taxonID	type	description	language	source
0D3AF096430D50118E26907C30E8D0C7.taxon	diagnosis	Diagnosis. Uniformly coloured brownish with smokey yellowish tegmina compared to the similar B. sandrangatensis which has a red head, thorax and base of tegmina. It also differs from this species by the length of its subgenital plates, 1.18 times longer than its pygofer height compared to 1.48 for B. sandrangatensis.	en	Bouteille, Emilien, Cesne, Maxime Le, Soulier-Perkins, Adeline (2021): Cercopidae spittle-bugs (Hemiptera, Cicadomorpha) of Madagascar: a new species of Bourgoinrana and revision of the Locris species. ZooKeys 1023: 29-47, DOI: http://dx.doi.org/10.3897/zookeys.1023.58733, URL: http://dx.doi.org/10.3897/zookeys.1023.58733
0D3AF096430D50118E26907C30E8D0C7.taxon	distribution	Distribution. Mount Beondroka in the natural reserve of Marojejy, Madagascar (Fig. 9 E).	en	Bouteille, Emilien, Cesne, Maxime Le, Soulier-Perkins, Adeline (2021): Cercopidae spittle-bugs (Hemiptera, Cicadomorpha) of Madagascar: a new species of Bourgoinrana and revision of the Locris species. ZooKeys 1023: 29-47, DOI: http://dx.doi.org/10.3897/zookeys.1023.58733, URL: http://dx.doi.org/10.3897/zookeys.1023.58733
0D3AF096430D50118E26907C30E8D0C7.taxon	description	Description. Total length of male holotype 7.9 mm (tegmina included), paratypes 8.4 and 8.6 mm. Flattened ventro-dorsally. Head in dorsal view, 1.6 times wider between eyes than long in midline, anterior and posterior margins gently and regularly curved. Ocelli very close to each other with distance between eye and ocellus 9 times greater than between ocelli, located close to head posterior margin. Pronotum slightly convex, 1.8 times wider than long in midline, posterior margin wave-shaped, concave in middle. Tegmina 3.8 times longer than wide, M and CuA with a common stem at base and forking around 1 / 3 of tegmen length, ScP + R (+ MA) forking after mid-length of tegmen, M and CuA both forking in apical third of tegmen. Metatibia bearing 1 lateral spine at 2 / 3 of its length.	en	Bouteille, Emilien, Cesne, Maxime Le, Soulier-Perkins, Adeline (2021): Cercopidae spittle-bugs (Hemiptera, Cicadomorpha) of Madagascar: a new species of Bourgoinrana and revision of the Locris species. ZooKeys 1023: 29-47, DOI: http://dx.doi.org/10.3897/zookeys.1023.58733, URL: http://dx.doi.org/10.3897/zookeys.1023.58733
0D3AF096430D50118E26907C30E8D0C7.taxon	description	Description of the male terminalia (Fig. 9 A-D). Pygofer height 1.6 mm (Fig. 9 A) in lateral view, dorsal margin almost straight, posterior margin almost straight 2 / 3 of its dorsal length then curving strongly anteriorly before having a final straight section, ventral margin straight and shorter than dorsal margin, anterior margin S-shaped. Subgenital plates 1.18 longer than pygofer height, gently and regularly curving dorsally with a rounded apex pointing dorso-posteriorly, thickness gradually diminishing from base to apex with a slight constriction before apex. Parameres in lateral view with a small hump in the middle of dorsal margin ending in a narrow elongate finger-shape apex slightly curved inward (Fig. 9 C, D), ventral margin ending in a finger-shape apex shorter and less narrow than dorsal one; in dorsal