taxonID	type	description	language	source
0643895EE9D85CF339494D346F2C0978.taxon	description	Description. Body narrowly elongate (Figs 1 - 6, 9 - 12) (broader in subalatus group, Figs 7 - 8); glabrous dorsally; color ranging from light to dark reddish-brown, often with yellow and / or piceous markings on elytra. Surface varying from smooth, shining, with very fine microsculpture (visible only under high magnification) to dull with dense, moderately fine, visible microsculpture. Length 6 - 13 mm. Head moderately sparsely to moderately densely, evenly punctate on vertex; punctures separated by average distance equal to or slightly greater than diameter of a puncture. Eyes moderate in size, separated by distances ranging from equal to or slightly greater than diameter of eye (OI ranging from 30 to 47); with distinct, well-developed nuchal-constriction (Figs 2, 4). Maxillary palpus (Fig. 16) with apical segment broadly securiform; apex subequal in length to outer side; mandible with apex shallowly notched medially. Antennae narrowly elongate (Figs 1 - 6); antennomere 2 very short, antennomere 3 much longer than 2, slightly shorter than to slightly longer than 4; 4 - 10 each elongate, at least two times longer than wide; sides only slightly widened from base to apex; antennal sensoriae small, visible only under high magnification, evenly distributed on segments 4 - 11. Pronotum with base distinctly narrower than base of elytra (Figs 1 - 10); sides variable, ranging from evenly narrowed from base to near apex to widest near middle and curved both anteriad and posteriad; width greater than length (PI ranging from 67 to 95); anterior margin truncate to slightly convex; anterior angles distinct, narrowly rounded. Basal foveae small, moderately deeply impressed, connected across base of pronotum by distinct transverse, prebasal groove. Prosternum elongate (Figs 17 - 18), horizontal anteriad of procoxae, prosternal process evenly rounded, abruptly declivous anteriad and posteriad of procoxae. Mesoventrite elongate (Fig. 17), distinctly more elongate than mesocoxal cavities except in species of subalatus group (Fig. 18); with shallow to moderately deep, V-shaped mesoventral cavity; intercoxal process gradually sloped to prosternum. Metaventrite with surface finely, sparsely to moderately coarsely punctate; disc distinctly more elongate between coxae than length of mesocoxal cavities (except in species of subalatus group). Third and fourth segments of anterior and intermediate tarsi and penultimate segment of posterior tarsi lobed ventrally, in addition, basal two segments of protarsus lobed ventrally in male (except Charisius salvini). Elytra elongate (Figs 1 - 6, 9 - 12); sides parallel for basal half; then evenly narrowed to apex except Charisius apterus and Charisius subalatus (Figs 7 - 8); striae moderately shallowly impressed near base, becoming more deeply impressed approaching apex (striae unimpressed between strial punctures in Charisius howdenorum (Fig. 19); strial interstices usually convex or rarely flat; impunctate or with a row of fine, median punctures visible only under high magnification. Elytral epipleurae ending just before apex of elytra; evenly arched from base to apex. Ventrites finely, sparsely punctate or impunctate. Male: Eighth sternite divided into two large, well developed lobes (Figs 21 - 24); apex of each lobe appearing glabrous, actually bearing small, densely placed, dentiform setae which extend along inner margin to near base (visible only under high magnification). Ninth sternite bilobed; lobes small, not joined medially, reaching only to base of eighth sternal lobes. Apicale of aedeagus variable (Figs 25 - 28); ranging from 2.8 to 3.5 times as long as basale (apicale very short in salvini group, basale 6.9 to 7.9 times longer than apicale).	en	Campbell, J. M. (2014): New species and records of Charisius Champion from Mexico and Central America (Coleoptera, Tenebrionidae, Alleculinae). ZooKeys 415: 269-293, DOI: http://dx.doi.org/10.3897/zookeys.415.6794, URL: http://dx.doi.org/10.3897/zookeys.415.6794
