identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
CE0587C302350F6BD79CF96646CBFF22.text	CE0587C302350F6BD79CF96646CBFF22.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eoparanaxia Ferratges & Domínguez & Ossó & Zamora 2023	<div><p>Genus EOPARANAXIA n. gen.</p> <p>Figure 3</p> <p>zoobank.org/ 07B9F785-4E4A-4018-9D7B-50B3576421F4</p> <p>Spain 0 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 1 1 0 0 1 1 0 0 0 0</p> <p>Italy 0 0 0 1 0 0 0 0 1 0</p> <p>Egypt 0 0 1 0 0 0 0 0 0 0</p> <p>Hungary 0 0 0 1 0 0 0 0 0 0</p> <p>UK 0 0 0 1 0 0 0 0 0 0</p> <p>Germany 0 0 0 0 0 0 0 0 0 0</p> <p>Lutetian Senegal Pakistan 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Venezuela 0 0 0 0 0 0 0 0 0 0</p> <p>Argentina 0 0 0 0 0 0 1 0 0 0</p> <p>Washington 0 0 0 0 0 0 0 0 0 0</p> <p>Carolina 0 0 0 0 0 0 0 0 0 0</p> <p>Florida 0 0 0 0 0 0 0 0 0 0</p> <p>New 0 0 0 0 0 0 0 0 0 0 Zealand</p> <p>0</p> <p>0</p> <p>0</p> <p>0</p> <p>0</p> <p>0</p> <p>0</p> <p>0</p> <p>0</p> <p>0</p> <p>0</p> <p>0</p> <p>0 0 0 0 0 1 0 1 1 0 1 1 1 0 0 0 0</p> <p>0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0</p> <p>0 0 0 0 0 0 1 0 0 0 1 0 0 0 0 0</p> <p>0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0</p> <p>0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0</p> <p>0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0</p> <p>0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0</p> <p>0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0</p> <p>0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0</p> <p>0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0</p> <p>0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0</p> <p>0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0</p> <p>0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0</p> <p>Type species. Eoparanaxia eocenica n. gen. n. sp. by monotypy and present designation. Gender feminine.</p> <p>Diagnosis. Carapace pyriform, longer than wide; rostrum axially sulcate, with two fused, long pseudorostral spines, length 0.53 times CW. Hepatic region with small spine, directed anterolaterally. Intestinal tubercle strong, protruding beyond posterior carapace margin; carapace regions well defined with metagastric and urogastric regions narrower than mesogastric and cardiac regions; mesobranchial region with posteriorly directed large spine. Dorsal regions armed with long spines.</p> <p>Etymology. The generic name derives from the prefix Eo- (from ɳως (gr.)= aurora), to generically indicate an ancestral form, in arbitrary combination with the generic name Paranaxia Rathbun, 1924, to refer to its morphological affinities with the new genus.</p> <p>Remarks. The material herein described is referred to the subfamily Pisinae Dana, 1851. Species of this subfamily possess elongated carapaces; elongated pseudorostral spines; orbits always with postorbital spine or lobe, usually cupped, sometimes with antorbital spine; carapace triangular, sometimes with posterior spine (see Schweitzer et al., 2020). The new genus possesses all these characteristics and is therefore referred to this subfamily.</p> <p>The new genus Eoparanaxia has similarities with Paranaxia, in the general outline of the carapace, showing a very elongated pseudorostral spines, parallel and distally bifid, slightly differentiated dorsal regions, with aligned spines in the dorsal part of the mesobranchial region, and well developed mesobranchial and intestinal spines (i.e., Windsor and Felder, 2014), sometimes beyond posterior carapace margin (see Rathbun, 1924, and Hosie and Hara, 2016, p. 128, figure 2; p. 129, figure 3).</p> <p>However, Paranaxia presents the pseudorostral spines separated from their base, a supraorbital eave with a pronounced spine at the preorbital lobe, postorbital angle with a spine separated from the anterior lobe by a notch. In addition, Eoparanaxia n. gen. exhibits a postorbital spine between abbreviations employed in Table 2.</p> <p>the gastric region and the hepatic region, which is not present in Paranaxia.</p> <p>The studied material has similar characteristics with the modern genus Sphenocarcinus A. Milne-Edwards, 1875, Oxypleurodon Miers, 1885, and Rhinocarcinus de Forges and Ng, 2009, including the shape of the rostrum with two long and coalescent cylindrical spines with slightly diverging sharp tips, and the shape and location of the orbits. However, Eoparanaxia n. gen. shows a different distribution of dorsal regions, with shallower dorsal grooves than in Sphenocarcinus, Oxypleurodon, and Rhinocacinus; intestinal region with a prominent conical spine unlike in Sphenocarcinus, Oxypleurodon, and Rhinocacinus, which lack such conical expansion; sinuous posterior margin; longitudinal antennal pits, parallel to the axis of the body unlike in the other three genera, which are oblique.