taxonID	type	description	language	source
8F682BCCD00D5DED925904BB1E03B143.taxon	description	Figs 4 A, 8 A, 10 D, 12 D, 15	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
8F682BCCD00D5DED925904BB1E03B143.taxon	description	Description. Size and form: Body length 1.7 - 2 mm. Body form elongate oval, moderately convex in lateral view (Fig. 4 A). Color and punctation: Dorsally bicolor, head brown, frons dark brown; pronotum yellow with two small black round spots along posterior margin; elytra dark brown, elytra margins paler (Fig. 4 A). Ventrally brown; maxillary palps, mouthparts, and antennae yellow (antennal club slightly darker), pro legs yellow, meso and meta legs pale brown. Clypeus and labrum with dense, fine, and weakly impressed ground punctation (punctures separated by 2 x their width); pronotum and elytra ground punctation fine, weakly impressed and sparser than on head (punctures separated by 3 - 4 x their width). Head: Clypeus and labrum shallowly emarginate anteromedially, lateral margins of the labrum bearing setae. Thorax: Prosternum carinate medially, strongly raised, acute and pointing anteriorly. Elevation of mesoventrite with two transversal ridges, elevated medially, lateral sides concave; longitudinal ridge broad anteriorly and sharp posteriorly, the point where the three ridges merged wide and blunt (Fig. 10 D); elevation flat in ventral view; mesoventrite with triangular shape in ventral view. Metaventrite convex in the median region, pubescent with narrow glabrous patch on the medial and posterolateral area; anterior margin extending to mesoventrite elevation. Metafemora with dense hydrofuge pubescence along basal three-quarters of the anterior margin and sparse pubescence along the entire basal posterior margin. Abdomen: Abdominal ventrites very densely pubescent. Aedeagus (Fig. 8 A) with basal piece 0.7 x the length of a paramere. Base of the parameres slightly narrower than the base of the median lobe; outer margin convex, inner margins along basal two-thirds slightly convex, apical third straight; apex of parameres slightly acute, pointing inwards. Median lobe shorter than the parameres, wide at basal region, narrowing in the midlength, then sinuate, apex slightly acute; gonopore situated at the apex of median lobe.	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
8F682BCCD00D5DED925904BB1E03B143.taxon	etymology	Etymology. The name derived from the Latin words bi meaning two and color meaning hue, referring to the dorsal coloration of the species. This species has yellow coloration in the pronotum and black in the elytra.	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
8F682BCCD00D5DED925904BB1E03B143.taxon	distribution	Distribution. Known from two localities in central Suriname: Kabalebo and the summit of Tafelberg Tepui (Fig. 15).	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
0D0CE7E52C3C530E95A1195899958AFD.taxon	description	Figs 4 J, 9 D, 12 A, 14	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
0D0CE7E52C3C530E95A1195899958AFD.taxon	description	Description. Size and form: Body length 1.5 - 1.7 mm. Body form elongate oval, moderately convex in lateral view (Fig. 4 J). Color and punctation: Dorsally yellow, head mostly dark brown, frons and medial region of the clypeus dark brown, lateral sides of clypeus yellow; pronotum yellow with two small dark brown round spots along posterior margin; elytra yellow (Fig. 4 J). Ventrally dark brown; maxillary palps, mouthparts, and antennae yellow. Pro legs yellow, meso and meta legs pale brown. Clypeus and labrum with dense, fine, and weakly impressed ground punctation (punctures separated by 2 x their width); pronotum and elytra ground punctation fine, weakly impressed and sparser than on head (punctures separated by 3 x their width). Head: Clypeus and labrum shallowly emarginate anteromedially, lateral margins of the labrum bearing setae. Thorax: Prosternum carinate medially, strongly raised, acute and pointing anteriorly. Elevation of mesoventrite with two transversal ridges, elevated medially, lateral sides concave; longitudinal ridge broad anteriorly and sharp posteriorly, the point where the three ridges merged wide and blunt (e. g., Fig. 10 C, D); elevation flat in ventral view; mesoventrite with triangular shape in ventral view. Metaventrite convex in the median region, pubescent with broad glabrous patch on the medial and posterolateral area; anterior margin extending to mesoventrite elevation. Metafemora with dense hydrofuge pubescence along basal three-quarters of the anterior margin and along basal one-quarter of posterior margin, then apical half of posterior margin with sparse setae. Abdomen: Abdominal ventrites very densely pubescent. Aedeagus (Fig. 9 D) with basal piece 0.6 x the length of a paramere. Base of the parameres narrower than the base of the median lobe; outer margins straight along basal two-thirds, then narrowing abruptly along apical third, inner margins convex along basal two-thirds and then apically slightly concave; apex of parameres rounded. Median lobe shorter than the parameres, approximately rectangular, narrow along apical fifth, apex bifurcated; gonopore drop-shaped and situated at apical fourth of median lobe.	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
0D0CE7E52C3C530E95A1195899958AFD.taxon	etymology	Etymology. The specific name comes from the Latin word bifidus meaning split into two parts, after the form of the median lobe " bifurcated apically " of the aedeagus.	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
0D0CE7E52C3C530E95A1195899958AFD.taxon	distribution	Distribution. Known only from the type locality in Venezuela (Fig. 14).	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
2333901953345BC580D9908E62640698.taxon	description	Figs 4 C, 8 C, 12 C, 15	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
2333901953345BC580D9908E62640698.taxon	description	Description. Size and form: Body length 1.7 - 1.9 mm. Body form elongate oval, moderately convex in lateral view (Fig. 4 C). Color and punctation: Dorsally dark reddish brown, lateral margins of clypeus, pronotum and elytra yellow brown (Fig. 4 C). Ventrally reddish brown, except for black abdominal ventrites; maxillary palps, mouth parts and antennae yellow (antennal club slightly darker). Clypeus and labrum with dense, fine, and weakly impressed ground punctation (punctures separated by 2 x their width); pronotum and elytra ground punctation fine, weakly impressed, and sparser than on head (punctures separated by 3 - 4 x their width). Head: Clypeus and labrum shallowly emarginate anteromedially, lateral margins of the labrum bearing setae. Thorax: Prosternum carinate medially, strongly raised, acute and pointing anteriorly. Elevation of mesoventrite with two transversal ridges, elevated medially, lateral sides concave; longitudinal ridge broad anteriorly and sharp posteriorly, the point where the three ridges merged wide and blunt (e. g., Fig. 10 C, D); elevation flat in ventral view; mesoventrite with triangular shape in ventral view. Metaventrite convex in the median region, pubescent with narrow glabrous patch on the medial and posterolateral area; anterior margin extending to mesoventrite elevation. Metafemora with dense hydrofuge pubescence along three-quarters of the anterior basal margin and sparse pubescence along the posterior basal margin. Abdomen: Abdominal ventrites very densely pubescent. Aedeagus (Fig. 8 C) with basal piece 0.7 x the length of a paramere. Base of the parameres narrower than the base of the median lobe; outer margins slightly convex, inner margins straight; apex of parameres acute and pointing outwards. Median lobe shorter than the parameres, approximately triangular, wide at the basal region, then slightly narrowing to the apex, apex acute, gonopore oval in shape and situated at the apex of the median lobe.	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
2333901953345BC580D9908E62640698.taxon	etymology	Etymology. The name is a combination of two Latin words brun meaning dark and badius meaning reddish brown, highlighting the distinguishable dark reddish brown color of this species.	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
