taxonID	type	description	language	source
D5088799B546C76BFF22DF418A0DFC39.taxon	type_taxon	Type species. Ananteris balzanii Thorell, 1891, by original designation.	en	Botero-Trujillo, Ricardo, Flórez, Eduardo (2011): A revisionary approach of Colombian Ananteris (Scorpiones, Buthidae): two new species, a new synonymy, and notes on the value of trichobothria and hemispermatophore for the taxonomy of the group. Zootaxa 2904 (1): 1-44, DOI: 10.11646/zootaxa.2904.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2904.1.1
D5088799B546C760FF22D8428B2BFB49.taxon	description	Figures 1 – 5, 32, 40, 48, 60; Tables 1, 2	en	Botero-Trujillo, Ricardo, Flórez, Eduardo (2011): A revisionary approach of Colombian Ananteris (Scorpiones, Buthidae): two new species, a new synonymy, and notes on the value of trichobothria and hemispermatophore for the taxonomy of the group. Zootaxa 2904 (1): 1-44, DOI: 10.11646/zootaxa.2904.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2904.1.1
D5088799B546C760FF22D8428B2BFB49.taxon	materials_examined	Type material. Holotype male: COLOMBIA: Meta Department: Puerto Gaitán, Altamira, Club Los Llaneros, 04 ° 19 ’ N 72 ° 05 ’ W, 140 m asl, into forest, direct collection, at night, 19 October 2006, I. Gélvez (MPUJ-SCO- 356) (examined). Further material examined. COLOMBIA: Meta Department: 3 adult males, 1 adult female, 1 subadult female, Puerto López, Remolinos, Cafam Llanos, 04 ° 16 ’ 28 ’’ N 72 ° 32 ’ 19 ’’ W, 183 m asl, UV light, 03 – 05 September 2008, J. A. Ochoa & R. Botero-Trujillo (MPUJ-SCO- 383). 2 adult males, 2 adult females, Puerto López, Remolinos, Cafam Llanos, 04 ° 16 ’ 28 ’’ N 72 ° 32 ’ 19 ’’ W, 183 m asl, UV light, 03 – 05 September 2008, J. A. Ochoa & R. Botero-Trujillo (ICN-As- 773). 6 adult males, 2 adult females, between Villavicencio and Restrepo, Farm Con Esto Tengo, 04 ° 11 ’ 51 ’’ N 73 ° 35 ’ 37 ’’ W, 407 m asl, UV light, 07 September 2008, R. Botero-Trujillo & J. A. Ochoa (MPUJ-SCO- 384). 7 adult males, 1 adult female, 1 juvenile female, between Villavicencio and Restrepo, Farm Con Esto Tengo, 04 ° 11 ’ 51 ’’ N 73 ° 35 ’ 37 ’’ W, 407 m asl, UV light, 07 September 2008, R. Botero-Trujillo & J. A. Ochoa (ICN-As- 774). 1 adult male, Serranía de La Macarena, 05 March 1992 (ICN-As- 641). 1 adult male, 2 adult females, San Martín, Rey Zamuro Natural Reserve, 03 ° 31 ’ 57.46 ’’ N 73 ° 30 ’ 52.35 ’’ W, 317 m asl, UV light, March 2010, J. A. Noriega (MPUJ-SCO- 398). Revised diagnosis. Carapace with well-developed anteromedian projection (Fig. 32); metasomal carinal formula 10: 10: 10: 6: 5, with ventrosubmedian and median lateral carinae absent on segment IV; median lateral carinae on metasomal segment III only weakly developed, evident in females but vestigial in males, especially posteriorly; V 1 and V 2 trichobothria unaligned axially, with V 2 located on an external position in relation to V 1 (Fig. 40); fixed finger trichobothria in the order eb: esb: est / db: et: dt, with est located beside db or nearly so (Fig. 48); dorsal surface of chelicerae with incomplete reticular pattern (Fig. 32); pectines with 17 – 21 teeth on males (mode = 18), 16 – 18 on females (mode = 18); sternite VII with paramedian longitudinal carinae vestigial and incomplete; pedipalp hand base color yellow most usually with conspicuous brown areas ventrally, internally and externally (brown may be absent on the external surface), but infrequently completely yellow; coxosternal region entirely yellow and spotless. Hemispermatophore: capsular region with median, external and internal lobes; flagellum with long and strongly coiled pars reflexa (Fig. 60). Male. Holotype described by Botero-Trujillo (2008 a) (Figs. 2 – 3). Hemispermatophore: Description based on specimens from MPUJ-SCO- 383 and ICN-As- 774: Flagelliform, thin and poorly sclerotized. Foot narrow and flat. Pedal flexure inconspicuous but movable. Body very long, wider on basal third. Capsular region with median, external and internal lobes, subequal and strong. Flagellum long, with short pars recta and long, coiled pars reflexa (Fig. 60). Carapace length to hemispermatophore body length ratio = 1: 1.08 to 1: 1.04. There was no variation between the four hemispermatophores examined. Female. Description based on adult female from MPUJ-SCO- 383. Measurements: see Table 2. Coloration: general coloration yellowish with variegated pigmentation over almost the entire body and appendages (Figs. 4 – 5). Carapace predominantly brown with some yellow spots and stripes; anterior and posterior margins brown; arising posteriorly to each lateral ocular tubercle there is a thin and almost straight yellow stripe that ends behind the median ocular tubercle; median ocular tubercle black, surrounded on its base by thin yellow lines on anterior, anterolateral and posterolateral margins. Chelicerae with coxa predominantly yellow with abundant pale-brown mottling; hand with incomplete reticular pattern but abundant reticulations on dorsal surface (Fig. 32); fixed finger yellow with reddish teeth, movable finger with a basal brown area and reddish teeth. Coxosternal region, genital operculum, pectinal basal piece, pectines and sternite III entirely yellow and spotless; sternites IV – VI predominantly yellow, with some minute and inconspicuous brownish spots paramedially; sternite VII with large darkbrown lateral areas; sternite V with a small flattened posterior median hyaline area. Tergites predominantly brown; twο ¹ οngitudina ¹ ye ¹¹ οw StripeS crοSSing tergiteS I — VI are οn ¹ y evident οn the pοSteriοr ha ¹ f οf each Segment beSide the midline; each side of tergites I – VI with two transverse yellow lines converging near the longitudinal lines, arrow-like; tergite VII with variegated pigmentation; lateral margins of tergites I – VII yellow, with an anterior small brown area. Metasoma predominantly yellowish; dorsal intercarinal spaces of segments I – IV with a median brown design wider anteriorly, triangle-like on segments I – III but arrow-like on segment IV; segments I – V with variegated pigmentation on all surfaces; segment V reddish with brownish areas. Telson vesicle yellowish-red with brown areas ventrally and laterally; subaculear tubercle yellowish. Pedipalp femur and patella predominantly brown dorsally and externally, with conspicuous yellow areas; primarily yellow ventrally and externally with abundant brown areas; both segments with trichobothrial pits yellow. Pedipalp coxa and trochanter yellow with large brown regions dorsally; hand with base color yellow and conspicuous brown areas ventrally, internally and externally; fingers entirely brown. Legs with variegated pigmentation. Carapace: densely covered with rounded granules especially on the brown areas; lateral margins converging anteriorly; anterior margin with well-developed median projection (Fig. 32); ocular carinae moderately strong and granulose, others inconspicuous; with a median depression anterior to the ocular tubercle; postocular furrow evident; median ocular tubercle low, located on the posterior half of the anterior third of carapace; lateral ocular tubercles each with three evident ocelli. Chelicerae: with abundant fine white setae on the internal and ventral surfaces; cheliceral dentition characteristic of the family Buthidae (Vachon 1963). Movable finger dorsally with two small basal teeth, one median pronounced, one subdistal slightly shorter than the median, and one distal tooth. Ventrally with three strong teeth, one basal, one median, and one distal tooth larger than its dorsal counterpart. Fixed finger dorsally with one basal and one median tooth forming a bicuspid, one subdistal, and one distal tooth. Ventrally with only one subdistal tooth slightly basal to the dorsal subdistal. Coxosternal region: sternum subtriangular, with deep median depression and two anterolateral furrows; all the components of this region smooth, with few setae; coxapophyses I – II densely setose anteriorly. Genital operculum and pectines: genital operculum completely divided longitudinally; pectinal basal piece as long as wide, pentagonal and deeply notched anteriorly; pectines small, not reaching posterior end of coxae IV; count of pieces on the pectines: marginal lamellae 3: 3, middle lamellae 8: 10, teeth 17: 18, fulcra absent. Sternites: with abundant setae; III – VI smooth, except for some granulations on the lateral margins of V – VI; VII with vestigial and incomplete paramedian longitudinal carinae and some granules on the brown regions, lateral carinae completely absent; V with a small flattened posterior median smooth area; spiracles oval elongated, approximately twice longer than wide. Tergites: with similar granulation to that of carapace; axial carina only evident on the posterior half of tergites II – VI; obsolete dorsolateral carinae present on tergites II – VI; tergite VII tetracarinate, with incomplete paired dorsolateral and lateral carinae, and low granulose median elevation on the position of the axial carina. Metasoma: with abundant setae; segments I – III with ten carinae (paired ventrosubmedian, ventrolateral, median lateral, dorsolateral and dorsosubmedian carinae); segment IV with six (ventrosubmedian and median lateral carinae absent); segment V with five (ventromedian, paired ventrolateral and dorsolateral carinae); ventrolateral and median lateral carinae converge distally in segments I – II; all carinae serrulate; intercarinal spaces with abundant granulation. Telson: vesicle elongated, smooth dorsally, with three ventral longitudinal carinae made up of conspicuous granulation; subaculear tubercle strong and spine-like; aculeus long and curved. Pedipalps: femur with five vestigial longitudinal carinae (dorsoexternal, dorsointernal, ventroexternal, ventrointernal and internal median carinae); patella without distinct carinae but with few small granules on the position of the dorsointernal and ventrointernal carinae; chela acarinate; fixed finger with six almost linear rows of granules, being the basal the longest; movable finger with seven rows (including the short apical row). Trichobothriotaxy type A, femur with β configuration (Vachon 1974, 1975). Legs: tibia, basitarsus and telotarsus with numerous ventral setae; tibial spur present on legs III – IV; prolateral pedal spur single on legs I – II, bifid on legs III – IV; retrolateral pedal spur present on all legs. Variability. Fixed finger trichobothria (n = 62): 37 (59.7 %) fingers with trichobothria in the order eb: esb: est / db: et: dt; 17 (27.4 %) in the order eb: esb: est: db: et: dt, with est very slightly basal to db; seven (11.3 %) in the order eb: esb: db: est: et: dt, with db very slightly basal to est; one (1.6 %) lacking est (anomalous condition). Cheliceral reticulation (n = 62): incomplete reticular pattern present on all chelicerae studied, but slightly less reticulated in the holotype (3.2 %), probably due to normal interpopulational variability. Pectinal teeth count: 16 to 18 on females (n = 20; mode = 18); 17 to 21 on males (n = 42; mode = 18). Coloration of the pedipalp hand (n = 62): 50 (80.6 %) hands with brown areas ventrally, internally and externally; ten (16.1 %) completely yellow; two (3.2 %) with brown areas ventrally and internally. Total body length (including telson): male 21 – 26 mm, female 27 – 33 mm. Anomalies. See ‘ Fixed finger trichobothria’ in the ‘ variability’ section above.	en	Botero-Trujillo, Ricardo, Flórez, Eduardo (2011): A revisionary approach of Colombian Ananteris (Scorpiones, Buthidae): two new species, a new synonymy, and notes on the value of trichobothria and hemispermatophore for the taxonomy of the group. Zootaxa 2904 (1): 1-44, DOI: 10.11646/zootaxa.2904.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2904.1.1