view, internal margin cut out in two small extension facing each other (Fig. 9 C). Lateral plates present with dorsal margin straight, making a rounded acute angle with the postero-ventral margin. Aedeagus long, shaped as a circumvented tube with a thumb-shaped extension on its dorsal margin, oriented dorso-anteriorly, small constriction before thumb extension; apical part bifid, prolonging ventral margin a lateral toothed extension folding posteriorly with apex pointing antero-ventrally and prolonging dorsal margin a thin ending in a lanceolate shape pointing anteriorly (Fig. 9 B).	en	Bouteille, Emilien, Cesne, Maxime Le, Soulier-Perkins, Adeline (2021): Cercopidae spittle-bugs (Hemiptera, Cicadomorpha) of Madagascar: a new species of Bourgoinrana and revision of the Locris species. ZooKeys 1023: 29-47, DOI: http://dx.doi.org/10.3897/zookeys.1023.58733, URL: http://dx.doi.org/10.3897/zookeys.1023.58733
0D3AF096430D50118E26907C30E8D0C7.taxon	etymology	Etymology. The species is named after the type locality, Beondroka.	en	Bouteille, Emilien, Cesne, Maxime Le, Soulier-Perkins, Adeline (2021): Cercopidae spittle-bugs (Hemiptera, Cicadomorpha) of Madagascar: a new species of Bourgoinrana and revision of the Locris species. ZooKeys 1023: 29-47, DOI: http://dx.doi.org/10.3897/zookeys.1023.58733, URL: http://dx.doi.org/10.3897/zookeys.1023.58733
C8A5D46397875A1CB267B1B54DE2FB9B.taxon	distribution	Distribution. Madagascar (Fig. 3 E), L. bipunctata bipunctata in northern Madagascar, L. bipunctata atra in western Madagascar.	en	Bouteille, Emilien, Cesne, Maxime Le, Soulier-Perkins, Adeline (2021): Cercopidae spittle-bugs (Hemiptera, Cicadomorpha) of Madagascar: a new species of Bourgoinrana and revision of the Locris species. ZooKeys 1023: 29-47, DOI: http://dx.doi.org/10.3897/zookeys.1023.58733, URL: http://dx.doi.org/10.3897/zookeys.1023.58733
C8A5D46397875A1CB267B1B54DE2FB9B.taxon	description	Description of the male terminalia (Fig. 3 A-D). Pygofer (Fig. 3 A), in lateral view almost trapezoidal with posterior margin being the longest side and slightly cut out in its first dorsal third. Aedeagus consists of two parts, first basal part, representing 2 / 3 of total length, tubular and almost same thickness over its entire length, curving regularly down then dorsally, and bearing on its apical dorsal part a small hump directed dorso-anteriorly; second part, hanging from first, narrowing in its middle then developing posteriorly a pair of pointed processes and finishing in a smooth and flattened apex slightly digit shaped ventro-posteriorly (Fig. 3 B). Parameres smoothly widening from pygofer's attachment and finishing posteriorly by a curved spine (Fig. 3 C). Subgenital plates large at base, narrowing quickly in a long and fine structure curving gently dorsally on its apical half (Fig. 3 D). Body length: 7.5 - 10 mm.	en	Bouteille, Emilien, Cesne, Maxime Le, Soulier-Perkins, Adeline (2021): Cercopidae spittle-bugs (Hemiptera, Cicadomorpha) of Madagascar: a new species of Bourgoinrana and revision of the Locris species. ZooKeys 1023: 29-47, DOI: http://dx.doi.org/10.3897/zookeys.1023.58733, URL: http://dx.doi.org/10.3897/zookeys.1023.58733