2333901953345BC580D9908E62640698.taxon	distribution	Distribution. Known only from the type locality near Manaus, Brazil (Fig. 15).	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
CAA1758E780E5BE7BBD01A536AFE704F.taxon	description	Figs 5 A-J, 10 A, 15	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
0F11712912915924B9FD43329AF6AD68.taxon	description	Figs 4 G, 8 H, 13 D, 15	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
0F11712912915924B9FD43329AF6AD68.taxon	description	Description. Size and form: Body length 1.7 - 1.8 mm. Body form elongate oval, convex in lateral view (Fig. 4 G). Color and punctation: Dorsally bicolor, head mostly brown, frons dark brown, posteromedial margin of the clypeus pale brown and lateral sides and anterior margin yellow; pronotum yellow with two small black round spots along posterior margin; elytra brown, lateral and posterior side of the elytra yellow (Fig. 4 G). Ventrally dark brown; maxillary palps, mouthparts, and antennae yellow. Pro and meso legs yellow, meta legs pale brown. Clypeus and labrum with dense, fine, and weakly impressed ground punctation (punctures separated by 2 x their width); pronotum and elytra ground punctation weakly impressed, fine, and sparser than on head (punctures separated by 3 x their width). Head: Clypeus and labrum shallowly emarginate anteromedially, lateral margins of the labrum bearing setae. Thorax: Prosternum carinate medially, strongly raised, acute and pointing anteriorly. Elevation of mesoventrite with two transversal ridges, elevated medially, lateral sides concave; longitudinal ridge broad anteriorly and sharp posteriorly, the point where the three ridges merged wide and blunt (e. g., Fig. 10 C, D); elevation flat in ventral view; mesoventrite with triangular shape in ventral view. Metaventrite convex in the median region, pubescent with narrow glabrous patch on the medial and posterolateral area; anterior margin extending to mesoventrite elevation. Metafemora with dense hydrofuge pubescence along basal three-quarters of the anterior margin and sparse pubescence along the apical posterior margin. Abdomen: Abdominal ventrites very densely pubescent. Aedeagus (Fig. 8 H) with basal piece 0.4 x the length of a paramere. Base of the parameres narrower than the base of the median lobe; outer margins nearly convex along basal two-thirds, then curved inwards along apex, inner margins nearly straight; apex of parameres acute. Median lobe shorter than the parameres, wide at basal region, narrowing in the midlength and then slightly broadening to apex, apex rounded and wide; gonopore ovate in shape and situated at apical fourth of median lobe.	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
0F11712912915924B9FD43329AF6AD68.taxon	etymology	Etymology. This species is named after Andrea Lorena Garcia Hernandez curator at the Coleccion de Insectos de la Universidad del Quindio (CIUQ) in recognition to her passion and contribution of the knowledge of the insects, specially hydrophilids in Colombia.	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
0F11712912915924B9FD43329AF6AD68.taxon	distribution	Distribution. This species is known from several collecting events at the Ducke Reserve near Manaus, Brazil (Fig. 15).	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
DE5FE26820BA5CAAB1C52315B54C204C.taxon	description	Figs 4 I, 7 D, 13 A, 14	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
DE5FE26820BA5CAAB1C52315B54C204C.taxon	description	Description. Size and form: Body length 1.7 - 1.9 mm. Body form elongate oval, convex in lateral view (Fig. 4 I). Color and punctation: Dorsally yellow, head yellow; pronotum paler than elytra, with two small black round spots along posterior margin (Fig. 4 I). Ventrally brown; maxillary palps, mouthparts, antennae (antennal club slightly darker) and legs yellow. Clypeus and labrum with dense, fine, and weakly impressed ground punctation (punctures separated by 2 x their width); pronotum and elytra with ground punctation fine, weakly impressed and sparser than on head (punctures separated by 3 x their width). Head: Clypeus and labrum shallowly emarginate anteromedially, lateral margins of the labrum bearing setae. Thorax: Prosternum carinate medially, strongly raised, pointing anteriorly and acute. Elevation of mesoventrite with one transversal ridge, elevated medially, lateral sides concave; longitudinal ridge sharp, the point where the two ridges merged rounded and obtuse (e. g., Fig. 10 A, B); elevation flat in lateral view; mesoventrite with triangular shape in ventral view. Metaventrite convex in the median region, pubescent with narrow glabrous patch on the medial and posterolateral area, medial region patch drop-shaped; anterior margin extending to mesoventrite elevation. Metafemora densely covered with hydrofuge pubescence on basal three-quarters. Abdomen: Abdominal ventrites very densely pubescent. Aedeagus (Fig. 7 D) with basal piece nearly the same length of a paramere. Base of the parameres slightly narrower than the base of the median lobe; outer margins sinuate, inner margins depressed in the midlength, depression extending to apex without reaching it; apex of parameres wide and blunt. Median lobe length almost equal to the parameres, wide at basal region, narrowing abruptly in the midlength, apical third slender, narrow, and rounded.	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
DE5FE26820BA5CAAB1C52315B54C204C.taxon	etymology	Etymology. L. M. Gonzalez-Rodriguez names this species in honor of Juan Jose Giraldo Gutierrez in gratitude for the encouragement and support in her career.	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
DE5FE26820BA5CAAB1C52315B54C204C.taxon	distribution	Distribution. Known from several localities in the Brazilian States of Rondonia and Mato Grosso do Sul (Fig. 14).	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
F684E1D524025BCEA17AD5836C0061D6.taxon	description	Fig. 14	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
F684E1D524025BCEA17AD5836C0061D6.taxon	materials_examined	Material examined (2 exs.). Bolivia: Santa Cruz: Amboro National Park, Guarda Parque Mataracu, 21 - 27. xi. 2004, malaise trap, leg. Robertson, Garcia, & Vidaurre (1 female, UMSP). Peru: Cusco: Quispicanchi Province, streams 1 km N Quince Mil, 13 ° 13.335 ' S, 70 ° 46.035 ' W, 730 m, 9. i. 2020, leg. Baca., Slow rivulets & pools w / saturated detritus next to stream, PE 20 - 0109 - 02 A (1 female, SEMC, DNA voucher SLE 2140).	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
FB77835A978C598989315BBB32322C1F.taxon	description	Figs 4 D, 8 B, 10 C, 12 B, 15	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
FB77835A978C598989315BBB32322C1F.taxon	description	Description. Size and form: Body length 1.6 - 1.8 mm. Body form elongate oval, convex in lateral view (Fig. 4 D). Color and punctation: Dorsally bicolor, head mostly brown, frons and middle region of the clypeus dark brown, lateral sides of the clypeus pale brown; pronotum yellow with two small black round spots along posterior margin; elytra dark brown, with a yellow spot on the anterior third of the elytra, lateral and posterior margins of the elytra yellow (Fig. 4 D). Ventrally dark brown; maxillary palps, mouthparts, and antennae yellow (antennal club slightly darker). Pro and meso legs yellow, meta legs pale brown. Clypeus and labrum with dense, fine, and weakly impressed ground punctation (punctures separated by 2 x their width); pronotum and elytra ground punctation fine, weakly impressed and sparser than on head (punctures separated by 3 x their width). Head: Clypeus and labrum shallowly emarginate anteromedially, lateral margins of the labrum bearing setae. Thorax: Prosternum carinate medially, strongly raised, acute and pointing anteriorly. Elevation of mesoventrite with two transversal ridges, elevated medially, lateral sides concave; longitudinal ridge broad anteriorly and sharp posteriorly, the point where the three ridges merged wide and blunt (Fig. 10 C); elevation flat in ventral view; mesoventrite with triangular shape in ventral view. Metaventrite convex in the median region, pubescent with narrow glabrous patch on the medial and posterolateral area; anterior margin extending to mesoventrite elevation. Metafemora with dense hydrofuge pubescence along basal three-quarters of the anterior margin and along basal one-quarter of the posterior margin, then anterior half of posterior margin with sparse setae. Abdomen: Abdominal ventrites very densely pubescent. Aedeagus (Fig. 8 B) with basal piece 0.6 x the length of a paramere. Base of the parameres narrower than the base of the median lobe; outer margin straight along basal two-thirds, then apically slightly convex, inner margins convex along basal two-thirds and then apically straight; basal two-thirds of the parameres broad then apical third narrower; apex of parameres rounded and pointing inwards. Median lobe shorter than the parameres, approximately triangular, with acute apex; gonopore triangular and situated at apex of median lobe.	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
