taxonID	type	description	language	source
D61087F05164323BFF1EE9D02A9FFAD9.taxon	description	(Figs 1, 18, 22, 25 – 27)	en	Bezděk, Jan, Lee, Chi-Feng (2019): Revision of Pyrrhalta (Coleoptera: Chrysomelidae: Galerucinae) species with maculate elytra. Zootaxa 4664 (1): 518-534, DOI: 10.11646/zootaxa.4664.4.4
D61087F05164323BFF1EE9D02A9FFAD9.taxon	materials_examined	Type locality. “ Ta-chu-lan, Shaowu Dist., Fukien Prov. ”. Types examined. Holotype: ♂, “ Fukien, S. China [w, p] // Shaowu, Tachulan [w, p] // Pyrrhalta / maculata / G + K [h] / J. L. Gressitt det. [w, p] // HOLOTYPE [p] ♂ / Pyrrhalta / maculata [h] / Gressitt & Kimoto [r, h] // NO 23 [w, p] // ILL [w, p] // 3212 [w, h] ” (BPBM). Additional material examined. 11 specimens — TAIWAN: Meifeng, Nantou Hsien, 2130 m, 2 – 4. vi. 1980, 1 ♂, L. Y. Chou & C. C. Chen leg. (TARI); Meifeng, Nantou Hsien, 2150 m, 7 – 9. v. 1981, 1 ♀, K. S. Lin & S. C. Lin leg. (TARI); Meifeng, Nantou Hsien, 2150 m, 28 – 29. viii. 1981, 1 ♀, L. Y. Chou & S. C. Lin leg. (TARI); Meifeng, Nantou Hsien, 2150 m, 19 – 21. iv. 1983, 2 ♀♀, K. C. Chou & S. P. Huang leg. (TARI); Tsuifeng, Nantou Hsien, 2300 m, 27. viii. 1981, 1 ♂, L. Y. Chou & S. C. Lin leg. (TARI); Tsuifeng, Nantou Hsien, 2300 m, 1 – 3. viii. 1981, 1 ♂ 3 ♀♀, T. Lin & W. S. Tang leg. (TARI); Taichung, Fushoushan, 26. vii. 2010, 1 ♀, T – Y. Liu leg. (TARI).	en	Bezděk, Jan, Lee, Chi-Feng (2019): Revision of Pyrrhalta (Coleoptera: Chrysomelidae: Galerucinae) species with maculate elytra. Zootaxa 4664 (1): 518-534, DOI: 10.11646/zootaxa.4664.4.4
D61087F05164323BFF1EE9D02A9FFAD9.taxon	description	Redescription. Body length: ♂♂ 4.0 – 4.5 mm (holotype 4.5 mm); ♀♀ 3.9 – 4.4 mm. Male (Figs 25, 27). Dorsally brown except following parts black: middle part of vertex, three spots on pronotum (one elongate median and two smaller round lateral), scutellum, except apex, each elytron with five spots: one round spot between humerus and scutellum, not touching anterior elytral margin; two spots along lateral elytral margin; two spots along suture. Legs brown with apical halves of tibiae and tarsi often infuscate. Antennae brown with infuscate six apical antennomeres. Ventral side: head and prosternum brown, mesoventrite black with brown middle part, metaventrite black with brown posterior margin, abdomen pale brown with darker terminal ventrite and lateral parts of all other ventrites. Head. Labrum transverse with rounded anterior margin, surface with transverse row of several punctures bearing long pale setae. Anterior clypeal margin straight. Anterior part of head semiopaque, sparsely covered with punctures and longer pale setae. Interantennal space narrow, 0.66 times as wide as transverse diameter of antennal insertion. Interocular space 2.30 as wide as transverse diameter of eye. Frontal tubercles subquadrangular, in posterior half distinctly elevated, semiopaque, covered with microsculpture, tubercles separated from each other by thin sulcus, laterally demarcated by thin furrow, posterior margins continuously connected with vertex. Vertex dull, cov- ered with microsculpture, small punctures and short setae. Antennae thin, 0.62 times as long as body, length ratios of antennomeres I – XI equal to 15 - 9 - 19 - 8 - 7 - 8 - 7 - 8 - 8 - 8 - 13, in lateral view antennomere III concave dorsally, with large triangular apical tooth ventraly, antennomeres IV – V with sharp ventral keel (Fig. 18). Pronotum transverse, 1.76 times as wide as long, widest at anterior third. Anterior margin weakly concave, lateral margins obtuse angulate with rounded middle part, anteriorly and posteriorly convergent and nearly straight, posterior margin straight, lateral parts of posterior margin oblique and concave. All margins with thin borders. Anterior and posterior angles toothed and bearing long pale seta. Surface with distinct elevation along anterior margin, laterally broader and curved posteriorly, disc transversely impressed. Anterior elevation lustrous and almost glabrous, disc densely covered with small punctures and short setae. Scutellum triangular, with rounded apex, surface densely covered with microsculpture, small punctures and short setae. Elytra moderately convex, slightly extended posteriorly, 0.75 as long as body, 1.70 times as long as wide, dull, covered with small confused punctures with interspaces as wide as or narrower than puncture diameters and dense short pale setae, surface slightly impressed in subscutellar area, remainder of surface even. Humeral calli well developed. Epipleura wide, in apical quarter gradually narrowing towards apex, densely covered with short pale setae. Macropterous. Ventrally semiopaque, covered with longer pale setae. Last abdominal ventrite with deep U-shaped incision. Legs moderately narrow. Protarsi: protarsomere I narrow, nearly subparallel, without modified ventral surface, II subtriangular, short, length ratios of protarsomeres I – IV equal to 7 - 6 - 5 - 9. Mesotarsi: mesotarsomere I narrow, subparallel, in lateral view axe-shaped, with narrow basal part and expanded apical half, angles of expanded part pointed, length ratios of mesotarsomeres I – IV equal to 8 - 7 - 5 - 10. Metatarsi: metatarsomere I narrow, subparallel, length ratios of metatarsomeres I – IV equal to 10 - 6 - 6 - 10. Claws bifid. Aedeagus (Fig. 1) curved in its midlength in axis ca. 90 °. Basal half near the midlength with two short, blunt lateral spines (right spine closer to midlength than left one). Apical half asymmetrical with right side slightly rounded and apex curved left. Female. Antennomere III long, tubular, without triangular extension in apical part of ventral side, antennomeres IV – V without sharp ventral keel. Pro- and mesotarsomeres I not modified. Last abdominal ventrite entire, without deep U-shaped incision. Spermatheca (Fig. 22) with C-shaped cornu with slightly wider basal part and subsphaerical nodulus, cornu slightly wider than nodulus, basal part of ductus spermathecae subconical.	en	Bezděk, Jan, Lee, Chi-Feng (2019): Revision of Pyrrhalta (Coleoptera: Chrysomelidae: Galerucinae) species with maculate elytra. Zootaxa 4664 (1): 518-534, DOI: 10.11646/zootaxa.4664.4.4
D61087F05164323BFF1EE9D02A9FFAD9.taxon	distribution	Distribution. China: Fujian (Gressitt & Kimoto 1963), Taiwan (Gressitt & Kimoto 1963, present study). The distribution in other countries and Chinese provinces refer either to other species or their identity cannot be confirmed based on available specimens.	en	Bezděk, Jan, Lee, Chi-Feng (2019): Revision of Pyrrhalta (Coleoptera: Chrysomelidae: Galerucinae) species with maculate elytra. Zootaxa 4664 (1): 518-534, DOI: 10.11646/zootaxa.4664.4.4
