taxonID	type	description	language	source
ED8009D762547C3DE26FF2F89B72FAEB.taxon	materials_examined	Material examined. HOLOTYPE (♂): USA, New Mexico, Catron County, Glenwood; Whitewater Picnic Area, 8 km east of Rt. 180, (33 ° 22 ' 22 " N, 108 ° 50 ' 50 " W), 12 July 1987, by IM Smith, IMS 870084. PARATYPES (0 ♀; 0 ♂):	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
ED8009D762547C3DE26FF2F89B72FAEB.taxon	diagnosis	Diagnosis. Torrenticola anoplopalpa are similar to other members of the Rala Group (T. boettgeri, T. dolichodactyla, T. kurtvietsi, T. keesdavidsi, T. lamellipalpis, and T. rala) by being colorless, having incomplete hind coxal margins and being distributed in the southwest. Torrenticola anoplopalpa can be differentiated from all other Rala Group by having a more elongate subcapitulum (ventral length / width ♂ = 4.16 in T. anoplopalpa, 2.04 - 3.56 in others). Additionally, T. anoplopalpa can be differentiated from all other Rala Group by femur / genu (♂ 1.94 in T. anoplopalpa, 0.98 - 1.86 in others), except T. keesdavidsi (1.84 - 1.96).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
ED8009D762547C3DE26FF2F89B72FAEB.taxon	description	Description. Female unknown. Male (Figure 20) (n = 1) (holotype only) with characters of the genus with following specifications. Dorsum - (640 long; 440 wide) ellipsoid and colorless. Anterio-medial platelets (152.5 long; 60 wide). Anterio-lateral platelets (205 long; 72.5 wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 375). Dorsal plate proportions: dorsum length / width 1.45; dorsal width / distance between Dgl- 4 1.17; anterio-medial platelet length / width 2.54; anterio-lateral platelet length / width 2.83; anterio-lateral / anterio-medial length 1.34. Gnathosoma - Subcapitulum (322.5 long (ventral); 235 long (dorsal); 77.5 tall) elongate and colorless. Rostrum (110 long; 32.5 wide). Chelicerae (275 long) with curved fangs (40 long). Subcapitular proportions: ventral length / height 4.16; rostrum length / width 3.38. Pedipalps short and stocky (especially tibiae) without extensions on femora and genua. Palpomeres: trochanter (35 long); femur (77.5 long); genu (40 long); tibia (47.5 long; 17.5 wide); tarsus (12.5 long). Palpomere proportions: femur / genu 1.94; tibia / femur 0.61; tibia length / width 2.71. Venter - (775 long; 510 wide) colorless. Gnathosomal bay (127.5 long; 70 wide). Cxgl- 4 subapical. Medial suture (70 long). Genital plates (160 long; 120 wide). Additional measurements: Cx- 1 (310 long (total); 180 long (medial )); Cx- 3 (330 wide); anterior venter (270 long). Ventral proportions: gnathosomal bay length / width 1.82; anterior venter / genital field length 1.69; anterior venter length / genital field width 2.25; anterior venter / medial suture 3.86. Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
ED8009D762547C3DE26FF2F89B72FAEB.taxon	etymology	Etymology. Specific epithet (anoplopalpa) refers to the pedipalps, which lack tubercles on the femora and genua, an uncommon condition in Torrenticola, which usually have tuberculate ventral extensions (anoplos, G. unarmed; palpus, L. hand, feeler).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
ED8009D762547C3DE26FF2F89B72FAEB.taxon	distribution	Distribution. Southwest. New Mexico (probably also Arizona) (Figure 19).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
F98FA07FC712BD0E83FAAC141E21D40E.taxon	materials_examined	Material examined. HOLOTYPE (♀): USA, North Carolina, Macon County, Rainbow Springs; beside Forest Route 67, 2.0 km south of road to Standing Indian Campground, (35 ° 3 ' 3 " N, 83 ° 31 ' 31 " W), 1 July 1990, by IM Smith, IMS 900072 PARATYPES (4 ♀; 5 ♂): Georgia, USA: 1 ♀ from White County, Helen; beside Road to Anna Ruby Falls just north of Unicoi State Park, (34 ° 44 ' 44 " N, 83 ° 43 ' 43 " W), 24 September 1992, by IM Smith, IMS 920051 * North Carolina, USA: 1 ♀ from Haywood County, Great Smokey Mountains National Park, Big Creek downstream of the bridge at picnic area, (35 ° 45 ' 45 " N, 83 ° 6 ' 6 " W), 15 September 2009, by AJ Radwell, AJR 090008 A * 1 ♀ and 2 ♂ from Haywood County, Great Smoky Mountains National Park; Cataloochee; beside Mt. Sterling Rd. near bridge 1.7 km n. of road to Campground, (35 ° 38 ' 38 " N, 83 ° 4 ' 4 " W), 6 September 2009, by IM Smith, IMS 090099 * 2 ♂ from Macon County, Rainbow Springs; beside Forest Route 67, 2.0 km south of road to Standing Indian Campground, (35 ° 3 ' 3 " N, 83 ° 31 ' 31 " W), 20 September 1991, by IM Smith, IMS 910054 * 1 ♂ (ALLOTYPE) from Macon County, Rainbow Springs; beside Forest Route 67, 2.0 km south of road to Standing Indian Campground, (35 ° 3 ' 3 " N, 83 ° 31 ' 31 " W), 1 July 1990, by IM Smith, IMS 900072 * Tennessee, USA: 1 ♀ from Blount County, Middle Prong of the Little River at Tremont, (35 ° 38 ' 38 " N, 83 ° 41 ' 41 " W), 16 September 2009, by AJ Radwell, AJR 090009	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
F98FA07FC712BD0E83FAAC141E21D40E.taxon	diagnosis	Diagnosis. Torrenticola arktonyx are similar to species with similar dorsal patterning, such as the Rusetria " 4 - Plate " group (T. dunni, T. glomerabilis, T. kittatinniana, T. pollani, T. rufoalba, and T. shubini), Elongata Group (T. elongata, T. gorti, and T. reduncarostra), Neoanomala Group (T. interiorensis and T. neoanomala), and T. erectirostra, T. robisoni, T. irapalpa, T. racupalpa, T. skvarlai, and T. bondi. Torrenticola arktonyx can be differentiated from all other Torrenticola by having distinctive longitudinal dark markings on the anterior portion of the dorsal plate that fade posteriorly.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
F98FA07FC712BD0E83FAAC141E21D40E.taxon	description	Description. Female (Figure 22) (n = 4) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (645 - 680 (670) long; 480 - 520 (510) wide) ovoid with purple coloration separated into anterior and posterior portions with faint orange medially, also with distinctive longitudinal dark markings on the anterior portion of the dorsal plate that fade posteriorly. Anterio-medial platelets (137.5 - 145 (137.5) long; 65 - 70 (65) wide). Anterio-lateral platelets (200 - 207.5 (200) long; 80 - 90 (80) wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 335 - 375 (375 )). Dorsal plate proportions: dorsum length / width 1.29 - 1.34 (1.31); dorsal width / distance between Dgl- 4 1.36 - 1.43 (1.36); anterio-medial platelet length / width 2.07 - 2.23 (2.12); anterio-lateral platelet length / width 2.31 - 2.56 (2.50); anterio-lateral / anterio-medial length 1.38 - 1.48 (1.45). Gnathosoma - Subcapitulum (345 - 362.5 (355) long (ventral); 260 - 270 (265) long (dorsal); 117.5 - 120 (117.5) tall) with purple coloration. Rostrum (135 - 135 (135) long; 45 - 47.5 (45) wide). Chelicerae (335 - 350 (350) long) with curved fangs (45 - 55 (45) long). Subcapitular proportions: ventral length / height 2.88 - 3.09 (3.02); rostrum length / width 2.84 - 3.00 (3.00). Pedipalps with dentate, flanged ventral extensions on femora and genua. Palpomeres: trochanter (40 - 42.5 (41.25) long); femur (107.5 - 111.25 (111.25) long); genu (75 - 80 (77.5) long); tibia (92.5 - 95 (95) long; 25 - 27.5 (25) wide); tarsus (25 - 25 (25) long). Palpomere proportions: femur / genu 1.38 - 1.44 (1.44); tibia / femur 0.85 - 0.86 (0.85); tibia length / width 3.36 - 3.80 (3.80). Venter - (815 - 840 (815) long; 510 - 600 (600) wide) with purple coloration. Gnathosomal bay (137.5 - 155 (137.5) long; 82.5 - 92.5 (92.5) wide). Cxgl- 4 subapical. Medial suture (50 - 60 (50) long). Genital plates (180 - 187.5 (180) long; 155 - 162.5 (155) wide). Additional measurements: Cx- 1 (300 - 310 (300) long (total); 150 - 165 (165) long (medial )); Cx- 3 (350 - 385 (385) wide); anterior venter (230 - 232.5 (230) long). Ventral proportions: gnathosomal bay length / width 1.49 - 1.82 (1.49); anterior venter / genital field length 1.24 - 1.28 (1.28); anterior venter length / genital field width 1.42 - 1.48 (1.48); anterior venter / medial suture 3.83 - 4.60 (4.60). Male (Figure 23) (n = 5) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (500 - 570 (570) long; 400 - 450 (450) wide) ovoid with purple coloration separated into anterior and posterior portions with faint orange medially, also with distinctive longitudinal dark markings on the anterior portion of the dorsal plate that fade posteriorly. Anterio-medial platelets (110 - 125 (125) long; 55 - 62.5 (62.5) wide). Anterio-lateral platelets (155 - 180 (180) long; 65 - 75 (75) wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 305 - 345 (345 )). Dorsal plate proportions: dorsum length / width 1.23 - 1.35 (1.27); dorsal width / distance between Dgl- 4 1.30 - 1.35 (1.30); anterio-medial platelet length / width 2.00 - 2.18 (2.00); anterio-lateral platelet length / width 2.21 - 2.69 (2.40); anterio-lateral / anterio-medial length 1.41 - 1.59 (1.44). Gnathosoma - Subcapitulum (275 - 300 (300) long (ventral); 210 - 235 (235) long (dorsal); 92.5 - 102.5 (102.5) tall) with purple coloration. Rostrum (110 - 125 (125) long; 35 - 40 (40) wide). Chelicerae (270 - 305 (305) long) with curved fangs (40 - 45 (40) long). Subcapitular proportions: ventral length / height 2.75 - 3.08 (2.93); rostrum length / width 3.00 - 3.20 (3.13). Pedipalps with dentate, flanged ventral extensions on femora and genua. Palpomeres: trochanter (32.5 - 37.5 (35) long); femur (85 - 97.5 (97.5) long); genu (60 - 68.75 (65) long); tibia (75 - 85 (85) long; 22.5 - 26.25 (25) wide); tarsus (21.25 - 25 (22.5) long). Palpomere proportions: femur / genu 1.42 - 1.50 (1.50); tibia / femur 0.81 - 0.91 (0.87); tibia length / width 3.10 - 3.40 (3.40). Venter - (650 - 725 (725) long; 430 - 490 (490) wide) with purple coloration. Gnathosomal bay (125 - 135 (135) long; 65 - 72.5 (72.5) wide). Cxgl- 4 subapical. Medial suture (75 - 90 (80) long). Genital plates (160 - 172.5 (172.5) long; 117.5 - 130 (130) wide). Additional measurements: Cx- 1 (250 - 290 (290) long (total); 130 - 150 (145) long (medial )); Cx- 3 (295 - 350 (350) wide); anterior venter (220 - 260 (260) long). Ventral proportions: gnathosomal bay length / width 1.85 - 2.08 (1.86); anterior venter / genital field length 1.38 - 1.55 (1.51); anterior venter length / genital field width 1.87 - 2.17 (2.00); anterior venter / medial suture 2.72 - 3.25 (3.25). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
F98FA07FC712BD0E83FAAC141E21D40E.taxon	etymology	Etymology. Specific epithet (arktonyx) refers to the distinctive longitudinal markings on the anterior dorsal plate, which resemble claw marks from a bear (arktos, G. bear; onyx, G. claw).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
F98FA07FC712BD0E83FAAC141E21D40E.taxon	distribution	Distribution. Southern Appalachians (Figure 21).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
3FA5D85712680D2788437B6402D344D7.taxon	materials_examined	Material examined. HOLOTYPE (♀): from USA, Arkansas, Montgomery County, South Fork Ouachita River, access off County Road 17 at Forest Road 903, 29 Jul 2011, by IM Smith, IMS 110040, DNA 1263. PARATYPES (4 ♀; 3 ♂): Arkansas, USA: 1 ♂ (ALLOTYPE) from Montgomery County, Ouachita River (34 ° 34 ' 53.20 " N, 93 ° 53 ' 0.16 " W), 5 Oct 2007, by AJ Radwell, & HW Robison, AJR 070300 A * 2 ♂ Montgomery County, Ouachita River (34 ° 34 ' 53.20 " N, 93 ° 53 ' 0.16 " W), 5 Oct 2007, by AJ Radwell, & HW Robison, AJR 070300 A * 3 ♀ from Montgomery County, South Fork Ouachita River, access off County Road 17 at Forest Road 903, 29 Jul 2011, by IM Smith, IMS 110040 * Missouri, USA: 1 ♀ from Crawford County, Huzzah Creek, Red Bluff campground, off Road V east of Davisville, 23 Jul 2011, by IM Smith, IMS 110029.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
3FA5D85712680D2788437B6402D344D7.taxon	diagnosis	Diagnosis. Torrenticola biscutella are similar to other members of the Rusetria " Eastern 2 - Plates " group (T. caerulea, T. delicatexa, T. feminellai, T. indistincta, T. malarkeyorum, T. microbiscutella, T. pendula, T. sellersorum, T. tysoni, T. ululata, and T. whitneyae) in having anterio-lateral platelets fused to the dorsal plate, having dorsal coloration separated into anterior and posterior portions (except T. indistincta and T. ululata), and being distributed in the east. It is one of only four Eastern 2 - Plates that have dark, bold, bluish-purple coloration (also T. pendula, T. sellersorum, and T. tysoni). Torrenticola biscutella can be differentiated from T. caerulea, T. ululata, T. indistincta, and T. feminellai by dorsal coloration and pattern. T. biscutella can be differentiated from T. tysoni by having a stockier rostrum (length / width = 2.55 - 2.83 in T. biscutella, 3.06 - 3.50 in T. tysoni). Female T. biscutella can be differentiated from female T. malarkeyorum by having a shorter subcapitulum (ventral length = 290 - 315 in T. biscutella, 317.5 - 335 in T. malarkeyorum). Male T. biscutella can be differentiated from male T. malarkeyorum by having a slightly rounder dorsum (length / width 1.37 - 1.42 in T. biscutella, 1.42 - 1.56 in T. malarkeyorum). Additionally, although T. biscutella and T. malarkeyorum have the same dorsal coloration and pattern, often the coloration is bold in T. biscutella and faint in T. malarkeyorum. Female T. biscutella can be differentiated from female T. delicatexa by having a shorter genital field (152.5 - 167.5 in T. biscutella, 175 - 198 in T. delicatexa) and male T. biscutella can be differentiated from male T. delicatexa by having a slightly rounder dorsum (length / width = 1.37 - 1.42 in T. biscutella, 1.44 - 1.56 in T. delicatexa). Female T. biscutella can be differentiated from female T. sellersorum by anterior venter / genital field length (0.82 - 0.88 in T. biscutella, 0.69 - 0.77 in T. sellersorum). Male T. biscutella can be differentiated from male T. sellersorum by having slightly stockier anterio-lateral platelets (length / width = 2.58 - 2.74 in T. biscutella, 2.76 - 3.00 in T. sellersorum). T. biscutella can be differentiated from T. pendula by having a stockier gnathosomal bay (1.55 - 1.85 in T. biscutella, 2.42 - 2.9 in T. pendula); more elongate tibiae (3.11 - 3.45 in T. biscutella, 2.78 - 3.05 in T. biscutella); and by dorsal pattern. T. biscutella can be differentiated from T. microbiscutella by having a more ovoid dorsum (length / width = 1.33 - 1.42 in T. biscutella, 1.63 - 1.75 in T. microbiscutella) and by anterior venter / genital field width (♀ = 0.84 - 0.91 in T. biscutella, 1.25 - 1.33 in T. microbiscutella; ♂ = 1.68 - 1.80 in T. biscutella and 1.95 - 2.29 in T. microbiscutella). T. biscutella can be differentiated from T. whitneyae by having more elongate pedipalpal tibiae (3.11 - 3.45 in T. biscutella, 2.42 - 2.95 in T. whitneyae) and by anterior venter / genital field length (♀ = 0.82 - 0.88 in T. biscutella, 0.59 - 0.75 in T. whitneyae; ♂ = 1.55 - 1.76 in T. biscutella and 1.37 - 1.43 in T. whitneyae).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
3FA5D85712680D2788437B6402D344D7.taxon	description	Description. Female (Figure 25) (n = 4) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (560 - 630 (560) long; 420 - 455 (420) wide) ovoid with bluish-purple coloration separated into anterior and posterior portions, and bold or faint orange medially. Anterio-medial platelets (122.5 - 135 (122.5) long; 40 - 45 (40) wide). Anterio-lateral platelets (140 - 170 (140) long; 62.5 - 75 (62.5) wide) fused to dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 300 - 330 (300 )). Dorsal plate proportions: dorsum length / width 1.33 - 1.38 (1.33); dorsal width / distance between Dgl- 4 1.38 - 1.40 (1.40); anterio-medial platelet length / width 3.00 - 3.31 (3.06); anterio-lateral platelet length / width 2.24 - 2.48 (2.24); anterio-lateral / anterio-medial length 1.14 - 1.28 (1.14). Gnathosoma - Subcapitulum (290 - 315 (290) long (ventral); 207 - 240 (208) long (dorsal); 137.5 - 155 (137.5) tall) colorless. Rostrum (110 - 125 (110) long; 42.5 - 47.5 (42.5) wide). Chelicerae (286 - 335 (286) long) with curved fangs (55 - 70 (56) long). Subcapitular proportions: ventral length / height 2.02 - 2.11 (2.11); rostrum length / width 2.56 - 2.67 (2.59). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (43.75 - 50 (43.75) long); femur (107.5 - 122.5 (107.5) long); genu (65 - 72.5 (65) long); tibia (80 - 86.25 (80) long; 23.75 - 25 (23.75) wide); tarsus (20 - 20 (20) long). Palpomere proportions: femur / genu 1.59 - 1.74 (1.65); tibia / femur 0.69 - 0.74 (0.74); tibia length / width 3.35 - 3.45 (3.37). Venter - (660 - 740 (660) long; 488 - 544 (489) wide) with faint bluish-purple coloration. Gnathosomal bay (151.25 - 172.5 (151.25) long; 97.5 - 100 (100) wide). Cxgl- 4 subapical. Medial suture absent. Genital plates (152.5 - 167.5 (152.5) long; 142.5 - 160 (142.5) wide). Additional measurements: Cx- 1 (274 - 309 (275) long (total); 118 - 135 (121) long (medial )); Cx- 3 (319 - 392 (319) wide); anterior venter (130 - 147.5 (130) long). Ventral proportions: gnathosomal bay length / width 1.55 - 1.73 (1.55); anterior venter / genital field length 0.82 - 0.88 (0.85); anterior venter length / genital field width 0.84 - 0.91 (0.91). Male (Figure 26) (n = 3) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (430 - 445 (440) long; 310 - 315 (310) wide) ovoid with bluish-purple coloration separated into anterior and posterior portions, and bold or faint orange medially. Anterio-medial platelets (97.5 - 97.5 (97.5) long; 33.75 - 36.25 (35) wide). Anterio-lateral platelets (122.5 - 130 (130) long; 45 - 50 (47.5) wide) fused to dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 230 - 242.5 (230 )). Dorsal plate proportions: dorsum length / width 1.37 - 1.42 (1.42); dorsal width / distance between Dgl- 4 1.30 - 1.35 (1.35); anterio-medial platelet length / width 2.69 - 2.89 (2.79); anterio-lateral platelet length / width 2.58 - 2.74 (2.74); anterio-lateral / anterio-medial length 1.26 - 1.33 (1.33). Gnathosoma - Subcapitulum (230 - 235 (235) long (ventral); 175 - 177.5 (177) long (dorsal); 20 - 20 (20) tall) colorless. Rostrum (85 - 92.5 (92.5) long; 30 - 36.25 (36.25) wide). Chelicerae (225 - 241 (241) long) with curved fangs (45 - 50 (46) long). Subcapitular proportions: ventral length / height 2.29 - 2.47 (2.29); rostrum length / width 2.55 - 2.83 (2.55). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (23.75 - 37.5 (37.5) long); femur (85 - 90 (90) long); genu (52.5 - 55 (55) long); tibia (68.75 - 72.5 (72.5) long; 21.25 - 22.5 (22.5) wide); tarsus (17.5 - 20 (20) long). Palpomere proportions: femur / genu 1.62 - 1.64 (1.64); tibia / femur 0.81 - 0.82 (0.81); tibia length / width 3.11 - 3.24 (3.22). Venter - (510 - 525 (525) long; 335 - 380 (336) wide) with faint bluish-purple coloration. Gnathosomal bay (115 - 122.5 (122.5) long; 65 - 67.5 (67.5) wide). Cxgl- 4 subapical. Medial suture (60 - 65 (65) long). Genital plates (102.5 - 110 (102.5) long; 100 - 100 (100) wide). Additional measurements: Cx- 1 (215 - 226 (226) long (total); 99 - 110 (100) long (medial )); Cx- 3 (252 - 275 (252) wide); anterior venter (167.5 - 180 (180) long). Ventral proportions: gnathosomal bay length / width 1.70 - 1.85 (1.81); anterior venter / genital field length 1.55 - 1.76 (1.76); anterior venter length / genital field width 1.68 - 1.80 (1.80); anterior venter / medial suture 2.68 - 2.83 (2.77). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
3FA5D85712680D2788437B6402D344D7.taxon	etymology	Etymology. Specific epithet (biscutella) refers to the appearance of only two anterio-dorsal platelets due to the fusion of lateral platelets with the dorsal shield (bi-, L. two; scutella, L. little plate).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
3FA5D85712680D2788437B6402D344D7.taxon	distribution	Distribution. Interior Highlands (both Ozarks and Ouachitas), likely endemic (Figure 24).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
D4803272947FB50C0C695F65BBFD724D.taxon	materials_examined	Material examined. PARATYPES (0 ♀; 2 ♂): Ohio, USA: 2 ♂ from Ottawa County, Middle Bass Island, rubble beach, 29 June 1954, by R Crowell.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
D4803272947FB50C0C695F65BBFD724D.taxon	diagnosis	Diagnosis. Torrenticola bittikoferae are similar to other members of the Tricolor Group (T. cardia, T. dimorpha, T. hoosieri, T. kringi, T. larvata, T. mohawk, T. olliei, T. pearsoni, T. sierrensis, T. tricolor, T. trimaculata, and T. unimaculata) in having a short, conical rostrum. Torrenticola bittikoferae can be differentiated from most other Tricolor Complex (except T. hoosieri, T. pearsoni, and T. dimorpha) by being colorless, whereas most other members have bold patterning. T. bittikoferae can be differentiated from T. hoosieri by having ventral extensions on the pedipalp femora and genua (lacking in T. hoosieri) and having stockier pedipalp tibiae (length / width = 2.7 - 2.8 in T. bittikoferae, 3.6 - 4.4 in T. hoosieri). T. bittikoferae can be differentiated from T. pearsoni by having Dgl- 4 further from the dorsal edge (dorsal width / distance between Dgl- 4 = 1.6 - 1.7 in T. bittikoferae, 1.2 - 1.3 in T. pearsoni); stockier pedipalp tibiae (length / width = 2.7 - 2.8 in T. bittikoferae, 3.0 - 3.3 in T. pearsoni); and a more elongate rostrum (length / width = 1.8 - 1.9 in T. bittikoferae, 2.0 - 2.4 in T. pearsoni). Torrenticola bittikoferae can be differentiated from T. dimorpha by having an unmodified dorsal plate (T. dimorpha has a dorsal plate medial extension covering nearly half the length of the anterio-medial platelets) and by males having unmodified pedipalps (male T. dimorpha have large, highly modified pedipalps which are expanded vertically and laterally).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
D4803272947FB50C0C695F65BBFD724D.taxon	description	Re-description. Male (Figure 28) (n = 2) with characters of the genus with following specifications. Dorsum - (620 - 670 long; 500 - 530 wide) circular and colorless. Anterio-medial platelets (132.5 - 137.5 long; 70 - 70 wide). Anterio-lateral platelets (192.5 - 202.5 long; 90 - 92.5 wide) free from dorsal plate. Dgl- 4 approaching midway between muscle scars and dorsum edge (distance between Dgl- 4 305 - 330). Dorsal plate proportions: dorsum length / width 1.24 - 1.26; dorsal width / distance between Dgl- 4 1.61 - 1.64; anterio-medial platelet length / width 1.89 - 1.96; anterio-lateral platelet length / width 2.14 - 2.19; anterio-lateral / anterio-medial length 1.40 - 1.53. Gnathosoma - Subcapitulum (265 long (ventral); 202.5 long (dorsal); 125 tall) colorless. Rostrum (95 - 100 long; 52.5 - 52.5 wide). Chelicerae (260 long) with curved fangs (50 long) short and conical. Subcapitular proportions: ventral length / height 2.12; rostrum length / width 1.81 - 1.90. Pedipalps with tuberculate ventral extensions with dentate tip on femora and tuberculate ventral extensions on genua. Palpomeres: trochanter (42.5 - 42.5 long); femur (101.25 - 107.5 long); genu (72.5 - 75 long); tibia (87.5 - 90 long; 32.5 - 32.5 wide); tarsus (25 - 35 long). Palpomere proportions: femur / genu 1.40 - 1.43; tibia / femur 0.81 - 0.89; tibia length / width 2.69 - 2.77. Venter - (790 - 800 long; 610 - 680 wide) colorless. Gnathosomal bay (122.5 - 125 long; 87.5 - 100 wide). Medial suture (102.5 - 117.5 long). Genital plates (137.5 - 142.5 long; 115 - 115 wide). Additional measurements: Cx- 1 (270 - 280 long (total); 152.5 - 152.5 long (medial )); Cx- 3 (405 - 410 wide); anterior venter (270 - 287.5 long). Ventral proportions: gnathosomal bay length / width 1.23 - 1.43; anterior venter / genital field length 1.96 - 2.02; anterior venter length / genital field width 2.35 - 2.50; anterior venter / medial suture 102.5 - 117.5. Female type specimens unavailable for present study. Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
D4803272947FB50C0C695F65BBFD724D.taxon	etymology	Etymology. Robert Crowell (1960) named the specific epithet (bittikoferae) after Lelia Bittikofer, his high school biology teacher.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
D4803272947FB50C0C695F65BBFD724D.taxon	distribution	Distribution. Known only from type locality: Lake Erie, Ohio (Figure 27).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
8DFEBA068F1C2285E3B7B3D27257049B.taxon	materials_examined	Material examined (1 ♀; 4 ♂). New Mexico, USA: 1 ♀ and 1 ♂ from Catron County, Glenwood; Whitewater Picnic Area 8 km east of Rt. 180, (33 ° 22 ' 22 " N, 108 ° 50 ' 50 " W), 12 July 1987, by IM Smith, IMS 870084 * 3 ♂ from Catron County, beside Rt. 15, 65 km north of Rt. 180, Silver City, (33 ° 12 ' 12 " N, 108 ° 13 ' 13 " W), 10 July 1987, by IM Smith, IMS 870081 A	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
8DFEBA068F1C2285E3B7B3D27257049B.taxon	diagnosis	Diagnosis. Torrenticola boettgeri are similar to other members of the Rala Group (T. anoplopalpa, T. dolichodactyla, T. keesdavidsi, T. kurtvietsi, T. lamellipalpis, and T. rala) by being colorless, having incomplete hind coxal margins and being distributed in the southwest. T. boettgeri can be differentiated from all other Rala Group by having a more elongate dorsum (length / width ♀ = 1.74 - 1.82 in T. boettgeri, 1.21 - 1.60 in others) and a stockier subcapitulum (ventral length / width = 1.96 in T. boettgeri, 2.06 - 3.52 in others; ♂ = 2.04 - 2.07 in T. boettgeri, 2.14 - 4.16 in others).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
8DFEBA068F1C2285E3B7B3D27257049B.taxon	description	Re-description. Female (Figure 30) (n = 3: one specimen examined from New Mexico; measurements from two additional specimens are included based upon those listed in Goldschmidt (2007) for K. O. Viets's (1977 a) specimen from Guatemala and Cramer's (1992) specimen from Mexico) with characters of the genus with following specifications. Note: measurements below are from the above three combined sources; those in parentheses are from the Guatemalan holotype (Viets 1977 a) as listed in Goldschmidt (2007). Dorsum - (668 - 800 (675) long; 367 - 440 (440) wide) ovoid, elongate, and colorless. Anterio-medial platelets (103 - 135 (126) long; 38 - 52.5 wide). Anterio-lateral platelets (179 - 200 (199) long; 47 - 70 wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 370). Dorsal plate proportions: dorsum length / width 1.53 - 1.82 (1.82); dorsal width / distance between Dgl- 4 1.19; anterio-medial platelet length / width 2.57 - 2.71; anterio-lateral platelet length / width 2.86 - 3.81; anterio-lateral / anterio-medial length 1.48 - 1.74 (1.58). Gnathosoma - Subcapitulum (245 - 280 (245) long (ventral); 185 long (dorsal); 142.5 tall) colorless. Rostrum (70 - 87.5 (70) long; 40 wide). Chelicerae with curved fangs (65 long). Subcapitular proportions: ventral length / height 1.96; rostrum length / width 2.19. Pedipalps short and stocky (especially tibiae) without extensions on femora and genua. Palpomeres: trochanter (32.5 long); femur (76.25 long); genu (65 long); tibia (32.5 long; 17.5 wide); tarsus (15 long). Palpomere proportions: femur / genu 1.17; tibia / femur 0.43; tibia length / width 1.86. Venter - (668 - 955 (845) long; 452 - 570 (500) wide) colorless. Gnathosomal bay (164 - 200 (164) long; 45 - 50 (47) wide). Cxgl- 4 apical. Medial suture (65 - 73 (73) long). Genital plates (170 - 188 (170) long; 148 - 160 (148) wide). Additional measurements: Cx- 1 (340 long (total); 133 - 135 (133) long (medial )); Cx- 3 (350 wide); anterior venter (212.5 long). Ventral proportions: gnathosomal bay length / width 3.49 - 4.44 (3.49); anterior venter / genital field length 1.15; anterior venter length / genital field width 1.33; anterior venter / medial suture 3.27. Male (Figure 31) (n = 4: new specimens from New Mexico) Dorsum - (710 - 780 long; 400 - 430 wide) ovoid, elongate, and colorless. Anterio-medial platelets (122.5 - 140 long; 45 - 50 wide). Anterio-lateral platelets (175 - 192.5 long; 55 - 62.5 wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 335 - 365). Dorsal plate proportions: dorsum length / width 1.74 - 1.81; dorsal width / distance between Dgl- 4 1.16 - 1.19; anterio-medial platelet length / width 2.68 - 2.95; anterio-lateral platelet length / width 2.96 - 3.27 (3.08); anterio-lateral / anterio-medial length 1.25 - 1.47. Gnathosoma - Subcapitulum (255 - 285 long (ventral); 165 - 187.5 long (dorsal); 125 - 137.5 tall) colorless. Rostrum (80 - 92.5 long; 35 - 40 wide). Chelicerae (285 - 310 long) with curved fangs (62.5 - 65 long). Subcapitular proportions: ventral length / height 2.04 - 2.07; rostrum length / width 2.13 - 2.36. Pedipalps short and stocky (especially tibiae) without extensions on femora and genua. Palpomeres: trochanter (30 - 31.25 long); femur (58.75 - 72.5 long); genu (60 - 62.5 long); tibia (32.5 - 37.5 long; 17.5 - 17.5 wide); tarsus (15 - 15 long). Palpomere proportions: femur / genu 0.98 - 1.16; tibia / femur 0.48 - 0.55; tibia length / width 1.86 - 2.14. Venter - (800 - 960 long; 485 - 510 wide) colorless. Gnathosomal bay (192.5 - 205 long; 40 - 50 wide). Cxgl- 4 apical. Medial suture (105 - 125 long). Genital plates (160 - 177.5 long; 120 - 125 wide). Additional measurements: Cx- 1 (305 - 330 long (total); 115 - 130 long (medial )); Cx- 3 (320 - 325 wide); anterior venter (235 - 262.5 long). Ventral proportions: gnathosomal bay length / width 3.85 - 5.00; anterior venter / genital field length 1.38 - 1.59; anterior venter length / genital field width 1.96 - 2.13; anterior venter / medial suture 2.00 - 2.24. Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
8DFEBA068F1C2285E3B7B3D27257049B.taxon	etymology	Etymology. Viets (1977 a) named the specific epithet (boettgeri) in honor of Klaus Boettger of the University of Kiel, Germany, who collected the type specimen in Rio Chilax near Coban, Guatemala.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
8DFEBA068F1C2285E3B7B3D27257049B.taxon	distribution	Distribution. New Mexico (probably also Arizona) and extending southward into Mexico and Guatemala (Figure 29).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
FF73350BBA9140CDF4D1F89BAFB217B6.taxon	materials_examined	Material examined. HOLOTYPE (♀): from USA, North Carolina, Haywood County, Great Smokey Mountains National Park, Cataloochee (35 ° 37 ' 31 " N, 83 ° 6 ' 46 " W), 20 Sep 2010, by IM Smith, IMS 100148, DNA 1431. PARATYPES (4 ♀; 5 ♂): North Carolina, USA: 1 ♂ (ALLOTYPE) from Haywood County, Great Smoky Mountains National Park; Cataloochee; beside road to Nellie 0.2 km west of Pretty Hollow Gap Trailhead, (35 ° 37 ' 37 " N, 83 ° 6 ' 6 " W), 16 June 2006, by IM Smith, IMS 060012 * 1 ♀ from Haywood County, Great Smoky Mountains National Park; Cataloochee; beside road to Nellie 0.2 km west of Pretty Hollow Gap Trailhead, (35 ° 37 ' 37 " N, 83 ° 6 ' 6 " W), 16 June 2006, by IM Smith, IMS 060012 * 1 ♀ and 1 ♂ from Haywood County, Great Smoky Mountains National Park; Cataloochee; beside road to Nellie 0.8 km west of Pretty Hollow Gap Trailhead, (35 ° 38 ' 38 " N, 83 ° 4 ' 4 " W), 20 June 2006, by IM Smith, IMS 060022 * 1 ♂ from Haywood County, Great Smoky Mountains National Park; Cataloochee; beside road to Nellie 0.4 km west of Pretty Hollow Gap Trailhead, (35 ° 37 ' 37 " N, 83 ° 6 ' 6 " W), 3 October 2007, by IM Smith, IMS 0701002 * ♀ and 2 ♂ from Haywood County, Great Smoky Mountains National Park; Waterville; Big Creek Picnic Area, (35 ° 45 ' 45 " N, 83 ° 6 ' 6 " W), 26 September 2007, by IM Smith, IMS 070089	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
FF73350BBA9140CDF4D1F89BAFB217B6.taxon	diagnosis	Diagnosis. Torrenticola bondi are similar to species with similar dorsal patterning, such as the Rusetria " 4 - Plate " group (T. dunni, T. glomerabilis, T. kittatinniana, T. pollani, T. rufoalba, and T. shubini), Elongata Group (T. elongata, T. gorti, and T. reduncarostra), Neoanomala Group (T. interiorensis and T. neoanomala), and T. erectirostra, T. robisoni, T. irapalpa, T. racupalpa, T. skvarlai, and T. arktonyx. They can be differentiated from Rusetria 4 - Plates and T. skvarlai by having distinct hind coxal margins. T. bondi can be differentiated from T. erectirostra and T. robisoni by having a straight, anteriorly-directed rostrum (upturned in others). T. bondi can be differentiated from T. arktonyx by having an unmodified dorsal plate (T. arktonyx has distinctive longitudinal dark markings on the anterior portion of the dorsal plate that fade posteriorly). T. bondi can be differentiated from T. racupalpa and T. irapalpa by having a more ovoid dorsum (dorsum length / width ♀ = 1.35 - 1.41 in T. bondi, 1.17 - 1.28 in others; ♂ = 1.32 - 1.45 in T. bondi, 1.20 - 1.30 in others) and shorter pedipalpal tibiae (♀ = 90 - 98 in T. bondi, 100 - 125 in others, ♂ = 77 - 83 in T. bondi, 87 - 110 in others). T. bondi can be differentiated from the Elongata Group by having a less elongate dorsum (length / width ♀ = 1.35 - 1.41 in T. bondi, 1.45 - 2.08 in Elongata Group; ♂ = 1.32 - 1.45 in T. bondi, 1.51 - 1.70 in Elongata Group) and having a stockier rostrum (length / width = 2.76 - 3.13 in T. bondi, 3.24 - 4.00 in Elongata Group). T. bondi can be differentiated from the Neoanomala Group by having a shorter medial suture (♀ = 10 - 15 in T. bondi, 22 - 40 in Neoanomala Group; ♂ = 55 - 70 in T. bondi, 75 - 108 in Neoanomala Group) and anterior venter / genital field width (♀ = 1.15 - 1.25 in T. bondi, 1.31 - 1.45 in Neoanomala Group; ♂ = 1.95 - 2.05 in T. bondi, 2.09 - 2.66 in Neoanomala Group).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
FF73350BBA9140CDF4D1F89BAFB217B6.taxon	description	Description. Female (Figure 33) (n = 5) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (620 - 670 (620) long; 440 - 490 (440) wide) ovoid with bluish-purple coloration separated into anterior and posterior portions and faint orange medially. Anterio-medial platelets (132.5 - 147.5 (132.5) long; 55 - 62.5 (55) wide). Anterio-lateral platelets (192.5 - 200 (192.5) long; 67.5 - 77.5 (67.5) wide) free from dorsal plate. Dgl- 4 approaching midway between muscle scars and dorsum edge (distance between Dgl- 4 270 - 290 (270 )). Dorsal plate proportions: dorsum length / width 1.35 - 1.41 (1.41); dorsal width / distance between Dgl- 4 1.63 - 1.72 (1.63); anterio-medial platelet length / width 2.20 - 2.52 (2.41); anterio-lateral platelet length / width 2.58 - 2.85 (2.85); anterio-lateral / anterio-medial length 1.34 - 1.41 (1.45). Gnathosoma - Subcapitulum (355 - 380 (355) long (ventral); 255 - 295 (255) long (dorsal); 135 - 150 (135) tall) faint bluish purple coloration. Rostrum (145 - 155 (145) long; 50 - 55 (52.5) wide). Chelicerae (354 - 385 (354) long) with curved fangs 52 - 75 (52) long). Subcapitular proportions: ventral length / height 2.50 - 2.63 (2.63); rostrum length / width 2.76 - 3.00 (2.76). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter 47.5 - 52.5 (47.5) long); femur (120 - 140 (120) long); genu (67.5 - 77.5 (67.5) long); tibia (90 - 97.5 (92.5) long; 22.5 - 25 (22.5) wide); tarsus (17.5 - 20 (17.5) long). Palpomere proportions: femur / genu 1.74 - 1.83 (1.78); tibia / femur 0.67 - 0.77 (0.77); tibia length / width 3.90 - 4.11 (4.11). Venter - (760 - 840 (760) long; 509 - 580 (509) wide) with faint bluish-purple coloration. Gnathosomal bay (170 - 190 (170) long; 90 - 102.5 (102.5) wide). Cxgl- 4 subapical. Medial suture (10 - 15 (10) long). Genital plates (175 - 180 (175) long; 150 - 160 (150) wide). Additional measurements: Cx- 1 (306 - 330 (306) long (total); 108 - 155 (108) long (medial )); Cx- 3 (365 - 410 (382) wide); anterior venter (177.5 - 187.5 (187.5) long). Ventral proportions: gnathosomal bay length / width 1.66 - 2.11 (1.66); anterior venter / genital field length 0.99 - 1.07 (1.07); anterior venter length / genital field width 1.15 - 1.25 (1.25); anterior venter / medial suture 12 - 18.75 (18.75). Male (Figure 34) (n = 5) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (515 - 550 (550) long; 380 - 410 (380) wide) ovoid with bluish-purple coloration separated into anterior and posterior portions and faint orange medially. Anterio-medial platelets (112.5 - 122.5 (122.5) long; 47.5 - 57.5 (57.5) wide). Anterio-lateral platelets (162.5 - 190 (180) long; 55 - 67.5 (67.5) wide) free from dorsal plate. Dgl- 4 approaching midway between muscle scars and dorsum edge (distance between Dgl- 4 235 - 260 (250 )). Dorsal plate proportions: dorsum length / width 1.32 - 1.45 (1.45); dorsal width / distance between Dgl- 4 1.48 - 1.62 (1.52); anterio-medial platelet length / width 2.13 - 2.47 (2.13); anterio-lateral platelet length / width 2.65 - 3.17 (2.67); anterio-lateral / anterio-medial length 1.44 - 1.57 (1.47). Gnathosoma - Subcapitulum (295 - 305 (305) long (ventral); 220 - 230 (225) long (dorsal); 107.5 - 112.5 (112.5) tall) faint bluish purple coloration. Rostrum (120 - 125 (125) long; 40 - 40 (40) wide). Chelicerae (285 - 295 (290) long) with curved fangs 50 - 55 (55) long). Subcapitular proportions: ventral length / height 2.68 - 2.79 (2.71); rostrum length / width 3.00 - 3.13 (3.13). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter 40 - 42.5 (42.5) long); femur (102.5 - 107.5 (105) long); genu (60 - 62.5 (60) long); tibia (77.5 - 82.5 (77.5) long; 21.25 - 23.75 (22.5) wide); tarsus (17.5 - 20 (20) long). Palpomere proportions: femur / genu 1.68 - 1.75 (1.75); tibia / femur 0.74 - 0.80 (0.74); tibia length / width 3.44 - 3.88 (3.44). Venter - (660 - 670 (665) long; 420 - 490 (460) wide) with faint bluish-purple coloration. Gnathosomal bay (140 - 150 (142.5) long; 75 - 90 (80) wide). Cxgl- 4 subapical. Medial suture (55 - 70 (60) long). Genital plates (132.5 - 142.5 (140) long; 107.5 - 112.5 (110) wide). Additional measurements: Cx- 1 (270 - 290 (285) long (total); 125 - 140 (135) long (medial )); Cx- 3 (310 - 340 (335) wide); anterior venter (215 - 220 (220) long). Ventral proportions: gnathosomal bay length / width 1.61 - 1.93 (1.78); anterior venter / genital field length 1.54 - 1.66 (1.57); anterior venter length / genital field width 1.95 - 2.05 (2.00); anterior venter / medial suture 3.07 - 3.91 (3.67). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
FF73350BBA9140CDF4D1F89BAFB217B6.taxon	etymology	Etymology. Specific epithet (bondi) named in honor of arachnologist Jason Bond, for his research on species delimitation and integrative taxonomy, which has been an inspiration to JRF, and for his thoughtful career advice, which was greatly appreciated.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
FF73350BBA9140CDF4D1F89BAFB217B6.taxon	distribution	Distribution. Known only from Great Smoky Mountains National Park, Haywood County, North Carolina (Figure 32).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
B175A1CEB0AE884A79EBB2745545037E.taxon	materials_examined	Material examined. HOLOTYPE (♀): from USA, Tennessee, Wayne County, beside service road parallel to Natchez Trace Parkway (35 ° 15 ' 9 " N, 87 ° 37 ' 53 " W), 27 Sep 2010, by IM Smith, IMS 100160, DNA 1882. PARATYPES (5 ♀; 3 ♂): Tennessee, USA: 2 ♀ from Wayne County, beside Natchez Trace Parkway at Lower Glenrock Branch Picnic Area, (35 ° 15 ' 15 " N, 87 ° 37 ' 37 " W), 2 June 1992, by IM Smith, IMS 920021 * 1 ♂ (ALLOTYPE) from Wayne County, Lower Glenrock Picnic Area off Natchez Trace Parkway, (35 ° 15 ' 15 " N, 87 ° 37 ' 37 " W), 5 October 2005, by IM Smith, IMS 050119 A * 1 ♀ and 2 ♂ from Wayne County, Lower Glenrock Picnic Area off Natchez Trace Parkway, (35 ° 15 ' 15 " N, 87 ° 37 ' 37 " W), 5 October 2005, by IM Smith, IMS 050119 A * 2 ♀ from Wayne County, beside service road parallel to Natchez Trace Parkway (35 ° 15 ' 9 " N, 87 ° 37 ' 53 " W), 27 Sep 2010, by IM Smith, IMS 100160.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
B175A1CEB0AE884A79EBB2745545037E.taxon	diagnosis	Diagnosis. Torrenticola caerulea are similar to other members of the Rusetria " Eastern 2 - Plates " group (T. biscutella, T. delicatexa, T. feminellai, T. indistincta, T. malarkeyorum, T. microbiscutella, T. pendula, T. sellersorum, T. tysoni, T. ululata, and T. whitneyae) in having anterio-lateral platelets fused to the dorsal plate, having dorsal coloration separated into anterior and posterior portions (except T. ululata and T. indistincta), and being distributed in the east. T. caerulea can be differentiated from all other Eastern 2 - Plates by having faint blue coloration. T. caerulea can be differentiated from T. ululata and T. feminellai by dorsal coloration and pattern. T. caerulea can be differentiated from T. tysoni by having a less elongate rostrum (length / width = 2.67 - 2.96 in T. caerulea, 3.06 - 3.50 in T. tysoni). T. caerulea can be differentiated from T. pendula by having a more elongate gnathosomal bay (1.40 - 2.09 in T. caerulea, 2.42 - 2.90 in T. pendula) and more elongate pedipalpal tibiae (3.11 - 3.83 in T. caerulea, 2.78 - 3.05 in T. pendula). T. caerulea can be differentiated from T. microbiscutella by having a less elongate dorsum (length / width = 1.32 - 1.56 in T. caerulea, 1.63 - 1.75 in T. microbiscutella). T. caerulea can be differentiated from T. whitneyae by having more elongate pedipalpal tibiae (length / width = 3.11 - 3.83 in T. caerulea, 2.42 - 2.95 in T. whitneyae) and by anterior venter / genital field width (♀ = 0.9 - 1.04 in T. caerulea, 0.67 - 0.80 in T. whitneyae; ♂ = 1.71 - 1.83 in T. caerulea, 1.5 - 1.54 in T. whitneyae). Female T. caerulea can be differentiated from female T. biscutella by having slightly more elongate pedipalpal tibae (length / width = 3.5 - 3.83 in T. caerulea, 3.35 - 3.45 in T. biscutella). Male T. caerulea can be differentiated from male T. biscutella by anterior venter / medial suture (2.4 - 2.57 in T. caerulea, 2.68 - 2.83 in T. biscutella). Female T. caerulea can be differentiated from female T. malarkeyorum, T. sellersorum, T. delicatexa, and T. indistincta by having a thinner genital field (120 - 145 in T. caerulea, 150 - 205 in others). Male T. caerulea can be differentiated from male T. indistincta by having a smaller dorsum (length = 405 - 460 in T. caerulea, 480 - 645 in T. indistincta; width = 260 - 305 in T. caerulea, 315 - 470 in T. indistincta). Body proportions of male T. caerulea do not differ from male T. malarkeyorum, T. sellersorum, and T. delicatexa, but can be differentiated by dorsal coloration.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
B175A1CEB0AE884A79EBB2745545037E.taxon	description	Description. Female (Figure 36) (n = 5) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (550 - 600 (580) long; 400 - 440 (440) wide) ovoid with faint blue coloration anteriorly and posteriorly, broadly connected medially. Anterio-medial platelets (125 - 130 (128.75) long; 40 - 45 (42.5) wide). Anterio-lateral platelets (145 - 157.5 (145) long; 57.5 - 65 (62.5) wide) fused to dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 275 - 320 (320 )). Dorsal plate proportions: dorsum length / width 1.32 - 1.48 (1.32); dorsal width / distance between Dgl- 4 1.25 - 1.51 (1.38); anterio-medial platelet length / width 2.78 - 3.25 (3.03); anterio-lateral platelet length / width 2.32 - 2.48 (2.32); anterio-lateral / anterio-medial length 1.12 - 1.21 (1.13). Gnathosoma - Subcapitulum (310 - 330 (320) long (ventral); 225 - 245 (240) long (dorsal); 137.5 - 156.25 (155) tall) colorless. Rostrum (120 - 135 (125) long; 45 - 47.5 (45) wide). Chelicerae (310 - 330 (320) long) with curved fangs (62 - 70 (65) long). Subcapitular proportions: ventral length / height 2.06 - 2.25 (2.06); rostrum length / width 2.67 - 2.84 (2.78). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (40 - 48.75 (48.75) long); femur (115 - 120 (120) long); genu (67.5 - 72.5 (72.5) long); tibia (85 - 87.5 (87.5) long; 22.5 - 25 (25) wide); tarsus (17.5 - 20 (20) long). Palpomere proportions: femur / genu 1.66 - 1.78 (1.66); tibia / femur 0.71 - 0.76 (0.73); tibia length / width 3.50 - 3.83 (3.50). Venter - (620 - 750 (660) long; 400 - 580 (660) wide) with faint blue coloration. Gnathosomal bay (145 - 175 (145) long; 82.5 - 116.25 (95) wide). Cxgl- 4 subapical. Medial suture absent. Genital plates (155 - 165 (155) long; 140 - 145 (145) wide). Additional measurements: Cx- 1 (257.5 - 305 (257.5) long (total); 125 - 135 (125) long (medial )); Cx- 3 (310 - 380 (380) wide); anterior venter (130 - 150 (137.5) long). Ventral proportions: gnathosomal bay length / width 1.40 - 1.94 (1.53); anterior venter / genital field length 0.82 - 0.97 (0.89); anterior venter length / genital field width 0.90 - 1.04 (0.95). Male (Figure 36) (n = 3) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (405 - 460 (460) long; 260 - 305 (300) wide) ovoid with faint blue coloration anteriorly and posteriorly, broadly connected medially. Anterio-medial platelets (95 - 106.25 (106.25) long; 35 - 37.5 (35) wide). Anterio-lateral platelets (117.5 - 130 (125) long; 40 - 47.5 (45) wide) fused to dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 190 - 240 (230 )). Dorsal plate proportions: dorsum length / width 1.51 - 1.56 (1.53); dorsal width / distance between Dgl- 4 1.27 - 1.37 (1.30); anterio-medial platelet length / width 2.53 - 3.04 (3.04); anterio-lateral platelet length / width 2.74 - 2.94 (2.78); anterio-lateral / anterio-medial length 1.18 - 1.24 (1.18). Gnathosoma - Subcapitulum (220 - 242.5 (227.5) long (ventral); 165 - 185 (165) long (dorsal); 87.5 - 97.5 (97.5) tall) colorless. Rostrum (88.75 - 92.5 (92.5) long; 30 - 35 (33.75) wide). Chelicerae (225 - 232.5 (227.5) long) with curved fangs (42.5 - 50 (45) long). Subcapitular proportions: ventral length / height 2.33 - 2.51 (2.33); rostrum length / width 2.64 - 2.96 (2.74). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (37.5 - 37.5 (37.5) long); femur (83.75 - 87.5 (87.5) long); genu (52.5 - 55 (55) long); tibia (67.5 - 70 (67.5) long; 20 - 22.5 (20) wide); tarsus (15 - 20 (17.5) long). Palpomere proportions: femur / genu 1.59 - 1.60 (1.59); tibia / femur 0.77 - 0.81 (0.77); tibia length / width 3.11 - 3.38 (3.38). Venter - (485 - 550 (550) long; 305 - 340 (330) wide) with faint blue coloration. Gnathosomal bay (120 - 127.5 (125) long; 57.5 - 65 (60) wide). Cxgl- 4 subapical. Medial suture (65 - 75 (75 )). Genital plates (110 - 117.5 (117.5) long; 90 - 105 (102.5) wide). Additional measurements: Cx- 1 (220 - 220 (220) long (total); 85 - 95 (95) long (medial )); Cx- 3 (240 - 260 (260) wide); anterior venter (165 - 180 (180) long). Ventral proportions: gnathosomal bay length / width 1.96 - 2.09 (2.08); anterior venter / genital field length 1.50 - 1.57 (1.53); anterior venter length / genital field width 1.71 - 1.83 (1.76). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
B175A1CEB0AE884A79EBB2745545037E.taxon	etymology	Etymology. Specific epithet (caerulea) refers to the overall and diagnostic bluish appearance of this species (caeruleus, L. sky-blue).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
B175A1CEB0AE884A79EBB2745545037E.taxon	distribution	Distribution. Known only from Wayne County, Tennessee (Figure 35). T. caerulea has been collected so rarely that comments about distribution are speculative, but given our collection efforts, it is reasonable to speculate that this species is at least restricted to the southern Appalachians.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
