taxonID	type	description	language	source
D61387AEB606526B3DAAF944FF05FE4B.taxon	diagnosis	Diagnosis. A full description of the genus is available in Sinclair (1994). Trichoclinocera is distinguished from other clinocerine genera by the following combination of characters: R 1 setose, R 4 and R 5 normally nearly parallel, apex of cell dm not strongly produced, M 1 and M 2 arising separately from cell dm, fore femur with rows of ventral setae; female abdomen truncate, with tergite 10 setose (acanthophorite-like); apex of phallus membranous, lacking filament.	en	Saigusa, Toyohei, Sinclair, Bradley J. (2016): Revision of the Japanese species of Trichoclinocera Collin (Diptera: Empididae: Clinocerinae). Zootaxa 4103 (3): 201-229, DOI: 10.11646/zootaxa.4103.3.1
D61387AEB606526B3DAAF944FF05FE4B.taxon	discussion	Remarks. The appearance of the apex of the phallus is dependent on the method of clearing and state of the specimen. The membrane may be destroyed in strong acids or bases, and it may not expand as evenly from specimen to specimen.	en	Saigusa, Toyohei, Sinclair, Bradley J. (2016): Revision of the Japanese species of Trichoclinocera Collin (Diptera: Empididae: Clinocerinae). Zootaxa 4103 (3): 201-229, DOI: 10.11646/zootaxa.4103.3.1
D61387AEB606526B3DAAF944FF05FE4B.taxon	distribution	Distribution. The genus occurs in the Palearctic, Nearctic and Oriental regions, and there are more than 30 additional undescribed species known from Burma, Bhutan, China, India, Nepal, Pakistan, Russia, Thailand and Vietnam.	en	Saigusa, Toyohei, Sinclair, Bradley J. (2016): Revision of the Japanese species of Trichoclinocera Collin (Diptera: Empididae: Clinocerinae). Zootaxa 4103 (3): 201-229, DOI: 10.11646/zootaxa.4103.3.1
D61387AEB605526A3DAAF94CFC16FF02.taxon	description	Description. See Sinclair & Saigusa (2005).	en	Saigusa, Toyohei, Sinclair, Bradley J. (2016): Revision of the Japanese species of Trichoclinocera Collin (Diptera: Empididae: Clinocerinae). Zootaxa 4103 (3): 201-229, DOI: 10.11646/zootaxa.4103.3.1
D61387AEB605526A3DAAF94CFC16FF02.taxon	distribution	Distribution. This species is restricted to the Japanese island of Honshu (Sinclair & Saigusa 2005, fig. 7), with flight records from mid-June to mid-September.	en	Saigusa, Toyohei, Sinclair, Bradley J. (2016): Revision of the Japanese species of Trichoclinocera Collin (Diptera: Empididae: Clinocerinae). Zootaxa 4103 (3): 201-229, DOI: 10.11646/zootaxa.4103.3.1
D61387AEB605526A3DAAF94CFC16FF02.taxon	discussion	Remarks. This species is usually found in shallow streamlets in forests.	en	Saigusa, Toyohei, Sinclair, Bradley J. (2016): Revision of the Japanese species of Trichoclinocera Collin (Diptera: Empididae: Clinocerinae). Zootaxa 4103 (3): 201-229, DOI: 10.11646/zootaxa.4103.3.1
D61387AEB604526F3DAAFF31FDBBFBEA.taxon	materials_examined	Type material examined. HOLOTYPE ♂, labelled: “ [Kyushu] / Magaribuchi / Sawara / FUKUOKA / 22. IV. 1962 / T. SAIGUSA ” (KUMF). PARATYPES: same data as holotype (29 ♂, 46 ♀, KUMF); same locality as holotype, 2. v. 1962, (3 ♂, 1 ♀, CNC, USNM); same locality as holotype, 2. v. 1962 (16 ♂, 19 ♀, KUMF). Additional material examined. Some 360 specimens (all KUMF) from the following prefectures: JAPAN. HONSHU. Iwate: Hayachinesan, 2. viii. 1963, TS. Kyoto: Momoi [35 ° 9 ′ 16 ″ N 135 ° 47 ′ 18 ″ E], 30. v. 1999, H. Ohishi; Kibune, Sakyo [35 ° 7 ′ 15 ″ N 135 ° 45 ′ 47 ″ E], 5. v. 2012, H. Ohishi. Mie: Hirakura, 7. vi. 1965, A. Nakanishi. Nagano: Shimashimadani, Azumi-mura, 6 – 24. vi. 1975, TS. Nara: Totsukawa, 17. vi. 1963, I. Miyagi. Osaka: Myokensan Nose [34 ° 55 ′ 40 ″ N 135 ° 27 ′ 51 ″ E], 29. iv. 1962, TS; Settsuyabakei, Takatsuki, 30. iv. 1962, TS. Ya m an a shi: Kanayama, Sudama-chô, 2. vi. 1962, TS; Kanayama, Sudama-chô, 27. v. 1975, A. Nakanishi; Shôsenkyô, Kofu-shi, 5. vi. 1962, TS. KYUSHU. Fukuoka: Magaribuchi, 2. v. 1962, TS. Oita: Kyûsuikyô, Kujûsan, 20. iv. 1968, TS. SHIKOKU. Ehime: Omogokei, 27. v. 1967, A. Nakanishi. Kochi: Myoga, Monobe-mura, 19. v. 1988, TS. Tokushima: Tsurigisan, 4. vi. 1981, K. Ôhara; Tsurugisan, 1000 m, 7. v. 1987, TS; Tochikubadani, Tônomaru, 18 – 22. vi. 1988, TS; Inodani, Tsurugisan, 21. vi. 1988, TS; Tsurugisan, Higashiiya, 4. vi. 1981, TG. Recognition. Distinguished from other Japanese species by dark legs, bright blue pruinescence of the face, spine on hind tibia lacking, and stout anterodorsal and posterodorsal bristles on mid and hindlegs lacking, and generally darker wings. Re-description. Wing length 4.2 – 5.0 mm. Male. Clothed in blue pruinescence except scutum brown. Head rounded; compound eyes large, rounded (Fig. 3 A). Face with blue pruinescence and median light brown stripe; face narrowest just below antennae, slightly wider than width of antennal sockets; lower margin of face with shallow cleft and low carina; gena one-fifth as wide as height of eye. Ocellar bristles longest among cranial bristles; occiput with 2 pairs of bristles posterior to ocellar triangle; postocular with row of short bristles along margin of eye, upper half darker and more stout than lower bristles. Arista-like stylus long and slender, tapered apically. Palpus cylindrical, slender, length less than half height of eye. Pleura, postpronotal lobe, notopleuron and scutellum with pale blue pruinescence, much more faint on scutum; posterolateral corner of scutum near postalar ridge concolorous with surrounding region; prescutellar depression brown; scutum faintly bivittate. Thoracic setae long, some longer than width of eye; acrostichal setulae lacking; 6 pairs dc; 1 pprn; 1 presut spal; 2 npl; 2 psut spal; 1 pal; 1 pair sctl, lacking setulae on disc; several setulae on notopleural depression and postpronotal lobe. Laterotergite with patch of pale setae; proepisternum with long pale setae, anepisternum with dark setulae along posterior margin; setae lacking on katepisternum. Wing narrow, infuscate (Fig. 16 A); pterostigma very narrow, faint, parallel-sided or absent. Basal costal seta subequal to length of posterior dc. Row of setulae from base of R to tip of R 1; cell dm produced apically; R 4 and R 5 parallel except for extreme apex. Halter brown with pale base. Legs generally dark towards tips, with coxae and ventral surface of femora with blue pruinescence. Coxae with long, pale marginal setae. Fore femur (Figs 17 A, B) with 1 – 2 stout anterior setae in oblique row on apical third; anteroventral margin with row of stout setae of variable lengths, sometimes expanded into 2 – 3 rows on basal half of femur; posteroventral margin with row of 5 – 7 long, stout dark setae nearly subequal to width of femur on basal two-thirds; base of femur with row of 4 – 6 long, pale setae, some longer than width of femur. Fore tibia with biserial row of short, stout setae beneath. Mid and hind femora with several pairs of preapical antero- and posteroventral setae; lacking antero- and posterodorsal bristles. Hind tibia lacking posterodorsal and anterodorsal basal setae. Tarsus length longer than corresponding fore tibia, shorter than mid and hind tibiae. Empodium pulvilliform, slightly shorter than claw, longer than half length of tarsomere. Hypandrium short, quadrate, slightly longer than epandrium (Figs 3 B, 4 A, 9 A). Phallus extended slightly beyond apex of clasping cercus, slightly arched at mid-length; apex with slender, arched anterior extension enclosing membrane, posterior extension expanded; ejaculatory apodeme strongly curved at basal third. Surstylus with narrow apical process bearing short microtrichia; anterior margin with tooth-like projections. Clasping cercus parallel-sided, slender, finger-like, bearing peg-like setae on inner dorsal margin; apex mostly truncate with tapered anteroapical corner.	en	Saigusa, Toyohei, Sinclair, Bradley J. (2016): Revision of the Japanese species of Trichoclinocera Collin (Diptera: Empididae: Clinocerinae). Zootaxa 4103 (3): 201-229, DOI: 10.11646/zootaxa.4103.3.1