C8A5D46397875A1CB267B1B54DE2FB9B.taxon	materials_examined	Material examined. Nominotypical subspecies- 10 males. [Ambabalame, Madagascar], [Coll. Mus. Congo, Coll. V. Lallemand], [♂], [R. Det., 6664 H], [Museum Paris, MNHN (EH) 24628]; [Madagascar], [Lavaudon], [Collectic E de Bergev], [Museum Paris], [Museum Paris, MNHN (EH) 24115], [Museum Paris, MNHN (EH) 24625]; [Madagascar Nord, distr. d'Ambanja, N. de Beangona-Ambevy, Vallee d'Antremade, 400 m, II- 1964, P. Soga], [Museum Paris]; [Museum Paris, Sainte-Marie-de-Madagascar, Coll. Noualhier, 1898]; [Museum Paris, Madagascar, Baie d'Antongil, A. Mocquerys, Coll. Noualhier, 1898], [Museum Paris, MNHN (EH), 24720]; [Museum Paris, Madagascar, Baie d'Antongil, A. Mocquerys, Coll. Noualhier, 1898], [Locris bipunctata Sign.], [Museum Paris, MNHN (EH), 24722]; [Madagascar Nord, distr. d'Ambanja, N. de Beangona-Ambevy, Vallee d'Antremabe, 400, II - 1964, P. Songa], [Museum Paris], [Museum Paris, MNHN (EH), 24719]; [Coll. Mus. Tervuren, Madagascar: Fampanambo, 1962, J. Vadon], [Locris bipunctata Signoret, H. Synave det., 1965], [Locris bipunctata Sign ♂♂], [Museum Paris, MNHN (EH), 24721]; [Madagascar Amber-Geb.], [Locris vicina Sign. ♂ Edm. Schmidt determ. 1911], [Miz Pan Warszawa 12 / 1945, 2487]; [Madagascar Amber-Geb.], [Locris vicina Sign. ♂ Edm. Schmidt determ. 1911], [970], [Miz Pan Warszawa. 12 / 1945, 2488]; and [Madagascar Amber-Geb.], [Locris vicina Sign. ♂ Edm. Schmidt determ. 1911], [Miz Pan Warszawa. 12 / 1945, 2489]. Subspecies Locris vicina atra - Paratype (male). [Entre Ambakaka et Bekopaka sur la Manambolo, Inst. Scient. Madagascar, VII - 49, RP], [Museum Paris], [Paratype], [Museum Paris MNHN (EH) 1474], [Locris bipunctata v. atra Lall.], [Museum Paris MNHN (EH) 24634] - 5 males. [CASENT 3004533], [Madagascar: Mahajanga, Prov: Parc National Tsingy de Bemaraha, 3.4 km 93 ° E, Bekopaka, Tombeau Vazimba, Elev 50 m, 6 - 10 Nov. 2001], [19 ° 8 ' 31 " S, 44 ° 49 ' 41 " E, coll: Fischer, Griswold et al., California Acad. Of Sciences malaise trap, in tropical dry forest, coll. Code: BLF 4233], [♂], [Locris bipunctata atra Lallemand, 1950, A. Soulier-Perkins det 2018]; [Madagascar Ouest, S-P. Antsalova Antsingy, Res. Nat. 9, A. Peyrieras, I- 1975], [Museum Paris]; [Madagascar Lambomakandro, Tulear], [Museum Paris, 1935, B. Catala], [H. Synave det., 1979, Locris bipunctata var. atra Lall.]; [Andobo 190 m, foret Antsingy, det. Antsalova, II - 57, P. Guiv], [Institut Scientifique, Madagascar], [Museum Paris, MNHN (EH), 24718]; and [Madagascar, Lambomakandro, Tulear], [Museum Paris, 1935, R. Catala], [H. Synave., 1979, Locris bipunctata var. Locris bipunctata atra Stall.], [Museum Paris, MNHN (EH), 24717]; 2 females. [♀], [Madagascar, province de Toliara, massif du Makay, 159 m, 21 ° 40 ' 29.4 " S, 44 ° 59 ' 36.2 " E], [Museum Paris, ft Ambalamanga, rv Mangoky, 18 - I- 2011, A. Soulier-Perkins rec.], [Locris bipunctata atra Lallemand, 1950, A. Soulier-Perkins det. 2020], [Museum Paris, MNHN (EH), 24759]; [♀], [Madagascar, province de Toliara, massif du Makay, 159 m, 21 ° 40 ' 29.4 " S, 44 ° 59 ' 36.2 " E], [Museum Paris, Pres riviere, PL, 20 - I- 2011, A. Soulier-Perkins rec.], [Locris bipunctata atra Lallemand, 1950, A. Soulier-Perkins det. 2020], [Museum Paris, MNHN (EH), 24760].	en	Bouteille, Emilien, Cesne, Maxime Le, Soulier-Perkins, Adeline (2021): Cercopidae spittle-bugs (Hemiptera, Cicadomorpha) of Madagascar: a new species of Bourgoinrana and revision of the Locris species. ZooKeys 1023: 29-47, DOI: http://dx.doi.org/10.3897/zookeys.1023.58733, URL: http://dx.doi.org/10.3897/zookeys.1023.58733
