taxonID	type	description	language	source
D06A385E5729BB4B02BCFB6C5EA76F85.taxon	type_taxon	Type species. Pseudharpinia dentata Schellenberg, 1931	en	Andrade, Luiz F., Senna, André R. (2020): Four new species of Pseudharpinia Schellenberg, 1931 (Crustacea: Amphipoda Phoxocephalidae) from southwestern Atlantic and new records of P. tupinamba Senna & Souza-Filho, 2011. Zootaxa 4763 (4): 501-537, DOI: 10.11646/zootaxa.4763.4.3
D06A385E5729BB4B02BCFB6C5EA76F85.taxon	diagnosis	Amended diagnosis. Head with dorsal keel or ventrolateral lines present in some species. Rostrum unconstricted. Eyes absent. Antennae 1 – 2 flagella unreduced in females. Antenna 1 peduncle article 2 short, with ventral setae widely spread. Antenna 2 peduncle article 1 ensiform, article 4 with one main row of facial stout setae, article 5 slender and short. Mandible molar pillow-shaped as a small hump, not triturative, bearing splayed stout setae; palpar hump small. Maxilla 1 palp 2 - articulate. Maxilla 2 weakly setose. Maxilliped palp article 4 not elongated, with long and distinct apical nail. Gnathopods 1 – 2 small, similar, carpus short and free, with weak eusirid attachment; propodus ovatorectangular, weakly elongated and weakly setose anteriorly; palms acute. Gnathopod 2 slightly larger than gnathopod 1. Pereopods 3 – 4 carpus moderately to strongly setose. Pereopod 5 basis narrow. Pereopods 5 – 6 merus and carpus narrow. Pereopod 7 basis often setose anterodistally, posteroventral lobe often serrate; ischium enlarged. Epimeral plates 1 – 2 without long posterior setae. Urosomite 3 smooth. Uropod 1 peduncle elongated, without ventroapical spike or lateroapical enlarged seta. Uropod 3 outer ramus article 2 short, with 1 or 2 long setae apically. Telson with one to four apical or subapical setae on each lobe, often with dorsolateral setae.	en	Andrade, Luiz F., Senna, André R. (2020): Four new species of Pseudharpinia Schellenberg, 1931 (Crustacea: Amphipoda Phoxocephalidae) from southwestern Atlantic and new records of P. tupinamba Senna & Souza-Filho, 2011. Zootaxa 4763 (4): 501-537, DOI: 10.11646/zootaxa.4763.4.3
D06A385E5729BB4B02BCFB6C5EA76F85.taxon	discussion	Composition of the genus. Pseudharpinia abyssalis (Pirlot, 1932); P. antarctica Ren in Ren & Huang, 1991; P. ayutlanta (J. L. Barnard, 1964); P. berardo Senna, 2010; P. birjulini (Gurjanova, 1953); P. bonhami sp. nov.; P. brevirostris Chevreux, 1920; P. calcariaria Bushueva, 1982; P. cariniceps (K. H. Barnard, 1932); P. cinca (J. L. Barnard, 1961); P. dentata (Schellenberg, 1931); P. excavata (Chevreux, 1887); P. inexpectata Jarret & Bousfield, 1994; P. jonesyi sp. nov.; P. latipes (Norman, 1900); P. macrospinosa Valério-Berardo & Piera, 2006; P. obtusifrons (Stebbing, 1888); P. ovata Senna, 2010; P. pagei sp. nov.; P. planti sp. nov.; P. tupinamba Senna & Souza-Filho, 2011; P. vallini (Dahl, 1954);	en	Andrade, Luiz F., Senna, André R. (2020): Four new species of Pseudharpinia Schellenberg, 1931 (Crustacea: Amphipoda Phoxocephalidae) from southwestern Atlantic and new records of P. tupinamba Senna & Souza-Filho, 2011. Zootaxa 4763 (4): 501-537, DOI: 10.11646/zootaxa.4763.4.3
D06A385E572ABB4E02BCFEB05D0D6ED9.taxon	description	(Figs 1 – 5)	en	Andrade, Luiz F., Senna, André R. (2020): Four new species of Pseudharpinia Schellenberg, 1931 (Crustacea: Amphipoda Phoxocephalidae) from southwestern Atlantic and new records of P. tupinamba Senna & Souza-Filho, 2011. Zootaxa 4763 (4): 501-537, DOI: 10.11646/zootaxa.4763.4.3
D06A385E572ABB4E02BCFEB05D0D6ED9.taxon	materials_examined	Type material. Holotype: female, 2.8 mm length, dissected and drawn, MBT 188, 24 ° 58 ’ S, 45 ° 26 ’ W, 98 m depth, São Paulo state, 30 May 1971, UERJ 441. Paratypes: 2 females, same data as for the holotype, UERJ 442; 1 female, MBT 219, 100 m depth, Rio de Janeiro state, 6 June 1971, UERJ 443.	en	Andrade, Luiz F., Senna, André R. (2020): Four new species of Pseudharpinia Schellenberg, 1931 (Crustacea: Amphipoda Phoxocephalidae) from southwestern Atlantic and new records of P. tupinamba Senna & Souza-Filho, 2011. Zootaxa 4763 (4): 501-537, DOI: 10.11646/zootaxa.4763.4.3
D06A385E572ABB4E02BCFEB05D0D6ED9.taxon	diagnosis	Diagnosis. Head with dorsal keel. Rostrum entire. Antenna 2 peduncle article 1 with a subacute ensiform process. Mandible molar with 3 multicuspidate stout setae, right lacinia mobilis apically multicuspidate. Maxilla 1 inner plate setose apically. Maxilliped inner plate with 1 stout seta apically; palp article 3 with long setae apically, article 4 bearing elongated apical nail. Gnathopods 1 – 2 ischium anterior margin with row of setules. Pereopod 6 basis with a weak acute projection proximally. Pereopod 7 basis reaching half of merus, strongly dentate, with 6 acute stout spines posteroventrally; merus posterior margin produced distally. Epimeral plate 1 anterior and ventral margins covered with setules. Epimeral plate 3 posteroventral corner produced into a large hook. Uropods 1 – 2, rami continuously setose to apex. Telson short, apex rounded, with 3 setae apically on each lobe.	en	Andrade, Luiz F., Senna, André R. (2020): Four new species of Pseudharpinia Schellenberg, 1931 (Crustacea: Amphipoda Phoxocephalidae) from southwestern Atlantic and new records of P. tupinamba Senna & Souza-Filho, 2011. Zootaxa 4763 (4): 501-537, DOI: 10.11646/zootaxa.4763.4.3
D06A385E572ABB4E02BCFEB05D0D6ED9.taxon	description	Description. Based on female holotype, UERJ 441. Head. Antenna 1 peduncle article 1 about 1.4 × longer than wide, ventral margin distal lobe with brush setae, dorsal margin with 1 pappose seta distally; article 2 ventral margin with 6 long pappose setae; article 3 short, ventral margin with 1 short and 2 long setae, facial margin with 3 brush setae distally; primary flagellum 7 - articulate; accessory flagellum 5 - articulate. Antenna 2 peduncle article 2 short; article 3 ventral margin with 1 short and 4 long setae, dorsal margin with 1 seta distally; article 4 about 1.3 × longer than wide, ventral margin with long pappose setae and 6 brush setae, facial margin with 1 brush seta medially, 5 setae distally and a row of 10 setae extending to dorsal apex; article 5 about 0.6 × the length of article 4, ventral margin with long pappose setae, dorsal margin with 1 brush seta distally; flagellum 8 - articulate. Upper lip and lower lip ordinary. Mandible incisor with 6 teeth, lacinia mobilis with 4 (left) – 6 (right) teeth, accessory setal row with 7 (left) – 6 (right) multicuspidate stout setae, palpar hump small; palp 3 - articulate, article 1 short, article 2 with 6 (left) – 9 (right) setae medially, article 3 slightly shorter than article 2, apex oblique, with 7 (left) – 6 (right) medium to long setae. Maxilla 1 inner plate setose apically; outer plate with 9 stout setae apically, being 2 smooth; palp article 2 with 1 long and 2 stout pectinate setae apically, lateral and medial margins covered with setules. Maxilla 2 inner plate with 6 plumose setae extending from distomedial margin to apex, lateral and medial margins covered with setules; outer plate slightly longer than inner, setose apically, lateral margin covered with setules. Maxilliped inner plate with 6 plumose setae, outer plate with 9 papposerrate stout setae increasing in length to apex, lateral margin with 5 long plumose setae; palp article 2 medial margin moderately setose, article 3 moderately setose, article 4 about 3.4 × longer than wide, with 2 accessory setules. Pereon. Gnathopod 1 coxa expanded anteroventrally, ventral margin with 19 long pappose setae; basis about 2.1 × longer than wide, anterior margin with 6 setae, posterior margin with 1 long plumose seta, posterodistal corner with 3 pappose setae; ischium short and subtriangular, posterior margin with 3 pappose setae; merus posterior margin with short to long simple and pappose setae; carpus anterior margin with 2 pappose setae distally, posterior margin with 3 setae, facial margin with a row of 5 setae; propodus weakly setose, anterodistal margin with a tuft of setae, posterior margin with 1 stout seta defining palm; palm covered with setules on its margin, palmar hump large; dactylus reaching the palmar corner, outer margin with 1 short seta distally. Gnathopod 2 coxa subrectangular, ventral margin with 9 long pappose setae; basis about 2.8 × longer than wide, anterior margin with short to medium setae, posterior margin with 1 long plumose