taxonID	type	description	language	source
C84287C40C3AEB6BFDF424D3183E7904.taxon	type_taxon	Type species Fusus niponicus E. A. Smith, 1879, by original designation. Genus composition In addition to species previously assigned to the genus (Hadorn & Fraussen 2005), we include sequenced species Granulifusus discrepans, G. williami comb. nov., G. jeanpierrevezzaroi comb. nov., G. annae sp. nov. and G. norfolkensis sp. nov. In addition to those species we allocate to Granulifusus, based on conchological similarities to sequenced species, G. tatianae sp. nov., G. kurodai comb. nov., and G. guidoi sp. nov., known from empty shells only.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C3AEB68FE4222A918817B10.taxon	description	Figs 3 A – E, 4 A	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C3AEB68FE4222A918817B10.taxon	diagnosis	Molecular diagnosis A molecular diagnosis is given in Table 2.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C3AEB68FE4222A918817B10.taxon	materials_examined	Type material Holotype JAPAN: off Cape Ashizuri, southwestern Shikoku (NSMT Mo 40377) (not seen). Sequenced material SOLOMON SEA: 1 lv, MADEEP, stn CP 4337, SE Ainto Bay, New Britain, 06 ° 07 ’ S, 149 ° 17 ’ E, 287 – 447 m (MNHN IM- 2013 - 46327). PHILIPPINES: AURORA 2007, 1 lv, stn CP 2657, 16 ° 01 ′ N, 121 ° 51 ′ E, 342 – 358 m (MNHN IM- 2007 - 34604); 2 lv, stn CP 2673, 15 ° 01 ′ N, 121 ° 45 ′ E, 431 – 493 m (MNHN IM- 2007 - 34583, MNHN IM- 2007 - 34488); 2 lv, stn CP 2732, 15 ° 27 ′ N, 121 ° 36 ′ E, 556 m (MNHN IM- 2007 - 34580, MNHN IM- 2007 - 34581). Other examined material PAPUA NEW GUINEA: 1 lv, PAPUA NIUGINI, stn CP 3949, 5 ° 12 ′ S, 145 ° 51 ′ E, 380 – 407 m (MHNH 2013 - 15374, not sequenced); 1 lv, stn CP 4023, 5 ° 22 ′ S; 145 ° 48 ′ E, 340 – 385 m (MNHN IM- 2013 - 9777, not sequenced).	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C3AEB68FE4222A918817B10.taxon	description	Description Shell large, with multispiral protoconch of 2.75 conical, smooth, convex, grayish whorls, last ¼ whorl with 6 – 7 rounded axial riblets progressing in strength, protoconch / teleoconch boundary without varix, defined by appearance of spiral sculpture. Protoconch diameter 960 µm, exposed height 890 µm (Fig. 3 E). Teleoconch with regularly convex whorls, upper ones slightly angulated, with long slightly sinuous to nearly straight siphonal canal; aperture in young specimens with 3 – 4 plicae, in adult with a single columellar plica, delimitating siphonal canal, and sometimes with indistinct parietal tooth. Axial sculpture of distinct axial ribs, 7 – 8 per whorl on upper whorls, obsolete on posterior whorls in shells over 50 mm. Spiral sculpture of 2 – 3 raised cords on periphery of upper whorls and numerous thin riblets of even strength. On posterior whorls, cords obsolete and some riblets more pronounced. Background colour tan, with darker brown spiral bands along suture. Radula (MNHN IM- 2013 - 46327, SL 18.2 mm, Fig. 4 A) 1.83 mm in length (32 % of AL without canal), of 75 transverse rows of teeth, of which 35 nascent. Radula narrow, distance between outer edges of lateral teeth about 65 µm (1.12 % of AL). Lateral teeth medium broad, arcuate, with attenuated outer anterior corner, producing short ‘ handle’. Lateral teeth with five closely spaced, slightly curved cusps similar in length, small outermost cusp (seen on lower right lateral tooth on Fig. 4 A), and much smaller but distinct innermost cusp. Central tooth small, very narrow, trapezoid, with three subequal cusps. Narrow but distinct transverse folds of radular membrane situated serially between rows of lateral teeth. Couto (2016: fig. 58) illustrated the radula of an adult specimen (SL 66 mm). The lateral teeth were relatively much broader, with 6 main cusps nearly identical in size and an additional innermost cusp. The radula had a similar width (1.1 % of AL, measurements taken from photo) to the one studied herein; the differences in teeth morphology can possibly be explained by ontogenetic variability.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C3AEB68FE4222A918817B10.taxon	distribution	Distribution From Japan throughout the East China Sea to the Philippines and the Solomons.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C3AEB68FE4222A918817B10.taxon	discussion	Remarks This is the largest species of Granulifusus, which attains a shell length over 90 mm (http: // www. conchology. be, last accessed 6 Mar. 2017). Due to the ontogenic change in sculpture pattern, the shell morphology differs from that in other Recent members of the genus, but the radula is similar. Young specimens bear a superficial resemblance to specimens of the Miocene species Pseudolatirus bilineatus of similar size (compare Fig. 3 C, G with Fig. 3 D’, F’).	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C39EB6DFE0920BD1E0B7A8C.taxon	description	Figs 3 H – K, 4 B	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C39EB6DFE0920BD1E0B7A8C.taxon	diagnosis	Molecular diagnosis A molecular diagnosis is given in Table 2.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C39EB6DFE0920BD1E0B7A8C.taxon	materials_examined	Type material Holotype PHILIPPINES: off Dipolog, Aliguay Island, from 60 to 150 m, in the National Museum of the Philippines, Manila (not seen). Paratypes PHILIPPINES: 4 specimens in the former collection of E. de Suduiraut and Conchology, Inc., Mactan Island (not seen). Sequenced material PHILIPPINES: AURORA 2007, 1 lv, stn CP 2709, 15 ° 12 ′ N, 121 ° 34 ′ E, 244 – 296 m (MNHN IM- 2007 - 34608). NEW CALEDONIA: CONCALIS, 1 lv, stn CP 2960, North of New Caledonia, Grand Passage, (19 ° 05 ′ S, 163 ° 13 ′ E, 382 – 387 m (MNHN IM- 2007 - 35734). SOLOMON ISLANDS: SALOMON 2, 1 lv, stn DW 2169, 09 ° 01 ′ S, 159 ° 06 ′ E, 100 – 200 m (MNHN IM- 2007 - 32612). SOCIETY ISLANDS: TARASOC, 2 lv, stn DW 3441, 16 ° 43 ′ S, 151 ° 26 ′ W, 350 – 360 m (MNHN IM- 2007 - 39387, MNHN IM- 2007 - 39389); 1 lv, stn DW 3491, 17 ° 29 ′ S, 149 ° 26 ′ W, 440 – 500 m (MNHN IM- 2007 - 39262). MADAGASCAR: ATIMO VATAE, 1 lv, stn DW 3515, south Madagascar, 24 ° 53 ′ S, 47 ° 28 ′ E, 184 – 203 m (MNHN IM- 2009 - 15090). Other material examined (in MNHN if not otherwise mentioned) TAIWAN: 3 dd, TAIWAN 2000, R / V Fishery Researcher No. 1, stn DW 34, 22 ° 02 ′ N, 120 ° 36 ′ E, 246 m; 1 dd, stn CP 35, 22 ° 02 ′ N, 120 ° 27 ′ E, 246 m; 1 dd, stn DW 36, 21 ° 55 ′ N, 120 ° 36 ′ E, 305 m. PHILIPPINES: 1 spm, Punta Engano, Mactan, in nets at 80 – 100 m (ANSP 456858); 1 spm, Olango, in lumun lumun nets, 40 – 100 m (ANSP 456859); 1 spm, Aliguay Island, trawled at 200 m (ANSP 456857); 4 spms, ‘ by boats from the Philippines’ (ANSP 456854); 5 spms, same data as preceding (ANSP 456861); 2 spms, Balicasag Island, in tangle nets, 80 – 120 m (ANSP 456855); 2 spms, Aliguay Island, in coral rubble, 140 m (ANSP 456856); 2 spms, Aliguay Island, dredged on sandy / rocky mud, 160 m (ANSP 456860). – MUSORSTOM 3: 1 dd, stn CP 96, 14 ° 00 ′ N, 120 ° 18 ′ E, 190 – 194 m. ARAFURA SEA: KARUBAR: 5 dd, stn DW 29, 05 ° 36 ′ S, 132 ° 56 ′ E, 181 – 184 m; 1 dd, stn DW 32, 05 ° 47 ′ S, 132 ° 51 ′ E, 170 – 206 m. SOLOMON ISLANDS: SALOMON 1: 6 dd, stn DW 1745, 09 ° 23 ′ S, 159 ° 59 ′ E, 253 – 356 m; 1 dd, stn DW 1758, 08 ° 49 ′ S, 159 ° 52 ′ E, 180 – 187 m; 1 lv, 8 dd, stn DW 1768, 08 ° 21 ′ S, 160 ° 42 ′ E, 194 – 286 m; 3 dd, stn DW 1778, 08 ° 19 ′ S, 160 ° 34 ′ E, 157 – 253 m; 1 lv, 6 dd, stn DW 1788, 09 ° 19 ′ S, 160 ° 15 ′ E, 341 – 343 m; 1 dd, stn DW 1811, 09 ° 46 ′ S, 160 ° 51 ′ E, 182 – 203 m; 1 dd, stn DW 1813, 09 ° 47 ′ S, 160 ° 53 ′ E, 201 – 227 m; 3 dd, stn DW 1820, 09 ° 52 ′ S, 160 ° 51 ′ E, 256 – 329 m; 1 juv. dd, stn DW 1834, 10 ° 12 ′ S, 161 ° 18 ′ E, 225 – 281 m. PAPUA NEW GUINEA: KAVIENG 2014: 1 lv, stn DW 4471, 02 ° 44 ′ S, 150 ° 38 ′ E, 126 – 220 m (MNHN IM- 2013 - 58726, not sequenced); 1 lv, stn DW 4484, 02 ° 26 ′ S, 149 ° 55 ′ E, 229 m (MNHN IM- 2013 - 58866, not sequenced). NEW CALEDONIA: BIOCAL: 1 dd, stn DW 43, 22 ° 46 ′ S, 167 ° 15 ′ E, 400 m; 1 dd, stn DW 82, 20 ° 31 ′ S, 166 ° 50 ′ E, 440 – 460 m; 1 dd, stn DW 83, 20 ° 35 ′ S, 166 ° 54 ′ E, 460 m. – BIOGEOCAL: 5 dd, stn KG 252, 21 ° 31 ′ S, 166 ° 21 ′ E, 330 m; 16 juv. dd, stn DW 253, 21 ° 32 ′ S, 166 ° 29 ′ E, 310 – 315 m; 1 dd, stn DW 292, 20 ° 28 ′ S, 166 ° 48 ′ E, 465 – 470 m; 3 dd, stn DW 308, 20 ° 40 ′ S, 166 ° 58 ′ E, 510 – 590 m. – SMIB 2: 2 dd, stn DW 23, 22 ° 31 ′ S, 167 ° 37 ′ E, 410 – 420 m. – SMIB 5: 3 dd, stn DW 88, 22 ° 19 ′ S, 166 ° 40 ′ E, 350 m; 1 dd, stn DW 98, 23 ° 02 ′ S, 168 ° 16 ′ E, 335 m. – SMIB 6: 1 lv, stn DW 126, 18 ° 59 ′ S, 163 ° 23 ′ E, 320 – 330 m. – SMIB 8: 3 dd, stn DW 165, 24 ° 47 ′ S, 168 ° 10 ′ E, 372 – 660 m; 1 dd, stn DW 187, 23 ° 17 ′ S, 168 ° 06 ′ E, 390 – 540 m; 3 dd, stn DW 189, 23 ° 18 ′ S, 168 ° 06 ′ E, 400 – 402 m. – CHALCAL 2: 1 lv, 2 dd, stn DW 69, 24 ° 44 ′ S, 168 ° 08 ′ E, 260 m. – BATHUS 1: 2 dd, stn DW 683, 20 ° 35 ′ S, 165 ° 07 ′ E, 380 – 400 m; 1 dd, stn DW 687, 20 ° 35 ′ S, 165 ° 07 ′ E, 408 – 440 m; 1 dd, stn DW 688, 20 ° 33 ′ S, 165 ° 00 ′ E, 270 – 282 m; 1 juv. lv, stn CP 698, 20 ° 34 ′ S, 164 ° 57 ′ E, 491 – 533 m; 1 dd, stn CP 701, 20 ° 58 ′ S, 165 ° 36 ′ E, 302 – 335 m. – BATHUS 2: 1 lv, 3 dd, stn DW 724, 22 ° 48 ′ S, 167 ° 26 ′ E, 344 – 358 m; 2 lv, 7 dd, stn DW 730, 23 ° 03 ′ S, 166 ° 58 ′ E, 397 – 400 m; 3 lv, stn DW 731, 22 ° 49 ′ S, 166 ° 45 ′ E, 300 – 370 m; 1 dd, stn CP 759, 22 ° 18 ′ S, 166 ° 10 ′ E, 370 – 420 m. – BATHUS 3: 8 lv + dd, stn DW 827, 23 ° 22 ′ S, 168 ° 01 ′ E, 381 – 469 m; 1 lv, stn DW 838, 23 ° 01 ′ S, 166 ° 56 ′ E, 400 – 402 m. – BATHUS 4: 2 dd, stn DW 885, 22 ° 05 ′ S, 165 ° 58 ′ E, 250 – 300 m; 3 dd, stn DW 887, 21 ° 07 ′ S, 164 ° 28 ′ E, 320 – 344 m; 1 lv, stn CP 889, 21 ° 01 ′ S, 164 ° 27 ′ E, 416 – 433 m; 6 dd, stn DW 902, 19 ° 01 ′ S, 163 ° 15 ′ E, 341 – 351 m; 2 dd, stn CP 907, 19 ° 01 ′ S, 163 ° 13 ′ E, 370 – 394 m; 1 dd, stn DW 943, 20 ° 12 ′ S, 164 ° 31 ′ E, 