D61387AEB604526F3DAAFF31FDBBFBEA.taxon	description	Female. Similar to male except as follows: long, dark posteroventral setae of fore femur less stout, more widely separated, intermixed with short, stout setae, longer than width of femur (Figs 18 A, B). Posterior marginal setae of tergum 7 rather sparse and fine, evenly distributed, not shortened on dorsal surface; sternum 7 not emarginated posteriorly, with rather fine posterior marginal setae evenly distributed but somewhat sparser midventrally; tergum 8 with rather short, straight more or less stiff posterior marginal setae, sparser laterally than dorsally; sternum 8 not emarginated posteriorly, densely clothed with minute pile-like setulae, sparsely clothed with minute setulae on lateral surface; tergite 10 with 30 – 35 rather short, apically curved setae, setae along dorsal margin towards apex mostly stronger and only slightly tapered apically, setae on posterolateral and basal areas shorter and more slender, occasionally with several straight setae basally; sternite 10 with short setae as long as sternum (Figs 12 A – C).	en	Saigusa, Toyohei, Sinclair, Bradley J. (2016): Revision of the Japanese species of Trichoclinocera Collin (Diptera: Empididae: Clinocerinae). Zootaxa 4103 (3): 201-229, DOI: 10.11646/zootaxa.4103.3.1
D61387AEB604526F3DAAFF31FDBBFBEA.taxon	distribution	Distribution. This species is known from the Japanese islands of Honshu, Shikoku and Kyushu (Fig. 5), with adults collected from April to August.	en	Saigusa, Toyohei, Sinclair, Bradley J. (2016): Revision of the Japanese species of Trichoclinocera Collin (Diptera: Empididae: Clinocerinae). Zootaxa 4103 (3): 201-229, DOI: 10.11646/zootaxa.4103.3.1
D61387AEB60152613DAAFB64FE02FC8A.taxon	materials_examined	Type material. HOLOTYPE ♂, labelled: “ (KYUSHU) / Kumamoto / Minamata / 6. vii. 1968 / T. Saigusa ”; “ [HOLOTYPE] / Trichoclinocera / gracilis / Saigusa et / Sinclair, 2016 [red label] ” (KUMF). PARATYPES: JAPAN. HONSHU. Hyogo: Oginosen [35 ° 26 ′ 21 ″ N 134 ° 26 ′ 26 ″ E], 20. vii. 1962, A. Nakanishi (5 ♂, 1 ♀, KUMF, CNC). Kyoto: Kibune, 8. vi. 1963, TS (9 ♂, 8 ♀, BMNH, CNC, KUMF). Nagano: Hakuba, trib of Himekawa River, 5 – 6. viii. 1998, BJS (1 ♂, CNC). Tokyo: Takaosan, 27. vi. 1965, A. Katô (1 ♂, 2 ♀, KUMF, CNC). Yamanashi: Kanayama, Sudama, 19. viii. 1962, TS (1 ♀, KUMF); Sekisuiji, Kofu, 1. vi. 1960, TS (1 ♂, KUMF); Shosenkyo, 6. vi. 1962, TS (18 ♂, 14 ♀, KUMF, BMNH, CNC). KYUSHU. Kumamoto: same data as holotype (8 ♂, 8 ♂, CNC, KUMF). SHIKOKU. Ehime: Nametoko, Uwajima, 24. v. 1988, TS (1 ♀, KUMF). Recognition. Distinguished from other Japanese species by its smaller size and more delicate appearance, ventral setae of fore femur mostly slender and pale, lacking anterodorsal and posterodorsal bristles on mid and hindlegs, and darker face.	en	Saigusa, Toyohei, Sinclair, Bradley J. (2016): Revision of the Japanese species of Trichoclinocera Collin (Diptera: Empididae: Clinocerinae). Zootaxa 4103 (3): 201-229, DOI: 10.11646/zootaxa.4103.3.1
D61387AEB60152613DAAFB64FE02FC8A.taxon	description	Description. Wing length 3.8 – 4.0 mm. Male. Clothed in blue pruinescence except head, scutum, and dorsum of abdomen brown. Head rounded; compound eyes large, rounded. Face mostly brown to greyish, with faint median brown stripe; face narrowest just below antennae, slightly wider than width of antennal sockets; lower margin of face with shallow cleft and low carina; gena one-fifth as wide as height of eye. Ocellar bristles longest among cranial bristles; occiput with 2 pairs of bristles posterior to ocellar triangle; postocular with row of short bristles along margin of eye, upper half darker and more stout than lower bristles. Arista-like stylus long and slender, slightly tapered apically. Palpus cylindrical, slender, length less than half height of eye. Pleura, postpronotal lobe and notopleuron with pale blue pruinescence, much more faint on scutum; posterolateral corner of scutum near postalar ridge concolorous with surrounding region; prescutellar depression brown; scutum faintly bivittate. Thoracic setae long, some longer than width of eye; acrostichal setulae lacking; 6 pairs dc; 1 pprn; 1 presut spal; 2 – 3 npl; 2 psut spal; 1 pal; 1 pair sctl, lacking setulae on disc; several setulae on notopleural depression and postpronotal lobe. Laterotergite with patch of pale setae; prothoracic episternum with long pale setae, setae lacking on katepisternum, metepisternum with pale setae on posterodorsal margin. Wing narrow, infuscate (Fig. 16 B); pterostigma very narrow, faint, parallel-sided. Basal costal seta subequal to length of posterior dc. Row of setulae from base of R to tip of R 1; cell dm produced apically; R 4 and R 5 generally parallel, divergent beyond mid-length. Halter brown with pale base. Legs uniformly dark, coxae with blue pruinescence. Coxae with long pale, marginal setae. Fore femur (Figs 17 C, D) with 2 stout anterior bristles in oblique row on apical third; anteroventral and posteroventral margins with continuous row of setae, increasing in length basally, increasing in stoutness on apical third; basal setae longer than width of femur. Fore tibia with biserial row of short, stout setae beneath. Foreleg tarsomeres with pale ventral pubescence. Mid and hind femora with several pairs of preapical antero- and posteroventral setae; anterodorsal bristles on apical half; lacking posterodorsal bristles. Hind tibia lacking pair of posterodorsal and anterodorsal basal setae; row of short bristles or preapical bristles lacking. Tarsus length longer than corresponding fore and mid tibiae, shorter than hind tibia. Empodium pulvilliform, slightly shorter than claw, longer than half length of tarsomere. Hypandrium short, broad, quadrate, slightly larger than epandrium (Figs 4 B, 9 B). Phallus extended slightly beyond apex of clasping cercus, slightly arched at mid-length; apex greatly expanded into 3 membranous lobes as seen laterally: anterior lobe single, posterior lobes paired; ejaculatory apodeme gently bent at anterior third. Surstylus with pair of teeth-like processes on anterior margin, each process bearing seta. Clasping cercus wide medially, strongly tapered or more slender, not attenuated on apical half, bearing peg-like setae on inner dorsal margin. Female. Similar to male except as follows: fore femur with only a few slender, pale posteroventral setae near base; fore tibia lacking biserial rows of erect setae beneath (Figs 18 C, D). Posterior marginal setae of tergum 7 moderately fine and longish, slightly shorter dorsally; sternum 7 with straight posterior margin bearing evenly distributed setae similar to setae on terga, slightly longer laterally, shorter on midventral area; tergum 8 with posterior marginal setae moderately dense, longish and weakly curved ventrally on dorsal surface; sternum 8 not distinctly emarginated posteriorly, densely clothed with minute pile-like setulae, sparsely with minute setulae on lateral surface; tergite 10 with 20 – 25 long and rather slender apically curved setae, somewhat stouter and shorter towards apex of tergite; sternite 10 with longish setae, longest 1.5 times as long as sternum (Figs 12 D – F).	en	Saigusa, Toyohei, Sinclair, Bradley J. (2016): Revision of the Japanese species of Trichoclinocera Collin (Diptera: Empididae: Clinocerinae). Zootaxa 4103 (3): 201-229, DOI: 10.11646/zootaxa.4103.3.1
D61387AEB60152613DAAFB64FE02FC8A.taxon	distribution	Distribution. This species is known from central Honshu and Shikoku (Fig. 6).	en	Saigusa, Toyohei, Sinclair, Bradley J. (2016): Revision of the Japanese species of Trichoclinocera Collin (Diptera: Empididae: Clinocerinae). Zootaxa 4103 (3): 201-229, DOI: 10.11646/zootaxa.4103.3.1
D61387AEB60152613DAAFB64FE02FC8A.taxon	etymology	Etymology. The specific epithet “ gracilis ” refers to the delicate long legs and foreleg chaetotaxy compared to other Japanese species of Trichoclinocera.	en	Saigusa, Toyohei, Sinclair, Bradley J. (2016): Revision of the Japanese species of Trichoclinocera Collin (Diptera: Empididae: Clinocerinae). Zootaxa 4103 (3): 201-229, DOI: 10.11646/zootaxa.4103.3.1