seta, posterodistal corner with 3 long pappose setae; ischium short and subrectangular, posterodistal corner with 6 setae, being 3 pappose; merus posterior margin with long pappose setae, facial margin with 1 seta; carpus anterior margin with 1 seta distally, posterior margin with 5 pappose setae; propodus weakly setose, anterodistal margin with a tuft of setae, posterior margin with 1 stout seta defining palm; palm covered with setules on its margin, palmar hump large; dactylus slightly longer than palm, outer margin with 1 short seta distally. Pereopod 3 coxa subrectangular, ventral margin with 8 long pappose setae; basis about 2.8 × longer than wide, anterior margin with 4 pappose setae, posterior margin with 2 long plumose setae, posterodistal corner with 5 long pappose setae; ischium short and subrectangular, posterodistal corner with a row of 6 long pappose setae; merus anterior margin produced distally, with 1 simple and 3 pappose setae, posterior margin moderately setose with pappose setae, facial margin with 6 setae distally; carpus anterior margin with 1 short seta distally, posterior margin with long pappose setae and stout setae progressively more robust and longer, facial margin with a row of 7 setae; propodus anterior margin with 2 setae distally, posterior margin with 6 long setae; dactylus stout, about 70 % the length of propodus. Pereopod 4 coxa posterior margin strongly expanded, posterodorsal margin excavate, ventral margin with 11 pappose setae; basis about 3 × longer than wide, posterior margin with 2 long plumose setae, posterodistal corner with 1 pappose seta; ischium short and subrectangular, posterodistal corner with 3 pappose setae; merus anterior margin produced distally, with 4 pappose setae, posterior margin moderately setose with pappose setae; carpus anterior margin with 1 short seta distally, posterior margin with long pappose setae and stout setae progressively more robust and longer, facial margin with a row of 6 setae; propodus anterior margin with 2 setae distally, posterior margin with 5 long setae; dactylus stout, about 65 % the length of propodus. Pereopod 5 coxa bilobate, posteroventral lobe produced, with 6 pappose setae; basis about 2.8 × longer than wide, anterior margin with long pappose setae, posterior margin with 1 plumose seta distally; ischium short, anterodistal corner with 4 pappose setae; merus anterior margin moderately setose, with posterior margin with 5 plumose setae; carpus anterior and posterior margins moderately and weakly setose respectively; propodus anterior and posterior margins moderately setose; dactylus about 60 % the length of propodus. Pereopod 6 damaged, describable to merus only; coxa posterior margin with 1 short seta; basis about 1.5 × longer than wide, anterior margin with 3 stout and 14 long pappose setae, anterodistal corner with 3 pappose setae, posterior margin with pappose setae and a row of setules; ischium short and subrectangular, anterior margin with 4 pappose setae; merus anterior margin with 7 stout setae, posterior margin with 6 pappose setae proximally, 1 seta medially and 4 setae distally. Pereopod 7 coxa subtriangular, posterior margin with 1 short seta; basis about 1.1 × longer than wide, posteroventral margin expanded and strongly dentate, with 6 large spines posteroventrally, reaching half of merus, anterodistal corner with 1 stout seta and 2 plumose setae; ischium distofacial lobe produced, anterior margin with 1 stout and 4 long plumose setae; merus anterior margin with 1 short and 2 stout setae, anterodistal corner with 4 stout setae; carpus anterior and posterior margin weakly setose; propodus anterior margin with 3 setae, posterior margin with 1 seta distally; dactylus about 85 % the length of propodus, inner margin with a row of setules distally. Pleon. Epimeral plate 1 margins rounded, anterodistal angle with 1 plumose seta. Epimeral plate 2 anterior margin concave, ventral margin with 6 plumose setae, posterior margin convex, with 1 short seta. Epimeral plate 3 anteroventral corner expanded, ventral margin weakly concave, with 8 plumose setae, posteroventral corner strongly produced dorsally into a hook, posterior margin covered with setules. Urosomite 1 ventral margin with a tuft of setae. Uropod 1 peduncle about 3.1 × longer than wide, dorsomedial margin with 8 setae, dorsolateral margin with 3 setae; outer ramus 1.1 × longer than inner, dorsomedial margin with 5 setae, dorsolateral margin with 6 setae; inner ramus with 7 setae continuously to apex. Uropod 2 peduncle about 2.4 × longer than wide, dorsomedial margin with 4 setae, dorsolateral margin with 6 setae, being the distalmost stout; outer ramus slightly longer than inner, dorsomedial and dorsolateral margins with 8 setae each; inner ramus with 8 setae continuously to apex. Uropod 3 peduncle about 1.2 × longer than wide, dorsoapical margin with 7 setae; outer ramus slightly longer than inner, article 1 dorsomedial margin with 7 setae, ventrolateral margin with 3 setae, article 2 twice longer than wide, with 2 long setae apically. Telson about 85 % cleft, lobes rounded with 3 setae each, dorsolateral margin with 3 plumose setae on each side.	en	Andrade, Luiz F., Senna, André R. (2020): Four new species of Pseudharpinia Schellenberg, 1931 (Crustacea: Amphipoda Phoxocephalidae) from southwestern Atlantic and new records of P. tupinamba Senna & Souza-Filho, 2011. Zootaxa 4763 (4): 501-537, DOI: 10.11646/zootaxa.4763.4.3
D06A385E572ABB4E02BCFEB05D0D6ED9.taxon	etymology	Etymology. Named after John Bonham (1948 – 1980), drummer of the band Led Zeppelin, who inspired many generations of musicians with his outstanding talent and unique way of playing.	en	Andrade, Luiz F., Senna, André R. (2020): Four new species of Pseudharpinia Schellenberg, 1931 (Crustacea: Amphipoda Phoxocephalidae) from southwestern Atlantic and new records of P. tupinamba Senna & Souza-Filho, 2011. Zootaxa 4763 (4): 501-537, DOI: 10.11646/zootaxa.4763.4.3
D06A385E572ABB4E02BCFEB05D0D6ED9.taxon	distribution	Type locality. São Paulo state coast (24 ° 58 ’ S, 45 ° 26 ’ W).	en	Andrade, Luiz F., Senna, André R. (2020): Four new species of Pseudharpinia Schellenberg, 1931 (Crustacea: Amphipoda Phoxocephalidae) from southwestern Atlantic and new records of P. tupinamba Senna & Souza-Filho, 2011. Zootaxa 4763 (4): 501-537, DOI: 10.11646/zootaxa.4763.4.3
D06A385E572ABB4E02BCFEB05D0D6ED9.taxon	discussion	Remarks. Due to a notable character as the strong projections on the posteroventral margin of pereopod 7 basis, Pseudharpinia bonhami sp. nov. can be considered very similar to P. birjulini and P. latipes. However, the new species can be distinguished from the former by the following characters (characters of P. birjulini in parentheses): head with entire rostrum (excavate laterally); pereopod 7 basis with 6 large spines posteroventrally (4 large spines confined posteriorly), ventral margin not serrated (serrated); pleonites naked dorsally (setose dorsally); epimeral plates 1 – 2 posteroventral corner rounded (acute); epimeral plate 3 posteroventral corner strongly produced into a hook (weakly produced); telson lobes with 3 setae each (2 setae). Pseudharpinia bonhami sp. nov. differs from the latter species by the following characters (characters of P. latipes in parentheses): gnathopod 1 palm shorter than the posterior margin of propodus (longer); pereopod 7 basis with 6 large spines posteroventrally (7 spines confined mostly ventrally), reaching half of merus (reaching apex of merus), anterodistal corner weakly setose (strongly setose).	en	Andrade, Luiz F., Senna, André R. (2020): Four new species of Pseudharpinia Schellenberg, 1931 (Crustacea: Amphipoda Phoxocephalidae) from southwestern Atlantic and new records of P. tupinamba Senna & Souza-Filho, 2011. Zootaxa 4763 (4): 501-537, DOI: 10.11646/zootaxa.4763.4.3
D06A385E572ABB4E02BCFEB05D0D6ED9.taxon	distribution	Geographic distribution. The species is known from the coast of the Brazilian states of Rio de Janeiro and São Paulo (Fig 24).	en	Andrade, Luiz F., Senna, André R. (2020): Four new species of Pseudharpinia Schellenberg, 1931 (Crustacea: Amphipoda Phoxocephalidae) from southwestern Atlantic and new records of P. tupinamba Senna & Souza-Filho, 2011. Zootaxa 4763 (4): 501-537, DOI: 10.11646/zootaxa.4763.4.3