ACE7D255F725CEB1D7B76DEE5C70EC02.taxon	materials_examined	Material examined. HOLOTYPE (♀): from USA, New York, Greene County, beside Rt. 23 A, 9.6 km west of Rt. 296 (Hunter), (42 ° 14 ' 14 " N, 74 ° 19 ' 19 " W), 22 June 1990, by IM Smith, IMS 900052 PARATYPES (9 ♀; 8 ♂): New York, USA: 3 ♀ and 3 ♂ from Cayuga County, beside Route 38 A at Niles, (42 ° 50 ' 50 " N, 76 ° 25 ' 25 " W), 22 July 1990, by IM Smith, IMS 900113 A * (allotype) 1 ♂ from Greene, beside Rt. 23 A, 9.6 km west of Rt. 296 (Hunter), (42 ° 14 ' 14 " N, 74 ° 19 ' 19 " W), 22 June 1990, by IM Smith, IMS 900052 * 2 ♀ and 1 ♂ from Greene County, beside Rt. 23 A, 9.6 km west of Rt. 296 (Hunter), (42 ° 14 ' 14 " N, 74 ° 19 ' 19 " W), 22 June 1990, by IM Smith, IMS 900052 * 1 ♀ from Schuyler County, beside Town Line Road off Route 228, 0.6 km south of Perry City, (42 ° 29 ' 29 " N, 76 ° 42 ' 42 " W), 21 July 1990, by IM Smith, IMS 900112 A * Ohio, USA: 1 ♀ and 1 ♂ from Hocking County, beside road near Ash Cave, (39 ° 24 ' 24 " N, 82 ° 33 ' 33 " W), 5 May 1993, by IM Smith, DR Cook, IMS 930001 A * Virginia, USA: 2 ♀ and 2 ♂ from Bath County, beside Rt. 687, 2.4 km south of Bacova, (38 ° 2 ' 2 " N, 79 ° 51 ' 51 " W), 15 July 1990, by IM Smith, IMS 900097	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
ACE7D255F725CEB1D7B76DEE5C70EC02.taxon	diagnosis	Diagnosis. Torrenticola cardia are similar to other members of the Tricolor Complex (T. bittikoferae, T. dimorpha, T. hoosieri, T. kringi, T. larvata, T. mohawk, T. pearsoni, T. olliei, T. sierrensis, T. tricolor, T. trimaculata, and T. unimaculata,) in having a short, conical rostrum. T. cardia can be differentiated from most Torrenticola, including other members of the Tricolor Complex, by having a distinct dorsal pattern. T. cardia are most similar to other members of the Tricolor Complex that have bold patterning (T. larvata, T. tricolor, T. unimaculata, T. trimaculata, T. kringi, and T. mohawk). T. cardia can be differentiated from T. tricolor, T. trimaculata, T. kringi, and T. mohawk by having a more ovoid dorsum (length / width ♀ = 1.39 - 1.47 in T. cardia, 1.15 - 1.35 in others; ♂ = 1.43 - 1.54 in T. cardia, 1.19 - 1.39 in others). T. cardia can be differentiated from T. unimaculata by dorsal pattern. Female T. cardia can be differentiated from female T. larvata by having a shorter subcapitulum (♀ = 265 - 273 in T. cardia, 275 - 288 in T. larvata) and a larger genital field (length ♀ = 190 - 198 in T. cardia, 182 - 188 in T. larvata; width ♀ = 160 - 175 in T. cardia, 145 - 153 in T. larvata). Male T. cardia can be differentiated from male T. larvata by having less elongate pedipalpal tibiae (length / width ♂ = 2.82 - 3.05 in T. cardia, 3.10 - 3.20 in T. larvata) and a larger dorsum (length ♂ = 625 - 670 in T. cardia, 550 - 610 in T. larvata; width ♂ = 405 - 445 in T. cardia, 350 - 400 in T. larvata).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
ACE7D255F725CEB1D7B76DEE5C70EC02.taxon	description	Description. Female (Figure 39) (n = 5) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (710 - 785 (750) long; 510 - 555 (510) wide) ellipsoid with reddish-purple, bluish-purple or bright orange spot medially extending in a strip anteriorly often to the anterior-medial platelets. Anterio-medial platelets (130 - 140 (140) long; 70 - 75 (75) wide). Anterio-lateral platelets (195 - 207.5 (197.5) long; 70 - 80 (80) wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 390 - 410 (395 )). Dorsal plate proportions: dorsum length / width 1.39 - 1.47 (1.47); dorsal width / distance between Dgl- 4 1.29 - 1.35 (1.29); anterio-medial platelet length / width 1.86 - 2.00 (1.87); anterio-lateral platelet length / width X = 2.47 - 2.93 (2.47); anterio-lateral / anterio-medial length 1.39 - 1.50 (1.41). Gnathosoma - Subcapitulum (265 - 272.5 (272.5) long (ventral); 180 - 190 (190) long (dorsal); 125 - 132.5 (130) tall) with reddish-purple or bluish purple coloration. Rostrum (90 - 100 (100) long; 40 - 42.5 (40) wide) short and conical. Chelicerae (250 - 265 (260) long) with curved fangs (60 - 60 (60) long). Subcapitular proportions: ventral length / height 2.04 - 2.12 (2.10); rostrum length / width 2.24 - 2.50 (2.50). Pedipalps with stocky, tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (38.75 - 42.5 41.25) long); femur (97.5 - 107.5 (102.5) long); genu (65 - 70 (67.5) long); tibia (80 - 91.25 (80) long; 26.25 - 27.5 (27.5) wide); tarsus (22.5 - 25 (22.5) long). Palpomere proportions: femur / genu 1.50 - 1.56 (1.52); tibia / femur 0.78 - 0.87 (0.78); tibia length / width 2.91 - 3.32 (2.91). Venter - (830 - 925 (850) long; 565 - 600 (565) wide) with reddish-purple or bluish-purple coloration restricted to the edges of the gnathosomal bay, coxal plates, and genital plates. Gnathosomal bay (150 - 162.5 (155) long; 72.5 - 80 (72.5) wide). Cxgl- 4 subapical. Medial suture (25 - 50 (50) long). Genital plates (190 - 197.5 (190) long; 160 - 175 (162.5) wide). Additional measurements: Cx- 1 (290 - 310 (290) long (total); 135 - 165 (135) long (medial )); Cx- 3 (355 - 380 (355) wide); anterior venter (185 - 212.5 (205) long). Ventral proportions: gnathosomal bay length / width 1.94 - 2.17 (2.14); anterior venter / genital field length 0.94 - 1.09 (1.08); anterior venter length / genital field width 1.12 - 1.26 (1.26); anterior venter / medial suture 4.10 - 7.60 (4.10). Male (Figure 40) (n = 5) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (625 - 670 (670) long; 405 - 445 (445) wide) ellipsoid with reddish-purple, bluish-purple or bright orange spot medially extending in a strip anteriorly often to the anterior-medial platelets. Anterio-medial platelets (120 - 130 (125) long; 65 - 72.5 (70) wide). Anterio-lateral platelets (170 - 197.5 (197.5) long; 75 - 80 (75) wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 325 - 380 (380 )). Dorsal plate proportions: dorsum length / width 1.43 - 1.54 (1.51); dorsal width / distance between Dgl- 4 1.17 - 1.28 (1.17); anterio-medial platelet length / width 1.72 - 2.00 (1.79); anterio-lateral platelet length / width 2.25 - 2.63 (2.63); anterio-lateral / anterio-medial length 1.35 - 1.58 (1.58). Gnathosoma - Subcapitulum (227.5 - 250 (250) long (ventral); 165 - 180 (170) long (dorsal); 90 - 112.5 (110) tall) with reddish-purple or bluish purple coloration. Rostrum (77.5 - 92.5 (90) long; 32.5 - 37.5 (37.5) wide) short and conical. Chelicerae (212.5 - 230 (225) long) with curved fangs (50 - 55 (50) long). Subcapitular proportions: ventral length / height 2.17 - 2.56 (2.27); rostrum length / width 2.27 - 2.47 (2.40). Pedipalps with stocky, tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (36.25 - 38.75 (36.25) long); femur (87.5 - 97.5 (92.5) long); genu (60 - 67.5 (62.5) long); tibia (73.75 - 82.5 (77.5) long; 25 - 27.5 (27.5) wide); tarsus (20 - 25 (22.5) long). Palpomere proportions: femur / genu 1.38 - 1.48 (1.48); tibia / femur 0.82 - 0.90 (0.84); tibia length / width 2.82 - 3.05 (2.82). Venter - (710 - 780 (780) long; 460 - 495 (490) wide) with reddish-purple or bluish-purple coloration restricted to the edges of the gnathosomal bay, coxal plates, and genital plates. Gnathosomal bay (125 - 137.5 (135) long; 67.5 - 72.5 (70) wide). Cxgl- 4 subapical. Medial suture (107.5 - 130 (125) long). Genital plates (150 - 175 (175) long; 97.5 - 115 (105) wide). Additional measurements: Cx- 1 (265 - 280 (280) long (total); 140 - 150 (145) long (medial )); Cx- 3 (325 - 355 (350) wide); anterior venter (265 - 290 (290) long). Ventral proportions: gnathosomal bay length / width 1.85 - 1.93 (1.93); anterior venter / genital field length 1.66 - 1.81 (1.66); anterior venter length / genital field width 2.52 - 2.77 (2.76); anterior venter / medial suture 2.21 - 2.51 (2.32). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
ACE7D255F725CEB1D7B76DEE5C70EC02.taxon	etymology	Etymology. Specific epithet (cardia) refers to the dorsal coloration, which is either a heart-shaped or resembles a bleeding heart (kardia, G. heart).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
ACE7D255F725CEB1D7B76DEE5C70EC02.taxon	distribution	Distribution. Appalachians (Figure 38).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
E94C7E0838592BCF2EFE6F4D10A5B0E5.taxon	materials_examined	Material examined. HOLOTYPE (♀): from USA, Oregon, Lane County, Gate Creek (44 ° 8 ' 48 " N, 122 ° 34 ' 20 " W), 11 Aug 2013, by JC O'Neill, & WA Nelson, JNOW 13 - 0811 - 001. PARATYPES (20 ♀; 26 ♂): California, USA: 1 ♀ from Alpine County, Markleeville Creek (38 ° 41 ' 39 " N, 119 ° 46 ' 41 " W), 30 Aug 2013, by JR Fisher, JRF 13 - 0830 - 001 * 1 ♀ from Del Norte County, Six Rivers National Forest, Middle Fork Smith River (41 ° 51 ' 20 " N, 123 ° 53 ' 10 " W), 15 Aug 2013, by JR Fisher, JRF 13 - 0815 - 002 * 5 ♂ from El Dorado County, El Dorado National Forest, Taylor Creek (38 ° 55 ' 59 " N, 120 ° 3 ' 21 " W), 27 Aug 2013, by JR Fisher, JRF 13 - 0827 - 003 * 1 ♀ from Mendocino County, Cottaneva Creek, beside Route 1, 21.8 kilometers southwest of Route 101, 5 Aug 1987, by IM Smith, IMS 870129 A * 2 ♂ from Trinity County, small cascading trickle beside Route 36, 5.2 kilometers west of Forest Glen Station, 6 Aug 1987, by IM Smith, IMS 870132 * 1 ♂ from Trinity County, South Fork of Trinity River, beside Route 36 at Forest Glen campground, 6 Aug 1987, by IM Smith, IMS 870131 * Oregon, USA: 2 ♂ from Coos County, Gaylord, Coquille Myrtle Grove State Park, Coquille River, 2 Jul 1983, by IM Smith, IMS 830014 * 3 ♀ and 3 ♂ from Coos County, Siskiyou National Forest, Road 33 between Powers & Agness, Coal Creek, 2 Jul 1983, by IM Smith, IMS 830015 * 2 ♂ from Coos County, Siskiyou National Forest, Road 33 between Powers & Agness, Daphne Grove campground, Coquille River, 2 Jul 1983, by IM Smith, IMS 830016 * 1 ♂ from Coos County, Siskiyou National Forest, Road 33 between Powers & Agness, Daphne Grove campground, 2 Jul 1983, by IM Smith, IMS 830017 * 1 ♀ and 1 ♂ from Curry County, Port Orford, Butler Bar campground, Elk River, 25 Jun 1976, by IM Smith, IMS 760162 * 1 ♀ from Curry County, Port Orford, Butler Bar campground, Elk River, 25 - 26 Jun 1976, by IM Smith, IMS 760163 * 2 ♂ from Curry County, Port Orford, small spring run beside road from Humbug Mountain State Park to McGribble campground, 25 Jun 1976, by IM Smith, IMS 760161 * 3 ♂ from Curry County, Port Orford, Humbug Mountain State Park Picnic Area, beside Route 1, Brush Creek, 1 Jul 1983, by IM Smith, IMS 830012 * 1 ♀ from Curry County, Port Orford, Humbug Mountain State Park Picnic Area, beside Route 1, Brush Creek, 3 Jul 1983, by IM Smith, IMS 830020 A * 2 ♀ from Curry County, Quosatana Creek (42 ° 29 ' 21 " N, 124 ° 14 ' 2 " W), 14 Aug 2013, JR Fisher, JRF 13 - 0814 - 003 * 1 ♀ from Curry County, Rogue River National Forest, Elk River (42 ° 42 ' 46 " N, 124 ° 18 ' 41 " W), 13 Aug 2013, by JR Fisher, JRF 13 - 0813 - 003 * 4 ♀ and 3 ♂ from Curry County, Sixes, Sixes River, beside road at mouth of Edson Creek, 4 Jul 1983, by IM Smith, IMS 830021 A * 1 ♂ (ALLOTYPE) from Lane County, Gate Creek (44 ° 8 ' 48 " N, 122 ° 34 ' 20 " W), 11 Aug 2013, by JC O'Neill, & WA Nelson, JNOW 13 - 0811 - 001 * 4 ♀ from Lane County, Gate Creek (44 ° 8 ' 48 " N, 122 ° 34 ' 20 " W), 11 Aug 2013, by JC O'Neill, & WA Nelson, JNOW 13 - 0811 - 001.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
E94C7E0838592BCF2EFE6F4D10A5B0E5.taxon	diagnosis	Diagnosis. Torrenticola copipalpa are similar to members of the Miniforma group (T. manni, T. miniforma, T. oliveri, T. pacificensis, T. pinocchio, and T. rockyensis) in having short, stocky pedipalps (except T. oliveri and T. pinocchio); similar pedipalpal extensions (unique to members of this group); and being among the smallest Torrenticola in the west (dorsum 500 - 625 long) (except T. oliveri). T. copipalpa are best differentiated from all other Miniforma group (except T. pinocchio) by having broad, flat pedipalp femoral tubercles (conical / tuberculate in all others). T. copipalpa can be differentiated from T. pinocchio by having a less elongate rostrum (length / width = 2.5 - 3.0 in T. copipalpa, 4.5 - 4.9 in T. pinocchio) and less elongate pedipalpal tibiae (length / width = 2.4 - 2.9 in T. copipalpa, 3.1 - 3.5 in T. pinocchio).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
E94C7E0838592BCF2EFE6F4D10A5B0E5.taxon	description	Description. Female (Figure 42) (n = 6) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (555 - 605 (605) long; 380 - 420 (420) wide) ovoid and usually colorless, occasionally with faint purple coloration without distinct pattern. Anterio-medial platelets (115 - 127.5 (127.5) long; 47.5 - 57.5 (57.5) wide). Anterio-lateral platelets (162.5 - 180 (180) long; 53.75 - 62.5 (62.5) wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 290 - 335 (335 )). Dorsal plate proportions: dorsum length / width 1.39 - 1.47 (1.44); dorsal width / distance between Dgl- 4 1.25 - 1.32 (1.25); anterio-medial platelet length / width 2.13 - 2.42 (2.22); anterio-lateral platelet length / width 2.87 - 3.04 (2.88); anterio-lateral / anterio-medial length 1.35 - 1.46 (1.41). Gnathosoma - Subcapitulum (312.5 - 337.5 (337.5) long (ventral); 228 - 257.5 (257.5) long (dorsal); 117.5 - 130 (125) tall) colorless. Rostrum (122.5 - 135 (130) long; 42.5 - 47.5 (45) wide). Chelicerae (313 - 341 (340) long) with curved fangs (50 - 59 (55) long). Subcapitular proportions: ventral length / height 2.58 - 2.70 (2.70); rostrum length / width 2.72 - 2.94 (2.89). Pedipalps short and stocky (especially tibiae) with broad, dentate, and anteriorly-directed ventral extensions on femora and dentate, flanged ventral extensions on genua. Palpomeres: trochanter (30 - 35 (35) long); femur (90 - 100 (97.5) long); genu (62.5 - 67.5 (67.5) long); tibia (52.5 - 58.75 (57.5) long; 20 - 22.5 (21.25) wide); tarsus (15 - 17.5 (15) long). Palpomere proportions: femur / genu 1.44 - 1.51 (1.41); tibia / femur 0.55 - 0.61 (0.59); tibia length / width 2.59 - 2.71 (2.71). Venter - (690 - 760 (760) long; 438 - 520 (520) wide) colorless. Gnathosomal bay (136.25 - 152.5 (152.5) long; 75 - 82.5 (80) wide). Cxgl- 4 subapical. Medial suture (40 - 45 (45) long). Genital plates (152.5 - 165 (165) long; 137.5 - 160 (160) wide). Additional measurements: Cx- 1 (261 - 290 (290) long (total); 108 - 155 (155) long (medial )); Cx- 3 (268 - 320 (320) wide); anterior venter (187.5 - 210 (210) long). Ventral proportions: gnathosomal bay length / width 1.65 - 1.97 (1.91); anterior venter / genital field length 1.19 - 1.28 (1.27); anterior venter length / genital field width 1.31 - 1.40 (1.31); anterior venter / medial suture 4.53 - 4.88 (4.67). Male (Figure 43) (n = 6) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (500 - 570 (520) long; 355 - 390 (360) wide) ovoid and usually colorless, occasionally with faint purple coloration without distinct pattern. Anterio-medial platelets (105 - 117.5 (105) long; 45 - 56.25 (45) wide). Anterio-lateral platelets (155 - 167.5 (155) long; 50 - 60 (50) wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 285 - 315 (290 )). Dorsal plate proportions: dorsum length / width 1.39 - 1.54 (1.44); dorsal width / distance between Dgl- 4 1.18 - 1.37 (1.24); anterio-medial platelet length / width 2.09 - 2.37 (2.33); anterio-lateral platelet length / width 2.79 - 3.10 (3.10); anterio-lateral / anterio-medial length 1.32 - 1.48 (1.48). Gnathosoma - Subcapitulum (280 - 307.5 (295) long (ventral); 215 - 253 (220) long (dorsal); 105 - 115 (105) tall) colorless. Rostrum (110 - 120 (115) long; 40 - 46.25 (40) wide). Chelicerae (280 - 328 (295) long) with curved fangs (45 - 65 (55) long). Subcapitular proportions: ventral length / height 2.67 - 2.81 (2.81); rostrum length / width 2.54 - 2.88 (2.88). Pedipalps short and stocky (especially tibiae) with broad, dentate, and anteriorly-directed ventral extensions on femora and dentate, flanged ventral extensions on genua. Palpomeres: trochanter (32.5 - 32.5 (32.5) long); femur (82.5 - 92.5 (87.5) long); genu (57.5 - 65 (60) long); tibia (52.5 - 57.5 (52.5) long; 18.75 - 21.25 (21.25) wide); tarsus (15 - 17.5 (15) long). Palpomere proportions: femur / genu 1.35 - 1.54 (1.46); tibia / femur 0.60 - 0.64 (0.60); tibia length / width 2.47 - 2.88 (2.47). Venter - (610 - 700 (670) long; 420 - 496 (440) wide) colorless. Gnathosomal bay (110 - 140 (130) long; 67.5 - 75 (75) wide). Cxgl- 4 subapical. Medial suture (77.5 - 97.5 (77.5) long). Genital plates (130 - 137.5 (130) long; 100 - 112.5 (105) wide). Additional measurements: Cx- 1 (235 - 263 (245) long (total); 115 - 130 (130) long (medial )); Cx- 3 (270 - 300 (280) wide); anterior venter (210 - 232.5 (217.5) long). Ventral proportions: gnathosomal bay length / width 1.63 - 1.87 (1.73); anterior venter / genital field length 1.62 - 1.77 (1.67); anterior venter length / genital field width 2.04 - 2.21 (2.07); anterior venter / medial suture 2.36 - 2.88 (2.81). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
E94C7E0838592BCF2EFE6F4D10A5B0E5.taxon	etymology	Etymology. Specific epithet (copipalpa) refers to the blade-like pedipalp femoral tubercles (copis, L. small knife; palpus, L. hand, feeler), which distinguish them from similar, co-occurring species.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
E94C7E0838592BCF2EFE6F4D10A5B0E5.taxon	distribution	Distribution. Northern California and western Oregon (Figure 41).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
9C4F7DD6DD4CBF79B95D64B295346E81.taxon	materials_examined	Material examined. HOLOTYPE (♀): from USA, Alabama, Clay County, beside Forest Route 649, 0.8 km northeast of road from Campbell Springs to Forest Route 600, (33 ° 22 ' 22 " N, 85 ° 52 ' 52 " W), 3 July 1990, by IM Smith, IMS 900075 A. PARATYPES (4 ♀; 4 ♂): Alabama, USA: 1 ♂ (ALLOTYPE) from Clay County, beside Forest Route 649, 0.8 km northeast of road from Campbell Springs to Forest Route 600, (33 ° 22 ' 22 " N, 85 ° 52 ' 52 " W), 3 July 1990, by IM Smith, IMS 900075 A * 4 ♀ and 3 ♂ from Clay County, beside Forest Route 649, 0.8 km northeast of road from Campbell Springs to Forest Route 600, (33 ° 22 ' 22 " N, 85 ° 52 ' 52 " W), 3 July 1990, by IM Smith, IMS 900075 A	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
9C4F7DD6DD4CBF79B95D64B295346E81.taxon	diagnosis	Diagnosis. Torrenticola daemon are similar to other members of the Raptor Group (T. danielleae, T. elusiva, T. gnoma, T. irapalpa, T. ivyae, T. longitibia, T. mjolniri, T. racupalpa, and T. raptor) in having round bodies; Dgl- 4 close to muscles scars; long, thin subcapitular rostra; and long, thin pedipalp tibiae. T. daemon can be differentiated from all other Raptor Group by having Dgl- 4 closer to the edge of the dorsum (dorsum width / distance between Dgl- 4 ♀ = 1.59 - 1.67 in T. daemon, 1.80 - 3.29 in others; ♂ = 1.45 - 1.65 in T. daemon, 1.66 - 2.73 in others), except T. irapalpa (♀ = 1.81 - 2.09, ♂ = 1.58 - 1.86) and T. danielleae (♀ = 1.57 - 1.70, ♂ = 1.42 - 1.52). T. daemon can be differentiated from T. longitibia, T. mjolniri, T. elusiva, T. racupalpa, T. raptor, T. danielleae, and T. ivyae by having a less elongate rostrum (length / width = 2.91 - 3.31 in T. daemon, 3.43 - 4.40 in others). Female T. daemon can be differentiated from female T. irapalpa by having Dgl- 4 closer to the dorsal edge (dorsal width / distance between Dgl- 4 ♀ = 1.59 - 1.67 in T. daemon, 1.81 - 2.09 in T. irapalpa) and a more elongate gnathosomal bay (length / width ♀ = 1.95 - 2.42 in T. daemon, 1.35 - 1.86 in T. irapalpa). Additionally, T. daemon can be differentiated from T. irapalpa by dorsal coloration and pattern.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
9C4F7DD6DD4CBF79B95D64B295346E81.taxon	description	Description. Female (Figure 45) (n = 5) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (590 - 655 (640) long; 460 - 500 (490) wide) circular with faint reddish-purple coloration separated into anterior and posterior portions, with bright reddish-purple coloration on the anterior-medial platelets, occasionally extending onto the dorsal plate. Anterio-medial platelets (127.5 - 147.5 (145) long; 57.5 - 67.5 (65) wide). Anterio-lateral platelets (182.5 - 210 (210) long; 72.5 - 82.5 (82.5) wide) free from dorsal plate. Dgl- 4 approximately halfway between the edge of the dorsum and the muscle scars (distance between Dgl- 4 285 - 300 (300 )). Dorsal plate proportions: dorsum length / width 1.26 - 1.34 (1.31); dorsal width / distance between Dgl- 4 1.59 - 1.67 (1.63); anterio-medial platelet length / width 2.04 - 2.28 (2.23); anterio-lateral platelet length / width 2.28 - 2.66 (2.55); anterio-lateral / anterio-medial length 1.29 - 1.51 (1.45). Gnathosoma - Subcapitulum (370 - 395 (395) long (ventral); 285 - 300 (300) long (dorsal); 145 - 157.5 (157.5) tall) colorless, occasionally with faint reddish-purple coloration. Rostrum (155 - 165 (165) long; 50 - 55 (55) wide). Chelicerae (375 - 390 (390) long) with curved fangs (65 - 75 (75) long). Subcapitular proportions: ventral length / height 2.39 - 2.57 (2.51); rostrum length / width 2.91 - 3.20 (3.00). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (50 - 52.5 (50) long); femur (137.5 - 146.25 (145) long); genu (75 - 80 (80) long); tibia (97.5 - 105 (105) long; 22.5 - 25 (25) wide); tarsus (20 - 22.5 (20) long). Palpomere proportions: femur / genu 1.81 - 1.90 (1.81); tibia / femur 0.67 - 0.72 (0.72); tibia length / width 4.05 - 4.33 (4.20). Venter - (710 - 800 (800) long; 515 - 570 (545) wide) colorless. Gnathosomal bay (182.5 - 200 (190) long; 80 - 97.5 (97.5) wide). Cxgl- 4 subapical. Medial suture (10 - 15 (15) long). Genital plates (160 - 175 (175) long; 145 - 150 (150) wide). Additional measurements: Cx- 1 (310 - 345 (345) long (total); 125 - 155 (155) long (medial )); Cx- 3 (350 - 375 (375) wide); anterior venter (155 - 185 (185) long). Ventral proportions: gnathosomal bay length / width 1.95 - 2.42 (1.95); anterior venter / genital field length 0.97 - 1.09 (1.06); anterior venter length / genital field width 1.07 - 1.23 (1.23); anterior venter / medial suture 10.33 - 17.50 (12.33). Male (Figure 46) (n = 4) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (545 - 570 (570) long; 410 - 425 (425) wide) circular with faint reddish-purple coloration separated into anterior and posterior portions. Anterio-medial platelets (115 - 125 (122.5) long; 52.5 - 58.75 (58.75) wide). Anterio-lateral platelets (175 - 188.75 (188.75) long; 67.5 - 75 (75) wide) free from dorsal plate. Dgl- 4 approximately halfway between the edge of the dorsum and the muscle scars (distance between Dgl- 4 255 - 290 (275 )). Dorsal plate proportions: dorsum length / width 1.31 - 1.34 (1.34); dorsal width / distance between Dgl- 4 1.45 - 1.65 (1.55); anterio-medial platelet length / width 2.09 - 2.22 (2.09); anterio-lateral platelet length / width 2.50 - 2.78 (2.52); anterio-lateral / anterio-medial length 1.40 - 1.63 (1.54). Gnathosoma - Subcapitulum (320 - 335 (335) long (ventral); 247.5 - 255 (255) long (dorsal); 120 - 122.5 (120) tall) colorless, occasionally with faint reddish-purple coloration. Rostrum (132.5 - 137.5 (137.5) long; 40 - 45 (45) wide). Chelicerae (310 - 330 (330) long) with curved fangs (60 - 65 (60) long). Subcapitular proportions: ventral length / height 2.67 - 2.79 (2.79); rostrum length / width 3.06 - 3.31 (3.06). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (42.5 - 46.25 (45) long); femur (117.5 - 122.5 (120) long); genu (67.5 - 70 (67.5) long); tibia (90 - 92.5 (91.25) long; 21.25 - 22.5 (22.5) wide); tarsus (20 - 20 (20) long). Palpomere proportions: femur / genu 1.74 - 1.78 (1.78); tibia / femur 0.76 - 0.77 (0.76); tibia length / width 4.00 - 4.35 (4.06). Venter - (655 - 705 (705) long; 460 - 470 (470) wide) colorless. Gnathosomal bay (165 - 165 (165) long; 70 - 80 (75) wide). Cxgl- 4 subapical. Medial suture (50 - 55 (55) long). Genital plates (145 - 155 (155) long; 115 - 120 (120) wide). Additional measurements: Cx- 1 (290 - 315 (290) long (total); 130 - 150 (130) long (medial )); Cx- 3 (320 - 340 (340) wide); anterior venter (210 - 225 (225) long). Ventral proportions: gnathosomal bay length / width 2.06 - 2.36 (2.20); anterior venter / genital field length 1.42 - 1.55 (1.45); anterior venter length / genital field width 1.75 - 1.91 (1.88); anterior venter / medial suture 4.00 - 4.20 (4.09). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
9C4F7DD6DD4CBF79B95D64B295346E81.taxon	etymology	Etymology. Specific epithet (daemon) refers to the diagnostic red coloration on the anterio-medial platelets, which resemble the red eyes of an evil demon (daemon, L. originally benevolent or benign nature spirits, but were characterized as dangerous or evil by the writings of Plato and later used in Christian literature, popularizing the idea of demons as evil; noun in apposition).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
9C4F7DD6DD4CBF79B95D64B295346E81.taxon	distribution	Distribution. Alabama (Figure 44).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
FE4E93ED2087CDC24C91688A17EFA5AC.taxon	materials_examined	Material examined. HOLOTYPE (♀): from USA, Georgia, Floyd County, The Pocket Campground beside road from Everett Springs to Villanow, (34 ° 35 ' 35 " N, 85 ° 5 ' 5 " W), 2 July 1990, by IM Smith, IMS 900073 A. PARATYPES (4 ♀; 5 ♂): Georgia, USA: 2 ♀ and 1 ♂ from Chattooga County, beside road from Everett Springs to Villanow 1.8 km south of The Pocket Recreation Area, (34 ° 34 ' 34 " N, 80 ° 5 ' 5 " W), 4 July 1990, by IM Smith, IMS 900076 * 1 ♂ (ALLOTYPE) from Floyd County, The Pocket Campground beside road from Everett Springs to Villanow, (34 ° 35 ' 35 " N, 85 ° 5 ' 5 " W), 2 July 1990, by IM Smith, IMS 900073 A * 2 ♀ and 3 ♂ from Floyd County, The Pocket Campground beside road from Everett Springs to Villanow, (34 ° 35 ' 35 " N, 85 ° 5 ' 5 " W), 2 July 1990, by IM Smith, IMS 900073 A	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
FE4E93ED2087CDC24C91688A17EFA5AC.taxon	diagnosis	Diagnosis. Torrenticola danielleae are similar to other members of the Raptor Group (T. daemon, T. elusiva, T. gnoma, T. irapalpa, T. ivyae, T. longitibia, T. mjolniri, T. racupalpa, and T. raptor) in having round bodies; Dgl- 4 close to muscles scars; long, thin subcapitular rostra; and long, thin pedipalp tibiae. T. danielleae can be differentiated from all other Raptor Group by having Dgl- 4 closer to the edge of the dorsum (dorsum width / distance between Dgl- 4 ♀ = 1.57 - 1.70 in T. danielleae, 1.80 - 3.29 in others; ♂ = 1.42 - 1.52 in T. danielleae, 1.58 - 2.73 in others), except T. daemon (♀ = 1.59 - 1.67 ♂ = 1.45 - 1.65). T. danielleae can be differentiated from T. daemon by having a more elongate rostrum (length / width = 3.43 - 3.75 in T. danielleae, 2.91 - 3.31 in T. daemon) and dorsal pattern.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
FE4E93ED2087CDC24C91688A17EFA5AC.taxon	description	Description. Female (Figure 48) (n = 5) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (635 - 680 (635) long; 510 - 545 (510) wide) circular with reddish-purple coloration posteriorly extending in a strip anteriorly to the edge of the dorsal plate. Anterio-medial platelets (135 - 155 (147.5) long; 55 - 65 (62.5) wide). Anterio-lateral platelets (170 - 195 (180) long; 70 - 80 (80) wide) free from dorsal plate. Dgl- 4 approximately halfway between the edge of the dorsum and the muscle scars (distance between Dgl- 4 320 - 335 (320 )). Dorsal plate proportions: dorsum length / width 1.20 - 1.30 (1.25); dorsal width / distance between Dgl- 4 1.57 - 1.7 (1.59); anterio-medial platelet length / width 2.25 - 2.73 (2.36); anterio-lateral platelet length / width 2.25 - 2.79 (2.25); anterio-lateral / anterio-medial length 1.15 - 1.44 (1.22). Gnathosoma - Subcapitulum (340 - 370 (365) long (ventral); 260 - 280 (270) long (dorsal); 135 - 145 (145) tall) colorless. Rostrum (150 - 160 (155) long; 40 - 45 (42.5) wide). Chelicerae (350 - 380 (380) long) with curved fangs (50 - 62.5 (62.5) long). Subcapitular proportions: ventral length / height 2.52 - 2.65 (2.52); rostrum length / width 3.44 - 3.75 (3.65). Pedipalps with long, tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (40 - 45 (42.5) long); femur (127.5 - 141.25 (137.5) long); genu (67.5 - 75 (75) long); tibia (92.5 - 107.5 (102.5) long; 20 - 22.5 (21.25) wide); tarsus (17.5 - 20 (20) long). Palpomere proportions: femur / genu 1.77 - 1.89 (1.83); tibia / femur 0.73 - 0.76 (0.75); tibia length / width 4.63 - 5.00 (4.82). Venter - (780 - 835 (780) long; 560 - 610 (560) wide) with reddish-purple coloration, occasionally faint. Gnathosomal bay (170 - 185 (185) long; 75 - 85 (80) wide). Cxgl- 4 subapical. Medial suture (25 - 27.5 (27.5) long). Genital plates (165 - 185 (185) long; 155 - 160 (160) wide). Additional measurements: Cx- 1 (300 - 340 (315) long (total); 135 - 155 (135) long (medial )); Cx- 3 (345 - 360 (355) wide); anterior venter (170 - 195 (185) long). Ventral proportions: gnathosomal bay length / width 2.00 - 2.31 (2.31); anterior venter / genital field length 1.00 - 1.12 (1.00); anterior venter length / genital field width 1.10 - 1.24 (1.16); anterior venter / medial suture 6.73 - 7.80 (6.73). Male (Figure 49) (n = 5) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (490 - 530 (510) long; 370 - 385 (370) wide) circular with reddish-purple coloration posteriorly extending in a strip anteriorly to the edge of the dorsal plate. Anterio-medial platelets (115 - 125 (120) long; 45 - 52.5 (52.5) wide). Anterio-lateral platelets (150 - 175 (150) long; 57.5 - 65 (65) wide) free from dorsal plate. Dgl- 4 approximately halfway between the edge of the dorsum and the muscle scars (distance between Dgl- 4 250 - 265 (260 )). Dorsal plate proportions: dorsum length / width 1.27 - 1.39 (1.38); dorsal width / distance between Dgl- 4 1.42 - 1.52 (1.42); anterio-medial platelet length / width 2.29 - 2.72 (2.29); anterio-lateral platelet length / width 2.31 - 2.92 (2.31); anterio-lateral / anterio-medial length 1.25 - 1.43 (1.25). Gnathosoma - Subcapitulum (285 - 295 (290) long (ventral); 210 - 220 (215) long (dorsal); 100 - 105 (105) tall) colorless. Rostrum (120 - 125 (120) long; 35 - 35 (35) wide). Chelicerae (270 - 300 (280) long) with curved fangs (40 - 50 (50) long). Subcapitular proportions: ventral length / height 2.74 - 2.85 (2.76); rostrum length / width 3.43 - 3.57 (3.43). Pedipalps with long, tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (33.75 - 37.5 (36.25) long); femur (100 - 110 (107.5) long); genu (60 - 62.5 (62.5) long); tibia (80 - 82.5 (82.5) long; 20 - 20 (20) wide); tarsus (17.5 - 20 (17.5) long). Palpomere proportions: femur / genu 1.67 - 1.79 (1.72); tibia / femur 0.74 - 0.83 (0.77); tibia length / width 4.00 - 4.13 (4.13). Venter - (620 - 670 (650) long; 410 - 430 (420) wide) with reddish-purple coloration, occasionally faint. Gnathosomal bay (135 - 145 (142.5) long; 62.5 - 70 (67.5) wide). Cxgl- 4 subapical. Medial suture (60 - 65 (60) long). Genital plates (132.5 - 145 (140) long; 102.5 - 107.5 (105) wide). Additional measurements: Cx- 1 (260 - 280 (280) long (total); 125 - 135 (135) long (medial )); Cx- 3 (290 - 310 (310) wide); anterior venter (205 - 220 (205) long). Ventral proportions: gnathosomal bay length / width 2.07 - 2.23 (2.11); anterior venter / genital field length 1.46 - 1.57 (1.46); anterior venter length / genital field width 1.91 - 2.05 (1.95); anterior venter / medial suture 3.15 - 3.52 (3.42). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
FE4E93ED2087CDC24C91688A17EFA5AC.taxon	etymology	Etymology. Specific epithet (danielleae) named in honor of Danielle Fisher-lab technician, environmental educator, colleague, friend, wife (of JRF), and mother of Ivy, our beautiful daughter-for her tireless and immense contributions to this research, and for bettering the lives of all those around her. Thank you, Danielle.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
FE4E93ED2087CDC24C91688A17EFA5AC.taxon	distribution	Distribution. Southern Appalachians, northeastern Georgia (Figure 47).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
F5BE8B8C021C7058E076E9FBE122648B.taxon	materials_examined	Material examined. Type series. HOLOTYPE (♂): from USA, New Jersey, Sussex County, Flatbrookeville, Flatbrook, 12 Oct 1953, by H Habeeb, HH 530113. PARATYPES (1 ♀; 0 ♂): New Jersey, USA: 1 ♀ (ALLOTYPE) from Sussex County, Flatbrookeville, Flatbrook, 12 Oct 1953, by H Habeeb, HH 530113. OTHER MATERIAL (19 ♀; 7 ♂): North Carolina, USA: 1 ♀ from Haywood County, Great Smokey Mountains National Park, Cataloochee River (35 ° 38 ' 45 " N, 83 ° 4 ' 34 " W), 6 Sep 2009, by IM Smith, IMS 090099 * Maine, USA: 1 ♀ and 1 ♂ from Franklin County, Small Falls picnic area beside Route 4, Sandy River (44 ° 52 ' N, 70 ° 31 ' W), 5 Jul 1989, by IM Smith, IMS 890069 * New Hampshire, USA: 1 ♀ and 1 ♂ from Woodstock County, beside Route 118, Jackman Brook (44 ° 0 ' N, 71 ° 45 ' W), 11 Sep 1992, by IM Smith, IMS 920036 * Pennsylvania, USA: 1 ♀ from Fayette County, Dunbar Creek (39 ° 57 ' 50 " N, 79 ° 35 ' 8.70 " W), 10 Aug 2014, by MJ Skvarla, MS 14 - 0810 - 001 * 2 ♀ and 2 ♂ from Fayette County, Ohiopyle State Park, Laurel Run (39 ° 50 ' 58 " N, 79 ° 30 ' 51 " W), 10 Aug 2014, by MJ Skvarla, MS 14 - 0810 - 005 * 2 ♀ from Somerset County, Laurel Hill State Park, Laurel Hill Creek (40 ° 1 ' 6 " N, 79 ° 14 ' 4 " W), 8 Aug 2014, by MJ Skvarla, MS 14 - 0808 - 001 * Quebec, Canada: 1 ♀ from Stanstead County, 1 kilometer south of Rock Island, Tomifobia River, Tompkin Stream, (45 ° 0 ' 31 " N, 72 ° 7 ' 6 " W), 20 Aug 1996, by IM Smith & M MacKenzie, IMS 960056 * Tennessee, USA: 1 ♀ from Blount County, Great Smokey Mountains National Park, Abrams River (35 ° 35 ' 31 " N, 83 ° 51 ' 21 " W), 17 Sep 2010, by IM Smith, IMS 100141 * 1 ♀ from Blount County, Great Smokey Mountains National Park, Little River (35 ° 40 ' 55 " N, 83 ° 39 ' 6 " W), 8 Sep 2009, by IM Smith, IMS 090102 * 2 ♀ from Sevier County, Great Smokey Mountains National Park, middle prong of Little Pigeon River (35 ° 43 ' 34 " N, 83 ° 24 ' 2 " W), 10 Sep 2010, by IM Smith, IMS 100127 * 1 ♀ from Sevier County, Great Smokey Mountains National Park, middle prong of Little Pigeon River (35 ° 43 ' 34 " N, 83 ° 24 ' 2 " W), 10 Sep 2010, by IM Smith, IMS 100128 * 1 ♀ from Sevier County, Great Smokey Mountains National Park, Sugarlands Nature Trail, spring (35 ° 40 ' 47 " N, 83 ° 31 ' 52 " W), 18 Sep 2010, by IM Smith, IMS 100147 * South Carolina, USA: 1 ♀ from Greenville County, Matthews Creek, 24 Apr 2014, by D Eargle, JRF 14 - 0424 - 001 * Vermont, USA: 1 ♀ and 1 ♂ from Addison County, beside road from Lincoln, Middlebury River, (44 ° 0 ' N, 73 ° 1 ' W), 6 Jul 1989, by IM Smith, IMS 890075 * Virginia, USA: 1 ♀ and 1 ♂ from Montgomery County, Blacksburg, beside Route 321 at Caldwell, Craig Creek (37 ° 20 ' 0 " N, 80 ° 20 ' 0 " W), 12 Jul 1990, by IM Smith, IMS 900089 A * 1 ♀ and 1 ♂ from Patrick County, Round Meadow Creek (36 ° 42 ' 59 " N, 80 ° 25 ' 29 " W), 10 Jun 2006, by IM Smith, IMS 060005 A.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
F5BE8B8C021C7058E076E9FBE122648B.taxon	diagnosis	Diagnosis. Torrenticola delicatexa are similar to other members of Rusetria " Eastern 2 - Plates " group (T. biscutella, T. caerulea, T. feminellai, T. indistincta, T. malarkeyorum, T. microbiscutella, T. pendula, T. sellersorum, T. tysoni, T. ululata, and T. whitneyae) in having anterio-lateral platelets fused to the dorsal plate, having dorsal coloration separated into anterior and posterior portions (except T. indistincta and T. ululata), and being distributed in the east. T. delicatexa can be differentiated from T. ululata, T. indistincta, and T. feminellai by dorsal coloration and pattern. T. delicatexa can be differentiated from T. tysoni by having a stockier rostrum (length / width = 2.33 - 3.00 in T. delicatexa, 3.06 - 3.50 in A 34). T. delicatexa can be differentiated from T. pendula by having a stockier gnathosomal bay (length / width = 1.28 - 2.22 in T. delicatexa, 2.42 - 2.90 in T. pendula), and a longer dorsum (♀ = 560 - 620 in T. delicatexa, 630 - 650 in T. pendula; ♂ = 420 - 465 in T. delicatexa, 500 in T. pendula). T. delicatexa can be differentiated from T. microbiscutella by having a less elongate dorsum (length / width = 1.38 - 1.56 in T. delicatexa, 1.63 - 1.75 in T. microbiscutella). Female T. delicatexa can be differentiated from female T. malarkeyorum, T. biscutella, and T. caerulea by having a longer genital field (175 - 185 in T. delicatexa, 153 - 170 in others). Male T. delicatexa can be differentiated from male T. biscutella by having a more ovoid dorsum (length / width = 1.44 - 1.56 in T. delicatexa; 1.37 - 1.42 in T. biscutella). Female T. delicatexa can be differentiated from female T. sellersorum by having a slightly more ovoid dorsum (length / width = 1.38 - 1.44 in T. delicatexa, 1.23 - 1.37 in T. sellersorum). Male T. delicatexa do not have any measurement differences with male T. malarkeyorum, T. caerulea, and T. sellersorum; however, they can be differentiated by dorsal coloration. T. delicatexa can be differentiated from T. whitneyae by having a slightly more ovoid dorsum (length / width ♀ = 1.38 - 1.44 in T. delicatexa, 1.26 - 1.38 in T. whitneyae; ♂ = 1.44 - 1.56 in T. delicatexa, 1.35 - 1.37 in T. whitneyae) and by dorsal coloration. Additionally, male T. delicatexa can be differentiated from male T. whitneyae by having more elongate pedipalpal tibiae (length / width = 2.89 - 3.63 in T. delicatexa, 2.48 - 2.70 in T. whitneyae).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
F5BE8B8C021C7058E076E9FBE122648B.taxon	description	Re-description. Male (Figure 51) (n = 5) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (410 - 465 (465) long; 270 - 320 (320) wide) ovoid with highly variable coloration, reddish-purple to purple (occasionally bluish-purple) separated into anterior and posterior portions. Anterio-medial platelets (83.75 - 102.5 (102.5) long; 30 - 32.5 (32.5) wide). Anterio-lateral platelets (122.5 - 132.5 (132.5) long; 40 - 55 (52.5) wide) fused to dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 205 - 255 (255 )). Dorsal plate proportions: dorsum length / width 1.44 - 1.56 (1.45); dorsal width / distance between Dgl- 4 1.23 - 1.37 (1.25); anterio-medial platelet length / width 2.79 - 3.17 (3.15); anterio-lateral platelet length / width 2.36 - 3.06 (2.52); anterio-lateral / anterio-medial length 1.29 - 1.46 (1.29). Gnathosoma - Subcapitulum (235 - 247.5 (247.5) long (ventral); 172.5 - 193.75 (194) long (dorsal); 97.5 - 107.5 (107.5) tall) colorless. Rostrum (90 - 100 (100) long; 32.5 - 37.5 (37.5) wide). Chelicerae (220 - 230 long) with curved fangs (45 - 52.5 (50) long). Subcapitular proportions: ventral length / height 2.30 - 2.45 (2.30); rostrum length / width 2.67 - 2.81 (2.67). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (33.75 - 40 (40) long); femur (82.5 - 92.5 (92.5) long); genu (50 - 55 (55) long); tibia (62.5 - 72.5 (65) long; 20 - 22.5 (22.5) wide); tarsus (15 - 17.5 (17.5) long). Palpomere proportions: femur / genu 1.59 - 1.75 (1.68); tibia / femur 0.70 - 0.83 (0.70); tibia length / width 2.89 - 3.63 (2.89). Venter - (490 - 540 (540) long; 311 - 435 (435) wide) usually colorless; occasionally with faint reddish-purple coloration. Gnathosomal bay (120 - 135 (135) long; 57.5 - 72.5 (72.5) wide). Cxgl- 4 subapical. Medial suture (55 - 75 (62.5) long). Genital plates (105 - 110 (106.25) long; 95 - 102.5 (102.5) wide). Additional measurements: Cx- 1 (220 - 235 (235) long (total); 78 - 100 (100) long (medial )); Cx- 3 (240 - 297.5 (297.5) wide); anterior venter (160 - 175 (170) long). Ventral proportions: gnathosomal bay length / width 1.86 - 2.22 (1.86); anterior venter / genital field length 1.52 - 1.60 (1.60); anterior venter length / genital field width 1.64 - 1.79 (1.66); anterior venter / medial suture 2.33 - 2.91 (2.72). Female (Figure 52) (n = 6) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (560 - 620 (620) long; 390 - 440 (435) wide) ovoid with highly variable coloration, reddish-purple to purple (occasionally bluish-purple) separated into anterior and posterior portions. Anterio-medial platelets (125 - 140 (140) long; 36.25 - 45 (42.5) wide). Anterio-lateral platelets (152.5 - 172.5 (162.5) long; 57.5 - 70 (70) wide) fused to dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 295 - 350 (330 )). Dorsal plate proportions: dorsum length / width 1.38 - 1.44 (1.43); dorsal width / distance between Dgl- 4 1.26 - 1.33 (1.32); anterio-medial platelet length / width 2.83 - 3.52 (3.29); anterio-lateral platelet length / width 2.26 - 2.88 (2.32); anterio-lateral / anterio-medial length 1.16 - 1.33 (1.16). Gnathosoma - Subcapitulum (305 - 345 (345) long (ventral); 216 - 260 (260) long (dorsal); 145 - 165 (160) tall) colorless. Rostrum (115 - 135 (135) long; 45 - 55 (45) wide). Chelicerae (312 - 350 (350) long) with curved fangs (54 - 75 (70) long). Subcapitular proportions: ventral length / height 1.97 - 2.22 (2.16); rostrum length / width 2.33 - 3.00 (3.00). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (42.5 - 52.5 (47.5) long); femur (105 - 130 (128.75) long); genu (67.5 - 75 (72.5) long); tibia (81.25 - 90 (87.5) long; 22.5 - 30 (30) wide); tarsus (20 - 22.5 (22.5) long). Palpomere proportions: femur / genu 1.56 - 1.78 (1.78); tibia / femur 0.65 - 0.77 (0.68); tibia length / width 2.92 - 3.61 (2.92). Venter - (640 - 690 (690) long; 431 - 540 (540) wide) usually colorless; occasionally with faint reddish-purple coloration. Gnathosomal bay (132.5 - 175 (157.5) long; 85 - 117.5 (117.5) wide). Cxgl- 4 subapical. Medial suture absent. Genital plates (175 - 197.5 (197.5) long; 150 - 172.5 (170) wide). Additional measurements: Cx- 1 (216 - 295 (295) long (total); 93 - 135 (135) long (medial )); Cx- 3 (304 - 400 (400) wide); anterior venter (115 - 135 (135) long). Ventral proportions: gnathosomal bay length / width 1.28 - 2.06 (1.34); anterior venter / genital field length 0.64 - 0.76 (0.68); anterior venter length / genital field width 0.71 - 0.83 (0.79). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
F5BE8B8C021C7058E076E9FBE122648B.taxon	etymology	Etymology. Habeeb (1955) did not offer an explanation for the specific epithet (delicatexa) and we are unable to offer helpful speculation.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
F5BE8B8C021C7058E076E9FBE122648B.taxon	distribution	Distribution. Appalachians (Figure 50).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
83DE0C600FCF3267273E7A349FF1BFDD.taxon	materials_examined	Material examined. HOLOTYPE (♀): from USA, Georgia, Chattooga County, beside road from Everett Springs to Villanow 1.4 km south of The Pocket Recreation Area, 4 July 1990, by IM Smith, IMS 900077. PARATYPES (4 ♀; 2 ♂): Georgia, USA: 1 ♂ (ALLOTYPE) from Chattooga County, beside road from Everett Springs to Villanow 1.4 km south of The Pocket Recreation Area, 4 July 1990, by IM Smith, IMS 900077 * 2 ♀ and 1 ♂ from Chattooga County, beside road from Everett Springs to Villanow 1.4 km south of The Pocket Recreation Area, 4 July 1990, by IM Smith, IMS 900077 * Tennessee, USA: 1 ♀ from Monroe County, beside Forest Route 35, 2.0 km northeast of road from Rt. 165 to Miller Chapel Baptist Church, (35 ° 21 ' 21 " N, 84 ° 9 ' 9 " W), 5 July 1990, by IM Smith, IMS 900078 * Virginia, USA: 1 ♀ from Alleghany County, Covington; beside Rt. 18, 0.5 km north of Rt. 657, (37 ° 44 ' 44 " N, 80 ° 2 ' 2 " W), 13 July 1990, by IM Smith, IMS 900091 A.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
83DE0C600FCF3267273E7A349FF1BFDD.taxon	diagnosis	Diagnosis. Torrenticola dentirostra are similar to other members of the Nigroalba Group (T. flangipalpa, T. nigroalba, and T. solisorta) in being small, slightly elongate, and having purple dorsal coloration restricted posteriorly. T. dentirostra can be differentiated from most Torrenticola (except T. erectirostra, T. karambita, and T. robisoni) by having a dentate bump midway on the dorsal edge of the rostrum. T. dentirostra can be differentiated from T. karambita, T. erectirostra, and T. robisoni by having a straight rostrum (others have upturned rostra) and by dorsal pattern.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
83DE0C600FCF3267273E7A349FF1BFDD.taxon	description	Description. Female (Figure 54) (n = 5) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (470 - 560 (470) long; 340 - 390 (340) wide) ovoid with faint purple coloration restricted posteriorly. Anterio-medial platelets (107.5 - 120 (107.5) long; 45 - 47.5 (45) wide). Anterio-lateral platelets (157.5 - 177.5 (157.5) long; 55 - 62.5 (55) wide) free from dorsal plate. Dgl- 4 closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 240 - 275 (240 )). Dorsal plate proportions: dorsum length / width 1.38 - 1.45 (1.38); dorsal width / distance between Dgl- 4 1.38 - 1.43 (1.42); anterio-medial platelet length / width 2.32 - 2.67 (2.39); anterio-lateral platelet length / width 2.84 - 3.05 (2.86); anterio-lateral / anterio-medial length 1.43 - 1.59 (1.47). Gnathosoma - Subcapitulum (275 - 330 (275) long (ventral); 200 - 247.5 (200) long (dorsal); 77.5 - 102.5 (77.5) tall) colorless. Rostrum (100 - 120 (100) long; 35 - 40 (35) wide) with dentate bump midway on dorsal edge. Chelicerae (255 - 320 (255) long) with curved fangs (45 - 55 (45) long). Subcapitular proportions: ventral length / height 3.22 - 3.73 (3.55); rostrum length / width 2.80 - 3.00 (2.86). Pedipalps with tuberculate ventral extensions dentate apically on femora and genua. Palpomeres: trochanter (30 - 35 (30) long); femur (92.5 - 110 (92.5) long); genu (55 - 67.5 (55) long); tibia (80 - 92.5 (80) long; 17.5 - 20 (17.5) wide); tarsus (12.5 - 15 (13.75) long). Palpomere proportions: femur / genu 1.62 - 1.68 (1.68); tibia / femur 0.84 - 0.88 (0.86); tibia length / width 4.57 - 4.93 (4.57). Venter - (610 - 685 (610) long; 390 - 425 (390) wide) mostly colorless with faint purple genital plates. Gnathosomal bay (85 - 110 (85) long; 60 - 80 (60) wide). Cxgl- 4 far from apex and ventral. Medial suture (60 - 70 (60) long). Genital plates (142.5 - 165 (142.5) long; 120 - 135 (120) wide). Additional measurements: Cx- 1 (230 - 260 (230) long (total); 140 - 160 (140) long (medial )); Cx- 3 (270 - 300 (270) wide); anterior venter (210 - 240 (210) long). Ventral proportions: gnathosomal bay length / width 1.30 - 1.50 (1.42); anterior venter / genital field length 1.42 - 1.64 (1.47); anterior venter length / genital field width 1.70 - 1.83 (1.75); anterior venter / medial suture 3.29 - 3.69 (3.50). Male (Figure 55) (n = 2) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (480 - 505 (505) long; 360 - 360 (360) wide ovoid with faint purple coloration restricted posteriorly. Anterio-medial platelets (110 - 110 (110) long; 40 - 42.5 (42.5) wide). Anterio-lateral platelets (162.5 - 165 (165) long; 50 - 52.5 (52.5) wide) free from dorsal plate. Dgl- 4 closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 240 - 245 (240 )). Dorsal plate proportions: dorsum length / width 1.33 - 1.40 (1.40); dorsal width / distance between Dgl- 4 1.47 - 1.50 (1.50); anterio-medial platelet length / width 2.59 - 2.75 (2.59); anterio-lateral platelet length / width 3.14 - 3.25 (3.14); anterio-lateral / anterio-medial length 1.48 - 1.50 (1.50). Gnathosoma - Subcapitulum (285 - 287.5 (287.5) long (ventral); 210 - 217.5 (217.5) long (dorsal); 77.5 - 80 (77.5) tall) colorless. Rostrum (100 - 105 (100) long; 32.5 - 35 (32.5) wide) with dentate bump midway on dorsal edge. Chelicerae (255 - 265 (265) long) with curved fangs (40 - 45 (45) long). Subcapitular proportions: ventral length / height 3.56 - 3.71 (3.71); rostrum length / width 3.00 - 3.08 (3.08). Pedipalps with tuberculate ventral extensions dentate apically on femora and genua. Palpomeres: trochanter (30 - 32.5 (32.5) long); femur (90 - 92.5 (90) long); genu (57.5 - 57.5 (57.5) long); tibia (82.5 - 85 (85) long; 17.5 - 18.75 (18.75) wide); tarsus (12.5 - 13.75 (13.75) long). Palpomere proportions: femur / genu 1.57 - 1.61 (1.57); tibia / femur 0.89 - 0.94 (0.94); tibia length / width 4.53 - 4.71 (4.53). Venter - (610 - 645 (645) long; 390 - 400 (400) wide) mostly colorless with faint purple genital plates. Gnathosomal bay (100 - 107.5 (407.5) long; 70 - 70 (70) wide). Cxgl- 4 far from apex and ventral. Medial suture (75 - 85 (85) long). Genital plates (130 - 135 (135) long; 102.5 - 102.5 (102.5) wide). Additional measurements: Cx- 1 (240 - 250 (250) long (total); 140 - 147.5 (147.5) long (medial )); Cx- 3 (275 - 280 (275) wide); anterior venter (235 - 247.5 (247.5) long). Ventral proportions: gnathosomal bay length / width 1.43 - 1.54 (1.54); anterior venter / genital field length 1.81 - 1.83 (1.83); anterior venter length / genital field width 2.29 - 2.41 (2.41); anterior venter / medial suture 2.91 - 3.13 (2.91). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