D61387AEB60152613DAAFB64FE02FC8A.taxon	discussion	Remarks. This species appears to be much rarer in collections than other Japanese species of Trichoclinocera, with only short series collected.	en	Saigusa, Toyohei, Sinclair, Bradley J. (2016): Revision of the Japanese species of Trichoclinocera Collin (Diptera: Empididae: Clinocerinae). Zootaxa 4103 (3): 201-229, DOI: 10.11646/zootaxa.4103.3.1
D61387AEB60F52633DAAFC44FC23F808.taxon	materials_examined	Type material. HOLOTYPE ♂, labelled: “ (YAMANASHI) / Kanayama / Sudama / 19. viii. 1962 / T. Saigusa ”; “ [HOLOTYPE] / Trichoclinocera / grandis / Saigusa et / Sinclair, 2016 [red label] ” (KUMF). PARATYPES: JAPAN. HOKKAIDO: Asiribetu, Sapporo, 30. vi. 1962, S. Takagi (4 ♂, HUS); Berabonai, Ashoro, 24. vii. 1967, TS (6 ♂, KUMF); Fujikawa, Nukabira, 19. vii. 1967, HS (2 ♂, 1 ♀, KUMF); Hattari, Atsuta R, 14. vi. 1998, BJS (8 ♂, 2 ♀, CNC); Kamishibinai, Biei-chô, 27. vi. 1967, TS (2 ♀, KUMF); Kamiwashippu, Ashoro-machi, 19. vi. 1967, TS (3 ♂, 2 ♀, KUMF); Kiyokawa, Ashoro, 27. vii. 1962, 23. vii. 1967, TS (19 ♂, 18 ♀, KUMF), same locality, 29. vii. 1967, HS (3 ♂, 3 ♀, KUMF); Mt. Daisetsu, 30. vii. 1967, K. Kusigemati (2 ♂, 4 ♀, HUS); Mt. Izari, 8. vii. 1990, A. Iwasaki (2 ♂, 1 ♀, KUMF); Nishinosawa, Eniwa-dake, 30. vi. 1986, TS (6 ♂, 4 ♀, KUMF); Pirikapetan, Satsunaidake, 27. vii. 1967, TS (2 ♂, 1 ♀, KUMF); Rausudake, 5. viii. 1967, TS (15 ♂, 13 ♀, KUMF); Satsunai-dake, 700 – 1000 m, 28. vii. 1967, TS (1 ♂, 3 ♀, KUMF); NW of Shikotsu-ko, 30. vi. 1986, TS (1 ♂, KUMF); Siretoko, 10 – 17. vii. 1965, T. Kumata, I. Miyagi (9 ♂, 4 ♀, HUS); Mt. Soronuma, 23. ix. 1966, K. Kusigemati (2 ♂, 2 ♀, HUS); Soranumadake, Sapporo, 8. viii. 1962, TS (4 ♂, 10 ♀, KUMF); Soranumadake, 8. viii. 1962, 18. viii. 1970, S. Takagi (11 ♂, 6 ♀, HUS); Syokanbetsu, 21. vii. 1964, I. Miyagi (1 ♂, 3 ♀, HUS); Takimo, Sapporo, Ashiribetsu Falls, 15. vi. 1998, BJS (5 ♂, 5 ♀, CNC); Tenninkyo, Taisetsuzan, 27. vi. 1976, TS (21 ♂, 24 ♀, KUMF); Tenninkyo, Daisetsuzan, 800 m, 10 – 14. vii. 1986, D. M. Wood (1 ♀, CNC); Toikanbetsu, 12. viii. 1965, T. Kumata, K. Kusige (9 ♂, 10 ♀, HUS); Yukomanpetsu, Taisetsuzan, 26. vi. 1967, TS (2 ♂, 9 ♀, KUMF). HONSHU. Aomori: Oirasekeikoku nr. Towadako, 14. vii. 1962, TS (5 ♂, 6 ♀, KUMF). Gifu: Futatsuya, Kawai-mura, 11. vi. 1977, JE (1 ♂, 6 ♀, KUMF). Iwate: Hayatine, 4. viii. 1964, S. Takagi (12 ♂, 5 ♀, HUS). Kyoto: Kibune, 8. vi. 1963, TS (6 ♂, 4 ♀, KUMF); same locality, 8. v. 1968, J. Yukawa (1 ♂, KUMF). Mie: Hirakura, 7. vi. 1965, A. Nakanishi (2 ♂, KUMF). Nagano: Akagawara, Senjôdake, 8. vii. 1963, TS (5 ♂, 2 ♀, KUMF); Hakuba, trib & Himekawa R, 5 – 6. viii. 1998, BJS (9 ♂, 6 ♀, CNC); Nakabusa-onsen, Tsubakurodake, 17. viii. 1962, TS (1 ♂, 3 ♀, KUMF); Shimashimadani, 9. vi. 1975, JE (1 ♂, KUMF); Spa. Nakabusa, 21 – 24. viii. 1961, A. Kato (2 ♂, 3 ♀, KUMF). Niigata: Mt. Donden, Sado, 18. vii. 1955, K. Baba (1 ♀, KUMF). Shiga: Hirasan, 300 – 700 m, 4. vi. 1969, K. Kanmiya (7 ♂, 6 ♀, KUMF). Tottori: Daisen, Daisen-chô, 9. vi. 1981, TG (2 ♂, 5 ♀, KUMF); Motodani, Daisen, 12. vi. 1963, TS (1 ♀, KUMF). Ya m a na sh i: Amarizawa, Nirazaki, 8. vi. 1962, 5. vi. 1975, TS (3 ♂, 3 ♀, KUMF); Hirogawara, Kitadake, 19. viii. 1978, JE (1 ♂, 1 ♀, KUMF); Hontanigawa, Kanayama, 23. viii. 1978, JE (2 ♀, KUMF); Kanayama, Sudama, 10 – 12. viii. 1961, 5. vi., 19. viii. 1962, 30. vi. 1963, 12 – 13. viii. 1973, 1 – 3. vi. 1975, TS (51 ♂, 42 ♀, KUMF, 1 ♂, BMNH), same except, 27. v. 1975, A. Nakanishi (6 ♂, 9 ♀, KUMF), same except, 3, 16, 23. vi., 28. vii., 2, 9, 11. viii., 1, 16. ix. 1975, JE (57 ♂, 36 ♀, KUMF), same except, MT, 16, 17, 18, 20 – 21, 24 – 25, 30. vi., 26. vii., 7, 24. viii., 1. ix. 1975, JE (24 ♂, 18 ♀, KUMF), same except, LT, 29, 31. viii. 1975, JE (31 ♂, 27 ♀, KUMF); Kurobera, 29. vii. 1975, JE (3 ♂, 2 ♀, KUMF); Ohkanbazawa, Kitadake, 20. viii. 1978, TS (3 ♂, 2 ♀, CNC); same except, 19. viii. 1978, TS (22 ♂, 17 ♀, KUMF); Shôsenkyô, Kôfu-shi, 5. vi. 1962, TS (1 ♂, 2 ♀, KUMF); Tokusatôge, Sudama, 23. vi. 1975, JE (12 ♂, 7 ♀, KUMF); Yanagidaira, Sudama, 2. viii. 1975, JE (1 ♂, KUMF). KYUSHU. Fukuoka: Mizunashi Cave, Mt. Ihara, Maebaru-shi, 14. vi. 1997, BJS (2 ♂, 1 ♀, CNC); Magaribuchi, Fukuoka, 2. v. 1962, TS (1 ♂, 4 ♀, KUMF); Nogochi, 1. v., 7. vi. 1997, 24. iv. 1998, BJS (8 ♂, 7 ♀, CNC); Shiraito Falls, 28. viii. 1999, BJS (2 ♂, CNC); Tainan Pass, Soeda-machi, 5. v. 1998, BJS (2 ♂, CNC). Kumamoto: Haki, Izumi-mura, 11. vii. 1978, TS (3 ♂, 3 ♀, KUMF); Haki, Izumi-mura, 23. v. 1997, BJS (1 ♂, 2 ♀, CNC); Hirasawado, Gokanoshô, 3. viii. 1975, TS (4 ♂, 7 ♀, KUMF); Momiki, Izumi-mura, 2. viii. 1975, TS (8 ♂, 3 ♀, KUMF); Momiki, Izumi-mura, 20. viii. 1999, BJS (3 ♂, 4 ♀, CNC); Shiratoriyama, Izumi-mura, 12 – 14. v. 1983, K. Ôhara (1 ♂, KUMF). Miyazaki: Okueyama, Kitagawa-cho, 3. vi. 1979, TG (4 ♂, 11 ♀, KUMF). Oita: Handakôgen, Kujû, 23. viii. 1962, T. Shirôzu (2 ♀, KUMF); Kujûsan, 28. v. 1967, TS (18 ♂, 20 ♀, KUMF); Kyûsuikei, Kujû-san, 21. v. 1962, TS (6 ♂, 3 ♀, BMNH); Kyûsuikyô, Kujûsan, 19, v. 1962, TS (20 ♂, 23 ♀, KUMF); same locality, 21. v. 1962, TS (50 ♂, 66 ♀; KUMF, CNC). SHIKOKU. Ehime: 1 ♂, 4 ♀, Omogokei, 28. v. 1967, A. Nakanishi (KUMF). Tokushima: Inodani, Tsurugisan, 21. vi. 1988, TS (17 ♂, 21 ♀, KUMF); Ichiu, 3. vi. 1981, TG (82 ♂, 58 ♀, KUMF); Tsurugisan, 4. vi. 1981, K. Ôhara (17 ♂, 12 ♀, KUMF); Tsurugisan, Higashiya, 4. vi. 1981, TG (2 ♂, 1 ♀, KUMF); Tochikubodani, Tomaru, 22. vi. 1988, TS (2 ♀, KUMF). Recognition. Distinguished from other Japanese species by its large size, presence of stout anterodorsal bristles on mid and hindlegs, posterodorsal bristles absent, and face with bright blue pruinescence.	en	Saigusa, Toyohei, Sinclair, Bradley J. (2016): Revision of the Japanese species of Trichoclinocera Collin (Diptera: Empididae: Clinocerinae). Zootaxa 4103 (3): 201-229, DOI: 10.11646/zootaxa.4103.3.1
D61387AEB60F52633DAAFC44FC23F808.taxon	description	Description. Wing length 5.6 – 6.4 mm. Male. Clothed in blue pruinescence except vertex of head and scutum brown. Head rounded; compound eyes large, rounded (Fig. 3 C). Face with bright blue pruinescence, with broad, median brown patch on carina and narrow dorsal stripe; face narrowest at middle, slightly wider than antennal sockets, lower margin of face with deep cleft and distinct carina; gena one-fifth as wide as height of eye. Ocellar bristles longest among cranial bristles; occiput with 2 – 4 pairs of bristles posterior to ocellar triangle; postocular with row of short bristles along margin of eye, upper third darker and more stout than lower bristles. Arista-like stylus long and slender, tapered apically. Palpus clavate, length greater than half height of eye. Pleura, postpronotal lobe, notopleuron and postalar, prescutellar depression, and scutellum with bright blue pruinescence; posterolateral corner of scutum near postalar ridge bluish; scutum faintly bivittate. Thoracic setae long, some ca. as long as width of eye; acrostichal setulae lacking; 6 – 8 pairs dc; 1 pprn; 1 presut spal; 3 npl; 2 psut spal; 1 pal; 1 pair sctl, lacking setulae on disc; several setulae on postpronotal lobe and notopleural depression. Laterotergite with patch of pale setae; prothoracic episternum with long pale setae, setae lacking on katepisternum, and hind margins of metepisternum. Wing narrow, infuscate (Fig. 16 C); pterostigma very narrow, faint, parallel-sided. Basal costal seta shorter than posterior dc. Row of setulae from base of R to tip of R 1; cell dm produced somewhat apically; R 4 and R 5 somewhat curved, parallel except extreme apex divergent. Halter brown with pale base. Legs greyish, generally darker towards tips, with coxae and trochanters clothed in blue pruinescence. Coxae with long pale, marginal setae. Fore femur with 2 stout anterior setae in horizontal row on apical third (Figs 17 C, D); row of 4 – 5 stout anteroventral setae on apical half, length ca. half width of femur, intermixed with 2 – 3 short