D06A385E572ABB4E02BCFEB05D0D6ED9.taxon	discussion	Bathymetric range. 98 – 100 m depth.	en	Andrade, Luiz F., Senna, André R. (2020): Four new species of Pseudharpinia Schellenberg, 1931 (Crustacea: Amphipoda Phoxocephalidae) from southwestern Atlantic and new records of P. tupinamba Senna & Souza-Filho, 2011. Zootaxa 4763 (4): 501-537, DOI: 10.11646/zootaxa.4763.4.3
D06A385E5720BB5902BCFF005D0D6CBC.taxon	description	(Figs 6 – 12)	en	Andrade, Luiz F., Senna, André R. (2020): Four new species of Pseudharpinia Schellenberg, 1931 (Crustacea: Amphipoda Phoxocephalidae) from southwestern Atlantic and new records of P. tupinamba Senna & Souza-Filho, 2011. Zootaxa 4763 (4): 501-537, DOI: 10.11646/zootaxa.4763.4.3
D06A385E5720BB5902BCFF005D0D6CBC.taxon	materials_examined	Type material. Holotype: female, 5.5 mm length, dissected and drawn, MBT 104, 26 ° 34 ’ S, 47 ° 59 ’ W, 58 m depth, Santa Catarina state, 20 June 1970, UERJ 444. Paratype: male, dissected and drawn, same data as for the holotype, UERJ 445. Additional material examined. 61 females, same data as for the holotype, UERJ 446; 20 females and 1 male, MBT 106, 26 ° 45 ’ S, 47 ° 16 ’ W, 130 m depth, Santa Catarina state, 20 June 1970, UERJ 447; 11 females, MBT 81, 24 ° 46 ’ S, 45 ° 58 ’ W, 72 m depth, São Paulo state, 26 June 1970, UERJ 448; 11 females, MBT 128, 29 ° 23 ’ S, 49 ° 10 ’ W, 54 m depth, Santa Catarina state, 25 June 1970, UERJ 449; 4 females, MBT 219, 23 ° 03 ’ S, 42 ° 00 ’ W, 100 m depth, Rio de Janeiro state, 6 June 1971, UERJ 450; 1 female and 1 male, MBT 188, 24 ° 58 ’ S, 45 ° 26 ’ W, 98 m depth, São Paulo state, 30 May 1971, UERJ 451; 1 female, MBT 218, 23 ° 03 ’ S, 42 ° 19 ’ W, 75 m depth, Rio de Janeiro state, 5 June 1971, UERJ 452; 1 female, MBT 137, 30 ° 03 ’ S, 50 ° 04 ’ W, 25 m depth, Rio Grande do Sul state, 26 June 1971, UERJ 453.	en	Andrade, Luiz F., Senna, André R. (2020): Four new species of Pseudharpinia Schellenberg, 1931 (Crustacea: Amphipoda Phoxocephalidae) from southwestern Atlantic and new records of P. tupinamba Senna & Souza-Filho, 2011. Zootaxa 4763 (4): 501-537, DOI: 10.11646/zootaxa.4763.4.3
D06A385E5720BB5902BCFF005D0D6CBC.taxon	diagnosis	Diagnosis. Head with dorsal keel. Rostrum entire. Antenna 2 peduncle article 1 with a rounded ensiform process. Mandible molar with 3 multicuspidate stout setae. Maxilliped inner plate with 1 stout seta apically; outer plate with stout bipectinate setae; palp article 4 bearing elongated apical nail. Gnathopods 1 – 2 basis stout; carpus subtriangular; propodus weakly setose. Pereopod 6 basis strongly expanded posteriorly as an acute spine. Pereopod 7 basis expanded posteroventrally, reaching half of ischium. Epimeral plate 3 ventral margin concave, posteroventral corner produced into a large hook. Uropods 1 – 2 rami continuously setose to apex. Uropod 3 outer ramus continuously setose to apex. Telson short, apex rounded, with 3 setae apically on each lobe.	en	Andrade, Luiz F., Senna, André R. (2020): Four new species of Pseudharpinia Schellenberg, 1931 (Crustacea: Amphipoda Phoxocephalidae) from southwestern Atlantic and new records of P. tupinamba Senna & Souza-Filho, 2011. Zootaxa 4763 (4): 501-537, DOI: 10.11646/zootaxa.4763.4.3
D06A385E5720BB5902BCFF005D0D6CBC.taxon	description	Description. Based on female holotype, UERJ 445. Head. Antenna 1 peduncle article 1 stout, about 1.3 × longer than wide, ventral margin with brush setae distally, dorsal margin with 1 long pappose seta distally; article 2 ventral margin with 9 long pappose and 2 brush setae, dorsal margin with 2 setae distally; article 3 short, ventral margin with 3 long setae; primary flagellum 7 - articulate; accessory flagellum 5 - articulate. Antenna 2 peduncle article 2 subrectangular; article 3 ventral margin with 2 setae distally, dorsal margin with 1 seta; article 4 slightly longer than wide, ventral margin with long pappose and brush setae, facial margin with 1 brush seta medially and a row of 8 long setae extending to dorsal apex; article 5 about 0.7 × the length of article 4, ventral margin with long simple and pappose setae, dorsal margin with 2 setae subapically; flagellum 7 - articulate. Upper lip and lower lip ordinary. Mandible incisor with 6 (left) – 5 (right) teeth, lacinia mobilis with 4 (left) – 2 (right) teeth, accessory setal row with 8 (left) – 11 (right) multicuspidate stout setae, palpar hump small; palp 3 - articulate, article 1 short, article 2 weakly setose, with setae on medial margin only, article 3 slightly shorter than article 2, apex oblique, with setae extending from medial margin to apex. Maxilla 1 inner plate with 1 plumose seta apically; outer plate with 9 multicuspidate stout setae; palp article 2 medial and lateral margins with setules, apical and subapical margins with sparse setae. Maxilla 2 inner plate subequal to outer, medial margin with setules and a row with 8 plumose setae extending to apical margin; outer plate setose apically, lateral margin with 2 setae distally. Maxilliped inner plate with 5 plumose setae; outer plate medial margin with 7 bipectinate stout setae progressively increasing in length to apex, lateral margin with 1 simple and 2 plumose setae; palp article 2 medial margin moderately setose, article 3 margins with medium to long setae, article 4 about 3.2 × longer than wide, with 2 accessory setae. Pereon. Gnathopod 1 coxa expanded anteroventrally, ventral margin with 13 pappose setae; basis about 1.8 × longer than wide, posterodistal corner with 3 setae; ischium short and subrectangular, posterior margin with 1 pappose and 1 simple setae; merus posterior margin with 7 pappose setae; carpus anterior margin with 1 long seta distally, posterior margin with 4 pappose and 4 simple setae; propodus weakly setose, anterodistal margin with a tuft of setae, posterior margin with 1 stout seta defining palm; palm covered with setules on its margin, palmar hump large; dactylus subequal to palm, outer margin with 1 short seta distally. Gnathopod 2 coxa subrectangular, anterior margin slightly convex, posterior margin almost straight, ventral margin with 10 plumose setae; basis about 2.4 × longer than wide, posterior margin with 2 pappose setae, posterodistal corner with 3 pappose setae; ischium short and subrectangular, posterior margin with 2 pappose setae; merus posterior margin with 9 pappose setae; carpus anterior margin with 1 pappose seta distally, posterior margin with 2 pappose setae; propodus weakly setose, anterodistal margin with a tuft setae, posterior margin with 1 stout seta defining palm; palm covered with setules on its margin, palmar hump large; dactylus slightly longer than palm. Pereopod 3 coxa subrectangular, ventral margin with 11 plumose setae; basis about 2.4 × longer than wide, posterior margin with 1 pappose seta, posterodistal corner with a row of 5 pappose setae; ischium short and subrectangular, posterodistal corner with a row of 5 pappose setae; merus stout, anterior margin produced distally, with 5 pappose setae distally, posterior margin moderately setose with pappose setae, facial margin with a row of 6 setae distally; carpus anterior margin with 1 short seta distally, posterior margin with long pappose setae and stout setae progressively more robust and longer, facial margin with row of 6 setae; propodus anterior margin with 2 setae distally, posterior margin with 7 setae; dactylus about 85 % the length of propodus. Pereopod 4 coxa posterior margin strongly expanded, posterodorsal margin excavate, ventral margin with 10 plumose setae; remaining articles are very similar to pereopod 3, differing only by a narrower merus and some arrangements of setae. Pereopod 5 coxa bilobate, posteroventral lobe produced, with 6 pappose setae; basis about 3.9 × longer than wide, anterior margin with 4 plumose setae, anterodistal corner with a row of 4 plumose setae; ischium short and subrectangular, anterodistal corner with a row of 4 plumose setae; merus margins moderately setose, facial margin with simple setae distally; carpus anterior and posterior margins strongly setose, facial margin with row of setae distally; propodus margins strongly setose; dactylus about 65 % the length of propodus. Pereopod 6 coxa small, posterior margin with 1 short seta; basis strongly expanded posteriorly as an acute spine proximally, anterior margin with plumose setae, posterior margin with 2 plumose setae proximally and covered with setules; ischium short and subrectangular, anterior margin with 5 setae; merus anterior margin with 10 stout setae, posterior