316 – 347 m. – BERYX 11: 15 lv + dd, stn DW 11 / CP 23, 24 ° 43 ′ – 24 ° 44 ′ S, 168 ° 08 ′ – 168 ° 10 ′ E, 270 – 350 m. – MUSORSTOM 5: 2 lv, 3 dd, stn 301, 22 ° 07 ′ S, 159 ° 25 ′ E, 487 – 610 m, Nova Bank. – MUSORSTOM 6: 3 lv, 3 dd, stn DW 391, 20 ° 47 ′ S, 167 ° 06 ′ E, 390 m; 1 lv, 2 dd, stn DW 392, 20 ° 47 ′ S, 167 ° 05 ′ E, 340 m; 1 dd, stn DW 393, 20 ° 48 ′ S, 167 ° 10 ′ E, 420 m; 3 dd, stn DW 397, 20 ° 47 ′ S, 167 ° 05 ′ E, 380 m; 1 lv, 2 dd, stn DW 398, 20 ° 47 ′ S, 167 ° 06 ′ E, 370 m; 12 lv + dd, stn DW 406, 20 ° 41 ′ S, 167 ° 07 ′ E, 373 m; 3 dd, stn DW 411, 20 ° 40 ′ S, 167 ° 03 ′ E, 424 m; 1 lv, 1 dd, stn DW 413, 20 ° 40 ′ S, 167 ° 03 ′ E, 463 m; 3 dd, stn DW 416, 20 ° 42 ′ S, 167 ° 00 ′ E, 343 m; 1 lv, stn DW 421, 20 ° 26 ′ S, 166 ° 40 ′ E, 245 m; 1 dd, stn DW 428, 20 ° 24 ′ S, 166 ° 13 ′ E, 420 m; 1 dd, stn DW 451, 20 ° 59 ′ S, 167 ° 25 ′ E, 330 m; 2 lv, 5 dd, stn DW 457, 21 ° 00 ′ S, 167 ° 29 ′ E, 353 m; 2 lv, 3 dd, stn DW 479, 21 ° 09 ′ S, 167 ° 55 ′ E, 310 m; 1 lv, stn DW 480, 21 ° 08 ′ S, 167 ° 56 ′ E, 380 m; 2 lv, 2 dd, stn DW 481, 21 ° 22 ′ S, 167 ° 50 ′ E, 300 m. – VOLSMAR: 1 dd, stn DW 9, 22 ° 23 ′ S, 171 ° 41 ′ E, 275 – 300 m; 1 dd, stn DW 38, 22 ° 22 ′ S, 168 ° 44 ′ E, 380 – 420 m; 1 dd, stn DW 51, 20 ° 59 ′ S, 170 ° 03 ′ E, 450 m. – NORFOLK 1: 3 lv, stn DW 1680, 24 ° 45 ′ S, 168 ° 10 ′ E, 385 – 392 m. VANUATU: MUSORSTOM 8: 1 lv, stn DW 963, 20 ° 20 ′ S, 169 ° 49 ′ E, 400 – 440 m; 1 dd, stn DW 964, 20 ° 20 ′ S, 169 ° 49 ′ E, 360 – 408 m; 1 dd, stn DW 967, 20 ° 19 ′ S, 169 ° 53 ′ E, 295 – 334 m; 3 dd, stn DW 977, 19 ° 25 ′ S, 169 ° 29 ′ E, 410 – 505 m; 8 dd, stn DW 978, 19 ° 23 ′ S, 169 ° 27 ′ E, 408 – 413 m; 3 dd, stn DW 988, 19 ° 16 ′ S, 169 ° 24 ′ E, 372 – 466 m; 1 dd, stn DW 1060, 16 ° 14 ′ S, 167 ° 21 ′ E, 375 – 397 m; 3 dd, stn DW 1065, 16 ° 16 ′ S, 167 ° 21 ′ E, 360 – 419 m. WALLIS AND FUTUNA: MUSORSTOM 7: 3 lv (?), stn DW 510, 14 ° 14 ′ S, 178 ° 11 ′ W, 280 – 370 m. FIJI: MUSORSTOM 10: 9 lv + dd, stn CP 1325, 17 ° 16 ′ S, 177 ° 50 ′ E, 282 – 322 m; 2 lv, stn CP 1326, 17 ° 14 ′ S, 177 ° 50 ′ E, 265 – 300 m; 7 lv + dd, stn CP 1327, 17 ° 13 ′ S, 177 ° 52 ′ E, 370 – 389 m; 1 dd, stn DW 1384, 18 ° 19 ′ S, 178 ° 06 ′ E, 260 – 305 m; 7 dd, stn DW 1388, 18 ° 19 ′ S, 178 ° 02 ′ E, 313 – 446 m; 2 lv, stn CP 1389, 18 ° 19 ′ S, 178 ° 05 ′ E, 241 – 417 m; 1 lv, 2 dd, stn CP 1390, 18 ° 19 ′ S, 178 ° 05 ′ E, 234 – 361 m. – BORDAU 1: 1 dd, stn DW 1417, 16 ° 27 ′ S, 178 ° 55 ′ W, 353 m; 1 lv, 1 dd, stn DW 1450, 16 ° 44 ′ S, 179 ° 58 ′ E, 327 – 420 m; 1 lv, stn CP 1476, 19 ° 41 ′ S, 178 ° 11 ′ W, 310 – 420 m; 2 lv, 3 dd, stn DW 1479, 20 ° 58 ′ S, 178 ° 45 ′ W, 450 – 460 m; 2 lv, stn CP 1481, 20 ° 57 ′ S, 178 ° 45 ′ W, 441 – 506 m; 2 lv, stn DW 1496, 18 ° 43 ′ S, 178 ° 23 ′ W, 392 – 407 m; 3 dd, stn DW 1499, 18 ° 40 ′ S, 178 ° 27 ′ W, 389 – 400 m; 1 lv, stn CP 1500, 18 ° 42 ′ S, 178 ° 26 ′ W, 366 – 389 m. TONGA: BORDAU 2: 2 dd, stn DW 1518, 21 ° 21 ′ S, 175 ° 07 ′ W, 336 – 347 m; 2 dd, stn DW 1534, 21 ° 43 ′ S, 175 ° 19 ′ W, 302 – 327 m; 2 dd, stn DW 1535, 21 ° 43 ′ S, 175 ° 18 ′ W, 268 m; 1 lv, stn DW 1536, 21 ° 45 ′ S, 175 ° 21 ′ W, 320 – 323 m; 1 dd, stn DW 1567, 21 ° 02 ′ S, 175 ° 19 ′ W, 351 – 356 m; 1 dd, stn DW 1583, 18 ° 37 ′ S, 174 ° 03 ′ W, 327 – 360 m; 1 dd, stn DW 1585, 18 ° 33 ′ S, 173 ° 57 ′ W, 578 m; 1 lv, stn DW 1586, 18 ° 34 ′ S, 173 ° 55 ′ W, 440 – 487 m; 1 lv, 3 dd, stn DW 1607, 22 ° 15 ′ S, 175 ° 23 ′ W, 356 – 367 m; 4 dd, stn DW 1631, 23 ° 23 ′ S, 176 ° 18 ′ W, 407 – 443 m; 2 dd, stn DW 1634, 21 ° 45 ′ S, 175 ° 20 ′ W, 321 – 322 m; 2 dd, stn DW 1635, 21 ° 44 ′ S, 175 ° 20 ′ W, 320 – 323 m; 2 dd, stn DW 1636, 21 ° 44 ′ S, 175 ° 20 ′ W, 321 – 331 m; 1 lv, stn CP 1641, 21 ° 09 ′ S, 175 ° 22 ′ W, 395 m,. RÉUNION: MD 32: 9 dd, stn DC 128, 20 ° 51 ′ S, 55 ° 36 ′ E, 280 – 340 m; 10 dd, stn CP 129, 20 ° 51 ′ S, 55 ° 36 ′ E, 290 – 300 m; 1 dd, stn DC 136, 20 ° 46 ′ S, 55 ° 36 ′ E, 915 – 922 m.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C39EB6DFE0920BD1E0B7A8C.taxon	description	Description Shell slender, fusiform, small to medium-sized for genus (attaining 30.8 mm, ANSP 456861). Protoconch multispiral of 4.25 whorls (Fig. 3 K), conical, smooth, except scattered microgranules more dense along suture, flesh coloured with reddish thin line along suture. Whorls somewhat convex, last ½ whorl with 11 – 12 rounded axial riblets progressing in strength, protoconch / teleoconch boundary without varix, defined by appearance of spiral sculpture. Protoconch diameter 1000 µm, exposed height 1240 µm. Teleoconch whorls convex, suture deep, adpressed. Axial sculpture of very prominent ribs, 7 – 9 per whorl. Spiral sculpture of evenly spaced strong cords, 2 on first whorl, 3 – 4 on penultimate whorl, 8 – 9 on last whorl and shell base, becoming much weaker on canal. Weaker but distinct secondary cords between primary cords, one on earliest whorls increasing to 5 on last whorl, primary and secondary cords becoming similar in size on canal. Siphonal canal long, open, straight, about ¼ of shell length. Aperture with 7 – 8 smooth teeth periodically appearing and corresponding to interspaces between axial ribs. Inner lip adherent, forming smooth, white columellar callus, 2 – 3 distinct columellar rounded plicae, adapical two more prominent than abapical one. Teleoconch light brown, usually with lighter spiral band on whorls periphery. Radula (MNHN IM- 2013 - 58726, SL 14.1, AL 3.1 mm, Fig. 4 B) 2.13 mm in length (69 % of AL), of about 125 transverse rows of teeth, of which 30 nascent. Radula narrow, distance between outer edges of lateral teeth about 50 µm (1.6 % of AL without canal). Lateral teeth medium broad, arcuate, with attenuated outer anterior corner, producing long ‘ handle’. Lateral teeth with three cusps on right side and four on left side, subequal in length, closely spaced, slightly curved and an additional small but distinct innermost cusp. Inner cusp (s) slightly narrower than lateral ones. Central tooth small, long trapezoid with narrower base, with three cusps, central one nearly twice as long as outer ones.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C39EB6DFE0920BD1E0B7A8C.taxon	distribution	Distribution Very widely distributed: in Indo-Pacific, from Taiwan and the Philippines, to French Polynesia, Madagascar and off Réunion. Recorded at depths from 170 to 920 m, alive in 225 – 610 m. Unconfirmed depths from commercial sources in the Philippines from 40 m.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C39EB6DFE0920BD1E0B7A8C.taxon	discussion	Remarks The species has a smooth columella and therefore is well differentiated from its congeners with columellar plicae. Besides, G. williami comb. nov. has a small, rather slender shell with attenuated straight canal. It is most similar to Granulifusus norfolkensis sp. nov., differing in details of colouration and in having a straight siphonal canal rather than inclined to left. Molecular data clearly separate the two species.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C3DEB61FE28270319C67C92.taxon	description	Figs 4 C – D, 5	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C3DEB61FE28270319C67C92.taxon	diagnosis	Molecular diagnosis A molecular diagnosis is given in Table 2.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C3DEB61FE28270319C67C92.taxon	materials_examined	Type material Holotype VIETNAM: off Nha Trang, SL 66.3 mm (MMM) (not seen). Sequenced material NEW CALEDONIA: TERRASSES: 1 lv, stn CP 3095, 22 ° 02 ′ S, 167 ° 06 ′ E, 320 – 380 m (MNHN IM- 2007 - 36886). – CONCALIS: 1 lv, stn CP 3026, Grand Passage, 20 ° 17 ′ S, 163 ° 50 ′ E, 590 – 809 m (MNHN IM- 2007 - 35083). PHILLIPPINES: AURORA 2007: 1 lv, stn CP 2748, 15 º 56 ′ N, 121 º 465 ′ E, 247 – 249 m (MNHN IM- 2007 - 34593). Other material examined SOUTH AFRICA: 2 spms, Agulhas Bank, 160 m (ANSP 425332); 1 spm, Agulhas Bank, 160 m (ANSP 425328); 3 spms, off Durban (ANSP 425336). TAIWAN: 2 spms, trawled (ANSP 425339); 2 spms, ‘ by divers under rocks, southeast Sea of Formosa’ (ANSP 425338); 2 spms, SW of Taiwan, 140 – 200 m (ANSP 425335); 2 spms, 200 – 300 m (ANSP 425333); 1 spm, China Sea, off Taiwan (ANSP 425329); 4 spms, trawled (ANSP 425334). MOZAMBIQUE: 3 spms (ANSP 425340). VIETNAM: 1 spm, Nha Trang region, 20 – 35 m (ANSP 425341). SOUTH CHINA SEA: 2 spms, on sand bottom (ANSP 422786); 1 spm, ‘ deep water’ (ANSP 422785). PHILIPPINES: 3 spms, off Bohol, tangle nets (ANSP 425330); 1 spm, Philippine Is (ANSP 425331); 1 dd, off Mindanao (ANSP 425337). – MUSORSTOM 2: 5 lv, stn CP 02, 14 ° 01 ′ N, 120 ° 17 ′ E, 184 – 186 m; 37 lv + dd, stn CP 26, 13 ° 49 ′ N, 120 ° 50 ′ E, 299 – 320 m; 18 lv + dd, stn CP 68, 14 ° 01 ′ N, 120 ° 18 ′ E, 195 – 199 m; 1 lv, stn CP 80, 13 ° 45 ′ N, 120 ° 37 ′ E, 178 – 205 m. – MUSORSTOM 3: 1 lv?, stn CP 97, 14 ° 00 ′ N, 120 ° 18 ′ E, 189 – 194 m; 5 lv, stn CP 112, 14 ° 00 ′ N, 120 ° 18 ′ E, 187 – 199 m. NEW CALEDONIA: ‘ Vauban ’ 1978 – 79: 1 dd, stn DR 34, 22 ° 32 ′ S, 166 ° 26 ′ E, 350 – 420 m; 1 lv, stn DR 40, 22 ° 30 ′ S, 166 ° 24 ′ E, 250 – 350 m. – LAGON: 1 lv, stn 1062, 20 ° 15 ′ S, 163 ° 53 ′ E, 300 – 320 m. – BATHUS 1: 2 lv, 4 dd, stn CP 656, 21 ° 13 ′ S, 165 ° 54 ′ E, 452 – 460 m; 2 dd, stn CP 657, 21 ° 14 ′ S, 165 ° 55 ′ E, 490 – 530 m; 1 dd, stn CP 669, 20 ° 57 ′ S, 165 ° 35 ′ E, 255 – 280 m; 1 lv, 1 dd, stn CP 670, 20 ° 54 ′ S, 165 ° 53 ′ E, 394 – 397 m; 1 dd, stn DW 683, 20 ° 35 ′ S, 165 ° 07 ′ E, 380 – 400 m; 8 lv, 1 dd, stn CP 695, 20 ° 35 ′ S, 164 ° 58 ′ E, 410 – 430 m; 1 dd, stn CP 701, 20 ° 58 ′ S, 165 ° 36 ′ E, 302 – 335 m; 1 lv, stn DE 705, 21 ° 02 ′ S, 165 ° 38 ′ E, 350 – 400 m; 5 lv, stn CP 707, 21 ° 43 ′ S, 166 ° 36 ′ E, 347 – 375 m; 2 dd, stn CP 710, 21 ° 43 ′ S, 166 ° 36 ′ E, 320 – 386 m; 1 lv, stn CP 711, 21 ° 43 ′ S, 166 ° 36 ′ E, 315 – 327 m; 1 lv, stn CP 712, 21 ° 44 ′ S, 166 ° 35 ′ E, 210 m; 1 lv, stn CP 713, 21 ° 45 ′ S, 166 ° 37 ′ E, 250 m. – BATHUS 4: 3 lv, stn CP 946, 20 ° 34 ′ S, 164 ° 58 ′ E, 386 – 430 m; 3 lv, stn CP 947, 20 ° 34 ′ S, 164 ° 58 ′ E, 470 – 490 m. FIJI: MUSORSTOM 10: 6 dd, stn CP 1320, 17 ° 17 ′ S, 177 ° 54 ′ E, 290 – 300 m; 1 lv, 2 dd, stn CP 1325, 17 ° 16 ′ S, 177 ° 50 ′ E, 282 – 322 m; 1 lv, stn CP 1326, 17 ° 14 ′ S, 177 ° 50 ′ E, 265 – 300 m; 1 lv, stn DW 1347, 17 ° 31 ′ S, 178 ° 40 ′ E, 344 – 349 m; 5 lv, 4 dd, stn CP 1348, 17 ° 30 ′ S, 178 ° 40 ′ E, 353 – 390 m; 1 dd, stn CP 1351, 17 ° 31 ′ S, 178 ° 40 ′ E, 292 – 311 m; 1 lv, stn CP 1355, 17 ° 50 ′ S, 178 ° 49 ′ E, 302 – 310 m; 2 lv, 1 dd, stn CP 1360, 18 ° 00 ′ S, 178 ° 48 ′ E, 402 – 444 m; 3 lv, stn CP 1368, 18 ° 11 ′ S, 178 ° 24 ′ E, 380 – 469 m; 1 dd, stn CP 1369, 18 ° 11 ′ S, 178 ° 23 ′ E, 392 – 433 m. TONGA: BORDAU 2: 3 dd, stn CP 1510, 21 ° 05 ′ S, 175 ° 23 ′ W, 461 – 497 m.