D61087F05164323BFF1EE9D02A9FFAD9.taxon	diagnosis	Differential diagnosis. Pyrrhalta maculata is characterised by long antennomere III, which is more than twice longer than IV. Antennomere III with large triangular subapical extension ventrally in males, whereas it is cylindri- cal in females. Similarly modified antennomere III can be found only in P. carolusi sp. nov. (cf. Figs 18 – 19). All other species have antennomere III much shorter, 1.0 – 1.3 times as long as IV. Pyrrhalta maculata and P. carolusi sp. nov. differ in the structure of aedeagus, which is strongly curved dorsally at midlength in P. maculata and flat and straight in P. carolusi sp. nov. (Figs 1 – 2).	en	Bezděk, Jan, Lee, Chi-Feng (2019): Revision of Pyrrhalta (Coleoptera: Chrysomelidae: Galerucinae) species with maculate elytra. Zootaxa 4664 (1): 518-534, DOI: 10.11646/zootaxa.4664.4.4
D61087F051613239FF1EEFAC2AA1FDD5.taxon	description	(Figs 2, 12, 19, 31)	en	Bezděk, Jan, Lee, Chi-Feng (2019): Revision of Pyrrhalta (Coleoptera: Chrysomelidae: Galerucinae) species with maculate elytra. Zootaxa 4664 (1): 518-534, DOI: 10.11646/zootaxa.4664.4.4
D61087F051613239FF1EEFAC2AA1FDD5.taxon	materials_examined	Type locality. Laos, Houa Phan prov., Ban Saluei, Phou Pane Mt., 20 ° 12´N 104 ° 01´E. Types. Holotype: ♂, “ NE LAOS, Hua Phan prov. / Ban Saluei, Phou Pan (Mt.), / N 20 ° 12´E 104 ° 01´, / 1300 - 1900 m, / 11. iv – 15. v. 2012 [w, p] // BMNH {E} / 2012 - 14 / C. Holzschuh [w, p] // BMNH {E} / 1217219 [w, p] ” (BMNH). Paratypes: 1 ♂, “ NE LAOS, Hua Phan prov. / Ban Saluei, Phou Pan (Mt.), / 1300 - 1900 m, 7. iv – 25.2010 / 20 ° 12´N, 104 ° 01´E / leg. C. Holzschuh [w, p] // BMNH {E} / 2012 - 14 / C. Holzschuh [w, p] // BMNH {E} / 1211996 [w, p] ” (BMNH); 1 ♂, “ Thailand NW / Mae Hong Son prov. / pass Soppong – Pai, / 20 km from Soppong / 29.4 – 17.5.2007 / P. Viktora lgt. [w, p] ” (NMPC). The specimens are provided with one additional printed red label: “ HOLOTYPUS, [or PARATYPUS, respectively], / Pyrrhalta / carolusi sp. nov., / Bezděk & Lee, 2019 ”.	en	Bezděk, Jan, Lee, Chi-Feng (2019): Revision of Pyrrhalta (Coleoptera: Chrysomelidae: Galerucinae) species with maculate elytra. Zootaxa 4664 (1): 518-534, DOI: 10.11646/zootaxa.4664.4.4
D61087F051613239FF1EEFAC2AA1FDD5.taxon	description	Description. Body length: ♂♂ 4.0 – 4.3 mm (holotype 4.3 mm). Male (holotype, Fig. 31). Body and antennae with following parts black: middle part of vertex and base of head in ventral and lateral views, three spots on pronotum (one elongate median and two round lateraly), scutellum, except apex, each elytron with five round spots (one between humerus and scutellum, not touching anterior elytral margin; two along lateral elytral margin; two along suture), mesoventrite except middle part, metaventrite except posterior margin, abdomen with darker lateral parts. Legs brown with infuscate tarsi and apical halves of tibiae. Head. Labrum transverse with rounded anterior margin, surface with transverse row of several punctures bearing long pale setae. Anterior clypeal margin slightly concave. Anterior part of head semiopaque, sparsely covered with punctures and longer setae. Interantennal space narrow, 0.5 times as wide as transverse diameter of antennal insertion. Interocular space 2.33 as wide as transverse diameter of eye. Frontal tubercles quadrate, lustrous, smooth, distinctly elevated, separated each other by thin sulcus, laterally demarcated by thin furrow. Vertex dull, covered with microsculpture, small punctures and short setae. Antennae thin, 0.64 times as long as body, length ratios of antennomeres I – XI equal to 16 - 7 - 20 - 7 - 9 - 8 - 8 - 8 - 8 - 8 - 12, antennomere III subapically with large tooth directed downwards, IV-VI ventrally with sharp keel forming small subtriangular tooth (Fig. 19). Pronotum transverse, 1.89 times as wide as long, widest at anterior third. Anterior margin broadly, weakly concave, lateral margins obtuse angulate with widely rounded middle part, anteriorly and posteriorly convergent and nearly straight, posterior margin weakly concave, lateral part of posterior margin oblique and concave. All margins thinly bordered very thinly bordered. Anterior and posterior angles toothed and bearing long pale seta. Surface elevated along anterior margin, laterally curved posteriorly, disc transversely impressed. Elevation almost glabrous, disc densely covered with small punctures and short setae. Scutellum subtriangular, with widely rounded apex, surface very densely covered with microsculpture, small punctures and short setae. Elytra moderately convex, parallel, 0.72 as long as body, 1.63 times as long as wide, dull, covered with small confused punctures with interspaces as wide as or narrower than puncture diameters and dense short pale setae, surface with very slightly impressed subscutellar area, remainder of surface even. Humeral calli well developed. Epipleura wide, in apical quarter gradually narrowing towards apex, densely covered with short pale setae. Macropterous. Ventrally semiopaque, covered with longer pale setae. Terminal abdominal ventrite with deep V-shaped inci- sion. Legs moderately narrow. Protarsi: protarsomeres I and II subtriangular, short, protarsomeres I not modified ventrally, length ratios of protarsomeres I – IV equal to 6 - 6 - 5 - 9. Mesotarsi: mesotarsomere I narrow, subparallel, in lateral view axe-shaped, with narrow basal part and expanded apical two thirds, angles of expanded part pointed (Fig. 12), length ratios of mesotarsomeres I – IV equal to 7 - 6 - 5 - 11. Metatarsi: metatarsomere I narrow, subtriangular, length ratios of metatarsomeres I – IV equal to 8 - 7 - 5 - 11. Claws bifid. Aedeagus (Fig. 2) relatively narrow, hockey stick shaped, with apex curved left, in lateral view flat, not curved dorsally. Female unknown. Variability. The paratype from Thailand has smaller black spots on the pronotum.	en	Bezděk, Jan, Lee, Chi-Feng (2019): Revision of Pyrrhalta (Coleoptera: Chrysomelidae: Galerucinae) species with maculate elytra. Zootaxa 4664 (1): 518-534, DOI: 10.11646/zootaxa.4664.4.4