9A21FCC9F7C946DE89F900FA9DA3AE39.taxon	description	Description. Dark reddish-brown; elytra yellow with broad basal, large medial, and smaller V-shaped piceous markings (Fig. 6). Length 9.9 - 10.9 mm. Head coarsely, densely, evenly punctate; punctures separated by average distance less than diameter of a puncture. Eyes moderate in size, mean ocular index of 5 specimens 35.4 (30 - 40). Pronotum distinctly wider than long, mean pronotal index of 5 specimens 82.6, ranging from 81 to 84; surface microsculpture densely, coarsely granulate, opaque; punctures coarse (Fig. 15), moderately dense, separated on center of disc by average distance equal to or slightly greater than diameter of a puncture, punctures becoming finer and distinctly more widely separated on sides of disc; sides straight, subparallel from base to apical fourth then convexly narrowed to apex; transverse groove broad, moderately deeply impressed, disc shallowly, but distinctly impressed along midline. Prosternum and hypomeron with a few widely scattered, moderately coarse punctures. Metaventrite normally elongate, length between meso- and metacoxae distinctly longer than length of mesocoxal cavity; surface finely, moderately sparsely punctate medially, punctures becoming coarser approaching sides. Ventrites with punctures fine; last two ventrites slightly more coarsely and densely punctate. Elytra with striae moderately impressed basally, becoming deeply impressed towards apex; strial interstices convex. Male. Anterior tibiae not sexually modified; anterior tarsal claws each with 7 - 9 teeth. Ventrite five not impressed medially. Lobes of eighth sternum (Fig. 21) broad, slightly curved medially, apices broadly, evenly rounded; apical and inner margins with row of very fine, short, dentiform setae; viewed laterally, lobes only slightly deflexed. Lobes of ninth sternum short, moderately broad, with apices evenly convex. Aedeagus with apicale (Fig. 25) narrow, with sides narrowed from base to narrowly rounded apex; sides moderately strongly sinuate just before apex. Female. Anterior tarsal claws each with 7 teeth. Elytra with sutural margin and apex entire.	en	Campbell, J. M. (2014): New species and records of Charisius Champion from Mexico and Central America (Coleoptera, Tenebrionidae, Alleculinae). ZooKeys 415: 269-293, DOI: http://dx.doi.org/10.3897/zookeys.415.6794, URL: http://dx.doi.org/10.3897/zookeys.415.6794
9A21FCC9F7C946DE89F900FA9DA3AE39.taxon	etymology	Etymology. The species name granulatus is derived from the unique granulate microsculpture on the pronotal disc.	en	Campbell, J. M. (2014): New species and records of Charisius Champion from Mexico and Central America (Coleoptera, Tenebrionidae, Alleculinae). ZooKeys 415: 269-293, DOI: http://dx.doi.org/10.3897/zookeys.415.6794, URL: http://dx.doi.org/10.3897/zookeys.415.6794
006C05AFE3A3B7B39DCCA24E9B759D5A.taxon	description	Description. Dark reddish-brown; elytron yellow with extensive brunneous markings as follows: moderately broad, transverse bands across basal fourth and across middle; narrow, jagged band across apical fourth and extreme apical portion; suture brunneous (Fig. 7). Length 9.8 mm. Head coarsely, densely punctate, punctures separated by distance distinctly less than diameter of a puncture. Eyes small, widely separated dorsally, ocular index of female holotype 47. Pronotum slightly wider than long, PI of holotype 88; surface with microsculpture finely, uniformly granulate, not shining; moderately densely and coarsely punctate, punctures separated by distance slightly greater than diameter of a puncture; punctures evenly distributed except becoming sparser on narrow band at anterior lateral angles of pronotum; sides distinctly convex, widest at apical third, distinctly narrowed towards base; transverse groove broad, shallowly impressed; midline broadly, shallowly impressed on basal half. Prosternum and hypomeron sparsely, finely punctate; postcoxal extension of sides of pronotum with row of coarse, moderately shallow impressions. Metaventrite short, length between meso- and metacoxae subequal in length to length of mesocoxal cavity; moderately coarsely and densely, evenly punctate. Abdomen with basal three ventrites with a few scattered, fine punctures; apical two ventrites more densely punctate with punctures separated by distance approximately two to three times diameter of puncture. Elytra 3.2 times longer than pronotum (in unique holotype) with striae shallowly, evenly impressed throughout; strial punctures coarse, narrowly separated along striae; intervals slightly convex. Wings reduced, flightless. Male. Unknown. Female. Anterior tarsal claws each with 6 teeth. Elytra with sutural margin near apex and apex emarginate as in Fig. 20. Fifth visible sternite broadly, shallowly impressed medially.	en	Campbell, J. M. (2014): New species and records of Charisius Champion from Mexico and Central America (Coleoptera, Tenebrionidae, Alleculinae). ZooKeys 415: 269-293, DOI: http://dx.doi.org/10.3897/zookeys.415.6794, URL: http://dx.doi.org/10.3897/zookeys.415.6794