D5088799B546C760FF22D8428B2BFB49.taxon	distribution	Distribution. So far known from five localities in Meta Department (Fig. 1). Probably widely distributed within this department’s limits and a typical element of open savanna-like formations. New locality records: COLOMBIA: Meta Department: i) Puerto López; ii) Villavicencio – Restrepo; iii) Serranía de La Macarena; iv) San Martín. Ecological notes. All specimens of A. arcadioi from ‘ Cafam Llanos’ and ‘ Con Esto Tengo’ were hand-captured at night, after detection with UV light when actively moving. In the former locality, most specimens were found in leaf litter or sandy walls of small streams inside gallery forests, whereas few were collected in borders of forests surrounded by extensive grasslands. In the latter locality, some specimens were found on the walls of a stream or the leaf litter close to it, but most were found on the border of a road in an area strongly influenced by human activities. Even though A. arcadioi was easily encountered in both places, it was noticeably less abundant than Tityus bastosi Lourenço, 1984 in ‘ Cafam Llanos’ and Tityus nematochirus Mello-Leitão, 1940 in ‘ Con Esto Tengo’, species with which it was found in sympatry. However, Ananteris specimens were very scarce in the places where Tityus Koch, 1836 was abundant.	en	Botero-Trujillo, Ricardo, Flórez, Eduardo (2011): A revisionary approach of Colombian Ananteris (Scorpiones, Buthidae): two new species, a new synonymy, and notes on the value of trichobothria and hemispermatophore for the taxonomy of the group. Zootaxa 2904 (1): 1-44, DOI: 10.11646/zootaxa.2904.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2904.1.1
D5088799B54DC760FF22D8838A34FA45.taxon	description	Figures 1, 6 – 9, 33, 41, 49, 61; Table 1	en	Botero-Trujillo, Ricardo, Flórez, Eduardo (2011): A revisionary approach of Colombian Ananteris (Scorpiones, Buthidae): two new species, a new synonymy, and notes on the value of trichobothria and hemispermatophore for the taxonomy of the group. Zootaxa 2904 (1): 1-44, DOI: 10.11646/zootaxa.2904.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2904.1.1
D5088799B54BC779FF22DEA08BD0FCEC.taxon	description	Figures 1, 10 – 13, 34, 42, 50, 56 – 57, 62; Tables 1, 2	en	Botero-Trujillo, Ricardo, Flórez, Eduardo (2011): A revisionary approach of Colombian Ananteris (Scorpiones, Buthidae): two new species, a new synonymy, and notes on the value of trichobothria and hemispermatophore for the taxonomy of the group. Zootaxa 2904 (1): 1-44, DOI: 10.11646/zootaxa.2904.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2904.1.1
D5088799B54BC779FF22DEA08BD0FCEC.taxon	materials_examined	Type material. Holotype female: COLOMBIA: Nariño Department: La Planada Natural Reserve, permanent plot, 01 ° 15 ’ N 78 ° 15 ’ W, 1885 m asl, pitfall trap, 02 – 04 May 2001, G. Oliva, M. 2369 (IAvH-E 100763) (examined). Further material examined. COLOMBIA: Nariño Department: 1 adult male, Barbacoas, Path El Barro, Río Ñambi Natural Reserve, 01 ° 17 ’ 10 ’’ N 78 ° 04 ’ 28 ’’ W, 1397 m asl, UV light, 11 September 2008, J. A. Ochoa, R. Botero-Trujillo, J. P. Botero, J. Souza & O. Tovar (MPUJ-SCO- 385). 1 adult female, 1 juvenile female, Barbacoas, Path El Barro, Río Ñambi Natural Reserve, 01 ° 17 ’ 10 ’’ N 78 ° 04 ’ 2 ’’ ' W, 1397 m asl, UV light, 11 September 2008, O. Tovar, J. Souza, R. Botero-Trujillo, J. A. Ochoa & J. P. Botero (ICN-As- 775). 1 adult female, 3 adult males, Barbacoas, Altaquer, Río Ñambi Natural Reserve, 1440 m asl, UV light, 13 October 2009, E. Flórez & D. Luna (ICN- As- 816). 4 adult females, Ricaurte, Path Alto Cartagena, Farm Nueva Estrella, 01 ° 13 ’ 16 ’’ N 77 ° 58 ’ 08 ’’ W, 1617 m asl, UV light, 12 September 2008, R. Botero-Trujillo, J. A. Ochoa, J. P. Botero, N. Rosales & C. Sevillano (MPUJ- SCO- 386). 3 adult females, 1 subadult female, Ricaurte, Path Alto Cartagena, Farm Nueva Estrella, 01 ° 13 ’ 16 ’’ N 77 ° 58 ’ 08 ’’ W, 1617 m asl, UV light, 12 September 2008, N. Rosales, C. Sevillano, R. Botero-Trujillo, J. P. Botero & J. A. Ochoa (ICN-As- 776). Revised diagnosis. Unique metasomal carinal formula 10: 10: 10: 6: 3, with median lateral carinae on segment III less pronounced than others and sometimes apparently absent, ventrosubmedian and median lateral carinae absent on segment IV, and ventrolateral carinae absent on segment V. Other features include: carapace weakly, evenly concave and lacking anteromedian projection (Fig. 34); metasomal segments with abundant and strong granulation that obscures the carinae and make them difficult to distinguish; V 1 and V 2 trichobothria unaligned axially, with V 2 located on an external position in relation to V 1 (Fig. 42); fixed finger trichobothria in the order eb: esb: db: est: et: dt (Fig. 50); dorsal surface of chelicerae with dense and complete reticular pattern (Fig. 34); pectines with 13 – 14 teeth on males (mode = 13), 12 – 13 on females (mode = 13); sternite VII with paramedian longitudinal carinae obsolete and incomplete, formed by scarcely aggregated granules that may eventually suggest they are absent; pedipalp hand uniformly yellow and spotless; coxosternal region base color yellow with diffuse brownish pigmentation that is darker and more conspicuous in coxa and coxapophysis of legs I. Hemispermatophore: capsular region with three lobes, the basal armed with three minute spines; flagellum with very short pars reflexa (Fig. 62). Male. Description based on adult male from ICN-As- 816. Measurements: see Table 2. Coloration: general coloration brownish with variegated pigmentation over almost the entire body and appendages (see Figs. 10 – 11 for MPUJ-SCO- 385). Carapace predominantly brown with some yellow spots and stripes; anterior and posterior margins dark-brown; arising posteriorly to each lateral ocular tubercle there is a thin and almost straight yellow stripe that ends behind the median ocular tubercle; median ocular tubercle black, surrounded on its base by thin yellow lines on anterior, anterolateral and posterolateral margins. Chelicerae with yellow coxa; hand with dense and complete brown reticular pattern on dorsal view (see Fig. 34 for MPUJ-SCO- 385); fixed and movable fingers entirely brown with reddish teeth. Coxosternal region and genital operculum predominantly yellow, with diffuse brownish pigmentation throughout; pectinal basal piece, pectines and sternite III completely yellow; sternites IV – VI with inconspicuous lateral minute brownish spots; sternite VII with light-brown lateral regions; sternite V with a flattened posterior median hyaline area. Tergites predominantly brown; tergites I – VI each with two paramedian yellow spots in anterior position and two transverse yellow lines converging near the midline (arrow-like); tergite VII predominantly brownish but with large yellow areas; lateral margins of tergites I – VII brown. Metasoma predominantly yellowish to reddish; dorsal intercarinal spaces of segments I – IV with a median brown design wider anteriorly, triangle-like on segments I – III but arrow-like on segment IV; all segments with variegated pigmentation on every surfaces; segment V reddish, darker than the preceding segments. Telson vesicle and base of aculeus yellowish, with one thin brown longitudinal band ventromedially, one large ventrolaterally on each side, and a broad one dorsally; distal half of aculeus red; subaculear tubercle yellowish. Pedipalp femur and patella completely brown, with some slightly lighter regions in all surfaces and trichobothrial pits yellow; coxa and trochanter with variegated pigmentation; chela with hand uniformly yellow, fingers predominantly brown (except basally) with yellow trichobothrial pits. Legs with trochanter and telotarsi predominantly yellow, other segments primarily brown. Carapace: densely covered with conspicuous rounded granules especially on the brown areas; lateral margins converging anteriorly; anterior margin weakly, evenly concave and lacking anteromedian projection (see Fig. 34 for MPUJ-SCO- 385); ocular carinae evident, others inconspicuous; with a median depression anterior to the ocular tubercle; postocular furrow evident; median ocular tubercle low, located on the posterior half of the anterior third of carapace; lateral ocular tubercles each with three evident ocelli. Chelicerae: with abundant fine white setae on the internal and ventral surfaces; cheliceral dentition characteristic of the family Buthidae (Vachon 1963). Movable finger dorsally with two small basal teeth, one median pronounced, one subdistal slightly shorter than the median, and one distal tooth. Ventrally with three strong teeth, one basal, one median, and one distal tooth larger than its dorsal counterpart. Fixed finger dorsally with one basal and one median tooth forming a bicuspid, one subdistal, and one distal tooth. Ventrally with only one subdistal tooth slightly basal to the dorsal subdistal. Coxosternal region: sternum subpentagonal, with deep median depression and two anterolateral furrows; all the components of this region smooth, with few setae; coxapophyses I – II densely setose anteriorly. Genital operculum and pectines: genital operculum completely divided longitudinally; pectinal basal piece heart-shaped, slightly longer than wide and deeply notched anteriorly; pectines long, with distal tip of marginal