taxonID	type	description	language	source
C853F73124130C5DFF34FEF4B03AFF6F.taxon	description	Revised diagnosis. Male and female: head short, with vertex not expanded dorso-caudad; thick and long bristles absent on head but present on sides of prothorax; fronto-clypeal groove absent; maxillary palpomere III strongly thickened and stout; mandible without mesal sub-median tooth; antennae with club composed of antennomeres IX – XI; pronotum with rounded sides and weakly arcuate anterior and posterior margins; base of pronotum with shallow and indistinctly demarcated pair of sub-median ante-basal pits, without lateral and sub-lateral carinae; basisternal part of prosternum much shorter than procoxal cavities; prosternum without intercoxal process or carina; prothoracic hypomeral ridges incomplete; mesoventral intercoxal process long, narrow and strongly expanding ventrally (keel-shaped); mesoventrite with asetose lateral impressions behind anterior ridge, without setose impressions; mesothorax without lateral foveae; mesocoxal projection with posterior lobe; metacoxae narrowly separated by subtrapezoidal metaventral intercoxal process; each elytron with single rudimentary and asetose basal fovea. Male: parameres slender, not fused with median lobe, with apical setae; internal armature of aedeagus asymmetrical. Redescription. Body of male (Fig. 1) strongly convex, elongate but moderately slender, with moderately long appendages, BL below 1 mm; cuticle glossy, brown, moderately densely setose. Head (Figs. 1 – 2, 4 – 5) only slightly elongate, rounded, with large eyes; occipital constriction (Figs. 4 – 5; occ) in the narrowest place much wider than half HW; tempora (Fig. 4; tm) long and convergent caudad, without bristles; vertex (Fig. 4; vt) broader than long, rounded, convex, not projecting dorso-caudad; frons (Fig. 4; fr) transverse and subtriangular; fronto-clypeal groove absent; antennal insertions broadly separated. Labrum transverse with rounded anterior margin. Mandibles (Figs. 4 – 5; md) symmetrical, each with broad basal part, without noticeable prostheca, and with slender and curved distal part, without mesal tooth. Each maxilla (Fig. 5) with subtriangular basistipes (Fig. 5; bst), elongate galea (Fig. 5; gal) and lacinia (Fig. 5; lac) and long maxillary palp (Fig. 5; mxp) composed of relatively long palpomere I, strongly elongate, pedunculate palpomere II, broad and stout palpomere III broadest near middle, and small, subconical and pointed palpomere IV. Labium (Fig. 5) with large and transverse submentum (Fig. 5; smn) sharply demarcated from gular plate (Fig. 5; gp) and laterally fused with postcardinal parts of hypostomae (Fig. 52; h), subtrapezoidal mentum (Fig. 5; mn); and short prementum bearing narrowly separated at bases small 3 - segmented labial palps (Fig. 5; lp). Hypostomal ridges (Fig. 5; hr) long and connecting between base of submentum and gular plate. Gular plate (Fig. 5; gp) large and strongly narrowing anterad; gular sutures (Fig. 5; gs) superficial; posterior tentorial pits (Fig. 5; ptp) distinct, located at base of submentum. Antennae (Figs. 1 – 2) with distinct club composed of antennomeres IX – XI. Pronotum (Figs. 1 – 2) in dorsal view oval with strongly rounded lateral margins, indistinct anterior and posterior corners; without marginal carinae or edges and without sub-lateral carinae; base of pronotum with shallow and indistinctly demarcated pair of sub-median ante-basal pits; sides of pronotum with dense, thick and long bristles. Prosternum (Fig. 6) with short basisternal part (Fig. 6; bs) indistinctly demarcated from procoxal cavities (Fig. 6; pcc); median part of sternum without intercoxal carina or process; procoxal sockets (Fig. 6; pcs) closed by lateral lobes of sternum; hypomera (Fig. 6; hy) elongate; hypomeral ridge (Fig. 6; hyr) visible only in posterior half of hypomeron; pronotosternal sutures (Fig. 6; nss) entire. Mesoscutellum very small, heart-shaped, broader than long, barely visible between bases of elytra; mesoscutoscutellar suture indiscernible in slide preparation. Mesoventrite (Fig. 7) with narrow anterior ridge (Fig. 7; ar) and median projection of anterior ridge (Fig. 7; par); mesoventral intercoxal process (Fig. 7; msvp) narrow and keel-shaped; mesanepisternum with long prepectus (Fig. 7; pre) and posterior part not visible in ventral view; mesepimeron not visible in ventral view; sides of mesothorax without foveae; mesoventrite with indistinctly delimited subtriangular lateral asetose impressions (Fig. 7; ai); mesocoxal projections (Fig. 7; mcp) strongly projecting, with mesocoxal sockets (Fig. 7; mscs) located on their mesal surface and with distinct asetose posterior lobes (Fig. 7; pl). Metaventrite (Fig. 7; vIII) strongly transverse, anteriorly fused with mesoventrite, posteriorly moderately deeply bisinuate and with narrow median subtrapezoidal metaventral intercoxal process (Fig. 7; mtvp) bearing median notch. Metanepisterna and metepimera not visible in ventral view, narrow. Metafurca (Fig. 7) with very short and broad stem and divergent lateral furcal arms (Fig. 7; lmfa). Elytra (Figs. 1 – 2) oval, each with single rudimentary and asetose basal fovea located in shallow basal impression; humeral calli well-marked and developed as longitudinal protuberances; elytral apices unmodified, separately rounded. Legs (Figs. 1 – 2, 7) moderately long and slender; procoxae subglobose, mesocoxae slightly elongate, metacoxae transverse, stout; all trochanters short; all femora weakly clavate; tibiae short and slightly expanded near middle; tarsi short and stout. Abdominal sternites (Fig. 7) unmodified, suture between VII and VIII barely marked. Aedeagus (Figs. 8 – 9) strongly elongate, thin-walled, with internal armature composed of moderately darkly sclerotized tubular structure, and with free (i. e., not fused with median lobe) and slender parameres bearing apical setae.	en	Jałoszyński, Paweł (2013): Taxonomy of ' Euconnus complex'. Part II. Revision of Archiconnus Franz, Parapseudoconnus Franz and Mexiconnus gen. nov. (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 3666 (4): 523-543, DOI: 10.11646/zootaxa.3666.4.7
C853F73124130C5DFF34FEF4B03AFF6F.taxon	distribution	Distribution and composition. Archiconnus is represented by a single species known from the northern part of Peru.	en	Jałoszyński, Paweł (2013): Taxonomy of ' Euconnus complex'. Part II. Revision of Archiconnus Franz, Parapseudoconnus Franz and Mexiconnus gen. nov. (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 3666 (4): 523-543, DOI: 10.11646/zootaxa.3666.4.7
C853F73124130C5DFF34FEF4B03AFF6F.taxon	discussion	Remarks. Two species of Archiconnus were described by Franz (1980, 1994), but the other one, known from Mexico, is in the present paper placed in a separate genus. Archiconnus was described as a genus similar to Euconnus (Napochus), with a diagnostic set of characters including contiguous hind coxae, pronotum with rounded sides, strongly enlarged maxillary palpomere III, round head with bristles similar to those on pronotum, large and anteriorly located eyes, narrow occipital constriction, antennal club composed of three antennomeres, pronotum without lateral carinae near hind angles, with a pair of approximate ante-basal foveae, elytra with a single basal fovea, and aedeagal parameres free (i. e., not fused with the median lobe). However, one of these characters is in fact not present in Archiconnus: there are no stiff and thick bristles on the head, which is covered with setae as thin as those on elytra, in contrast to pronotal bristles.	en	Jałoszyński, Paweł (2013): Taxonomy of ' Euconnus complex'. Part II. Revision of Archiconnus Franz, Parapseudoconnus Franz and Mexiconnus gen. nov. (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 3666 (4): 523-543, DOI: 10.11646/zootaxa.3666.4.7
