taxonID	type	description	language	source
C55B87B4C33EFFE039F2FF04BE50FD5D.taxon	description	(Fig. 1)	en	Hespenheide, Henry A. (2007): The identity of Agrilus impexus Horn, a new species, and taxonomic notes and records for other Agrilus Curtis species (Coleoptera: Buprestidae). Zootaxa 1617 (1): 57-66, DOI: 10.11646/zootaxa.1617.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1617.1.3
C55B87B4C33EFFE039F2FF04BE50FD5D.taxon	description	Head with front shallowly but distinctly depressed along midline, surface rugose, more coarsely so above middle; epistoma 1 / 3 width of distance between inner margins of eyes between antennal insertions, shallowly emarginate below, eyes oval, slightly emarginate on inner margins; antennae serrate from segment 5, segments 5 – 11 rounded-triangular. Pronotum subequal to elytra at posterior margin, with sides shallowly rounded outward, widest near middle; marginal and submarginal carinae undulate, separated and subparallel for entire length when viewed from side; from above anterior margin regularly shallowly rounded; basal margin slightly emarginate at middle of each elytron, nearly transverse before scutellum; disk moderately convex, with distinct depression along midline and oblique depressions along lateral margins; prehumeral carinae strong, nearly straight, 1 / 3 length of pronotum; surface coarsely transversely rugose; two tiny pores along basal margin just medial to prehumeral carinae. Scutellum pentagonal with transverse ridge. Elytra subequal in width at humeri and beyond middle, lateral margins shallowly emarginate between, apices narrowly, separately rounded, minutely toothed; disk convex each elytron with narrow oblique depression at base and broader, shallower one along suture for apical 4 / 5; surface transversely imbricate-punctate. Prosternum with sides of prosternal process slightly narrowing between coxae, truncate and emarginate at apex, prosternal lobe nearly transverse. Posterior coxae with posterior margin shallowly, regularly emarginate, upper angles obtusely rounded. Abdomen with suture nearly obsolete between ventrites 1 and 2, narrowly depressed along midline of ventrites 1 – 2. Legs with first metatarsal segment equal in length to next two combined, tarsal claws similar on all feet, cleft with short inner tooth. Genitalia as in Fig. 1. Female: As male except coloration reddish coppery throughout, more golden beneath; setae on prosternum and along midline of metasternum sparse and appressed; abdominal ventrites 1 – 2 convex, without medial depression or line of setae. Specimens examined. Colorado, Colo. Spr., 6000 – 7000 ft, 15 – 30.06.1896, H. F. Wickham (USNM). Iowa, County 5, 16.07.1936 (USNM). Kansas, [without locality] (USNM), Madison, M. C. V [anDuzee]. coll. (CASC), Onaga, Crevecoeur (USNM), Onaga, 08.07.1901 (SGWC), VI / 24, Wickham Coll. (USNM), Snow, Coll Hubbard & Schwarz (USNM), Topeka, 03 Aug (USNM); Meade Co., 30.06.1986, G. A. Salsbury, sweep rangeland (TCMC); Riley Co., Jun, Popenoe (USNM), Riley Co., July 13, 14, 17, Popenoe (SGWC), Riley Co., July 19, G. A. Dean (SGWC). Minnesota, Ft. Snelling, 28.06.1923, high prairie assn. (USNM). Missouri: Camden Co., Ha Ha Tonka State Park, Lodge Glade, 7.06.2001, D. G. LeDoux & F. Lloyd (TCMC); Nebraska, J. B. Smith “ # 1 C. T. ” (USNM). New Mexico: Jemez Springs, 18. VI. 1947, B. E. White, on Chrysothamnus sp. (CASC). Oklahoma: Paroner (?), 14.07.1932, A. E. P. (SGWC). Wyoming, Sundance, 31.07.1935 (USNM), Crook Co., 08.07.1938 (USNM).	en	Hespenheide, Henry A. (2007): The identity of Agrilus impexus Horn, a new species, and taxonomic notes and records for other Agrilus Curtis species (Coleoptera: Buprestidae). Zootaxa 1617 (1): 57-66, DOI: 10.11646/zootaxa.1617.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1617.1.3
C55B87B4C33EFFE039F2FF04BE50FD5D.taxon	biology_ecology	Hosts. Adults have been collected on Chrysothamnus (Asteraceae) in New Mexico. Species of Chrysothamnus are the adult and presumed larval hosts of at least three other Agrilus: A. duncani Knull, col- lected commonly on C. nauseosus (Pall.) Britton at Portal, Arizona; A. pubifrons Fisher, collected on C. viscidiflorus (Hook.) Nutt. by W. F. Barr in Idaho; and A. walsinghami Crotch.	en	Hespenheide, Henry A. (2007): The identity of Agrilus impexus Horn, a new species, and taxonomic notes and records for other Agrilus Curtis species (Coleoptera: Buprestidae). Zootaxa 1617 (1): 57-66, DOI: 10.11646/zootaxa.1617.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1617.1.3