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C3DEB61FE28270319C67C92.taxon	description	Description Shell slender, fusiform, large for genus (to 90.0 mm, ANSP 425335). Protoconch multispiral, conical, smooth, translucent pale grey, of 3.25 somewhat convex whorls, last ½ whorl with 4 – 6 rounded axial riblets progressing in strength, protoconch / teleoconch boundary without varix. Protoconch diameter 1100 µm, exposed height 1280 µm (Fig. 5 E). Teleoconch whorls convex, suture deep, adpressed. Axial sculpture of very prominent ribs, 7 – 9 per whorl. Spiral sculpture of evenly spaced strong cords, 2 – 3 on first whorl, 6 – 8 on penultimate whorl, about 23 – 25 on last whorls, becoming evanescent towards tip of canal. Weaker secondary cords between primary strong cord pairs, 1 on earliest whorls increasing to 3 on later whorls. Siphonal canal long, open, straight, about ¼ of shell length. Aperture with 7 – 12 interior lirae. Inner lip adherent, forming smooth, white columellar callus, with 3 rounded columellar plicae, adapical two more prominent than abapical one. In side view additional adapicalmost weak plicae can be observed. Teleoconch pale flesh to pale orange-brown with lighter prominent axial cords. Periostracum medium to dark brown. Radula (MNHN IM- 2007 - 34593, 62.2 mm, Fig. 4 C – D) 2.6 mm in length (13 % of AL), of more than 105 transverse rows of teeth, of which 37 nascent. Radula narrow, distance between outer edges of lateral teeth about 130 µm (0.67 % of AL without canal). Lateral teeth medium broad, arcuate, with attenuated outer anterior corner, producing short ‘ handle’, and with six cusps, subequal in length, closely spaced, slightly curved and one additional small but distinct innermost cusp. Central tooth small, trapezoid with narrower base, with three cusps, central narrowest. Narrow but distinct transverse folds of the membrane situated serially between rows of lateral teeth (arrow). The radula of an additional specimen was examined and it is generally similar, although the left lateral teeth have five cusps, while vs six on the right teeth; also the central teeth are relatively narrower and longer, tricuspid with central cusp longest. Body morphology and digestive system anatomy are in all details similar to Angulofusus nedae Fedosov & Kantor, 2012, the major difference being the longer and thinner proboscis, which attains 20 mm in studied specimen (2.2 AL without canal) and is coiled within rhynchodeum.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C3DEB61FE28270319C67C92.taxon	distribution	Distribution Throughout the tropical Indo-West Pacific, at depths 140 – 809 m, one specimen from commercial sources said to have been collected in 20 – 35 m (Vietnam).	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C3DEB61FE28270319C67C92.taxon	discussion	Remarks Conchologically the species is most similar to Granulifusus annae sp. nov. which has a similar size, but G. jeanpierrevezzaroi comb. nov. differs in having fewer axial ribs per whorl and being nearly uniformly coloured with lighter spiral cords, while in Granulifusus annae sp. nov. the spiral cords are darker than the background. These two species also differ in radular morphology. Granulifusus jeanpierrevezzaroi comb. nov. is a very common species in the Philippines and is found on soft bottoms throughout the tropical Indo-West Pacific. It is remarkable that the species still had to be named, although it had been illustrated several times under incorrect names. Shells of many specimens are covered with zoantharians (e. g., Fig. 5 F – G).	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C30EB66FDD8243B19A97C96.taxon	description	Figs 4 E, 6	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C30EB66FDD8243B19A97C96.taxon	diagnosis	Molecular diagnosis A molecular diagnosis is given in Table 2. Diagnosis Shell thin but solid, slender, fusiform, up to 55 mm, protoconch multispiral of 2.25 whorls. Teleoconch whorls convex, axial sculpture of broad, raised axial ribs, spiral sculpture of evenly spaced, strong cords, 35 on last whorl. Aperture lirate inside, columella with several weak plicae. Teleoconch light tan with darker spiral cords.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C30EB66FDD8243B19A97C96.taxon	etymology	Etymology The specific epithet pays homage to Ann Snyder, in fitting tribute for this elegant and beautiful species.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C30EB66FDD8243B19A97C96.taxon	materials_examined	Type material Holotype PAPUA NEW GUINEA: Rempi Area, 05 ° 03 ′ S, 145 ° 49 ′ E, 120 m, expedition PAPUA NIUGINI, stn PP 08, sequenced (Fig. 6 A – C) (MNHN IM- 2013 - 14709). Sequenced material NEW CALEDONIA: EXBODI: 1 lv, stn CP 3823, 21 ° 55 ′ S, 166 ° 55 ′ E, 246 – 255 m (MNHN IM- 2013 - 42520). PHILIPPINES: AURORA 2007: 3 lv, stn CP 2716, 14 ° 30 ′ N, 121 ° 41 ′ E, 335 – 356 m (MNHN IM- 2007 - 34487, MNHN IM- 2007 - 34585, MNHN IM- 2007 - 34609); 2 lv, stn CP 2717, 14 ° 29 ′ N, 121 ° 43 ′ E, 311 – 361 m (MNHN IM- 2007 - 34486, MNHN IM- 2007 - 34586). Other material examined (all in MNHN) VANUATU: MUSORSTOM 8: 2 dd, stn DW 1003, 18 ° 49 ′ S, 168 ° 59 ′ E, 200 – 327 m; 1 lv, stn CP 1123, 15 ° 07 ′ S, 166 ° 55 ′ E, 262 – 352 m. NEW CALEDONIA: HALIPRO 1: 1 dd, stn CP 853, 21 ° 45 ′ S, 166 ° 37 ′ E, 241 – 250 m; 2 juv. dd, stn CC 855, 21 ° 45 ′ S, 166 ° 37 ′ E, 204 – 220 m; 2 dd, 1 juv. dd, stn CP 866, 21 ° 26 ′ S, 166 ° 17 ′ E, 550 – 600 m. – BATHUS 1: 1 lv (radula examined), stn CP 669, 20 ° 57 ′ S, 165 ° 35 ′ E, 255 – 280 m; 2 dd, stn CP 695, 20 ° 35 ′ S, 164 ° 58 ′ E, 410 – 430 m; 1 dd, stn CP 710, 21 ° 43 ′ S, 166 ° 36 ′ E, 320 – 386 m; 1 dd, stn CP 713, 21 ° 45 ′ S, 166 ° 37 ′ E, 250 m. LOYALTY RIDGE: MUSORSTOM 6: 1 lv (?), stn DW 449, 20 ° 54 ′ S, 167 ° 18 ′ E, 300 m. FIJI: MUSORSTOM 10: 1 dd, stn CP 1320, 17 ° 16.8 ′ S, 177 ° 53.6 ′ E, 290 – 300 m; 1 dd, stn DW 1384, 18 ° 18.5 ′ S, 178 ° 05.8 ′ E, 260 – 305 m. – BORDAU 1: 1 lv (?), 2 dd, stn CP 1402, 16 ° 38 ′ S, 179 ° 36 ′ E, 260 – 279 m; 1 lv, 1 dd, stn CP 1403, 16 ° 40 ′ S, 179 ° 36 ′ E, 220 – 224 m.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C30EB66FDD8243B19A97C96.taxon	description	Description Shell thin, slender, fusiform, consisting of 2.25 protoconch and 8.5 teleoconch whorls (in holotype). Protoconch bluntly conical (Fig. 6 C), with moderately convex whorls, last ½ whorl with 6 axial riblets, protoconch / teleoconch discontinuity distinct. Protoconch diameter 930 µm, exposed height 850 µm. Teleoconch whorls convex, suture deep, impressed. Axial sculpture of broad, raised axial ribs, running from suture to suture, in holotype 9 on first whorl, 8 on 2 nd – 6 th whorls and 9 on last whorl. Spiral sculpture of evenly spaced, strong, raised cords, 3 – 4 on earlier whorls, abapicalmost strongest, 6 on antepenultimate and penultimate whorls, 35 on last whorl. Between major cords, 1 – 3 smaller secondary cords, slightly nodulose at intersection with raised growth lines. Siphonal canal open, long, straight. Aperture with 12 weak lirae inside. Inner lip completely adherent to columella, forming very thin, shiny columellar callus with 5 weak columellar plicae. No umbilicus. Protoconch light brown, interior of aperture white. Teleoconch light-tan with still lighter spiral band on shell periphery, spiral cords tan, usually darker than background, upper spiral whorls brownish adapically. Operculum small, length in holotype 5.3 mm, occupying slightly more than ¼ of aperture without canal, oval, nucleus supposingly terminal, older part of operculum missing. Holotype measurements: SL 50.3 mm, AL (with canal) 30.0 mm, SW 15.9 mm. Radula (Fig. 4 E, New Caledonia, BATHUS 1, stn CP 669, 20 ° 57 ′ S, 165 ° 35 ′ E, 255 – 280 m, shell on Fig. 6 F) narrow, distance between outer edges of lateral teeth about 60 µm (0.48 % of AL without canal). Lateral teeth medium broad, arcuate, with attenuated outer anterior corner, producing medium long ‘ handle’. Lateral teeth with outermost very short small cusp (Fig. 4 E, black arrows), four broadly spaced and slightly recurved, long cusps, subequal in length. Inner anterior corner also attenuated, forming a triangular outgrowth, sometimes bearing indistinct an additional innermost small cusp. Central tooth small, very narrow (length to width ratio 5), with three short and blunt cusps, central longest. Narrow but distinct transverse folds of membrane situated serially between rows of lateral teeth, on both sides of central teeth (white arrow). The radula of the holotype was examined and illustrated by Couto (2016: fig. 28) under the name Pseudolatirus kuroseanus. It is in all details similar to the radula figured here, the minor differences are that the outermost small cusp is more distinct, the innermost outgrowth bears a distinct small cusp, and the cusps of the central teeth are sharp rather than blunt.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C30EB66FDD8243B19A97C96.taxon	distribution	Distribution Philippines, New Caledonia, Papua New Guinea, Vanuatu, Fiji, 120 – 600 m, alive in 120 – 360 m.