C853F73124130C5DFF34FEF4B03AFF6F.taxon	description	Archiconnus is highly similar to Euconnus s. str., from which it differs in lacking the fronto-clypeal groove (present in Euconnus s. str.); mandibles without subapical mesal tooth (with tooth in Euconnus s. str.); absent sublateral pronotal carinae (present in Euconnus s. str.); internal parts of prothoracic hypomera demarcated laterally by an incomplete hypomeral ridge (complete in Euconnus s. str.) and strongly expanding mesally over sides of prosternum and therefore concealing a large part of procoxal sockets (largely visible in Euconnus s. str.); lateral impressions of mesoventrite asetose (setose in Euconnus s. str.); absent ventral and dorsal foveae on mesoventrite (present in Euconnus s. str.); presence of distinct posterior lobe of mesocoxal projection (absent in Euconnus s. str.); nearly contiguous metacoxae (moderately broadly separated in Euconnus s. str.); and each elytron with a single rudimentary fovea barely discernible even in transparent mounts (two deep foveae in Euconnus s. str.). The number of these differences justifies the placement of Archiconnus as at least a separate subgenus within Euconnus, but since the character states within the enormously large Euconnus and its many subgenera are still very poorly studied, for the time being this taxon should be treated as a genus. The indistinctly demarcated basisternal part of prosternum, lack of prosternal intercoxal carina, procoxal sockets largely concealed under the internal part of hypomeron, which is laterally demarcated by an incomplete hypomeral ridge, and a single rudimentary elytral basal fovea make Archiconnus similar to previously revised Euconnomorphus (Jałoszyński 2012). However, Euconnomorphus clearly differs from Archiconnus in a slender, strongly elongate maxillary palpomere III (strongly broadened and stout in Archiconnus); subconically expanded vertex (not expanded in Archiconnus); two pairs of deep foveae on sides of the mesoventrite (absent in Archiconnus); and in absence of posterior lobes of mesocoxal projections (present in Archiconnus). Previously revised Venezolanoconnus (Jałoszyński 2012) is highly similar to Archiconnus in the general body form, the shape of vertex (not subconical), the eyes large in males and rudimentary in females, lack of bristles on sides of the head, the maxillary palpomere III strongly broadened and stout, mandibles without subapical mesal tooth, absent frontoclypeal groove, pronotal base without sub-lateral carinae and with a rudimentary pair of ante-basal pits (only in some species, in others entirely absent), sides of pronotum with dense and long bristles, lack of the prosternal intercoxal carina, presence of asetose impressions of mesoventrite and a similar shape and width of the metaventral intercoxal process. Venezolanoconnus differs from Archiconnus in hypostomal ridges not connected posteromesally (connected in Archiconnus), antennae gradually thickened distally (with distinct club in Archiconnus), prothoracic hypomera with the internal part demarcated laterally by complete hypomeral ridges (incomplete in Archiconnus), absent posterior lobes of mesocoxal projections (present in Archiconnus) and extremely broad, hyaline parameres laterally embracing median lobe (slender parameres in Archiconnus). Archiconnus is also similar to Parapseudoconnus, from which it differs in the hypostomal ridges connected in middle behind the submentum (short, not connected ridges in Parapseudoconnus), absent prosternal intercoxal carina (present in Parapseudoconnus), incomplete prothoracic hypomeral ridges (complete or nearly complete in Parapseudoconnus), postero-lateral (postcoxal) parts of prosternum separated from internal parts of prothoracic hypomera (fused in Parapseudoconnus), absent ventro-lateral foveae of the mesoventrite (present in Parapseudoconnus), and the aedeagus with parameres (parameres absent in Parapseudoconnus). Archiconnus mexicanus Franz is here separated from A. aberrans and placed in its own genus, Mexiconnus. The differences between these taxa are numerous and commented on in Remarks under the latter genus. Archiconnus huallaganus Franz (Figs. 1 – 9) Archiconnus huallaganus Franz, 1980: 189, Fig. 173.	en	Jałoszyński, Paweł (2013): Taxonomy of ' Euconnus complex'. Part II. Revision of Archiconnus Franz, Parapseudoconnus Franz and Mexiconnus gen. nov. (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 3666 (4): 523-543, DOI: 10.11646/zootaxa.3666.4.7
C853F73124130C5DFF34FEF4B03AFF6F.taxon	materials_examined	Material studied. Holotype: 3: three labels (Fig. 3): " Umg. Tarapoto / Peru, lg. Franz " with " SA 137 [or " / 37 '] " on the reverse side [white, printed; reverse handwritten], " Euconnus [sic!] / (Archiconnus) / huallague [sic!] m. / det. H. Franz " [white, handwritten and printed], " Typus " [red, handwritten] (NHMW). Paratype: Ƥ: same data as for holotype (NHMW).	en	Jałoszyński, Paweł (2013): Taxonomy of ' Euconnus complex'. Part II. Revision of Archiconnus Franz, Parapseudoconnus Franz and Mexiconnus gen. nov. (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 3666 (4): 523-543, DOI: 10.11646/zootaxa.3666.4.7
C853F73124130C5DFF34FEF4B03AFF6F.taxon	diagnosis	Diagnosis. This is the only known species of re-defined Archiconnus and can be identified on the basis of the generic characters and the aedeagus. Redescription. Body of male (Fig. 1) strongly convex, elongate and relatively slender, with moderately long appendages, BL 0.81 mm; glossy, uniformly brown with slightly lighter legs and palps; vestiture slightly lighter than cuticle. Head (Fig. 1) roundly rhomboidal and slightly elongate, broadest at eyes, HL 0.18 mm, HW 0.15 mm; occipital constriction broader than half HW; tempora slightly shorter than eyes, rounded and gradually convergent caudad; vertex nearly as long as broad, convex, only slightly (not conically) projected dorso-caudad; frons modified, with pair of large lateral median tubercles located antero-mesally to distinct supraantennal tubercles; eyes large and strongly convex, moderately coarsely faceted, nearly circular with shallow posterior emargination. Punctures on head dorsum fine and sparse, inconspicuous; setae long, moderately dense, erect, those on vertex mostly directed caudad, thick bristles absent. Antennae (Fig. 1) slender, with indistinctly demarcated club composed of antennomeres IX – XI, AnL 0.30 mm; antennomeres I – II elongate, III – VII each about as broad as long or only slightly transverse; VIII strongly transverse; IX and X each strongly transverse; XI 1.7 x as long as broad, slightly asymmetrical, with subconical, blunt apex. Pronotum (Fig. 1) in dorsal view elongate oval, with broadly rounded and indistinct front and hind angles, broadest near posterior fourth, PL 0.24 mm, PW 0.21 mm; anterior, posterior and lateral margins rounded; base of pronotum with shallow and indistinctly demarcated pair of sub-median ante-basal pits. Punctures on pronotal disc fine and inconspicuous; setae long, moderately dense and strongly erect, sides of pronotum with dense thick and long bristles. Elytra (Fig. 1) oval and more convex than pronotum, broadest distinctly behind middle, EL 0.40 mm, EW 0.30 mm, EI 1.33; humeral calli distinct, developed as short longitudinal protuberances; basal impressions indistinct, basal pits barely discernible; elytral apices separately rounded. Punctures on elytral disc as fine as those on pronotum; setae much shorter than pronotal bristles, moderately sparse and suberect. Hind wings well-developed, about twice as long as elytra. Legs (Fig. 1) moderately long and slender, without modifications. Aedeagus (Figs. 8 – 9) elongate, AeL 0.13 mm, median lobe subtriangular, strongly narrowing from base to apex; distal part with truncate apex, in lateral view strongly curved dorsally; internal armature composed of tubular structures located in basal part of median lobe; parameres slender, each with two long and thick apical setae. Female (Figs. 2, 4 – 7). Similar to male but clearly differing in non-modified frons and rudimentary eyes (Fig. 4), each composed of a single ommatidium. BL 0.79 mm; HL 0.16 mm, HW 0.14 mm, AnL 0.28 mm; PL 0.23 mm, PW 0.20 mm; EL 0.40 mm, EW 0.30 mm, EI 1.33.	en	Jałoszyński, Paweł (2013): Taxonomy of ' Euconnus complex'. Part II. Revision of Archiconnus Franz, Parapseudoconnus Franz and Mexiconnus gen. nov. (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 3666 (4): 523-543, DOI: 10.11646/zootaxa.3666.4.7
C853F73124130C5DFF34FEF4B03AFF6F.taxon	distribution	Distribution. Northern Peru, San Martin Province.	en	Jałoszyński, Paweł (2013): Taxonomy of ' Euconnus complex'. Part II. Revision of Archiconnus Franz, Parapseudoconnus Franz and Mexiconnus gen. nov. (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 3666 (4): 523-543, DOI: 10.11646/zootaxa.3666.4.7