lamella reaching posterior end of coxae IV; count of pieces on the pectines: marginal lamellae 3: 3, middle lamellae 7: 7, teeth 13: 14, fulcra absent. Sternites: III – VI smooth; VII with conspicuous but weak granulation and paramedian longitudinal carinae obsolete and incomplete, lateral carinae completely absent; III – VII with abundant setae, especially on lateral margins; V with flattened posterior median smooth area; spiracles elongated, approximately twice longer than wide in sternites III – IV and three times longer than wide in V – VI. Tergites: with similar granulation to that of carapace; axial carina only evident on the posterior half of tergites II – VI; dorsolateral carinae completely absent on tergites I – VI; tergite VII tetracarinate, with incomplete paired dorsolateral and lateral carinae, and median elevation on the position of the axial carina. Metasoma: with abundant setae; segments I – III with ten carinae (paired ventrosubmedian, ventrolateral, median lateral, dorsolateral and dorsosubmedian carinae); segment IV with six (ventrosubmedian and median lateral carinae absent); segment V with three (ventromedian, paired dorsolateral carinae); ventrolateral and median lateral carinae converge distally in segments I – II; ventrosubmedian carinae not parallel in segment I but arranged into S-shape, separated from each other by a greater distance anteriorly than posteriorly; ventrosubmedian carinae connected to each other and to ventrolateral carinae by a transverse row of granules in segments II – III; all carinae granulose, formed by unconnected granules (except for dorsolateral and dorsosubmedian carinae in segments I – IV whose granules are close together); intercarinal spaces with abundant strong granulation. Telson: vesicle moderately elongated, smooth dorsally, with three granulose ventral carinae and lateral granulations; subaculear tubercle strong and spine-like; aculeus long and curved. Pedipalps: femur with five vestigial longitudinal carinae (dorsoexternal, dorsointernal, ventroexternal, ventrointernal and internal median carinae); patella without distinct carinae but with few granules on the position of the dorsointernal carina; chela acarinate; fixed finger with six almost linear rows of granules, being the basal the longest; movable finger with seven rows (including the short apical row). Trichobothriotaxy type A, femur with β configuration (Vachon 1974, 1975). Legs: tibia, basitarsus and telotarsus with numerous ventral setae; tibial spur present on legs III – IV; prolateral pedal spur single on legs I – II, bifid on legs III – IV; retrolateral pedal spur present on all legs. Hemispermatophore: Description based on one specimen from ICN-As- 816: Flagelliform, thin and poorly sclerotized. Foot narrow and flat. Pedal flexure inconspicuous but movable. Body very long, slightly wider on basal third. Capsular region with a large basal lobe armed with three minute spines distally, an external lobe, and a distinctly curved and well-developed internal lobe. Flagellum with long pars recta and very short pars reflexa (Fig. 62). Carapace length to hemispermatophore body length ratio = 1: 1.24. There was no variation between both hemispermatophores examined. Female. Holotype described by Botero-Trujillo (2008 a). Variability. Pectinal teeth count: 12 to 13 on females (n = 23; mode = 13); 13 to 14 on males (n = 7; mode = 13). Note that one female has ten teeth on the left pectine, but this low number is not included in the count because such pectine lacks the distal piece of the marginal lamella (anomalous condition). Note also that one male has the left pectine damaged and incomplete, with only four basal teeth, and therefore is excluded from the count. Carination of sternite VII (n = 16): paramedian longitudinal carinae are obsolete in most specimens studied (93.8 %), but are apparently absent in the holotype female (6.2 %). Total body length (including telson): male 14 – 16 mm, female 19 – 25 mm. Anomalies. One female lacks V 2 trichobothrium in the right hand. See ‘ Pectinal teeth count’ in the ‘ variability’ section above for further anomalies.	en	Botero-Trujillo, Ricardo, Flórez, Eduardo (2011): A revisionary approach of Colombian Ananteris (Scorpiones, Buthidae): two new species, a new synonymy, and notes on the value of trichobothria and hemispermatophore for the taxonomy of the group. Zootaxa 2904 (1): 1-44, DOI: 10.11646/zootaxa.2904.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2904.1.1
D5088799B54BC779FF22DEA08BD0FCEC.taxon	distribution	Distribution. So far known only from three localities in Nariño Department (Fig. 1). New locality records: COLOMBIA: Nariño Department: i) Barbacoas (Figs. 56 – 57); ii) Ricaurte. Ecological notes. All specimens of A. dorae from ‘ Río Ñambi’ and ‘ Nueva Estrella’ were hand-captured at night, after detection with UV light when actively moving in the leaf litter or fallen trunks. In the former locality, A. dorae was found in sympatry with A. ochoai sp. nov., Tityus sp. and Teuthraustes sp. In the latter, it was found with Troglotayosicus humiculum Botero-Trujillo & Francke, 2009, Tityus sp. (presumably Tityus cuellari Lourenço, 1994) and Teuthraustes sp. (Ochoa et al. 2010).	en	Botero-Trujillo, Ricardo, Flórez, Eduardo (2011): A revisionary approach of Colombian Ananteris (Scorpiones, Buthidae): two new species, a new synonymy, and notes on the value of trichobothria and hemispermatophore for the taxonomy of the group. Zootaxa 2904 (1): 1-44, DOI: 10.11646/zootaxa.2904.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2904.1.1
D5088799B554C77CFF22DFF98A92FD71.taxon	description	Figures 1, 14 – 17, 35, 43, 51, 63; Tables 1, 2	en	Botero-Trujillo, Ricardo, Flórez, Eduardo (2011): A revisionary approach of Colombian Ananteris (Scorpiones, Buthidae): two new species, a new synonymy, and notes on the value of trichobothria and hemispermatophore for the taxonomy of the group. Zootaxa 2904 (1): 1-44, DOI: 10.11646/zootaxa.2904.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2904.1.1
D5088799B554C77CFF22DFF98A92FD71.taxon	materials_examined	Type material. Holotype female: COLOMBIA: Caquetá Department: Florencia, Path El Paraiso, School El Paraiso, 850 m asl, June 1990, P. Ruiz-C. (ICN-As- 007) (examined). Further material examined. COLOMBIA: Caquetá Department: 1 adult female, La Montañita, Santuario Las Iglesias, Itarca, 330 m asl, 25 April 2004, M. Agudelo (ICN-As- 579). 1 adult female, Chiribiquete Natural National Park, River Mesay, 20 January 2000, F. Quevedo (ICN-As- 361). 1 adult female, Chiribiquete Natural National Park, River Sararamano, April 2000, F. Quevedo (ICN-As- 342). 1 adult female, Solano, Chiribiquete Natural National Park, Bosque Amarillo, pitfall trap with decomposing fish, 24 – 26 February 2001, Cuñare Expedition (IAvH-E 100813). 1 adult male, Solano, Chiribiquete Natural National Park, River Sararamano, Bosque Verde Oscuro, pitfall trap, 11 April 2000 (IAvH-E 100814). 1 adult male, Solano, Chiribiquete Natural National Park, River Sararamano, Bosque Verde Militar, 0 ° 10 ’ 47 ’’ N 72 ° 37 ’ 24 ’’ W, 300 m asl, Winkler trap, 03 – 07 April 2000, E. González (IAvH-E 100815). 1 adult male, Solano, Chiribiquete Natural National Park, River Amú, Bosque Naranja, trap for dung beetles, 01 – 03 March 2001 (IAvH-E 100816). Revised diagnosis. Carapace weakly, evenly concave and lacking anteromedian projection (Fig. 35); metasomal carinal formula 10: 10: 8: 8: 5; telson vesicle with three ventral longitudinal carinae made up of weak granulation; males with telson noticeably elongated; V 1 and V 2 trichobothria arranged linearly parallel to the external surface of the chela (aligned axially) (Fig. 43); fixed finger trichobothria in the order eb: esb: est / db: et: dt, with est located beside db or nearly so (Fig. 51); dorsal surface of chelicerae with dense and almost complete reticular pattern, except on anterior margin where immaculately yellow and spotless (Fig. 35); pectines with 23 – 24 teeth on males (mode = 24), 21 – 23 on females (mode = 22); sternite VII with incomplete but strong paramedian longitudinal carinae; pedipalp hand base color yellow with conspicuous brown areas ventrally, internally, dorsally and externally; coxosternal region almost entirely yellow and spotless, except for coxapophyses I brownish anteriorly. Hemispermatophore: capsular region with two lobes; flagellum with long pars recta and coiled pars reflexa (Fig. 63). Male. Description based on IAvH-E 100814. Measurements: see Table 2. Coloration: general coloration yellowish to reddish-brown with variegated pigmentation over almost the entire body and appendages (Figs. 14 – 15). Carapace predominantly brown with some yellow spots and stripes; anterior margin yellow, posterior margin brown; arising posteriorly to each lateral ocular tubercle there is a thin and almost straight yellow stripe that ends behind the median ocular tubercle; median ocular tubercle black, surrounded on its base by thin yellow lines on anterolateral and posterolateral margins. Chelicerae with yellow coxa; hand with dense and almost complete reticular pattern on dorsal surface, except anteriorly where immaculately yellow and spotless (Fig. 35); fixed finger yellow and movable finger predominantly brown, each with reddish teeth. Coxosternal region, genital operculum, pectinal basal piece, pectines and sternite III entirely yellow, except for coxapophyses I with anterior brownish spots; sternites IV – VII with inconspicuous brownish areas posteriorly; sternite V with a subtriangular-to-flattened posterior median hyaline area. Tergites predominantly brown; two longitudinal yellow stripes crossing tergites I – VI are only evident on the posterior half of each segment beside the midline; each side of tergites I – VI with two transverse yellow lines converging near the longitudinal lines, arrow-like; tergite VII predominantly yellow, with large brown regions laterally; lateral margins of tergites I – VII completely yellow. Metasoma predominantly yellowish-red; dorsal intercarinal spaces of segments I – IV with a median brown design wider anteriorly, triangle-like on segments I – II, arrow-like on segments III – IV; segments I – IV with variegated pigmentation on all surfaces; segment V dark-red. Telson vesicle reddish, lighter than segment V and with inconspicuous brownish spots ventroexternally; aculeus dark-red distally, yellowish basally; subaculear tubercle yellowish-red. Pedipalps predominantly brown; coxa and trochanter with variegated pigmentation; femur