D61087F051613239FF1EEFAC2AA1FDD5.taxon	diagnosis	Differential diagnosis. Pyrrhalta carolusi sp. nov. has an elongate antennomere III which is about 2.8 - 2.9 times longer than IV and has a large, triangular subapical expansion. This character is known only in males of P. maculata (Figs 18 – 19). All other maculate Pyrrhalta species has antennomere III much shorter, 1.0 – 1.3 times as long as IV. Pyrrhalta carolusi sp. nov. and P. maculata can be distinguished by the structure of the aedeagus, which is flat and not curved in P. carolusi sp. nov. while strongly curved dorsally in midlength in P. maculata (Figs 1 – 2).	en	Bezděk, Jan, Lee, Chi-Feng (2019): Revision of Pyrrhalta (Coleoptera: Chrysomelidae: Galerucinae) species with maculate elytra. Zootaxa 4664 (1): 518-534, DOI: 10.11646/zootaxa.4664.4.4
D61087F051613239FF1EEFAC2AA1FDD5.taxon	distribution	Distribution. Laos, Thailand.	en	Bezděk, Jan, Lee, Chi-Feng (2019): Revision of Pyrrhalta (Coleoptera: Chrysomelidae: Galerucinae) species with maculate elytra. Zootaxa 4664 (1): 518-534, DOI: 10.11646/zootaxa.4664.4.4
D61087F051613239FF1EEFAC2AA1FDD5.taxon	etymology	Etymology. Dedicated to Carolus Holzschuh (Austria), specialist in Cerambycidae and the collector of two of the three known specimens of this new species.	en	Bezděk, Jan, Lee, Chi-Feng (2019): Revision of Pyrrhalta (Coleoptera: Chrysomelidae: Galerucinae) species with maculate elytra. Zootaxa 4664 (1): 518-534, DOI: 10.11646/zootaxa.4664.4.4
D61087F051633237FF1EE8F0290AFCFD.taxon	description	(Figs 3, 8, 13, 17, 32)	en	Bezděk, Jan, Lee, Chi-Feng (2019): Revision of Pyrrhalta (Coleoptera: Chrysomelidae: Galerucinae) species with maculate elytra. Zootaxa 4664 (1): 518-534, DOI: 10.11646/zootaxa.4664.4.4
D61087F051633237FF1EE8F0290AFCFD.taxon	materials_examined	Type locality. Indonesia, Brastagi, Sibayak Mt. Types. Holotype: ♂, “ N. SUMATRA – Brastagi / Mt Sibayak, 1500 - 2000 m / 20 – 26. iv. 1998 / lgt. Vít Kabou- rek [w, p] ” (NMPC). The holotype is provided with one additional printed red label: “ HOLOTYPUS, / Pyrrhalta / kaboureki sp. nov., / Bezděk & Lee, 2019 ”.	en	Bezděk, Jan, Lee, Chi-Feng (2019): Revision of Pyrrhalta (Coleoptera: Chrysomelidae: Galerucinae) species with maculate elytra. Zootaxa 4664 (1): 518-534, DOI: 10.11646/zootaxa.4664.4.4
D61087F051633237FF1EE8F0290AFCFD.taxon	description	Description. Body length: ♂ 3.9 mm (holotype). Male (holotype, Fig. 32). Head pale brown with slightly darker labrum, frontal tubercles and middle part of vertex. Pronotum pale brown with small indistinct darker spot medially, close to posterior margin. Antennomeres I-VI pale brown, gradually darker dorsally, VII-XI black. Scutellum pale brown with darker basal and middle parts. Elytra pale brown, each elytron with five brownish-black spots: one small rounded spot between humerus and scutellum, not touching anterior elytral margin; two larger longitudinaly elongate spots placed slightly before elytral midlength; two larger connected spots placed at posterior third. Legs pale brown with darker apical halves of tibiae and brownish-black tarsi. Ventral side: prosternum pale brown, mesoventrite black with brown middle part, meta- ventrite black, abdomen dark brown. Head. Labrum transverse with rounded anterior margin, surface with transverse row of several punctures bearing long pale setae. Anterior clypeal margin with small triangular impression. Anterior part of head semiopaque, sparsely covered with punctures and setae, surfaces below antennal insertions obliquely impressed. Interantennal space narrow, 0.3 times as wide as transverse diameter of antennal insertion. Interocular space 2.0 as wide as transverse diameter of eye. Frontal tubercles triangular, semiopaque, covered with microsculpture, separated each other by thin sulcus, laterally separated from frons by shallow oblique channel, posterior margins indistinctly separated from vertex. Vertex dull, covered with microsculpture, small punctures, and short setae. Antennae long, filiform, 0.71 times as long as body, length ratios of antennomeres I – XI equal to 15 - 6 - 13 - 11 - 10 - 10 - 10 - 9 - 9 - 9 - 12. Pronotum transverse, 2.0 times as wide as long, widest at anterior third. Anterior margin widely, weakly concave, lateral margins rounded at widest part, anteriorly and posteriorly convergent and nearly straight, posterior margin nearly straight, lateral part of posterior margin oblique. All margins thinly bordered. Anterior and posterior angles toothed and bearing long pale seta. Surface with distinct elevation along anterior margin, laterally curved posteriorly, disc transversely impressed. Elevation and anterior angles areas semiopaque, almost glabrous, covered with larger punctures, disc densely covered with small punctures and short setae. Scutellum slightly convex, subtriangular, with widely rounded apex, surface very densely covered with small punctures and short pale setae. Elytra semicylindrical, parallel, 0.69 as long as body, 1.50 times as long as wide, dull, covered with small confused punctures with interspaces wider than puncture diameters and dense short pale setae (setae darker on elytral spots), surface slightly uneven in middle part and in subscutellar area. Apical angle of each elytron rounded. Epipleura wide, in apical quarter gradually narrowing towards apex, densely covered with short pale setae. Ventrally semiopaque, covered with longer pale setae. Last abdominal ventrite wide and curved with deep Ushaped incision (Fig. 8). Legs narrow, inner apical margin of mesotibiae emarginate, with short sharp spine. Protarsi: protarsomere I short, narrow, parallel, II subtriangular, slightly wider than I, length ratios of protarsomeres I – IV equal to 5 - 6 - 4 - 7. Mesotarsi: mesotarsomere I short, narrow and subparallel, ventrally with distinct tooth in basal part (Fig. 13), length ratios of mesotarsomeres I – IV equal to 4 - 5 - 4 - 8. Metatarsi: metatarsomere I enlarged, triangular, inner apical angle more prolonged than outer one (Fig. 17), II small and subtriangular, length ratios of metatarsomeres I – IV equal to 11 - 8 - 5 - 8. Claws bifid. Aedeagus (Fig. 3) narrow and asymmetrical, arched right at anterior third, apex subtriangular and slightly asymmetrical. Ventrally with sharp, long median keel, in apical half surrounded by elongate, anteriorly wider fur- row. Female unknown.	en	Bezděk, Jan, Lee, Chi-Feng (2019): Revision of Pyrrhalta (Coleoptera: Chrysomelidae: Galerucinae) species with maculate elytra. Zootaxa 4664 (1): 518-534, DOI: 10.11646/zootaxa.4664.4.4