C853F73124130C5DFF34FEF4B03AFF6F.taxon	discussion	Remarks. Franz (1980) included in the type series of A. huallaganus a holotype male and two paratype females (one labeled as allotype). The allotype female, however, clearly differs from the other paratype and the holotype in the body shape and proportions of body parts and it certainly belongs to a different species. Details of the ventral side of this specimen were not studied and it may belong to Archiconnus or Euconnus. In the original description Franz (1980) gives further collecting data for the holotype as a road to Yurimagua, 80 km from Tarapoto, bog or swamp forest (" Sumpfwald "), and collecting data for the paratype female as 40 km from Tarapoto, montane forest, 0 1.10.1968 (it is unclear whether the collecting date refers only to the paratype or both holotype and paratype).	en	Jałoszyński, Paweł (2013): Taxonomy of ' Euconnus complex'. Part II. Revision of Archiconnus Franz, Parapseudoconnus Franz and Mexiconnus gen. nov. (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 3666 (4): 523-543, DOI: 10.11646/zootaxa.3666.4.7
C853F73124160C53FF34FEA5B513FD1C.taxon	diagnosis	Diagnosis. Male: head short, with vertex not expanded dorso-caudad; thick and long bristles absent on head but present on sides of prothorax; fronto-clypeal groove absent; maxillary palpomere III strongly thickened and stout; mandible with mesal sub-median tooth; antennae with club composed of antennomeres IX – XI; pronotum with rounded sides and weakly arcuate anterior and posterior margins; base of pronotum with deep and distinct transverse ante-basal groove slightly deepened at each end and not reaching lateral pronotal margins, without lateral and sub-lateral carinae; basisternal part of prosternum much shorter than procoxal cavities; prosternum with fine intercoxal carina; prothoracic hypomeral ridges complete; mesoventral intercoxal process long, narrow and strongly expanding ventrally (keel-shaped); mesoventrite with asetose lateral impressions behind anterior ridge, without setose impressions; mesothorax with a pair of deep ventro-lateral foveae; mesocoxal projection with very short, barely discernible posterior lobe; metacoxae narrowly separated by subtrapezoidal metaventral intercoxal process; each elytron with two rudimentary and asetose basal foveae; parameres not fused with median lobe, with broad basal and slender distal parts, without apical setae; internal armature of aedeagus asymmetrical. Females and their diagnostic characters unknown.	en	Jałoszyński, Paweł (2013): Taxonomy of ' Euconnus complex'. Part II. Revision of Archiconnus Franz, Parapseudoconnus Franz and Mexiconnus gen. nov. (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 3666 (4): 523-543, DOI: 10.11646/zootaxa.3666.4.7
C853F73124160C53FF34FEA5B513FD1C.taxon	description	Description. Body of male (Fig. 10) moderately strongly convex, slender, with moderately long appendages, BL below 1 mm; cuticle glossy, brown, moderately setose. Head (Figs. 10, 12) short and approximately rhomboidal, with large eyes; occipital constriction (Fig. 12; occ) in the narrowest place much broader than half HW; tempora long and strongly convergent caudad, without bristles; vertex rounded and convex, broader than long, not expanded dorso-caudad; frons transverse and subtriangular; fronto-clypeal groove absent; antennal insertions broadly separated. Labrum transverse with rounded sides and nearly straight anterior margin. Mandibles (Fig. 12; md) symmetrical, each with broad basal part, without noticeable prostheca, and with slender and curved distal part with one small subapical mesal tooth. Each maxilla (Fig. 12) with subtriangular basistipes (Fig. 12; bst), elongate galea (Fig. 12; gal) and lacinia (Fig. 12; lac) and moderately long maxillary palp (Fig. 12; mxp) composed of relatively long palpomere I, strongly elongate, pedunculate palpomere II, moderately elongate, very broad palpomere III broadest near middle, and small, subconical palpomere IV. Labium (Fig. 12) with approximately hexagonal, elongate submentum (Fig. 12; smn) indistinctly delimited from gular plate (Fig. 12; gp) and laterally demarcated from hypostomae by long and recurved lateral sutures of submentum (Fig. 12; lss); subrectangular mentum (Fig. 12; mn); and short prementum with small 3 - segmented labial palps (Fig. 12; lp) narrowly separated at bases. Hypostomal ridges (Fig. 12; hr) short, not reaching lateral margins of submentum. Gular plate (Fig. 12; gp) indistinctly demarcated, strongly narrowing anterad; gular sutures (Fig. 12; gs) superficial; posterior tentorial pits (Fig. 12; ptp) distinct and located at base of submentum. Antennae (Fig. 10) with club composed of antennomeres IX – XI. Pronotum (Fig. 10) in dorsal view oval, with indistinct front angles and well-marked hind angles; sides strongly rounded; base of pronotum with deep transverse ante-basal groove slightly deepened at each end and not reaching lateral pronotal margins, without lateral carinae or edges and without sub-lateral carinae; sides of pronotum with long and thick bristles. Prosternum (Fig. 13) with basisternal part (Fig. 13; bs) relatively short and indistinctly demarcated from procoxal cavities (Fig. 13; pcc); median part of sternum with fine prosternal intercoxal carina (Fig. 13; pc); procoxal sockets (Fig. 13; pcs) closed by lateral lobes of sternum; hypomera (Fig. 12; hy) elongate, each with elongate internal (adcoxal) part demarcated laterally by complete hypomeral ridge (Fig. 13; hyr); pronotosternal sutures (Fig. 13; nss) entire. Mesoscutellum very small, subtriangular and elongate, barely visible between bases of elytra; mesoscutoscutellar suture fine but discernible. Mesoventrite (Fig. 14) with narrow anterior ridge (Fig. 14; ar) with small median projection (Fig. 14; par); mesoventral intercoxal process (Fig. 14; msvp) narrow and keel-shaped; mesoventrite behind anterior ridge with sharply delimited lateral asetose impressions (Fig. 14; ai); prepectus (Fig. 14; pre) long, posterior portion of mesanepisternum only partly visible in ventral view; mesepimeron not visible in ventral view; mesothorax with deep ventro-lateral fovea at each side (Fig. 14; vlf); mesocoxal projections (Fig. 14; mcp) with mesocoxal sockets (Fig. 14; mscs) located on their meso-ventral surface, without posterior lobes. Metaventrite (Fig. 14; vIII) strongly transverse, anteriorly fused with mesoventrite, posteriorly moderately deeply bisinuate and with narrow median subtrapezoidal metaventral intercoxal process (Fig. 14; mtvp) bearing median notch. Metanepisterna and metepimera narrow, only posterior parts of episterna visible in ventral view. Metafurca (Fig. 14) with very short and broad stem and divergent lateral furcal arms (Fig. 14; lmfa). Elytra (Fig. 10) oval, each with two rudimentary and asetose basal foveae barely discernible even in transparent mounts; humeral calli well-marked and developed as longitudinal protuberances; elytral apices unmodified, separately rounded. Legs (Figs. 10, 13 – 16) long and slender; procoxae subglobose, mesocoxae slightly elongate, metacoxae transverse, stout; all trochanters short; all femora weakly clavate; tibiae short and slightly expanded near middle; tarsi short and stout, protarsi (Figs. 15 – 16) strongly broadened at base and narrowing distally, with long and dense setae on ventral surface, especially tarsomere I with dense cushion of long setae. Abdominal sternites (Fig. 14) only partly preserved in holotype; sternites III and IV unmodified. Aedeagus (Figs. 17 – 18) strongly elongate, thin-walled, with internal armature composed of moderately darkly sclerotized and tubular structure forming an open loop, and with free (i. e., not fused with median lobe) parameres without apical setae; basal part of parameres very broad and embracing laterally median lobe, distal parts slender and narrowing to pointed apices.	en	Jałoszyński, Paweł (2013): Taxonomy of ' Euconnus complex'. Part II. Revision of Archiconnus Franz, Parapseudoconnus Franz and Mexiconnus gen. nov. (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 3666 (4): 523-543, DOI: 10.11646/zootaxa.3666.4.7
C853F73124160C53FF34FEA5B513FD1C.taxon	distribution	Distribution and composition. Mexiconnus is represented by a single species known from the southernmost part of Mexico.	en	Jałoszyński, Paweł (2013): Taxonomy of ' Euconnus complex'. Part II. Revision of Archiconnus Franz, Parapseudoconnus Franz and Mexiconnus gen. nov. (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 3666 (4): 523-543, DOI: 10.11646/zootaxa.3666.4.7