FB77835A978C598989315BBB32322C1F.taxon	etymology	Etymology. The specific name comes from the Latin word insignitus meaning marked and refers to the distinctive yellow spot in the elytra of this species.	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
FB77835A978C598989315BBB32322C1F.taxon	distribution	Distribution. Known only from the type locality in southeastern Venezuela (Fig. 15).	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
92D0546CE5C7510E8DDE602E9471C202.taxon	description	Figs 4 E, 8 G, 13 E, 15	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
92D0546CE5C7510E8DDE602E9471C202.taxon	description	Description. Size and form: Body length 1.6 - 1.9 mm. Body form elongate oval, convex in lateral view (Fig. 4 E). Color and punctation: Dorsally bicolor, head pale brown, pronotum yellow with two small black round spots along posterior margin; elytra dark brown (Fig. 4 E). Ventrally brown; maxillary palps, mouthparts, and antennae yellow (antennal club slightly darker). Pro and meso legs yellow, meta legs pale brown. Clypeus and labrum with dense, coarse, and moderately impressed ground punctation (punctures separated by 2 x their width); elytra and pronotum ground punctation coarse, moderately impressed and less dense than on head (punctures separated by 3 x their width). Head: Clypeus and labrum shallowly emarginate anteromedially, lateral margins of the labrum bearing setae. Thorax: Prosternum carinate medially, strongly raised, acute and pointing anteriorly. Elevation of mesoventrite with two transversal ridges, elevated medially, lateral sides concave; longitudinal ridge broad anteriorly and sharp posteriorly, the point where the three ridges merged wide and blunt (e. g., Fig. 10 C, D); elevation flat in ventral view; mesoventrite with triangular shape in ventral view. Metaventrite convex in the median region, pubescent with narrow glabrous patch on the medial and posterolateral area; anterior margin extending to mesoventrite elevation. Metafemora densely covered with hydrofuge pubescence on basal three-quarters. Abdomen: Abdominal ventrites very densely pubescent. Aedeagus (Fig. 8 G) with basal piece nearly the same length as a paramere. Base of the parameres as wide as the base of the median lobe; outer margins convex, then curved inwards along apex, inner margins convex along basal two-thirds, and concave at apical third; apex of parameres rounded. Median lobe shorter than the parameres, wide at basal region, slightly narrowing in the midlength, apex wide and emarginate medially; gonopore with an oval shape and situated at apex of the median lobe.	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
92D0546CE5C7510E8DDE602E9471C202.taxon	etymology	Etymology. This species is named after the Juma, an indigenous tribe located in the Acua River, in the southern part of the state of Amazonas-Brazil.	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
92D0546CE5C7510E8DDE602E9471C202.taxon	distribution	Distribution. Known only from the Ducke Reserve near Manaus, Brazil (Fig. 15).	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
C6FB4C8FF4BD572D8E23E641A3FBE08A.taxon	description	Figs 2 A-C, 7 A, 11 A, B, 14	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
C6FB4C8FF4BD572D8E23E641A3FBE08A.taxon	materials_examined	Additional material examined (139 exs.). Venezuela: Aragua State: Henri Pittier National Park, Rio Curucuruma, 10 ° 21.070 ' N, 67 ° 34.920 ' W, 11. i. 2006, leg. A. E. Z. Short, waterfall / seep, AS- 06 - 023 (75 exs., MIZA, SEMC, including DNA Voucher MSC 1820); Henri Pittier National Park, Rio Castano Regresiva del Diablo, 10.35669 ° N, 67.60645 ° W, 6. i. 2009, leg. A. E. Z. Short, log in stream, VZ 09 - 0106 - 01 A (5 exs., SEMC); same data except seeps / wet rock, VZ 09 - 0106 - 01 B (19 exs., SEMC); Henri Pittier National Park, Ranch Grande, 10. i. 2006, leg. Short, stream and seep at Toma, AS- 06 - 021, (8 exs., SEMC), same data except 2. vii. 2020, VZ 10 - 0702 - 01 A (1 ex., SEMC, DNA voucher SLE 2111). Barinas State: 13 km NW Barinitas, 8 ° 48.424 ' N, 70 ° 31.139 ' W, 992 m, 24. i. 2012, leg. A. Short & Gustafson, seepage by road, VZ 12 - 0124 - 02 A (22 exs., SEMC, including DNA voucher SLE 2123); same data except small stream pool, VZ 12 - 0124 - 02 B (1 ex., SEMC). Merida State: ca. 2.5 km S. La Azulita, 8 ° 44.335 ' N, 70 ° 37.131 ' W, 842 m, 28. i. 2012, leg. G. T. Gustafson, stream pools, VZ 12 - 0128 - 02 B (1 ex., SEMC); ca. 12 km, SE of Santo Domingo, 8 ° 51.933 ' N, 70 ° 37.131 ' W, 1682 m, 22. i. 2012, leg. Short & Arias, wall seep 1, VZ 12 - 0122 - 03 A (7 exs., SEMC, including DNA voucher SLE 2124). Trujillo State: ca. 3 km NE Laguna Agua Negro, 9 ° 19.371 ' N, 70 ° 9.303 ' W, 1770 m, 21. i. 2009, leg. Short, Garcia & Camacho, small mountain stream w / detritus, VZ 09 - 0121 - 03 x (1 ex., SEMC).	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
C6FB4C8FF4BD572D8E23E641A3FBE08A.taxon	description	Description. Size and form: Body length 1.6 - 1.8 mm. Body form elongate oval, convex in lateral view (Fig. 2 A). Color and punctation: Dorsally black, with lateral margins of the pronotum and elytra reddish brown (Fig. 2 A). Ventrally reddish brown, except for black abdominal ventrites; maxillary palps and antennae yellow (antennal club slightly darker) (Fig. 2 B). Clypeus and labrum with dense, coarse, and moderately impressed ground punctation (punctures separated by 2 x their width); pronotum and elytra ground punctation dense, coarse, and moderately impressed and sparser than on head (punctures separated by 3 x their width). Head: Clypeus and labrum shallowly emarginate anteromedially, lateral margins of the labrum bearing setae. Thorax: Prosternum carinate medially, strongly raised, pointing anteriorly and acute. Elevation of mesoventrite with one transversal ridge, elevated medially, lateral sides concave; longitudinal ridge narrowed anteriorly and broadening posteriorly, the point where the two ridges merged blunt (e. g., Fig. 10 A, B); elevation concave in lateral view; mesoventrite with triangular shape in ventral view. Metaventrite convex in the median region, pubescent with narrow glabrous patch on the medial and posterolateral area; anterior margin extending to mesoventrite elevation. Metafemora with dense hydrofuge pubescence along basal three-quarters of the anterior margin and along basal one-quarter of posterior margin, then apical half of the posterior margin with sparse setae (Fig. 2 B). Abdomen: Abdominal ventrites very densely pubescent. Aedeagus (Fig. 7 A) with basal piece 1.3 x the length of a paramere. Base of the parameres slightly narrower than the base of the median lobe; outer margin sinuate, inner margin nearly straight, tapering along apical third, parameres thin and rounded at apex. Median lobe shorter than the parameres, broad at the base and gradually widening to the apex, with the apex acute.	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