D61087F051633237FF1EE8F0290AFCFD.taxon	diagnosis	Differential diagnosis. Pyrrhalta kaboureki sp. nov. differs from other maculate Pyrrhalta species by the combination of the following characters: elytral spots placed closer to elytral disc (two outer spots closer to lateral and apical margins in other species, compare Figs. 27 – 36), structure of aedeagus (Fig. 3), metatarsomere I triangular and enlarged (Fig. 17) (not modified in other maculate Pyrrhalta species), mesotarsus I with small subbasal tooth ventrally (Fig. 13), subapical antennomeres about twice as long as wide (about 1.0 – 1.3 times as long as wide in other maculate Pyrrhalta species), interspaces between elytral punctation wider than puncture diameters (interspaces ca. as wide as or less than puncture diameter in other species).	en	Bezděk, Jan, Lee, Chi-Feng (2019): Revision of Pyrrhalta (Coleoptera: Chrysomelidae: Galerucinae) species with maculate elytra. Zootaxa 4664 (1): 518-534, DOI: 10.11646/zootaxa.4664.4.4
D61087F051633237FF1EE8F0290AFCFD.taxon	distribution	Distribution. Indonesia (Sumatra).	en	Bezděk, Jan, Lee, Chi-Feng (2019): Revision of Pyrrhalta (Coleoptera: Chrysomelidae: Galerucinae) species with maculate elytra. Zootaxa 4664 (1): 518-534, DOI: 10.11646/zootaxa.4664.4.4
D61087F051633237FF1EE8F0290AFCFD.taxon	etymology	Etymology. Dedicated to Vít Kabourek (Czech Republic, Zlín), entomologist, publisher, and collector of the holotype.	en	Bezděk, Jan, Lee, Chi-Feng (2019): Revision of Pyrrhalta (Coleoptera: Chrysomelidae: Galerucinae) species with maculate elytra. Zootaxa 4664 (1): 518-534, DOI: 10.11646/zootaxa.4664.4.4
D61087F0516D3235FF1EE9082B92FD41.taxon	description	(Figs 4, 14, 33)	en	Bezděk, Jan, Lee, Chi-Feng (2019): Revision of Pyrrhalta (Coleoptera: Chrysomelidae: Galerucinae) species with maculate elytra. Zootaxa 4664 (1): 518-534, DOI: 10.11646/zootaxa.4664.4.4
D61087F0516D3235FF1EE9082B92FD41.taxon	materials_examined	Type locality. Myanmar, Kambaiti. Types examined. Holotype: ♂, “ N. E. BURMA / Kambaiti, 7000 ft. [p] / 3 / 5 1934 [h] / R. MALAISE [w, p] ” (NHRS). The holotype is provided with one additional printed red label: “ HOLOTYPUS, / Pyrrhalta / kambaitien- sis sp. nov., / Bezděk & Lee, 2019 ”.	en	Bezděk, Jan, Lee, Chi-Feng (2019): Revision of Pyrrhalta (Coleoptera: Chrysomelidae: Galerucinae) species with maculate elytra. Zootaxa 4664 (1): 518-534, DOI: 10.11646/zootaxa.4664.4.4
D61087F0516D3235FF1EE9082B92FD41.taxon	description	Description. Body length: ♂ 4.5 mm (holotype). Male (holotype, Fig. 33). Dorsally brown, relatively lustrous, the following parts black with green metallic tint middle part of vertex, three elongate spots on pronotum, scutellum except extreme apex and four spots on each elytron (one round spot beween humerus and scutellum, touching anterior elytral margin, transverse broad stripe with irregular anterior and posterior margins, not touching suture at lateral elytral margins, and two preapical rounded spots. Antennae brown. Legs brown with slightly infuscate middle part of tibiae. Ventral side: head and prosternum brown, mesoventrite black with brown middle part, metaventrite brown with black lateral margins, abdomen completely brown. Head. Labrum transverse with rounded anterior margin, surface with transverse row of several punctures bearing long pale setae. Anterior clypeal margin straight. Anterior part of head lustrous, sparsely covered with punctures and short pale setae. Interantennal space narrow, 0.5 times as wide as transverse diameter of antennal insertion. Interocular space 2.45 as wide as transverse diameter of eye. Frontal tubercles subquadrate, strongly bulging, semiopaque, separated each other by thin sulcus, laterally demarcated by thin furrow, in apical half covered with microsculpture, in posterior half covered with microsculpture mixed with small punctures. Vertex semiopaque, covered with microsculpture, small punctures and short setae. Antennae thin with slightly more elongate apical antennomeres, 0.64 times as long as body, length ratios of antennomeres I – XI equal to 14 - 8 - 12 - 10 - 9 - 8 - 9 - 8 - 8 - 10 - 17, antennomeres VIII-XI slightly flattened on ventral side. Pronotum transverse, twice as wide as long, widest at anterior third. Anterior margin widely, weakly concave, lateral margins obtuse angulate with rounded middle part, anteriorly and posteriorly convergent and nearly straight, posterior margin slightly concave, lateral part of posterior margin oblique and concave. All margins thinly bordered. Anterior and posterior angles toothed, with setigerous pores (setae broken in holotype). Surface with distinct elevation along anterior margin, laterally curved posteriorly, disc transversely impressed. Elevation glabrous and lustrous in middle part, laterally sparsely covered with punctures, disc densely covered with large punctures and short setae. Scutellum subtriangular, with rounded apex, surface covered with small punctures and short setae. Elytra elongate and relatively flat, slightly extended posteriorly, 0.75 as long as body, 1.80 times as long as wide, lustrous, covered with small confused punctures with interspaces as wide as or narrower than puncture diameters and dense short pale setae, surface even. Humeral calli well developed. Epipleura wide, in apical quarter gradually narrowing towards apex, glabrous, smooth. Macropterous. Ventrally semiopaque, covered with longer pale setae. Last abdominal ventrite with deep V-shaped incision. Legs moderately narrow. Mesotarsi: mesotarsomere I triangular, in lateral view with very small tooth in midlength ventrally (Fig. 14), length ratios of mesotarsomeres I – III equal to 7 - 7 - 5. Metatarsi: metatarsomere I elongate triangular, length ratios of metatarsomeres I – III equal to 10 - 8 - 5. Protarsi, all tarsomeres IV and claws missing in holotype. Aedeagus (Fig. 4) with apical quarter narrow, finger shaped, strongly curved dorsally and slightly left. Basal half of aedeagus nearly parallel. Female unknown.	en	Bezděk, Jan, Lee, Chi-Feng (2019): Revision of Pyrrhalta (Coleoptera: Chrysomelidae: Galerucinae) species with maculate elytra. Zootaxa 4664 (1): 518-534, DOI: 10.11646/zootaxa.4664.4.4
D61087F0516D3235FF1EE9082B92FD41.taxon	diagnosis	Differential diagnosis. Pyrrhalta kambaitiensis sp. nov. can be distiguished from other maculate Pyrrhalta species by the combination of following characters: dorsally relatively lustrous, black spots with distinct metallic sheen, middle pair of elytral spots connected to transverse stripe (elytra dull, without metallic sheen on black spots and middle pair of spots separated in other species), pronotum covered with large punctures (punctures distinctly smaller in other species), aedeagus with thin finger-like apical process curved upwards (Fig. 4).	en	Bezděk, Jan, Lee, Chi-Feng (2019): Revision of Pyrrhalta (Coleoptera: Chrysomelidae: Galerucinae) species with maculate elytra. Zootaxa 4664 (1): 518-534, DOI: 10.11646/zootaxa.4664.4.4