C853F73124160C53FF34FEA5B513FD1C.taxon	discussion	Remarks. Mexiconnus differs from Archiconnus in several important characters that justify its separate placement. The submentum in Mexiconnus is demarcated from post-cardinal parts of hypostomae by complete lateral sutures and hypostomal ridges are short and posteriorly reaching only to half length of submentum, while in Archiconnus lateral sutures of submentum are absent and the hypostomal ridges are not only long, but connected in middle between the base of submentum and the gular plate. In Mexiconnus the prosternal intercoxal carina is present and the internal parts of prothoracic hypomera are laterally demarcated by complete hypomeral ridges, while in Archiconnus the prosternal carina is absent and the hypomeral ridges are incomplete. In Mexiconnus the mesoventrite has deep ventro-lateral foveae, entirely missing in Archiconnus. Mexiconnus has two rudimentary and barely discernible in transparent mounts basal elytral foveae, while only a single, also rudimentary fovea can be seen in Archiconnus. Also the aedeagi, described by Franz (1994) as similar, differ significantly in the shape of parameres. In Mexiconnus the parameres are massive and their broad basal parts embrace the median lobe laterally, while distal parts are slender and gradually narrowing towards pointed apices devoid of setae. In Archiconnus the parameres are slender also in their basal halves, each with rounded apex bearing two very long and thick setae. Strongly broadened protarsi in the only known male of Mexiconnus mexicanus may constitute another difference between these two genera, as in Archiconnus male and females do not differ in the shape of protarsomeres. Mexiconnus clearly differs from other members of the hitherto revised genera of the ' Euconnus complex'. From Euconnus s. str. (Jałoszyński 2012) it differs in more slender body, with elytra not so much broader than the pronotum, the lack of fronto-clypeal groove (present in Euconnus s. str.), presence of lateral sutures of submentum (absent in Euconnus s. str.), short hypostomal ridges (long in Euconnus s. str.), pronotum in dorsal view strongly rounded (bell-shaped in Euconnus s. str.), presence of prosternal intercoxal carina (absent in Euconnus s. str.), lack of sub-lateral pronotal carinae (present in Euconnus s. str.), asetose impressions of the mesoventrite (setose in Euconnus s. str.), nearly contiguous metacoxae (moderately broadly separated in Euconnus s. str.), rudimentary basal elytral foveae (deep in Euconnus s. str.), and the broad parameres devoid of apical setae (slender and with setae in Euconnus s. str.).	en	Jałoszyński, Paweł (2013): Taxonomy of ' Euconnus complex'. Part II. Revision of Archiconnus Franz, Parapseudoconnus Franz and Mexiconnus gen. nov. (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 3666 (4): 523-543, DOI: 10.11646/zootaxa.3666.4.7
C853F73124160C53FF34FEA5B513FD1C.taxon	description	Mexiconnus differs from Euconnomorphus (Jałoszyński 2012) in the vertex not expanded dorso-caudad (strongly subconical in Euconnomorphus), short and stout maxillary palpomere III (strongly elongate in Euconnomorphus), antennae with distinct club (gradually thickened distally in Euconnomorphus), presence of lateral sutures of submentum (absent in Euconnomorphus), short hypostomal ridges (long in Euconnomorphus), pronotum in dorsal view strongly rounded (bell-shaped in Euconnus s. str.), presence of prosternal intercoxal carina (absent in Euconnomorphus), complete prothoracic hypomeral ridges (incomplete in Euconnomorphus), welldemarcated asetose impressions of the mesoventrite (not demarcated in Euconnomorphus), and two (and not one) rudiments of basal elytral foveae. Mexiconnus differs from Venezolanoconnus in much more slender body, antennal club composed of three antennomeres (antennae gradually thickening distally in Venezolanoconnus), presence of lateral sutures of submentum (absent in Venezolanoconnus), short hypostomal ridges (long in Venezolanoconnus), pronotum in dorsal view strongly rounded (in Venezolanoconnus strongly rounded in anterior half and weakly in posterior half )), presence of prosternal intercoxal carina (absent in Venezolanoconnus), two (and not one) rudiments of basal elytral foveae, and strongly elongate distal halves of parameres (broad and short in Venezolanoconnus). Mexiconnus differs from Parapseudoconnus in the presence of lateral sutures of submentum (absent in Parapseudoconnus), postero-lateral (postcoxal) parts of prosternum separated from internal parts of prothoracic hypomera (fused in Parapseudoconnus), two rudiments of basal elytral foveae (one deep and setose fovea in Parapseudoconnus), and presence of parameres (absent in Parapseudoconnus).	en	Jałoszyński, Paweł (2013): Taxonomy of ' Euconnus complex'. Part II. Revision of Archiconnus Franz, Parapseudoconnus Franz and Mexiconnus gen. nov. (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 3666 (4): 523-543, DOI: 10.11646/zootaxa.3666.4.7
C853F73124180C50FF34FD4CB5B5FE6C.taxon	materials_examined	Material studied. Holotype: 3: three labels (Fig. 11): " Umg. San-Cristobal / Chiapas, Mexico " with " Mx 16 " on the reverse side [white, printed; reverse handwritten], " Archiconnus / mexicanus m. / det. H. Franz " [white, handwritten and printed], " Holotypus " [red, handwritten] (NHMW). A white printed label was added during the present study: " MEXICONNUS / mexicanus / (Franz, 1994) / det. P. JAŁOSZYŃSKI, ' 13 ".	en	Jałoszyński, Paweł (2013): Taxonomy of ' Euconnus complex'. Part II. Revision of Archiconnus Franz, Parapseudoconnus Franz and Mexiconnus gen. nov. (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 3666 (4): 523-543, DOI: 10.11646/zootaxa.3666.4.7
C853F73124180C50FF34FD4CB5B5FE6C.taxon	diagnosis	Diagnosis. This is the only known species of Mexiconnus and can be identified on the basis of the generic characters and the aedeagus.	en	Jałoszyński, Paweł (2013): Taxonomy of ' Euconnus complex'. Part II. Revision of Archiconnus Franz, Parapseudoconnus Franz and Mexiconnus gen. nov. (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 3666 (4): 523-543, DOI: 10.11646/zootaxa.3666.4.7
C853F73124180C50FF34FD4CB5B5FE6C.taxon	description	Description. Body of male (Fig. 10) slightly flattened, elongate and slender, with long appendages, BL 0.93 mm; glossy, uniformly moderately dark brown with slightly lighter appendages; vestiture slightly lighter than cuticle. Head (Figs. 10, 12) short and rhomboidal in shape, broadest at eyes, HL 0.18 mm, HW 0.20 mm; occipital constriction slightly broader than half HW; tempora slightly longer than eyes, strongly convergent caudad and only weakly rounded; vertex strongly transverse, convex, not expanded dorso-caudad; frons confluent with vertex, slightly convex, subtrapezoidal and steeply but not abruptly lowering towards strongly transverse clypeus; supraantennal tubercles feebly marked; eyes large and strongly convex, kidney-shaped and transverse in relation to the long axis of the head, finely faceted. Punctures on head dorsum fine and sparse, inconspicuous; setae long, moderately dense, suberect to erect, those on vertex and tempora directed caudad, thick bristles absent. Antennae (Fig. 10) long and slender, with moderately distinctly delimited club composed of antennomeres IX – XI, AnL 0.50 mm; antennomeres I – IX elongate; X distinctly transverse; XI only 1.2 x as long as broad, symmetrical, with broadly rounded apex. Pronotum (Fig. 10) in dorsal view elongate oval, broadest in anterior fourth, PL 0.25 mm, PW 0.23 mm; anterior margin slightly arcuate; front angles indistinct; lateral margins strongly rounded and strongly convergent towards well-marked, obtuse and blunt hind angles; posterior margin slightly arcuate; base of pronotum with deep and distinct transverse ante-basal groove slightly deepened at each end and not reaching lateral pronotal margins. Punctures on pronotal disc fine and inconspicuous; setae moderately long and dense, suberect, sides of pronotum with thick and long bristles. Elytra (Fig. 10) oval and only slightly more convex than pronotum, broadest near middle, EL 0.50 mm, EW 0.35 mm, EI 1.43; humeral calli distinct, each developed as short longitudinal protuberance; basal elytral impressions distinct; basal pits indiscernible in a dry-mounted specimen; elytral apices separately rounded. Punctures on median circumsutural part of elytral disc more distinct and denser than those on pronotum but only superficial and with diffused margins, punctures on remaining surface of elytra fine and inconspicuous; setae short, moderately sparse and suberect. Hind wings well-developed, about twice as long as elytra. Legs (Figs. 10, 15 – 16) long and slender; protarsi strongly broadened at base and with dense cushion of long setae on ventral surface, especially on tarsomere I. Aedeagus (Figs. 17 – 18) elongate, AeL 0.14 mm, median lobe oval, strongly narrowing from base to subapical region where the distal third of aedeagus is rapidly narrowing and forming slender apical projection; internal armature composed of tubular structures forming an open loop; parameres with broad basal and slender distal parts, each with pointed apex and without setae. Female. Unknown.	