taxonID	type	description	language	source
CF548A3AFFD0E173FF07C982FE18FB94.taxon	description	(Figs 1, 3) Chinese common name: 九AEDz沟Dzm	en	Zhang, Yong-Qin, Wei, Guo-Hao, Yin, Zi-Wei (2024): Two new species of ant-loving beetles (Coleoptera: Staphylinidae: Pselaphinae) from Jiulong National Wetland Park, China. Zootaxa 5406 (2): 336-342, DOI: 10.11646/zootaxa.5406.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5406.2.6
CF548A3AFFD0E173FF07C982FE18FB94.taxon	materials_examined	Type material (10 exx.). HOLOTYPE: CHINA: ♂: ‘ China: Zhejiang, Lishui, Jiulong National Wetland Park, 28 ° 23 ’ 4.86 ” N, 119 ° 49 ’ 29.15 ” E, 50 m, 22. ix. 2023, fern leaves, Zhang, Wei, Ming & Jia leg. (浙江丽水九AE国NJae 地公园) ’ (SNUC). PARATYPES: CHINA: 1 ♂, 1 ♀, same collecting data as for holotype; 4 ♀♀, same collecting data as for holotype, except ‘ 23. ix. 23 ’ (浙江丽水九AE国NJae地公园) ’; 1 ♂, 2 ♀♀, ‘ China: Zhejiang, Lishui, Bihu To. Hongyu Vill., Jiulong Wetland Park, 28 ° 23 ’ 6 ” N, 119 ° 49 ’ 29 ” E, 50 m, 2. vi. 2023, riverbank, sifted, Z-W. Yin leg. (丽水九AE国NJae地公园河岸 ® 丛) ’ (1 ♂, 1 ♀ in IZCAS, others in SNUC).	en	Zhang, Yong-Qin, Wei, Guo-Hao, Yin, Zi-Wei (2024): Two new species of ant-loving beetles (Coleoptera: Staphylinidae: Pselaphinae) from Jiulong National Wetland Park, China. Zootaxa 5406 (2): 336-342, DOI: 10.11646/zootaxa.5406.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5406.2.6
CF548A3AFFD0E173FF07C982FE18FB94.taxon	diagnosis	Diagnosis. Male. Body length approximately 2.0 mm. Head sub-rectangular at base; vertex finely punctate, with short, transverse sulcus at anterior portion; antenna elongate; antennomeres more or less elongate, lacking obvious modifications, antennomere 10 moderately swollen and distinctly longer than 9. Pronotum lacking median longitudinal sulcus, with smooth disc, basal color roughened. Discal striae of elytra extending posteriorly to approximately apical 3 / 4 of elytral length. Metaventrite with pair of admesal denticulate projections. Mesotibia with moderately long apical spine. Tergite 1 (IV) predominantly large, dorsally longer than 2 – 4 (V – VII) combined, lacking modifications. Aedeagus strongly asymmetric; median lobe with restricted basal capsule and subtriangular foramen, ventral stalk slightly shorter than dorsal lobe, narrowed at apex. Female. Body length approximately 1.8 mm, legs and metaventrite lacking modifications, genitalia as in Fig. 1 G.	en	Zhang, Yong-Qin, Wei, Guo-Hao, Yin, Zi-Wei (2024): Two new species of ant-loving beetles (Coleoptera: Staphylinidae: Pselaphinae) from Jiulong National Wetland Park, China. Zootaxa 5406 (2): 336-342, DOI: 10.11646/zootaxa.5406.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5406.2.6
CF548A3AFFD0E173FF07C982FE18FB94.taxon	description	Description. Male. Body (Fig. 1 A) length 1.90 – 2.01 mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface finely punctate, covered with short pubescence. Head (Fig. 1 B) sub-rectangular, truncate at base, slightly wider than long, length 0.38 – 0.40 mm, width across eyes 0.42 – 0.45 mm; vertex finely punctate, with moderately separated foveae (dorsal tentorial pits), with short, transverse sulcus at apical portion of vertex; tempora much shorter than eyes, convergent posteriorly; lacking mediobasal carina; antennal tubercles weakly raised; frons slightly impressed medially, confluent with clypeus; clypeus smooth, its anterior margin carinate and moderately raised; ocular-mandibular carinae complete, distinct. Venter with small gular foveae (posterior tentorial pits) in single pit, with distinct median carina extending from foveae anteriorly to mouthparts. Eyes greatly prominent, each composed of approximately 29 – 35 ommatidia. Maxillary palpus with palpomere 1 minute, 2 elongate, curved, pedunculate basally and enlarged apically, 3 short, sub-trapezoidal, 4 fusiform, widest near middle. Antenna moderately elongate, length 0.81 – 0.93 mm; club loosely formed by enlarged apical three antennomeres (Fig. 1 C); antennomere 1 thick, subcylindrical, 2 – 7 each elongate, successively longer, 8 shortest, 9 much longer and broader than 8, 10 longer than 9, 11 longest, slightly shorter than (21: 25), sub-conical. Pronotum (Fig. 1 B) almost as long as wide, length 0.43 – 0.46 mm, width 0.42 – 0.45 mm, widest at middle; semi-circular lateral sulci extending from dorsal surface laterally and posteriorly and then fused with lateral ends of antebasal sulcus; disc moderately convex, finely punctate, median longitudinal sulcus absent, lacking median antebasal fovea, with short mediobasal impression, lateral antebasal foveae connected by transverse antebasal sulcus. Prosternum with anterior part slightly shorter than coxal part, with small lateral procoxal foveae; hypomeral grooves short, present only on basal halves, lacking lateral antebasal hypomeral impression; hypomeral ridges close to coxal cavities. Elytra much wider than long, length 0.61 – 0.66 mm, width 0.70 – 0.76 mm; suboval, dorsal surface with dense, short pubescence; each elytron with two large, asetose, basal foveae; discal striae long, extending from outer basal foveae posteriorly to 7.6 / 10 of elytral length; humeral region moderately raised, subhumeral fovea absent, with sulcate marginal stria extending from approximately middle to posterior margin. Metathoracic wings fully developed. Mesoventrite short, demarcated from metaventrite by ridged anterior edges of impressed areas where lateral mesocoxal foveae situated at mesal ends of impressions, with pair of admesal carinae; median mesoventral foveae widely separated, originating from shared setose, transverse opening, lateral mesoventral foveae large and setose, broadly forked internally, with short intercoxal process. Metaventrite finely punctate, broadly impressed at middle, lateral margin of impression with pair of denticulate projections; with large, setose lateral mesocoxal foveae and pair of smaller, setose lateral metaventral foveae, posterior margin with small and narrow split at middle. Legs moderately elongate; protibia slightly arcuate; mesotibia with distinct spine at apex (Fig. 1 D); mesofemur widened to middle; each tarsus with one major and one reduced setiform pretarsal claw. Abdomen widest at lateral margins of tergite 1 (IV), length 0.78 – 0.82 mm, width 0.62 – 0.66 mm. Tergite 1 (IV) in dorsal view longer than 2 – 4 (V – VII) combined, with inner basolateral foveae embracing thin sulcus and pair of outer basolateral foveae, short, triangular discal carinae present; tergites 2 (V) and 3 (VI) lacking foveae and sulcus, 4 (VII) with pair of small basolateral foveae, slightly shorter than 2 (V) and 3 (VI) combined along middle; tergite 5 (VIII) semicircular, posterior margin narrowly and deeply emarginate at middle. Sternite 2 (IV) with mediobasal and basolateral foveae; midlength of sternite 2 approximately as long as 3 – 4 (V – VI) combined, 3 and 4 each short at middle, combined approximately longer than 5 (VII), 3 – 5 (V – VII) lacking foveae, 6 (VIII) transverse, posterior