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C30EB66FDD8243B19A97C96.taxon	discussion	Remarks This species is moderately variable both in slenderness of the shell and in whorl convexity. Rarely the spiral cords are lighter than the shell background (observed in sequenced juvenile specimen MNHN IM- 2007 - 34609). This new species is similar to G. kurodai comb. nov. and has been confused with it, but can be distinguished by a different protoconch, consisting of 2.25 whorls in G. annae sp. nov. rather than only 1.75 whorls in G. kurodai comb. nov., as well as by a straight siphonal canal, darker shell coloration, and more pronounced and less numerous primary spiral cords (6 on penultimate whorl in G. annae sp. nov. and 10 in paratype of G. kurodai comb. nov.). The species is also conchologically similar to G. jeanpierrevezzaroi comb. nov., differing in the more brownish background shell colour with more contrastingly coloured spiral cords, more dense axial ribs and a smaller protoconch. The molecular analysis clearly differentiates these species. The radula of the new species is somewhat different from its congeners in having longer and more broadly spaced cusps on the lateral teeth.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C37EB64FDFD242B192D7F12.taxon	description	Fig. 7 A – D	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C37EB64FDFD242B192D7F12.taxon	diagnosis	Diagnosis Shell thin, slender, fusiform, up to 14 mm, protoconch of 4 whorls. Teleoconch whorls weakly convex, axial sculpture of narrow, broadly spaced, axial ribs, spiral sculpture of evenly spaced, strong primary cords, 7 on last whorl and weaker secondary cords between them. Aperture weakly lirate inside, columella with several distinct columellar plicae. Teleoconch light tan with several brown spiral bands present, also on siphonal canal.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C37EB64FDFD242B192D7F12.taxon	etymology	Etymology The specific epithet refers to the collecting locality, the Norfolk Ridge, where the first known specimen of this species was collected.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C37EB64FDFD242B192D7F12.taxon	materials_examined	Type material Holotype NEW CALEDONIA: north of Ile des Pins, 22 ° 28 ′ S, 167 ° 29 ′ E, 404 – 405 m, expedition KANACONO, stn DW 4685, sequenced (MNHN IM- 2013 - 68811). Other material examined NEW CALEDONIA: 1 lv, TERRASSES, stn DW 3063, Norfolk Ridge, 23 ° 23 ′ S, 168 ° 00 ′ E, 430 – 480 m, sequenced (MNHN IM- 2007 - 36921).	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C37EB64FDFD242B192D7F12.taxon	description	Description Shell thin, slender, fusiform, of nearly 4 protoconch and 5 teleoconch whorls (in holotype). Protoconch acutely conical (Fig. 7 C), with moderately convex whorls, last ½ whorl with 6 axial, progressively thickening, riblets, protoconch / teleoconch transition abrupt, marked by onset of distinctive sculpture. Protoconch diameter 1130 µm, exposed height 1150 µm. Teleoconch whorls weakly convex, suture deeply impressed. Axial sculpture of broad, prominent ribs extending from suture to suture (7 per whorl in holotype), including on last and penultimate whorls. Spiral sculpture of evenly spaced, strong, raised cords, nodulous at crossings with axial ribs, in holotype 2 on first whorl, 3 on other whorls, 7 on last whorl before transition to canal. Three or four weaker secondary cords situated between primary cords, slightly nodulous at intersections with raised growth lines. Two brown secondary cords situated in each interspace between primary cords on last whorl. Siphonal canal open, long, slightly turned to left and recurved, sculptured by 25 (in holotype) weaker cords of slightly uneven strength. Aperture with 8 weak lirae inside. Inner lip completely adherent to columella, forming very thin, shiny columellar callus with 4 columellar plicae, first very weak. No umbilicus. Protoconch transparent flesh colour, interior of aperture white. Teleoconch light tan with several brown spiral bands also present on canal. Major spiral cords slightly lighter than background, secondary spiral cords mostly brown. Holotype measurements: SL 13.0 mm, AL (without canal) 3.5 mm, SW 4.7 mm.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C37EB64FDFD242B192D7F12.taxon	distribution	Distribution Known only from southern New Caledonia, off Ile des Pins and Norfolk Ridge, in 404 – 480 m. The multispiral protoconch suggests a planktonic dispersal phase and a possibly broader distribution.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C37EB64FDFD242B192D7F12.taxon	discussion	Remarks The species is known from just two specimens, the holotype and a slightly larger specimen attaining SL 13.8 mm (Fig. 7 D). Granulifusus norfolkensis sp. nov. can be recognized by its small size, coloration and constricted aperture. Conchologically it is most similar to Granulifusus tatianae sp. nov., also from southern New Caledonia, but differs in having a smaller and more slender shell with longer siphonal canal and brown spiral bands.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C35EB5BFDD324BB1A767B8B.taxon	description	Figs 4 F, 7 E – J	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C35EB5BFDD324BB1A767B8B.taxon	diagnosis	Diagnosis Shell strong, broadly fusiform, up to 18 mm, protoconch of 3.5 whorls. Teleoconch whorls strongly convex, axial sculpture of broad opisthocline axial ribs, spiral sculpture of strong primary cords and numerous finer secondary cords between them. Aperture lirate inside, columella with several distinct columellar plicae. Teleoconch light tan, axial ribs brown.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C35EB5BFDD324BB1A767B8B.taxon	etymology	Etymology The specific epithet is named in honour of Tatiana Steyker, wife of Yuri Kantor, patient permanent companion in all travels and many fieldtrips.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C35EB5BFDD324BB1A767B8B.taxon	materials_examined	Type material Holotype NORFOLK RIDGE: south of New Caledonia, 23 ° 17 ′ S, 168 ° 14 ′ E, 190 – 212 m, expedition NORFOLK 1, stn DW 1727 (MNHN IM- 2000 - 33595). Paratype NORFOLK RIDGE: 1 specimen, same station as for holotype (MNHN IM- 2000 - 33596); 1 specimen (ANSP, ex Snyder collection). Other material examined (all in MNHN) NEW CALEDONIA: BIOCAL: 1 dd, stn CP 84, 20 ° 43 ′ S, 167 ° 01 ′ E, 150 – 210 m. – SMIB 3: 1 dd, stn DW 08, 24 ° 45 ′ S, 168 ° 08 ′ E, 233 m. – SMIB 8: 1 dd, stn DW 157, 24 ° 46 ′ S, 168 ° 08 ′ E, 251 – 257 m; CHALCAL 2: 3 dd, stn DW 84, 23 ° 24 ′ S, 168 ° 07 ′ E, 170 m. – NORFOLK 1: 1 lv, 2 dd; stn DW 1724, 23 ° 17 ′ S, 168 ° 14 ′ E, 200 – 291 m; 1 lv, 1 dd, stn DW 1727, 23 ° 17 ′ S, 168 ° 14 ′ E, 190 – 212 m. LOYALTY RIDGE: MUSORSTOM 6: 1 dd, stn DW 442, 20 ° 54 ′ S, 167 ° 17 ′ E, 200 m; 1 lv, 1 dd, stn DW 473, 21 ° 09 ′ S, 167 ° 55 ′ E, 236 m. – LIFOU 2000: 2 dd, stn DW 1650, 20 ° 54.2 ′ S, 167 ° 01.7 ′ E, 120 – 250 m. WALLIS AND FUTUNA: MUSORSTOM 7: 1 dd, stn CP 517, 14 ° 13 ′ S, 178 ° 10 ′ W, 233 – 235 m.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C35EB5BFDD324BB1A767B8B.taxon	description	Description Shell fusiform, solid, of 3.5 protoconch and 7 weakly convex teleoconch whorls in holotype. Protoconch bluntly conical (Fig. 7 G), last ¾ whorl with axial riblets, protoconch / teleoconch discontinuity distinct, marked by appearance of teleoconch sculpture. Protoconch smooth except scattered microgranules, more dense along suture, where they are arranged in oblique lines. Protoconch diameter 820 – 910 µm (latter in holotype), exposed height 860 – 920 µm (former in holotype). Teleoconch whorls convex with impressed suture. Axial sculpture of broad, prominent axial ribs, aligned axially on spire from one whorl to the next, 7 ribs per whorl, and of very weak incremental lines. Spiral sculpture of evenly spaced, strong, raised primary cords, in holotype 3 on spire whorls, 13 on last whorl, and numerous, much weaker secondary threads, about 5 – 7 between two adjacent primary cords. Intersection of secondary spiral threads and incremental lines forming finely reticulate microsculpture. Siphonal canal open, short, recurved. Aperture with 8 strong lirae inside, adapicalmost stronger, forming spirally elongated tooth constricting base of siphonal canal. Inner lip partially detached, with three prominent columellar plicae and one strong parietal tooth, forming well-defined posterior canal. No umbilicus. Colour of protoconch and interior of aperture white. Teleoconch background colour creamy white, axial ribs light brown, spiral cords lighter than ribs. Holotype measurements: SL 14.3 mm, AL (without canal) 4.5 mm, SW 6.1 mm. Radula (MUSORSTOM 6, stn DW 473, 21 ° 09 ′ S, 167 ° 55 ′ E, 236 m, Fig. 4 F) narrow, distance between outer edges of lateral teeth about 55 µm, with slender central tooth, about twice as long as broad, bearing three strong cusps. Lateral tooth medium broad, with long basal ‘ handle’ on outer edge, with five strong, subequal cusps, the outermost more clearly separated from the rest, and one smaller inner cusp. Narrow but distinct transverse folds of the membrane situated serially between rows of lateral teeth.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C35EB5BFDD324BB1A767B8B.taxon	distribution	Distribution Southwest Pacific, known only from Futuna and the New Caledonia region, alive in 200 – 236 m, empty shells in 170 – 251 m.