identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
C67D87F00C2DFFEBFF0A4EAB8FEC06B7.text	C67D87F00C2DFFEBFF0A4EAB8FEC06B7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zodarion Walckenaer 1826	<div><p>Genus Zodarion Walckenaer, 1826</p><p>Type species. Enyo nitida Audouin, 1826</p><p>Regarding male palps and epigynes, the Zodarion species of Morocco can be classified in the following groups:</p><p>italicum group</p><p>Diagnosis. Males: Tibial apophysis short, triangular, as wide or wider than long; median apophysis with basal part broader than triangular distal part; embolus without subterminal tooth, basal part often with ridges. Females: Epigyne without pit or slit-like apertures, with two parallel or converging chitinous sutures, limiting a square, trapezoid or triangular postero-median plate.</p><p>Species in Morocco. Zodarion isabellinum: (Simon, 1870) and Z. pallidum Denis, 1952 .</p><p>elegans group</p><p>Diagnosis. Males: tibial apophysis 2–3 longer than wide, terminally recurved and with one or more lobes, teeth or grooves; median apophysis flat and compact, with medio-lateral incision; embolus oblique and straigth, with subterminal and terminal teeth. Females: Epigyne with deep, transverse pit, at posterior margin mostly with pockets.</p><p>Species in Morocco. Zodarion maculatum (Simon, 1870), Z. maghrebense sp. nov. and Zodarion valentii Bosmans, Loverre &amp; Adante, 2019 .</p><p>mostafai group</p><p>Diagnosis. This is a newly established group of which only Z. trianguliferum was previously known. Since this species is only known from the female, Z. mostafai sp. nov., with both sexes known, is selected to give a name to the new group.</p><p>Males with tibial apophysis strongly elongated, 3–4 times longer than wide, with terminal peculiar knob; median apophysis flat, with wide proximal part and pointed distal part; embolus without subterminal tooth (difference with elegans group), tip curved. Females: epigynes with paired pockets, with only posterior margin chitinised; spermathecae small, separated by more than four times their diameter.</p><p>Species in Morocco. Zodarion azrouense sp. nov., Z. ericorum sp. nov., Z. jeanclaudeledouxi sp. nov., Z. mostafai sp. nov., Z. ogeri sp. nov., Z. trianguliferum Denis, 1952 and Z. wesolowskae sp. nov.</p></div>	https://treatment.plazi.org/id/C67D87F00C2DFFEBFF0A4EAB8FEC06B7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Benhalima, Souâd;Bosmans, Robert	Benhalima, Souâd, Bosmans, Robert (2020): Revision of the genus Zodarion Walckenaer, 1833 (part IV). The species of Morocco (Araneae: Zodariidae). Zootaxa 4899 (1): 93-114, DOI: 10.11646/zootaxa.4899.1.5
C67D87F00C2EFFEAFF0A4BBC8ED104FD.text	C67D87F00C2EFFEAFF0A4BBC8ED104FD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zodarion Walckenaer 1826	<div><p>Key to the Zodarion species of the Moroccan region</p><p>Males (male of Z. trianguliferum unknown)</p><p>1 Tibial apophysis equal to or shorter than the tibia’s diameter (Figs 3, 9, 15, 19).................................... 2</p><p>- Tibial apophysis much longer than the tibia’s diameter (Figs 32–33, 37–38, 42–43, 72–73, 76–77, 80–81, 84–85, 88–89, 92–93).............................................................................................. 3</p><p>2 Embolus gradually narrowing; median apophysis small (Figs 10, 20).................................... Z. pallidum</p><p>- Distal part of embolus needle-like; median apophysis large (Figs 4, 16)................................ Z. isabellinum</p><p>3 Embolus terminally with two teeth (Figs 29–30, 34–36, 39–41)................................................. 4</p><p>- Embolus terminally with one tooth (Figs 70–71, 74–75, 78–79, 82–83, 86–87, 90–91)............................... 6</p><p>4 Tibial apophysis with oblique retrolateral groove (Figs 32–33).......................................... Z. valentii</p><p>- Tibial apophysis without oblique retrolateral groove (Figs 37–38, 42–43)......................................... 5</p><p>5 Tibial apophysis elongated, with median blunt tooth (Figs 37–38)..................................... Z. maculatum</p><p>- Tibial apophysis shorter, with recurved tip (Figs 42–43)..................................... Z. maghrebense sp. nov.</p><p>6 Tibial apophysis narrow from its base (Figs 72–73, 76–77, 92–93).............................................. 7</p><p>- Tibial apophysis wide (Figs 80–81, 84–85, 88–89)........................................................... 9</p><p>7 Distal part of tibial apophysis well developed, bent in dorso-lateral direction (Figs 92–93)......... Z. wesolowskae sp. nov.</p><p>- Distal part of tibial apophysis small (Figs 72–73, 76–77)...................................................... 8</p><p>8 Tibial apophysis gradually narrowing, with recurved tip (Figs 76–77)............................. Z. ericorum sp. nov.</p><p>- Tibial apophysis terminally widened into a spoonlike tip (Figs 72–73)........................... Z. azrouense sp. nov.</p><p>9 Tibial apophysis truncate terminally, tip slightly recurved (Figs 88–89)............................... Z. ogeri sp. nov.</p><p>- Tibial apophysis rounded terminally, tip distinctly recurved (Figs 80–81, 84–85).................................. 10</p><p>10 Tibial apophysis less elongated, with wide recurved tip (Figs 78–79)..................... Z. jeanclaudeledouxi sp. nov.</p><p>- Tibial apophysis elongated, with small recurved tip (Figs 82–83)................................. Z. mostafai sp. nov.</p><p>Females (females of Z. jeanclaudeledouxi sp. nov., Z. ogeri sp. nov. and Z. wesolowskae sp. nov. unknown)</p><p>1 Epigyne flat, without large pit (Figs 5, 12, 17, 21)............................................................ 2</p><p>- Epigyne with large pit, mostly near the posterior border (Figs 44–58), rarely in the median part (Figs 94–101) or in the anterior part of the epigyne and accompanied by a chitinised knob (Figs 102–109)........................................ 