D61087F0516D3235FF1EE9082B92FD41.taxon	distribution	Distribution. Myanmar.	en	Bezděk, Jan, Lee, Chi-Feng (2019): Revision of Pyrrhalta (Coleoptera: Chrysomelidae: Galerucinae) species with maculate elytra. Zootaxa 4664 (1): 518-534, DOI: 10.11646/zootaxa.4664.4.4
D61087F0516D3235FF1EE9082B92FD41.taxon	etymology	Etymology. Named after the type locality Kambaiti (Myanmar).	en	Bezděk, Jan, Lee, Chi-Feng (2019): Revision of Pyrrhalta (Coleoptera: Chrysomelidae: Galerucinae) species with maculate elytra. Zootaxa 4664 (1): 518-534, DOI: 10.11646/zootaxa.4664.4.4
D61087F0516F3233FF1EE89C2D55FE39.taxon	description	(Figs 5, 9 – 10, 15, 20, 23, 34)	en	Bezděk, Jan, Lee, Chi-Feng (2019): Revision of Pyrrhalta (Coleoptera: Chrysomelidae: Galerucinae) species with maculate elytra. Zootaxa 4664 (1): 518-534, DOI: 10.11646/zootaxa.4664.4.4
D61087F0516F3233FF1EE89C2D55FE39.taxon	materials_examined	Type locality. Laos, Houa Phan prov., Ban Saluei → Phou Pane Mt., 20 ° 12 – 13.5´N 103 ° 59.5´– 104 ° 01´E. Types. Holotype: ♂, “ Laos-NE, Houa Phan prov., / 20 ° 12 – 13.5´N 103 ° 59.5´– 104 ° 01´E, / Ban Saluei → Phou Pane Mt., / 1340 – 1870 m, 2 – 22. vi. 2011, / Vít Kubáň & Lao coll. Lgt. [w, p] // Primary mountain forest, / at light + individual collecting, / Laos 2008 Expedition / National Muzeum Prague, / Czech Republic [w, p] ” (NMPC). Paratypes: LAOS: 4 ♂♂ 15 ♀♀, same labels as holotype (NMPC, 1 ♂ 1 ♀ TARI); 1 ♀, “ LAOS – NE; HUA PHAN prov.; / BAN SALUEI; Phu Phan Mt.; / 20 ° 15´N 104 ° 02´E; 1500 – 2000 m; / J. Bezděk leg.; 26. iv – 11. v. 2001 [w, p] ” (JBCB); 2 ♂♂ 7 ♀♀, “ NE LAOS, Hua Phan prov. / Ban Saluei, Phou Pan (Mt.) / ~ N 20 ° 12´E 104 ° 01´/ 1300 – 1900 m, 3 – 30. iv. 2014 / C. Holzschuh leg. [w, p] // BMNH {E} / 2015 - 38 / C. Holzschuh [w, p] ” (BMNH); 2 ♀♀, “ LAOS-NE, Houa Phan prov. / 20 ° 12 – 13.5´N, 103 ° 59.5 - 104.01´E, / Ban Saluei → Phou Pane Mt., / 1340 – 1870 m, 22. iv – 15. v. 2008 / Vít. Kubáň & Lao coll. leg. [w, p] // Primary mountain forest, / individual collecting. / Laos 2008 Expedition / National Museum Prague, / Czech Republic [w, p] ” (NMPC); 58 ♂♂ 40 ♀♀, “ NE LAOS, Hua Phan prov. / Ban Saluei, Phou Pan (Mt.) / ~ N 20 ° 12´E 104 ° 01´/ 1300 – 1900 m, 3 – 30. iv. 2014 / C. Holzschuh leg. [w, p] // B. M. 2015 - 38 / C. Holzschuh [w, p] ” (BMNH); 1 ♂ 4 ♀♀, “ NE LAOS, Hua Phan prov. / Ban Saluei, Phou Pan (Mt.), / N 20 ° 12´E 104 ° 01´, / 1300 – 1900 m, / 11. iv – 15. v. 2012 [w, p] // BMNH {E} / 2012 - 14 / C. Holzschuh [w, p] ” (BMNH); 3 ♂♂ 4 ♀♀, “ Collection / Naturhistorisches / Museum Basel [w, p] // LAO, Phongsaly prov., / 21 ° 41 – 2´N 102 ° 06 - 8´E, / 28. v – 20. vi. 2003, / PHONGSALY env., / ~ 1500 m, Vít Kubáň leg. [w, p] ” (NHMB); 1 ♀, “ LAO, Phongsaly prov., / 21 ° 41 - 2´N 102 ° 06 - 8´E, / 28. v – 20. vi. 2003, / PHONGSALY env., / ~ 1500 m, Brancucci leg. [w, p] ” (NHMB); 1 ♂ 8 ♀♀, “ Collection / Naturhistorisches / Museum Basel [w, p] // LAO, Phongsaly prov., / 21 ° 41 - 2´N 102 ° 06 - 8´E, / 28. v – 20. vi. 2003, / PHONGSALY env., / ~ 1500 m, Pacholátko leg. [w, p] ” (NHMB). THAILAND: 1 ♂ 1 ♀, “ THAILAND, Chiang Mai prov., / Chiang Mai env., NP Doi / Suthep-Pui, 18 ° 49´34.25 ´´ N, [w, p] // 98 ° 53´43.40 ´´ E; sweeping in / forest around a trail margin, / 9. iv. 2015, Albert Damaška lgt. [w, p] ” (JBCB); 5 ♀♀, “ THAILAND, Nan prov., / Doi Phuka Nat. Park, / 28. IV – 12. V. 2002 / P. Průdek & M. Obořil leg. [w, p] ” (JBCB); 1 ♂ 5 ♀♀, “ Thailand, Mae Hong Son pr., / Kiwlom-pass near Soppong, / 23. vi – 2. vii. 2002, 1400 m, / WGS 84: 19 ° 26´N, 98 ° 19´E, / lgt. Fouqué R. + H. [w, p] ” (JBCB); 3 ♀♀, “ NW THAILAND, 19 – 23. / Chiang Mai, 4.1991 / Doi Suthep to Doi Pui / J. Horák leg. [w, p] ” (JBCB); 1 ♀, “ THAILAND Nan pr. / Doi Phuka NP / May 2000 / local collector lgt. / BMNH {E} 2016 - 44 [w, p] ” (BMNH); 1 ♀, “ NE Thailand, 19 - 22.4 / 1991, Chieng Mai prov. / Doi Suthep, S. Bily leg [w, p] // Pyrrhalta / maculata Gr. / Kim. [h] / det. L. Medvedev [p] 1997 [h] ” (NHMB); 1 ♂, “ Thailand NW, 29. IV – 17. V. 2007 / Mae Hong Son prov. / pass Soppong-Pai, 20 km from / Soppong P. Viktora leg. [y, p] ” (JBCB); 1 ♀, “ NW Thai 9 – 16.5.1991 / MAE HONG SON / Ban Huai Po 1600 m / leg. P. Pacholátko [w, p] ” (JBCB); 1 ♀, “ S THAILAND, BETONG / GUNUNG CANG DUN VILL / YALA distr. / 25.3 – 22.4.1993 / LGT. HORAK [w, p] ” (JBCB); 1 ♀, “ NW Thailand, 1991 / Chiang Mai, 18.49 N, 99.00 E / Doi Suthep to Doi Pui / 19 – 23.4., L. Dembický leg. [w, p] ” (NMW). CHINA: Yunnan: 1 ♀, “ CHINA: S-YUNNAN / (Xishuangbanna) / 45 km SW Jinghong, vic. Bangzhang vill. [w, p] // N 21 ° 44´37 ´´, E 100 ° 27´02 ´´, / 16 – 1700 m, 03 – 05. V. 2009 / leg. A. Weigel, blüh. Cast. [w, p] // Pyrrhalta / maculata G / K [h] / L. Medvedev det. [w, p] ” (NMEG); 1 ♂ 6 ♀♀, “ CHINA: Yannan / Honghe / Qibaofeng, 1350 m, / 23 ° 4´4.2 ´´ N, 103 ° 23´/ 46.9 ´´ E, 17. VI. 2018 / leg. A. Weigel KL / KF [w, p] // collection / NATURKUNDE- / MUSEUM ERFURT [y, p] ” (NMEG). The specimens are provided with one ad- ditional printed red label: “ HOLOTYPUS, [or PARATYPUS, respectively], / Pyrrhalta / lucka sp. nov., / Bezděk & Lee, 2019 ”.	en	Bezděk, Jan, Lee, Chi-Feng (2019): Revision of Pyrrhalta (Coleoptera: Chrysomelidae: Galerucinae) species with maculate elytra. Zootaxa 4664 (1): 518-534, DOI: 10.11646/zootaxa.4664.4.4
D61087F0516F3233FF1EE89C2D55FE39.taxon	description	Description. Body length: ♂♂ 3.5 – 4.4 mm (holotype 3.7 mm); ♀♀ 3.5 – 4.5 mm. Male (holotype, Fig. 34). Dorsally pale brown except following parts black: middle part of vertex, three spots on pronotum (one elongate median and two smaller round spots laterally), scutellum, each elytron with five spots: one small round spot between humerus and scutellum, not touching anterior elytral margin; two round spots along lateral elytral margin; two elongate spots along suture. Legs pale brown with tarsi and apical halves of all tibiae infuscate. Antennae with pale brown antennomeres I-III, darker brown IV-VI and brown VII-XI. Ventral side: head and prosternum pale brown, mesoventrite black with brown middle part, metaventrite black with brown posterior margin, abdomen pale brown with lateral parts of each abdominal ventrite black. Head. Labrum transverse with rounded anterior margin, surface with transverse row of several punctures bearing long pale setae. Anterior clypeal margin slightly concave. Anterior part of head semiopaque, sparsely covered with punctures and longer pale setae. Interantennal space narrow, 0.65 times as wide as transverse diameter of antennal insertion. Interocular space 