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C35EB5BFDD324BB1A767B8B.taxon	discussion	Remarks Granulifusus tatianae sp. nov. is superficially similar to Okutanius kuroseanus gen. et comb. nov. Both species have brown axial ribs aligned along the spire against a lighter, creamy background, and both have a multispiral protoconch indicating planktotrophic larval development. Okutanius kuroseanus gen. et comb. nov., however, reaches a much larger adult size and is proportionally broader at comparable size; it has a longer siphonal canal, weaker columellar plicae, and spiral cords of uneven strength, without the very regular pattern of evenly spaced primary cords separated by 5 – 7 very fine, even, spiral threads of G. tatianae sp. nov. The species also resembles G. norfolkensis sp. nov. in sculpture and particularly in apertural armature. Nevertheless, the shell shape in Granulifusus tatianae sp. nov. is rather different, and the species can be easily distinguished by size at maturity: G. tatianae sp. nov. attains a significantly larger size, up to 18.1 mm, versus 13.6 mm in G. norfolkensis sp. nov. The new species was not examined molecularly and we tentatively assign it to Granulifusus based on the similarities of shell sculpture and apertural ornamentation to G. norfolkensis sp. nov. and G. williami comb. nov.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C0AEB5AFE0621221ABC7B44.taxon	description	Fig. 8 A – B	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C0AEB5AFE0621221ABC7B44.taxon	materials_examined	Type material Holotype JAPAN: off Gotō Islands, Nagasaki Prefecture, Western Kyūshū (NSMT Mo 38547), type locality corrected (Okutani & Sakurai 1966) [not seen]. Paratypes JAPAN: paratype 1, Hyotanse Bank (NSMT-Mo 58524) (examined); paratype 2, off Gotō Islands (whereabouts unknown) (not seen); paratype 3, Kashiwajima (NSMT-Mo 70270) (examined).	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C0AEB5AFE0621221ABC7B44.taxon	description	Description Shell medium-sized, fusiform, with long, slightly sinuous siphonal canal. Protoconch paucispiral, domeshaped (Fig. 8 B), of 1.75 whorls, with strongly convex first whorl and much flatter last ¾ of whorl, with two closely spaced and rather indistinct axial riblets at transition to teleoconch, protoconch / teleoconch discontinuity distinct. Protoconch diameter 900 µm, exposed height 750 µm. Teleoconch with regularly convex whorls, aperture with two very weak columellar plicae, and indistinct parietal tooth; spiral sculpture of unevenly spaced narrow cords of similar strength; axial ribs not aligned along the spire. Background colour cream to yellowish, primary spiral cords light tan.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C0AEB5AFE0621221ABC7B44.taxon	distribution	Distribution Known from Japan, in 100 – 200 m, records from other localities may refer to G. annae sp. nov. and G. guidoi sp. nov.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C0AEB5AFE0621221ABC7B44.taxon	discussion	Remarks We were able to study two paratypes NSMT-Mo 58524 (paratype 1) and NSMT-Mo 70270 (paratype 3). They are clearly not conspecific, paratype 3 differing in its much larger protoconch and general shell shape and sculpture; it represents a juvenile Fusolatirus, resembling F. luteus Snyder & Bouchet, 2006 (see Snyder & Bouchet 2006: fig. 2 D – E). Conchologically, paratype 1 is very similar to the holotype and is the largest known specimen (SL 35.1 mm). Conchologically Granulifusus kurodai comb. nov. is similar to G. annae sp. nov.; for differences see the description of the latter. In the absence of data on radula and molecular phylogeny, the assignment of this species to Granulifusus is tentative, based on its shell resemblance to sequenced species such as G. annae sp. nov.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C0BEB58FDD9206A1F8D7B24.taxon	description	Fig. 8 E – H	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C0BEB58FDD9206A1F8D7B24.taxon	diagnosis	Diagnosis Shell strong, slender, fusiform, height up to 31 mm, protoconch of 2.5 whorls. Teleoconch whorls strongly convex, axial sculpture of medium broad opisthocline axial ribs, spiral sculpture of thin, distinct, primary cords, varying slightly in size. Aperture lirate inside, columella with distinct to weak columellar plicae. Teleoconch light tan with light orange-brown canal and caramel-colored spiral cords.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C0BEB58FDD9206A1F8D7B24.taxon	etymology	Etymology Named for Guido Poppe, the author of the landmark Philippine Marine Molluscs and discoverer of many new molluscs in the Philippines and elsewhere, who generously provided some of the type material.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C0BEB58FDD9206A1F8D7B24.taxon	materials_examined	Type material Holotype PHILIPPINES: off Aliguay Island, northeast of Mindanao, in 50 – 150 m (MNHN IM- 2000 - 33597, ex Snyder collection). Paratypes PHILIPPINES: 1 specimen, same data as for holotype, SL 30.7 mm (MNHN- 2000 - 33598); 1 specimen, trawled off Aliguay Island, 80 – 150 m, SL 30.7 mm (ANSP 456853); 1 specimen, trawled off Aliguay Island, 100 – 150 m, 25.5 mm (NMP).	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C0BEB58FDD9206A1F8D7B24.taxon	description	Description Shell slender, fusiform, of average size for genus. Protoconch multispiral, of 2.5 smooth whorls in holotype, with 4 axial riblets on last whorl, pale grey-orange. Transition to teleoconch marked by groove between last riblet and first teleoconch axial rib. Teleoconch of 8 – 9 convex whorls, suture deep, adpressed. Axial sculpture of distinct ribs, closely spaced, 10 on earlier whorls, increasing to 12 on later whorls. Spiral sculpture of dark cords, often with less prominent and paler secondary cords between them. First whorl with single dark cord, adding roughly one cord per whorl, with 7 cords on penultimate whorl. Cords continue to anterior end of canal, crowding closer together at tip; in holotype, last whorl with about 25 cords. Siphonal canal recurved back from aperture, moderately long, open. Dark spiral cords matched on inner lip by low, light brown lirae. Columellar callus white, with a few spiral lines continuing inward at adapical end of shield. Two stronger rounded columellar plicae situated towards aperture and one much weaker towards canal. Teleoconch white to light tan with light orange-brown neck; caramel-colored spiral cords on entire shell. Holotype measurements: SL 30.2 mm, AL (without canal) 8.1 mm, SW 10.1 mm.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C0BEB58FDD9206A1F8D7B24.taxon	distribution	Distribution Known only from the type locality.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C0BEB58FDD9206A1F8D7B24.taxon	discussion	Remarks Granulifusus guidoi sp. nov. is most similar to G. kurodai comb. nov., with which it is often confused. It differs by a multispiral protoconch with more whorls, and a narrower shell with narrower aperture. It is also similar to Okutanius ellenae gen. et sp. nov., which has white cords with dark interspaces on the teleoconch whereas G. guidoi sp. nov. has dark, caramel-colored cords with light tan interspaces. In the absence of data on radula and molecular phylogeny, we tentatively assign this species to Granulifusus based on shell similarities to such sequenced species as G. annae sp. nov.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C09EB5FFD1620891AD879DA.taxon	type_taxon	Type species Fusolatirus kuroseanus Okutani, 1975, here designated.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C09EB5FFD1620891AD879DA.taxon	diagnosis	Diagnosis Shell medium-size, reaching 30 mm, fusiform to broad fusiform, strong, with spiral colour bands. Teleoconch whorls evenly strongly convex, shell base sharply narrowing towards long, weakly sinuous siphonal canal. Protoconch multispiral, of 3 – 3.75 whorls. Axial sculpture of distinct broad, weakly opisthocline, closely spaced ribs. Spiral sculpture of distinct primary cords, subequal in size, and much thinner cordlets between them. Aperture narrow, oval, relatively small, columella with several weak plicae. No distinct parietal knob. Operculum spans entire aperture, with terminal nucleus. Lateral teeth with five closely spaced, slightly curved cusps and much smaller, but distinct, innermost cusp. Outermost very short small cusp present, but usually overlaid by large next cusp and not seen in dorsal view. Central tooth small, narrow, trapezoid, with three cusps. Narrow but distinct transverse folds of membrane situated serially between rows of lateral teeth.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C09EB5FFD1620891AD879DA.taxon	etymology	Etymology The genus is named after Takashi Okutani in recognition of his lifelong contribution to malacology, including the description of the type species of the genus. The gender is masculine.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C09EB5FFD1620891AD879DA.taxon	diagnosis	Differential diagnosis Okutanius gen. nov. is very similar to Granulifusus and conchological and morphological characters overlap at least in some species. Species of Okutanius gen. nov. do not reach as large size as in Granulifusus. Nevertheless, without molecular data, it would be difficult to suggest the existence of two clearly separate genera. Genus composition In addition to two sequenced species, O. kuroseanus and O. ellenae sp. nov., we allocate Pseudolatirus aikeni Lussi, 2014, known from empty shell only, to Okutanius gen. nov. based on shell similarity to sequenced O. kuroseanus.