3</p><p>2 Median plate rectangular (Figs 5, 17)............................................................ Z. isabellinum</p><p>- Median plate trapezoid (Figs 12, 21).............................................................. Z. pallidum 3 Epigyne with pit situated at posterior border (Figs 44–45, 49–50, 54–55)......................................... 4</p><p>- Epigyne with pit in the median or anterior part (Figs 94–95, 98–99, 102–103, 106–107)............................. 6</p><p>4 Epigynal pit with postero-median angularity (Figs 49–50)........................................... Z. maculatum</p><p>- Epigynal pit without postero-median angularity (Figs 44–45, 54–55)............................................. 5</p><p>5 Epigynal pit twice as wide as high (Figs 44–45)...................................................... Z. valentii</p><p>- Epigynal pit only slightly wider than high (Figs 54–55)..................................... Z. maghrebense sp. nov.</p><p>6 Epigyne anteriorly with triangular plate (Figs 106–107).......................................... Z. trianguliferum</p><p>- Epigyne anteriorly without triangular plate (Figs 94–95, 98–99, 102–103)........................................ 7</p><p>7 Epigynal pit positioned anteriorly (Figs 102–103)............................................. Z. mostafai sp. nov.</p><p>- Epigynal pit positioned medially (Figs 94–95, 98–99)........................................................ 8</p><p>8 Epigynal pit hardly chitinised (Figs 94–95)................................................. Z. azrouense sp. nov.</p><p>- Epigynal pit with posterior border strongly chitinised (Figs 98–99)............................... Z. ericorum sp. nov.</p></div>	https://treatment.plazi.org/id/C67D87F00C2EFFEAFF0A4BBC8ED104FD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Benhalima, Souâd;Bosmans, Robert	Benhalima, Souâd, Bosmans, Robert (2020): Revision of the genus Zodarion Walckenaer, 1833 (part IV). The species of Morocco (Araneae: Zodariidae). Zootaxa 4899 (1): 93-114, DOI: 10.11646/zootaxa.4899.1.5
C67D87F00C2FFFEAFF0A4A998F7A01A3.text	C67D87F00C2FFFEAFF0A4A998F7A01A3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zodarion isabellinum (Simon 1870)	<div><p>Zodarion isabellinum (Simon, 1870)</p><p>Figures 1–6, 15–18</p><p>Enyo isabellina Simon, 1870: 43 (Lectotype Ƌ, paralectotype ♀, designated by Bosmans 1994, deposited in MNHN AR1522: SPAIN: Granada, Cadiz—examined).</p><p>Zodarion isabellinum Denis, 1935: 19; Bosmans 1994: 136, figs 77–78, 129–130, 147.</p><p>Description and diagnosis. See Bosmans (1994).</p><p>Other material examined. MOROCCO: Rabat-Salé-Kénitra, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-6.7197223&amp;materialsCitation.latitude=34.051666" title="Search Plazi for locations around (long -6.7197223/lat 34.051666)">Forêt de Maâmora</a>, Sidi Amira, 34°03’06’’N, 06°43’11’’W, leg. S. Benhalima (in litter layer of Quercus suber forest), 15.IX.1989, 1♀, 6.XI.1989, 1♀ , and 5.XII.1989, 1♀ 4 juveniles (CSB); Tanger-Tétouan-Al <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-6.1508336&amp;materialsCitation.latitude=35.121387" title="Search Plazi for locations around (long -6.1508336/lat 35.121387)">Hoceima</a>, Larâche, Rhaba El Khalifa, 35°07’17’’N, 06°09’03’’W), leg. S. Benhalima (litter in Quercus suber forest), 7.III.1989, 1♀, 25.IX.1989 , 2ƋƋ, and 10.XI.1989, 2♀♀ (CSB) .</p><p>Distribution. Spain, and here cited for the first time in NW Morocco (Map 1).</p></div>	https://treatment.plazi.org/id/C67D87F00C2FFFEAFF0A4A998F7A01A3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Benhalima, Souâd;Bosmans, Robert	Benhalima, Souâd, Bosmans, Robert (2020): Revision of the genus Zodarion Walckenaer, 1833 (part IV). The species of Morocco (Araneae: Zodariidae). Zootaxa 4899 (1): 93-114, DOI: 10.11646/zootaxa.4899.1.5
C67D87F00C2FFFEDFF0A4F4D8FA2027C.text	C67D87F00C2FFFEDFF0A4F4D8FA2027C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zodarion pallidum Denis 1952	<div><p>Zodarion pallidum Denis, 1952</p><p>Figures 7–14, 19–22</p><p>Zodarion pallidum Denis, 1952: 61, figs 7–9 (Holotype Ƌ and paratype ♀, deposited in MNHN: MOROCCO: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-8.378889&amp;materialsCitation.latitude=30.868055" title="Search Plazi for locations around (long -8.378889/lat 30.868055)">Marrakech-Safi</a>, Tizi n’Test, 30°52’05’’N, 08°22’44’’W), 2100 m a.s.l., 1939, leg. L. Berland—examined).</p><p>Remark. A label with the name of the species was not present in the tube, but as the name of the locality fits with the name of the type locality and the number of specimens also agrees with the original description, so this material is considered to be the type material.</p><p>Diagnosis. Zodarion pallidum differs clearly from all other species of the italicum group by its smaller size and pale colour. Males are furthermore distinguished by the rounded, proximal lobe of the tibial apophysis and the small median apophysis, and females by the epigyne with a trapezoid median plate.</p><p>Description. Measurements: Male (n=7): Total length 1.6–2.0; carapace 0.72–1.02 long, 0.58–0.87 wide. Female (n=3): Total length 2.6–2.7; carapace 1.02–1.03 long, 0.68–0.70 wide. Colour (Figs 7–8): male: carapace yellowish orange, ocular quadrangle nearly black; chelicerae, sternum and legs somewhat paler than carapace; abdomen dorsally sepia, with very small dots and large postero-dorsal spot whitish, venter whitish; female: decoloured. Eyes: AM=1 (0.75); AL=0.66; PM=0.6; PL=0.7; a=0.53; b=P.27; c=1.33; d=0.2; MOQ: AW= PW; L=0.84PW.</p><p>Male palp (Figs 9–11, 19–20): tibial apophysis triangular with narrow, slightly curved tip, base with large rounded lobe; median apophysis small, triangular, with pointed distal part; embolus gradually narrowing, pointed, accompanied by a large embolic membrane.</p><p>Epigyne (Figs 12–14, 21–22): with trapezoid median plate 3 times as wide as long, with chitinised anterior margin and chitinisation spreading laterally. Spermathecae small, separated by almost 4 diameters, at their mesal side accompanied by bladder-like structures.