultimate whorl is almost flat, (like a platform) but is much more steep (sloping) on the last whorl. Last whorl adnate to the penultimate and slightly ascending near the aperture (~ 5–10 ° compared to the shell axis). Peristome not very strong but expanded, not reflected. It is forming a weak callus on the parietal side. Aperture equipped with five main barriers (angulo-parietal, upper palatal, lower palatal, basal and columellar) and one small infraparietal lamella which is knob-like (although it can be absent as well). Angulo-parietal lamella is the strongest in the aperture, consisting of larger parietal part and indistinct angular part so that it appears slightly bilobed. Upper palatal plica short, weaker than the lower palatal and slightly bent towards it. Lower palatal plica roughly as strong as the columellar, situated halfway between upper palatal and basal plica. Basal plica small, slightly weaker than the upper palatal. Columellar lamella strong, directed towards the upper palatal plica and almost reaching the expanding peristome. Peristome is weakly swollen in front of the columellar lamella. Surface of all apertural barriers is finely granulated to almost smooth. Sinulus wide and not distinctly separated from the rest of the aperture. Umbilicus initially very narrow and then abruptly widening at the last whorl, measuring ~ 1 / 5–1 / 6 of the shell width. A weak groove is visible running alongside the umbilicus but it is getting lost near the middle part of the last whorl.</p><p>Differential diagnosis.</p><p>See under H. sculpturatum and H. khmerianum .</p><p>Measurements</p><p>(in mm, n = 4). SH = 1.93–2.46; SW = 2.09–2.24; AH = 1.02–1.16; AW = 1.05–1.18.</p><p>Etymology.</p><p>The two largest whorls (last and the penultimate) are keeled which gives the initial impression that the whole shell is keeled, hence the specific epithet.</p><p>Distribution.</p><p>This species is known only from the type locality.</p><p>Remarks.</p><p>Some specimens had a more depressed shell which was even slightly wider than high.</p></div>	https://treatment.plazi.org/id/44A493DFBA625C4FB85B476735F7175E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
6625028B828659898D58D5F30D7FF4F1.text	6625028B828659898D58D5F30D7FF4F1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma coriaceum Gojsina & Pall-Gergely 2025	<div><p>Hypselostoma coriaceum Gojšina &amp; Páll-Gergely sp. nov.</p><p>Figs 186 D, 191, 192, 223</p><p>Type material.</p><p>Holotype. Malaysia • 1 shell (SH: 2.11 mm; SW 1: 3.35 mm); Langkawi Island, Tanjung Rhu; 23 Aug. 1998; Hemmen, Wiesbaden ex. coll.; CUMZ 14464 . Paratypes. Malaysia • 30 shells; same data as for holotype; coll. PGB • 1 shell; same data as for holotype; coll. VG • 1 shell; same data as for holotype; coll. HA • 4 shells; Kedah, limestone outcrop near Tanjung Rhu, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.81081&amp;materialsCitation.latitude=6.4476333" title="Search Plazi for locations around (long 99.81081/lat 6.4476333)">Langkawi Island</a>; 6°26′51.48″N, 99°48′38.89″E; 16. May 2009; M. E. Marzuki leg.; coll. PGB .</p><p>Type locality.</p><p>Malaysia, Langkawi Island, Tanjung Rhu.</p><p>Diagnosis.</p><p>Shell depressed-conical, yellowish, and glossy. Last whorl bluntly keeled. Aperture detached from the penultimate whorl and slightly ascending upwards. Teleoconch finely granulated and radially striated. Spiral striation absent. Aperture equipped with five main strong barriers (parietal, upper and lower palatal, basal, and columellar) and additional smaller one of variable number.</p><p>Description.</p><p>The yellowish, glossy, and weakly translucent, depressed-conical shell is consisting of 4–4.5 whorls separated by a deep suture. The finely pitted protoconch is showing a weak spiralling pattern, consisting of ~ 1.5 whorls without a clear boundary with teleoconch. Teleoconch finely granulated, sandpaper-like (surface microsculpture similar to H. hungerfordianum) and inconspicuously regularly radially striated. Last whorl bluntly keeled, slightly detached from the penultimate and slightly to moderately ascending upwards (~ 25–35 ° compared to the shell axis). Peristome expanded and not reflected, with sharp edges. Aperture equipped with five main barriers (parietal, angular, upper palatal, lower palatal, and columellar) and variable number of smaller ones. Angular lamella is the weakest among the main five, the rest of the main barriers are developed roughly to the same level. They are all high and straight. Parietal and angular lamellae are directed towards the palatal wall. In front of the upper palatal plica, a small swelling is present which is probably homologous with the palatal tubercle present in other genera (e. g., Bensonella, see Páll-Gergely and White 2023). Between the angular lamella and upper palatal plica, one or two additional plicae are present. There are also usually two interpalatal plicae, two between lower palatal and columellar (including basal plica) and two between the columellar and parietal lamellae. None of these smaller barriers reach the profiles of the main ones, and none of the apertural barriers reach the peristome. Surface of all apertural barriers is very finely spiniferous medially and very finely granulated laterally. Sinulus parabolic and not very strongly separated from the rest of the aperture. Umbilicus open, initially narrow, and then abruptly widening at the last whorl, measuring ~ ¼ of the shell width and showing the penultimate whorl. A strong groove can be observed running from the peristome towards the inner side of the umbilicus.</p><p>Differential diagnosis.</p><p>This species is larger than both H. salpinx and H. saxicola and it additionally differs from them by the absence of spiral striation and a narrower umbilicus.</p><p>Measurements</p><p>. SH = 2.05–2.32; SW 1 = 3.12–3.37; SW 2 = 1.85–2.09; AH = 1.33–1.58; AW = 1.28–1.40.</p><p>Etymology.</p><p>The specific epithet is due to its “ leathery ” surface sculpture.</p><p>Distribution.</p><p>Known only from Tanjung Rhu, Langkawi Island, Malaysia.</p></div>	https://treatment.plazi.org/id/6625028B828659898D58D5F30D7FF4F1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
9CCDDBD20882511E8F199CB8F5C96193.text	9CCDDBD20882511E8F199CB8F5C96193.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma crossei Morlet 1886	<div><p>Hypselostoma crossei Morlet, 1886</p><p>Figs 242 B, 244, 245, 246, 247, 248, 249, 250, 251</p><p>Hypselostoma crossei Morlet, 1886: 2–3.</p><p>Hypselostoma crossei — Möllendorff 1891: 338.</p><p>Hypselostoma crossei brevituba Möllendorff, 1901 b: 76 . syn. nov.</p><p>Hypselostoma crossei endodonta Möllendorff, 1901 b: 76 . syn. nov.</p><p>Hypselostoma brevituba — Fischer and Dautzenberg 1904: 408.</p><p>Hypselostoma crossei endodonta — Fischer and Dautzenberg 1904: 408.</p><p>Gyliotrachela crossei — Pilsbry 1917: 215–216; van Benthem Jutting 1950: 44; Inkhavilay et al. 2019: 147, fig. 17 I; Thompson and Upatham 1997: 233.</p><p>Gyliotrachela crossei brevituba — Pilsbry 1917: 217, pl. 36, figs 9–13; van Benthem Jutting 1950: 45; Zilch 1984: 165; Schileyko 2011: 3.</p><p>Boysidia (Bensonella) xingyinensis Guo, Zhou &amp; Luo, 2006: 541–542 (Chinese description), 542–543 (English description), figs 1–4. syn. nov.</p><p>Gyliotrachela crossei endodonta — Pilsbry 1917: 217; van Benthem Jutting 1950: 45; Zilch 1984: 165; Schileyko 2011: 3.</p><p>Boysidia (Bensonella) tianxingqiaoensis Luo, Chen &amp; Zhang, 2000: 147 (Chinese description), 150 (English description), figs 1–4. syn. nov.</p><p>Boysidia (Bensonella) tianxingqiaoensis — Chen and Zhang 2002: 693.</p><p>Gyliotrachela crossei crossei — Schileyko 2011: 3.</p><p>Gyliotrachela plesiolopa Inkhavilay &amp; Panha in Inkhavilay et al. 2016: 222, figs 1, 3 D – F, 4 D. syn. nov.</p><p>Gyliotrachela plesiolopa — Inkhavilay et al. 2019: 60, fig. 26 D.</p><p>Boysidia tianxingqiaoensis — Dumrongrojwattana and Assawawattagee 2018: 216.</p><p>Boysidia xingyinensis — Dumrongrojwattana and Assawawttagee 2018: 216.</p><p>Gyliotrachela tianxingqiaoensis – Gojšina et al. 2022: 132, figs 2–5; Tongkerd et al. 2024: 182, fig. 13 N.</p><p>Type material examined.</p><p>Vietnam • syntype of H. crossei crossei; coll. Morlet; MNHN -IM-2000-35155 • lectotype of G. crossei brevituba; ex. coll. Fruhstorfer; coll. Möllendorff; SMF 4582 • lectotype of G. crossei endodonta; ex. coll. Fruhstorfer; coll. Möllendorff; SMF 4584 . China • holotype of B. tianxingqiaoensis; 13 Aug. 1997; D. - N. Chen leg.; IZCAS 025075 • holotype of B. xingyinensis; 02 Oct. 2004; Guo Y. - H, Zhou W. - C., Luo T. - C. leg.; IZCAS 069205 . Laos • 2 paratypes of G. plesiolopa; from the type locality; ex. coll. S. Panha; SMF 357824 .</p><p>Additional material examined.</p><p>Vietnam • 1 shell; Cao Bang Province, wet limestone wall; Jul. 2006.; M. Calo leg.; coll. PGB • 1 shell; Ninh Binh Province, Ninh Binh, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.96477&amp;materialsCitation.latitude=20.2623" title="Search Plazi for locations around (long 105.96477/lat 20.2623)">Hoa Lu Temple</a>; 20°15.738'N, 105°57.886'E; 8 m a. s. l.; 01 Oct. 2009; Hemmen leg; coll. PGB • 1 shell; Hoa Binh Province, ca 15 km rd. Tang Dao to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.03175&amp;materialsCitation.latitude=20.599184" title="Search Plazi for locations around (long 105.03175/lat 20.599184)">Co Luong</a> (rd # 15 left side); 20°35.951'N, 105°01.905'E; 16 Oct. 2011.; Hemmen leg; coll. PGB • 5 shells; Quang Ninh Province, Ha Long Bay area, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.12217&amp;materialsCitation.latitude=20.816" title="Search Plazi for locations around (long 107.12217/lat 20.816)">Tien Ong Cave on Hang Trai Island</a>; 20°48.96'N, 107°07.33'E; 6 Sept. 2003.; Hemmen leg; coll. PGB • 1 shell; Quang Ninh Province, near Dau Go Isl., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.01375&amp;materialsCitation.latitude=20.89755" title="Search Plazi for locations around (long 107.01375/lat 20.89755)">Halong Bay</a>; 20°53.853'N, 107°00.825'E; 2 m a. s. l.; 18 Nov. 2005; J. U. Otani leg.; coll. PGB • 23 shells; Ninh Binh Province, Nho Quan District, Cuc Phuong N. P., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.66772&amp;materialsCitation.latitude=20.29355" title="Search Plazi for locations around (long 105.66772/lat 20.29355)">near kiosk at entrance to Cave of the prehistoric man (Dong Nguoi Xua)</a>; 20°17.613'N, 105°40.063'E; 247 m a. s. l.; 12 Apr. 2007; K. Okubo &amp; J. U. Otani leg.; coll. PGB • 2 shells; Quang Ninh Province, Halong Bay area, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.12217&amp;materialsCitation.latitude=20.816" title="Search Plazi for locations around (long 107.12217/lat 20.816)">Hang Trai Id.</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.12217&amp;materialsCitation.latitude=20.816" title="Search Plazi for locations around (long 107.12217/lat 20.816)">Tieng Ong cave</a>; 20°48.96'N, 107°07.33'E; Hemmen leg; coll. PGB • 15 shells; Hai Phong city, Cat Ba island, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.98268&amp;materialsCitation.latitude=20.8567" title="Search Plazi for locations around (long 106.98268/lat 20.8567)">Gia Luan Commune</a>; 20°51.402'N, 106°58.961'E; 4 m a. s. l.; 18 Nov. 2005; J. U. Otani leg.; Coll, PGB • 3 shells; Cao Bang Province, Pac Rao, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.51237&amp;materialsCitation.latitude=22.823084" title="Search Plazi for locations around (long 106.51237/lat 22.823084)">300 m from junction to Xa Canh Tien towards Trung Khanh, right side of the road</a>; 22°49.385'N, 106°30.742'E; 530 m a. s. l.; 28 May 2012; A. Hunyadi leg.; coll. HA • 2 shells; Lang Son Province, Lang Son, NNE side of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.74917&amp;materialsCitation.latitude=21.85305" title="Search Plazi for locations around (long 106.74917/lat 21.85305)">Nui Vong Phu</a>; 21°51.183'N, 106°44.950'E; 11 Nov. 2011; A. Hunyadi leg.; coll. HA • 1 shell; Lang Son Province, north of Chi Lang, pass along the path (north of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.50945&amp;materialsCitation.latitude=21.582417" title="Search Plazi for locations around (long 106.50945/lat 21.582417)">Đong Banh</a>); 21°34.945'N, 106°30.567'E; 75 m a. s. l.; 13 Nov. 2011; A. Hunyadi leg.; coll. HA • 5 shells; Lang Son, vicinity of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.74682&amp;materialsCitation.latitude=21.855883" title="Search Plazi for locations around (long 106.74682/lat 21.855883)">Chua Tam Thanh</a>; 21°51.353'N, 106°44.809'E; 265 m a. s. l.; 21 Feb. 2020; A. Hunyadi leg.; coll. HA • 1 shell; Thai Nguyen Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.65325&amp;materialsCitation.latitude=21.910217" title="Search Plazi for locations around (long 105.65325/lat 21.910217)">Cho Chu</a>, rock wall at the northeastern part of the village; 21°54.613'N, 105°39.195'E; 90 m a. s. l.; 21 Nov. 2011; A. Hunyadi leg.; coll. HA • 34 shells: Lang Son Province, Lang Son, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.75058&amp;materialsCitation.latitude=21.855967" title="Search Plazi for locations around (long 106.75058/lat 21.855967)">Thanh Nha Mac</a>, southern group of rocks; 21°51.358'N, 106°45.035'E; 295 m a. s. l.; 26 May 2012; leg. A. Hunyadi; coll. HA • 7 shells; Lang Son Province, Huu Lung district, Minh Tien, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.29045&amp;materialsCitation.latitude=21.560083" title="Search Plazi for locations around (long 106.29045/lat 21.560083)">Cau Cheo Minh Tien</a>, southern side of the bridge; 21°33.605'N, 106°17.427'E; 20 m a. s. l.; 20 Feb. 2020; A. Hunyadi leg.; coll. HA • 7 shells; Thanh Hoa Province, north of Thanh Hoa, Ham Rong Eny, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.77972&amp;materialsCitation.latitude=19.8518" title="Search Plazi for locations around (long 105.77972/lat 19.8518)">Đong Tien Son</a>; 19°51.108'N, 105°46.783'E; 35 m a. s. l.; 24 Nov. 2011; A. Hunyadi leg.; coll. HA • 60 shells; Hoa Binh Province, Tan Lac district, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.26955&amp;materialsCitation.latitude=20.63222" title="Search Plazi for locations around (long 105.26955/lat 20.63222)">Quy Hau</a>, 1300 m west on road no. 6, rock wall; 20.63222°N, 105.26955°E; 150 m a. s. l.; 11 Feb. 2020; A. Hunyadi, H. V. Luong, J. U. Otani &amp; S. V. Pham leg.; coll. HA • 54 shells; Son La Province, 32.2 km northwest from centre of Son La towards Tuan Giao, Phong Lang, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.73275&amp;materialsCitation.latitude=21.43805" title="Search Plazi for locations around (long 103.73275/lat 21.43805)">Tong Lanh</a>, northeastern part of the village; 21°26.283'N, 103°43.965'E; 550 m a. s. l.; 07 Feb. 2020; A. Hunyadi, H. V. Luong, J. U. Otani &amp; S. V. Pham leg.; coll. HA • 180 shells; Son La Province, Quynh Nhai district, 20 km north from junction of Thuan Chau, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.681816&amp;materialsCitation.latitude=21.55735" title="Search Plazi for locations around (long 103.681816/lat 21.55735)">Chieng Khoang</a>, cave above the village; 21°33.441'N, 103°40.909'E; 315 m a. s. l.; 07 Feb. 2020; A. Hunyadi, H. V. Luong, J. U. Otani &amp; S. V. Pham leg.; coll. HA • 155 shells; Thanh Hoa Province, Quan Hoa district, Khu di tich <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.06693&amp;materialsCitation.latitude=20.398134" title="Search Plazi for locations around (long 105.06693/lat 20.398134)">Hang Ma</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.06693&amp;materialsCitation.latitude=20.398134" title="Search Plazi for locations around (long 105.06693/lat 20.398134)">Song Luong</a> gorge; 20°23.888'N, 105°4.016'E; 100 m a. s. l.; 12 Feb. 2020; A. Hunyadi, H. V. Luong, J. U. Otani &amp; S. V. Pham leg.; coll. HA • 386 shells; Thanh Hoa Province, 3.6 km northwest from centre of Ngoc Lac, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.359634&amp;materialsCitation.latitude=20.098284" title="Search Plazi for locations around (long 105.359634/lat 20.098284)">Lang Sat, 600 m north on the road no 15</a>, under rock wall; 20°5.897'N, 105°21.578'E; 55 m a. s. l.; 13 Feb. 2020; A. Hunyadi, H. V. Luong, J. U. Otani &amp; S. V. Pham leg.; coll. HA • 51 shells; Hoa Binh Province, Mai Chau, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.07804&amp;materialsCitation.latitude=20.66029" title="Search Plazi for locations around (long 105.07804/lat 20.66029)">550 m west of Cho Mai Chau</a>, stream bank; 20.66029°N, 105.07804°E; 265 m a. s. l.; 11 Feb. 2020; A. Hunyadi, H. V. Luong, J. U. Otani &amp; S. V. Pham leg.; coll. HA . China • 1 shell; Guangxi, Guigang Shi, Guzhang Xiang, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=109.31948&amp;materialsCitation.latitude=23.34775" title="Search Plazi for locations around (long 109.31948/lat 23.34775)">Chuanshancun</a>, road leading to north, limestone rock wall on the east; 23°20.865'N, 109°19.169'E; 150 m a. s. l.; 10 Sept. 2009; A. Hunyadi leg.; coll. HA • 17 shells; Guangxi, Chongzuo Shi, Longzhou Xian, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.768265&amp;materialsCitation.latitude=22.587317" title="Search Plazi for locations around (long 106.768265/lat 22.587317)">Wude Xiang</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.768265&amp;materialsCitation.latitude=22.587317" title="Search Plazi for locations around (long 106.768265/lat 22.587317)">Banxintun</a>, vicinity of the junction; 22°35.239'N, 106°46.096'E; 350 m a. s. l.; 24 Sept. 2013; A. Hunyadi &amp; M. Szekeres leg.; coll. HA • 2 shells; Yunnan, Yuxi shi, Chengjiang Xian, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.84357&amp;materialsCitation.latitude=24.564034" title="Search Plazi for locations around (long 102.84357/lat 24.564034)">Luchong Ziran Fengjingqu</a>, under <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.84357&amp;materialsCitation.latitude=24.564034" title="Search Plazi for locations around (long 102.84357/lat 24.564034)">Liushanting</a>; 24°33.842'N, 102°50.614'E; 1778 m a. s. l.; 18 Mar. 2011; A. Hunyadi leg.; coll. HA • 2 shells; Guangxi, Guigang Shi, Guzhang Xiang, rocks above <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=109.320786&amp;materialsCitation.latitude=23.350933" title="Search Plazi for locations around (long 109.320786/lat 23.350933)">Chuanshancun</a>; 23°21.056'N, 109°19.247'E; 200 m a. s. l.; 10 Sept. 2009; A. Hunyadi leg.; coll. HA • 9 shells; Guangxi, Laibin Shi, Wuxuan Xian, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=109.642235&amp;materialsCitation.latitude=23.497116" title="Search Plazi for locations around (long 109.642235/lat 23.497116)">Wuxuan Tongling Zhen</a> road, north of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=109.642235&amp;materialsCitation.latitude=23.497116" title="Search Plazi for locations around (long 109.642235/lat 23.497116)">Yagangcun</a>, rock wall; 23°29.827'N, 109°38.534'E; 21 Sept. 2013; A. Hunyadi &amp; M. Szekeres leg.; coll. HA • 3 shells; Guangxi, Hechi Shi, Du an <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.0215&amp;materialsCitation.latitude=24.053284" title="Search Plazi for locations around (long 108.0215/lat 24.053284)">Yaozu Zizhixian</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.0215&amp;materialsCitation.latitude=24.053284" title="Search Plazi for locations around (long 108.0215/lat 24.053284)">Gaoling Xiang</a>, 2 km west from <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.0215&amp;materialsCitation.latitude=24.053284" title="Search Plazi for locations around (long 108.0215/lat 24.053284)">Dingfucun</a>; 24°03.197'N, 108°01.290'E; 320 m a. s. l.; 10 Aug. 2009; A. Hunyadi leg.; coll. HA • 17 shells; Yunnan, Wenshan Zhuangzu Miaozu Zizhizhou, Guangnan Xian, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.41458&amp;materialsCitation.latitude=23.7447" title="Search Plazi for locations around (long 105.41458/lat 23.7447)">Babao Zhen</a>, southeastern edge of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.41458&amp;materialsCitation.latitude=23.7447" title="Search Plazi for locations around (long 105.41458/lat 23.7447)">Babao</a>; 23°44.682'N, 105°24.875'E; 1130 m; 24 Mar. 2011; A. Hunyadi leg.; coll. HA • 33 shells; Guangxi, Laibin Shi, Wuxuan Xian, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=109.63958&amp;materialsCitation.latitude=23.48585" title="Search Plazi for locations around (long 109.63958/lat 23.48585)">Wuxuan Tongling Zhen</a> road, west from <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=109.63958&amp;materialsCitation.latitude=23.48585" title="Search Plazi for locations around (long 109.63958/lat 23.48585)">Yagangcun</a>, rocks; 23°29.151'N, 109°38.375'E; 80 m a. s. l.; 21 Sep 2013; A. Hunyadi &amp; M. Szekeres leg.; coll. HA • 11 shells; Guangxi, Bose Shi, Leye Xian, southern edge of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.72075&amp;materialsCitation.latitude=24.657267" title="Search Plazi for locations around (long 106.72075/lat 24.657267)">Molicun</a>, left bank of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.72075&amp;materialsCitation.latitude=24.657267" title="Search Plazi for locations around (long 106.72075/lat 24.657267)">Buliu He</a>; 24°39.436'N, 106°43.245'E; 540 m a. s. l.; 08 Sep. 2013; A. Hunyadi &amp; M. Szekeres leg.; coll. HA • 17 shells; Guangxi, Bose Shi, Lingyun Xian, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.6396&amp;materialsCitation.latitude=24.2918" title="Search Plazi for locations around (long 106.6396/lat 24.2918)">Xiajia Xiang</a>, southeastern edge of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.6396&amp;materialsCitation.latitude=24.2918" title="Search Plazi for locations around (long 106.6396/lat 24.2918)">Xiajia</a>, rock wall above the graves; 24°17.508'N, 106°38.376'E; 440 m a. s. l.; 22 Oct. 2009; A. Hunyadi leg.; coll. HA • 5 shells; village across from Yangchangshilin, Moyangzhen, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.82754&amp;materialsCitation.latitude=25.544167" title="Search Plazi for locations around (long 106.82754/lat 25.544167)">Luodianxian</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.82754&amp;materialsCitation.latitude=25.544167" title="Search Plazi for locations around (long 106.82754/lat 25.544167)">Guizhousheng</a>; 25°32′39″N, 106°49′39.14″E; 908 m a. s. l.; T. Ishibe, K. Okubo, J. U. Otani leg.; coll. PGB • 4 shells; same data as previous; coll. PGB • 9 shells; Daxiaojingfengjingqu, Moyangzhen, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.86083&amp;materialsCitation.latitude=25.560833" title="Search Plazi for locations around (long 106.86083/lat 25.560833)">Lodianxian</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.86083&amp;materialsCitation.latitude=25.560833" title="Search Plazi for locations around (long 106.86083/lat 25.560833)">Guizhousheng</a>; 25°33′39″N, 106°51′39″E; 450 m a. s. l.; T. Ishibe, K. Okubo, J. U. Otani leg.; coll. PGB • 8 shells; same data as previous; T. Ishibe, K. Okubo, J. U. Otani leg.; coll. PGB • 1 shell; Xianshanyuan, Shuangqiao-zhen, Wuming-xian, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.29476&amp;materialsCitation.latitude=23.03467" title="Search Plazi for locations around (long 108.29476/lat 23.03467)">Guangxi Zhuanzu Zizhiou</a>; 23°02.08015'N, 108°17.68551'E; 177 m a. s. l.; Ohai leg.; coll. PGB .</p><p>Type localities.</p><p>“ Tonkin ” (Montagne de ľÉléphant, Tonkin, Vietnam) ( H. crossei crossei); “ Lang-son ” (Tonkin, N Vietnam) ( H. crossei brevituba); “ Lang-son ” (Tonkin, N Vietnam) ( H. crossei endodonta); “ collected from Tianxingqiao Town, Zhenning Bouyeizu Miaozu Zizhixian, Guizhou Province (26°N, 105°07'E), Guizhou Province ”, China ( G. tianxingqiaoensis); “ collected from Nahui Town, Xingyi County (25°1'N, 104°8'E), Guizhou Province, China ” ( G. xingyinensis); “ Limestone outcrop in Nawit Village, Viengxay District, Houaphane Province, Laos (20°22'37.3"N, 104°16'43.2"E), 695 m amsl ” ( G. plesiolopa).</p><p>Differential diagnosis.</p><p>This species differs from all its congeners by the combination of the dark brown shell, which is moderately umbilicate, numerous and variable apertural barriers, almost smooth shell surface (pasty), as well as the absence of the spiral striation.</p><p>Distribution.</p><p>This species is known from a wide area covering China, N Vietnam, Myanmar, and Laos (from where it is known as G. plesiolopa).</p><p>Remarks.</p><p>The examination of type specimens of H. crossei brevituba and H. crossei endodonta and numerous additional samples of H. crossei, led us to the conclusion that this species is very variable regarding the appearance of the last whorl and also regarding barrier numbers, as was also observed by Pilsbry (1917). These were the characters used to describe the subspecies and there are no reasons to support their distinctness. Gyliotrachela plesiolopa, G. xingyinensis and G. tianxingqiaoensis are junior synonyms of this species since no major morphological differences were observed. The latter two are originally described in exactly the same way, using exactly the same sentences, and are not compared mutually (or to H. crossei) but to other very distant species (Luo et al. 2000; Guo et al. 2006). Inkhavilay et al. (2016) pointed out that G. plesiolopa has fewer whorls (four) than H. crossei (five). However, this is not true due to the fact that we examined the types of the nominotypical subspecies of H. crossei and the other two described subspecies all of which showed between 3.5–4 whorls. According to them, H. crossei is also larger, but there is a clear continuum in size variation up to 1 mm. Therefore, we do not consider this character as relevant.</p></div>	https://treatment.plazi.org/id/9CCDDBD20882511E8F199CB8F5C96193	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
A15C56E7CB1B581C8B04FC2C5095E548.text	A15C56E7CB1B581C8B04FC2C5095E548.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma cucumense Panha 1998	<div><p>Hypselostoma cucumense Panha, 1998</p><p>Figs 111 L, 128, 154</p><p>Hypselostoma cucumensis Panha, 1998 b: 65–66, figs 1 b, 3.</p><p>Hypselostoma cucumensis — Hemmen and Hemmen 2001: 41; Panha and Burch 2008: 90, fig. 77; Tongkerd et al. 2004: 143, fig. 2; Dumrongrojwattana and Wongkamhaeng 2024: 324, fig. 8.</p><p>Material examined.</p><p>Thailand • 35 shells; Phetchaburi Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.82488&amp;materialsCitation.latitude=13.2386" title="Search Plazi for locations around (long 99.82488/lat 13.2386)">Khao Yoi</a>, northeastern side of the mountain; 13°14.316'N, 99°49.493'E; 30 m a. s. l.; 24 Feb. 2015; A. Hunyadi leg.; coll. HA .</p><p>Type locality.</p><p>“ Limestone hills at Kangkrajan National Park, Petchaburi Province, 13°14′9″N, 99°49′43″E, 80 meters elevation ” (Thailand).</p><p>Differential diagnosis.</p><p>This species resembles H. adela in shell size, surface sculpture and colouration. However, in H. adela, the angular and parietal lamellae are separated, the barriers are generally stronger and the last whorl is always at least slightly descending (slightly ascending in H. cucumense). See also under H. populare sp. nov. and H. khaochongpran .</p><p>Distribution.</p><p>This species is known from the type locality and its immediate surroundings.</p></div>	https://treatment.plazi.org/id/A15C56E7CB1B581C8B04FC2C5095E548	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
4F79B0374FEE57BFA78B9D8FC032481A.text	4F79B0374FEE57BFA78B9D8FC032481A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma cultura (Tanmuangpak & Dumrongrojwattana 2022) Gojšina & Hunyadi & Sutcharit & Tongkerd & Auffenberg & Grego & Vermeulen & Reischütz & Páll-Gergely 2025	<div><p>Hypselostoma cultura (Tanmuangpak &amp; Dumrongrojwattana, 2022) comb. nov.</p><p>Figs 111 M, 127, 154</p><p>Gyliotrachela cultura Tanmuangpak &amp; Dumrongrojwattana, 2022: 408–415, fig. 2.</p><p>Material examined.</p><p>Thailand • 52 shells; Chonburi Province, Ko Sichang, Tha Thewawong, vicinity of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.805084&amp;materialsCitation.latitude=13.170934" title="Search Plazi for locations around (long 100.805084/lat 13.170934)">Mondop Roi Phraphutthabat</a>; 13°10.256'N, 100°48.305'E; 140 m a. s. l.; 11 Mar. 2023; A. Hunyadi leg.; coll. HA • 3 shells; Chonburi Province, Si Racha, Ko Si Chang; 25 Oct. 2019; J. U. Otani leg.; coll. PGB • 2 shells; Lampang Province, Maemoh district, Wat Tam Innaeramitr; 10 Aug. 2014; J. U. Otani leg.; coll. PGB .</p><p>Type locality.</p><p>“ Agricultural areas in Mueang Loei District, Loei Province, Thailand (17°34′43.020″N, 101°51′04.020″E) ”.</p><p>Differential diagnosis.</p><p>See under H. khaowongense .</p><p>Distribution.</p><p>This species is, apart from the type locality in Loei province, known from two more localities: one in Lampang and one in Chonburi province.</p><p>Remarks.</p><p>This species might be conspecific with H. khaowongense . The differences mentioned by Tanmuangpak and Dumrongrojwattana (2022) between this species and H. khaowongense (therein under Gyliotrachela saraburiensis) are only regarding the number of apertural barriers, which are very variable in H. khaowongense . We have noticed additionally that the shells of H. cultura are strongly shouldered when observed laterally (e. g., as in Tanmuangpak and Dumrongrojwattana 2022: Fig. 2 E, K) which is not typical for H. khaowongense . Due to the absence of relevant comparative material, we do not formally synonymise the two species. Specimens examined by us from Chonburi (Fig. 127) and Lampang provinces show very similar shell morphology as figured in Tanmuangpak and Dumrongrojwattana (2022), just with a wider umbilicus. We do not intend to describe this species as separate from H. cultura until more comparative material becomes available.</p></div>	https://treatment.plazi.org/id/4F79B0374FEE57BFA78B9D8FC032481A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