brown dorsally and internally but predominantly yellow ventrally and externally, trichobothrial pits yellow; patella with variegated pigmentation on dorsal, external and internal surfaces where predominantly brown, yellowish ventrally, with trichobothrial pits yellow; hand primarily yellow but with conspicuous brown areas on all surfaces; fixed and movable fingers entirely brown. Legs with variegated pigmentation from femur to basitarsi, other segments yellow. Carapace: densely covered with minute rounded granules especially on the brown areas; lateral margins converging anteriorly; anterior margin weakly, evenly concave and lacking anteromedian projection (Fig. 35); ocular carinae strong and granulose, others inconspicuous; with a median depression anterior to the ocular tubercle; postocular furrow evident; median ocular tubercle low, located on the posterior half of the anterior third of carapace; lateral ocular tubercles each with four ocelli, being the posteriormost very reduced and unaligned with the remaining three pairs. Chelicerae: with abundant fine white setae on the internal and ventral surfaces; cheliceral dentition characteristic of the family Buthidae (Vachon 1963). Movable finger dorsally with two small basal teeth, one median pronounced, one subdistal slightly shorter than the median, and one distal tooth. Ventrally with three strong teeth, one basal, one median, and one distal tooth larger than its dorsal counterpart. Fixed finger dorsally with one basal and one median tooth forming a bicuspid, one subdistal, and one distal tooth. Ventrally with only one subdistal tooth located at the same level of the dorsal subdistal. Coxosternal region: sternum subtriangular, with deep median depression and two anterolateral furrows; all the components of this region scarcely granulose, with abundant setae; coxapophyses I – II densely setose anteriorly. Genital operculum and pectines: genital operculum completely divided longitudinally; pectinal basal piece as long as wide, deeply notched anteriorly; pectines long, with distal tip of marginal lamella reaching posterior end of coxae IV; count of pieces on the pectines: marginal lamellae 3: 3, middle lamellae 8: 8, teeth 23: 24, fulcra absent. Sternites: with dense granulation throughout, primarily on lateral margins, and abundant setae; VII with incomplete but strong paramedian longitudinal carinae, lateral carinae completely absent; V with subtriangular-to-flattened posterior median smooth area; spiracles elongated, approximately three times longer than wide in sternites III – V and four times longer than wide in VI. Tergites: with similar granulation to that of carapace; axial carina only evident on the posterior half of tergites I – VI; dorsolateral carinae completely absent on tergites I – VI; tergite VII tetracarinate, with incomplete paired dorsolateral and lateral carinae, and granulose median elevation on the position of the axial carina. Metasoma: with few long setae; segments I – II with ten carinae (paired ventrosubmedian, ventrolateral, median lateral, dorsolateral and dorsosubmedian carinae); segments III – IV with eight (median lateral carinae absent); segment V with five (ventromedian, paired ventrolateral and dorsolateral carinae); ventrolateral and median lateral carinae converge distally in segments I – II; ventrosubmedian and ventrolateral carinae connected anteriorly by a transverse row of granules in segments II – IV; all carinae serrulate; intercarinal spaces with abundant rounded granules. Telson: vesicle noticeably elongated; tegument almost completely smooth except for three ventral longitudinal carinae made up of weak granulation; subaculear tubercle strong and spine-like; aculeus long and curved. Pedipalps: with fine granulation on dorsal and internal surfaces of femur and patella; femur with five longitudinal carinae (dorsoexternal, dorsointernal, ventroexternal, ventrointernal and internal median carinae); patella without distinct carinae but with few granules on the position of dorsointernal and ventrointernal carinae; chela acarinate; fixed finger with six almost linear rows of granules, being the basal the longest; movable finger with seven rows (including the short apical row). Trichobothriotaxy type A, femur with β configuration (Vachon 1974, 1975). Legs: tibia, basitarsus and telotarsus with numerous ventral setae; tibial spur present on legs III – IV; prolateral pedal spur single on legs I – II, bifid on legs III – IV; retrolateral pedal spur present on all legs. Hemispermatophore: Description based on IAvH-E 100815 and IAvH-E 100816: Flagelliform, thin and sclerotized. Foot narrow and flat. Pedal flexure inconspicuous but movable. Body long, slightly wider on basal third. Capsular region with a large, markedly sclerotized basal lobe, and an internal lobe. Flagellum with long pars recta and long, coiled pars reflexa (Fig. 63). Carapace length to hemispermatophore body length ratio = 1: 1.45 to 1: 1.53. There was no variation between the four hemispermatophores examined. Female. Holotype described by Lourenço (1994). Variability. Fixed finger trichobothria (n = 16): six (37.5 %) fingers with trichobothria in the order eb: esb: est / db: et: dt; six (37.5 %) in the order eb: esb: db: est: et: dt, with db very slightly basal to est; four (25 %) in the order eb: esb: est: db: et: dt, with est very slightly basal to db. Pectinal teeth count: 21 to 23 on females (n = 10; mode = 22); 23 to 24 on males (n = 6; mode = 24). Total body length (including telson): male 31 – 32 mm, female 29 – 36 mm.	en	Botero-Trujillo, Ricardo, Flórez, Eduardo (2011): A revisionary approach of Colombian Ananteris (Scorpiones, Buthidae): two new species, a new synonymy, and notes on the value of trichobothria and hemispermatophore for the taxonomy of the group. Zootaxa 2904 (1): 1-44, DOI: 10.11646/zootaxa.2904.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2904.1.1
D5088799B554C77CFF22DFF98A92FD71.taxon	distribution	Distribution. So far known from four localities in Caquetá Department (Fig. 1), probably widely distributed within this department’s limits. Ecological notes. No information hitherto exists to date for this species.	en	Botero-Trujillo, Ricardo, Flórez, Eduardo (2011): A revisionary approach of Colombian Ananteris (Scorpiones, Buthidae): two new species, a new synonymy, and notes on the value of trichobothria and hemispermatophore for the taxonomy of the group. Zootaxa 2904 (1): 1-44, DOI: 10.11646/zootaxa.2904.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2904.1.1
D5088799B551C77EFF22DE8B8B44FB31.taxon	description	Figures 1, 18 – 21, 36, 44, 52, 64	en	Botero-Trujillo, Ricardo, Flórez, Eduardo (2011): A revisionary approach of Colombian Ananteris (Scorpiones, Buthidae): two new species, a new synonymy, and notes on the value of trichobothria and hemispermatophore for the taxonomy of the group. Zootaxa 2904 (1): 1-44, DOI: 10.11646/zootaxa.2904.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2904.1.1
D5088799B551C77EFF22DE8B8B44FB31.taxon	materials_examined	Type material. Holotype female: VENEZUELA: Mérida State: Antonio Pinto Salinas, 2 km from Mesa Bolívar to Libertad, 08 ° 28 ’ 36 ’’ N 71 ° 35 ’ 30 ’’ W, 1000 m asl, 23 December 1982, A. R. Delgado de González, J. A. González D. & M. A. González-Sponga (MAGS- 4420) (not examined). Paratypes: 1 male (MAGS- 3333) and 1 subadult female (MAGS- 4421) with identical data to the holotype (not examined). MAGS: Manuel Angel González- Sponga’s collection, Venezuela. Material examined. COLOMBIA: Norte de Santander Department: 1 adult male, Toledo, Tamá Natural National Park, 1000 m asl, direct collection, 09 October 1999, V. Rodríguez (ICN-As- 291). 1 adult male, Toledo, Tamá Natural National Park, 1430 m asl, 29 September 1999, V. Rodríguez (ICN-As- 435). 1 adult female, Toledo, Tamá Natural National Park, 1000 m asl, direct collection, 26 September 1999, Valeria (ICN-As- 320). 2 adult females, Toledo, Tamá Natural National Park, Path El Diamante, Alto de La Herrera, 1000 m asl, 28 September 1999, V. R. Mayaso (ICN-As- 321). 1 juvenile male, Toledo, Tamá Natural National Park, forest, 1000 m asl, September 1999, E. González (IAvH-E 100817). Revised diagnosis. Carapace with well-developed anteromedian projection (Fig. 36); metasomal carinal formula 10: 10: 10: 6: 5, with ventrosubmedian and median lateral carinae absent on segment IV; median lateral carinae on metasomal segment III only weakly developed, evident in females but vestigial in males, especially posteriorly; ventrosubmedian carinae on metasomal segment III only present in the anterior half; V 1 and V 2 trichobothria markedly unaligned axially, with V 2 located on an external position in relation to V 1 (Fig. 44); fixed finger trichobothria in the order eb: esb: est / db: et: dt, with est located beside db or nearly so (Fig. 52); dorsal surface of chelicerae predominantly yellow, almost without brown reticulations except anteriorly (Fig. 36); pectines with 17 – 19 teeth on both males (mode = 18) and females (mode = 18); sternite VII with paramedian longitudinal carinae vestigial and incomplete; pedipalp hand uniformly yellow and spotless; coxosternal region base color yellow with diffuse brownish pigmentation that is more conspicuous in coxapophyses I and coxa of legs I – II. Hemispermatophore: capsular region with median, external and internal lobes; flagellum with long and strongly coiled pars reflexa (Fig. 64).	en	Botero-Trujillo, Ricardo, Flórez, Eduardo (2011): A revisionary approach of Colombian Ananteris (Scorpiones, Buthidae): two new species, a new synonymy, and notes on the value of trichobothria and hemispermatophore for the taxonomy of the group. Zootaxa 2904 (1): 1-44, DOI: 10.11646/zootaxa.2904.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2904.1.1