setae; basal half with row of ca. 10 very short, stout setae and 2 – 4 anteroventral setae, less stout than apical setae, length ca. one-third width of femur; posteroventral margin with 3 – 5 setae on apical half, less stout than anteroventral setae, with 2 – 6 short setulae intermixed; basal half with row of pale setae, increasing in length basally to longer than basal width of femur. Fore tibia with biserial row of short, stout setae beneath. Foreleg tarsomeres with white, dense ventral pubescence, length ca. half width of tarsomeres. Mid and hind femora with widely spaced pairs of antero- and posteroventral, and anterodorsal bristles on apical two-thirds; posterodorsal bristles absent. Base of hind tibia with pair of divergent dorsal setae (Fig. 3 D) (1 posterodorsal and 1 anterodorsal); anteroventral margin with row of short bristles, less than width of tibia. Tarsus length longer than corresponding fore and mid tibiae, shorter than hind tibia. Empodium pulvilliform, slightly shorter than claw, ca. half length of tarsomere. Hypandrium quadrate, shorter than epandrium (Figs 4 C, 9 C). Phallus not extended beyond apex of clasping cercus, slightly arched at mid-length; apex with anterior, slender sickle-shaped extension enclosing membrane, posterior extension, expanded; ejaculatory apodeme nearly straight at apical third. Surstylus narrow, short, with membranous posterior margin and 2 short anterior teeth-like processes. Clasping cercus broad basally, strongly tapered to rounded apex on apical half; bearing peg-like setae on inner dorsal margin. Female. Similar to male except as follows: basal anteroventral bristles of fore femur absent, replaced with long, pale setae; posteroventral basal setae less pronounced. Fore tibiae lacking biserial row of erect setae beneath (Figs 18 E, F). Posterior marginal setae of tergum 7 long, strong, dense and evenly distributed; sternum 7 weakly emarginated posteriorly, bearing strong posterior marginal setae somewhat indistinctly gathered in midventral and sublateral groups, but not so clearly grouped as in T. takagii sp. nov., 1 – 2 most lateral setae usually more or less isolated from sublateral group; tergum 8 with posterior marginal setae long, dense and weakly curved ventrally on dorsal area, slightly shorter and straight on lateral areas; sternum 8 moderately emarginated posteriorly, densely clothed with minute pile-like setulae, sparsely clothed with minute setulae on lateral surface; tergite 10 with 25 – 30 long and rather stout apically curved setae, slightly shorter towards tip, with 1 – 3 setae stouter than others; sternite 10 with setae almost as long as sternite (Figs 13 A – C).	en	Saigusa, Toyohei, Sinclair, Bradley J. (2016): Revision of the Japanese species of Trichoclinocera Collin (Diptera: Empididae: Clinocerinae). Zootaxa 4103 (3): 201-229, DOI: 10.11646/zootaxa.4103.3.1
D61387AEB60F52633DAAFC44FC23F808.taxon	distribution	Distribution. This is a wide ranging species, known from the Japanese islands of Hokkaido, Honshu, Shikoku, and Kyushu (Fig. 7). Adults are collected from April to September.	en	Saigusa, Toyohei, Sinclair, Bradley J. (2016): Revision of the Japanese species of Trichoclinocera Collin (Diptera: Empididae: Clinocerinae). Zootaxa 4103 (3): 201-229, DOI: 10.11646/zootaxa.4103.3.1
D61387AEB60F52633DAAFC44FC23F808.taxon	etymology	Etymology. The specific epithet “ grandis ” refers to the larger size compared to other Japanese species of Trichoclinocera.	en	Saigusa, Toyohei, Sinclair, Bradley J. (2016): Revision of the Japanese species of Trichoclinocera Collin (Diptera: Empididae: Clinocerinae). Zootaxa 4103 (3): 201-229, DOI: 10.11646/zootaxa.4103.3.1
D61387AEB60F52633DAAFC44FC23F808.taxon	discussion	Remarks. This species is the most common Trichoclinocera in Japan.	en	Saigusa, Toyohei, Sinclair, Bradley J. (2016): Revision of the Japanese species of Trichoclinocera Collin (Diptera: Empididae: Clinocerinae). Zootaxa 4103 (3): 201-229, DOI: 10.11646/zootaxa.4103.3.1
D61387AEB60C52653DAAFF50FE11FE84.taxon	materials_examined	Type material. HOLOTYPE ♂, labelled: “ [MIE: Matsusaka] / Kushidagawa / Shôchô / 10 m alt., 34 ° 30 ' 9 " N / 136 ° 30 ' 45 " E / April 14, 2014 / T. SAIGUSA col. ”; [HOLOTYPE] / Trichoclinocera / miranda / Saigusa et / Sinclair, 2016 [red label] ” (KUMF). PARATYPES: JAPAN. HONSHU. Fukui: Namamori, Obama City [35 ° 28 ′ 55 ″ N 135 ° 44 ′ 52 ″ E], 2. v. 2010, Hisashi Ohishi (1 ♀, KUMF). Mie: same as holotype (50 ♂, 50 ♀, KUMF, CNC); same locality, 30. iv. 2013, Y. Shinogi (3 ♂, 4 ♀, KUMF, CNC); 2 ♂, 2 ♀, same locality, 1. v. 2013, Y. Shinogi; 1 ♂, 1 ♀, same locality, 3. v. 2013, Y. Shinogi (KUMF); Kushidagawa, Kayumi, Iinan-chô, Matsusaka-shi, 34 ° 27 ′ 54.9252 ″ N 136 ° 23 ′ 40.3218 ″ E, 7. v. 2013, Y. Shinogi (4 ♂, 7 ♀, CNC, KUMF). Wakayama: Arida Rv. (estuary), Arida [34 ° 4 ′ 58 ″ N 135 ° 06 ′ 27 ″ E], 5. iv. 2012, Hiroshi Ohishi (5 ♂, KUMF). Recognition. Distinguished from other Japanese species by the divergent radial fork, face with median brownish stripe, male fore femur with anteroventral spine-like setae widespread and of various lengths, and clasping cercus parallel-sided.	en	Saigusa, Toyohei, Sinclair, Bradley J. (2016): Revision of the Japanese species of Trichoclinocera Collin (Diptera: Empididae: Clinocerinae). Zootaxa 4103 (3): 201-229, DOI: 10.11646/zootaxa.4103.3.1
D61387AEB60C52653DAAFF50FE11FE84.taxon	description	Description. Wing length 2.9 – 3.5 mm. Male. Clothed in blue pruinescence except vertex of head, scutum, and dorsum of abdomen brown. Head rounded; compound eyes large, rounded. Face with blue pruinescence, except for slender median brownish stripe; face parallel-sided, slightly wider than width of antennal sockets; lower margin of face with shallow cleft and low carina; gena one-fourth as wide as height of eye. Ocellar bristles longest among cranial bristles; occiput with 2 pairs of bristles posterior to ocellar triangle; postocular with row of short bristles along margin of eye, upper half darker and more stout than lower bristles. Arista-like stylus long and thick. Palpus cylindrical, slender, length less than half height of eye. Pleura with pale blue pruinescence, lacking distinct pruinescence on scutum; posterolateral corner of scutum near postalar ridge concolorous with surrounding region; prescutellar depression brown; scutum not bivittate. Thoracic setae long, not longer than width of eye; acrostichal setulae lacking; 5 pairs dc; 1 pprn; 1 presut spal; 2 npl; 2 psut spal; 1 pal; 1 pair sctl, lacking setulae on disc; several setae on postpronotal lobe. Laterotergite with patch of pale setae; prothoracic episternum with long pale setae, dark setae on posterior margin of anepisternum; katepisternum with pale setae. Wing narrow, infuscate (Fig. 16 D); pterostigma very narrow, faint, parallel-sided. Basal costal seta slightly shorter than length of posterior dc. Row of setulae from base of R to tip of R 1; cell dm produced apically; R 4 and R 5 divergent, basal portion of R 4 gently curved, not angularly curved. Halter brown. Legs uniformly dark, coxae with blue pruinescence. Coxae with long, pale marginal setae. Fore femur (Figs 17 G, H) with 1 stout anterior bristle in on apical fourth; anteroventral margin with continuous row of stout setae, longest in middle, more slender and pale at base; posteroventral margin with stout seta at apical fourth, 3 – 4 stout setae at middle, longer than width of femur, base with row of long slender setae, similar in length to middle setae. Fore tibia with biserial row of short, stout setae beneath. Foreleg tarsomeres with pale ventral pubescence. Mid and hind femora with several pairs of preapical antero- and posteroventral setae; lacking anterodorsal and posterodorsal bristles. Hind tibia lacking pair of posterodorsal and anterodorsal basal setae; row of short bristles or preapical bristles lacking. Tarsus length shorter than corresponding tibia. Empodium pulvilliform, slightly shorter than claw, longer than