margin with 4 plumose setae proximally and 2 stout distally; carpus anterior margin with stout and simple setae, distofacial lobe produced; propodus anterior margin covered with long setae; dactylus about 65 % the length of propodus. Pereopod 7 coxa rounded; basis slightly longer than wide, posteroventral margin expanded, reaching half of ischium, anterodistal angle with a row of 5 setae, posterior margin serrated, with 2 plumose setae distally; ischium distofacial lobe produced, anterior margin with 2 setae, anterodistal angle with a row of 3 setae; merus anterior margin with 5 setae, posterior margin produced distally; carpus anterior margin with 3 setae, posterior margin with 2 setae distally; propodus anterior margin with 2 setae, posterior margin with 1 seta distally; dactylus about 90 % the length of propodus. Pleon. Epimeral plate 1 anterior margin concave, anteroventral corner with 1 seta. Epimeral plate 2 anterior margin concave, anteroventral corner produced, ventral margin with 4 setae. Epimeral plate 3 anterior margin concave, ventral margin with 2 setae, posterior margin with 3 short setae, posteroventral corner strongly produced dorsally into an acute hook. Uropod 1 peduncle 4 × longer than wide, dorsomedial margin with 9 setae, dorsolateral margin with 4 setae; outer ramus slightly longer than inner, dorsomedial and dorsolateral margins with 6 and 8 setae respectively; inner ramus dorsomedial margin with 7 setae. Uropod 2 peduncle 2.3 × longer than wide, dorsomedial margin with 4 setae, dorsolateral margin with 5 setae; outer ramus slightly longer than inner, dorsomedial and dorsolateral margins with 8 and 7 setae respectively; inner ramus dorsomedial margin with 9 setae. Uropod 3 peduncle slightly longer than wide, dorsoapical margin with a row of 7 stout setae; outer ramus slightly longer than inner, article 1 dorsomedial margin with 10 long setae continuously to apex, article 2 about 1.7 × longer than wide, with 2 long setae apically; inner ramus dorsomedial margin with 1 long seta subapically and 2 setae apically, dorsolateral margin with 4 setae. Telson about 75 % cleft, lobes rounded with 3 setae each. Sexual dimorphism. Based on male paratype, UERJ 445. Body smaller in size. Head. Antenna 1 peduncle article 1 stout, about 1.1 × longer than wide, ventral margin with brush setae and long setules distally, dorsal margin with a row of setules and 1 long plumose setae distally; article 2 ventral margin with long setae and 1 brush seta distally, dorsal margin with 1 seta distally; article 3 ventral margin with 2 setae; primary flagellum 8 - articulate; accessory flagellum 5 - articulate. Antenna 2 peduncle article 3 ventral margin with 6 setae, dorsal margin covered with setules; article 4 ventral margin with long pappose setae and 5 brush setae, facial margin with 1 brush setae and a row of 10 long setae extending to dorsal apex; article 5 slightly longer than article 4, ventral margin weakly setose, dorsal margin with 6 tufts of setules and 3 calceoli; flagellum 36 - articulate, with setules and 14 calceoli. Pereon. Gnathopods 1 – 2, similar to female, but more robust; propodus stouter; palms longer and more acute. Pereopod 6, basis, strongly expanded posteriorly as an acute, but shorter spine. Pleon. Uropod 3 peduncle 1.6 × longer than wide, dorsoapical margin with a row of 9 stout setae; outer ramus subequal to inner, article 1 margins covered with long plumose setae, article 2 about 1.6 × longer than wide, with 1 long plumose seta on each margin and 2 long setae apically; inner ramus margins covered with long plumose setae, with 2 long plumose setae apically.	en	Andrade, Luiz F., Senna, André R. (2020): Four new species of Pseudharpinia Schellenberg, 1931 (Crustacea: Amphipoda Phoxocephalidae) from southwestern Atlantic and new records of P. tupinamba Senna & Souza-Filho, 2011. Zootaxa 4763 (4): 501-537, DOI: 10.11646/zootaxa.4763.4.3
D06A385E5720BB5902BCFF005D0D6CBC.taxon	etymology	Etymology. Named after John Paul Jones, artistic name of John Richard Baldwin also known as Jonesy, bassist / keyboardist of the band Led Zeppelin, as a homage to his brilliance and contributions to music as a multi-instrumentalist.	en	Andrade, Luiz F., Senna, André R. (2020): Four new species of Pseudharpinia Schellenberg, 1931 (Crustacea: Amphipoda Phoxocephalidae) from southwestern Atlantic and new records of P. tupinamba Senna & Souza-Filho, 2011. Zootaxa 4763 (4): 501-537, DOI: 10.11646/zootaxa.4763.4.3
D06A385E5720BB5902BCFF005D0D6CBC.taxon	distribution	Type locality. Santa Catarina state coast (26 ° 34 ’ S, 47 ° 59 ’ W).	en	Andrade, Luiz F., Senna, André R. (2020): Four new species of Pseudharpinia Schellenberg, 1931 (Crustacea: Amphipoda Phoxocephalidae) from southwestern Atlantic and new records of P. tupinamba Senna & Souza-Filho, 2011. Zootaxa 4763 (4): 501-537, DOI: 10.11646/zootaxa.4763.4.3
D06A385E5720BB5902BCFF005D0D6CBC.taxon	discussion	Remarks. This species presents a very remarkable character, so far considered unique for Pseudharpinia. The presence of an acute spine projected posteriorly on the basis of pereopod 6 is an exclusive character that makes Pseudharpinia jonesyi sp. nov. easily distinguishable not only within the genus, but in Harpiniinae as well. Geographic distribution. The species is known from the coast of the Brazilian states of Rio de Janeiro, São Paulo, Santa Catarina and Rio Grande do Sul (Fig 24). Bathymetric range. 25 – 130 m depth.	en	Andrade, Luiz F., Senna, André R. (2020): Four new species of Pseudharpinia Schellenberg, 1931 (Crustacea: Amphipoda Phoxocephalidae) from southwestern Atlantic and new records of P. tupinamba Senna & Souza-Filho, 2011. Zootaxa 4763 (4): 501-537, DOI: 10.11646/zootaxa.4763.4.3
D06A385E5738BB5F02BCFBD95D336B41.taxon	description	(Figs 13 – 17)	en	Andrade, Luiz F., Senna, André R. (2020): Four new species of Pseudharpinia Schellenberg, 1931 (Crustacea: Amphipoda Phoxocephalidae) from southwestern Atlantic and new records of P. tupinamba Senna & Souza-Filho, 2011. Zootaxa 4763 (4): 501-537, DOI: 10.11646/zootaxa.4763.4.3
D06A385E5738BB5F02BCFBD95D336B41.taxon	materials_examined	Type material. Holotype: female, 4.8 mm length, dissected and drawn, MBT 144, 23 ° 53 ’ S, 42 ° 28 ’ W, 500 m depth, Rio de Janeiro state, 3 September 1970, UERJ 454.	en	Andrade, Luiz F., Senna, André R. (2020): Four new species of Pseudharpinia Schellenberg, 1931 (Crustacea: Amphipoda Phoxocephalidae) from southwestern Atlantic and new records of P. tupinamba Senna & Souza-Filho, 2011. Zootaxa 4763 (4): 501-537, DOI: 10.11646/zootaxa.4763.4.3
D06A385E5738BB5F02BCFBD95D336B41.taxon	diagnosis	Diagnosis. Head with ventrolateral lines. Rostrum entire. Antenna 1 peduncle article 3 very short; primary flagellum article 1 stout, about twice longer than wide. Antenna 2 peduncle with a narrow ensiform process. Mandible molar with 4 pectinate setae, lacinia mobilis stout and crown-shaped. Maxilliped inner plate with 1 stout seta apically; palp article 4 bearing medium apical nail. Gnathopod 1 palm sinuous, palmar hump slightly serrate. Gna- thopod 2 carpus almost cryptic; palm excavate in “ V ” shape, palmar hump slightly serrate. Pereopod 6 basis facial margin with a vertical row of setules. Pereopod 7 basis expanded posteroventrally, reaching the apex of ischium, anterodistal angle with a row of plumose setae. Epimeral plate 1 subquadrate. Epimeral plate 3 posteroventral corner produced into a medium hook. Uropod 1 peduncle dorsomedial margin with a stout curved seta distally. Telson short, apex rounded, with 1 stout seta apically on each lobe.	en	Andrade, Luiz F., Senna, André R. (2020): Four new species of Pseudharpinia Schellenberg, 1931 (Crustacea: Amphipoda Phoxocephalidae) from southwestern Atlantic and new records of P. tupinamba Senna & Souza-Filho, 2011. Zootaxa 4763 (4): 501-537, DOI: 10.11646/zootaxa.4763.4.3