</p><p>Other material examined. MOROCCO: Fès-Meknès, Missour, 33°02’24’’, 03°59’52’’W, 22.III–3.IV.2002 (pitfalls in steppe), 1Ƌ 1♀ (CRB); Marrakech- <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-7.6775&amp;materialsCitation.latitude=31.225" title="Search Plazi for locations around (long -7.6775/lat 31.225)">Safi</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-7.6775&amp;materialsCitation.latitude=31.225" title="Search Plazi for locations around (long -7.6775/lat 31.225)">Marrakech</a>, Oued Tensift, 31°41’33’’N, 07°59’17’’W, 370 m a.s.l., leg. J. Van Keer, 19.II.1996 (dry river bed), 1Ƌ, (CJVK); Setti Fatma, 31°13’30’’N, 07°40’39’’W, leg. V. &amp; B. Roth, I.1987, 1Ƌ (RMCA 172503); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-10.006667&amp;materialsCitation.latitude=29.6325" title="Search Plazi for locations around (long -10.006667/lat 29.6325)">Souss-Massa</a>, Gourizim, 5 km E, 29°37’57’’N, 10°00’24’’W), 15 m a.s.l., leg. R. Bosmans, 26.IV.2012 (stones in Argania steppe), 1Ƌ (CRB); Tamri Estuary, 30°42’35’’N, 09°51’16’’W, 5 m a.s.l., leg. J. Van Keer, 15–28.IV.2012 (pitfalls in Tamarisk marsh), 1♀ (CJVK); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-7.280278&amp;materialsCitation.latitude=30.724445" title="Search Plazi for locations around (long -7.280278/lat 30.724445)">Tizi</a> ‘n Bachkoum, 30°43’28’’N, 07°16’49’’W, 1650 m a.s.l., leg. R . Bosmans, 4.II.1996 (stones along rivulet), 1Ƌ (CRB); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-9.494445&amp;materialsCitation.latitude=29.845556" title="Search Plazi for locations around (long -9.494445/lat 29.845556)">Barrage</a> Youssef-Ben- Tachfine, 29°50’44’’N, 09°29’40’’W, 100 m a.s.l., leg. R . Bosmans, 27.IV.2012 (stones in steppe), 1Ƌ (CRB) .</p><p>Distribution. Central part of Morocco (Map 1), occurring from sea level to more than 2000 m.</p></div>	https://treatment.plazi.org/id/C67D87F00C2FFFEDFF0A4F4D8FA2027C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Benhalima, Souâd;Bosmans, Robert	Benhalima, Souâd, Bosmans, Robert (2020): Revision of the genus Zodarion Walckenaer, 1833 (part IV). The species of Morocco (Araneae: Zodariidae). Zootaxa 4899 (1): 93-114, DOI: 10.11646/zootaxa.4899.1.5
C67D87F00C29FFEFFF0A4E6A8FD70797.text	C67D87F00C29FFEFFF0A4E6A8FD70797.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zodarion valentii Bosmans, Loverre & Addante 1999	<div><p>Zodarion valentii Bosmans, Loverre &amp; Addante, 1999</p><p>Figures 23–24, 29–33, 44–48</p><p>Enyo algirica, Simon 1870: 98 (misidentification, not Zodarion algiricum (Lucas, 1846)); Simon, 1873: 65, 172, pl. II, fig. 12 (misidentification).</p><p>Zodarion algiricum: Denis, 1935: 67; Denis, 1937: 14 (all misidentifications).</p><p>Zodarion cfr. affine: Bosmans, 1994: 126 .</p><p>Zodarion valentii Bosmans, Loverre &amp; Addante, in Bosmans et al., 2019: 10, figs 4a–f, 5a–e (Holotype Ƌ and 3 ƋƋ 3♀♀ paratypes, deposited in MCSNB: ITALY: Puglia, Valenzano—examined).</p><p>Description and diagnosis. See Bosmans et al. (2019).</p><p>Comments. Simon (1870, 1873) and Denis (1935, 1937) misidentied this species as Z. algiricum . Bosmans (1994) mentioned females from South Spain as a species related to Z. affine (Simon, 1870) .</p><p>Previous records. ALGERIA: Tlemcen, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-1.7275001&amp;materialsCitation.latitude=35.095833" title="Search Plazi for locations around (long -1.7275001/lat 35.095833)">Marnia</a> (34°50’58’’N, 01°43’39’’W), Nemours (35°05′45″N, 01°51′28″W), Tlemcen (34°43’14’’N, 01°04’28’’W) (Denis 1937 sub Z. algiricum, misidentifications). MOROCCO: without precise locality (Simon 1870; Denis 1937, sub Z. algiricum, misidentifications). SPAIN: Cadiz, Algeciras (Bosmans 1994; sub Z. cfr . affine).</p><p>Other material examined. ALGERIA: Tlemcen, Tafna, along <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-1.4683334&amp;materialsCitation.latitude=35.297222" title="Search Plazi for locations around (long -1.4683334/lat 35.297222)">Oued Tafna</a>, 35°17’50’’N, 01°28’06’’W, 350 m a.s.l., leg. R . Bosmans, 23.IV.1984 (in Oxalis vegetation), 1Ƌ (CRB); Monts de Traras, between El Arba and <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-1.6666667&amp;materialsCitation.latitude=35.033333" title="Search Plazi for locations around (long -1.6666667/lat 35.033333)">El Arabienne</a>, 35°02’00’’N, 01°40’00’’W, leg. R . Bosmans, 24.V.1990, 2ƋƋ (CRB). MOROCCO: Without precise locality, labeled “ Zodarion algiricum obscurum E. S. AR 1549 ”, 13ƋƋ 9♀♀ (MNHN). Fès-Meknès, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-3.9977777&amp;materialsCitation.latitude=33.04" title="Search Plazi for locations around (long -3.9977777/lat 33.04)">Missour</a>, 33°02’24”N, 03°59’52”W, 605 m a.s.l. , 27.VI.2002, 1Ƌ, 16.II.2002, 1♀ (CRB), Oriental, N. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-3.2316668&amp;materialsCitation.latitude=34.048332" title="Search Plazi for locations around (long -3.2316668/lat 34.048332)">Mahiriya</a>, 34°02’54’’N, 03°13’54’’W, leg. J. Van Keer , 30.V .2018 (stones in steppe), 1♀ (CJVK); Oujda, 15 km W., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-1.9952778&amp;materialsCitation.latitude=34.710278" title="Search Plazi for locations around (long -1.9952778/lat 34.710278)">Plaine du Amgad</a>, 34°42’37’’N, 01°59’43’’W, 590 m as.s.l., leg. J. Van Keer, 24.V .2018 (stones bordering wheat fields), 1Ƌ (CJVK); Saidia, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-2.3205554&amp;materialsCitation.latitude=35.115555" title="Search Plazi for locations around (long -2.3205554/lat 35.115555)">Mouloudja National Park</a>, 35°06’56’’N, 02°19’14’’W, leg. R. Bosmans, 25.V–2.VI .2018 (pitfalls in coastal dunes), 1Ƌ 1♀ (CRB); Rabat-Salé-Kénitra, Forêt de Maâmora, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-6.590278&amp;materialsCitation.latitude=34.044724" title="Search Plazi for locations around (long -6.590278/lat 34.044724)">Sidi Allal El Bahraoui</a>, 34°02’41’’N, 06°35’25’’W, 185 m a.s.l., leg. S. Benhalima, 3.VII.1988, 1Ƌ (ISRM) .</p><p>Distribution. Italy (Puglia and Sicily), S Spain, NE Morocco and NW Algeria (Map 2).</p></div>	https://treatment.plazi.org/id/C67D87F00C29FFEFFF0A4E6A8FD70797	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Benhalima, Souâd;Bosmans, Robert	Benhalima, Souâd, Bosmans, Robert (2020): Revision of the genus Zodarion Walckenaer, 1833 (part IV). The species of Morocco (Araneae: Zodariidae). Zootaxa 4899 (1): 93-114, DOI: 10.11646/zootaxa.4899.1.5