half length of tarsomere. Hypandrium short, broad, quadrate, slightly larger than epandrium (Figs 4 D, 9 D). Phallus extended to apex of clasping cercus, slightly arched at mid-length; apex with small membranous expansion, not divided into lobes; ejaculatory apodeme nearly straight. Surstylus with membranous apex with fine setae. Clasping cercus finger-like, parallel-sided, with broad, rounded apex; bearing peg-like setae on inner anterodorsal margin. Female. Similar to male except as follows: setae of fore femur not as stout as in male; lacking long, stout posteroventral setae at middle (Figs 18 G, H). Similar to T. shinogii sp. nov., but differing as follows: posterior marginal setae of abdominal segment 7 weaker; those of sternum 7 weak and rather pale, not so distinctly grouped sublaterally as in T. shinogii sp. nov.; posterior marginal setae of tergum 8 sparser and those on lateral portions 2 – 3 in number and mixing several short setulae; sternum 8 as in shinogii; tergite 10 with 13 – 17 setae, of which 3 – 4 setae much stronger than others and distributed on apical half (not exclusively confined to apex), other setae rather weak but most of them weakly curved apically (Figs 13 D – F).	en	Saigusa, Toyohei, Sinclair, Bradley J. (2016): Revision of the Japanese species of Trichoclinocera Collin (Diptera: Empididae: Clinocerinae). Zootaxa 4103 (3): 201-229, DOI: 10.11646/zootaxa.4103.3.1
D61387AEB60C52653DAAFF50FE11FE84.taxon	distribution	Distribution. This species is known only from a few localities in central Honshu (Fig. 6).	en	Saigusa, Toyohei, Sinclair, Bradley J. (2016): Revision of the Japanese species of Trichoclinocera Collin (Diptera: Empididae: Clinocerinae). Zootaxa 4103 (3): 201-229, DOI: 10.11646/zootaxa.4103.3.1
D61387AEB60C52653DAAFF50FE11FE84.taxon	etymology	Etymology. The specific epithet is from the Latin “ mirandus ” (wonderful, strange) in reference to the surprising discovery of this species when the Trichoclinocera fauna of Japan was thought to be well understood. Remarks. See T. shinogii sp. nov. The habitat of this species is the same as that of T. shinogii sp. nov. The adults sit on moist sand close to water of slowly moving rivers, or on pebbles in shallow streamlets partially separated from the main river.	en	Saigusa, Toyohei, Sinclair, Bradley J. (2016): Revision of the Japanese species of Trichoclinocera Collin (Diptera: Empididae: Clinocerinae). Zootaxa 4103 (3): 201-229, DOI: 10.11646/zootaxa.4103.3.1
D61387AEB60B52673DAAFEB4FB7FF8A3.taxon	materials_examined	Type material. HOLOTYPE ♂, labelled: “ (YAMANASHI) / Hirogawara / Kitadake / 19. viii. 1978 / T. Saigusa ”; “ [HOLOTYPE] / Trichoclinocera / setigera / Saigusa et / Sinclair, 2016 [red label] ” (KUMF). PARATYPES: JAPAN. HONSHU. Akita: 1 ♂, Kuroyu, Hachimantai, 9. vii. 1962, TS (1 ♂, KUMF). Nagano: Ise-daki, 2100 m, Mt. Kiso-koma, 19. vii. 1963, TS (2 ♂, 1 ♀, KUMF); Shiroumdake, 2600 m, 15. vii. 1963, TS (1 ♂, KUMF); Spa Nakabusa, 22 – 24. viii. 1961, A. Kato (7 ♂, 10 ♀, KUMF). Yamagata: Asahidake, 5. viii. 1975, O. Yata (1 ♂, KUMF). Yamanahi: Kitadake, 1500 – 1800 m, 23. vii. 1981, TG (1 ♂, KUMF); Ôkanbazawa, Kitadake, 20. viii. 1978, TS (4 ♂, KUMF); same as holotype, TS, JE (6 ♂, 2 ♀, KUMF, CNC). Recognition. This species is distinguished from other Japanese species by lacking antero- and posterodorsal bristles on mid and hindlegs, possessing a largely dark scutum, and pair of apical phallic filaments.	en	Saigusa, Toyohei, Sinclair, Bradley J. (2016): Revision of the Japanese species of Trichoclinocera Collin (Diptera: Empididae: Clinocerinae). Zootaxa 4103 (3): 201-229, DOI: 10.11646/zootaxa.4103.3.1
D61387AEB60B52673DAAFEB4FB7FF8A3.taxon	description	Description. Wing length 5.2 – 5.6 mm. Male. Clothed in blue pruinescence except vertex and scutum brown. Head rounded; compound eyes large, rounded. Face with blue pruinescence, with broad, median brown patch on carina and narrow dorsal stripe; face narrowest just below antennae, greater than width of antennal sockets, lower margin of face with deep cleft and distinct carina; gena one-fifth as wide as height of eye. Ocellar bristles longest among cranial bristles; occiput with 2 pairs of bristles posterior to ocellar triangle; postocular with row of short bristles along margin of eye, upper third darker and stouter than lower bristles. Arista-like stylus long and slender, tapered apically. Palpus clavate, slender, length greater than half height of eye. Pleura, postpronotal lobe and notopleuron with bright blue pruinescence; posterolateral corner of scutum near postalar ridge, scutellum pale brown; prescutellar depression with narrow band of blue pruiescence along anterior margin of scutellum, rarely extending to posterior dc; scutum faintly bivittate. Thoracic setae long, somewhat shorter than width of eye; acrostichal setulae lacking; 6 pairs dc; 1 pprn; 1 presut spal; 3 npl; 2 psut spal; 1 pal; 1 pair sctl, lacking setulae on disc; several setulae sometimes present on postpronotal lobe and notopleural depression. Laterotergite with patch of pale setae; prothoracic episternum with long pale setae, setae lacking on katepisternum, and hind margins of metepisternum. Wing narrow, infuscate (Fig. 16 E); pterostigma very narrow, faint, parallel-sided. Basal costal seta ca. subequal to length of posterior dc. Row of setulae from base of R to tip of R 1; cell dm produced somewhat apically; R 4 and R 5 somewhat curved, parallel except extreme apex divergent. Halter brown with pale base. Legs greyish, generally darker towards tips, with coxae and trochanters clothed in blue pruinescence. Coxae with long, pale marginal setae. Fore femur (Figs 17 I, J) with 2 stout anterior setae in horizontal row on apical third; distal half with anteroventral row of short, stout setae, length less than half width of femur, setae more closely approximated near mid-length; proximal half with anteroventral row of pale setulae; 2 – 6 stout posteroventral setae on apical third; basal half with posteroventral row of ca. 8 widely-spaced, slender setae, some longer than width of femur. Fore tibia with biserial row of short, stout setae beneath. Foreleg tarsomeres with dense, whitish ventral pubescence, length ca. half width of tarsomeres. Mid and hind femora with several pairs of preapical antero- and posteroventral setae; lacking antero- and posterodorsal bristles. Base of hind tibia lacking pair of posterodorsal and anterodorsal setae; anteroventral margin lacking short bristles. Tarsus length longer than corresponding fore tibia, shorter than mid and hind tibiae. Empodium pulvilliform, slightly shorter than claw, longer than half length of tarsomere. Hypandrium short, oval, slightly longer than epandrium (Figs 8 A, 9 E). Phallus short, not extended beyond apex of clasping cercus, slightly bent at mid-length; apex with anterior, slender sickle-shaped extension enclosing membrane, posterior margin with pair of subapical, curved slender filaments, ca. one-third length of phallus; ejaculatory apodeme nearly straight at apical third. Width of surstylus nearly subequal to width of clasping cercus; anterior margin with pair of teeth-like projections; apically with pair of short, finger-like lobes. Clasping cercus finger-like, parallel-sided with truncate tip; slightly arched; bearing peg-like setae on inner dorsal margin, with small inner basal ridge. Female. Similar to male except as follows: anteroventral setae of fore femur absent or reduced; posteroventral basal setae less pronounced or absent. Fore tibiae lacking biserial row of erect setae beneath (Figs 18 I, J). Posterior marginal setae of tergum 7 moderately thick and long, evenly distributed, shorter middorsally; sternum 7 not emarginated posteriorly, bearing moderately thick posterior marginal setae evenly distributed but short and weaker midventrally, no distinct isolated setae posterolaterally; tergum 8 with posterior marginal setae long and weakly curved ventrally, short laterally and middorsally; sternum 8 not emarginated posteriorly, densely clothed with minute pile-like setulae, sparsely clothed with minute setulae on lateral surface; tergite 10 with 25 – 28 moderately long, thick and apically curved setae, those at apex of tergite not especially thickened, those along lateral margin and basal part somewhat finer; sternite 10 with short setae (Figs 14 A, C).	en	Saigusa, Toyohei, Sinclair, Bradley J. (2016): Revision of the Japanese species of Trichoclinocera Collin (Diptera: Empididae: Clinocerinae). Zootaxa 4103 (3): 201-229, DOI: 10.11646/zootaxa.4103.3.1