margin convex at middle. Aedeagus (Fig. 1 E, F) 0.29 mm in length, moderately sclerotized, dorso-ventrally strongly asymmetric; median lobe with restricted basal capsule and small, subtriangular foramen, ventral stalk elongate, widest at approximately apical 1 / 4 and then tapering apically; dorsal lobe broadened at base, narrowing towards apex; parameres fused and reduced to ventral membrane. Female. General external morphology similar to male; antennomeres simple; each eye composed of about 28 – 30 ommatidia; humeri flat; metathoracic wings reduced. Measurements (as for male): body length 1.80 – 1.83 mm, length / width of head 0.37 – 0.38 mm / 0.44 mm, length / width of pronotum 0.43 – 0.46 mm / 0.42 – 0.44 mm, length / width of elytra 0.58 – 0.59 mm / 0.71 – 0.73 mm, length / width of abdomen 0.52 – 0.56 mm / 0.64 – 0.67 mm; length of antenna 0.82 – 0.92 mm; genital complex (Fig. 1 G) 0.22 mm wide, genital plate approximately as wide as sternite 7 (IX), with lateral arms slender and narrowing apically. Comparative notes. The male of this species is most similar to Trisiniotus taoismus Feng & Yin, distributed in Jiangsu, eastern China by the simple male antennomeres 10. They can be separated by antennomeres 10 being distinctly longer than 9 (as long as 9 in T. taoismus), a finely punctate pronotum (pronotum with much more distinct punctures in T. taoismus), much larger apical spines of the mesotibiae, and a different aedeagus. Within the genus, T. taoismus is the only other species that has the female genital complex illustrated (Feng & Yin 2022: fig. 1 F). The genital complex (Fig. 1 G) of T. jiulong sp. nov. is less transverse, and the lateral arms of the genital plate are relatively much slenderer than those of T. taoismus.	en	Zhang, Yong-Qin, Wei, Guo-Hao, Yin, Zi-Wei (2024): Two new species of ant-loving beetles (Coleoptera: Staphylinidae: Pselaphinae) from Jiulong National Wetland Park, China. Zootaxa 5406 (2): 336-342, DOI: 10.11646/zootaxa.5406.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5406.2.6
CF548A3AFFD0E173FF07C982FE18FB94.taxon	biology_ecology	Biology. All individuals were collected from sifted fern leaf samples along or near the riverbank (Fig. 3).	en	Zhang, Yong-Qin, Wei, Guo-Hao, Yin, Zi-Wei (2024): Two new species of ant-loving beetles (Coleoptera: Staphylinidae: Pselaphinae) from Jiulong National Wetland Park, China. Zootaxa 5406 (2): 336-342, DOI: 10.11646/zootaxa.5406.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5406.2.6
CF548A3AFFD0E173FF07C982FE18FB94.taxon	distribution	Distribution. Eastern China: Zhejiang.	en	Zhang, Yong-Qin, Wei, Guo-Hao, Yin, Zi-Wei (2024): Two new species of ant-loving beetles (Coleoptera: Staphylinidae: Pselaphinae) from Jiulong National Wetland Park, China. Zootaxa 5406 (2): 336-342, DOI: 10.11646/zootaxa.5406.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5406.2.6
CF548A3AFFD0E173FF07C982FE18FB94.taxon	etymology	Etymology. The name is a noun in apposition, taken from the type locality of the new species, i. e., Jiulong National Wetland Park.	en	Zhang, Yong-Qin, Wei, Guo-Hao, Yin, Zi-Wei (2024): Two new species of ant-loving beetles (Coleoptera: Staphylinidae: Pselaphinae) from Jiulong National Wetland Park, China. Zootaxa 5406 (2): 336-342, DOI: 10.11646/zootaxa.5406.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5406.2.6