D06A385E5738BB5F02BCFBD95D336B41.taxon	description	Description. Based on female holotype, UERJ 454. Head. Antenna 1 peduncle article 1 about 1.5 × longer than wide, ventral margin with brush setae distally, dorsal margin with 1 long seta distally; article 2 ventral margin setose distally with pappose setae, dorsal margin with 3 setae distally; article 3 ventral margin with 1 short and 1 long seta, dorsal margin with 1 seta; primary flagellum 7 - articulate; accessory flagellum 5 - articulate. Antenna 2 peduncle article 2 rounded; article 3 ventral margin with 1 seta, dorsal margin with 1 seta distally, facial margin with a row of 6 pappose setae; article 4 about 1.2 × longer than wide, ventral margin with medium to long pappose setae, dorsal margin with 3 setae distally, facial margin with 1 brush seta medially and a row of 7 long setae extending to dorsal apex; article 5 about 0.6 × the length of article 4, ventral margin with 7 pappose setae, dorsal margin with 3 setae distally; flagellum 7 - articulate. Upper lip and lower lip ordinary. Mandible incisor with 5 (left) – 3 (right) teeth, lacinia mobilis with 4 (left) – 2 (right) teeth, accessory setal row with 8 (left) – 7 (right) stout multicuspidate setae, palpar hump small; palp 3 - articulate, article 1 short, article 2 weakly setose, with setae on medial margin only, article 3 slightly longer than article 2, apex oblique, with setae extending from medial margin to apex. Maxilla 1 inner plate with 4 plumose setae; outer plate with 7 stout multicuspidate and 2 smooth setae; palp article 2 medial and lateral margins with setules, apical margin with 2 stout setae, distomedial margin with 2 stout and 5 slender setae. Maxilla 2 inner plate subequal to outer, medial margin with setules and a row of 10 plumose setae extending to apical margin; outer plate setose apically, lateral margin covered with setules. Maxilliped inner plate with 5 plumose setae; outer plate with 9 bipectinate stout setae progressively increasing in length to apex, lateral margin with 4 long setae; palp article 2 medial margin moderately setose, article 3 magins with medium to long setae, facial margin with 3 setae, article 4 about 3.7 × longer than wide, with 1 accessory seta. Pereon. Gnathopod 1 coxa weakly expanded anteroventrally, ventral margin with 10 pappose setae; basis about 2.5 × longer than wide, anterior margin with 5 pappose setae, posterodistal corner with 3 setae, facial margin with 3 setae; ischium short and subquadrate, posterodistal corner with 4 setae, being 2 pappose; merus posterior margin with 9 setae; carpus posterior margin with 4 setae; propodus weakly setose, anterodistal margin with a tuft of setae, posterior margin with 1 stout seta defining palm; palm covered with setules on its margin, palmar hump short; dactylus slightly shorter than palm, outer margin with 1 short seta medially. Gnathopod 2 coxa subrectangular, ventral margin with 6 pappose setae; basis about 3 × longer than wide, anterior margin with 11 setae, posterior margin with 4 setae, posterodistal corner with 5 long pappose setae; ischium short and subtriangular, posterodistal corner with a row of 7 pappose setae extending to facial margin; merus posterior margin with 10 setae, being 3 pappose, facial margin with a row of 5 setae distally; carpus posterior margin with 5 setae; propodus weakly setose, anterodistal margin with a tuft of setae, posterior margin with 1 stout seta defining palm; palm covered with setules on its margin, palmar hump large; dactylus subequal to palm, outer margin with a leaf-shaped seta distally. Pereopod 3 coxa subrectangular, ventral margin with 8 pappose setae; basis about 2.4 × longer than wide, anterior margin with 12 setae, posterior margin with 3 long setae, posterodistal corner with 5 long pappose setae, facial margin with 4 setae; ischium short and subrectangular, posterodistal corner with a row of 7 pappose setae extending to facial margin; merus anterior margin with 7 setae distally, posterior margin moderately setose, facial margin with 2 rows of 3 setae; carpus posterior margin with long slender and stout setae progressively more robust and longer, facial margin with a row of 4 setae; propodus anterior margin with 2 setae distally, posterior margin with 8 setae extending to facial margin apex; dactylus about 65 % the length of propodus. Pereopod 4 coxa posterior margin strongly expanded, posterodorsal margin excavate, ventral margin with 11 plumose setae; remaining articles are very similar to pereopod 3, differing only by some arrangements of setae. Pereopod 5 coxa bilobate, posteroventral lobe produced, with 3 pappose setae; basis about 2.1 × longer than wide, anterior margin with 7 setae, anterodistal corner with 2 long pappose setae, posterodistal corner with 1 plumose seta; ischium short and subquadrate, anterodistal angle with 4 pappose setae; merus anterior margin moderately setose, posterior margin weakly setose, facial margin with 3 stout setae distally; carpus margins moderately setose with long pappose and stout setae; propodus margins moderately setose, anterior margin with 1 stout seta distally; dactylus about 90 % the length of propodus. Pereopod 6 coxa bilobate and naked; basis about 1.3 × longer than wide, anterior margin with slender setae and 2 stout setae medially, anterodistal corner with 1 long and 2 stout setae; ischium short and subrectangular, anterodistal corner with 2 stout and 2 long setae; merus anterior margin with 4 sets of stout setae (1 - 3 - 3 - 3), posterior margin with 5 setae, being 3 pappose and a row of 3 stout and 3 simple setae distally; carpus anterior margin with 3 sets of stout setae (3 - 2 - 1) anterior margin with 1 stout and 1 short seta distally; propodus and dactylus missin. Pereopod 7 coxa subtriangular, posteroventral margin with 2 short setae; basis slightly wider than long, posteroventral margin expanded, serrate posteroventrally, almost reaching the apex of merus, anterodistal corner with a row of 9 plumose setae; ischium distofacial lobe produced, anterior margin with 9 setae, being 4 stout; merus anterior margin with 2 setae, anterodistal angle with 2 stout and 2 simple setae, posterodistal corner produced, with 5 setae; carpus anterior margin with 3 setae, anterodistal corner with 4 setae, posterodistal corner with 3 setae; propodus anterior margin with 3 setae, posterodistal corner with 2 setae; dactylus about 1.2 × longer than propodus. Pleon. Epimeral plate 1 anterior margin concave, ventral margin almost straight, posterior margin convex, with 1 short seta. Epimeral plate 2 anterior margin concave, ventral margin with 2 plumose setae, facial margin with 1 plumose seta. Epimeral plate 3 anteroventral corner with 1 seta, ventral margin with 4 setae, posteroventral corner produced into a medium hook, posterior margin covered with setules, with 2 short setae. Uropod 1 peduncle about 2.8 × longer than wide, dorsomedial margin with 5 setae and 1 stout curved seta distally, dorsolateral margin with 3 setae, being the distalmost stout; outer ramus slightly longer than inner, dorsolateral margin with 6 setae; inner ramus dorsomedial margin with 4 setae. Uropod 2 peduncle 2.4 × longer than wide, dorsomedial margin with a row of 6 setae extending to dorsolateral apex, being the distalmost stout; outer ramus slightly longer than inner, with 10 setae continuously to apex; inner ramus dorsomedial margin with 3 setae, apical margin with 1 seta. Uropod 3 missing. Telson about 80 % cleft, with an acclivity and 1 stout seta on each lobe, dorsolateral margins with 1 plumose seta.	en	Andrade, Luiz F., Senna, André R. (2020): Four new species of Pseudharpinia Schellenberg, 1931 (Crustacea: Amphipoda Phoxocephalidae) from southwestern Atlantic and new records of P. tupinamba Senna & Souza-Filho, 2011. Zootaxa 4763 (4): 501-537, DOI: 10.11646/zootaxa.4763.4.3
D06A385E5738BB5F02BCFBD95D336B41.taxon	etymology	Etymology. Named after Jimmy Page, artistic name of James Patrick Page, guitarist and founder of the band Led Zeppelin, considered one of the most influential guitarists in the world, inspiring not only musicians but we both authors during scientific discussions.	en	Andrade, Luiz F., Senna, André R. (2020): Four new species of Pseudharpinia Schellenberg, 1931 (Crustacea: Amphipoda Phoxocephalidae) from southwestern Atlantic and new records of P. tupinamba Senna & Souza-Filho, 2011. Zootaxa 4763 (4): 501-537, DOI: 10.11646/zootaxa.4763.4.3
D06A385E5738BB5F02BCFBD95D336B41.taxon	distribution	Type locality. Rio de Janeiro state coast (23 ° 53 ’ S, 42 ° 28 ’ W).	en	Andrade, Luiz F., Senna, André R. (2020): Four new species of Pseudharpinia Schellenberg, 1931 (Crustacea: Amphipoda Phoxocephalidae) from southwestern Atlantic and new records of P. tupinamba Senna & Souza-Filho, 2011. Zootaxa 4763 (4): 501-537, DOI: 10.11646/zootaxa.4763.4.3