</p> <p>The modern genera Pisa Leach, 1815, Leptopisa Stimpson, 1871 (both included in Pisinae Dana, 1851), and Oregonia Dana, 1851 (Oregoniidae Garth, 1958) also show similarities with the new genus in the shape of the rostrum and orbits (i.e., Zariquiey-Álvarez, 1968; p.449, figure 151; Carmona-Suárez and Poupin, 2016, p. 369, figure 5). However, the shape and distribution of dorsal regions, posterior margin, dorsal surface with elevated regions, without spines, are clearly different in the modern genus. Furthermore, Eoparanaxia n. gen. presents the pseudorostrum fused throughout its length, only diverging at the tip, (while in the three mentioned taxa it separates from the base); presents a strong and long intestinal spine, a long branchial spines and spiny ridges on the dorsal surface (of which are absent in the three taxa). In addition, Pisa has anterolateral margins generally straight or slightly concave, with fewer or without spines and lacks the strong and prominent spine in the intestinal region that the new genus possesses.</p> <p>Other modern genera, Rochinia A. Milne-Edwards, 1875, Scyramathia A. Milne-Edwards, 1880, Minyorhyncha Tavares and Santana, 2018, and Anamathia Smith, 1885 share with the new genus the general outline of the carapace, distribution of spines in the posterolateral and posterior margins, especially in juvenile stages (see Tavares and Santana, 2018; p. 206-214, figures 1-9). However, all these genera have clearly separated and divergent pseudorostral spines unlike in Eoparanaxia n. gen., which presents parallel and fused spines. Some species of Doclea Leach, 1815, show similarities with the new genus, for instance a fused bilobed pseudorostrum only diverging at the tip, a long and acute intestinal spine, a row of axial spines, oblique carinae, paralleling anterolateral margin, and also in the branchial regions. However, the modern genus Doclea differs from Eoparanaxia n. gen. in having a shorter pseudorostrum, a notch in the supraorbital eave, and mostly rounded or less elongated outline.</p> <p>The material studied here also shows a certain resemblance to some representatives of Inachoididae Dana, 1851, in view of the general outline of the carapace (see Santana, 2008; Lima et al., 2022). However, Inachoididae have usually a shorter pseudorostral spine, strongly fused, and generally ended in a single tip, concave posterior margin, and less spinose/tuberculate dorsal surface.</p> </div>	https://treatment.plazi.org/id/CE0587C302350F6BD79CF96646CBFF22	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ferratges, Fernando A.;Domínguez, Josep Lluis;Ossó, Àlex;Zamora, Samuel	Ferratges, Fernando A., Domínguez, Josep Lluis, Ossó, Àlex, Zamora, Samuel (2023): Spider crabs (Decapoda: Brachyura: Majoidea) from the upper Eocene of south Pyrenees (Huesca, Spain). Palaeontologia Electronica (a 27) 26 (2): 1-29, DOI: 10.26879/1270, URL: http://dx.doi.org/10.26879/1270
CE0587C3023A0F6BD566FECA44E0F8A7.text	CE0587C3023A0F6BD566FECA44E0F8A7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eoparanaxia eocenica Ferratges & Domínguez & Ossó & Zamora 2023	<div><p>Eoparanaxia eocenica n. sp.</p> <p>Figures 3 and 4</p> <p>zoobank.org/ 1AAD44D5-0D9C-4642-8FC2-85193465EFDF</p> <p>Type material. Holotype (MPZ 2023 /1), a near-complete carapace, partially decorticated. Paratypes (MPZ 2023 /2 and MPZ 2023 /3) that correspond to one rostrum and half of a posterior carapace.</p> <p>Diagnosis. As for the genus, by monotypy.</p> <p>Description. Carapace pyriform, longer than wide, with spines and conical tubercles; pseudorostral spines parallel and fused, distally bifid, 0.53 times CW, with a row of aligned granules on each spine; orbits not well preserved, small, rounded, obliquely directed. Regions well defined by shallow grooves and deep branchiocardiac groove, with spaced and large spines at the top of the dorsal regions. Gastric region prominent, slightly higher than other regions, with pointed tubercles distributed in anterior gastric region, becoming long gastric spines on the axis of the carapace; urogastric region more depressed. Hepatic region slightly swollen, with one acute stout spine directed anterolaterally. Branchial regions inflated; epibranchial region inflated; mesobranchial region crossed by oblique ridge with five tubercles/spines. Cardiac region elevated, with two tubercles in the anterior part and one in the posterior margin. Intestinal tubercle large, protruding beyond posterior margin of carapace, conical, apex pointed.