1006C0BD8D015D499CF0FC8C8121A780.text	1006C0BD8D015D499CF0FC8C8121A780.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma depressispira (van Benthem Jutting 1949) Gojšina & Hunyadi & Sutcharit & Tongkerd & Auffenberg & Grego & Vermeulen & Reischütz & Páll-Gergely 2025	<div><p>Hypselostoma depressispira (van Benthem Jutting, 1949) comb. nov.</p><p>Figs 242 C, 252, 259</p><p>Gyliotrachela depressispira van Benthem Jutting, 1949 b: 60, pl. 3.</p><p>Gyliotrachela depressispira — van Benthem Jutting 1950: 27, 45, fig. 15; Berry 1963: 361; Maassen 2001: 75.</p><p>Type material examined.</p><p>Malaysia • 12 paratypes; Pahang, Bukit Chintamani; ex. coll. Raffles Museum, Singapore; RMNH.Moll.137152 .</p><p>Additional material examined.</p><p>Malaysia • 2 shells; Pahang, 25 km northwest from Kuantan, north of Kampung Panching, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.147316&amp;materialsCitation.latitude=3.9115334" title="Search Plazi for locations around (long 103.147316/lat 3.9115334)">Gua Charas</a>, rock temple; 03°54.692'N, 103°08.839'E; 140 m a. s. l.; 24 Jan. 2013; A. Hunyadi leg.; coll. HA • 28 shells; Kelantan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.9686&amp;materialsCitation.latitude=4.8829" title="Search Plazi for locations around (long 101.9686/lat 4.8829)">Gua Musang</a>, vicinity of cave entrance; 04°52.974'N, 101°58.116'E; 135 m a. s. l.; 17 Jan. 2013; A. Hunyadi leg.; coll. HA • 37 shells; Kelantan, Gua Musang, 6 km towards Pulai, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.9497&amp;materialsCitation.latitude=4.8368835" title="Search Plazi for locations around (long 101.9497/lat 4.8368835)">Gua Madu</a>, vicinity of cave, 04°50.213'N, 101°56.982'E; 120 m a. s. l.; 17 Jan. 2013; A. Hunyadi leg.; coll. HA • 25 shells; Pahang, Gua Bama, Kuala Lipis 9 km, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.9656&amp;materialsCitation.latitude=4.1942" title="Search Plazi for locations around (long 101.9656/lat 4.1942)">Padang Tungku</a>; 04°11.652'N, 101°57.936'E; 120 m a. s. l.; 19 Jan. 2013; A. Hunyadi leg.; coll. HA • 201 shells; Pahang, Bukit Cinta Manis, south-southeastern side, Lebuhraya Karak 800 m towards <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.013565&amp;materialsCitation.latitude=3.4452333" title="Search Plazi for locations around (long 102.013565/lat 3.4452333)">Kampung Cinta Manis</a>; 03°26.714'N, 102°00.814'E; 22 Jan. 2013; A. Hunyadi leg.; coll. HA • 3 shells; Pahang, Kota Gelanggi, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.47314&amp;materialsCitation.latitude=3.8912637" title="Search Plazi for locations around (long 102.47314/lat 3.8912637)">Jengka-Jerantut</a> road, on limestone surfaces covered with mosses; 03°53′28.55″N, 102°28′23.28″E; 29. May 2011; M. E. Marzuki leg.; coll. PGB • 4 shells; Pahang, Gua Bama at Kp. Relong (~ 14 km N of Kuala Lipis); Oct. 1998; Hemmen leg; coll. PGB • 6 shells; Pahang, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.965416&amp;materialsCitation.latitude=4.19405" title="Search Plazi for locations around (long 101.965416/lat 4.19405)">Gua Bama</a> (~ 12 / 13 km N of Kuala Lipis); 04°11.643'N, 101°57.925'E; Nov. 2000; Hemmen leg; coll. PGB .</p><p>Type locality.</p><p>“ Bukit Chintamani, Pahang ”, Malaysia.</p><p>Differential diagnosis.</p><p>This species bears some resemblance to the more depressed form of H. fortunatum sp. nov. However, H. depressispira has much more apertural barriers as well as separated angular and parietal lamellae. The last whorl is also more prominently keeled in H. depressispira and the umbilicus is slightly wider. See also under H. tubiferum .</p><p>Distribution.</p><p>This species is known from Peninsular Malaysia.</p><p>Remarks.</p><p>While examining the types, variability was observed in the smaller barriers. In some specimens, there were two and in others three interpalatal plicae. The number of lamellae between the columellar and parietal ranged between three and four. Number of barriers between lower palatal plica and columellar lamella ranged from two to three. Some specimens from Gua Musang (Kelantan) had only one barrier between the lower palatal and columellar and only one dot-like barrier between the columellar and parietal. Otherwise, the apertural barrier variability of other samples examined by us matches that of the type series.</p></div>	https://treatment.plazi.org/id/1006C0BD8D015D499CF0FC8C8121A780	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
5708339FAFB45FC3B90AB009FD182773.text	5708339FAFB45FC3B90AB009FD182773.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma depressum (Vermeulen, Luu, Theary & Anker 2019) Gojšina & Hunyadi & Sutcharit & Tongkerd & Auffenberg & Grego & Vermeulen & Reischütz & Páll-Gergely 2025	<div><p>Hypselostoma depressum (Vermeulen, Luu, Theary &amp; Anker, 2019) comb. nov.</p><p>Figs 111 N, 129, 182</p><p>Anauchen depressus Vermeulen et al., 2019: 35–37, figs 60–63.</p><p>Anauchen depressus — Sutcharit et al. 2023: 1290.</p><p>Type material examined.</p><p>Cambodia • holotype; NHMUK 20180574 .</p><p>Type locality.</p><p>“ Cambodia: Kampot Province, Kampot area: Phnom La’Ang, cave with shrine at its entrance, first chamber with collapsed roof … southeast end … ”.</p><p>Differential diagnosis.</p><p>This species differs from H. chaunosalpinx by the keeled last whorl and less numerous apertural barriers, none of which are hooked. The umbilicus is also wider in H. chaunosalpinx . See also under H. taehwani .</p><p>Distribution.</p><p>This species is known only from the type locality.</p><p>Remarks.</p><p>The type locality of this species was inaccessible during collecting efforts in 2023 due to the restricted access and the presence of concrete walls around the locality. This indicates that the species could be seriously threatened by quarrying.</p></div>	https://treatment.plazi.org/id/5708339FAFB45FC3B90AB009FD182773	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
9D3638C42F885881A13CF3B751B4DEE6.text	9D3638C42F885881A13CF3B751B4DEE6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma diarmaidi (Panha & J. B. Burch 2003) Gojšina & Hunyadi & Sutcharit & Tongkerd & Auffenberg & Grego & Vermeulen & Reischütz & Páll-Gergely 2025	<div><p>Hypselostoma diarmaidi (Panha &amp; J. B. Burch, 2003) comb. nov.</p><p>Figs 111 O, 130, 131, 154</p><p>Gyliotrachela diarmaidi Panha &amp; Burch in Burch et al. 2003: 157–162, figs 10, 11.</p><p>Gyliotrachela diarmaidi — Tongkerd et al. 2004: 143, fig. 2; Panha and Burch 2008: 67–69, fig. 59; Dumrongrojwattana and Wongkamhaeng 2024: 323, fig. 8.</p><p>Material examined.</p><p>Thailand • 2 shells; Rayong Province, Wang Chan; 10 Aug. 2013 • 9 shells; Songkhla Province, 31.3 km NW Hat Yai 1.2 km W of Hwy. 43. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2667&amp;materialsCitation.latitude=7.1667" title="Search Plazi for locations around (long 100.2667/lat 7.1667)">Limestone</a> outcrop, evergreen forest, leaf litter, roots and sticks, small solution pits; 7.1667°N, 100.2667°E; 80 m a. s. l.; 08 Apr. 1988; K. Auffenberg leg.; UF 344927 • 1 shell; same data as previous; locality code KA-0636 B; UF 344944 .</p><p>Type locality.</p><p>“ Pluangthong Mountain, west of Pluangthong Temple, Botong District, Chonburi Province, 13°11'5"N, 101°34'59"E, 110 meters elevation, Thailand ”.</p><p>Differential diagnosis.</p><p>See under H. kohrin and H. surakiti .</p><p>Distribution.</p><p>Apart from the type locality, this species is known from one other sampling site near the type locality as well as from the Rayong, Sa Kaeo, Prachinburi, and Songkhla provinces (Dumrongrojwattana and Wongkamhaeng 2024).</p></div>	https://treatment.plazi.org/id/9D3638C42F885881A13CF3B751B4DEE6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
B839F2EC620B5114979CA2D3A83BB218.text	B839F2EC620B5114979CA2D3A83BB218.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma dilatatum van Benthem Jutting 1962	<div><p>Hypselostoma dilatatum van Benthem Jutting, 1962</p><p>Figs 111 P, 132, 182</p><p>Hypselostoma dilatatum van Benthem Jutting, 1962: 5–6, fig. 2.</p><p>Hypselostoma dilatatum — Schileyko 2011: 3; Vermeulen et al. 2019: 39.</p><p>Type material examined.</p><p>Vietnam • 2 paratypes; Nui Xo Ngach, 16 km SE of Haiten; 1960; E. Saurin leg.; RMNH.Moll.137167 .</p><p>Additional material examined.</p><p>Vietnam • 4 shells; Kien Giang Province, Kien Luong district, Binh An commune, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.61625&amp;materialsCitation.latitude=10.22765" title="Search Plazi for locations around (long 104.61625/lat 10.22765)">Ba Nui village</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.61625&amp;materialsCitation.latitude=10.22765" title="Search Plazi for locations around (long 104.61625/lat 10.22765)">cave Mo So</a>, limestone rocks; 10°13.659'N, 104°36.975'E; 18 Sept. 2018; I. Dedov, N. Simov, R. Bekchiev, P. Beron leg., coll. PGB .</p><p>Type locality.</p><p>“ Xa Ngach, à 16 km. au S. - E. de Hatien, Sud Vietnam ”.</p><p>Differential diagnosis.</p><p>This species is similar to H. cambodjense from which it differs by its hooked apertural barriers and a narrower umbilicus. See also under H. rupestre .</p><p>Distribution.</p><p>This species is known from the Vietnam part of the Mekong Delta limestone.</p><p>Remarks.</p><p>One paratype is figured, the second paratype was corroded and with broken hooked barriers in the aperture. It was also slightly larger than the figured specimen.</p></div>	https://treatment.plazi.org/id/B839F2EC620B5114979CA2D3A83BB218	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
DE7A437EA5D75E6890195C1AB86ED722.text	DE7A437EA5D75E6890195C1AB86ED722.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma discobasis Vermeulen, Luu, Theary & Anker 2019	<div><p>Hypselostoma discobasis Vermeulen, Luu, Theary &amp; Anker, 2019</p><p>Figs 111 Q, 133, 182</p><p>Hypselostoma discobasis Vermeulen et al., 2019: 39–40, figs 66–68.</p><p>Hypselostoma discobasis — Sutcharit et al. 2023: 1290.</p><p>Type material examined.</p><p>Cambodia • holotype; NHMUK 20180576 .</p><p>Type locality.</p><p>“ Cambodia: Kampot Province, Kampot area: Phnom La’Ang, southeast-end ”.</p><p>Differential diagnosis.</p><p>See under H. sorormajor sp. nov. and H. sororminor sp. nov.</p><p>Distribution.</p><p>This species in known only from the type locality.</p><p>Remarks.</p><p>The type locality of this species was inaccessible during collecting efforts in 2023 due to the restricted access and the presence of concrete walls around the locality. This indicates that the species could be seriously threatened by quarrying.</p></div>	https://treatment.plazi.org/id/DE7A437EA5D75E6890195C1AB86ED722	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
1F3670D619EB5212998F3831EF24233F.text	1F3670D619EB5212998F3831EF24233F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma edentatum (Panha & J. B. Burch 2002)	<div><p>Hypselostoma edentatum (Panha &amp; J. B. Burch, 2002)</p><p>Figs 111 R, 134, 154</p><p>Systenostoma edentatum Panha &amp; Burch, 2002 d: 121–124, fig. 4.</p><p>Systenostoma edentata — Panha and Burch 2008: 119–120, fig. 102.</p><p>Hypselostoma edentata — Jochum et al. 2014: 31; Dumrongrojwattana and Wongkamhaeng 2024: 324, fig. 8.</p><p>Type material examined.</p><p>Thailand • holotype; 1997; S. Panha leg.; CUMZ ver. 022 .</p><p>Additional material examined.</p><p>Thailand • 12 shells; Chiang Rai Province, limestone knoll, 4 km NE of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.98722&amp;materialsCitation.latitude=19.571945" title="Search Plazi for locations around (long 99.98722/lat 19.571945)">Ban Pa Ngae</a> limestone knoll, base of limestone ledge; 19°34.3167'N, 99°59.2333'E; 410 m a. s. l.; F. G. Thompson leg.; 12. May 1988; locality code FGT-4429; UF 380304 .</p><p>Type locality.</p><p>“ Tamphatai National Park …, Phrae Province, 18 ° 36 ” 20 ' N, 99 ° 53 ” 49 ' E, 650 meters elevation …, Thailand … ”.</p><p>Differential diagnosis.</p><p>This species differs from all of its congeners by the absence of apertural barriers, which makes it superficially more similar to some Tonkinospira species. From all similar toothless Aulacospira species from Thailand, this species differs by the presence of raised spiral striation. See also under H. panhai .</p><p>Distribution.</p><p>This species in known from Lampang and Chiang Rai provinces in Thailand.</p><p>Remarks.</p><p>The type locality of this species is not in Phrae but in Lampang province. Specimens from Chiang Rai province shared the same shell morphology but were slightly more depressed than the holotype.</p></div>	https://treatment.plazi.org/id/1F3670D619EB5212998F3831EF24233F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
90B0F1AA1BE2571ABB7A656C2E813548.text	90B0F1AA1BE2571ABB7A656C2E813548.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma elephas van Benthem Jutting 1950	<div><p>Hypselostoma elephas van Benthem Jutting, 1950</p><p>Figs 224 A, 225, 241</p><p>Hypselostoma elephas van Benthem Jutting, 1950: 23–24, fig. 12.</p><p>Hypselostoma elephas — van Benthem Jutting 1950: 41; Maassen 2001: 77.</p><p>Boysidia (Dasypupa) elephas — Thompson and Dance 1983: 109.</p><p>Type material examined.</p><p>Malaysia • holotype; RMNH.Moll.137159 .</p><p>Type locality.</p><p>“ Bukit Tenggek, Pahang ”, Malaysia.</p><p>Differential diagnosis.</p><p>See under H. terae .</p><p>Distribution.</p><p>This species in known from Bukit Tenggek and Gunung Sagu (IUCN, 2015).</p><p>Remarks.</p><p>This species is critically endangered due to the heavy quarrying of localities where it occurs (Clements 2009; IUCN 2015).</p></div>	https://treatment.plazi.org/id/90B0F1AA1BE2571ABB7A656C2E813548	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
956C9AEC0FC25B8AABB2932BFBB74568.text	956C9AEC0FC25B8AABB2932BFBB74568.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma emergens (van Benthem Jutting 1950) Gojšina & Hunyadi & Sutcharit & Tongkerd & Auffenberg & Grego & Vermeulen & Reischütz & Páll-Gergely 2025	<div><p>Hypselostoma emergens (van Benthem Jutting, 1950) comb. nov.</p><p>Figs 224 B, 226, 241</p><p>Gyliotrachela emergens van Benthem Jutting, 1950: 34–35, fig. 20.</p><p>Gyliotrachela emergens — van Benthem Jutting 1950: 45; Zilch 1984: 165; Maassen 2001: 75.</p><p>Type material examined.</p><p>Malaysia • 54 paratypes; from the type locality; 1939; Raffles Museum, Singapore ex. coll.; RMNH.Moll.137150 .</p><p>Additional material examined.</p><p>Malaysia • 2 shells; Perlis, ca 2 km S of Bukit Keteri (NE of Kangar); 22 Mar. 1998; Hemmen leg.; coll. PGB .</p><p>Type locality.</p><p>“ Bukit Chuping, Perlis ”, Malaysia.</p><p>Differential diagnosis.</p><p>See under H. modestum .</p><p>Distribution.</p><p>This species is known from Bukit Chuping (type locality) and Bukit Keteri (this study), both located in Perlis, N Peninsular Malaysia.</p><p>Remarks.</p><p>Some of the paratypes have more or less protruding and detached last whorl. Also, variability is observed in the keel near the aperture, it can be more or less sharp / pronounced.</p></div>	https://treatment.plazi.org/id/956C9AEC0FC25B8AABB2932BFBB74568	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
988ED4596B5B5B2588653E486C6D3832.text	988ED4596B5B5B2588653E486C6D3832.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma erawan Panha & J. B. Burch 2003	<div><p>Hypselostoma erawan Panha &amp; J. B. Burch, 2003</p><p>Figs 111 S, 135, 154</p><p>Hypselostoma erawan Panha &amp; Burch in Burch et al. 2003: 169–174, figs 14, 15.</p><p>Hypselostoma erawan — Tongkerd et al. 2004: 145, fig. 4 A – C; Panha and Burch 2008: 91–92, fig. 78; Dumrongrojwattana and Wongkamhaeng 2024: 324, fig. 8.</p><p>Type material examined.</p><p>Thailand • 3 paratypes; from the type locality; SMF 331455 .</p><p>Type locality.</p><p>“ A limestone mountain in the southern part of Erawan National Park, Triyoke District, Karnchanaburi Province, 14°12'15"N, 99°7'58"E, 70 meters elevation, Thailand ”.</p><p>Differential diagnosis.</p><p>See under H. populare sp. nov.</p><p>Distribution.</p><p>This species is known only from the type locality.</p></div>	https://treatment.plazi.org/id/988ED4596B5B5B2588653E486C6D3832	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
7A5D2E449F7D5849B4C7E64BDBD5A197.text	7A5D2E449F7D5849B4C7E64BDBD5A197.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma everetti E. A. Smith 1896	<div><p>Hypselostoma everetti E. A. Smith, 1896</p><p>Figs 111 T, 136, 137, 140</p><p>Hypselostoma everetti Smith, 1896: 148, Plate 10, fig. 9–9 b.</p><p>Hypselostoma dohertyi Fulton, 1899: 215, pl. 11, fig. 17. syn. nov.</p><p>Gyliauchen everetti — Pilsbry 1917: 218–219, pl. 37, figs 4, 6, 10.</p><p>Gyliauchen dohertyi — Pilsbry 1917: 219–220, pl. 37, figs 7–9.</p><p>Gyliotrachela everetti — van Benthem Jutting 1950: 45; Vermeulen and Whitten 1998: 80–81, fig. 64.</p><p>Gyliotrachela dohertyi — van Benthem Jutting 1950: 45; Zilch 1984: 165.</p><p>Type material examined.</p><p>Indonesia • syntype of H. everetti; NHMUK 1896.5. 17.1–6; syntype of H. dohertyi; SMF 4586 / 1.</p><p>Additional material examined.</p><p>Indonesia • 2 shells; Kalao island, Celebes; SMF 227446 • 2 shells; Kalao island, T. H. Aldrich coll.; UF 00112280 . • 1 shell; Timor island, 1 km W. Baucau; 18. Aug. 1971; F. G. Thompson leg.; locality code FGT-1729; UF 210107 .</p><p>Type localities.</p><p>“ Kalao island ”, Indonesia ( H. everetti); “ Tenimber I ”, (= Tanimbar islands, Indonesia) ( H. dohertyi).</p><p>Differential diagnosis.</p><p>This species differs from other congeners by the presence of strong keel (almost like a shoulder) positioned slightly above the centre of the periphery and a very wide umbilicus.</p><p>Distribution.</p><p>This species is known from Timor island, Kalao island, Sulawesi, Java, Bali, Lesser Sunda islands and Tanimbar islands (latter is the type locality of H. dohertyi).</p><p>Remarks.</p><p>During examination of the type material, we found no clear differences between H. everetti and H. dohertyi . The differences mentioned by Fulton (1899) in the original description include the narrower shell, less detached aperture, “ not perspectively umbilicated ” shell and that the shell is “ more closely coiled below ”. The umbilicus appears very slightly narrower in H. dohertyi but this falls within the intraspecific variability. Other characters mentioned are also very similar between the two species and also fall within the intraspecific variability, which is especially true for the level of the detachment of the last whorl.</p></div>	https://treatment.plazi.org/id/7A5D2E449F7D5849B4C7E64BDBD5A197	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
F491D1DE91CC5D1A9CD30E7865390DA9.text	F491D1DE91CC5D1A9CD30E7865390DA9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma fortunatum Gojsina, Hunyadi & Pall-Gergely 2025	<div><p>Hypselostoma fortunatum Gojšina, Hunyadi &amp; Páll-Gergely sp. nov.</p><p>Figs 186 E, 193, 194, 195, 223</p><p>Type material.</p><p>Holotype. Thailand • 1 shell (SH: 1.57 mm; SW 1: 2.83 mm); Phattalung Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.08884&amp;materialsCitation.latitude=7.6251" title="Search Plazi for locations around (long 100.08884/lat 7.6251)">Khao Ok Thalu</a>, rock wall; 07°37.506'N 100°05.330'E; 19 Feb. 2015; A. Hunyadi leg.; CUMZ 14465 . Paratypes. Thailand • 20 shells; same data as for holotype; coll. HA .</p><p>Additional material examined.</p><p>Thailand • 6 shells (juveniles / damaged, not paratypes); same data as for holotype; coll. HA • 11 shells; Songkhla Province, 31.3 km NW <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.26667&amp;materialsCitation.latitude=7.1666665" title="Search Plazi for locations around (long 100.26667/lat 7.1666665)">Hat Yai</a>, 1.2 km W of Hwy. 43; 7°10'N, 100°16'E; 80 m a. s. l.; 08 Apr. 1988; K. Auffenberg leg.; locality code KA-0636 A; UF 344932 • 1 shell; same locality data as previous; 08 Apr. 1988; K. Auffenberg leg.; locality code KA-0635; UF 591324 .</p><p>Type locality.</p><p>Thailand, Phattalung Province, Khao Ok Thalu, rock wall; 07°37.506'N, 100°05.330'E.</p><p>Diagnosis.</p><p>Hypselostoma with depressed-conical shell, teleoconch with finely sandpaper-like surface and radially but not spirally striated. Aperture equipped with only four relatively weak barriers (angulo-parietal, upper palatal, lower palatal and columellar).</p><p>Description.</p><p>Shell depressed-conical, chestnut brown, glossy, not translucent. It is consisting of 3.5–4 regularly increasing, convex whorls separated by a deep suture. Protoconch of the same colour as the teleoconch, finely pitted, not showing spiralling pattern and consisting of ~ 1.25 whorls. Teleoconch sculpture finely granulated (sandpaper-like) with additional radial growth lines which are more or less regularly spaced. Last whorl bluntly keeled at the centre of the periphery, slightly to moderately detached from the penultimate whorl and strongly ascending near the aperture (usually ranging from 65 to almost 90 ° compared to the shell axis). The keel is the most prominent on the last whorl behind the peristome. Peristome brown, weakly expanded, not reflected. Aperture equipped with four main barriers (angulo-parietal, upper palatal, lower palatal and columellar), all of which are relatively weak. Angulo-parietal lamella is the strongest in the aperture, consisting of a high parietal part and weak, tubercle-like angular part which is sometimes weakly sinuated in the middle. The angular part is sometimes also more distinctly separated from the parietal. Upper palatal plica relatively weak and straight. Lower palatal plica only slightly stronger (broader and higher) than the upper palatal. Columellar lamella horizontal, developed to the same extent as the lower palatal plica. Weak basal plica and infraparietal lamella sometimes present. All barriers weaker than in the majority of the congeners, all finely spiniferous medially but indistinctly granulated to almost smoot laterally. Sinulus wide and not strongly separated from the rest of the aperture. Umbilicus moderately wide to wide, measuring ~ 1 / 3–1 / 5 of the shell width and clearly showing the penultimate whorl. A relatively shallow groove runs towards the inner side of the umbilicus, terminating at the penultimate whorl.</p><p>Differential diagnosis.</p><p>See under H. pattalungense and H. depressispira .</p><p>Measurements</p><p>(in mm, n = 5). SH = 1.46–1.58; SW 1 = 2.67–2.90; SW 2 = 1.74–1.92; AH = 1.06–1.26; AW = 1.05–1.23.</p><p>Etymology.</p><p>The shape of the aperture, between the barriers, resembles a four-leaf clover which is a common symbol of luck, hence the specific epithet which is derived from the Latin word for lucky (fortunatus).</p><p>Distribution.</p><p>This species is known from Phattalung and Songkhla provinces (S Thailand).</p><p>Remarks.</p><p>This species is present in one more distinct form superficially resembling H. depressispira . Namely, in this form the spire is much more depressed and the umbilicus is noticeably wider. We decided not to treat this form a separate species since other characters (apertural dentition, general shell shape, and its surface sculpture) are fully shared with the “ typical form ” of H. fortunatum sp. nov.</p></div>	https://treatment.plazi.org/id/F491D1DE91CC5D1A9CD30E7865390DA9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
7ABF7A5A9DF350338C9846886C710393.text	7ABF7A5A9DF350338C9846886C710393.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma frequens (van Benthem Jutting 1950) Gojšina & Hunyadi & Sutcharit & Tongkerd & Auffenberg & Grego & Vermeulen & Reischütz & Páll-Gergely 2025	<div><p>Hypselostoma frequens (van Benthem Jutting, 1950) comb. nov.</p><p>Figs 224 C, 227, 241</p><p>Paraboysidia frequens van Benthem Jutting, 1950: 17, fig. 7.</p><p>Paraboysidia frequens — van Benthem Jutting 1950: 38; van Benthem Jutting 1960: 14; van Benthem Jutting 1961: 35; Berry 1963: pl. 5, fig. 40; Zilch 1984: 165; Maassen 2001: 73–74; Foon and Marzuki 2023: 142, fig. 1 AQ.</p><p>Paraboysidia oreia van Benthem Jutting, 1961: 37, pl. 9, fig. 3. syn. nov.</p><p>Gyliotrachela frequens — Schilthuizen et al. 1999: 284, fig. 2.</p><p>Paraboysidia oreia — Maassen 2001: 75; Foon et al. 2017: 79–80, fig. 30 D.</p><p>Type localities.</p><p>“ Kota Tongkat, Pahang ”, Malaysia ( H. frequens); “ Gunong Batu Kurau, Perak ”, Malaysia ( P. oreia).</p><p>Type material examined.</p><p>Malaysia • 48 paratypes of Paraboysidia frequens; from the type locality; 1947; Raffles Museum, Singapore ex. coll.; RMNH.Moll.137114 • 23 paratypes of Paraboysidia oreia; from the type locality; RMNH.Moll.137132 .</p><p>Additional material examined.</p><p>Malaysia • 1 shell; Perak, Ipoh, Perak Tong Temple; Oct. 1998; Hemmen leg.; coll. PGB • 5 shells; Selangor, Batu Caves (N of Kuala Lumpur); Nov. 1998; Hemmen leg.; coll. PGB • 1 shell; Selangor, Templer Park, Bukit Takun (NNW of Kuala Lumpur); Nov. 1998; Hemmen leg.; coll. PGB • 4 shells; Kedah, Gunung Keriang (N of Alor Setar); Sept. 1999; Hemmen leg.; coll. PGB .</p><p>Differential diagnosis.</p><p>See under H. kelantanense, H. modestum, H. muaklekense and H. serpa . This species also similar to the distant Bensonella perfecta sp. nov. from which it differs by less numerous barriers on the palatal side, the absence of the palatal tubercle on the peristome edge, non-spiniferous apertural barrier surface, less conical shell, and very slightly narrower umbilicus.</p><p>Distribution.</p><p>This species is known from Pahang, Kedah, Perak, and Selangor provinces of Peninsular Malaysia. Possibly more widely spread.</p><p>Remarks.</p><p>In the original description of Paraboysidia oreia, van Benthem-Jutting (1961) mentioned that it differs from H. frequens by the less numerous barriers, less inflated and more rounded last whorl. During our examination of the type material, we found no differences between the two species: the last whorl was not more inflated nor more rounded. The apertural barriers were sometimes more numerous in H. frequens but this is only true for smaller and more variable barriers which are of none or very limited taxonomic importance. Therefore, we treat P. oreia as a junior synonym of H. frequens .</p></div>	https://treatment.plazi.org/id/7ABF7A5A9DF350338C9846886C710393	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
CB7BF46F98FE5A1089D4556B076B7402.text	CB7BF46F98FE5A1089D4556B076B7402.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma fruhstorferi Mollendorff 1897	<div><p>Hypselostoma fruhstorferi Möllendorff, 1897</p><p>Figs 111 U, 138, 139, 140</p><p>Hypselostoma fruhstorferi Möllendorff, 1897: 70–71.</p><p>Gyliauchen fruhstorferi — Pilsbry 1917: 217, pl. 37, figs 1–3; van Benthem Jutting 1950: 46.</p><p>Gyliotrachela concreta van Benthem Jutting, 1949 a: 64–65, fig. 1. syn. nov.</p><p>Gyliotrachela concreta — van Benthem Jutting 1950: 44.</p><p>Gyliotrachela fruhstorferi — van Benthem Jutting 1952: 318; Zilch 1984: 165.</p><p>Type material examined.</p><p>Indonesia • lectotype of H. fruhstorferi; SMF 4580 • 5 paralectotypes of H. fruhstorferi; SMF 4581 • 16 paratypes of G. concreta; from the type locality; Sept. 1948; G. A. Tammes-Bolt leg.; RMNH. Moll. 137155 .</p><p>Additional material examined.</p><p>Indonesia • 5 shells; Sulawesi, South-Sulawesi, Lemo SE of Rantepao; 700 m a. s. l.; May 1995; W. J. M. Maassen leg.; coll. PGB • 2 shells; Lesser Sunda islands, Rintja island, ca 1 mile S Rintja village, mouth of cave; Apr. 1994; W. Auffenberg leg.; UF 220736 .</p><p>Type localities.</p><p>“ Java ” (Djampang, Java, Indonesia) ( H. fruhstorferi); “ Sulawesi, Main road from Makalé to Kalossi, South Celebes, 700–800 m alt. ” ( G. concreta).</p><p>Differential diagnosis.</p><p>This species is probably most similar to the geographically distant H. khaowongense and for differences, see under that species.</p><p>Distribution.</p><p>This species in known from Lesser Sunda islands, Java and Sulawesi islands, Indonesia.</p><p>Remarks.</p><p>Slightly variable smaller barriers. Some specimens of G. concreta had two and some three barriers between the lower palatal and columellar one. Interpalatal plica can be well developed or weaker.</p><p>We observed no differences between the type specimens of G. concreta and H. fruhstorferi . Van Benthem Jutting (1949 a) compared the newly described G. concreta with H. fruhstorferi but her decision was based on variable characters such as the level of detachment of the last whorl, broadness of the last whorl (which is, by us, not observed as different between the types) and level of keel development (namely slightly weaker in G. concreta, but this difference is very slight). We treat all these differences as intraspecific variability.</p></div>	https://treatment.plazi.org/id/CB7BF46F98FE5A1089D4556B076B7402	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
F3E4D81E73B55F01BAD8C70631BE810A.text	F3E4D81E73B55F01BAD8C70631BE810A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma fungus Gojsina, Hunyadi & Pall-Gergely 2025	<div><p>Hypselostoma fungus Gojšina, Hunyadi &amp; Páll-Gergely sp. nov.</p><p>Figs 111 V, 141, 142, 177</p><p>Material examined.</p><p>Holotype. Cambodia • 1 shell (SH 2: 2.94 mm; SW: 3.09 mm); Steung Treng Province, 36.3 km northwest and 5 km north from Stung Treng Mekong Bridge, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.74796&amp;materialsCitation.latitude=13.7762165" title="Search Plazi for locations around (long 105.74796/lat 13.7762165)">Phnom Chhnok</a>; 13°46.573'N, 105°44.878'E; 120 m a. s. l.; 25 Oct. 2023, A. Hunyadi &amp; J. U. Otani leg.; CUMZ 14453.</p><p>Paratypes. Cambodia • 219 shells; same data as for holotype; coll. HA • 1 shell; same data as for holotype; coll. VG .</p><p>Additional material examined.</p><p>Cambodia • 3 shells (broken, not paratypes); same data as for holotype; coll. HA .</p><p>Type locality.</p><p>Steung Treng Province, 36.3 km northwest and 5 km north from Stung Treng Mekong Bridge, Phnom Chhnok; 13°46.573'N, 105°44.878'E; 120 m a. s. l.</p><p>Diagnosis.</p><p>Hypselostoma species with a large last whorl (previous whorls sunken into it) and spirally striated teleoconch. Last whorl strongly detached from the penultimate and descending at almost 80 ° compared to the shell axis. Aperture equipped with four main barriers (angulo-parietal, upper palatal, lower palatal and columellar) and some smaller ones. Basal furrow present. Umbilicus extremely wide, almost as wide as the whole shell.</p><p>Description.</p><p>Shell depressed-conical, brown, and not glossy, consisting of 3.5–4 whorls separated by a deep suture. Protoconch rounded, roughly pitted and consisting of one whorl. Initial teleoconch whorls bluntly keeled and sunken into the last whorl (but sometimes they appear more elevated). Last whorl contributing to almost 100 % of the shell height. It is extending obliquely around the previous whorls and is strongly shouldered both on its upper and lower sides, which gives the shell a unique appearance. Surface of the teleoconch is irregularly, very finely and densely spirally striated. Whitish radial streaks are numerous and present on all teleoconch whorls but unevenly spaced (rather randomly positioned). Last whorl is strongly detached from the penultimate and strongly descending (~ 80 ° compared to the shell axis). This makes the aperture profile almost perpendicular to the shell axis. The so called “ trumpet ” is quite broad (similar to e. g., H. chaunosalpinx). Peristome is dirty white (or light brown), thick and strongly expanded, not reflected. Aperture is equipped with four main barriers (angulo-parietal, upper palatal, lower palatal and columellar) and additional smaller ones. The parietal part of the angulo-parietal lamella is strong and very high (blade-like), but the angular is very short and pointed towards the palatal side. Upper palatal plica is directed towards the lower palatal plica but is weaker. Lower palatal plica is long and low in its inner part and much higher in its outer part (closer to the peristome). This plica is similar to the columellar lamella but more slender. Basal (?) plica is situated closer to the lower palatal. It is not clear whether this plica actually represents the infrapalatal or the basal. In the region where usually the basal plica is present, there is a deep basal furrow which is also rarely encountered in this group (similar structure is also present in H. torta sp. nov.). This furrow is stretching all the way to the expanding peristome. Columellar lamella almost horizontal, thick, and long. A distinct infraparietal lamella is present (sometimes absent) between the columellar and parietal lamellae. Of all these barriers, only the angulo-parietal and columellar lamellae are reaching the expanding peristome. Surface of all apertural barriers is finely granulated. Sinulus narrow, well separated from the rest of the aperture. Umbilicus extremely wide (almost as wide as the whole shell). The lower edge of the last whorl (towards the umbilicus) is very strongly ridged (almost forming a cliff-like structure), leaving the walls of the umbilicus very steep (like a playground slide). The walls of the umbilicus are especially densely sculptured with white radial streaks.</p><p>Differential diagnosis.</p><p>See under H. srakeoense, H. pendulum, H. torticollis, and H. transitans .</p><p>Measurements</p><p>(in mm, n = 5). SH 1 = 2.83–2.97; SH 2 = 1.62–1.85; SW = 3.09–3.85; AH = 1.82–2; AW = 1.77–2.05.</p><p>Etymology.</p><p>This species’ shape resembles a mushroom, which gave its specific epithet (Lat. fungus). The detached last whorl resembles a stalk while the rest of the shell resembles a cap. To be used as a noun in apposition.</p><p>Distribution.</p><p>This species is known only from its type locality.</p><p>Remarks.</p><p>The largest, and still unexplored limestone hill in Steung Treng Province (13°47.941'N, 105°43.7671'E) was inaccessible during the collecting efforts in 2023 due to the works of a cement factory. Due to the proximity of this hill to the type locality of H. fungus sp. nov., it is possible that this species can be found here as well and be threatened by quarrying.</p></div>	https://treatment.plazi.org/id/F3E4D81E73B55F01BAD8C70631BE810A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