D06A385E5738BB5F02BCFBD95D336B41.taxon	discussion	Remarks. Pseudharpinia pagei sp. nov. presents some resemblance to the Brazilian species P. berardo due to the following characters: antenna 1 peduncle article 3 short, primary flagellum article 1 stout; maxilla 1 inner plate with 4 plumose setae; gnathopod 1 palm sinuous; gnathopods 1 – 2 carpus short and almost cryptic; pereopod 7 basis wider than long, anteroventral corner with a row of plumose setae. However, the new species can be distinguished from P. berardo by the following characters (characters of P. berardo in parentheses): head with ventrolateral line present (absent); mandible incisors with 3 or 5 teeth (2 or 3), molar with 4 pectinate setae (2 setae); gnathopods 1 – 2 palmar hump distinct and slightly serrate (smooth); pereopod 6 basis facial margin with a vertical row of setules (without); pereopod 7 basis expanded posteroventrally almost reaching the apex of merus (reaching half of ischium), ischium distofacial lobe well developed (poorly developed); epimeral plate 3 posteroventral corner produced into a medium hook (produced into a larger hook), posterior margin covered with setules (without setules); telson with 1 stout seta on each lobe (2 setae), dorsolateral margins with 1 plumose seta (2 setae, being 1 plumose).	en	Andrade, Luiz F., Senna, André R. (2020): Four new species of Pseudharpinia Schellenberg, 1931 (Crustacea: Amphipoda Phoxocephalidae) from southwestern Atlantic and new records of P. tupinamba Senna & Souza-Filho, 2011. Zootaxa 4763 (4): 501-537, DOI: 10.11646/zootaxa.4763.4.3
D06A385E5738BB5F02BCFBD95D336B41.taxon	distribution	Geographic distribution. The species is known only from the type locality, Rio de Janeiro state coast (Fig 24).	en	Andrade, Luiz F., Senna, André R. (2020): Four new species of Pseudharpinia Schellenberg, 1931 (Crustacea: Amphipoda Phoxocephalidae) from southwestern Atlantic and new records of P. tupinamba Senna & Souza-Filho, 2011. Zootaxa 4763 (4): 501-537, DOI: 10.11646/zootaxa.4763.4.3
D06A385E5738BB5F02BCFBD95D336B41.taxon	discussion	Bathymetric range. 500 m depth.	en	Andrade, Luiz F., Senna, André R. (2020): Four new species of Pseudharpinia Schellenberg, 1931 (Crustacea: Amphipoda Phoxocephalidae) from southwestern Atlantic and new records of P. tupinamba Senna & Souza-Filho, 2011. Zootaxa 4763 (4): 501-537, DOI: 10.11646/zootaxa.4763.4.3
D06A385E573EBB5702BCFBFC5D456E35.taxon	description	(Figs 18 – 23)	en	Andrade, Luiz F., Senna, André R. (2020): Four new species of Pseudharpinia Schellenberg, 1931 (Crustacea: Amphipoda Phoxocephalidae) from southwestern Atlantic and new records of P. tupinamba Senna & Souza-Filho, 2011. Zootaxa 4763 (4): 501-537, DOI: 10.11646/zootaxa.4763.4.3
D06A385E573EBB5702BCFBFC5D456E35.taxon	materials_examined	Type material. Holotype: female, 3.5 mm length, dissected and drawn, MBT 218, 23 ° 03 ’ S, 42 ° 19 ’ W, 75 m depth, Rio de Janeiro state, 5 June 1971, UERJ 455. Paratype: male, dissected and drawn, same data as for the holotype, UERJ 456. Additional material examined. 4 females, same data as for the holotype, UERJ 457.	en	Andrade, Luiz F., Senna, André R. (2020): Four new species of Pseudharpinia Schellenberg, 1931 (Crustacea: Amphipoda Phoxocephalidae) from southwestern Atlantic and new records of P. tupinamba Senna & Souza-Filho, 2011. Zootaxa 4763 (4): 501-537, DOI: 10.11646/zootaxa.4763.4.3
D06A385E573EBB5702BCFBFC5D456E35.taxon	diagnosis	Diagnosis. Head with dorsal keel and ventrolateral line. Rostrum entire. Antenna 1 primary and accessory flagella article 1 elongate. Antenna 2 peduncle article 1 with a subacute ensiform process. Mandible molar with 3 pectinate stout setae, left lacinia mobilis stout. Maxilla 1 inner plate with 4 setae. Maxilla 2 inner plate slightly longer than outer. Maxilliped inner plate with 1 stout seta apically; outer plate medial margin with stout blunt brush setae extending to apical margin; palp article bearing elongated apical nail. Pereopods 3 – 4 carpus anterior margin with a comb-like structure distally. Pereopod 6 carpus with a distofacial rounded lobe produced and covered with setules. Pereopod 7 basis expanded posteroventrally, reaching half of merus; merus strongly produced posteroventrally. Uropods 1 – 2 peduncle ventroapical corner combed. Telson with 2 long setae on each lobe.	en	Andrade, Luiz F., Senna, André R. (2020): Four new species of Pseudharpinia Schellenberg, 1931 (Crustacea: Amphipoda Phoxocephalidae) from southwestern Atlantic and new records of P. tupinamba Senna & Souza-Filho, 2011. Zootaxa 4763 (4): 501-537, DOI: 10.11646/zootaxa.4763.4.3
D06A385E573EBB5702BCFBFC5D456E35.taxon	description	Description. Based on female holotype, UERJ 455. Head. Antenna 1 peduncle article 1 about 1.5 × longer than wide, ventral margin with brush setae distally, dorsal margin with 1 long seta distally; article 2 ventral margin with 1 brush, 1 long and 5 pappose setae, dorsal margin with 1 seta distally; article 3 short, ventral margin with 1 short and 1 stout seta distally, facial margin with 1 brush and 2 short setae distally; primary flagellum 6 - articulate, article 1 about 3 × longer than wide; accessory flagellum 4 - articulate, article 1 about 3.8 × longer than wide. Antenna 2 peduncle article 2 short and subrectangular; article 3 facial margin with 2 setae; article 4 slightly longer than wide, ventral margin with long pappose and brush setae, facial margin with 1 brush seta medially and a row of 8 long setae extending to dorsal apex; article 5 about 0.6 × the length of article 4, ventral margin with 1 brush and 7 pappose setae, dorsal margin with 1 seta distally; flagellum 7 - articulate. Upper lip and lower lip ordinary. Mandible incisor with 3 (left) – 4 (right) teeth, lacinia mobilis with 9 teeth (left) — bifid and cuspidate (right), accessory setal row with 8 (left) — 4 (right) multicuspidate stout setae, palpar hump small; palp 3 - articulate, article 1 short, article 2 weakly setose with 1 short seta proximally and 2 long distally, article 3 slightly shorter than article 2, apex oblique, with setae extending from medial margin to apex. Maxilla 1 outer plate with stout setae, being 7 multicuspidate; palp article 2 medial margin with 3 setae distally, apical margin with 3 setae, lateral margin with setules distally. Maxilla 2 inner plate medial margin covered with set- ules, apicomedial corner with larger plumose setae, apical margin with 5 setae, lateral margin with setules distally; outer plate apical margin setose, lateral margin with setules and 2 simple setae. Maxilliped inner plate with 5 long plumose setae; outer plate medial margin with 6 stout blunt brush setae progressively increasing in length to apex, lateral margin with 2 long stout setae, being 1 plumose; palp article 2 medial margin weakly setose, article 3 margins weakly setose, with long setae apically, article 4 about 2.1 × longer than wide, with 2 accessory setae. Pereon. Gnathopod 1 coxa expanded anteroventrally, ventral margin with 1 short and 16 long pappose setae; basis about 2.3 × longer than wide, anterior and posterior margins with 3 setae, posterodistal corner with 1 seta; ischium short and subquadrate, posterior margin with 2 setae, being 1 pappose; merus posterior margin with 2 simple and 2 pappose setae; carpus posterior margin with 8 setae; propodus weakly setose, anterodistal margin with a tuft of setae, posterior margin with 1 stout seta defining palm; palm covered with setules on its margin, palmar hump medium; dactylus reaching the palmar corner, outer margin with 1 short seta medially and 1 leaf-shaped seta distally. Gnathopod 2 coxa subrectangular, ventral margin with 10 pappose setae; basis about 3 × longer than wide, anterior margin with 9 setae, posterior margin with 2 setae, posterodistal corner with 3 pappose setae; ischium short and subquadrate, posterodistal corner with 2 setae; merus posterior margin with 6 pappose setae; carpus posterior margin with 5 setae; propodus weakly setose, anterodistal margin with a tuft of setae, posterior margin with 1 stout seta defining palm; palm covered with setules on its margin, palmar hump medium; dactylus reaching the palmar hump, outer margin with 1 short seta medially and 1 leaf-shaped seta distally. Pereopod 3 coxa subrectangular, ventral margin with 10 pappose setae; basis about 2.8 × longer than wide, posterior margin with 4 long setae, posterodistal corner with 4 long pappose setae; ischium short and subrectangular, posterodistal