83DE0C600FCF3267273E7A349FF1BFDD.taxon	etymology	Etymology. Specific epithet (dentirostra) refers to the diagnostic tooth-like serrations on the dorsal surface of the rostrum (dentis, L. tooth; rostrum, L. snout).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
83DE0C600FCF3267273E7A349FF1BFDD.taxon	distribution	Distribution. Southern Appalachians (Figure 53).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
7366960139AEC05906F653B7A2111112.taxon	materials_examined	Material examined. HOLOTYPE (♀): from USA, Texas, Bandera County, Lost Maples State Natural Area, north of Vanderpool; picnic area, (29 ° 48 ' 48 " N, 99 ° 34 ' 34 " W), 10 October 2010, by IM Smith, IMS 100184. PARATYPES (8 ♀; 5 ♂): Texas, USA: 1 ♂ (ALLOTYPE) from Bandera County, Lost Maples State Natural Area, north of Vanderpool; picnic area, (29 ° 48 ' 48 " N, 99 ° 34 ' 34 " W), 10 October 2010, by IM Smith, IMS 100184 * 1 ♀ and 1 ♂ from Bandera County, Lost Maples State Natural Area, north of Vanderpool; picnic area, (29 ° 48 ' 48 " N, 99 ° 34 ' 34 " W), 10 October 2010, by IM Smith, IMS 100184 * 2 ♀ and 3 ♂ from Bandera County, Lost Maples State Natural Area north of Vanderpool; picnic area, (29 ° 48 ' 48 " N, 99 ° 34 ' 34 " W), 7 October 2010, by IM Smith, IMS 100178 * 2 ♀ from Bandera County, Lost Maples State Natural Area; picnic area, (29 ° 48 ' 48 " N, 99 ° 34 ' 34 " W), 27 May 1997, by IM Smith, IMS 970008 * 1 ♀ from Bandera County, Vanderpool; beside Rt. 187, 0.7 km south of entrance to Lost Maples State Natural Area, (29 ° 48 ' 48 " N, 99 ° 34 ' 34 " W), 2 May 2009, by IM Smith, IMS 090007 * 1 ♀ from Kinney County, Brackettville; beside Rt. 90, 12.1 km west of Rt. 131, (29 ° 20 ' 20 " N, 100 ° 32 ' 32 " W), 4 May 2003, by IM Smith, IMS 030007 * 1 ♀ from Uvalde County, Garner State Park; river crossing site, (29 ° 35 ' 35 " N, 99 ° 44 ' 44 " W), 28 May 1998, by IM Smith, IMS 980027 A.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
7366960139AEC05906F653B7A2111112.taxon	diagnosis	Diagnosis. Torrenticola dimorpha are similar to other members of the Tricolor Complex (T. bittikoferae, T. cardia, T. hoosieri, T. kringi, T. larvata, T. mohawk, T. pearsoni, T. olliei, T. sierrensis, T. tricolor, T. trimaculata, and T. unimaculata) in having a short, conical rostrum. T. dimorpha can be differentiated from all other Torrenticola by having a dorsal plate with a medial extension covering nearly half the length of the anterio-medial platelets and by males having large, highly modified pedipalps, which are expanded vertically and laterally. Additionally, T. dimorpha can be differentiated from most other Tricolor Complex (except T. bittikoferae, T. hoosieri, and T. pearsoni) by being colorless, whereas most other members have bold patterning.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
7366960139AEC05906F653B7A2111112.taxon	description	Description. Female (Figure 57) (n = 5) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (590 - 670 (650) long; 455 - 520 (490) wide) colorless and ovoid. Anterior dorsal plate with medial extension covering nearly half the length of the anterio-medial platelets. Anterio-medial platelets (117.5 - 135 (135) long; 52.5 - 57.5 (57.5) wide). Anterio-lateral platelets (145 - 172.5 (172.5 )) long; 55 - 65 (62.5) wide) free from dorsal plate. Dgl- 4 approximately halfway in between the edge of the dorsum and the muscle scars (distance between Dgl- 4 320 - 360 (360 )). Dorsal plate proportions: dorsum length / width 1.29 - 1.33 (1.33); dorsal width / distance between Dgl- 4 1.36 - 1.48 (1.36); anterio-medial platelet length / width 2.24 - 2.35 (2.35); anterio-lateral platelet length / width 2.46 - 2.78 (2.76); anterio-lateral / anterio-medial length 1.19 - 1.33 (1.28). Gnathosoma - Subcapitulum (230 - 260 (255) long (ventral); 150 - 170 (170) long (dorsal); 110 - 132.5 (132.5) tall) colorless. Posterior dorsal apodeme long. Rostrum (65 - 70 (70) long; 42.5 - 50 (45) wide) very short. Chelicerae (210 - 245 (235) long) with curved fangs (60 - 70 (70) long). Subcapitular proportions: ventral length / height 1.90 - 2.09 (1.92); rostrum length / width 1.40 - 1.56 (1.56). Pedipalps with long, tuberculate ventral extensions on femora and no ventral extensions on genua. Palpomeres: trochanter (37.5 - 40 (37.5) long); femur (76.25 - 85 (82.5) long); genu (52.5 - 62.5 (60) long); tibia (65 - 72.5 (70) long; 22.5 - 22.5 (22.5) wide); tarsus (20 - 22.5 (22.5) long). Palpomere proportions: femur / genu 1.36 - 1.48 (1.38); tibia / femur 0.84 - 0.87 (0.85); tibia length / width 2.89 - 3.22 (3.11). Venter - (720 - 800 (770) long; 505 - 600 (580) wide) colorless. Gnathosomal bay (120 - 145 (145) long; 72.5 - 85 (85) wide). Cxgl- 4 apical. Medial suture (25 - 30 (30 )). Genital plates (170 - 195 (185) long; 162.5 - 180 (180) wide). Additional measurements: Cx- 1 (240 - 270 (270) long (total); 110 - 130 (125) long (medial )); Cx- 3 (315 - 350 (350) wide); anterior venter (137.5 - 155 (155) long). Ventral proportions: gnathosomal bay length / width 1.60 - 1.79 (1.71); anterior venter / genital field length 0.78 - 0.84 (0.84); anterior venter length / genital field width 0.85 - 0.86 (0.86). Male (Figure 58) (n = 5) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (465 - 510 (500) long; 310 - 330 (320) wide) colorless and ovoid. Anterior dorsal plate with medial extension covering nearly half the length of the anterio-medial platelets. Muscle scars absent or very faint. Anterio-medial platelets (102.5 - 115 (107.5) long; 42.5 - 45 (45) wide). Anterio-lateral platelets (177.5 - 197.5 (195) long; 42.5 - 55 (55) wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 225 - 245 (245 )). Dorsal plate proportions: dorsum length / width 1.50 - 1.58 (1.56); dorsal width / distance between Dgl- 4 1.31 - 1.41 (1.31); anterio-medial platelet length / width 2.34 - 2.63 (2.39); anterio-lateral platelet length / width 3.38 - 4.65 (3.55); anterio-lateral / anterio-medial length 1.64 - 1.81 (1.81). Gnathosoma - Subcapitulum (227.5 - 255 (240) long (ventral); 160 - 177.5 (165) long (dorsal); 100 - 110 (100) tall) colorless. Rostrum (45 - 50 (47.5) long; 32.5 - 37.5 (32.5) wide) very short. Chelicerae (195 - 235 (205) long) with curved fangs (55 - 65 (60) long). Subcapitular proportions: ventral length / height 2.17 - 2.40 (2.40); rostrum length / width 1.29 - 1.46 (1.46). Pedipalps highly modified and expanded, with long, tuberculate ventral extensions on femora and no ventral extensions on genua. Palpomeres: trochanter (53.75 - 57.5 (55) long); femur (107.5 - 112.5 (112.5) long); genu (92.5 - 103.75 (103.75) long); tibia (77.5 - 85 (85) long; 30 - 35 (35) wide); tarsus (20 - 25 (22.5) long). Palpomere proportions: femur / genu 1.08 - 1.22 (1.08); tibia / femur 0.69 - 0.77 (0.76); tibia length / width 2.36 - 2.67 (2.43). Venter - (550 - 620 (580) long; 350 - 370 (370) wide) colorless and highly modified with coxae II-IV forming a large ventral plate that covers the insertions of legs IV. Suture dividing coxae III and IV incomplete. Apodemes expanded internally. Gnathosomal bay (110 - 115 (112.5) long; 90 - 110 (90) wide) expanded to accommodate large pedipalps. Cxgl- 4 apical. Medial suture (155 - 165 (155) long). Genital plates (100 - 120 (105) long; 105 - 110 (105) wide) triangular. Additional measurements: Cx- 1 (230 - 260 (250) long (total); 130 - 145 (130) long (medial )); Cx- 3 (335 - 345 (335) wide); anterior venter (297.5 - 315 (300) long). Ventral proportions: gnathosomal bay length / width 1.05 - 1.25 (1.25); anterior venter / genital field length 2.63 - 2.98 (2.86); anterior venter length / genital field width 2.70 - 2.86 (2.86); anterior venter / medial suture 1.88 - 2.03 (1.94). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
7366960139AEC05906F653B7A2111112.taxon	etymology	Etymology. Specific epithet (dimorpha) refers to the sexual dimorphism in terms of size and morphology (di, G. two; morph ḗ, G. form), the resulting morphology is so unlike all other Torrenticolidae that upon first glance specimens appear to be a different family altogether.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
7366960139AEC05906F653B7A2111112.taxon	distribution	Distribution. Texas (probably also extending southward into Mexico) (Figure 56).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
617B9D2BE3F758981A203218BE3C1C3D.taxon	materials_examined	Material examined. HOLOTYPE (♂): from USA, New Mexico, Catron County, Cottonwood Campground beside Rt. 180 south of Rt. 12, (33 ° 37 ' 37 " N, 108 ° 54 ' 54 " W), 12 July 1987, by IM Smith, IMS 870086. PARATYPES (0 ♀; 0 ♂):	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
617B9D2BE3F758981A203218BE3C1C3D.taxon	diagnosis	Diagnosis. Torrenticola dolichodactyla are similar to other members of the Rala Group (T. rala, T. lamellipalpis, T. boettgeri, T. kurtvietsi, T. keesdavidsi, and T. anoplopalpa) by being colorless, having incomplete hind coxal margins and being distributed in the southwest. T. dolichodactyla can be differentiated from all other Rala Group by having longer pedipalpal tarsi (♂ = 52.5 in T. dolichodactyla, 10 - 20 in others) and Dgl- 4 closer to the muscle scars (dorsum width / distance between Dgl- 4 ♂ = 1.74 in T. dolichodactyla, 1.16 - 1.48 in others).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
617B9D2BE3F758981A203218BE3C1C3D.taxon	description	Description. Female unknown. Male (Figure 60) (n = 1) (holotype only) with characters of the genus with following specifications. Dorsum - (735 long; 590 wide) circular and colorless. Anterio-medial platelets (177.5 long; 77.5 wide). Anterio-lateral platelets (227.5 long; 97.5 wide) free from dorsal plate. Dgl- 4 halfway between the muscle scars and the edge of the dorsum (distance between Dgl- 4 340). Dorsal plate proportions: dorsum length / width 1.25; dorsal width / distance between Dgl- 4 1.74; anterio-medial platelet length / width 2.29; anterio-lateral platelet length / width 2.33; anterio-lateral / anterio-medial length 1.28. Gnathosoma - Subcapitulum (325 long (ventral); 235 long (dorsal); 135 tall) colorless. Rostrum (120 long; 50 wide). Chelicerae (325 long) with curved fangs (65 long). Subcapitular proportions: ventral length / height 2.41; rostrum length / width 2.40. Pedipalps with elongate tarsi and tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (55 long); femur (140 long); genu (80 long); tibia (110 long; 32.5 wide); tarsus (52.5 long). Palpomere proportions: femur / genu 1.75; tibia / femur 0.79; tibia length / width 3.38. Venter - (850 long; 720 wide) colorless. Gnathosomal bay (170 long; 115 wide). Cxgl- 4 apical. Medial suture (95 long). Genital plates (220 long; 170 wide). Additional measurements: Cx- 1 (350 long (total); 175 long (medial )); Cx- 3 (480 wide); anterior venter (305 long). Ventral proportions: gnathosomal bay length / width 1.48; anterior venter / genital field length 1.39; anterior venter length / genital field width 1.79; anterior venter / medial suture 3.21. Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
617B9D2BE3F758981A203218BE3C1C3D.taxon	etymology	Etymology. Specific epithet (dolichodactyla) refers to elongate pedipalp tarsus (dolichos, G. long; daktylos, G. finger), which is the most elongate of all Torrenticolidae.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
617B9D2BE3F758981A203218BE3C1C3D.taxon	distribution	Distribution. New Mexico (probably also Arizona) (Figure 59).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
63B364F0307B6BFE78A59B64434E4215.taxon	materials_examined	Material examined. Type series. HOLOTYPE (♀): from USA, Tennessee, Sevier County, Great Smokey Mountains National Park, Cosby Recreation Area (35 ° 46 ' 54 " N, 83 ° 13 ' 2 " W), 16 Sep 2010, by IM Smith, IMS 100140, DNA 1289. PARATYPES (9 ♀; 5 ♂): North Carolina, USA: 1 ♂ (ALLOTYPE) from Haywood County, Great Smokey Mountains National Park, Rough Fork Creek (35 ° 37 ' 31 " N, 83 ° 6 ' 46 " W), 20 Sep 2010, by IM Smith, IMS 100148 * 2 ♀ and 2 ♂ from Haywood County, Great Smokey Mountains National Park, Rough Fork Creek (35 ° 37 ' 31 " N, 83 ° 6 ' 46 " W), 20 Sep 2010, by IM Smith, IMS 100148 * 1 ♀ from Haywood County, Great Smokey Mountains National Park, Waterville (35 ° 44 ' 59 " N, 83 ° 6 ' 42 " W), 16 Sep 2010, by IM Smith, IMS 100138 * South Carolina, USA: 2 ♀ from Greenville County, Matthews Creek, 24 Apr 2014, by D Eargle, JRF 14 - 0424 - 001 * Tennessee, USA: 1 ♀ from Blount County, Great Smokey Mountains National Park, Abrams River (35 ° 35 ' 31 " N, 83 ° 51 ' 21 " W), 17 Sep 2010, by IM Smith, IMS 100141 * 1 ♀ and 1 ♂ from Sevier County, Great Smokey Mountains National Park (35 ° 40 ' 47 " N, 83 ° 31 ' 48 " W), 3 Sep 2009, by IM Smith, IMS 090096 * 2 ♀ and 1 ♂ from Sevier County, Great Smokey Mountains National Park (35 ° 43 ' 33 " N, 83 ° 24 ' 1 " W), 12 Sep 2010, by IM Smith, IMS 100131 * Virginia, USA: 1 ♀ from Smyth County, Mount Rogers National Recreation Area, beside Route 600, Little Laurel Creek, 10 Jul 1990, by IM Smith, IMS 900086.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
63B364F0307B6BFE78A59B64434E4215.taxon	diagnosis	Diagnosis. Torrenticola dunni are similar to other members of the Rusetria " 4 - Plates " group (T. dunni, T. glomerabilis, T. kittatinniana, T. pollani, T. rufoalba and T. shubini) and T. skvarlai in having anterio-lateral platelets free from the dorsal plate, dorsal coloration separated into anterior and posterior portions, and indistinct hind coxal margins. T. dunni can be differentiated from T. pollani by having a larger dorsum (length ♀ = 605 - 680 in T. dunni, 535 - 560 in T. pollani; ♂ = 500 - 540 in T. dunni, 440 - 492 in T. pollani; width ♀ = 440 - 490 in T. dunni, 410 - 420 in T. pollani; ♂ = 350 - 370 in T. dunni, 310 - 340 in T. pollani); and a stockier rostrum (length / width = 2.80 - 3.14 in T. dunni, 3.27 - 3.82 in T. pollani). Female T. dunni can be differentiated from female T. shubini by having a thinner rostrum (length / width = 2.8 - 3.0 in T. dunni, 2.5 - 2.7 in T. shubini). Male T. dunni can be differentiated from male T. shubini by having a longer anterior venter (277 - 285 in T. dunni, 215 - 238 in T. shubini). T. dunni can be differentiated from T. glomerabilis by having Dgl- 4 closer to the dorsal edge (dorsal width / distance between Dgl- 4 = 1.2 - 1.4 in T. dunni, 1.5 - 1.7 in T. glomerabilis) and stockier tibiae (length / width ♀ = 3.27 - 3.50 in T. dunni, 4.11 - 4.50 in T. glomerabilis, ♂ = 3.25 - 3.44 in T. dunni, 3.55 - 4.38 in T. glomerabilis). Female T. dunni can be differentiated from female T. kittatinniana by having a longer pedipalp genu (70 - 75 in T. dunni, 64 in T. kittatinniana); a longer subcapitulum (ventral length = 330 - 355 in T. dunni, 310 in T. kittatinniana); and anterio-medial platelets more elongate (length / width = 2.33 - 2.54 in T. dunni, 2.83 in T. kittatinniana). Male T. dunni can be differentiated from male T. kittatinniana by having a longer anterior venter (277 - 285 in T. dunni, 235 in T. kittatinniana) and wider dorsum (350 - 370 in T. dunni, 340 in T. kittatinniana). T. dunni can be differentiated from T. rufoalba by having a larger dorsum (length ♀ = 605 - 680 in T. dunni, 550 in T. rufoalba; ♂ = 500 - 540 in T. dunni, 440 in T. rufoalba; width ♀ = 440 - 490 in T. dunni, 400 in T. rufoalba; ♂ = 350 - 370 in T. dunni; 320 in T. rufoalba). T. dunni can be differentiated from T. skvarlai by having a conical pedipalpal femoral tubercle, whereas T. skvarlai has a broad and flat pedipalpal femoral tubercle, and by having a longer anterior venter (♀ = 160 - 190 in T. dunni, 140 - 152.5 in T. skvarlai; ♂ = 277.5 - 285 in T. dunni, 177.5 - 205 in T. skvarlai).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
63B364F0307B6BFE78A59B64434E4215.taxon	description	Description. Female (Figure 62) (n = 5) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum- (605 - 680 (655) long; 440 - 490 (460) wide) ovoid with purple or reddish-purple coloration separated into anterior and posterior portions, and occasionally with faint orange medially. Anterio-medial platelets (117.5 - 125 (125) long; 46.25 - 52.5 (50) wide). Anterio-lateral platelets (172.5 - 197.5 (192.5) long; 62.5 - 68.75 (62.5) wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 315 - 380 (340 )). Dorsal plate proportions: dorsum length / width 1.37 - 1.42 (1.42); dorsal width / distance between Dgl- 4 1.29 - 1.40 (1.35); anterio-medial platelet length / width 2.33 - 2.54 (2.50); anterio-lateral platelet length / width 2.76 - 3.08 (3.08); anterio-lateral / anterio-medial length 1.44 - 1.61 (1.54). Gnathosoma - Subcapitulum (330 - 355 (345) long (ventral); 250 - 265 (255) long (dorsal); 132.5 - 150 (150) tall) colorless. Rostrum (130 - 140 (135) long; 45 - 50 (45) wide). Chelicerae (325 - 355 (350) long) with curved fangs (60 - 65 (60) long). Subcapitular proportions: ventral length / height 2.30 - 2.53 (2.30); rostrum length / width 2.80 - 3.00 (3.00). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (42.5 - 50 (48.75) long); femur (117.5 - 132.5 (131.25) long); genu (70 - 75 (75) long); tibia (85 - 95 (90) long; 25 - 27.5 (27.5) wide); tarsus (20 - 20 (20) long). Palpomere proportions: femur / genu 1.68 - 1.83 (1.75); tibia / femur 0.69 - 0.72 (0.69); tibia length / width 3.27 - 3.50 (3.27). Venter - (710 - 810 (780) long; 540 - 600 (600) wide) with faint bluish-purple or reddish purple coloration or colorless. Gnathosomal bay (157.5 - 175 (175) long; 92.5 - 115 (115) wide). Cxgl- 4 subapical. Medial suture (20 - 25 (20) long). Genital plates (160 - 185 (177.5) long; 145 - 160 (152.5) wide). Additional measurements: Cx- 1 (290 - 330 (330) long (total); 140 - 160 (155) long (medial )); Cx- 3 (365 - 410 (400) wide); anterior venter (160 - 190 (190) long). Ventral proportions: gnathosomal bay length / width 1.50 - 1.70 (1.52); anterior venter / genital field length 0.99 - 1.07 (1.07); anterior venter length / genital field width 1.08 - 1.25 (1.25); anterior venter / medial suture 7.60 - 9.50 (9.50). Male (Figure 63) (n = 5) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (500 - 540 (540) long; 350 - 370 (360) wide) ovoid with purple or reddish-purple coloration separated into anterior and posterior portions, and occasionally with faint orange medially. Anterio-medial platelets (95 - 102.5 (100) long; 37.5 - 42.5 (41.25) wide). Anterio-lateral platelets (165 - 172.5 (172.5) long; 55 - 60 (55) wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 265 - 295 (285 )). Dorsal plate proportions: dorsum length / width 1.35 - 1.53 (1.50); dorsal width / distance between Dgl- 4 1.24 - 1.32 (1.26); anterio-medial platelet length / width 2.41 - 2.56 (2.42); anterio-lateral platelet length / width 2.88 - 3.14 (3.14); anterio-lateral / anterio-medial length 1.66 - 1.74 (1.73). Gnathosoma - Subcapitulum (275 - 285 (285) long (ventral); 205 - 215 (215) long (dorsal); 102.5 - 115 (105) tall) colorless. Rostrum (105 - 112.5 (110) long; 35 - 38.75 (35) wide). Chelicerae (265 - 280 (275) long) with curved fangs (50 - 55 (52.5) long). Subcapitular proportions: ventral length / height 2.43 - 2.71 (2.71); rostrum length / width 2.90 - 3.14 (3.14). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (40 - 47.5 (40) long); femur (105 - 107.5 (107.5) long); genu (62.5 - 66.25 (65) long); tibia 77.5 - 85 (77.5) long; 22.5 - 25 (22.5) wide); tarsus (17.5 - 20 (20) long). Palpomere proportions: femur / genu 1.58 - 1.72 (1.65); tibia / femur 0.72 - 0.79 (0.72); tibia length / width 3.25 - 3.44 (3.44). Venter - (640 - 660 (655) long; 440 - 470 (460) wide) with faint bluish-purple or reddish purple coloration or colorless. Gnathosomal bay (125 - 135 (130) long; 75 - 82.5 (82.5) wide). Cxgl- 4 subapical. Medial suture (120 - 135 (125) long). Genital plates (130 - 137.5 (135) long; 85 - 90 (87.5) wide). Additional measurements: Cx- 1 (260 - 275 (275) long (total); 135 - 145 (145) long (medial )); Cx- 3 (330 - 350 (335) wide); anterior venter (277.5 - 285 (285) long). Ventral proportions: gnathosomal bay length / width 1.56 - 1.80 (1.58); anterior venter / genital field length 2.04 - 2.19 (2.11); anterior venter length / genital field width 3.11 - 3.35 (3.26); anterior venter / medial suture 2.06 - 2.33 (2.28). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
63B364F0307B6BFE78A59B64434E4215.taxon	etymology	Etymology. Specific epithet (dunni) named in honor of Rob Dunn of North Carolina State University, for his exceptional writings and research that personalize ecology by bringing nature indoors; and particularly for his storytelling ability, in which he wonderfully conveys that we humans, rather than being separate from nature, are indeed just as wild as what we perceive outdoors-a sentiment exemplified by his book, Wild Life of Our Bodies: Predators, Parasites, and Partners That Shape Who We Are Today (2011).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
63B364F0307B6BFE78A59B64434E4215.taxon	distribution	Distribution. Southeastern Appalachians (Figure 61).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
8427BE98437F2F0F98DAA453CA33D221.taxon	materials_examined	Material examined. Type series. HOLOTYPE (♀): from USA, California, Nevada County, north of Lake Tahoe, Martis Creek, Jun 1933, by PR Needham, RM 330010. PARATYPES (1 ♀; 0 ♂): California, USA: 1 ♀ from Nevada County, north of Lake Tahoe, Martis Creek, Jun 1933, by PR Needham, RM 330010. OTHER MATERIAL (94 ♀; 94 ♂): Alberta, Canada: 1 ♀ and 2 ♂ from Waterton Lakes National Park, Cameron Creek, beside Akamina Parkway (49 ° 5 ' N, 113 ° 52 ' W), 2 Aug 1985, by IM Smith, IMS 850133 * 2 ♀ and 2 ♂ from Waterton Lakes National Park, Cameron Creek, beside Akamina Parkway, west of Rowe Creek, 27 Jun 1980, by IM Smith, IMS 800086 A * 1 ♀ and 2 ♂ from Waterton Lakes National Park, Cameron Creek, beside Akamina Parkway, 12 - 15 Jun 1980, by IM Smith, IMS 800069 A & IMS 800069 B * 2 ♀ and 2 ♂ from Waterton Lakes National Park, Little Prairie Picnic Area, Cameron Creek, 5 - 9 Jun 1980, by IM Smith, IMS 800053 B * British Columbia, Canada: 1 ♀ and 1 ♂ from Bella Coola Valley, Tweedsmuir Provincial Park, Atnarko River, at campground, 28 Jul 1983, by IM Smith & AB Smith, IMS 830054 A * 1 ♀ and 1 ♂ from Bella Coola Valley, Tweedsmuir Provincial Park, Atnarko Slough, beside Highway 20, west of Youngs Creek Picnic Area, 4 Aug 1983, by IM Smith, IMS 830064 * 1 ♀ from Bella Coola Valley, Tweedsmuir Provincial Park, Belarko, Atnarko River, 24 - 26 Jul 1983, by IM Smith, IMS 830049 B * 1 ♀ and 2 ♂ from Bella Coola Valley, Tweedsmuir Provincial Park, Hotnarko River, at end of Atnarko tote road, 27 Jul 1983, by IM Smith, IMS 830052 * 2 ♀ and 3 ♂ from Bella Coola Valley, Tweedsmuir Provincial Park, Hotnarko River, at end of Atnarko tote road, 31 Jul 1983, by IM Smith, IMS 830059 A & IMS 830059 B * 3 ♂ from Bella Coola Valley, Tweedsmuir Provincial Park, Youngs Creek Picnic Area, Atnarko Slough, 24 - 27 Jul 1983, by IM Smith, IMS 830048 A * 2 ♀ and 1 ♂ from Fernie, Lizard Creek, beside Highway 3, 1.8 km west of Fernie Mountain Provincial Park, 16 Aug 2012, by IM Smith, IMS 120073 * 1 ♀ and 1 ♂ from Tweedsmuir Provincial Park, Youngs Creek, beside Highway 20, between Heckman Pass & Bella Coola Valley, 5 Aug 1983, by IM Smith, IMS 830065 * 2 ♀ and 2 ♂ from Vancouver Island, Caycuse, Nixon Creek, 8 Jul 1976, by IM Smith, IMS 760197 & IMS 760198 * 1 ♀ and 2 ♂ from Vancouver Island, beside Highway 4, 35.6 kilometers east of Pacific Rim Road, 9 Jul 1976, by IM Smith, IMS 760206 * 1 ♀ and 1 ♂ from Vancouver Island, Lake Cowichan, Cowichan River, above Skutz Falls, 9 Jul 1979, by IM Smith, IMS 790035 * 1 ♀ and 1 ♂ from Vancouver Island, Lake Cowichan, Skutz Falls, Skutz Creek, near Cowichan River, 9 Jul 1979, by IM Smith, IMS 790036 A & IMS 790036 B * 1 ♀ and 2 ♂ from Vancouver Island, Lake Cowichan, South Shore Road, north of Mesachie Lake, Robertson River, 4 Jul 1976, by IM Smith, IMS 760183 A * 1 ♀ from Vancouver Island, Lake Cowichan, South Shore Road, north of Mesachie Lake, tributary of Robertson River, 4 - 10 Jul 1976, by IM Smith, IMS 760182 * 1 ♀ and 2 ♂ from Vancouver Island, Lost Shoe Creek, beside Highway 4, 1.3 kilometers east of Pacific Rim Road, 9 Jul 1976, by IM Smith, IMS 760202 * 1 ♀ and 2 ♂ from Vancouver Island, Malahat, Goldstream Provincial Park, Goldstream River, 26 Jun 1979, by IM Smith, IMS 790028 A * 1 ♂ from Vancouver Island, spring run beside North Shore Road, 1.7 kilometers north of Lake Cowichan, 7 Jun 1979, by IM Smith, IMS 790008 A * 3 ♀ and 2 ♂ from Vancouver Island, North Shore Road, 3.2 kilometers south of Youbou, 4 Jul 1976, by IM Smith, IMS 760190 * 1 ♀ from Vancouver Island, Youbou, Shaw Creek, North Shore Road, 4.3 kilometers south of north end of Cowichan Lake, 8 Jul 1976, by IM Smith, IMS 760196 * 1 ♀ from Vancouver Island, spring run beside South Shore Road, 2.3 kilometers north of Lake Cowichan, 6 Jun 1979, by IM Smith, IMS 790007 * 1 ♀ and 1 ♂ from Vancouver Island, Ucluelet, beside Highway 4, 16.6 kilometers east of Pacific Rim Road, 18 - 19 Jul 1979, by IM Smith, IMS 790047 * California, USA: 1 ♀ and 1 ♂ from Humboldt County, Prairie Creek State Park, Prairie Creek, 12 Jul 1964, by H Habeeb, HH 640021 * 1 ♀ from Humboldt County, Trinidad, stream beside Patrick Point Road, near Bishop Pine Lodge, 7 Aug 1987, by IM Smith, IMS 870134 * 1 ♀ from Inyo County, Inyo National Forest, Bishop Creek (37 ° 17 ' 23 " N, 118 ° 33 ' 14 " W), 2 Sep 2013, by JR Fisher, JRF 13 - 0902 - 003 * 1 ♂ from Mendocino County, Cottaneva Creek, beside Route 1, 21.8 kilometers southwest of Route 101, 5 Aug 1987, by IM Smith, IMS 870129 A * 2 ♀ and 3 ♂ from Mendocino County, small stream at beach access road, off Route 1, 2.6 kilometers south of Westport, 5 Aug 1987, by IM Smith, IMS 870128 A * 1 ♀ from Mono County, Humboldt-Toiyabe National Forest, Leavitt Creek (38 ° 18 ' 40 " N, 119 ° 34 ' 49 " W), 31 Aug 2013, by JR Fisher, JRF 13 - 0831 - 004 * 1 ♂ from Monterey County, Los Padres National Forest, Salmon Creek (35 ° 48 ' 57 " N, 121 ° 21 ' 29 " W), 6 Sep 2013, by JR Fisher, JRF 13 - 0906 - 003 * 2 ♀ and 2 ♂ from Monterey County, Nacimiento River, beside Nacimiento-Ferguson Road at Nacimiento campground, 30 Jul 1987, by IM Smith, IMS 870120 A * 2 ♀ and 2 ♂ from Monterey County, Salmon Creek, beside Route 1, south of Gorda, 28 Jul 1987, by IM Smith, IMS 870114 A * 1 ♀ from Nevada County, beside Route 89, north of Hobart Mills, 13 Jun 1976, by IM Smith, IMS 760109 * 1 ♀ and 2 ♂ from San Bernardino County, Claremont, Mt Baldy, 3.5 kilometers above Mt Baldy Village, 24 Jul 1987, by IM Smith, IMS 870107 * 1 ♀ from San Bernardino County, Claremont, Mt Baldy, San Antonio Falls, above Monker Flats, 24 Jul 1987, by IM Smith, IMS 870105 * 3 ♀ and 2 ♂ from San Bernardino County, Claremont, Mt Baldy, stream below San Antonio Falls, above Monker Flats, 24 Jul 1987, by IM Smith, IMS 870106 * 2 ♀ and 2 ♂ from Shasta County, Battle Creek, beside Route 44, 5.6 kilometers west of Viola, 10 Aug 1987, by IM Smith, IMS 870139 A * 1 ♀ and 1 ♂ from Trinity County, small cascading trickle beside Route 36, 5.2 kilometers west of Forest Glen Station, 6 Aug 1987, by IM Smith, IMS 870132 * 3 ♂ from Tulare County, Stony Creek at Stony Creek Picnic Area, east of Sequoia National Park, 1 Aug 1987, by IM Smith, IMS 870124 A * 1 ♀ and 1 ♂ from Ventura County, Ojai, North Fork of Ventura River, beside Route 33, just above Wheeler Gorge, 25 - 26 Jul 1987, by IM Smith, IMS 870109 A * Idaho, USA: 1 ♀ from Custer County, Challis National Forest, Stanley Creek (44 ° 15 ' 12 " N, 115 ° 0 ' 19 " W), 30 Jul 2012, by JR Fisher, WA Nelson, & JC O'Neill, ROW 12 - 0730 - 005 * 1 ♂ from Custer County, Salmon River (44 ° 12 ' 31 " N, 114 ° 55 ' 51 " W), 29 Jul 2012, by JR Fisher, WA Nelson, & JC O'Neill, ROW 12 - 0729 - 003 * 2 ♀ and 3 ♂ from Lemhi County, North Fork of Salmon River, beside Route 93, 15 kilometers north of North Fork, 1 Jul 1985, by IM Smith, IMS 850062 * Montana, USA: 2 ♀ from Flathead County, stream beside Route 487 near north end of Whitefish Lake, 29 Jun 1985, by IM Smith, IMS 850057 * 3 ♀ and 4 ♂ from Lake County, stream beside Route 83, 39.5 kilometers north of Condon, 30 Jun 1985, by IM Smith, IMS 850059 A * 1 ♀ from Missoula County, Lolo National Forest, Lolo Creek (46 ° 46 ' 7 " N, 114 ° 27 ' 53 " W), 7 Aug 2012, by JR Fisher, WA Nelson, & JC O'Neill, ROW 12 - 0807 - 003 * 1 ♂ from Ravalli County, Bitterroot National Forest, East Fork Bitterroot River (45 ° 51 ' 40 " N, 114 ° 1 ' 46 " W), 3 Aug 2012, by JR Fisher, WA Nelson, & JC O'Neill, ROW 12 - 0803 - 005 * Oregon, USA: 1 ♀ and 1 ♂ from Benton County, Marys Peak near Philomath, Parker Creek, 27 - 28 Jun 1983, by IM Smith & AB Smith, IMS 830006 * 1 ♀ from Clackamas County, Rhododendron Pioneer Tollgate campground, Zigzag River, 27 Jun 1976, by IM Smith, IMS 760164 * 1 ♀ and 2 ♂ from Curry County, Port Orford, Butler Bar campground, Elk River, 25 Jun 1976, by IM Smith, IMS 760162 * 1 ♂ from Coos County, Siskiyou National Forest, Road 33 between Powers & Agness, Daphne Grove campground, Coquille River, 2 Jul 1983, by IM Smith, IMS 830016 * 1 ♀ and 1 ♂ from Coos County, Siskiyou National Forest, Road 33 between Powers & Agness, Daphne Grove campground, 2 Jul 1983, by IM Smith, IMS 830017 * 2 ♀ and 5 ♂ from Curry County, Port Orford, Humbug Mountain State Park Picnic Area, beside Route 1, Brush Creek, 1 Jul 1983, by IM Smith, IMS 830012 * 4 ♀ and 4 ♂ from Curry County, Port Orford, Humbug Mountain State Park Picnic Area, Brush Creek, beside Route 1, 3 Jul 1983, by IM Smith, IMS 830020 A & IMS 830020 B * 1 ♀ from Curry County, Port Orford, Humbug Mountain State Park Picnic Area, beside Route 1, 1 Jul 1983, by IM Smith, IMS 830013 * 1 ♀ from Curry County, Rogue River National Forest, Elk River (42 ° 42 ' 46 " N, 124 ° 18 ' 41 " W), 13 Aug 2013, by JR Fisher, JRF 13 - 0813 - 003 * 2 ♀ and 2 ♂ from Curry County, Siskiyou National Forest, road 33 between Powers and Agness, North Fork of Foster Creek, 2 Jul 1983, by IM Smith, IMS 830019 * 1 ♀ from Curry County, Sixes, Sixes River, beside road at mouth of Edson Creek, 4 Jul 1983, by IM Smith, IMS 830021 A * 1 ♀ from Grant County, Prairie City, Lunch Creek, beside Route 26. east of Dixie Pass, 17 - 20 Jun 1976, by IM Smith, IMS 760125 * 1 ♀ and 1 ♂ from Grant County, Prairie City, Strawberry Forest Camp, Strawberry Creek, 17 - 20 Jun 1976, by IM Smith, IMS 760126 * 2 ♀ from Lane County, Gate Creek (44 ° 8 ' 48 " N, 122 ° 34 ' 20 " W), 11 Aug 2013, by JC O'Neill, & WA Nelson, JNOW 13 - 0811 - 001 * 3 ♀ and 1 ♂ from Marion County, Marion Forks Riverside campground, North Fork of Santiam River, 22 Jun 1976, by IM Smith, IMS 760145 * 3 ♀ and 4 ♂ from Multnomah County, Columbia River Scenic Highway, Horsetail Falls, 27 Jun 1983, by IM Smith, IMS 830005 * 1 ♂ from Tillamook County, Siuslaw National Forest, Alder Creek (45 ° 9 ' 27 " N, 123 ° 47 ' 60 " W), 6 Aug 2013, by JC O'Neill, & WA Nelson, JNOW 13 - 0806 - 002 * Washington, USA: 2 ♀ from Lewis County, Gifford Pinchot National Forest (46 ° 39 ' 49 " N, 121 ° 41 ' 11 " W), 23 Jul 2013, by JC O'Neill, & WA Nelson, JNOW 13 - 0723 - 005 * 2 ♀ from Mason County, Olympic National Forest, Cabin Creek (47 ° 35 ' 44 " N, 123 ° 7 ' 39 " W), 22 Jul 2013, by JC O'Neill, & WA Nelson, JNOW 13 - 0722 - 004 * 2 ♀ from Snohomish County, Mount Baker National Forest, Marten River (48 ° 4 ' 19 " N, 121 ° 36 ' 24 " W), 28 Jul 2013, by JC O'Neill, & WA Nelson, JNOW 13 - 0728 - 002 * Wyoming, USA: 1 ♂ from Johnson County, Bighorn Mountains, Clear Creek, west of Buffalo Mosier Gulch Picnic Area, 28 Jul 2012, by IM Smith, IMS 120041 * 1 ♀ and 1 ♂ from Johnson County, Crazy Woman Creek, beside Route 16, 14.9 kilometers west of road to South Fork campground, 18 Aug 1987, by IM Smith, IMS 870158 A * 1 ♀ and 2 ♂ from Washakie County, Ten Sleep, Ten Sleep Creek, beside Route 16, 4 kilometers west of Route 435, 18 Aug 1987, by IM Smith, IMS 870157 * 2 ♀ and 1 ♂ from Washakie County, Ten Sleep Creek, Ten Sleep Wigwam Rearing Station, 26 Jul 2012, by IM Smith, IMS 120044.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
8427BE98437F2F0F98DAA453CA33D221.taxon	diagnosis	Diagnosis. Torrenticola ellipsoidalis are similar to other members of the Ellipsoidalis Group (T. multiforma, T. occidentalis, and T. leviathan), in being among the largest Torrenticola in the west (dorsum length ♀ = 700 - 885; ♂ = 665 - 850), although T. sierrensis are also large (dorsum length ♀ = 700 - 880; ♂ = 590 - 735) but can easily be distinguished from the Ellipsoidalis Group by being circular instead of ellipsoid or rectangular (dorsum length / width = 1.17 - 1.28 in T. sierrensis, 1.30 - 1.67 in Ellipsoidalis Group). T. ellipsoidalis can be differentiated from T. multiforma by having stockier subcapitular rostra (length / width = 1.8 - 2.1 in T. ellipsoidalis, 2.5 - 2.8 in T. multiforma). T. ellipsoidalis can be differentiated from T. occidentalis (only known from females) by having a longer medial suture (40 - 57.5 in T. ellipsoidalis, 20 in T. occidentalis) and by having stockier anterio-lateral platelets (length / width = 2.00 - 2.39 in T. ellipsoidalis, 2.54 in T. occidentalis). T. ellipsoidalis can be differentiated from T. leviathan by having less elongate pedipalpal tibiae (length / width = 2.6 - 3.3 in T. ellipsoidalis, 3.4 - 4.2 in T. leviathan) and stockier anterio-medial platelets (length / width ♀ = 1.43 - 1.72 in T. ellipsoidalis, 1.94 - 2.14 in T. leviathan; ♂ = 1.53 - 2.00 in T. ellipsoidalis, 2.15 in T. leviathan).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
8427BE98437F2F0F98DAA453CA33D221.taxon	description	Re-description. Female (Figure 65) (n = 8) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (765 - 885 (800) long; 520 - 605 (540) wide) rectangular and usually colorless, occasionally with faint purple coloration without distinct pattern. Anterio-medial platelets (127.5 - 147.5 137.5) long; 80 - 97.5 (80) wide). Anterio-lateral platelets (207.5 - 235 (217.5) long; 90 - 115 (92.5) wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 390 - 470 (410 )). Dorsal plate proportions: dorsum length / width 1.41 - 1.64 (1.48); dorsal width / distance between Dgl- 4 1.27 - 1.40 (1.32); anterio-medial platelet length / width 1.43 - 1.72 (1.72); anterio-lateral platelet length / width 2.00 - 2.39 (2.35); anterio-lateral / anterio-medial length 1.48 - 1.77 (1.58). Gnathosoma - Subcapitulum (285 - 315 (310) long (ventral); 194 - 219 (215) long (dorsal); 145 - 165 (165) tall) colorless. Rostrum (115 - 127.5 (117.5) long; 57.5 - 62.5 (60) wide) short and conical. Chelicerae (261 - 289 long) with curved fangs (61 - 74 long). Subcapitular proportions: ventral length / height 1.82 - 2.07 (1.88); rostrum length / width 1.84 - 2.09 (1.96). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (37.5 - 42.5 (40) long); femur (101.25 - 107.5 (102.5) long); genu (70 - 77.5 (77.5) long); tibia (80 - 90 (85) long; 26.25 - 27.5 (27.5) wide); tarsus (17.5 - 21.25 (20) long). Palpomere proportions: femur / genu 1.32 - 1.46 (1.32); tibia / femur 0.76 - 0.85 (0.83); tibia length / width 2.91 - 3.27 (3.09). Venter - (885 - 1000 (935) long; 605 - 700 (605) wide) colorless. Gnathosomal bay (157.5 - 180 (177.5) long; 80 - 105 (85) wide). Cxgl- 4 subapical. Medial suture (40 - 57.5 (47.5) long). Genital plates (201.25 - 222.5 (205) long; 167.5 - 195 (172.5) wide). Additional measurements: Cx- 1 (308 - 337.5 (335) long (total); 122 - 162.5 (162.5) long (medial )); Cx- 3 (393 - 440 (405) wide); anterior venter (210 - 237.5 (225) long). Ventral proportions: gnathosomal bay length / width 1.50 - 2.12 (2.09); anterior venter / genital field length 1.01 - 1.13 (1.10); anterior venter length / genital field width 1.13 - 1.30 (1.30); anterior venter / medial suture 3.83 - 5.94 (4.74). Male (Figure 66) (n = 6) with characters of the genus with following specifications. Dorsum - (725 - 850 long; 450 - 565 wide) rectangular and usually colorless, occasionally with faint purple coloration without distinct pattern. Anterio-medial platelets (122.5 - 165 long; 72.5 - 95 wide). Anterio-lateral platelets (195 - 230 long; 85 - 107.5 wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 350 - 460). Dorsal plate proportions: dorsum length / width 1.38 - 1.67; dorsal width / distance between Dgl- 4 1.22 - 1.43; anterio-medial platelet length / width 1.53 - 2.00; anterio-lateral platelet length / width 2.07 - 2.36; anterio-lateral / anterio-medial length 1.39 - 1.67. Gnathosoma - Subcapitulum (280 - 290 long (ventral); 196 - 203.75 long (dorsal); 138.75 - 155 tall) colorless. Rostrum (102.5 - 115 long; 52.5 - 60 wide) short and conical. Chelicerae (263 - 280 long) with curved fangs (60 - 74 long). Subcapitular proportions: ventral length / height 1.87 - 2.04; rostrum length / width 1.86 - 2.02. Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (35 - 41.25 long); femur 92.5 - 100 long); genu (65 - 72.5 long); tibia (72.5 - 80 long; 25 - 30 wide); tarsus (17.5 - 20 long). Palpomere proportions: femur / genu 1.31 - 1.46; tibia / femur 0.74 - 0.84; tibia length / width 2.64 - 3.1. Venter - (840 - 980 long; 469 - 653 wide) colorless. Gnathosomal bay (147.5 - 177.5 long; 77.5 - 90 wide). Cxgl- 4 subapical. Medial suture (70 - 90 long). Genital plates (177.5 - 236.25 long; 131.25 - 162.5 wide). Additional measurements: Cx- 1 (283 - 345 long (total); 117 - 167.5 long (medial )); Cx- 3 (348 - 432.5 wide); anterior venter (245 - 270 long). Ventral proportions: gnathosomal bay length / width 1.69 - 2.15; anterior venter / genital field length 1.14 - 1.44; anterior venter length / genital field width 1.66 - 1.96; anterior venter / medial suture 2.72 - 3.71. Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
8427BE98437F2F0F98DAA453CA33D221.taxon	etymology	Etymology. Marshall (1943) presumably named the specific epithet (ellipsoidalis) after the elongate body of this species, as she wrote, " the body is an ellipse. "	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
8427BE98437F2F0F98DAA453CA33D221.taxon	distribution	Distribution. Western (Figure 64). T. ellipsoidalis was previously recorded from Martis Creek and Gibbon River, Wyoming (Marshall 1943); and from Torch River, Saskatchewan (Quaglia & Conroy 1984). We expand the range into most of western North America. However, T. ellipsoidalis is not known from the southwest.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
273AF2EABE1CDCA5B1425560FC3B0666.taxon	materials_examined	Material examined. HOLOTYPE (♀): from USA, Mississippi, Tishomingo County, Tishomingo State Park, Rock Quarry Branch (34 ° 36 ' N, 88 ° 11 ' W), 18 Sep 1991, by IM Smith, IMS 910049. PARATYPES (1 ♀; 2 ♂): Mississippi, USA: 1 ♂ (ALLOTYPE) from Tishomingo County, Tishomingo State Park, Rock Quarry Branch (34 ° 36 ' 43 " N, 88 ° 12 ' 4 " W), 20 Sep 2009, by IM Smith, IMS 090115, DNA 1593 * 1 ♀ and 1 ♂ from Tishomingo County, Tishomingo State Park, Rock Quarry Branch (34 ° 36 ' 43 " N, 88 ° 12 ' 4 " W), 20 Sep 2009, by IM Smith, IMS 090115.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
273AF2EABE1CDCA5B1425560FC3B0666.taxon	diagnosis	Diagnosis. Torrenticola elongata are similar to species with similar dorsal patterning, such as the Rusetria " 4 - Plate " group (T. dunni, T. glomerabilis, T. kittatinniana, T. pollani, T. rufoalba and T. shubini), Neoanomala Group (T. interiorensis and T. neoanomala), and T. bondi, T. erectirostra, T. robisoni, T. gorti, T. reduncarostra, T. irapalpa, T. racupalpa, T. skvarlai, and T. arktonyx. T. elongata can be differentiated from all other Torrenticola with similar dorsal patterning by having a more elongate dorsum (length / width ♀ = 1.92 - 2.08 in T. elongata, 1.17 - 1.58 in others; ♂ = 1.70 - 1.70 in T. elongata, 1.20 - 1.68 in others). Additionally, they can be differentiated from Rusetria 4 - Plates and T. skvarlai by having distinct hind coxal margins, they can be differentiated from T. erectirostra and T. robisoni by having a straight, anteriorly-directed rostrum (upturned in others), and they can be differentiated from T. arktonyx by having an unmodified dorsal plate (T. arktonyx has distinctive longitudinal dark markings on the anterior portion of the dorsal plate that fade posteriorly).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
273AF2EABE1CDCA5B1425560FC3B0666.taxon	description	Description. Female (Figure 68) (n = 2) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (540 - 565 (540) long; 260 - 295 (260) wide) ovoid and elongate with purple coloration separated into anterior and posterior portions. Anterio-medial platelets (105 - 105 (105) long; 47.5 - 50 (47.5) wide). Anterio-lateral platelets (150 - 157.5 (150) long; 42.5 - 45 (45) wide) free from dorsal plate. Dgl- 4 closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 180 - 200 (180 )). Dorsal plate proportions: dorsum length / width 1.92 - 2.08 (1.92); dorsal width / distance between Dgl- 4 1.44 - 1.48 (1.48); anterio-medial platelet length / width 2.10 - 2.21 (2.21); anterio-lateral platelet length / width 3.33 - 3.71 (3.33); anterio-lateral / anterio-medial length 1.43 - 1.50 (1.43). Gnathosoma - Subcapitulum (285 - 295 (285) long (ventral); 210 - 222 (210) long (dorsal); 101.25 - 102.5 (101.25) tall) colorless. Rostrum (115 - 117.5 (117.5) long; 32.5 - 36.25 (32.5) wide) elongate. Chelicerae (( 290) long) with curved fangs (45 - 47.5 (45) long). Subcapitular proportions: ventral length / height 2.81 - 2.88 (2.81); rostrum length / width 3.24 - 3.54 (3.24). Pedipalps with tuberculate ventral extensions on femora and on genua. Palpomeres: trochanter (35 - 37.5 (37.5) long); femur (96.25 - 100 (96.25) long); genu (55 - 57.5 (55) long); tibia (61.25 - 62.5 (61.25) long; 20 - 21.25 (20) wide); tarsus (15 - 17.5 (17.5) long). Palpomere proportions: femur / genu 1.74 - 1.75 (1.75); tibia / femur 0.63 - 0.64 (0.64); tibia length / width 2.94 - 3.06 (3.06). Venter - (690 - 690 (690) long; 300 - 350 (300) wide) colorless. Gnathosomal bay (130 - 135 (130) long; 56.25 - 60 (56.25) wide). Cxgl- 4 subapical. Medial suture (52.5 - 60 (60) long). Genital plates (142.5 - 145 (142.5) long; 120 - 122.5 (120) wide). Additional measurements: Cx- 1 (250 - 270 (250) long (total); 120 - 125 (120) long (medial )); Cx- 3 (240 - 258 (240) wide); anterior venter (205 - 207.5 (205) long). Ventral proportions: gnathosomal bay length / width 2.25 - 2.31 (2.31); anterior venter / genital field length 1.43 - 1.44 (1.44); anterior venter length / genital field width 1.69 - 1.71 (1.71); anterior venter / medial suture 3.42 - 3.95 (3.42). Male (Figure 69) (n = 2) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (450 - 460 (460) long; 265 - 270 (270) wide) ovoid and elongate with purple coloration separated into anterior and posterior portions. Anterio-medial platelets (92.5 - 100 (100) long; 45 - 47.5 (45) wide). Anterio-lateral platelets (130 - 142.5 (130) long; 43.75 - 45 (43.75) wide) free from dorsal plate. Dgl- 4 closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 175 - 180 (180 )). Dorsal plate proportions: dorsum length / width 1.70 - 1.70 (1.70); dorsal width / distance between Dgl- 4 1.50 - 1.51 (1.50); anterio-medial platelet length / width 1.95 - 2.22 (2.22); anterio-lateral platelet length / width 2.97 - 3.17 (2.97); anterio-lateral / anterio-medial length 1.30 - 1.54 (1.30). Gnathosoma - Subcapitulum (250 - 255 (255) long (ventral); 184 - 187 (184) long (dorsal); 85 - 87.5 (85) tall) colorless. Rostrum (97.5 - 102.5 (102.5) long; 30 - 30 (30) wide) elongate. Chelicerae (243 - 262 (243) long) with curved fangs (34 - 42 (42) long). Subcapitular proportions: ventral length / height 2.83 - 3.00 (3.00); rostrum length / width 3.25 - 3.42 (3.42). Pedipalps with tuberculate ventral extensions on femora and on genua. Palpomeres: trochanter (31.25 - 37.5 (31.25) long); femur (83.75 - 85 (83.75) long); genu (47.5 - 47.5 (47.5) long); tibia (57.5 - 60 (57.5) long; 18.75 - 20 (20) wide); tarsus (15 - 15 (15) long). Palpomere proportions: femur / genu 1.76 - 1.79 (1.76); tibia / femur 0.69 - 0.71 (0.69); tibia length / width 2.88 - 3.20 (2.88). Venter - (565 - 570 (570) long; 325 - 329 (329) wide) colorless. Gnathosomal bay (105 - 110 (105) long; 50 - 55 (55) wide). Cxgl- 4 subapical. Medial suture (55 - 80 (55) long). Genital plates (107.5 - 113.75 (113.75) long; 92.5 - 92.5 (92.5) wide). Additional measurements: Cx- 1 (207 - 232 (231) long (total); 82 - 98 (98) long (medial )); Cx- 3 (257 - 266 (266) wide); anterior venter (215 - 220 (215) long). Ventral proportions: gnathosomal bay length / width 1.91 - 2.20 (1.91); anterior venter / genital field length 1.89 - 2.05 (1.89); anterior venter length / genital field width 2.32 - 2.38 (2.32); anterior venter / medial suture 2.75 - 3.91 (3.91). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
273AF2EABE1CDCA5B1425560FC3B0666.taxon	etymology	Etymology. Specific epithet (elongata) refers to elongated bodies of this species, which is more pronounced than in all other North American Torrenticola (elongatus, L. prolonged).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