D61387AEB60B52673DAAFEB4FB7FF8A3.taxon	distribution	Distribution. This species is known only from the northern half of the island of Honshu (Fig. 10).	en	Saigusa, Toyohei, Sinclair, Bradley J. (2016): Revision of the Japanese species of Trichoclinocera Collin (Diptera: Empididae: Clinocerinae). Zootaxa 4103 (3): 201-229, DOI: 10.11646/zootaxa.4103.3.1
D61387AEB60B52673DAAFEB4FB7FF8A3.taxon	etymology	Etymology. The specific epithet “ setigera ” is from the Latin meaning bearing bristles, in reference to pair of long apical phallic filaments.	en	Saigusa, Toyohei, Sinclair, Bradley J. (2016): Revision of the Japanese species of Trichoclinocera Collin (Diptera: Empididae: Clinocerinae). Zootaxa 4103 (3): 201-229, DOI: 10.11646/zootaxa.4103.3.1
D61387AEB60B52673DAAFEB4FB7FF8A3.taxon	discussion	Remarks. This species appears to be confined to mountainous and subalpine regions.	en	Saigusa, Toyohei, Sinclair, Bradley J. (2016): Revision of the Japanese species of Trichoclinocera Collin (Diptera: Empididae: Clinocerinae). Zootaxa 4103 (3): 201-229, DOI: 10.11646/zootaxa.4103.3.1
D61387AEB60952793DAAF8ADFCCCFEF2.taxon	materials_examined	Type material. HOLOTYPE ♂, labelled: “ [MIE: Matsusaka] / Kushidagawa / Shôchô / 10 m alt., 34 ° 30 ' 9 " N / 136 ° 30 ' 45 " E / April 15, 2014 / T. SAIGUSA col. ”; [HOLOTYPE] / Trichoclinocera / shinogii / Saigusa et / Sinclair, 2016 [red label] ” (KUMF). PARATYPES: JAPAN. HONSHU. Aichi: Yahagi-Kago River, Ochiai, Toyota City, 35 ° 6 ′ 3 ″ N 137 ° 10 ′ 17 ″ E, 27. iv. 2008, H. Ohishi (16 ♂, 12 ♀, KUMF, CNC). Kyoto: Kizugawa, Tonosyo, Joyo, 20, 22, 30. v. 2010, H. Ohishi (1 ♂, 13 ♀, KUMF, CNC). Mie: same locality as holotype, 21, 23. iv. 2013, Y. Shinogi (2 ♀, KUMF); same locality as holotype, 1. v. 2013, Y. Shinogi (19 ♂, 14 ♀, KUMF, CNC); same locality as holotype, 3. v. 2013, Y. Shinogi (41 ♂, 28 ♀, KUMF, CNC); same locality as holotype, 14. iv. 2014, TS (16 ♂, 15 ♀, KUMF); same data as holotype (50 ♂, 50 ♀, KUMF, CNC); Kushidagawa, Iwachô, Matsusaka, 21. iv. 2013, Y. Shinogi (4 ♂, 5 ♀, KUMF); Kushidagawa, Saekinaka, Taki-chô, 34 ° 30 ′ 21 ″ N 136 ° 31 ′ 3 ″ E, 1. v. 2013, Y. Shinogi (3 ♂, 7 ♀, KUMF). Recognition. Distinguished from other Japanese species by the divergent radial fork, face entirely blue pruinescent, without median stripe, lateral margin of scutum and postpronotal lobe with broad band of blue pruinescence, male fore femur with anteroventral spine-like setae in tight row and clasping cercus expanded in middle.	en	Saigusa, Toyohei, Sinclair, Bradley J. (2016): Revision of the Japanese species of Trichoclinocera Collin (Diptera: Empididae: Clinocerinae). Zootaxa 4103 (3): 201-229, DOI: 10.11646/zootaxa.4103.3.1
D61387AEB60952793DAAF8ADFCCCFEF2.taxon	description	Description. Wing length 3.2 – 3.7 mm. Male. Clothed in blue pruinescence except vertex of head and scutum brown. Head rounded; compound eyes large, rounded. Face completely clothed in blue pruinescence; face parallelsided, slightly wider than width of antennal sockets; lower margin of face with shallow cleft and low carina; gena one-fourth as wide as height of eye. Ocellar bristles longest among cranial bristles; occiput with 2 pairs of bristles posterior to ocellar triangle; postocular with row of short bristles along margin of eye, upper half darker and more stout than lower bristles. Arista-like stylus long and thick. Palpus cylindrical, slender, length less than half height of eye. Pleura with pale blue pruinescence, with distinct band of lateral pruinescence on scutum; posterolateral corner of scutum near postalar ridge concolorous with surrounding region; prescutellar depression brown; scutum not bivittate. Thoracic setae long, not longer than width of eye; acrostichal setulae lacking; 5 – 7 pairs dc; 1 pprn, with several short setae; 1 presut spal; 2 npl; 2 psut spal; 1 pal; 1 pair sctl, lacking setulae on disc. Laterotergite with patch of pale setae; prothoracic episternum with long pale setae; anepisternum and katepisternum bare. Wing narrow, infuscate (Fig. 16 F); pterostigma indistinct. Basal costal seta slightly shorter than length of posterior dc. Row of setulae from base of R to tip of R 1; cell dm produced apically; R 4 and R 5 divergent, basal portion of R 4 gently curved, not angularly curved. Halter brown, with pale base. Legs dark, femora and coxae with blue pruinescence. Coxae with long pale, marginal setae. Fore femur (Figs 17 K, L) with 2 stout anterior bristle in on apical fourth; anteroventral margin with continuous row of stout setae, narrowly spaced in middle, without setae at base; posteroventral margin with 3 stout setae at apical third, 2 stout setae proximal to middle, longer than width of femur, base with row of long slender setae, decreasing in length basally. Fore tibia with biserial row of short, stout setae beneath. Foreleg tarsomeres with pale ventral pubescence. Mid and hind femora with several pairs of preapical antero- and posteroventral setae; anterodorsal bristles widely spaced; posterodorsal bristles lacking. Hind tibia lacking pair of posterodorsal and anterodorsal basal setae; preapical bristles present. Tarsus length longer than corresponding fore and mid tibiae, shorter than hind tibia. Empodium pulvilliform, slightly shorter than claw, longer than half length of tarsomere. Hypandrium short, broad, quadrate, slightly larger than epandrium (Figs 8 B, 9 F). Phallus extended slightly beyond apex of clasping cercus, slightly arched at mid-length; apex with small membranous expansion, not divided into lobes; ejaculatory apodeme nearly straight. Surstylus with membranous apex with fine setae. Clasping cercus expanded in middle, gradually tapered to narrow, rounded apex; bearing peg-like setae on inner anterodorsal margin. Female. Similar to male except as follows: lacking stout anteroventral and posteroventral setae of fore femur. Fore tibiae lacking biserial row of erect setae beneath (Figs 18 K, L). Posterior marginal setae of tergum 7 rather weak and sparsely distributed; sternum 7 not emarginated posteriorly, with sparse posterior marginal setae, of which several stronger setae almost confined to sublateral groups, 1 – 2 isolated stronger setae occasionally appearing midventrally; tergum 8 bearing straight posterior marginal setae, sparse, but longish dorsally, shorter and sparse laterally; sternum 8 with midventral group of minute pile-like setulae mixing some longish erect setulae, lateral setulae extremely minute; tergite 10 sparsely bearing 12 – 16 setae, of which 2 apical ones are very strong and apically curved, other setae shorter, weaker and almost straight or only very weakly curved apically; sternite 10 with short setae (Figs 14 D – F).	en	Saigusa, Toyohei, Sinclair, Bradley J. (2016): Revision of the Japanese species of Trichoclinocera Collin (Diptera: Empididae: Clinocerinae). Zootaxa 4103 (3): 201-229, DOI: 10.11646/zootaxa.4103.3.1
D61387AEB60952793DAAF8ADFCCCFEF2.taxon	distribution	Distribution. This species is known only from a few sites in Honshu (Fig. 10).	en	Saigusa, Toyohei, Sinclair, Bradley J. (2016): Revision of the Japanese species of Trichoclinocera Collin (Diptera: Empididae: Clinocerinae). Zootaxa 4103 (3): 201-229, DOI: 10.11646/zootaxa.4103.3.1
D61387AEB60952793DAAF8ADFCCCFEF2.taxon	etymology	Etymology. The specific epithet is named after the first collector of this species.	en	Saigusa, Toyohei, Sinclair, Bradley J. (2016): Revision of the Japanese species of Trichoclinocera Collin (Diptera: Empididae: Clinocerinae). Zootaxa 4103 (3): 201-229, DOI: 10.11646/zootaxa.4103.3.1