D5088799B551C77EFF22DE8B8B44FB31.taxon	discussion	Remarks. Taking into account that certain morphological features were not appropriately described (or were not described at all) in González-Sponga’s (2006) original description of A. meridana, some information for this species was obtained from the contributions of Rojas-Runjaic et al. (2007, 2008), who examined the holotype female of this species. The features obtained from Rojas-Runjaic et al. (2007, 2008) are the ‘ relative position of the ventral trichobothria on pedipalp chela’ and the ‘ degree of pigment reticulation on the dorsal surface of chelicerae’. Likewise, some other features mentioned in the original description were confirmed after these authors’ contributions, i. e., the number of carinae on metasomal segment IV and the relative position of db and est trichobothria on the fixed finger. The ‘ length and degree of development of paramedian carinae on sternite VII’, which has never been previously mentioned for this species, was tested in the holotype female from photographs that were made available to R. B. - T. by F. J. M. Rojas-Runjaic. The very high correspondence obtained between i) the Colombian specimens, ii) the information of this species provided by González-Sponga (2006) and Rojas-Runjaic et al. (2007, 2008), and iii) photographs of the holotype female and one (presumably female) topotype, suffices to assign the Colombian specimens to A. meridana. This decision is supported by the fact that the Colombian population is located only some 100 straight Miles southwestward the type locality of A. meridana (Fig. 1). It is important to note that, even though the holotype female of A. meridana seems to have lost some coloration details (due to preservation effects and because the specimen was collected nearly three decades ago), the coxosternal region and pedipalp hand seem to have been predominantly (at least for the former) or completely (for the latter) yellow, as in the Colombian specimens. These features are included in the above diagnosis because they have proved to be useful for assisting species definitions, and in order to help distinguishing A. meridana from other Colombian species of the genus. Even though González-Sponga (2006: fig. 98) and Rojas-Runjaic et al. (2008: 70) agreed in that est trichobothrium is located basally to db in the fixed finger of the holotype female of A. meridana, the variation in the relative position of this pair of trichobothria observed in the Colombian specimens (see below) indicates there is no reason to think that both populations are not conspecific. This is more so since, judging from González-Sponga’s (2006: 98) illustration, est is not considerably but only slightly basal to db, something herein deemed to be normal intraspecific variability. Male. Paratype not fully described but measured by González-Sponga (2006). Male is not fully described herein because that is the aim of an ongoing, still unpublished work by F. J. M. Rojas-Runjaic. Hemispermatophore: Description based on ICN-As- 435: Flagelliform, thin and poorly sclerotized. Foot narrow and flat. Pedal flexure inconspicuous but movable. Body very long, wider on basal third. Capsular region with median, external and internal lobes, subequal and strong. Flagellum long, with short pars recta and long, coiled pars reflexa (Fig. 64). Carapace length to hemispermatophore body length ratio = 1: 1.22. There was no variation between both hemispermatophores examined. Female. Holotype described by González-Sponga (2006). Variability. Fixed finger trichobothria (n = 12): six (50 %) fingers with trichobothria in the order eb: esb: est / db: et: dt; five (41.7 %) in the order eb: esb: db: est: et: dt, with db very slightly basal to est; one (8.3 %) in the order eb: esb: est: db: et: dt, with est very slightly basal to db. Pectinal teeth count: 17 to 19 on females (n = 6; mode = 18); 17 to 19 on males (n = 6; mode = 18). Coloration of the coxosternal region: the juvenile male has this region uniformly yellow, possibly an effect of immaturity. Total body length (including telson): male 19 – 25 mm, female 28 – 33 mm.	en	Botero-Trujillo, Ricardo, Flórez, Eduardo (2011): A revisionary approach of Colombian Ananteris (Scorpiones, Buthidae): two new species, a new synonymy, and notes on the value of trichobothria and hemispermatophore for the taxonomy of the group. Zootaxa 2904 (1): 1-44, DOI: 10.11646/zootaxa.2904.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2904.1.1
D5088799B551C77EFF22DE8B8B44FB31.taxon	distribution	Distribution. Known from two localities, one in Venezuela (Mérida State) and one in Colombia (Norte de Santander Department) (Fig. 1). The specimens herein recorded from the Tamá Natural National Park represent the first records of A. meridana from Colombia and the first of the genus from Norte de Santander Department. Ecological notes. According to González-Sponga (2006), this species was found living in cracks beside a road. No further information exists to date.	en	Botero-Trujillo, Ricardo, Flórez, Eduardo (2011): A revisionary approach of Colombian Ananteris (Scorpiones, Buthidae): two new species, a new synonymy, and notes on the value of trichobothria and hemispermatophore for the taxonomy of the group. Zootaxa 2904 (1): 1-44, DOI: 10.11646/zootaxa.2904.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2904.1.1
D5088799B553C771FF22D94B8896FF21.taxon	description	Figures 1, 22 – 23, 37, 45, 53, 65; Tables 1, 2	en	Botero-Trujillo, Ricardo, Flórez, Eduardo (2011): A revisionary approach of Colombian Ananteris (Scorpiones, Buthidae): two new species, a new synonymy, and notes on the value of trichobothria and hemispermatophore for the taxonomy of the group. Zootaxa 2904 (1): 1-44, DOI: 10.11646/zootaxa.2904.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2904.1.1
D5088799B553C771FF22D94B8896FF21.taxon	materials_examined	Type material. Holotype female: COLOMBIA: Meta Department: Villavicencio, Path El Carmen, 850 – 1000 m asl, into forest, pitfall trap, 23 December 2005, M. Viola (MPUJ-SCO- 245) (examined). Paratype female: COLOMBIA: Meta Department: Villavicencio, Path El Carmen, 850 – 1000 m asl, River Caño Blanco, under litter, direct collection, at night, 18 April 2005, R. Botero-Trujillo (MPUJ-SCO- 039) (examined). Further material examined. COLOMBIA: Meta Department: 1 adult male, Villavicencio, Path El Carmen, 04 ° 08 ’ 27 ’’ N 73 ° 40 ’ 12 ’’ W, 794 m asl, UV light, 06 September 2008, J. A. Ochoa & R. Botero-Trujillo (MPUJ- SCO- 382). Cundinamarca Department: 1 adult male, Medina, Toquiza, 1000 m asl, pitfall trap, 30 January 2000, C. Cortez (ICN-As- 385). Revised diagnosis. Carapace with well-developed anteromedian projection (Fig. 37); metasomal carinal formula 10: 10: 10: 6: 5, with median lateral carinae on segment III only weakly developed, and ventrosubmedian and median lateral carinae absent on segment IV; V 1 and V 2 trichobothria unaligned axially, with V 2 located on an external position in relation to V 1 (Fig. 45); fixed finger trichobothria in the order eb: esb: est / db: et: dt (Fig. 53); dorsal surface of chelicerae with dense and complete reticular pattern (Fig. 37); pectines with 17 teeth on males, 15 – 16 on females; sternite VII with paramedian longitudinal carinae vestigial and incomplete; pedipalp hand uniformly yellow and spotless; coxosternal region base color yellow with conspicuous brown areas. Hemispermatophore: capsular region with median, external and internal lobes; flagellum with long and strongly coiled pars reflexa (Fig. 65). Male. Description based on MPUJ-SCO- 382. Measurements: see Table 2. Coloration: general coloration yellowish with variegated pigmentation over almost the entire body and appendages (see Figs. 22 – 23 for ICN-As- 385). Carapace predominantly brown with some yellow spots and stripes; anterior and posterior margins brown; arising posteriorly to each lateral ocular tubercle there is a thin and almost straight yellow stripe that ends behind the median ocular tubercle; median ocular tubercle black, surrounded on its base by thin yellow lines on anterior and posterolateral margins. Chelicerae with yellow coxa; hand with dense and complete reticular pattern on dorsal surface (see Fig. 37 for ICN-As- 385); fixed and movable fingers predominantly yellow and brown, respectively, each with reddish teeth. Coxosternal region base color yellow with large and conspicuous brown area in all the components, except for coxapophyses II which predominantly yellow; genital operculum yellow; pectinal basal piece with few brown mottling; pectines almost entirely yellow, with base of basalmost piece of both marginal and median lamellae brown-colored; sternites III – VII predominantly yellow, with some posterior brown regions located halfway between the midline and the lateral margins in sternites IV – VII; sternite V with a large subtriangular-to-oval posterior median hyaline area. Tergites predominantly brown; two longitudinal yellow stripes crossing tergites I – VI are only evident on the posterior half of each segment beside the midline; these lines are more conspicuous in tergites III – VI and appear as rounded spots in tergites I – II; each side of tergites I – VI with two transverse yellow lines converging near the longitudinal lines, arrow-like; tergite VII with variegated pigmentation; lateral margins of tergites I – VII yellow. Metasoma predominantly yellowish to reddish; dorsal intercarinal spaces of segments I – IV with a median brown design wider anteriorly, triangle-like on segments I – III but arrow-like on segment IV; segments I – IV with variegated pigmentation on all surfaces; segment V reddish, with variegated darker pigmentation ventrally and laterally. Telson vesicle yellowish-red, lighter than segment V; aculeus yellowish basally, dark-red distally; subaculear tubercle yellowish. Pedipalp femur and patella completely brown, lighter ventrally, with variegated pigmentation on dorsal and external surfaces of patella, and both segments with trichobothrial pits yellow; coxa and trochanter with variegated pigmentation; chela with hand uniformly yellow and spotless, fingers entirely brown with trichobothrial pits yellow. Legs with variegated pigmentation, brown spots in all segments except for the telotarsus which completely yellow. Carapace: densely covered with rounded granules especially on the brown areas; lateral margins converging anteriorly; anterior margin slightly bi-concave (with well-developed anteromedian projection in ICN-As- 385, Fig. 37); ocular carinae moderately strong and granulose, others inconspicuous; with a median depression anterior to the ocular tubercle; postocular furrow evident; median ocular tubercle low, located on the posterior half of the anterior third of carapace; lateral ocular tubercles each with three evident ocelli. Chelicerae: with abundant fine white setae on the internal and ventral surfaces; cheliceral dentition characteristic of the family Buthidae (Vachon 1963). Movable finger dorsally with two small basal teeth, one median pronounced, one subdistal slightly shorter than the median, and one distal tooth. Ventrally with three strong teeth, one basal, one median, and one distal tooth larger than its dorsal