FD90B3CA36CF5905B48509C7BCA2100D.text	FD90B3CA36CF5905B48509C7BCA2100D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma geckophilum Gojsina, Hunyadi & Pall-Gergely 2025	<div><p>Hypselostoma geckophilum Gojšina, Hunyadi &amp; Páll-Gergely sp. nov.</p><p>Figs 111 W, 143, 144, 154</p><p>Type material.</p><p>Holotype. Thailand • 1 shell (SH: 2.92 mm; SW 1: 2.74 mm); Chanthaburi Province, Kaeng Hang Maeo district, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.81775&amp;materialsCitation.latitude=12.887267" title="Search Plazi for locations around (long 101.81775/lat 12.887267)">Tham Khao Wongkot</a>; 12°53.236'N, 101°49.065'E; 60 m a. s. l.; 08 Mar. 2023; A. Hunyadi leg.; CUMZ 14451 . Paratypes. Thailand • 105 shells; same data as for holotype; coll. HA • 1 shell; same data as for holotype; coll. VG .</p><p>Additional material examined.</p><p>Thailand • 14 shells (juveniles / damaged, not paratypes); same data as for holotype; coll. HA .</p><p>Type locality.</p><p>Thailand, Chanthaburi Province, Kaeng Hang Maeo district, Tham Khao Wongkot; 12°53.236'N, 101°49.065'E; 60 m a. s. l.</p><p>Diagnosis.</p><p>Shell concave-conical, last whorl shouldered, previous whorls weakly convex. Teleoconch with raised spiral striae and strong whitish streaks. Last whorl detached from the penultimate, slightly turned downwards. Aperture equipped with five barriers (angulo-parietal, upper palatal, lower palatal, basal, and columellar). Umbilicus initially narrow but suddenly expanding at the last whorl.</p><p>Description.</p><p>Shell concave-conical (due to the strongly expanded last whorl), light brown, weakly glossy, opaque. It is consisting of 4.5–5 step-like, regularly increasing whorls separated by a very deep suture. All whorls except for the last one are rounded, convex. Protoconch slightly lighter than the rest of the shell, spirally striated (~ 25 densely arranged spiral striae) and consisting of ~ 1.25 whorls. Boundary between protoconch and teleoconch not particularly clear but visible as stronger spiral striation and darker surface. Teleoconch densely and strongly spirally striated, especially on the last whorl. Spiral striae slightly almost regularly spaced, leaving the space between the two striae up to the width of three or rarely two striae. Spiral striae get more densely arranged at the lower part of the last whorl than those at the centre of the periphery. There are ~ 33 spiral striae on the last whorl in standard view. Strong and wide radial white streaks cross the spiral striae. These streaks vary in number and are mostly present on the last three whorls. Last whorl shouldered, slightly detached from the penultimate and slightly descending near the aperture (~ 5–10 ° compared to the shell axis). Peristome of the same colour as the rest of the shell, or very slightly lighter. It is expanded and not reflected. Aperture equipped with four main barriers (angulo-parietal, upper palatal, lower palatal and columellar). Angular part of the angulo-parietal lamella reaching the peristome, very small and pointed, leaned towards the upper palatal plica. Parietal part of the angulo-parietal lamella is much stronger and high, blade shaped. Upper palatal plica moderately developed, slightly curved towards the lower palatal and similarly strong. Lower palatal plica high and narrow, making it particularly slender. Columellar lamella strong as the palatal plicae, almost horizontal and very slightly leaned towards the lower palatal plica. Between these main barriers, only a small and weak basal one is additionally observed. Surface of all apertural barriers is smooth. Sinulus wide, not strongly separated from the rest of the aperture. Umbilicus initially very narrow, suddenly expanded at the last whorl, measuring between 1 / 3 and ¼ of the shell width. A deep groove is running from the columellar side of the peristome towards the inner walls of the umbilicus.</p><p>Differential diagnosis.</p><p>See under H. platybasis sp. nov.</p><p>Measurements</p><p>(in mm, n = 5). SW 1 = 2.48–2.74; SW 2 = 1.68–1.75; SH = 2.62–2.92; AH = 1.09–1.21; AW = 1.12–1.25.</p><p>Etymology.</p><p>The specific epithet is due to the fact that there were many geckos at its habitat.</p><p>Distribution.</p><p>This species is known only from the type locality.</p><p>Remarks.</p><p>The concrescent angulo-parietal lamella can have the appearance of the typical form (with strong parietal and weaker, pointed angular part) or can have the appearance of only a single, parietal lamella with a very slightly visible swelling which represents the angular part. Infrapalatal lamella can be well visible or very weakly developed.</p></div>	https://treatment.plazi.org/id/FD90B3CA36CF5905B48509C7BCA2100D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
BBEC67F6B35455BFB0DBB1AD4A6CD73C.text	BBEC67F6B35455BFB0DBB1AD4A6CD73C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma holimanae F. G. Thompson & H. G. Lee 1988	<div><p>Hypselostoma holimanae F. G. Thompson &amp; H. G. Lee, 1988</p><p>Figs 111 X, 145, 154</p><p>Hypselostoma holimanae Thompson &amp; Lee, 1988: 78, figs 1–6.</p><p>Hypselostoma holimanae — Thompson and Upatham 1997: 223; Hemmen and Hemmen 2001: 41; Tongkerd et al. 2004: 143, fig. 2; Panha and Burch 2008: 92–93, fig. 79; Dumrongrojwattana and Wongkamhaeng 2024: 324, fig. 8.</p><p>Type material examined.</p><p>Thailand • holotype; UF 00113427 • 14 paratypes; from the type locality; UF 00113483 / 1; UF 00113428 / 13 .</p><p>Additional material examined.</p><p>Thailand • 10 shells; Kanchanaburi province, 15 km W <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.291&amp;materialsCitation.latitude=14.154" title="Search Plazi for locations around (long 99.291/lat 14.154)">Kanchanaburi</a>; 14.154°N, 99.291°E; 15 Mar. 1987; S. Holiman leg.; UF 540579 .</p><p>Type locality.</p><p>“ Thailand, Kanchanaburi province, small limestone range on the west border of Kanchanaburi Agricultural College, about 15 km west of Kanchanaburi ”.</p><p>Differential diagnosis.</p><p>See under H. khaowongense .</p><p>Distribution.</p><p>This species is known only from the type locality.</p></div>	https://treatment.plazi.org/id/BBEC67F6B35455BFB0DBB1AD4A6CD73C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
FEDD6A30FC4F5729BA22BE5005C13FA0.text	FEDD6A30FC4F5729BA22BE5005C13FA0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma hungerfordianum Moellendorff 1891	<div><p>Hypselostoma hungerfordianum Möllendorff, 1891</p><p>Figs 186 F, 196, 197, 198, 199, 200, 201</p><p>Hypselostoma hungerfordianum Möllendorff, 1891: 337, pl. 30, figs 7, 7 a.</p><p>Hypselostoma striolatum Möllendorff, 1894: 152. syn. nov.</p><p>Hypselostoma hungerfordianum — Sykes 1902: 61.</p><p>Hypselostoma striolatum — Fischer and Dautzenberg 1904: 408.</p><p>Gyliauchen hungerfordianus — Pilsbry 1917: 212, pl. 36, figs 1–4.</p><p>Gyliauchen striolatus — Pilsbry 1917: 215.</p><p>Gliotrachela hungerfordiana [sic] — Laidlaw 1933: 214.</p><p>Gliotrachela striolatus [sic] — Laidlaw 1933: 214.</p><p>Gyliotrachela hungerfordiana — van Benthem Jutting 1949 b: 60; van Benthem Jutting 1950: 46; van Benthem Jutting 1950: 26; Zilch 1959: 164, fig. 563; van Benthem Jutting 1960: 14; Zilch 1984: 166; Davison 1995: 239; Schileyko 1998: 142, fig. 162; Maassen 1999: 122–123; Schilthuizen 1999: 283; Maassen 2001: 75–76; Foon et al. 2017: 79, fig. 30 B; Hoekstra and Schilthuizen 2011: 267–272, fig. 2; Foon and Marzuki 2023: 142, fig. 1 AP; Tongkerd et al. 2024: 180–182, figs 10, 13 M.</p><p>Gyliotrachela striolata — van Benthem Jutting 1950: 46; Zilch 1984: 166; Thompson and Upatham 1997: 233; Hemmen and Hemmen 2001: 41.</p><p>Gyliotrachela khaochongensis Panha in Panha 1998 a: 53 –56, fig. 2.</p><p>Gyliotrachela khaochongensis — Hemmen and Hemmen 2001: 41; Panha and Burch 2008: 70–71, fig. 61. Tongkerd et al. 2024: 180, fig. 10 B.</p><p>Hypselostoma satulensis Panha &amp; Burch in Panha et al. 2004: 75–76, fig. 12. syn. nov.</p><p>Hypselostoma satulensis — Panha and Burch 2008: 98–99, fig. 84; Dumrongrojwattana and Wongkamhaeng 2024: 324, fig. 8.</p><p>Gyliotrachela striolatus [sic] — Panha and Burch 2008: 81–82, fig. 71.</p><p>Gyliotrachela phoca Tongkerd &amp; Panha, 2013 in Tongkerd et al. 2013: 71–75, figs 5–7.</p><p>Gyliotrachela phoca — Tongkerd et al. 2024: 180. Dumrongrojwattana and Wongkamhaeng 2024: 324, fig. 8.</p><p>Type material examined.</p><p>Malaysia • lectotype of H. hungerfordianum; SMF 4587. Thailand • holotype of H. striolatum; SMF 4591 • holotype of H. satulensis; S. Panha leg.; CUMZ ver. 44028; paratype of G. khaochongensis; from the type locality; CUMZ ver. 010.</p><p>Additional material examined.</p><p>Thailand-South • 6 shells; Suratthani Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.398&amp;materialsCitation.latitude=8.66" title="Search Plazi for locations around (long 99.398/lat 8.66)">6 km S of Na San</a>, limestone mountain; 8.660°N, 99.398°E; 03 June 1987; F. Thompson leg.; locality code FGT-4311; UF 530647 • 2 shells; Suratthani Province, Hwy. 401, 2.7 km W junction <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.666664&amp;materialsCitation.latitude=9.166667" title="Search Plazi for locations around (long 99.666664/lat 9.166667)">Hwy</a>. 4142 and 401, evergreen forest on rocky hillside; 9°10'N, 99°40'E; 90 m a. s. l.; 19 Apr. 1988; K. Auffenberg leg.; locality code KA-0674; UF 591325 • 34 shells; Suratthani Province, 15.3 km NE Ban Na San off road to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.433334&amp;materialsCitation.latitude=8.883333" title="Search Plazi for locations around (long 99.433334/lat 8.883333)">Tha Rom Yen National Park</a>; 8°53'N, 99°26'E; 40 m a. s. l.; 20 Apr. 1988; K. Auffenberg leg.; locality code KA-0678; UF 345386 • 12 shells; Suratthani Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.382&amp;materialsCitation.latitude=8.812" title="Search Plazi for locations around (long 99.382/lat 8.812)">1 km NE of Na San</a>, 100 m limestone mountain; 8.812°N, 99.382°E; 02 June 1987; F. G. Thompson leg.; locality code FGT-4309; UF 547905 • 3 shells; Suratthani Province, Angthong islands, Wia Talap island, steep path to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.92443&amp;materialsCitation.latitude=8.126717" title="Search Plazi for locations around (long 98.92443/lat 8.126717)">Buaboke cave</a>, at the base of limestone rocks; 8°7.603'N, 98°55.466'E; 50 m a. s. l.; Sep. 2007; A. Reischütz leg.; coll. REI • 1 shell; Suratthani Province, 15.3 km NE Ban Na San off road to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.433334&amp;materialsCitation.latitude=8.883333" title="Search Plazi for locations around (long 99.433334/lat 8.883333)">Tha Rom Yen National Park</a>; 8°53'N, 99°26'E; 40 m a. s. l.; 20 Apr. 1988; K. Auffenberg leg.; locality code KA-0678; UF 591326 • 3 shells; Phangnga Province, 32 km NE Khok Kloi, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.45&amp;materialsCitation.latitude=8.45" title="Search Plazi for locations around (long 98.45/lat 8.45)">Wat Tham Suwan Khuha</a>; 08°27'N, 98°27'E; 20 m a. s. l.; 27 Mar. 1988; K. Auffenberg leg.; locality code KA-0601; UF 345751 • 1 shell; Phangnga Province, limestone knoll, 5.7 km SW intersection <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.48333&amp;materialsCitation.latitude=8.45" title="Search Plazi for locations around (long 98.48333/lat 8.45)">Hwys</a> 4 &amp; 4152; 8°27'N, 98°29'E; 20 m a. s. l.; 28 Mar. 1988; K. Auffenberg leg.; locality code KA-0602; UF 345771 • 1 shell; Krabi Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.6828&amp;materialsCitation.latitude=8.3462" title="Search Plazi for locations around (long 98.6828/lat 8.3462)">5.6 km SW of Ao Luek Tai</a>, on uneven surface of large boulder at the base of limestone cliff; 8°20.772'N, 98°40.968'E; 20–25 m a. s. l.; 16 Mar. 2020; ex. coll. S. Aiken; coll. HA • 8 shells; Krabi Province, 9.2 km northwest from <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.870514&amp;materialsCitation.latitude=8.12175" title="Search Plazi for locations around (long 98.870514/lat 8.12175)">Phang Nga</a>; 08°07.305'N, 98°52.231'E; 40 m a. s. l.; 21 Feb. 2015; A. Hunyadi leg.; coll. HA • 3 shells; Krabi Province, Viewpoint Hill, Railay (= <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.83746&amp;materialsCitation.latitude=8.008516" title="Search Plazi for locations around (long 98.83746/lat 8.008516)">Rai Leh</a>) <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.83746&amp;materialsCitation.latitude=8.008516" title="Search Plazi for locations around (long 98.83746/lat 8.008516)">Beach West</a>, at the base of limestone rocks; 8°0.511'N, 98°50.248'E; 40 m a. s. l.; Sep. 2007; A. Reischütz leg.; coll. REI • 4 shells; Nakhon Si Thammarat Province, Lan Saka, N side of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.75&amp;materialsCitation.latitude=8.416667" title="Search Plazi for locations around (long 99.75/lat 8.416667)">Hwy</a>. 4015; 08°25'N, 99°45'E; 16 Apr. 1988; K. Auffenberg leg.; locality code KA-0665; UF 345226 • 8 shells; Nakhon Si Thammarat Province, circa 38 km WNW of Tha Sala, outcrop 1.5 km W of junction <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.78333&amp;materialsCitation.latitude=8.716666" title="Search Plazi for locations around (long 99.78333/lat 8.716666)">Hwys</a> 4186 and 4188; 8°43'N, 99°47'E; 80 m a. s. l.; 16 Apr. 1988; K. Auffenberg leg.; locality code KA-0661; UF 345194 • 16 shells; Nakhon Si Thammarat Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.379364&amp;materialsCitation.latitude=8.4213" title="Search Plazi for locations around (long 99.379364/lat 8.4213)">Wat Tham Thong Panara</a>, vicinity of the rock temple; 08°25.278'N, 99°22.762'E; 19 Feb. 2015; A. Hunyadi leg.; coll. HA • 23 shells; Nakhon Si Thammarat Province, circa 38 km WNW of Tha Sala, outcrop 1.5 km W of junction <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.78333&amp;materialsCitation.latitude=8.716666" title="Search Plazi for locations around (long 99.78333/lat 8.716666)">Hwys</a> 4186 and 4188; 08°43'N, 99°47'E; 80 m a. s. l.; 16 Apr. 1998; K. Auffenberg leg.; locality code KA-0661; UF 345193 • 20 shells; Phattalung Province, 6 km NW of Phattalung, 1 km NE of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.05&amp;materialsCitation.latitude=7.6666665" title="Search Plazi for locations around (long 100.05/lat 7.6666665)">Hwy.</a> 4048; 7°40'N, 100°3'E; 60 m a. s. l.; 13 Apr. 1988; K. Auffenberg leg.; locality code KA-0653; UF 345102 • 54 shells; Phatthalung Province, Khao Pu-Khao Ya Nat. Park, Si Banphot district, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.83604&amp;materialsCitation.latitude=7.6966166" title="Search Plazi for locations around (long 99.83604/lat 7.6966166)">Ruesi Cave</a>; 7°41.797'N, 99°50.162'E; 25 Feb. 2023; A. Hunyadi, K. Okubo &amp; J. U. Otani leg.; coll. PGB • 25 shells; Phattalung Province, 6.4 km E of Hwy. 4 on Hwy. 4081, 1 km SW of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.183334&amp;materialsCitation.latitude=7.45" title="Search Plazi for locations around (long 100.183334/lat 7.45)">Khao Chai Son</a>; 7°27'N, 100°11'E; 50 m a. s. l.; 11 Apr. 1988; K. Auffenberg leg.; locality code KA-0648; UF 345063 • 14 shells; Phattalung Province, 6.4 km E of Hwy. 4 on Hwy. 4081, 1 km SW of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.183334&amp;materialsCitation.latitude=7.45" title="Search Plazi for locations around (long 100.183334/lat 7.45)">Khao Chai Son</a>; 7°27'N, 100°11E; 11 Apr. 1988; K. Auffenberg leg.; locality code KA-0648; UF 345064 • 7 shells; Phattalung Province, 16.2 km S of junction of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.98333&amp;materialsCitation.latitude=7.4166665" title="Search Plazi for locations around (long 99.98333/lat 7.4166665)">Hwy</a>. s 4 and 4122; 7°25'N, 99°59'E; 150 m a. s. l.; 09 Apr. 1988; K. Auffenberg leg.; locality code KA-0637; UF 344971 • 4 shells; Phattalung Province, 6 km NW of Phattalung, 1 km NE of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.05&amp;materialsCitation.latitude=7.667" title="Search Plazi for locations around (long 100.05/lat 7.667)">Hwy.</a> 404; 7°40.02'N, 100°3'E; 60 m a. s. l.; 13 Apr. 1988; K. Auffenberg leg.; locality code KA-0653; UF 436278 • 18 shells; Phattalung Province, 15.1 km W Hwy. 41 on Hwy. 4164, Khaopu and <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.933334&amp;materialsCitation.latitude=7.7833333" title="Search Plazi for locations around (long 99.933334/lat 7.7833333)">Khaoya National Park</a>; 7°47'N, 99°56'E; 160 m a. s. l.; 14 Apr. 1988; K. Auffenberg leg.; locality code KA-0657; UF 345154 • 6 shells; Phattalung Province, 6.4 km E Hwy. 4 on Hwy. 4081, 1 km SW <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.183&amp;materialsCitation.latitude=7.45" title="Search Plazi for locations around (long 100.183/lat 7.45)">Khao Chai Son</a>; 7°27'N, 100°10.98'E; 11 Apr. 1988; K. Auffenberg leg.; locality code KA-0649; UF 345073 • 17 shells; Phattalung Province, 16.2 km S of junction of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.98333&amp;materialsCitation.latitude=7.4166665" title="Search Plazi for locations around (long 99.98333/lat 7.4166665)">Hwys</a> 4 and 4122; 7°25'N, 99°59'E; 150 m a. s. l.; 09 Apr. 1988; K. Auffenberg leg.; locality code KA-0637; UF 344970 • 3 shells; Phattalung Province, Khao Pu Khao Ya National Park, vicinity of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.87238&amp;materialsCitation.latitude=7.666333" title="Search Plazi for locations around (long 99.87238/lat 7.666333)">Matcha cave</a>; 7°39.980'N, 99°52.343'E; 125 m a. s. l.; 25 Feb. 2023; A. Hunyadi leg.; coll. HA • 23 shells; Phattalung Province, Khao Pu Khao Ya National Park, Si Banphot district, ca 6 km north-northwest from <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.84703&amp;materialsCitation.latitude=7.6789665" title="Search Plazi for locations around (long 99.84703/lat 7.6789665)">Taphaen</a>; 7°40.738'N, 99°50.822'E; 85 m a. s. l.; 24 Feb. 2023; A. Hunyadi, K. Okubo &amp; J. U. Otani leg.; coll. HA • 1 shell; Yala Province, 6.3 km W of Yala, Ban Na Tham, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.25&amp;materialsCitation.latitude=6.55" title="Search Plazi for locations around (long 101.25/lat 6.55)">Wat Khuha Phi Muk</a>; 6°33'N, 101°15'E; 40 m a. s. l.; 01 Apr. 1988; K. Auffenberg leg.; locality code KA-0612; UF 344665 • 3 shells; Yala Province, 6.3 km W of Yala, Ban Na Tham, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.25&amp;materialsCitation.latitude=6.55" title="Search Plazi for locations around (long 101.25/lat 6.55)">Wat Khuha Phi Muk</a>; 6°33'N, 101°15'E; 40 m a. s. l.; 01 Apr. 1988; K. Auffenberg leg.; locality code KA-0612; UF 344664 • 25 shells; Yala Province, Mueang Yala district, Na Tham, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.23303&amp;materialsCitation.latitude=6.5219336" title="Search Plazi for locations around (long 101.23303/lat 6.5219336)">Silp Cave</a> (~ 9 km from centre of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.23303&amp;materialsCitation.latitude=6.5219336" title="Search Plazi for locations around (long 101.23303/lat 6.5219336)">Yala</a>); 6°31.316'N. 101°13.982'E; 21 Feb. 2023; A. Hunyadi, K. Okubo &amp; J. U. Otani leg.; coll. HA. Malaysia • 2 shells; Selangor, Templer Park, Bukit Takun; NNW of Kuala Lumpur; Nov. 1998; ex. coll. Hemmen; coll. PGB • 46 shells; Kedah, Baling, eastern side of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.91338&amp;materialsCitation.latitude=5.67625" title="Search Plazi for locations around (long 100.91338/lat 5.67625)">Bukit Baling</a>; 55 m a. s. l.; 05°40.575'N, 100°54.803'E; 11 Jan. 2013; A. Hunyadi leg.; coll. HA • 9 shells; Kelantan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.9686&amp;materialsCitation.latitude=4.8829" title="Search Plazi for locations around (long 101.9686/lat 4.8829)">Gua Musang</a>, environment of the cave entrance; 04°52.974'N, 101°58.116'E; 135 m a. s. l.; 17 Jan. 2013; A. Hunyadi leg.; coll. HA • 7 shells; Perlis, Kaki Bukit, vicinity of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.202835&amp;materialsCitation.latitude=6.6445665" title="Search Plazi for locations around (long 100.202835/lat 6.6445665)">Gua Kelam</a>; 06°38.674'N, 100°12.170'E; 45–55 m a. s. l.; 13 Jan. 2013; A. Hunyadi leg.; coll. HA • 11 shells; Pahang, 15 km west from Bandar Pusat Jengka, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.433&amp;materialsCitation.latitude=3.6977" title="Search Plazi for locations around (long 102.433/lat 3.6977)">Hutan Lipur Gunung Senyum</a>; 03°41.862'N, 102°25.980'E; 85 m a. s. l.; 23 Jan. 2013; A. Hunyadi leg.; coll. HA • 3 shells; Perak, 800 m north from Padang Rengas on the road no. A 107, eastern side of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.84155&amp;materialsCitation.latitude=4.7865834" title="Search Plazi for locations around (long 100.84155/lat 4.7865834)">Gunung Pondok</a>; 04°47.195'N, 100°50.493'E; 90 m a. s. l.; 08 Jan. 2013; A. Hunyadi leg.; coll. HA • 12 shells; Kelantan, Gua Musang, 6 km Pulai, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.9497&amp;materialsCitation.latitude=4.8368835" title="Search Plazi for locations around (long 101.9497/lat 4.8368835)">Gua Madu</a>, vicinity of the cave; 04°50.213'N, 101°56.982'E; 120 m a. s. l.; 17 Jan. 2013; A. Hunyadi leg.; coll. HA • 3 shells; Selangor, Kuala Lumpur E - Kanching, Templer Park, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.64063&amp;materialsCitation.latitude=3.2979667" title="Search Plazi for locations around (long 101.64063/lat 3.2979667)">Perangsang Templer Golf Club</a>, rock; 03°17.878'N, 101°38.438'E; 100 m a. s. l.; 25 Jan. 2013; A. Hunyadi leg.; coll. HA • 22 shells; Perak, Lenggong, Taman Awam Gua Tok Giring; 05°07.128'N, 100°.351'E; 90 m a. s. l.; 06 Jan. 2013; A. Hunyadi leg.; coll. HA • 131 shells; Perak, northeast from Chemor, 2 km east from <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.1299&amp;materialsCitation.latitude=4.7466497" title="Search Plazi for locations around (long 101.1299/lat 4.7466497)">Kampung Kanthan</a>, western side of the mountain; 04°44.799'N, 101°07.794'E; 105 m a. s. l.; 08 Jan. 2013; A. Hunyadi leg.; coll. HA • 304 shells; Pahang, Bukit Cinta Manis, south-southeastern side, 800 m from Lebuhraya Karak towards <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.013565&amp;materialsCitation.latitude=3.4452333" title="Search Plazi for locations around (long 102.013565/lat 3.4452333)">Kampung Cinta Manis</a>; 03°26.714'N, 102°00.814'E; 130 m a. s. l.; 22 Jan. 2013; A. Hunyadi leg.; coll. HA • 18 shells; Perlis, Kangar, Bukit Kaya / <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.19115&amp;materialsCitation.latitude=6.4346" title="Search Plazi for locations around (long 100.19115/lat 6.4346)">Bukit Lagi</a>, northeastern edge of the mountain; 06°26.076'N, 100°11.469'E; 5 m a. s. l.; 12 Jan. 2013; A. Hunyadi leg.; coll. HA • 67 shells; Pahang, Gua Bama, Kuala Lipis 9 km, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.9656&amp;materialsCitation.latitude=4.1942" title="Search Plazi for locations around (long 101.9656/lat 4.1942)">Padang Tungku</a>; 04°11.652'N, 101°57.936'E; 120 m a. s. l.; 19 Jan. 2013; A. Hunyadi leg.; coll. HA • 180 shells; Pahang, 20 km southeast from Jerantut, Gua Kota Gelanggi, below <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.47353&amp;materialsCitation.latitude=3.9" title="Search Plazi for locations around (long 102.47353/lat 3.9)">Gua Balai</a>; 03°54.000'N, 102°28.412'E; 115 m a. s. l.; 21 Jan. 2013; A. Hunyadi leg.; coll. HA • 4 shells; Kelantan Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.969635&amp;materialsCitation.latitude=4.8838835" title="Search Plazi for locations around (long 101.969635/lat 4.8838835)">Gua Musang</a>, behind railway station; 04°53.033'N, 101°58.178'E; J. U. Otani leg.; coll. PGB • 6 shells; Pahang Province, Lipis, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.18771&amp;materialsCitation.latitude=4.2158666" title="Search Plazi for locations around (long 102.18771/lat 4.2158666)">Kenong Rimba</a> park; 04°12.952'N, 102°11.263'E; 83 m a. s. l.; J. U. Otani leg.; coll. PGB • 2 shells; Pahang Province, Jerantut, Taman Negara, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.44247&amp;materialsCitation.latitude=4.510667" title="Search Plazi for locations around (long 102.44247/lat 4.510667)">Kuala Tahan</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.44247&amp;materialsCitation.latitude=4.510667" title="Search Plazi for locations around (long 102.44247/lat 4.510667)">Gua Luas</a>; 04°30.640'N, 102°26.548'E; 75 m a. s. l.; J. U. Otani leg.; coll. PGB • 1 shell; Pahang Province, Jerantut, Kg. Baheru, Gua Kota Gelanggi, Gua Terang Bulan; J. U. Otani leg.; coll. PGB • 8 shells; Kedah, 1.5 km east from Kodiang, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.31506&amp;materialsCitation.latitude=6.3905168" title="Search Plazi for locations around (long 100.31506/lat 6.3905168)">Gua Kerbau</a>; 06°23.431'N, 100°18.904'E, 14 Jan. 2013; A. Hunyadi leg.; coll. HA • 5 shells; Kelantan Province, Gua Musang, 264 km from KL, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.00159&amp;materialsCitation.latitude=4.7780833" title="Search Plazi for locations around (long 102.00159/lat 4.7780833)">Route</a> 8; 191 m a. s. l.; 04°46.685'N, 102°00.095'E; J. U. Otani leg.; coll. PGB • 5 shells; Selangor, Templer Park, Bukit Takun; NNW of Kuala Lumpur; Nov. 1999; J. Hemmen leg.; coll. PGB • 5 shells; Kelantan Province, Gua Madu, south of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.965416&amp;materialsCitation.latitude=4.873417" title="Search Plazi for locations around (long 101.965416/lat 4.873417)">Gua Musang</a>; 04°52.405'N, 101°57.925'E; 02 Nov. 2000; J. Hemmen leg.; coll. PGB • 8 shells; Pahang, Kota Gelanggi, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.47314&amp;materialsCitation.latitude=3.8912632" title="Search Plazi for locations around (long 102.47314/lat 3.8912632)">Jengka-Jerantut</a> road, on limestone surfaces covered with mosses; 03°53.4758'N, 102°28.388'E; 29. May 2011; M. E. Marzuki leg.; coll. PGB .</p><p>Type localities.</p><p>“ Perak ” (Bukit Pondong, Malaysia) ( H. hungerfordianum); “ Kuankalong Limestone Hill, Satul Province, 6°52'59"N, 100°07'38"E, 110 meters elevation ”, Thailand ( H. satulensis); “ Samui islands, Gulf of Siam ”, Thailand ( H. striolatum).</p><p>Differential diagnosis.</p><p>See under H. sichang and H. loei .</p><p>Distribution.</p><p>This species is widely spread in Malaysia and adjacent regions of Thailand. Also known from Myanmar where the northernmost locality of the species is known (Tongkerd et al. 2024). The southernmost localities are positioned in and near the city of Kuala Lumpur (Malaysia).</p><p>Remarks.</p><p>This species can have a more or less elongated spire. The umbilicus is usually consistent in its width but we have observed slight variations. The apertural dentition is variable in terms of the number of smaller barriers between the five main ones (angular, parietal, upper palatal, lower palatal and columellar). Gyliotrachela striolata was known only from a single damaged type, which retained its last whorl only. Even in damaged condition, it was clear that there are no differences between G. striolata and H. hungerfordianum which is why they are treated conspecific. Hypselostoma satulensis was described from Satun province in S Thailand and shows no morphological differences from H. hungerfordianum . Its type locality is practically located in the middle of the distribution area of H. hungerfordianum . We have examined the material from northern Thailand hosted in UF with the following data: 2 shells; Mae Hong Son Province, Tham Lod Cave, 1.5 km SE Ban Tham Lod Village; 19°34'N, 98°9'E; 720 m a. s. l.; 19 Mar. 1988; K. Auffenberg leg.; locality code KA-0576; UF 345527. Since this locality is quite far from the actual distribution area of this species, we believe that this sample was mislabelled and that this species actually does not occur in Mae Hong Son Province.</p></div>	https://treatment.plazi.org/id/FEDD6A30FC4F5729BA22BE5005C13FA0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
54099BE0415F52958A15FEBE8D8920BB.text	54099BE0415F52958A15FEBE8D8920BB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma insularum Pilsbry 1908	<div><p>Hypselostoma insularum Pilsbry, 1908</p><p>Figs 111 Y, 146, 147</p><p>Hypselostoma insularum Pilsbry, 1908: 41, fig. 2.</p><p>Hypselostoma insularum — Pilsbry 1917: 182–183, pl. 32, figs 1–4, 6; van Benthem Jutting 1950: 41.</p><p>Hypselostoma kentingensis Hwang, 2014: 30–32, fig. 2. syn. nov.</p><p>Type material examined.</p><p>Japan • lectotype of H. insularum; ANSP 95252 .</p><p>Type localities.</p><p>“ Yonakunijima, Ryuyku ”, Japan ( H. insularum); “ TAIWAN: Kenting Tropical Botanical Garden, Kenting, Hengchun, Pingdung, 21°58'04.5"N 120°48'55.3"E, alt. 295 m ” ( H. kentingensis).</p><p>Differential diagnosis.</p><p>This species is unique by its shell shape which resembles a pyramid (due to the sharp keel located at the base of the last whorl and moderately deep suture). All apertural barriers are strong but this is especially true for the angulo-parietal lamella.</p><p>Distribution.</p><p>This species is known from the type locality and Kenting Tropical Botanical Garden, Kenting, Hengchun, Pingdung, Taiwan (21°58'04.5"N, 120°48'55.3"E). The latter is the type locality of H. kentingensis .</p><p>Remarks.</p><p>Hypselostoma kentingensis is considered a junior synonym of this species since no major differences were observed between the two. All differences mentioned by Hwang (2014) in the original description are considered intraspecific variability. The statement in Hwang (2014) that H. kentingensis is smaller is not correct given the fact that the dimensions of the two species largely overlap (see Pilsbry 1917).</p></div>	https://treatment.plazi.org/id/54099BE0415F52958A15FEBE8D8920BB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