755EF0FD9DBF5338877ED22EEC802D64.taxon	distribution	Distribution. Madagascar and eastern Africa	en	Bouteille, Emilien, Cesne, Maxime Le, Soulier-Perkins, Adeline (2021): Cercopidae spittle-bugs (Hemiptera, Cicadomorpha) of Madagascar: a new species of Bourgoinrana and revision of the Locris species. ZooKeys 1023: 29-47, DOI: http://dx.doi.org/10.3897/zookeys.1023.58733, URL: http://dx.doi.org/10.3897/zookeys.1023.58733
755EF0FD9DBF5338877ED22EEC802D64.taxon	description	Description of the male terminalia (Fig. 5). Pygofer (Fig. 5 A), in lateral view, dorsal margin straight and perpendicular to anterior margin, posterior margin generally convex. Aedeagus consist of two parts, first basal part, representing 3 / 4 of total length with a base elbow shaped before widening ventrally then curving up dorsally and narrowing into a tubular structure almost of same thickness to its regular rounded dorsal apex; second part, hanging from first, its width is regular for most of its length with apex in shape of a swan, neck oriented posteriorly (Fig. 5 B). Parameres prolonged apically by two little structures, inner one curved into a spine and external one rounded (Fig. 5 C). Subgenital plates wide at base then abruptly narrowing in a long filament shape curved downward at its apex (Fig. 5 D). Body length: 8 - 10.5 mm.	en	Bouteille, Emilien, Cesne, Maxime Le, Soulier-Perkins, Adeline (2021): Cercopidae spittle-bugs (Hemiptera, Cicadomorpha) of Madagascar: a new species of Bourgoinrana and revision of the Locris species. ZooKeys 1023: 29-47, DOI: http://dx.doi.org/10.3897/zookeys.1023.58733, URL: http://dx.doi.org/10.3897/zookeys.1023.58733
755EF0FD9DBF5338877ED22EEC802D64.taxon	materials_examined	Material examined. Holotype (male). [Holotypus], [Madagascar], [Coll. Mus. Congo, Coll. V. Lallemand], [R. DET. 6664 z z], [Locris johannae ssp. nigrolimbata Lall.], [H. Synave det .. 195 L. johannae f. nigrolimbata Lall.], [Holotype ♂ L. johannae ssp. nigrolimbata Lall. Michel Boulard det. 1989] - 4 males. [Afrique orient. Anglaise Voi Alluaud & Jeannel Mars 1911. 600 m. St. 60], [Coll. Mus. Congo Coll. V. Lallemand], [R. Det. 6665]; [Afr. Or. Angl. (Wa-Kikuyu), Fort-Hall Alluaud & Jeannel Janv. 1912 - 1330 m - St. 80], [Coll. Mus. Congo Coll. V. Lallemand], [R. Det. 6665]; [♂], [Nairobi B. E. A.], [G. Babault, avril 1923], [Locris johannae ssn. Locris nigrolimbata], [Museum Paris, MNHN (EH), 24629]; and [Tanzanie: Mts Uluguru, Kimboza ofr. Heliophile, alt. 600 M 24 - 30 / VII / 71], [Coll. Mus. Tervuren, Mission Mts. Uluguru, L. Berger, N. Leleup, J. Debecker V / VIII / 71], [H. Synave det. 1957, Locris johannae Lall.], [Museum Paris, MNHN (EH), 24635].	en	Bouteille, Emilien, Cesne, Maxime Le, Soulier-Perkins, Adeline (2021): Cercopidae spittle-bugs (Hemiptera, Cicadomorpha) of Madagascar: a new species of Bourgoinrana and revision of the Locris species. ZooKeys 1023: 29-47, DOI: http://dx.doi.org/10.3897/zookeys.1023.58733, URL: http://dx.doi.org/10.3897/zookeys.1023.58733