C55B87B4C33EFFE039F2FF04BE50FD5D.taxon	discussion	Discussion. The geographic distribution of A. impexus appears to coincide roughly with the Great Plains. In his description Horn (1891) listed specimens from both Arkansas and Nebraska. Fisher (1928) also listed specimens from Mississippi which I have not seen. The species has not been collected frequently, with only three specimens in the last 60 years known to me. As noted in the introduction, the name has been misapplied to the much more common and widespread species associated with woody legumes treated below; this practice dates from Fisher’s (1928) inclusion under A. impexus of two specimens from Arizona, reared from “ Palo Verde, ” that are the following species. Agrilus impexus is interesting for its weak sexual color dimorphism. In the series from New Mexico males are usually more or less greenish on the head, pronotum, and beneath, with a distinctly black front, and reddish-coppery on the elytra, whereas females are usually reddish-coppery throughout, although more golden beneath. Agrilus impexus is significantly larger than the following species - males measure 6.2 – 7.7 mm long (mean = 7.00 mm for 16 specimens), and females measure 6.7 – 8.0 mm long (mean = 7.31 mm for 22 specimens).	en	Hespenheide, Henry A. (2007): The identity of Agrilus impexus Horn, a new species, and taxonomic notes and records for other Agrilus Curtis species (Coleoptera: Buprestidae). Zootaxa 1617 (1): 57-66, DOI: 10.11646/zootaxa.1617.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1617.1.3
C55B87B4C33DFFE439F2FC84BE27F9B5.taxon	description	(Fig. 2) Description. Holotype male: Slender, subcylindrical, 6.00 mm long, 1.55 mm wide; dark reddish-coppery throughout, except epistoma between antennal insertions metallic green; setae pale and recumbent, short, sparse and relatively inconspicuous, except denser on front and sides of pronotum above and below prehumeral carina, denser and conspicuous in three patches on each elytron - a small spot in basal depressions, a narrow, more elongate spot along suture at 1 / 3. and a nearly isometric spot at 2 / 3 - on dorsal portions of abdominal ventrites 1 – 2, in spots on lateral portions of ventrites 3 – 5, in narrow line at middle of ventrites 1 – 2, on outer ½ of posterior coxae, in upper posterior corners of metasternum and epimeron, setae dense and semierect on prosternum and continuing along midline of anterior ½ of metasternum. Head with front convex, but narrowly, distinctly depressed along midline, surface rugose, more coarsely so above middle; epistoma between antennal insertions 1 / 4 width of distance between inner margins of eyes, emarginate below, eyes oval, slightly emarginate on inner margins; antennae serrate from segment 5, segments 5 – 11 rounded-triangular. Pronotum subequal to elytra at posterior margin, with sides shallowly expanded outward then nearly parallel to apex; marginal and submarginal carinae slightly undulate, separated for entire length and converging slightly toward base when viewed from side; from above anterior margin produced as angulately-rounded lobe; basal margin angulately emarginate at middle of each elytron, nearly transverse before scutellum; disk moderately convex, with distinct depression along midline and oblique depressions interior to and in front of prehumeral carinae to lateral margins; prehumeral carinae strong, nearly straight, 2 / 5 length of pronotum; surface coarsely transversely rugose. Scutellum pentagonal with transverse ridge. Elytra subequal in width at humeri and beyond middle, lateral margins shallowly emarginate between, apices narrowly, separately rounded, minutely toothed; disk convex, each elytron with oval depression at base and narrower, almost indistinct one along suture to apical setal spots, then narrower and stronger to apex; surface transversely imbricate-punctate, more rugose at base. Prosternum with sides of prosternal process slightly narrowing between coxae, truncate and emarginate at apex, prosternal lobe nearly transverse. Posterior coxae with posterior margin nearly straight, upper angles somewhat produced and obtusely rounded. Abdomen with suture nearly obsolete between ventrites 1 and 2, narrowly depressed along midline of ventrite 1 and anterior 2 / 3 of ventrite 2. Hind legs with first metatarsal segment equal in length to next two combined, tarsal claws similar on all feet, cleft with short inner tooth. Genitalia as in Figure 2. Allotype Female: As male except 6.10 mm long, 1.60 mm wide; entirely dark reddish-coppery, setae on front, prosternum and along midline of metasternum sparse and appressed; abdominal ventrites 1 – 2 convex, without medial depression or line of setae. Males from the type series from New Mexico and Arizona measure 3.70 – 6.10 mm long (mean = 5.38 mm for 51 specimens), and females measure 4.50 – 6.70 mm long (mean = 5.66 mm for 41 specimens).	en	Hespenheide, Henry A. (2007): The identity of Agrilus impexus Horn, a new species, and taxonomic notes and records for other Agrilus Curtis species (Coleoptera: Buprestidae). Zootaxa 1617 (1): 57-66, DOI: 10.11646/zootaxa.1617.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1617.1.3