en	Jałoszyński, Paweł (2013): Taxonomy of ' Euconnus complex'. Part II. Revision of Archiconnus Franz, Parapseudoconnus Franz and Mexiconnus gen. nov. (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 3666 (4): 523-543, DOI: 10.11646/zootaxa.3666.4.7
C853F73124180C50FF34FD4CB5B5FE6C.taxon	distribution	Distribution. Southern Mexico, Chiapas State.	en	Jałoszyński, Paweł (2013): Taxonomy of ' Euconnus complex'. Part II. Revision of Archiconnus Franz, Parapseudoconnus Franz and Mexiconnus gen. nov. (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 3666 (4): 523-543, DOI: 10.11646/zootaxa.3666.4.7
C853F73124180C50FF34FD4CB5B5FE6C.taxon	discussion	Remarks. In the original description Franz (1994) gives further collecting data: Los Alcanfores (misspelled as " Alcanfones "; a place located west of San Cristóbal de Las Casas), old oak forest, 17.06.1992.	en	Jałoszyński, Paweł (2013): Taxonomy of ' Euconnus complex'. Part II. Revision of Archiconnus Franz, Parapseudoconnus Franz and Mexiconnus gen. nov. (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 3666 (4): 523-543, DOI: 10.11646/zootaxa.3666.4.7
C853F731241A0C57FF34F81AB059F9B5.taxon	description	Parapseudoconnus Franz, 1980: 218 (as subgenus of Pseudoconnus Franz, 1980). Type species: Pseudoconnus aberrans Franz, 1980 (monotypy). Revised diagnosis. Male and female: head short, with vertex not expanded dorso-caudad; thick and long bristles absent on head but present on sides of prothorax; fronto-clypeal groove absent; maxillary palpomere III strongly thickened and stout; mandible without mesal sub-median tooth; antennae with club composed of antennomeres IX – XI; pronotum with rounded sides and weakly arcuate anterior and posterior margins, base of pronotum with one pair of external lateral pits and short longitudinal sub-lateral carina; basisternal part of prosternum much shorter than procoxal cavities; prosternum with fine intercoxal carina; prothoracic hypomeral ridges complete or nearly complete; postero-lateral (postcoxal) parts of prosternum fused with internal parts of prothoracic hypomera; mesoventral intercoxal process long, narrow and strongly expanding ventrally (keel-shaped); mesoventrite with asetose lateral impressions behind anterior ridge, without setose impressions; mesothorax with ventro-lateral foveae; metacoxae narrowly separated by subtrapezoidal or subtriangular metaventral intercoxal process; each elytron with single deep and setose basal fovea. Male: aedeagus without parameres, with asymmetrical internal armature. Redescription. Body of male (Figs. 19, 21) strongly convex, elongate but moderately slender, with moderately long appendages, BL below 1 mm; cuticle glossy, brown, moderately densely setose. Head (Figs. 19 – 21, 26 – 27, 32) short, approximately rhomboidal or rounded, with large eyes; occipital constriction (Figs. 27, 32; occ) in the narrowest place much wider than half HW; tempora (Fig. 26; tm) long and convergent caudad, without bristles; vertex (Fig. 26; vt) broader than long, rounded, convex, not projecting dorsocaudad; frons (Fig. 26; fr) transverse and subtriangular; fronto-clypeal groove absent; antennal insertions broadly separated. Labrum (Fig. 26; lb) transverse with rounded anterior margin. Mandibles (Figs. 26 – 27, 32; md) symmetrical, each with broad basal part, without noticeable prostheca, and with slender and curved distal part, without mesal tooth. Each maxilla (Figs. 27, 32) with subtriangular basistipes (Figs. 27, 32; bst), elongate galea (Figs. 27, 32; gal) and lacinia (Figs. 27, 32; lac) and long maxillary palp (Figs. 27, 32; mxp) composed of relatively long palpomere I, strongly elongate, pedunculate palpomere II, broad and stout palpomere III broadest between middle and basal third, and small, subconical and pointed palpomere IV. Labium (Figs. 27, 32) with large and transverse submentum (Figs. 27, 32; smn) not demarcated from gular plate (Figs. 27, 32; gp) and laterally fused with postcardinal parts of hypostomae (Figs. 27, 32; h), subtrapezoidal mentum (Figs. 27, 32; mn); and short prementum with small 3 - segmented labial palps narrowly separated at bases (Figs. 27, 32; lp). Hypostomal ridges (Figs. 27, 32; hr) posteriorly reaching to half length of submentum. Gular plate (Figs. 27, 32; gp) large and strongly narrowing anterad; gular sutures (Figs. 27, 32; gs) superficial; posterior tentorial pits (Figs. 27, 32; ptp) distinct, located at base of submentum. Antennae (Figs. 19 – 21, 26) with distinct club composed of antennomeres IX – XI. Pronotum (Figs. 19 – 21) in dorsal view approximately oval with strongly rounded lateral margins, moderately distinct anterior and distinct posterior corners; without marginal carinae or edges but with short sub-lateral carinae visible near posterior pronotal corners; base of pronotum with one pair of shallow and small external lateral pits; sides of pronotum with dense, thick and long bristles. Prosternum (Figs. 28, 33) with short basisternal part (Figs. 28, 33; bs) distinctly demarcated from procoxal cavities (Figs. 28, 33; pcc); median part of sternum with fine intercoxal carina; procoxal sockets (Figs. 28, 33; pcs) closed by lateral lobes of sternum which are fused with internal parts of prothoracic hypomera; hypomera (Figs. 28, 33; hy) elongate; hypomeral ridge (Figs. 28, 33; hyr) complete or nearly complete; pronotosternal sutures (Figs. 28, 33; nss) entire. Mesoscutellum very small, subtriangular, barely visible between bases of elytra; mesoscuto-scutellar suture indiscernible in slide preparation. Mesoventrite (Figs. 29, 34) with narrow anterior ridge (Figs. 29, 34; ar) and median projection of anterior ridge (Figs. 29, 34; par); mesoventral intercoxal process (Figs. 29, 34; msvp) narrow and keel-shaped; mesanepisternum with long prepectus (Figs. 29, 34; pre) and posterior part in ventral view visible only near to ventro-lateral fovea; mesepimeron not visible in ventral view; sides of mesothorax with deep ventro-lateral foveae (Figs. 29, 34; vlf); mesoventrite with variously distinctly delimited lateral asetose impressions (Figs. 29, 34; ai), without setose impressions; mesocoxal projections (Figs. 29, 34; mcp) with mesocoxal sockets (Figs. 29, 34; mscs) located on their mesal or meso-ventral surface and with or without posterior lobes (Fig. 29; pl). Metaventrite (Figs. 29, 34; vIII) strongly transverse, anteriorly fused with mesoventrite, posteriorly moderately deeply bisinuate and with narrow median subtrapezoidal metaventral intercoxal process (Figs. 29, 34; mtvp) bearing median notch. Metanepisterna and metepimera narrow, only posterior parts of episterna partly visible in ventral view. Metafurca (Fig. 34) with very short and broad stem and divergent lateral furcal arms (Fig. 34; lmfa). Elytra (Figs. 19 – 21) oval, each with single deep and setose basal fovea located in shallow basal impression; humeral calli well-marked and developed as longitudinal protuberances; elytral apices unmodified, separately rounded. Legs (Figs. 19 – 21, 28 - 29, 33 – 34) moderately long and slender; procoxae subglobose, mesocoxae slightly elongate, metacoxae transverse, stout; all trochanters short; all femora weakly clavate; tibiae short and slightly expanded near middle or nearly parallel-sided; tarsi short and stout. Abdominal sternites (Fig. 29) unmodified, suture between VII and VIII barely marked. Aedeagus (Figs. 30 – 31, 35 – 36) elongate, thin-walled, with asymmetrical internal armature composed of moderately darkly sclerotized set of complicated sclerites; parameres absent.	en	Jałoszyński, Paweł (2013): Taxonomy of ' Euconnus complex'. Part II. Revision of Archiconnus Franz, Parapseudoconnus Franz and Mexiconnus gen. nov. (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 3666 (4): 523-543, DOI: 10.11646/zootaxa.3666.4.7