91E7A29EF6305C338AA67E80FF872994.taxon	distribution	Distribution. Madagascar (Fig. 7 E)	en	Bouteille, Emilien, Cesne, Maxime Le, Soulier-Perkins, Adeline (2021): Cercopidae spittle-bugs (Hemiptera, Cicadomorpha) of Madagascar: a new species of Bourgoinrana and revision of the Locris species. ZooKeys 1023: 29-47, DOI: http://dx.doi.org/10.3897/zookeys.1023.58733, URL: http://dx.doi.org/10.3897/zookeys.1023.58733
91E7A29EF6305C338AA67E80FF872994.taxon	description	Description of the male terminalia (Fig. 7 A-D). Pygofer (Fig. 7 A) in lateral view, dorsal and anterior margins perpendicular, posterior margin S-shaped and making an acute angle with the ventral margin. Aedeagus consists of two parts, first basal part, representing a small 2 / 3 of total length with a very regular tubular shape almost curving as half a circle, its dorsal apical part bearing two little bumps oriented dorso-anteriorly; second part is hanging from first, it is widening smoothly, two processes are pointing posteriorly at mid-length, apex bulbous and covered in a dense padding (Fig. 7 B), genital duct passes through aedeagus and open in the centre of padded apex (Fig. 7 B). Parameres with a dorsal margin S-shaped giving it a cup shape prolonged apically by a spine curved downward (Fig. 7 C). Subgenital plates wide at base and narrowing abruptly in a long filament structure curving abruptly dorsally on last third (Fig. 7 D). Body length: 10.5 - 12.5 mm.	en	Bouteille, Emilien, Cesne, Maxime Le, Soulier-Perkins, Adeline (2021): Cercopidae spittle-bugs (Hemiptera, Cicadomorpha) of Madagascar: a new species of Bourgoinrana and revision of the Locris species. ZooKeys 1023: 29-47, DOI: http://dx.doi.org/10.3897/zookeys.1023.58733, URL: http://dx.doi.org/10.3897/zookeys.1023.58733
91E7A29EF6305C338AA67E80FF872994.taxon	materials_examined	Material examined. MNHN Collection- 1 female [♀], [Madagascar. Tam.], [Soanierana-Ivongo, 8. XI. 57, F. Keiser], [R. I. Sc. N. B., I. G. 23.285], [Museum Paris, MNHN (EH), 24723]; - 10 males. [20 / III / 2006, 633 m, parc de Zombitse Leobondro, brd riviere, 22 ° 40.460 ' S, 44 ° 51.633 ' E], [Museum Paris, Madagascar 2006, reg. Atsimo-andrefana, A. Soulier-Perkins rec.], [Locris vicina (Signoret, 1860), A. Soulier-Perkins det. 2007], [♂], [Museum Paris, MNHN (EH), 24659]; [♂], [Museum Paris Madagascar, Antanambe, Mocquerys, Coll. Noualhier 1898]; [♂], [Museum Paris, Madagascar, catat 1844 - 91]; [Madagascar Nord, distr. d'Ambanja, N. de Beangona-Ambevy, Vallee d'Antremabe, 400 m, II - 1964, P. Soga], [Mueum Paris], [Museum Paris, MNHN (EH), 24724]; [Fort. Dauphin], [Museum Paris, 1933, A. Seyrig], [Museum Paris, MNHN (EH), 24725]; [Maeyatanana, Madagscar], [Collection le Moult, Naturaliste. Paris], [R. Mus. Hist. Nat., Belg. I. G. 12.595], [Museum Paris, MNHN (EH), 24726]; [Madagascar, province de Mahajanga, P N Tsingy de Namoroka, 16 ° 28 ' 08 " S, 45 ° 20 ' 52 " E], [Museum Paris, 09 - IX- 2012, Th. Bourgoin rec.], [Museum Paris, MNHN (EH), 24727]; [21 / XI / 2005, region lac Alaotra, bord riv. Mavolava, 17 ° 40.357 ' S, 47 ° 54.289 ' E], [904 m, entre Ambakireny et Morano-Chrome], [Museum Paris, Madagascar 2005, Bourgoin, Ouvrard, Attie, Soulier-Perkins], [Locris