C55B87B4C33DFFE439F2FC84BE27F9B5.taxon	materials_examined	Holotype: Arizona: [Pima Co.,] Mt. Lemmon, 23.08.1937, Drake & Andre (USNM). Allotype: Arizona: [Pima Co.,] Mt. Lemon, Santa Car. Mts., 22.08.1934, C. J. Drake (USNM). Paratypes: Arizona: Catal [ina?] Spgs, 10.4, Hubbard & Schwarz, in Parkinsonia microphylla (1 F, USNM), 11.4, Hubbard & Schwarz, on Parkinsonia microphylla (1 F, USNM); Cochise Co., SW Cochise Co., 6 mi W of Montezuma Pass, (± 5500 ’), Bear Crk., 21.08.1992, A. & N. McF., on Mimosa dysocarpa (1 M, 3 F, WFBM); Huachuca Mts., 6.8 mi W Montezuma Pass, 06.08.1978, D. S. Verity, on Acacia (1 M, DSVC); Parker Lake, 05.08.1978, D. S. Verity, on Acacia angustissima (2 M, DSVC); 2 mi E Coronado National Memorial, 07.08.1978, D. S. Verity, Calliandra (1 M, 3 F, DSVC); 2 mi NW Portal, 20.08.1970, D. S. Verity (1 M, DSVC); 5.4 mi E Dragoon, 02.05.1993, D. S. Verity, Prosopis (1 F, DSVC); Copper Canyon, 10.08.2001, J. & J. Davidson (2 M, CLBC); Pinery Cyn., 1.6 mi E State 181 on 42, 02.09.1976, F. G. Andrews, ex Calliandra humilius (1 M, 1 F, CSCA); “ D ” Hill 2 mi NE Douglas, 16.08.1978, M. Wasbauer, J. Slansky (2 F, CSCA); Paradise Rd., 1 mi W Portal, 4900 ’, 31 ° 55 ’ N 109 ° 9 – 10 ’ W, 06.06.1983, H. A. Hespenheide, on Prosopis juliflora (Sw.) DC (3 M, CHAH); 2 mi ESE Portal, 4500 ’, 31 ° 54 ’ N 109 ° 6 ’ W, 18.05.1985, H. A. Hespenheide (1 F, CHAH); Cave Ck. Cyn., Chiricahua Mts., Sunny Flat, 5100 ’, 31 ° 53 ’ N 109 ° 10 ’ W, 28.08.1980, H. A. Hespenheide, Prosopis (1 M, CHAH); Maricopa Co., 5 mi N Sunflower, 17.07.1967, W. F. Barr, on Mimosa biuncifera (1 M, WFBM), 21.06.1985, W. F. Barr, beating Mimosa (1 F, WFBM); Pima Co., same data as allotype, (1 M, USNM); same data as holotype, 23.08.1937, Drake & Andre (2 M, USNM; 1 M, WFBC); Tucson, Sabino Canyon, 16.05.1953, A. & H. Dietrich (2 M, DSVC); Tucson, Houghton Rd. at Agua Caliente Wash, 20.04.1990, W. F. Barr, beat Prosopis (1 M, WFBC); Baboquivari Cyn., Baboquivari Mts., 16.08.1988, W. F. Barr, beating Acacia greggii (1 M, 1 F, WFBM); Baboquivari Mts., 20.08.1937, Drake & Andre (3 F, USNM; 1 F, WFBM); Santa Rita Mts., Box, Cyn., 5000 ’, 20.08.1937, C. D. Johnson (1 M, WFBM), 1700 m, 03.08.1996, M. W. Gates G 96 / 045 (1 F, USNM); Madera Cn., 27.08.1964, G. Walters, beating mimosa (1 M, CHAH); 17 mi E Sells, 05.05.1963, D. S. Verity, Prosopis juliflora (1 M, DSVC); Brown Canyon, Baboquivari Mts., 01 – 02.09.1957, D. S. Verity (1 M, DSVC); Ariz., Pinal CO., 7 – 1 / 2 mi SE Oracle, 01.06.1973, D. G. Marqua (1 F, DSVC); Santa Cruz Co., Patagonia, 23.08.1937, Drake & Andre (1 F, WFBM); 1 mi W White Rock Camp, 13.08.1989, W. F. Barr, beat Prosopis (7 M, 4 F WFBM), 04.08.1990, W. F. Barr, beat Prosopis (5 M, WFBM); Montosa Cyn., 14 km E Amado, 04.08.1989, W. F. Barr (1 M, 2 F, WFBM), 13 km E Amado, 21.04.1990, W. F. Barr (1 M, 2 F, WFBM), 2 mi W Pena Blanca L., 23.07.1987, W. F. Barr, on mesquite (1 F, WFBM); 1 mi S Pena Blanca Lake, 4000 ’, 19.05.1983, R. Anderson, oak grasssland (1 M, RLWE); 12 mi N Sonoita, 26.07.1984, W. F. Barr, on mesquite (1 F, WFBC), Nogales, 17.08.1906, Schffr (1 M, USNM), 26.08.1933 (1 M, 2 F, WFBM), 17.08.1906, F. W. Nunenmacher (1 F, WFBM), 23.08.1937, Drake & Andre (1 F, WFBM); Sycamore Canyon, 9 mi W Pena Blanca Lake, 4100 ’, 12.08.1983, R. Anderson, oak savan. (1 M, RLWE). New Mexico: Dona Ana Co, Aguirre sprgs RA, 5600 ’, 26 – 27.08.1997, Wappes & Turnbow (1 F, USNM); Hidalgo Co., Peloncillo Mt, 16.08.1968, V. Roth (1 F, U. Idaho); 1 mi N Rodeo, 24.07.1989, T. Eichlin & F. Andrews, reared from Hoffmanseggia (1 M, 1 F, CSCA); 2 mi S Animas, 4.08.1994, F. Andrews & T. Eichlin (1 M, CSCA); 12.2 mi W Animas, 29.07.19987, F. Andrews & T. Eichlin, ex Prosopis juliflora (1 F, CSCA); 4 km NNW Rodeo, 4130 ’, 31 ° 52 ’ N 109 ° 03 ’ W, 06.06.1983, H. A. Hespenheide, mesquite (1 M, CHAH); 30 mi S Animas, 19.08.1980, H. A. Hespenheide (2 M, CHAH). Additional specimens examined. California: Imperial Co., Algodones Dunes 2 mi SW Cactus, 32 ° 50 ’ 00 ” N 114 ° 54 ’ 48 ” W, Site 5, 12.04.1979, collected on Olneya tesota (2 F, CSCA); Riverside Co., Palm Sprs., 22, 23.06.1957, R. L. Westcott, collected on mesquite (1 F, RLWE); Pinyon Flat, 12.06.1965, D. S. Verity (1 M, DSVC); Santa Rosa Mts., Hwy 74, alt 2000 ’, 22.06.1968, D. S. Verity, Acacia greggii (1 F, DSVC). Honduras: El Par, El Zamora, 25.071977, C. W. & L. O’Brien & Marshall (1 F, CLBC). Mexico: Nogales 71328, Ise bunker, 51 - 3289, 3.04.1951 (1 M, USNM); at Nogales, 1.03.1967, C. H. Spitzer, w. Capsicum sp. (bell pepper), 67 - 6344 (1 F, USNM); Baja California, Bahia de Los Angeles, 19.06.1968, N. Leppla et al. (2 M, WFBC); Parador Punta Prieta, 24.06.1979, D. S. Verity, Prosopis juliflora (5 M, 2 F, DSVC); 3 mi S San Augostin, 25.06.1980, D. S. Verity, on Prosopis (1 F, DSVC). Baja California Norte, 31.7 mi WNW Bahia de Los Angeles, 12.07.1979, Hardy, Andrews & Giuliani, collected on Prosopis juliflora (1 M, CSCA); vic. “ km 103, ” E El Rosario, 25.04.1985, R. L. Westcott (1 M, RLWE); 3 mi SSW Desengano, 24.04.1985, R. L. Westcott, beating mesquite, Prosopis glandulosa var. torreyana (23 M, 24 F, RLWE); San Matias Pass, Hwy. 3, km. 149, 27.06.1983, R. L. Westcott, Eriogonum fasciculatum var. polifolium (1 M, RLWE); N., Faro de San Jose Rd. 27 km (air) WSW Catavina, 365 m, 29.34.13 N 115.54. 