CF548A3AFFD2E175FF07CCBBFD8DF875.taxon	description	(Figs 2, 3) Chinese common name: 梁氏丽Dzm	en	Zhang, Yong-Qin, Wei, Guo-Hao, Yin, Zi-Wei (2024): Two new species of ant-loving beetles (Coleoptera: Staphylinidae: Pselaphinae) from Jiulong National Wetland Park, China. Zootaxa 5406 (2): 336-342, DOI: 10.11646/zootaxa.5406.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5406.2.6
CF548A3AFFD2E175FF07CCBBFD8DF875.taxon	materials_examined	Type material (56 exx.). HOLOTYPE: CHINA: ♂: ‘ China: Zhejiang, Lishui, Jiulong National Wetland Park, 28 ° 23 ′ 4.86 ″ N, 119 ° 49 ’ 29.15 ” E, 50 m, 23. ix. 2023, fern leaves, Zhang, Wei, Ming & Jia leg. (浙江丽水九AE国NJ ae 地公园) ’ (SNUC). PARATYPES: CHINA: 25 ♂♂, 16 ♀♀, same collecting data as for holotype; 5 ♂♂, 4 ♀♀, same collecting data as for holotype, except ‘ 22. ix. 2023 ’ (浙江丽水九AE国NJae地公园) ’ (SNUC). 2 ♂♂, 3 ♀♀, ‘ China: Zhejiang, Lishui, Bihu To Hongyu Vill., Jiulong Wetland Park, 28 ° 23 ’ 6 ” N, 119 ° 49 ’ 29 ” E, 50 m, 2. vi. 2023, riverbank, Z – W. Yin leg. (丽水九AEae地公园) ’ (2 ♂♂, 2 ♀♀ in IZCAS, others in SNUC).	en	Zhang, Yong-Qin, Wei, Guo-Hao, Yin, Zi-Wei (2024): Two new species of ant-loving beetles (Coleoptera: Staphylinidae: Pselaphinae) from Jiulong National Wetland Park, China. Zootaxa 5406 (2): 336-342, DOI: 10.11646/zootaxa.5406.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5406.2.6
CF548A3AFFD2E175FF07CCBBFD8DF875.taxon	diagnosis	Diagnosis. Male. Body length approximately 2.2 mm. Head sub-rectangular at base; vertex with transverse sulcus between antennal tubercles and long mediobasal carina, vertexal foveae large and asetose; antennae elongate, lacking distinct club, antennomere 11 largest. Pronotum with distinct median and lateral longitudinal sulci, with complete transverse antebasal sulcus. Discal striae of elytra extending posteriorly to approximately apical 4 / 5 of elytral length, humeri denticulate. Mesotrochanter with pencil-like protuberance on ventral margin, mesotibia with large apical spine. Metatrochanter with large, axe-like projection. Tergite 1 (IV) with large cavity in posterior half, lateral setiferous patches composed of short setae. Aedeagus strongly asymmetric; median lobe with relatively small basal capsule and broad ventral stalk; dorsal lobe narrowed near middle; parameres fused to broad membrane. Female. Body length approximately 2.2 mm, legs and abdomen lacking modifications, genitalia as in Fig. 2 J.	en	Zhang, Yong-Qin, Wei, Guo-Hao, Yin, Zi-Wei (2024): Two new species of ant-loving beetles (Coleoptera: Staphylinidae: Pselaphinae) from Jiulong National Wetland Park, China. Zootaxa 5406 (2): 336-342, DOI: 10.11646/zootaxa.5406.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5406.2.6
CF548A3AFFD2E175FF07CCBBFD8DF875.taxon	description	Description. Male. Body (Fig. 2 A) length 2.20 – 2.25 mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface covered with long pubescence. Head (Fig. 2 B) sub-rectangular, rounded at base, much wider than long, length 0.44 – 0.45 mm, width across eyes 0.51 – 0.50 mm; vertex finely punctate at middle, area anterior to eyes coarsely punctate, with large asetose vertexal foveae (dorsal tentorial pits) and curved transverse sulcus between antennal tubercles, distinct mediobasal carina extending from head base anteriorly to sulcus, antennal tubercles moderately raised; frons slightly impressed medially, confluent with clypeus; clypeus with carinate and moderately raised anterior margin, both frons and clypeus covered with distinct punctures; ocular-mandibular carinae complete, branched below eye, obliquely extended to ventral surface. Venter with tiny gular foveae (posterior tentorial pits) in