D61387AEB60952793DAAF8ADFCCCFEF2.taxon	discussion	Remarks. This species was collected together with T. miranda sp. nov. standing on the water surface or wet sand at the edge of the Kushida River (Figs 1 C, D), a behaviour similar to species of Hydrophorus Fallén (Dolichopodidae). This is very unusual behaviour for Trichoclincera, which almost always sit on the wet surface of large stones or boulders in rapids close to the water surface.	en	Saigusa, Toyohei, Sinclair, Bradley J. (2016): Revision of the Japanese species of Trichoclinocera Collin (Diptera: Empididae: Clinocerinae). Zootaxa 4103 (3): 201-229, DOI: 10.11646/zootaxa.4103.3.1
D61387AEB61752783DAAF991FEA8F808.taxon	materials_examined	Type material. HOLOTYPE ♂, labelled: “ (OITA) / Kyûsuikei / Kujûsan / 28. v. 1972 / T. Saigusa ”; [HOLOTYPE] / Trichoclinocera / stigmatica / Saigusa et / Sinclair, 2016 [red label] ” (KUMF). PARATYPES: JAPAN. KYUSHU. Fukuoka: Fukuoka-shi, Nogochi, 7. vi., 17. viii. 1997, 24. iv. 1998, BJS (4 ♂, 5 ♀, CNC); Magaribuchi (Nogochi), Sawara, 5. x. 1961, TS (11 ♂, 4 ♀, KUMF, CNC). Kumamoto: Hae, Itsuki village, 25. viii. 1997, BJS (1 ♂, 2 ♀, CNC); Hirasawado, Gokanoshô, 3. viii. 1975, TS (10 ♂, 2 ♀, KUMF); Nakamichi-bashi, 400 m, Izumi-mura, 24. v., 25. viii. 1997, BJS (26 ♂, 22 ♀, CNC); Izumi-mura, Momiki, 20. viii. 1999, BJS (2 ♂, 2 ♀, CNC); Shiratoriyama, Izumi-mura, 12 – 14. v. 1983, K. Ôhara (1 ♂, KUMF). Miyazaki: Seki-no-O, 28. iv. 1963, S. Takagi (2 ♂, HUS). Oita: Kyûsuikei, Kujûsan, 21. v. 1962, TS (42 ♂, 74 ♀, KUMF, BMNH, CNC); same locality, 19. v. 1963, TS (9 ♂, 19 ♀, KUMF); same data as holotype (108 ♂, 55 ♀, KUMF). Recognition. Distinguished from other Japanese species by its distinct pterostigma, presence of antero- and posterodorsal bristles on mid and hind femora, and face with bright blue pruinescence.	en	Saigusa, Toyohei, Sinclair, Bradley J. (2016): Revision of the Japanese species of Trichoclinocera Collin (Diptera: Empididae: Clinocerinae). Zootaxa 4103 (3): 201-229, DOI: 10.11646/zootaxa.4103.3.1
D61387AEB61752783DAAF991FEA8F808.taxon	description	Description. Wing length 4.8 – 5.4 mm. Male. Clothed in blue pruinescence except vertex of head, scutum, and dorsum of abdomen brown. Head rounded; compound eyes large, rounded. Face with blue pruinescence, with broad, median brown patch on carina and narrow dorsal stripe; face narrowest just below antennae, slightly wider than antennal sockets, lower margin of face with deep cleft and distinct carina; gena one-fifth as wide as height of eye. Ocellar bristles longest among cranial bristles; occiput with 2 pairs of bristles posterior to ocellar triangle; postocular with row of short bristles along margin of eye, upper third darker and more stout than lower bristles. Arista-like stylus long and slender, tapered apically. Palpus clavate, length greater than half height of eye. Pleura, postpronotal lobe, notopleuron, postalar, prescutellar depression, and scutellum with bright blue pruinescence; posterolateral corner of scutum near postalar ridge bluish; scutum faintly bivittate. Thoracic setae long, some ca. as long as width of eye; acrostichal setulae lacking; 6 – 8 pairs dc; 1 pprn; 1 presut spal; 3 npl; 2 psut spal; 1 pal; 1 pair sctl, lacking setulae on disc; several setulae on postpronotal lobe and notopleural depression. Laterotergite with patch of pale setae; prothoracic episternum with long pale setae; katepisternum with pale setulae near middle, metepisternum with pale setulae at posterodorsal margin. Wing narrow, infuscate (Fig. 16 G); pterostigma narrow, distinct, elliptical. Basal costal seta ca. subequal to posterior dc. Row of setulae from base of R to tip of R 1; cell dm produced apically; R 4 and R 5 curved, parallel except extreme apex divergent. Halter brown with pale base. Legs greyish, generally darker towards tips, with coxae and trochanters clothed in blue pruinescence. Coxae with long pale, marginal setae. Fore femur (Figs 17 M, N) with 2 stout, oblique anterior setae on apical third; widely spaced row of 3 – 5 anterodorsal setae on apical two-thirds, less than half width of femur; row of 6 – 8 stout anteroventral setae on apical half, short and long setae intermixed, some longer than half width of femur; basal half with row of ca. 9 – 12 very short, stout setae and 4 – 6 stout, sub-basal setae, some longer than half width of femur; posteroventral margin with 4 – 5 stout, widely spaced setae on apical half, less stout apically; basal half with row 9 – 13 short, slender setae and 2 – 3 stout, sub-basal setae some longer than basal width of femur. Fore tibia with biserial row of short, stout setae beneath. Foreleg tarsomeres with white, dense ventral pubescence, length ca. half width of tarsomeres. Mid and hind femora with widely spaced pairs of antero- and posteroventral; widely spaced anterodorsal bristles on apical three-quarters; widely spaced posterodorsal bristles on apical half. Base of hind tibia with pair of divergent dorsal setae (1 posterodorsal and 1 anterodorsal); anteroventral margin lacking bristles; 1 dorsal and 1 ventral preapical bristles present. Tarsus length longer than corresponding fore tibia, shorter than mid and hind tibiae. Empodium pulvilliform, slightly shorter than claw, longer than half length of tarsomere. Hypandrium quadrate, broad, shorter than epandrium (Figs 8 C, 9 G). Phallus not extended beyond apex of clasping cercus, strongly arched at mid-length; apex with anterior, slender, sickle-shaped extension enclosing membrane, posterior similar to ventral process; ejaculatory apodeme nearly straight at anterior third. Surstylus narrow, short, inconspicuous with membranous posteroapical margin; anterior margin with several short teeth-like projections. Clasping cercus broadest in middle, expanded also subapically, strongly tapered to rounded apex; bearing peg-like setae on inner dorsal face. Female. Similar to male except as follows: basal anteroventral bristles of fore femur absent; posteroventral basal setae less pronounced. Fore tibiae lacking biserial row of erect setae beneath (Figs 18 M, N). Posterior marginal setae of tergum 7 fairly strong, slightly denser than in takagii; sternum 7 weakly emarginated posteriorly, with midventral length 3 / 4 as long as lateral length, bearing fairly strong hind marginal setae evenly distributed, posterolateral extension of sternum 7 with rather strong isolated seta directed posteroventrally; tergum 8 with posterior marginal setae fairly long and dense dorsally, moderately long laterally; sternum 8 weakly emarginated posteriorly, densely clothed with minute pile-like setulae, sparsely clothed with minute setulae on lateral surface; tergite 10 bearing some 50 apically curved setae evenly distributed except basal marginal area, 2 apical setae only slightly stronger than others on disc; sternite 10 with short setae mixed with 2 – 3 longer setae 1.5 times as long as sternum (Figs 15 A – C).	en	Saigusa, Toyohei, Sinclair, Bradley J. (2016): Revision of the Japanese species of Trichoclinocera Collin (Diptera: Empididae: Clinocerinae). Zootaxa 4103 (3): 201-229, DOI: 10.11646/zootaxa.4103.3.1
D61387AEB61752783DAAF991FEA8F808.taxon	distribution	Distribution. This species is restricted to the southern island of Kyushu (Fig. 10). Adults are collected from May to October.	en	Saigusa, Toyohei, Sinclair, Bradley J. (2016): Revision of the Japanese species of Trichoclinocera Collin (Diptera: Empididae: Clinocerinae). Zootaxa 4103 (3): 201-229, DOI: 10.11646/zootaxa.4103.3.1
D61387AEB61752783DAAF991FEA8F808.taxon	etymology	Etymology. In reference to the distinct pterostigma, which is weakly differentiated in other Japanese species of Trichoclinocera.	en	Saigusa, Toyohei, Sinclair, Bradley J. (2016): Revision of the Japanese species of Trichoclinocera Collin (Diptera: Empididae: Clinocerinae). Zootaxa 4103 (3): 201-229, DOI: 10.11646/zootaxa.4103.3.1