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C0EEB5CFE7722F5185E78B2.taxon	description	Figs 4 G, 9 A – G	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C0EEB5CFE7722F5185E78B2.taxon	materials_examined	Type material JAPAN: holotype of of F. kuroseanus, Kurose Bank, off Izu islands, south of Honshū, 33 ° 23.5 ′ N, 139 ° 36.2 ′ E, 260 m (NSMT Mo 69575) (illustrated by Higo et al. 2001: 87); holotype of P. leucostriatus (said to be off Midway Island, 32 ° 02 ′ N, 173 ° 06 ′ W, 365 m, but label of holotype says 29 ° 46.4 ′ N, 179 ° 02.9 ′ E, 265 m) in the Institute of Malacology, Tokyo (IMT 79 - 18) [examined]; paratype, same location as holotype, said to be donated to USNM. Sequenced material PAPUA NEW GUINEA, NEW IRELAND: KAVIENG 2014: 1 lv, stn CP 4503, 02 ° 30 ′ S, 150 ° 44 ′ E, 191 – 200 m (MNHN IM- 2013 - 59070). Other material examined TAIWAN: TAIWAN 2000: 2 dd, stn DW 05, 23 ° 41 ′ N, 119 ° 56 ′ E, 213 – 236 m; 2 dd, stn DW 34, 22 ° 01 ′ N, 120 ° 36 ′ E, 240 – 246 m; 25 dd, stn DW 36, 21 ° 54 ′ N, 120 ° 36 ′ E, 300 – 331 m. PHILIPPINES: 2 spms, Panglao, Bohol, tangle nets, 160 m, (ANSP 425326); 2 spms, off Aliguay I., Mindanao, 150 m (ANSP 425322); 1 spm, Bohol, Balicasag, 240 m (ANSP 425324); 1 spm, Philippine Is, tangle nets (ANSP 425325); 1 spm, Balicasag I., Bohol, tangle nets, 240 m (ANSP 425327); 1 spm, Siquijor I., tangle nets, 150 m (ANSP 425323). ARAFURA SEA: KARUBAR: 1 lv, 13 dd, stn DW 15, 05 ° 17 ′ S, 132 ° 41 ′ E, 212 – 221 m. SOLOMON ISLANDS: SALOMON 1: 1 dd, stn DW 1765, 08 ° 43 ′ S, 160 ° 07 ′ E, 325 – 380 m; 1 dd, stn DW 1812, 09 ° 46 ′ S, 160 ° 52 ′ E, 222 – 246 m. NEW CALEDONIA: “ Vauban ” 1978 – 79: 1 dd, stn DR 37, 22 ° 32 ′ S, 166 ° 26 ′ E, 175 – 250 m. – BIOCAL: 1 dd, stn DW 81, 20 ° 29 ′ S, 166 ° 47 ′ E, 430 – 470 m; 1 juv. Lv, stn CP 105, 21 ° 31 ′ S, 166 ° 22 ′ E, 330 – 335 m. – BIOGEOCAL: 3 dd, stn DW 253, 21 ° 32 ′ S, 166 ° 29 ′ E, 310 – 315 m. – MUSORSTOM 4: 1 dd, stn DW 181, 18 ° 57 ′ S, 163 ° 22 ′ E, 350 m. – BATHUS 1: 1 dd, stn DW 655, 21 ° 17 ′ S, 165 ° 57 ′ E, 375 m; 4 dd, stn DW 688, 20 ° 33 ′ S, 165 ° 00 ′ E, 270 – 282 m; 2 dd, stn DW 689, 20 ° 33 ′ S, 165 ° 00 ′ E, 260 – 265 m. – BATHUS 4: 1 dd, stn DW 886, 21 ° 01 ′ S, 164 ° 28 ′ E, 250 – 300 m; 1 dd, stn DW 887, 21 ° 07 ′ S, 164 ° 28 ′ E, 320 – 344 m. – SMIB 4: 1 dd, stn DW 44, 24 ° 46 ′ S, 168 ° 08 ′ E, 270 – 300 m. – SMIB 5: 1 dd, stn DW 90, 22 ° 19 ′ S, 168 ° 42 ′ E, 340 m. – SMIB 8: 1 dd, stn DW 160, 24 ° 47 ′ S, 168 ° 08 ′ E, 280 – 282 m. LOYALTY RIDGE: MUSORSTOM 6: 2 lv, 3 dd, stn DW 391, 20 ° 47 ′ S, 167 ° 06 ′ E, 390 m; 1 dd, stn DW 397, 20 ° 47 ′ S, 167 ° 05 ′ E, 380 m; 1 lv, 1 dd, stn DW 398, 20 ° 47 ′ S, 167 ° 06 ′ E, 370 m; 1 dd, stn DW 418, 20 ° 42 ′ S, 167 ° 03 ′ E, 283 m; 3 dd, stn DW 459, 21 ° 01 ′ S, 167 ° 31 ′ E, 425 m; 3 dd, stn DW 480, 21 ° 08 ′ S, 167 ° 56 ′ E, 380 m; 1 dd, stn DW 481, 21 ° 22 ′ S, 167 ° 50 ′ E, 300 m; 1 dd, stn DW 482, 21 ° 21 ′ S, 167 ° 47 ′ E, 375 m; 2 dd, stn DW 487, 21 ° 23 ′ S, 167 ° 46 ′ E, 500 m. VANUATU: MUSORSTOM 8: 1 dd, stn DW 1059, 16 ° 13 ′ S, 167 ° 21 ′ E, 408 – 430 m; 1 dd, stn DW 1065, 16 ° 16 ′ S, 167 ° 21 ′ E, 360 – 419 m. WALLIS AND FUTUNA: MUSORSTOM 7: 1 dd, stn DW 496, 14 ° 20 ′ S, 178 ° 04 ′ W, 250 – 330 m (Futuna I.); 1 dd, stn DW 509, 14 ° 15 ′ S, 178 ° 11 ′ W, 200 – 240 m (Futuna I.); 1 dd, stn DW 523, 13 ° 12 ′ S, 176 ° 16 ′ W, 455 – 515 m (Wallis I.); 1 dd, stn DW 569, 12 ° 30 ′ S, 176 ° 51 ′ W, 300 – 305 m (Waterwitch Bank); 1 dd, stn DW 571, 12 ° 31 ′ S, 176 ° 52 ′ W, 502 – 508 m (Waterwitch Bank); 1 dd, stn DW 610, 13 ° 21 ′ S, 176 ° 09 ′ W, 286 m (Wallis I.); 1 dd, stn DW 626, 11 ° 54 ′ S, 179 ° 32 ′ W, 597 – 600 m (Bayonnaise Bank). FIJI: BORDAU 1: 1 lv, 2 dd, stn DW 1469, 19 ° 40 ′ S, 178 ° 10 ′ W, 314 – 377 m; 1 lv; stn DW 1497, 18 ° 44 ′ S, 178 ° 25 ′ W, 335 – 350 m. TONGA: BORDAU 2: 2 dd, stn DW 1535, 21 ° 43 ′ S, 175 ° 18 ′ W, 268 m; 3 dd, stn DW 1567, 21 ° 02 ′ S, 175 ° 19 ′ W, 351 – 356 m; 1 dd, stn DW 1583, 18 ° 37 ′ S, 174 ° 03 ′ W, 327 – 360 m; 1 lv (?), 1 dd, stn DW 1587, 18 ° 37 ′ S, 173 ° 54 ′ W, 309 – 400 m; 1 dd, stn DW 1589, 18 ° 39 ′ S, 173 ° 54 ′ W, 281 m; 1 lv (?), stn DW 1595, 19 ° 03 ′ S, 174 ° 19 ′ W, 523 – 806 m; 1 dd, stn DW 1602, 20 ° 49 ′ S, 174 ° 57 ′ W, 263 – 320 m; 1 dd, stn DW 1605, 22 ° 17 ′ S, 175 ° 16 ′ W, 441 m; 1 lv, stn DW 1607, 22 ° 15 ′ S, 175 ° 23 ′ W, 356 – 367 m; 1 lv (?), stn DW 1630, 23 ° 23 ′ S, 176 ° 18 ′ W, 360 m; 1 lv (?), stn DW 1636, 21 ° 44 ′ S, 175 ° 20 ′ W, 321 – 331 m. CHRISTMAS ISLAND: 2 dd, off NE point, 10 ° 25 ′ S, 105 ° 42 ′ E, 183 m, coll. R. Kirkpatrick 1916 (AMS C 324273). RÉUNION: MD 32: 1 dd, stn CP 129, 20 ° 51 ′ S, 55 ° 36 ′ E, 290 – 300 m; 1 dd worn, stn DC 136, 20 ° 46 ′ S, 55 ° 36 ′ E, 915 – 922 m. SOUTH AFRICA: 1 dd, former Transkei, off Port Grosvenor, 29 ° 57.4 ′ S, 31 ° 26.4 ′ E, 105 m (NM C 636); 1 lv (?), former Transkei, off Mtamvuna River, 31 ° 08.4 ′ S, 30 ° 16.6 ′ E, 160 m (NM C 9652); 2 dd, Zululand, SE of Port Durnford, 29 ° 05.8 ′ S, 32 ° 09.4 ′ E, 165 m (NM S 903); 1 dd, Zululand, SE of Port Durnford, 29 ° 04.6 ′ S, 32 ° 09.0 ′ E, 142 m (NM D 7822).	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C0EEB5CFE7722F5185E78B2.taxon	description	Description Shell fusiform, up to 28 mm in length. Protoconch acutely conical, multispiral, of 3 smooth, weakly convex whorls, with several slightly arcuate axial riblets on last ½ whorl (Fig. 9 C). Protoconch diameter 870 µm, exposed height 1000 µm. Teleoconch with regularly convex whorls, without imbricated subsutural spiral ridge, with long siphonal canal; aperture with weak columellar plication, and indistinct parietal tooth; spiral sculpture of unevenly arranged cords of uneven strength; axial ribs aligned along the spire. Background colour from creamy light to yellowish brown, axial ribs brown, primary spiral cords white. Radula (Fig. 4 G) examined in two specimens, from Fiji (BORDAU 1, stn DW 1497) and from the New Ireland (MNHN IM- 2013 - 59070, 18.1 mm) (shell: Fig. 9 A). They are very similar and that from the New Ireland specimen is illustrated and described. Radula narrow, distance between outer edges of lateral teeth about 70 µm (1.34 % of AL). Lateral teeth broad, slightly arcuate, with strongly attenuated outer anterior corner, producing long ‘ handle’. Lateral teeth with outermost small cusp, on most rows overlaid by following major cusp and not seen on ventral view (marked by white arrow), five almost equal, closely spaced, cusps, outer one more clearly separated from the rest, and one smaller inner cusp. Central tooth trapezoid, with three short blunt cusps, central slightly longer. Distinct transverse folds of the membrane situated serially between rows of lateral teeth, reaching anterior margin of central tooth.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C0EEB5CFE7722F5185E78B2.taxon	distribution	Distribution Southwest Pacific Ocean, Indian Ocean to KwaZulu-Natal, in 150 – 620 m, alive in 150 – 400 m.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C0EEB5CFE7722F5185E78B2.taxon	discussion	Remarks This distinct, medium-sized species is similar to Granulifusus tatianae sp. nov.; for differences, see the description of the latter. It also resembles Okutanius ellenae gen. et sp. nov., but can be easily differentiated from the latter species by the colour pattern.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C0DEB50FDE7201B1A067E2C.taxon	description	Figs 4 H, 10	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C0DEB50FDE7201B1A067E2C.taxon	diagnosis	Diagnosis Shell strong, slender, narrowly fusiform, up to 30 mm, protoconch of 3.75 whorls. Teleoconch whorls strongly convex, axial sculpture of broad, nearly orthocline, axial ribs, spiral sculpture of dense, evenly spaced, strong, raised cords. Aperture lirate inside, columella with weak columellar plicae to nearly smooth. Teleoconch with alternating white spiral cords and orange brown interspaces, and three white bands, one subsutural, one just below periphery of last whorl and one at tip of siphonal canal. Molecular diagnosis See Table 3.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C0DEB50FDE7201B1A067E2C.taxon	etymology	Etymology The specific epithet pays homage to Ellen Strong, in fitting tribute for this elegant and handsome species.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C0DEB50FDE7201B1A067E2C.taxon	materials_examined	Type material Holotype NEW CALEDONIA: South of Ile des Pins, 22 ° 53 ′ S, 167 ° 35 ′ E, 376 – 390 m, expedition KANACONO, stn DW 4677, sequenced (MNHN IM- 2013 - 68819). Paratypes NEW CALEDONIA: 1 sequenced specimen from the type locality (MNHN IM- 2013 - 68876); 2 specimens, 24 ° 44 ′ S, 168 ° 10 ′ E, 270 – 350 m, expedition BERYX 11, stn DW 11 (AMS C. 572407 and NSMT-Mo 79000); 1 specimen, 22 ° 58 ′ S, 167 ° 33 ′ E, 410 - 440 m, expedition MUSORSTOM 4, stn DW 222 (ANSP 476089). Sequenced material NEW CALEDONIA: holotype and paratype, TERRASSES: 1 lv, stn DW 3082, 22 ° 29 ′ S; 167 ° 23 ′ E, 290 m (MNHN IM- 2007 - 38356). CORAL SEA: EBISCO: 1 lv, stn DW 2520, Banc Kelso, 24 ° 06 ′ S, 159 ° 41 ′ E, 350 – 400 m (MNHN IM- 2007 - 32522). Other material examined CORAL SEA: MUSORSTOM 5: 2 dd, stn DW 299, 22 ° 48 ′ S, 159 ° 24 ′ E, 360 – 390 m; 5 lv, 10 dd, stn DW 300, 22 ° 48 ′ S, 159 ° 24 ′ E, 450 m; 18 dd, stn DW 301, 22 ° 07 ′ S, 159 ° 25 ′ E, 487 – 610 m; 1 dd, stn DW 304, 22 ° 10 ′ S, 159 ° 26 ′ E, 385 – 420 m; 2 dd, stn DW 306, 22 ° 08 ′ S, 159 ° 21 ′ E, 375 – 415 m; 1 lv, 3 dd, stn DC 362, 19 ° 53 ′ S, 158 ° 40 ′ E, 410 m. NEW CALEDONIA: LAGON: 1 lv, stn 444, 18 ° 15 ′ S, 162 ° 59 ′ E, 300 – 350 m. – BIOCAL: 1 dd, stn DW 38, 23 ° 00 ′ S, 167 ° 15 ′ E, 360 m; 8 lv + dd, stn DW 64, 24 ° 48 ′ S, 168 ° 09 ′ E, 250 m; 1 lv, 3 dd stn DW 65, 24 ° 48 ′ S, 168 ° 09 ′ E, 245 – 275 m. – BIOGEOCAL: 2 dd, stn KG 252, 21 ° 31 ′ S, 166 ° 21 ′ E, 330 m; 5 dd, stn DW 253, 21 ° 32 ′ S, 166 ° 29 ′ E, 310 – 315 m. – SMIB 5: 1 lv, stn DW 98, 23 ° 02 ′ S, 168 ° 16 ′ E, 335 m. – 1 dd, S. of Ile des Pins, 22 ° 52 ′ S, 167 ° 35.5 ′ E, 370 m (AMS C 324274). – BATHUS 2: 1 dd, stn DW 731, 22 ° 49 ′ S, 166 ° 45 ′ E, 300 – 370 m. – BATHUS 3: 1 dd, stn CP 804, 23 ° 41 ′ S, 168 ° 00 ′ E, 244 – 278 m. – MUSORSTOM 4: 1 lv, stn DW 222, 22 ° 58 ′ S, 167 ° 33 ′ E, 410 - 440 m (paratype ANSP); 2 dd, stn DW 234, 22 ° 15 ′ S, 167 ° 08 ′ E, 350 – 365 m. – CHALCAL 2: 14 dd, stn DW 69, 24 ° 44 ′ S, 168 ° 08 ′ E, 260 m; 1 lv, 3 dd, stn DW 70, 24 ° 46 ′ S, 168 ° 09 ′ E, 232 m; 2 dd, stn DW 71, 24 ° 42 ′ S, 168 ° 10 ′ E, 230 m; 1 dd, stn DW 78, 23 ° 41 ′ S, 168 ° 00 ′ E, 233 m; 3 dd, stn DW 79, 23 ° 41 ′ S, 168 ° 00 ′ E, 243 m, 3 dd. – BERYX 11: 30 lv + dd, stn DW 11, 24 ° 44 ′ S, 168 ° 10 ′ E, 270 – 350 m; 5 spms, stn DW 18, 24 ° 48 ′ S, 168 ° 09 ′ E, 250 – 270 m. – HALICAL 1: 1 dd, stn 1 – 4, 18 ° 53 ′ – 18 ° 56 ′ S, 163 ° 24 ′ E, 350 – 400 m. – LITHIST: 1 dd, stn CP 10, 24 ° 48 ′ S, 168 ° 09 ′ E, 245 – 261 m; 3 dd, stn DW 11, 24 ° 47 ′ S, 168 ° 08 ′ E, 254 – 283 m; 4 dd, stn DW 12, 24 ° 46 ′ S, 168 ° 08 ′ E, 235 – 280 m; 1 dd, stn DW 13, 23 ° 45 ′ S, 168 ° 17 ′ E, 400 m. – NORFOLK 1: 15 lv + dd, stn DW 1651, 23 ° 27 ′ S, 167 ° 50 ′ E, 276 – 350 m; 1 dd, stn DW 1652, 23 ° 26 ′ S, 167 ° 50 ′ E, 290 – 378 m; 3 dd, stn DW 1653, 23 ° 28 ′ S, 167 ° 51 ′ E, 328 – 340 m; 1 dd, stn DW 1654, 23 ° 28 ′ S, 167 ° 52 ′ E, 366 – 560 m; 1 lv, 2 dd, stn DW 1658, 23 ° 26 ′ S, 167 ° 50 ′ E, 320 – 336 m; 1 dd, stn DW 1672, 23 ° 43 ′ S, 168 ° 01 ′ E, 267 – 324 m; 2 dd, stn DW 1673, 23 ° 39 ′ S, 168 ° 00 ′ E, 278 m; 1 dd, stn DW 1674, 23 ° 40 ′ S, 168 ° 00 ′ E, 245 – 253 m; 3 dd, stn DW 1675, 24 ° 45 ′ S, 168 ° 09 ′ E, 231 – 233 m; 27 lv + dd, stn DW 1679, 24 ° 43 ′ S, 168 ° 10 ′ E, 298 – 324 m; 1 dd, stn DW 1680, 24 ° 45 ′ S, 168 ° 10 ′ E, 385 – 392 m; 4 dd, stn DW 1704, 23 ° 45 ′ S, 168 ° 16 ′ E, 400 – 420 m; 6 dd, stn DW 1707, 23 ° 43 ′ S, 168 ° 16 ′ E, 381 – 493 m; 3 dd, stn DW 1709, 23 ° 42 ′ S, 168 ° 16 ′ E, 380 – 389 m; 1 dd, stn DW 1710, 23 ° 44 ′ S, 168 ° 17 ′ E, 386 – 426 m; 1 dd, stn DW 1722, 23 ° 18 ′ S, 168 ° 01 ′ E, 540 m; 8 lv + dd, stn DW 1723, 23 ° 18 ′ S, 168 ° 15 ′ E, 266 – 267 m; 6 dd, stn DW 1729, 23 ° 20 ′ S, 168 ° 16 ′ E, 340 – 619 m; 2 dd, stn DW 1732, 23 ° 20 ′ S, 168 ° 16 ′ E, 347 – 1063 m. VANUATU: VOLSMAR: 2 dd, stn DW 07, 22 ° 26 ′ S, 171 ° 44 ′ E, 325 – 400 m; 1 dd, stn DW 16, 22 ° 25 ′ S, 171 ° 41 ′ E, 420 – 500 m; 1 dd, stn DW 39, 22 ° 20 ′ S, 168 ° 44 ′ E, 280 – 305 m; 1 dd, stn DW 49, 21 ° 00 ′ S, 170 ° 04 ′ E, 285 m; 1 dd, stn DW 50, 20 ° 59 ′ S, 170 ° 04 ′ E, 425 m. – MUSORSTOM 8: 1 dd, stn DW 967, 20 ° 19 ′ S, 169 ° 53 ′ E, 295 – 334 m. TONGA: BORDAU 2: 2 dd, stn DW 1607, 22 ° 15 ′ S, 175 ° 23 ′ W, 356 – 367 m.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C0DEB50FDE7201B1A067E2C.taxon	description	Description Shell thin, slender, fusiform, consisting in holotype of 3.75 protoconch and 7 teleoconch whorls. Protoconch conical (Fig. 10 C), with weakly convex whorls, last ½ whorl with 6 axial riblets, protoconch / teleoconch discontinuity distinct. Protoconch diameter 1070 µm, exposed height 1110 µm. Teleoconch whorls convex, suture deep, adpressed. Axial sculpture of broad, prominent ribs, aligned axially on spire from one whorl to the next, 6 ribs per whorl. Spiral sculpture of evenly spaced, strong, raised cords, in holotype 2 on first whorl, 3 on subsequent whorls, abapicalmost stronger, 5 on antepenultimate and penultimate whorls, 22 on last whorl. No second order axial or spiral sculpture. Siphonal canal open, long, slightly recurved. Aperture with 11 weak lirae inside, base of siphonal canal not clearly demarcated. Inner lip completely adherent to columella, forming smooth, white, shiny columellar shield, with three weak columellar plicae. No umbilicus. Colour of protoconch and interior of aperture white or fleshcoloured. Teleoconch with alternating white spiral cords and orange brown interspaces, and three white bands, one subsutural, one just below periphery of last whorl and one at tip of siphonal canal. Holotype measurements: SL 22.7, LWL 15.4 mm, AL (without canal) 5.7 mm, SW 7.8 mm. Live animal has orange body (Fig. 10 D) with irregular white spots, more numerous on body than on foot. Operculum transparent, yellow, with terminal nucleus shifted leftwards, closing aperture completely. Radula (Fig. 4 H) examined in two specimens from Norfolk Ridge (NORFOLK 1, stn DW 1658, and stn DW 1679), very similar in both specimens. Radula narrow, distance between outer edges of lateral teeth about 80 µm (1.6 % of AL). Lateral teeth medium broad, arcuate, with attenuated outer anterior corner, producing medium long ‘ handle’. Lateral teeth with outermost small cusp, on most rows overlaid by following major cusp and not seen on ventral view (marked by white arrow), five slightly recurved, short cusps, subequal in length, outer more distant from others, and one additional innermost short, but distinct, cusp. Central tooth small, trapezoid, with three short cusps, central longest. Narrow but distinct transverse folds of the membrane situated serially between rows of lateral teeth, reaching upper corners of central teeth.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C0DEB50FDE7201B1A067E2C.taxon	distribution	Distribution Coral Sea, New Caledonia to Tonga, in 245 – 610 m, alive in 245 – 450 m.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C0DEB50FDE7201B1A067E2C.taxon	discussion	Remarks Okutanius ellenae gen. et sp. nov. reaches 29.6 mm in length, showing little variation in shell shape and colouration. It resembles O. kuroseanus gen. et comb. nov., but is easily separated by its sculpture of even spiral cords and distinct colour pattern of alternating white cords and dark interspaces. O. kuroseanus gen. et comb. nov. reaches a much larger adult size, and is also broader at comparable size. O. ellenae gen. et sp. nov. also superficially resembles Granulifusus tatianae gen. et sp. nov., which differs by its short siphonal canal, strong apertural barriers, and the presence of second order spiral and axial sculpture.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C01EB50FE4725851F987A8C.taxon	description	Fig. 9 H – I	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C01EB50FE4725851F987A8C.taxon	materials_examined	Type material Holotype South Africa: off Park Rynie, southern KwaZulu – Natal, 110 m, SL 25.6 mm, SW 10.3 mm (NMSA W 9698 / T 3896).	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C01EB50FE4725851F987A8C.taxon	description	Description Shell strong, fusiform, up to 25.6 mm. Protoconch and uppermost teleoconch whorls eroded, protoconch conical, multispiral, of 3 + whorls (protoconch tip broken off). Teleoconch with regularly strongly convex whorls, with medium long siphonal canal. Aperture broad, oval, with nearly smooth columella with very weak ridges in the lower part, corresponding to spiral cords not completely overlaid by callus. Spiral sculpture of unevenly spaced distinct cords of varying strength; axial ribs strong, weakly opisthocline, and closely spaced, not aligned along the spire. Background colour creamy, with two darker brown spiral bands and more dark shell base. Primary spiral cords white.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C01EB50FE4725851F987A8C.taxon	distribution	Distribution Only known from the type locality.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C01EB50FE4725851F987A8C.taxon	discussion	Remarks The dead collected and partially distorted holotype is the only known specimen. In ventral view it is clearly seen that the shell axis is not straight, and the strongly recurved leftwards siphon is the result of abnormal shell growth. Pending knowledge on the body and radula, we tentatively allocate this species to Okutanius gen. nov., based on shell resemblance to sequenced species of the genus.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C06EB57FD17270418DB7B84.taxon	type_taxon	Type species Pseudolatirus pallidus Kuroda & Habe, 1961, here designated.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C06EB57FD17270418DB7B84.taxon	diagnosis	Diagnosis Shell medium-sized, adult size from 27 mm (V. wallacei gen. et comb. nov.) to 60 mm (V. pallidus gen. et comb. nov.), fusiform, strong but light, usually offwhite, with evenly convex to inflated teleoconch whorls, constricted shell base and long, straight or weakly sinuous canal. Protoconch paucispiral or multispiral (1.5 – 3.25 whorls). Axial sculpture of distinct narrow ribs, nearly coaxial, separated by broader (1 – 2 times) interspaces. Spiral sculpture of distinct cords, unequal in size, covering entire shell surface, major cords nodulose at intersection with axial ribs. Aperture broad, oval, relatively small, with smooth or weakly plaited columella (plaits correspond to spiral cords covered by columellar callus). Parietal knob present in some specimens. Operculum spans entire aperture, with terminal nucleus. Lateral teeth with 5 – 7 closely spaced, slightly curved cusps and much smaller, but distinct, innermost cusp. Outer anterior corner forms long distinct ‘ handle’, tooth outline convex at transition from cusp side to ‘ handle’. Central tooth small, rectangular, with three cusps. Narrow but distinct transverse folds of membrane situated serially between rows of lateral teeth. Differential diagnosis Vermeijius gen. nov. differs from Granulifusus and Okutanius gen. nov. in its uniformly coloured light shell with smooth or nearly smooth columella, more abruptly constricted shell base, producing a relatively narrow canal. The radula differs in having a proportionally broader central tooth. The molecular phylogenetic analysis clearly demonstrates the isolated position of the new genus as sister taxon to Chryseofusus.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C06EB57FD17270418DB7B84.taxon	etymology	Etymology The genus is named after Geerat J. Vermeij from University of California at Davis in recognition of his immense contribution to malacology, including the taxonomy and biogeography of the Fasciolariidae. Gender masculine. Genus composition We include in the genus Pseudolatirus pallidus, Fusinus virginiae and Fusus? retiarius; additionally, Fusinus (Simplicifusus) guidonis Delsaerdt, 1995 from Somalia, Fusinus palmarium Hadorn & Fraussen, 2006 from eastern Australia and Fusinus wallacei Hadorn & Fraussen, 2006 were not sequenced, but are attributed to Vermeijius gen. nov. based on conchological similarities.