single pit, with distinct median carina extending from pit anteriorly to mouthparts. Eyes moderately prominent, each composed of approximately 28 – 30 ommatidia. Antenna moderately elongate, length 0.99 – 1.00 mm; lacking distinct club; antennomere 1 thick, subcylindrical, 2 much smaller than 1, elongate, 3 slightly longer than 4, antennomeres 4, 6 and 8 small, of similar size, 5 and 7 slightly larger than 8, 9 – 11 each moderately enlarged, 11 largest, approximately as long as antennomeres 9 and 10 combined (1: 1), sub-fusiform. Pronotum (Fig. 2 B) approximately as long as wide, length 0.48 – 0.49 mm, width 0.49 – 0.50 mm, widest at middle; lateral margins rounded, convergent basally, parallel in basal 1 / 5; disc moderately convex, finely punctate, with median longitudinal sulcus as long as semi-circular lateral sulci in dorsal view; lacking median antebasal fovea, with complete, transverse antebasal sulcus connecting lateral antebasal foveae; small outer and inner pair of basolateral foveae present. Prosternum with anterior part slightly longer than coxal part, with small lateral procoxal foveae; thick hypomeral grooves extending from base to middle of anterior part, antebasal portion impressed, hypomeral ridges close to coxal cavities. Elytra much wider than long, length 0.65 – 0.67 mm, width 0.80 – 0.82 mm; roundly trapezoidal, dorsal surface with dense, long pubescence; each elytron with two large, asetose, basal foveae; discal striae extending from outer basal foveae posteriorly for 4 / 5 elytral length; humeri denticulate, subhumeral foveae present, carinate marginal striae extending from foveae to posterior margin of elytra. Metathoracic wings fully developed. Mesoventrite short, demarcated from metaventrite by transverse carina; median mesoventral foveae widely separated, in shared transverse opening, lateral mesoventral foveae large and setose, forked, posterior pair of forks much broader than anterior pair; marginal striae complete. Metaventrite slightly prominent, weakly impressed at middle, with well-developed lateral mesocoxal and two lateral metaventral foveae; posterior margin with small and narrow split at middle. Legs moderately elongate; foreleg simple; mesotrochanter (Fig. 2 C) with pencil-like extension on ventral margin, apex of extension with thin tuft of setae; mesotibia with distinct, subtriangular apical spine; metatrochanter (Fig. 2 D) with large, axe-like projection on ventral margin; each tarsus with one major and one reduced setiform pretarsal claw. Abdomen widest at mesal margins of tergite 1 (IV), length 0.67 – 0.69 mm, width 0.71 – 0.73 mm. Tergite 1 (IV) (Fig. 2 E) greatly modified, approximately three times as long as 2 – 4 (V – VII) combined; with large central cavity in posterior half, with oval median nodule at posterior margin of cavity, with three pairs (lateral, medial, anteromedial) of setose tufts inside cavity, with large setiferous patches lateral to the cavity, basal sulcus separated by mediobasal and large basolateral foveae, with short, thin triangular discal carinae; tergites 2 and 3 (V and VI) lacking foveae, 4 (VII) longer than 2 and 3 combined along middle, with one pair of basolateral foveae, tergite 5 (VIII) semicircular, transverse, posterior margin roundly emarginate at middle. Sternite 2 (IV) with two mediobasal foveae, and two pairs of basolateral foveae in short sulci, with pair of short lateral carinae; midlength of sternites 3 – 5 (V – VII) short, lacking foveae, sternite 6 (VIII) transverse, posterior