AF42A8DDD59656FDA54909F2206CF774.text	AF42A8DDD59656FDA54909F2206CF774.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma iunior Gojsina & Pall-Gergely 2025	<div><p>Hypselostoma iunior Gojšina &amp; Páll-Gergely sp. nov.</p><p>Figs 111 Z, 148, 149, 154</p><p>Type material.</p><p>Holotype. Thailand • 1 shell (SH: 2.44 mm, SW: 1.70 mm); Nan Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.73583&amp;materialsCitation.latitude=18.85" title="Search Plazi for locations around (long 100.73583/lat 18.85)">Pha Tup cave</a>, Ban Pha Tup, 12 km N of Nan; 18°51′0″N, 100°44′9″E; 260 m a. s. l.; 29 Apr. 1988; F. G. Thompson leg.; UF 347376 . Paratypes. Thailand • 1 shell; same data as for holotype; CUMZ 14454 • 3 shells; same data as for holotype; UF 591327 • 3 shells; Nan Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.75&amp;materialsCitation.latitude=18.87389" title="Search Plazi for locations around (long 100.75/lat 18.87389)">Ban Pha Sing</a>, 16 km N of Nan; 18°52′26″N, 100°45′0″E; 270 m a. s. l.; 28 Apr. 1988; F. G. Thompson leg.; locality code FGT-4349, UF 347358 .</p><p>Additional material examined.</p><p>Thailand • 7 shells (juveniles, not paratypes); same data as for holotype; UF 583729 .</p><p>Type locality.</p><p>Thailand, Nan Province, Pha Tup cave, Ban Pha Tun, 12 km N of Nan; 18°51′0″N, 100°44′9″E; 260 m a. s. l.</p><p>Diagnosis.</p><p>Hypselostoma with very dense spiral striation on the teleoconch. Apertural barriers very weak and four (parietal, upper palatal, lower palatal and columellar). Umbilicus narrow, widening at the last whorl.</p><p>Description.</p><p>Shell conical, light yellow to brown, consisting of 5–5.5 convex, rounded whorls separated by a deep suture. Protoconch slightly lighter than the rest of the shell, consisting of ~ 1.5 whorls, densely spirally striated (striae clearer terminally). Teleoconch very finely, densely spirally striated crossed by stronger and much coarser, irregularly spaced radial growth lines. Last whorl rounded to slightly shouldered, adnate to the penultimate and slightly to moderately descending (~ 25–35 ° compared to the shell axis) making the aperture profile prosocline to the shell axis. Peristome white but not much expanded, not reflected. It is only particularly expanded on the columellar side where it forms a strong margin which is, however, not covering the umbilicus. The peristome is discontinuous on the parietal side where it is completely merged with the penultimate whorl. Aperture almost as wide as high. Apertural barriers few and much weaker than in the majority of congeners. There are four barriers: parietal, upper palatal, lower palatal and columellar. Parietal lamella is the strongest in the aperture but still somehow weak, moderately high and curved towards the palatal side. Two palatal plicae (upper and lower palatal) are very weak, like small thickenings. Columellar lamella also weak and shares the same appearance as the palatal plicae but situated slightly closer to the peristome edge. None of the apertural barriers are reaching the thin peristome. Surface of all apertural barriers is almost smooth but very finely granulated. Sinulus wide and not strongly isolated from the rest of the aperture. Umbilicus moderately wide, measuring 1 / 5 of the shell width, initially narrow and suddenly widening at the last whorl (which gives it a slightly elongated appearance).</p><p>Differential diagnosis.</p><p>This species is similar to A. eotvosi with which is also geographically adjacent. It differs by the much larger sizes in A. eotvosi, narrower umbilicus and the lack of spiral striation. The shell of A. eotvosi is also more pointed, especially near the apex. See also under H. panhai .</p><p>Measurements</p><p>(in mm, n = 6). SH = 2.27–2.48; SW = 1.67–1.72; AH = 0.81–0.92; AW = 0.84–0.96.</p><p>Etymology.</p><p>This species is named iunior for the very weak teeth and weakly expanded peristome which is common in subadult hypselostomatid specimens.</p><p>Distribution.</p><p>This species is known only from its type locality.</p><p>Remarks.</p><p>Since the most similar species ( H. panhai) belongs to the genus Hypselostoma (as shown in a phylogenetic study by Tongkerd et al. 2004), this species is also included in this genus even though it fits in the diagnosis of Anauchen .</p></div>	https://treatment.plazi.org/id/AF42A8DDD59656FDA54909F2206CF774	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
67FBD2C26B6E5F399C99D0BC2039C6A9.text	67FBD2C26B6E5F399C99D0BC2039C6A9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma kelantanense (Sykes 1902) Gojšina & Hunyadi & Sutcharit & Tongkerd & Auffenberg & Grego & Vermeulen & Reischütz & Páll-Gergely 2025	<div><p>Hypselostoma kelantanense (Sykes, 1902) comb. nov.</p><p>Figs 224 D, 228, 229, 230, 241</p><p>Boysidia kelantanense Sykes, 1902: 61, pl. 3, fig. 7.</p><p>Boysidia kelantanensis — Möllendorff 1902: 139; Pilsbry 1917: 208, pl. 35, figs 10–12.</p><p>Paraboysidia kelantanensis — Laidlaw 1933: 213–214.</p><p>Paraboysidia kelantanensis kelantanensis — van Benthem Jutting 1950: 12–13, 39, fig. 4; Maassen 2001: 74.</p><p>Paraboysidia kelantanensis rafflesi van Benthem Jutting, 1950: 13–14, 39, fig. 5. syn. nov.</p><p>Paraboysidia kelantanensis tenuidentata van Benthem Jutting, 1950: 15–17, 39, fig. 6. syn. nov.</p><p>Paraboysidia kelantanensis rafflesi — Maassen 2001: 74.</p><p>Paraboysidia kelantanensis tenuidentata — Maassen 2001: 74.</p><p>Type material examined.</p><p>Malaysia • 1 syntype of Paraboysidia kelantanensis kelantanensis; NHMUK 1930.9. 12.13–15 • syntypes of Paraboysidia kelantanensis kelantanensis; Kelantan; UF 00113222 • 4 paratypes of Paraboysidia kelantanensis rafflesi; from the type locality; 1947; Raffles Museum, Singapore ex. coll.; RMNH.Moll.137125 • 1 paratype of Paraboysidia kelantanensis tenuidentata; from the type locality; Mar. 1939; Raffles Museum, Singapore ex. coll.; RMNH.Moll.137127. Syntypes of Paraboysidia kelantanensis kelantanensis; ANSP 98191.</p><p>Type localities.</p><p>“ Kelantan ”, Malaysia ( H. kelantanensis kelantanensis); “ Kota Tongkat, Pahang ”, Malaysia ( H. kelantanensis rafflesi); “ Kramat Pulai, Perak ”, Malaysia ( H. kelantanensis tenuidentata).</p><p>Differential diagnosis.</p><p>This species is in terms of apertural dentition and surface sculpture similar to H. frequens . However, it has a more rounded shell with more bulging whorls and narrower umbilicus than previous species. See also under H. serpa .</p><p>Distribution.</p><p>This species is known from Kelantan, Pahang, and Perak provinces in Peninsular Malaysia.</p><p>Remarks.</p><p>We observed no crucial differences between the two subspecies described by van Benthem Jutting (1950) and the nominotypical subspecies. Furthermore, one paratype of P. kelantanensis rafflesi had the identical shell morphology as that of P. kelantanensis tenuidentata which indicates that the differences mentioned by van Benthem Jutting (1950) (strength of the apertural barriers and overall shell shape) can be considered as a part of intraspecific variability. Some of the examined type specimens are more slender than others. Additionally, some of them had a small lamella between parietal and columellar, while some of them did not.</p></div>	https://treatment.plazi.org/id/67FBD2C26B6E5F399C99D0BC2039C6A9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
61815C3DF1905356966A2C30D2E9D12C.text	61815C3DF1905356966A2C30D2E9D12C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma khaochakan (Panha & J. B. Burch 2003) Gojšina & Hunyadi & Sutcharit & Tongkerd & Auffenberg & Grego & Vermeulen & Reischütz & Páll-Gergely 2025	<div><p>Hypselostoma khaochakan (Panha &amp; J. B. Burch, 2003) comb. nov.</p><p>Gyliotrachela khaochakan Panha &amp; Burch in Burch et al. 2003: 138–143, figs 4–5.</p><p>Gyliotrachela srirachaensis Panha &amp; Burch in Panha et al. 2004: 69–70, fig. 8. syn. nov.</p><p>Gyliotrachela khaochakan — Tongkerd et al. 2004: 143, fig. 2; Panha and Burch 2008: 69–70, fig. 60; Dumrongrojwattana and Wongkamhaeng 2024: 323, fig. 8.</p><p>Gyliotrachela srirachaensis — Panha and Burch 2008: 81, fig. 70; Dumrongrojwattana and Wongkamhaeng 2024: 324, fig. 8.</p><p>Type material examined.</p><p>Thailand • 2 paratypes of H. khaochakan; from the type locality; SMF 331445 • 1 paratype of H. khaochakan; from the type locality; CUMZ 44041 • 1 paratype of G. srirachaensis; from the type locality; S. Panha leg.; CUMZ ver. 44008 .</p><p>Additional material examined.</p><p>4 shells; Sa Kaeo Province, Khao Chakan district, vicinity of Wat Tham Khao Chakan; 13°39.668'N 102°05.195'E; 80 m a. s. l.; 05 Mar. 2023, A. Hunyadi &amp; J. U. Otani leg.; coll. HA.</p><p>Type localities.</p><p>“ Chakan Mountain, east of Khaochakan Temple, Khaochakan District, Srakeaw Province, 13°39'17"N, 102°5'12"E, 70 meters elevation, Thailand ” ( H. khaochakan); “ Sichang Island, Sriracha, Chonburi Province, 13°20'05"N, 100°55'24"E, 70 meters elevation ” ( H. srirachaensis).</p><p>Differential diagnosis.</p><p>This species differs from H. chedi by the separated angular and parietal lamellae. See also under H. smokon .</p><p>Distribution.</p><p>This species is, apart from the type locality, known also from Sichang Island (type locality of H. srirachaensis).</p><p>Remarks.</p><p>Gyliotrachela srirachaensis is a junior synonym of H. khaochakan since we could not find any morphological differences. The two species were not compared in the original description of G. srirachaensis .</p></div>	https://treatment.plazi.org/id/61815C3DF1905356966A2C30D2E9D12C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
9D27F6D2C6B2533BBAFB726332D252B5.text	9D27F6D2C6B2533BBAFB726332D252B5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma khaochongpran (Panha & J. B. Burch 2002)	<div><p>Hypselostoma khaochongpran (Panha &amp; J. B. Burch, 2002)</p><p>Anauchen khaochongpran Panha &amp; Burch, 2002 e: 22, fig. 3.</p><p>Hypselostoma khaochongpran — Panha and Burch 2008: 94, fig. 80; Dumrongrojwattana and Wongkamhaeng 2024: 324, fig. 8.</p><p>Type material examined.</p><p>Thailand • holotype; 1999; S. Panha leg; CUMZ 4022 .</p><p>Type locality.</p><p>“ Khaochongpran, Ratchaburi province, 13°35′38″N, 99°40′0″E, 60 meters elevation ” (Thailand).</p><p>Differential diagnosis.</p><p>This species is strikingly similar to H. cucumense but differs by the much weaker apertural barriers and less distinctly ascending last whorl.</p><p>Distribution.</p><p>This species is known only from the type locality.</p></div>	https://treatment.plazi.org/id/9D27F6D2C6B2533BBAFB726332D252B5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
5FE53108D50858959AEB2A12B43BEA2A.text	5FE53108D50858959AEB2A12B43BEA2A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma khaowongense Panha 1998	<div><p>Hypselostoma khaowongense Panha, 1998</p><p>Hypselostoma khaowongensis Panha, 1998 b: 66, fig. 4.</p><p>Hypselostoma khaowongensis — Hemmen and Hemmen 2001: 41.</p><p>Gyliotrachela saraburiensis Panha &amp; Burch in Burch et al. 2003: 143–151, figs 6–7. syn. nov.</p><p>Gyliotrachela muangon Panha &amp; Burch in Panha et al. 2004: 67–68, fig. 7. syn. nov.</p><p>Gyliotrachela khaowongensis — Panha and Burch 2008: 71–72, fig. 62; Dumrongrojwattana and Wongkamhaeng 2024: 323, fig. 8.</p><p>Gyliotrachela saraburiensis — Tongkerd et al. 2004: 143, fig. 2; Panha and Burch 2008: 78–80, fig. 68; Dumrongrojwattana and Wongkamhaeng 2024: 324, fig. 8.</p><p>Gyliotrachela muangon — Panha and Burch 2008: 76–77, fig. 66; Dumrongrojwattana and Wongkamhaeng 2024: 324, fig. 8; Gojšina et al. 2022: 137, fig. 6; Tongkerd et al. 2024: 182–184, figs 11,13O.</p><p>Type material examined.</p><p>Thailand • holotype; S. Panha leg.; CUMZ ver. 001 • paratype of G. saraburiensis; from the type locality; CUMZ 44047 • 2 paratypes of H. khaowongense; from the type locality; SMF 331439 • 3 paratypes of G. saraburiensis; from the type locality; SMF 331446 • 2 paratypes of G. muangon; from the type locality; S. Panha leg.; SMF 331459 .</p><p>Additional material examined.</p><p>Thailand-North • 24 shells; Chiang Rai Province, Chiang Rai ca 5 km west from centre of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.77737&amp;materialsCitation.latitude=9.913667" title="Search Plazi for locations around (long 99.77737/lat 9.913667)">Wat Phra That Tham Doi</a> Kong Khao; 9°54.820'N, 99°46.642'E; 11 Feb. 2015; A. Hunyadi leg.; coll. HA • 33 shells; Chiang Rai Province, ca 5 km WNW of Chiang Rai, Tham Phra cave by <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.78903&amp;materialsCitation.latitude=19.91745" title="Search Plazi for locations around (long 99.78903/lat 19.91745)">Mae Nam Kok river</a>, limestone rocks near the cave entrance; 19°55.047'N, 99°47.342'E; 420 m a. s. l; Sep. 2007; A. Reischütz leg.; coll. REI • 12 shells; Chiang Mai Province, 5 km northwest from Chiang Dao, vicinity of Wat Tham <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.92852&amp;materialsCitation.latitude=19.3938" title="Search Plazi for locations around (long 98.92852/lat 19.3938)">Chiang</a> Dao; 19°23.628'N, 98°55.711'E; 450 m a. s. l.; 07 Feb. 2015; A. Hunyadi leg.; coll. HA • 13 shells; Chiang Mai Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.058&amp;materialsCitation.latitude=19.433" title="Search Plazi for locations around (long 99.058/lat 19.433)">Doi Pha Sao Mountain</a>, 3 km W of Ban Prang Mao; 19°25.98'N, 99°3.48'E; 500 m a. s. l.; 19 Jun. 1987; F. G. Thompson leg.; locality code FGT-4338; UF 528706 • 21 shells; Chiang Mai Province, NW side of Doi Pha Sam Sao ravine with <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.04889&amp;materialsCitation.latitude=19.4075" title="Search Plazi for locations around (long 99.04889/lat 19.4075)">Dipterocarp forest</a>, leaf litter; 19°24.45'N, 99°2.9333'E; 800 m a. s. l.; 20 May 1988; F. G. Thompson leg.; locality code FGT-4453; UF 347006 • 59 shells; Chiang Mai Province, NW side of Doi Pha Sam Sao ravine with <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.04889&amp;materialsCitation.latitude=19.4075" title="Search Plazi for locations around (long 99.04889/lat 19.4075)">Dipterocarp forest</a>, leaf litter; 19°24.45'N, 99°2.9333'E; 800 m a. s. l.; 20. May 1988; F. G. Thompson leg.; locality code FGT-4452; UF 346992 • 150 shells; Mae Hong Son Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.38333&amp;materialsCitation.latitude=19.383333" title="Search Plazi for locations around (long 98.38333/lat 19.383333)">13.9 km NW Pai</a>, along road 1095; 19°23'N, 98°23'E; 780 m a. s. l.; 18 Mar. 1988; K. Auffenberg leg.; locality code KA-0571; UF 345938 • 3 shells; Mae Hong Son Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.15&amp;materialsCitation.latitude=19.566668" title="Search Plazi for locations around (long 98.15/lat 19.566668)">Ban Tham Lod Village</a>, 7 km N Soppong on Road 1095; 19°34'N, 98°9'E; 790 m a. s. l.; 18 Mar. 1988; K. Auffenberg leg.; locality code KA-0574; UF 591328 • 20 shells; Mae Hong Son Province, 21 km N Mae Hong Son, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.96667&amp;materialsCitation.latitude=19.433332" title="Search Plazi for locations around (long 97.96667/lat 19.433332)">Fish Cave</a>; 19°26'N, 97°58'E; 380 m a. s. l.; 22 Mar. 1988; K. Auffenberg leg.; locality code KA-0593; UF 345672 • 40 shells; Mae Hong Son Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.98333&amp;materialsCitation.latitude=19.45" title="Search Plazi for locations around (long 97.98333/lat 19.45)">40 km N of Mae Hong Son</a>; 19°27'N, 97°59'E; 550 m a. s. l.; 22 Mar. 1988; K. Auffenberg leg.; locality code KA-0588; UF 345643 • 1 shell; Mae Hong Son Province, 21 km N Mae Hong Son, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.96667&amp;materialsCitation.latitude=19.433332" title="Search Plazi for locations around (long 97.96667/lat 19.433332)">Fish Cave</a>; 19°26'N, 97°58'E; 380 m a. s. l.; 22 Mar. 1988; K. Auffenberg leg.; locality code KA-0595 A; UF 345702 • 7 shells; Mae Hong Son Province, Mae Hong Son, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.96667&amp;materialsCitation.latitude=19.416666" title="Search Plazi for locations around (long 97.96667/lat 19.416666)">Road</a> 1095; 19°25'N, 97°58'E; 390 m a. s. l.; 22 Mar. 1988; K. Auffenberg leg.; locality code KA-0595 B; UF 591329 • 3 shells; Mae Hong Son Province, Mae Hong Son, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.96667&amp;materialsCitation.latitude=19.416666" title="Search Plazi for locations around (long 97.96667/lat 19.416666)">Road</a> 1095; 19°25'N, 97°58'E; 390 m a. s. l.; 22 Mar. 1988; K. Auffenberg leg.; locality code KA-0595; UF 345699 • 1 shell; Lampang Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.851944&amp;materialsCitation.latitude=17.964722" title="Search Plazi for locations around (long 98.851944/lat 17.964722)">6 km SW Mae Pak</a>, limestone hill; 17°57.8833'N, 98°51.1167'E; 600 m a. s. l.; 13 June 1987; F. G. Thompson leg.; locality code FGT-4328; UF 591330 • 11 shells; Lampang Province, limestone dome at pass <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.80639&amp;materialsCitation.latitude=18.466112" title="Search Plazi for locations around (long 99.80639/lat 18.466112)">11 km SSW of Ban Pang La</a>; 18°27.9667'N, 99°48.3833'E; 610 m a. s. l.; 14 May 1988; F. G. Thompson leg.; locality code FGT-4441; UF 591331 • 2 ethanol preserved shells; same locality data as previous; F. G. Thompson leg.; locality code FGT-4440; UF 346908 • 43 shells; Phrae Province, limestone knoll <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.944725&amp;materialsCitation.latitude=18.168888" title="Search Plazi for locations around (long 99.944725/lat 18.168888)">4 km N of Ban Nam Rin</a>, deep rocks and leaves; 18°10.1333'N, 99°56.6833'E; 200 m a. s. l.; 16. May 1988; F. G. Thompson leg.; locality code FGT-4444; UF 346931 • 1 shell; Phrae Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.346115&amp;materialsCitation.latitude=18.385" title="Search Plazi for locations around (long 100.346115/lat 18.385)">Than Pha Nan Cave</a>, 6 km NE of Rang Kwang; 18°23.1'N, 100°20.7667'E; 300 m a. s. l.; 30 Apr. 1988; F. G. Thompson leg.; locality code FGT-4400; UF 347403 • 2 shells; Phrae Province., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.944725&amp;materialsCitation.latitude=18.168888" title="Search Plazi for locations around (long 99.944725/lat 18.168888)">4 km N of Ban Nam Rin valley</a> with limestone ridges, leaf litter at base of boulders; 18°10.1333'N, 99°56.6833'E; 200 m a. s. l.; 16. May 1988; F. G. Thompson leg.; locality code FGT-4445; UF 346935 • 5 ethanol preserved shells; Phrae Province, 11 km W of Phrae, on <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.07056&amp;materialsCitation.latitude=18.178055" title="Search Plazi for locations around (long 100.07056/lat 18.178055)">Road</a> 1023 teak plantation and cut forest; 18°10.6833'N, 100°4.2333'E; 290 m a. s. l.; 16. May 1988; F. G. Thompson leg.; locality code FGT-4446; UF 346967 • 19 shells; same locality data as previous; F. G. Thompson leg.; locality code FGT-4447; UF 591332. Thailand-Northeast • 28 shells; Petchabun Province, 24.4 km S. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.166664&amp;materialsCitation.latitude=16.266666" title="Search Plazi for locations around (long 101.166664/lat 16.266666)">Petchabun on Hwy.</a> 2275, evergreen forest in back of outcrop, leaf litter next to rocks; 16°16'N, 101°10'E; 120 m a. s. l.; 02. May 1988; K. Auffenberg leg.; locality code KA-0693; UF 346096 • 1 shell; Nakhon Ratchasima Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.38&amp;materialsCitation.latitude=14.534" title="Search Plazi for locations around (long 101.38/lat 14.534)">3.4 km W of Ban Mu Si</a>; 14.534°N, 101.380°E; 380 m a. s. l.; 05. May 1987; F. G. Thompson leg.; locality code FGT-4241; UF 534868 • 9 shells; Nakhon Ratchasima Province, Pak Chong district, mountain above <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.265434&amp;materialsCitation.latitude=14.614417" title="Search Plazi for locations around (long 101.265434/lat 14.614417)">Wat Thep Phithak Punnaram</a>; 14°36.865'N, 101°15.926'E; 500 m a. s. l.; 04 Mar. 2023; A. Hunyadi leg.; coll. HA. Thailand-West • 16 shells; Kanchanaburi Province, Erawan Nat. Park, Erawan Falls Trail; 14°22.310'N, 99°08.699'E; 85 m a. s. l.; 17 Feb. 2015; A. Hunyadi leg.; coll. HA • 10 shells; Petchabun Province, 24.4 km S. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.166664&amp;materialsCitation.latitude=16.266666" title="Search Plazi for locations around (long 101.166664/lat 16.266666)">Petchabun on Hwy.</a> 2275, evergreen forest in back of outcrop, leaf litter next to rocks; 16°16'N, 101°10'E; 120 m a. s. l.; 02. May 1988; K. Auffenberg leg.; locality code KA-0694; UF 346100. Thailand-Central • 10 shells; Phitsanulok Province, Noen Maprang district, Chomphu, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.68812&amp;materialsCitation.latitude=16.677517" title="Search Plazi for locations around (long 100.68812/lat 16.677517)">Tham Phra Wang Daeng</a>; 16°40.651'N, 100°41.287'E; 02 Mar. 2023; A. Hunyadi, K. Okubo &amp; J. U. Otani leg.; coll. HA • 37 shells; Nakhon Sawan Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.26667&amp;materialsCitation.latitude=15.233334" title="Search Plazi for locations around (long 100.26667/lat 15.233334)">19.5 km NE intersection of Hwys 11 and 32</a>, limestone outcrop, base of cliff under Georissa zone; 15°14'N, 100°16'E (provided coordinates point in Chai Nat Province); 50 m a. s. l.; 02. May 1988; K. Auffenberg leg.; locality code KA-0697; UF 346135 • 1 shell; Nakhon Sawan Province, 19.5 km NE intersection of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.26667&amp;materialsCitation.latitude=16.233334" title="Search Plazi for locations around (long 100.26667/lat 16.233334)">Hwys</a> 11 and 32; 16°14'N, 100°16'E; 50 m a. s. l.; 02. May 1988; K. Auffenberg leg.; locality code KA-0697; UF 346126 • 1 shell; Lopburi Province, Tha Wung district, Khao Samo Khon, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.50252&amp;materialsCitation.latitude=14.902317" title="Search Plazi for locations around (long 100.50252/lat 14.902317)">Wat Tham Chang Pueak</a>; 14°54.139'N, 100°30.151'E; 30 m a. s. l.; 03 Mar. 2023; A. Hunyadi leg.; coll. HA • 2 shells; Saraburi Province, behind <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.8&amp;materialsCitation.latitude=14.733" title="Search Plazi for locations around (long 100.8/lat 14.733)">Phra Phutthabat</a>, limestone hill with scrubby evergreen forest, leaf litter among rocks; 14.733°N, 100.800°E; 50 m a. s. l.; 03. May 1988; K. Auffenberg leg.; locality code KA-0699; UF 508979 . Laos • 27 shells; Luang Prabang Province, old forest on Phousi Mountain in Luang Prabang, clay, black soil, on and under rocks; 19°53.411'N, 102°08.216'E; 374 m a. s. l.; 13 Nov. 2004; A. Abdou &amp; I. Muratov leg.; MNHN -IM-2012-25387 • 6 shells; Luang Prabang Province, 3.1 km from centre of Nong Khiaw towards Pak Xeng, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.6287&amp;materialsCitation.latitude=20.553583" title="Search Plazi for locations around (long 102.6287/lat 20.553583)">Tham Pha Toke</a>, vicinity of the cave; 20°33.215'N, 102°37.722'E; 345 m a. s. l.; 05 Oct. 2019; A. Hunyadi, K. Okubo &amp; J. U. Otani leg.; coll. HA • 24 shells; Luang Prabang Province, 12 km from centre of Nong Khiaw towards Pak Xeng, northeast from <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.68602&amp;materialsCitation.latitude=20.543083" title="Search Plazi for locations around (long 102.68602/lat 20.543083)">Ban</a> Huai Lek, left side of the road; 20°32.585'N, 102°41.161'E; 345 m a. s. l.; 05 Oct. 2019; A. Hunyadi, K. Okubo &amp; J. U. Otani leg.; coll. HA . Myanmar • 3 shells; Shan State, c. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.268616&amp;materialsCitation.latitude=20.1875" title="Search Plazi for locations around (long 97.268616/lat 20.1875)">3.6 km NNE of Hsihseng</a> centre, limestone hill; 20°11′15″N, 97°16′07″E; 1040 m a. s. l.; 07 Oct. 2018; A. Hunyadi, K. Okubo &amp; J. U. Otani leg.; coll. HA • 7 shells; Shan State, 9.5 km from centre of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.25857&amp;materialsCitation.latitude=20.234434" title="Search Plazi for locations around (long 97.25857/lat 20.234434)">Hsihseng</a>; 20°14.066'N, 97°15.514'E; 1140 m a. s. l.; 07 Oct. 2018; A. Hunyadi, K. Okubo &amp; J. U. Otani leg.; coll. HA • 21 shells; Shan State, 3.6 km north-northeast from centre of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.26857&amp;materialsCitation.latitude=20.187468" title="Search Plazi for locations around (long 97.26857/lat 20.187468)">Hsihseng</a>; 20°11.248'N, 97°16.114'E; 1040 m a. s. l.; 07 Oct. 2018; A. Hunyadi, K. Okubo &amp; J. U. Otani leg.; coll. HA • 63 shells; Shan State, ca 16 km from centre of Taunggyi centre towards <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.137314&amp;materialsCitation.latitude=20.787716" title="Search Plazi for locations around (long 97.137314/lat 20.787716)">Hopong</a>, 1.5 km north on rd. # 4, „ Shwe Pyi Aunchonda ” monastery; 20°47.263'N, 97°8.239'E; 1110 m a. s. l.; 08 Oct. 2018; A. Hunyadi, K. Okubo &amp; J. U. Otani leg.; coll. HA • 57 shells; Shan State, ca 6 km east from centre of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.3004&amp;materialsCitation.latitude=20.133368" title="Search Plazi for locations around (long 97.3004/lat 20.133368)">Hsihseng</a>; 20°8.002'N, 97°18.024'E; 500 m a. s. l.; 07 Oct. 2018; A. Hunyadi, K. Okubo &amp; J. U. Otani leg.; coll. HA .</p><p>Type localities.</p><p>“ A limestone hill near Takli District, Nakornsawan Province, 15°16′52″N, 100°22′38″E, 114 meters elevation ”, Thailand ( H. khaowongense); “ East of Tepitak Tamaram Temple, Tepitak Mountain, Muang Saraburi District, Saraburi Province, 14°36'57"N, 101°15'50"E, 510 meters elevation. Thailand ” ( G. saraburiensis); “ Muangon Cave, San Kam Pang District, Chiangmai Province, 18°47'11"N, 99°14'16"E, 280 meters elevation ” ( G. muangon).</p><p>Differential diagnosis.</p><p>This species is similar to H. holimanae but is clearly different by the separated angular and parietal lamellae (fused in H. holimanae) as well as the presence of numerous and variable apertural barriers (only four in H. holimanae). Hypselostoma fruhstorferi has a narrower umbilicus and fewer apertural barriers. This species is also similar to H. cultura from which it can be separated by the less prominently shouldered last whorl and a narrower umbilicus.</p><p>Distribution.</p><p>Clearly a widespread species: its distribution stretches from Chiang Rai province in N Thailand to the southernmost known localities in Ratchaburi and Chonburi provinces. Westwards, this species stretches to Taunggyi in Myanmar and the easternmost locality is known from Luang Prabang province in Laos.</p><p>Remarks.</p><p>Examined specimens had the usual main barriers and a larger number of smaller barriers between them of very variable number (usually 2–5 between each of the main barriers). Umbilicus was always wide but sometimes more or less so (specimens from Chiang Mai (Thailand) had the narrowest umbilicus). Last whorl is always at least very slightly keeled, in some specimens more prominent. Spiral striation is present, in some populations hardly observable and in some very rough. During comparison of the type material of G. saraburiensis and G. muangon, we have not observed any difference between these species and H. khaowongense . The former two are not compared with the latter in the respective original descriptions. Because of no difference observed, we treat both G. saraburiensis and G. muangon as junior synonyms of H. khaowongense .</p></div>	https://treatment.plazi.org/id/5FE53108D50858959AEB2A12B43BEA2A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
A0D2A6E194C752C9A7AC44D38F0BCDD3.text	A0D2A6E194C752C9A7AC44D38F0BCDD3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma khaowongkot (Panha & J. B. Burch 2004) Gojšina & Hunyadi & Sutcharit & Tongkerd & Auffenberg & Grego & Vermeulen & Reischütz & Páll-Gergely 2025	<div><p>Hypselostoma khaowongkot (Panha &amp; J. B. Burch, 2004) comb. nov.</p><p>Fig. 185</p><p>Gyliotrachela khaowongkot Panha &amp; Burch in Panha et al. 2004: 65–67, fig. 6.</p><p>Gyliotrachela khaowongkot — Panha and Burch 2008: 72–73, fig. 63; Dumrongrojwattana and Wongkamhaeng 2024: 323, fig. 8.</p><p>Material examined.</p><p>Thailand • holotype; S. Panha leg.; CUMZ ver. 44031 .</p><p>Type locality.</p><p>“ Khaowongkot, Ban Mee District, Lopburi Province, 15°1'13"N, 100°32'44"E, 60 meters elevation ”.</p><p>Differential diagnosis.</p><p>See under H. tridentatum and H. utongense .</p><p>Distribution.</p><p>This species is known only from the type locality.</p></div>	https://treatment.plazi.org/id/A0D2A6E194C752C9A7AC44D38F0BCDD3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
68759A089ECD5694B83ABEF1F1366DBD.text	68759A089ECD5694B83ABEF1F1366DBD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma khmerianum (Sutcharit & Panha 2023) Gojšina & Hunyadi & Sutcharit & Tongkerd & Auffenberg & Grego & Vermeulen & Reischütz & Páll-Gergely 2025	<div><p>Hypselostoma khmerianum (Sutcharit &amp; Panha, 2023) comb. nov.</p><p>Figs 186 G, 202, 223</p><p>Gyliotrachela khmeriana Sutcharit &amp; Panha in Sutcharit et al. 2023: 1295–1299, fig. 4.</p><p>Type material examined.</p><p>Cambodia • holotype; CUMZ 14206 .</p><p>Additional material examined.</p><p>Thailand • 15 shells (4 adult, 1 of them broken); Sa Kaeo Province, Khlong Hat district, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.328186&amp;materialsCitation.latitude=13.415767" title="Search Plazi for locations around (long 102.328186/lat 13.415767)">Tham Phet Pho Thong</a>; 13°24.946'N, 102°19.691'E; 03 June 2023; A. Hunyadi leg.; coll. HA .</p><p>Type locality.</p><p>“ The limestone hills at Phnom Sampeov Mountain, Banan District, Battambang Province, Cambodia (13.026000°N, 103.101011°E) ”.</p><p>Differential diagnosis.</p><p>This species is not similar to any hitherto described congener due to the combination of conical shell, absence of spiral striation, all whorls keeled, strong but not numerous apertural barriers, and separated angular and parietal lamellae. The shell shape somehow resembles H. sculpturatum but it differs from it by its adnate last whorl, separated angular and parietal lamellae and not spirally striated teleoconch. Hypselostoma circumcarinatum sp. nov. has its whorls keeled, a merged angulo-parietal lamella and spirally striated whorls. It is different from H. chatnareeae by the separated angular and parietal lamellae, weaker keel, and the absence of a deep groove above the keel.</p><p>Distribution.</p><p>This species is known from the type locality in Cambodia and is now reported for the first time in Thailand (Sa Kaeo province).</p></div>	https://treatment.plazi.org/id/68759A089ECD5694B83ABEF1F1366DBD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
651CF96A72E658D6A9D44462F99BCC7C.text	651CF96A72E658D6A9D44462F99BCC7C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma kohrin (Panha & J. B. Burch 2003) Gojšina & Hunyadi & Sutcharit & Tongkerd & Auffenberg & Grego & Vermeulen & Reischütz & Páll-Gergely 2025	<div><p>Hypselostoma kohrin (Panha &amp; J. B. Burch, 2003) comb. nov.</p><p>Gyliotrachela kohrin Panha &amp; Burch in Burch et al. 2003: 151–157, figs 8–9.</p><p>Gyliotrachela kohrin — Tongkerd et al. 2004: 143, fig. 2; Panha and Burch 2008: 74–75, fig. 64; Dumrongrojwattana and Wongkamhaeng 2024: 323, fig. 8.</p><p>Material examined.</p><p>Thailand • 1 shell; Chonburi Province, Koh Rin, 25 km from eastern coast of Thailand; 26 Oct. 2013; J. U. Otani leg.; coll. PGB • 2 shells; Chonburi Province, Pattaya, Koh Rin, 28 June 2014; J. U. Otani leg.; coll. PGB .</p><p>Type locality.</p><p>“ On a limestone hill on Kohrin (Rin Island), which is a small island located in the upper Gulf of Thailand, south of Pattya Beach, eastern coast of Thailand, about 25 kilometers from the coast, Satthip District, Chonberi Province, 12°48'5"N, 100°42'4"E, 30 meters elevation, Thailand ”.</p><p>Differential diagnosis.</p><p>This species is similar to H. diarmaidi from which it can be separated by the last whorl distally shortly detached as well as the weaker keel and not so concave surface above it. See also under H. surakiti .</p><p>Distribution.</p><p>This species is known only from the type locality.</p></div>	https://treatment.plazi.org/id/651CF96A72E658D6A9D44462F99BCC7C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