counterpart. Fixed finger dorsally with one basal and one median tooth forming a bicuspid, one subdistal, and one distal tooth. Ventrally with only one subdistal tooth slightly basal to the dorsal subdistal. Coxosternal region: sternum subtriangular, with deep median depression and two anterolateral furrows; all the components of this region very scarcely granulose, with abundant setae; coxapophyses I – II densely setose anteriorly. Genital operculum and pectines: genital operculum completely divided longitudinally; pectinal basal piece as long as wide, deeply notched anteriorly; pectines long, with distal tip of marginal lamella surpassing posterior end of coxae IV; count of pieces on the pectines: marginal lamellae 3: 3, middle lamellae 8: 9, teeth 17: 17, fulcra absent. Sternites: with abundant setae; III – VI completely smooth; VII with vestigial and incomplete paramedian longitudinal carinae, lateral carinae completely absent; V with large subtriangular-to-oval posterior median smooth area; spiracles elongated, approximately three times longer than wide in sternites III – V and four times longer than wide in VI. Tergites: with similar granulation to that of carapace; axial carina only evident on the posterior half of tergites I – VI; obsolete dorsolateral carinae present on tergites III – VI; tergite VII tetracarinate, with incomplete paired dorsolateral and lateral carinae, and granulose median elevation on the position of the axial carina. Metasoma: with abundant setae; segments I – III with ten carinae (paired ventrosubmedian, ventrolateral, median lateral, dorsolateral and dorsosubmedian carinae); segment IV with six (ventrosubmedian and median lateral carinae absent); segment V with five (ventromedian, paired ventrolateral and dorsolateral carinae); ventrolateral and median lateral carinae converge distally in segments I – II; all carinae serrulate; intercarinal spaces with abundant granulation. Telson: vesicle markedly elongated; tegument almost completely smooth except for three ventral longitudinal carinae made up of weak granulation; subaculear tubercle strong and spine-like; aculeus long and curved. Pedipalps: femur with five vestigial longitudinal carinae (dorsoexternal, dorsointernal, ventroexternal, ventrointernal and internal median carinae); patella without distinct carinae but with few granules on the position of the dorsointernal and ventrointernal carinae; chela acarinate; fixed finger with six almost linear rows of granules, being the basal the longest; movable finger with seven rows (including the short apical row). Trichobothriotaxy type A, femur with β configuration (Vachon 1974, 1975). Legs: tibia, basitarsus and telotarsus with numerous ventral setae; tibial spur present on legs III – IV; prolateral pedal spur single on legs I – II, bifid on legs III – IV; retrolateral pedal spur present on all legs. Hemispermatophore: Description based on MPUJ-SCO- 382: Flagelliform, thin and poorly sclerotized. Foot narrow and flat. Pedal flexure inconspicuous but movable. Body very long, wider on basal third. Capsular region with median, external and internal lobes, subequal and strong. Flagellum long, with short pars recta and long, coiled pars reflexa (Fig. 65). Carapace length to hemispermatophore body length ratio = 1: 1.04. There was no variation between both hemispermatophores examined. Female. Holotype described by Botero-Trujillo (2007). Variability. Pectinal teeth count: 15 to 16 on females (n = 4); 17 on males (n = 4). Total body length (including telson): male 21 – 25 mm, female 25 – 30 mm.	en	Botero-Trujillo, Ricardo, Flórez, Eduardo (2011): A revisionary approach of Colombian Ananteris (Scorpiones, Buthidae): two new species, a new synonymy, and notes on the value of trichobothria and hemispermatophore for the taxonomy of the group. Zootaxa 2904 (1): 1-44, DOI: 10.11646/zootaxa.2904.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2904.1.1
D5088799B553C771FF22D94B8896FF21.taxon	distribution	Distribution. So far known only from two localities, one each in Meta and Cundinamarca departments (Fig. 1). Probably an element typical of the eastern foothills of the Andes. New locality records: COLOMBIA: Cundinamarca Department: Medina. Ecological notes. All specimens of A. myriamae from Villavicencio (including the types) were collected at night, one of them after detection with UV light when actively moving. Two specimens from the type locality were hand-captured in sandy walls of small streams, whereas the third one and that from Medina were collected with pitfall traps, suggesting this species inhabits the leaf litter. Ananteris myriamae was not easily encountered in the type locality, where it was found in sympatry with the abundant T. nematochirus and T. bastosi, and the rarely found Tityus gaffini Lourenço, 2000.	en	Botero-Trujillo, Ricardo, Flórez, Eduardo (2011): A revisionary approach of Colombian Ananteris (Scorpiones, Buthidae): two new species, a new synonymy, and notes on the value of trichobothria and hemispermatophore for the taxonomy of the group. Zootaxa 2904 (1): 1-44, DOI: 10.11646/zootaxa.2904.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2904.1.1
D5088799B55CC773FF22DD3B8CE7F845.taxon	description	Figures 1, 24 – 27, 38, 46, 54, 56 – 57, 66; Tables 1, 2	en	Botero-Trujillo, Ricardo, Flórez, Eduardo (2011): A revisionary approach of Colombian Ananteris (Scorpiones, Buthidae): two new species, a new synonymy, and notes on the value of trichobothria and hemispermatophore for the taxonomy of the group. Zootaxa 2904 (1): 1-44, DOI: 10.11646/zootaxa.2904.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2904.1.1
D5088799B55CC773FF22DD3B8CE7F845.taxon	materials_examined	Type material. Holotype male (adult): COLOMBIA: Nariño Department: Barbacoas, Path El Barro, Río Ñambi Natural Reserve, 01 ° 17 ’ 10 ’’ N 78 ° 04 ’ 28 ’’ W, 1397 m asl, UV light, 11 September 2008, O. Tovar, J. P. Botero, R. Botero-Trujillo, J. Souza & J. A. Ochoa (MPUJ-SCO- 388). Allotype female (adult): COLOMBIA: identical data to the holotype (MPUJ-SCO- 389). Paratypes: 1 subadult male and 2 adult females with identical data to the holotype (ICN-As- 799). 3 adult males, 1 small (?) male, 1 subadult male and 1 juvenile male, Barbacoas, Altaquer, Río Ñambi Natural Reserve, 1440 m asl, UV light, 13 October 2009, E. Flórez & D. Luna (ICN-As- 815).	en	Botero-Trujillo, Ricardo, Flórez, Eduardo (2011): A revisionary approach of Colombian Ananteris (Scorpiones, Buthidae): two new species, a new synonymy, and notes on the value of trichobothria and hemispermatophore for the taxonomy of the group. Zootaxa 2904 (1): 1-44, DOI: 10.11646/zootaxa.2904.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2904.1.1
D5088799B55CC773FF22DD3B8CE7F845.taxon	etymology	Etymology. This species is named after our colleague and friend José A. Ochoa, currently at the Universidade de São Paulo (Brazil), in recognition to his contributions to the knowledge of South American (primarily Peruvian) scorpions.	en	Botero-Trujillo, Ricardo, Flórez, Eduardo (2011): A revisionary approach of Colombian Ananteris (Scorpiones, Buthidae): two new species, a new synonymy, and notes on the value of trichobothria and hemispermatophore for the taxonomy of the group. Zootaxa 2904 (1): 1-44, DOI: 10.11646/zootaxa.2904.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2904.1.1
D5088799B55CC773FF22DD3B8CE7F845.taxon	diagnosis	Diagnosis. Anterior margin of the carapace very slightly bi-concave, without anteromedian projection (Fig. 38); metasomal carinal formula 10: 10: 10: 8: 5, with median lateral carinae on segment III obsolete; telson vesicle densely granulose ventrally and laterally (more conspicuous in females), with three ventral longitudinal carinae armed with strong granulation, and one noticeably larger granule (tubercle-like) under the subaculear tubercle; V 1 and V 2 trichobothria very slightly unaligned axially, with V 2 located only slightly external to V 1 (Fig. 46); fixed finger trichobothria in the order eb: esb: est: db: et: dt (Fig. 54); dorsal surface of chelicerae with dense and complete reticular pattern, and small yellowish rounded area anteriorly (Fig. 38); pectines with 20 – 23 teeth on males (mode = 21), 22 – 23 on females (mode = 22); sternite VII with incomplete but strong paramedian longitudinal carinae; pedipalp hand uniformly yellow and spotless; movable and fixed fingers yellow in the basal third / quarter, the rest brownish; coxosternal region predominantly yellow and spotless, except for brownish regions on coxapophyses (and coxae on males) I and anterior tip of coxapophyses II. Hemispermatophore: capsular region with two lobes; flagellum with long pars recta and coiled pars reflexa (Fig. 66). Male. Description based on holotype. Measurements: see Table 2. Coloration: general coloration yellowish to reddish-brown with variegated pigmentation over almost the entire body and appendages (Figs. 24 – 25). Carapace predominantly brown with some yellow spots and stripes; anterior and posterior margins brown; median ocular tubercle black, surrounded on its base by thin yellow lines on anterolateral and posterolateral margins. Chelicerae with yellow coxa; hand with dense and complete reticular pattern on dorsal surface, and small yellowish rounded area anteriorly (Fig. 38); fixed and movable fingers dark-brown, each with reddish teeth. Coxosternal region almost entirely yellow and spotless, except for brownish regions on coxapophyses I, coxae I and anterior tip of coxapophyses II; genital operculum, pectinal basal piece, pectines and sternite III entirely yellow; sternites IV – VI with lateral margins brown; sternites V – VI with inconspicuous brownish areas located posteriorly midway between the midline and the lateral margins; sternite VII with broad brown longitudinal bands between the paramedian carinae and the lateral margins; sternite V with a subtriangular posterior median hyaline area. Tergites predominantly brown; two longitudinal yellow stripes crossing tergites II – VI are only evident on the posterior half of each segment beside the midline; each side of tergites I – VI with two transverse yellow lines converging near the longitudinal lines, arrow-like; tergite VII predominantly brown, with large yellow regions dorsally and laterally; lateral margins of tergites I – VII completely yellow. Metasoma predominantly yellowish-red; dorsal intercarinal spaces of segments I – IV with a median brown design wider anteriorly, triangle-like on segments I – II but arrow-like on