C6FB4C8FF4BD572D8E23E641A3FBE08A.taxon	distribution	Distribution. This species is widespread in the Merida Andes and Coastal Mountains of Venezuela (Fig. 14). Originally described from localities in the Venezuelan states of Barinas and Merida (Spangler 1972, it was later recorded from the state of Aragua (Garcia 2000). Here, we report additional localities in these three states as well as report it from the state of Trujillo for the first time.	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
C7942A8C21C05F87921DE469A5A301AA.taxon	description	Figs 8 F, 15	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
C7942A8C21C05F87921DE469A5A301AA.taxon	materials_examined	Additional material examined (2 exs.). French Guiana: Savane Roche Virginie, near RN 2, 4.1883, - 52.13982, 64 m, Crique Chauve-souris, leg. Short, 10. iii. 2020, solid granite substrate, detritus along margins on granite, FG 20 - 0310 - 01 A (2 exs., SEMC, including DNA vouchers SLE 2337 and 2387).	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
C7942A8C21C05F87921DE469A5A301AA.taxon	description	Description. Size and form: Body length 1.6 - 1.8 mm. Body form elongate oval, convex in lateral view (e. g., Fig. 4 A). Color and punctation: Dorsally bicolor, head bicolor, frons dark brown, clypeus yellow; pronotum yellow with two small black round spots along posterior margin; elytra dark brown, elytra margins paler (e. g., Fig. 4 A). Ventrally dark brown; maxillary palps, mouthparts, and antennae yellow (antennal club slightly darker), legs brown. Clypeus and labrum with dense, fine, and weakly impressed ground punctation (punctures separated by 2 x their width); pronotum and elytra ground punctation fine, weakly impressed and sparser than on head (punctures separated by 3 x their width). Head: Clypeus and labrum shallowly emarginate anteromedially, lateral margins of the labrum bearing setae. Thorax: Prosternum carinate medially, strongly raised, acute and pointing anteriorly. Elevation of mesoventrite with two transversal ridges, elevated medially, lateral sides concave; longitudinal ridge broad anteriorly and sharp posteriorly, the point where the three ridges merged wide and blunt (e. g., Fig. 10 C, D); elevation flat in ventral view; mesoventrite with triangular shape in ventral view. Metaventrite convex in the median region, pubescent with narrow glabrous patch on the medial and posterolateral area; anterior margin extending to mesoventrite elevation. Metafemora with dense hydrofuge pubescence along basal three-quarters of the anterior margin and along basal one-quarter of the posterior margin, then apical half of the posterior margin with sparse setae. Abdomen: abdominal ventrites very densely pubescent. Aedeagus (Fig. 8 F) with basal piece 0.8 x the length of the paramere. Base of the parameres slightly narrower than the base of the median lobe, inner margins straight, then convex reaching the apex, outer margins convex, and rounded apex pointing inwards. Median lobe shorter than the parameres, approximately rectangular with apex blunt; gonopore situated at the apex of the median lobe.	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
C7942A8C21C05F87921DE469A5A301AA.taxon	distribution	Distribution. This species is only known from a few localities in French Guiana (Fig. 15).	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
ED05D427A3D45C48951553F49EDB5A8B.taxon	description	Figs 2 D-F, 7 B, 14	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
ED05D427A3D45C48951553F49EDB5A8B.taxon	description	Description. Size and form: Body length 1.8 mm. Body form elongate oval, convex in lateral view (Fig. 2 D). Color and punctation: Dorsally yellow, head mostly yellow, frons pale brown; pronotum with two small black round spots along posterior margin (Fig. 2 D). Ventrally brown; maxillary palps, mouthparts, antennae, pro and meso legs yellow, meta legs pale brown (Fig. 2 E). Clypeus and labrum with dense, fine, and weakly impressed ground punctation (punctures separated by 2 x their width); pronotum and elytra ground punctation fine, weakly impressed and sparser than on head (punctures separated by 3 x their width). Head: Clypeus and labrum shallowly emarginate anteromedially, lateral margins of the labrum bearing setae. Thorax: Prosternum carinate medially, strongly raised, pointing anteriorly and acute. Elevation of mesoventrite with one transversal ridge, elevated medially, lateral sides concave; longitudinal ridge sharp, the point where the two ridges merged rounded and obtuse (e. g., Fig. 10 A, B); elevation concave in lateral view; mesoventrite with triangular shape in ventral view. Metaventrite convex in the median region, pubescent with narrow glabrous patch on the medial and posterolateral area; anterior margin extending to mesoventrite elevation. Metafemora densely covered with hydrofuge pubescence on basal three-quarters (Fig. 2 E). Abdomen: Abdominal ventrites very densely pubescent. Aedeagus (Fig. 7 B) with basal piece 1.1 x the length of a paramere. Base of the parameres broader than the base of the median lobe; outer margin slightly convex along basal two-thirds, then slightly sinuate apically, inner margins nearly straight along basal two-thirds and then sinuate apically; apex of parameres rounded. Median lobe shorter than the parameres, approximately triangular, with acute apex.	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
ED05D427A3D45C48951553F49EDB5A8B.taxon	distribution	Distribution. Only known from the type locality in Mexico (Fig. 14).	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
26E62B4CA6645394A7736CD2A7E2457E.taxon	description	Figs 4 F, 9 E, 15	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
26E62B4CA6645394A7736CD2A7E2457E.taxon	description	Description. Size and form: Body length 1.7 - 1.9 mm. Body form elongate oval, convex in lateral view (Fig. 4 F). Color and punctation: Dorsally reddish brown, head and pronotum pale reddish yellow; pronotum with two small black round spots along posterior margin; elytra dark reddish brown (Fig. 4 F). Ventrally reddish brown; maxillary palps and mouthparts light yellow reddish, and antennae yellow (antennal club slightly darker). Pro and meso legs yellow, meta legs pale reddish brown. Clypeus and labrum with dense, fine, and weakly impressed ground punctation (punctures separated by 2 x their width); pronotum and elytra ground punctation fine, weakly impressed and sparser than on head (punctures separated by 3 x their width). Head: Clypeus and labrum shallowly emarginate anteromedially, lateral margins of the labrum bearing setae. Thorax: Prosternum carinate medially, strongly raised, acute and pointing anteriorly. Elevation of mesoventrite with two transversal ridges, elevated medially, lateral sides concave; longitudinal ridge broad anteriorly and sharp posteriorly, the point where the three ridges merged wide and blunt (e. g., Fig. 10 C, D); elevation slightly convex in ventral view; mesoventrite with triangular shape in ventral view. Metaventrite convex in the median region, pubescent with narrow glabrous patch on the medial and posterolateral area; anterior margin extending to mesoventrite elevation. Metafemora with dense hydrofuge pubescence along basal three-quarters of the anterior margin and sparse pubescence along the apical posterior margin. Abdomen: Abdominal ventrites very densely pubescent. Aedeagus (Fig. 9 E) with basal piece 0.5 x the length of a paramere. Base of the parameres narrower than the base of the median lobe; parameres tubular shape, outer margins sinuate and covered by lanceolate appendages that cover three-quarters of the margin; inner margins sinuate, apex of the parameres rounded. Median lobe shorter than the parameres, wide at basal region, narrowing apically, apex rounded and wide; gonopore crown-shaped and situated at apex of median lobe.	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
26E62B4CA6645394A7736CD2A7E2457E.taxon	etymology	Etymology. The species name is derived from the Latin word parvus meaning little or small in reference to the small aedeagus size of this species.	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