53 W, 19.03.1999, R. L. Westcott, beating mesquite (4 M, RLWE); vic. Las Arrastas, 425 m ,, 25.05.1989, R. L. Westcott (1 M, RLWE); 17 mi E Valle de Trinidad, 700 m, 02 – 03.07.1975, R. L. Westcott (1 F, RLWE); San Matias Pass, Hwy 3, km 149, 27.06.1983, D. S. Verity, Prosopis juliflora (2 F, DSVC); 14 km S Rosarito, 28 ° 31 ’ 41 ” N 114 ° 02 ’ 14 ” W, 8 / 12.07.1988, C. L. Bellamy, beating Prosopis glandulosa var. torreyana (1 F, CLBC); 31.7 mi WNW Bahia de Los Angeles, 12.07.1979, Hardy, Andrews & Giuliani, collected on Prosopis juliflora (4 M, CSCA); 13 mi W San Felipe, 01.04.1993, F. Andrews, A. Gilbert (1 M, CSCA); 2.6 mi SE Catavina, 23.03.1981, D. Faulkner, F. Andrews, collected on Calliandra californica (1 M, CSCA); 16 mi N Puertecitos, 650 ’, 30.03.1993, F. Andrews, A. Gilbert (1 M, CSCA); Baja California Sur, Bahia Concepción, vic. El Requesón, 09.07.1988, em 25.08.1988, R. L. Westcott, reared ex Ditaxis b. brandegeei (1 M, RLWE); Bahia Concepción, 40 km along Hwy. 1, 23.05.1989, em 22.06.1989 – 27.02.1990, R. L. Westcott, reared ex wood Ditaxis brandegeei (5 M, 5 F, RLWE); Ramal Los Naranjos, km 12 – 14, 490 – 610 m, 23 ° 17 ’ N 109 ° 51 ’ W, 02.10.1987, R. L. Westcott (1 F, RLWE); 7 mi S Loreto, 25 ° 55 ’ N, 20.05.1989, R. L. Westcott (1 M, RLWE); 1 mi NE La Burrera, 500 m, 23 ° 30 ’ 38 ” N 110 ° 02 ’ 40 ” W, 05 / 06.10.1987, R. L. Westcott & A. D. Mudge, on living twigs & small limbs of Acacia farnesiana (1 M, RLWE); 31 – 32 km W La Paz, 23 / 24.07.1977, Dozier & Westcott, beating Prosopis glandulosa var. torreyana (11 M, 2 F, RLWE); 33 km W La Paz, 24.07.1977, Dozier & Westcott, beating Prosopis glandulosa var. torreyana (4 M, 2 F, RLWE); 3 mi S San Pedro, 23.07.1977, R. L. Westcott, beating mesquite, Prosopis glandulosa var. torreyana (2 M, 1 F, RLWE); 1 mi N, 6 mi W San Jose del Cabo, 400 m, 15 / 16.10.1978, Dozier & Westcott, on Acacia (1 F, RLWE); Hwy. 1, 7 – 1 / 2 km SW Tropic of Cancer, 215 m, 17.10.1978, Dozier & Westcott (1 M, 4 F, RLWE); 3 km E La Burrera, 515 m, 02 / 03.09.1977, Fisher & Westcott (1 F, RLWE); 1 km SW La Burrera, 470 m, 27.08.1977, Fisher & Westcott (1 M, RLWE); 6 / 7 mi NW San Jose del Cabo, 13 – 1400 ’, 21 / 22.09.1976, R. L. Westcott, Mimosa xantii, celosa (4 M, 1 F, RLWE); 2 mi E El Triunfo, 1700 ’, 17.09.1976, R. L. Westcott, beating Prosopis sp. (2 M, RLWE), D. S. Verity, Prosopis (1 F, DSVC); 4 – 1 / 2 mi SW Miraflores, 1200 ’, 20.09.1976, R. L. Westcott (1 M, RLWE); 31.8 km W La Paz, 22.04.1985, R. L. Westcott (20 M, 7 F, RLWE); 3.3 mi S Col. Calles, 17.04.1985, R. L. Westcott (1 M, RLWE); 8 mi S Pescadero, 16, 17.04.1985, R. L. Westcott (1 M, 1 F, RLWE); 3 – 1 / 2 mi SE Rancho Buena Vista (110 °), 1300 ’, 18.04.1985, R. L. Westcott (1 M, 1 F, RLWE); 6 – 7 mi NW San Jose del Cabo, 13 – 1400 ’, 21 – 22.09.1976, D. S. Verity, Mimosa xantii (1 M, 3 F, DSVC); 36 km S Mulegé, 11.08.1992, H. & A. Howden (1 M, CMNC); 57 km SW Loreto (jct Rd. to Agua Verde), 360 m, 12.08.1992, H. & A. Howden (2 M, CMNC); Ramal Naranjos Rd., 7.3 mi W Hwy 1, 1500 ’, 01.09.1990, F. Andrews, T. Eichlin, A. Gilbert (3 M, CSCA); Ramal Naranjos Rd., 2.9 mi W Hwy 1, 650 ’, 01.09.1990, F. Andrews, T. Eichlin, A. Gilbert (1 F, CSCA); Ramal de Naranjas 6 mi W Hwy 1 nr Santa Anita, 10.10.1983, F. Andrews, D. Faulkner (1 F, CSCA); 7.4 mi E Hwy 19, nr El Mezquitillo, 1000 ’, 03.09.1990, F. Andrews, T. Eichlin, A. Gilbert (2 M, CSCA); 36.6 mi SE Todos Santos, 10.10.1983, Faulkner & Andrews (1 M, 2 F, CSCA); 12.2 mi SE San Perdito near Rancho Saucito, 08.10.1983, F. Andrews, D. Faulkner (1 F, CSCA); 7 mi E San Ignacio, 23.09.1981, F. Andrews, D. Faulkner (1 M, CSCA); 29.5 mi S Loreto, 25.09.1981, F. Andrews, D. Faulkner (1 F, CSCA); San Bartolo, 03.10.1981, F. Andrews, D. Faulkner (1 M, CSCA); 8.7 mi N Santa Anita, 650 ’, 05.09.1990, F. Andrews, T. Eichlin, A. Gilbert (1 F, CSCA); 14 mi N Cabo San Lucas nr San Ramon, 1200 ’, 04.09.1990, F. Andrews, T. Eichlin, A. Gilbert (1 F, CSCA); 6.4 mi SE San Perdito, 09.10.1981, F. Andrews, D. Faulkner (1 F, CSCA); 14 mi N Todos Santos, 04.10.1981, F. Andrews, D. Faulkner (1 M, CSCA); Guerrero., 4.7 km WSW Xochipala, NNW