C853F731241A0C57FF34F81AB059F9B5.taxon	distribution	Distribution and composition. One species of Parapseudoconnus is known to occur in the north-western part of Brazil and three others in the northern part of Peru.	en	Jałoszyński, Paweł (2013): Taxonomy of ' Euconnus complex'. Part II. Revision of Archiconnus Franz, Parapseudoconnus Franz and Mexiconnus gen. nov. (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 3666 (4): 523-543, DOI: 10.11646/zootaxa.3666.4.7
C853F731241A0C57FF34F81AB059F9B5.taxon	discussion	Remarks. Parapseudoconnus differs from Euconnus s. str. in distinct antennal club composed of three antennomeres (antennae gradually thickening distally in Euconnus s. str.), lack of fronto-clypeal groove (present in Euconnus s. str.), short hypostomal ridges (long in Euconnus s. str.), presence of prosternal intercoxal carina (absent in Euconnus s. str.), postero-lateral (postcoxal) parts of prosternum fused with internal parts of prothoracic hypomera (separated in Euconnus s. str.), asetose impressions of mesoventrite (setose in Euconnus s. str.), nearly contiguous metacoxae (moderately broadly separated in Euconnus s. str.), and the aedeagus without parameres (with parameres in Euconnus s. str.). Parapseudoconnus differs from Euconnomorphus in distinct antennal club composed of three antennomeres (antennae gradually thickening distally in Euconnomorphus), short hypostomal ridges (long in Euconnomorphus), short head with the vertex not expanded dorso-caudad (long head with the vertex subconical in Euconnomorphus), presence of prosternal intercoxal carina (absent in Euconnomorphus), postero-lateral (postcoxal) parts of prosternum fused with internal parts of prothoracic hypomera (separated in Euconnomorphus), deep and setose basal elytral fovea (rudimentary and asetose in Euconnomorphus), and the aedeagus without parameres (with parameres in Euconnomorphus).	en	Jałoszyński, Paweł (2013): Taxonomy of ' Euconnus complex'. Part II. Revision of Archiconnus Franz, Parapseudoconnus Franz and Mexiconnus gen. nov. (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 3666 (4): 523-543, DOI: 10.11646/zootaxa.3666.4.7
C853F731241A0C57FF34F81AB059F9B5.taxon	description	Parapseudoconnus differs from Venezolanoconnus in distinct antennal club composed of three antennomeres (antennae gradually thickening distally in Venezolanoconnus), short hypostomal ridges (long in Venezolanoconnus), presence of prosternal intercoxal carina (absent in Venezolanoconnus), postero-lateral (postcoxal) parts of prosternum fused with internal parts of prothoracic hypomera (separated in Venezolanoconnus), deep and setose basal elytral fovea (rudimentary and asetose in Venezolanoconnus), and the aedeagus without parameres (with parameres in Venezolanoconnus). Differences between Parapseudoconnus, Archiconnus and Mexiconnus were described in remarks under the two latter genera.	en	Jałoszyński, Paweł (2013): Taxonomy of ' Euconnus complex'. Part II. Revision of Archiconnus Franz, Parapseudoconnus Franz and Mexiconnus gen. nov. (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 3666 (4): 523-543, DOI: 10.11646/zootaxa.3666.4.7
C853F731241C0C4AFF34F9E4B3B9FF4C.taxon	description	Pseudoconnus (Parapseudoconnus) aberrans Franz, 1980: 218, Fig. 206.	en	Jałoszyński, Paweł (2013): Taxonomy of ' Euconnus complex'. Part II. Revision of Archiconnus Franz, Parapseudoconnus Franz and Mexiconnus gen. nov. (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 3666 (4): 523-543, DOI: 10.11646/zootaxa.3666.4.7
C853F731241C0C4AFF34F9E4B3B9FF4C.taxon	materials_examined	Material studied. Holotype: 3: four labels (Fig. 22): " Umg. Manau / Amazongebiet / Brasil, lg. L. Beck " with " 45 " on the reverse side [white, printed; reverse handwritten], " Parapseudocon- / nus aberrans / m. / det. H. Franz " [white, handwritten and printed], " 3 " [white, printed], " Typus " [red, handwritten] (NHMW). Paratypes (6 exx.: 2 33, 4 ƤƤ): same data as for the holotype, one female laballed " Allotypus " [red, handwritten] (NHMW).	en	Jałoszyński, Paweł (2013): Taxonomy of ' Euconnus complex'. Part II. Revision of Archiconnus Franz, Parapseudoconnus Franz and Mexiconnus gen. nov. (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 3666 (4): 523-543, DOI: 10.11646/zootaxa.3666.4.7
C853F731241C0C4AFF34F9E4B3B9FF4C.taxon	diagnosis	Diagnosis. Vertex in males and females with distinct postero-median impression; aedeagus with three elongate and curved sclerites. Redescription. Body of male (Fig. 19) strongly convex and moderately elongate, with moderately long appendages, BL 0.84 – 0.86 mm (mean 0.85 mm); glossy, uniformly light brown with slightly lighter appendages; vestiture slightly lighter than cuticle. Head (Fig. 19) approximately pentagonal and not elongate, broadest at eyes, HL 0.16 – 0.18 mm (mean 0.17 mm), HW 0.18 – 0.20 mm (mean 0.19 mm); occipital constriction much broader than half HW; tempora slightly shorter than eyes, rounded and gradually convergent caudad; vertex not projected dorso-caudad; broader than long, convex except for shallow postero-median impression; frons subtriangular and confluent with vertex; supraantennal tubercles weakly developed; eyes large and strongly convex, moderately coarsely faceted, kidneyshaped with deep posterior emargination, nearly transverse in relation to the long axis of head. Punctures on head dorsum fine and sparse, inconspicuous; setae long, sparse, suberect to erect, those on vertex mostly directed caudad, thick bristles absent. Antennae (Fig. 19) slender, with distinctly delimited club composed of antennomeres IX – XI, AnL 0.30 mm; antennomeres I – II elongate, III – VII each about as broad as long; VIII – X each strongly transverse; XI 1.4 x as long as broad, slightly asymmetrical, with subconical, blunt apex. Pronotum (Fig. 19) in dorsal view sub-oval, broadest between middle and posterior third; PL 0.23 mm, PW 0.23 – 0.25 mm (mean 0.24 mm); anterior margin weakly arcuate; front angles rounded and indistinct; sides broadly rounded; hind angles well-marked and obtuse; posterior margin weakly arcuate; base of pronotum with shallow and small but distinct pair of external lateral pits, each adjacent to short longitudinal sub-lateral carina. Punctures on pronotal disc fine and inconspicuous; setae long, moderately dense and suberect, sides of pronotum with dense thick and long bristles, especially in posterior half. Elytra (Fig. 19) oval and only slightly more convex than pronotum, broadest slightly anterior to middle, EL 0.45 – 0.46 mm (mean 0.45 mm), EW 0.38 mm, EI 1.20 – 1.23 (mean 1.21); humeral calli distinct, developed as short longitudinal protuberances; basal impressions short but distinct, basal pits well-visible; elytral apices separately rounded. Punctures on elytral disc nearly as fine as those on pronotum; setae short and sparse, suberect. Hind wings well-developed, about twice as long as elytra. Legs (Fig. 19) moderately long and slender, without modifications. Aedeagus (Figs. 30 – 31) elongate, AeL 0.13 mm, thin-walled, in two studied and previously dissected males slightly distorted during previous preparations, oval in shape, with internal armature composed of three elongate and curved sclerites. Female (Figs. 20, 26 – 29). Similar to male but clearly differing in rudimentary eyes, each composed of a single ommatidium. BL 0.86 – 0.88 mm (mean 0.87 mm); HL 0.18 mm, HW 0.19 mm, AnL 0.28 mm; PL 0.24 mm, PW 0.24 – 0.25 mm (mean 0.25 mm); EL 0.45 – 0.46 mm (mean 0.45 mm), EW 0.38 – 0.39 mm (mean 0.38 mm), EI 1.16 – 1.23 (mean 1.18).	en	Jałoszyński, Paweł (2013): Taxonomy of ' Euconnus complex'. Part II. Revision of Archiconnus Franz, Parapseudoconnus Franz and Mexiconnus gen. nov. (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 3666 (4): 523-543, DOI: 10.11646/zootaxa.3666.4.7