26E62B4CA6645394A7736CD2A7E2457E.taxon	distribution	Distribution. Only known from the type locality in southern Suriname (Fig. 15).	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
468569EEC5E158F99A09D8B66D975E7C.taxon	description	Figs 4 B, 8 D, 12 C, 15	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
468569EEC5E158F99A09D8B66D975E7C.taxon	description	Description. Size and form: Body length 1.6 - 1.8 mm. Body form elongate oval, convex in lateral view (Fig. 4 B). Color and punctation: Dorsally bicolor, head brown, frons dark brown, clypeus pale brown; pronotum yellow with two small black round spots along posterior margin; elytra dark brown, elytra margins paler (Fig. 4 B). Ventrally brown; maxillary palps, mouthparts, antennae, and legs yellow. Clypeus and labrum with dense, fine, and weakly impressed ground punctation (punctures separated by 2 x their width); pronotum and elytra ground punctation fine, weakly impressed and sparser than on head (punctures separated by 3 x their width). Head: Clypeus and labrum shallowly emarginate anteromedially, lateral margins of the labrum bearing setae. Thorax: Prosternum carinate medially, strongly raised, acute and pointing anteriorly. Elevation of mesoventrite with two transversal ridges, elevated medially, lateral sides concave; longitudinal ridge broad anteriorly and sharp posteriorly, the point where the three ridges merged wide and blunt (e. g., Fig. 10 C, D); elevation flat in ventral view; mesoventrite with triangular shape in ventral view. Metaventrite convex in the median region, pubescent with narrow glabrous patch on the medial and posterolateral area; anterior margin extending to mesoventrite elevation. Metafemora with dense hydrofuge pubescence along basal three-quarters of the anterior margin and along basal one-quarter of the posterior margin, then apical half of the posterior margin with sparse setae. Abdomen: Abdominal ventrites very densely pubescent. Aedeagus (Fig. 4 B) with basal piece 0.7 x the length of a paramere. Base of the parameres narrower than the base of the median lobe; outer margins straight along basal two-thirds, then apically slightly convex, inner margins straight along basal two-thirds and then convex and tapering apically; apex of parameres rounded and pointing inwards. Median lobe shorter than the parameres, approximately triangular, gradually narrowing from the base, broad and rounded apex; gonopore oval-shaped and situated at apex of median lobe.	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
468569EEC5E158F99A09D8B66D975E7C.taxon	etymology	Etymology. This species is named after the Patamona, an indigenous tribe located in the mountainous region from which this species is known.	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
468569EEC5E158F99A09D8B66D975E7C.taxon	distribution	Distribution. Known from several closely situated localities in western Guyana (Fig. 15).	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
5F0EB05A0AD25303AF591B6F6E87EF8E.taxon	materials_examined	Additional material examined (173 exs.). Costa Rica: Cartago Province: Tapanti National Park, Building by Rio Villegas, 29. v. 2006, leg. A. E. Z. Short, HG-vapor light, AS- 06 - 066 (5 exs., SEMC, INBio, including DNA voucher SLE 2397). Panama: Panama province: Barrio Colorado, 9 ° 11 ' N, 79 ° 51 ' W, 40 m, 22 - 25 - VI- 2000, leg. S. Chatzimanolis, flight intercept trap, PAN 1 C 00 022 (3 exs., SEMC); same data except PAN 1 C 00 024 (5 exs., SEMC); same data except PAN 1 C 00 025 (2 exs., SEMC); same data except PAN 1 C 00 033 (1 ex., SEMC); same data except PAN 1 C 00 0234 (1 ex., SEMC); same data except PAN 1 C 00 014 (1 ex., SEMC); same data except 07 - VI- 1994, leg. D. Banks (2 exs., SEMC); same data except 08 - VIII- 1994, leg. D. Banks (1 ex., SEMC), same data except 08 - VIII- 1994, leg. D. Banks (1 ex., SEMC); same data except 04 - VIII- 1994, leg. D. Banks (1 ex., SEMC); same data except 01 - VIII- 1994, leg. D. Banks (1 ex., SEMC); Old plantation Rd. 6.9 km S Gamboa, 09 ° 05 ' N, 79 ° 40 ' W, 80 m, 04 - 07 - VI- 1995, leg. J, Ashe & R. Brooks, # 137 flight intercept trap (1 ex., SEMC), same data except 07 - 22 - VI- 1995, # 266 flight intercept trap (1 ex., SEMC); Colon, Parque Nacional Soberania, Pipeline Rd km 6.1, 09 ° 07 ' N, 79 ° 45 ' W, 40 m, 07 - 21 - VI- 1995, leg. J, Ashe & R. Brooks, # 265 flight intercept trap (2 exs., SEMC); Colon, Escobal & Pina Rds, 14 km N jct. 02 - 11 - VI- 1996, leg. J, Ashe & R. Brooks flight intercept trap PAN 1 AB 96 181 B (1 ex., SEMC). Venezuela: Aragua State: Henri Pittier Natural Park Rio Cumboto, 10.39376 ° N, 67.79597 ° W, 130 m, 4. i. 2009, leg. Short, Garcia & Miller, river side pools, VZ 09 - 0104 - 02 B (18 exs., SEMC, including DNA voucher SLE 2381); Rio La Trilla 10.37319 ° N, 67.74250 ° W, 295 m, leg. Short, Garcia & Miller, pools, VZ 09 - 0104 - 01 A (1 ex., SEMC). Barinas State: Rio Santa Barbara, E. Santa Barbara. 7 ° 50.028 N, 71 ° 11.188 W, 177 m, 26. i. 2012, leg. Short, Arias, Gustafson, sandy sidepool in floodplain, VZ 12 - 0126 - 01 B, (1 ex., SEMC). Portuguesa State: Trib. of Rio Guanare, S. Biscucuy, 9 ° 14.457 ' N, 69 ° 55.994 ' W, 370 m, 19. i. 2009, leg. Short, Garcia & Miller, gravel stream, VZ 09 - 0119 - 03 X (15 exs., SEMC, including DNA vouchers SLE 2391 and SLE 2392). Zulia State: Perija Natural Park Tukuko: Rio Manantial, 9 ° 50.490 ' N, 72 ° 49.310 ' W, 270 m, 29. i. 2009, leg. Short, Garcia & Camacho, gravel margin, VZ 09 - 0129 - 01 A (91 exs., MIZA, SEMC, including DNA vouchers SLE 1112 and SLE 2371); same data except 29. i. 2009, leg. Short, Garcia & Miller, detrital pool, VZ 09 - 0129 - 01 B (1 ex., SEMC); same data except 22. ix. 2007, leg. A. E. Z. Short, rock pools / margin, AS- 07 - 020 b (8 exs., SEMC); Toromo, 10 ° 03.058 ' N, 72 ° 49.974 ' W, 435 m, 31. xii. 2005, leg. A. E. Z. Short, small stream and seep, AS- 06 - 001 (6 exs., SEMC); same data except 28. i. 2009, detrital pool, VZ 09 - 0128 - 01 A (3 exs., SEMC).	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
5F0EB05A0AD25303AF591B6F6E87EF8E.taxon	description	Description. Size and form: Body length 1.6 - 1.9 mm. Body form elongate oval, strongly convex in lateral view (Fig. 3 A). Color and punctation: Dorsally yellow, head mostly pale brown or yellow, frons brown or dark brown; pronotum paler than elytra, with two small black round spots along posterior margin (Fig. 3 A, D, G). Ventrally brown; maxillary palps, mouthparts, antennae yellow (antennal club slightly darker), legs pale brown (Fig. 3 B, E, H). Clypeus and labrum with dense, fine, and weakly ground punctation (punctures separated by 2 x their width); pronotum and elytra ground punctation fine, weakly impressed and sparser than on head (punctures separated by 3 x their width). Head: Clypeus and labrum shallowly emarginate anteromedially, lateral margins of the labrum bearing setae. Thorax: Prosternum carinate medially, strongly raised, pointing anteriorly and acute. Elevation of mesoventrite with one transversal ridge, elevated medially, lateral sides concave; longitudinal ridge sharp and broadening posteriorly almost to the end, the point where the two ridges merged rounded and obtuse (e. g., Fig. 10 A, B); elevation concave in lateral view; mesoventrite with triangular shape in ventral view. Metaventrite convex in the median region, pubescent with narrow glabrous patch on the medial and posterolateral area; anterior margin extending to mesoventrite elevation. Metafemora densely covered with hydrofuge pubescence on basal three-quarters (Fig. 3 B, E, H). Abdomen: Abdominal ventrites very densely pubescent. Aedeagus (Fig. 6 A-I) basal piece 1.2 x the length of a paramere. Parameres broad, base wider than the base of the median lobe, outer and inner margins convex, pinched at the apex, broad and rounded apex. Length of the median lobe can vary (median lobe as long as the parameres (Fig. 6 F, I), slightly shorter (Fig. 6 A, B) or longer than the parameres (Fig. 6 H), median lobe with triangular shape, wide at base and gradually tapering to apical third, with rounded apex.	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