2.62 as wide as transverse diameter of eye. Frontal tubercles subtriangular semiopaque, covered with microsculpture, separated each other by thin sulcus, laterally demarcated by thin furrow, posterior margins continuous with vertex. Vertex dull, covered with microsculpture, small punctures and short setae. Antennae (Fig. 20) subclavate, 0.70 times as long as body, length ratios of antennomeres I – XI equal to 13 - 7 - 10 - 9 - 8 - 8 - 8 - 8 - 9 - 9 - 15, last five antennomeres distinctly dilated, antennomeres IX-X as wide as long, XI 1.85 times as long as wide, antennomeres VIII-XI distinctly flattened on ventral side. Pronotum transverse, 1.75 times as wide as long, widest at anterior third. Anterior margin widely, weakly concave, lateral margins obtuse angulate with broadly rounded middle part, anteriorly and posteriorly convergent and nearly straight, posterior margin weakly concave, lateral parts of posterior margin oblique and concave. All margins thinly bordered. Anterior and posterior angles toothed and bearing long pale seta. Surface with distinct elevation along anterior margin, laterally wider and curved posteriorly, disc transversely impressed. Elevation almost glabrous and lustrous, disc densely covered with small punctures and short setae. Scutellum triangular, with rounded apex, surface densely covered with microsculpture, small punctures and short setae. Elytra moderately convex, slightly extended posteriorly, 0.73 as long as body, 1.65 times as long as wide, dull, covered with small confused punctures with interspaces as wide as or narrower than puncture diameters and dense, short, pale setae, surface weakly impressed in subscutellar area, rest of surface even. Humeral calli well developed. Epipleura wide, in apical quarter gradually narrowed towards apex, densely covered with short pale setae. Macropterous. Ventrally semiopaque, covered with longer pale setae. Last abdominal ventrite with deep U-shaped incision (Fig. 9). Legs moderately narrow. Protarsi: protarsomere I narrow, subparallel, II subtriangular, short, ventrally, protarsomeres I concave in lateral view, basally with distinct tooth (Fig. 10), length ratios of protarsomeres I – IV equal to 5 - 6 - 4 - 9. Mesotarsi: mesotarsomere I narrow, subparallel, in lateral view axe-shaped, with very narrow basal part and extended apical two thirds, angles of extended part pointed (Fig. 15), length ratios of mesotarsomeres I – IV equal to 6 - 6 - 5 - 9. Metatarsi: metatarsomere I narrow, subparallel, length ratios of metatarsomeres I – IV equal to 7 - 6 - 5 - 10. Claws bifid. Aedeagus (Fig. 5) with wide flat apical half, in dorsal view sublanceolate with large triangular incision on left side, anterior part of incision with membranous structure, basal half of aedeagus narrower than apical half but robust in lateral view. Female. Antennae with brown antennomeres I-III, antennomeres IV and V gradually darkened, VI-XI black. Antennae subclavate but apical five antennomeres narrower than in male (ca 1.3 times as long as wide). Pro- and mesotarsomeres I not modified. Last abdominal ventrite entire, without deep U-shaped incision. Spermatheca (Fig. 23) with C-shaped cornu, basal part slightly broader, and subsphaerical nodulus, cornu slightly wider than nodulus, basal part of ductus spermathecae finger-like where thin ductus is attached.	en	Bezděk, Jan, Lee, Chi-Feng (2019): Revision of Pyrrhalta (Coleoptera: Chrysomelidae: Galerucinae) species with maculate elytra. Zootaxa 4664 (1): 518-534, DOI: 10.11646/zootaxa.4664.4.4
D61087F0516F3233FF1EE89C2D55FE39.taxon	diagnosis	Differential diagnosis. In habitus, Pyrrhalta lucka sp. nov. is similar to P. schillhammeri sp. nov. from Laos. The males of both species possess subclavate antennae and modified ventrally, pro- and mesotarsomeres I (Figs 10 – 11, 15 – 16, 20 – 21). Both species differ in the structure of the aedeagus. The aedeagus of P. lucka sp. nov. has an extended, flat, lanceolate apical half, while the aedeagus of P. schillhammeri sp. nov. has the apical half narrow and slightly curved left (Figs 5 – 6). In the additional similar species, Pyrrhalta tsoui sp. nov. from Taiwan and Laos, the elytral spots near the suture are round (elongate in P. lucka sp. nov.), the antennae lack distinctly enlarged apical antennomeres (antennae subclavate in P. lucka sp. nov.), and ventrally, pro- and mesotarsomeres I are not modified (modified in P. lucka sp. nov.).	en	Bezděk, Jan, Lee, Chi-Feng (2019): Revision of Pyrrhalta (Coleoptera: Chrysomelidae: Galerucinae) species with maculate elytra. Zootaxa 4664 (1): 518-534, DOI: 10.11646/zootaxa.4664.4.4
D61087F0516F3233FF1EE89C2D55FE39.taxon	distribution	Distribution. Laos, Thailand, China (Yunnan).	en	Bezděk, Jan, Lee, Chi-Feng (2019): Revision of Pyrrhalta (Coleoptera: Chrysomelidae: Galerucinae) species with maculate elytra. Zootaxa 4664 (1): 518-534, DOI: 10.11646/zootaxa.4664.4.4
D61087F0516F3233FF1EE89C2D55FE39.taxon	etymology	Etymology. Dedicated to our dear friend and colleague Lucie “ Lucka ” Havlová (Czech Republic, Brno).	en	Bezděk, Jan, Lee, Chi-Feng (2019): Revision of Pyrrhalta (Coleoptera: Chrysomelidae: Galerucinae) species with maculate elytra. Zootaxa 4664 (1): 518-534, DOI: 10.11646/zootaxa.4664.4.4
D61087F051693232FF1EE8762C72FD89.taxon	description	(Figs 6, 11, 16, 21, 35)	en	Bezděk, Jan, Lee, Chi-Feng (2019): Revision of Pyrrhalta (Coleoptera: Chrysomelidae: Galerucinae) species with maculate elytra. Zootaxa 4664 (1): 518-534, DOI: 10.11646/zootaxa.4664.4.4
D61087F051693232FF1EE8762C72FD89.taxon	materials_examined	Type locality. Laos, Champasak prov., Bolavens Plateau, Muang Paksong — B. Nam Thang. Types examined. Holotype: ♂, “ S-LAOS: Prov. Champasak / Bolavens-Plateau / M. Paksong — B. Nam Thang / 26. / 27.5.1996, 800 – 1100 m / leg. Schillhammer (6) [w, p] ” (NMW). The holotype is provided with one additional printed red label: “ HOLOTYPUS, / Pyrrhalta / schillhammeri sp. nov., / Bezděk & Lee, 2019 ”.	en	Bezděk, Jan, Lee, Chi-Feng (2019): Revision of Pyrrhalta (Coleoptera: Chrysomelidae: Galerucinae) species with maculate elytra. Zootaxa 4664 (1): 518-534, DOI: 10.11646/zootaxa.4664.4.4