margin broadly and roundly prominent at middle; sternite 7 (IX) (Fig. 2 F) with moderately sclerotized, rounded apex and few long setae along apical margin. Aedeagus (Fig. 2 G, H) 0.40 mm in length, dorso-ventrally asymmetric; median lobe with relatively small basal capsule and roundly triangular basal foramen, ventral stalk broad, trilobed, lateral lobes curved and longer than subtriangular one; dorsal lobe broadened near base, narrowing apically and with small hook-like structure near apex; parameres fused and reduced to ventral membrane. Female. Similar to male in external morphology; legs and abdomen lacking spines / projections or modifications; each compound eye composed of approximately 28 ommatidia. Measurements (as for male): body length 2.16 mm; length / width of head 0.42 – 0.45 / 0.51 mm, pronotum 0.47 – 0.48 / 0.48 – 0.49 mm, elytra 0.61 – 0.63 / 0.78 – 0.79 mm; abdomen 0.64 – 0.66 / 0.70 – 0.74 mm; length of antenna 0.93 – 0.98 mm; genital complex (Fig. 2 I) 0.30 mm wide, genital plate approximately as wide as sternite 7 (IX), lateral arms broadened through whole length. Comparative notes. This species is morphologically similar to a number of congeners from Japan (Nomura 1991; Jałoszyński 2023) and A. choui Yin from Taiwan (Yin 2018) in sharing a similar abdominal cavity on male tergite 1 (IV); and these species may form a monophyletic complex. Arthromelodes lianghongbini sp. nov. may be easily separated by the unique, broad and trilobed median lobe of the aedeagus. Arthromelodes tokushigei Jałoszyński from Okinawa, Japan is a wingless species, and A. choui Yin of Taiwan has more elongate antennae, and a modified metaventrite of the male. The genital complex (Jałoszyński 2023: fig. 41) of A. tokushigei has sternite 7 (IX) with a pair of long lateral extensions, and the genital plate is short, only about half as wide as sternite 7, with slender lateral arms.	en	Zhang, Yong-Qin, Wei, Guo-Hao, Yin, Zi-Wei (2024): Two new species of ant-loving beetles (Coleoptera: Staphylinidae: Pselaphinae) from Jiulong National Wetland Park, China. Zootaxa 5406 (2): 336-342, DOI: 10.11646/zootaxa.5406.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5406.2.6
CF548A3AFFD2E175FF07CCBBFD8DF875.taxon	biology_ecology	Biology. All individuals were collected from sifted fern leaf samples along or near the riverbank (Fig. 3).	en	Zhang, Yong-Qin, Wei, Guo-Hao, Yin, Zi-Wei (2024): Two new species of ant-loving beetles (Coleoptera: Staphylinidae: Pselaphinae) from Jiulong National Wetland Park, China. Zootaxa 5406 (2): 336-342, DOI: 10.11646/zootaxa.5406.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5406.2.6
CF548A3AFFD2E175FF07CCBBFD8DF875.taxon	distribution	Distribution. Eastern China: Zhejiang.	en	Zhang, Yong-Qin, Wei, Guo-Hao, Yin, Zi-Wei (2024): Two new species of ant-loving beetles (Coleoptera: Staphylinidae: Pselaphinae) from Jiulong National Wetland Park, China. Zootaxa 5406 (2): 336-342, DOI: 10.11646/zootaxa.5406.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5406.2.6
CF548A3AFFD2E175FF07CCBBFD8DF875.taxon	etymology	Etymology. This species is named in honor of Hong-Bin Liang, who led the project and provided various kinds of support during our field work.	en	Zhang, Yong-Qin, Wei, Guo-Hao, Yin, Zi-Wei (2024): Two new species of ant-loving beetles (Coleoptera: Staphylinidae: Pselaphinae) from Jiulong National Wetland Park, China. Zootaxa 5406 (2): 336-342, DOI: 10.11646/zootaxa.5406.2.6, URL: http://dx.doi.org/10.11646/zootaxa.5406.2.6