</p> <p>Anterolateral margin slightly sinuous. Posterolateral margin convex, with a long spine laterally directed obliquely. Posterolateral and posterior margin rimmed.</p> <p>Etymology. The specific name refers to the Eocene.</p> <p>Remarks. Some species of the genus Paranaxia like P. keesingi Hosie and Hara, 2016, and P. serpulifera (Guérin, 1832, in Guérin-Méneville 1829- 1837) show similarities with the material studied here. Nevertheless, these taxa have unfused pseudorostral spines, clearly divergent. The new taxon bears some similarities with Macrocheira longirostra Schweitzer and Feldmann, 1999, with a similar outline of the carapace, bifid rostrum, very long pseudorostral spines fused at the base, and divergent at the tip. However, M. longirostra show some differences: 1) lacks spines at the posterolateral margins; 2) has a posterior margin almost straight without intestinal spine; and 3) a dorsal surface much more tuberculated than the new taxon.</p> </div>	https://treatment.plazi.org/id/CE0587C3023A0F6BD566FECA44E0F8A7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ferratges, Fernando A.;Domínguez, Josep Lluis;Ossó, Àlex;Zamora, Samuel	Ferratges, Fernando A., Domínguez, Josep Lluis, Ossó, Àlex, Zamora, Samuel (2023): Spider crabs (Decapoda: Brachyura: Majoidea) from the upper Eocene of south Pyrenees (Huesca, Spain). Palaeontologia Electronica (a 27) 26 (2): 1-29, DOI: 10.26879/1270, URL: http://dx.doi.org/10.26879/1270
CE0587C3023A0F6CD75AFF7D44BBFC3F.text	CE0587C3023A0F6CD75AFF7D44BBFC3F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Planobranchia Schweitzer and Feldmann 2010	<div><p>Genus PLANOBRANCHIA Schweitzer and Feldmann, 2010</p> <p>Type species. Micromaia laevis Lőrenthey, 1909, by original designation of Schweitzer and Feldmann (2010, p. 407).</p> <p>Fossil included species.? Planobranchia egyptensis Feldmann, Schweitzer, Bennett, Franţescu, Resar, and Trudeau, 2011; P. elongata n. sp. (herein); P. laevis (Lőrenthey, 1909); P. palmuelleri Artal, Van Bakel, and Onetti, 2014; P. simplex (Remy in Gorodiski and Remy, 1959).</p> <p>Emended diagnosis. Carapace pyriform, widest at midlength of branchial region; moderately vaulted transversely and longitudinally, front produced, singular, sulcate longitudinally. Orbits small, laterally situated, with strong and subtriangular orbital spines. Gastric regions only weakly differentiated; defined laterally by prominent V-shaped Groove converging from anterior margin of orbits to urogastric region, the narrowest part of axial regions. Cardiac region nearly as wide as widest part of gastric regions, hexagonal to ovoid in outline; bearing two nodes on medial transverse ridge. Intestinal region well defined, long, approximately as wide as urogastric region. Epibranchial and mesobranchial regions strongly inflated, separated from one another by subtle arcuate attachment scar expressed on mold of the interior of the carapace; widest part of these regions converge as angular projections toward urogastric region. Metabranchial region extends from widest part of cardiac region posterolaterally around posterior margin of metabranchial region and clearly defined axially by posterior margin of cardiac region and intestinal region; depressed below other regions. Surface of carapace weakly ornamented by fine granules or pits; lacking strong tubercles, posterior margin convex, rimmed (new additions to the original diagnosis of Schweitzer and Feldmann, 2010).</p> <p>Remarks. The studied specimen can be assigned to Planobranchia Schweitzer and Feldmann, 2010, because it shares the diagnostic characteristics of the genus (see Schweitzer and Feldmann, 2010) like: 1) the moderately vaulted transversely and longitudinally pyriform carapace; 2) weakly differentiated gastric regions, defined laterally by prominent V-shaped groove converging from anterior margin of orbits to urogastric region; 3) hexagonal to ovoid cardiac region, bearing two nodes on medial transverse ridge; 4) strongly inflated epibranchial and mesobranchial regions, separated from one another by subtle arcuate attachment scar; 4) weakly ornamented surface of the carapace by fine pits, lacking strong tubercles.