corner with 5 pappose setae; merus anterior margin with 4 setae distally, posterior margin moderately setose, facial margin with 4 setae distally; carpus anterior margin with 1 short seta distally, posterior margin with long slender and stout setae progressively more robust and longer, facial margin with a row of 4 setae; propodus anterior margin with 2 setae distally, posterior margin with 1 stout seta distally and a row of 6 setae extending to facial margin; dactylus about 85 % the length of propodus. Pereopod 4 coxa posterior margin strongly expanded, posterodorsal margin excavate, ventral margin with 12 pappose setae; remaining articles are very similar to pereopod 3, differing only by some arrangements of setae. Pereopod 5 coxa bilobate, posteroventral lobe strongly produced, with 4 pappose setae and a row of setules; basis about 2.8 × longer than wide, anterior margin with a row of setules proximally, 2 setae and 1 pappose seta distally, anterodistal corner with 4 setae, posterior margin with 1 plumose seta distally; ischium short and subrectangular, anterodistal corner with 5 pappose setae; merus anterior margin moderately setose, with a row of 7 setae distally, posterior margin with 5 pappose setae and a row of 5 setae distally; carpus anterior margin moderately setose, with 2 stout setae and a row of 4 setae distally, posterior margin with 5 plumose setae and a row of 5 setae distally; propodus anterior margin moderately setose, with 2 stout setae, posterior margin setose; dactylus about 65 % the length of propodus. Pereopod 6 coxa subtriangular, posterior margin with 1 short seta; basis about 1.4 × longer than wide, anterior margin with plumose setae extending to facial apex and 2 stout setae, posterior margin with 1 seta and a row of setules; ischium short and subrectangular, anterodistal corner with 1 short and 2 plumose setae, posterior margin with setules; merus anterior margin with 3 stout setae, posterior margin with 2 stout setae distally; carpus anterior margin with 4 setae, posterior margin with 2 setae distally; propodus anterior margin with 8 long setae; dactylus about 60 % the length of propodus. Pereopod 7 coxa trapezoidal, posterior margin with 1 short seta; basis about 1.3 × longer than wide, posteroventral margin expanded and serrate, reaching half of merus, ventral margin with 5 plumose setae, anterodistal corner with 1 stout and 4 long plumose setae; ischium distofacial lobe produced, anterior margin with 2 stout and 4 long plumose setae; merus anterior margin with 5 setae, posterior margin with 1 seta distally; carpus anterior margin with 3 seta, posterior margin with 2 setae; propodus posterior margin with 1 seta distally; dactylus slightly longer than propodus. Pleon. Epimeral plate 1 anterior margin covered with setules, ventral margin and posteroventral corner rounded. Epimeral plate 2 ventral margin weakly concave, with 3 plumose setae, posterior margin convex. Epimeral plate 3 ventral margin weakly concave, with 7 plumose setae, posteroventral corner strongly produced dorsally into a hook, posterior margin with 3 setae. Uropod 1 peduncle 2.9 × longer than wide, dorsomedial and dorsolateral margins with 3 stout setae distally each; outer ramus slightly longer than inner, dorsomedial margin with 8 setae, dorsolateral margin with 5 setae; inner ramus dorsomedial margin with 7 setae. Uropod 2 peduncle about 2.5 × longer than wide, dorsomedial margin with 1 seta distally, dorsolateral margin with 3 setae; outer ramus slightly longer than inner, dorsomedial margin with 5 setae, dorsolateral margin with 2 setae; inner ramus dorsomedial margin with 4 setae. Uropod 3 peduncle slightly longer than wide, dorsoapical margin with a row of 8 setae; outer ramus 1.4 × longer than inner, article 1 dorsomedial margin with 7 long setae, dorsolateral margin with 2 setae, article 2 about 1.4 × longer than wide, with 2 long setae apically. Telson about 70 % cleft, lobes rounded with 2 long setae apically each, dorsolateral margin with 2 plumose setae on each side. Sexual dimorphism. Based on male paratype, UERJ 456. Body smaller in size. Head. Antenna 1 peduncle article 1 about 1.3 × longer than wide, ventral margin with brush setae and long setules distally, dorsal margin with a row of setules and 1 seta distally; article 2 ventral margin with 6 long setae, dorsal margin with 1 seta distally, facial margin with 1 brush seta; article 3 ventral and facial margins with 2 setae each; primary flagellum 6 - articulate; accessory flagellum 4 - articulate. Antenna 2 peduncle article 2 subtriangular; article 3 ventral margin with 1 seta, dorsal margin covered with setules, facial margin with 2 setae; article 4 ventral margin with long pappose and brush setae, dorsal margin covered with setules, facial margin with 1 brush seta and a row of 7 long setae extending to dorsal apex; article 5 slightly longer than article 4, ventral margin with simple and brush setae, dorsal margin with 6 tufts of setules and 4 calceoli; flagellum 36 - articulate, with setules and 15 calceoli. Pereon. Gnathopods 1 – 2, similar to female, but more robust; palms margins minutely pectinate, palmar humps slightly larger. Pleon. Uropod 3 peduncle 1.2 × longer than wide, dorsomedial margin with 2 setae distally, dorsoapical margin with a row of 5 setae; outer ramus slightly longer than inner, article 1 margins covered with plumose setae, article 2 about 1.3 × longer than wide, with 2 long plumose setae apically; inner ramus margins covered with plumose setae, with 2 long plumose setae apically.	en	Andrade, Luiz F., Senna, André R. (2020): Four new species of Pseudharpinia Schellenberg, 1931 (Crustacea: Amphipoda Phoxocephalidae) from southwestern Atlantic and new records of P. tupinamba Senna & Souza-Filho, 2011. Zootaxa 4763 (4): 501-537, DOI: 10.11646/zootaxa.4763.4.3
D06A385E573EBB5702BCFBFC5D456E35.taxon	etymology	Etymology. Named after Robert Plant, artistic name of Robert Anthony Plant, lead singer of the band Led Zeppelin and owner of one of the greatest voices in the world.	en	Andrade, Luiz F., Senna, André R. (2020): Four new species of Pseudharpinia Schellenberg, 1931 (Crustacea: Amphipoda Phoxocephalidae) from southwestern Atlantic and new records of P. tupinamba Senna & Souza-Filho, 2011. Zootaxa 4763 (4): 501-537, DOI: 10.11646/zootaxa.4763.4.3
D06A385E573EBB5702BCFBFC5D456E35.taxon	distribution	Type locality. Rio de Janeiro state coast (23 ° 03 ’ S, 42 ° 19 ’ W).	en	Andrade, Luiz F., Senna, André R. (2020): Four new species of Pseudharpinia Schellenberg, 1931 (Crustacea: Amphipoda Phoxocephalidae) from southwestern Atlantic and new records of P. tupinamba Senna & Souza-Filho, 2011. Zootaxa 4763 (4): 501-537, DOI: 10.11646/zootaxa.4763.4.3
D06A385E573EBB5702BCFBFC5D456E35.taxon	discussion	Remarks. Pseudharpinia planti sp. nov. presents the following new combination of character states within the genus: head with dorsal keel and ventrolateral line; antenna 1 flagella article 1 elongate; left mandible with stout la- cinia mobilis, with 9 teeth; maxilliped outer plate with stout blunt brush setae; pereopods 3 – 4 carpus anterior margin with a comb-like structure distally; pereopod 6 carpus with a distofacial rounded lobe produced and covered with setules; telson with 2 long setae on each lobe apically each.	en	Andrade, Luiz F., Senna, André R. (2020): Four new species of Pseudharpinia Schellenberg, 1931 (Crustacea: Amphipoda Phoxocephalidae) from southwestern Atlantic and new records of P. tupinamba Senna & Souza-Filho, 2011. Zootaxa 4763 (4): 501-537, DOI: 10.11646/zootaxa.4763.4.3
D06A385E573EBB5702BCFBFC5D456E35.taxon	distribution	Geographic distribution. The species is known only from the type locality, Rio de Janeiro state coast (Fig 24).	en	Andrade, Luiz F., Senna, André R. (2020): Four new species of Pseudharpinia Schellenberg, 1931 (Crustacea: Amphipoda Phoxocephalidae) from southwestern Atlantic and new records of P. tupinamba Senna & Souza-Filho, 2011. Zootaxa 4763 (4): 501-537, DOI: 10.11646/zootaxa.4763.4.3
D06A385E573EBB5702BCFBFC5D456E35.taxon	discussion	Bathymetric range. 75 m depth.	en	Andrade, Luiz F., Senna, André R. (2020): Four new species of Pseudharpinia Schellenberg, 1931 (Crustacea: Amphipoda Phoxocephalidae) from southwestern Atlantic and new records of P. tupinamba Senna & Souza-Filho, 2011. Zootaxa 4763 (4): 501-537, DOI: 10.11646/zootaxa.4763.4.3