C67D87F00C2AFFE0FF0A4E9588B2040F.text	C67D87F00C2AFFE0FF0A4E9588B2040F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zodarion maculatum (Simon 1870)	<div><p>Zodarion maculatum (Simon, 1870)</p><p>Figures 25–26, 34–38, 49–53</p><p>Enyo maculata Simon, 1870: 146 (Lectotype ♀ and paralectotype ♀, designated by Bosmans 1994, deposited in MNHN:</p><p>ITALY: Sicily—examined).</p><p>Zodarium maculatum: Denis 1935: 67, fig. 7; Denis 1937: 14.</p><p>Zodarion variegatum Denis, 1956: 201, figs 12–14 (Holotype ♀, deposited in MNHN: MOROCCO: Casablanca, Aïn Sbaâ, leg. Gattefossé, 2.VIII.1951 —examined). syn. nov.</p><p>Zodarion maculatum: Bosmans 1994: 127, figs 39–41, 103–104.</p><p>Remarks. The tube with the type of Zodarion variegatum includes a label Zodarion fusciventris, with Denis’ handwriting, but a species with this name has never been described. Moreover, the specimen fits completely Denis’ (1956) description and the locality is the same as the type locality of Z. variegatum, so it is obviously the holotype of Z. variegatum .</p><p>Examination of the holotype female of Z. variegatum and comparison with our material of Z. maculatum shows they are the same species. Denis’ excellent fig. 13 of the epigyne of Z. variegatum shows clearly the postero-median process in the epigynal pit, as mentioned in the diagnosis below.</p><p>Diagnosis. The species cannot be mistaken for any other species of the elegans group by the long, curved and terminally pointed tibial apophysis in the male (Fig. 34), and by the shape of the rounded epigynal pit with a posteromedian triangular process in the female (Fig. 50).</p><p>Description. See Bosmans (1994), and Figs 25–26, 34–38, 49– 53 in the present paper.</p><p>Previous records. MOROCCO: Casablanca-Settat, Casablanca, Ain-Sbaâ, 33°37’45’’N, 07°29’56’’W, leg. Gattefossé, 2.VIII.1951, 6Ƌ, 1♀ (Denis 1956; MNHN, holotype of Z. variegatum); Marrakech-Safi, Marrakech, Oued Tensift, 31°41’33’’N, 07°59’17’’W, IV.1930, 1♀ (Caporiaco 1932); Tanger-Tétouan-Al Houceima, Tanger, 35°37’00’’N, 05°47’00’’W, E. leg. Simon, 1868, 2♀♀ (Simon 1870; MNHN).</p><p>Other material examined. None.</p><p>Distribution. NW Morocco, Algeria, S Spain, S Italy and Sicily (Map 2).</p></div>	https://treatment.plazi.org/id/C67D87F00C2AFFE0FF0A4E9588B2040F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Benhalima, Souâd;Bosmans, Robert	Benhalima, Souâd, Bosmans, Robert (2020): Revision of the genus Zodarion Walckenaer, 1833 (part IV). The species of Morocco (Araneae: Zodariidae). Zootaxa 4899 (1): 93-114, DOI: 10.11646/zootaxa.4899.1.5
C67D87F00C25FFE0FF0A492888D103C4.text	C67D87F00C25FFE0FF0A492888D103C4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zodarion maghrebense Bosmans & Benhalima 2020	<div><p>Zodarion maghrebense Bosmans &amp; Benhalima sp. nov.</p><p>Figures 27–28, 39–43, 54–58</p><p>Etymology. The name is derived from the Maghreb region, which includes Algeria and Morocco, the two countries in which the species occurs.</p><p>Diagnosis. Males are recognised by the elongated tibial apophysis, terminally with anterior and posterior blunt tooth of equal length, females by the epigyne with deep postero-median incision.</p><p>Description. Measurements: male (n=4): Total length 2.8–3.8; carapace 1.34–1.96 long, 1.04–1.38 wide. Female (n=4): total length 4.0–5.0; carapace 1.86–2.48 long, 1.25–1.58 wide. Colour (Figs 27–28): carapace dark blackish brown, posterior margin partly yellowish brown; legs of male with black femora, contrasting with pale yellowish patellae, tibiae brown, metarsi and tarsi yellowish brown; femora of females only with distal third darkened; abdomen dark sepia, venter and postero-median dorsal spot whitish. Eyes (holotype): AM=1 (0.1); AL=0.6; PM=0.5; PL=0.65; a=0.6; b=0.3; c=2; d=0.25; MOQ: AW=0.8PW; L=0.8PW.</p><p>Male palp (Figs 39–43): tibial apophysis twice as long as wide, proximal part straight, distal part curving upwards in an angle of 60°, tip split into 2 blunt teeth of about equal length, one directed anteriorly, the other posteriorly; tegular base protruding in posterior direction; median apophysis U-shaped, distal part truncate; embolus with 2 teeth, ventral tooth rounded, dorsal tooth truncate.</p><p>Epigyne (Figs 54–55): with large, oval, transverse pit; twice as wide as high, postero-laterally with large, paired pockets; with deep postero-median incision in antero-ventral view, as shown in Fig. 57 Vulva (Figs 56–58): spermathecae small, separated by almost 5 diameters.</p><p>Type material. Holotype Ƌ and 2ƋƋ paratypes from ALGERIA: SE Tlemcen, Forêt d’Hafir, 34°46’56”N, 01°30’50’’W), 1350 m a.s.l., leg. R. Bosmans, 6. V.1984 (stones in mixed Quercus ilex and Q. suber forest) (RBINS).