</p> <p>Some authors assigned Planobranchia to the subfamily Majinae (Schweitzer and Feldmann, 2010; Feldmann et al., 2011; and Schweitzer et al., 2020), and justify this inclusion by the supraorbital margin with an “eave orbital” and a postorbital spine. Subsequently, Artal et al. (2014) proposed to include the genus Planobranchia in Inachidae, justifying its inclusion by similarities in the frontal and orbital construction. Nevertheless, due to the bad preservation, specifically of the pseudorostrum and part of the supraorbital margin, Artal et al. (2014) have misinterpreted the fronto-orbital conformation and the anterolateral spines. Due to the lack of the anterior part of the pseudorostrum spines, these authors have suggested that Planobranchia could have a short pseudorostrum like many inachids, placing the orbits laterally on the sides of the pseudorostrum. Also, the antorbital spine has been interpreted as the postorbital spine, and the two following spines (intercalated spine and postorbital spine) as anterolateral spines. Instead, Planobranchia has a rather elongated rostrum and an orbital construction consisting of three spines.</p> <p>The characteristics of the new material and the reanalysis of previously known taxa suggest that this genus has more affinity with the subfamily Pisinae, so its inclusion in this group is suggested here. Placement in Pisinae is supported by the morphology of the carapace outline, the distribution and shape of the dorsal regions, supraorbital margin formed by a prominent antorbital spine, a small intercalated spine, and a well-developed postorbital spine; axial regions separated from the periphery by deep grooves; hepatic lobe marked by a lump or spine; highly developed branchial regions; thickened cardiac region; mesogastric region in which a prominent lump stands out (see Griffin and Tranter, 1986, Schweitzer et al., 2020).</p> </div>	https://treatment.plazi.org/id/CE0587C3023A0F6CD75AFF7D44BBFC3F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ferratges, Fernando A.;Domínguez, Josep Lluis;Ossó, Àlex;Zamora, Samuel	Ferratges, Fernando A., Domínguez, Josep Lluis, Ossó, Àlex, Zamora, Samuel (2023): Spider crabs (Decapoda: Brachyura: Majoidea) from the upper Eocene of south Pyrenees (Huesca, Spain). Palaeontologia Electronica (a 27) 26 (2): 1-29, DOI: 10.26879/1270, URL: http://dx.doi.org/10.26879/1270
CE0587C3023D0F6CD51EFC2843AEFA9F.text	CE0587C3023D0F6CD51EFC2843AEFA9F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Planobranchia elongata Ferratges & Domínguez & Ossó & Zamora 2023	<div><p>Planobranchia elongata n. sp.</p> <p>Figures 5 A-B and 6</p> <p>zoobank.org/ 8FE98FE3-0E49-4C45-A30A-73324B9512B3</p> <p>Type material. One partial specimen, partially decorticated: MPZ 2023 /4.</p> <p>Diagnosis. Carapace pyriform, longer than wide, maximum width in mesobranchial region; front singular, sulcate longitudinally, pseudorostral spines short, fused; orbits small, laterally situated, with strong and subtriangular orbital spines; dorsal regions swollen, distinct, bounded by grooves; mesogastric region inflated and smooth, narrow and elongated anteriorly, bounded by two elongated ridges; metagastric region narrow, Ushaped; branchial regions differentiated; mesobranchial region arched; posterior margin broad, rimmed.</p> <p>Description. Carapace pyriform in outline, longer than wide; dorsal surface covered by small pits, convex in both directions; maximum width in mesobranchial region; front produced, straight, directed forwards, with two fused spines, with longitudinal ridges; orbits anterolaterally directed, supraorbital margin with strong spines: antorbital spine is the largest, triangular in shape and slightly forward; intercalated spine shortest, conical, separated by supraorbital sutures; postorbital spine triangular, medium sized, facing out. Lateral margins smooth, arched; hepatic region slightly inflated, defined by shallow groves; notable subtriangular spine in the postorbital lobe; mesogastric region inflated, ushaped, apparently smooth and anterior portion ridged; protogastric region weakly defined; urogastric region bounded by shallow grooves; branchial regions delimited from axial regions by grooves; cardiac region triangular; epibranchial region inflated, oblique, arched posteriorly; mesobranchial region broadly inflated; metabranchial region depressed; intestinal region not preserved; posterior margin not preserved, appears broad.