273AF2EABE1CDCA5B1425560FC3B0666.taxon	distribution	Distribution. Known only from Tishomingo County, Mississippi (Figure 67).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
2196443FDADF7B9F6993A16D2E6358B2.taxon	materials_examined	Material examined. HOLOTYPE (♀): from Canada, New Brunswick, Charlotte County, Rollingham, Whittier Ridge, Highway 770, 6.6 km east of covered bridge, 3 Oct 2011, by IM Smith, IMS 110120, DNA 1857.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
2196443FDADF7B9F6993A16D2E6358B2.taxon	diagnosis	Diagnosis. Torrenticola elusiva are similar to other members of the Raptor Group (T. gnoma, T. irapalpa, T. longitibia, T. mjolniri, T. racupalpa, T. raptor, T. danielleae, T. daemon, and T. ivyae) in having round bodies; Dgl- 4 close to muscles scars; long, thin subcapitular rostra; and long, thin pedipalp tibiae. T. elusiva can be differentiated from T. racupalpa by having a stockier subcapitulum (ventral length / height = 2.39 in T. elusiva, 2.48 - 2.73 in T. racupalpa); and by dorsal pattern. T. elusiva can be differentiated from T. irapalpa and T. daemon by having Dgl- 4 closer to the muscle scars (dorsal width / distance between Dgl- 4 = 2.5 in T. elusiva, 1.59 - 2.09 in others); a more elongate rostrum (length / width = 3.65 in T. elusiva, 2.66 - 3.39 in others); and by dorsal coloration and pattern. T. elusiva can be differentiated from T. gnoma by being larger (dorsum length = 645 in T. elusiva, 540 - 595 in T. gnoma); having a more elongate rostrum (length / width = 3.65 in T. elusiva, 2.74 - 3.13 in T. gnoma); and dorsal coloration. T. elusiva can be differentiated from T. mjolniri and T. ivyae by having stockier pedipalp tibiae (length / width = 4.42 in T. elusiva, 5.00 - 6.00 in others); and a stockier rostrum (length / width = 3.65 in T. elusiva, 3.81 - 4.32 in others). T. elusiva can be differentiated from T. raptor by having Dgl- 4 closer to the muscle scars (dorsal width / distance between Dgl- 4 = 2.50 in T. elusiva, 1.8 - 2.02 in T. raptor); shorter anterior venter (163.75 in T. elusiva, 205 - 240 in T. raptor); and stockier pedipalp tibiae (length / width = 4.42 in T. elusiva, 6 - 7.54 in T. raptor). T. elusiva can be differentiated from T. danielleae by having Dgl- 4 closer to the muscle scars (dorsal width / distance between Dgl- 4 = 2.5 in T. elusiva, 1.57 - 1.70 in T. danielleae) and by dorsal coloration. T. elusiva cannot be confidently differentiated from T. longitibia because T. elusiva is only known from a single female and T. longitibia is only known from two males; however, T. elusiva is only known from Charlotte County, New Brunswick, whereas T. longitibia is only known from Monroe County, Tennessee. Additionally, two character systems that vary minimally between sexes are rostrum and pedipalp tibiae proportions, which do differ between T. elusiva and T. longitibia as follows: pedipalp tibia stockier (4.42 in T. elusiva, 5.5 - 5.5 in T. longitibia) and rostrum stockier (3.65 in T. elusiva, 4.15 - 4.23 in T. longitibia).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
2196443FDADF7B9F6993A16D2E6358B2.taxon	description	Description. Female (Figure 71) (n = 1) (holotype only) with characters of the genus with following specifications. Dorsum - (645 long; 500 wide) circular with bluish-purple coloration posteriorly with a broad anterior extension reaching the anterior edge of the dorsal plate. Anterio-medial platelets (152.5 long; 70 wide). Anterio-lateral platelets (182.5 long; 87.5 wide) free from dorsal plate. Dgl- 4 much closer to the muscle scars than to edge of dorsum (distance between Dgl- 4 200). Dorsal plate proportions: dorsum length / width 1.29; dorsal width / distance between Dgl- 4 2.50; anterio-medial platelet length / width 2.18; anterio-lateral platelet length / width 2.09; anterio-lateral / anterio-medial length 1.20. Gnathosoma - Subcapitulum (340 long (ventral); 259 long (dorsal); 142.5 tall) colorless. Rostrum (155 long; 42.5 wide) elongate. Chelicerae (333 long) with curved fangs (59 long). Subcapitular proportions: ventral length / height 2.39; rostrum length / width 3.65. Pedipalps elongate (especially tibiae) with tuberculate ventral extensions with dentate tip on femora and tuberculate ventral extensions on genua. Palpomeres: trochanter (48.75 long); femur (132.5 long); genu (72.5 long); tibia (105 long; 23.75 wide); tarsus (17.5 long). Palpomere proportions: femur / genu 1.83; tibia / femur 0.79; tibia length / width 4.42. Venter - (730 long; 554 wide) colorless. Gnathosomal bay (176.25 long; 87.5 wide). Cxgl- 4 subapical. Medial suture (17.5 long). Genital plates (167.5 long; 150 wide). Additional measurements: Cx- 1 (288 long (total); 115 long (medial )); Cx- 3 (384 wide); anterior venter (163.75 long). Ventral proportions: gnathosomal bay length / width 2.01; anterior venter / genital field length 0.98; anterior venter length / genital field width 1.09; anterior venter / medial suture 9.36. Male unknown. Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
2196443FDADF7B9F6993A16D2E6358B2.taxon	etymology	Etymology. Specific epithet (elusiva) refers to the fact that we were only able to find a single specimen of this species, despite extensive searching among the abundant samples taken from the type locality in New Brunswick.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
2196443FDADF7B9F6993A16D2E6358B2.taxon	distribution	Distribution. Known only from Charlotte County, New Brunswick (Figure 70).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
1B8D1B082E8E51399530882E406DBC56.taxon	materials_examined	Material examined. HOLOTYPE (♀): from Canada, New Brunswick, York County, Stanley, Nashwaak River, Stanley Municipal Park, 19 Jun 2012, by IM Smith, IMS 120031, DNA 2962. PARATYPES (9 ♀; 9 ♂): New Brunswick, Canada: 3 ♂ from Charlotte County, Rollingham, Digdeguash River, beside Highway 770 at covered bridge, 30 Jun 1989, by IM Smith, IMS 890053 * 1 ♀ and 2 ♂ from Charlotte County, Digdeguash River, beside Sorrel Ridge Road west of Whittier Road, 10 Jun 2012, by IM Smith, IMS 120015 * 4 ♀ and 4 ♂ from York County, Magaguadavic River, beside Highway 3 just east of Thomaston Corners, 1 Jul 1989, by IM Smith, IMS 890055 A * 1 ♂ (ALLOTYPE) from York County, Stanley, Davis Brook, beside Highway 3, 3.5 km south of Highway 4 at Thomaston Corner, 11 Jun 2012, by IM Smith, IMS 120017, DNA 2964 * 2 ♀ and 1 ♂ from New Brunswick, York County, Stanley, Davis Brook, beside Highway 3, 3.5 km south of Highway 4 at Thomaston Corner, 11 Jun 2012, by IM Smith, IMS 120017 * Maine, USA: 1 ♀ from Aroostook County, Ashland, beside Route 11, Aroostook River (46 ° 38 ' N 68 ° 24 ' W), 4 Jul 1989, by IM Smith, IMS 890067 * New York, USA: 1 ♂ from Cayuga County, Dutch Hollow Brook, beside Route 38 A at Niles, 22 Jul 1990, by IM Smith, IMS 900113 A * Virginia, USA: 1 ♀ from Amherst County, beside Blue Ridge, Otter Creek (37 ° 36 ' 57 " N, 79 ° 19 ' 27 " W), 7 Sep 2007, by IM Smith, IMS 070056 A.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
1B8D1B082E8E51399530882E406DBC56.taxon	diagnosis	Diagnosis. Torrenticola erectirostra are similar to other members of the Erectirostra Group (T. karambita and T. robisoni), species with similar dorsal patterning, such as Rusetria " 4 - Plate " group (T. dunni, T. glomerabilis, T. kittatinniana, T. pollani, T. rufoalba and T. shubini), Elongata Group (T. gorti and T. elongata), Neoanomala Group (T. interiorensis and T. neoanomala), T. bondi, T. irapalpa, T. racupalpa, and T. skvarlai. They can be differentiated from all other Torrenticola, except T. karambita and T. robisoni, by having a dentate, upturned rostrum that is wide when viewed ventrally. T. erectirostra can be differentiated from T. karambita by having dorsal coloration (T. karambita is colorless) and a slightly more elongate rostrum (length / width ♀ = 1.72 - 1.91 in T. erectirostra, 1.57 - 1.62 in T. karambita; ♂ = 2.0 - 2.2 in T. erectirostra, 1.6 - 1.95 in T. karambita). T. erectirostra can be differentiated from T. robisoni by having less elongate anterio-lateral platelets (length / width ♀ = 2.52 - 2.69 in T. erectirostra, 2.96 - 3.00 in T. robisoni) and by being distributed in the Appalachians, while T. robisoni is in the Interior Highlands.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
1B8D1B082E8E51399530882E406DBC56.taxon	description	Description. Female (Figure 73) (n = 5) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum- (690 - 750 (735) long; 480 - 510 (510) wide) ovoid with bluish-purple or purple coloration separated into anterior and posterior portions with orange medially. Anterio-medial platelets (150 - 165 (162.5) long; 62.5 - 75 (75) wide). Anterio-lateral platelets (195 - 225 (220) long; 77.5 - 83.75 (82.5) wide) free from dorsal plate. Dgl- 4 closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 325 - 370 (370 )). Dorsal plate proportions: dorsum length / width 1.41 - 1.47 (1.44); dorsal width / distance between Dgl- 4 1.38 - 1.51 (1.38); anterio-medial platelet length / width 2.17 - 2.48 (2.17); anterio-lateral platelet length / width 2.52 - 2.69 (2.67); anterio-lateral / anterio-medial length 1.26 - 1.36 (1.35). Gnathosoma - Subcapitulum (315 - 350 (350) long (ventral); 225 - 247.5 (247.5) long (dorsal); 130 - 140 (130) tall) colorless. Rostrum (105 - 125 125) long; 55 - 72.5 (72.5) wide) wide and upturned with dentation. Chelicerae (320 - 345 (345) long) with curved fangs (45 - 55 (45) long). Subcapitular proportions: ventral length / height 2.42 - 2.69 (2.69); rostrum length / width 1.72 - 1.91 (1.72). Pedipalps short and stocky (especially tibiae) with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (45 - 55 (50) long); femur (100 - 107.5 (102.5) long); genu (57.5 - 67.5 (57.5) long); tibia (50 - 65 (62.5) long; 27.5 - 30 (28.75) wide); tarsus (17.5 - 20 (20) long). Palpomere proportions: femur / genu 1.48 - 1.78 (1.78); tibia / femur 0.50 - 0.61 (0.61); tibia length / width 1.82 - 2.17 (2.17). Venter - (860 - 920 (900) long; 580 - 650 (650) wide) colorless. Gnathosomal bay (210 - 220 long; 105 - 150 wide). Cxgl- 4 far from apex. Medial suture (17.5 - 27.5 (22.5) long). Genital plates (187.5 - 202.5 (202.5) long; 162.5 - 180 (180) wide). Additional measurements: Cx- 1 (330 - 360 (350) long (total); 140 - 160 (160) long (medial )); Cx- 3 (410 - 460 (460) wide); anterior venter (192.5 - 220 (220) long). Ventral proportions: gnathosomal bay length / width 1.47 - 2.00; anterior venter / genital field length 0.98 - 1.09 (1.09); anterior venter length / genital field width 1.13 - 1.25 (1.22); anterior venter / medial suture 7.09 - 12.14 (9.78). Male (Figure 74) (n = 5) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (580 - 640 (620) long; 400 - 430 (400) wide) ovoid with bluish-purple or purple coloration separated into anterior and posterior portions with orange medially. Anterio-medial platelets (130 - 150 (138.75) long; 52.5 - 58.75 (58.75) wide). Anterio-lateral platelets (187.5 - 205 (205) long; 62.5 - 70 (68.75) wide) free from dorsal plate. Dgl- 4 closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 275 - 305 (300 )). Dorsal plate proportions: dorsum length / width 1.41 - 1.55 (1.55); dorsal width / distance between Dgl- 4 1.33 - 1.45 (1.33); anterio-medial platelet length / width 2.26 - 2.73 (2.36); anterio-lateral platelet length / width 2.78 - 3.20 (2.98); anterio-lateral / anterio-medial length 1.37 - 1.48 (1.48). Gnathosoma - Subcapitulum (270 - 292.5 (285) long (ventral); 175 - 215 (197.5) long (dorsal); 96.25 - 110 (105) tall) colorless. Rostrum (90 - 107.5 (98.75) long; 45 - 50 (46.25) wide) wide and upturned with dentation. Chelicerae (265 - 285 (265) long) with curved fangs (45 - 50 (50) long). Subcapitular proportions: ventral length / height 2.45 - 2.86 (2.71); rostrum length / width 2.00 - 2.17 (2.14). Pedipalps short and stocky (especially tibiae) with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (42.5 - 47.5 (42.5) long); femur (80 - 91.25 (87.5) long); genu (55 - 57.5 (55) long); tibia (50 - 57.5 (50) long; 23.75 - 27.5 (23.75) wide); tarsus (15 - 17.5 (15) long). Palpomere proportions: femur / genu 1.39 - 1.59 (1.59); tibia / femur 0.57 - 0.63 (0.57); tibia length / width 2.00 - 2.11 (2.11). Venter - (720 - 780 (750) long; 470 - 495 (470) wide) colorless. Gnathosomal bay (167.5 - 177.5 (172.5) long; 100 - 105 (105) wide). Cxgl- 4 far from apex. Medial suture (75 - 82.5 (75) long). Genital plates (152.5 - 165 (157.5) long; 112.5 - 125 (112.5) wide). Additional measurements: Cx- 1 (290 - 330 (310) long (total); 125 - 150 (140) long (medial )); Cx- 3 (360 - 390 (360) wide); anterior venter (232.5 - 250 (250) long). Ventral proportions: gnathosomal bay length / width 1.60 - 1.78 (1.64); anterior venter / genital field length 1.47 - 1.64 (1.59); anterior venter length / genital field width 1.94 - 2.22 (2.22); anterior venter / medial suture 2.94 - 3.33 (3.33). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
1B8D1B082E8E51399530882E406DBC56.taxon	etymology	Etymology. Specific epithet (erectirostra) refers to the upturned rostrum characteristic of members of the Erectirostra Group (erectus, raised up; rostrum, L. snout).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
1B8D1B082E8E51399530882E406DBC56.taxon	distribution	Distribution. Appalachians (Figure 72).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
5DC77C9DD3D0E31A062356B6DDA67172.taxon	materials_examined	Material examined. HOLOTYPE (♀): from USA, Georgia, Chattooga County, Cloudland; beside Rt. 48 just east of Alabama state line, (34 ° 31 ' 31 " N, 85 ° 30 ' 30 " W), 29 September 2005, by IM Smith, IMS 050110 A. PARATYPES (4 ♀; 6 ♂): Georgia, USA: 2 ♀ and 4 ♂ from Chattooga County, Cloudland; beside Rt. 48 just east of Alabama State line, (34 ° 31 ' 31 " N, 85 ° 30 ' 30 " W), 28 September 1992, by IM Smith, IMS 920056 A * 1 ♂ (ALLOTYPE) from Chattooga County, Cloudland; beside Rt. 48 just east of Alabama state line, (34 ° 31 ' 31 " N, 85 ° 30 ' 30 " W), 29 September 2005, by IM Smith, IMS 050110 A * 2 ♀ and 1 ♂ from Chattooga County, Cloudland; beside Rt. 48 just east of Alabama state line, (34 ° 31 ' 31 " N, 85 ° 30 ' 30 " W), 29 September 2005, by IM Smith, IMS 050110 A.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
5DC77C9DD3D0E31A062356B6DDA67172.taxon	diagnosis	Diagnosis. Torrenticola feminellai are similar to other members of the Rusetria " Eastern 2 - Plates " group (T. biscutella, T. caerulea, T. delicatexa, T. indistincta, T. malarkeyorum, T. pendula, T. sellersorum, T. tysoni, T. ululata, T. whitneyae, and T. microbiscutella) in having anterio-lateral platelets fused to the dorsal plate, having dorsal coloration separated into anterior and posterior portions (except T. ululata and T. indistincta), and being distributed in the east. T. feminellai can be differentiated from all other Eastern 2 - Plates by having a more elongate rostrum (length / width ♀ = 3.05 - 3.38 in T. feminellai, 2.33 - 3.00 in others; ♂ = 3.14 - 3.38 in T. feminellai, 2.50 - 3.05 in others), except T. tysoni (3.06 - 3.50). Additionally, T. feminellai can be differentiated from all other Eastern 2 - Plates by having a distinct dorsal pattern.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
5DC77C9DD3D0E31A062356B6DDA67172.taxon	description	Description. Female (Figure 76) (n = 5) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (590 - 690 (640) long; 470 - 540 (500) wide) circular with reddish-purple coloration in the shape of an hourglass. Anterio-medial platelets (137.5 - 150 (150) long; 52.5 - 55 (55) wide). Anterio-lateral platelets (185 - 202.5 (202.5) long; 75 - 80 (75) wide) partially fused to dorsal plate. Dgl- 4 approximately halfway between the edge of the dorsum and the muscle scars (distance between Dgl- 4 290 - 310 (290 )). Dorsal plate proportions: dorsum length / width 1.24 - 1.33 (1.28); dorsal width / distance between Dgl- 4 1.59 - 1.74 (1.72); anterio-medial platelet length / width 2.62 - 2.76 (2.73); anterio-lateral platelet length / width 2.34 - 2.70 (2.70); anterio-lateral / anterio-medial length 1.25 - 1.36 (1.35). Gnathosoma - Subcapitulum (320 - 357.5 (352.5) long (ventral); 245 - 275 (267.5) long (dorsal); 140 - 160 (160) tall) colorless. Rostrum (135 - 157.5 (145) long; 40 - 47.5 (47.5) wide). Chelicerae (335 - 375 (375) long) with curved fangs (65 - 75 (70) long). Subcapitular proportions: ventral length / height 2.17 - 2.30 (2.20); rostrum length / width 3.05 - 3.38 (3.05). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (46.25 - 52.5 (52.5) long); femur (117.5 - 140 (140) long); genu (65 - 77.5 (75) long); tibia (90 - 100 (100) long; 22.5 - 25 (25) wide); tarsus (17.5 - 20 (20) long). Palpomere proportions: femur / genu 1.73 - 1.87 (1.87); tibia / femur 0.71 - 0.77 (0.71); tibia length / width 3.85 - 4.11 (4.00). Venter - (670 - 760 (700) long; 550 - 690 (550) wide) colorless. Gnathosomal bay (177.5 - 195 (195) long; 80 - 115 (80) wide). Cxgl- 4 subapical. Medial suture absent. Genital plates (185 - 195 (195) long; 167.5 - 180 (170) wide). Additional measurements: Cx- 1 (310 - 320 (320) long (total); 120 - 140 (125) long (medial )); Cx- 3 (340 - 410 (360) wide); anterior venter (130 - 140 (130) long). Ventral proportions: gnathosomal bay length / width 1.61 - 2.44 (2.44); anterior venter / genital field length 0.67 - 0.76 (0.67); anterior venter length / genital field width 0.74 - 0.82 (0.76). Male (Figure 77) (n = 5) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (520 - 565 (545) long; 370 - 410 (390) wide) circular with reddish-purple coloration in the shape of an hourglass. Anterio-medial platelets (115 - 126.25 (122.5) long; 42.5 - 50 (45) wide). Anterio-lateral platelets (150 - 180 (155) long; 55 - 67.5 (60) wide) partially fused to dorsal plate. Dgl- 4 approximately halfway between the edge of the dorsum and the muscle scars (distance between Dgl- 4 215 - 250 (245 )). Dorsal plate proportions: dorsum length / width 1.36 - 1.42 (1.40); dorsal width / distance between Dgl- 4 1.54 - 1.72 (1.59); anterio-medial platelet length / width 2.42 - 2.82 (2.72); anterio-lateral platelet length / width 2.58 - 2.77 (2.58); anterio-lateral / anterio-medial length 1.27 - 1.43 (1.27). Gnathosoma - Subcapitulum (270 - 300 (290) long (ventral); 200 - 227.5 (215) long (dorsal); 105 - 120 (115) tall) colorless. Rostrum (110 - 120 (120) long; 32.5 - 37.5 (37.5) wide). Chelicerae (270 - 310 (295) long) with curved fangs (50 - 60 (55) long). Subcapitular proportions: ventral length / height 2.50 - 2.59 (2.52); rostrum length / width 3.14 - 3.38 (3.20). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (35 - 45 (42.5) long); femur (95 - 112.5 (105) long); genu (60 - 65 (62.5) long); tibia (77.5 - 86.25 (85) long; 20 - 23.75 (21.25) wide); tarsus (15 - 17.5 (17.5) long). Palpomere proportions: femur / genu 1.58 - 1.75 (1.68); tibia / femur 0.76 - 0.82 (0.81); tibia length / width 3.63 - 4.00 (4.00). Venter - (610 - 685 (640) long; 415 - 470 (470) colorless. Gnathosomal bay (135 - 160 (155) long; 62.5 - 70 (62.5) wide). Cxgl- 4 subapical. Medial suture (65 - 75 (70) long). Genital plates (130 - 140 (140) long; 115 - 125 (120) wide). Additional measurements: Cx- 1 (245 - 290 (270) long (total); 105 - 130 (120) long (medial )); Cx- 3 (285 - 315 (315) wide); anterior venter (190 - 205 (195) long). Ventral proportions: gnathosomal bay length / width 2.00 - 2.48 (2.48); anterior venter / genital field length 1.39 - 1.49 (1.39); anterior venter length / genital field width 1.58 - 1.72 (1.63); anterior venter / medial suture 2.60 - 2.93 (2.79). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
5DC77C9DD3D0E31A062356B6DDA67172.taxon	etymology	Etymology. Specific epithet (feminellai) named in honor of Jack Feminella, professor of biology at Auburn University, who believed in me (JRF) enough to employ me as a lab technician in stream ecology, write a winning recommendation letter for graduate school, and was the first to teach me how to conduct self-directed research.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
5DC77C9DD3D0E31A062356B6DDA67172.taxon	distribution	Distribution. Southern Appalachians, Georgia (Figure 75).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
4C6DA6035B6B30B95C2C1B284D2AD5B4.taxon	materials_examined	Material examined. HOLOTYPE (♀): from USA, Alabama, Lauderdale County, Natchez Trace Parkway, (34 ° 56 ' 31 " N, 87 ° 49 ' 41 " W), 27 Sep 2010, by IM Smith, IMS 100162, DNA 1310. PARATYPES (5 ♀; 7 ♂): Alabama, USA: 3 ♂ from Lauderdale County, Natchez Trace Parkway, (34 ° 56 ' 31 " N, 87 ° 49 ' 41 " W), 24 Sep 2009, by IM Smith, IMS 090121 * 1 ♂ (ALLOTYPE) from Lauderdale County, Natchez Trace Parkway, (34 ° 56 ' 31 " N, 87 ° 49 ' 41 " W), 27 Sep 2010, by IM Smith, IMS 100162, DNA 1309 * 1 ♀ and 2 ♂ from Lauderdale County, Natchez Trace Parkway, (34 ° 56 ' 31 " N, 87 ° 49 ' 41 " W), 27 Sep 2010, by IM Smith, IMS 100162 * Arkansas, USA: 1 ♀ from Carroll County, beside Route 62, Kings River, 21 Jul 1960, by DR Cook, DRC 600026 * Kentucky, USA: 1 ♀ from Larue County, beside Route 31 E, 4 kilometers south of Route 61 at Hodgenville (37 ° 35 ' N, 85 ° 42 ' W), 28 May 1992, by IM Smith, IMS 920014 * North Carolina, USA: 1 ♀ and 1 ♂ from Haywood County, Great Smokey Mountains National Park, Big Creek (35 ° 45 ' 3.92 " N, 83 ° 6 ' 31.67 " W), 15 Sep 2009, by AJ Radwell, AJR 090008 A * Tennessee, USA: 1 ♀ from Sevier County, Great Smokey Mountains National Park, Sugarlands Nature Trail (35 ° 40 ' 47 " N, 83 ° 31 ' 52 " W), 7 Sep 2009, by IM Smith, IMS 090101.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
4C6DA6035B6B30B95C2C1B284D2AD5B4.taxon	diagnosis	Diagnosis. Torrenticola flangipalpa are similar to other members of the Nigroalba Group (T. nigroalba, T. solisorta, and T. dentirostra) in being small, slightly elongate, having purple dorsal coloration restricted posteriorly, and having distinct yet poorly-defined hind coxal margins (can be indistinct, at least in males). T. flangipalpa are best differentiated from other members of the Nigroalba Group by having a flange-like, anteriorly-directed pedipalp femoral extension (this extension is tuberculate in other members of the Nigroalba Group). Additionally, T. flangipalpa can be differentiated from T. nigroalba and T. solisorta by having a longer anterior venter (235 - 265 in T. flangipalpa; 192 - 225 in others) and stockier pedipalpal tibiae (length / width ♀ = 4.79 - 5.0 in T. flangipalpa, 5.38 - 5.83 in others; length / width ♂ = 4.4 - 4.86 in T. flangipalpa, 5.08 - 5.33 in others). T. flangipalpa can be differentiated from T. dentirostra by having a smooth rostrum (T. dentirostra has a dentate bump midway on the dorsal edge of the rostrum). Other Torrenticola with purple dorsal coloration restricted posteriorly, such as T. tahoei and T. oregonensis are larger (dorsum length ♀ = 530 - 565 in T. flangipalpa, 600 - 810 in others; ♂ = 480 - 510 in T. flangipalpa, 560 - 820 in others) and distributed in the west (T. flangipalpa is only known from Alabama and Tennessee).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
4C6DA6035B6B30B95C2C1B284D2AD5B4.taxon	description	Description. Female (Figure 79) (n = 5) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (530 - 580 (545) long; 365 - 425 (380) wide) ovoid with purple or bluish-purple coloration restricted posteriorly. Anterio-medial platelets (112.5130 (127.5) long; 47.5 - 57.5 (47.5) wide). Anterio-lateral platelets (157.5 - 175 (172.5) long; 55 - 62.5 (55) wide) free from dorsal plate. Dgl- 4 closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 255 - 295 (255 )). Dorsal plate proportions: dorsum length / width 1.33 - 1.45 (1.43); dorsal width / distance between Dgl- 4 1.39 - 1.49 (1.49); anterio-medial platelet length / width 2.25 - 2.74 (2.68); anterio-lateral platelet length / width 2.86 - 3.14 (3.14); anterio-lateral / anterio-medial length 1.29 - 1.40 (1.35). Gnathosoma - Subcapitulum (305 - 330 (307.5) long (ventral); 225 - 255 (225) long (dorsal); 90 - 95 (90) tall) elongate and colorless. Rostrum (112.5 - 130 (117.5) long; 37.5 - 42.5 (37.5) wide) elongate. Chelicerae (285 - 320 (286) long) with curved fangs (43 - 50 (44) long). Subcapitular proportions: ventral length / height 3.39 - 3.47 (3.42); rostrum length / width 2.82 - 3.06 (3.00). Pedipalps elongate (especially tibiae) with broad, dentate, anteriorly-directed flange on femora and with variable, dentate flange-like extension on genua. Palpomeres: trochanter (32.5 - 35 (35) long); femur (93.75 - 102.5 (93.75) long); genu (55 - 62.5 (55) long); tibia (80 - 90 (83.75) long; 16.25 - 18.75 (17.5) wide); tarsus (15 - 17.5 (15) long). Palpomere proportions: femur / genu 1.60 - 1.70 (1.70); tibia / femur 0.82 - 0.90 (0.89); tibia length / width 4.79 - 5.00 (4.79). Venter - (680 - 750 (680) long; 430 - 495 (436) wide) with faint purple or bluish-purple coloration. Gnathosomal bay (112.5 - 140 (120) long; 67.5 - 75 (67.5) wide). Cxgl- 4 far from apex. Medial suture (67.5 - 80 (67.5) long). Genital plates (145 - 160 (151.25) long; 125 - 132.5 (125) wide). Additional measurements: Cx- 1 (266 - 310 (266) long (total); 150 - 170 (156) long (medial )); Cx- 3 (278 - 321 (278) wide); anterior venter (235 - 255 (245) long). Ventral proportions: gnathosomal bay length / width 1.50 - 2.00 (1.78); anterior venter / genital field length 1.55 - 1.76 (1.62); anterior venter length / genital field width 1.86 - 2.04 (1.96); anterior venter / medial suture 3.06 - 3.78 (3.63). Male (Figure 80) (n = 6) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (480 - 510 (480) long; 330 - 370 (330) wide) ovoid with purple or bluish-purple coloration restricted posteriorly. Anterio-medial platelets (112.5 - 122.5 (115) long; 41.25 - 47.5 (41.25) wide). Anterio-lateral platelets (152.5 - 162.5 (152.5) long; 50 - 53.75 (50) wide) free from dorsal plate. Dgl- 4 closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 235 - 260 (235 )). Dorsal plate proportions: dorsum length / width 1.38 - 1.45 (1.45); dorsal width / distance between Dgl- 4 1.39 - 1.44 (1.40); anterio-medial platelet length / width 2.42 - 2.79 (2.79); anterio-lateral platelet length / width 3.02 - 3.10 (3.05); anterio-lateral / anterio-medial length 1.33 - 1.44 (1.33). Gnathosoma - Subcapitulum (272.5 - 290 (272.5) long (ventral); 200 - 209 (200) long (dorsal); 75 - 87.5 (75) tall) elongate and colorless. Rostrum (102.5 - 107.5 (105) long; 32.5 - 35 (35) wide) elongate. Chelicerae (250 - 272 (260) long) with curved fangs (39 - 62 (40) long). Subcapitular proportions: ventral length / height 3.31 - 3.68 (3.63); rostrum length / width 2.93 - 3.31 (3.00). Pedipalps elongate (especially tibiae) with broad, dentate, anteriorly-directed flange on femora and with variable, dentate flange-like extension on genua. Palpomeres: trochanter (27.5 - 31.25 (27.5) long); femur (85 - 90 (87.5) long); genu 47.5 - 57.5 (55) long); tibia (77.5 - 85 (80) long; 17.5 - 18.75 (17.5) wide); tarsus (12.5 - 15 (12.5) long). Palpomere proportions: femur / genu 1.52 - 1.79 (1.59); tibia / femur 0.91 - 0.94 (0.91); tibia length / width 4.40 - 4.86 (4.57). Venter - (600 - 640 (600) long; 356 - 420 (380) wide) with faint purple or bluish-purple coloration. Gnathosomal bay (100 - 112.5 (105) long; 65 - 72.5 (65) wide). Cxgl- 4 far from apex. Medial suture (82.5 - 107.5 (95) long). Genital plates (122.5 - 127.5 (122.5) long; 92.5 - 100 (95) wide). Additional measurements: Cx- 1 (240 - 258 (240) long (total); 122 - 160 (135) long (medial )); Cx- 3 (251 - 291 (265) wide); anterior venter (245 - 265 (245) long). Ventral proportions: gnathosomal bay length / width 1.48 - 1.62 (1.62); anterior venter / genital field length 1.98 - 2.12 (2.00); anterior venter length / genital field width 2.55 - 2.68 (2.58); anterior venter / medial suture 2.42 - 3.00 (2.58). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
4C6DA6035B6B30B95C2C1B284D2AD5B4.taxon	etymology	Etymology. Specific epithet (flangipalpa) refers the expanded (i. e., flanged) pedipalp femoral tubercle (flange, English; palpus, L. hand, feeler), which distinguishes this species from other members of the Nigroalba Group.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
4C6DA6035B6B30B95C2C1B284D2AD5B4.taxon	distribution	Distribution. Southeastern (Figure 78).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
D86D58F4FFAA077A3716A312FA89185F.taxon	materials_examined	Material examined. HOLOTYPE (♀): from USA, New Hampshire, Coos County, picnic area beside Rt. 110 ca. 1 km east of Stark, (44 ° 36 ' 36 " N, 71 ° 24 ' 24 " W), 5 July 1989, by IM Smith, IMS 890071 PARATYPES (4 ♀; 4 ♂): 1 ♂ (ALLOTYPE) from Coos County, New Hampshire picnic area beside Rt. 110 ca. 1 km east of Stark, (44 ° 36 ' 36 " N, 71 ° 24 ' 24 " W), 5 July 1989, by IM Smith, IMS 890071 * 4 ♀ and 4 ♂ from Coos County, picnic area beside Rt. 110 ca. 1 km east of Stark, (44 ° 36 ' 36 " N, 71 ° 24 ' 24 " W), 5 July 1989, by IM Smith, IMS 890071.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
D86D58F4FFAA077A3716A312FA89185F.taxon	diagnosis	Diagnosis. Torrenticola folkertsae are similar to other members of the Partial 2 - Plate Group (T. magnexa, T. priapus, and T. pulchra) in having anterio-lateral platelets partially fused to the dorsal plate and being distributed in the east. T. folkertsae can be differentiated from T. magnexa and T. pulchra by having more elongate pedipalpal tibiae (length / width = 4.05 - 4.83 in T. folkertsae, 2.61 - 4.00 in others) and by dorsal coloration and pattern. T. folkertsae can be differentiated from T. priapus by having more elongate pedipalpal tibiae (length / width ♀ = 4.5 - 4.83 in T. folkertsae, 3.9 - 4.22 in T. priapus ♂ = 4.05 - 4.33 in T. folkertsae, 3.5 - 3.78 in T. priapus) and a less elongate rostrum (length / width = 2.55 - 3.00 in T. folkertsae, 3.17 - 3.39 in T. priapus).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
D86D58F4FFAA077A3716A312FA89185F.taxon	description	Description. Female (Figure 82) (n = 5) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (600 - 720 (720) long; 505 - 600 (580) wide) ovoid with faint reddish-purple medially. Anterio-medial platelets (140 - 167.5 (156.25) long; 57.5 - 70 (65) wide). Anterio-lateral platelets (180 - 205 (185) long; 85 - 95 (85) wide) partially fused to dorsal plate. Dgl- 4 closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 340 - 380 (380 )). Dorsal plate proportions: dorsum length / width 1.11 - 1.24 (1.24); dorsal width / distance between Dgl- 4 1.40 - 1.75 (1.53); anterio-medial platelet length / width 2.32 - 2.50 (2.40); anterio-lateral platelet length / width 2.12 - 2.31 (2.18); anterio-lateral / anterio-medial length 1.12 - 1.41 (1.18). Gnathosoma - Subcapitulum (310 - 345 (345) long (ventral); 240 - 270 (270) long (dorsal); 145 - 162.5 (157.5) tall) colorless. Rostrum (135 - 150 (140) long; 50 - 55 (55) wide). Chelicerae (320 - 415 (415) long) with curved fangs (65 - 70 (70) long). Subcapitular proportions: ventral length / height 2.07 - 2.19 (2.19); rostrum length / width 2.55 - 2.86 (2.55). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (48.75 - 52.5 (52.5) long); femur (130 - 145 (145) long); genu (77.5 - 87.5 (87.5) long); tibia (108.75 - 120 (118.75) long; 22.5 - 26.25 (26.25) wide); tarsus (17.5 - 22.5 (22.5) long). Palpomere proportions: femur / genu 1.65 - 1.68 (1.66); tibia / femur 0.82 - 0.87 (0.82); tibia length / width 4.50 - 4.83 (4.52). Venter - (680 - 880 (860) long; 560 - 675 (650) wide) colorless. Gnathosomal bay (167.5 - 185 (182.5) long; 80 - 95 (95) wide). Cxgl- 4 far from apex. Medial suture (15 - 20 (20) long). Genital plates (175 - 197.5 (197.5) long; 167.5 - 175 (167.5) wide). Additional measurements: Cx- 1 (300 - 340 (330) long (total); 127.5 - 157.5 (145) long (medial )); Cx- 3 (380 - 400 (400) wide); anterior venter (155 - 180 (180) long). Ventral proportions: gnathosomal bay length / width 1.92 - 2.13 (1.92); anterior venter / genital field length 0.79 - 1.00 (0.91); anterior venter length / genital field width 0.91 - 1.07 (1.07); anterior venter / medial suture 8.38 - 12.00 (9.00). Male (Figure 83) (n = 5) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (535 - 580 (570) long; 420 - 450 (445) wide) ovoid with faint reddish-purple medially. Anterio-medial platelets (125 - 135 (135) long; 55 - 65 (55) wide). Anterio-lateral platelets (180 - 187.5 (180) long; 62.5 - 77.5 (77.5) wide) partially fused to dorsal plate. Dgl- 4 closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 280 - 310 (310 )). Dorsal plate proportions: dorsum length / width 1.27 - 1.29 (1.28); dorsal width / distance between Dgl- 4 1.44 - 1.54 (1.44); anterio-medial platelet length / width 2.04 - 2.45 (2.45); anterio-lateral platelet length / width 2.32 - 2.88 (2.32); anterio-lateral / anterio-medial length 1.33 - 1.48 (1.33). Gnathosoma - Subcapitulum (265 - 292.5 (292.5) long (ventral); 210 - 227.5 (227.5) long (dorsal); 110 - 115 (110) tall) colorless. Rostrum (110 - 120 (120) long; 40 - 42.5 (40) wide). Chelicerae (275 - 295 (295) long) with curved fangs (55 - 57.5 (55) long). Subcapitular proportions: ventral length / height 2.36 - 2.66 (2.66); rostrum length / width 2.75 - 3.00 (3.00). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (42.5 - 46.25 (45) long); femur (110 - 116.25 (116.25) long); genu (67.5 - 72.5 (72.5) long); tibia (97.5 - 102.5 (102.5) long; 22.5 - 25 (23.75) wide); tarsus (17.5 - 20 (17.5) long). Palpomere proportions: femur / genu 1.55 - 1.67 (1.60); tibia / femur 0.88 - 0.90 (0.88); tibia length / width 4.05 - 4.33 (4.32). Venter - (640 - 685 (680) long; 470 - 570 (510) wide) colorless. Gnathosomal bay (132.5 - 150 (142.5) long; 70 - 80 (75) wide). Cxgl- 4 subapical. Medial suture (105 - 125 (115) long). Genital plates (130 - 145 (145) long; 125 - 130 (130) wide). Additional measurements: Cx- 1 (255 - 295 (295) long (total); 130 - 150 (150) long (medial )); Cx- 3 (330 - 365 (340) wide); anterior venter (250 - 275 (267.5) long). Ventral proportions: gnathosomal bay length / width 1.81 - 2.00 (1.90); anterior venter / genital field length 1.79 - 1.96 (1.84); anterior venter length / genital field width 2.00 - 2.16 (2.06); anterior venter / medial suture 2.17 - 2.45 (2.33). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
D86D58F4FFAA077A3716A312FA89185F.taxon	etymology	Etymology. Specific epithet (folkertsae) named in honor of Debbie Folkerts, professor of biology at Auburn University, who, together with her late husband George Folkerts, were instrumental to JRF in channeling his passion for natural history and teaching into a career path.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
D86D58F4FFAA077A3716A312FA89185F.taxon	distribution	Distribution. New Hampshire (Figure 81).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
92837E14246876E623F6B4E49F2E72DF.taxon	materials_examined	Material examined. HOLOTYPE (♀): from USA, Tennessee, Sevier County, Great Smokey Mountains National Park, Sugarlands Nature Trail (35 ° 40 ' 47 " N, 83 ° 31 ' 51 " W), 10 Sep 2010, by IM Smith, IMS 100125. PARATYPES (23 ♀; 25 ♂): Georgia, USA: 2 ♀ and 2 ♂ from Floyd County, beside road from Everett Spring to Villanow, 1.4 kilometers south of The Pocket Campground, tributary of Johns Creek, 4 Jul 1990, by IM Smith, IMS 900077 * Kentucky, USA: 2 ♀ and 3 ♂ from Bell County, Middlesboro, near north boundary of Cumberland Gap National Historical Park, Sugar Run, 9 Jul 1990, by IM Smith, IMS 900084 * 2 ♀ and 1 ♂ from Bell County, Pineville, Pine Mountain State Resort Park, Laurel Cove, Lower shelter Picnic Area, 9 Jul 1990, by IM Smith, IMS 900083 * Pennsylvania, USA: 1 ♀ from Fayette County, Dunbar Creek (39 ° 57 ' 50 " N, 79 ° 35 ' 8.70 " W), 10 Aug 2014, by MJ Skvarla, MS 14 - 0810 - 001 * 1 ♀ from Huntingdon County, Alan Seeger Natural Area, beside road from McAlevys Fort to Route 322, Stone Creek, 19 Jul 1990, by IM Smith, IMS 900107 * Tennessee, USA: 1 ♂ (ALLOTYPE) from Sevier County, Great Smokey Mountains National Park, Sugarlands Nature Trail (35 ° 40 ' 47 " N, 83 ° 31 ' 51 " W), 10 Sep 2010, by IM Smith, IMS 100125 * 1 ♂ from Sevier County, Great Smokey Mountains National Park, Laurel Creek (35 ° 39 ' 7 " N, 83 ° 42 ' 32 " W), 17 Sep 2010, by IM Smith, IMS 100145 * 4 ♀ and 5 ♂ from Sevier County, Great Smokey Mountains National Park, Sugarlands Nature Trail (35 ° 40 ' 47 " N, 83 ° 31 ' 51 " W), 10 Sep 2010, by IM Smith, IMS 100125 * Virginia, USA: 1 ♀ and 1 ♂ from Alleghany County, Clifton Forge, beside Route 606, 1.2 kilometers southeast of Forest Route 125, Smith Creek, 13 Jul 1990, by IM Smith, IMS 900093 * 1 ♀ and 1 ♂ from Alleghany County, Longdale Furnace, beside Forest Route 108, 1.7 kilometers west of Route 850, Simpson Creek, 14 Jul 1990, by IM Smith, IMS 900094 * 1 ♀ and 1 ♂ from Amherst County, Upper Otter Creek Overlook beside Blue Ridge, Otter Creek (37 ° 36 ' 57 " N, 79 ° 19 ' 27 " W), 7 Sep 2007, by IM Smith, IMS 070056 A * 2 ♀ and 2 ♂ from Augusta County, beside Forest Road 42, 15.9 kilometers east of Vesuvius, Coles Run, 26 Jul 1990, by IM Smith, IMS 900060 * 2 ♂ from Bath County, beside Forest Route 1744 at Route 39, between Warm Springs and Mountain Grove, O'Roarke Draw, 15 Jul 1990, by IM Smith, IMS 900098 * 1 ♀ and 2 ♂ from Bath County, beside Forest Route 364, off Route 39 east of Warm Springs, Panther Run, 15 Jul 1990, by IM Smith, IMS 900099 * 1 ♀ and 1 ♂ from Giles County, Mechanicsburg, beside Dismal Creek Road, Standrock Brook (37 ° 11 ' 38 " N, 80 ° 53 ' 26 " W), 9 Sep 2005, by IM Smith, IMS 050066 * 2 ♀ and 1 ♂ from Giles County, Mechanicsburg, beside Dismal Creek Road, Standrock Brook (37 ° 11 ' 38 " N, 80 ° 53 ' 26 " W), 11 Jul 1990, IMS 900088 * 3 ♂ from Montgomery County, Blacksburg, beside Route 621 at Caldwell Fields Campground, Craig Creek (37 ° 20 ' N, 80 ° 20 ' W), 12 Jul 1990, by IM Smith, IMS 900089 A * 1 ♀ and 1 ♂ from Page County, beside Route 730, 0.2 kilometers west of Route 675, Passage Creek, 25 Jun 1990, by IM Smith, IMS 900059 * 1 ♀ from Rock Bridge County, Vesuvius, beside Route 56, 2.2 kilometers west of Blue Ridge Parkway, Little Marys Creek, 26 Jun 1990, by IM Smith, IMS 900062.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
92837E14246876E623F6B4E49F2E72DF.taxon	diagnosis	Diagnosis. Torrenticola glomerabilis are similar to other members of the Rusetria " 4 - Plate " group (T. dunni, T. glomerabilis, T. kittatinniana, T. pollani, T. rufoalba and T. shubini) and T. skvarlai in having anterio-lateral platelets free from the dorsal plate, dorsal coloration separated into anterior and posterior portions, and indistinct hind coxal margins. T. glomerabilis can be differentiated from T. dunni, T. shubini, T. kittatinniana, by having Dgl- 4 closer to the dorsal edge (dorsal width / distance between Dgl- 4 = 1.53 - 1.66 in T. glomerabilis, 1.20 - 1.42 in others) and stockier tibiae (length / width ♀ = 4.11 - 4.50 in T. glomerabilis, 3.27 - 3.60 in others; ♂ = 3.55 - 4.38 in T. glomerabilis, 2.80 - 3.45 in others). T. glomerabilis can be differentiated from T. pollani and T. rufoalba by having stockier anterio-medial platelets (length / width ♀ = 1.9 - 2.3 in T. glomerabilis, 2.5 - 3.0 in others; ♂ = 1.9 - 2.2 in T. glomerabilis, 2.3 - 2.9 in others) and wider dorsum (♀ = 460 - 490 in T. glomerabilis, 400 - 420 in others; ♂ = 395 - 430 in T. glomerabilis, 310 - 340 in others). T. glomerabilis can be differentiated from T. skvarlai by having a conical pedipalpal femoral tubercle, whereas T. skvarlai has a broad and flat pedipalpal femoral tubercle, and by having a longer anterior venter (♀ = 202 - 213 in T. glomerabilis, 140 - 153 in T. skvarlai; ♂ = 240 - 280 in T. glomerabilis, 177 - 205 in T. skvarlai).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
92837E14246876E623F6B4E49F2E72DF.taxon	description	Description. Female (Figure 85) (n = 5) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (580 - 615 (605) long; 460 - 490 (475) wide) circular with bold bluish-purple or reddish-purple coloration separated into anterior and posterior portions. Anterio-medial platelets (125 - 132.5 (132.5) long; 55 - 65 (60) wide). Anterio-lateral platelets (172.5 - 195 (180) long; 70 - 82.5 (82.5) wide) free from dorsal plate. Dgl- 4 approaching midway between muscle scars and dorsum edge (distance between Dgl- 4 280 - 310 (310 )). Dorsal plate proportions: dorsum length / width 1.23 - 1.32 (1.27); dorsal width / distance between Dgl- 4 1.53 - 1.66 (1.53); anterio-medial platelet length / width 1.96 - 2.27 (2.21); anterio-lateral platelet length / width 2.17 - 2.48 (2.18); anterio-lateral / anterio-medial length 1.36 - 1.53 (1.36). Gnathosoma - Subcapitulum (320 - 330 (320) long (ventral); 223 - 243 (223) long (dorsal); 112.5 - 120 (117.5) tall) colorless. Rostrum (132.5 - 137.5 (132.5) long; 40 - 47.5 (40) wide). Chelicerae (320 - 330 (321) long) with curved fangs (50 - 55 (53) long). Subcapitular proportions: ventral length / height 2.72 - 2.89 (2.72); rostrum length / width 2.89 - 3.34 (3.31). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (37.5 - 42.5 (40) long); femur (112.5 - 117.5 (116.25) long); genu (67.5 - 70 (67.5) long); tibia (88.75 - 95 (88.75) long; 20 - 22.5 (20) wide); tarsus (17.5 - 17.5 (17.5) long). Palpomere proportions: femur / genu 1.67 - 1.74 (1.72); tibia / femur 0.76 - 0.83 (0.76); tibia length / width 4.11 - 4.50 (4.44). Venter - (710 - 730 (730) long; 512 - 550 (513) wide) bold bluish-purple or reddish-purple coloration. Gnathosomal bay (130 - 155 (155) long; 72.5 - 95 (72.5) wide). Cxgl- 4 subapical. Medial suture (22.5 - 50 (27.5) long). Genital plates (167.5 - 177.5 (175) long; 150 - 157.5 (157.5) wide). Additional measurements: Cx- 1 (276 - 305 (291) long (total); 122 - 160 (149) long (medial )); Cx- 3 (320 - 370 (321) wide); anterior venter (202.5 - 212.5 (202.5) long). Ventral proportions: gnathosomal bay length / width 1.37 - 2.14 (2.14); anterior venter / genital field length 1.14 - 1.24 (1.16); anterior venter length / genital field width 1.29 - 1.40 (1.29); anterior venter / medial suture 4.15 - 9.00 (7.36). Male (Figure 86) (n = 6) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum- (495 - 575 (530) long; 395 - 430 (420) wide) circular with bold bluish-purple or reddish-purple coloration separated into anterior and posterior portions. Anterio-medial platelets (112.5 - 120 (120) long; 52.5 - 60 (55) wide). Anterio-lateral platelets (165 - 187.5 (167.5) long; 60 - 67.5 (60) wide) free from dorsal plate. Dgl- 4 approaching midway between muscle scars and dorsum edge (distance between Dgl- 4 235 - 280 (275 )). Dorsal plate proportions: dorsum length / width 1.25 - 1.34 (1.26); dorsal width / distance between Dgl- 4 1.50 - 1.68 (1.53); anterio-medial platelet length / width 1.96 - 2.18 (2.18); anterio-lateral platelet length / width 2.54 - 2.88 (2.79); anterio-lateral / anterio-medial length 1.40 - 1.67 (1.40). Gnathosoma - Subcapitulum (260 - 297.5 (290) long (ventral); 188 - 225 (212.5) long (dorsal); 93.75 - 103.75 (93.75) tall) colorless. Rostrum (105 - 120 (120) long; 35 - 40 (35) wide). Chelicerae (249 - 298 (285) long) with curved fangs (40 - 50 (50) long). Subcapitular proportions: ventral length / height 2.77 - 3.09 (3.09); rostrum length / width 2.80 - 3.43 (3.43). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (35 - 35 (35) long); femur (95 - 105 (102.5) long); genu (57.5 - 65 (60) long); tibia (80 - 88.75 (82.5) long; 20 - 25 (20) wide); tarsus (15 - 17.5 (15) long). Palpomere proportions: femur / genu 1.58 - 1.71 (1.71); tibia / femur 0.80 - 0.87 (0.80); tibia length / width 3.55 - 4.38 (4.13). Venter - (600 - 690 (670) long; 443 - 540 (460) wide) bold bluish-purple or reddish-purple coloration. Gnathosomal bay (107.5 - 135 (132.5) long; 70 - 80 (70) wide). Cxgl- 4 subapical. Medial suture (85 - 107.5 (92.5) long). Genital plates (135 - 147.5 (140) long; 110 - 120 (117.5) wide). Additional measurements: Cx- 1 (224 - 280 (280) long (total); 88 - 160 (150) long (medial )); Cx- 3 (292 - 342 (300) wide); anterior venter (240 - 280 (260) long). Ventral proportions: gnathosomal bay length / width 1.48 - 1.93 (1.89); anterior venter / genital field length 1.78 - 1.90 (1.86); anterior venter length / genital field width 2.13 - 2.41 (2.21); anterior venter / medial suture 2.47 - 2.87 (2.81). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
92837E14246876E623F6B4E49F2E72DF.taxon	etymology	Etymology. Specific epithet (glomerabilis) refers to the rounded body of this species compared to all other members of the Rusetria Complex (glomerabilis, L. round).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
92837E14246876E623F6B4E49F2E72DF.taxon	distribution	Distribution. Appalachians (Figure 84).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
0FFB04A8A1F141C972A64A79BFC7615E.taxon	materials_examined	Material examined. HOLOTYPE (♀): from USA, Mississippi, Attala County, Hurricane Creek (33 ° 4 ' 57 " N, 89 ° 31 ' 29 " W), 12 Sep 2008, by IM Smith, IMS 080052. PARATYPES (6 ♀; 10 ♂): Georgia, USA: 1 ♂ from Lowndes County, Withlacoochee River, beside Route 84 at Brooks County line, 13 Sep 1968, by DR Cook, DRC 680075 * Illinois, USA: 1 ♀ and 2 ♂ from Clark County, Big Creek (32 ° 25 ' 59 " N, 87 ° 41 ' 15 " W), 30 Jul 2014, by MJ Skvarla, MS 14 - 0730 - 001 * 1 ♂ from Vermilion County, Fairmount, beside Vermilion County Road 680 E, Jordan Creek (40 ° 4 ' N, 87 ° 50 ' W), 10 Sep 1991, by IM Smith, IMS 910030 * Mississippi, USA: 1 ♂ (ALLOTYPE) from USA, Mississippi, Attala County, Hurricane Creek (33 ° 4 ' 57 " N, 89 ° 31 ' 29 " W), 12 Sep 2008, by IM Smith, IMS 080052 * 1 ♀ and 1 ♂ from Attala County, Hurricane Creek (33 ° 4 ' 58 " N, 89 ° 31 ' 31 " W), 30 Sep 2010, by IM Smith, IMS 100168 * 1 ♀ and 1 ♂ from Grenada County, Leflore, beside Black Creek Road, Black Creek (33 ° 43 ' N, 90 ° 3 ' W), 16 Sep 1991, by IM Smith, IMS 910044 * 1 ♀ and 1 ♂ from Jefferson County, off Natchez Trace Parkway, Coles Creek (31 ° 41 ' 26 " N, 91 ° 10 ' 52 " W), 2 Oct 1994, by IM Smith, IMS 940029 A * 1 ♀ and 1 ♂ from Tishomingo County, Tishomingo State Park, Bear Creek, (34 ° 36 ' N, 88 ° 11 ' W), 18 Sep 1991, by IM Smith, IMS 910047 A * Oklahoma, USA: 2 ♀ and 2 ♂ from Pushmataha County, beside Route 271, South of Albion, Walnut Creek (34 ° 39 ' N, 95 ° 7 ' W), 1 Jul 1987, by IM Smith, IMS 870063 A.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
0FFB04A8A1F141C972A64A79BFC7615E.taxon	diagnosis	Diagnosis. Torrenticola gnoma are similar to other members of the Raptor Group (T. irapalpa, T. longitibia, T. mjolniri, T. elusiva, T. racupalpa, T. raptor, T. danielleae, T. daemon, and T. ivyae) in having round bodies; Dgl- 4 close to muscles scars; long, thin subcapitular rostra; and long, thin pedipalp tibiae. T. gnoma can be differentiated from T. elusiva by being smaller (dorsum length = 540 - 595 in T. gnoma, 645 in T. elusiva); having a stockier rostrum (length / width = 2.74 - 3.13 in T. gnoma, 3.65 in T. elusiva); and by dorsal coloration. T. gnoma can be differentiated from T. racupalpa by having a stockier rostrum (length / width = 2.74 - 3.13 in T. gnoma, 3.56 - 3.88 in T. racupalpa) and by dorsal coloration and pattern. T. gnoma can be differentiated from T. irapalpa, T. danielleae, and T. daemon by dorsal coloration and pattern. Additionally, female T. gnoma can be differentiated from female T. irapalpa, T. danielleae, and T. daemon by having Dgl- 4 closer to the muscle scars (dorsal width / distance between Dgl- 4 ♀ = 2.65 - 3.29 in T. gnoma, 1.57 - 2.09 in others). T. gnoma can be differentiated from T. mjolniri, T. longitibia, T. raptor, and T. ivyae by having stockier pedipalp tibiae (length / width = 3.88 - 4.67 in T. gnoma, 4.75 - 7.54 in others) and a stockier rostrum (length / width = 2.56 - 3.23 in T. gnoma, 3.44 - 4.4 in others).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