D61087F051693232FF1EE8762C72FD89.taxon	description	Description. Body length: ♂ 3.7 (holotype). Male (holotype, Fig. 35). Head pale brown with middle part of vertex black. Pronotum pale brown with one elongate median and two round lateral black spots. Antennae pale brown with darkened antennomeres IV-VII. Scu- tellum black. Elytra pale brown, each elytron with five brownish-black spots: one small round spot between humerus and scutellum, not touching anterior elytral margin; two round spots along lateral elytral margin; two elongate spots along suture. Legs pale brown. Ventral side: prosternum pale brown, mesoventrite black with brown middle part, metaventrite black with brown posterior margin, abdomen pale brown with lateral parts of each abdominal ventrite black. Head. Labrum transverse with rounded anterior margin, surface with transverse row of several punctures bearing long pale setae. Anterior clypeal margin weakly concave. Anterior part of head semiopaque, sparsely covered with punctures. Interantennal space narrow, 0.6 times as wide as transverse diameter of antennal insertion. Interocular space 2.5 as wide as transverse diameter of eye. Frontal tubercles subtriangular semiopaque, covered with microsculpture, separated each other by thin sulcus, laterally demarcated by thin furrow, posterior margins continuous with vertex. Vertex dull, covered with microsculpture, small punctures and short setae. Antennae (Fig. 21) subclavate, 0.62 times as long as body, length ratios of antennomeres I – XI equal to 13 - 6 - 8 - 8 - 6 - 6 - 7 - 6 - 7 - 7 - 13, last five antennomeres distinctly dilated, antennomeres VIII-X as wide as long, XI twice as long as wide, antennomeres VIII-XI distinctly flattened on ventral sides. Pronotum transverse, 1.77 times as wide as long, widest at anterior third. Anterior margin widely shallowly concave, lateral margins obtuse angulate with widely rounded middle part, anteriorly and posteriorly convergent and nearly straight, posterior margin weakly concave, lateral part of posterior margin oblique and concave. All margins thinly bordered. Anterior and posterior angles toothed and bearing long pale seta. Surface with distinct elevation along anterior margin, laterally curved posteriorly, disc transversely impressed. Elevation almost glabrous, with irregular small lustrous parts, disc densely covered with small punctures and short setae. Scutellum subtriangular, with widely rounded apex, surface densely covered with microsculpture, small punctures and short setae. Elytra moderately convex, parallel, 0.72 as long as body, 1.62 times as long as wide, dull, covered with small confused punctures with interspaces as wide as or narrower than puncture diameters and dense, short pale setae, surface with weakly impressed subscutellar area, rest of surface even. Humeral calli well developed. Epipleura wide, in apical quarter gradually narrowing towards apex, densely covered with short pale setae. Macropterous. Ventrally semiopaque, covered with longer pale setae. Last abdominal ventrite with deep U-shaped incision. Legs moderately narrow. Protarsi: protarsomeres I and II subtriangular, short, protarsomere I concave in lateral view, basally with distinct tooth (Fig. 11), length ratios of protarsomeres I – IV equal to 6 - 6 - 4 - 7. Mesotarsi: meso- tarsomere I narrow, subparallel, in lateral view axe-shaped, with narrow basal part and expanded apical two thirds, angles of expanded parts pointed (Fig. 16), length ratios of mesotarsomeres I – IV equal to 7 - 6 - 4 - 7. Metatarsi: meta- tarsomere I narrow, subtriangular, length ratios of metatarsomeres I – IV equal to 7 - 6 - 5 - 8. Claws bifid. Aedeagus (Fig. 6) relatively narrow, with apical fifth curved left and tip straight, right side of aedeagus nearly straight, subapical part of left side broadly emarginate. Female unknown.	en	Bezděk, Jan, Lee, Chi-Feng (2019): Revision of Pyrrhalta (Coleoptera: Chrysomelidae: Galerucinae) species with maculate elytra. Zootaxa 4664 (1): 518-534, DOI: 10.11646/zootaxa.4664.4.4
D61087F051693232FF1EE8762C72FD89.taxon	diagnosis	Differential diagnosis. The habitus of Pyrrhalta schillhammeri sp. nov. is similar to that of P. lucka sp. nov. The males of both species possess subclavate antennae and modified ventral sides of pro- and mesotarsomeres I. Both species differ in the structure of aedeagus. The aedeagus of P. schillhammeri sp. nov. has the apical half narrow and slightly curved left while the aedeagus of P. lucka sp. nov. has an elongate, flat, lanceolate apical half (Figs 5 – 6). The additional similar species, P. tsoui sp. nov. from Taiwan and Laos, has both elytral spots near the suture round (elongate in P. schillhammeri sp. nov.), antennae without distinctly enlarged apical antennomeres (antennae subclavate in P. schillhammeri sp. nov.), and ventral sides of pro- and mesotarsomeres I not modified (modified in P. schillhammeri sp. nov., see Figs 11, 16).	en	Bezděk, Jan, Lee, Chi-Feng (2019): Revision of Pyrrhalta (Coleoptera: Chrysomelidae: Galerucinae) species with maculate elytra. Zootaxa 4664 (1): 518-534, DOI: 10.11646/zootaxa.4664.4.4
D61087F051693232FF1EE8762C72FD89.taxon	distribution	Distribution. Laos.	en	Bezděk, Jan, Lee, Chi-Feng (2019): Revision of Pyrrhalta (Coleoptera: Chrysomelidae: Galerucinae) species with maculate elytra. Zootaxa 4664 (1): 518-534, DOI: 10.11646/zootaxa.4664.4.4
D61087F051693232FF1EE8762C72FD89.taxon	etymology	Etymology. Dedicated to Harald Schillhammer (NMW) who collected the holotype.	en	Bezděk, Jan, Lee, Chi-Feng (2019): Revision of Pyrrhalta (Coleoptera: Chrysomelidae: Galerucinae) species with maculate elytra. Zootaxa 4664 (1): 518-534, DOI: 10.11646/zootaxa.4664.4.4
D61087F051683231FF1EE8D428FBFB01.taxon	description	(Figs 7, 24, 36)	en	Bezděk, Jan, Lee, Chi-Feng (2019): Revision of Pyrrhalta (Coleoptera: Chrysomelidae: Galerucinae) species with maculate elytra. Zootaxa 4664 (1): 518-534, DOI: 10.11646/zootaxa.4664.4.4
D61087F051683231FF1EE8D428FBFB01.taxon	materials_examined	Type locality. Taiwan, Taitung County, Wulu. Types examined. Holotype: ♂, “ Taiwan: Taitung / Wulu (Îů) / 09. VI. 2011, leg. Uika Ong [w, p] ” (TARI). Paratypes: 2 ♂♂ 9 ♀♀, same label data as in holotype (TARI); 1 ♂, “ Taiwan: Hualien (19551) / Pilu (Ḃñ) / 31. V. 2011, leg. M. - H. Tsou [w, p] ” (TARI); 6 ♀♀, “ Taiwan: Taitung (20420) / Litao (" ũ) / 20. VI. 2011, leg. C. - F. Lee [w, p] ” (TARI); 2 ♀♀, “ Taiwan: Hsinchu / Chienshih (ẊƂ) / 05. VIII. 2012, leg. Y. - L. Lin [w, p] ” (TARI); 2 ♀♀, “ Taiwan: Taitung (# 24657) / Haituan (Ā ầ) / 18. VI. 2013, leg. J. - C. Chen [w, p] ” (TARI); 1 ♀, “ Taiwan: Taoyuan (22817) / Hsuanyuan (Ṙã) / 09. V. 2012, leg. H. Lee [w, p] ” (TARI); 1 ♂, “ Taiwan: Taipei (20713) / Hsinhsien (ª Ễ) / 08. VII. 2011, leg. M. - H. Tsou [w, p] ” (TARI); 1 ♀, “ Taiwan: Taipei (20712) / Fushan (ÑƜ) / 08. VII. 2011, leg. M. - H. Tsou [w, p] ” (TARI); 2 ♀♀, “ C. TAIWAN: Tungpu / 1200 m, Nantou Hsien / 23 - 27. VII. 1984 / K. C. Chou & C. H. Yang [o, p] ” (TARI); 1 ♀, “ C. TAIWAN: Meifeng / 2250 m, Nantou Hsien / 31. VIII- 2. IX. 1982 / L. Y. Chou & K. C. Chou [o, p] ” (TARI); 1 ♀, “ C. TAIWAN: Sungkang / 2100 m, Nantou Hsien / 13 - 15. IX. 1984 / K. S. Lin & S. C. Lin [o, p] ” (TARI); 1 ♂, “ Laos-NE, Houa Phan prov., / 20 ° 12 - 13.5´N 103 ° 59.5´104.01´E, / Ban Saluei → Phou Pane Mt., / 1340 - 1870 m, 2 – 22. vi. 2011, / Vít Kubáň & Lao coll. Lgt. [w, p] // Primary mountain forest, / at light + individual collecting, / Laos 2008 Expedition / National Muzeum Prague, / Czech Republic [w, p] ” (NMPC). The specimens are provided with one additional printed red label: “ HOLOTYPUS, [or PARATYPUS, respectively], / Pyrrhalta / tsoui sp. nov., / Bezděk & Lee, 2019 ”.	en	Bezděk, Jan, Lee, Chi-Feng (2019): Revision of Pyrrhalta (Coleoptera: Chrysomelidae: Galerucinae) species with maculate elytra. Zootaxa 4664 (1): 518-534, DOI: 10.11646/zootaxa.4664.4.4