</p> <p>Etymology. The specific name refers to its elongated morphology.</p> <p>Remarks. The type species, Planobranchia laevis, from the Lutetian of Egypt, shows clear affinity with P. elongata n. sp., but has some differences: 1) distinct hepatic region, with smaller hepatic spine than the new species; 2) gastric regions less pronounced and more elongated and dorsal surface ornated with very small granules, instead of the smooth surface with setal pits that the new species has. Planobranchia palmuelleri from the Lutetian of Catalonia is clearly different from P. elongata n. sp. in some aspects like: 1) its carapace outline, being much wider and shorter than the new species; and 2) by its shorter rostrum. Planobranchia simplex from the Lutetian of Senegal, shows certain similarities with the new species, such as an arched margin, defined by a thin rim that does not appear to be separated by strong depressions on the branchial regions, however; P. simplex clearly differs from the new species in: 1) the outline of the carapace, much wider than the new species and 2) the deeper axial grooves. The new taxon differs from? P. egyptensis from the Lutetian of Egypt, in having a slimmer outline and more inflated dorsal regions.</p> </div>	https://treatment.plazi.org/id/CE0587C3023D0F6CD51EFC2843AEFA9F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ferratges, Fernando A.;Domínguez, Josep Lluis;Ossó, Àlex;Zamora, Samuel	Ferratges, Fernando A., Domínguez, Josep Lluis, Ossó, Àlex, Zamora, Samuel (2023): Spider crabs (Decapoda: Brachyura: Majoidea) from the upper Eocene of south Pyrenees (Huesca, Spain). Palaeontologia Electronica (a 27) 26 (2): 1-29, DOI: 10.26879/1270, URL: http://dx.doi.org/10.26879/1270
CE0587C302200F71D4F1FF7D473CFDB7.text	CE0587C302200F71D4F1FF7D473CFDB7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Spinirostrimaia Beschin, De Angeli, Checchi, and Zarantonello 2012	<div><p>Genus SPINIROSTRIMAIA Beschin, De Angeli, Checchi, and Zarantonello, 2012</p> <p>Type species. Micromaia margaritata Fabiani, 1910.</p> <p>Fossil included species. Spinirostrimaia margaritata (Fabiani, 1910) (from Lutetian of Italy); S. echinate n. sp. (herein).</p> <p>Remarks. The studied material can be assigned to Spinirostrimaia based on the carapace general outline elongate, distribution of dorsal regions, orbital position and shape of the orbits, and the long pseudorostral spines with three lateral spines on the outer margins (see diagnosis in Beschin et al., 2012).</p> </div>	https://treatment.plazi.org/id/CE0587C302200F71D4F1FF7D473CFDB7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ferratges, Fernando A.;Domínguez, Josep Lluis;Ossó, Àlex;Zamora, Samuel	Ferratges, Fernando A., Domínguez, Josep Lluis, Ossó, Àlex, Zamora, Samuel (2023): Spider crabs (Decapoda: Brachyura: Majoidea) from the upper Eocene of south Pyrenees (Huesca, Spain). Palaeontologia Electronica (a 27) 26 (2): 1-29, DOI: 10.26879/1270, URL: http://dx.doi.org/10.26879/1270
CE0587C302200F75D518FDA04719FDC2.text	CE0587C302200F75D518FDA04719FDC2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Spinirostrimaia echinata Ferratges & Domínguez & Ossó & Zamora 2023	<div><p>Spinirostrimaia echinata n. sp.</p> <p>Figures 7-9</p> <p>zoobank.org/ B169EC2C-D88A-465D-A140-14B3A9395D48</p> <p>Type material. The holotype is a female specimen, near-complete carapace, partially decorticated, with one cheliped and thoracic sternum (MPZ 2023 /6) (Figure 7). There are four paratypes (MPZ 2023 /7-2023/10) (Figure 8).</p> <p>Additional material. Thirteen additional specimens composed by near-complete carapaces (MPZ 2023/11-2023/23).</p> <p>Diagnosis. Carapace pyriform, longer than wide, convex; regions separated by shallow grooves; dorsal surface covered by small spines. Frontal margin narrow, sulcate, with two long, subparallel pseudorostral spines with spiny outer margin proximally situated; dorsal surface covered by spines (pearl-shaped tubercles if partially decorticated); orbits with prominent supraorbital eaves, without orbital spines; anterolateral margins elongated, interrupted with hepatic and branchial grooves; hepatic lobe with three lateral spines; mesobranchial margin strongly convex, with numerous spines (10-13); posterior margin with small spines.