D61387AEB615527A3DAAFF50FEEAFD42.taxon	materials_examined	Type material. HOLOTYPE ♂, labelled: “ (TOKUSHIMA) / Ichiu / 3. vi. 1981 / T. Gotô ”; “ [HOLOTYPE] / Trichoclinocera / takagii / Saigusa et / Sinclair, 2016 [red label] ” (KUMF). PARATYPES: JAPAN. HOKKAIDO: 2 ♀, Akan, Kushiro, 3. vii. 1958, M. Takahashi (KUMF); Sapporo, 2. vii. 1960, Takagi (6 ♂, 12 ♀, HSU, KUMF, CNC); Sapporo, Maru-Y., 28. vi. 1961, S. Takagi (2 ♂, 3 ♀, HUS); Simamatu, 12. vi. 1964, I. Miyagi (1 ♂, 4 ♀, HUS); Titose, 14. viii. 1962, S. Takagi (4 ♂, HUS, KUMF, CNC). HONSHU. Kyoto: 1 ♂, Kibune [35 ° 7 ′ 15 ″ N 135 ° 45 ′ 47 ″ E], 8. vi. 1963, TS (1 ♂, KUMF). Mie: Kushidagawa, Shô-chô, Matsusaka [34 ° 30 ′ 9 ″ N 136 ° 30 ′ 45 ″ E], 26. iv. 2013, Y. Shinogi (1 ♂, 1 ♀, KUMF); Kushidagawa, Hirose-machi, Matsusaka, 30. iv. 2013, Y. Shinogi (1 ♂, 1 ♀, KUMF). Osaka: Myôkensan, Nose, 29. iv. 1962, TS (1 ♂, KUMF). Wakayama: Kôyasan, Kôya-machi, 6. vi. 1981, TG (15 ♂, 29 ♀, KUMF). Ya m a gu ch i: Chyomonkyo [34 ° 21 ′ 41 ″ N 131 ° 33 ′ 1 ″ E], 4. viii. 1963. I. Miyagi (4 ♂, 1 ♀, HUS). KYUSHU. Fukuoka: Magaribuchi, Sagara, 2. v. 1962, TS (12 ♂, 17 ♀, KUMF, BMNH, CNC); same locality, 22. iv. 1962, TS (8 ♀, KUMF). Miyazaki: Seki-no-O, 28. iv. 1963, S. Takagi (8 ♂, HUS, KUMF, CNC). Oita: Kyûsuikyô, Kujûsan, 21. v. 1962, TS (2 ♂, KUMF) same locality, 19. v. 1962, TS (1 ♀, KUMF). SHIKOKU. Ehime: Omogo, Iyo, 14. vii. 1952, S. Miyamoto (2 ♂, 3 ♀, KUMF); Omogokei, 27 – 28. v. 1967, A. Nakanishi (6 ♂, 11 ♀, KUMF). Tokushima: same data as holotype (28 ♂, 57 ♀, KUMF, CNC). Recognition. Distinguished from other Japanese species by lack of a distinct pterostigma, presence of antero- and posterodorsal bristles on mid and hind femora, and pale brownish face.	en	Saigusa, Toyohei, Sinclair, Bradley J. (2016): Revision of the Japanese species of Trichoclinocera Collin (Diptera: Empididae: Clinocerinae). Zootaxa 4103 (3): 201-229, DOI: 10.11646/zootaxa.4103.3.1
D61387AEB615527A3DAAFF50FEEAFD42.taxon	description	Description. Wing length 4.2 – 4.7 mm. Male. Clothed in blue pruinescence except vertex of head, scutum, and dorsum of abdomen brown. Head rounded; compound eyes large, rounded. Face with blue pruinescence, with median brown patch on carina and narrow dorsal stripe; face narrowest just below antennae, slightly wider than antennal sockets, lower margin of face with deep cleft and distinct carina; gena one-fifth as wide as height of eye. Ocellar bristles longest among cranial bristles; occiput with 2 pairs of bristles posterior to ocellar triangle; postocular with row of short bristles along margin of eye, upper half darker and more stout than lower bristles. Arista-like stylus long and slender, tapered apically. Palpus clavate, length greater than half height of eye. Pleura, postpronotal lobe, notopleuron, postalar, prescutellar depression, and scutellum with bright blue pruinescence; posterolateral corner of scutum near postalar ridge bluish; scutum faintly bivittate. Thoracic setae long, some ca. as long as width of eye; acrostichal setulae lacking; 6 – 7 pairs dc; 1 pprn; 1 presut spal; 3 npl; 2 psut spal; 1 pal; 1 pair sctl, lacking setulae on disc; several setulae on postpronotal lobe and notopleural depression. Laterotergite with patch of pale setae; prothoracic episternum with long pale setae; katepisternum with pale setulae near middle, metepisternum with pale setulae at posterodorsal margin. Wing narrow, infuscate (Fig. 16 H); pterostigma lacking. Basal costal seta longer than posterior dc. Row of setulae from base of R to tip of R 1; cell dm produced apically; R 4 and R 5 slightly curved, somewhat divergent throughout. Halter brown with pale base. Legs greyish, generally darker towards tips, with coxae and trochanters clothed in blue pruinescence. Coxae with long, pale marginal setae. Fore femur (Figs 17 O, P) with 2 stout, oblique anterior setae on apical third, separated by 1 – 2 x length of setae; widely spaced row of 3 – 5 anterodorsal setae on apical two-thirds, less than half width of femur; row of 3 stout, widely spaced anteroventral setae on apical half, with short setae intermixed and apical row of ca. 6 similar sized setae; basal half with row of ca. 12 – 15 short, stout setae increasing in length basally; posteroventral margin with 3 stout, widely spaced setae on apical half, length ca. subequal to width of femur; basal fourth with row of 1 dark and several long, pale setae, longer than width of femur. Fore tibia with biserial row of short, stout setae beneath. Foreleg tarsomeres with white, dense ventral pubescence, length ca. half width of tarsomeres. Mid and hind femora with widely spaced pairs of antero- and posteroventral; widely spaced anterodorsal bristles on apical three-quarter; widely spaced posterodorsal bristles on apical half. Base of hind tibia with pair of divergent dorsal setae (1 posterodorsal and 1 anterodorsal); anteroventral margin lacking bristles; 1 dorsal and 1 ventral preapical bristles present. Tarsus length longer than corresponding fore tibia, shorter than mid and hind tibiae. Empodium pulvilliform, slightly shorter than claw, longer than half length of tarsomere. Hypandrium quadrate, broad, shorter than epandrium (Figs 8 D, 9 H). Phallus extended slightly beyond apex of clasping cercus, gently arched at mid-length; apex with anterior, slender sickle-shaped extension enclosing membrane, ejaculatory apodeme nearly straight at anterior third. Surstylus narrow, short, inconspicuous with membranous posteroapical margin; anterior margin with short 1 – 2 teeth-like projections. Clasping cercus thumblike, broadest subapically, tapered to rounded apex; bearing peg-like setae on inner dorsal and apical face; long setae on anterior margin. Female. Similar to male except as follows: basal anteroventral bristles of fore femur absent; posteroventral basal of setae less pronounced. Fore tibiae lacking biserial row of erect setae beneath (Figs 18 O, P). Posterior marginal setae of tergum 7 fairly strong and evenly distributed, shorter dorsally; sternum 7 deeply emarginated posteriorly, with midventral length almost 1 / 2 as long as lateral length, bearing strong hind marginal setae mostly represented by three groups of closely set setae, one midventral and 2 sublateral, most other posterior marginal setae very short and pale, except 1 stronger seta occasionally appearing between midventral and lateral groups of setae; posterolateral part of sternum 7 with 1 – 2 moderately long isolated setae; tergum 8 with posterior marginal setae long and rather sparse dorsally, short laterally mixed with several moderately long setae; sternum 8 deeply emarginated posteriorly, densely clothed with minute pile-like setulae, sparsely clothed with minute setulae on lateral surface; tergite 10 with 2 strong apically curved setae and some 25 weaker curved setae on disc, basal ones only weakly curved apically; sternite 10 with short setae (Figs 15 D – F).	en	Saigusa, Toyohei, Sinclair, Bradley J. (2016): Revision of the Japanese species of Trichoclinocera Collin (Diptera: Empididae: Clinocerinae). Zootaxa 4103 (3): 201-229, DOI: 10.11646/zootaxa.4103.3.1
D61387AEB615527A3DAAFF50FEEAFD42.taxon	distribution	Distribution. This is a wide ranging species, known from the Japanese islands of Hokkaido, Honshu, Shikoku, and Kyushu (Fig. 11). Adults are collected in from May to July.	en	Saigusa, Toyohei, Sinclair, Bradley J. (2016): Revision of the Japanese species of Trichoclinocera Collin (Diptera: Empididae: Clinocerinae). Zootaxa 4103 (3): 201-229, DOI: 10.11646/zootaxa.4103.3.1
D61387AEB615527A3DAAFF50FEEAFD42.taxon	etymology	Etymology. Named in honour of the first collector of this species, Prof. Emeritus Sadao Takagi of Hokkaido University.	en	Saigusa, Toyohei, Sinclair, Bradley J. (2016): Revision of the Japanese species of Trichoclinocera Collin (Diptera: Empididae: Clinocerinae). Zootaxa 4103 (3): 201-229, DOI: 10.11646/zootaxa.4103.3.1