D61087F051683231FF1EE8D428FBFB01.taxon	description	Description. Body length: ♂♂ 3.6 – 4.1 mm (holotype 3.8 mm); ♀♀ 4.0 – 4.8 mm. Male (holotype). Dorsally reddish brown except following parts black: middle part of vertex, three spots on pronotum (one very small median and two small round lateraly), basal half of scutellum, each elytron with five spots: one round spot between humerus and scutellum, not touching anterior elytral margin; two spots before middle (outer small elongate, inner larger round); two spots subapically. Legs and antennae reddish brown. Ventral side: head and prosternum reddish brown, mesoventrite black, brown medially, metaventrite black with brown posterior margin, abdomen reddish brown. Head. Labrum transverse with rounded anterior margin, surface with transverse row of several punctures bearing long pale setae. Anterior clypeal margin straight. Anterior part of head semiopaque, sparsely covered with punctures and longer pale setae. Interantennal space narrow, 0.66 times as wide as transverse diameter of antennal insertion. Interocular space 2.10 as wide as transverse diameter of eye. Frontal tubercles subquadrate, semiopaque, covered with microsculpture, separated by thin sulcus, laterally demarcated by thin furrow, posterior margins separated by shallow impressed line from vertex. Vertex dull, covered with microsculpture, small punctures and short setae. Antennae thin, 0.66 times as long as body, length ratios of antennomeres I – XI equal to 13 - 6 - 11 - 10 - 8 - 9 - 7 - 7 - 7 - 7 - 12, last five antennomeres slightly dilated, antennomere IV with thin elevated ridge ventrally, antennomeres VIII-XI slightly flattened ventrally. Pronotum transverse, 1.96 times as wide as long, widest at anterior third. Anterior margin widely shallowly concave, lateral margins with rounded middle part, anteriorly and posteriorly convergent and nearly straight, posterior margin slightly concave, lateral parts of posterior margin oblique and concave. All margins thinly bordered. Anterior and posterior angles pointed and bearing long pale seta. Surface with distinct elevation along anterior margin, laterally widened and curved posteriorly, disc transversely impressed. Elevation glabrous and lustrous, disc densely covered with small punctures and short setae. Scutellum triangular, with rounded apex, surface very densely covered with microsculpture, small punctures and short setae. Elytra moderately convex, posteriorly slightly widened, 0.71 as long as body, 1.59 times as long as wide, dull, covered with small confused punctures with interspaces as wide as or narrower than puncture diameters and dense short pale setae, surface slightly impressed in subscutellar area, rest of surface even. Humeral calli well developed. Epipleura wide, in apical quarter gradually narrowing towards apex, densely covered with short pale setae. Macropterous. Ventrally semiopaque, covered with longer pale setae. Last abdominal ventrite transverse, with deep V-shaped incision. Legs moderately narrow. Protarsi: protarsomere I subtriangular, not modified ventrally, II subtriangular, short, length ratios of protarsomeres I – IV equal to 6 - 5 - 4 - 8. Mesotarsi: mesotarsomere I elongate subtriangular, not modi- fied ventrally, length ratios of mesotarsomeres I – IV equal to 7 - 6 - 5 - 9. Metatarsi: metatarsomere I elongate subtrian- gular, length ratios of metatarsomeres I – IV equal to 8 - 7 - 6 - 10. Claws bifid. Aedeagus (Fig. 7) with apical half regularly curved in lateral view. In dorsal view, aedeagus asymmetrical, relatively narrow, with quadrate apex turned left and dorsally. Female. Antennomere IV without ventral ridge. Last abdominal ventrite entire, without V-shaped incision. Spermatheca (Fig. 24) with C-shaped cornu and subsphaerical nodulus, cornu and nodulus of the same width, basal part of ductus spermathecae finger-like with oblique edge where thin ductus is attached. Variability. Middle pronotal black spot variable, usually elongate and as large as lateral spots.	en	Bezděk, Jan, Lee, Chi-Feng (2019): Revision of Pyrrhalta (Coleoptera: Chrysomelidae: Galerucinae) species with maculate elytra. Zootaxa 4664 (1): 518-534, DOI: 10.11646/zootaxa.4664.4.4
D61087F051683231FF1EE8D428FBFB01.taxon	diagnosis	Differential diagnosis. The habitus of Pyrrhalta tsoui sp. nov. (Fig. 36) is similar to that of P. lucka sp. nov. and P. schillhammeri sp. nov., but P. tsoui sp. nov. can be distinguished by the following characters: antennae not subclavate in males (subclavate in P. schillhammeri sp. nov. and P. lucka sp. nov.), pro- and mesotarsomeres I not modified (modified in P. schillhammeri sp. nov. and P. lucka sp. nov., Figs 10 – 11, 15 – 16), and aedeagus with apical part curved upwards (aedeagus flat in P. schillhammeri sp. nov. and P. lucka sp. nov., Figs 5 – 7).	en	Bezděk, Jan, Lee, Chi-Feng (2019): Revision of Pyrrhalta (Coleoptera: Chrysomelidae: Galerucinae) species with maculate elytra. Zootaxa 4664 (1): 518-534, DOI: 10.11646/zootaxa.4664.4.4
D61087F051683231FF1EE8D428FBFB01.taxon	distribution	Distribution. Taiwan, Laos.	en	Bezděk, Jan, Lee, Chi-Feng (2019): Revision of Pyrrhalta (Coleoptera: Chrysomelidae: Galerucinae) species with maculate elytra. Zootaxa 4664 (1): 518-534, DOI: 10.11646/zootaxa.4664.4.4
D61087F051683231FF1EE8D428FBFB01.taxon	etymology	Etymology. Dedicated to Mr. Mei-Hua Tsou, the first member of the TARI research team to collect this new species.	en	Bezděk, Jan, Lee, Chi-Feng (2019): Revision of Pyrrhalta (Coleoptera: Chrysomelidae: Galerucinae) species with maculate elytra. Zootaxa 4664 (1): 518-534, DOI: 10.11646/zootaxa.4664.4.4