Chilpancingo, 1310 m, 11.10.1989, R. L. Westcott, collected from Acacia sp. (1 M, RLWE); 8 mi N Iguala, 23.08.1958, R. B. Selander (1 F, DSVC); vic. Ahuacatitlan, 1540 m, 18 ° 21.745 ’ N 99 ° 48.922 ’ W, 21.07.2003, C. L. Bellamy, CLB 856, misc. beating (1 M, CLBC); 3 km SE Xalitla, 610 m, N 18.00 W 98.24, 16.07.1992, C. L. Bellamy (2 M, 1 F, CLBC); Carr. Iguala-Taxco, 1133 m, 18 ° 25 ’ 30 ” N 99 ° 36 ’ 06 ” W, 08.07.2005, H. Brailovsky, E. Barrera (1 M, UNAM); Michoacán, 30.3 km N Arteaga, 945 m, 31.07.1988, R. S. Anderson ,, Acacia woodland (1 M, RLWE); Morelos, El Limon, Sierra de Huautla, 610 m, 18.32 N - 98.57 W, 11.07.1996, C. L. Bellamy No. 577 (1 [sex?], CLBC); 13 mi E Cuernavaca, 12.05.1983, C. W. O’Brien (1 M, CLBC); vic. Contlalco above Rio Amacuzac, 1050 m, N 18.39 W 99.27 W, 16.07.1999, C. L. Bellamy, CLB 712 (1 F, CLBC); Oaxaca, 10 km NNE Huajuapan de Leon, 27.10.1990, R. L. Westcott, collected from Acacia (14, RLWE); 10 km S S. Fco Huapanapan, 29.10, 01.11.1990, C. L. Bellamy (1 F, CLBC); 2.5 km S S. Fco Huapanapan, 1830 m, 18.06 N 97.40 W, 29.10.1990, C. L. Bellamy (1 F, CLBC); 4 km SE Totolapan, Hwy 190, 915 m, N 16 ° 40 ’ 06 ” W 96 ° 16 ’ 28 ”, 18.07.2003, C. L. Bellamy, CLB 845, misc. beating (1 M, CLBC); 5 km N Tezoatlan, 1625 m, 06.07.1992, C. L. Bellamy (1 F, CLBC); 8 mi SE Tecomavaca, Hwy 131, 1500 ’, 31.08.1982, C. & L. O’Brien & G. Wibmer (1 F, CLBC); 4 km W Tamazulapan, 27.10.1990, C. L. Bellamy, beating Prosopis (2 M, 3 F, CLBC); 3 km N Cacaloxtepec, 06.07.1992, C. L. Bellamy (1 F, CLBC); 6 km SW Cacaloxtepec, 1725 m, 17.40 N 97.46 W, 28.10.1990, C. L. Bellamy (4 M, 2 F, CLBC); 7 km SW Cacaloxtepec, 1725 m, N 17.46 “ E ” 97.46 W, 06.07.1992, C. L. Bellamy (3 M, 2 F, CLBC); 10 km NNW Huajuapan de Leon, Hwy 190, 02.11.1990, C. L. Bellamy (2 M, 1 F, CLBC); 3 – 5 km N Huajuapan de Leon, 1770 m, 28.10, 01.11.1990, C. L. Bellamy (1 F, CLBC); 6 km SE Huajuapan de Leon, 27,28.10, 01.11.1990, C. L. Bellamy (2 M, 7 F, CLBC); 0.5 km NE Cacaloxtepec, 5484 ft gps, N 17 ° 44 ’ 29 ” W 97 ° 46 ’ 36 ”, 15.10.2004, C. L. Bellamy, CLB 889, beating white-thorn legume trees (1 F, CLBC); Puebla, Puerto del Gato, 11.5 km NW Tehuitzingo, 1220 m, 13.10.1989, R. L. Westcott, collected from Acacia sp. (1 F, RLWE); 2.7 km N Chietla, 1200 m, 18 ° 30 98 ° 30, 12.10.1989, R. L. Westcott (2 M, RLWE); 2.7 km NW Petlalcingo, 1570 m, 18 ° 04 ’ N 97 ° 58 ’ W, 07.07.2003, 1990, R. L. Westcott (1 F, RLWE); near Puebla, 06.07.1959, D. S. Verity (1 M, DSVC); 6 km SW San Bartolo Teontepec, 1890 m, 18.29 N 97.34 W, 14.07.1999, C. L. Bellamy, CLB 710 (1 F, CLBC); 6 km N Las Cidras, Hwy 190, 08.09.1985, W. Opitz (2 F, CLBC); 7 km SSE Acatlan, 1280 m, 05.07.1992, C. L. Bellamy (2 M, 7 F, CLBC); 14 mi SE Petlalcingo, 5800 ’, 01.06.1974, O’Briens & Marshall (2 M, CLBC); 3 km N Calipan, 1890 m, 18.17 N 97.11 W, 30.10.1990, C. L. Bellamy (1 M, 1 F CLBC); 2.7 km NW Petlalcingo, 1500 m, 18 ° 05 ’ 49 ” N 97 ° 56 ’ 49 ” W, 20.07.2003, C. L. Bellamy, CLB 852, misc. beating legume trees (2 M, 2 F, CLBC), 2.7 km NW Petlalcingo, Hwy 190, 4860 ft gps 18 ° 05 ’ 52 ” N 97 ° 56 ’ 45 ” W, 20.10.2006, 1990, C. L. Bellamy, CLB 985, beating misc. Acacia spp. (1 F, CLBC); Puerto del Gato, 12 km NW Tehuitzingo, N 18 ° 23 ’ 29 ” W 98 ° 20 ’ 01 ”, 14.10.2004, C. L. Bellamy, CLB 886, beating Acacia sp. (4 M, 1 F, CLBC); 11 km SE Izucar de Matamoros, Hwy 190, (Microondas Xuchapa road), 1390 m, N 18 ° 31 ’ 20 ” W 98 ° 24 ’ 38 ”, 14.10.2004, C. L. Bellamy, CLB 884, beating Acacia sp. (2 M, CLBC); 4 km SSE Chila, 1770 m, 17 ° 57 ’ 98 ° 07 ’, 05.07.1992, R. L. Westcott (1 M, RLWE); 6 km W Teontepec, 1865 msm, 18.28.34 N 97.34.30 W, 26.06.1997, J. Romero Napoles (1 M, CEAM); Sinaloa, Mazatlan vic. 8.5 km SW Noria, 600 ’, 02.12.1989, Rifkind, Gum, Williams (1 M, WFBC); 7 mi S Culiacan, 23.08.1960, R. L. Westcott (1 F, RLWE); 20 mi S Culiacan, 22.07.1965, D. S. Verity (1 M, DSVC); 5.3 mi SW Soquitlan, 30.06.1982, F. G. Andrews (3 M, 2 F, CSCA); Sonora, 2 mi E Alamos, 24.08.1959, R. L. Westcott (1 M, RLWE); 8 mi S Nogales, 14.08.1973, D. S. Verity (1 F, DSVC); 20 – 31 km N Alamos, 380 – 420 m, 08.08.1992, J. Rifkind (3 F, CLBC); 18 km E Navojoa, 284 m, 08.08.1992, J. Rifkind (2 F, CLBC); San Carlos, 16.07.1983, E. Fisher, mal. trap (1 F, CSCA); near Benjamin Hill, 06.1968, M. J. Wargo, F. G. Andrews, J. W. Smith (1 M, CSCA). Costa Rica: Guanacaste Pr., 6 mi S, 6 mi W Cañas, Taboga, 10 o 19 ’ N 85 o 09 ’ W, 06, 11.02.1967, H. A. Hespenheide, Mimosa (1 M, 2 F, CHAH), 06, 08.02.1967, H. A. Hespenheide, Inga sp. (4 M, 1 F, CHAH), 07, 08.02.1967, H. A. Hespenheide, Mimosa albida (1 M, 3 F, CHAH), 11.02.1967, H. A. Hespenheide, Pithecellobium dulce (1 M, 3 F, CHAH). Note: Over the years, many additional specimens of A. paraimpexus have been seen and annotated by me as “ A. impexus. ” I have included here only material seen in the last two years.	en	Hespenheide, Henry A. (2007): The identity of Agrilus impexus Horn, a new species, and taxonomic notes and records for other Agrilus Curtis species (Coleoptera: Buprestidae). Zootaxa 1617 (1): 57-66, DOI: 10.11646/zootaxa.1617.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1617.1.3