006C05AFE3A3B7B39DCCA24E9B759D5A.taxon	etymology	Etymology. The species name apterus is derived from the species having the flight wings reduced to short stubs.	en	Campbell, J. M. (2014): New species and records of Charisius Champion from Mexico and Central America (Coleoptera, Tenebrionidae, Alleculinae). ZooKeys 415: 269-293, DOI: http://dx.doi.org/10.3897/zookeys.415.6794, URL: http://dx.doi.org/10.3897/zookeys.415.6794
25E6D1AD9F3C417EC061D50F84AEF8A9.taxon	description	Description. Uniformly reddish-brown; without elytral markings (Fig. 8). Length 7.6 - 8.7 mm. Head coarsely, densely, evenly punctate on vertex; punctures separated by average distance less than diameter of a puncture. Eyes moderate in size, widely separated dorsally; mean OI of 9 specimens 45.2 (range 42 - 48). Pronotum slightly wider than long, average PI of 9 specimens 90.5 (range 87 - 96); with surface very finely granulate, opaque; finely, shallowly, sparsely punctate, punctures separated by average distance at least two times diameter of a puncture; punctures evenly distributed over most of disc except becoming nearly impunctate near sides; sides distinctly curved, widest across middle; with midline shallowly impressed in basal half. Prosternum and hypomeron with a few widely scattered, moderately coarse punctures; postcoxal extension of sides of pronotum with a few coarse, moderately shallow impressions. Metaventrite (Fig. 18) short, length between meso- and metacoxae shorter than length of mesocoxal cavity; moderately coarsely, densely, contiguously punctate medially in males, punctures becoming coarser and sparser approaching sides; coarsely punctate medially in female with punctures distinctly separated, punctures becoming coarser approaching sides. Ventrites finely, moderately sparsely punctate. Elytra 3.1 - 3.3 times longer than pronotum; striae shallowly, evenly impressed throughout (in specimens from Miramundo, Guatemala and El Salvador striae completely unimpressed between punctures); strial punctures coarse, narrowly separated along striae; intervals flat. Wings reduced, distinctly shorter than elytra; flightless. Male. Anterior femora with patch of fine, dense pubescence on middle of ventral margin. Anterior tibiae slightly widened near middle on inner margin. Fifth ventrite unimpressed medially. Eighth sternal lobes (Fig. 22) broad, only slightly curved medially and distinctly curved ventrally near apices; apices of lobes narrowly rounded; viewed laterally, apices of lobes distinctly deflexed. Lobes of ninth sternum short, broad, with apices almost truncate. Aedeagus with apicale (Fig. 26) moderately narrow, with sides converging from base to near apex and distinctly constricted just before apex; apex narrowly rounded; viewed laterally, apicale slightly curved dorsally with apex very narrowly rounded; basale 2.8 to 2.9 times longer than apicale. Female. Elytra with sutural margin and apex entire.	en	Campbell, J. M. (2014): New species and records of Charisius Champion from Mexico and Central America (Coleoptera, Tenebrionidae, Alleculinae). ZooKeys 415: 269-293, DOI: http://dx.doi.org/10.3897/zookeys.415.6794, URL: http://dx.doi.org/10.3897/zookeys.415.6794
46F36BA3CDCFCF92B847F37CD5BEE81D.taxon	description	Description. Dark reddish-brown to dark brown (Fig. 10); antennae and legs slightly paler than body; elytra without markings. Length 6.5 - 8.6 mm. Head moderately coarsely, densely, evenly punctate on vertex; punctures separated by average distance less than diameter of a puncture. Eyes moderate in size, mean OI of 10 specimens 43.9 (range 41 - 47). Pronotum distinctly wider than long, mean PI of 10 specimens 82.6, ranging from 77 to 86; surface with microsculpture moderately coarse, granulate, only slightly shining; punctures moderately coarse, moderately dense, separated on center of disc by average distance equal to or slightly greater than diameter of a puncture, punctures finer, distinctly sparser on sides; sides straight or slightly sinuate on basal two-thirds, then evenly, convexly narrowed to apex; con vex, widest near middle then evenly narrowed to base and apex; disc often faintly impressed along midline. Prosternum and hypomeron moderately sparsely, coarsely, irregularly punctate; punctures