0FFB04A8A1F141C972A64A79BFC7615E.taxon	description	Description. Female (Figure 88) (n = 5) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (540 - 595 (550) long; 440 - 500 (455) wide) circular with a large spot of coloration medially extending in a thin strip anteriorly to the edge of the dorsal plate, coloration variable from navy blue to purple to pink. Anterio-medial platelets (122.5 - 137.5 (131.25) long; 55 - 62.5 (60) wide). Anterio-lateral platelets (152.5 - 187.5 (167.5) long; 67.5 - 75 (70) wide) free from dorsal plate. Dgl- 4 much closer to the muscle scar than to dorsum edge (distance between Dgl- 4 140 - 185 (155 )). Dorsal plate proportions: dorsum length / width 1.17 - 1.30 (1.21); dorsal width / distance between Dgl- 4 2.65 - 3.29 (2.94); anterio-medial platelet length / width 2.06 - 2.39 (2.19); anterio-lateral platelet length / width 2.26 - 2.59 (2.39); anterio-lateral / anterio-medial length 1.11 - 1.46 (1.28). Gnathosoma - Subcapitulum (285 - 305 (290) long (ventral); 225 - 239 (226) long (dorsal); 115 - 135 (120) tall) colorless. Rostrum (122.5 - 130 (125) long; 40 - 47.5 (40) wide) elongate. Chelicerae (285 - 310 (286) long) with curved fangs (53 - 60 (54) long). Subcapitular proportions: ventral length / height 2.26 - 2.52 (2.42); rostrum length / width 2.74 - 3.13 (3.13). Pedipalps elongate with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (40 - 45 (40) long); femur (110 - 120 (112.5) long); genu (60 - 67.5 (62.5) long); tibia (87.5 - 105 (92.5) long; 20 - 22.5 (20) wide); tarsus (17.5 - 20 (17.5) long). Palpomere proportions: femur / genu 1.63 - 1.88 (1.80); tibia / femur 0.76 - 0.88 (0.82); tibia length / width 4.17 - 4.67 (4.63). Venter - (660 - 730 (680) long; 500 - 575 (500) wide) colorless. Gnathosomal bay (142.5 - 172.5 (152.5) long; 75 - 92.5 (75) wide). Cxgl- 4 subapical. Medial suture (15 - 27.5 (27.5) long). Genital plates (152.5 - 165 (157.5) long; 142.5 - 152.5 (142.5) wide). Additional measurements: Cx- 1 (252 - 285 (257) long (total); 84 - 122 (108) long (medial )); Cx- 3 (317 - 377 (318) wide); anterior venter (160 - 167.5 (165) long). Ventral proportions: gnathosomal bay length / width 1.57 - 2.16 (2.03); anterior venter / genital field length 0.97 - 1.10 (1.05); anterior venter length / genital field width 1.05 - 1.16 (1.16); anterior venter / medial suture 6.00 - 10.67 (6.00). Male (Figure 89) (n = 5) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (420 - 495 (450) long; 355 - 375 (375) wide) circular with a large spot of coloration medially extending in a thin strip anteriorly to the edge of the dorsal plate, coloration variable from navy blue to purple to pink. Anterio-medial platelets (1.80 - 2.30 (2.18) long; 50 - 62.5 (55) wide). Anterio-lateral platelets (135 - 152.5 (152.5) long; 60 - 65 (60) wide) free from dorsal plate. Dgl- 4 much closer to the muscle scar than to dorsum edge (distance between Dgl- 4 130 - 180 (165 )). Dorsal plate proportions: dorsum length / width 1.17 - 1.32 (1.20); dorsal width / distance between Dgl- 4 2.06 - 2.73 (2.27); anterio-medial platelet length / width 1.80 - 2.30 (2.18); anterio-lateral platelet length / width 2.16 - 2.54 (2.54); anterio-lateral / anterio-medial length 1.17 - 1.36 (1.27). Gnathosoma- Subcapitulum (240 - 265 (265) long (ventral); 175 - 196 (196) long (dorsal); 97.5 - 105 (105) tall) colorless. Rostrum (98.75 - 107.5 (107.5) long; 35 - 40 (35) wide) elongate. Chelicerae (225 - 257 (256) long) with curved fangs (41 - 51 (50) long). Subcapitular proportions: ventral length / height 2.29 - 2.56 (2.52); rostrum length / width 2.56 - 3.07 (3.07). Pedipalps elongate with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (35 - 47.5 (47.5) long); femur (96.25 - 103.75 (103.75) long); genu (52.5 - 57.5 (57.5) long); tibia (77.5 - 90 (88.75) long; 20 - 22.5 (20) wide); tarsus (17.5 - 20 (18.75) long). Palpomere proportions: femur / genu 1.72 - 1.83 (1.80); tibia / femur 0.81 - 0.91 (0.86); tibia length / width 3.88 - 4.44 (4.44). Venter - (560 - 590 (581) long; 354 - 440 (395) wide) colorless. Gnathosomal bay (105 - 130 (127.5) long; 62.5 - 77.5 (62.5) wide). Cxgl- 4 subapical. Medial suture (67.5 - 80 (70) long). Genital plates (122.5 - 135 (127.5) long; 100 - 110 (105) wide). Additional measurements: Cx- 1 (217 - 269 (255) long (total); 90 - 119 (115) long (medial )); Cx- 3 (264 - 312 (295) wide); anterior venter (197.5 - 222.5 (207.5) long). Ventral proportions: gnathosomal bay length / width 1.40 - 2.04 (2.04); anterior venter / genital field length 1.49 - 1.65 (1.63); anterior venter length / genital field width 1.80 - 2.12 (1.98); anterior venter / medial suture 2.53 - 2.96 (2.96). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
0FFB04A8A1F141C972A64A79BFC7615E.taxon	etymology	Etymology. Specific epithet (gnoma) refers to the dorsal pattern, which, although variable, resembles the head and cap of a gnome (gnoma, L. diminutive fabled being, dwarf).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
0FFB04A8A1F141C972A64A79BFC7615E.taxon	distribution	Distribution. Eastern, but apparently absent from Appalachians and Northeast (Figure 87).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
D0E295A135CB58788B83B80A9FFC6D37.taxon	materials_examined	Material examined. HOLOTYPE (♀): from Canada, New Brunswick, York County, Davis Brook, beside Hwy 3, 3.5 km south of Hwy 4 at Thomaston Corner, 11 Jun 2012, by IM Smith, IMS 120017, DNA 2970. PARATYPES (24 ♀; 22 ♂): Alabama, USA: 1 ♀ and 1 ♂ from Cleburne County, beside Route 431, 3.3 kilometers southeast of Calhoun, Jackson Creek (33 ° 36 ' N, 85 ° 42 ' W), 2 Jul 1990, by IM Smith, IMS 900074 * 2 ♀ and 1 ♂ from DeKalb County, Desoto State Park, beside Trail Y, West Fork of Little River (34 ° 29 ' N, 85 ° 32 ' W), 26 Sep 1992, by IM Smith, IMS 920053 A * Georgia, USA: 2 ♀ and 2 ♂ from Chattooga County, Cloudland, beside Route 48, East Fork of Little River (34 ° 31 ' 25 " N, 85 ° 30 ' 23 " W), 28 Sep 1992, by IM Smith, IMS 920056 A * 4 ♀ and 1 ♂ from Floyd County, Johns Creek, beside road from Everett Springs to Villanow, south of The Pocket campground, 4 Jul 1990, by IM Smith, IMS 900076 & IMS 900077 * Kentucky, USA: 3 ♀ and 2 ♂ from McCreary County, Rock Creek, White Oak Junction, beside Forest Route 556, south of Route 1363, 8 Jul 1990, by IM Smith, IMS 900082 A & IMS 900082 B * Maine, USA: 2 ♀ and 3 ♂ from Aroostook County, Ashland, beside Route 11 at bridge, Aroostook River (46 ° 38 ' N, 68 ° 24 ' W), 4 Jul 1989, by IM Smith, IMS 890067 * New Brunswick, Canada: 1 ♂ (ALLOTYPE) from York County, Davis Brook, beside Hwy 3, 3.5 km south of Hwy 4 at Thomaston Corner, 11 Jun 2012, by IM Smith, IMS 120017, DNA 2972 * 2 ♀ and 2 ♂ from York County, Davis Brook, beside Hwy 3, 3.5 km south of Hwy 4 at Thomaston Corner, 11 Jun 2012, by IM Smith, IMS 120017 * 1 ♀ and 1 ♂ from York County, Magaguadavic River, beside Highway 3 just east of Thomaston Corners, 1 Jul 1989, by IM Smith, IMS 890055 A * South Carolina, USA: 1 ♂ from Greenville County, Matthews Creek, 24 Apr 2014, by D Eargle, JRF 14 - 0424 - 001 * Tennessee, USA: 4 ♀ and 1 ♂ from Monroe County, Tellico River (35 ° 19 ' N, 84 ° 10 ' W), 5 Jun 1990, by IM Smith, IMS 900079 * 2 ♂ from Monroe County, Turkey Creek, beside Forest Route 35, northeast of road from Route 165 to Miller Chapel Church, 5 Jul 1990, by IM Smith, IMS 900078 * 2 ♀ and 4 ♂ from Monroe County, Tellico River (35 ° 20 ' 27 " N, 84 ° 11 ' 31 " W), 12 Sep 2009, by IM Smith, IMS 090111 * Virginia, USA: 1 ♀ from Scott County, beside Route 58 / 421,). 9 kilometers east of Route 709, North Fork of Hoiston River (36 ° 39 ' N, 82 ° 28 ' W), 7 Jul 1990, by IM Smith, IMS 900080.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
D0E295A135CB58788B83B80A9FFC6D37.taxon	diagnosis	Diagnosis. Torrenticola gorti specimens from Tellico River system in Monroe County (Tennessee) can be differentiated from all other Torrenticola by the distinctively dark coloration with a red spot dorsally. Other color morphs are similar to species with similar dorsal patterning, such as the Rusetria " 4 - Plate " group (T. dunni, T. glomerabilis, T. kittatinniana, T. pollani, T. rufoalba and T. shubini), Neoanomala Group (T. interiorensis and T. neoanomala), and T. bondi, T. elongata, T. reduncarostra, T. erectirostra, T. robisoni, T. irapalpa, T. racupalpa, T. skvarlai, and T. arktonyx. They can be differentiated from Rusetria 4 - Plates and T. skvarlai by having distinct hind coxal margins. T. gorti can be differentiated from T. erectirostra, T. robisoni, and T. reduncarostra by having a straight, anteriorly-directed rostrum (upturned in others). T. gorti can be differentiated from T. arktonyx by having an unmodified dorsal plate (T. arktonyx has distinctive longitudinal dark markings on the anterior portion of the dorsal plate that fade posteriorly). T. gorti can be differentiated from T. racupalpa and T. irapalpa by being more elongate (dorsum length / width = 1.47 - 1.6 in T. gorti, 1.15 - 1.3 in others) and tibia / femur (0.65 - 0.73 in T. gorti, 0.77 - 0.91 in others). T. gorti can be differentiated from T. elongata by having a more ovoid dorsum (length / width = 1.47 - 1.58 in T. gorti, 1.7 - 2.1 in T. elongata) and larger dorsum (length ♀ = 570 - 600 in T. gorti, 540 - 565 in T. elongata; ♂ = 500 - 525 in T. gorti, 450 - 460 in T. elongata). T. gorti can be differentiated from the Neoanomala Group by having a more elongate dorsum (length / width ♀ = 1.47 - 1.58 in T. gorti, 1.29 - 1.43 in Neoanomala Group; ♂ = 1.54 - 1.58 in T. gorti, 1.34 - 1.50 in Neoanomala Group) and having a more elongate rostrum (length / width = 3.29 - 3.73 in T. gorti, 2.59 - 2.90 in Neoanomala Group). T. gorti can be differentiated from T. bondi by having a more elongate dorsum (length / width = 1.47 - 1.58 in T. gorti, 1.32 - 1.45 in T. bondi) and having a more elongate rostrum (length / width = 3.32 - 3.73 in T. gorti, 2.76 - 3.13 in T. bondi).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
D0E295A135CB58788B83B80A9FFC6D37.taxon	description	Description. Female (Figure 91) (n = 5) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (570 - 600 (600) long; 380 - 390 (380) wide) ovoid and elongate with three distinct color morphs: 1) navy blue coloration separated into anterior and posterior portions that meet or nearly meet laterally, and with bold orange coloration in between, from Tellico River, Monroe County, Tennessee; 2) purple coloration separated into anterior and posterior portions; 3) purple coloration separated into anterior and posterior portions with a strip of bold orange medially. Anterio-medial platelets (117.5 - 137.5 (130) long; 52.5 - 57.5 (57.5) wide). Anterio-lateral platelets (162.5 - 172.5 (172.5) long; 55 - 60 (60) wide) free from dorsal plate. Dgl- 4 approaching midway between muscle scars and dorsum edge (distance between Dgl- 4 220 - 260 (260 )). Dorsal plate proportions: dorsum length / width 1.47 - 1.58 (1.58); dorsal width / distance between Dgl- 4 1.46 - 1.73 (1.46); anterio-medial platelet length / width 2.24 - 2.45 (2.26); anterio-lateral platelet length / width 2.75 - 3.09 (2.88); anterio-lateral / anterio-medial length 1.20 - 1.45 (1.33). Gnathosoma - Subcapitulum (327.5 - 342.5 (337.5) long (ventral); 247.75 - 265 (252.5) long (dorsal); 122.5 - 127.5 (122.5) tall) colorless. Rostrum (137.5 - 142.5 (142.5) long; 37.5 - 42.5 (40) wide) elongate. Chelicerae 330 - 345 (340) long) with curved fangs (56 - 60 (60) long). Subcapitular proportions: ventral length / height 2.59 - 2.76 (2.76); rostrum length / width 3.32 - 3.73 (3.56). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (37.5 - 45 (45) long); femur (117.5 - 123.75 (120) long); genu (62.5 - 67.5 (67.5) long); tibia (77.5 - 82.5 (80) long; 20 - 25 (25) wide); tarsus (17.5 - 20 (20) long). Palpomere proportions: femur / genu 1.78 - 1.90 (1.78); tibia / femur 0.65 - 0.68 (0.67); tibia length / width 3.20 - 4.13 (3.20). Venter (Figure 57) - (695 - 750 (750) long; 410 - 494 (430) wide) with three distinct color morphs: 1) navy-blue coloration; 2) colorless; 3) purple coloration. Gnathosomal bay (142.5 - 172.5 (172.5) long; 65 - 77.5 (70) wide). Cxgl- 4 subapical. Medial suture (25 - 37.5 (27.5) long). Genital plates (160 - 167.5 (167.5) long; 135 - 140 (137.5) wide). Additional measurements: Cx- 1 (285 - 300 (300) long (total); 107 - 140 (130) long (medial )); Cx- 3 (290 - 353 (300) wide); anterior venter (187.5 - 195 (187.5) long). Ventral proportions: gnathosomal bay length / width 1.90 - 2.46 (2.46); anterior venter / genital field length 1.12 - 1.22 (1.12); anterior venter length / genital field width 1.36 - 1.44 (1.36); anterior venter / medial suture 5.20 - 7.80 (6.82). Male (Figure 92) (n = 6) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (500 - 525 (520) long; 320 - 340 (330) wide) ovoid and elongate with three distinct color morphs: 1) navy blue coloration separated into anterior and posterior portions that meet or nearly meet laterally, and with bold orange coloration in between; 2) purple coloration separated into anterior and posterior portions; 3) purple coloration separated into anterior and posterior portions with a strip of bold orange medially. Anterio-medial platelets (108.75 - 117.5 (115) long; 47.5 - 57.5 (50) wide). Anterio-lateral platelets (156.25 - 165 (157.5) long; 52.5 - 57.5 (53.75) wide) free from dorsal plate. Dgl- 4 approaching midway between muscle scars and dorsum edge (distance between Dgl- 4 210 - 230 (230 )). Dorsal plate proportions: dorsum length / width 1.54 - 1.58 (1.58); dorsal width / distance between Dgl- 4 1.43 - 1.58 (1.43); anterio-medial platelet length / width 2.04 - 2.35 (2.30); anterio-lateral platelet length / width 2.78 - 2.98 (2.93); anterio-lateral / anterio-medial length 1.36 - 1.44 (1.37). Gnathosoma - Subcapitulum (280 - 292.5 (282.5) long (ventral); 211 - 231 (215) long (dorsal); 92.5 - 1.5 (95) tall) colorless. Rostrum (115 - 127.5 (122.5) long; 33.75 - 37.5 (35) wide) elongate. Chelicerae (275 - 302 (280) long) with curved fangs (36 - 50 (50) long). Subcapitular proportions: ventral length / height 2.76 - 3.16 (2.97); rostrum length / width 3.29 - 3.50 (3.50). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (35 - 37.5 (35) long); femur (87.5 - 105 (95) long); genu (52.5 - 58.75 (56.25) long); tibia (62.5 - 72.5 (67.5) long; 18.75 - 22.5 (18.75) wide); tarsus (15 - 17.5 (15) long). Palpomere proportions: femur / genu 1.67 - 1.79 (1.69); tibia / femur 0.67 - 0.73 (0.71); tibia length / width 3.13 - 3.63 (3.60). Venter - (630 - 680 (640) long; 380 - 438 (380) wide) with three distinct color morphs: 1) navy-blue coloration; 2) colorless; 3) purple coloration. Gnathosomal bay (122.5 - 142.5 (142.5) long; 60 - 67.5 (60) wide). Cxgl- 4 subapical. Medial suture (62.5 - 87.5 (65) long). Genital plates (125 - 135 (135) long; 102.5 - 111.25 (102.5) wide). Additional measurements: Cx- 1 (255 - 280 (280) long (total); 100 - 140 (140) long (medial )); Cx- 3 (285 - 304 (290) wide); anterior venter (215 - 250 (215) long). Ventral proportions: gnathosomal bay length / width 1.89 - 2.38 (2.38); anterior venter / genital field length 1.59 - 1.89 (1.59); anterior venter length / genital field width 1.96 - 2.27 (2.10); anterior venter / medial suture 2.86 - 3.48 (3.31). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
D0E295A135CB58788B83B80A9FFC6D37.taxon	etymology	Etymology. Specific epithet (gorti) refers to Gort, the fictional giant robot of The Day the Earth Stood Still. In the 2008 film, Gort was depicted with a dark body and a single red eye that shot a destructive beam. This species is named for the resemblance that the distinctive specimens from Tennessee have to Gort's red eye.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
D0E295A135CB58788B83B80A9FFC6D37.taxon	distribution	Distribution. Appalachians (Figure 90).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
1A004CD99F8EDC0E851EF02F1D3BE6D8.taxon	materials_examined	Material examined. HOLOTYPE (♀): from USA, Indiana, Wayne County, south of I- 70 (39 ° 51 ' 13 " N, 85 ° 8 ' 4 " W), 31 Jul 2014, by MJ Skvarla, MS 14 - 0731 - 001. PARATYPES (1 ♀; 5 ♂): Indiana: 1 ♂ (ALLOTYPE) from Wayne County, south of I- 70 (39 ° 51 ' 13 " N, 85 ° 8 ' 4 " W), 31 Jul 2014, by MJ Skvarla, MS 14 - 0731 - 001 * 1 ♀ and 4 ♂ from Wayne County, south of I- 70 (39 ° 51 ' 13 " N, 85 ° 8 ' 4 " W), 31 Jul 2014, by MJ Skvarla, MS 14 - 0731 - 001.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
1A004CD99F8EDC0E851EF02F1D3BE6D8.taxon	diagnosis	Diagnosis. Torrenticola hoosieri are similar to other members of the Tricolor Complex (T. bittikoferae, T. larvata, T. pearsoni, T. olliei, T. sierrensis, T. tricolor, T. trimaculata, and T. unimaculata, T. cardia, T. kringi, T. dimorpha, and T. mohawk) in having a short, conical rostrum. T. hoosieri can be differentiated from all other Tricolor Complex, and nearly all other Torrenticola, by lacking pedipalp ventral extensions on femora and genua. Additionally, T. hoosieri can be differentiated from all other Tricolor Complex by having more elongate pedipalp tibiae (3.67 - 4.33 in T. hoosieri, 2.65 - 3.55 in others) and by being colorless (rarely with diffuse pink dorsal coloration), except T. bittikoferae, T. pearsoni, and T. dimorpha.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
1A004CD99F8EDC0E851EF02F1D3BE6D8.taxon	description	Description. Female (Figure 94) (n = 2) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (650 - 700 (650) long; 450 - 470 (450) wide) ellipsoid and colorless, occasionally with pink coloration without a distinct pattern. Anterio-medial platelets (115 - 125 (115) long; 55 - 60 (55) wide). Anterio-lateral platelets (177.5 - 180 (177.5) long; 67.5 - 67.5 (67.5) wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 345 - 375 (345 )). Dorsal plate proportions: dorsum length / width 1.44 - 1.49 (1.44); dorsal width / distance between Dgl- 4 1.25 - 1.30 (1.30); anterio-medial platelet length / width 2.08 - 2.09 (2.09); anterio-lateral platelet length / width 2.63 - 2.67 (2.63); anterio-lateral / anterio-medial length 1.44 - 1.54 (1.54). Gnathosoma - Subcapitulum (277.5 - 300 (277.5) long (ventral); 202.5 - 221.23 (202.5) long (dorsal); 127.5 - 138.75 (127.5) tall) colorless. Rostrum (120 - 132.5 (120) long; 52.5 - 55 (52.5) wide) short and conical. Chelicerae (260 - 291 (260) long) with curved fangs (75 - 77 (75) long). Subcapitular proportions: ventral length / height 2.16 - 2.18 (2.18); rostrum length / width 2.29 - 2.41 (2.29). Pedipalps without extensions on femora and genua. Palpomeres: trochanter (50 - 55 (50) long); femur (130 - 137.5 (130) long); genu (72.5 - 80 (72.5) long); tibia (102.5 - 110 (102.5) long; 27.5 - 30 (27.5) wide); tarsus (25 - 27.5 (27.5) long). Palpomere proportions: femur / genu 1.72 - 1.79 (1.79); tibia / femur 0.79 - 0.80 (0.79); tibia length / width 3.67 - 3.73 (3.73). Venter - (790 - 800 (800) long; 480 - 551 (480) wide) colorless. Gnathosomal bay (122.5 - 130 (130) long; 85 - 105 (85) wide). Cxgl- 4 subapical. Medial suture (30 - 30 (30) long). Genital plates (182.5 - 188.75 (188.75) long; 150 - 152.5 (150) wide). Additional measurements: Cx- 1 (257 - 260 (260) long (total); 120 - 129 (120) long (medial )); Cx- 3 (330 - 390 (330) wide); anterior venter (162.5 - 167.5 (162.5) long). Ventral proportions: gnathosomal bay length / width 1.17 - 1.53 (1.53); anterior venter / genital field length 0.86 - 0.92 (0.86); anterior venter length / genital field width 1.08 - 1.10 (1.08); anterior venter / medial suture 5.42 - 5.58 (5.42). Male (Figure 95) (n = 5) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (580 - 640 (640) long; 390 - 410 (400) wide) ellipsoid and colorless. Anterio-medial platelets (110 - 117.5 (115) long; 55 - 60 (60) wide). Anterio-lateral platelets (167.5 - 175 (175) long; 65 - 72.5 (70) wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 320 - 360 (325 )). Dorsal plate proportions: dorsum length / width 1.43 - 1.60 (1.60); dorsal width / distance between Dgl- 4 1.14 - 1.23 (1.23); anterio-medial platelet length / width 1.91 - 2.05 (1.92); anterio-lateral platelet length / width 2.31 - 2.58 (2.50); anterio-lateral / anterio-medial length 1.43 - 1.59 (1.52). Gnathosoma - Subcapitulum (245 - 280 (280) long (ventral); 185 - 202.5 (200) long (dorsal); 107.5 - 120 (120) tall) colorless. Rostrum (107.5 - 110 (110) long; 42.5 - 45 (45) wide). Chelicerae (245 - 260 (260) long) with curved fangs (65 - 70 (67.5) long). Subcapitular proportions: ventral length / height 2.23 - 2.36 (2.33); rostrum length / width 2.44 - 2.59 (2.44). Pedipalps without extensions on femora and genua. Palpomeres: trochanter (37.5 - 47.5 (45) long); femur (117.5 - 125 (125) long); genu (70 - 75 (75) long); tibia (97.5 - 100 (100) long; 22.5 - 25 (25) wide); tarsus (22.5 - 27.5 (27.5) long). Palpomere proportions: femur / genu 1.67 - 1.73 (1.67); tibia / femur 0.79 - 0.83 (0.80); tibia length / width 3.90 - 4.33 (4.00). Venter - (670 - 740 (740) long; 450 - 495 (460) wide) colorless. Gnathosomal bay (120 - 130 (125) long; 80 - 85 (85) wide). Cxgl- 4 subapical. Medial suture (102.5 - 122.5 (117.5) long). Genital plates (140 - 150 (147.5) long; 102.5 - 110 (105) wide). Additional measurements: Cx- 1 (240 - 260 (260) long (total); 120 - 150 (130) long (medial )); Cx- 3 (330 - 350 (345) wide); anterior venter (237.5 - 270 (270) long). Ventral proportions: gnathosomal bay length / width 1.47 - 1.59 (1.47); anterior venter / genital field length 1.70 - 1.84 (1.83); anterior venter length / genital field width 2.21 - 2.57 (2.57); anterior venter / medial suture 2.14 - 2.51 (2.30). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
1A004CD99F8EDC0E851EF02F1D3BE6D8.taxon	etymology	Etymology. Specific epithet (hoosieri) refers to Hoosier, the English demonym for a person from Indiana, the type locality.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
1A004CD99F8EDC0E851EF02F1D3BE6D8.taxon	distribution	Distribution. Known only from Wayne County, Indiana (Figure 93).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
5966A4325168B2AD6FDC633309C6EC31.taxon	materials_examined	Material examined. LECTOTYPE (1 ♀): from USA, Wisconsin, Green Lake County, Green Lake, Aug 1921, by C Juday, RM 210013. PARALECTOTYPE (1 ♂): from USA, Wisconsin, Green Lake County, Green Lake, Aug 1921, by C Juday, RM 210013. OTHER MATERIAL (5 ♀; 4 ♂): Manitoba, Canada: 1 ♂ from Fidler Lake; Station 7., (57 ° 11 ' 11 " N, 96 ° 56 ' 56 " W), 20 June 1977, by Freshwater Institute, IMS 770231 * 1 ♀ from Northern Indian Lake; Station 9., (56 ° 47 ' 47 " N, 98 ° 56 ' 56 " W), 20 June 1977, by Freshwater Institute, IMS 770224 * 1 ♂ from Northern Indian Lake; Station 1., (56 ° 47 ' 47 " N, 98 ° 56 ' 56 " W), 20 June 1977, by Freshwater Institute, IMS 770232 * 1 ♀ from North Pine River near Pine River, 29 May 1981, by P Schefter, E Fuller, ROM 810578 * 1 ♀ and 1 ♂ from Southern Indian Lake, (57 ° 10 ' 10 " N, 98 ° 29 ' 29 " W), 1 July 1977, by Freshwater Institute, IMS 770234 * 1 ♀ and 1 ♂ from Southern Indian Lake, (56 ° 47 ' 47 " N, 98 ° 56 ' 56 " W), 27 July 1977, by Freshwater Institute., IMS 770215 * 1 ♀ from Southern Indian Lake, (57 ° 10 ' 10 " N, 98 ° 29 ' 29 " W), 5 September 1978, by Freshwater Institute., IMS 780049	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
5966A4325168B2AD6FDC633309C6EC31.taxon	diagnosis	Diagnosis. Torrenticola indistincta are similar to other members of the Rusetria " Eastern 2 - Plates " group (T. biscutella, T. caerulea, T. delicatexa, T. malarkeyorum, T. pendula, T. sellersorum, T. tysoni, T. ululata, T. whitneyae, T. microbiscutella, and T. feminellai) in having anterio-lateral platelets fused to the dorsal plate, and being distributed in the east. T. indistincta can be differentiated from other Eastern 2 - Plates by having faint coloration separated into anterior and posterior portions connected medially. Female T. indistincta can be differentiated from female T. caerulea by having a larger genital field (length = 185 - 225 in T. indistincta, 155 - 165 in T. caerulea; width = 185 - 205 in T. indistincta, 120 - 145 in T. caerulea). Male T. indistincta can be differentiated from male T. caerulea by having a larger dorsum (length = 480 - 645 in T. indistincta, 405 - 460 in T. caerulea; width = 315 - 470 in T. indistincta, 260 - 305 in T. caerulea). T. indistincta can be differentiated from T. microbiscutella by having a less elongate dorsum (length / width = 1.21 - 1.52 in T. indistincta; 1.63 - 1.75 in T. microbiscutella).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
5966A4325168B2AD6FDC633309C6EC31.taxon	description	Re-description. Female (Figure 97) (n = 5) (lectotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (590 - 880 (640) long; 460 - 720 (485) wide) ovoid with faint coloration separated into anterior and posterior portions and connected medially. Anterio-medial platelets (125 - 195 (125) long; 45 - 77.5 (47.5) wide). Anterio-lateral platelets (172.5 - 235 (172.5) long; 75 - 115 (80) wide) fused to dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 340 - 485 (340 )). Dorsal plate proportions: dorsum length / width 1.21 - 1.32 (1.32); dorsal width / distance between Dgl- 4 1.32 - 1.48 (1.43); anterio-medial platelet length / width 2.35 - 2.94 (2.63); anterio-lateral platelet length / width 2.02 - 2.33 (2.16); anterio-lateral / anterio-medial length 1.19 - 1.38 (1.38). Gnathosoma - Subcapitulum (300 - 395 (322.5) long (ventral); 225 - 295 (237.5) long (dorsal); 145 - 195 (150) tall) tall and colorless. Rostrum (120 - 160 (125) long; 45 - 62.5 (47.5) wide). Chelicerae (300 - 405 (335) long) with curved fangs (60 - 80 (62.5) long). Subcapitular proportions: ventral length / height 1.97 - 2.15 (2.15); rostrum length / width 2.50 - 2.78 (2.63). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (45 - 65 (47.5) long); femur (112.5 - 160 (117.5) long); genu (62.5 - 90 (62.5) long); tibia (87.5 - 122.5 (87.5) long; 25 - 31.25 (25) wide); tarsus (17.5 - 30 (17.5) long). Palpomere proportions: femur / genu 1.67 - 1.88 (1.88); tibia / femur 0.74 - 0.80 (0.74); tibia length / width 3.50 - 3.92 (3.50). Venter - (650 - 995 (800) long; 535 - 880 (565) wide) colorless. Gnathosomal bay (175 - 237.5 (175) long; 90 - 130 (97.5) wide). Cxgl- 4 subapical. Medial suture absent. Genital plates (185 - 225 (205) long; 185 - 205 (190) wide). Additional measurements: Cx- 1 (295 - 385 (305) long (total); 120 - 155 (135) long (medial )); Cx- 3 (345 - 505 (345) wide); anterior venter (125 - 167.5 (135) long). Ventral proportions: gnathosomal bay length / width 1.79 - 2.02 (1.79); anterior venter / genital field length 0.66 - 0.83 (0.66); anterior venter length / genital field width 0.66 - 0.83 (0.71). Male (Figure 98) (n = 5) (lectotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (480 - 645 (480) long; 315 - 470 (315) wide) ovoid with faint coloration separated into anterior and posterior portions and connected medially. Anterio-medial platelets (102.5 - 147.5 (102.5) long; 35 - 55 (35) wide). Anterio-lateral platelets (137.5 - 180 (150) long; 55 - 82.5 (55) wide) fused to dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 235 - 350 (235 )). Dorsal plate proportions: dorsum length / width 1.37 - 1.52 (1.52); dorsal width / distance between Dgl- 4 1.31 - 1.34 (1.34); anterio-medial platelet length / width 2.33 - 3.05 (2.93); anterio-lateral platelet length / width 2.12 - 2.73 (2.73); anterio-lateral / anterio-medial length 1.19 - 1.46 (1.46). Gnathosoma - Subcapitulum (238.75 - 317.5 (238.75) long (ventral); 167.5 - 232.5 (167.5) long (dorsal); 102.5 - 137.5 (102.5) tall) tall and colorless. Rostrum (85 - 125 (85) long; 32.5 - 50 (32.5) wide). Chelicerae (232.5 - 320 (232.5) long) with curved fangs (47.5 - 65 (47.5) long). Subcapitular proportions: ventral length / height 2.22 - 2.33 (2.33); rostrum length / width 2.50 - 2.86 (2.62). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (37.5 - 52.5 (37.5) long); femur (86.25 - 117.5 (86.25) long); genu (50 - 77.5 (50) long); tibia (70 - 100 (70) long; 22.5 - 28.75 (22.5) wide); tarsus (17.5 - 22.5 (17.5) long). Palpomere proportions: femur / genu 1.52 - 1.73 (1.73); tibia / femur 0.80 - 0.85 (0.81); tibia length / width 3.11 - 3.48 (3.11). Venter - (570 - 780 (570) long; 370 - 510 (370) wide) colorless. Gnathosomal bay (127.5 - 180 (127.5) long; 65 - 100 (65) wide). Cxgl- 4 subapical. Medial suture (65 - 75 (75) long). Genital plates (117.5 - 165 (117.5) long; 112.5 - 150 (112.5) wide). Additional measurements: Cx- 1 (235 - 315 (235) long (total); 107.5 - 150 (107.5) long (medial )); Cx- 3 (250 - 395 (277.5) wide); anterior venter (190 - 235 (190) long). Ventral proportions: gnathosomal bay length / width 1.80 - 1.96 (1.96); anterior venter / genital field length 1.38 - 1.68 (1.62); anterior venter length / genital field width 1.53 - 1.69 (1.69); anterior venter / medial suture 2.53 - 3.54 (2.53). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
5966A4325168B2AD6FDC633309C6EC31.taxon	etymology	Etymology. Marshall (1929) named the specific epithet (indistincta) in reference to the Rusetria Complex character of fused (" indistinct ") lateral platelets.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
5966A4325168B2AD6FDC633309C6EC31.taxon	distribution	Distribution. Midwest and into Manitoba (Figure 96). Young (1969) reported this species from Colorado, but this likely represents T. mulleni rather than T. indistincta.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
5DB52660B32A1C59C587D04F897B7826.taxon	materials_examined	Material examined. HOLOTYPE (♂): from USA, Arkansas, Montgomery County, Caddo Gap, access track off Manfred Road, 0.3 km west of Route 8, 29 Jul 2011, by IM Smith, IMS 110037. PARATYPES (7 ♀; 11 ♂): Arkansas, USA: 1 ♀ (ALLOTYPE) from Montgomery County, Caddo Gap, access track off Manfred Road, 0.3 km west of Route 8, 29 Jul 2011, by IM Smith, IMS 110037 * 2 ♂ from Montgomery County, Caddo Gap, access track off Manfred Road, 0.3 km west of Route 8, 29 Jul 2011, by IM Smith, IMS 110037 * 1 ♀ and 1 ♂ from Newton County, Ozark-St. Francis National Forest, Little Buffalo River, 11 Jul 2012, by TD Edwards, TDE 12 - 0711 - 004 * Missouri, USA: 3 ♀ and 5 ♂ from Crawford County, Huzzah Creek, Red Bluff campground, off Road V, east of Davisville, 23 Jul 2011, by IM Smith, IMS 110029 * Oklahoma, USA: 2 ♀ and 3 ♂ from Pushmataha County, Walnut Creek, beside Route 271, south of Albion, 1 Jul 1987, by IM Smith, IMS 870063 A.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
5DB52660B32A1C59C587D04F897B7826.taxon	diagnosis	Diagnosis. Torrenticola interiorensis are similar to species with similar dorsal patterning, such as the Rusetria 4 - Plate Group (T. dunni, T. glomerabilis, T. kittatinniana, T. pollani, T. rufoalba and T. shubini), Elongata Group (T. gorti, T. elongata, and T. reduncarostra), and T. bondi, T. erectirostra, T. robisoni, T. irapalpa, T. neoanomala, T. racupalpa, T. skvarlai, and T. arktonyx. They can be differentiated from Rusetria 4 - Plates and T. skvarlai by having distinct hind coxal margins. T. interiorensis can be differentiated from T. erectirostra, T. robisoni and T. reduncarostra by having a straight, anteriorly-directed rostrum (upturned in T. erectirostra and T. reduncarostra). T. interiorensis can be differentiated from T. arktonyx by having an unmodified dorsal plate (T. arktonyx has distinctive longitudinal dark markings on the anterior portion of the dorsal plate). T. interiorensis can be differentiated from T. racupalpa by having less elongate pedipalpal tibiae (length / width = 3.76 - 4.22 in T. interiorensis; 4.44 - 5.50 in T. racupalpa) and less elongate rostrum (length / width = 2.63 - 2.88 in T. interiorensis; 3.56 - 3.88 in T. racupalpa). T. interiorensis can be differentiated from T. irapalpa by having Dgl- 4 closer to the dorsal edge (dorsal width / distance between Dgl- 4 ♀ = 1.48 - 1.61 in T. interiorensis, 1.81 - 2.09 in T. irapalpa; ♂ = 1.42 - 1.45 in T. interiorensis, 1.58 - 1.86 in T. irapalpa) and more ovoid dorsum (length / width ♀ = 1.29 - 1.38 in T. interiorensis, 1.20 - 1.28 in T. irapalpa; ♂ = 1.34 - 1.47 in T. interiorensis, 1.26 - 1.30 in T. irapalpa). T. interiorensis can be differentiated from Elongata Group by being slightly more ovoid (dorsum length / width ♀ = 1.29 - 1.38 in T. interiorensis, 1.45 - 2.08 in Elongata Group; ♂ = 1.34 - 1.47 in T. interiorensis, 1.51 - 1.7 in Elongata Group) and having a stockier rostrum (length / width = 2.63 - 2.88 in T. interiorensis, 3.24 - 4.00 in Elongata Group). T. interiorensis can be differentiated from T. bondi by having a longer medial suture (♀ = 25 - 30 in T. interiorensis, 10 - 15 in T. bondi; ♂ = 75 - 83 in T. interiorensis, 55 - 70 in T. bondi), anterior venter / genital field width (♀ = 1.31 - 1.38 in T. interiorensis, 1.15 - 1.25 in T. bondi; ♂ = 2.09 - 2.27 in T. interiorensis, 1.95 - 2.05 in T. bondi), and Dgl- 4 closer to edge of dorsum (dorsum width / distance between Dgl- 4 ♀ = 1.48 - 1.61 in T. interiorensis, 1.63 - 1.72 in T. bondi; ♂ = 1.42 - 1.45 in T. interiorensis, 1.48 - 1.62 in T. bondi). Female T. interiorensis can be differentiated from female T. neoanomala by having stockier anterio-lateral platelets (length / width = 2.62 - 2.67 in T. interiorensis, 2.86 - 3.09 in T. neoanomala). Male T. interiorensis can be differentiated from male T. neoanomala by having a shorter anterior venter (220 - 240 in T. interiorensis, 267.5 - 290 in T. neoanomala) and a shorter genital field (132 - 138 in T. interiorensis, 145 - 160 in T. neoanomala).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
5DB52660B32A1C59C587D04F897B7826.taxon	description	Description. Male (Figure 100) (n = 5) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (510 - 545 (510) long; 350 - 405 (360) wide) ovoid with purple coloration separated into anterior and posterior portions and orange medially. Anterio-medial platelets (115 - 122.5 (115) long; 45 - 50 (45) wide). Anterio-lateral platelets (150 - 167.5 (150) long; 50 - 52.5 (50) wide) free from dorsal plate. Dgl- 4 closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 245 - 285 (250 )). Dorsal plate proportions: dorsum length / width 1.34 - 1.47 (1.42); dorsal width / distance between Dgl- 4 1.42 - 1.45 (1.44); anterio-medial platelet length / width 2.45 - 2.72 (2.56); anterio-lateral platelet length / width 3.00 - 3.19 (3.00); anterio-lateral / anterio-medial length 1.30 - 1.37 (1.30). Gnathosoma - Subcapitulum (265 - 282.5 (265) long (ventral); 200 - 212 (200) long (dorsal); 105 - 112.5 (105) tall) colorless. Rostrum (112.5 - 115 (112.5) long; 40 - 40 (40) wide). Chelicerae (260 - 282 (260) long) with curved fangs (45 - 58 (45) long). Subcapitular proportions: ventral length / height 2.51 - 2.60 (2.52); rostrum length / width 2.81 - 2.88 (2.81). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (32.5 - 37.5 (32.5) long); femur (97.5 - 105 (97.5) long); genu (55 - 62.5 (55) long); tibia (77.5 - 81.25 (77.5) long; 20 - 21.25 (20) wide); tarsus (17.5 - 18.75 (17.5) long). Palpomere proportions: femur / genu 1.68 - 1.78 (1.77); tibia / femur 0.77 - 0.79 (0.79); tibia length / width 3.76 - 3.88 (3.88). Venter - (608 - 660 (640) long; 424 - 480 (480) wide) mostly colorless with faint purple in areas surrounding coxae. Gnathosomal bay (102.5 - 137.5 (110) long; 65 - 85 (75) wide). Cxgl- 4 subapical. Medial suture (75 - 82.5 (75) long). Genital plates (132.5 - 137.5 (132.5) long; 100 - 107.5 (100) wide). Additional measurements: Cx- 1 (231 - 270 (260) long (total); 123.25 - 134 (130) long (medial )); Cx- 3 (300 - 338 (315) wide); anterior venter (220 - 240 (220) long). Ventral proportions: gnathosomal bay length / width 1.37 - 2.12 (1.47); anterior venter / genital field length 1.60 - 1.80 (1.66); anterior venter length / genital field width 2.09 - 2.27 (2.20); anterior venter / medial suture 2.73 - 2.98 (2.93). Female (Figure 101) (n = 5) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (550 - 620 (550) long; 410 - 480 (415) wide) ovoid with purple coloration separated into anterior and posterior portions and orange coloration. Anterio-medial platelets (123.75 - 142.5 (123.75) long; 50 - 60 (50) wide). Anterio-lateral platelets (160 - 187.5 (160) long; 60 - 70 (60) wide) free from dorsal plate. Dgl- 4 closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 255 - 305 (280 )). Dorsal plate proportions: dorsum length / width 1.29 - 1.38 (1.33); dorsal width / distance between Dgl- 4 1.48 - 1.61 (1.48); anterio-medial platelet length / width 2.36 - 2.52 (2.48); anterio-lateral platelet length / width 2.62 - 2.68 (2.67); anterio-lateral / anterio-medial length 1.21 - 1.33 (1.29). Gnathosoma - Subcapitulum (307.5 - 325 (310) long (ventral); 220 - 240 (232.5) long (dorsal); 120 - 130 (120) tall) colorless. Rostrum (122.5 - 140 (125) long; 45 - 50 (45) wide). Chelicerae (300 - 340 (310) long) with curved fangs (50 - 65 (55) long). Subcapitular proportions: ventral length / height 2.44 - 2.58 (2.58); rostrum length / width 2.63 - 2.83 (2.78). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (40 - 45 (40) long); femur (112.5 - 127.5 (115) long); genu (65 - 70 (65) long); tibia (85 - 95 (85) long; 21.25 - 22.5 (21.25) wide); tarsus (17.5 - 22.5 (20) long). Palpomere proportions: femur / genu 1.73 - 1.82 (1.77); tibia / femur 0.74 - 0.76 (0.74); tibia length / width 3.78 - 4.22 (4.00). Venter - (680 - 780 (680) long; 470 - 580 (510) wide) mostly colorless with purple in areas surrounding coxae. Gnathosomal bay (112.5 - 152.5 (145) long; 70 - 100 (90) wide). Cxgl- 4 subapical. Medial suture (25 - 30 (30) long). Genital plates (155 - 175 (155) long; 137.5 - 152.5 (145) wide). Additional measurements: Cx- 1 (242 - 310 (280) long (total); 125 - 160 (140) long (medial )); Cx- 3 (320 - 390 (350) wide); anterior venter (190 - 205 (190) long). Ventral proportions: gnathosomal bay length / width 1.22 - 2.07 (1.61); anterior venter / genital field length 1.17 - 1.23 (1.23); anterior venter length / genital field width 1.31 - 1.38 (1.31); anterior venter / medial suture 6.33 - 8.20 (6.33). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
5DB52660B32A1C59C587D04F897B7826.taxon	etymology	Etymology. Specific epithet (interiorensis) refers to the Interior Highlands, where this species was found within both major regions (Ozarks and Ouachitas), but not found outside these regions, which suggests it is endemic to the region.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
5DB52660B32A1C59C587D04F897B7826.taxon	distribution	Distribution. Interior Highlands (Ozarks and Ouachitas), likely endemic (Figure 99).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
1A4005F3A3258B383069F3A7EDCCB0A6.taxon	materials_examined	Material examined. HOLOTYPE (♀): from New Brunswick, Canada, York County, Stanley, Nashwaak River, Stanley Municipal Park, 19 Jun 2012, by IM Smith, IMS 120031, DNA 2956. PARATYPES (39 ♀; 26 ♂): Arkansas, USA: 1 ♀ from Newton County, Buffalo National River, Cecil Creek (36 ° 5 ' 15.72 " N, 93 ° 13 ' 23.28 " W), 13 Jun 2012, by TD Edwards, TDE 12 - 0613 - 010 * 1 ♂ from Newton County, Ozark-St. Francis National Forest, Little Buffalo River, 11 Jun 2012, by TD Edwards, TDE 12 - 0711 - 004 * 2 ♀ from Polk County, beside Forest Route 38, north of Shady Lake Recreation Area, East Saline Creek (34 ° 22 ' 23.39 " N, 94 ° 1 ' 51.22 " W), 30 Jul 2011, by IM Smith, IMS 110041 * Kentucky, USA: 2 ♀ and 1 ♂ from McCreary County, White Oak Junction, Rock Creek, beside Forest Route 556, 2.3 kilometers south of Route 1363, 8 Jul 1990, by IM Smith, IMS 900082 A & IMS 900082 B * Illinois, USA: 3 ♀ and 1 ♂ from Kankakee County, Kankakee River State Park, beside Route 102, Rock Creek (41 ° 12 ' N, 88 ° 0 ' W), 9 Sep 1991, by IM Smith, IMS 910027 A * Indiana, USA: 1 ♀ from Wayne County, south of intersection of Interstate 70 and Route 1 (39 ° 51 ' 13 " N, 85 ° 8 ' 4 " W), 31 Jul 2014, by MJ Skvarla, MS 14 - 0731 - 001 * Missouri, USA: 3 ♀ Wayne County, beside Road 143 near Sam A Baker State Park, 8 Jul 1960, by DR Cook, DRC 600010 * New Brunswick, Canada: 2 ♀ and 1 ♂ from Charlotte County, Rollingdam, Digdeguash River, beside Highway 770 at covered bridge, 30 Jun 1989, by IM Smith, IMS 890053 * 1 ♀ and 1 ♂ from Charlotte County, Digdeguash River, beside Sorrel Ridge Road, west of Whittier Road, 10 Jun 2012, by IM Smith, IMS 120015 * 2 ♀ and 1 ♂ from Charlotte County, Rollingham, Digdegaush River, beside Highway 770, 3 Oct 2011, by IM Smith, IMS 110118 * 1 ♂ (ALLOTYPE) from York County, Stanley, Nashwaak River, Stanley Municipal Park, 19 Jun 2012, by IM Smith, IMS 120031, DNA 2957 * Ohio, USA: 1 ♂ from Montgomery County, Engelwood Metro Park (39 ° 52 ' 58 " N, 84 ° 17 ' 33 " W), 31 Jul 2014, by MJ Skvarla, MS 14 - 0731 - 002 * 1 ♀ and 2 ♂ from Pickaway County, beside Scioto-Darby Road, just north of Darby, Big Darby Creek (40 ° 2 ' N, 83 ° 15 ' W), 30 Jun 1997, by IM Smith & M MacKenzie, IMS 970016 * Ontario, Canada: 1 ♀ and 1 ♂ from Grey County, Durham, Saugeen River, beside County Road 27 near Durham Conservation Area, 9 Jun 1989, by IM Smith, IMS 890028 A * Pennsylvania, USA: 1 ♀ from Bedford County, Chaneysville, Sweet Root Picnic Area beside Route 326, 18 Jul 1990, by IM Smith, IMS 900105 * 1 ♀ from Huntingdon County, Alan Seeger Natural Area, Stone Creek, beside road from McAlevys Fort to Route 322, 19 Jul 1990, by IM Smith, IMS 900107 * 1 ♂ from Tioga County, Straight Run, beside Straight Run Road, 2.1 kilometers north of Route 6 between Ansonia & Wellsboro, 20 Jul 1990, by IM Smith, IMS 900111 * Saskatchewan, Canada: 2 ♀ and 2 ♂ from Torch River beside Highway 106 at access road, 30 kilometers north of Highway 55 at Smeaton, 30 Jul 1988, by IM Smith, IMS 880054 A * 2 ♀ and 2 ♂ from Torch River at end of forest access road, 10 kilometers north of Highway 55 at Love, 30 Jul 1988, by IM Smith, IMS 880056 * Texas, USA: 2 ♀ from Bandera County, Vanderpool, beside Route 187, Sabinal River (29 ° 48 ' 10 " N, 99 ° 34 ' 30 " W), 2 May 2009, by IM Smith, IMS 090007 * 1 ♀ and 1 ♂ from Kinney County, Brackettville, beside Route 90, 12.1 kilometers west of Route 131, Pinto Creek (29 ° 20 ' 6 " N, 100 ° 32 ' 5 " W), 28 Sep 1994, by IM Smith, IMS 940025 * 1 ♀ from Uvalde County, Garner State Park, Frio River (29 ° 35 ' 22 " N, 99 ° 44 ' 12 " W), 28 May 1998, by IM Smith, IMS 980027 A * 1 ♀ and 1 ♂ from Val Verde County, Bakers Crossing Campground off Route 163, Devils River (29 ° 58 ' N, 101 ° 9 ' W), 5 Oct 1999, by IM Smith, IMS 990061 A * 2 ♀ from Val Verde County, Dolan Falls Preserve, Devils River (29 ° 53 ' 12 " N, 100 ° 59 ' 37 " W), 24 May 2011, by IM Smith, IMS 110013 * 4 ♀ and 4 ♂ from Val Verde County, Dolan Falls Preserve, Snake Spring (29 ° 53 ' 43 " N, 100 ° 58 ' 58 " W), 25 May 2011, by IM Smith, IMS 110015 * Virginia, USA: 1 ♀ and 1 ♂ from Bath County, Jackson River, beside Forest Route 1843 (continuation of Route 623), 3.5 kilometers south of Route 220, 16 Jul 1990, by IM Smith, IMS 900100 * 2 ♀ and 2 ♂ from Scott County, North Fork Holston River, beside Route 58 / 421, 0.9 kilometers east of Route 709 at Hiltons, 7 Jul 1990, by IM Smith, IMS 900080 * West Virginia, USA: 1 ♀ and 1 ♂ from Pendleton County, South Branch of the Potomac River, beside Route 28 / 55, 20.8 kilometers southwest of Route 42, 17 Jul 1990, by IM Smith, IMS 900104.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
1A4005F3A3258B383069F3A7EDCCB0A6.taxon	diagnosis	Diagnosis. Torrenticola irapalpa are similar to other members of the Raptor Group (T. gnoma, T. longitibia, T. mjolniri, T. elusiva, T. racupalpa, T. raptor, T. danielleae, T. daemon, and T. ivyae) in having round bodies; Dgl- 4 close to muscles scars; long, thin subcapitular rostra; and long, thin pedipalp tibiae. T. irapalpa can be differentiated from T. longitibia, T. mjolniri, T. elusiva, T. racupalpa, and T. ivyae by having Dgl- 4 closer to the dorsal edge (dorsal width / distance between Dgl- 4 = 1.81 - 2.09 in T. irapalpa, 2.19 - 2.77 in others) and a less elongate rostrum (length / width = 2.66 - 3.39 in T. irapalpa; 3.56 - 4.32 in others). T. irapalpa can be differentiated from T. gnoma by having Dgl- 4 closer to the dorsal edge (dorsal width / distance between Dgl- 4 ♀ = 1.81 - 2.09 in T. irapalpa, 2.65 - 3.29 in T. gnoma; ♂ = 1.58 - 1.86 in T. irapalpa, 2.06 - 2.73 in T. gnoma) and by dorsal coloration and pattern. T. irapalpa can be differentiated from T. raptor by having less elongate tibiae (length / width ♀ = 4.09 - 5.67 in T. irapalpa, 6.00 - 7.54 in T. raptor; ♂ = 4.25 - 4.75 in T. irapalpa, 5.29 - 5.63 in T. raptor); less elongate subcapitulum (ventral length / height = 2.52 - 2.90 in T. irapalpa, 2.98 - 3.27 in T. raptor); and by dorsal pattern. T. irapalpa can be differentiated from T. danielleae by having Dgl- 4 further from the dorsal edge (dorsal width / distance between Dgl- 4 = ♀ = 1.81 - 2.09 in T. irapalpa, 1.57 - 1.70 in T. danielleae; ♂ = 1.58 - 1.86 in T. irapalpa, 1.42 - 1.52 in T. danielleae); a less elongate rostrum (length / width = 2.66 - 3.39 in T. irapalpa, 3.43 - 3.75 in T. danielleae); and by dorsal coloration and pattern. Female T. irapalpa can be differentiated from female T. daemon by having Dgl- 4 further from the dorsal edge (dorsal width / distance between Dgl- 4 ♀ = 1.81 - 2.09 in T. irapalpa, 1.59 - 1.67 in T. daemon) and a less elongate gnathosomal bay (length / width ♀ = 1.35 - 1.86 in T. irapalpa, 1.95 - 2.42 in T. daemon). Male T. irapalpa can be differentiated from male T. daemon by dorsal coloration and pattern.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