5F0EB05A0AD25303AF591B6F6E87EF8E.taxon	distribution	Distribution. Known from Guatemala, Costa Rica, Panama, and Venezuela (Fig. 14).	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
404A20425109524CB4D7B2D2F84D3860.taxon	description	Figs 4 K, 9 F, 14	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
404A20425109524CB4D7B2D2F84D3860.taxon	description	Description. Size and form: Body length 1.6 mm. Body form elongate oval, convex in lateral view (Fig. 4 K). Color and punctation: Dorsally yellow, head mostly yellow and frons pale brown; pronotum with two small black round spots along posterior margin (Fig. 4 K). Ventrally brown; maxillary palps, mouthparts, antennae (antennal club slightly darker) and legs yellow. Clypeus and labrum with dense, fine, and weakly impressed ground punctation (punctures separated by 2 x their width); pronotum and elytra ground punctation fine, weakly impressed and sparser than on head (punctures separated by 3 x their width). Head: Clypeus and labrum shallowly emarginate anteromedially, lateral margins of the labrum bearing setae. Thorax: Prosternum carinate medially, strongly raised, pointing anteriorly and acute. Elevation of mesoventrite with one transversal ridge, elevated medially, lateral sides concave; longitudinal ridge sharp, the point where the two ridges merged acute (e. g., Fig. 10 B); elevation flat in lateral view; mesoventrite with triangular shape in ventral view. Metaventrite convex in the median region, pubescent with narrow glabrous patch on the medial and posterolateral area, medial region patch drop-shaped; anterior margin extending to mesoventrite elevation. Metafemora densely covered with hydrofuge pubescence on basal three-quarters. Abdomen: Abdominal ventrites very densely pubescent. Aedeagus (Fig. 9 F) with basal piece 0.7 x the length of a paramere. Base of the parameres wider than the base of the median lobe; outer margins straight along basal two-thirds, then apically sinuate, inner margins straight along basal two-thirds and then convex and tapering apically; apex of parameres rounded and pointing outwards. Median lobe much shorter than the parameres, basal half rectangular, apical half narrowing abruptly, apex rounded.	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
404A20425109524CB4D7B2D2F84D3860.taxon	etymology	Etymology. The species is named after Peru, the country where it was collected, as well as for being the first species described for the genus in this country.	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
404A20425109524CB4D7B2D2F84D3860.taxon	distribution	Distribution. Known only from the type locality in Peru (Fig. 14).	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
33F55D59C29C52838EC6CDB4E3BD0670.taxon	description	Figs 8 E, 13 F, 15	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
33F55D59C29C52838EC6CDB4E3BD0670.taxon	description	Description. Size and form: Body length 1.8 mm. Body form elongate oval, moderately convex in lateral view. Color and punctation: Dorsally bicolor, head mostly brown, frons brown, clypeus pale brown; pronotum yellow with two small black round spots along posterior margin; elytra dark brown. Ventrally brown; maxillary palps, mouthparts, antennae (antennal club slightly darker), legs brown. Clypeus and labrum with dense, fine, and weakly impressed ground punctation (punctures separated by 2 x their width); pronotum and elytra ground punctation fine, weakly impressed, and sparser than on head (punctures separated by 3 x their width). Head: Clypeus and labrum shallowly emarginate anteromedially, lateral margins of the labrum bearing setae. Thorax: Prosternum carinate medially, strongly raised, acute and pointing anteriorly. Elevation of mesoventrite with two transversal ridges, elevated medially, lateral sides concave; longitudinal ridge broad anteriorly and sharp posteriorly, the point where the three ridges merged wide and blunt (e. g., Fig. 10 C, D); elevation flat in ventral view; mesoventrite with triangular shape in ventral view. Metaventrite convex in the median region, pubescent with narrow glabrous patch on the medial and posterolateral area; anterior margin extending to mesoventrite elevation. Metafemora with dense hydrofuge pubescence along basal three-quarters of the anterior margin and along basal one-quarter of posterior margin, then apical half of the posterior margin with sparse setae. Abdomen: Abdominal ventrites very densely pubescent. Aedeagus (Fig. 8 E) with basal piece 0.7 x the length of a paramere. Base of the parameres nearly the same as the base of the median lobe; outer margins convex, inner margins nearly straight; apex of parameres broad, rounded and slightly pointing inwards. Median lobe shorter than the parameres, wide at basal region, narrowing in basal third, margins straight and apex rounded and nearly notched; gonopore with an oval shape and covering approximately two-thirds of the length of the median lobe.	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
33F55D59C29C52838EC6CDB4E3BD0670.taxon	etymology	Etymology. The specific name comes from the Latin word retusus meaning rounded and notched, after the form of the apex of the median lobe of the aedeagus.	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
33F55D59C29C52838EC6CDB4E3BD0670.taxon	distribution	Distribution. This species is only known from the type locality in western Guyana (Fig. 15).	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
FC97C583DD215ACB89CCE8A5BE6E3D01.taxon	description	Figs 2 G-I, 9 A-C, 14	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
FC97C583DD215ACB89CCE8A5BE6E3D01.taxon	materials_examined	Additional material examined. Trinidad: Guanapo State: 4.1 km up Guanapo Valley, trib of Guanapo River, 460 ft, 11 - VII- 2005 (1 ex., SEMC); Verdant vale, Arima River, 10 ° 42 ' N, 61 ° 18 ' W, 570 ft, 9 - VII- 2005 (1 ex., SEMC). Venezuela: Aragua: Rancho Grande Biol. Stn. 1150 m, 10 ° 21 ' N, 67 ° 41 ' W, 25 - 28 II 1995, S. Marshall, yellow pan trap (1 ex., SEMC).	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
FC97C583DD215ACB89CCE8A5BE6E3D01.taxon	description	Description. Size and form: Body length 1.8 - 1.9 mm. Body form elongate oval, moderately convex in lateral view (Fig. 2 G). Color and punctation: Dorsally bicolor, head mostly brown, frons brown, medially region of the clypeus pale brown with lateral margins yellow; pronotum yellow with two small black round spots along posterior margin; elytra yellow except by a brown wideband on the anterior third of the elytra (Fig. 2 G). Ventrally brown; maxillary palps, mouthparts, antennae, and legs yellow (antennal club slightly darker) (Fig. 2 H). Clypeus and labrum with dense, fine, and weakly impressed ground punctation (punctures separated by 5 x their width); pronotum and elytra ground punctation fine, weakly impressed and sparser than on head (punctures separated by 3 x their width). Head: Clypeus and labrum shallowly emarginate anteromedially, lateral margins of the labrum bearing setae. Thorax: Prosternum carinate medially, strongly raised, pointing anteriorly and acute. Elevation of mesoventrite with one transversal ridge, elevated medially, lateral sides concave; longitudinal ridge narrowed anteriorly and broadening posteriorly, the point where the two ridges merged acute (e. g., Fig. 10 B); elevation concave in lateral view; mesoventrite with triangular shape in ventral view. Metaventrite convex in the median region, pubescent with narrow glabrous patch on the medial and posterolateral area; anterior margin extending to mesoventrite elevation. Metafemora with dense hydrofuge pubescence along basal three-quarters of the anterior margin and along basal half of the posterior margin. Abdomen: Abdominal ventrites very densely pubescent. Aedeagus (Fig. 9 A-C) basal piece 0.4 x the length of a paramere; broad parameres, base of the parameres much wider than the base of the median lobe, base of the parameres with two accessories with ovate shape, outer margins of parameres strongly sinuate, inner margins slightly sinuate, with membranous acuminate apex, bending outwards; median lobe shorter than the parameres, approximately triangular, with apex rounded.	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