</p><p>Other material examined. ALGERIA: Tlemcen, S Tlemcen, Col de Zarifète, 34°50’21’’N, 01°22’37’’W, 1150 m a.s.l., leg. R. Bosmans, 24.IV.1984 (stones in Quercus ilex forest), 3ƋƋ (CRB); S Tlemcen, forêt de Tal Terny, 34°47’00’’N, 01°22’00’’W, 1300 m a.s.l., leg. R. Bosmans, 6–24. V.1990 (pitfalls in Quercus ilex forest), 1Ƌ (CRB). MOROCCO: Béni Mellal-Khénifra, between Azilal and Béni Mellal, 47 km from Azilal, 32°13’33’’N, 06°32’59’’W, leg. R. Bosmans, 19. VI.1984 (in herbs), 1♀ (CRB); Kasba Tadla E, SW Aït Rouadi, 32°36’03’’N, 06°09’14’’W, 500 m a.s.l., leg. J. Van Keer, 17.IV.2013, 1♀ (CJVK); Casablanca-Settat, Mechra-Benabbou, 32°41’26’’N, 07°49’14’’W, 400 m a.s.l., leg. R. Bosmans &amp; J. Van Keer, 9.II.1996 (recent mixed forest plantation in steppe), 1Ƌ 3♀♀ (CJVK, CRB); Marrakech-Safi, Ben Guerir 10 km N, 32°14’32’N, 07°56’46’’W, 500 m a.s.l., leg. J. Van Keer, 9.II.1996 (Ampelosma grassland), 2♀♀ (CJVK); Chichaoua 10 km S, 31°15’49’’N, 08°50’38’’W, 400 m a.s.l., leg. J. Van Keer, 9.II.1996 (stones in steppe), 1♀ (CJVK); Oriental, Debdou S, Aïn Tafrant, 33°57’59’’N, 03°02’14’’W, 1550 m a.s.l., leg. J. Van Keer, 31. V.2018 (stones in mixed Pinus-Thuya forest), 1Ƌ (CJVK); Debdou S, 33°57’35’’N, 03°02’57’’W, 1550 m a.s.l., leg. J. Van Keer, 31. V.2018 (litter in Quercus ilex forest), 1♀ (CJVK); Tanger-Tétouan-Al Houceima, Tétouan, 10 km E Chefchaouen, 35°10’02’’N, 05°16’17’’W, 500 m a.s.l., leg. R. Bosmans, 15. V.1984 (litter in Quercus suber forest), 2♀♀ (CRB); 4 km E Bab Bered, 34°59’40’’N, 04°50’43’’W, 1400 m a.s.l., leg. R. Bosmans, 15. V.1984 (stones in mixed Quercus ilex and Q. suber forest), 1♀ (CRB).</p><p>Distribution. North and Central Morocco, NW Algeria (Map 2).</p></div>	https://treatment.plazi.org/id/C67D87F00C25FFE0FF0A492888D103C4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Benhalima, Souâd;Bosmans, Robert	Benhalima, Souâd, Bosmans, Robert (2020): Revision of the genus Zodarion Walckenaer, 1833 (part IV). The species of Morocco (Araneae: Zodariidae). Zootaxa 4899 (1): 93-114, DOI: 10.11646/zootaxa.4899.1.5
C67D87F00C25FFE3FF0A4F24881906B7.text	C67D87F00C25FFE3FF0A4F24881906B7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zodarion azrouense Bosmans & Benhalima 2020	<div><p>Zodarion azrouense Bosmans &amp; Benhalima sp. nov.</p><p>Figures 59–60, 70–73, 94–97</p><p>Etymology. The species is named after the beautiful city of Azrou in the Middle Atlas.</p><p>Diagnosis. Males differ from all other species of this group by the short, relatively thicker tip of the embolus, much longer and more curved in the other species. The distal part of the tibial apophysis is curved upwards and swollen, spoon-like, simply rounded in other species of the group. Females are recognized by the presence of a poorly chitinised antero-median epigynal pit, which is differently shaped in other species.</p><p>Description. Measurements (n=1): Male: total length 3.5; carapace 1.22 long, 0.88 wide. Female (n=2). Total length 3.5–4.7; prosoma 1.85–2.48 long, 1.30–1.54 wide. Colour (Figs 59–60): carapace dark violet brown, fovea and radiating striae darkened; legs: coxae white, contrasting with dark brown to black femora, femora II–IV whith basal part white; tibia brown, with pale lateral stripes; metatarsi and tarsi yellowish white; abdomen black with postero-dorsal, lateral and ventral white spots. Eyes: AM=1 (0.12); AL=0.89; PM=1.12; PL=0.83; a=0.61; b=d=0.22; c=1.61; MOQ: AW=0.81PW; L=0.88PW.</p><p>Male palp (Figs 70–73): tibial apophysis elongated, 3 times as long as wide, proximal part 4 times as long as distal part, slightly curved, distally widening and curved in dorso-lateral direction; tip rounded anteriorly, posteriorly bluntly pointed; median apophysis with wide proximal part and small, curved distal part; tip of embolus subterminally bent in anterior direction.</p><p>Epigyne (Figs 94–97): with large, shallow pit, occupying one third of epigyne’s width, anteriorly rounded, widening posteriorly, posterior margin unsclerotized; spermathecae very small, separated by more than 5 diameters.</p><p>Type material. Holotype Ƌ and paratype 1♀ from MOROCCO: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-5.1636114&amp;materialsCitation.latitude=33.466667" title="Search Plazi for locations around (long -5.1636114/lat 33.466667)">Fès-Meknès</a>, Azrou NE, 33°28’00’’N, 05°09’49’’W, 1400 m a.s.l., leg. R. Bosmans &amp; J. Van Keer, 18.IV.2012 (litter and stones in Quercus ilex forest) (RBINS).</p><p>Paratype: MOROCCO: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-5.220556&amp;materialsCitation.latitude=33.438057" title="Search Plazi for locations around (long -5.220556/lat 33.438057)">Azrou</a>, 33°26’17’’N, 05°13’14’’W, 1450 m a.s.l., leg. R . Bosmans, 7.II.1996 (litter and stones in dense Quercus ilex forest), 1♀ (RBINS) .</p><p>Distribution. Only known from the type locality (Map 3).</p></div>	https://treatment.plazi.org/id/C67D87F00C25FFE3FF0A4F24881906B7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Benhalima, Souâd;Bosmans, Robert	Benhalima, Souâd, Bosmans, Robert (2020): Revision of the genus Zodarion Walckenaer, 1833 (part IV). The species of Morocco (Araneae: Zodariidae). Zootaxa 4899 (1): 93-114, DOI: 10.11646/zootaxa.4899.1.5
C67D87F00C26FFE2FF0A4BBD89B104C3.text	C67D87F00C26FFE2FF0A4BBD89B104C3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zodarion ericorum Benhalima & Bosmans 2020	<div><p>Zodarion ericorum Bosmans sp. nov.</p><p>Figures 61–62, 74–77, 98–101</p><p>Etymology. The first author dedicates this species to Eric Gobbers (Gent, Belgium) and Erika Vanden Bergh (Instituut voor Natuur- en Bosonderzoek, Belgium), two Belgian biologists, for helping him collect material during his first trip to Morocco in 1984.</p><p>Diagnosis. Males are recognized by the large basal knob in the tegulum, which is absent in other species or only poorly developed in Z. jeanclaudeledouxi sp. nov. Females are distinguished by the shape of the transverse oval pit in the epigyne, which is not the same in all other species.</p><p>Description. Measurements. Male (n=4): total length 2.4–3.8; carapace 1.30–1.57 long, 0.90–1.1 wide. Female (n=7): total length 3.8–5.5; prosoma 1.76–3.4 long, 1.25–2.49 wide. Colour (Figs 61–62): carapace dark reddish brown to black, posterior part sometimes yellowish orange of variable size; coxae pale yellowish, femora dark brown to black, basal part white; tibiae brown to pale brown, with paler stripes laterally; metatarsi yellowish brown, base often infuscated; tarsi yellowish brown; abdomen in male with scutum, dark sepia, dorsally with two triangular spots and one oval postero-median spot whitish, in some specimens triangular spots absent; venter and oblique lateral stripe whitish. Female as male, but abdomen without scutum. Eyes: AM=1 (0.13); AL=0.78; PM=PL=0.67; a=0.67; b=0.23; c=1.56; d=0.33; MOQ: AW=0.88PW; L=0.81PW.