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C06EB4AFE10212C1A077EDF.taxon	description	Figs 11 A – E, 12 A – B	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C06EB4AFE10212C1A077EDF.taxon	diagnosis	Molecular diagnosis A molecular diagnosis is given in Table 4.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C06EB4AFE10212C1A077EDF.taxon	materials_examined	Type material Holotype JAPAN: off Cape Ashizuri, Kochi Prefecture (NSMT Mo 40378) [not seen]. Sequenced material SOLOMON ISLANDS: SALOMON 2: 1 lv, stn CP 2290, Tetepare, 8 ° 40 ′ S, 157 ° 32 ′ E, 384 – 418 m (MNHN IM- 2007 - 32537). NEW CALEDONIA: CONCALIS: 1 lv, stn CP 2962, Grand Passage, 18 ° 59 ′ S, 163 ° 08 ′ E, 580 – 647 m (MNHN IM- 2007 - 35093). SOUTH CHINA SEA: DongSha 2014, 2 lv, stn CP 4128, 20 ° 42 ′ N, 116 ° 08 ′ E, 420 – 444 m (MNHN IM- 2013 - 50173, MNHN IM- 2013 - 50174).	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C06EB4AFE10212C1A077EDF.taxon	description	Description Shell up to 60 mm in length, solid, fusiform, with strongly constricted base with long, nearly straight, siphonal canal, slightly inclined to left. Protoconch obtusely conical (Fig. 11 E) of 3.25 whorls, with evenly convex, brown, smooth whorls, last half whorl with seven distinct, arcuate, axial ribs, protoconch / teleoconch discontinuity distinct, marked by appearance of spiral sculpture of teleoconch. Protoconch diameter 1160 µm, exposed height 1100 µm. Teleoconch with strongly convex whorls. Aperture wide, oval, with weak columellar plicae, corresponding to primary spiral cords overlaid by callus, from nearly absent to 4 – 5 on lower half of columella. Spiral sculpture of unevenly arranged cords, alternating in strength, strongly nodulous at intersection with axial ribs. Axial ribs broadly spaced, narrow, not aligned along spire, 10 – 12 on last whorl. Growth lines usually thickened, forming reticulated structure at intersection with spiral cords. Background colour pale flesh to nearly white. Operculum elongate, leaf-shaped, nearly spanning the entire aperture. Nucleus terminal, often abraded. Radula (Fig. 12 A – B, MNHN IM- 2007 - 35093, SL 42.5 mm) long, of about 105 rows of teeth, 30 nascent, 4.3 mm long (0.41 of AL without canal); narrow, distance between outer edges of lateral teeth about 140 µm wide (1.33 % of AL without canal). Lateral teeth broad, slightly arcuate, with strongly attenuated outer anterior corner, producing long ‘ handle’. Lateral teeth with six, almost equal, recurved, closely spaced, major cusps and one smaller inner cusp. Teeth outline convex at transition of outer cusp side into ‘ handle’. Central tooth subrectangular, with three short, sharp cusps, central slightly longer. Distinct transverse folds of the membrane situated serially between rows of lateral teeth. The radula illustrated by line drawing by Hadorn & Fraussen (2002: fig. 22) is very similar, differing in the presence of 7 major cusps on lateral teeth versus 6 in our specimen.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C06EB4AFE10212C1A077EDF.taxon	distribution	Distribution Japan to New Caledonia, in 384 – 647 m.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C06EB4AFE10212C1A077EDF.taxon	discussion	Remarks Callomon & Snyder (2009) discussed and illustrated the type of the species stored in NSMT (Mo 40378) and concluded that it does not match the dimensions of the holotype (52.2 vs 56.8 mm for the latter according to the original description); the specimen may be a figured paratype. Vermeijius pallidus gen. et comb. nov. is somewhat similar to V. virginiae gen. et comb. nov.; for differences, see the description of the latter. Vermeijius virginiae (Hadorn & Fraussen, 2002) gen. et comb. nov. Figs 11 F – J; 12 C – D	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C06EB4AFE10212C1A077EDF.taxon	diagnosis	Molecular diagnosis A molecular diagnosis is given in Table 4.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C06EB4AFE10212C1A077EDF.taxon	materials_examined	Type material Holotype MOZAMBIQUE CHANNEL: off Southwest Madagascar, 22 ° 10 ′ S, 43 ° 05 ′ E, 525 m, Campagne Crevettière 1986, stn CH 79 (MNHN IM- 2000 - 9626). Sequenced material SOUTH MADAGASCAR: ATIMO VATAE: 7 lv, stn CP 3585, 25 ° 32 ′ S, 44 ° 16 ′ E, 549 – 576 m (MNHN IM- 2009 - 15020, MNHN IM- 2009 - 15025, MNHN IM- 2009 - 15026, MNHN IM- 2009 - 15030, MNHN IM- 2009 - 15031, MNHN IM- 2009 - 15033, MNHN IM- 2009 - 15034); 2 lv, stn CP 3591, 25 ° 03 ′ S; 43 ° 58 ′ E, 400 – 402 m (MNHN IM- 2009 - 15021, MNHN IM- 2009 - 15084).	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C06EB4AFE10212C1A077EDF.taxon	description	Description Shell large for genus, up to 56 mm, strong, fusiform, with strongly constricted base with long, nearly straight, siphonal canal, slightly inclined to left. Protoconch (Fig. 11 J) conical of 2.25 moderately convex, smooth, greyish and semitransparent whorls, with elevated first whorl, and 2 – 5 indistinct axial ribs at transition to teleoconch. Protoconch / teleoconch discontinuity distinct, marked by appearance of teleoconch spiral sculpture. Protoconch diameter 960 µm, exposed height 800 µm. Teleoconch with regularly strongly convex whorls. Aperture broad, oval, columella smooth, or with 1 or 2 weak, inconspicuous plicae, corresponding to underlying spiral sculpture of preceding whorl. Spiral sculpture of unevenly spaced cords, alternating in strength, slightly nodulous at intersection with axial ribs. Growth lines usually thickened, forming reticulated structure at intersection with spiral cords. Axial ribs not aligned along spire, 9 – 11 on last whorl. Background colour pale flesh to nearly white. Operculum elongate, leaf-shaped, spanning nearly the entire aperture. Nucleus terminal. Radula (Fig. 12 C – D, MNHN IM- 2009 - 15030, SL 51.5 mm) long, of about 150 rows of teeth, 40 nascent, 4.5 mm long (0.37 of AL without canal), narrow, distance between outer edges of lateral teeth about 130 µm (1.06 % of AL without canal). Lateral teeth broad, slightly arcuate, with strongly attenuated outer anterior corner, producing long ‘ handle’. Lateral teeth with six, almost equal, recurved, closely spaced, major cusps and one smaller inner cusp. Teeth outline convex at transition of outer cusp side into ‘ handle’. Central tooth subrectangular, with three very short, sharp cusps, central slightly longer. The radula illustrated by line drawing by Hadorn & Fraussen (2002: fig. 19) is very similar to that of our specimen, but has 7 major cusps on lateral teeth versus 6 in our specimen.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C06EB4AFE10212C1A077EDF.taxon	distribution	Distribution Southern Madagascar, in 400 – 576 m.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C06EB4AFE10212C1A077EDF.taxon	discussion	Remarks The species is rather similar conchologically and is a sister to V. pallidus gen. et comb. nov. It differs in having fewer protoconch whorls, thinner spiral cords, forming less defined beads at intersection with axial ribs.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C1BEB49FE6625FF19F97E67.taxon	description	Fig. 11 K	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C1BEB49FE6625FF19F97E67.taxon	materials_examined	Type material Lectotype SOMALIA: dd, 38.5 × 15.7 mm, off southern Somalia, 01 º 49 ′ N, 45 ° 29 ′ E, 1134 m, Valdivia, stn 256, designated by Hadorn & Fraussen (2002) (ZMB 61012). Sequenced material SOUTH MADAGASCAR: ATIMO VATAE: 1 lv, stn CP 3595, South of Pointe Barrow, 25 ° 35 ′ S, 44 ° 15 ′ E, 821 – 910 m (MNHN IM- 2009 - 15087).	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C1BEB49FE6625FF19F97E67.taxon	description	Description Shell large for genus, up to 57 mm in length, strong, broadly fusiform, gradually constricting to medium long, sinuous, coaxial siphonal canal. Protoconch dome-shaped, paucispiral, of 1.5 convex smooth whorls, greyish, semitransparent, with 3 weak arcuate axial ribs at transition to teleoconch. Protoconch / teleoconch discontinuity distinct, marked by appearance of teleoconch spiral sculpture. Protoconch diameter 1000 µm, exposed height 700 µm. Teleoconch with strongly convex whorls. Aperture broad, oval, columella smooth. Spiral sculpture of moderately strong cords, beaded at intersection with axial ribs and much thinner, although distinct, secondary cords, covering interspaces between primary cords; 4 – 5 primary spiral cords on spire whorls, including penultimate. Axial ribs not aligned along spire, 11 – 12 per whorl. Background colour pale flesh to nearly white. Operculum elongate, leaf-shaped, spanning nearly entire aperture. Nucleus terminal. Radula (Hadorn & Fraussen 2002: fig. 21) with medium broad, arcuate, lateral teeth, with attenuated outer anterior corner, producing medium long ‘ handle’. Lateral teeth with five slightly recurved cusps, outermost longest, more distant from others. Additional innermost short, but distinct, cusp present. Central tooth small, subrectangular, with three very short cusps, medium longest.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C1BEB49FE6625FF19F97E67.taxon	distribution	Distribution East Africa, from Somalia to South Madagascar, in 500 – 1134 m.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
C84287C40C1BEB49FE6625FF19F97E67.taxon	discussion	Remarks In the COI-based tree Vermeijius retiarius gen. et comb. nov. did not cluster with V. pallidus gen. et comb. nov. and V. virginiae gen. et comb. nov., but in the three genes tree it was recovered as a sister group to the clade combining these two species. The relationships are highly supported in the Bayesian analysis, therefore we include it in Vermeijius gen. nov. It differs from the two sequenced species by a more stout shell with paucispiral protoconch.	en	Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery, Bouchet, Philippe (2018): Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae). European Journal of Taxonomy 433: 1-57, DOI: 10.5852/ejt.2018.433