segments III – IV; segments I – V with variegated pigmentation on all surfaces. Telson vesicle yellowish-red, lighter than segment V and with a longitudinal red band located on each side; aculeus yellowish basally, reddish distally; subaculear tubercle yellow. Pedipalp coxa and trochanter with variegated pigmentation; femur brown on all surfaces but lighter ventrally, trichobothrial pits yellow; patella brown with variegated pigmentation on dorsal, external and ventral surfaces, trichobothrial pits yellow; hand uniformly yellow and spotless; movable and fixed fingers yellow on the basal quarter and brownish on the rest of their length. Legs with variegated pigmentation from trochanter to basitarsi, telotarsi yellow. Carapace: densely covered with rounded granules especially on the brown areas; lateral margins converging anteriorly; anterior margin very slightly bi-concave, without anteromedian projection (Fig. 38); ocular carinae strong and granulose, others inconspicuous; with a median depression anterior to the ocular tubercle; postocular furrow evident; median ocular tubercle low, located on the posterior half of the anterior third of carapace; lateral ocular tubercles each with four ocelli, being the posteriormost very reduced and unaligned with the remaining three pairs. Chelicerae: with abundant fine white setae on the internal and ventral surfaces; cheliceral dentition characteristic of the family Buthidae (Vachon 1963). Movable finger dorsally with two small basal teeth, one median pronounced, one subdistal slightly shorter than the median, and one distal tooth. Ventrally with three strong teeth, one basal, one median, and one distal tooth larger than its dorsal counterpart. Fixed finger dorsally with one basal and one median tooth forming a bicuspid, one subdistal, and one distal tooth. Ventrally with only one subdistal tooth slightly basal to the dorsal subdistal. Coxosternal region: sternum subtriangular, with deep median depression and two anterolateral furrows; all the components of this region with few setae, almost completely smooth except for few granulations on the anterior margin of coxae; coxapophyses I – II densely setose anteriorly. Genital operculum and pectines: genital operculum completely divided longitudinally; pectinal basal piece as long as wide, deeply notched anteriorly; pectines long, with distal tip of marginal lamella reaching posterior end of coxae IV; count of pieces on the pectines: marginal lamellae 3: 3, middle lamellae 8: 9, teeth 21: 21, fulcra absent. Sternites: III completely smooth, lateral margins of IV – VI and posterior margin of VI with few granulations, VII densely granulose and with incomplete but strong paramedian longitudinal carinae, lateral carinae absent; V with subtriangular posterior median smooth area; spiracles elongated, approximately twice longer than wide in sternites III – IV and three times longer than wide in V – VI. Tergites: with similar granulation to that of carapace; axial carina only evident on the posterior half of tergites I – VI; dorsolateral carinae absent on tergites I – VI; tergite VII tetracarinate, with incomplete paired dorsolateral and lateral carinae, and granulose median elevation on the position of the axial carina. Metasoma: with few setae; segments I – III with ten carinae (paired ventrosubmedian, ventrolateral, median lateral, dorsolateral and dorsosubmedian carinae); segment IV with eight (median lateral carinae absent); segment V with five (ventromedian, paired ventrolateral and dorsolateral carinae); ventrolateral and median lateral carinae converge distally on segments I – II; ventrosubmedian and ventrolateral carinae are connected anteriorly by a transverse row of granules on segments II – IV; median lateral carinae on segment III obsolete; all carinae serrulate; dorsosubmedian carinae on segments II – IV with the distal granules well-defined and larger than the preceding; intercarinal spaces with minute dispersed granulation. Telson: vesicle densely granulose laterally and (primarily) ventrally, with three ventral longitudinal carinae armed with strong granulation, and one noticeably larger granule (tubercle-like) under the subaculear tubercle; subaculear tubercle strong and spine-like; aculeus long and curved. Pedipalps: with fine granulation on dorsal and internal surfaces of femur; femur with five longitudinal carinae (dorsoexternal, dorsointernal, ventroexternal, ventrointernal and internal median carinae); patella without distinct carinae but with few spine-like tubercles on the position of the dorsointernal and ventrointernal carinae; chela acarinate; fixed finger with six almost linear rows of granules, being the basal the longest; movable finger with seven rows (including the short apical row). Trichobothriotaxy type A, femur with β configuration (Vachon 1974, 1975). Legs: tibia, basitarsus and telotarsus with numerous ventral setae; tibial spur present on legs III – IV; prolateral pedal spur single on legs I – II, bifid on legs III – IV; retrolateral pedal spur present on all legs. Hemispermatophore: Description based on one specimen from ICN-As- 815: Flagelliform, thin and sclerotized. Foot narrow and flat. Pedal flexure inconspicuous but movable. Body long, slightly wider on basal third. Capsular region with a large, markedly sclerotized basal lobe, and an internal lobe. Flagellum with long pars recta and long, coiled pars reflexa (Fig. 66). Carapace length to hemispermatophore body length ratio = 1: 1.28. There was no variation between both hemispermatophores examined. Allotype female. Measurements: see Table 2. Figs. 26 – 27. Similar to holotype male but with the following differences deemed to be due to sexual dimorphism: body size larger; posterior median hyaline and smooth area of sternite V smaller and less evident; pectines markedly smaller, not reaching posterior end of coxae IV; telson not elongated. Aside from these primarily sexual differences, others include: general coloration darker; coxosternal region with minute brownish spots restricted to anterior half of coxapophyses I and anterior tip of coxapophyses II; anterior margin of coxae without any granulations; count of pieces on the pectines: marginal lamellae 3: 3, middle lamellae 8: 10, teeth 22: 23, vestigial fulcra present; sternite IV with the same inconspicuous brownish areas that are found on V – VI. Variability. Pectinal teeth count: 22 to 23 on females (n = 6; mode = 22); 20 to 23 on males (n = 16; mode = 21). Total body length (including telson): male 29 – 32 mm, female 41 – 43 mm.	en	Botero-Trujillo, Ricardo, Flórez, Eduardo (2011): A revisionary approach of Colombian Ananteris (Scorpiones, Buthidae): two new species, a new synonymy, and notes on the value of trichobothria and hemispermatophore for the taxonomy of the group. Zootaxa 2904 (1): 1-44, DOI: 10.11646/zootaxa.2904.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2904.1.1
D5088799B55CC773FF22DD3B8CE7F845.taxon	distribution	Distribution. Known only from the type locality in Barbacoas municipality (Nariño Department), extreme southwestern Colombia in the western slope of the Andes (Figs. 1, 56 – 57). Ecological notes. All specimens of A. ochoai sp. nov. were hand-captured at night after detection with UV light. Some individuals were detected when actively moving in the leaf litter, whereas others were found under decomposing logs. Ananteris ochoai sp. nov. was found in sympatry with A. dorae, Tityus sp. and Teuthraustes sp.	en	Botero-Trujillo, Ricardo, Flórez, Eduardo (2011): A revisionary approach of Colombian Ananteris (Scorpiones, Buthidae): two new species, a new synonymy, and notes on the value of trichobothria and hemispermatophore for the taxonomy of the group. Zootaxa 2904 (1): 1-44, DOI: 10.11646/zootaxa.2904.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2904.1.1
D5088799B559C74BFF22DCE08D86FA41.taxon	description	Figures 1, 28 – 31, 39, 47, 55, 58 – 59, 67; Tables 1, 2	en	Botero-Trujillo, Ricardo, Flórez, Eduardo (2011): A revisionary approach of Colombian Ananteris (Scorpiones, Buthidae): two new species, a new synonymy, and notes on the value of trichobothria and hemispermatophore for the taxonomy of the group. Zootaxa 2904 (1): 1-44, DOI: 10.11646/zootaxa.2904.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2904.1.1
D5088799B559C74BFF22DCE08D86FA41.taxon	materials_examined	Type material. Holotype male (adult): COLOMBIA: Santander Department: Girón, Quintas del Llanito, at night, 07 ° 04 ’ 15 ’’ N 73 ° 10 ’ 23 ’’ W, 703 m asl, UV light, February 2009, S. García (MPUJ-SCO- 390). Allotype female (adult): same location, 2011, S. García (MPUJ-SCO- 400). Paratypes: 4 adult males, same location, February 2009, S. García (MPUJ-SCO- 391 to MPUJ-SCO- 394). 5 adult males and 1 juvenile female, same location, June 2009, S. García (ICN-As- 779). 1 juvenile female, same location, June 2009, S. García (ICN-As- 777). 1 juvenile female, same location, June 2009, S. García (MPUJ-SCO- 395). 1 juvenile female, same location, June 2009, S. García (UIS-ART- 3233). 1 juvenile female, same location, June 2009, S. García (MPUJ-SCO- 396). Further material examined (non-type). COLOMBIA: Santander Department: 1 juvenile female, same data as above, June 2009, S. García (ICN-As- 800) (preserved in 96 % ethanol for DNA isolation).	en	Botero-Trujillo, Ricardo, Flórez, Eduardo (2011): A revisionary approach of Colombian Ananteris (Scorpiones, Buthidae): two new species, a new synonymy, and notes on the value of trichobothria and hemispermatophore for the taxonomy of the group. Zootaxa 2904 (1): 1-44, DOI: 10.11646/zootaxa.2904.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2904.1.1
D5088799B559C74BFF22DCE08D86FA41.taxon	etymology	Etymology. This species is named after Solimary García, enthusiast student of Colombian arachnids who collected the type series.	en	Botero-Trujillo, Ricardo, Flórez, Eduardo (2011): A revisionary approach of Colombian Ananteris (Scorpiones, Buthidae): two new species, a new synonymy, and notes on the value of trichobothria and hemispermatophore for the taxonomy of the group. Zootaxa 2904 (1): 1-44, DOI: 10.11646/zootaxa.2904.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2904.1.1