C55B87B4C33DFFE439F2FC84BE27F9B5.taxon	distribution	Distribution. As treated here, from southwestern New Mexico to Southern California, south through western México, including Baja California, to Costa Rica (see also record in Hespenheide 1989, Westcott et al. 1990).	en	Hespenheide, Henry A. (2007): The identity of Agrilus impexus Horn, a new species, and taxonomic notes and records for other Agrilus Curtis species (Coleoptera: Buprestidae). Zootaxa 1617 (1): 57-66, DOI: 10.11646/zootaxa.1617.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1617.1.3
C55B87B4C33DFFE439F2FC84BE27F9B5.taxon	biology_ecology	Hosts. Adults of this species have been collected on a variety of mimosaceous shrubs - Inga sp., mesquite (Prosopis spp., including P. glandulosa var. torreyana (L. D. Benson) M. C. Johnst. and P. juliflora (Sw.) DC), Mimosa albida Humb. & Bonpl. ex Willd., M. biuncifera Benth., M. dysocarpa Benth., M. xantii Gray, Acacia angustissima (Mill.) Kuntze, A. farnesiana (L.) Willd., A. greggii Gray, Calliandra californica Benth., C. humilius Benth., Pithecellobium dulce (Roxb.) Benth. - and Olneya tesota Gray (Fabaceae, s. str.) Cercidium (“ Parkinsonia ”) microphyllum (Torr.) Rose & Standl. (Caesalpinacae). Specimens included here as A. paraimpexus have been reared from tubers of the caesalpinoid legume Hoffmanseggia jamesii Torrey & Gray in extreme southwestern New Mexico (Eichlin & Andrews; Fred Andrews, personal communication) and from twigs of Ditaxis b. brandegei (Millsp.) Rose & Standl. (Euphorbiaceae) in Baja California (Westcott) - see below.	en	Hespenheide, Henry A. (2007): The identity of Agrilus impexus Horn, a new species, and taxonomic notes and records for other Agrilus Curtis species (Coleoptera: Buprestidae). Zootaxa 1617 (1): 57-66, DOI: 10.11646/zootaxa.1617.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1617.1.3
C55B87B4C33DFFE439F2FC84BE27F9B5.taxon	etymology	Etymology. This species is named for its similarity to and confusion with Agrilus impexus.	en	Hespenheide, Henry A. (2007): The identity of Agrilus impexus Horn, a new species, and taxonomic notes and records for other Agrilus Curtis species (Coleoptera: Buprestidae). Zootaxa 1617 (1): 57-66, DOI: 10.11646/zootaxa.1617.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1617.1.3
C55B87B4C33DFFE439F2FC84BE27F9B5.taxon	discussion	Discussion. Because of the distinct elytral spots, this species keys to Agrilus impexus in couplet 104 of Fisher’s (1928) monograph, and, as noted above, specimens from Arizona were included by him under that name although they were smaller (4.9 and 5.3 mm) than the size range given for A. impexus (“ 6.5 – 7.5 mm ”). In addition to size and coloration, discussed under A. impexus, A. impexus has stronger and more elongate medial and posterior elytral spots, a more convex anterior portion of the pronotum, and prehumeral carinae which are stronger, straighter, and diverge less from the marginal carinae of the pronotum. Overall, those two species differ considerably more than do A. paraimpexus and A. addendus. In A. addendus the elongate medial elytral spot is typically more conspicuous than the posterior spot (the opposite of many A. paraimpexus), and is often joined to the basal spot, although frequently all of the elytral spots are indistinct. In general, the elytral spots of setae are more distinct and conspicuous on females than on males. As Fisher (1928) noted in his key, A. addendus typically lacks a medial depression on the pronotum, which is like that of A. impexus in being more convex on the anterior portion. The genitalia of all three species are very similar to one another (Figures 1 – 3), as well as to those of the closely-related A. illectus Fall. There is some variation in the form of the genitalia in all three species, but those of A. paraimpexus are typically distinct in having the apices of the parameres apically truncate rather than rounded. Overall, A. paraimpexus and A. addendus appear to be closely related allopatric species with A. addendus occurring in Texas and eastern México and A. paraimpexus occurring along the Pacific side of Central America and México into the southwestern U. S. Because of variation over the extensive range of A. paraimpexus, especially in Baja California, and the relatively small differences between A. paraimpexus and A. addendus, only specimens from New Mexico and Arizona are designated types of A. paraimpexus.	en	Hespenheide, Henry A. (2007): The identity of Agrilus impexus Horn, a new species, and taxonomic notes and records for other Agrilus Curtis species (Coleoptera: Buprestidae). Zootaxa 1617 (1): 57-66, DOI: 10.11646/zootaxa.1617.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1617.1.3