C853F731241C0C4AFF34F9E4B3B9FF4C.taxon	distribution	Distribution. North-western Brazil, Amazonas State.	en	Jałoszyński, Paweł (2013): Taxonomy of ' Euconnus complex'. Part II. Revision of Archiconnus Franz, Parapseudoconnus Franz and Mexiconnus gen. nov. (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 3666 (4): 523-543, DOI: 10.11646/zootaxa.3666.4.7
C853F731241C0C4AFF34F9E4B3B9FF4C.taxon	discussion	Remarks. Two specimens must be excluded from the type series of Parapseudoconnus aberrans due to misidentifications: one female belongs to the same genus but clearly differs in the general body shape and certainly is not conspecific with the holotype male; another specimen has not only a different body shape but also a distinct transverse groove on the pronotal base and belongs either to Euconnus or Protoconnus Franz. Both females were collected in a different site than P. aberrans, as evident from the numbers handwritten on the reverse side of locality labels (45 for P. aberrans and 32 and 38 for the two misidentified females). In the original description Franz (1980) indeed gave further collecting data that confirm this suspicion: the holotype and six paratypes were collected on the road to Pacaás Novas, 9.5 km from Guayara-Mirim, while two remaining specimens come from Reserve Ducke and Lago Janauaca near Manaus.	en	Jałoszyński, Paweł (2013): Taxonomy of ' Euconnus complex'. Part II. Revision of Archiconnus Franz, Parapseudoconnus Franz and Mexiconnus gen. nov. (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 3666 (4): 523-543, DOI: 10.11646/zootaxa.3666.4.7
C853F73124000C4BFF34FF0CB26FF9A1.taxon	description	Vetusteconnus Franz, 1993: 101 (unnecessary replacement name for Pseudoconnus Franz). Revised diagnosis. A subgenus of Parapseudoconnus showing the following differences in relation to Parapseudoconnus s. str.: prothoracic hypomeral ridges complete; anterior margins of mesocoxal cavities distinctly demarcated by carina; mesocoxal sockets located on mesal surface of mesocoxal projections and therefore not visible in ventral view; posterior lobes of mesocoxal projections absent; and the internal armature of the aedeagus with long and slender flagellum projecting distally beyond the median lobe. Redescription. Body of male (Fig. 45) strongly convex, elongate and relatively slender, with long appendages, vestiture distinct but unremarkable. Head (Fig. 42) as in Parapseudoconnus s. str., without important differences. Prothorax (Fig. 33) differs from that of Parapseudoconnus s. str. in ventral structures: internal parts of hypomera demarcated laterally by complete hypomeral ridges (Fig. 33; hyr), and pronotosternal sutures (Fig. 33; nss) strongly curved laterally, so that the basisternal part of prosternum is expanded laterally. Mesoventrite (Fig. 33) as in Parapseudoconnus s. str., except for indistinctly demarcated asetose impressions (Fig. 34; ai), presence of distinct carinae demarcating antrerior margins of mesocoxal cavities (Fig. 34; mscc), and mesocoxal projections (Fig. 34; mcp), which conceal mesocoxal sockets (Fig. 34; mscs) located on their mesal margins; posterior lobes of mesocoxal projections absent. Metaventrite (Fig. 34) as in Parapseudoconnus s. str., except for broader metaventral intercoxal process (Fig. 34: mtvp). Elytra (Fig. 45) as in Parapseudoconnus s. str. Legs (Figs. 21, 33 – 34) long; femora clavate, tibiae slender, tarsi moderately elongate. Abdominal sternites in the only studied specimen not preserved. Aedeagus (Figs. 35 – 36) partly damaged or distorted in studied specimens, so the shape of its distal part is not possible to describe, with thin walls and complex, asymmetrical internal armature composed of a system of irregular sclerites, with long flagellum projecting distally beyond median lobe.	en	Jałoszyński, Paweł (2013): Taxonomy of ' Euconnus complex'. Part II. Revision of Archiconnus Franz, Parapseudoconnus Franz and Mexiconnus gen. nov. (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 3666 (4): 523-543, DOI: 10.11646/zootaxa.3666.4.7
C853F73124000C4BFF34FF0CB26FF9A1.taxon	distribution	Distribution and composition. Three species are known to occur in the northern part of Peru.	en	Jałoszyński, Paweł (2013): Taxonomy of ' Euconnus complex'. Part II. Revision of Archiconnus Franz, Parapseudoconnus Franz and Mexiconnus gen. nov. (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 3666 (4): 523-543, DOI: 10.11646/zootaxa.3666.4.7
C853F73124000C4BFF34FF0CB26FF9A1.taxon	discussion	Remarks. Specimens of Parapseudoconnus belonging to Neuraphomimus are poorly preserved in NHMW and only the male of P. simulator is nearly intact, while the type specimens of the remaining two species are partly or nearly completely damaged. The aedeagi, partly damaged or distorted during previous studies, are fragile and they were remounted only to ensure their safety and visibility in the mounting medium (previously the aedeagi were mounted in such a way that they were barely visible), as described in the Materials and Methods section. No attempts were made to obtain preparations with the aedeagi in a perfectly dorsal or ventral position, which would have required additional manipulations and might have caused further damage. Therefore, the aedeagi are here illustrated as they are embedded, not necessarily in dorsal or lateral aspects. In order to identify new specimens their aedeagi must be studied in various positions to carefully examine the shape of internal armature and compare it with that showed in Figs. 35 – 36 and in Franz (1980; Fig. 205).	en	Jałoszyński, Paweł (2013): Taxonomy of ' Euconnus complex'. Part II. Revision of Archiconnus Franz, Parapseudoconnus Franz and Mexiconnus gen. nov. (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 3666 (4): 523-543, DOI: 10.11646/zootaxa.3666.4.7
C853F73124000C49FF34F916B44BFB63.taxon	materials_examined	Material studied. Holotype: 3: three labels (Fig. 23): " Umg. Tarapoto / Peru, lg. Franz " with " SA 137 [or / 37] " on the reverse side [white, printed; reverse handwritten], " Pseudoconnus / simulator / 3 m. / det. H. Franz " [white, handwritten and printed], " Typus " [red, handwritten] (NHMW).	en	Jałoszyński, Paweł (2013): Taxonomy of ' Euconnus complex'. Part II. Revision of Archiconnus Franz, Parapseudoconnus Franz and Mexiconnus gen. nov. (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 3666 (4): 523-543, DOI: 10.11646/zootaxa.3666.4.7
C853F73124000C49FF34F916B44BFB63.taxon	diagnosis	Diagnosis. Aedeagus with internal armature as in Fig. 35. Redescription. Body of male (Fig. 21) moderately strongly convex and elongate, with moderately long appendages, BL 0.75 mm; glossy, uniformly light brown with slightly lighter appendages; vestiture slightly lighter than cuticle. Head (Figs. 21, 32) approximately rhomboidal and slightly transverse, broadest at eyes, HL 0.15 mm, HW 0.18 mm; occipital constriction slightly broader than half HW; tempora slightly shorter than eyes, only slightly rounded and strongly convergent caudad; vertex distinctly broader than long, convex, only slightly (not conically) projected dorso-caudad, without postero-median impression; frons subtriangular, confluent with vertex; supraantennal tubercles weakly developed; eyes large and strongly convex, moderately coarsely faceted, kidney-shaped with deep posterior emargination, nearly transverse in relation to the long axis of head. Punctures on head dorsum fine and sparse, inconspicuous; setae moderately long, sparse, suberect to erect, those on vertex mostly directed caudad, thick bristles absent. Antennae (Fig. 21) slender, with distinctly delimited club composed of antennomeres IX – XI, AnL 0.30 mm; antennomeres I – II elongate, III – VIII about as broad as long or only slightly elongate; VIII – X distinctly transverse; XI 1.7 x as long as broad, slightly asymmetrical, with subconical, blunt apex. Pronotum (Fig. 21) in dorsal view sub-oval, broadest between middle and posterior third; PL 0.20 mm, PW 0.20 mm; anterior margin strongly arcuate; front angles rounded and indistinct; sides broadly rounded; hind angles moderately distinct and obtuse; posterior margin weakly arcuate; base of pronotum with shallow and small but distinct pair of external lateral pits, each adjacent to barely marked longitudinal sub-lateral carina. Punctures on pronotal disc fine and inconspicuous; setae long, sparse and suberect to erect, sides of pronotum with dense thick and very long bristles. Elytra (Fig. 21) oval and only slightly more convex than pronotum, broadest distinctly anterior to middle, EL 0.40 mm, EW 0.33 mm, EI 1.23; humeral calli distinct, developed as short longitudinal protuberances; basal impressions short but distinct, basal pits well-visible; elytral apices separately rounded. Punctures on elytral disc nearly as fine as those on pronotum; setae short and sparse, only slightly suberect. Hind wings well-developed, about twice as long as elytra. Legs (Fig. 21) moderately long and slender, without modifications. Aedeagus elongate, AeL 0.11 mm, as in Fig. 35. Female. Unknown (see Remarks).	en	Jałoszyński, Paweł (2013): Taxonomy of ' Euconnus complex'. Part II. Revision of Archiconnus Franz, Parapseudoconnus Franz and Mexiconnus gen. nov. (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 3666 (4): 523-543, DOI: 10.11646/zootaxa.3666.4.7