D5088799B559C74BFF22DCE08D86FA41.taxon	diagnosis	Diagnosis. Carapace with well-developed anteromedian projection (Fig. 39); metasomal carinal formula 10: 10: 10: 6: 5, with ventrosubmedian and median lateral carinae absent on segment IV; median lateral carinae on metasomal segment III vestigial, especially posteriorly; V 1 and V 2 trichobothria markedly unaligned axially, with V 2 located on an external position in relation to V 1 (Fig. 47); fixed finger trichobothria in the order eb: esb: est / db: et: dt, with est located beside or slightly basal to db (Fig. 55); dorsal surface of chelicerae predominantly yellow, almost without brown reticulations except anteriorly (Fig. 39); pectines with 19 – 21 teeth on males (mode = 19), 18 – 20 on females (mode = 19); sternite VII with paramedian longitudinal carinae vestigial and incomplete; pedipalp hand uniformly yellow and spotless; coxosternal region almost entirely yellow, except for some brownish areas on the anterior margin of all coxae. Hemispermatophore: capsular region with median, external and internal lobes; flagellum with long and strongly coiled pars reflexa (Fig. 67). Male. Description based on holotype. Measurements: see Table 2. Coloration: general coloration yellowish with variegated pigmentation over almost the entire body and appendages (Figs. 28 – 29). Carapace predominantly brown with some yellow spots and stripes; anterior and posterior margins brown; arising posteriorly to each lateral ocular tubercle there is a thin and almost straight yellow stripe that ends behind the median ocular tubercle; median ocular tubercle black, surrounded by yellow lines. Chelicerae with yellow coxa; hand almost completely yellow except for some anterior reticulations on dorsal, internal and external surfaces (Fig. 39); fixed finger completely yellow with reddish teeth, movable finger with the basal half brown and reddish teeth. Coxosternal region predominantly yellow, with some brownish areas along the anterior margin of all coxae; genital operculum, pectinal basal piece, pectines and sternites III – V entirely yellow and spotless; sternite VI with two paramedian, brown, rounded areas posteriorly; sternite VII with two large longitudinal brownish areas paramedially; sternite V with a large subtriangular-to-oval posterior median smooth area. Tergites primarily brown; two longitudinal yellow stripes crossing tergites I – VI are only evident on the posterior half of each segment beside the midline; each side of tergites I – VI with two transverse yellow lines converging near the longitudinal lines, arrow-like; tergite VII mostly yellow dorsally, brownish dorsolaterally; lateral margins of tergites I – VII yellow. Metasoma predominantly yellowish to reddish; dorsal intercarinal spaces of segments II – IV with a median brown design wider anteriorly, triangle-like on segments II – III but arrow-like on segment IV; all segments with variegated pigmentation on all surfaces; segment V dark-red. Telson vesicle reddish, lighter than segment V; aculeus dark-red distally. Pedipalp femur brownish dorsally, almost completely yellow externally, ventrally and internally; patella predominantly brown dorsally and externally with variegated pigmentation, yellow ventrally; trichobothrial pits surrounded by yellow areas on both femur and patella; coxa with variegated pigmentation; trochanter predominantly yellow; chela with hand uniformly yellow and spotless; fingers brown except for the distal quarter which is yellow. Legs with variegated pigmentation, brown spots on all segments except for the telotarsus that is completely yellow. Carapace: densely covered with rounded granules especially on the brown areas; lateral margins converging anteriorly; anterior margin with well-developed median projection (Fig. 39); ocular carinae moderately strong and granulose, others inconspicuous; with a median depression anterior to the ocular tubercle; postocular furrow evident; median ocular tubercle low, located on the posterior half of the anterior third of carapace; lateral ocular tubercles each with three ocelli. Chelicerae: with abundant fine white setae on the internal and ventral surfaces; cheliceral dentition characteristic of the family Buthidae (Vachon 1963). Movable finger dorsally with two small basal teeth, one median pronounced, one subdistal slightly shorter than the median, and one distal tooth. Ventrally with three strong teeth, one basal, one median, and one distal tooth larger than its dorsal counterpart. Fixed finger dorsally with one basal and one median tooth forming a bicuspid, one subdistal, and one distal tooth. Ventrally with only one subdistal tooth slightly basal to the dorsal subdistal. Coxosternal region: sternum subtriangular, with deep median depression and two anterolateral furrows; all the components of this region smooth, with abundant setae; coxapophyses I – II densely setose anteriorly. Genital operculum and pectines: genital operculum completely divided longitudinally; pectinal basal piece slightly longer than wide, deeply notched anteriorly; pectines long, with distal tip of marginal lamella reaching posterior end of coxae IV; count of pieces on the pectines: marginal lamellae 3: 3, middle lamellae 9: 9, teeth 19: 19, vestigial fulcra present. Sternites: with abundant setae; III – VI smooth; VII with vestigial and incomplete paramedian longitudinal carinae, lateral carinae absent; V with large subtriangular-to-oval posterior median smooth area; spiracles elongated, approximately twice longer than wide in sternites III – IV and three times longer than wide in V – VI. Tergites: with similar granulation to that of carapace; tergites II – VI with axial carina only evident on the posterior half and dorsolateral carinae absent; tergite VII tetracarinate, with incomplete paired dorsolateral and lateral carinae, and granulose median elevation on the position of the axial carina. Metasoma: with abundant setae; segments I – III with ten carinae (paired ventrosubmedian, ventrolateral, median lateral, dorsolateral and dorsosubmedian carinae); segment IV with six (ventrosubmedian and median lateral carinae absent); segment V with five (ventromedian, paired ventrolateral and dorsolateral carinae); median lateral carinae on metasomal segment III vestigial, especially posteriorly; ventrolateral and median lateral carinae converge distally on segments I – II; all carinae serrulate; intercarinal spaces with granulations. Telson: vesicle markedly elongated, almost completely smooth except for three vestigial ventral longitudinal carinae made up of weak granulation; subaculear tubercle strong and spine-like; aculeus long and curved. Pedipalps: femur with five longitudinal carinae (dorsoexternal, dorsointernal, ventroexternal, ventrointernal and internal median carinae); patella without distinct carinae but with some granules on the position of the dorsointernal and ventrointernal carinae; chela acarinate; fixed finger with six almost linear rows of granules, being the basal the longest; movable finger with seven rows (including the short apical row). Trichobothriotaxy type A, femur with β configuration (Vachon 1974, 1975). Legs: tibia, basitarsus and telotarsus with numerous ventral setae; tibial spur present on legs III – IV; prolateral pedal spur single on legs I – II, bifid on legs III – IV; retrolateral pedal spur present on all legs. Hemispermatophore: Description based on ICN-As- 779: Flagelliform, thin and poorly sclerotized. Foot narrow and flat. Pedal flexure inconspicuous but movable. Body very long, wider on basal third. Capsular region with median, external and internal lobes, subequal and strong. Flagellum long, with short pars recta and long, coiled pars reflexa (Fig. 67). Carapace length to hemispermatophore body length ratio = 1.02: 1. There was no variation between both hemispermatophores examined. Allotype female. Measurements: see Table 2. Figs. 30 – 31. Similar to holotype male but with the following differences deemed to be due to sexual dimorphism: body size larger; posterior median hyaline and smooth area of sternite V smaller and less evident; pectines smaller, not reaching posterior end of coxae IV; telson not elongated. Aside from these primarily sexual differences, others include: general coloration darker; count of pieces on the pectines: marginal lamellae 3: 3, middle lamellae 8: 8, teeth 18: 19, fulcra absent; telotarsi II – IV with brownish regions; six rows of granules in movable finger of left pedipalp. Variability. Fixed finger trichobothria (n = 33): 19 (57.6 %) in the order eb: esb: est: db: et: dt, with est very slightly basal to db; 14 (42.4 %) fingers with trichobothria in the order eb: esb: est / db: et: dt. Pectinal teeth count: 18 to 20 on females (n = 14; mode = 19); 19 to 21 on males (n = 19; mode = 19). Note that one male has 13 teeth on the left pectine, but this low number is not included in the count because such pectine lacks the distal piece of the marginal lamella (anomalous condition). Total body length (including telson): male 25 – 27 mm, female 34.32 mm. Anomalies. One male has only two rows of granules in the movable finger of the right pedipalp. The allotype female has six rows of granules in the movable finger of left pedipalp (lacks the short apical row). See ‘ Pectinal teeth count’ in the ‘ variability’ section above for further anomalies.	en	Botero-Trujillo, Ricardo, Flórez, Eduardo (2011): A revisionary approach of Colombian Ananteris (Scorpiones, Buthidae): two new species, a new synonymy, and notes on the value of trichobothria and hemispermatophore for the taxonomy of the group. Zootaxa 2904 (1): 1-44, DOI: 10.11646/zootaxa.2904.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2904.1.1
D5088799B559C74BFF22DCE08D86FA41.taxon	distribution	Distribution. Known only from the type locality in Girón municipality (Santander Department), north of the western slope of the Colombian eastern Andes (Figs. 1, 58 – 59). Ecological notes. All the specimens of A. solimariae sp. nov. were hand-captured at night with UV light between 19: 00 and 21: 00 hrs (not searched for after that hour). The area of collection is some 10000 m 2 and is located in an urban zone with marked human intervention (Figs. 58 – 59). The area is dominated by native vegetation and shrubs. Most specimens were found when actively moving in the leaf litter. No other scorpion species was detected in the type locality of A. solimariae sp. nov., where this species seems to be very abundant.	en	Botero-Trujillo, Ricardo, Flórez, Eduardo (2011): A revisionary approach of Colombian Ananteris (Scorpiones, Buthidae): two new species, a new synonymy, and notes on the value of trichobothria and hemispermatophore for the taxonomy of the group. Zootaxa 2904 (1): 1-44, DOI: 10.11646/zootaxa.2904.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2904.1.1