</p><p>Male palp (Figs 74–77): tibial apophysis elongated, proximal part nearly rectangular and straight, distal part curved in postero-dorsal direction, terminally rounded; retrolateral margin of cymbium with indentation; tegulum with rounded knob at its base; median apophysis with broad base, rectangular, distal part having the form a bird’s beak; tip of embolus slender, terminally curved in antero-prolateral direction.</p><p>Epigyne (Figs 98–101): with large, transverse, oval pit, somewhat more than 2 times as wide as high, posterolaterally acccompied by semi-circular pockets; spermathecae small, separated by more than 3 diameters.</p><p>Type material. Holotype Ƌ and 1Ƌ 1♀ paratypes from MOROCCO: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-6.57&amp;materialsCitation.latitude=33.864166" title="Search Plazi for locations around (long -6.57/lat 33.864166)">Beni Mellal-Khénifra</a>, Oued Zem, 33°51’51’’N, 06°34’12’’W, leg. E. Vanden Bergh, 12.V–23.VI.1984 (pitfalls in young Pinus plantation) (RBINS).</p><p>Other material examined. MOROCCO: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-7.946111&amp;materialsCitation.latitude=32.24222" title="Search Plazi for locations around (long -7.946111/lat 32.24222)">Marrakech-Safi</a>, Ben Guerir 10 km N, 32°14’32’’N, 07°56’46’’W, 500 m a.s.l., leg. R . Bosmans, 9.II.1996 (grassland with Asphodelus), 1♀ (CRB); 10 km S Chichaoua, 31°15’49’’N, 08°50’38’’W, leg. R . Bosmans, 9.II.1996 (stones in steppe), 2♀♀ (CRB); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-8.005834&amp;materialsCitation.latitude=31.693611" title="Search Plazi for locations around (long -8.005834/lat 31.693611)">Marrakech</a>, Gueliz, 31°41’37’’N, 08°00’21’’W, 445 m a.s.l., leg. R . Bosmans, 7.VII.1999 (grass field in flooded hotel garden), 1Ƌ (CRB); 3 km S Imi-’n-Tanoute, 31°09’35’’N, 08°50’38’W, 1200 m a.s.l., leg. J. Van Keer, 9.II.1996 (stones in olive groove), 1Ƌ 1♀ (CJVK); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-9.513056&amp;materialsCitation.latitude=31.5425" title="Search Plazi for locations around (long -9.513056/lat 31.5425)">Ounara</a> E, 31°32’33’’N, 09°30’47’’W, 250 m a.s.l., leg. R . Bosmans, 8.VII.1999 (stones in Arganus steppe), 1♀ (CRB); Rabat-Salé-Kénitra, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-6.590278&amp;materialsCitation.latitude=34.044724" title="Search Plazi for locations around (long -6.590278/lat 34.044724)">Forêt de Maâmora</a>, Sidi Amira, 34°03’06’’N, 06°43’11’’W, leg. S. Benhalima, 5.XII.1989 (litter in Quercus suber forest), 1Ƌ 1♀ (CSB); idem, 2 km S Sidi Allal El Bahraoui, 34°02’41’’N, 06°35’25’’W, leg. R . Bosmans, 8.II.1996, 1♀ (CRB); idem, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-8.053056&amp;materialsCitation.latitude=30.578056" title="Search Plazi for locations around (long -8.053056/lat 30.578056)">Sidi Allal El Bahraoui</a>, leg. S. Benhalima, 15.IX.1989, 1Ƌ, 2.X.1989, 1Ƌ, 6.XI.1989, 1Ƌ and 27.II.1990, 1Ƌ (CSB); Souss-Massa, between Aoulous and Taliouine, 30°34’41’’N, 08°03’11’’W, 600 m a.s.l., leg. R . Bosmans, 4.II.1996 (stones in Arganus steppe), 1♀ (CRB) .</p><p>Distribution. Central and West Morocco (Map 3).</p></div>	https://treatment.plazi.org/id/C67D87F00C26FFE2FF0A4BBD89B104C3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Benhalima, Souâd;Bosmans, Robert	Benhalima, Souâd, Bosmans, Robert (2020): Revision of the genus Zodarion Walckenaer, 1833 (part IV). The species of Morocco (Araneae: Zodariidae). Zootaxa 4899 (1): 93-114, DOI: 10.11646/zootaxa.4899.1.5
C67D87F00C27FFE2FF0A4A69881901C3.text	C67D87F00C27FFE2FF0A4A69881901C3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zodarion jeanclaudeledouxi Bosmans & Benhalima 2020	<div><p>Zodarion jeanclaudeledouxi Bosmans &amp; Benhalima sp. nov.</p><p>Figures 63, 78–81</p><p>Etymology. The species is dedicated to the late Jean-Claude Ledoux (†), a well-known French arachnologist and author of many papers on Mediterranean spiders.</p><p>Diagnosis. This species is recognized by the wide basal part of the tibial apophysis, suddenly narrowing into the distal part, a character shared with Z. mostafai sp. nov. . Both species differ from other members of this group by the larger embolar tooth, which is curved in the prolateral direction in Z. jeanclaudeledouxi sp. nov., while smaller and curved in the anterior direction in Z. mostafai sp. nov. .</p><p>Description. Measurements: Male (n=1): total length 2.6; carapace 1.39 long, 1.01 wide. Colour (Fig. 63): as in Z. ericorum sp. nov. . Eyes: AM=1 (0.1); AL=0.62; PM=0.63; PL=0.75; a=0.5; b=d=0.25; c=1.62; MOQ: AW=0.81PW; L= 0.88PW.</p><p>Male palp (Figs 78–81): tibial apophysis elongated, proximal part wide, rectangular, 3 times longer than distal part, suddenly narrowing into distal part, curved in retrolateral direction, terminaly rounded; median apophysis with wide proximal part and somewhat smaller, curved distal part; tip of embolus distinctly curved in antero-prolateral direction.</p><p>Female: Unknown.</p><p>Types. Holotype Ƌ from MOROCCO: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-5.8408337&amp;materialsCitation.latitude=32.700832" title="Search Plazi for locations around (long -5.8408337/lat 32.700832)">Béni Mellal-Khénifra</a>, Khénifra, Ouaoumana, 32°42’03’’N, 05°50’27’’W, 800 m a.s.l., leg. R. Bosmans, 17.IV.2012 (stones in wasteland) (RBINS).</p><p>Distribution. Only known from the type locality (Map 3).</p></div>	https://treatment.plazi.org/id/C67D87F00C27FFE2FF0A4A69881901C3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Benhalima, Souâd;Bosmans, Robert	Benhalima, Souâd, Bosmans, Robert (2020): Revision of the genus Zodarion Walckenaer, 1833 (part IV). The species of Morocco (Araneae: Zodariidae). Zootaxa 4899 (1): 93-114, DOI: 10.11646/zootaxa.4899.1.5