C55B87B4C339FFEA39F2F93CBF75FEE5.taxon	description	(Fig. 3)	en	Hespenheide, Henry A. (2007): The identity of Agrilus impexus Horn, a new species, and taxonomic notes and records for other Agrilus Curtis species (Coleoptera: Buprestidae). Zootaxa 1617 (1): 57-66, DOI: 10.11646/zootaxa.1617.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1617.1.3
C55B87B4C339FFEA39F2F93CBF75FEE5.taxon	diagnosis	Diagnosis. As Agrilus paraimpexus, but pronotum with disk more regularly convex on anterior ½, depressed only very slightly along midline on basal ½, lateral margins shallowly arcuately rounded; elytra with spots of setae in sutural depression indistinct and usually inconspicuous, basal and median spots often connected, api- cal spots often elongate, not distinct. Genitalia as in Fig. 3.	en	Hespenheide, Henry A. (2007): The identity of Agrilus impexus Horn, a new species, and taxonomic notes and records for other Agrilus Curtis species (Coleoptera: Buprestidae). Zootaxa 1617 (1): 57-66, DOI: 10.11646/zootaxa.1617.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1617.1.3
C55B87B4C339FFEA39F2F93CBF75FEE5.taxon	distribution	Distribution: Texas to Veracruz, México. Specimens examined. Texas: [Texas, Belfrage] “ 221. // J. L. Leconte / Coll. // Type 5079 // addendus / Cr., ” (Lectotype, MCZ); “ Tex // addendus / Cr. // Horn Coll / H 4737 ” Lectoallotype (MCZ); “ Tex // 4737 // Type No 56659 USNM // Agrilus frisoni Fisher ” (USNM). Brownsville, 22.05.1935, J. N. Knull (7, NMPC, CHAH); Davis Mts., 24.06.1956, D. J. & J. N. Knull (RLWE); Del Rio, 25 – 26.04.1959, Becker & Howden, beating Acacia sp. (1, CHAH); Rocksprings, 25.04.1959, Becker & Howden (1, CHAH); Blanco Co., 8 mi S Sandy, 03.05.1988, W. F. Barr (2, WFBC); Coke Co., 3 mi SE Bronte, 30.03.1972, C. W. O’Brien (1, RLWE); Dimmit Co., 7 mi N Carrizo Springs, 29.03.1972, C. W. O’Brien (1, RLWE); Kenedy Co., 49 mi N Raymondville, 30.06.1961, R. L. Westcott (5, RLWE); Kimble Co., Segovia, 12.04.1996, C. L. & S. L. Staines, taken on mesquite (1, CHAH); Pecos Co., Fort Stockton, 30.08.2003, C. W. & L. B. O’Brien (1, CLBC); Sutton Co., 8 mi SW Roosevelt, 18.04.1976, F. T. Hovore (1, RLWE), R. Penrose (6, RLWE); Upton Co., 9 mi SW Rankin, 26.03.1972, C. W. O’Brien & Marshall (1, RLWE); 25.03.1972, C. W. O’Brien & Marshall, night (3, RLWE); Uvalde Co., Garner St. Pk., 14.06.1983, B. K. Dozier (1, CHAH), Speir Rch., 3 mi NW Uvalde, 1 – 7.05.1977, T. Eichlin, M. Wasbauer (2, RLWE), 1.05.1977, Malaise trap 10 A- 5 P (1, RLWE), 4.05.1977, Malaise trap 9 A- 5 P (1, RLWE), 6.05.1977, Malaise trap 10 A- 6 P (1, RLWE), 7.05.1977, beating vegetation (1, RLWE); 15.4 mi W Cline, 3.07.1972, R. L. Westcott (5, RLWE). Mexico: Nuevo Leon, 29 km W Linares on # 60 Santa Rosa Canyon, 2400 ’, 3.06.1983, R. Anderson, dry oak forest (1, RLWE); Tamaulipas, Nacimiento del Rio Frio, 5 mi S Gomez Farias, 400 ’, 5 - 6.061983, R. Anderson, tropical decid. forest (1, RLWE); Hwy 101, 11 mi SW Cd Victoria 4000 ’, 22.07.1982, O’Brien & Wibmer (1, CLBC); Hwy 101, 81 mi NW Cd Victoria 1250 ’, 21.07.1982, O’Brien & Wibmer (1, CLBC); Veracruz, 45 mi S Tampico, Moldenke, from Myrmecodendron [= Acacia] (1, CHAH); [Mexico], 20.03.1943, with lettuce, Laredo, No. 31677, Lot No 43 - 2577 (1, USNM).	en	Hespenheide, Henry A. (2007): The identity of Agrilus impexus Horn, a new species, and taxonomic notes and records for other Agrilus Curtis species (Coleoptera: Buprestidae). Zootaxa 1617 (1): 57-66, DOI: 10.11646/zootaxa.1617.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1617.1.3
C55B87B4C339FFEA39F2F93CBF75FEE5.taxon	discussion	Discussion. This species is very close to A. paraimpexus in morphology and shares mesquite (Prosopis spp.) as an adult host, but is allopatrically distributed and morphologically distinct as given in the diagnosis and as discussed under A. paraimpexus. There are two specimens in the Museum of Comparative Zoology labeled as “ Cotypes ” of A. addendus. Fisher (1928: 269) clearly considered Cotype 1 to be the type of A. addendus (“ Redescribed from the male cotype No. 1 in the Museum of Comparative Zoology ”) although he did not formally designate it to be the lectotype. Fisher also wrote (1928: 270): “ The two cotypes in the LeConte and Horn collections were both collected by Belfrage in Texas. Specimen No. 2 in the LeConte collection labeled “ addendus Cr. Type 5079 - 2 ” is A. abjectus Horn, and not A. addendus. ” On the preceding page Fisher (1928: 269) also says his description of the female of