separated by distance greater than diameter of a puncture. Metaventrite finely, moderately densely punctate medially in male; punctures becoming coarser approaching sides; finely and sparsely punctate medially in female. Abdomen with fine, scattered punctures on basal three ventrites, last two visible ventrites more coarsely and slightly more densely punctate. Elytra (Fig. 19) with striae unimpressed; strial interstices flat. Male. Anterior tibia not widened on inner side. Fifth ventrite unimpressed medially. Lobes of eighth sternum (Fig. 23) broad, straight, apices evenly convex; viewed laterally, lobes slightly deflexed apically. Lobes of ninth sternum short, moderately broad, with apices broadly convex. Aedeagus with apicale (Fig. 27) moderately narrow; sides narrowed from base to near middle then subparallel to evenly convex apex; viewed laterally, apicale straight, basale 3.2 - 3.3 times longer than apicale. Female. Elytra with sutural margin and apex entire.	en	Campbell, J. M. (2014): New species and records of Charisius Champion from Mexico and Central America (Coleoptera, Tenebrionidae, Alleculinae). ZooKeys 415: 269-293, DOI: http://dx.doi.org/10.3897/zookeys.415.6794, URL: http://dx.doi.org/10.3897/zookeys.415.6794
46F36BA3CDCFCF92B847F37CD5BEE81D.taxon	etymology	Etymology. This species is named in honor of Dr. Henry and Anne Howden, Canadian Museum of Nature, Alymer, Quebec, Canada who have facilitated several of my trips to Mexico and Guatemala; and have collected many of the specimens described in this paper.	en	Campbell, J. M. (2014): New species and records of Charisius Champion from Mexico and Central America (Coleoptera, Tenebrionidae, Alleculinae). ZooKeys 415: 269-293, DOI: http://dx.doi.org/10.3897/zookeys.415.6794, URL: http://dx.doi.org/10.3897/zookeys.415.6794
81042338C0C58984B9BE99E3E10A7207.taxon	description	Description. Body reddish-brown; elytra testaceus with each elytron having piceous markings as follows (Fig. 12): a small circle near middle, a narrow, crescent-shaped band at apical fourth and extreme apex piceous to black (Fig. 12). Length 7.0 - 10.9 mm. Head moderately coarsely punctate on vertex; punctures separated by distance subequal to diameter of a puncture. Eyes moderately small, mean OI of 5 males 32.7 (range 30 - 35) and of five females 38.6 (range 37 - 40). Pronotum distinctly wider than long, mean PI of 11 specimens 84.6 (range 83 to 88); surface smooth, shining; punctures coarse, moderately densely, evenly distributed, separated by average distance distinctly greater than diameter of a puncture; sides sinuate near base, widest near middle than evenly convex to just before apex; disc evenly convex in cross section. Prosternum and hypomeron coarsely, densely punctate; punctures separated by distance less than diameter of a puncture. Metaventrite finely, sparsely punctate medially, punctures becoming coarser and denser approaching sides. Ventrites with punctures moderately fine, sparsely distributed; last two visible ventrites more coarsely and densely punctate. Elytra with striae moderately deeply, evenly impressed throughout; strial interstices moderately convex. Male. Anterior tibia triangularly widened on inner side near middle. Fifth ventrite broadly, deeply, triangularly impressed medially. Lobes of eighth sternum (Fig. 24) broad, with apical third broadly expanded, apices obliquely truncate; viewed laterally, distinctly deflexed apically. Lobes of ninth sternum short, moderately broad, with apices evenly convex. Aedeagus with apicale (Fig. 28) short, moderately narrow; sides narrowed from base to narrowly, evenly convex apex, slightly constricted medially; viewed laterally, apicale strongly curved dorsally; basale 7.1 - 7.9 times longer than apicale. Female. Elytra with sutural margin and apex entire.	en	Campbell, J. M. (2014): New species and records of Charisius Champion from Mexico and Central America (Coleoptera, Tenebrionidae, Alleculinae). ZooKeys 415: 269-293, DOI: http://dx.doi.org/10.3897/zookeys.415.6794, URL: http://dx.doi.org/10.3897/zookeys.415.6794
81042338C0C58984B9BE99E3E10A7207.taxon	etymology	Etymology. The species name punctatus refers to the coarse punctation of the pronotum.	en	Campbell, J. M. (2014): New species and records of Charisius Champion from Mexico and Central America (Coleoptera, Tenebrionidae, Alleculinae). ZooKeys 415: 269-293, DOI: http://dx.doi.org/10.3897/zookeys.415.6794, URL: http://dx.doi.org/10.3897/zookeys.415.6794