D06A385E5737BB6802BCFF005D0D6D31.taxon	description	(Fig 25)	en	Andrade, Luiz F., Senna, André R. (2020): Four new species of Pseudharpinia Schellenberg, 1931 (Crustacea: Amphipoda Phoxocephalidae) from southwestern Atlantic and new records of P. tupinamba Senna & Souza-Filho, 2011. Zootaxa 4763 (4): 501-537, DOI: 10.11646/zootaxa.4763.4.3
D06A385E5737BB6802BCFF005D0D6D31.taxon	materials_examined	Material examined. 1 female, dissected and drawn, MBT 188, 24 ° 58 ’ S, 45 ° 26 ’ W, 98 m depth, São Paulo state, 30 May 1971, UERJ 458. 7 females, MBT 219, 23 ° 03 ’ S, 42 ° 00 ’ W, 100 m depth, Rio de Janeiro state, 6 June 1971, UERJ 459.	en	Andrade, Luiz F., Senna, André R. (2020): Four new species of Pseudharpinia Schellenberg, 1931 (Crustacea: Amphipoda Phoxocephalidae) from southwestern Atlantic and new records of P. tupinamba Senna & Souza-Filho, 2011. Zootaxa 4763 (4): 501-537, DOI: 10.11646/zootaxa.4763.4.3
D06A385E5737BB6802BCFF005D0D6D31.taxon	diagnosis	Amended diagnosis (after Senna & Souza-Filho, 2011). Antenna 1 peduncle article 1 dorsally smooth or with setules; article 2 anteroventral corner with 5 – 7 pappose or plumose setae. Antenna 2 peduncle article 4 ventral margin with 7 – 9 pappose or plumose setae and 3 – 5 brush setae, facial margin with 1 brush seta and a row of 8 – 9 setae extending to dorsal apex. Mandible incisors with 6 teeth, accessory setal row with 6 pectinate setae; right lacinia mobilis multicuspidate apically. Maxilla 1 inner plate with 4 short plumose setae apically. Maxilla 2 inner plate with 7 plumose setae apically. Gnathopod 1 coxa ventral margin with 14 – 16 pappose or plumose setae; propodus about 2.5 × longer than wide, posterior margin about 1.5 – 2 × longer than palm; palm sinuous. Gnathopod 2 coxa ventral margin with 9 – 13 pappose or plumose setae; propodus about 2.5 × longer than wide, posterior margin about 1.5 – 2 × longer than palm. Pereopod 3 carpus, posterior margin with 7 long curved stout setae. Pereopod 6 basis anterior margin with a facial row of long plumose setae and 4 short stout setae, posterior margin slightly concave, with 3 blunt lobes proximally and 4 – 5 plumose setae; carpus anterior margin with 3 stout setae. Pereopod 7 basis posteroventral lobe reaching the apex of merus; dactyl strong, about 1.1 × longer than propodus. Epimeral plate ventral margin with 5 short plumose setae. Epimeral plate 3 ventral margin with 3 small and 2 long plumose setae, posterior margin with 2 short setae. Uropod 1 peduncle with dorsoapical corner bearing 3 stout setae. Uropod 2 peduncle dorsodistal margin with corona of cuticular spines, rami continuously bearing trifid stout setae. Uropod 3 inner ramus about 1.1 × longer than peduncle; outer ramus article 2 about 0.1 × as long as article 1, bearing 2 long slender setae apically. Telson with 2 apicolateral stout setae on each lobe.	en	Andrade, Luiz F., Senna, André R. (2020): Four new species of Pseudharpinia Schellenberg, 1931 (Crustacea: Amphipoda Phoxocephalidae) from southwestern Atlantic and new records of P. tupinamba Senna & Souza-Filho, 2011. Zootaxa 4763 (4): 501-537, DOI: 10.11646/zootaxa.4763.4.3
D06A385E5737BB6802BCFF005D0D6D31.taxon	description	Description. Based on material examined female UERJ 458. Head. Antenna 1 peduncle article 1 about 1.3 × longer than wide, ventral margin with 6 plumose setae distally, dorsal margin with 1 seta distally; article 2 ventral margin with 7 long pappose setae and 2 brush setae, dorsal margin with 1 seta distally; article 3 short, ventral margin with 2 setae, facial margin with 1 brush and 2 short setae; primary flagellum 7 - articulate, presenting aestethasc on articles 5 – 6; accessory flagellum 5 - articulate. Antenna 2 peduncle article 1 with a well-developed ensiform process; article 2 short and subrectangular; article 3 ventral margin with 2 short setae, dorsal margin with 1 seta distally, facial margin with 3 setae; article 4 about 1.2 × longer than wide, ventral margin with 5 long pappose and 5 brush setae, dorsal margin with 2 setae proximally and 3 long setae distally, facial margin with 1 brush seta and a row of 9 long setae extending to dorsal apex; article 5 about 0.6 × the length of article 4, ventral margin with 6 long pappose setae; flagellum 7 - articulate. Pereon. Gnathopod 1 coxa weakly expanded anteroventrally, ventral margin with 16 long pappose setae; basis about 2.5 × longer than wide, anterior margin with 3 setae, posterior margin with 3 long setae, posterodistal corner with 3 pappose setae; ischium short and subrectangular, posterior margin with 3 pappose setae; merus anterior margin with 1 seta distally, posterior margin with 5 pappose setae; carpus anterior margin with 1 seta distally posterior margin with 6 pappose setae; propodus weakly setose, anterodistal margin with a tuft of setae, posterior margin with 1 stout seta defining palm; palm covered with setules on its margin, palmar hump medium; dactylus reaching the palmar corner, outer margin with 1 short seta medially. Gnathopod 2 coxa subrectangular, ventral margin with 13 long pappose setae; basis about 3.4 × longer than wide anterior margin with 9 setae, posterior margin with 2 long setae, posterodistal corner with 5 long pappose setae; ischium short and subrectangular, posterior margin with 3 pappose setae; merus posterior margin with 8 pappose setae; carpus anterior margin with 1 seta distally, posterior margin with 6 pappose setae; propodus weakly setose, anterodistal margin with a tuft of setae, posterior margin with 1 stout seta defining palm; palm covered with setules on its margin, palmar hump large; dactylus reaching the palmar corner. Pereopod 6 coxa weakly expanded posteroventrally, posterior margin with 1 short seta; basis about 2 × longer than wide, anterior margin with 4 stout setae and a row of long plumose setae extending to facial margin, posterior margin with 3 blunt lobes and 5 plumose setae proximally and a row of setules distally; ischium short and subrectangular, naked; merus anterior margin with 1 simple and 3 stout setae, posterior margin with 2 stout setae distally; carpus anterior margin with 3 stout and 2 simple setae, posterior margin with 1 seta distally; propodus anterior margin with 13 long setae; dactylus about 50 % the length of propodus.	en	Andrade, Luiz F., Senna, André R. (2020): Four new species of Pseudharpinia Schellenberg, 1931 (Crustacea: Amphipoda Phoxocephalidae) from southwestern Atlantic and new records of P. tupinamba Senna & Souza-Filho, 2011. Zootaxa 4763 (4): 501-537, DOI: 10.11646/zootaxa.4763.4.3
D06A385E5737BB6802BCFF005D0D6D31.taxon	materials_examined	Type locality. Off the estuarine complex of Santos, north coast of São Paulo state (24 º 10 ’ 32 ” S, 45 º 29 ’ 10 ” W).	en	Andrade, Luiz F., Senna, André R. (2020): Four new species of Pseudharpinia Schellenberg, 1931 (Crustacea: Amphipoda Phoxocephalidae) from southwestern Atlantic and new records of P. tupinamba Senna & Souza-Filho, 2011. Zootaxa 4763 (4): 501-537, DOI: 10.11646/zootaxa.4763.4.3
D06A385E5737BB6802BCFF005D0D6D31.taxon	discussion	Remarks. Pseudharpinia tupinamba was described for the estuarine complex of Santos (São Paulo state) and	en	Andrade, Luiz F., Senna, André R. (2020): Four new species of Pseudharpinia Schellenberg, 1931 (Crustacea: Amphipoda Phoxocephalidae) from southwestern Atlantic and new records of P. tupinamba Senna & Souza-Filho, 2011. Zootaxa 4763 (4): 501-537, DOI: 10.11646/zootaxa.4763.4.3
D06A385E5737BB6802BCFF005D0D6D31.taxon	distribution	Geographic distribution. The species is known for the type locality (São Paulo state), Ilha Grande Bay (Rio de Janeiro state) and it is now recorded from new localities at both states (Fig 26). There are no geographic coordinates available for the Ilha Grande Bay record.	en	Andrade, Luiz F., Senna, André R. (2020): Four new species of Pseudharpinia Schellenberg, 1931 (Crustacea: Amphipoda Phoxocephalidae) from southwestern Atlantic and new records of P. tupinamba Senna & Souza-Filho, 2011. Zootaxa 4763 (4): 501-537, DOI: 10.11646/zootaxa.4763.4.3
D06A385E5737BB6802BCFF005D0D6D31.taxon	discussion	Bathymetric range. 58 – 100 m depth.	en	Andrade, Luiz F., Senna, André R. (2020): Four new species of Pseudharpinia Schellenberg, 1931 (Crustacea: Amphipoda Phoxocephalidae) from southwestern Atlantic and new records of P. tupinamba Senna & Souza-Filho, 2011. Zootaxa 4763 (4): 501-537, DOI: 10.11646/zootaxa.4763.4.3