FC97C583DD215ACB89CCE8A5BE6E3D01.taxon	distribution	Distribution. Originally described from the Venezuelan states of Aragua and Barinas (Spangler 1972), it was later reported from the states of Trujillo and Falcon (Garcia 2000). Here we report it for the first time from Trinidad (Fig. 14).	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
5BD081ADE949546CA1603FEFBEAAACE9.taxon	materials_examined	Additional material examined (362 exs.). Brazil: Roraima State: Serra do Tepequem, 3 ° 47.334 ' N, 61 ° 42.570 ' W, 14. i. 2018, leg. Short, Benetti, & Santana, small forested stream, BR 18 - 0114 - 02 A (1 ex., SEMC, DNA voucher SLE 2377); Caroebe, Rio Caroebe, ca. 13 km NE of Caroebe, 00 ° 54.786 ' N, 59 ° 34.397 ' W, 150 m, 17. i. 2018, leg. Short and Benetti, margins of sandy river, BR 18 - 0117 - 04 A (26 exs., INPA, SEMC, including DNA voucher SLE 2330); Rio Jatapu nr. Usina de Jatapu, 00 ° 50.939 ' N, 59 ° 18.262 ' W, 145 m, 17. i. 2018, leg. A. Short, marginal pools of river, BR 18 - 0117 - 02 A (1 ex., SEMC); Amajari, Rio Cocal, 3 ° 44.135 ' N, 61 ° 43.542 ' W, 237 m, 15. i. 2018, leg. A. Short, clearwater creek, BR 18 - 0115 - 01 A (6 exs., SEMC, including DNA voucher SLE 2374). French Guiana: St. Laurent du Maroni ca. 15 km SW, Crique des cascades, 5.34662, - 54.10539, 17 m, 4. iii. 2020; leg. Short and Neff, stream margin, FG 20 - 0304 - 02 A (1 ex., SEMC); same data except leg. Short, leaf packs on rock, FG 20 - 0304 - 02 B (2 exs., SEMC, including DNA voucher SLE 2339); same data except small rock pools in granite FG 20 - 0304 - 02 C (1 ex., SEMC); Carbet ONF Grillon, Piste Belizon, Crique Grillon, 4.28219, - 52.45163, 65 m, 11. iii. 2020; leg. Short, rock pools by waterfall, FG 20 - 0311 - 01 C (2 exs., SEMC, including DNA voucher SLE 2383); Crique a l'Est above Crique Eau Chire, 3.66383, - 53.22193, 156 m, 10. xi. 2016, leg. D. Post (1 ex., SEMC, DNA voucher SLE 2113). Guyana: Region 6: Upper Berbice, ca. 1 km, W basecamp, 4 ° 09.143 ' N, 58 ° 11.207 ' W, 170 m, 22. ix. 2014, leg. Short, Salisbury, La Cruz, margins of creek, GY 14 - 0921 - 03 H (6 exs., SEMC); Upper Berbice, Basecamp 1, creek next to basecamp (upstream), 4.154817, - 58.178616, 24. ix. 2014, leg. Short, creek margins, GY 14 - 0924 - 01 A (1 ex., SEMC, DNA voucher SLE 2108); Upper Berbice, Basecamp 1, 4 ° 09.289 ' N, 58 ° 10.717 ' W, 96 m, 22. ix. 2014, leg. Short, Salisbury, La Cruz, GY 14 - 0924 - 01 A (2 ex., SEMC). Region 8: Upper Potaro Camp I (ca. 7 km NW Chenapau), 5 ° 0.673 ' N, 59 ° 38.358 ' W, 500 m, 14. iii. 2014, leg. Short, Salisbury, La Cruz, clear water creek rapids, GY 14 - 0314 - 01 A (7 exs., SEMC); same data except 5 ° 0.660 ' N, 59 ° 38.283 ' W, 484 m, 11. iii. 2014, leg. Short, Baca, Salisbury, La Cruz, clear water creek rapids, GY 14 - 0311 - 04 A (6 exs., SEMC, including DNA voucher SLE 2368); same data except Potaro margin trail, 5 ° 0.571 ' N, 59 ° 38.202 ' W, 524 m, 11. iii. 2014, leg. Short and Baca, leaf packs in rocky stream, GY 14 - 0311 - 05 A (1 ex., SEMC). Region 9: Kusad, Mts., Mokoro Creek, 2 ° 48.531 ' N, 59 ° 51.900 ' W, 170 m, 27. x. 2013, leg. Short, Isaacs and Salisbury, main seepage area, GY 13 - 1027 - 03 B (2 exs., SEMC); same data except basecamp, leg. A. Short and Washington, on wet rocks, GY 13 - 1024 - 03 C (4 exs., SEMC); same data except 24. x. 2013, leg. Salisbury, small rock pool with detritus, GY 13 - 1024 - 03 A (1 ex., SEMC); Kusad, Mts., basecamp area, 2 ° 48.588 ' N, 59 ° 51.931 ' W, 194 m, 23. x. 2013, leg. A. Short, leaf packs in flow of creek, GY 13 - 1023 - 02 B (9 exs., CDBG, SEMC, including DNA voucher SLE 2335); Kusad, Mts., Taraara Wao, 2 ° 47.417 ' N, 59 ° 53.986 ' W, 113 m, 28. x. 2013, leg. Short, Isaacs and Salisbury, margin and isolated side pools, GY 13 - 1028 - 01 A (5 exs., SEMC, including DNA voucher SLE 2107); N. Parabara, creek by basecamp (Bototo wau creek), 2 ° 10.908 ' N, 59 ° 20.306 ' W, leg. Short, 31. x. 2013, stream margins, GY 13 - 1031 - 01 A (1 ex., SEMC, DNA voucher SLE 2380). Region Mazaruni-Potaro: Takutu Mountains, 6 ° 15 ' N, 59 ° 5 ' W, 2 - 14. xii. 1983, leg. P. D. Perkins, stream ex. leaf packs and twigs (8 exs., SEMC) Suriname: Brokopondo District: Brownsberg Nature Park, near Capaci House, 4 ° 56.934 ' N, 55 ° 10.825 ' W, 460 m, 17. iii. 2017, leg. A. Short, small stream, SR 17 - 0317 - 01 A (1 ex., SEMC); Leo Val / lrene Val return loop trail, 4.95069 ' N, - 55.18599 ' W, 470 m, 18. iii. 2017, leg. Baca and Johnson, stream, SR 17 - 0318 - 01 B (1 ex., SEMC); Leo Val, 4 ° 57 ' 16.08 " N, - 55 ° 11 ' 26.82 " W, 317 m, 19. iii. 2017, leg. Short, leaf packs / detritus from behind waterfall, SR 17 - 0319 - 01 C (5 exs., SEMC); same data except SR 17 - 0323 - 01 C (6 exs., SEMC); same data except 23. iii. 2017, leg. Baca, submerged woody debris, SR 17 - 0323 - 01 A (1 ex., SEMC); same data except side pools in creek, SR 17 - 0323 - 01 D (1 ex., SEMC); Witti Kreek, 4 ° 55.674 ' N, 55 ° 09.874 ' W, 84 m, 21. iii. 2017, leg. Short and Baca, small side stream, SR 17 - 0321 - 01 D (2 exs., SEMC, including DNA voucher SLE 2121); Brownsberg Nature Park, Mazaroni Val, 04 ° 56.351 ' N, 55 ° 12.108 ' W, 394 m, 5. viii. 2012, leg. Short, Maier, Mcintosh, forested waterfall and stream, SR 12 - 0805 - 01 A (22 exs., NZCS, SEMC); Brownsberg Nature Park, 04 ° 57.268 ' N, 55 ° 11.447 ' W, 317 m, 4. viii. 2012, leg. Short, Maier, Mcintosh, forested waterfall and stream, SR 12 - 0804 - 02 A (21 exs., SEMC); Brownsberg Nature Park, 04 ° 56.871 ' N, 55 ° 10.911 ' W, 462 m, 4. viii. 2012, leg. Short, Maier, McIntosh, SR 12 - 0804 - 01 A (2 exs., SEMC including SLE 505 and SLE 506). Sipaliwini District: Kabalebo Nature Resort: Bwkw rapids, 4.40041 ° N, 57.24658 ° W, 90 m, 9 - 10.9 iii. 2019, leg. Short and class, large isolated muddy pool by river, SR 19 - 0309 - 02 A (1 ex., SEMC); Moi Moi creek, 4.42313 ° N, 57.19198 ° W, 104 m, 10 - 14. iii. 2019, leg. Short and class, rock and detrital pools along creek, SR 19 - 0310 - 01 A (3 exs., SEMC); same data except leg. Short and Baca, small seeps, SR 19 - 0310 - 01 F (10 exs., SEMC); same data except leg. Short, margin of detrital pool in drying creekbed, SR 19 - 0310 - 01 G (5 exs., SEMC); same data except leg. Short, Baca and class, SR 19 - 0310 - 01 J (1 ex., SEMC); same data except leg. Baca, margin of stream pool with root mats, SR 19 - 0310 - 01 L (5 exs., SEMC); same data except leg. Short and class, SR 19 - 0310 - 01 M (10 exs., SEMC, including DNA voucher SLE 1799); same data except Leg. S. Baca, upstream riparian habitats, SR 19 - 0310 - 01 N (2 exs., SEMC); Sand Crk, 4.38476 ° N, 57.24636 ° W, 72 m, 13 - 15. iii. 2019, leg. Short and Baca, upper marginal pool along river, SR 19 - 0313 - 01 D (2 exs., SEMC); Charlie Falls, 4.38302 ° N, 57.21161 ° W, 174 m, 11. iii. 2019, leg. Short and class, rock pools in creekbed, SR 19 - 0311 - 01 A (39 exs., NZCS, SEMC, including DNA voucher SLE 1811); same data except leg. Short, seepage, SR 19 - 0311 - 01 B (7 exs., SEMC); Sipaliwini Savanna Nature Res. 4 - Brothers Mts, 2 ° 00.342 ' N, 55 ° 58.149 ' W, 337 m, 31. iii. 2017, leg. Short and Baca, clear water stream sandy with detritus, SR 17 - 0331 - 01 B (31 exs., SEMC, including DNA Voucher SLE 2114); same data except leg. Baca, small rocky creek, SR 17 - 0331 - 01 A (1 ex., SEMC); Raleighvallen Nature Reserve, Fungu Island, 4 ° 43.459 ' N, 56 ° 12.658 ' W, 30 m, 14. iii. 2016, leg. A. Short, isolated river margin pools, rocky bottom, SR 16 - 0314 - 01 E (1 ex., SEMC); Coppename Rvr-Voltzberg trail, 15. iii. 2016, leg. A. Short, small sandy stream, SR 16 - 0315 - 02 A (1 ex., SEMC); Lolopaise Area, 4 ° 42.48 ' N, 56 ° 13.15908 ' W, 24 m, 18. iii. 2016, leg. Short et al., intermittent stream margins, flotation, SR 16 - 0318 - 01 D (1 ex., SEMC); Brownsberg Nature Park, 04 ° 56.871 ' N, 55 ° 10.911 ' W, 462 m, 4. viii. 2012, leg. Short, Maier, Mcintosh, forested stream with lots of detritus, SR 12 - 0804 - 01 A (8 exs., SEMC); Raleighfallen Nature