1A4005F3A3258B383069F3A7EDCCB0A6.taxon	description	Description. Female (Figure 103) (n = 9) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (575 - 710 (610) long; 465 - 580 (475) wide) circular with navy blue to bluish-purple coloration separated into anterior and posterior portions with bold orange medially. Anterio-medial platelets (132.5 - 160 (137.5) long; 52.5 - 67.5 (62.5) wide). Anterio-lateral platelets (155 - 197.5 (177.5) long; 67.5 - 87.5 (77.5) wide) free from dorsal plate. Dgl- 4 midway between muscle scars and dorsum edge (distance between Dgl- 4 465 - 580 (475 )). Dorsal plate proportions: dorsum length / width 1.20 - 1.28 (1.28); dorsal width / distance between Dgl- 4 1.81 - 2.09 (1.90); anterio-medial platelet length / width 2.12 - 2.52 (2.20); anterio-lateral platelet length / width 2.17 - 2.39 (2.29); anterio-lateral / anterio-medial length 1.17 - 1.42 (1.29). Gnathosoma - Subcapitulum (300 - 360 (340) long (ventral); 219 - 270 (270) long (dorsal); 125 - 157.5 (145) tall) with faint navy blue to bluish-purple coloration, sometimes colorless. Rostrum (126.25 - 152.5 (142.5) long; 55 - 76 (65) wide) elongate. Chelicerae (245 - 360 (360) long) with curved fangs (55 - 75 (65) long). Subcapitular proportions: ventral length / height 2.29 - 2.57 (2.34); rostrum length / width 2.66 - 3.39 (3.17). Pedipalps elongate (especially tibiae) with long tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (45 - 52.5 (45) long); femur (115 - 132.5 (132.5) long); genu (65 - 80 (72.5) long); tibia (100 - 117.5 (102.5) long; 17.5 - 20 (20) wide); tarsus (17.5 - 20 (20) long). Palpomere proportions: femur / genu 1.66 - 1.89 (1.83); tibia / femur 0.77 - 0.89 (0.77); tibia length / width 4.09 - 5.67 (4.82). Venter - (690 - 870 (770) long; 508 - 743 (550) wide) with navy blue to bluish purple coloration. Gnathosomal bay (132.5 - 165 (165) long; 87.5 - 120 (90) wide). Cxgl- 4 far from apex. Medial suture (10 - 22.5 (15) long). Genital plates (155 - 175 (175) long; 140 - 162.5 (150) wide). Additional measurements: Cx- 1 (266 - 334 (310) long (total); 103 - 156 (140) long (medial )); Cx- 3 (332 - 487 (375) wide); anterior venter (156 - 197.5 (182.5) long). Ventral proportions: gnathosomal bay length / width 1.35 - 1.86 (1.83); anterior venter / genital field length 0.99 - 1.18 (1.04); anterior venter length / genital field width 1.06 - 1.27 (1.22); anterior venter / medial suture 8.56 - 15.63 (12.17). Male (Figure 104) (n = 8) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (520 - 620 (560) long; 400 - 490 (435) wide) circular with navy blue to bluish-purple coloration separated into anterior and posterior portions with bold orange medially. Anterio-medial platelets (115 - 150 (140) long; 55 - 67.5 (62.5) wide). Anterio-lateral platelets (160 - 188.75 (187.5) long; 62.5 - 77.5 (75) wide) free from dorsal plate. Dgl- 4 midway between muscle scars and dorsum edge (distance between Dgl- 4 220 - 310 (250 )). Dorsal plate proportions: dorsum length / width 1.26 - 1.30 (1.29); dorsal width / distance between Dgl- 41.58 - 1.86 (1.74); anterio-medial platelet length / width 2.00 - 2.48 (2.24); anterio-lateral platelet length / width 2.34 - 2.60 (2.50); anterio-lateral / anterio-medial length 1.22 - 1.41 (1.34). Gnathosoma - Subcapitulum (275 - 340 (310) long (ventral); 207 - 240 (240) long (dorsal); 105 - 132.5 (107.5) tall) with faint navy blue to bluish-purple coloration, sometimes colorless. Rostrum (112.5 - 137.5 (135) long; 40 - 47.5 (40) wide) elongate. Chelicerae (252 - 305 (305) long) with curved fangs (51 - 60 (55) long). Subcapitular proportions: ventral length / height 2.52 - 2.90 (2.88); rostrum length / width 2.81 - 3.38 (3.38). Pedipalps elongate (especially tibiae) with long tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (35 - 50 (42.5) long); femur (107.5 - 123.75 (115) long); genu (60 - 70 (67.5) long); tibia (87.5 - 108.75 (95) long; 20 - 25 (20) wide); tarsus (18.75 - 20 (20) long). Palpomere proportions: femur / genu 1.70 - 1.83 (1.70); tibia / femur 0.81 - 0.91 (0.83); tibia length / width 4.25 - 4.75 (4.75). Venter - (630 - 705 (690) long; 490 - 540 (500) wide) with navy blue to bluish purple coloration. Gnathosomal bay (135 - 177.5 (146.25) long; 75 - 95 (75) wide). Cxgl- 4 far from apex. Medial suture (50 - 75 (75) long). Genital plates (127.5 - 158.75 (143.75) long; 100 - 137.5 (122.5) wide). Additional measurements: Cx- 1 (260 - 290 (290) long (total); 137 - 155 (155) long (medial )); Cx- 3 (323 - 360 (345) wide); anterior venter (215 - 255 (245) long). Ventral proportions: gnathosomal bay length / width 1.42 - 2.06 (1.95); anterior venter / genital field length 1.51 - 1.75 (1.70); anterior venter length / genital field width 1.78 - 2.20 (2.00); anterior venter / medial suture 3.27 - 4.40 (3.27). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
1A4005F3A3258B383069F3A7EDCCB0A6.taxon	etymology	Etymology. Specific epithet (irapalpa) refers to the pedipalps of this species, which resemble ferocious weapons of wrath (ira, L. fury, rage, wrath; palpus, L. hand, feeler).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
1A4005F3A3258B383069F3A7EDCCB0A6.taxon	distribution	Distribution. Primarily eastern (Figure 102), but note that T. irapalpa is one of few Torrenticola found in the Great Plains, at least in the south (Texas) and north (Saskatchewan).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
7F32B68934AC97A78B27EFFE72817CDA.taxon	materials_examined	Material examined. HOLOTYPE (♀): from USA, Florida, Polk County, beside Rt. 471 at junction of Pasco, Polk and Sumter County lines, (28 ° 19 ' 19 " N, 82 ° 4 ' 4 " W), 24 April 1992, by IM Smith, IMS 920003 PARATYPES (5 ♀; 5 ♂): Florida, USA: 2 ♀ from Hillsborough County, beside Rt. 41 / 301 near Hillsborough River State Park, south of Zephyrhills, (28 ° 10 ' 10 " N, 82 ° 12 ' 12 " W), 24 April 1992, by IM Smith, IMS 920001 * 2 ♀ and 3 ♂ from Pasco County, beside Rt. 39 just north of Crystal Springs, (28 ° 11 ' 11 " N, 82 ° 10 ' 10 " W), 27 April 1992, by IM Smith, IMS 920011 * 1 ♂ (ALLOTYPE) from Polk County, beside Rt. 471 at junction of Pasco, Polk and Sumter County lines, (28 ° 19 ' 19 " N, 82 ° 4 ' 4 " W), 24 April 1992, by IM Smith, IMS 920003 * 1 ♀ and 1 ♂ from Polk County, beside Rucks Dairy Road 1.3 km south of Lake Arbuckle Road, (27 ° 42 ' 42 " N, 81 ° 26 ' 26 " W), 25 April 1992, by IM Smith, IMS 920004	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
7F32B68934AC97A78B27EFFE72817CDA.taxon	diagnosis	Diagnosis. Torrenticola ivyae are similar to other members of the Raptor Group (T. gnoma, T. irapalpa, T. longitibia, T. mjolniri, T. elusiva, T. racupalpa, T. raptor, T. danielleae, and T. daemon) in having round bodies; Dgl- 4 close to muscles scars; long, thin subcapitular rostra; and long, thin pedipalp tibiae. T. ivyae can be differentiated from T. gnoma, T. elusiva, T. danielleae, and T. daemon by having a more elongate rostrum (length / width ♀ = 4.00 - 4.15 in T. ivyae, 2.74 - 3.75 in others; ♂ = 3.85 - 4.08 in T. ivyae, 2.56 - 3.57 in others) and more elongate pedipalpal tibiae (length / width ♀ = 5.07 - 5.64 in T. ivyae, 4.05 - 5.00 in others, ♂ = 4.75 - 5.20 in T. ivyae, 3.88 - 4.44 in others). T. ivyae can be differentiated from T. irapalpa, T. raptor, T. danielleae, and T. daemon by having Dgl- 4 closer to the muscle scars (dorsum width / distance between Dgl- 4 = 2.20 - 2.75 in T. ivyae, 1.42 - 2.09 in others). T. ivyae can be differentiated from T. longitibia (known only from males) by femur / genu (1.83 - 1.88 in T. ivyae, 2.1 - 2.17 in T. longitibia) and having less elongate tibiae (length / width = 4.75 - 5.20 in T. ivyae, 5.50 - 5.50 in T. longitibia). Female T. ivyae can be differentiated from female T. racupalpa by having a more elongate rostrum (length / width = 4.00 - 4.15 in T. ivyae, 3.56 - 3.82 in T. racupalpa) and more elongate pedipalpal tibiae (length / width = 5.07 - 5.64 in T. ivyae, 4.44 - 5.00 in T. racupalpa). Male T. ivyae can be differentiated from male T. racupalpa by having a longer anterior venter (♂ 220 - 230 in T. ivyae, 200 - 205 in T. racupalpa) and a shorter genital field (♂ 142 - 148 in T. ivyae, 160 - 165 in T. racupalpa). Female T. ivyae can be differentiated from female T. mjolniri by having a smaller dorsum (length ♀ = 550 - 590 in T. ivyae, 605 - 640 in T. mjolniri; width ♀ = 460 - 500 in T. ivyae, 510 - 545 in T. mjolniri) and a shorter anterior venter (♀ 155 - 170 in T. ivyae, 180 - 195 in T. mjolniri). Male T. ivyae can be differentiated from male T. mjolniri by having a less elongate subcapitulum (ventral length / width = 2.57 - 2.75 in T. ivyae, 2.82 - 3.00 in T. mjolniri). Additionally, T. ivyae can be differentiated from T. mjolniri by being found in Florida (T. ivyae is known from the northeast).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
7F32B68934AC97A78B27EFFE72817CDA.taxon	description	Description. Female (Figure 106) (n = 5) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (550 - 590 (580) long; 460 - 500 (495) wide) circular with reddish-purple coloration anteriorly and posteriorly connected medially. Anterio-medial platelets (120 - 135 (120) long; 60 - 62.5 (62.5) wide). Anterio-lateral platelets (165 - 180 (165) long; 65 - 75 (75) wide) free from dorsal plate. Dgl- 4 much closer to the muscle scars than to the edge of the dorsum (distance between Dgl- 4 180 - 195 (180 )). Dorsal plate proportions: dorsum length / width 1.16 - 1.20 (1.17); dorsal width / distance between Dgl- 4 2.42 - 2.75 (2.75); anterio-medial platelet length / width 1.92 - 2.25 (1.92); anterio-lateral platelet length / width 2.20 - 2.62 (2.20); anterio-lateral / anterio-medial length 1.26 - 1.38 (1.38). Gnathosoma - Subcapitulum (300 - 325 (325) long (ventral); 232.5 - 257.5 (247.5) long (dorsal); 115 - 127.5 (120) tall) colorless. Rostrum (130 - 145 (145) long; 32.5 - 35 (35) wide) elongate. Chelicerae (315 - 340 (335) long) with curved fangs (55 - 60 (60) long). Subcapitular proportions: ventral length / height 2.55 - 2.71 (2.71); rostrum length / width 4.00 - 4.15 (4.14). Pedipalps elongate with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (42.5 - 46.25 (45) long); femur (117.5 - 125 (125) long); genu (65 - 67.5 (65) long); tibia (91.25 - 100 (95) long; 17.5 - 18.75 (18.75) wide); tarsus (15 - 17.5 (17.5) long). Palpomere proportions: femur / genu 1.81 - 1.92 (1.92); tibia / femur 0.76 - 0.83 (0.76); tibia length / width 5.07 - 5.64 (5.07). Venter - (630 - 755 (750) long; 490 - 540 (540) wide) with reddish-purple coloration. Gnathosomal bay (150 - 175 (165) long; 67.5 - 75 (72.5) wide). Cxgl- 4 far from apex. Medial suture (25 - 25 (25) long). Genital plates (150 - 165 (157.5) long; 142.5 - 150 (150) wide). Additional measurements: Cx- 1 (280 - 300 (300) long (total); 120 - 145 (145) long (medial )); Cx- 3 (320 - 335 (325) wide); anterior venter (155 - 170 (170) long). Ventral proportions: gnathosomal bay length / width 2.00 - 2.44 (2.28); anterior venter / genital field length 0.94 - 1.13 (1.08); anterior venter length / genital field width 1.03 - 1.19 (1.13); anterior venter / medial suture 6.20 - 6.80 (6.80). Male (Figure 107) (n = 4) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (530 - 550 (550) long; 390 - 450 (420) wide) circular with reddish-purple coloration anteriorly and posteriorly connected medially. Anterio-medial platelets (125 - 135 (130) long; 55 - 62.5 (57.5) wide). Anterio-lateral platelets (160 - 190 (160) long; 60 - 75 (60) wide) free from dorsal plate. Dgl- 4 much closer to the muscle scars than to the edge of the dorsum (distance between Dgl- 4 165 - 205 (180 )). Dorsal plate proportions: dorsum length / width 1.19 - 1.36 (1.31); dorsal width / distance between Dgl- 4 2.20 - 2.39 (2.33); anterio-medial platelet length / width 2.00 - 2.45 (2.26); anterio-lateral platelet length / width 2.27 - 2.92 (2.67); anterio-lateral / anterio-medial length 1.23 - 1.46 (1.23). Gnathosoma - Subcapitulum (295 - 302.5 (300) long (ventral); 220 - 230 (220) long (dorsal); 110 - 115 (110) tall) colorless. Rostrum (125 - 132.5 (130) long; 32.5 - 32.5 (32.5) wide) elongate. Chelicerae (295 - 305 (295) long) with curved fangs (45 - 50 (50) long). Subcapitular proportions: ventral length / height 2.57 - 2.75 (2.73); rostrum length / width 3.85 - 4.08 (4.00). Pedipalps elongate with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (40 - 42.5 (40) long); femur (110 - 112.5 (110) long); genu (60 - 60 (60) long); tibia (90 - 97.5 (90) long; 17.5 - 20 (17.5) wide); tarsus (15 - 16.25 (16.25) long). Palpomere proportions: femur / genu 1.83 - 1.88 (1.83); tibia / femur 0.82 - 0.87 (0.82); tibia length / width 4.75 - 5.20 (5.14). Venter - (625 - 700 (700) long; 470 - 500 (470) wide) with reddish-purple coloration. Gnathosomal bay (140 - 150 (142.5) long; 62.5 - 72.5 (65) wide). Cxgl- 4 far from apex. Medial suture (60 - 80 (75) long). Genital plates (142.5 - 147.5 (145) long; 112.5 - 125 (120) wide). Additional measurements: Cx- 1 (280 - 290 (280) long (total); 125 - 145 (145) long (medial )); Cx- 3 (310 - 330 (310) wide); anterior venter (220 - 230 (230) long). Ventral proportions: gnathosomal bay length / width 1.93 - 2.36 (2.19); anterior venter / genital field length 1.52 - 1.59 (1.59); anterior venter length / genital field width 1.76 - 2.00 (1.92); anterior venter / medial suture 2.75 - 3.67 (3.07). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
7F32B68934AC97A78B27EFFE72817CDA.taxon	etymology	Etymology. Specific epithet (ivyae) named in honor of Ivy Fisher, our (JRF and DMF) beautiful daughter, who was born in Florida, the type locality.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
7F32B68934AC97A78B27EFFE72817CDA.taxon	distribution	Distribution. Florida (Figure 105).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
CE1105A2125662200413D784B3DE7C1D.taxon	materials_examined	Material examined. HOLOTYPE (♀): from USA, Tennessee, Sevier County, Great Smokey Mountains National Park, Sugarlands Nature Trail (35 ° 40 ' 47 " N, 83 ° 31 ' 51 " W), 10 Sep 2010, by IM Smith, IMS 100125, DNA 1758. PARATYPES (1 ♀; 3 ♂): 1 ♂ (ALLOTYPE) from Sevier County, Great Smokey Mountains National Park, Sugarlands Nature Trail (35 ° 40 ' 47 " N, 83 ° 31 ' 51 " W), 10 Sep 2010, by IM Smith, IMS 100125, DNA 1846 * 1 ♀ and 2 ♂ from Sevier County, Great Smokey Mountains National Park, Sugarlands Nature Trail (35 ° 40 ' 47 " N, 83 ° 31 ' 51 " W), 10 Sep 2010, by IM Smith, IMS 100125	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
CE1105A2125662200413D784B3DE7C1D.taxon	diagnosis	Diagnosis. Torrenticola karambita is similar to other members of the Erectirostra Group (T. erectirostra and T. robisoni) in having an upturned rostrum that is wide when viewed ventrally. T. karambita can be differentiated from T. erectirostra and T. robisoni by lacking coloration (T. erectirostra has purplish dorsal coloration) and a slightly stockier rostrum (length / width ♀ = 1.57 - 1.62 in T. karambita, 1.72 - 2.09 in others; ♂ = 1.6 - 1.95 in T. karambita, 2.0 - 2.2 in T. erectirostra).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
CE1105A2125662200413D784B3DE7C1D.taxon	description	Description. Female (Figure 109) (n = 2) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (690 - 725 (690) long; 490 - 500 (490) wide) ovoid and colorless. Anterio-medial platelets (145 - 162.5 (145) long; 65 - 75 (65) wide). Anterio-lateral platelets (217.5 - 220 (217.5) long; 80 - 82.5 (82.5) wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 340 - 360 (340 )). Dorsal plate proportions: dorsum length / width 1.41 - 1.45 (1.41); dorsal width / distance between Dgl- 4 1.39 - 1.44 (1.44); anterio-medial platelet length / width 2.17 - 2.23 (2.23); anterio-lateral platelet length / width 2.64 - 2.75 (2.64); anterio-lateral / anterio-medial length 1.35 - 1.50 (1.50). Gnathosoma - Subcapitulum (325 - 350 (325) long (ventral); 229 - 254 (229) long (dorsal); 135 - 135 (135) tall) colorless. Rostrum (110 - 117.5 (110) long; 70 - 72.5 (70) wide) wide and unturned with dentation. Chelicerae (320 - 329 (321) long) with curved fangs (40 - 52 (41) long). Subcapitular proportions: ventral length / height 2.41 - 2.59 (2.41); rostrum length / width 1.57 - 1.62 (1.57). Pedipalps short and stocky (especially tibiae) with short tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (52.5 - 55 (52.5) long); femur (101.25 - 105 (101.25) long); genu (65 - 67.5 (67.5) long); tibia (55 - 57.5 (55) long; 30 - 30 (30) wide); tarsus (17.5 - 17.5 (17.5) long). Palpomere proportions: femur / genu 1.50 - 1.62 (1.50); tibia / femur 0.54 - 0.55 (0.54); tibia length / width 1.83 - 1.92 (1.83). Venter - (850 - 850 (850) long; 595 - 607 (607) wide) colorless. Gnathosomal bay (170 - 172.5 (172.5) long; 125 - 130 (130) wide). Cxgl- 4 far from apex. Medial suture (15 - 17.5 (15) long). Genital plates (195 - 197.5 (195) long; 165 - 175 (165) wide). Additional measurements: Cx- 1 (336 - 348 (336) long (total); 159 - 175 (174) long (medial )); Cx- 3 (415 - 441 (415) wide); anterior venter (202.5 - 222.5 (202.5) long). Ventral proportions: gnathosomal bay length / width 1.33 - 1.36 (1.33); anterior venter / genital field length 1.04 - 1.13 (1.04); anterior venter length / genital field width 1.23 - 1.27 (1.23); anterior venter / medial suture 12.71 - 13.50 (13.50). Male (Figure 110) (n = 3) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum- (610 - 655 (610) long; 410 - 440 (420) wide) ovoid and colorless. Anterio-medial platelets (133.75 - 145 (133.75) long; 65 - 67.5 (65) wide). Anterio-lateral platelets (205 - 217.5 (207.5) long; 75 - 77.5 (75) wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 290 - 330 (290 )). Dorsal plate proportions: dorsum length / width 1.44 - 1.60 (1.45); dorsal width / distance between Dgl- 4 1.33 - 1.45 (1.45); anterio-medial platelet length / width 2.06 - 2.19 (2.06); anterio-lateral platelet length / width 2.65 - 2.81 (2.77); anterio-lateral / anterio-medial length 1.44 - 1.55 (1.55). Gnathosoma - Subcapitulum (285 - 290 (290) long (ventral); 202 - 210 (202) long (dorsal); 108.75 - 115 (112.5) tall) colorless. Rostrum (97.5 - 102.5 (97.5) long; 52.5 - 60 (60) wide) wide and unturned with dentation. Chelicerae (268 - 269 (269) long) with curved fangs (43 - 52 (47) long). Subcapitular proportions: ventral length / height 2.48 - 2.67 (2.58); rostrum length / width 1.63 - 1.95 (1.63). Pedipalps short and stocky (especially tibiae) with short tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (47.5 - 48.75 (47.5) long); femur (87.5 - 95 (95) long); genu (57.5 - 62.5 (62.5) long); tibia (55 - 57.5 (57.5) long; 27.5 - 27.5 (27.5) wide); tarsus (15 - 17.5 (17.5) long). Palpomere proportions: femur / genu 1.44 - 1.52 (1.52); tibia / femur 0.61 - 0.63 (0.61); tibia length / width 2.00 - 2.09 (2.09). Venter - (760 - 785 (760) long; 490 - 521 (512) wide) colorless. Gnathosomal bay (165 - 175 (165) long; 105 - 107.5 (107.5) wide). Cxgl- 4 far from apex. Medial suture (70 - 77.5 (77.5) long). Genital plates (156.25 - 167.5 (157.5) long; 120 - 125 (125) wide). Additional measurements: Cx- 1 (284 - 320 (284) long (total); 115 - 154 (116) long (medial )); Cx- 3 (366 - 373 (372) wide); anterior venter (247.5 - 248.75 (248.75) long). Ventral proportions: gnathosomal bay length / width 1.53 - 1.67 (1.53); anterior venter / genital field length 1.48 - 1.58 (1.58); anterior venter length / genital field width 1.99 - 2.06 (1.99); anterior venter / medial suture 3.19 - 3.54 (3.21). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
CE1105A2125662200413D784B3DE7C1D.taxon	etymology	Etymology. Specific epithet (karambita) refers to the upturned rostrum that has dentation on both sides in females, which resembles a karambit-small, recurved knives used in the Filipino martial arts practiced by JRF. The karambit is thought to have originated with the Minangkabau people of West Sumatra based upon a similarity to a tiger's claws.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
CE1105A2125662200413D784B3DE7C1D.taxon	distribution	Distribution. Known only from Great Smokey Mountains National Park, Sevier County, Tennessee (Figure 108).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
2DFA012C5FE5127F31DB73395C4E76C3.taxon	materials_examined	Material examined (6 ♀; 10 ♂). Arizona, USA: 1 ♀ and 1 ♂ from Cochise County, Chiricahua Mountains; beside Forest Road 42 near junction with Forest Road 42 B, (31 ° 55 ' 55 " N, 109 ° 15 ' 15 " W), 16 July 1987, by IM Smith, IMS 870093 A * 1 ♀ and 1 ♂ from Cochise County, Chiricahua Mountains; Sycamore Campground east of Sunizona, (31 ° 52 ' 52 " N, 109 ° 20 ' 20 " W), 15 July 1987, by IM Smith, IMS 870091 * 3 ♂ from Cochise County, Chiricahua Mountains; Cave Creek Recreation Area; John Hands Picnic Area off Forest Road 42 A west of Portal, (31 ° 53 ' 53 " N, 109 ° 13 ' 13 " W), 15 July 1987, by IM Smith, IMS 870092 A * 1 ♀ and 1 ♂ from Coconino County, Oak Creek Canyon; beside Rt. 89 A just north of Pine Flat Campground, (35 ° 1 ' 1 " N, 111 ° 44 ' 44 " W), 21 July 1987, by IM Smith, IMS 870099 A * 1 ♀ from Coconino County, Oak Creek Canyon; beside Rt. 89 A between Bootlegger and Banjo Bill Campgrounds, (34 ° 58 ' 58 " N, 111 ° 45 ' 45 " W), 21 July 1987, by IM Smith, IMS 870100 B * 1 ♂ from Coconino County, Oak Creek Canyon; beside Rt. 89 A just north of Pine Flat Campground, (35 ° 1 ' 1 " N, 111 ° 44 ' 44 " W), 21 July 1987, by IM Smith, IMS 870099 B * New Mexico, USA: 2 ♀ from Catron County, Glenwood; Whitewater Picnic Area 8 km east of Rt. 180, (33 ° 22 ' 22 " N, 108 ° 50 ' 50 " W), 12 July 1987, by IM Smith, IMS 870084 * 2 ♂ from Catron County, beside Rt. 15, 65 km north of Rt. 180 (Silver City), (33 ° 12 ' 12 " N, 108 ° 13 ' 13 " W), 10 July 1987, by IM Smith, IMS 870081 A * 1 ♂ from Catron County, beside Rt. 15, 65 km north of Rt. 180 (Silver City), (33 ° 12 ' 12 " N, 108 ° 13 ' 13 " W), 10 July 1987, by IM Smith, IMS 870081 B.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
2DFA012C5FE5127F31DB73395C4E76C3.taxon	diagnosis	Diagnosis. Torrenticola keesdavidsi are similar to other members of the Rala Group (T. rala, T. lamellipalpis, T. boettgeri, T. kurtvietsi, T. dolichodactyla, and T. anoplopalpa) by being colorless, having incomplete hind coxal margins and being distributed in the southwest. T. keesdavidsi can be differentiated from all other Rala Group by having a shorter dorsum (length ♀ = 555 - 605 in T. keesdavidsi, 630 - 800 in others; ♂ = 500 - 590 in T. keesdavidsi, 600 - 780 in others) and dentate, flanged ventral extensions on the femora (others are lacking extensions, have tuberculate extensions, or flat, wide lamellate extensions). Additionally, T. keesdavidsi can be differentiated from all other Rala Group by having more elongate pedipalpal tibiae (length / width = 4.50 - 5.20 in T. keesdavidsi, 1.75 - 3.38 in others), except T. lamellipalpis (4.32 - 4.94).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
2DFA012C5FE5127F31DB73395C4E76C3.taxon	description	Re-description. Female (Figure 112) (n = 5) with characters of the genus with following specifications. Dorsum - (555 - 605 long; 450 - 495 wide) circular and colorless. Anterio-medial platelets (135 - 137.5 long; 50 - 60 wide). Anterio-lateral platelets (180 - 192.5 long; 70 - 80 wide) free from dorsal plate. Dgl- 4 closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 315 - 335). Dorsal plate proportions: dorsum length / width 1.21 - 1.31; dorsal width / distance between Dgl- 4 1.41 - 1.48; anterio-medial platelet length / width 2.25 - 2.70; anterio-lateral platelet length / width 2.38 - 2.75; anterio-lateral / anterio-medial length 1.33 - 1.43. Gnathosoma - Subcapitulum (312.5 - 327.5 long (ventral); 222.5 - 240 long (dorsal); 97.5 - 105 tall) colorless. Rostrum (120 - 130 long; 35 - 37.5 wide). Chelicerae (295 - 310 long) with curved fangs (45 - 50 long). Subcapitular proportions: ventral length / height 3.07 - 3.21; rostrum length / width 3.33 - 3.57. Pedipalps with dentate, flanged ventral extensions on femora and genua. Palpomeres: trochanter (35 - 37.5 long); femur (105 - 110 long); genu (55 - 57.5 long); tibia (90 - 95 long; 18.75 - 20 wide); tarsus (15 - 17.5 long). Palpomere proportions: femur / genu 1.91 - 2.00; tibia / femur 0.84 - 0.88; tibia length / width 4.50 - 4.93. Venter - (695 - 755 long; 510 - 560 wide) colorless. Gnathosomal bay (145 - 150 long; 67.5 - 80 wide). Cxgl- 4 far from apex. Medial suture (35 - 55 long). Genital plates (162.5 - 175 long; 152.5 - 160 wide). Additional measurements: Cx- 1 (285 - 295 long (total); 140 - 150 long (medial )); Cx- 3 (320 - 330 wide); anterior venter (192.5 - 222.5 long). Ventral proportions: gnathosomal bay length / width 1.88 - 2.19; anterior venter / genital field length 1.18 - 1.29; anterior venter length / genital field width 1.26 - 1.44; anterior venter / medial suture 4.05 - 5.50. Male (Figure 113) (n = 5) with characters of the genus with following specifications. Dorsum - (500 - 590 long; 410 - 450 wide) circular and colorless. Anterio-medial platelets (120 - 130 long; 55 - 60 wide). Anterio-lateral platelets (165 - 190 long; 65 - 75 wide) free from dorsal plate. Dgl- 4 closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 280 - 320). Dorsal plate proportions: dorsum length / width 1.21 - 1.33; dorsal width / distance between Dgl- 4 1.40 - 1.48; anterio-medial platelet length / width 2.09 - 2.27; anterio-lateral platelet length / width 2.33 - 2.71; anterio-lateral / anterio-medial length 1.32 - 1.48. Gnathosoma - Subcapitulum (280 - 310 long (ventral); 202.5 - 230 long (dorsal); 85 - 105 tall) colorless. Rostrum (107.5 - 125 long; 32.5 - 35 wide). Chelicerae (260 - 297.5 long) with curved fangs (40 - 50 long). Subcapitular proportions: ventral length / height 2.81 - 3.29; rostrum length / width 3.29 - 3.57. Pedipalps with dentate, flanged ventral extensions on femora and genua. Palpomeres: trochanter (32.5 - 35 long); femur (97.5 - 110 long); genu (50 - 56.25 long); tibia (87.5 - 97.5 long; 17.5 - 20 wide); tarsus (15 - 17.5 long). Palpomere proportions: femur / genu 1.84 - 1.96; tibia / femur 0.86 - 0.95; tibia length / width 4.81 - 5.20. Venter - (630 - 720 long; 460 - 510 wide) colorless. Gnathosomal bay (130 - 152.5 long; 60 - 75 wide). Cxgl- 4 far from apex. Medial suture (80 - 95 long). Genital plates (140 - 155 long; 115 - 125 wide). Additional measurements: Cx- 1 (260 - 280 long (total); 130 - 150 long (medial )); Cx- 3 (280 - 340 wide); anterior venter (225 - 260 long). Ventral proportions: gnathosomal bay length / width 1.93 - 2.17; anterior venter / genital field length 1.55 - 1.86; anterior venter length / genital field width 1.96 - 2.17; anterior venter / medial suture 2.61 - 2.89. Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
2DFA012C5FE5127F31DB73395C4E76C3.taxon	etymology	Etymology. Cramer (1992) named this species in honor of Kees Davids " as an acknowledgement of his great contributions to acarology and aquatic ecology " (translated from Spanish).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
2DFA012C5FE5127F31DB73395C4E76C3.taxon	distribution	Distribution. Southwest (Arizona and New Mexico), extending southward into Mexico (Figure 111).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
3568A3D5D2E55937C17429283D9881A8.taxon	materials_examined	Material examined. HOLOTYPE (♂): from USA, New Jersey, Sussex County, Little Flatbrook, north of Bevans, 12 Oct 1953, by H Habeeb, HH 530110. PARATYPES (1 ♀ and 0 ♂): New Jersey, USA: 1 ♀ (ALLOTYPE) from Morris County, Brook, Brookside, 20 May 1953, by H Habeeb, HH 530045.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
3568A3D5D2E55937C17429283D9881A8.taxon	diagnosis	Diagnosis. Torrenticola kittatinniana are similar to other members of the Rusetria " 4 - Plate " group (T. dunni, T. glomerabilis, T. pollani, T. rufoalba and T. shubini) and T. skvarlai in having anterio-lateral platelets free from the dorsal plate, dorsal coloration separated into anterior and posterior portions, and indistinct hind coxal margins. Female T. kittatinniana can be differentiated from female T. dunni by having shorter pedipalpal genua (64 in T. kittatinniana, 70 - 75 in T. dunni); a shorter subcapitulum (ventral length = 310 in T. kittatinniana, 330 - 355 in T. dunni); and stockier anterio-medial platelets (length / width = 2.83 in T. kittatinniana, 2.33 - 2.54 in T. dunni). Male T. kittatinniana can be differentiated from male T. dunni by having a shorter anterior venter (235 in T. kittatinniana, 277 - 285 in T. dunni) and thinner dorsum (340 in T. kittatinniana, 350 - 370 in T. dunni). T. kittatinniana can be differentiated from T. pollani by having a stockier rostrum (length / width = 2.71 - 3.16 in T. kittatinniana, 3.27 - 3.82 in T. pollani) and stockier tibiae (length / width ♀ = 3.3 in T. kittatinniana, 3.8 - 4.2 in T. pollani; ♂ = 2.80 in T. kittatinniana, 3.4 - 3.8 in T. pollani). Female T. kittatinniana can be differentiated from female T. shubini by having a more elongate rostrum (length / width = 3.16 in T. kittatinniana, 2.5 - 2.7 in T. shubini) and a shorter subcapitulum (125 in T. kittatinniana, 140 - 145 in T. shubini). Male T. kittatinniana can be differentiated from male T. shubini by having a longer dorsum (500 in T. kittatinniana, 400 - 465 in T. shubini) and a longer genital field (115 in T. kittatinniana, 90 - 108 in T. shubini). T. kittatinniana can be differentiated from T. glomerabilis by having Dgl- 4 closer to the dorsal edge (dorsal width / distance between Dgl- 4 = 1.2 - 1.4 in T. kittatinniana, 1.5 - 1.7 in T. glomerabilis) and stockier tibiae (length / width ♀ = 3.3 in T. kittatinniana, 4.11 - 4.5 in T. glomerabilis; ♂ = 2.8 in T. kittatinniana, 3.5 - 4.4 in T. glomerabilis). T. kittatinniana can be differentiated from T. rufoalba by having a longer dorsum (♀ = 640 in T. kittatinniana, 550 in T. rufoalba; ♂ = 500 in T. kittatinniana, 440 in T. rufoalba) and more elongate anterio-medial platelets (length / width = 2.83 - 2.88 in T. kittatinniana, 2.45 - 2.61 in T. rufoalba). Additionally, male T. kittatinniana have a longer anterior venter (235 in T. kittatinniana, 195 in T. rufoalba). T. kittatinniana can be differentiated from T. skvarlai by having a conical pedipalpal femoral tubercle, whereas T. skvarlai has a broad and flat pedipalpal femoral tubercle, and by having a longer anterior venter (♀ = 165 in T. kittatinniana, 140 - 152.5 in T. skvarlai; ♂ = 235 in T. kittatinniana, 177.5 - 205 in T. skvarlai).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
3568A3D5D2E55937C17429283D9881A8.taxon	description	Re-description. Female (Figure 115) (n = 1) (allotype only) with characters of the genus with following specifications. Dorsum - (550 long; 400 wide) ovoid with purple coloration separated into anterior and posterior portions bordered with orange. Anterio-medial platelets (107.5 long; 41.25 wide). Anterio-lateral platelets (168.75 long; 55 wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 255). Dorsal plate proportions: dorsum length / width 1.38; dorsal width / distance between Dgl- 4 1.57; anterio-medial platelet length / width 2.61; anterio-lateral platelet length / width 3.07; anterio-lateral / anterio-medial length 1.57. Gnathosoma- Subcapitulum (310 long (ventral); 235 long (dorsal); 127.5 tall) colorless. Rostrum (130 long; 42.5 wide). Chelicerae (315 long) with curved fangs (62.5 long). Subcapitular proportions: ventral length / height 2.43; rostrum length / width 3.06. Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (42.5 long); femur (115 long); genu (65 long); tibia (87.5 long; 25 wide); tarsus (17.5 long). Palpomere proportions: femur / genu 1.77; tibia / femur 0.76; tibia length / width 3.50. Venter - (640 long; 450 wide) mostly colorless with faint purple coloration in areas surrounding coxae. Gnathosomal bay (142.5 long; 92.5 wide). Cxgl- 4 subapical. Medial suture (17.5 long). Genital plates (167.5 long; 155 wide). Additional measurements: Cx- 1 (125 long (total); 125 long (medial )); Cx- 3 (335 wide); anterior venter (155 long). Ventral proportions: gnathosomal bay length / width 1.54; anterior venter / genital field length 0.93; anterior venter length / genital field width 1.00; anterior venter / medial suture 8.86. Male (Figure 116) (n = 1) (holotype only) with characters of the genus with following specifications. Dorsum - (500 long; 340 wide) ovoid with purple coloration separated into anterior and posterior portions bordered with orange. Anterio-medial platelets (90 long; 31.25 wide). Anterio-lateral platelets (140 long; 47.5 wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 280). Dorsal plate proportions: dorsum length / width 1.47; dorsal width / distance between Dgl- 4 1.21; anterio-medial platelet length / width 2.88; anterio-lateral platelet length / width 2.95; anterio-lateral / anterio-medial length 1.56. Gnathosoma - Subcapitulum (237.5 long (ventral); 180 long (dorsal); 100 tall) colorless. Rostrum (95 long; 35 wide). Chelicerae (222.5 long) with curved fangs (45 long). Subcapitular proportions: ventral length / height 2.38; rostrum length / width 2.71. Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (37.5 long); femur (87.5 long); genu (57.5 long); tibia (70 long; 25 wide); tarsus (17.5 long). Palpomere proportions: femur / genu 1.52; tibia / femur 0.80; tibia length / width 2.80. Venter - (600 long; 435 wide) mostly colorless with faint purple coloration in areas surrounding coxae. Gnathosomal bay (107.5 long; 72.5) wide). Cxgl- 4 subapical. Medial suture (102.5 long). Genital plates (115 long; 82.5 wide). Additional measurements: Cx- 1 (235 long (total); 125 long (medial )); Cx- 3 (285 wide); anterior venter (235 long). Ventral proportions: gnathosomal bay length / width 1.48; anterior venter / genital field length 2.04; anterior venter length / genital field width 2.85; anterior venter / medial suture 2.29. Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
3568A3D5D2E55937C17429283D9881A8.taxon	etymology	Etymology. Habeeb (1955) did not specify an explanation for the specific epithet (kittatinniana). However, we suspect it refers to the type locality, which is located in the Great Kittatinny Valley, which is named for Kittatinny Mountain in northwestern New Jersey, a northern extension of the Appalachian Ridge and Valley province.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
3568A3D5D2E55937C17429283D9881A8.taxon	distribution	Distribution. Northern New Jersey (Figure 114).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
B18D1A316B2968C9B3E6E7DA203477D1.taxon	materials_examined	Material examined. HOLOTYPE (♀): from USA, Texas, Tyler County, Spurger; beside Farm Road 1013, 8.2 km west of Rt. 92, (30 ° 41 ' 41 " N, 94 ° 15 ' 15 " W), 30 September 1994, by IM Smith, IMS 940027 A. PARATYPES (4 ♀; 5 ♂): Texas, USA: 1 ♂ (ALLOTYPE) from Tyler County, Spurger; beside Farm Road 1013, 8.2 km west of Rt. 92, (30 ° 41 ' 41 " N, 94 ° 15 ' 15 " W), 30 September 1994, by IM Smith, IMS 940027 A * 4 ♀ and 4 ♂ from Tyler, Spurger; beside Farm Road 1013, 8.2 km west of Rt. 92, (30 ° 41 ' 41 " N, 94 ° 15 ' 15 " W), 30 September 1994, by IM Smith, IMS 940027 A.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
B18D1A316B2968C9B3E6E7DA203477D1.taxon	diagnosis	Diagnosis. Torrenticola kringi are similar to other members of the Tricolor Complex (T. bittikoferae, T. hoosieri, T. larvata, T. pearsoni, T. olliei, T. sierrensis, T. tricolor, T. trimaculata, T. unimaculata, T. cardia, T. dimorpha, and T. mohawk) in having a short, conical rostrum. T. kringi can be differentiated from most Torrenticola, including other members of the Tricolor Complex, by having a distinct dorsal pattern of a large anterior dorsal spot. The only other species with this pattern is T. ululata, which, like all Rusetria 2 - Plates, have anterio-lateral platelets fused to the dorsal plate and T. unimaculata, which has a stockier rostrum (length / width = 2.6 - 3.2 in T. kringi, 1.9 - 2.2 in T. unimaculata). T. kringi are most similar to other members of the Tricolor Complex that have bold patterning (T. larvata, T. tricolor, T. trimaculata, T. unimaculata, T. cardia, and T. mohawk). T. kringi can be further differentiated from all other members of the Tricolor Complex by having a more elongate rostrum (length / width = 2.67 - 3.13 in T. kringi, 1.29 - 2.59).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
B18D1A316B2968C9B3E6E7DA203477D1.taxon	description	Description. Female (Figure 118) (n = 5) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (550 - 580 (580) long; 460 - 500 (490) wide) circular with coloration restricted to a single dark spot anteriorly (occasionally extending medially), with an orange spot posterior to the dark spot. Anterio-medial platelets (125 - 145 (145) long; 60 - 70 (70) wide). Anterio-lateral platelets (160 - 185 (185) long; 65 - 75 (75) wide) free to dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 335 - 365 (350 )). Dorsal plate proportions: dorsum length / width 1.15 - 1.20 (1.18); dorsal width / distance between Dgl- 4 1.26 - 1.46 (1.40); anterio-medial platelet length / width 2.00 - 2.25 (2.07); anterio-lateral platelet length / width 2.29 - 2.69 (2.47); anterio-lateral / anterio-medial length 1.19 - 1.40 (1.28). Gnathosoma - Subcapitulum (290 - 302.5 (300) long (ventral); 215 - 227.5 (227.5) long (dorsal); 117.5 - 125 (125) tall) colorless. Rostrum (120 - 130 (125) long; 40 - 45 (42.5) wide). Chelicerae (275 - 300 (290) long) with curved fangs (50 - 55 (50) long). Subcapitular proportions: ventral length / height 2.34 - 2.47 (2.40); rostrum length / width 2.67 - 3.13 (2.94). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (40 - 42.5 (41.25) long); femur (107.5 - 112.5 (112.5) long); genu (72.5 - 77.5 (77.5) long); tibia (91.25 - 97.5 (96.25) long; 27.5 - 28.75 (28.75) wide); tarsus (25 - 26.25 (25) long). Palpomere proportions: femur / genu 1.43 - 1.48 (1.45); tibia / femur 0.85 - 0.89 (0.86); tibia length / width 3.32 - 3.55 (3.35). Venter - (630 - 690 (680) long; 500 - 530 (525) wide) colorless. Gnathosomal bay (132.5 - 152.5 (145) long; 75 - 82.5 (82.5) wide). Cxgl- 4 subapical. Medial suture (22.5 - 30 (30) long). Genital plates (165 - 180 (180) long; 150 - 160 (160) wide). Additional measurements: Cx- 1 (240 - 270 (270) long (total); 110 - 130 (130) long (medial )); Cx- 3 (320 - 345 (345) wide); anterior venter (145 - 170 (170) long). Ventral proportions: gnathosomal bay length / width 1.67 - 1.94 (1.76); anterior venter / genital field length 0.85 - 0.97 (0.94); anterior venter length / genital field width 0.92 - 1.07 (1.06); anterior venter / medial suture 5.17 - 6.44 (5.67). Male (Figure 119) (n = 5) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (480 - 560 (510) long; 395 - 465 (420) wide) circular with coloration restricted to a single dark spot anteriorly (occasionally extending medially), with an orange spot posterior to the dark spot. Anterio-medial platelets (110 - 130 (117.5) long; 52.5 - 65 (57.5) wide). Anterio-lateral platelets (140 - 170 (150) long; 50 - 75 (60) wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 285 - 350 (315 )). Dorsal plate proportions: dorsum length / width 1.19 - 1.30 (1.21); dorsal width / distance between Dgl- 4 1.29 - 1.39 (1.33); anterio-medial platelet length / width 1.96 - 2.27 (2.04); anterio-lateral platelet length / width 2.27 - 2.80 (2.50); anterio-lateral / anterio-medial length 1.27 - 1.31 (1.28). Gnathosoma - Subcapitulum (240 - 275 (260) long (ventral); 177.5 - 210 (195) long (dorsal); 90 - 105 (100) tall) colorless. Rostrum (100 - 115 (110) long; 35 - 40 (37.5) wide). Chelicerae (225 - 270 (260) long) with curved fangs (45 - 50 (50) long). Subcapitular proportions: ventral length / height 2.55 - 2.75 (2.60); rostrum length / width 2.81 - 2.93 (2.93). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (35 - 38.75 (37.5) long); femur (85 - 101.25 (97.5) long); genu (60 - 71.25 (67.5) long); tibia (70 - 82.5 (77.5) long; 23.75 - 27.5 (26.25) wide); tarsus (22.5 - 25 (23.75) long). Palpomere proportions: femur / genu 1.36 - 1.44 (1.44); tibia / femur 0.79 - 0.87 (0.79); tibia length / width 2.95 - 3.14 (2.95). Venter - (590 - 695 (640) long; 440 - 510 (470) colorless. Gnathosomal bay (100 - 130 (115) long; 62.5 - 77.5 (75) wide). Cxgl- 4 subapical. Medial suture (85 - 100 (85) long). Genital plates (115 - 130 (125) long; 95 - 105 (100) wide). Additional measurements: Cx- 1 (220 - 270 (250) long (total); 120 - 150 (145) long (medial )); Cx- 3 (275 - 325 (305) wide); anterior venter (220 - 255 (240) long). Ventral proportions: gnathosomal bay length / width 1.53 - 1.79 (1.53); anterior venter / genital field length 1.87 - 1.96 (1.92); anterior venter length / genital field width 2.32 - 2.43 (2.40); anterior venter / medial suture 2.44 - 2.83 (2.82). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
B18D1A316B2968C9B3E6E7DA203477D1.taxon	etymology	Etymology. Specific epithet (kringi) named in honor of Tim Kring, Department Head of Entomology at Virginia Tech, who was a member of JRF's Ph. D. committee. His friendship, humor, advice, and student-focused approach has been a great influence to JRF.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
B18D1A316B2968C9B3E6E7DA203477D1.taxon	distribution	Distribution. Eastern Texas (Figure 117).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
746F1C46554494C6236037EB4300F692.taxon	materials_examined	Material examined (9 ♀; 3 ♂). Arizona, USA: 1 ♀ and 1 ♂ from Cochise County, Chiricahua Mountains; Cave Creek Recreation Area; John Hands Picnic Area off Forest Road 42 A west of Portal, (31 ° 53 ' 53 " N, 109 ° 13 ' 13 " W), 15 July 1987, by IM Smith, IMS 870092 A * 2 ♀ from Cochise County, Chiricahua Mountains; Cave Creek Recreation Area; John Hands Picnic Area off Forest Road 42 A west of Portal, (31 ° 53 ' 53 " N, 109 ° 13 ' 13 " W), 15 July 1987, by IM Smith, IMS 870092 B * 1 ♀ and 1 ♂ from Cochise County, Chiricahua Mountains; beside Forest Road 42 near junction with Forest Road 42 B, (31 ° 55 ' 55 " N, 109 ° 15 ' 15 " W), 16 July 1987, by IM Smith, IMS 870093 A * 2 ♀ and 1 ♂ from Cochise County, Chiricahua Mountains; Cave Creek Recreation Area; Stewart Campground beside Forest Road 42 west of Portal, (31 ° 53 ' 53 " N, 109 ° 10 ' 10 " W), 16 July 1987, by IM Smith, IMS 870094 * 1 ♀ and 1 ♂ from Cochise County, Chiricahua Mountains; Sycamore Campground east of Sunizona, (31 ° 52 ' 52 " N, 109 ° 20 ' 20 " W), 15 July 1987, by IM Smith, IMS 870091 * Mexico, Mexico: 1 ♀ from beside Hwy. 134 at Km 29, (19 ° 59 ' 59 " N, 103 ° 30 ' 30 " W), 7 May 1985, by IM Smith & C Cramer, IMS 850138 * 1 ♀ from beside Hwy. 134 at Km 30, (21 ° 0 ' 0 " N, 101 ° 51 ' 51 " W), 14 May 1985, by IM Smith & C Cramer, IMS 850139	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
746F1C46554494C6236037EB4300F692.taxon	diagnosis	Diagnosis. Torrenticola kurtvietsi are similar to other members of the Rala Group (T. rala, T. keesdavidsi, T. boettgeri, T. lamellipalpis, T. dolichodactyla, and T. anoplopalpa) by being colorless, having incomplete hind coxal margins and being distributed in the southwest. T. kurtvietsi can be differentiated from all other Rala Group by having a more elongate gnathosomal bay (length / width = 4.47 - 5.43 in T. kurtvietsi, 1.48 - 2.73 in others), except T. boettgeri (3.85 - 5.00); and stockier pedipalpal tibiae (length / width = 1.75 - 2.00 in T. kurtvietsi, 2.71 - 5.20 in others), except T. boettgeri (1.86 - 2.14). T. kurtvietsi can be differentiated from T. boettgeri by having a less elongate dorsum (length / width = 1.29 - 1.42 in T. kurtvietsi, 1.74 - 1.82 in T. boettgeri) and a more elongate subcapitulum (ventral length / width = 2.51 - 2.70 in T. kurtvietsi, 1.96 - 2.07 in T. boettgeri).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
746F1C46554494C6236037EB4300F692.taxon	description	Re-description. Female (Figure 121) (n = 5) with characters of the genus with following specifications. Dorsum - (680 - 730 long; 510 - 560 wide) circular with light pink coloration without a distinct pattern or colorless. Anterio-medial platelets (135 - 150 long; 60 - 70 wide). Anterio-lateral platelets (185 - 215 long; 75 - 80 wide) free from dorsal plate. Dgl- 4 closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 360 - 410). Dorsal plate proportions: dorsum length / width 1.29 - 1.34; dorsal width / distance between Dgl- 4 1.34 - 1.44; anterio-medial platelet length / width 2.07 - 2.25; anterio-lateral platelet length / width 2.47 - 2.80; anterio-lateral / anterio-medial length 1.33 - 1.48. Gnathosoma - Subcapitulum (310 - 332.5 long (ventral); 205 - 230 long (dorsal); 115 - 132.5 tall) colorless. Rostrum (105 - 112.5 long; 35 - 35 wide). Chelicerae (325 - 360 long) with curved fangs (45 - 55 long). Subcapitular proportions: ventral length / height 2.51 - 2.70; rostrum length / width 3.00 - 3.21. Pedipalps short and stocky (especially tibiae) without extensions on femora and genua. Palpomeres: trochanter (30 - 35 long); femur 78.75 - 87.5 long); genu (57.5 - 62.5 long); tibia (35 - 40 long; 20 - 20 wide); tarsus (12.5 - 15 long). Palpomere proportions: femur / genu 1.37 - 1.48; tibia / femur 0.41 - 0.47; tibia length / width 1.75 - 2.00. Venter - (840 - 915 long; 610 - 650 wide) colorless. Gnathosomal bay (180 - 205 long; 35 - 42.5 wide). Cxgl- 4 apical. Medial suture (40 - 47.5 long). Genital plates (185 - 195 long; 165 - 177.5 wide). Additional measurements: Cx- 1 (340 - 360 long (total); 150 - 180 long (medial )); Cx- 3 (370 - 390 wide); anterior venter (210 - 225 long). Ventral proportions: gnathosomal bay length / width 4.75 - 5.43; anterior venter / genital field length 1.08 - 1.18; anterior venter length / genital field width 1.18 - 1.33; anterior venter / medial suture 4.53 - 5.25. Male (Figure 122) (n = 4) with characters of the genus with following specifications. Dorsum - (610 - 650 long; 450 - 470 wide) circular with light pink coloration without a distinct pattern or colorless. Anterio-medial platelets (140 - 150 long; 62.5 - 65 wide). Anterio-lateral platelets (195 - 205 long; 70 - 75 wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 345 - 360). Dorsal plate proportions: dorsum length / width 1.31 - 1.42; dorsal width / distance between Dgl- 4 1.30 - 1.35; anterio-medial platelet length / width 2.19 - 2.31; anterio-lateral platelet length / width 2.67 - 2.79; anterio-lateral / anterio-medial length 1.33 - 1.46. Gnathosoma - Subcapitulum (297.5 - 310 long (ventral); 205 - 215 long (dorsal); 112.5 - 115 tall) colorless. Rostrum (100 - 107.5 long; 35 - 35 wide). Chelicerae (325 - 355 long) with curved fangs (50 - 50 long). Subcapitular proportions: ventral length / height 2.64 - 2.70; rostrum length / width 2.86 - 3.07. Pedipalps short and stocky (especially tibiae) without extensions on femora and genua. Palpomeres: trochanter (27.5 - 35 long); femur (75 - 82.5 long); genu (57.5 - 60 long); tibia (36.25 - 38.75 long; 18.75 - 21.25 wide); tarsus (10 - 15 long). Palpomere proportions: femur / genu 1.29 - 1.43; tibia / femur 0.45 - 0.48; tibia length / width 1.81 - 2.00. Venter - (785 - 815 long; 530 - 540 wide) colorless. Gnathosomal bay (185 - 195 long; 40 - 42.5 wide). Cxgl- 4 apical. Medial suture (100 - 120 long). Genital plates (175 - 185 long; 135 - 140 wide). Additional measurements: Cx- 1 (340 - 350 long (total); 150 - 170 long (medial )); Cx- 3 (360 - 370 wide); anterior venter (287.5 - 307.5 long). Ventral proportions: gnathosomal bay length / width 4.47 - 4.81; anterior venter / genital field length 1.58 - 1.71; anterior venter length / genital field width 135 - 140; anterior venter / medial suture 2.40 - 3.00. Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