75F14CD8BADC5C099EDD1807B489DA0F.text	75F14CD8BADC5C099EDD1807B489DA0F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma loei Panha & Prateespasen 2005	<div><p>Hypselostoma loei Panha &amp; Prateespasen, 2005</p><p>Figs 186 H, 203, 223</p><p>Hypselostoma loei Panha &amp; Prateespasen, 2005: 99, fig. 2.</p><p>Hypselostoma loei — Panha and Burch 2008: 94–95, fig. 81; Dumrongrojwattana and Wongkamhaeng 2024: 324, fig. 8.</p><p>Type material examined.</p><p>Thailand • 1 paratype; from the type locality; CUMZ ver. 025 .</p><p>Additional material examined.</p><p>Thailand-Central • 3 shells; Phitsanulok Province, Noen Maprang district, Chomphu, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.68812&amp;materialsCitation.latitude=16.677517" title="Search Plazi for locations around (long 100.68812/lat 16.677517)">Tham Phra Wang Daeng</a>; 16°40.651'N, 100°41.287'E; 150 m a. s. l.; 02 Mar. 2023; A. Hunyadi &amp; J. U. Otani leg.; coll. HA • Thailand- Northeast • 54 shells; Loei Province, Nong Hin district, 20.3 km southwest from the centre of Nong Hin towards <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.727585&amp;materialsCitation.latitude=17.041183" title="Search Plazi for locations around (long 101.727585/lat 17.041183)">Pha Wai</a>, left side of the road no. 3029; 17°2.471'N, 101°43.655'E; 705 m a. s. l.; 28 Feb. 2023; A. Hunyadi &amp; J. U. Otani leg.; coll. HA • 46 shells; Loei Province, Mueang Loei district, rock wall above <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.859634&amp;materialsCitation.latitude=17.464933" title="Search Plazi for locations around (long 101.859634/lat 17.464933)">Wat Tham Piya Thammarangsi</a>; 17°27.896'N, 101°51.578'E; 27 Feb. 2023; A. Hunyadi &amp; J. U. Otani leg.; coll. HA • 9 shells; Loei Province, 9.2 km NW Loei Buddhist temple, up and N of main cave entrance, among leaves; 17°35'N, 101°44'E; 28 Apr. 1988; K. Auffenberg leg.; locality code KA-0685, UF 345859 • 1 shell; Loei Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.889&amp;materialsCitation.latitude=17.474" title="Search Plazi for locations around (long 101.889/lat 17.474)">1 km E Ban Huai Muang</a>; 17.474°N, 101.889°E; 375 m a. s. l.; 21. May 1987; F. G. Thompson leg.; locality code FGT-4274, UF 527260 • 11 shells; Nong Bua Lamphu Province, Na Klang district, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.11736&amp;materialsCitation.latitude=17.315517" title="Search Plazi for locations around (long 102.11736/lat 17.315517)">Wat Phuttha Banphot</a> - Tham Pha Choo; 17°18.931'N, 102°7.042'E; 27 Feb. 2023; A. Hunyadi &amp; J. U. Otani leg.; coll. HA • 13 shells; Khon Kaen Province, Phu Pha Man district, rocks above <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.80157&amp;materialsCitation.latitude=16.652834" title="Search Plazi for locations around (long 101.80157/lat 16.652834)">Wat Phu Hua Chang</a>; 16°39.170'N, 101°48.094'E; 345 m a. s. l.; 01 Mar. 2023; A. Hunyadi &amp; J. U. Otani leg.; coll. HA • 10 shells; Khon Kaen Province, Chum Phae district, Na Nong Tum, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.95637&amp;materialsCitation.latitude=16.817017" title="Search Plazi for locations around (long 101.95637/lat 16.817017)">Kled Kaew cave</a>; 16°49.021'N, 101°57.382'E; 580 m a. s. l.; 01 Mar. 2023; A. Hunyadi &amp; J. U. Otani leg.; coll. HA</p><p>Type locality.</p><p>“ Limestone hills in Loei Province, 16°45'47"N. 101°59'8"E, 110 meters elevation … Thailand … ”.</p><p>Differential diagnosis.</p><p>This species is similar to H. hungerfordianum but the latter has separate angular and parietal lamellae and more numerous barriers.</p><p>Distribution.</p><p>Known from Loei, Phitsanulok, Khon Kaen and Nong Bua Lamphu provinces in Thailand.</p><p>Remarks.</p><p>The striking similarity of this species with H. hungerfordianum suggests that Gyliotrachela is a junior synonym of Hypselostoma .</p></div>	https://treatment.plazi.org/id/75F14CD8BADC5C099EDD1807B489DA0F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
63EBE0B33D4D577BA5158FC59B135BDD.text	63EBE0B33D4D577BA5158FC59B135BDD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma luctans (van Benthem Jutting 1950) Gojšina & Hunyadi & Sutcharit & Tongkerd & Auffenberg & Grego & Vermeulen & Reischütz & Páll-Gergely 2025	<div><p>Hypselostoma luctans (van Benthem Jutting, 1950) comb. nov.</p><p>Figs 186 I, 204, 205, 223</p><p>Gyliotrachela luctans van Benthem Jutting, 1950: 33–34, fig. 19.</p><p>Gyliotrachela luctans — van Benthem Jutting 1950: 46; Zilch 1984: 166; Maassen 2001: 76; Foon et al. 2017: 79, fig. 30 C.</p><p>Type material examined.</p><p>Malaysia • 23 paratypes; from the type locality; 1938; Raffles Museum Singapore ex. coll.; RMNH.Moll.137158 .</p><p>Additional material examined.</p><p>Thailand • 7 shells; Yala Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.21892&amp;materialsCitation.latitude=6.33005" title="Search Plazi for locations around (long 101.21892/lat 6.33005)">9.5 km from the junction to Bannang Sata, 1700 m north from the road no 4077, left side of the side road</a>; 6°19.803'N, 101°13.135'E; 120 m a. s. l.; 22 Feb. 2023; A. Hunyadi leg.; coll. HA .</p><p>Type locality.</p><p>“ Gunong Pondok, Padang Rengas, Perak ”, Malaysia.</p><p>Differential diagnosis.</p><p>This species is similar to H. bubalus sp. nov. with which it is also geographically approximate. It differs from it by a clearly narrower umbilicus. Hypselostoma bubalus sp. nov. is also spirally striated on some isolated parts of the shell (above or below the keel), but H. luctans is not. See also under H. piconis .</p><p>Distribution.</p><p>This species is known from Peninsular Malaysia and Yala Province in Thailand.</p><p>Remarks.</p><p>There can be two interpalatal plicae, and in some cases, none are visible. Umbilicus can sometimes be slightly wider, with a stronger groove inside (than the figured specimen).</p></div>	https://treatment.plazi.org/id/63EBE0B33D4D577BA5158FC59B135BDD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
E46B9263724853E2947D2A28B51E5DDC.text	E46B9263724853E2947D2A28B51E5DDC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma modestum (van Benthem Jutting 1950) Gojšina & Hunyadi & Sutcharit & Tongkerd & Auffenberg & Grego & Vermeulen & Reischütz & Páll-Gergely 2025	<div><p>Hypselostoma modestum (van Benthem Jutting, 1950) comb. nov.</p><p>Figs 224 E, 231, 241</p><p>Gyliotrachela modesta van Benthem Jutting, 1950: 31–32, fig. 18.</p><p>Gyliotrachela modesta — van Benthem Jutting 1950: 46; Davison and Kiew 1990: 25; Maassen 2001: 76.</p><p>Type material examined.</p><p>Malaysia • 2 paratypes; from the type locality; 1939; Raffles Museum, Singapore ex. coll.; RMNH.Moll.137148 .</p><p>Type locality.</p><p>“ Gua Musang, Kelantan ”, Malaysia.</p><p>Differential diagnosis.</p><p>This species is very similar to H. emergens from which it differs in its narrower last whorl (but also the entire shell) which is also more detached and protruding. The keel on the last whorl is also stronger in H. emergens (best seen in the lateral view). Hypselostoma frequens has a narrower umbilicus and more enlarged last whorl when compared to the previous ones, which makes the shell less slender. See also under H. muaklekense and H. burchi .</p><p>Distribution.</p><p>This species is known only from the type locality.</p></div>	https://treatment.plazi.org/id/E46B9263724853E2947D2A28B51E5DDC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
0E054234EDE15B4F921E1FBBCFC7B938.text	0E054234EDE15B4F921E1FBBCFC7B938.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma muaklekense (Panha & J. B. Burch 2002) Gojšina & Hunyadi & Sutcharit & Tongkerd & Auffenberg & Grego & Vermeulen & Reischütz & Páll-Gergely 2025	<div><p>Hypselostoma muaklekense (Panha &amp; J. B. Burch, 2002) comb. nov.</p><p>Figs 242 D, 253, 259</p><p>Paraboysidia muaklekensis Panha &amp; Burch, 2002 c: 79–81, fig. 2.</p><p>Gyliotrachela muaklekensis — Panha and Burch 2008: 75–76, fig. 65; Dumrongrojwattana and Wongkamhaeng 2024: 324, fig. 8.</p><p>Type material examined.</p><p>Thailand • holotype; 1997; S. Panha leg.; CUMZ ver. 020 .</p><p>Type locality.</p><p>“ Tepitak mountain, Muaklek District, Saraburi Province, 14°36'57"N, 101 ' 15 ' 50 " E, 700 meters elevation ” (Thailand).</p><p>Differential diagnosis.</p><p>This species is most similar to H. modestum from which it can be separated by the wider umbilicus, three strong apertural barriers on the palatal side (two in H. modestum) and smaller dimensions (SH = 2.9–3.1 and SW = 2.3–2.7 in H. modestum while SH = 1.8–2.0 and SW = 1.4–1.5 in H. muaklekense). Hypselostoma frequens also has only two strong barriers on the palatal side, is less slender and larger (SH = 2.8–3.5, SW = 2.4–2.6).</p><p>Distribution.</p><p>This species is known only from the type locality.</p></div>	https://treatment.plazi.org/id/0E054234EDE15B4F921E1FBBCFC7B938	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
94691362CCF75175A3DC2CE31A574849.text	94691362CCF75175A3DC2CE31A574849.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma ophis Gojsina, Hunyadi & Pall-Gergely 2025	<div><p>Hypselostoma ophis Gojšina, Hunyadi &amp; Páll-Gergely sp. nov.</p><p>Figs 186 J, 206, 207, 223</p><p>Type material.</p><p>Holotype. Thailand • 1 shell (SH: 2.1 mm; SW: 2.6 mm); Phattalung Province, Phattalung, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.08884&amp;materialsCitation.latitude=7.6251" title="Search Plazi for locations around (long 100.08884/lat 7.6251)">Khao Ok Thalu</a>, rock wall; 07°37.506'N, 100°05.330'E; 19 Feb. 2015; A. Hunyadi leg.; CUMZ 14467 . Paratypes. Thailand • 5 shells (3 whole and 2 last whorls with retained apertures); same data as for holotype; coll. HA .</p><p>Additional material examined.</p><p>Thailand • 1 shell (fragment, not paratype); same data as for holotype; coll. HA .</p><p>Type locality.</p><p>Thailand, Phattalung Province, Phattalung, Khao Ok Thalu, rock wall; 07°37.506'N, 100°05.330'E.</p><p>Diagnosis.</p><p>A Hypselostoma species with sandpaper-like shell surface and a bluntly keeled last whorl which is detached from the penultimate and slightly ascending upwards. Five main barriers in the aperture (parietal, angular, upper palatal, lower palatal and columellar). Parietal lamella very long and wavy. Umbilicus moderately wide.</p><p>Description.</p><p>Shell shape concave-conical (due to the strongly enlarged last whorl), consisting of 4.5–5 convex whorls separated by a deep suture. Colouration dark brown, weakly glossy and opaque. Protoconch finely pitted, showing very weak spiralling pattern, consisting of ~ 1.25–1.5 whorls, coloured the same as the teleoconch. Teleoconch very finely granulated (sandpaper-like) with additional radial growth lines. Also, some irregularly spaced radial white streaks are occasionally visible. Last whorl bluntly keeled, moderately detached from the penultimate and slightly ascending upwards (~ 20 ° compared to the shell axis). In lateral view, two strong grooves are visible near the aperture, one above and one below the central keel. Peristome expanded and not reflected, slightly lighter than the rest of the shell. Aperture equipped with five strong and thick barriers (parietal, angular, upper palatal, lower palatal, and columellar) and several smaller ones. Parietal lamella very strong, wavy, and very long. Angular lamella short and sometimes separated in inner and outer part. Inner part is weaker. Both palatal plicae very strong, nearly as strong as the parietal lamella or even stronger but clearly shorter. In front of the upper palatal plica, a small swelling is present which is probably homologous with the palatal tubercle present in other genera (e. g., Bensonella, see Páll-Gergely and White 2023) Columellar lamella nearly horizontal, stronger only than the angular among the main barriers. Additionally, smaller barriers are present between the main ones. There is usually one in the interpalatal region, one basal and one between the columellar and parietal. None of these barriers reach the peristome. Surface of all barriers is very finely spiniferous medially and very finely granulated laterally. Sinulus rounded and visibly separated from the rest of the aperture. Umbilicus open and moderately wide, measuring 1 / 3 of the shell width and showing the penultimate whorl. A strong groove running from the peristome towards the umbilicus is visible. This groove is getting lost near the penultimate whorl.</p><p>Differential diagnosis.</p><p>See under H. venustum .</p><p>Measurements</p><p>(in mm, n = 4). SH = 1.76–2.1; SW 1 = 2.38–2.6; SW 2 = 1.48–1.54; AH = 0.88–0.95; AW = 0.85–0.89.</p><p>Etymology.</p><p>The specific epithet is derived from the Greek work ophis (snake) which is in special reference to the curved and long parietal lamella (which is also one of the longest in all hitherto described Hypselostoma species). To be used as a noun in apposition.</p><p>Distribution.</p><p>This species is known only from the type locality.</p></div>	https://treatment.plazi.org/id/94691362CCF75175A3DC2CE31A574849	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
5676833304895855ADBD69D06A8CB2DC.text	5676833304895855ADBD69D06A8CB2DC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma paini (F. G. Thompson & Dance 1983) Gojšina & Hunyadi & Sutcharit & Tongkerd & Auffenberg & Grego & Vermeulen & Reischütz & Páll-Gergely 2025	<div><p>Hypselostoma paini (F. G. Thompson &amp; Dance, 1983) comb. nov.</p><p>Figs 224 F, 232, 239</p><p>Boysidia (Dasypupa) paini Thompson &amp; Dance, 1983: 107–108, figs 9, 10.</p><p>Boysidia paini — Dumrongrojwattana and Assawawattagee 2018: 216.</p><p>Type material examined.</p><p>Malaysia • holotype; UF 35949 • 6 paratypes; from the type locality; UF 35951 .</p><p>Additional material examined.</p><p>Malaysia • 4 shells; Borneo, Sarawak, Fifth division, SW flank Gunong Budah, Medalam Valley, trib. Of Limbang R.; UF 35950 .</p><p>Type locality.</p><p>“ Foot of cliff about 400 yards north of the main to Deer Cave, Melinau Paku Valley, Fourth Div., Sarawak, Borneo, 04°5'N, 114°53'E ”.</p><p>Differential diagnosis.</p><p>See under H. procerum and H. tertiusfrater nom. nov.</p><p>Distribution.</p><p>This species is known only from Sarawak, Borneo.</p></div>	https://treatment.plazi.org/id/5676833304895855ADBD69D06A8CB2DC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
4D3E43294DE85AF594AC5CA1C0254D1E.text	4D3E43294DE85AF594AC5CA1C0254D1E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma panhai J. B. Burch & Tongkerd 2003	<div><p>Hypselostoma panhai J. B. Burch &amp; Tongkerd, 2003</p><p>Fig. 185</p><p>Hypselostoma panhai Burch &amp; Tongkerd in Burch et al. 2003: 178–183, fig. 18.</p><p>Hypselostoma panhai — Tongkerd et al. 2004: 145, fig. 4 D – H; Panha and Burch 2008: 95–97, fig. 82; Dumrongrojwattana and Wongkamhaeng 2024: 324, fig. 8.</p><p>Type material examined.</p><p>Thailand • holotype; CUMZ 44055 .</p><p>Type locality.</p><p>“ Chongkhaokad, west of the World War II museum, near the famous bridge of the now abandoned railroad of World War II, Triyoke District, Kanchanaburi Province, 14°24'8"N, 98°53'23"E, 140 meters elevation, Thailand ”.</p><p>Differential diagnosis.</p><p>This species is similar to H. edentatum but the latter lacks apertural barriers. However, this species is most similar to H. iunior sp. nov. from which it can be separated by the less conical shell, deeper sutures, more convex whorls, clearly weaker parietal callus, slightly wider umbilicus, and much coarser spiral striation (dense in H. iunior sp. nov.).</p><p>Distribution.</p><p>This species is known only from the type locality.</p></div>	https://treatment.plazi.org/id/4D3E43294DE85AF594AC5CA1C0254D1E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
4338866150C656CF8C5239786E1CA04A.text	4338866150C656CF8C5239786E1CA04A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma pattalungense Panha & J. B. Burch 2004	<div><p>Hypselostoma pattalungense Panha &amp; J. B. Burch, 2004</p><p>Figs 242 E, 254, 259</p><p>Hypselostoma pattalungensis Panha &amp; Burch in Panha et al. 2004: 73–75, fig. 11.</p><p>Hypselostoma pattalungensis — Panha and Burch 2008: 97–98, fig. 83; Dumrongrojwattana and Wongkamhaeng 2024: 324, fig. 8.</p><p>Type locality.</p><p>“ Ko Si Ko Ha (Ko Na Thewada), Limestone Hill, Pattalung Province ” (Thailand).</p><p>Type material examined.</p><p>Thailand • 3 paratypes; from the type locality; S. Panha leg.; SMF 331456 .</p><p>Additional material examined.</p><p>Thailand • 3 shells; Phattalung Province, 6.4 km E of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.183334&amp;materialsCitation.latitude=7.45" title="Search Plazi for locations around (long 100.183334/lat 7.45)">Hwy.</a> 4081, 1 km SW of Khao Chai Son; 7°27'N, 100°11'E; 50 m a. s. l.; 11 Apr. 1988; K. Auffenberg leg.; locality code KA-0648; UF 345066 • 1 shell; same locality data as previous; UF 591338 • 13 shells; Songkhla Province, 31.3 km NW <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.26667&amp;materialsCitation.latitude=7.1666665" title="Search Plazi for locations around (long 100.26667/lat 7.1666665)">Hat Yai</a>, 1.2 km W of Hwy. 43; 7°10'N, 100°16'E; 80 m a. s. l.; 08 Apr. 1988; K. Auffenberg leg.; locality code KA-0636; UF 344933 • 1 shell; same locality data as previous; 08 Apr. 1988; K. Auffenberg leg.; locality code KA-0635; UF 591337 .</p><p>Differential diagnosis.</p><p>This species is most similar to H. fortunatum sp. nov., from which it differs by stronger and more numerous apertural barriers, separated angular and parietal lamellae as well as the higher spire.</p><p>Distribution.</p><p>This species is known from Phattalung and Songkhla provinces, Thailand.</p><p>Remarks.</p><p>Intraspecific variability is noticed on the appearance of the subcolumellar lamella, it can be present or completely absent.</p></div>	https://treatment.plazi.org/id/4338866150C656CF8C5239786E1CA04A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
70BC59E38C9A575A9A6FEC2897365365.text	70BC59E38C9A575A9A6FEC2897365365.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma pendulum (Panha & J. B Burch 2002) Gojšina & Hunyadi & Sutcharit & Tongkerd & Auffenberg & Grego & Vermeulen & Reischütz & Páll-Gergely 2025	<div><p>Hypselostoma pendulum (Panha &amp; J. B Burch, 2002) comb. nov.</p><p>Figs 242 F, 255, 259</p><p>Antroapiculus pendulus Panha &amp; Burch, 2002 a: 144–148, figs 2, 3.</p><p>Anthroapiculus pendulus [sic] — Dumrongrojwattana and Wongkamhaeng 2024: 323, fig. 7.</p><p>Type material examined.</p><p>Thailand • 1 paratype; from the type locality; May 1998.; S. Panha, J. B. Burch, P. Dumrongrojwattana, C. Sutcharit, S. Tumpeesuwan, P. Tongkerd, W. Wanarat, S. Klomtong leg.; CUMZ ver. 077 .</p><p>Type locality.</p><p>“ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.93917&amp;materialsCitation.latitude=17.2475" title="Search Plazi for locations around (long 98.93917/lat 17.2475)">Chonglom Mountain</a>, Bhumiphol Dam reservoir, Samngao District, Tak province 17°14′51″N, 98°56′21″E ” (Thailand).</p><p>Differential diagnosis.</p><p>This species is not similar to any other congener due to the completely flat but high and broad last whorl which is also strongly descending near the aperture (almost 90 ° angle). The parietal side of the aperture is with only a single, parietal lamella while the angular is absent. Hypselostoma srakeoense, H. torticollis, and H. fungus sp. nov. are superficially similar but none of them are flat and the latter two are also spirally striated. Hypselostoma torticollis additionally has separated parietal and angular lamellae on the parietal side.</p><p>Distribution.</p><p>This species is known only from the type locality.</p></div>	https://treatment.plazi.org/id/70BC59E38C9A575A9A6FEC2897365365	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
A03D76F2657950ABBD54A01D5609C6CF.text	A03D76F2657950ABBD54A01D5609C6CF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma phupaman (Panha & J. B. Burch 2002) Gojšina & Hunyadi & Sutcharit & Tongkerd & Auffenberg & Grego & Vermeulen & Reischütz & Páll-Gergely 2025	<div><p>Hypselostoma phupaman (Panha &amp; J. B. Burch, 2002) comb. nov.</p><p>Figs 242 G, 256, 259</p><p>Paraboysidia phupaman Panha &amp; Burch, 2002 c: 90, fig. 6.</p><p>Gyliotrachela phupaman — Panha and Burch 2008: 77–78, fig. 67; Dumrongrojwattana and Wongkamhaeng 2024: 324, fig. 8.</p><p>Type material examined.</p><p>Thailand • 1 paratype; from the type locality; 1998; S. Panha, P. Dumrongrojwattana, C. Sutcharit, S. Tumpeesuwan leg.; CUMZ ver. 086 .</p><p>Type locality.</p><p>“ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.904724&amp;materialsCitation.latitude=16.664444" title="Search Plazi for locations around (long 101.904724/lat 16.664444)">Phupaman mountains</a>, Petchaboon Province, Thailand, 16°39′52″N, 101°54′17″E, 60 meters elevation ”.</p><p>Differential diagnosis.</p><p>This species is not similar to any of its congeners due to the unique, double-keeled last whorl.</p><p>Distribution.</p><p>This species is known only from the type locality.</p></div>	https://treatment.plazi.org/id/A03D76F2657950ABBD54A01D5609C6CF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
6BC13778D39F59229375461E690CAAB2.text	6BC13778D39F59229375461E690CAAB2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma piconis van Benthem Jutting 1949	<div><p>Hypselostoma piconis van Benthem Jutting, 1949</p><p>Figs 186 K, 208, 223</p><p>Hypselostoma piconis van Benthem Jutting, 1949 b: 59, pl. 2., figs a – d.</p><p>Hypselostoma piconis — van Benthem Jutting 1950: 24, 42, fig. 13; Maassen 2001: 77.</p><p>Type material examined.</p><p>Malaysia • 1 paratype; from the type locality; Raffles Museum, Singapore ex. coll.; RMNH.Moll.140161 .</p><p>Additional material examined.</p><p>Malaysia • 9 shells; Perak, 8–10 km northeast from Sungai Siput, east from <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.122215&amp;materialsCitation.latitude=4.866417" title="Search Plazi for locations around (long 101.122215/lat 4.866417)">Batu Lima</a>, cave temple; 04°51.985'N, 101°07.333'E; 70 m a. s. l.; 07 Jan. 2013; A. Hunyadi leg.; coll. HA .</p><p>Type locality.</p><p>“ Sungei Siput, Perak ”, Malaysia.</p><p>Differential diagnosis.</p><p>Hypselostoma piconis is strikingly similar to H. luctans from which it can be separated by the appearance of the apertural barriers, most importantly the concrescent angular and parietal lamellae, which are separate in H. luctans .</p><p>Distribution.</p><p>This species is known only from the type locality and its surroundings.</p><p>Remarks.</p><p>The fact that the major difference between H. piconis and H. luctans is the appearance of the barriers on the parietal side further supports the indistinctness of Gyliotrachela and its synonymy with Hypselostoma .</p></div>	https://treatment.plazi.org/id/6BC13778D39F59229375461E690CAAB2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
DF28882580A65621A850B69ACDD03977.text	DF28882580A65621A850B69ACDD03977.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma platybasis Gojsina, Hunyadi & Pall-Gergely 2025	<div><p>Hypselostoma platybasis Gojšina, Hunyadi &amp; Páll-Gergely sp. nov.</p><p>Type material.</p><p>Holotype. Cambodia • 1 shell (SH: 2.60 mm; SW: 2.56 mm); Steung Treng Province, 55.7 km northwest + 2 km north from Stung Treng Mekong Bridge, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.60342&amp;materialsCitation.latitude=13.797017" title="Search Plazi for locations around (long 105.60342/lat 13.797017)">Chap Phleung Mt.</a> (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.60342&amp;materialsCitation.latitude=13.797017" title="Search Plazi for locations around (long 105.60342/lat 13.797017)">Neak Khiev Mt.</a>); 13°47.821'N, 105°36.205'E; 135 m a. s. l.; 26 Oct. 2023, A. Hunyadi &amp; J. U. Otani leg.; CUMZ 14457 . Paratypes. Cambodia • 160 shells; same data as for holotype; coll. HA • 1 shell; same data as for holotype; coll. VG .</p><p>Additional material examined.</p><p>Cambodia • 10 shells (juveniles, not paratypes); same data as holotype; coll. HA .</p><p>Type locality.</p><p>Steung Treng Province, 55.7 km northwest + 2 km north from Stung Treng Mekong Bridge, Chap Phleung Mt. (Neak Khiev Mt.); 13°47.821'N, 105°36.205'E; 135 m a. s. l.</p><p>Diagnosis.</p><p>Hypselostoma species with bluntly keeled penultimate (at the centre of the periphery) and shouldered last whorl. Teleoconch densely spirally striated. Last whorl very slightly detached from the penultimate and with a sharp shoulder. Apertural barriers few (angulo-parietal (parietal), upper palatal, lower palatal and columellar) and relatively weak. Umbilicus initially narrow and then abruptly widening at the last whorl.</p><p>Description.</p><p>Shell conical or slightly concave-conical, dark brown, consisting of 4–4.5 whorls separated by a deep suture. Protoconch rounded, slightly glossy, roughly pitted, consisting of ~ 1.5 whorls. Initial teleoconch whorls rounded, penultimate whorl bluntly keeled. Teleoconch sculptured with strong, raised spiral striae which are crossed by weaker radial growth lines usually in form of whitish radial streaks. There are ~ 30 spiral striae on the last whorl in standard view. The spacing between two striae ranges to ~ 2–3 × the width of one stria. Basis of the last whorl flat. Last whorl with a sharp shoulder (almost like a keel), adnate or very slightly detached from the penultimate, very slightly descending near the aperture (less than 5 ° compared to the shell axis), almost straight. Peristome brown, not very strong but still expanded and not reflected. Aperture equipped with four main barriers (parietal (angulo-parietal), upper palatal, lower palatal and columellar), none of which reach the expanding peristome. Angulo-parietal lamella is the strongest, longest and the highest in the aperture. There is usually no sign of the angular part. Upper and lower palatal plicae both low and relatively weak, almost equally strong. Basal plica clearly present and moderately strong or absent (with only a slight swelling in its place). Columellar lamella developed to a similar extent as both palatal plicae. There is one additional swelling in the columello-parietal embayment which corresponds to the infraparietal lamella. Surface of all apertural barriers is finely granulated. Sinulus wide and not distinctly separated from the rest of the aperture. Umbilicus initially narrow, but abruptly widening at the last whorl, becoming very wide, and measuring slightly&gt; ½ of the shell width. There is a deep groove running from the peristome edge towards the umbilicus along the inner side of the last whorl. This groove is terminating at the transition from the last whorl to penultimate.</p><p>Differential diagnosis.</p><p>This species is the most similar to H. geckophilum sp. nov. from which it differs in the more rectangular shape of the last whorl and a stronger shoulder. Umbilicus is also much narrower in H. geckophilum sp. nov. and the barriers are more slender.</p><p>Measurements</p><p>(in mm, n = 5). SH = 2.36–2.75; SW = 2.56–2.85; AH = 1.20–1.32; AW = 1.25–1.47.</p><p>Etymology.</p><p>This species is named for the flat base of the shell.</p><p>Distribution.</p><p>This species is only known from the type locality.</p><p>Remarks.</p><p>The largest, and still unexplored limestone hill in Steung Treng Province (13°47.941'N, 105°43.7671'E) was inaccessible during the collecting efforts in 2023 due to the works of a cement factory. Due to the proximity of this hill to the type locality of H. platybasis sp. nov., it is possible that this species can be found here as well and be threatened by quarrying.</p></div>	https://treatment.plazi.org/id/DF28882580A65621A850B69ACDD03977	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
8506BD4320BD50BDB23729928BE771F5.text	8506BD4320BD50BDB23729928BE771F5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma populare Gojsina, Hunyadi & Pall-Gergely 2025	<div><p>Hypselostoma populare Gojšina, Hunyadi &amp; Páll-Gergely sp. nov.</p><p>Type material.</p><p>Holotype. Thailand • 1 shell (SH: 1.4 mm; SW: 2.2 mm); Krabi Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.870514&amp;materialsCitation.latitude=8.12175" title="Search Plazi for locations around (long 98.870514/lat 8.12175)">9.2 km northwest from Krabi towards Phang Nga, right side of the road</a>; 08°07.305'N, 98°52.231'E; 40 m a. s. l.; 21 Feb. 2015, A. Hunyadi leg.; CUMZ 14455 . Paratypes. Thailand • 5 shells; same data as for holotype; coll. HA • 1 shell; Chumphon Province, Pathio, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.49838&amp;materialsCitation.latitude=10.905467" title="Search Plazi for locations around (long 99.49838/lat 10.905467)">500 m from the junction Ban Tham Tong - Thung Yang Beach</a>, large rock; 10°54.328'N, 99°29.903'E; 20 m a. s. l.; 23 Feb. 2015; A. Hunyadi leg.; coll. HA • 10 shells; Ranong Province, south of Kra Buri, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.76555&amp;materialsCitation.latitude=10.324667" title="Search Plazi for locations around (long 98.76555/lat 10.324667)">Tham Phra Kayang</a>, southeastern part of the rock, 10°19.480'N, 98°45.933'E; 22 Feb. 2015; A. Hunyadi leg.; coll. HA • 17 shells; Chumphon Province, 2.5 km northeast from Pathio, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.3207&amp;materialsCitation.latitude=10.73085" title="Search Plazi for locations around (long 99.3207/lat 10.73085)">Tham Khao Phlu</a>, 10°43.851'N, 99°19.242'E; 30 m a. s. l.; 23 Feb. 2015; A. Hunyadi leg.; coll. HA • 2 shells; Krabi Province, Railay (= Rai Leh) <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.83698&amp;materialsCitation.latitude=8.014417" title="Search Plazi for locations around (long 98.83698/lat 8.014417)">Beach West</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.83698&amp;materialsCitation.latitude=8.014417" title="Search Plazi for locations around (long 98.83698/lat 8.014417)">Railay Highlands</a>, at the base of limestone rocks; 8°0.865'N, 98°50.219'E; 30 m a. s. l.; Sep. 2007; A. Reischütz leg.; coll. REI • 3 shells; Krabi Province, N of Krabi, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.92443&amp;materialsCitation.latitude=8.126717" title="Search Plazi for locations around (long 98.92443/lat 8.126717)">Tiger Cave Temple (= Wat Tham Suea), along the ~ 200 steps to the cave</a>, at the base of limestone rocks; 8°7.603'N, 98°55.466'E; 90 m a. s. l.; Sep. 2007; A. Reischütz leg.; coll. REI .</p><p>Additional material examined.</p><p>Thailand • 8 shells (damaged / juveniles, not paratypes); Chumphon Province, 2.5 km northeast from Pathio, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.3207&amp;materialsCitation.latitude=10.73085" title="Search Plazi for locations around (long 99.3207/lat 10.73085)">Tham Khao Phlu</a>, 10°43.851'N, 99°19.242'E; 30 m a. s. l.; 23 Feb. 2015; A. Hunyadi leg.; coll. HA • 5 shells (damaged / juveniles, not paratypes); Ranong Province, south of Kra Buri, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.76555&amp;materialsCitation.latitude=10.324667" title="Search Plazi for locations around (long 98.76555/lat 10.324667)">Tham Phra Kayang</a>, southeastern part of the rock, 10°19.480'N, 98°45.933'E; 22 Feb. 2015; A. Hunyadi leg.; coll. HA .</p><p>Type locality.</p><p>Krabi Province, 9.2 km northwest from Krabi towards Phang Nga, right side of the road; 08°07.305'N, 98°52.231'E; 40 m a. s. l.</p><p>Diagnosis.</p><p>Shell depressed, concave-conical, last whorl strongly detached from the penultimate and ascending at nearly 60–70 ° angle. Teleoconch strongly spirally striated. Aperture equipped with four blunt barriers (angulo-parietal, upper palatal, lower palatal, and columellar). Umbilicus very wide.</p><p>Description.</p><p>Shell depressed, concave-conical (due to the strongly enlarged last whorl), consisting of 3.75–4.25 whorls separated by a very deep suture. Colouration yellowish or very pale brownish, opaque. Protoconch initially almost smooth and then finely pitted, showing spiralling pattern and consisting of 1.25–1.5 whorls. Boundary between the protoconch and teleoconch clearly visible due to the change of microsculpture. Teleoconch strongly and regularly spirally striated, spiral striae thread-like raised. Space between two spiral striae measuring approximately the width of two to four spiral striae. There are ~ 30 spiral striae on the last whorl in standard view. Weaker, irregularly spaced radial growth lines are also visible crossing the spiral striae. Some irregularly spaced radial white streaks are occasionally present. Last whorl shouldered, strongly detached from the penultimate and ascending strongly near the aperture (~ 60–70 ° compared to the shell axis). Peristome lighter than the rest of the shell, fragile, expanded and not reflected. Aperture equipped with four moderately strong barriers (angulo-parietal, upper palatal, lower palatal, and columellar). Angulo-parietal lamella is the strongest in the aperture, consisting of two parts corresponding to angular and parietal lamellae in the former Gyliotrachela . The angular part of the angulo-parietal lamella is very small and pointy, leaning towards the palatal wall. The parietal part of the angulo-parietal lamella is much stronger and positioned deeper in the aperture, leaned towards the columellar lamella. Both palatal plicae and a columellar lamella are developed to similar level, relatively short and blunt. Upper palatal plica nearly fully rectangular in shape, lower palatal plica and columellar lamella more rounded and positioned even deeper in the aperture than the upper palatal plica. The apertural barriers are not very variable. None of these barriers reach the peristome. Surface of all apertural barriers is finely granulated. Sinulus rounded and well separated from the rest of the aperture. Umbilicus open, very wide, measuring ~ 1 / 3 of the shell width and showing all previous whorls.</p><p>Differential diagnosis.</p><p>This species is nearly identical in external appearance to Hypselostoma erawan . However, H. erawan clearly differs in the appearance of the apertural barriers. Hypselostoma erawan has more barriers than the new species and palatal plicae are hooked (neither of the barriers in H. populare sp. nov. are hooked). Acinolaemus colpodon F. G. Thompson &amp; Upatham, 1997 is smaller and also has hooked barriers. Hypselostoma cucumense has the same appearance of the apertural barriers but the last whorl in H. populare sp. nov. is without a keel. The umbilicus is also initially narrow in H. cucumense and then widening at the last whorl, but it is wide even in its initial parts in H. populare sp. nov.</p><p>Measurements</p><p>(in mm, n = 5). SH = 1.33–1.41; SW = 2.13–2.22; AH = 0.79–0.86; AW = 0.7–0.87.</p><p>Etymology.</p><p>Named after the fact that many sampling sites of this species are popular touristic destinations.</p><p>Distribution.</p><p>This species is known from Krabi, Chumphon and Ranong provinces in Thailand.</p><p>Remarks.</p><p>This species is not very variable in terms of shell size and apertural dentition but in some specimens the spire is more depressed conical than in others.</p></div>	https://treatment.plazi.org/id/8506BD4320BD50BDB23729928BE771F5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