C853F73124000C49FF34F916B44BFB63.taxon	distribution	Distribution. Northern Peru, San Martin Province.	en	Jałoszyński, Paweł (2013): Taxonomy of ' Euconnus complex'. Part II. Revision of Archiconnus Franz, Parapseudoconnus Franz and Mexiconnus gen. nov. (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 3666 (4): 523-543, DOI: 10.11646/zootaxa.3666.4.7
C853F73124000C49FF34F916B44BFB63.taxon	discussion	Remarks. The holotype male is accompanied in NHMW by a highly similar, but microphthalmous female labeled as Pseudoconnus? simulator, and bearing the same collecting data. It is not possible to verify whether these specimens are conspecific or not without collecting a larger series composed of both sexes from the same spot. In the original description, Franz (1980) mentions that the type series is composed of a male holotype and female paratype, but in the Franz Coll. only this provisionally identified female is present, not labeled as paratype. In the original description Franz (1980) gives further collecting details: road Tarapoto-Yurimagua, 80 km from Tarapoto, between Pongo de Caimarachi and Shanussi, in litter of a bog or swamp forest (" Sumpfwald "), 0 1.10.1968.	en	Jałoszyński, Paweł (2013): Taxonomy of ' Euconnus complex'. Part II. Revision of Archiconnus Franz, Parapseudoconnus Franz and Mexiconnus gen. nov. (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 3666 (4): 523-543, DOI: 10.11646/zootaxa.3666.4.7
C853F73124020C4EFF34FADFB414FB93.taxon	materials_examined	Material studied. Remnants of the holotype preparation: 3: three labels (Fig. 24): " Umg. Tarapoto / Peru, lg. Franz " with " SA 137 [or / 37] " on the reverse side [white, printed; reverse handwritten], " Pseudoconnus / fraudulentus / 3 m. / det. H. Franz " [white, handwritten and printed], " Typus " [red, handwritten] (NHMW).	en	Jałoszyński, Paweł (2013): Taxonomy of ' Euconnus complex'. Part II. Revision of Archiconnus Franz, Parapseudoconnus Franz and Mexiconnus gen. nov. (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 3666 (4): 523-543, DOI: 10.11646/zootaxa.3666.4.7
C853F73124020C4EFF34FADFB414FB93.taxon	diagnosis	Diagnosis. Aedeagus with internal armature as in Fig. 36. Redescription. The only known specimen (male holotype) is damaged (the mounting card bears a droplet of glue with small fragments of the basal abdominal sternite and hind coxae) except for the aedeagus (Fig. 36), which is distorted, but with apparently intact internal armature; AeL 0.10.	en	Jałoszyński, Paweł (2013): Taxonomy of ' Euconnus complex'. Part II. Revision of Archiconnus Franz, Parapseudoconnus Franz and Mexiconnus gen. nov. (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 3666 (4): 523-543, DOI: 10.11646/zootaxa.3666.4.7
C853F73124020C4EFF34FADFB414FB93.taxon	description	Female. Unknown.	en	Jałoszyński, Paweł (2013): Taxonomy of ' Euconnus complex'. Part II. Revision of Archiconnus Franz, Parapseudoconnus Franz and Mexiconnus gen. nov. (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 3666 (4): 523-543, DOI: 10.11646/zootaxa.3666.4.7
C853F73124020C4EFF34FADFB414FB93.taxon	distribution	Distribution. Northern Peru, San Martin Province.	en	Jałoszyński, Paweł (2013): Taxonomy of ' Euconnus complex'. Part II. Revision of Archiconnus Franz, Parapseudoconnus Franz and Mexiconnus gen. nov. (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 3666 (4): 523-543, DOI: 10.11646/zootaxa.3666.4.7
C853F73124020C4EFF34FADFB414FB93.taxon	discussion	Remarks. In the original description Franz (1980) gives further collecting details: road Tarapoto-Yurimagua, 80 km from Tarapoto, between Pongo de Caimarachi and Shanussi, in litter of a bog or swamp forest (" Sumpfwald "), 0 1.10.1968. The damaged holotype, with only a partly distorted aedeagus left, makes it impossible to verify the generic and subgeneric status of P. fraudulentus. However, the internal armature of the aedeagus seems to be preserved in a non-distorted state and it may be possible to identify new specimens, preferably by direct comparisons of their aedeagi to the holotype preparation, and not only by comparing with the Fig. 36. A possibility of a synonymy between P. simulator and P. fraudulentus cannot be excluded, as their aedeagi are highly similar and both holotypes were probably collected in the same spot. Parapseudoconnus (Neuraphomimus) monticola Franz (Fig. 25)	en	Jałoszyński, Paweł (2013): Taxonomy of ' Euconnus complex'. Part II. Revision of Archiconnus Franz, Parapseudoconnus Franz and Mexiconnus gen. nov. (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 3666 (4): 523-543, DOI: 10.11646/zootaxa.3666.4.7
C853F73124020C4EFF34FADFB414FB93.taxon	description	Pseudoconnus (s. str.) monticola Franz, 1980: 217, Fig. 205.	en	Jałoszyński, Paweł (2013): Taxonomy of ' Euconnus complex'. Part II. Revision of Archiconnus Franz, Parapseudoconnus Franz and Mexiconnus gen. nov. (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 3666 (4): 523-543, DOI: 10.11646/zootaxa.3666.4.7
C853F73124020C4EFF34FADFB414FB93.taxon	materials_examined	Material studied. Remnants of the holotype preparation: three labels (Fig. 25): " Umg. Tarapoto / Peru, lg. Franz " with " SA 136 [or / 36] " on the reverse side [white, printed; reverse handwritten], " Pseudoconnus / monticola / 3 m. / det. H. Franz " [white, handwritten and printed], " Typus " [red, handwritten] (NHMW). This species is known from a single holotype male only (but see also Remarks), and the specimen preserved in the Franz Coll. (NHMW) is damaged to the extent that makes it impossible to identify the species. Only the head, a part of prothorax and partly also fore legs are preserved. The original genital preparation in euparal or Canada balsam on a plastic mounting card has been in the past pinned in such a way that the card folded in half longitudinally and the mounting medium cracked and most of it has fallen off. A careful examination revealed that the remnants of the mounting medium do not contain the aedeagus or any other structures. Franz (1980, Fig. 205) illustrated the aedeagus of P. m o n t i c o l a and it may be possible to identify new specimens using this illustration. Only then a neotype can be designated.	en	Jałoszyński, Paweł (2013): Taxonomy of ' Euconnus complex'. Part II. Revision of Archiconnus Franz, Parapseudoconnus Franz and Mexiconnus gen. nov. (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 3666 (4): 523-543, DOI: 10.11646/zootaxa.3666.4.7
C853F73124020C4EFF34FADFB414FB93.taxon	discussion	Remarks. Franz (1980) in the original description mentions that the type series of P. m o n t i c o l a is composed of the holotype male and a paratype female. The male in the Franz Coll. (NHMW) is accompanied by another partly damaged specimen (only abdomen and the ventral part of pterothorax with three legs survived), but identified by Franz as a female of P. monticola with a question mark and not labeled as paratype. The identity (even to genus level) of these remnants is not possible. In the original description Franz (1980) gives further collecting details for P. monticola: road Tarapoto-Yurimagua, mountains 40 km from Tarapoto, in a forest litter, 0 1.10.1968.	en	Jałoszyński, Paweł (2013): Taxonomy of ' Euconnus complex'. Part II. Revision of Archiconnus Franz, Parapseudoconnus Franz and Mexiconnus gen. nov. (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 3666 (4): 523-543, DOI: 10.11646/zootaxa.3666.4.7