</p> <p>Description. Carapace pyriform, twice as long as wide, (not counting the pseudorostrum), ovate; pseudorostrum long, bifid, composed of two spines which have three tiny spines on outer margin; frontal region with two longitudinal crests, tuberculate. Almost complete orbits, with prominent supraorbital eave, without antorbital spine, but marked preorbital lobe; intercalated spine triangular; postorbital spine elongated, slightly curved forward; hepatic lobe with three pointed spines, slightly curved forward. Lateral margins convex, notched by the cervical groove. Dorsal regions ornamented with sharp spines, with pearl-shaped tubercles appearance if partially decorticated; carapace regions well defined by relatively shallow grooves; axial regions elevate above other regions. Proto- and mesogastric regions inflated; meta- and urogastric regions narrower than mesogastric and cardiac regions; hepatic region inflated; branchial regions wide; epi- and mesobranchial regions inflated, poorly differentiated by a shallow groove; metabranchial region slightly depressed; cardiac region inflated, with two lateral subtriangular extensions defined by shallow grooves; intestinal region small, slightly depressed; posterior margin broad, rimmed, with small spines. Branchiocardiac grooves deep.</p> <p>Epistome wide, smooth, and rimmed. Female thoracic sternum strongly concave, with interrupted sutures (only preserved 1/2 to 5/6, see Figure 7B). Female chelipeds elongated and thin; merus elongated, with longitudinal depression on the ventral surface, surrounded by well-separated blunt spines; carpus slightly elongated, ornamented with small spines; palm slightly compressed, oval in cross section; fingers acute, relatively short, square in section, with longitudinal striae. Basal antennal article moderately wide, broader at the base than at its distal extremity.</p> <p>Etymology. The specific epithet make reference to its spinose carapace.</p> <p>Remarks. The new species shows similarities with the type species Spinirostrimaia margaritata in the general shape of the carapace, piriform and elongated, and long pseudorostral spines. However, the new species differs in some aspects: 1) subparallel pseudorostral spines, not convergent as in S. margaritata; 2) slightly smaller postorbital spine, without ornamentation; 3) the dorsal ornamentation, more spinose in the new species (instead of pearl-shaped tubercles of S. margaritata); 4) prominent spine in the margin of the hepatic region, that is absent in S. margaritata; 5) small, sharp spines covering the dorsal surface, lacking pearl-shaped tubercles as in S. margaritata; 6) wider posterior margin than S. margaritata, and less convex and rimmed; and 7) more spiny margins in the carapace of the new species than S. margaritata (see Beschin et al., 2012, figures 41, 80; t. 6, figures 3, 6a, b, 7a, b).</p> <p>The fossil species Cromimaia meneguzzoi (Beschin, Busulini, De Angeli, and Tessier, 1985) bears some resemblance to the new species. However, S. echinata n. sp. differs in some aspects: 1) the urogastric region is narrower in the new species, with three tubercles forming a triangle, and not aligned as in C. meneguzzoi; 2) narrower cardiac region, better delimited by branchiocardiac grooves in the new species; 3) the pseudorostral spines are exceedingly shorter in C. meneguzzoi, in contrast to the long spines of the new species (see Beschin et al., 2012, figure 40; t. 6, figures 4ac). The new species also shows similarities with Micromaia tuberculata Bittner, 1875, but has differences in the branchiocardiac groove and both the urogastric region and the beginning of the cardiac region are narrower; the intestinal region is slightly more swollen; and chiefly in having two long and parallel pseudorostral spines, whereas in M. tuberculata they are much shorter and flattened subtriangular (see Beschin et al., 2012, figure 38; t. 6, figure 2).</p> </div>	https://treatment.plazi.org/id/CE0587C302200F75D518FDA04719FDC2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ferratges, Fernando A.;Domínguez, Josep Lluis;Ossó, Àlex;Zamora, Samuel	Ferratges, Fernando A., Domínguez, Josep Lluis, Ossó, Àlex, Zamora, Samuel (2023): Spider crabs (Decapoda: Brachyura: Majoidea) from the upper Eocene of south Pyrenees (Huesca, Spain). Palaeontologia Electronica (a 27) 26 (2): 1-29, DOI: 10.26879/1270, URL: http://dx.doi.org/10.26879/1270