AF8B9E6C07B65A5A9511FBBE4CECAEAF.text	AF8B9E6C07B65A5A9511FBBE4CECAEAF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma procerum (F. G. Thompson & Dance 1983) Gojšina & Hunyadi & Sutcharit & Tongkerd & Auffenberg & Grego & Vermeulen & Reischütz & Páll-Gergely 2025	<div><p>Hypselostoma procerum (F. G. Thompson &amp; Dance, 1983) comb. nov.</p><p>Figs 224 G, 233, 239</p><p>Boysidia (Dasypupa) procera Thompson &amp; Dance, 1983: 108–109, figs 11, 12.</p><p>Type material examined.</p><p>Malaysia • holotype; UF 38022 • 3 paratypes; from the type locality; UF 38023 .</p><p>Type locality.</p><p>“ Southwest flank of Gunong Budah, Medalam Valley, a tributary of the Limbang river, Fifth Division, Sarawak, Borneo 04°08'N, 115°5'E ”.</p><p>Differential diagnosis.</p><p>This species is most similar to H. paini which is also described from Borneo. However, this species is more elongated and has a wider umbilicus. See also under H. tertiusfrater nom. nov.</p><p>Distribution.</p><p>This species is known only from the type locality.</p></div>	https://treatment.plazi.org/id/AF8B9E6C07B65A5A9511FBBE4CECAEAF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
DC1C4FFC49A95C5F84B051C6EBCE57D1.text	DC1C4FFC49A95C5F84B051C6EBCE57D1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma rupestre van Benthem Jutting 1962	<div><p>Hypselostoma rupestre van Benthem Jutting, 1962</p><p>Hypselostoma rupestre van Benthem Jutting, 1962: 6–8, fig. 3.</p><p>Hypselostoma rupestre — Schileyko 2011: 3–4; Vermeulen et al. 2019: 40.</p><p>Material examined.</p><p>Cambodia • 32 shells; Kampot Province, Kampong Trach district, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.48037&amp;materialsCitation.latitude=10.438916" title="Search Plazi for locations around (long 104.48037/lat 10.438916)">Phnom Teuk Thom</a> (northwest from Wat Aranhatry); 10°26.335'N, 104°28.822'E; 400 m a. s. l.; 21 Oct. 2023; A. Hunyadi, T. Ishibe, K. Okubo, J. U. Otani leg.; coll. HA .</p><p>Type locality.</p><p>“ Nui Hon Chong (Cap de la Table), à 32 km au S. - E. de Hatien, Sud Vietnam ”.</p><p>Differential diagnosis.</p><p>This species differs from H. cambodjense by its rounded whorls. It is thus superficially more similar to H. dilatatum but the latter has several hooked apertural barriers (they are all blunt in H. rupestre).</p><p>Distribution.</p><p>This species in known from several localities in Kien Giang province (Vietnam). We have also found this species in Kampot province, Cambodia (see under Material examined).</p></div>	https://treatment.plazi.org/id/DC1C4FFC49A95C5F84B051C6EBCE57D1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
8DB4A8D817915734988534CE3BDB6270.text	8DB4A8D817915734988534CE3BDB6270.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma salpinx (van Benthem Jutting 1961) Gojšina & Hunyadi & Sutcharit & Tongkerd & Auffenberg & Grego & Vermeulen & Reischütz & Páll-Gergely 2025	<div><p>Hypselostoma salpinx (van Benthem Jutting, 1961) comb. nov.</p><p>Gyliotrachela salpinx van Benthem Jutting, 1961: 38–39, pl. 10, fig. 4.</p><p>Gyliotrachela salpinx — Maassen 2001: 76.</p><p>Type material examined.</p><p>Malaysia • 60 paratypes; from the type locality; Aug. 1950; Raffles Museum ex. coll.; RMNH.Moll.137154 .</p><p>Additional material examined.</p><p>Malaysia • 6 shells; Pahang, 10 km northeast from Raub, 1.5 km northwest from Sungai Ruan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.92312&amp;materialsCitation.latitude=3.8302834" title="Search Plazi for locations around (long 101.92312/lat 3.8302834)">vicinity of Bukit Serdam limestone rock</a>; 03°49.817'N, 101°55.387'E; 170 m a. s. l.; 20 Jan. 2013; A. Hunyadi leg.; coll. HA • 6 shells; Pahang, Tok Machang (end of Raub), road Raub-Sungai Ruan; Oct. 1998; ex. coll. Hemmen, Wiesbaden; coll. PGB . Thailand • 4 damaged shells; Thailand, Suratthani Province, Angthong islands, Mae Koh island, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.66627&amp;materialsCitation.latitude=9.657666" title="Search Plazi for locations around (long 99.66627/lat 9.657666)">footpath to the Lagoon</a>, at the base of limestone rocks; 9°39.460'N, 99°39.976'E; 20 m a. s. l.; Sep. 2007; A. Reischütz leg.; coll. REI ( cf. salpinx) • 1 shell; Phattalung Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.183334&amp;materialsCitation.latitude=7.45" title="Search Plazi for locations around (long 100.183334/lat 7.45)">6.4 km E of Hwy. 4081, 1 km SW of Khao Chai Son</a>; 7°27'N, 100°11'E; 11 Apr. 1988; K. Auffenberg leg.; locality code KA-0648; UF 591343 • 1 shell; same locality data as previous; 11 Apr. 1988; K. Auffenberg leg.; locality code KA-0648; UF 591342; 3 shells; Phattalung Province, 6 km NW of Phattalung, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.05&amp;materialsCitation.latitude=7.6666665" title="Search Plazi for locations around (long 100.05/lat 7.6666665)">1 km NE of Hwy. 4048</a>; 7°40'N, 100°3'E; 60 m a. s. l.; 13 Apr. 1988; K. Auffenberg leg.; locality code KA-0653; UF 591345 • 5 shells; Phattalung Province, 6.4 km E of Hwy. 4 on Hwy. 4081, 1 km SW of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.183&amp;materialsCitation.latitude=7.45" title="Search Plazi for locations around (long 100.183/lat 7.45)">Khao Chai Son</a>, large limestone outcrop with solution pits; 7.450°N, 100.183°E; 50 m a. s. l.; 11 Apr. 1988; K. Auffenberg leg.; locality code KA-0649; UF 591344 • 1 shell; Phattalung Province, Khao Pu Khao Ya National Park, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.87238&amp;materialsCitation.latitude=7.666333" title="Search Plazi for locations around (long 99.87238/lat 7.666333)">Matcha cave vicinity</a>; 7°39.980'N, 99°52.343'E; 125 m a. s. l.; 25 Feb. 2023; A. Hunyadi leg.; coll. HA • 35 shells; Songkhla Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.26667&amp;materialsCitation.latitude=7.1666665" title="Search Plazi for locations around (long 100.26667/lat 7.1666665)">31.3 km NW of Hat Yai, 1.2 km W of Hwy. 43</a>; 7°10'N, 100°16'E; 80 m a. s. l.; 08 Apr. 1988; K. Auffenberg leg.; locality code KA-0635; UF 344926 • 5 shells; same locality data as previous; 08 Apr. 1988; K. Auffenberg leg.; locality code KA-0636; UF 591341 .</p><p>Type locality.</p><p>“ Bukit Serdam, near Raub, Pahang ” (Malaysia).</p><p>Differential diagnosis.</p><p>See under H. saxicola and H. coriaceum sp. nov.</p><p>Distribution.</p><p>This species is known only from the limited area (Raub) in Malaysia and Suratthani, Phattalung and Songkhla provinces in Thailand.</p></div>	https://treatment.plazi.org/id/8DB4A8D817915734988534CE3BDB6270	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
FBA32FF5A19C5B32A91D752F4B11A18A.text	FBA32FF5A19C5B32A91D752F4B11A18A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma saxicola (van Benthem Jutting 1960) Gojšina & Hunyadi & Sutcharit & Tongkerd & Auffenberg & Grego & Vermeulen & Reischütz & Páll-Gergely 2025	<div><p>Hypselostoma saxicola (van Benthem Jutting, 1960) comb. nov.</p><p>Gyliotrachela saxicola van Benthem Jutting, 1960: 14–16, fig. 1.</p><p>Gyliotrachela saxicola — Maassen 2001: 76.</p><p>Type material examined.</p><p>Malaysia • 22 paratypes; Perlis, N of Kangar, limestone hill near Kampong Tebing Tinggi, on the face of the rock; 19 Dec. 1958; W. S. S. van der Feen-van Benthem Jutting leg.; RMNH.Moll.137139 .</p><p>Type locality.</p><p>“ Limestone hill near Kampong Tebing Tinggi, N. of Kangar, Perlis ”, Malaysia.</p><p>Differential diagnosis.</p><p>This species is most similar to H. salpinx from which it can be separated by smaller number of overall stronger barriers and wider umbilicus (which is initially narrower in H. salpinx). See also under H. coriaceum sp. nov.</p><p>Distribution.</p><p>This species is known only from the type locality.</p><p>Remarks.</p><p>In paratypes examined, besides the five barriers that are always present, there were additional 1–3 smaller. These are interpalatal, infrapalatal and infraparietal and they were present in some and absent in other specimens. As a consequence, the number of barriers varied from 6–8 (as there was always at least one of the smaller). This was also noted by van Benthem Jutting (1960) in the original description. She also noted that the last whorl can be more or less detached or completely adnate, but this variation is not useful for separation of the species.</p></div>	https://treatment.plazi.org/id/FBA32FF5A19C5B32A91D752F4B11A18A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
3570D9ADBB175A1DB6E8A3D70AE6E5B5.text	3570D9ADBB175A1DB6E8A3D70AE6E5B5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma sculpturatum Gojsina, Hunyadi & Pall-Gergely 2025	<div><p>Hypselostoma sculpturatum Gojšina, Hunyadi &amp; Páll-Gergely sp. nov.</p><p>Type material.</p><p>Holotype. Cambodia • 1 shell (SH: 2.70 mm; SW 1: 3.80 mm); Steung Treng Province, 36.3 km northwest + 5 km north from Stung Treng Mekong Bridge, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.74796&amp;materialsCitation.latitude=13.7762165" title="Search Plazi for locations around (long 105.74796/lat 13.7762165)">Phnom Chhnok</a>; 13°46.573'N, 105°44.878'E; 120 m a. s. l.; 25 Oct. 2023; A. Hunyadi &amp; J. U. Otani leg.; CUMZ 14458.</p><p>Paratypes. Cambodia • 51 shells; same data as for holotype; coll. HA • 1 shell; same data as for holotype; coll. VG .</p><p>Additional material examined.</p><p>Cambodia • 4 shells (juveniles, not paratypes); same data as holotype; coll. HA .</p><p>Type locality.</p><p>Cambodia, Steung Treng Province, 36.3 km northwest + 5 km north from Stung Treng Mekong Bridge, Phnom Chhnok; 13°46.573'N, 105°44.878'E; 120 m a. s. l.</p><p>Diagnosis.</p><p>Hypselostoma species with all teleoconch whorls keeled, detached last whorl and strongly spirally striated shell surface. There are five strong barriers in the aperture (angulo-parietal, upper palatal, lower palatal, basal and columellar). Umbilicus initially narrow, suddenly widening at the last whorl.</p><p>Description.</p><p>Shell conical-ovoid, brownish, not glossy. It is consisting of 4–4.5 whorls separated by a deep suture. Protoconch rounded, of ~ 1.75 finely pitted whorls of the same colour as the rest of the shell. Teleoconch sculptured with strong, raised spiral striae. Spacing between two striae equals the width of one stria or is twice as wide as a stria. Spiral striae are more densely arranged on the initial teleoconch whorls, and more coarsely on the last whorl. The number of striae on the last whorl is ~ 25 in standard view. There are several white, unevenly positioned, radial streaks crossing the spiral striae. All teleoconch whorls are keeled, and the strength of the keel is increasing towards the last whorl which is especially strongly keeled. Surfaces of the whorls above and below the centrally positioned keel are flat, to very slightly concave (above the keel on the last whorl). Last whorl moderately detached from the penultimate, distanced from the shell axis. From the lateral view (Fig. 168 B), it can be seen that the last whorl is initially slightly ascending and then straightening right near the aperture so that the aperture profile is almost parallel to the shell axis (leaving the aperture only slightly ascending at ~ 5–10 ° compared to the shell axis). Peristome light brownish, expanded and not reflected. Aperture equipped with five barriers (angulo-parietal, upper palatal, lower palatal, basal, and columellar). Angulo-parietal strong and high, lamella usually appears as a single barrier (parietal) but has a very short swollen part in front of it which most probably represents the remains of the angular part. Upper palatal plica moderately high, slightly curved towards the lower palatal plica (somehow similar to the upper palatal plica in H. serpa). Lower palatal plica slightly higher, longer, and more slender than the upper palatal but similar in width. It is pointing towards the embayment between the columellar and parietal lamellae. Basal plica more closely positioned to the lower palatal (not in its usual position between the lower palatal and columellar), ~ 2 × lower than the latter and short. Columellar lamella thick and broad, more closely resembling a very large swelling than a distinct barrier. There is also a small swelling between the columellar and parietal lamellae. Surface of all apertural barriers is finely granulated. Sinulus narrow, distinctly separated from the rest of the aperture. Umbilicus initially very narrow and suddenly widening at the last whorl. It is measuring between ½ and ¹ ⁄ 3 of the shell width. There is a deep groove running from the peristome edge towards the umbilicus along the inner side of the last whorl. This groove is terminating at the transition from the last whorl to penultimate.</p><p>Differential diagnosis.</p><p>This species superficially resembles H. circumcarinatum sp. nov. but the latter has its last whorl adnate from the penultimate, weaker barriers, narrower umbilicus, and less prominent shell sculpture. The new species is distinguished from H. khmerianum by the concrescent angular and parietal lamella as well as by the presence of spiral striation.</p><p>See under H. taehwani .</p><p>Measurements</p><p>(in mm, n = 5). SH = 2.35–2.70; SW 1 = 3.52–3.80; SW 2 = 2.08–2.18; AH = 1.32–1.62; AW = 1.40–1.79.</p><p>Etymology.</p><p>This species is named for the strong surface sculpture of its shell.</p><p>Distribution.</p><p>This species is only known from the type locality.</p><p>Remarks.</p><p>The largest, and still unexplored limestone hill in Steung Treng Province (13°47.941'N, 105°43.7671'E) was inaccessible during the collecting efforts in 2023 due to the works of a cement factory. Due to the proximity of this hill to the type locality of H. sculpturatum sp. nov., it is possible that this species can be found here as well and be threatened by quarrying.</p></div>	https://treatment.plazi.org/id/3570D9ADBB175A1DB6E8A3D70AE6E5B5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
7ED1CB55E4BB592083FE443726198F80.text	7ED1CB55E4BB592083FE443726198F80.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma serpa (van Benthem Jutting 1950) Gojšina & Hunyadi & Sutcharit & Tongkerd & Auffenberg & Grego & Vermeulen & Reischütz & Páll-Gergely 2025	<div><p>Hypselostoma serpa (van Benthem Jutting, 1950) comb. nov.</p><p>Figs 224 H, 234, 241</p><p>Paraboysidia serpa van Benthem Jutting, 1950: 18–19, 40, fig. 8.</p><p>Paraboysidia serpa — Maassen 2001: 75.</p><p>Paraboysidia tarutao Panha &amp; Burch, 2002 c: 85–87, fig. 4. syn. nov.</p><p>Gyliotrachela tarutoa [sic] — Panha and Burch 2008: 84, fig. 73.</p><p>Gyliotrachela tarutao — Dumrongrojwattana and Wongkamhaeng 2024: 324, fig. 8.</p><p>Type material examined.</p><p>Malaysia • 18 paratypes of H. serpa; from the type locality; Dec. 1938; Raffles Museum, Singapore ex. coll.; RMNH.Moll.140164 . Thailand • 1 paratype of G. tarutao; from the type locality; 1998; S. Panha leg.; CUMZ ver. 062 .</p><p>Additional material examined.</p><p>Thailand • 22 shells; Krabi Province, Railay (= Rai Leh) Beach West, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.83698&amp;materialsCitation.latitude=8.014417" title="Search Plazi for locations around (long 98.83698/lat 8.014417)">Railay Highlands</a>, at the base of limestone rocks; 8°0.865'N, 98°50.219'E; 30 m a. s. l.; Sep. 2007; A. Reischütz leg.; coll. REI • 1 shell; Krabi Province, Phi Phi islands, Phi Phi Don island, climbing rock above the western end of the beach in <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.765785&amp;materialsCitation.latitude=7.7343836" title="Search Plazi for locations around (long 98.765785/lat 7.7343836)">Ton Sai Bay</a>, at the base of limestone rocks in the forest; 7°44.063'N, 98°45.947'E; 50 m a. s. l.; Mar. 2010; A. Reischütz leg; coll. REI .</p><p>Type localities.</p><p>“ Baling, Kedah ”, Malaysia ( H. serpa); “ Tarutoa National Park, Satul Province, Thailand, 6 ° 41 ' 58 ' N, 99 ° 38 ' 48 ' E, 70 meters elevation ” ( P. tarutao).</p><p>Differential diagnosis.</p><p>Hypselostoma serpa is most similar to H. frequens, from which it can be separated by the keeled last whorl (rounded in H. frequens), the bent upper palatal plica and the wider umbilicus. Hypselostoma kelantanense has a more ovoid shell shape, more rounded and bulging whorls as well as narrower umbilicus.</p><p>Distribution.</p><p>This species is known from Kedah, Malaysia (the type locality) as well as from Krabi Province and Ko Tarutao Island, Satun province, Thailand (the type locality of G. tarutao).</p><p>Remarks.</p><p>In the paratypes, interpalatal plica is simple or bulge like, infraparietal lamella can be present or absent. All examined paratypes have upper palatal plica curved towards the lower one which could be characteristic for this species. Paraboysidia tarutao is treated as a junior synonym of this species since no morphological differences were noticed. In the original description (Panha and Burch 2002 c), P. tarutao was not compared to H. serpa which gives the impression that the latter was overlooked.</p></div>	https://treatment.plazi.org/id/7ED1CB55E4BB592083FE443726198F80	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
95DC87B3CABE5743B611F5C3C4BD9EF8.text	95DC87B3CABE5743B611F5C3C4BD9EF8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma sichang (Panha & J. B. Burch 2002) Gojšina & Hunyadi & Sutcharit & Tongkerd & Auffenberg & Grego & Vermeulen & Reischütz & Páll-Gergely 2025	<div><p>Hypselostoma sichang (Panha &amp; J. B. Burch, 2002) comb. nov.</p><p>Figs 186 L, 209, 210, 223</p><p>Anauchen sichang Panha &amp; Burch, 2002 e: 21, fig. 2.</p><p>Gyliotrachela sichang — Panha and Burch 2008: 80, fig. 69; Dumrongrojwattana and Wongkamhaeng 2024: 324, fig. 8.</p><p>Type material examined.</p><p>Thailand • holotype; 2001; S. Panha leg.; CUMZ 44001 • 2 paratypes; from the type locality; 2001; S. Panha leg.; SMF 331463 .</p><p>Additional material examined.</p><p>Thailand • 10 shells; Sa Kaeo Province, Wang Sombun district, vicinity of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.21785&amp;materialsCitation.latitude=13.447884" title="Search Plazi for locations around (long 102.21785/lat 13.447884)">Tham Khoa Phlapphueng Thong</a>; 13°26.873'N, 102°13.071'E; 06 Mar. 2023; A. Hunyadi leg.; coll. HA .</p><p>Type locality.</p><p>“ Sichang island, Si Racha, Chon Buri Province, 13°20′05″N, 100°55′24″E, 70 meters elevation ” (Thailand).</p><p>Differential diagnosis.</p><p>This species is similar to H. hungerfordianum from which it can be separated by the much stronger and lower peripheral keel, very rough, sandpaper-like shell surface sculpture as well as the narrower umbilicus. See also under H. vesovici sp. nov.</p><p>Distribution.</p><p>This species is, apart from the type locality, known from one more sampling site in Sa Kaeo Province.</p></div>	https://treatment.plazi.org/id/95DC87B3CABE5743B611F5C3C4BD9EF8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
7F69B785C94B5EDF9CC905798722236A.text	7F69B785C94B5EDF9CC905798722236A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma similare Gojsina, Hunyadi & Pall-Gergely 2025	<div><p>Hypselostoma similare Gojšina, Hunyadi &amp; Páll-Gergely sp. nov.</p><p>Figs 242 H, 257, 258, 259</p><p>Type material.</p><p>Holotype. Thailand • 1 shell (SH: 1.83 mm; SW 1: 2.51 mm); Rayong Province, Khao Chamao district, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.5975&amp;materialsCitation.latitude=13.123983" title="Search Plazi for locations around (long 101.5975/lat 13.123983)">Wat Tham Khao Prathun</a>, gorge above the temple; 13°07.439'N, 101°35.850'E; 115 m a. s. l.; 09 Mar. 2023, A. Hunyadi leg.; CUMZ 14470 . Paratypes. Thailand • 2 shells; same data as for holotype; coll. HA .</p><p>Type locality.</p><p>Thailand, Rayong Province, Khao Chamao district, Wat Tham Khao Prathun, gorge above the temple; 13°07.439'N, 101°35.850'E; 115 m a. s. l.</p><p>Diagnosis.</p><p>Shell triangular conical. Teleoconch almost smooth, finely radially but not spirally striated. Last whorl detached from the penultimate and ascending, keeled below the centre of the periphery. Aperture equipped with weak barriers, five main (angular, parietal, upper palatal, lower palatal and columellar) and several smaller ones. Umbilicus narrow, measuring 1 / 7–1 / 8 of the shell width.</p><p>Description.</p><p>Shell conical, light brownish, opaque, consisting of 3.75–4.25 weakly convex and regularly increasing whorls separated by a relatively shallow suture. Protoconch finely pitted, without spiral striae, consisting of ~ 1.5 whorls, same colour as the rest of the shell. Teleoconch sculpture delicate, very finely dimpled (pasty) and crossed by relatively weak radial growth lines. Spiral striae absent. Occasionally, a few stronger whitish radial streaks can be observed at the last whorl near the aperture. Last whorl weakly detached from the penultimate and slightly ascending near the aperture (~ 15–20 ° compared to the shell axis). It is bluntly but prominently keeled below the periphery, making the last whorl oblique to the shell axis and not convex but with a sloping, flat outline. There is a slight furrow above the keel. Peristome thick, whitish, expanded and not reflected. All apertural barriers are weak. Aperture equipped with five main barriers (angular, parietal, upper palatal, lower palatal, and columellar) and several smaller ones of variable number and appearance. There are usually no plicae in the sinulus, usually one plica in the interpalatal region, two or three plicae in the region of the basal plica and usually one or two lamellae in the columello-parietal region. Angular lamella tubercle-like, almost reaching the peristome. Behind it, more deeply situated, there is one additional part which also belongs to the angular lamella. Parietal lamella stronger and higher when compared to the others (except lower palatal plica), not reaching the profile of the angular tubercle. Lower palatal plica developed to the same extent as the parietal lamella and slightly higher and stronger than the upper palatal plica. Columellar lamella horizontal, developed equally as the upper palatal plica. Anterior to the upper palatal plica, there is one more tubercle-like part, possibly homologous with the palatal tubercle found in the genus Bensonella (see Páll-Gergely and White 2023) or it even might represent a part of the upper palatal plica. Weaker barriers are all positioned deeper in the aperture and never reaching the profiles of the main ones. A tubercle-like swelling is present in the columellar-parietal transition embayment. All barriers are strongly and coarsely spiniferous. Sinulus small, rounded and distinctly separated from the rest of the aperture. Umbilicus narrow, with almost perpendicular walls, showing only a part of the penultimate whorl and measuring ~ 1 / 7–1 / 8 of the shell width.</p><p>Differential diagnosis.</p><p>By its last whorl keeled at its base, this species in superficially most similar to H. insularum and Hypselostoma species inhabiting the Philippines. However, this species most clearly differs from these representatives by the presence of two lamellae on the parietal side (angular and parietal) and more numerous barriers in the aperture including smaller ones. Hypselostoma insularum further has strong spiral striae which are absent in the new species.</p><p>Etymology.</p><p>Named after the fact that this species is in shell shape very similar to some Hypselostoma species inhabiting the Philippines.</p><p>Measurements</p><p>(in mm, n = 3). SH = 1.82–1.91; SW 1 = 2.34–2.51; SW 2 = 1.45–1.51; AH = 0.98–1.01; AW = 0.9–0.98.</p><p>Distribution.</p><p>This species is known only from the type locality.</p><p>Remarks.</p><p>All barriers in the aperture very weak, smaller ones are hardly observable, only in form of very weak lines. Single interpalatal plica present or absent. Sinulus plicae present or absent.</p></div>	https://treatment.plazi.org/id/7F69B785C94B5EDF9CC905798722236A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
968874338F085E538FEF1899AD1850DA.text	968874338F085E538FEF1899AD1850DA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma smokon (Panha & J. B. Burch 2004)	<div><p>Hypselostoma smokon (Panha &amp; J. B. Burch, 2004)</p><p>Anauchen smokon Panha &amp; Burch in Panha et al. 2004: 60, fig. 3.</p><p>Hypselostoma smokon — Panha and Burch 2008: 99–100, fig. 85; Dumrongrojwattana and Wongkamhaeng 2024: 324, fig. 8.</p><p>Type material examined.</p><p>Thailand • 2 paratypes; from the type locality; S. Panha leg.; SMF 331457 .</p><p>Type locality.</p><p>“ Smokon Mountain, Banmi District, Lopburi Province, 14°54'54"N, 100°29'45"E, 30 meters elevation ”.</p><p>Differential diagnosis.</p><p>This species is similar in shell shape to H. chedi and H. khaochakan . From them, H. smokon clearly differs by the concrescent angular and parietal lamellae. It is most similar to H. chedi which has its last whorl bluntly keeled (shouldered in H. smokon) and narrower umbilicus.</p><p>Distribution.</p><p>This species is known only from the type locality.</p><p>Remarks.</p><p>In the original description, the species was placed in the genus Anauchen because of the absent angular lamella. However, it is present as an angular part of the joined angulo-parietal lamella which justifies its placement in the genus Hypselostoma .</p></div>	https://treatment.plazi.org/id/968874338F085E538FEF1899AD1850DA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
F0E153FF8E5E5689ADB24DF76E7EEE3F.text	F0E153FF8E5E5689ADB24DF76E7EEE3F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma sorormajor Gojsina, Hunyadi & Pall-Gergely 2025	<div><p>Hypselostoma sorormajor Gojšina, Hunyadi &amp; Páll-Gergely sp. nov.</p><p>Type material.</p><p>Holotype. Cambodia • 1 shell (SH: 2.70 mm; SW 1: 3.91 mm); Steung Treng Province, 55.7 km northwest + 2 km north from Stung Treng Mekong Bridge, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.60342&amp;materialsCitation.latitude=13.797017" title="Search Plazi for locations around (long 105.60342/lat 13.797017)">Chap Phleung Mt.</a> (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.60342&amp;materialsCitation.latitude=13.797017" title="Search Plazi for locations around (long 105.60342/lat 13.797017)">Neak Khiev Mt.</a>); 13°47.821'N, 105°36.205'E; 135 m a. s. l.; 26 Oct. 2023, A. Hunyadi &amp; J. U. Otani leg.; CUMZ 14459 . Paratypes. Cambodia • 28 shells; same data as for holotype; coll. HA .</p><p>Additional material examined.</p><p>Cambodia • 3 shells (2 damaged and 1 juvenile, not paratypes); same data as for holotype; coll. HA • 41 shells; Steung Treng Province, 40 km northwest + 2 km north from Stung Treng <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.7274&amp;materialsCitation.latitude=13.759467" title="Search Plazi for locations around (long 105.7274/lat 13.759467)">Mekong Bridge</a>, limestone mountain at the left side of the joining road; 13°45.568'N, 105°43.644'E; 135 m a. s. l.; 26 Oct. 2023; A. Hunyadi &amp; J. U. Otani leg.; coll. HA • 78 shells; Steung Treng Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.6859&amp;materialsCitation.latitude=13.764433" title="Search Plazi for locations around (long 105.6859/lat 13.764433)">Stung Treng Mekong Bridge</a>, 45 km NWN + 400 m N, limestone hill, right side of the road # 64; 13°45.866'N, 105°41.154'E; 180 m a. s. l.; 26 Oct. 2023; A. Hunyadi &amp; J. U. Otani leg.; coll. HA • 1 shell; same data as previous; coll. VG • 26 shells; Steung Treng Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.74883&amp;materialsCitation.latitude=13.75975" title="Search Plazi for locations around (long 105.74883/lat 13.75975)">36.3 km northwest + 3.1 km north from Stung Treng Mekong Bridge</a>, limestone mountain on the left side of the joining road; 13°45.585'N, 105°44.930'E; 130 m a. s. l.; 25 Oct. 2023; A. Hunyadi &amp; J. U. Otani leg.; coll. HA .</p><p>Type locality.</p><p>Cambodia, Steung Treng Province, 55.7 km northwest + 2 km north from Stung Treng Mekong Bridge, Chap Phleung Mt. (Neak Khiev Mt.); 13°47.821'N, 105°36.205'E; 135 m a. s. l.</p><p>Diagnosis.</p><p>A Hypselostoma species with depressed, concave-conical shell, bluntly and strongly keeled last whorl, raised spiral striation, hooked apertural barriers (upper palatal, interpalatal, lower palatal and basal) and very wide umbilicus. Angular and parietal lamellae fused. Shell width reaches 4 mm.</p><p>Description.</p><p>Shell depressed, concave-conical, brown to yellowish-brown, consisting of 4.75–5.5 whorls separated by a deep suture. Protoconch slightly lighter than the rest of the shell, finely pitted, showing spiralling pattern initially and clearer spiral striae terminally. It is consisting of ~ 1.25–1.5 whorls. Protoconch and initial teleoconch whorls rounded, penultimate weakly convex. Last whorl with strong but blunt keel positioned at the centre of the periphery. Above the keel there is a deep or rarely shallow groove. All whorls densely sculptured with strong, raised spiral striae crossed by less dense radial growth lines and occasionally by a few thick, whitish radial streaks. Spacing between the spiral striae irregular, but usually around the width of two to four spiral striae. Last whorl moderately detached from the penultimate and moderately ascending (~ 35 ° compared to the shell axis). Peristome expanded, not reflected, its surface finely pitted. Aperture equipped with four main barriers (angulo-parietal, upper palatal, lower palatal, and columellar) and several smaller ones. Angulo-parietal lamella strong, its angular part is leaned towards the palatal side while the parietal part is leaned towards the columellar side. Angular part is pointed and smaller than the parietal. It is the only barrier reaching the expanding peristome. Sometimes, there is a deep constriction between the angular and parietal part so that these lamellae appear almost separated (as in former Gyliotrachela). Upper and lower palatal plicae are hooked, pointing outside and roughly the same size. Columellar lamella thick and almost horizontal. Of smaller barriers, there is usually one hooked interpalatal, one infrapalatal, one basal plica (hooked or blunt) and one blunt lamella in the columello-parietal transition embayment. Of them, the strongest one is the basal, which is almost as strong as the palatals. A distinct swelling is also observed at the palatal side (in front of the upper palatal and interpalatal plicae) which is probably homologous to the palatal tubercle in the majority of Bensonella species. Surface of all apertural barriers is finely granulated. Sinulus rounded, distinctly separated from the rest of the aperture. Umbilicus very wide, measuring ~ ½ of the shell width. There is a periumbilical keel situated on the last ~ 0.75 whorl right above the umbilicus. Along this keel there is a deep groove on the umbilical side.</p><p>Differential diagnosis.</p><p>This species differs from H. discobasis in the less slender shell and hooked apertural barriers. Hypselostoma aquila sp. nov. is much smaller, more slender and has a narrower umbilicus. For differences from H. sororminor sp. nov. see under that species.</p><p>Measurements</p><p>(in mm, n = 15). SH = 1.62–2.78; SW 1 = 2.44–4.01; SW 2 = 1.57–2.56; AH = 0.85–1.71; AW = 0.81–1.46.</p><p>Etymology.</p><p>The name of this species is derived from two Latin words, soror meaning sister and major meaning greater, larger, or elder, which serves as a comparison with similar but smaller H. sororminor sp. nov.</p><p>Distribution.</p><p>This species is known from four closely situated localities in Steung Treng Province.</p><p>Remarks.</p><p>This is a very variable species in terms of shell size. These different “ forms ” were not regarded as distinct species since there were no other significant morphological differences noticed. The angulo-parietal lamellae were sometimes clearly fused, while sometimes almost completely separated. The largest, and still unexplored limestone hill in Steung Treng Province (13°47.941'N, 105°43.7671'E) was inaccessible during the collecting efforts in 2023 due to the works of a cement factory. Due to the proximity of this hill to the type locality of H. sorormajor sp. nov., it is possible that this species can be found here as well and be threatened by quarrying.</p></div>	https://treatment.plazi.org/id/F0E153FF8E5E5689ADB24DF76E7EEE3F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