vicina (Signoret, 1860), A. Soulier-Perkins det. 2007], [♂], [Museum Paris, MNHN (EH), 24728]; [Madagascar Mahajanga, P. N. Tsingy de Namoroka, wetland near village, 16 ° 23 ' 50 " S, 45 ° 17 ' 12 " E], [Museum Paris, 29 - X- 2016, PL., T. Bourgoin, G. Kunz & A. Soulier-Perkins rec.], [Museum Paris, MNHN (EH), 24729] - RIScNB [Ampijoro, Tsaramandroso], [Institut scientifique, Madagascar], [H. Synave det., 1957, Locris vicina Signoret], [R. I. Sc. N. B., I. G. 21.002]; MIIZ collection - 5 males [Madagasckar, Ambodimanga, Hammerstem S., I. 1906], [Locris bipunctata Sign. ♂ Edm. Schmidt determ. 1909], [Miz Pan Warszawa. 12 / 1945, 2490]; [Madagaskar, Ambodimanga, Hammerstem S., I. 1906], [Locris bipunctata Sign. ♂ Edm. Schmidt determ. 1909], [Miz Pan Warszawa. 12 / 1945, 2491]; [Madagaskar, Ambodimanga, Hammerstem S., I - II. 1906], [Locris bipunctata Sign. ♂ Edm. Schmidt determ. 1909], [Miz Pan Warszawa. 12 / 1945, 2492]; [Madagaskar, Ambodimanga, Hammerstem S., I - II. 1906], [Locris bipunctata Sign. ♂ Edm. Schmidt determ. 1909], [Miz Pan Warszawa. 12 / 1945, 2493]; and [Madagaskar, Ambodimanga, Hammerstem S., I - II. 1906], [Locris bipunctata Sign. ♂ Edm. Schmidt determ. 1909], [Miz Pan Warszawa. 12 / 1945, 2494]. Casent collection - 7 males [CASENT 8107240], [Madagascar: Majunga, Ampijoroa National Park, 160 km N of Maevatanana, on RN 04, elev 43 m, 25 Jan- 7 Feb 2005], [16 ° 19.16 ' S, 46 ° 48.80 ' E, California Acad of Sciences, coll: M. Irwin, R. Harin'Hala, malaise trap - in deciduous forest, MA- 25 - 42]; [CASENT 8107233], [Madagascar: Majunga, Ampijoroa National Park, 160 km N of Maevatanan, on RN 04, elev 43 m, 2 - 9 November 2003], [16 ° 19.16 ' S, 46 ° 48.80 ' E, California Acad of Sciences, coll: M. Irwin, R. Harin'Hala, malaise trap - in deciduous forest, MA- 25 - 22]; [CASENT 8107236], [Madagascar: Majunga, Ampijoroa National Park, 160 km N of Maevatanan, on RN 04, elev 43 m, 10 - 21 January 2004], [16 ° 19.16 ' S, 46 ° 48.80 ' E, California Acad of Sciences, coll: M. Irwin, R. Harin'Hala, malaise trap - in deciduous forest, MA- 25 - 29]; [CASENT 8077112], [Perinet, Madagascar, 14 Jul. 1966, Liusnan], [Locris vicina, det. Penny, 99]; [CASENT 8077114], [Madagascar, Hellville, Nossi-Be Isl., XI - 18 - 1959], [E. S. Ross, Collector], [Locris vicina]; [CASENT 3002067], [Madagscar: Province, d'Antsiranana, Montagne des Francais. 7.2 km 142 ° SE, Antsiranana (= Diego Suarez), Elev 180 m, 22 - 28 Feb 2001], [12 ° 19 ' 22 ° S, 49 ° 20 ' 17 ° E, colls: Fisher. Griswold et al. Calif. Academy of Sciences, malaise trap, in tropical dry forest. Code: BLF 3130]; and [CASENT 3008146], [Madagascar: Fianarantsoa, Province, Parc National d'Isalo, 9.1 km 354 ° N Ranohira, elev 725 m, 27 - 31 Jan 2003, 22 ° 28 ' 54 " S, 045 ° 27 ' 42 " E], [coll. Fisher, Griswold et al. California Acad. Of Sciences, collected at light-gallery forest, collection code: BLF 7304].	en	Bouteille, Emilien, Cesne, Maxime Le, Soulier-Perkins, Adeline (2021): Cercopidae spittle-bugs (Hemiptera, Cicadomorpha) of Madagascar: a new species of Bourgoinrana and revision of the Locris species. ZooKeys 1023: 29-47, DOI: http://dx.doi.org/10.3897/zookeys.1023.58733, URL: http://dx.doi.org/10.3897/zookeys.1023.58733