A. addendus was made from “ the female cotype No. H- 4737 in the Philadelphia Academy of Natural Sciences. ” The Horn collection is also now at the Museum of Comparative Zoology and there are five specimens labeled H- 4737 (labeled “ Tex ” or “ S. W. Tex ”), only one of which also bears the label “ addendus / Cr., ” apparently in the same hand as Cotype 1, and it is a female. Although Fisher refers to these specimens as cotypes, the specimens themselves are not so labeled, and Crotch’s description gives male characters, but does not specify the number of specimens or refer to a female. Interestingly, Horn’s (1891) treatment of A. addendus listed Texas and Arizona as the species’ range. Under “ Variations, ” Horn writes that “ two forms may be separated in this species, those with the pubescent spots well marked and those without them. The former variety shows some pubescence at the sides of the thorax and at the sides of the last three ventral segments. ” From this it seems clear that Horn is referring to A. paraimpexus and A. addendus, respectively, and that he recognized both the close relationship and the consistent differences between A. addendus and A. paraimpexus. Interestingly, the unique male type of Fisher’s Agrilus frisoni also has a penciled label with the number 4737 and may have come from the same series from which A. addendus was described. It differs from A. addendus only in the deep medial depression of the pronotum, which seems to be an individual deformity in that it is not perfectly bilaterally symmetrical. The only non-catalogue reference to A. frisoni since its description is apparently that of MacRae and Nelson (2003) which listed distributional records without comment. Specimens in that paper probably include both A. addendus (Tamaulipas) and A. paraimpexus (Puebla, Sinaloa), with the Chihuahua record uncertain as to which. Males from Texas measure 4.10 – 6.10 mm long (mean = 5.28 mm for 27 specimens), and females measure 4.40 – 6.20 mm long (mean = 5.34 mm for 23 specimens).	en	Hespenheide, Henry A. (2007): The identity of Agrilus impexus Horn, a new species, and taxonomic notes and records for other Agrilus Curtis species (Coleoptera: Buprestidae). Zootaxa 1617 (1): 57-66, DOI: 10.11646/zootaxa.1617.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1617.1.3
C55B87B4C337FFEB39F2F94EB827FD75.taxon	description	(Figs. 4 – 5)	en	Hespenheide, Henry A. (2007): The identity of Agrilus impexus Horn, a new species, and taxonomic notes and records for other Agrilus Curtis species (Coleoptera: Buprestidae). Zootaxa 1617 (1): 57-66, DOI: 10.11646/zootaxa.1617.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1617.1.3
C55B87B4C337FFEB39F2F94EB827FD75.taxon	diagnosis	Diagnosis. Slender (Fig. 4), coppery-red throughout; pronotum with disk transversely rugose, deeply depressed and sparsely setose along midline for entire length, lateral margins nearly parallel; elytra with stripes of setae in sutural depression on posterior 5 / 6, apices separately prolonged, acute, with small teeth; apex of vertical portion of first abdominal ventrite expanded, setose; spots of setae on upper anterior angles of abdominal ventrites 3 and 4 (Fig. 4). Genitalia as in Fig. 5. Specimens examined. Texas: Tom Green Co., San Angelo, 05.05.1982, T. W. Fuchs (TAMU). Guatemala: Alta Verapaz, Lanquin, 1000 ft., 02, 03, 11.061948, R. D. Mitchell (FMNH); El Progresso, 3 – 7 km N Estancia de la Virgen, 1000 – 1500 m, 24.06.1993, F. Génier (CMNC), 12 km N Estancia de la Virgen, 1400 m, 05.06.1991, R. Anderson (CMNC).	en	Hespenheide, Henry A. (2007): The identity of Agrilus impexus Horn, a new species, and taxonomic notes and records for other Agrilus Curtis species (Coleoptera: Buprestidae). Zootaxa 1617 (1): 57-66, DOI: 10.11646/zootaxa.1617.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1617.1.3
C55B87B4C337FFEB39F2F94EB827FD75.taxon	discussion	Discussion. This widespread species has been reported from Tabasco and Veracruz (Waterhouse 1889), as well as from the Mexican states of Chiapas, México, Oaxaca and San Luis Potosí (Westcott et al. 1989) and from Colima, Guerrero, Hidalgo, Michoacán, Nayarit and Puebla, and from Honduras (Westcott and Hespenheide 2006). The species keys to A. nevadensis Horn in Fisher’s key (1928), but differs in color, pronotal sculpture, pattern of setae and nature of the apices of the elytra, and pattern of setae on abdominal ventrites. Specimens vary in size from 7.6 – 10.8 mm in length (mean = 9.17 mm, N = 46).	en	Hespenheide, Henry A. (2007): The identity of Agrilus impexus Horn, a new species, and taxonomic notes and records for other Agrilus Curtis species (Coleoptera: Buprestidae). Zootaxa 1617 (1): 57-66, DOI: 10.11646/zootaxa.1617.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1617.1.3