B902884D17DF528C95464B957F1870A0.text	B902884D17DF528C95464B957F1870A0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma sororminor Gojsina, Hunyadi & Pall-Gergely 2025	<div><p>Hypselostoma sororminor Gojšina, Hunyadi &amp; Páll-Gergely sp. nov.</p><p>Type material.</p><p>Holotype. Cambodia • 1 shell (SH: 2.44 mm; SW 1: 3.57 mm); Steung Treng Province, 36.6 km northwest + 3.1 km north from Stung Treng <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.74883&amp;materialsCitation.latitude=13.75975" title="Search Plazi for locations around (long 105.74883/lat 13.75975)">Mekong Bridge</a>, limestone mountain on the left side of the joining road; 13°45.585'N, 105°44.930'E; 130 m a. s. l.; 25 Oct. 2023, A. Hunyadi &amp; J. U. Otani leg.; CUMZ 14460 . Paratypes. Cambodia • 32 shells; same data as for holotype; coll. HA • 1 shell; same data as for holotype; coll. VG .</p><p>Additional material examined.</p><p>Cambodia • 2 shells (juveniles / damaged, not paratypes); same data as for holotype; coll. HA .</p><p>Type locality.</p><p>Cambodia, Steung Treng Province, 36.6 km northwest + 3.1 km north from Stung Treng Mekong Bridge, limestone mountain on the left side of the joining road; 13°45.585'N, 105°44.930'E; 130 m a. s. l.</p><p>Diagnosis.</p><p>A Hypselostoma species with depressed, concave-conical shell, bluntly and strongly keeled last whorl, raised spiral striation, blunt apertural barriers (altogether 6 or 7) and very wide umbilicus. Angular and parietal lamellae fused.</p><p>Description.</p><p>Shell depressed, concave-conical, brown, consisting of 5–5.5 whorls separated by a deep suture. Protoconch lighter than the rest of the shell, coarsely spirally striated (~ 12 striae) and consisting of 1.25–1.5 whorls. Protoconch and initial teleoconch whorls rounded, penultimate weakly convex. Last whorl with strong but blunt keel positioned at the centre of the periphery. Above the keel there is a deep groove. All whorls densely sculptured with strong, raised spiral striae crossed by less dense radial growth lines and occasionally by a few thin, whitish radial streaks. Spacing between the spiral striae irregular, but usually around the width of two, three, or four spiral striae. Last whorl slightly to moderately detached from the penultimate and slightly to moderately ascending near the aperture (~ 25–35 ° compared to the shell axis). Peristome expanded, not reflected, its surface finely pitted. Aperture equipped with four strong, main barriers (angulo-parietal, upper palatal, lower palatal, and columellar) and several smaller ones. Angulo-parietal lamella strong, its angular part is leaned towards the palatal side while its parietal part is leaned towards the columellar side. Angular part is pointed and smaller than the parietal. It is the only barrier reaching the expanding peristome. Upper and lower palatal plicae approximately the same size, the lower palatal being longer and more slender. Upper palatal plica slightly bent towards the lower palatal. Columellar lamella very thick and almost horizontal. Of smaller barriers, a small and weak interpalatal plica is present, stronger (but still weak) basal and a moderate knob-like plica in the columello-parietal transition embayment. A weak swelling is also observed at the palatal side (in front of the upper palatal plicae) which is probably homologous to the palatal tubercle in the majority of Bensonella species. Surface of all apertural barriers is densely granulated. Sinulus rounded, distinctly separated from the rest of the aperture. Umbilicus very wide, measuring between 1 / 3 and ½ of the shell width. There is a periumbilical keel situated on the last ~ 0.75 whorl right above the umbilicus. Along this keel there is a deep groove on the umbilical side.</p><p>Differential diagnosis.</p><p>This species differs from H. discobasis in the less slender shell and less numerous apertural barriers which are also thicker. It differs from H. sorormajor sp. nov. in its smaller dimensions and apertural barrier arrangement: it has less numerous barriers, none of which are hooked (in contrast to H. sorormajor sp. nov.). Hypselostoma aquila sp. nov. is smaller, more slender, has a narrower umbilicus and hooked barriers.</p><p>Measurements</p><p>(in mm, n = 5). SH = 2.37–2.57; SW 1 = 3.20–3.61; SW 2 = 2.05–2.40; AH = 1.25–1.42; AW = 1.22–1.26.</p><p>Etymology.</p><p>The name of this species is derived from two Latin words, soror meaning sister and minor meaning smaller which serves as a comparison with similar but usually larger H. sorormajor sp. nov.</p><p>Distribution.</p><p>This species is only known from the type locality.</p><p>Remarks.</p><p>The largest, and still unexplored limestone hill in Steung Treng Province (13°47.941'N, 105°43.7671'E) was inaccessible during the collecting efforts in 2023 due to the works of a cement factory. Due to the proximity of this hill to the type locality of H. sororminor sp. nov., it is possible that this species can be found here as well and be threatened by quarrying.</p></div>	https://treatment.plazi.org/id/B902884D17DF528C95464B957F1870A0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
A887FB65FABB5BFDACD4C14C7083EBAF.text	A887FB65FABB5BFDACD4C14C7083EBAF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma srakeoense (Panha & J. B. Burch 2004)	<div><p>Hypselostoma srakeoense (Panha &amp; J. B. Burch, 2004)</p><p>Figs 186 M, 211, 223</p><p>Anauchen srakeoensis Panha &amp; Burch in Panha et al. 2004: 62–63, fig. 4.</p><p>Anauchen srakeoensis — Panha and Burch 2008: 52–53, fig. 48; Dumrongrojwattana and Wongkamhaeng 2024: 323, fig. 7.</p><p>Hypselostoma srakeoensis — Sutcharit et al. 2023: 1291–1293, fig. 2.</p><p>Type material examined.</p><p>Thailand • 1 paratype; from the type locality; S. Panha leg.; SMF 331473 .</p><p>Additional material examined.</p><p>Thailand • 1 shell; Srakeo Province, Khaochakan District, Wat Khao Chakan; 22 June 2014; P. Dumrongrojwattana leg.; ex. coll. Maassen; coll. PGB .</p><p>Type locality.</p><p>“ Plubpluengtong limestone hills, Srakeo Province, 13°48'02"N, 102°12'49"E, 110 meters elevation ” (Thailand).</p><p>Differential diagnosis.</p><p>This species is superficially similar to several congeners with strongly enlarged last whorl which is also sharply descending. The two most similar species are H. torticollis and H. fungus sp. nov. However, H. torticollis has separated lamellae on the parietal side and it is also less depressed with clearly narrower umbilicus. Hypselostoma fungus sp. nov. has more rounded last whorl, wider umbilicus, more apertural barriers and is spirally striated. See also under H. pendulum .</p><p>Distribution.</p><p>This species is known from four sampling sites, two in Sa Kaeo province (Thailand) and two in Banteay Meanchey province (Cambodia) (Sutcharit et al. 2023).</p></div>	https://treatment.plazi.org/id/A887FB65FABB5BFDACD4C14C7083EBAF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
C1EECA34E5E15C94A4A28728002CE654.text	C1EECA34E5E15C94A4A28728002CE654.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma surakiti (Panha & J. B. Burch 2003)	<div><p>Hypselostoma surakiti (Panha &amp; J. B. Burch, 2003)</p><p>Figs 186 N, 212, 223</p><p>Gyliotrachela surakiti Panha &amp; Burch in Burch et al. 2003: 162–169, figs 12, 13.</p><p>Gyliotrachela surakiti — Tongkerd et al. 2004: 143, fig. 2; Panha and Burch 2008: 82–84, fig. 72; Dumrongrojwattana and Wongkamhaeng 2024: 324, fig. 8.</p><p>Type material examined.</p><p>Thailand • 2 paratypes; from the type locality; SMF 331450 • 1 paratype; from the type locality; CUMZ ver 078 .</p><p>Type locality.</p><p>“ South of Puttabanpot Temple, Pajoh Village, Nawang District, Nongbualumpoo Province, 17°19'1"N, 102°7'3"E, 320 meters elevation, Thailand ”.</p><p>Differential diagnosis.</p><p>This species is by the level of its last whorl ascension similar to H. kohrin and H. diarmaidi . It can be separated from H. kohrin and H. diarmaidi by the absence of typical spiral striation on the shell which is present in the latter two.</p><p>Distribution.</p><p>This species is known only from the type locality.</p><p>Remarks.</p><p>In H. surakiti, we could observe some irregular and weak spiral striae on the last whorl although they are not in form of typical raised spiral striae.</p></div>	https://treatment.plazi.org/id/C1EECA34E5E15C94A4A28728002CE654	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
4C91D7CE8F855BCC907456E6CD0C0B92.text	4C91D7CE8F855BCC907456E6CD0C0B92.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma taehwani Panha & J. B. Burch 2003	<div><p>Hypselostoma taehwani Panha &amp; J. B. Burch, 2003</p><p>Hypselostoma taehwani Panha &amp; Burch in Burch et al. 2003: 174–178, figs 16, 17.</p><p>Hypselostoma taehwani — Panha and Burch 2008: 101–102, fig. 86.</p><p>Type material examined.</p><p>Thailand • 1 paratype; from the type locality; CUMZ 44053 .</p><p>Type locality.</p><p>“ Central area of Tamrong Temple, Muang Petchburi District, near the main Petchkasem artery of the Asian Hwy. to southern Thailand, Petchaburi Province, 13°01'32"N, 99°55'11"E, 10 meters elevation, Thailand ”.</p><p>Differential diagnosis.</p><p>This species is strikingly similar to H. sculpturatum sp. nov. They can be separated by the following characters: i) initial teleoconch whorls and the penultimate whorl are more angled in H. sculpturatum sp. nov.; ii) umbilicus is wider and has a wider, deeper, and longer umbilical groove; iii) basal plica is in H. taehwani situated closer to the columellar lamella but always closer to the lower palatal plica in H. sculpturatum sp. nov. Furthermore, the protoconch is spirally striated in H. taehwani but pitted in H. sculpturatum sp. nov. Hypselostoma depressum has a wider umbilicus, much less detached last whorl near the aperture as well as the narrower spire. See also under H. burchi .</p><p>Distribution.</p><p>This species is known only from the type locality.</p></div>	https://treatment.plazi.org/id/4C91D7CE8F855BCC907456E6CD0C0B92	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
DE00A42C28DD5827AF5C620726E1EE1F.text	DE00A42C28DD5827AF5C620726E1EE1F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma terae Tomlin 1939	<div><p>Hypselostoma terae Tomlin, 1939</p><p>Figs 224 I, 235, 236, 237, 241</p><p>Hypselostoma terae Tomlin, 1939: 146, pl. 12, fig. 2.</p><p>Hypselostoma terae — van Benthem Jutting 1949 b: 59; van Benthem Jutting 1950: 43; van Benthem Jutting 1950: 21–22, fig. 10; Berry 1963: pl. 5, fig. 42.</p><p>Hypselostoma megaphonum van Benthem Jutting, 1950: 21–23, fig. 11. syn. nov.</p><p>Hypselostoma megaphonum — van Benthem Jutting 1950: 42; Maassen 2001: 77.</p><p>Hypselostoma perigyra van Benthem Jutting, 1950: 25–26, fig. 14. syn. nov.</p><p>Hypselostoma perigyra — van Benthem Jutting 1950: 42; Maassen 2001: 77.</p><p>Boysidia (Dasypupa) terae — Thompson and Dance 1983: 109.</p><p>Boysidia (Dasypupa) megaphona — Thompson and Dance 1983: 109.</p><p>Boysidia (Dasypupa) perigyra — Thompson and Dance 1983: 109.</p><p>Type material examined.</p><p>Malaysia • 53 paratypes of H. terae; from the type locality; RMNH Moll.137160 • 23 paratypes of H. perigyra; from the type locality; RMNH Moll.137163 • 23 paratypes of H. megaphonum; from the type locality; RMNH Moll.137161 .</p><p>Additional material examined.</p><p>Malaysia • 2 shells; Pahang, Bukit Cinta Manis, SSE side, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.013565&amp;materialsCitation.latitude=3.4452333" title="Search Plazi for locations around (long 102.013565/lat 3.4452333)">Lebuhraya Karak 800 m - Kampung Cinta Manis</a>; 03°26.714'N, 102°00.814'E; 130 m a. s. l.; 22 Jan. 2013; A. Hunyadi leg.; coll. HA • 2 shells; Pahang, 25 km northwest from Kuantan, east from Kampung Panching, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.147316&amp;materialsCitation.latitude=3.9115334" title="Search Plazi for locations around (long 103.147316/lat 3.9115334)">Gua Charas</a>, rock temple; 03°54.692'N, 103°08.839'E; 140 m a. s. l.; 24 Jan. 2013; A. Hunyadi leg.; coll. HA • 1 shell; Selangor, Kual Lumpur E- Kanching, Templer Park, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.64063&amp;materialsCitation.latitude=3.2979667" title="Search Plazi for locations around (long 101.64063/lat 3.2979667)">Perangsang Templer Golf Club</a>, limestone rock; 03°17.878'N, 101°38.438'E; 100 m a. s. l.; 25 Jan. 2013; A. Hunyadi leg.; coll. HA • 25 shells; Pahang, 15 km west from Bandar Pusat Jengka, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.433&amp;materialsCitation.latitude=3.6977" title="Search Plazi for locations around (long 102.433/lat 3.6977)">Hutan Lipur Gunung Senyum</a>; 03°41.862'N, 102°25.980'E; 85 m a. s. l.; 23 Jan. 2023; A. Hunyadi leg.; coll. HA .</p><p>Type localities.</p><p>“ Bukit Chintamani, Pahang ” (Bukit Cinta Manis), Malaysia ( H. terae); “ Bukit Charas, near Kuantan, Pahang ”, Malaysia ( H. megaphonum); “ Bukit Takun, Kanching, Selangor ”, Malaysia ( H. perigyra).</p><p>Differential diagnosis.</p><p>This species differs from H. elephas by its less detached and protruding last whorl as well as the much more conical shell.</p><p>Distribution.</p><p>This species is known from Pahang and Selangor provinces in Peninsular Malaysia.</p><p>Remarks.</p><p>Hypselostoma megaphonum and H. perigyra are synonymised with this species based on the following: i) all three species share the same arrangement and morphology of apertural barriers; ii) umbilici of all three species are equally narrow (with very slight variability which can be considered intraspecific); iii) all three species have the same narrow and sharp peripheral keel and iv) all three species are of similar size and distributed geographically close. All differences mentioned by van Benthem Jutting (1950) in the original description (last whorl inflation level, umbilicus width and apertural barrier strength) are recognised as intraspecific variability. Hypselostoma megaphonum can be interpreted as an intermediary form between H. terae and H. perigyra based on the following characters: i) last whorl more inflated than in H. terae but less than in H. perigyra; ii) apertural barriers stronger than in H. terae but weaker than in H. perigyra; iii) umbilicus wider than in H. terae but narrower than in H. perigyra .</p></div>	https://treatment.plazi.org/id/DE00A42C28DD5827AF5C620726E1EE1F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
4133167011B95CA49DC7D3CF8F5B35DB.text	4133167011B95CA49DC7D3CF8F5B35DB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma tertiusfrater Gojsina & Pall-Gergely 2025	<div><p>Hypselostoma tertiusfrater Gojšina &amp; Páll-Gergely nom. nov.</p><p>Figs 224 J, 238, 239</p><p>Boysidia (Dasypupa) salpinx Thompson &amp; Dance, 1983: 106–107, figs 2–8.</p><p>Boysidia salpinx — Dumrongrojwattana and Assawawattagee 2018: 216; Marzuki et al. 2021: 103–104, figs 45 E, 50 B; Inkhavilay and Sutcharit 2024: 441, fig. 1 D.</p><p>Type material examined.</p><p>Malaysia • holotype; UF 35944 • 33 paratypes; from the type locality; UF 36281 / 2, used for SEM by F. G. Thompson, destroyed; UF 35946 / 31 .</p><p>Additional material examined.</p><p>Malaysia • 3 shells; Borneo, Sarawak, 4 th Div., Niah Caves Nat. Park, near Painted Cave; Aug. 2002; coll. PGB • 24 shells; Borneo, Sarawak, SE portion, Bau area, Gunong Kapor, entrance to main cave; UF 35945 • 397 shells; Borneo, Sabah, ca 20 km upstream from Simatuoh, E side of Sapulut R., Batu Punggul; 16. May 1988; D. K. Dorman leg; locality code DKD- 91, UF 196670 • 190 shells; Borneo, Sarawak, Fourth division, Niah Caves National park, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=113.76667&amp;materialsCitation.latitude=3.8" title="Search Plazi for locations around (long 113.76667/lat 3.8)">summit of Gunong Subis, SE Batu Niah</a>; 03°48'N, 113°46′E; 29 Apr. 1988; D. K. Dorman leg.; locality code DKD- 77, UF 196488 • 15 shells; same data as previous; UF 196487 • 1 shell; Borneo, Sabah, ca 20 km upstream from Simatuoh, E side of Sapulut R., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=116.611664&amp;materialsCitation.latitude=4.6466665" title="Search Plazi for locations around (long 116.611664/lat 4.6466665)">Batu Punggul</a>; 04°38.8'N, 116°36.7'E; 16. May 1988; D. K. Dorman leg.; locality code DKD- 90, UF 196667 • 91 shells; same data as previous; locality code DKD-091; UF 196669 • 26 shells; Borneo, Sarawak, SW portion, S of Serian, E side Gunong Selabor, main entrance Lobang Batu cave; UF 35948 • 25 shells; Borneo, Sarawak, SW portion S side Gunong Selabor, S of Serian; UF 35947 • 3 shells; Sabah, ca. 25 km W Kunak, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=118.15&amp;materialsCitation.latitude=4.616667" title="Search Plazi for locations around (long 118.15/lat 4.616667)">Baturong Caves</a>; 04°37'N, 118°09'E; 12. Jun. 1988; David K. Dorman leg.; locality code DKD-102; UF 196829 .</p><p>Type locality.</p><p>“ Gunong Subis, limestone massif about 40 mi SW of Mira, Niah area, Fourth Div., Sarawak, Borneo, 03°51'N, 113°45'E ”.</p><p>Differential diagnosis.</p><p>This species is clearly different from the two congeners from Borneo ( H. paini and H. procerum) by its depressed-conical shell (vs conical) and a wider umbilicus.</p><p>Etymology.</p><p>The specific epithet is made of tertius meaning third and frater meaning brother, given the fact that this is one of three Hypselostoma species described from Borneo.</p><p>Distribution.</p><p>This species in known from Sarawak and Sabah, Borneo.</p><p>Remarks.</p><p>Boysidia salpinx F. G. Thompson &amp; Dance, 1983 is moved to the genus Hypselostoma herein, and thus, becomes a secondary homonym of Hypselostoma salpinx (van Benthem Jutting, 1961) . Therefore, Hypselostoma tertiusfrater nom. nov. is proposed as a replacement name.</p></div>	https://treatment.plazi.org/id/4133167011B95CA49DC7D3CF8F5B35DB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
EDCFC5D2D0C25AF6B17E4596E8649E84.text	EDCFC5D2D0C25AF6B17E4596E8649E84.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma torta Gojsina, Auffenberg & Pall-Gergely 2025	<div><p>Hypselostoma torta Gojšina, Auffenberg &amp; Páll-Gergely sp. nov.</p><p>Type material.</p><p>Holotype. Thailand • 1 shell (SH: 3.8 mm, SW: 3.5 mm); Nakhon Sawan Province <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.26667&amp;materialsCitation.latitude=15.233334" title="Search Plazi for locations around (long 100.26667/lat 15.233334)">19.5 m NE intersection Hwys 11 and 32</a>; 15°14'N, 100°16'E; 50 m a. s. l.; 02. May 1988; K. Auffenberg leg.; locality code KA-0696; UF 591362 . Paratypes. Thailand • 2 shells; same data as for holotype; CUMZ 14462 • 20 shells; same data as for holotype; UF 346125 • 18 shells; same data as for holotype; locality code KA-0697; UF 346133 • 4 ethanol-preserved shells; same data as for holotype; locality code KA-0696; UF 591361 .</p><p>Additional material examined.</p><p>Thailand • 5 shells (juveniles, not paratypes); same data as for holotype; locality code KA-0697; UF 583727 .</p><p>Type locality.</p><p>Thailand, Nakhon Sawan Province 19.5 m NE intersection Hwys 11 and 32; 15°14'N, 100°16'E; 50 m a. s. l.</p><p>Diagnosis.</p><p>A Hypselostoma with all teleoconch whorls shouldered and very weakly spirally striated. Spiral striae not raised. Last whorl detached from the penultimate and slightly descending. Four weak barriers in the aperture. Columellar lamella is like a strong swelling, forming a basal furrow.</p><p>Description.</p><p>Shell concave-conical (due to the strongly enlarged last whorl), brownish, consisting of 5–5.5 whorls separated by a very deep suture. Protoconch finely pitted, not showing spiralling pattern, rounded, same colour as the initial teleoconch whorls but slightly lighter than the last whorl. First two teleoconch whorls weakly shouldered, penultimate strongly and the last whorl is very strongly shouldered. The outline of the last and the penultimate whorl above the shoulder is concave (forming a flat platform). This surface is on initial teleoconch whorls more sloping. Whole teleoconch surface is weakly spirally striated (these spiral striae are not thread-like raised as in many congeners, but very blunt and indistinct) and occasionally crossed by radial growth lines and several thicker, whitish radial streaks which are rather unevenly positioned. These whitish streaks seem to be the densest on the penultimate whorl. Last whorl convex at its base, towards the umbilicus. It is slightly to moderately detached from the penultimate whorl and slightly descending near the aperture (~ 15–25 ° compared to the shell axis). Peristome very strong, dirty white, expanded but not reflected. Aperture equipped with three or four barriers (parietal (angulo-parietal?), upper palatal, lower palatal (sometimes absent), and columellar). There are no smaller barriers between the main ones. Parietal lamella is strong, high, and triangular when observed in perpendicular view. There is a very small tubercle-like swelling in front of the parietal lamella which may be homologous with the angular part of the usually present angulo-parietal (or just angular) in other congeners. Upper palatal plica moderately strong, almost twice as weak as the parietal lamella. Lower palatal plica very weak and low, dot-like, sometimes absent. Columellar lamella not present in its usual form (its borders are not clearly delimited) but as a rather strong, swelled part of the aperture which forms a prominent basal furrow below it (similar to the one present in H. fungus sp. nov.). The shape of the columellar side of the aperture is also reminiscent of the playground slide. Surface of all apertural barriers is smooth. Sinulus wide, distinctly separated from the rest of the aperture. Umbilicus wide, measuring ~ 1 / 3 of the shell width, expanding at the last whorl. Inside the umbilicus, there is a strong groove present, stretching across the whole last whorl.</p><p>Differential diagnosis.</p><p>This species is not similar to any of its congeners. It most closely resembles some Aulacospira species (e. g., A. pluangtong, A. smaesarnensis). However, it has a much wider umbilicus than both species as well as spiral striation and way more strongly shouldered whorls.</p><p>Measurements</p><p>(in mm, n = 5). SH = 3.45–3.8; SW = 2.83–3.62; AH = 1.62–1.82; AW = 1.38–1.84.</p><p>Etymology.</p><p>Since all whorls of this species are shouldered, the shell has an appearance of the wedding cake. The specific epithet torta comes from a Serbian word torta meaning a cake. To be used as a noun in apposition.</p><p>Distribution.</p><p>This species is known only from the type locality.</p><p>Remarks.</p><p>The shell of this species can sometimes be more depressed so that it is almost as wide as high. This species may belong to a separate genus or it may be closely related to some Aulacospira species ( A. pluangtong, A. smaesarnensis). However, due to the different shell shape (concave-conical), height (&gt; 3.4 mm) and surface sculpture (delicate, non-raised spiral striae) from typical Aulacospira species (inhabiting the Philippines), we provisionally place this species in the genus Hypselostoma .</p></div>	https://treatment.plazi.org/id/EDCFC5D2D0C25AF6B17E4596E8649E84	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
B2D28E0DA1275D35B9CDD2BA3574FFE6.text	B2D28E0DA1275D35B9CDD2BA3574FFE6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma torticollis (van Benthem Jutting 1962) Gojšina & Hunyadi & Sutcharit & Tongkerd & Auffenberg & Grego & Vermeulen & Reischütz & Páll-Gergely 2025	<div><p>Hypselostoma torticollis (van Benthem Jutting, 1962) comb. nov.</p><p>Gyliotrachela torticollis van Benthem Jutting, 1962: 8–10, fig. 4.</p><p>Gyliotrachela torticollis — Sutcharit et al. 2023: 1293–1295, fig. 3.</p><p>Type material examined.</p><p>Cambodia • 3 paratypes; Battambang, Phuom Truong Mean; 1960; E. Saurin leg.; RMNH.Moll.137136 .</p><p>Type locality.</p><p>“ Phum Troung Mean, à 15 km au S. - O. de Battambang, Cambodge ” (Cambodia).</p><p>Differential diagnosis.</p><p>Hypselostoma fungus sp. nov. has merged angular and parietal lamellae, wider umbilicus, last whorl without a keel and a flatter shell. See also under H. transitans, H. pendulum, H. srakeoense .</p><p>Distribution.</p><p>This species is known only from the Battambang province, Cambodia.</p><p>Remarks.</p><p>Of three examined paratypes, one did not have the aperture as downturned as the others.</p></div>	https://treatment.plazi.org/id/B2D28E0DA1275D35B9CDD2BA3574FFE6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
D376B9833A165298A454CD46DC6DD0E3.text	D376B9833A165298A454CD46DC6DD0E3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma transitans Mollendorff 1894	<div><p>Hypselostoma transitans Möllendorff, 1894</p><p>Figs 186 O, 213, 214, 223</p><p>Hypselostoma transitans Möllendorff, 1894: 16, figs 12, 13.</p><p>Hypselostoma translucidum [sic] — Fischer and Dautzenberg 1904: 408.</p><p>Gyliauchen transitans — Pilsbry 1917: 214–215, pl. 36, figs 5–8.</p><p>Gliotrachela transitans [sic] — Laidlaw 1933: 214.</p><p>Gyliotrachela transitans transitans — van Benthem Jutting 1950: 47; van Benthem Jutting 1950: 28; Thompson and Upatham 1997: 233.</p><p>Gyliotrachela transitans — Zilch 1984: 166; Hemmen and Hemmen 2001: 41; Panha and Burch 2008: 85, fig. 74; Dumrongrojwattana and Wongkamhaeng 2024: 324, fig. 8.</p><p>Hypselostoma transitans — Fischer and Dautzenberg 1904: 408.</p><p>Type material examined.</p><p>Thailand • 1 lectotype; SMF 4589 • 8 paralectotypes; from the type locality; SMF 4590 .</p><p>Additional material examined.</p><p>Thailand • 2 shells; Chumphon Province, 2.5 km northeast from Pathio, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.3207&amp;materialsCitation.latitude=10.73085" title="Search Plazi for locations around (long 99.3207/lat 10.73085)">Tham Khao Phlu</a>; 10°43.851'N, 99°19.242'E; 23 Feb. 2015; A. Hunyadi leg.; coll. HA • 22 fragments and 1 complete shell; Suratthani Province, Angthong islands, Wia Talap island, steep path to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.92443&amp;materialsCitation.latitude=8.126717" title="Search Plazi for locations around (long 98.92443/lat 8.126717)">Buaboke cave</a>, at the base of limestone rocks; 8°7.603'N, 98°55.466'E; 50 m a. s. l.; Sep. 2007; A. Reischütz leg.; coll. REI .</p><p>Type locality.</p><p>“ Samui islands, Gulf of Siam ”, Thailand.</p><p>Differential diagnosis.</p><p>Hypselostoma torticollis is similar due to the down-turning aperture and same arrangement of the apertural barriers but this species has a much wider umbilicus, more enlarged last whorl and is spirally striated. Hypselostoma fungus sp. nov. is spirally striated, has merged angular and parietal lamellae, more down-turning last whorl and a much wider umbilicus. See also under H. venustum .</p><p>Distribution.</p><p>This species in known from Chumphon and Suratthani provinces in Thailand.</p><p>Remarks.</p><p>In some paralectotypes, a lamella between the columellar and parietal was positioned much deeper in the aperture. We have examined a distinct form of this species from Tham Khao Phlu (Chumphon province). This form had a wider umbilicus and a sharper keel as well as more enlarged last whorl in comparison to previous ones. However, we decided not to treat it as distinct species due to the fact that we have also found a transitional form between the typical H. transitans (close to its type locality) and the one from Tham Khao Phlu.</p></div>	https://treatment.plazi.org/id/D376B9833A165298A454CD46DC6DD0E3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
3F54769239FE575F881C38DB805CFDE1.text	3F54769239FE575F881C38DB805CFDE1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma tridentatum (Panha & J. B. Burch 2004) Gojšina & Hunyadi & Sutcharit & Tongkerd & Auffenberg & Grego & Vermeulen & Reischütz & Páll-Gergely 2025	<div><p>Hypselostoma tridentatum (Panha &amp; J. B. Burch, 2004) comb. nov.</p><p>Gyliotrachela tridentatus Panha &amp; Burch in Panha et al. 2004: 70–71, fig. 9.</p><p>Gyliotrachela tridentata — Panha and Burch 2008: 86–87, fig. 75.</p><p>Gyliotrachela tridentatus — Dumrongrojwattana and Wongkamhaeng 2024: 324, fig. 8.</p><p>Material examined.</p><p>Thailand • 2 shells; Srakeo Province, Plubpluengtong, Phloeng Thong Cave, Khao Phiap; 24 Dec. 2012; K. Okubo leg.; coll. PGB . Cambodia • Battambang Province, Phnum Proek District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.60817&amp;materialsCitation.latitude=13.3139" title="Search Plazi for locations around (long 102.60817/lat 13.3139)">Phnom Prampi Cave</a>; 13°18.834'N, 102°36.490'E; 90 m a. s. l.; 10 Oct. 2023; A. Hunyadi, J. U. Otani leg., coll. HA .</p><p>Type locality.</p><p>“ Plubpluengtong limestone hills, Srakeo Province, 13°27'07"N, 102°12'49"E, 80 meters elevation ” (Thailand).</p><p>Differential diagnosis.</p><p>This species has fewer barriers in the aperture than H. khaowongkot which is the most similar species. It also does not have columellar lamella (present in H. khaowongkot) and lacks the strongly spiniferous surface of apertural barriers (also present in H. khaowongkot). See also under H. utongense .</p><p>Distribution.</p><p>This species is, apart from the type locality, also known from Phnom Prampi Cave in Battambang Province which is a first record from Cambodia.</p></div>	https://treatment.plazi.org/id/3F54769239FE575F881C38DB805CFDE1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