C67D87F00C27FFE5FF0A4D5488360477.text	C67D87F00C27FFE5FF0A4D5488360477.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zodarion mostafai Benhalima & Bosmans 2020	<div><p>Zodarion mostafai Benhalima &amp; Bosmans sp. nov.</p><p>Figures 64–65, 82–85, 102–105</p><p>Etymology. The species is with pleasure and great love dedicated to Mostafa Bounouar (Rabat, Morocco), husband of the first author of this paper.</p><p>Diagnosis. For males, see Z. jeanclaudeledouxi sp. nov. . Females can be distinghuished from the other species of this group by the presence and the shape of a trapezoid knob in the anterior part of the epigyne, which is absent in the other species.</p><p>Description. Measurements: Male (n=3): total length 3.5–5.6; carapace 1.86–3.00 long, 1.2–2.2 wide. Female (n=2): total length 4.2–5.3; carapace 1.95–2.30 long, 1.25–1.55 wide. Colour (Figs 64–65): as in Z. ericorum sp. nov., but in males the tibiae are completely black and the pale part of legs more yellowish, and in the female only Ti IV with proximal part white. Eyes AM=1 (0.12); AL=PL=0.89; PM=0.63; a=0.63; b=0.2; c=0.16; d=0.32; MOQ: AW=0.93PW; L=0.96PW.</p><p>Male palp (Figs 82–85): tibial apophysis elongated, proximal part relatively wide compared to other species, nearly rectangular, distal part a short knob curved in retrolateral direction; median apophysis with trapezoid proximal part and small, pointed distal part; tip of embolus short, curved in antero-prolateral direction.</p><p>Epigyne (Figs 102–105): with anteromedian trapezoid structure; posterior margin of epigyne deeply incised; trapezoid structure covering 2 median pockets; spermathecae small, separated by 5 diameters.</p><p>Type material. Holotype Ƌ and 1♀ paratypes from MOROCCO: Rabat-Salé-Kénitra, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-6.7197223&amp;materialsCitation.latitude=34.051666" title="Search Plazi for locations around (long -6.7197223/lat 34.051666)">Forêt de Maâmora</a>, Sidi Amira, 34°03’06’’N, 06°43’11’’W, leg. S. Benhalima, 6.XI.1989 (litter in Quercus suber forest) (ISRM).</p><p>Paratypes: Same data as holotype, 1Ƌ 1♀ (MNHN) .</p><p>Other material examined. MOROCCO: Rabat-Salé-Kénitra, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-6.590278&amp;materialsCitation.latitude=34.044724" title="Search Plazi for locations around (long -6.590278/lat 34.044724)">Forêt de Maâmora</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-6.590278&amp;materialsCitation.latitude=34.044724" title="Search Plazi for locations around (long -6.590278/lat 34.044724)">Sidi</a> Allal-El Bahraoui, 34°07’36”N, 06°33’22”W, leg. S. Benhalima, 12.II.1988 (under bark of Quercus suber), 1Ƌ 1♀ (CSB); Forêt de Maâmora, 2 km N Sidi Allal-El Bahraoui, 34°02’41’’N, 06°35’25’’W, leg. S. Benhalima, 3.II.1989, 1Ƌ (CSB); idem, leg. R. Bosmans, 8.II.1996 (litter in Q. suber forest), 1♀ (CRB).</p><p>Distribution. Only known from the Maâmora forest (Map 3).</p></div>	https://treatment.plazi.org/id/C67D87F00C27FFE5FF0A4D5488360477	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Benhalima, Souâd;Bosmans, Robert	Benhalima, Souâd, Bosmans, Robert (2020): Revision of the genus Zodarion Walckenaer, 1833 (part IV). The species of Morocco (Araneae: Zodariidae). Zootaxa 4899 (1): 93-114, DOI: 10.11646/zootaxa.4899.1.5
C67D87F00C20FFE5FF0A49FD8F2A016F.text	C67D87F00C20FFE5FF0A49FD8F2A016F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zodarion ogeri Bosmans & Benhalima 2020	<div><p>Zodarion ogeri Bosmans &amp; Benhalima sp. nov.</p><p>Figures 66, 86–89</p><p>Etymology. The species is dedicated to our friend Pierre Oger (Waret L’Evêque, Belgium), eminent photographer of all species treated in this paper.</p><p>Diagnosis. Males are recognized by the tibial apophysis that is relatively wide, nearly straight, terminally truncate and with a small incision, while more curved or terminally pointed in the other species. Zodarion jeanclaudeledouxi sp. nov. is closely related but differs by the longer recurved tip of the tibial apophysis.</p><p>Description. Measurements: Male (n=1): total length 4.1; carapace 3.12 long, 1.23 wide. Colour (Fig. 66): Carapace dark brown to black; femora black; other segments white to pale yellowish, tibiae laterally striked with brown; abdomen black, with 3 dorsal spots and oblique lateral white stripes. Eyes: AM=1 (0.14); AL=PL=0.77; PM=0.55; a=0.73; b=0.36; c=1.55; d=0.27; MOQ: AW=0.97PW; AW=0.72L.</p><p>Male palp (Figs 86–89): tibial apophysis elongated as in all species of this group, but proximal part nearly rectangular, more than 4 times longer than wide, terminally incised and curved in posterior direction; proximal part of median apophysis with triangular base, distal part a slender, pointed tooth; embolus slender, curved in anterior direction.</p><p>Female: Unknown.</p><p>Type material. Holotype Ƌ from MOROCCO: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-5.9316664&amp;materialsCitation.latitude=33.878056" title="Search Plazi for locations around (long -5.9316664/lat 33.878056)">Meknès</a>, Oued Beth, 33°52’41’’N, 05°55’54’’W, 500 m a.s.l., leg. R. Bosmans, 7.II.1996 (stones in Pinus plantation) (RBINS).</p><p>Distribution. Only known from the type locality in the region of Meknès (Map 3).</p></div>	https://treatment.plazi.org/id/C67D87F00C20FFE5FF0A49FD8F2A016F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Benhalima, Souâd;Bosmans, Robert	Benhalima, Souâd, Bosmans, Robert (2020): Revision of the genus Zodarion Walckenaer, 1833 (part IV). The species of Morocco (Araneae: Zodariidae). Zootaxa 4899 (1): 93-114, DOI: 10.11646/zootaxa.4899.1.5