746F1C46554494C6236037EB4300F692.taxon	etymology	Etymology. Cramer (1992) named this species in honor of Kurt O. Viets " in recognition of his great contribution to the knowledge of aquatic mites and especially for his invaluable efforts in bringing together the world catalog on this subject " (translated from Spanish).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
746F1C46554494C6236037EB4300F692.taxon	distribution	Distribution. Southeastern Arizona (probably also western New Mexico), extending south into Mexico (Figure 120).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
27929BE26BCDF5D5DEF4FB80CD2FA939.taxon	materials_examined	Material examined (5 ♀; 4 ♂). New Mexico, USA: 2 ♀ and 2 ♂ from Catron County, beside Rt. 15, 65 km north of Rt. 180 (Silver City), (33 ° 12 ' 12 " N, 108 ° 13 ' 13 " W), 10 July 1987, by IM Smith, IMS 870081 A * 2 ♀ from Catron County, beside Rt. 15 just below mouth of Little Creek, (33 ° 12 ' 12 " N, 108 ° 13 ' 13 " W), 11 July 1987, by IM Smith, IMS 870083 A * 2 ♂ from Catron County, Glenwood; Whitewater Picnic Area 8 km east of Rt. 180, (33 ° 22 ' 22 " N, 108 ° 50 ' 50 " W), 12 July 1987, by IM Smith, IMS 870084 * 1 ♀ from Grant County, Gila River Recreation Area beside Rt. 15 at Grapevine Recreation Area north of Silver City, (33 ° 11 ' 11 " N, 108 ° 12 ' 12 " W), 11 July 1987, by IM Smith, IMS 870082 B.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
27929BE26BCDF5D5DEF4FB80CD2FA939.taxon	diagnosis	Diagnosis. Torrenticola lamellipalpis are similar to other members of the Rala Group (T. rala, T. keesdavidsi, T. boettgeri, T. kurtvietsi, T. dolichodactyla, and T. anoplopalpa) by being colorless, having incomplete hind coxal margins and being distributed in the southwest. T. lamellipalpis can be differentiated from all other Rala Group by having a wide, prominent lamellate extension on the pedipalpal femora that extend to the genua (others either without extensions or with tuberculate or flanged, dentate extensions).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
27929BE26BCDF5D5DEF4FB80CD2FA939.taxon	description	Re-description. Female (Figure 124) (n = 5) with characters of the genus with following specifications. Dorsum - (660 - 735 long; 425 - 490 wide) ovoid and colorless. Anterio-medial platelets (162.5 - 175 long; 55 - 57.5 wide). Anterio-lateral platelets (205 - 225 long; 65 - 75 wide) free from dorsal plate. Dgl- 4 closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 355 - 370). Dorsal plate proportions: dorsum length / width 1.46 - 1.60; dorsal width / distance between Dgl- 4 1.20 - 1.32; anterio-medial platelet length / width 2.87 - 3.05; anterio-lateral platelet length / width 2.80 - 3.35; anterio-lateral / anterio-medial length 1.17 - 1.36. Gnathosoma - Subcapitulum (440 - 465 long (ventral); 330 - 350 long (dorsal); 125 - 135 tall) colorless. Rostrum (175 - 187.5 long; 55 - 60 wide). Chelicerae (390 - 420 long) with curved fangs (60 - 65 long). Subcapitular proportions: ventral length / height 3.38 - 3.52; rostrum length / width 3.00 - 3.41. Pedipalps with wide, prominent lamellate extensions on femora that extends halfway into the genua. Palpomeres: trochanter (55 - 60 long); femur (145 - 152.5 long); genu (75 - 77.5 long); tibia (97.5 - 102.5 long; 22.5 - 23.75 wide); tarsus (17.5 - 20 long). Palpomere proportions: femur / genu 1.90 - 1.97; tibia / femur 0.66 - 0.69; tibia length / width 4.32 - 4.56. Venter - (815 - 930 long; 500 - 560 wide) colorless. Gnathosomal bay (180 - 195 long; 80 - 100 wide). Cxgl- 4 subapical. Medial suture (35 - 45 long). Genital plates (185 - 195 long; 155 - 165 wide). Additional measurements: Cx- 1 (380 - 390 long (total); 190 - 200 long (medial )); Cx- 3 (350 - 365 wide); anterior venter (240 - 260 long). Ventral proportions: gnathosomal bay length / width 1.90 - 2.31; anterior venter / genital field length 1.23 - 1.38; anterior venter length / genital field width 1.48 - 1.65; anterior venter / medial suture 5.67 - 6.86. Male (Figure 125) (n = 4) with characters of the genus with following specifications. Dorsum - (620 - 680 long; 400 - 460 wide) ovoid and colorless. Anterio-medial platelets (147.5 - 165 long; 52.5 - 57.5 wide). Anterio-lateral platelets (195 - 217.5 long; 70 - 77.5 wide) free from dorsal plate. Dgl- 4 closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 320 - 360). Dorsal plate proportions: dorsum length / width 1.48 - 1.58; dorsal width / distance between Dgl- 4 1.25 - 1.28; anterio-medial platelet length / width 2.77 - 3.14; anterio-lateral platelet length / width 2.69 - 3.04; anterio-lateral / anterio-medial length 1.28 - 1.32. Gnathosoma - Subcapitulum (392.5 - 427.5 long (ventral); 292.5 - 325 long (dorsal); 112.5 - 127.5 tall) colorless. Rostrum (155 - 170 long; 50 - 55 wide). Chelicerae (350 - 390 long) with curved fangs (55 - 55 long). Subcapitular proportions: ventral length / height 3.29 - 3.56; rostrum length / width 3.09 - 3.24. Pedipalps with wide, prominent lamellate extensions on femora that extends halfway into the genua. Palpomeres: trochanter (51.25 - 55 long); femur (130 - 135 long); genu (70 - 77.5 long); tibia (97.5 - 105 long; 20 - 22.5 wide); tarsus (17.5 - 20 long). Palpomere proportions: femur / genu 1.74 - 1.86; tibia / femur 0.74 - 0.79; tibia length / width 4.67 - 4.94. Venter - (780 - 840 long; 450 - 520 wide) colorless. Gnathosomal bay (170 - 195 long; 80 - 90 wide). Cxgl- 4 subapical. Medial suture (110 - 130 long). Genital plates (160 - 180 long; 107.5 - 127.5 wide). Additional measurements: Cx- 1 (335 - 370 long (total); 170 - 185 long (medial )); Cx- 3 (325 - 375 wide); anterior venter (295 - 325 long). Ventral proportions: gnathosomal bay length / width 2.00 - 2.29; anterior venter / genital field length 1.75 - 1.86; anterior venter length / genital field width 2.47 - 2.83; anterior venter / medial suture 2.42 - 2.83. Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
27929BE26BCDF5D5DEF4FB80CD2FA939.taxon	etymology	Etymology. Viets (1977 b) named the specific epithet (lamellipalpis) in reference to the " extensive ventral lamella " on the pedipalp femora (lamella, L. small plate or flake; palpus, L. hand, feeler).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
27929BE26BCDF5D5DEF4FB80CD2FA939.taxon	distribution	Distribution. Southwestern New Mexico (probably also found in southeastern Arizona), extending southward into Guatemala (Figure 123).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
FB27CB3A82F88DB8ACFA9B608733BEDB.taxon	materials_examined	Material examined. HOLOTYPE (♀): USA, Arkansas, Polk Co., Bard Springs, Ouachita National Forest, Blaylock Creek (34 ° 23 ' 28.3 " N, 94 ° 00 ' 31.8 " W), 11 Aug 2009, by AJ Radwell and BG Crump, AJR 090307 B. PARATYPES (5 ♀; 8 ♂): Arkansas, USA: 1 ♂ (ALLOTYPE) from Polk County, Bard Springs, Ouachita National Forest, Blaylock Creek (34 ° 23 ' 28.3 " N, 94 ° 00 ' 31.8 " W), 11 Aug 2009, by AJ Radwell and BG Crump, AJR 090307 B * 3 ♀ from Polk County, beside Forest Road 38, North of Shady Lake Rec Area, East Saline Creek (34 ° 22 ' 53.4 " N, 94 ° 01 ' 51.2 " W), 30 Jul 2011, by IM Smith, IMS 110041 * 1 ♀ and 6 ♂ from Montgomery County, Ouachita National Forest, Ouachita River at Mcguire (34 ° 22 ' 53.4 " N, 94 ° 1 ' 51.2 " W), 27 Aug 2011, by AJ Radwell, AJR 110307 * 1 ♀ and 1 ♂ from Garland County, beside Route 7, 3 miles south of Mountain Valley, South Fork of Saline River (34 ° 35 ' 43.3 " N, 93 ° 00 ' 45.3 " W), 11 May 1977, by DR Cook, DRC 770002 * 1 ♂ from Montgomery County, Ouachita National Forest, Ouachita River at Pine Ridge (34 ° 34 ' 53.5 " N, 93 ° 53 ' 00.9 " W), 5 Oct 2007, by AJ Radwell and HW Robison, AJR 070300 A.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
FB27CB3A82F88DB8ACFA9B608733BEDB.taxon	diagnosis	Diagnosis. Torrenticola larvata are similar to other members of the Tricolor Complex (T. bittikoferae, T. hoosieri, T. pearsoni, T. olliei, T. sierrensis, T. tricolor, T. trimaculata, T. unimaculata, T. cardia, T. kringi, T. dimorpha, and T. mohawk) in having a short, conical rostrum. T. larvata can be differentiated from all other Torrenticola, including other members of the Tricolor Complex, by having a distinct dorsal pattern. T. larvata can be further differentiated from T. bittikoferae, T. sierrensis, T. tricolor, T. trimaculata, T. kringi, and T. mohawk by being more elongate (length / width = 1.41 - 1.57 in T. larvata; 1.17 - 1.39 in others); and from T. bittikoferae, T. olliei, T. sierrensis, T. trimaculata, T. unimaculata, T. dimorpha, and T. mohawk by having a more elongate rostrum (length / width = 2.32 - 2.53 in T. larvata, 1.56 - 2.27 in others).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
FB27CB3A82F88DB8ACFA9B608733BEDB.taxon	description	Re-description (amended from Cherri et al. 2016). Female (Figure 127) (n = 6) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (650 - 725 (650) long; 450 - 475 (460) wide) ovoid and elongate with bluish-purple coloration restricted to the anterio-medial platelets and anterior-most portion of anterio-lateral platelets (rarely continuing to anterior border of the dorsal plate) and to the posterior dorsal plate within the area of primary sclerotization (posterior coloration is sometimes absent) with wide strip of orange medially. Anterio-medial platelets (127.5 - 132.5 (127.5) long; 175 - 190 (175) wide). Anterio-lateral platelets (175 - 190 (175) long; 70 - 77.5 (70) wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 340 - 375 (340 )). Dorsal plate proportions: dorsum length / width 1.41 - 1.54 (1.41); dorsal width / distance between Dgl- 4 1.27 - 1.35 (1.35); anterio-medial platelet length / width 1.83 - 2.04 (2.04); anterio-lateral platelet length / width 2.29 - 2.56 (2.50); anterio-lateral / anterio-medial length 1.37 - 1.43 (1.37). Gnathosoma - Subcapitulum (275 - 287.5 (275) long (ventral); 200 - 211 (200) long (dorsal); 125 - 140 (125) tall) with bluish-purple coloration. Rostrum (110 - 117.5 (112.5) long; 47.5 - 50 (47.5) wide) short and conical. Chelicerae (260 - 295 (265) long) with curved fangs (53 - 62 (60) long). Subcapitular proportions: ventral length / height 2.04 - 2.25 (2.20); rostrum length / width 2.32 - 2.42 (2.37). Pedipalps with tuberculate ventral extensions with dentate tips on femora and genua. Palpomeres: trochanter (40 - 42.5 (40) long); femur (105 - 107.5 (107.5) long); genu (67.5 - 72.5 (67.5) long); tibia (87.5 - 97.5 (90) long; 25 - 30 (26.25) wide); tarsus (25 - 27.5 (25) long). Palpomere proportions: femur / genu 1.48 - 1.59 (1.59); tibia / femur 0.81 - 0.91 (0.84); tibia length / width 3.25 - 3.50 (3.43). Venter - (720 - 850 (795) long; 525 - 604 (525) wide) colorless or with bluish-purple coloration, but always with bold bluish-purple on the dorsal coxal area. Gnathosomal bay (117.5 - 140 (140) long; 77.5 - 92.5 (77.5) wide). Cxgl- 4 subapical. Medial suture (25 - 35 (35) long). Genital plates (182.5 - 187.5 (185) long; 145 - 152.5 (150) wide). Additional measurements: Cx- 1 (260 - 358.5 (260) long (total); 125 - 183 (125) long (medial )); Cx- 3 (307 - 375 (325) wide); anterior venter (180 - 195 (180) long). Ventral proportions: gnathosomal bay length / width 1.31 - 1.81 (1.81); anterior venter / genital field length 0.97 - 1.04 (0.97); anterior venter length / genital field width 1.20 - 1.31 (1.20); anterior venter / medial suture 5.14 - 7.80 (5.14). Male (Figure 128) (n = 6) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (550 - 610 (560) long; 350 - 400 (360) wide) ovoid and elongate with bluish-purple coloration restricted to the anterio-medial platelets and anterior-most portion of anterio-lateral platelets (rarely continuing to anterior border of the dorsal plate) and to the posterior dorsal plate within the area of primary sclerotization (posterior coloration is sometimes absent) with wide strip of orange medially. Anterio-medial platelets (115 - 120 (120) long; 52.5 - 65 (55) wide). Anterio-lateral platelets (147.5 - 162.5 (155) long; 57.5 - 70 (70) wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 285 - 340 (295 )). Dorsal plate proportions: dorsum length / width 1.53 - 1.57 (1.56); dorsal width / distance between Dgl- 4 1.18 - 1.26 (1.22); anterio-medial platelet length / width 1.85 - 2.24 (2.18); anterio-lateral platelet length / width 2.21 - 2.83 (2.21); anterio-lateral / anterio-medial length 1.26 - 1.41 (1.29). Gnathosoma - Subcapitulum (230 - 247.5 (240) long (ventral); 167 - 182 (175) long (dorsal); 97.5 - 102.5 (100) tall) with bluish-purple coloration. Rostrum (91.25 - 100 (93.75) long; 37.5 - 40 (40) wide) short and conical. Chelicerae (221 - 239 (225) long) with curved fangs (45 - 52 (50) long). Subcapitular proportions: ventral length / height 2.30 - 2.46 (2.40); rostrum length / width 2.34 - 2.53 (2.34). Pedipalps with tuberculate ventral extensions with dentate tips on femora and genua. Palpomeres: trochanter (32.5 - 37.5 (35) long); femur (85 - 95 (90) long); genu (56.25 - 62.5 (60) long); tibia (75 - 82.5 (80) long; 23.75 - 26.25 (25) wide); tarsus (22.5 - 26.25 (25) long). Palpomere proportions: femur / genu 1.48 - 1.61 (1.50); tibia / femur 0.82 - 0.89 (0.89); tibia length / width 3.10 - 3.20 (3.20). Venter - (660 - 710 (680) long; 415 - 443 (420) wide) colorless or with bluish-purple coloration, but always with bold bluish-purple on the dorsal coxal area. Gnathosomal bay (112.5 - 130 (125) long; 65 - 70 (67.5) wide). Cxgl- 4 subapical. Medial suture (102.5 - 125 (110) long). Genital plates (138.75 - 147.5 (140) long; 97.5 - 110 (110) wide). Additional measurements: Cx- 1 (244 - 266 (250) long (total); 135 - 140 (140) long (medial )); Cx- 3 (283 - 305 (295) wide); anterior venter (245 - 270 (260) long). Ventral proportions: gnathosomal bay length / width 1.67 - 1.94 (1.85); anterior venter / genital field length 1.77 - 1.89 (1.86); anterior venter length / genital field width 2.36 - 2.77 (2.36); anterior venter / medial suture 2.16 - 2.41 (2.36). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
FB27CB3A82F88DB8ACFA9B608733BEDB.taxon	etymology	Etymology. Cherri et al. (2016) named the specific epithet (larvata) for the anterior coloration that gives adults a masked appearance (larvat, L. masked).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
FB27CB3A82F88DB8ACFA9B608733BEDB.taxon	distribution	Distribution. Ouachita Mountains of Arkansas (Figure 126).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
A54A46FE4C869928B1D2E950FAA49A77.taxon	materials_examined	Material examined. HOLOTYPE (♀): from USA, California, Mendocino County, beside Rt. 128 at Paul M. Dimmick Recreation Area, (39 ° 10 ' 10 " N, 123 ° 38 ' 38 " W), 4 August 1987, by IM Smith, IMS 870127 A PARATYPES (3 ♀; 1 ♂): California, USA: 1 ♂ (ALLOTYPE) from Mendocino, beside Rt. 128 at Paul M. Dimmick Recreation Area, (39 ° 10 ' 10 " N, 123 ° 38 ' 38 " W), 4 August 1987, by IM Smith, IMS 870127 A * 3 ♀ from Mendocino County, beside Rt. 128 at Paul M. Dimmick Recreation Area, (39 ° 10 ' 10 " N, 123 ° 38 ' 38 " W), 4 August 1987, by IM Smith, IMS 870127 A	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
A54A46FE4C869928B1D2E950FAA49A77.taxon	diagnosis	Diagnosis. Torrenticola leviathan are similar to other members of the Ellipsoidalis Group (T. multiforma, T. occidentalis, and T. ellipsoidalis), in being among the largest Torrenticola in the west (dorsum length ♀ = 700 - 885; ♂ = 665 - 850), although T. sierrensis are also large (dorsum length ♀ = 700 - 880; ♂ = 590 - 735) but can easily be distinguished from the Ellipsoidalis Group by being circular instead of ellipsoid or rectangular (dorsum length / width = 1.17 - 1.28 in T. sierrensis, 1.30 - 1.67 in Ellipsoidalis Group). T. leviathan can be differentiated from all other Ellipsoidalis Group by having more elongate pedipalpal tibiae (length / width = 3.43 - 4.20 in T. leviathan, 2.64 - 3.33 in others).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
A54A46FE4C869928B1D2E950FAA49A77.taxon	description	Description. Female (Figure 130) (n = 4) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (845 - 870 (870) long; 570 - 610 (580) wide) ellipsoid with faint orange coloration without a distinct pattern or colorless. Anterio-medial platelets (150 - 155 (155) long; 70 - 80 (75) wide). Anterio-lateral platelets (240 - 260 (245) long; 95 - 105 (95) wide) free from dorsal plate. Dgl- 4 closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 420 - 460 (425 )). Dorsal plate proportions: dorsum length / width 1.40 - 1.51 (1.50); dorsal width / distance between Dgl- 4 1.33 - 1.36 (1.36); anterio-medial platelet length / width 1.94 - 2.14 (2.07); anterio-lateral platelet length / width 2.40 - 2.58 (2.58); anterio-lateral / anterio-medial length 1.55 - 1.70 (1.58). Gnathosoma - Subcapitulum (330 - 350 (330) long (ventral); 230 - 230 (230) long (dorsal); 165 - 170 (170) tall) colorless. Rostrum (125 - 130 (125) long; 55 - 60 (55) wide) short and conical. Chelicerae (335 - 340 (340) long) with curved fangs (80 - 85 (80) long). Subcapitular proportions: ventral length / height 2.00 - 2.06 (2.00); rostrum length / width 2.17 - 2.27 (2.27). Pedipalps with short, tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (50 - 52.5 (52.5) long); femur (130 - 130 (130) long); genu (75 - 77.5 (77.5) long); tibia (105 - 107.5 (107.5) long; 25 - 27.5 (27.5) wide); tarsus (20 - 22.5 (22.5) long). Palpomere proportions: femur / genu 1.68 - 1.73 (1.68); tibia / femur 0.81 - 0.83 (0.83); tibia length / width 3.91 - 4.20 (3.91). Venter - (970 - 1035 (1035) long; 640 - 680 (640) wide) colorless. Gnathosomal bay (190 - 215 (190) long; 85 - 100 (100) wide). Cxgl- 4 subapical. Medial suture (25 - 30 (30) long). Genital plates (210 - 215 (210) long; 180 - 195 (180) wide). Additional measurements: Cx- 1 (390 - 400 (390) long (total); 180 - 205 (205) long (medial )); Cx- 3 (440 - 460 (440) wide); anterior venter (220 - 245 (245) long). Ventral proportions: gnathosomal bay length / width 1.90 - 2.53 (1.90); anterior venter / genital field length 1.05 - 1.17 (1.17); anterior venter length / genital field width 1.13 - 1.36 (1.36); anterior venter / medial suture 7.50 - 8.80 (8.17). Male (Figure 131) (n = 1) (allotype only) with characters of the genus with following specifications. Dorsum - (770 long; 510 wide) ellipsoid with faint orange coloration without a distinct pattern or colorless. Anterio-medial platelets (140 long; 65 wide). Anterio-lateral platelets (230 long; 85 wide) free from dorsal plate. Dgl- 4 closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 400). Dorsal plate proportions: dorsum length / width 1.51; dorsal width / distance between Dgl- 4 1.28; anterio-medial platelet length / width 2.15; anterio-lateral platelet length / width 2.71; anterio-lateral / anterio-medial length 1.64. Gnathosoma - Subcapitulum (285 long (ventral); 200 long (dorsal); 137.5 tall) colorless. Rostrum (95 long; 52.5 wide) short and conical. Chelicerae (275 long) with curved fangs (75 long). Subcapitular proportions: ventral length / height 2.07; rostrum length / width 1.81. Pedipalps with short, tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (47.5 long); femur (112.5 long); genu (70 long); tibia (90 long; 26.25 wide); tarsus (20 long). Palpomere proportions: femur / genu 1.61; tibia / femur 0.80; tibia length / width 3.43. Venter - (860 long; 570 wide) colorless. Gnathosomal bay (180 long; 90 wide). Cxgl- 4 subapical. Medial suture (110 long). Genital plates (160 long; 120 wide). Additional measurements: Cx- 1 (360 long (total); 185 long (medial )); Cx- 3 (420 wide); anterior venter (305 long). Ventral proportions: gnathosomal bay length / width 2.00; anterior venter / genital field length 1.91; anterior venter length / genital field width 2.54; anterior venter / medial suture 2.77. Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
A54A46FE4C869928B1D2E950FAA49A77.taxon	etymology	Etymology. Specific epithet (leviathan) refers to the large body size of this species, surpassed by few other Torrenticola (liwyatan, Hebrew, a large aquatic animal; noun in apposition). The Leviathan (modern spelling) is depicted in Jewish mythology (as written in the Tanakh) as one of two huge beasts-the Leviathan in the ocean and the Behemoth on land.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
A54A46FE4C869928B1D2E950FAA49A77.taxon	distribution	Distribution. Northwestern California (Figure 129).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
2B6ECB1D337DAEA27EBA9F1FBE67AD0C.taxon	materials_examined	Material examined. HOLOTYPE (♂): from USA, Tennessee, Monroe County, beside Forest Route 35 (35 ° 21 ' 47 " N, 84 ° 9 ' 47 " W), 12 Sep 2009, by IM Smith, IMS 090112, DNA 2979. PARATYPES (0 ♀; 1 ♂): Tennessee, USA: 1 ♂ from USA, Tennessee, Monroe County, beside Forest Route 35 (35 ° 21 ' 47 " N, 84 ° 9 ' 47 " W), 12 Sep 2009, by IM Smith, IMS 090112.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
2B6ECB1D337DAEA27EBA9F1FBE67AD0C.taxon	diagnosis	Diagnosis. Torrenticola longitibia are similar to other members of the Raptor Group (T. gnoma, T. irapalpa, T. mjolniri, T. elusiva, T. racupalpa, T. raptor, T. danielleae, T. daemon, and T. ivyae) in having round bodies; Dgl- 4 close to muscles scars; long, thin subcapitular rostra; and long, thin pedipalp tibiae. T. longitibia can be differentiated from all other members of the Raptor Group (both males and females) by having a longer femur with respect to the genu (femur / genu = 2.1 - 2.17 in T. longitibia, 1.66 - 2.00 in others).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
2B6ECB1D337DAEA27EBA9F1FBE67AD0C.taxon	description	Description. Female unknown. Male (Figure 133) (n = 2) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum- (530 - 560 (530) long; 420 - 430 (420) wide) circular with navy blue coloration posteriorly extending in a thin strip anteriorly to the edge of the dorsal plate. Anterio-medial platelets (122.5 - 132.5 (122.5) long; 62.5 - 72.5 (62.5) wide). Anterio-lateral platelets (172.5 - 180 (172.5) long; 78.75 - 80 (80) wide) free from dorsal plate. Dgl- 4 much closer to the muscle scars than to the dorsum edge (distance between Dgl- 4 155 - 175 (155 )). Dorsal plate proportions: dorsum length / width 1.26 - 1.30 (1.26); dorsal width / distance between Dgl- 4 2.46 - 2.71 (2.71); anterio-medial platelet length / width 1.83 - 1.96 (1.96); anterio-lateral platelet length / width 2.16 - 2.29 (2.16); anterio-lateral / anterio-medial length 1.36 - 1.41 (1.41). Gnathosoma - Subcapitulum (300 - 315 (300) long (ventral); 235 - 245 (235) long (dorsal); 110 - 117.5 (110) tall) colorless. Rostrum (135 - 137.5 (135) long; 32.5 - 32.5 (32.5) wide) elongate. Chelicerae (325 - 325 (325) long) with curved fangs (50 - 50 (50) long). Subcapitular proportions: ventral length / height 2.68 - 2.73 (2.73); rostrum length / width 4.15 - 4.23 (4.15). Pedipalps elongate (especially tibiae) with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (42.5 - 45 (42.5) long); femur (125 - 131.25 (125) long); genu (57.5 - 62.5 (57.5) long); tibia (110 - 110 (110) long; 20 - 20 (20) wide); tarsus (15 - 15 (15) long). Palpomere proportions: femur / genu 2.10 - 2.17 (2.17); tibia / femur 0.84 - 0.88 (0.88); tibia length / width 5.50 - 5.50 (5.50). Venter - (660 - 710 (660) long; 500 - 550 (500) wide) colorless. Gnathosomal bay (150 - 155 (150) long; 75 - 85 (85) wide). Cxgl- 4 far from apex. Medial suture (60 - 60 (60) long). Genital plates (137.5 - 150 (137.5) long; 115 - 122.5 (115) wide). Additional measurements: Cx- 1 (270 - 300 (270) long (total); 130 - 140 (130) long (medial )); Cx- 3 (320 - 325 (320) wide); anterior venter (205 - 220 (205) long). Ventral proportions: gnathosomal bay length / width 1.76 - 2.07 (1.76); anterior venter / genital field length 1.47 - 1.49 (1.49); anterior venter length / genital field width 1.78 - 1.80 (1.78); anterior venter / medial suture 3.42 - 3.67 (3.42). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
2B6ECB1D337DAEA27EBA9F1FBE67AD0C.taxon	etymology	Etymology. Specific epithet (longitibia) refers to the long, thin pedipalpal tibiae (longus, L. long; tibia, L. tibia).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
2B6ECB1D337DAEA27EBA9F1FBE67AD0C.taxon	distribution	Distribution. Known only from Monroe County, Tennessee (Figure 132).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
2BD62332D7E62961DD10D4764B0305D5.taxon	materials_examined	Material examined. HOLOTYPE (♀): from Canada, New Brunswick, Victoria County, Salmon River, 21 June 1953, by Habeeb, HH 530075. PARATYPES (0 ♀; 1 ♂): New Brunswick, Canada: 1 ♂ (ALLOTYPE) from Victoria County, Salmon River, 21 June 1953, by Habeeb, HH 530075. OTHER MATERIAL (29 ♀; 14 ♂): Maine, USA: 2 ♀ from Franklin County, Smalls Falls Picnic Area, beside Route 4, Sandy River (44 ° 52 ' N, 70 ° 31 ' W), 5 Jul 1989, by IM Smith, IMS 890069 * Missouri, USA: 1 ♀ from McDonald County, Tiff City, beside Route 43, Buffalo Creek (36 ° 40 ' 17 " N, 94 ° 36 ' 17 " W), 2 May 1996, by IM Smith, IMS 960004 * Nova Scotia, Canada: 1 ♀ from Inverness County, Cape Brenton Highland National Park, 10 Sep 2011, by IM Smith, IMS 110072 * 2 ♀ from Inverness County, Inervess, Cheticamp River, 10 Sep 2011, by IM Smith, IMS 110071 * New Brunswick, Canada: 1 ♀ and 1 ♂ from Victoria County, St. Quentin, beside Highway 17, 14 km southwest of Highway 180, 14 Sep 2011, by IM Smith, IMS 110059 * 2 ♀ from York County, Napadogan, beside Road J- 19, 4 km south of Hwy 107, 6 Oct 2011, by IM Smith, IMS 110126 * Newfoundland, Canada: 2 ♀ from Crooked Feeder, beside Highway 1, north of Deer Lake (49 ° 10 ' N, 57 ° 26 ' W), 25 Jul 1977, by IM Smith, IMS 770125 B * 1 ♀ and 1 ♂ from Deer Lake, beside Highway 430, 6.2 kilometers north of Highway 1 (49 ° 10 ' N, 57 ° 26 ' W), 23 Jul 1977, by IM Smith, IMS 770112 * 2 ♀ and 2 ♂ from Molly Chignic Brook, beside Highway 1 (47 ° 51 ' N, 59 ° 10 ' W), 7 Oct 1989, by IM Smith, IMS 890133 * Ohio, USA: 2 ♂ from Montgomery County, Engelwood Metro Park (39 ° 52 ' 58 " N, 84 ° 17 ' 33 " W), 31 Jul 2014, by MJ Skvarla, MS 14 - 0731 - 002 * Quebec, Canada: 1 ♀ from Gatineau County, Gatineau Park, Meech Lake (45 ° 32 ' 27 " N, 75 ° 54 ' 53 " W), 27 Jul 2010, by IM Smith, IMS 100105 * 2 ♀ and 2 ♂ from Gatineau County, Gatineau Park, tributary above Blanchet Beach, 6 Apr 2012, by IM Smith, IMS 120001 * 1 ♀ and 1 ♂ from Gatineau County, Gatineau Park, beside Wolf Trail above Blanchet (45 ° 32 ' 30 " N, 75 ° 55 ' 29 " W), 12 Jun 2008, by IM Smith, IMS 080009 A * 1 ♂ from Matane County, St. - Bernard-des-Lacs, Riviere Ste. Anne (48 ° 55 ' 45 " N, 66 ° 7 ' 0 " W), 25 Aug 1975, by BP Smith, DW Barr, & N Avruch, BPS 750562 * 2 ♀ and 1 ♂ from Matapedia County, beside secondary road, Riviere Matapedia (48 ° 17 ' 40 " N, 67 ° 15 ' 20 " W), 24 Aug 1975, by BP smith, DW Barr, & N Avruch, BPS 750557 * 1 ♀ from Stanstead County, Tompkin Stream, Tomifobia River (45 ° 0 ' 31 " N, 72 ° 7 ' 6 " W), 20 Aug 1996, by IM Smith & M MacKenzie, IMS 960056 * Tennessee, USA: 1 ♀ from Blount County, Great Smokey Mountains National Park, Abrams River (35 ° 35 ' 31 " N, 83 ° 51 ' 21 " W), 17 Sep 2010, by IM Smith, IMS 100141 * 1 ♀ from Sevier County, Great Smokey Mountains National Park (35 ° 46 ' 54 " N, 83 ° 13 ' 2 " W), 16 Sep 2010, by IM Smith, IMS 100140 * 1 ♀ from Sevier County, Great Smokey Mountains National Park, middle prong Little Pigeon River (35 ° 42 ' 38 " N, 83 ° 22 ' 59 " W), 10 Sep 2009, by IM Smith, IMS 090106 * Vermont, USA: 2 ♀ and 1 ♂ from Addison County, beside road from Lincoln to Ripton, Middlebury River (44 ° 0 ' N, 73 ° 1 ' W), 6 Jul 1989, by IM Smith, IMS 890075 * 2 ♀ from Addison County, Lincoln, beside US Forest Service Road # 54, New Haven River (44 ° 6 ' N, 72 ° 59 ' W), 6 Jul 1989, by IM Smith, IMS 890074 * 2 ♀ and 2 ♂ from Lamoille County, Stowe, beside Route 108, West Branch of Waterbury River (44 ° 30 ' N, 72 ° 46 ' W), 6 Jul 1989, by IM Smith, IMS 890072.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
2BD62332D7E62961DD10D4764B0305D5.taxon	diagnosis	Diagnosis. Torrenticola magnexa are similar to other members of the Partial 2 - Plate Group (T. folkertsae, T. pulchra, and T. priapus) in having anterio-lateral platelets partially fused to the dorsal plate and being distributed in the east. T. magnexa can be differentiated from other Partial 2 - Plate Group by dorsal coloration and pattern. T. magnexa can be further differentiated from T. New 23 A by having a less elongate rostrum (length / width = 2.25 - 3.00 in T. magnexa, 3.17 - 3.39 in T. priapus). T. magnexa can be further differentiated from T. folkertsae by having less elongate pedipalpal tibiae (length / width = 3.21 - 4.00 in T. magnexa, 4.05 - 4.83 in T. folkertsae). Male T. magnexa can be further differentiated from male T. pulchra by having a larger genital field (length ♂ = 125 - 148 in T. magnexa, 110 - 123 in T. pulchra; width ♂ = 115 - 125 in T. magnexa, 87 - 95 in T. pulchra) and more elongate pedipalpal tibiae (length / width ♂ = 3.78 - 4.00 in T. magnexa, 3.00 - 3.35 in T. pulchra). Female T. magnexa can be differentiated from female T. pulchra by having a wider genital field (♀ 170 - 188 in T. magnexa, 147 - 160 in T. pulchra) and longer pedipalpal tibiae (♀ 102 - 113 in T. magnexa, 82 - 93 in T. pulchra).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
2BD62332D7E62961DD10D4764B0305D5.taxon	description	Re-description. Female (Figure 135) (n = 6) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (680 - 750 (750) long; 500 - 560 (560) wide) ovoid with bluish-purple or reddish-purple coloration separated into anterior and posterior portions with red medially. Anterio-medial platelets 155 - 170 (162.5) long; 57.5 - 65 (60) wide). Anterio-lateral platelets (192.5 - 210 (210) long; 72.5 - 85 (85) wide) partially fused to dorsal plate (especially posteriorly). Dgl- 4 close to the edge of the dorsum (distance between Dgl- 4 390 - 410 (400 )). Dorsal plate proportions: dorsum length / width 1.28 - 1.42 (1.34); dorsal width / distance between Dgl- 4 1.28 - 1.40 (1.40); anterio-medial platelet length / width 2.42 - 2.72 (2.71); anterio-lateral platelet length / width 2.47 - 2.71 (2.47); anterio-lateral / anterio-medial length 1.13 - 1.33 (1.29). Gnathosoma - Subcapitulum (330 - 355 long (ventral); 251 - 277 long (dorsal); 150 - 170 tall) mostly colorless. Rostrum (137.5 - 150 long; 47.5 - 52.5 wide). Chelicerae (335 - 363 long) with curved fangs (55 - 78 long). Subcapitular proportions: ventral length / height 2.00 - 2.37; rostrum length / width 2.75 - 3.00. Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (47.5 - 52.5 (50) long); femur (127.5 - 135 (135) long); genu (75 - 82.5 (80) long); tibia 102.5 - 107.5 (102.5) long; 26.25 - 30 (28.75) wide); tarsus (17.5 - 25 (22.5) long). Palpomere proportions: femur / genu 1.58 - 1.71 (1.69); tibia / femur 0.76 - 0.83 (0.76); tibia length / width 3.57 - 4.00 (3.57). Venter - (770 - 900 (830) long; 588 - 672 (660) wide) colorless. Gnathosomal bay (165 - 190 (165) long; 95 - 125 (125) wide). Cxgl- 4 subapical. Medial suture (0 - 15 (7.5) long) occasionally absent. Genital plates (190 - 207.5 (207.5) long; 170 - 187.5 (187.5) wide). Additional measurements: Cx- 1 (299 - 359 (320) long (total); 96 - 155 (155) long (medial )); Cx- 3 (369 - 455 (455) wide); anterior venter (167.5 - 175 (175) long). Ventral proportions: gnathosomal bay length / width 1.32 - 1.87 (1.32); anterior venter / genital field length 0.84 - 0.91 (0.84); anterior venter length / genital field width 0.93 - 1.03 (0.93); anterior venter / medial suture (proportion cannot be calculated for specimens without a medial suture) 11.33 - 23.33 (23.33). Male (Figure 136) (n = 6) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (540 - 630 (600) long; 390 - 450 (430) wide) ovoid with bluish-purple or reddish-purple coloration separated into anterior and posterior portions with red medially. Anterio-medial platelets (125 - 140 (132.5) long; 47.5 - 52.5 (50) wide). Anterio-lateral platelets (158.75 - 192.5 (170) long; 60 - 70 (60) wide) partially fused to dorsal plate (especially posteriorly). Dgl- 4 close to the edge of the dorsum (distance between Dgl- 4 310 - 350 (310 )). Dorsal plate proportions: dorsum length / width 1.35 - 1.43 (1.40); dorsal width / distance between Dgl- 4 1.26 - 1.39 (1.39); anterio-medial platelet length / width 2.48 - 2.95 (2.65); anterio-lateral platelet length / width 2.44 - 2.85 (2.83); anterio-lateral / anterio-medial length 1.22 - 1.43 (1.28). Gnathosoma - Subcapitulum (272.5 - 300 (285) long (ventral); 200 - 229 (215) long (dorsal); 113 - 127.5 (122.5) tall) mostly colorless. Rostrum (110 - 122.5 (117.5) long; 40 - 45 (42.5) wide). Chelicerae (260 - 292 (290) long) with curved fangs (49 - 60 (55) long). Subcapitular proportions: ventral length / height 2.22 - 2.42 (2.33); rostrum length / width 2.59 - 2.84 (2.76). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (37.5 - 45 (40) long); femur (105 - 112.5 (110) long); genu (65 - 72.5 (67.5) long); tibia (85 - 97.5 (90) long; 22.5 - 25 (22.5) wide); tarsus (17.5 - 20 (20) long). Palpomere proportions: femur / genu 1.55 - 1.67 (1.63); tibia / femur 0.80 - 0.87 (0.82); tibia length / width 3.78 - 4.00 (4.00). Venter - (640 - 770 (710) long; 465 - 550 (550) wide) colorless. Gnathosomal bay (127.5 - 160 (127.5) long; 77.5 - 95 (77.5) wide). Cxgl- 4 subapical. Medial suture (62.5 - 85 (75) long). Genital plates (125 - 147.5 (140) long; 115 - 125 (115) wide). Additional measurements: Cx- 1 (251 - 285 (270) long (total); 102 - 140 (140) long (medial )); Cx- 3 (335 - 375 (375) wide); anterior venter (207.5 - 240 (220) long). Ventral proportions: gnathosomal bay length / width 1.37 - 1.94 (1.65); anterior venter / genital field length 1.57 - 1.78 (1.57); anterior venter length / genital field width 1.80 - 1.98 (1.91); anterior venter / medial suture 2.82 - 3.68 (2.93). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
2BD62332D7E62961DD10D4764B0305D5.taxon	etymology	Etymology. Habeeb (1957) did not specify an etymology for the specific epithet (magnexa). However, surely this name refers to the similarity of this species to the Palaearctic T. connexa (Koenike, 1908) (neos, G. new).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
2BD62332D7E62961DD10D4764B0305D5.taxon	distribution	Distribution. Eastern (Figure 134).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
1BCCBDA47084977C6BA7631AA07948A7.taxon	materials_examined	Material examined. HOLOTYPE (♀): from USA, Missouri, Ozark County, Bryant Creek, downstream of Highway 95 bridge, 10 Sep 2011, by AJ Radwell, AJR 110105 A, DNA 2120. PARATYPES (9 ♀; 7 ♂): Maine, USA: 1 ♂ from Aroostook County, Ashland, beside Route 11 at bridge, Aroostook River (46 ° 38 ' N, 68 ° 24 ' W), 4 Jul 1989, by IM Smith, IMS 890067 * Missouri, USA: 1 ♀ and 1 ♂ from Oregon County, beside Route 19, north of Greer, Eleven Point River (36 ° 48 ' N, 91 ° 20 ' W), 28 Jun 1987, by IM Smith, IMS 870056 * 1 ♂ (ALLOTYPE) from Ozark County, Bryant Creek, downstream of Highway 95 bridge, 10 Sep 2011, by AJ Radwell, AJR 110105 A * 1 ♀ from Ozark County, Bryant Creek, downstream of Highway 95 bridge, 10 Sep 2011, by AJ Radwell, AJR 110105 A * 1 ♀ and 1 ♂ from McDonald County, Tiff City, beside Route 43, Buffalo Creek (36 ° 40 ' 17 " N, 94 ° 36 ' 17 " W), 2 May 1996, by IM Smith, IMS 960004 * New Brunswick, Canada: 3 ♀ from Charlotte County, Rollingham, Digdegaush River, beside Highway 770, 3 Oct 2011, by IM Smith, IMS 110118 * New Hampshire, USA: 1 ♀ from Coos County, picnic area beside Route 110, Upper Ammonoosuc River (44 ° 36 ' N, 71 ° 24 ' W), 5 Jul 1989, by IM Smith, IMS 890071 * Tennessee, USA: 1 ♀ and 1 ♂ from Blount County, Great Smokey Mountains National Park, Cades Cove, Forge Creek (35 ° 35 ' 31 " N, 83 ° 51 ' 21 " W), 17 Sep 2010, by IM Smith, IMS 100141 * 1 ♂ from Sevier Co., Great Smokey Mountains National Park, Middle Prong Little Pigeon River (35 ° 43 ' 33 " N, 83 ° 24 ' 1 " W), 12 Sep 2010, by IM Smith, IMS 100131 * Virginia, USA: 1 ♀ and 1 ♂ from Amherst County, Upper Otter Creek Overlook beside Blue Ridge, Otter Creek (37 ° 36 ' 57 " N, 79 ° 19 ' 27 " W), 7 Sep 2007, by IM Smith, IMS 070056 A * 1 ♂ from Giles County, Mechanicsburg, beside Dismal Creek Road, Standrock Brook (37 ° 11 ' 38 " N, 80 ° 53 ' 26 " W), 9 Sep 2005, by IM Smith, IMS 050066.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
1BCCBDA47084977C6BA7631AA07948A7.taxon	diagnosis	Diagnosis. Torrenticola malarkeyorum are similar to other members of the Rusetria " Eastern 2 - Plates " group (T. biscutella, T. caerulea, T. delicatexa, T. indistincta, T. pendula, T. sellersorum, T. tysoni, T. ululata, T. whitneyae, T. microbiscutella, and T. feminellai) in having anterio-lateral platelets fused to the dorsal plate, having dorsal coloration separated into anterior and posterior portions (except T. ululata and T. indistincta), and being distributed in the east. T. malarkeyorum can have variable coloration, including light bluish purple and reddish purple. Although several other species are purplish, some T. malarkeyorum are easily recognizable because they have bluish-purple coloration similar to T. tysoni and T. biscutella, albeit much fainter than these species. T. malarkeyorum can be differentiated from T. ululata, T. indistincta, and T. feminellai by dorsal coloration and pattern. T. malarkeyorum can be differentiated from T. tysoni by having a stockier rostrum (length / width = 2.57 - 2.89 in T. malarkeyorum, 3.06 - 3.5 in T. tysoni). Female T. malarkeyorum can be differentiated from female T. biscutella by having a longer subcapitulum (ventral length = 317.5 - 335 in T. malarkeyorum, 290 - 315 in T. biscutella). Male T. malarkeyorum can be differentiated from male T. biscutella by having a slightly more ovoid dorsum (length / width = 1.42 - 1.56 in T. malarkeyorum; 1.37 - 1.42 in T. biscutella. Additionally, although T. malarkeyorum and T. biscutella have the same dorsal coloration and pattern, often the coloration is faint in T. malarkeyorum and bold in T. biscutella. Female T. malarkeyorum can be differentiated from female T. caerulea by having a wider genital field (152.5 - 165 in T. malarkeyorum, 120 - 145 in T. caerulea). Additionally, T. malarkeyorum can be differentiated from T. caerulea by dorsal coloration and pattern. T. malarkeyorum can be differentiated from T. pendula by having a stockier gnathosomal bay (length / width = 1.62 - 2.26 in T. malarkeyorum, 2.42 - 2.90 in T. pendula); more elongate pedipalp tibiae (length / width = 3.22 - 3.60 in T. malarkeyorum, 2.78 - 3.05 in T. pendula); and by dorsal pattern. T. malarkeyorum can be differentiated from T. whitneyae by having more elongate pedipalpal tibiae (length / width = 3.22 - 3.60 in T. malarkeyorum, 2.42 - 2.95 in T. whitneyae) and by anterior venter / genital field length (♀ = 0.85 - 0.89 in T. malarkeyorum, 0.59 - 0.75 in T. whitneyae; ♂ = 1.52 - 1.88 in T. malarkeyorum, 1.37 - 1.43 in T. whitneyae). T. malarkeyorum can be differentiated from T. microbiscutella by having a less elongate dorsum (length / width = 1.33 - 1.56 in T. malarkeyorum, 1.63 - 1.75 in T. microbiscutella). Female T. malarkeyorum can be differentiated from female T. delicatexa by having a shorter genital field (162.5 - 170 in T. malarkeyorum, 175 - 198 in T. delicatexa). Female T. malarkeyorum can be differentiated from female T. sellersorum by anterior venter / genital field length (0.85 - 0.89 in T. malarkeyorum, 0.69 - 0.77 in T. sellersorum). Male T. malarkeyorum do not have any measurement differences with male T. delicatexa, and T. sellersorum; however, they can be differentiated by dorsal coloration.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
1BCCBDA47084977C6BA7631AA07948A7.taxon	description	Description. Female (Figure 138) (n = 5) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (590 - 640 (635) long; 425 - 470 (470) wide) ovoid with highly variable coloration, usually faint (occasionally bold) bluish-purple or reddish-purple separated into anterior and posterior portions, and with faint orange medially. Anterio-medial platelets (121.25 - 135 (132.5) long; 41.25 - 47.5 (45) wide). Anterio-lateral platelets (150 - 170 (170) long; 55 - 70 (70) wide) fused to dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 320 - 335 (330 )). Dorsal plate proportions: dorsum length / width 1.33 - 1.41 (1.35); dorsal width / distance between Dgl- 4 1.27 - 1.44 (1.42); anterio-medial platelet length / width 2.68 - 3.18 (2.94); anterio-lateral platelet length / width 2.38 - 2.73 (2.43); anterio-lateral / anterio-medial length 1.11 - 1.36 (1.28). Gnathosoma - Subcapitulum (317.5 - 335 (335) long (ventral); 234 - 250 (250) long (dorsal); 135 - 155 (155) tall) colorless. Rostrum (123.75 - 130 (130) long; 45 - 47.5 (45) wide). Chelicerae (323.75 - 342.5 (342.5) long) with curved fangs (59 - 70 (70) long). Subcapitular proportions: ventral length / height 2.13 - 2.35 (2.16); rostrum length / width 2.61 - 2.89 (2.89). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (42.5 - 50 (47.5) long); femur (115 - 123.75 (120) long); genu (67.5 - 72.5 (72.5) long); tibia (90 - 92.5 (90) long; 25 - 27.5 (27.5) wide); tarsus (17.5 - 20 (20) long). Palpomere proportions: femur / genu 1.62 - 1.71 (1.66); tibia / femur 0.73 - 0.80 (0.75); tibia length / width 3.27 - 3.60 (3.27). Venter - (700 - 770 (760) long; 482 - 600 (600) wide) colorless (occasionally with faint bluish-purple coloration). Gnathosomal bay (165 - 182.5 (182.5) long; 97.5 - 110 (110) wide). Cxgl- 4 subapical. Medial suture absent. Genital plates (162.5 - 170 (170) long; 152.5 - 165 (165) wide). Additional measurements: Cx- 1 (284 - 330 (330) long (total); 116 - 145 (145) long (medial )); Cx- 3 (316 - 385 (385) wide); anterior venter (137.5 - 150 (150) long). Ventral proportions: gnathosomal bay length / width 1.65 - 1.82 (1.66); anterior venter / genital field length 0.85 - 0.89 (0.88); anterior venter length / genital field width 0.88 - 0.95 (0.91). Male (Figure 139) (n = 5) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (420 - 470 (470) long; 285 - 320 (320) wide) ovoid with highly variable coloration, usually faint (occasionally bold) bluish-purple or reddish-purple separated into anterior and posterior portions, and with faint orange medially. Anterio-medial platelets (92.5 - 110 (110) long; 32.5 - 37.5 (37.5) wide). Anterio-lateral platelets (117.5 - 127.5 (127.5) long; 45 - 52.5 (52.5) wide) fused to dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 210 - 240 (240 )). Dorsal plate proportions: dorsum length / width 1.42 - 1.56 (1.47); dorsal width / distance between Dgl- 4 1.29 - 1.36 (1.33); anterio-medial platelet length / width 2.80 - 3.15 (2.93); anterio-lateral platelet length / width 2.43 - 2.61 (2.43); anterio-lateral / anterio-medial length 1.12 - 1.32 (1.33). Gnathosoma - Subcapitulum (227.5 - 240 (230) long (ventral); 162 - 180 (163) long (dorsal); 95 - 100 (100) tall) colorless. Rostrum (90 - 95 (92.5) long; 32.5 - 35 (35) wide). Chelicerae (212.5 - 230 long) with curved fangs (40 - 50 long). Subcapitular proportions: ventral length / height 2.28 - 2.53 (2.30); rostrum length / width 2.57 - 2.85 (2.64). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (32.5 - 40 (35) long); femur (76.25 - 87.5 (87.5) long); genu (52.5 - 55 (55) long); tibia (65 - 75 (75) long; 20 - 22.5 (22.5) wide); tarsus (15 - 17.5 (17.5) long). Palpomere proportions: femur / genu 1.39 - 1.62 (1.59); tibia / femur 0.76 - 0.92 (0.86); tibia length / width 3.22 - 3.50 (3.33). Venter - (495 - 560 (560) long; 320 - 405 (396) wide) colorless (occasionally with faint bluish-purple coloration). Gnathosomal bay (97.5 - 130 (115) long; 55 - 65 (65) wide). Cxgl- 4 subapical. Medial suture (65 - 72.5 (72.5) long). Genital plates (97.5 - 115 (110) long; 95 - 120 (105) wide). Additional measurements: Cx- 1 (197.5 - 230 (230) long (total); 90 - 115 (107) long (medial )); Cx- 3 (250 - 300 (295) wide); anterior venter (160 - 200 (200) long). Ventral proportions: gnathosomal bay length / width 1.62 - 2.26 (1.77); anterior venter / genital field length 1.52 - 1.88 (1.82); anterior venter length / genital field width 1.53 - 2.00 (1.90); anterior venter / medial suture 2.37 - 2.92 (2.76). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
1BCCBDA47084977C6BA7631AA07948A7.taxon	etymology	Etymology. Specific epithet (malarkeyorum) named in honor of JRF's sister, Mayme Malarkey and her family-Andy, Jack, Molly, and Lucy-who are a constant joy to all whom they encounter.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
1BCCBDA47084977C6BA7631AA07948A7.taxon	distribution	Distribution. Eastern (Figure 137), especially highlands (Appalachians and Ozarks).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
D5FDE25E633432965886BFCAECE63429.taxon	materials_examined	Material examined. HOLOTYPE (♀): from USA, New Mexico Catron County, Whitewater Creek, Glenwood Whitewater Picnic Area, 5 May 2012, by IM Smith, IMS 120005, DNA 2906. PARATYPES (2 ♀; 4 ♂): New Mexico, USA: 1 ♂ (ALLOTYPE) from Catron County, Whitewater Creek, Glenwood Whitewater Picnic Area, 5 May 2012, by IM Smith, IMS 120005, DNA 2907 * 1 ♀ and 1 ♂ from Catron County, Whitewater Creek, Glenwood Whitewater Picnic Area, 5 May 2012, by IM Smith, IMS 120005 * 1 ♀ and 2 ♂ from Catron County, Whitewater Creek, Glenwood Whitewater Picnic Area, 12 Jul 1987, by IM Smith, IMS 870084.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
D5FDE25E633432965886BFCAECE63429.taxon	diagnosis	Diagnosis. Torrenticola manni are similar to members of the Miniforma group (T. copipalpa, T. miniforma, T. pacificensis, T. rockyensis, T. oliveri, and T. pinocchio) in having short, stocky pedipalps (except T. oliveri and T. pinocchio); similar pedipalpal extensions (unique to members of this group); and being among the smallest Torrenticola in the west (dorsum 500 - 625 long) (except T. oliveri). T. manni can be differentiated from all other Miniforma group by being from the southwest (all others are from the northwest or Rocky Mountains). T. manni are best differentiated from T. rockyensis by having more elongate pedipalp tibiae (length / width = 3.13 - 3.38 in T. manni, 2.47 - 3.11 in T. rockyensis). T. manni are best differentiated from T. copipalpa by having tuberculate pedipalp femoral extensions (broad and flat in T. copipalpa). T. manni are best differentiated from T. pacificensis by having more elongate tibiae (length / width = 3.24 - 3.38 in T. manni, 2.67 - 3.00 in T. pacificensis); having more elongate subcapitular rostra (length / width ♀ = 3.00 - 3.13 in T. manni, 2.59 - 2.68 in T. pacificensis; ♂ = 3.13 - 3.20 in T. manni, 2.76 - 3.07 in T. pacificensis). T. manni are best differentiated from T. miniforma by being larger (dorsum length ♀ = 565 - 620 in T. manni, 545 in T. miniforma; ♂ = 535 - 550 in T. manni, 485 in T. miniforma) and having more elongate pedipalp tibiae (length / width = 3.13 - 3.38 in T. manni, 2.38 - 2.88 in T. miniforma). T. manni can be differentiated from T. oliveri by having a shorter anterior venter (192 - 230 in T. manni, 250 - 310 in T. oliveri) and less elongate pedipalpal tibiae (length / width = 3.13 - 3.38 in T. manni, 3.68 - 4.13 in T. oliveri). T. manni can be differentiated from T. pinocchio by having a less elongate rostrum (length / width = 3.0 - 3.2 in T. manni, 4.5 - 4.9 in T. pinocchio) and a more ovoid dorsum (length / width = 1.25 - 1.36 in T. manni, 1.53 - 1.64 in T. pinocchio).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