64C84E2FAE2A5EC2AA14A4B042FB6EA4.text	64C84E2FAE2A5EC2AA14A4B042FB6EA4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma troglodytes (van Benthem Jutting 1950) Gojšina & Hunyadi & Sutcharit & Tongkerd & Auffenberg & Grego & Vermeulen & Reischütz & Páll-Gergely 2025	<div><p>Hypselostoma troglodytes (van Benthem Jutting, 1950) comb. nov.</p><p>Figs 224 K, 240, 241</p><p>Gyliotrachela troglodytes van Benthem Jutting, 1950: 35–36, 47, fig. 21.</p><p>Gyliotrachela troglodytes — Maassen 2001: 77.</p><p>Type material examined.</p><p>Malaysia • 3 paratypes; from the type locality; Sept. 1941; Raffles Museum, Singapore ex. coll.; RMNH.Moll.137138 .</p><p>Type locality.</p><p>“ Gua Bama, Padang Tengku, Pahang ”, Malaysia.</p><p>Differential diagnosis.</p><p>This species can be separated from H. vicinum sp. nov. by the notably narrower umbilicus, weaker peripheral keel, and the absence of the basal and one interpalatal plica. Furthermore, the shell surface sculpture seems to be less rough than in H. vicinum sp. nov.</p><p>Distribution.</p><p>This species is known only from the type locality.</p><p>Remarks.</p><p>In the original description, an infraparietal lamella is mentioned. We observed it in paratypes but it should be noted that it is dot-like and very small.</p><p>4. Hypselostoma tubiferum group</p><p>Diagnosis. Hypselostoma tubiferum group is devoid of any peculiar shell surface sculpture. It is smooth or finely pitted with some radial growth lines.</p><p>Remarks. This group includes nine species. Their distribution spans from Guangxi, Guizhou, and Yunnan provinces in China southwards to Peninsular Malaysia. Westernmost localities are known from central Myanmar.</p></div>	https://treatment.plazi.org/id/64C84E2FAE2A5EC2AA14A4B042FB6EA4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
F475DBCA40045DADA0BA4AF63CEA4EF7.text	F475DBCA40045DADA0BA4AF63CEA4EF7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma tubiferum (Benson 1856)	<div><p>Hypselostoma tubiferum (Benson, 1856)</p><p>Figs 242 I, 260, 261</p><p>Tanystoma tubiferum Benson, 1856 a: 130 .</p><p>Hypselostoma tubiferum — Pfeiffer 1859: 325; Blanford 1863: 326; Hanley and Theobald 1870: 4, pl. 8 fig. 3; Stoliczka 1871: 173, pl. 7 fig. 1; Pfeiffer 1876 in 1875–1876: 488; Möllendorff 1891: 338; Gude 1914: 298; Pilsbry 1917: 178–179, pl. 31 figs 1–5; van Benthem Jutting 1950: 43; Zilch 1984: 163; Gojšina et al. 2022: 137, fig. 7; Tongkerd et al. 2024: 188, fig. 13 Q.</p><p>Pupa (Hypselostoma) tubifera — Nevill 1878: 193.</p><p>Material examined.</p><p>Myanmar • 6 shells; Tondung / Thyet Mio; W. Theobald leg.; NHMUK 1888.12.4.17 –22 • 35 shells; Mandalay, Patheingyi, near <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=96.21515&amp;materialsCitation.latitude=21.988016" title="Search Plazi for locations around (long 96.21515/lat 21.988016)">Shin Bodh Mell Waterfall</a>; 21°59.281'N, 96°12.909'E; 225 m a. s. l.; 16 Oct. 2018; A. Hunyadi leg.; coll. HA • 31 shells; Mandalay, Kyaukse, Kho Nan Shin <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=96.33715&amp;materialsCitation.latitude=21.735884" title="Search Plazi for locations around (long 96.33715/lat 21.735884)">Cave Pagoda</a>; 21°44.153'N, 96°20.229'E; 110 m a. s. l.; 17 Oct. 2018; A. Hunyadi leg.; coll. HA • 45 shells; Mandalay, Dee Dote, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=96.3547&amp;materialsCitation.latitude=21.7094" title="Search Plazi for locations around (long 96.3547/lat 21.7094)">Waterfall</a>; 21°42.564'N, 96°21.282'E; 185 m a. s. l.; 17 Oct. 2018; A. Hunyadi leg.; coll. HA • 3 shells; " India " (probably referring to former British India); F. F. Linter coll.; UF 00112342 • 1 shell; India (probably referring to former British India), Tankarin; 01. Jun. 1998 – 30. Jun. 1998; UF 540729 .</p><p>Type locality.</p><p>“ Thyet-Mio prope ripas fluminis Irawadi Burmanici, saxis calcareis adhaerens ” (nowadays Thayet), Myanmar.</p><p>Differential diagnosis.</p><p>This species is superficially similar to H. depressispira from which it can be separated by the concrescent angulo-parietal lamella (separated in H. depressispira) and fewer barriers in the aperture which are developed to roughly the same extent.</p><p>Distribution.</p><p>This species is, apart from the type locality, known from several localities in Mandalay, Myanmar.</p><p>Remarks.</p><p>Colouration of the animal body was observed and described by Stoliczka (1871). It was described as pale grey with blackish tentacles. Foot is short and ovate-elongated. The striking similarity of this species with H. depressispira suggests that Gyliotrachela is a junior synonym of Hypselostoma .</p></div>	https://treatment.plazi.org/id/F475DBCA40045DADA0BA4AF63CEA4EF7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
71181B30F41C5C2AAC2AD8A7A05DAA17.text	71181B30F41C5C2AAC2AD8A7A05DAA17.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma undefined-1	<div><p>Hypselostoma sp. 1</p><p>Material examined.</p><p>Thailand • 2 shells; Trang Province, Kantang district, Yong Ling beach; 4 July 2014; J. U. Otani leg.; coll. PGB . Malaysia • 344 shells; Kelantan Province, 19 km south of Gua Musang; Oct. 1999; Hemmen leg.; coll. PGB .</p><p>Remarks.</p><p>Two samples from Thailand and Malaysia represent two different species, both of them probably most similar to H. burchi . Two specimens from Thailand are slightly more slender, lack interpalatal plica and have slightly narrower umbilicus. Specimens from Malaysia have slightly more detached and descending last whorl and they lack the interpalatal plica as well. Due to the absence of proper comparison material of H. burchi, we do not describe these taxa in this work.</p></div>	https://treatment.plazi.org/id/71181B30F41C5C2AAC2AD8A7A05DAA17	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
6392CEF432265ADC95BAD9B10781109B.text	6392CEF432265ADC95BAD9B10781109B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma undefined-2	<div><p>Hypselostoma sp. 2</p><p>Material examined.</p><p>Thailand • 1 shell; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.7&amp;materialsCitation.latitude=9.3" title="Search Plazi for locations around (long 99.7/lat 9.3)">Suratthani Province</a>, 4.5 km S Don Sak, E side of Hwy. 4142, evergreen forest, limestone outcrop, base of cliff; 9°18'N, 99°42'E; 18 Apr. 1988; K. Auffenberg leg.; locality code KA-0674; UF 345307 .</p><p>Remarks.</p><p>A single fragment, identification impossible.</p></div>	https://treatment.plazi.org/id/6392CEF432265ADC95BAD9B10781109B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
E032DFB56C205AAC9BADEEFA5BC7BC80.text	E032DFB56C205AAC9BADEEFA5BC7BC80.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma utongense Panha & J. B. Burch 2004	<div><p>Hypselostoma utongense Panha &amp; J. B. Burch, 2004</p><p>Hypselostoma utongensis Panha &amp; Burch in Panha et al. 2004: 72–73, fig. 10.</p><p>Hypselostoma utongensis — Panha and Burch 2008: 103, fig. 87; Dumrongrojwattana and Wongkamhaeng 2024: 324, fig. 8.</p><p>Material examined.</p><p>Thailand • 16 shells; Kanchanaburi Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.14498&amp;materialsCitation.latitude=14.371834" title="Search Plazi for locations around (long 99.14498/lat 14.371834)">Erawan National Park</a>, Erawan Falls Trail; 14°22.310'N, 99°08.699'E; 17 Feb. 2015; A. Hunyadi leg.; coll. HA .</p><p>Type locality.</p><p>“ Tam Sua Hill, Utong District, Supanburi Province, 14°34'27"N, 99°46'26"E, 80 meters elevation ”.</p><p>Differential diagnosis.</p><p>This species is similar to two Hypselostoma which share the completely (or almost completely) flat shell with strongly upturning last whorl ( H. khaowongkot and H. tridentatum). From them, it most obviously differs by its hooked upper and lower palatal plicae which are blunt in the former.</p><p>Distribution.</p><p>This species is known from two provinces in Thailand, Supanburi (type locality) and Kanchanaburi (this study).</p><p>Remarks.</p><p>In some specimens, previous whorls are partly sunken into the last while in others completely.</p><p>2. Hypselostoma hungerfordianum group</p><p>Diagnosis. Hypselostoma hungerfordianum group is characterised by a peculiar shell surface sculpture, as if “ scaly, granulated ”. This surface sculpture is unique and we assume that all the species which share this trait are possibly more closely related.</p><p>Remarks. This group has representatives across Southeast Asia and includes 19 species. The distribution spans from Kayin State in Myanmar and eastwards to western Cambodia (Battambang Province). Southernmost localities are known in Peninsular Malaysia while northernmost come from Loei Province in Thailand.</p></div>	https://treatment.plazi.org/id/E032DFB56C205AAC9BADEEFA5BC7BC80	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
A02A9D5E1C08555BA9BECCC1A11E5401.text	A02A9D5E1C08555BA9BECCC1A11E5401.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma venustum (van Benthem Jutting 1950)	<div><p>Hypselostoma venustum (van Benthem Jutting, 1950)</p><p>Figs 186 P, 215, 216, 223</p><p>Gyliotrachela transitans venusta van Benthem Jutting, 1950: 28–29, 47, fig. 16.</p><p>Gyliotrachela transitans venusta — Maassen, 2001: 76.</p><p>Gyliotrachela transitans helioscopia van Benthem Jutting, 1950: 29–31, fig. 17. syn. nov.</p><p>Gyliotrachela transitans heliscopia [sic] — Maassen, 2001: 76.</p><p>Type material examined.</p><p>Malaysia • holotype of G. transitans venusta; Raffles Museum, Singapore ex. coll.; RMNH.Moll.137140 • 8 paratypes of G. transitans helioscopia; from the type locality; Raffles Museum, Singapore ex. coll.; 1947; RMNH.Moll.137144 .</p><p>Additional material examined.</p><p>Malaysia • 3 shells; Pahang, 20 km southeast from Jerantut, Gua Kota Gelanggi, under <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.47353&amp;materialsCitation.latitude=3.9" title="Search Plazi for locations around (long 102.47353/lat 3.9)">Gua Balai</a>; 03°54.000'N, 102°28.412'E; 115 m a. s. l.; 21 Jan. 2013; A. Hunyadi leg.; coll. HA .</p><p>Type localities.</p><p>“ Gunong Pondok, Padang Rengas, Perak ”, Malaysia ( H. venustum); “ Kota Tongkat, Pahang ”, Malaysia ( G. transitans helioscopia).</p><p>Differential diagnosis.</p><p>This species is similar to H. ophis sp. nov., from which it can be separated by narrower umbilicus and shorter and wavy parietal lamella. Hypselostoma transitans has more rough shell sculpture as well as the descending last whorl and a wider umbilicus. See also under H. vesovici sp. nov.</p><p>Distribution.</p><p>This species is known only from three localities in Peninsular Malaysia (Gunong Pondok, Kota Tongkat and Gua Kota Gelanggi).</p><p>Remarks.</p><p>Van Benthem Jutting (1950) recognised two additional subspecies of G. transitans, namely G. transitans helioscopia and G. transitans venusta . However, these two taxa should not be considered as subspecies since they differ from the nominotypical subspecies in several very important characters: last whorl is in nominotypical subspecies strongly descending near the aperture while in G. transitans venusta it is slightly ascending; whorls are more depressed in nominotypical subspecies; shell surface is also different, being more sandpaper-like in nominotypical subspecies and almost smooth to very finely sculptured in other two subspecies. Finally, umbilicus is wider in G. transitans . Because of all aforementioned clear differences, we conclude that H. venusta is a good species separated from H. transitans . Gyliotrachela transitans helioscopia is its junior synonym due to the virtually identical shell morphology. Van Benthem Jutting (1950) noted that in G. transitans venusta, the shell is less elevated, peripheral keel sharper and longitudinal grooves deeper than in G. transitans helioscopia . These differences are only minor and fall within the intraspecific variability, thus there are no clear characters which allow the separation of these two taxa.</p></div>	https://treatment.plazi.org/id/A02A9D5E1C08555BA9BECCC1A11E5401	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
3E3B47F9AE5F59EA86FC7143B4614F37.text	3E3B47F9AE5F59EA86FC7143B4614F37.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma vesovici Gojsina & Pall-Gergely 2025	<div><p>Hypselostoma vesovici Gojšina &amp; Páll-Gergely sp. nov.</p><p>Figs 186 Q, 217, 218, 223</p><p>Type material.</p><p>Holotype. Malaysia • 1 shell (SH: 2.18 mm; SW 1: 3.12 mm); Gua Telinga, Kuala Tahan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.39578&amp;materialsCitation.latitude=4.3721" title="Search Plazi for locations around (long 102.39578/lat 4.3721)">Taman Negara</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.39578&amp;materialsCitation.latitude=4.3721" title="Search Plazi for locations around (long 102.39578/lat 4.3721)">Jerantut</a>, Pahang; 4°22.326'N, 102°23.747'E; 69 m a. s. l.; T. Ishibe, K. Ohara, K. Okubo &amp; J. U. Otani leg.; CUMZ 14463 . Paratypes. Malaysia • 10 shells; same data as for holotype; coll. PGB • 1 shell; same data as for holotype; coll. VG • 1 shell; same data as for holotype; coll. HA .</p><p>Type locality.</p><p>Malaysia, Gua Telinga, Kuala Tahan, Taman Negara, Jerantut, Pahang; 4°22.326'N, 102°23.747'E; 69 m a. s. l.</p><p>Diagnosis.</p><p>Shell concave-conical, with roughly granulated (sandpaper-like) surface sculpture. Last two whorls keeled below the periphery, keel especially strong on the last whorl. Last whorl detached from the penultimate, directed upwards. Aperture equipped with five main barriers and usually five smaller ones. All barriers are very broad and thick, very strongly spiniferous. Umbilicus very narrow, dot-like.</p><p>Description.</p><p>Shell concave-conical (due to the strongly enlarged last whorl), pinkish-brown in colour, corneous. It is consisting of 4.5–5 regularly increasing whorls separated by a deeply impressed suture. Protoconch finely pitted, not showing spiralling pattern, consisting of 1.25–1.5 whorls and slightly darker than the rest of the shell. Protoconch- teleoconch boundary clearly visible due to the change in shell surface sculpture. Teleoconch roughly granulated, sandpaper-like. On the last whorl near the keel, granules can in some specimens be arranged in a way to provoke a spiralling pattern although no regular spiral striae are developed. Radial lines are strong, widely and irregularly spaced, ~ 17 radial lines on the last whorl in standard view. They are more densely arranged on the initial teleoconch whorls. Protoconch and initial teleoconch whorls rounded and convex, last two whorls are keeled below the centre of the periphery. Last whorl especially strongly keeled, with a deep groove above the keel. It is also moderately detached from the penultimate near the aperture, and slightly ascending (~ 15–25 ° compared to the shell axis). Peristome whitish, strongly expanded but not reflected. Aperture large, equipped with five main barriers (parietal, angular, upper palatal, lower palatal and columellar). Angular lamella is the weakest and the shortest among the main five. All other main barriers are relatively thick and broad, almost equally developed. Parietal lamella is the longest in the aperture and surpassing the profile of the angular. Between the main barriers, several smaller ones could be observed. There are usually two interpalatal plicae, two between the lower palatal and the columellar and one knob-like infraparietal lamella. All barriers, as well as the aperture surface, are very strongly spiniferous. These spines are widely spaced (especially on the barriers) and much stronger than in other congeners. Sinulus not very distinctly separated from the rest of the aperture. Umbilicus is very narrow, dot-like, not showing previous whorls and measuring slightly&gt; 1 / 10 of the shell width. A very shallow groove is visible running alongside the umbilicus.</p><p>Differential diagnosis.</p><p>This species is by the general shell shape similar to H. venustum from which it clearly differs by a sharply keeled last whorl below the periphery (provoking a discoid basis of the shell and a concave surface above the keel), thicker apertural barriers with very strongly spiniferous surfaces and a slightly narrower umbilicus with a much shallower groove inside it. Furthermore, the penultimate whorl in H. venustum is rounded, not keeled as in H. vesovici sp. nov. and the aperture in H. vesovici sp. nov. is much larger when compared to the rest of the shell than in H. venustum . However, the species which has the same appearance of the last whorl, and thus most similar are H. sichang and H. discobasis . In contrast to the new species, these two representatives share the different shell surface (very roughly sandpaper-like in H. sichang and coarsely spirally striated in H. discobasis) and different arrangement of the apertural barriers (less strong and spiniferous, angular and parietal lamellae merged in H. discobasis), as well as much wider umbilicus especially in H. discobasis . Hypselostoma vesovici sp. nov. also has clearly much larger aperture when compared to the rest of the shell than in H. sichang .</p><p>Measurements</p><p>(in mm, n = 5). SH = 2.10–2.35; SW 1 = 2.53–3.12; SW 2 = 1.49–1.77; AH = 1.10–1.23; AW = 1.05–1.25.</p><p>Etymology.</p><p>Named after Dr. Nikola Vesović, a prominent Serbian entomologist and a friend of the first author.</p><p>Distribution.</p><p>Known only from the type locality.</p></div>	https://treatment.plazi.org/id/3E3B47F9AE5F59EA86FC7143B4614F37	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
80BF9D4EEC8F5050810CAE16FBD94350.text	80BF9D4EEC8F5050810CAE16FBD94350.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma vicinum Gojsina, Auffenberg & Pall-Gergely 2025	<div><p>Hypselostoma vicinum Gojšina, Auffenberg &amp; Páll-Gergely sp. nov.</p><p>Figs 186 R, 219, 220, 223</p><p>Type material.</p><p>Holotype. Thailand • 1 shell (SH: 1.9 mm, 2.6 mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.666664&amp;materialsCitation.latitude=9.166667" title="Search Plazi for locations around (long 99.666664/lat 9.166667)">Suratthani Province</a>, limestone outcrop along Hwy. 401, 2.7 km W junc Hwys. 4142 &amp; 401, evergreen forest on rocky hillside, below cliff, base of cliff; 9°10'N, 99°40'E; 90 m a. s. l.; 18 Apr. 1988; K. Auffenberg leg.; UF 345327.</p><p>Paratypes. Thailand • 1 shell; same data as for holotype; CUMZ 14468 • 51 shells; same data as for holotype; UF 591363 • 45 shells; same data as for holotype; locality code KA-0674, UF 345336 • 16 ethanol-preserved specimens; same data as for holotype; locality code KA-0673, UF 345328 .</p><p>Additional material examined.</p><p>Thailand • 12 shells (apical whorls / juveniles, not paratypes); same data as for holotype; UF 583724 • 3 shells (juveniles / damaged, not paratypes); same data as for holotype; locality code KA-0674; UF 583725 .</p><p>Type locality.</p><p>Thailand, Suratthani Province, limestone outcrop along Hwy. 401, 2.7 km W junc Hwys. 4142 &amp; 401, evergreen forest on rocky hillside, below cliff, base of cliff; 9°10'N, 99°40'E; 90 m a. s. l.</p><p>Diagnosis.</p><p>A Hypselostoma species with roughly granulated (not spirally striated) teleoconch, and keeled, detached, and ascending last whorl. Apertural barriers few (five main ones and usually three smaller ones).</p><p>Description.</p><p>Shell depressed, concave-conical (due to the strongly enlarged last whorl), chestnut brown and weakly glossy, consisting of 3.75–4.25 convex whorls separated by a deep suture. Protoconch is of the same colouration as the teleoconch, finely pitted and showing no spiralling pattern. Boundary between the protoconch and the teleoconch not clear under microscope because of the very similar surface sculpture of the two regions, but the former is ~ 1.5 whorls. Teleoconch roughly granulated (sandpaper-like), weakly sculptured with radial growth lines but devoid of spiral striation (these radial lines are much harder to observe than the prominent granulation). Occasionally, thicker whitish radial streaks are unevenly placed across the surface of the shell. Last whorl with a very blunt keel (which is getting stronger near the aperture) positioned at the centre of the periphery. Shell surface is immediately below and above the keel concave, and then again convex towards the suture and the umbilicus. Last whorl is also very slightly detached from the penultimate and moderately ascending upwards (~ 40–50 ° compared to the shell axis). Peristome dirty white, expanded but not reflected, its surface very finely pitted. There are altogether five main apertural barriers (angular, parietal, upper palatal, lower palatal and columellar) and several smaller ones of variable number. Parietal lamella is the strongest and highest in the aperture, straight but directed towards the palatal side. Angular lamella ~ 2 × lower and shorter than the parietal, but extends closer to the expanding peristome, almost reaching it. Upper and lower palatal plicae almost the same in height and length, the former may be slightly weaker than the latter. Columellar lamella horizontal, almost identical to the lower palatal plica. Among the smaller barriers, there is usually one interpalatal plica which is very small, dot-like. Slightly larger basal plica is present nearly halfway between the lower palatal and columellar. One dot-like swelling is present in the columello-parietal transition embayment. Surface of all apertural barriers is very finely spiniferous medially and very finely granulated laterally. Sinulus wide, not distinctly separated from the rest of the aperture. Umbilicus wide, measuring slightly&gt; 1 / 4 of the shell width. A shallow groove runs alongside the umbilicus but is visible only around the last quarter of the last whorl.</p><p>Differential diagnosis.</p><p>Less depressed forms of H. fortunatum sp. nov. can resemble this species but the barriers in the former are much weaker and angular and parietal lamellae are concrescent. See also under H. troglodytes .</p><p>Measurements</p><p>(in mm, n = 5). SH = 1.73–2.06; SW 1 = 2.49–2.85; SW 2 = 1.63–1.79; AH = 0.93–1.19; AW = 0.94–1.10.</p><p>Etymology.</p><p>The specific epithet vicinum comes from the Latin word for neighbour. This name is provided because the most similar species to the new one, H. troglodytes, lives in the neighbouring Peninsular Malaysia.</p><p>Distribution.</p><p>This species is known only from the type locality.</p></div>	https://treatment.plazi.org/id/80BF9D4EEC8F5050810CAE16FBD94350	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
BC7FB5AE88D6531187EFB079DAE51A20.text	BC7FB5AE88D6531187EFB079DAE51A20.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma vujici Gojsina & Pall-Gergely 2025	<div><p>Hypselostoma vujici Gojšina &amp; Páll-Gergely sp. nov.</p><p>Figs 186 S, 221, 222, 223</p><p>Type material.</p><p>Holotype. Thailand • 1 shell (SH: 2.47 mm; SW: 1.93 mm); Nakhon Sawan Province, 4.0 km NW of Ban Non San; 15 ° 942 ' N, 99 ° 873 ' E; 100 m a. s. l.; 12 June 1987; F. G. Thompson leg.; UF 528941 . Paratypes. Thailand • 1 shell; same data as for holotype; CUMZ 14469 • 2 shells; same data as for holotype; UF 591364 • 8 shells (2 whole, others badly damaged, paratypes); same locality data as for holotype; locality code FGT-4325, UF 530691 .</p><p>Additional material examined.</p><p>Thailand • 3 shells (2 juveniles, 1 last whorl with damaged aperture, not paratypes); same data as for holotype; UF 583723 .</p><p>Type locality.</p><p>Thailand, Nakhon Sawan Province, 4.0 km NW of Ban Non San; 15 ° 942 ' N, 99 ° 873 ' E; 100 m a. s. l. (provided coordinates are wrong, they cannot be understood by Google Earth).</p><p>Diagnosis.</p><p>A Hypselostoma species with 5.75–6 roughly granulated whorls and a very deep suture. Last whorl somehow canted and adnate to the penultimate near the aperture. Palatal plicae and basal plica hooked. Other barriers (angulo-parietal, columellar and infraparietal) blunt. Umbilicus narrow.</p><p>Description.</p><p>Shell high, concave-conical, shiny brown or even colourless, consisting of 5–5.5 whorls separated by a very deep suture. Protoconch almost smooth, with very weak spiralling pattern, consisting of ~ 1.5 whorls but its boundary with the teleoconch is not clearly visible due to the similar surface sculpture. Teleoconch with weak and coarse radial growth lines, which are on the last two whorls visible as white streaks, and rough granulation (sandpaper-like) but no spiral striation. Initial teleoconch whorls rounded, penultimate only very slightly less rounded. Last whorl adnate to slightly detached from the penultimate, with a very weak keel situated near the centre of the periphery. It is slightly descending near the aperture (~ 5–10 ° compared to the shell axis). Peristome dirty white, expanded but not reflected. Aperture rounded (almost as wide as high), equipped with six main barriers (angulo-parietal, upper palatal, interpalatal, lower palatal, basal and columellar) and one weak infraparietal swelling-like lamella. Angulo-parietal lamella is the strongest in the aperture, with clearly distinguishable angular and parietal part and a prominent sinuation between them. Parietal part stronger and higher than the angular, both curved towards the palatal wall. All palatal plicae hooked, pointing outside. Upper palatal and interpalatal plica positioned close together, almost equally as strong. Lower palatal plica stronger than others and situated some distance from the interpalatal. Basal plica also hooked but less clearly than the palatals, strong as the interpalatal and upper palatal but situated closer to the columellar lamella than to the lower palatal plica. Columellar lamella not hooked, long and almost horizontal. A small swelling-like infraparietal lamella is present. Angulo-parietal lamella is reaching the expanding peristome, columellar lamella almost reaching it. Surface of all apertural barriers is finely granulated. Sinulus wide, not distinctly separated from the rest of the aperture. Umbilicus always open but of variable width, measuring ~ 1 / 6 or ≤ 1 / 3 of the shell width. Umbilical groove is strong in specimens with wider and weak in specimens with narrower umbilicus.</p><p>Differential diagnosis.</p><p>This species can be separated from all other congeners by the combination of a slender shell with a larger number of whorls (usually ~ 6), roughly granulated teleoconch and hooked basal and three palatal plicae in the aperture.</p><p>Measurements</p><p>(in mm, n = 4). SH = 2.36–2.62; SW = 1.78–1.93; AH = 0.92–0.96; AW = 0.85–0.93.</p><p>Etymology.</p><p>Named after Mihailo Vujić, a prominent Serbian entomologist and a friend of the first author.</p><p>Distribution.</p><p>This species is known only from the type locality.</p><p>Remarks.</p><p>We have noticed significant intraspecific variability in this species: there were some specimens with elongated-conical, brown shell and some with more depressed spire or much lighter (sometimes even colourless) shell. Umbilicus varied from 1 / 6 to 1 / 3 of the shell width with weak or sometimes deep and strong groove inside it.</p><p>3. Hypselostoma terae group</p><p>Diagnosis. This species group is characterised by the fine, pasty surface sculpture (like floury dough). Apertural barriers are never spiniferous, never hooked and not numerous: apart from the main five barriers (angular, parietal, two palatals, and a columellar), there are usually up to two more small barriers.</p><p>Remarks. This group includes 11 species. All of them inhabit Borneo, Peninsular Malaysia, and adjacent regions of Thailand. The morphology of the species within this group agrees well with the distribution pattern.</p></div>	https://treatment.plazi.org/id/BC7FB5AE88D6531187EFB079DAE51A20	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
4B7B972D41A757539C4BD3938597E898.text	4B7B972D41A757539C4BD3938597E898.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostoma W. H. Benson 1856	<div><p>Genus Hypselostoma Benson, 1856</p><p>Hypselostoma Benson, 1856 b: 342 (replacement name for Tanystoma Benson, 1856, non Motschulsky 1845, Carabidae, Coleoptera).</p><p>Gyliotrachela Tomlin, 1930: 24. (replacement name for Gyliauchen Pilsbry 1917 non Nicoll 1915, Platyhelminthes, Trematoda) syn. nov.</p><p>Antroapiculus Panha &amp; Burch, 2002 a: 144–148 . syn. nov.</p><p>Diagnosis.</p><p>This genus is subdivided into four species groups: i) H. bensonianum group ii) H. hungerfordianum group; iii) H. terae group and iv) H. tubiferum group. This subdivision is based on the shell surface sculpture (coupled with apertural barrier morphology and geographic distribution in H. terae group) which is raised spirally striated in H. bensonianum group, granulated (sandpaper-like) in H. hungerfordianum group and fine, pasty in H. terae and H. tubiferum group. Most representatives share the depressed-conical or conical shell shape with last whorl adnate to or detached from the penultimate.</p><p>Remarks.</p><p>Hypselostoma aenigma sp. nov. is provisionally placed in this genus due to the specific apertural barrier arrangement. Hypselostoma torta sp. nov. is also not a typical representative since it has non-raised spiral striae which are probably not homologous to the raised ones as usual for the genus.</p><p>Genera Gyliotrachela and Antroapiculus are both treated as junior synonyms of Hypselostoma due to the following facts: i) Antroapiculus is described based on a unique shell (flat with strongly downwards descending aperture) and a single lamella on the parietal side (parietal lamella). This however does not seem to warrant separation of a new genus since the flat shell ( H. tridentatum, H. khaowongkot, H. utongense) and strongly descending aperture ( H. srakeoense, H. chaunosalpinx, H. fungus sp. nov., H. transitans, H. torticollis) are shared with other congeners. Furthermore, apertural barrier arrangement is also similar to H. torticollis, H. fungus sp. nov., H. depressum which indicates that the combination of characters in H. pendulum could just be an autapomorphic state which could be changed rapidly during the evolution; ii) Gyliotrachela is treated distinct from Hypselostoma solely based on the separated parietal and angular lamella on the parietal side (Pilsbry 1917). Since we have found several examples of species with almost exactly the same shell morphology, but with different apertural barrier arrangements (Fig. 262), we find it less likely that they belong to different genera and find this criterium insufficient.</p><p>1. Hypselostoma bensonianum group</p><p>Diagnosis. Hypselostoma bensonianum group is characterised by raised and usually widely spaced spiral striation (in form of threads).</p><p>Remarks. A total of 46 species belong to this group. This group has a wide distribution across Southeast Asia as well as in Australia where only one species can be found. In Southeast Asia, this group spans from central and eastern Myanmar to Taiwan as well as southwards all the way to Indonesia. Northernmost localities on mainland Southeast Asia are known from Luang Prabang Province in Laos.</p></div>	https://treatment.plazi.org/id/4B7B972D41A757539C4BD3938597E898	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
210A87A628B75980882D8C1D5113D8AD.text	210A87A628B75980882D8C1D5113D8AD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypselostomatidae Zilch 1959	<div><p>Family Hypselostomatidae Zilch, 1959</p><p>Remarks.</p><p>Hypselostomatidae were originally introduced as a subfamily of Chondrinidae (Zilch 1959) . In the same work, the subfamily Aulacospirinae Zilch, 1959 was erected, later synonymised by Schileyko (1998). Bouchet et al. (2017) treated Hypselostomatinae as a subfamily of Gastrocoptidae, while some authors used Hypselostomatidae (Páll-Gergely 2023 a, b; Páll-Gergely and White 2023; Gojšina et al. 2022) Vertiginidae (Panha 1998 a; Tanmuangpak and Dumrongrojwattana 2022) or Pupillidae (Burch and Panha 2002; Panha and Burch 2002 a) to classify these representatives.</p></div>	https://treatment.plazi.org/id/210A87A628B75980882D8C1D5113D8AD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Vermeulen, Jaap J.;Reischütz, Alexander;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Páll-Gergely, Barna (2025): A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species. ZooKeys 1235: 1-338, DOI: 10.3897/zookeys.1235.145281