C67D87F00C20FFE5FF0A4C8589BC0287.text	C67D87F00C20FFE5FF0A4C8589BC0287.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zodarion wesolowskae Bosmans & Benhalima 2020	<div><p>Zodarion wesolowskae Bosmans &amp; Benhalima sp. nov.</p><p>Figures 67, 90–93</p><p>Etymology. The species is dedicated to Wanda Wesołowska (Wrocław, Poland), in honour of her systematic and taxonomic work on Salticidae .</p><p>Diagnosis. Males of the species can be distinguished from all other Moroccan species of the mostafai group by the well-developed, sinuous distal part of the tibial apophysis, which is half as long as the proximal part, and much shorter in all other species of the group.</p><p>Description. Measurements: total length 3.4; carapace 1.83 long, 1.30 wide. Colour (Fig. 67): carapace brown to dark brown; femora black, base of femora III–IV white, other segments white to pale yellowish, tibiae laterally striked with brown; abdomen black, with postero-dorsal spot and oblique lateral whitish stripes. Eyes:AM=1 (0.14); AL=PL=0.64; PM=0.55; a=d=0.36; b=0.32; c=1; MOQ: AW=0.89PW; AW=0.89L.</p><p>Male palp (Figs 90–93): proximal part of tibial apophysis elongated, 4 times longer than distal part; distal part gently curved, first bent into retrolateral direction, than in antero-dorsal direction, terminally pointed; proximal part of median apophysis gradually narrowing, distal part a strong, pointed tooth; tip of embolus small, terminally curved in anterior direction.</p><p>Female: Unknown. It is possible that Z. trianguliferum is the unknown female of this species.</p><p>Type material. Holotype Ƌ from MOROCCO: Béni Mellal-Khénifra, N Khénifra, Source d’Oum Er Rbia, 33°03’04’’N, 05°24’44’’W, 1450 m a.s.l., leg. R. Bosmans, 13.V.1984 (among stones and grasses) (RBINS).</p><p>Distribution. Only known from the type locality.</p></div>	https://treatment.plazi.org/id/C67D87F00C20FFE5FF0A4C8589BC0287	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Benhalima, Souâd;Bosmans, Robert	Benhalima, Souâd, Bosmans, Robert (2020): Revision of the genus Zodarion Walckenaer, 1833 (part IV). The species of Morocco (Araneae: Zodariidae). Zootaxa 4899 (1): 93-114, DOI: 10.11646/zootaxa.4899.1.5
C67D87F00C3CFFF8FF0A4E188892072F.text	C67D87F00C3CFFF8FF0A4E188892072F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zodarion trianguliferum Denis 1952	<div><p>Zodarion trianguliferum Denis, 1952</p><p>Figures 68–69, 106–109</p><p>Zodarion trianguliferum Denis, 1952: 61, fig. 6 (Holotype ♀, deposited in MNHN: MOROCCO: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-8.377778&amp;materialsCitation.latitude=30.864445" title="Search Plazi for locations around (long -8.377778/lat 30.864445)">Marrakech-Safi</a>, Marrakech, Tizi n’Test, 30°51’52’’N, 08°22’40’’W, 2100 m a.s.l., leg. L. Berland, 1939—examined).</p><p>Remarks. At first, the type material of Z. trianguliferum was not found in the MNHN, but a tube containing one female and with a label corresponding with the name of the type locality of the species was present. The epigyne of the single specimen in the tube also corresponds with Denis’ figure and, as in similar cases, we therefore consider this to be the female holotype. Zodarion jeanclaudeledouxi sp. nov., Z. ogeri sp. nov. or Z. wesolowskae sp. nov. are possibly the unknown male of Z. trianguliferum .</p><p>Diagnosis. The species is recognized by the epigyne with a single antero-median pocket.</p><p>Description. Measurements. Female (n=2): total length 4.3–4.5; carapace 1.70–1.73 long, 1.10 wide. Colour (Figs 68–69): carapace dark reddish brown to black, with postero-median whitish area; femora dark brown to black, basal part of femora III–IV whitish; other segments white to pale yellowish, tibiae laterally striked with brown; abdomen dark sepia, dorsally with 2 triangular spots and one oval postero-median whitish spot, wide oblique lateral stripe and venter whitish. Eyes:AM=1 (0.13);AL=PL=0.76; PM=0.69; a=0.5; b=d=0.3; c=1.4; MOQ:AW=0.92PW; L=1.0PW.</p><p>Epigyne (Figs 106–109): poorly chitinised, only antero-median arch-like structure with one pocket; spermathecae small, separated by 5 diameters.</p><p>Male: Unknown.</p><p>Other material examined. MOROCCO: Béni Mellal-Khénifra, Kasba Tadla, SW Aït Roadi, 32°36’03’’N, 06°09’14’’W, 500 m a.s.l., leg. R . Bosmans, 17.IV.2012, 1♀ (CRB) .</p><p>Distribution. Known from two localities in Central Morocco (Map 3).</p></div>	https://treatment.plazi.org/id/C67D87F00C3CFFF8FF0A4E188892072F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Benhalima, Souâd;Bosmans, Robert	Benhalima, Souâd, Bosmans, Robert (2020): Revision of the genus Zodarion Walckenaer, 1833 (part IV). The species of Morocco (Araneae: Zodariidae). Zootaxa 4899 (1): 93-114, DOI: 10.11646/zootaxa.4899.1.5
C67D87F00C3DFFFBFF0A4AC58F0202D7.text	C67D87F00C3DFFFBFF0A4AC58F0202D7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zodarion abnorme Denis 1952	<div><p>Zodarion abnorme Denis, 1952</p><p>Zodarion abnorme Denis, 1952: 62, fig. 12 (Holotype ♀: MOROCCO: Fès-Meknès, Timahdit, Forêt d’Adim, 33°03’35”N, 04°58’26”W—not examined, unavailable).</p><p>Remarks. The holotype of Z. abnorme, described from Morocco, Forêt d’Adim, could not be traced in the MNHN, where Denis normally deposited his species. It is a very small species (1.8 mm), only known from the female. Denis (1952) gave the name abnorme by the “abnormal” shape of the eyes, the posterior lateral ones being “indistinct”. There are thus only 6 eyes, which is typical for the members of the genus Trygetus Simon, 1882 . The small size of the species also corresponds with the small size of the members of the genus Trygetus . Until new topotypic material is collected, we leave the species in the genus Zodarion .</p><p>Distribution. Only known from the type locality in Morocco (Map 3).</p><p>MAP 1. Distribution records of Zodarion species of the italicum group in Morocco.</p><p>MAP 2. Distribution records of Zodarion species of the elegans group in Morocco and Algeria.</p><p>MAP 3. Distribution records of Zodarion species of the jeanclaudeledouxi group in Morocco.</p></div>	https://treatment.plazi.org/id/C67D87F00C3DFFFBFF0A4AC58F0202D7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Benhalima, Souâd;Bosmans, Robert	Benhalima, Souâd, Bosmans, Robert (2020): Revision of the genus Zodarion Walckenaer, 1833 (part IV). The species of Morocco (Araneae: Zodariidae). Zootaxa 4899 (1): 93-114, DOI: 10.11646/zootaxa.4899.1.5