Reserve, trail to Raleighfallen, 04 ° 42.480 ' N, 56 ° 13.159 ' W, 24 m, 27. vii. 2012, leg. Short, Mcintosh and Kadosoe, creek margins, SR 12 - 0727 - 03 A (22 exs., SEMC, including DNA voucher SLE 2394); same data except leg. C. Mcintosh, detrital pools near creek in forest, SR 12 - 0727 - 03 D (1 ex., SEMC); Raleighvallen Nature Reserve Voltzberg Station, 04 ° 40.910 ' N, 56 ° 11.138 ' W, 78 m, 9. vii. 2012, leg. Short, Maier, McIntosh, and Kadosoe, stream margins, SR 12 - 0729 - 02 A (1 ex., SEMC, DNA voucher SLE 2389); Raleighfallen Nature Reserve, Voltzberg trail, 04 ° 40.910 ' N, 56 ° 11.138 ' W, 78 m, 30. vii. 2012, leg. C. Maier and V. Kadosoe, margin of stream, SR 12 - 0730 - 01 A (1 ex., SEMC); Raleighfallen Nature Reserve, Fungu island, 04 ° 43.459 ' N, 56 ° 12.658 ' W, 30 m, 1. viii. 2012, leg. Short, Maier, Mcintosh and Kadosoe, small creek, SR 12 - 0801 - 01 B (2 exs., SEMC); Camp 2, on Sipaliwini District, CI-RAP Survey, 2 ° 10.973 ' N, 56 ° 47.235 ' W, 210 m, 29 - 30. viii. 2010, leg. Short and Kadosoe, Inselberg, SR 10 - 0829 - 01 A (2 exs., SEMC); Camp 1, Upper Palumeu, CI-RAP Survey, 2.47700 ° N, 55.62941 ° W, 275 m, 11. iii. 2012, leg. Short and Kadosoe, around waterfall, SR 12 - 0311 - 03 A (2 exs., SEMC); Camp 3, Wehepai, 2 ° 21.776 ' N, 56 ° 41.861 ' W, 237 m, 4 - 6. ix. 2010, leg. Short and Kadosoe, sandy forest creek, SR 10 - 0904 - 01 A (36 exs., SEMC, including DNA voucher SLE 2375). Venezuela: Bolivar State: Piedra de la Virgen, 6 ° 5 ' 14.1 " N, 61 ° 23 ' 55.8 " W, 400 m, 31. vii. 2008, leg. A. Short M. Garcia, L. Joly, small forest stream, AS- 08 - 056 (7 exs., SEMC, including DNA voucher SLE 2378). Amazonas State: Tobogan de la Selva, 5 ° 23.207 ' N, 67 ° 36.922 ' W, 125 m, 14. i. 2009, leg. A. Short, clumps of wet leaves on rock, VZ 09 - 0114 - 01 D (6 exs., MIZA, SEMC, including DNA voucher SLE 2366).	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
5BD081ADE949546CA1603FEFBEAAACE9.taxon	description	Description. Size and form: Body length 1.6 - 1.8 mm. Body form elongate oval, moderately convex in lateral view (e. g., Fig. 2 D). Color and punctation: Dorsally yellow to pale brown, head mostly yellow and frons pale brown; pronotum paler than elytra, with two small black round spots along posterior margin (e. g., Fig. 2 D). Ventrally brown; maxillary palps and antennae yellow (antennal club slightly darker), mouth parts and legs pale brown. Clypeus and labrum with dense, fine, and weakly impressed ground punctation (punctures separated by 1 - 2 x their width); pronotum and elytra with ground punctation fine, weakly impressed and sparser than on head (punctures separated by 3 x their width). Head: Clypeus and labrum shallowly emarginate anteromedially, lateral margins of the labrum bearing setae. Thorax: Prosternum carinate medially, strongly raised, pointing anteriorly and acute. Elevation of mesoventrite with one transversal ridge, elevated medially, lateral sides concave; longitudinal ridge sharp, the point where the two ridges merged rounded and obtuse (Fig. 10 A); elevation flat in lateral view; mesoventrite with triangular shape in ventral view. Metaventrite convex in the median region, pubescent with narrow glabrous patch on the medial and posterolateral area, medial region patch drop-shaped; anterior margin extending to mesoventrite elevation. Metafemora densely covered with hydrofuge pubescence on basal three-quarters (e. g., Fig. 2 E). Abdomen: Abdominal ventrites very densely pubescent. Aedeagus (Fig. 5 A) basal piece nearly the same length of a paramere. Base of the parameres slightly narrower than the base of the median lobe; outer margins sinuate, tapering almost reaching the apex, inner margins sinuate, apex of the parameres slightly bending outwards, rounded in the outer margin with an indentation in the inner margin (the depth of the indentation varies, Fig. 5 A-J). Median lobe nearly as long as the parameres or a bit longer, broad at the base and gradually narrowing to the apex, apex of the median lobe rounded.	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
5BD081ADE949546CA1603FEFBEAAACE9.taxon	distribution	Distribution. This species is widely distributed throughout the Guiana Shield region, from Tobogan de la Selva in Venezuela east to the coast of French Guiana (Fig. 15). Reported from Suriname in Short and Kadosoe (2011) and Short (2013; as N. shorti) and from Guyana as Notionotus sp. B in Short et al. (2018).	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
90490B357B285A32B923C50F04517D50.taxon	diagnosis	Diagnosis. The species of this group can be diagnosed by the following combination of characters: (1) the shape of the elevation mesoventrite, having one transverse ridge and one longitudinal ridge (Fig. 10 A, B); (2) parameres nearly as long as the basal piece; the length of the median lobe and the length of the parameres is approximately subequal. (e. g., Notionotus liparus, Fig. 7 A).	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
756E9E936B7A51C4B06D1606A8AC451A.taxon	description	Figs 3 A-I, 6 A-I, 11 C, D, 14	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
BCB825EF80255E7EBCD5C4220D99CE9A.taxon	description	Figs 4 H, 7 C, 13 B, 14	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
BCB825EF80255E7EBCD5C4220D99CE9A.taxon	description	Description. Size and form: Body length 1.7 - 2.3 mm. Body form elongate oval, strongly convex in lateral view (Fig. 4 H). Color and punctation: Dorsally brown, head mostly brown, frons and medial region of the clypeus dark brown, lateral side of the clypeus pale brown; pronotum pale brown with two small black round spots along posterior margin, elytra dark brown (Fig. 4 H). Ventrally dark brown; maxilla, maxillary palps, antennae (antennal club slightly darker) yellow, legs pale brown. Clypeus and labrum with dense, fine, and weakly impressed ground punctation (punctures separated by 2 x their width); pronotum and elytra ground punctation fine, weakly impressed and sparser than on head (punctures separated by 3 x their width). Head: Clypeus and labrum shallowly emarginate anteromedially, lateral margins of the labrum bearing setae. Thorax: Prosternum carinate medially, strongly raised, pointing anteriorly and acute. Elevation of mesoventrite with one transversal ridge, elevated medially, lateral sides concave; longitudinal ridge sharp, the point where the two ridges merged rounded and obtuse (e. g., Fig. 10 B); elevation flat in lateral view; mesoventrite with triangular shape in ventral view. Metaventrite convex in the median region, pubescent with narrow glabrous patch on the medial and posterolateral area, medial region patch drop-shaped; anterior margin extending to mesoventrite elevation. Metafemora densely covered with hydrofuge pubescence on basal three-quarters. Abdomen: Abdominal ventrites very densely pubescent. Aedeagus (Fig. 7 C) with basal piece nearly the same length as a paramere. Base of the parameres slightly narrower than the base of the median lobe; outer and inner margins sinuate; apex of parameres wide and blunt, pointing outwards. Median lobe of the same length as the parameres, wide at basal region, narrowing in the midlength, then widening along at apical third, apex rounded; gonopore with rounded shape and situated at midlength of median lobe.	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
BCB825EF80255E7EBCD5C4220D99CE9A.taxon	etymology	Etymology. The name is derived from the Latin word vatius meaning bent outwards, after the form of the parameres slightly pointing outwards of the aedeagus.	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
BCB825EF80255E7EBCD5C4220D99CE9A.taxon	distribution	Distribution. This species is known from several localities in Bahia and Mato Grosso do Sul States in Brazil (Fig. 14).	en	Gonzalez-Rodriguez, Liza M., Short, Andrew Edward Z. (2022): Revision of the water scavenger beetle genus Notionotus Spangler, 1972 in the Neotropical Region (Coleoptera, Hydrophilidae, Enochrinae). ZooKeys 1109: 141-191, DOI: http://dx.doi.org/10.3897/zookeys.1109.80775, URL: http://dx.doi.org/10.3897/zookeys.1109.80775