D5FDE25E633432965886BFCAECE63429.taxon	description	Description. Female (Figure 141) (n = 3) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (565 - 620 (620) long; 390 - 450 (450) wide) ovoid with faint purple coloration restricted posteriorly. Anterio-medial platelets (110 - 125 (125) long; 52.5 - 56.25 (56.25) wide). Anterio-lateral platelets (170 - 182.5 (182.5) long; 62.5 - 70 (70) wide) free from dorsal plate. Dgl- 4 much closer to the edge of dorsum than to the muscle scars (distance between Dgl- 4 295 - 335 (335 )). Dorsal plate proportions: dorsum length / width 1.38 - 1.45 (1.38); dorsal width / distance between Dgl- 4 1.32 - 1.36 (1.34); anterio-medial platelet length / width 2.10 - 2.22 (2.22); anterio-lateral platelet length / width 2.61 - 2.81 (2.61); anterio-lateral / anterio-medial length 1.46 - 1.55 (1.46). Gnathosoma - Subcapitulum (305 - 337.5 (337.5) long (ventral); 225 - 255 (255) long (dorsal); 110 - 121.25 (121.25) tall) colorless. Rostrum (125 - 140 (140) long; 40 - 45 (45) wide). Chelicerae (305 - 332 (332) long) with curved fangs (50 - 64 (57) long). Subcapitular proportions: ventral length / height 2.77 - 2.82 (2.78); rostrum length / width 3.00 - 3.11 (3.11). Pedipalps short and stocky (especially tibiae) with broadly tuberculate ventral extensions with dentate tip on femora and dentate, flanged ventral extensions on genua. Palpomeres: trochanter (30 - 35 (35) long); femur (95 - 102.5 (102.5) long); genu (65 - 72.5 (72.5) long); tibia (62.5 - 70 (68.75) long; 20 - 21.25 (21.25) wide); tarsus (15 - 16.25 (16.25) long). Palpomere proportions: femur / genu 1.41 - 1.46 (1.41); tibia / femur 0.66 - 0.68 (0.67); tibia length / width 3.13 - 3.29 (3.24). Venter - (700 - 780 (780) long; 458 - 477 (477) wide) colorless. Gnathosomal bay (142.5 - 150 (150) long; 60 - 70 (70) wide). Cxgl- 4 subapical. Medial suture (45 - 50 (45) long). Genital plates (180 - 192.5 (192.5) long; 160 - 180 (180) wide). Additional measurements: Cx- 1 (255 - 281 (275) long (total); 120 - 134 (131) long (medial )); Cx- 3 (290 - 307 (307) wide); anterior venter (192.5 - 202.5 (202.5) long). Ventral proportions: gnathosomal bay length / width 2.11 - 2.38 (2.14); anterior venter / genital field length 1.05 - 1.07 (1.05); anterior venter length / genital field width 1.13 - 1.23 (1.13); anterior venter / medial suture 3.85 - 4.50 (4.50). Male (Figure 142) (n = 4) (allotypic measurements in parentheses when available )) with characters of the genus with following specifications. Dorsum - (535 - 550 (545) long; 375 - 400 (390) wide) ovoid with faint purple coloration restricted posteriorly. Anterio-medial platelets (105 - 112.5 (112.5) long; 50 - 52.5 (52.5) wide). Anterio-lateral platelets (157.5 - 175 (167.5) long; 62.5 - 65 (65) wide) free from dorsal plate. Dgl- 4 much closer to the edge of dorsum than to the muscle scars (distance between Dgl- 4 300 - 305 (305 )). Dorsal plate proportions: dorsum length / width 1.34 - 1.47 (1.40); dorsal width / distance between Dgl- 4 1.25 - 1.33 (1.28); anterio-medial platelet length / width 2.00 - 2.15 (2.14); anterio-lateral platelet length / width 2.52 - 2.80 (2.58); anterio-lateral / anterio-medial length 1.49 - 1.58 (1.49). Gnathosoma - Subcapitulum (290 - 305 (297.5) long (ventral); 220 - 230 (227.5) long (dorsal); 100 - 110 (107.5) tall) colorless. Rostrum (120 - 127.5 (127.5) long; 37.5 - 40 (40) wide). Chelicerae (285 - 305 (300) long) with curved fangs (45 - 50 (46) long). Subcapitular proportions: ventral length / height 2.73 - 2.90 (2.77); rostrum length / width 3.13 - 3.20 (3.19). Pedipalps short and stocky (especially tibiae) with broadly tuberculate ventral extensions with dentate tip on femora and dentate, flanged ventral extensions on genua. Palpomeres: trochanter (27.5 - 32.5 (32.5) long); femur (87.5 - 97.5 (92.5) long); genu (60 - 65 (65) long); tibia (62.5 - 67.5 (67.5) long; 18.75 - 20 (20) wide); tarsus (13.75 - 15 (15) long). Palpomere proportions: femur / genu 1.40 - 1.50 (1.42); tibia / femur 0.65 - 0.73 (0.73); tibia length / width 3.19 - 3.38 (3.38). Venter - (665 - 695 (695) long; 424.75 - 490 (424.75) wide) colorless. Gnathosomal bay (115 - 147.5 (140) long; 55 - 62.5 (62.5) wide). Cxgl- 4 subapical. Medial suture (87.5 - 90 (87.5) long). Genital plates (140 - 150 (150) long; 112.5 - 116.25 (116.25) wide). Additional measurements: Cx- 1 (240 - 272 (272) long (total); 115 - 125 (122) long (medial )); Cx- 3 (275 - 315 (278) wide); anterior venter (220 - 230 (230) long). Ventral proportions: gnathosomal bay length / width 1.92 - 2.68 (2.24); anterior venter / genital field length 1.52 - 1.58 (1.53); anterior venter length / genital field width 1.91 - 1.98 (1.98); anterior venter / medial suture 2.44 - 2.63 (2.63). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
D5FDE25E633432965886BFCAECE63429.taxon	etymology	Etymology. Specific epithet (manni) named in honor of author Charles Mann, whose books about the peopling of North America (e. g., 1491, 1493) breach history and venture into human ecology. They are an inspiration to confronting misconceptions and a reminder that even seemingly well-known history, whether archeological or evolutionary, is in fact usually not well-known. His books are a battle cry to never cease learning and to always question.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
D5FDE25E633432965886BFCAECE63429.taxon	distribution	Distribution. Known only from Catron County, New Mexico (Figure 140).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
54C4B243F42FD44984E9EB9A88DEA37D.taxon	materials_examined	Material examined. HOLOTYPE (♀): from USA, Georgia, White County, Helen; beside Road to Anna Ruby Falls just north of Unicoi State Park, (34 ° 44 ' 44 " N, 83 ° 43 ' 43 " W), 24 September 1992, by IM Smith, IMS 920051. PARATYPES (5 ♀; 6 ♂): Georgia, USA: 1 ♂ (ALLOTYPE) from White County, Helen; beside Road to Anna Ruby Falls just north of Unicoi State Park, (34 ° 44 ' 44 " N, 83 ° 43 ' 43 " W), 24 September 1992, by IM Smith, IMS 920051 * 3 ♀ and 4 ♂ from White County, Helen; beside Road to Anna Ruby Falls just north of Unicoi State Park, (34 ° 44 ' 44 " N, 83 ° 43 ' 43 " W), 24 September 1992, by IM Smith, IMS 920051 * North Carolina, USA: * 2 ♀ and 2 ♂ from Macon County, Rainbow Springs; beside Forest Route 67, 2.0 km south of road to Standing Indian Campground, (35 ° 3 ' 3 " N, 83 ° 31 ' 31 " W), 1 July 1990, by IM Smith, IMS 900072.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
54C4B243F42FD44984E9EB9A88DEA37D.taxon	diagnosis	Diagnosis. Torrenticola microbiscutella are similar to other members of the Rusetria " Eastern 2 - Plates " group (T. biscutella, T. caerulea, T. delicatexa, T. indistincta, T. malarkeyorum, T. pendula, T. sellersorum, T. tysoni, T. ululata, T. whitneyae, and T. feminellai) in having anterio-lateral platelets fused to the dorsal plate, having dorsal coloration separated into anterior and posterior portions (except T. ululata and T. indistincta), and being distributed in the east. T. microbiscutella can be differentiated from all other Eastern 2 - Plates by having an elongate dorsum (length / width = 1.63 - 1.75 in T. microbiscutella, 1.21 - 1.56 in others). T. microbiscutella can be differentiated from most other Eastern 2 - Plates by having faint dorsal coloration (most other Eastern 2 - Plates have bold coloration).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
54C4B243F42FD44984E9EB9A88DEA37D.taxon	description	Description. Female (Figure 144) (n = 5) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (490 - 540 (510) long; 290 - 325 (300) wide) ovoid and elongate with faint reddish-purple coloration separated into anterior and posterior portions. Anterio-medial platelets (105 - 112.5 (110) long; 32.5 - 36.25 (33.75) wide). Anterio-lateral platelets (130 - 147.5 (135) long; 35 - 45 (45) wide) fused to dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 235 - 250 (235 )). Dorsal plate proportions: dorsum length / width 1.65 - 1.75 (1.70); dorsal width / distance between Dgl- 4 1.23 - 1.30 (1.28); anterio-medial platelet length / width 3.00 - 3.46 (3.26); anterio-lateral platelet length / width 3.00 - 3.79 (3.00); anterio-lateral / anterio-medial length 1.20 - 1.34 (1.23). Gnathosoma - Subcapitulum (260 - 270 (270) long (ventral); 195 - 210 (210) long (dorsal); 105 - 110 (107.5) tall) colorless. Rostrum (102.5 - 107.5 (107.5) long; 40 - 42.5 (42.5) wide) conical. Chelicerae (255 - 265 (262.5) long) with curved fangs (55 - 55 (55) long). Subcapitular proportions: ventral length / height 2.45 - 2.57 (2.51); rostrum length / width 2.53 - 2.63 (2.53). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (35 - 37.5 (37.5) long); femur (92.5 - 97.5 (97.5) long); genu (55 - 60 (57.5) long); tibia (72.5 - 80 (75) long; 20 - 22.5 (21.25) wide); tarsus (15 - 17.5 (17.5) long). Palpomere proportions: femur / genu 1.57 - 1.70 (1.70); tibia / femur 0.77 - 0.84 (0.77); tibia length / width 3.53 - 3.65 (3.53). Venter - (610 - 660 (620) long; 335 - 370 (370) wide) colorless. Gnathosomal bay (120 - 142.5 (120) long; 65 - 75 (75) wide). Cxgl- 4 subapical. Medial suture (42.5 - 50 (45) long). Genital plates (135 - 150 (145) long; 117.5 - 125 (120) wide). Additional measurements: Cx- 1 (220 - 240 (230) long (total); 95 - 105 (105) long (medial )); Cx- 3 (235 - 270 (270) wide); anterior venter (150 - 160 (157.5) long). Ventral proportions: gnathosomal bay length / width 1.60 - 2.04 (1.60); anterior venter / genital field length 1.07 - 1.15 (1.09); anterior venter length / genital field width 1.25 - 1.33 (1.31); anterior venter / medial suture 3.10 - 3.76 (3.50). Male (Figure 145) (n = 5) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (430 - 455 (440) long; 260 - 280 (260) wide) ovoid and elongate with faint reddish-purple coloration separated into anterior and posterior portions. Anterio-medial platelets (82.5 - 101.25 (95) long; 31.25 - 35 (35) wide). Anterio-lateral platelets (115 - 127.5 (115) long; 35 - 40 (37.5) wide) fused to dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 205 - 215 (205 )). Dorsal plate proportions: dorsum length / width 1.63 - 1.69 (1.69); dorsal width / distance between Dgl- 4 1.21 - 1.30 (1.27); anterio-medial platelet length / width 2.64 - 3.12 (2.71); anterio-lateral platelet length / width 3.07 - 3.64 (3.07); anterio-lateral / anterio-medial length 1.21 - 1.45 (1.21). Gnathosoma - Subcapitulum (215 - 235 (232.5) long (ventral); 162.5 - 177.5 (176.25) long (dorsal); 77.5 - 90 (87.5) tall) colorless. Rostrum (82.5 - 95 (92.5) long; 30 - 36.25 (35) wide) conical. Chelicerae (197.5 - 232.5 (212.5) long) with curved fangs (45 - 47.5 (47.5) long). Subcapitular proportions: ventral length / height 2.58 - 2.77 (2.66); rostrum length / width 2.62 - 2.85 (2.64). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (30 - 32.5 (32.5) long); femur (75 - 81.25 (76.25) long); genu (50 - 53.75 (50) long); tibia (66.25 - 70 (67.5) long; 18.75 - 20 (20) wide); tarsus (15 - 17.5 (15) long). Palpomere proportions: femur / genu 1.50 - 1.60 (1.53); tibia / femur 0.84 - 0.90 (0.89); tibia length / width 3.31 - 3.60 (3.38). Venter - (530 - 575 (540) long; 270 - 310 (305) colorless. Gnathosomal bay (110 - 125 (110) long; 55 - 60 (60) wide). Cxgl- 4 subapical. Medial suture (75 - 90 (90) long). Genital plates (85 - 95 (90) long; 85 - 100 (85) wide). Additional measurements: Cx- 1 (200 - 220 (205) long (total); 95 - 100 (100) long (medial )); Cx- 3 (220 - 240 (240) wide); anterior venter (185 - 195 (195) long). Ventral proportions: gnathosomal bay length / width 1.83 - 2.18 (1.83); anterior venter / genital field length 2.00 - 2.29 (2.17); anterior venter length / genital field width 1.95 - 2.29 (2.29); anterior venter / medial suture 2.17 - 2.47 (2.17). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
54C4B243F42FD44984E9EB9A88DEA37D.taxon	etymology	Etymology. Specific epithet (microbiscutella) is named because this species is the smallest (at least in females) and most elongate of all members of the Rusetria Complex (mikros, G. small; bi, L. two; scutella, L. little plate).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
54C4B243F42FD44984E9EB9A88DEA37D.taxon	distribution	Distribution. Southern Appalachians (Figure 143).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
229C229D8155E936C156B0F740C9496A.taxon	materials_examined	Material examined. LECTOTYPE (1 ♀): from USA, California, Humboldt County, Prairie Creek Redwoods State Park, Prairie Creek, 12 Jul 1964, by H Habeeb, HH 640021. PARALECTOTYPE (1 ♂): from USA, California, Humboldt County, Prairie Creek Redwoods State Park, Prairie Creek, 12 Jul 1964, by H Habeeb, HH 640021.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
229C229D8155E936C156B0F740C9496A.taxon	diagnosis	Diagnosis. Torrenticola miniforma are similar to members of the Miniforma group (T. copipalpa, T. manni, T. pacificensis, T. rockyensis, T. oliveri, and T. pinocchio) in having short, stocky pedipalps (except T. oliveri and T. pinocchio); similar pedipalpal extensions (unique to members of this group); and being among the smallest Torrenticola in the west (dorsum 500 - 625 long) (except T. oliveri). T. miniforma are best differentiated from T. rockyensis by being smaller (dorsum length ♀ = 545 in T. miniforma, 570 - 620 in T. rockyensis; ♂ = 485 in T. miniforma, 525 - 545 in T. rockyensis); having more elongate subcapitular rostra (length / width ♀ = 3.13 in T. miniforma, 2.72 - 2.91 in T. rockyensis; ♂ = 3.19 in T. miniforma, 2.83 - 3.00 in T. rockyensis); and by being only known from Humboldt County, California, whereas T. rockyensis is distributed in the Rocky Mountains (Idaho & Montana). T. miniforma are best differentiated from T. copipalpa by having tuberculate pedipalp femoral extensions (broad and flat in T. copipalpa). T. miniforma are best differentiated from T. pacificensis by having more elongate anterio-medial platelets (length / width = 2.44 - 2.65 in T. miniforma, 2.00 - 2.16 in T. pacificensis); more elongate subcapitular rostra (length / width = 3.13 - 3.19 in T. miniforma, 2.59 - 3.07 in T. pacificensis); and by being only known from Humboldt County, California, whereas T. pacificensis is distributed in the Pacific Coast Ranges of Washington and Oregon. T. miniforma are best differentiated from T. manni by being smaller (dorsum length ♀ = 545 in T. miniforma, 565 - 620 in T. manni; ♂ = 485 in T. miniforma, 535 - 550 in T. manni); having stockier pedipalp tibiae (length / width = 2.38 - 2.88 in T. miniforma, 3.13 - 3.38 in T. manni); and by being only known from Humboldt County, California, whereas T. manni is only known from Catron County, New Mexico. T. miniforma can be differentiated from T. oliveri by having a shorter anterior venter (172 - 200 in T. miniforma, 250 - 310 in T. oliveri) and having less elongate pedipalpal tibiae (length / width = 2.3 - 2.9 in T. miniforma, 3.6 - 4.2 in T. oliveri). T. miniforma can be differentiated from T. pinocchio by having a less elongate rostrum (length / width = 3.1 - 3.2 in T. miniforma, 4.5 - 4.9 in T. pinocchio) and a more ovoid dorsum (length / width = 1.43 - 1.43 in T. miniforma, 1.53 - 1.64 in T. pinocchio).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
229C229D8155E936C156B0F740C9496A.taxon	description	Re-description. Female (Figure 147) (n = 1) (lectotype only) with characters of the genus with following specifications. Dorsum - (545 long; 380 wide) ovoid and colorless. Anterio-medial platelets (112.5 long; 42.5 wide). Anterio-lateral platelets (152.5 long; 50 wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 300). Dorsal plate proportions: dorsum length / width 1.43; dorsal width / distance between Dgl- 4 1.27; anterio-medial platelet length / width 2.65; anterio-lateral platelet length / width 3.05; anterio-lateral / anterio-medial length 1.36. Gnathosoma - Subcapitulum (295 long (ventral); 225 long (dorsal); 112.5 tall) colorless. Rostrum (117.5 long; 37.5 wide). Chelicerae (290 long) with curved fangs (52.5 long). Subcapitular proportions: ventral length / height 2.62; rostrum length / width 3.13. Pedipalps short and stocky (especially tibiae) with tuberculate ventral extensions on femora and dentate, flanged ventral extensions on genua. Palpomeres: trochanter (32.5 long); femur (90 long); genu (62.5 long); tibia (57.5 long; 20 wide); tarsus (15 long). Palpomere proportions: femur / genu 1.44; tibia / femur 0.64; tibia length / width 2.88. Venter - (680 long; 420 wide) colorless. Gnathosomal bay (145 long; 62.5 wide). Cxgl- 4 subapical. Medial suture (50 long). Genital plates (167.5 long; 165 wide). Additional measurements: Cx- 1 (260 long (total); 50 long (medial )); Cx- 3 (280 wide); anterior venter (172.5 long). Ventral proportions: gnathosomal bay length / width 2.32; anterior venter / genital field length 1.03; anterior venter length / genital field width 1.05; anterior venter / medial suture 3.45. Male (Figure 148) (n = 1) (paralectotype only) with characters of the genus with following specifications. Dorsum - (485 long; 340 wide) ovoid and colorless. Anterio-medial platelets (97.5 long; 40 wide). Anterio-lateral platelets (145 long; 47.5 wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 270). Dorsal plate proportions: dorsum length / width 1.43; dorsal width / distance between Dgl- 4 1.26; anterio-medial platelet length / width 2.44; anterio-lateral platelet length / width 3.05; anterio-lateral / anterio-medial length 1.49. Gnathosoma - Subcapitulum (265 long (ventral); 200 long (dorsal); 95 tall) colorless. Rostrum (107.5 long; 33.75 wide). Chelicerae (265 long) with curved fangs (47.5 long). Subcapitular proportions: ventral length / height 2.79; rostrum length / width 3.19. Pedipalps short and stocky (especially tibiae) with tuberculate ventral extensions on femora and dentate, flanged ventral extensions on genua. Palpomeres: trochanter (30 long); femur (77.5 long); genu (52.5 long); tibia (47.5 long; 20 wide); tarsus (15 long). Palpomere proportions: femur / genu 1.48; tibia / femur 0.61; tibia length / width 2.38. Venter - (610 long; 380 wide) colorless. Gnathosomal bay (125 long; 50 wide). Cxgl- 4 subapical. Medial suture (87.5 long). Genital plates (125 long; 50 wide). Additional measurements: Cx- 1 (227.5 long (total); 102.5 long (medial )); Cx- 3 (250 wide); anterior venter (200 long). Ventral proportions: gnathosomal bay length / width 2.50; anterior venter / genital field length 1.54; anterior venter length / genital field width 1.86; anterior venter / medial suture 2.29. Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
229C229D8155E936C156B0F740C9496A.taxon	etymology	Etymology. Habeeb (1974) did not specify an etymology for the specific epithet (miniforma). However, we suspect the specific epithet refers to the small body size of this species, as with others of the Miniforma Group (minor, L. small; forma, L. form), as Habeeb (1974) notes this species is " the smallest species that has come to the attention of the writer from the pacific area. "	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
229C229D8155E936C156B0F740C9496A.taxon	distribution	Distribution. Known only from Prairie Creek, Humboldt County, California (Habeeb 1974) (Figure 146).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
A49CA197E6B092BD7461CE3B00899E6A.taxon	materials_examined	Material examined. HOLOTYPE (♀): from Canada, Ontario, Hastings County, Maple Leaf, Papineau Creek, beside Hwy 62, 18 Aug 2011, by IM Smith, IMS 110054, DNA 2860. PARATYPES (12 ♀; 12 ♂): Maine, USA: 1 ♀ and 1 ♂ from Kennebec County, China, beside Causeway along northern shore of China Lake (44 ° 28 ' 45 " N, 69 ° 30 ' 43 " W), 16 Sep 2003, by IM Smith, IMS 030037 * Ontario, Canada: 1 ♂ from Frontenac County, Salmon River, beside Highway 7 north of Arden, 26 - 28 Aug 1982, by IM Smith, CJ Hill, & C Cramer, IMS 820005 * 1 ♂ from Hastings County, Duff Corners, Vanderwater Conservation (44 ° 23 ' 14 " N, 77 ° 19 ' 1 " W), 13 Aug 2009, by IM Smith & ML MacKenzie, IMS 090089 A * 1 ♂ (ALLOTYPE) from Hastings County, Maple Leaf, Papineau Creek, beside Hwy 62, 18 Aug 2011, by IM Smith, IMS 110054, DNA 2857 * 1 ♀ and 1 ♂ from Hastings County, Maple Leaf, Papineau Creek, beside Hwy 62, 18 Aug 2011, by IM Smith, IMS 110054 * 6 ♀ and 5 ♂ from Hastings County, Maynooth, Papineau Creek, beside Highway 127, 17 Aug 2011, by IM Smith, IMS 110050 & IMS 110051 * 1 ♀ and 1 ♂ from Hastings County, Price Conservation Area, Skootamata River, junction of Highways 7 and 37, 20 May 1980, by IM Smith, IMS 800009 * 1 ♀ from Muskoka District, Baysville, Echo Lake, bay near Lawsons Cottages, 9 - 12 Aug 1980, by IM Smith, IMS 800015 * 1 ♀ from Muskoka District, Baysville, Echo Lake, 18 Aug 1981, by IM Smith, IMS 810023 * Quebec, Canada: 1 ♀ and 1 ♂ from Gatineau County, Gatineau Park, Lac Philippe, at campground, 2 Jun 1981, by IM Smith & CJ Hill, IMS 810001.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
A49CA197E6B092BD7461CE3B00899E6A.taxon	diagnosis	Diagnosis. Torrenticola mjolniri are similar to other members of the Raptor Group (T. gnoma, T. irapalpa, T. longitibia, T. elusiva, T. racupalpa, T. raptor, T. danielleae, T. daemon, and T. ivyae) in having round bodies; Dgl- 4 close to muscles scars; long, thin subcapitular rostra; and long, thin pedipalp tibiae. T. mjolniri can be differentiated from T. elusiva, T. gnoma, and T. daemon by having more elongate pedipalp tibiae (length / width = 5.00 - 6.00 in T. mjolniri, 3.88 - 4.67 in others) and a more elongate rostrum (length / width = 3.81 - 4.32 in T. mjolniri, 2.56 - 3.65 in others). T. mjolniri can be differentiated from T. racupalpa by having a longer anterior venter (♀ = 180 - 195 in T. mjolniri, 152.5 - 165 in T. racupalpa; ♂ = 230 - 255 in T. mjolniri, 200 - 205 in T. racupalpa) and by dorsal pattern. T. mjolniri can be differentiated from T. irapalpa and T. danielleae by having Dgl- 4 closer to the muscle scars (dorsal width / distance between Dgl- 4 = 2.2 - 2.48 in T. mjolniri, 1.42 - 2.09 in others); having a more elongate rostrum (3.81 - 4.32 in T. mjolniri, 2.66 - 3.75 in others); and by dorsal pattern. Male T. mjolniri can be differentiated from T. longitibia (only males) by having Dgl- 4 closer to the edge of dorsum (dorsal width / distance between Dgl- 4 = 2.2 - 2.32 in T. mjolniri, 2.46 - 2.71 in T. longitibia); stockier pedipalp tibiae (length / width = 5.0 - 5.33 in T. mjolniri, 5.5 - 5.5 in T. longitibia); and a more elongate subcapitulum (ventral length / height = 2.82 - 3.0 in T. mjolniri, 2.68 - 2.73 in T. longitibia). T. mjolniri can be differentiated from T. raptor by having Dgl- 4 closer to the muscle scars (dorsal width / distance between Dgl- 4 = 2.2 - 2.48 in T. mjolniri, 1.68 - 2.02 in T. raptor) and a stockier subcapitulum (ventral length / width ♀ = 2.68 - 2.9 in T. mjolniri, 2.98 - 3.18 in T. raptor; ♂ = 2.82 - 3.0 in T. mjolniri, 3.13 - 3.27 in T. raptor). Additionally, female T. mjolniri have a shorter anterior venter (180 - 195 in ♀ T. mjolniri, 205 - 240 in ♀ T. raptor). Female T. mjolniri can be differentiated from female T. ivyae by having a larger dorsum (length ♀ = 605 - 640 in T. mjolniri, 550 - 590 in T. ivyae; width ♀ = 510 - 545 in T. mjolniri, 460 - 500 in T. ivyae) and a longer anterior venter (♀ 180 - 195 in T. mjolniri, 155 - 170 in T. ivyae). Male T. mjolniri can be differentiated from male T. ivyae by having a more elongate subcapitulum (ventral length / width = 2.82 - 3.00 in T. mjolniri, 2.57 - 2.75 in T. ivyae). Additionally, T. mjolniri can be differentiated from T. ivyae by being distributed in the northeast (T. ivyae is known from Florida).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
A49CA197E6B092BD7461CE3B00899E6A.taxon	description	Description. Female (Figure 150) (n = 5) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (605 - 640 (605) long; 510 - 545 (520) wide) circular with navy blue to purple coloration posteriorly and a small spot anteriorly often connected medially with a thin strip that is occasionally orange. Anterio-medial platelets (130 - 150 (135) long; 67.5 - 70 (67.5) wide). Anterio-lateral platelets (182.5 - 200 (187.5) long; 67.5 - 70 (67.5) wide) free from dorsal plate. Dgl- 4 closer to the muscle scars than to edge dorsum (distance between Dgl- 4 207.5 - 245 (210 )). Dorsal plate proportions: dorsum length / width 1.16 - 1.24 (1.16); dorsal width / distance between Dgl- 4 2.22 - 2.48 (2.48); anterio-medial platelet length / width 1.86 - 2.14 (2.00); anterio-lateral platelet length / width 2.11 - 2.38 (2.27); anterio-lateral / anterio-medial length 1.22 - 1.46 (1.39). Gnathosoma - Subcapitulum (342.5 - 355 (342.5) long (ventral); 255 - 277 (262.5) long (dorsal); 122.5 - 128.75 (126.25) tall) colorless. Rostrum (142.5 - 151.25 (151.25) long; 35 - 38.75 (35) wide). Chelicerae (355 - 383 (360) long) with curved fangs (52.5 - 55 (52.5) long) elongate. Subcapitular proportions: ventral length / height 2.68 - 2.90 (2.71); rostrum length / width 3.81 - 4.32 (4.32). Pedipalps elongate (especially tibiae) with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (47.5 - 47.5 (47.5) long); femur (130 - 135 (130) long); genu (65 - 70 (65) long); tibia (110 - 112.5 (112.5) long; 18.75 - 22.5 (18.75) wide); tarsus (15 - 17.5 (15) long). Palpomere proportions: femur / genu 1.86 - 2.00 (2.00); tibia / femur 0.83 - 0.87 (0.87); tibia length / width 5.00 - 6.00 (6.00). Venter - (780 - 825 (790) long; 536 - 621 (570) wide) often colorless, occasionally areas surrounding coxae are navy blue to purple. Gnathosomal bay (170 - 175 (175) long; 70 - 82.5 (70) wide). Cxgl- 4 far from apex. Medial suture (15 - 20 (15) long). Genital plates (160 - 173.75 (170) long; 142.5 - 152.5 (142.5) wide). Additional measurements: Cx- 1 (305 - 330 long (total); 132 - 164 (162.5) long (medial )); Cx- 3 (332 - 383 wide); anterior venter (180 - 195 (187.5) long). Ventral proportions: gnathosomal bay length / width 2.06 - 2.50 (2.50); anterior venter / genital field length 1.07 - 1.22 (1.10); anterior venter length / genital field width 1.18 - 1.32 (1.32); anterior venter / medial suture 9.00 - 12.50 (12.50). Male (Figure 151) (n = 6) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (520 - 580 (520) long; 430 - 450 (430) wide) circular with navy blue to purple coloration posteriorly and a small spot anteriorly often connected medially with a thin strip that is occasionally orange. Anterio-medial platelets (120 - 135 (122.5) long; 60 - 67.5 (60) wide). Anterio-lateral platelets (175 - 200 (185) long; 67.5 - 77.5 (70) wide) free from dorsal plate. Dgl- 4 closer to the muscle scars than to edge dorsum (distance between Dgl- 4 185 - 205 (195 )). Dorsal plate proportions: dorsum length / width 1.21 - 1.29 (1.21); dorsal width / distance between Dgl- 4 2.20 - 2.32 (2.21); anterio-medial platelet length / width 1.85 - 2.04 (2.04); anterio-lateral platelet length / width 2.34 - 2.67 (2.64) anterio-lateral / anterio-medial length 1.40 - 1.51 (1.51). Gnathosoma - Subcapitulum (305 - 325 (305) long (ventral); 233 - 241 (233) long (dorsal); 105 - 112.5 (107.5) tall) colorless. Rostrum (130 - 140 (130) long; 32.5 - 32.5 (32.5) wide). Chelicerae (297 - 322 (297) long) with curved fangs (48 - 55 (49) long) elongate. Subcapitular proportions: ventral length / height 2.82 - 3.00 (2.84); rostrum length / width 4.00 - 4.31 (4.00). Pedipalps elongate (especially tibiae) with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (42.5 - 42 (42.5) long); femur (115 - 122.5 (117.5) long); genu (60 - 65 (65) long); tibia (100 - 102.5 (102.5) long; 18.75 - 20 (20) wide); tarsus (17.5 - 17.5 (17.5) long). Palpomere proportions: femur / genu 1.81 - 1.96 (1.81); tibia / femur 0.82 - 0.87 (0.87); tibia length / width 5.00 - 5.33 (5.13). Venter - (670 - 740 (670) long; 459 - 504 (479) wide) often colorless, occasionally areas surrounding coxae are navy blue to purple. Gnathosomal bay (142.5 - 160 (145) long; 60 - 67.5 (67.5) wide). Cxgl- 4 far from apex. Medial suture (62.5 - 85 (62.5) long). Genital plates (137.5 - 147.5 (137.5) long; 115 - 142.5 (115) wide). Additional measurements: Cx- 1 (275 - 324 (287) long (total); 139 - 152 (139) long (medial )); Cx- 3 (307 - 371 (324) wide); anterior venter (230 - 255 (230) long). Ventral proportions: gnathosomal bay length / width 2.15 - 2.46 (2.15); anterior venter / genital field length 1.60 - 1.76 (1.67); anterior venter length / genital field width 1.79 - 2.00 (2.00); anterior venter / medial suture 3.00 - 3.68 (3.68). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
A49CA197E6B092BD7461CE3B00899E6A.taxon	etymology	Etymology. Specific epithet (mjolniri) named after Mjoelnir-the hammer of Thor in Norse mythology-the ancient symbol of which resembles the dorsal patterning this species.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
A49CA197E6B092BD7461CE3B00899E6A.taxon	distribution	Distribution. Northeastern (Figure 149).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
691A665838B42705F6D4247A2F27C89F.taxon	materials_examined	Material examined. HOLOTYPE (♀): from USA, Maine, Aroostook County, Ashland; beside Rt. 11 at bridge over Aroostook River, (46 ° 38 ' 38 " N, 68 ° 24 ' 24 " W), 4 July 1989, by IM Smith, IMS 890067. PARATYPES (4 ♀; 5 ♂): 1 ♂ (ALLOTYPE) from Aroostook County, Ashland; beside Rt. 11 at bridge over Aroostook River, (46 ° 38 ' 38 " N, 68 ° 24 ' 24 " W), 4 July 1989, by IM Smith, IMS 890067 * 4 ♀ and 4 ♂ from Aroostook County, Ashland; beside Rt. 11 at bridge over Aroostook River, (46 ° 38 ' 38 " N, 68 ° 24 ' 24 " W), 4 July 1989, by IM Smith, IMS 890067.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
691A665838B42705F6D4247A2F27C89F.taxon	diagnosis	Diagnosis. Torrenticola mohawk are similar to other members of the Tricolor Complex (T. bittikoferae, T. hoosieri, T. larvata, T. pearsoni, T. olliei, T. sierrensis, T. tricolor, T. trimaculata, T. unimaculata, T. cardia, T. kringi, and T. dimorpha) in having a short, conical rostrum. T. mohawk are most similar to other members of the Tricolor Complex that have bold patterning (T. larvata, T. tricolor, T. trimaculata, T. unimaculata, T. cardia, and T. kringi). T. mohawk can be differentiated from T. larvata, T. tricolor, T. cardia, and T. kringi by having a stockier rostrum (length / width ♀ = 1.80 - 2.00 in T. mohawk, 2.14 - 3.13 in others; ♂ = 2.00 - 2.13 in T. mohawk, 2.27 - 2.93 in others). T. mohawk can be differentiated from T. unimaculata and T. trimaculata by dorsal pattern.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
691A665838B42705F6D4247A2F27C89F.taxon	description	Description. Female (Figure 153) (n = 5) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (680 - 725 (680) long; 540 - 560 (560) wide) ellipsoid with faint reddish-purple coloration medially. Anterio-medial platelets (130 - 140 (132.5) long; 60 - 77.5 (60) wide). Anterio-lateral platelets (185 - 197.5 (190) long; 75 - 82.5 (75) wide) free from dorsal plate. Dgl- 4 closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 365 - 385 (365 )). Dorsal plate proportions: dorsum length / width 1.21 - 1.32 (1.21); dorsal width / distance between Dgl- 4 1.40 - 1.53 (1.53); anterio-medial platelet length / width 1.81 - 2.21 (2.21); anterio-lateral platelet length / width 2.31 - 2.53 (2.53); anterio-lateral / anterio-medial length 1.40 - 1.43 (1.43). Gnathosoma - Subcapitulum (240 - 267.5 (260) long (ventral); 170 - 190 (185) long (dorsal); 115 - 125 (120) tall) colorless. Rostrum (82.5 - 95 (90) long; 42.5 - 52.5 (45) wide) short and conical. Chelicerae (225 - 255 (240) long) with curved fangs (50 - 60 (60) long). Subcapitular proportions: ventral length / height 2.08 - 2.33 (2.17); rostrum length / width 1.80 - 2.00 (2.00). Pedipalps with short, tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (37.5 - 42.5 (41.25) long); femur (87.5 - 100 (95) long); genu (60 - 67.5 (65) long); tibia (77.5 - 87.5 (82.5) long; 25 - 27.5 (27.5) wide); tarsus (22.5 - 23.75 (23.75) long). Palpomere proportions: femur / genu 1.38 - 1.48 (1.46); tibia / femur 0.87 - 0.92 (0.87); tibia length / width 3.00 - 3.18 (3.00). Venter - (800 - 850 (825) long; 600 - 630 (605) wide) colorless. Gnathosomal bay (125 - 145 (125) long; 70 - 75 (70) wide). Cxgl- 4 subapical. Medial suture (30 - 45 (35) long). Genital plates (190 - 205 (195) long; 157.5 - 170 (160) wide). Additional measurements: Cx- 1 (260 - 285 (260) long (total); 130 - 140 (130) long (medial )); Cx- 3 (355 - 370 (360) wide); anterior venter (187.5 - 200 (190) long). Ventral proportions: gnathosomal bay length / width 1.70 - 2.07 (1.79); anterior venter / genital field length 0.93 - 0.99 (0.97); anterior venter length / genital field width 1.12 - 1.21 (1.19); anterior venter / medial suture 4.44 - 6.33 (5.43). Male (Figure 154) (n = 5) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (615 - 660 (655) long; 460 - 485 (470) wide) ellipsoid with faint reddish-purple coloration medially. Anterio-medial platelets (120 - 130 (122.5) long; 65 - 76.25 (67.5) wide). Anterio-lateral platelets (180 - 192.5 (187.5) long; 75 - 80 (75) wide) free from dorsal plate Dgl- 4 closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 355 - 375 (360 )). Dorsal plate proportions: dorsum length / width 1.33 - 1.39 (1.39); dorsal width / distance between Dgl- 4 1.27 - 1.31 (1.31); anterio-medial platelet length / width 1.70 - 1.85 (1.81); anterio-lateral platelet length / width 2.25 - 2.50 (2.50); anterio-lateral / anterio-medial length 1.44 - 1.54 (1.53). Gnathosoma - Subcapitulum (227.5 - 250 (250) long (ventral); 170 - 172.5 (172.5) long (dorsal); 105 - 107.5 (107.5) tall) colorless. Rostrum (80 - 90 (90) long; 40 - 45 (45) wide) short and conical. Chelicerae (210 - 225 (225) long) with curved fangs (50 - 55 (55) long). Subcapitular proportions: ventral length / height 2.17 - 2.33 (2.33); rostrum length / width 2.00 - 2.13 (2.00). Pedipalps with short, tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (36.25 - 37.5 (37.5) long); femur (87.5 - 92.5 (90) long); genu (60 - 67.5 (65) long); tibia (75 - 80 (77.5) long; 26.25 - 28.75 (27.5) wide); tarsus (20 - 22.5 (22.5) long). Palpomere proportions: femur / genu 1.37 - 1.50 (1.38); tibia / femur 0.83 - 0.89 (0.86); tibia length / width 2.73 - 2.95 (2.82). Venter - (735 - 780 (780) long; 510 - 530 (520) wide) colorless. Gnathosomal bay (107.5 - 127.5 (120) long; 65 - 70 (70) wide). Cxgl- 4 subapical. Medial suture (120 - 130 (130) long). Genital plates (145 - 155 (152.5) long; 105 - 107.5 (105) wide). Additional measurements: Cx- 1 (250 - 280 (280) long (total); 145 - 155 (155) long (medial )); Cx- 3 (350 - 360 (360) wide); anterior venter (290 - 307.5 (307.5) long). Ventral proportions: gnathosomal bay length / width 1.54 - 1.82 (1.71); anterior venter / genital field length 1.93 - 2.03 (2.02); anterior venter length / genital field width 2.74 - 2.93 (2.93); anterior venter / medial suture 2.27 - 2.42 (2.37). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
691A665838B42705F6D4247A2F27C89F.taxon	etymology	Etymology. Specific epithet (mohawk) named in honor of the Mohawk people (Kanien'keha: ka), in reference to the dorsal coloration of this species, which resembles the hair-style by the same name (as a noun in apposition). The modern hairstyle does not exactly match traditional Mohawk styles, which resembled that of other tribes of the Iroquois Confederacy. Nevertheless, mohawks the hairstyle, which were originally associated with the Mohawk people in the historically problematic film Drums Along the Mohawk (1939), remains a symbol of the Mohawk people.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
691A665838B42705F6D4247A2F27C89F.taxon	distribution	Distribution. Maine (Figure 152).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
74963862567792A3ECDF7BAF3E3B0F99.taxon	materials_examined	Material examined. HOLOTYPE (♀): from USA, Wyoming, Fremont County, Wind River Mountains, Sinks Canyon, Popo Agie River, south of Lander, 1 Aug 2012, by IM Smith, IMS 120049, DNA 2928. PARATYPES (6 ♀; 5 ♂): Idaho, USA: 1 ♀ from Blaine County, Little Wood River (43 ° 29 ' 51 " N, 114 ° 3 ' 28 " W), 27 Jul 2012, by JR Fisher, WA Nelson, & JC O'Neill, ROW 12 - 0727 - 001 * 1 ♂ from Custer County, Morgan Creek (44 ° 39 ' 20 " N, 114 ° 12 ' 56 " W), 31 Jul 2012, by JR Fisher, WA Nelson, & JC O'Neill, ROW 12 - 0731 - 004 * Montana, USA: 1 ♀ and 1 ♂ from Ravalli County, Bitterroot National Forest, East Fork Bitterroot River (45 ° 51 ' 40 " N, 114 ° 1 ' 46 " W), 3 Aug 2012, by JR Fisher, WA Nelson, & JC O'Neill, ROW 12 - 0803 - 005 * 1 ♀ and 1 ♂ from Missoula County, Lolo National Forest, off 12, Lolo Creek (46 ° 46 ' 7 " N, 114 ° 27 ' 53 " W), 7 Aug 2012, by JR Fisher, WA Nelson, & JC O'Neill, ROW 12 - 0807 - 003 * Utah, USA: 1 ♀ from Summit County, Wasatch-Cache National Forest, Slate Creek (40 ° 37 ' 45 " N, 111 ° 11 ' 46 " W), 23 Jul 2012, by JR Fisher, WA Nelson, & JC O'Neill, ROW 12 - 0723 - 006 * 1 ♂ from Wasatch County, Wasatch-Cache National Forest, Provo River (40 ° 35 ' 37 " N, 111 ° 5 ' 43 " W), 23 Jul 2012, by JR Fisher, WA Nelson, & JC O'Neill, ROW 12 - 0723 - 001 * Wyoming, USA: 1 ♂ (ALLOTYPE) from Fremont County, Wind River Mountains, Sinks Canyon, Popo Agie River, south of Lander, 1 Aug 2012, by IM Smith, IMS 120049, DNA 2929 * 1 ♀ from Fremont County, Wind River Mountains, Sinks Canyon, Popo Agie River, south of Lander, 1 Aug 2012, by IM Smith, IMS 120049 * 1 ♀ from Johnson County, Bighorn Mountains, Clear Creek, west of Buffalo Mosier Gulch Picnic Area, 28 Jul 2012, by IM Smith, IMS 120041.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
74963862567792A3ECDF7BAF3E3B0F99.taxon	diagnosis	Diagnosis. Torrenticola mulleni are similar to other members of the Rusetria " Western 2 - Plates " group (T. nortoni, T. walteri, and T. welbourni) in having anterio-lateral platelets fused to the dorsal plate, having faint dorsal coloration, and being distributed in the west. Female T. mulleni can be differentiated from other female Western 2 - Plates by having a longer medial suture (20 - 22.5 in T. mulleni, 10 - 12.5 in others). Male T. mulleni can be differentiated from other male Western 2 - Plates by having a longer genital field (130 - 140 in T. mulleni, 112 - 125 in others). Additionally, T. mulleni can be differentiated from other Western 2 - Plates by being distributed in the Rocky Mountains (Idaho, Montana, Utah and Wyoming) instead of California, Oregon and British Columbia.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
74963862567792A3ECDF7BAF3E3B0F99.taxon	description	Description. Female (Figure 156) (n = 5) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (595 - 645 (640) long; 415 - 470 (440) wide) ovoid with faint purple coloration separated into anterior and posterior portions. Anterio-medial platelets (127.5 - 145 (145) long; 40 - 47.5 (45) wide). Anterio-lateral platelets (151.25 - 165 (160) long; 50 - 57.5 (57.5) wide) fused to dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 300 - 330 (320 )). Dorsal plate proportions: dorsum length / width 1.37 - 1.45 (1.45); dorsal width / distance between Dgl- 4 1.34 - 1.47 (1.38); anterio-medial platelet length / width 2.95 - 3.24 (3.22); anterio-lateral platelet length / width 2.75 - 3.15 (2.78); anterio-lateral / anterio-medial length 1.10 - 1.29 (1.10). Gnathosoma - Subcapitulum (300 - 327.5 (322.5) long (ventral); 225 - 245 (245) long (dorsal); 125 - 150 (145) tall) colorless. Rostrum (120 - 135 (125) long; 40 - 45 (45) wide). Chelicerae (300 - 335 (320) long) with curved fangs (40 - 65 (65) long). Subcapitular proportions: ventral length / height 2.14 - 2.40 (2.22); rostrum length / width 2.78 - 3.18 (2.78). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (42.5 - 52.5 (42.5) long); femur (112.5 - 125 (117.5) long); genu (65 - 72.5 (67.5) long); tibia (82.5 - 90 (82.5) long; 26.25 - 28.75 (26.25) wide); tarsus (17.5 - 20 (17.5) long). Palpomere proportions: femur / genu 1.70 - 1.75 (1.74); tibia / femur 0.70 - 0.78 (0.70); tibia length / width 3.0 - 3.33 (3.14). Venter - (720 - 790 (770) long; 460 - 500 (490) wide) colorless. Gnathosomal bay (160 - 175 (175) long; 75 - 90 (85) wide). Cxgl- 4 subapical. Medial suture (20 - 22.5 (20) long). Genital plates (195 - 205 (205) long; 170 - 180 (175) wide). Additional measurements: Cx- 1 (285 - 300 (300) long (total); 115 - 135 (130) long (medial )); Cx- 3 (302.5 - 335 (320) wide); anterior venter (145 - 162.5 (155) long). Ventral proportions: gnathosomal bay length / width 1.89 - 2.16 (2.06); anterior venter / genital field length 0.74 - 0.80 (0.76); anterior venter length / genital field width 0.85 - 0.91 (0.89); anterior venter / medial suture 6.89 - 8.0 (7.75). Male (Figure 157) (n = 4) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (505 - 535 (530) long; 350 - 370 (360) wide) ovoid with faint purple coloration separated into anterior and posterior portions. Anterio-medial platelets (107.5 - 115 (110) long; 40 - 41.25 (40) wide). Anterio-lateral platelets (142.5 - 147.5 (142.5) long; 50 - 52.5 (52.5) wide) fused to dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 270 - 280 (280 )). Dorsal plate proportions: dorsum length / width 1.44 - 1.47 (1.47); dorsal width / distance between Dgl- 4 1.29 - 1.37 (1.29); anterio-medial platelet length / width 2.61 - 2.88 (2.75); anterio-lateral platelet length / width 2.71 - 2.95 (2.71); anterio-lateral / anterio-medial length 1.28 - 1.33 (1.30). Gnathosoma - Subcapitulum (262.5 - 277.5 (265) long (ventral); 195 - 210 (202.5) long (dorsal); 105 - 110 (105) tall) colorless. Rostrum (100 - 110 (110) long; 37.5 - 40 (40) wide). Chelicerae (265 - 270 (270) long) with curved fangs (50 - 55 (55) long). Subcapitular proportions: ventral length / height 2.39 - 2.58 (2.52); rostrum length / width 2.63 - 2.75 (2.75). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (40 - 40 (40) long); femur (97.5 - 102.5 (102.5) long); genu (60 - 65 (62.5) long); tibia (75 - 82.5 (82.5) long; 26.25 - 27.5 (27.5) wide); tarsus (17.5 - 17.5 (17.5) long). Palpomere proportions: femur / genu 1.58 - 1.64 (1.64); tibia / femur 0.75 - 0.80 (0.80); tibia length / width 2.73 - 3.00 (3.00). Venter - (620 - 650 (650) long; 390 - 425 (410) wide) colorless. Gnathosomal bay (130 - 135 (132.5) long; 70 - 75 (75) wide). Cxgl- 4 subapical. Medial suture (100 - 105 (102.5) long). Genital plates (130 - 140 (140) long; 100 - 110 (110) wide). Additional measurements: Cx- 1 (265 - 285 (285) long (total); 140 - 145 (140) long (medial )); Cx- 3 (290 - 310 (310) wide); anterior venter (250 - 260 (250) long). Ventral proportions: gnathosomal bay length / width 1.77 - 1.93 (1.77); anterior venter / genital field length 1.79 - 2.00 (1.79); anterior venter length / genital field width 2.27 - 2.60 (2.27); anterior venter / medial suture 2.38 - 2.60 (2.44). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
74963862567792A3ECDF7BAF3E3B0F99.taxon	etymology	Etymology. Specific epithet (mulleni) named in honor of Gary Mullen of Auburn University, who boldly devoted the entire second half of his Arachnology course to mites, and thus, in 2001, first introduced JRF to these creatures, sparking a life-long fascination.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
74963862567792A3ECDF7BAF3E3B0F99.taxon	distribution	Distribution. Rocky Mountain region (Figure 155).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
FB959FB0C31D58B9BF53CE481BAC972E.taxon	materials_examined	Material examined. LECTOTYPE (1 ♀): from USA, California, Humboldt County, Prairie Creek Redwoods State Park, Prairie Creek, 12 Jul 1964, by Habeeb, HH 640021. PARALECTOTYPE (1 ♂): from USA, California, Humboldt County, Prairie Creek Redwoods State Park, Prairie Creek, 12 Jul 1964, by Habeeb, HH 640021. OTHER MATERIAL (95 ♀; 84 ♂): Arizona, USA: 1 ♀ from Cochise County, Chiricahua Mountains, Turkey Creek at Sycamore campground, east of Sunizona, 15 Jul 1987, by IM Smith, IMS 870091 * British Columbia, Canada: 1 ♀ and 1 ♂ from Bella Coola Valley, Tweedsmuir Provincial Park, Atnarko River, at campground, 28 Jul 1983, by IM Smith & AB Smith, IMS 830054 A * 2 ♀ and 2 ♂ from Bella Coola Valley, Tweedsmuir Provincial Park, Atnarko Slough, beside Highway 20, west of Youngs Creek Picnic Area, 4 Aug 1983, by IM Smith, IMS 830064 * 3 ♀ and 2 ♂ from Bella Coola Valley, Tweedsmuir Provincial Park, Belarko, Atnarko River, 24 - 26 Jul 1983, by IM Smith, IMS 830049 B & IMS 830049 C * 1 ♀ and 1 ♂ from Bella Coola Valley, Tweedsmuir Provincial Park, Hotnarko River, at end of Atnarko tote road, 27 Jul 1983, by IM Smith, IMS 830052 * 5 ♀ and 6 ♂ from Bella Coola Valley, Tweedsmuir Provincial Park, Hotnarko River, at end of Atnarko tote road, 31 Jul 1983, by IM Smith, IMS 830059 A & IMS 830059 B * 1 ♀ and 1 ♂ from Bella Coola Valley, Tweedsmuir Provincial Park, Youngs Creek Picnic Area, Atnarko Slough, 24 - 27 Jul 1983, by IM Smith, IMS 830048 A * 1 ♀ and 1 ♂ from Bella Coola Valley, Tweedsmuir Provincial Park, Youngs Creek Picnic Area, tributary of Atnarko Slough, 2 Aug 1983, by IM Smith, IMS 830062 C * 2 ♀ and 1 ♂ from Fernie, Lizard Creek, beside Highway 3, 1.8 km west of Fernie Mountain Provincial Park, 16 Aug 2012, by IM Smith, IMS 120073 * 1 ♂ from McClinchy River, beside Highway 20, west of Kleena Kleene, 5 Aug 1983, by IM Smith, IMS 830068 A * 1 ♀ and 1 ♂ from Tweedsmuir Provincial Park, Youngs Creek, beside Highway 20, between Heckman Pass & Bella Coola Valley, 5 Aug 1983, by IM Smith, IMS 830065 * 5 ♀ from Vancouver Island, Caycuse, Nixon Creek, 8 Jul 1976, by IM Smith, IMS 760197 & IMS 760198 * 3 ♀ and 1 ♂ from Vancouver Island, beside Highway 4, 35.6 kilometers east of Pacific Rim Road, 9 Jul 1976, by IM Smith, IMS 760206 * 1 ♀ and 1 ♂ from Vancouver Island, Lake Cowichan, South Shore Road, north of Mesachie Lake, Robertson River, 4 Jul 1976, by IM Smith, IMS 760183 A * 1 ♂ from Vancouver Island, Lake Cowichan, South Shore Road, north of Mesachie Lake, tributary of Robertson River, 4 - 10 Jul 1976, by IM Smith, IMS 760182 * 1 ♀ and 2 ♂ from Vancouver Island, Lost Shoe Creek, beside Highway 4, 1.3 kilometers east of Pacific Rim Road, 9 Jul 1976, by IM Smith, IMS 760202 * 1 ♀ and 1 ♂ from Vancouver Island, Malahat, Goldstream Provincial Park, Goldstream River, 26 Jun 1979, by IM Smith, IMS 790028 A * 1 ♀ and 2 ♂ from Vancouver Island, North Shore Road, 3.2 kilometers south of Youbou, 4 Jul 1976, by IM Smith, IMS 760190 * 3 ♂ from Vancouver Island, Youbou, Shaw Creek, North Shore Road, 4.3 kilometers south of north end of Cowichan Lake, 8 Jul 1976, by IM Smith, IMS 760196 * 1 ♀ from Vancouver Island, spring-fed pool beside South Shore Road, 2.3 kilometers south of Caycuse, 26 Jul 1979, by IM Smith, IMS 790052 * 1 ♀ from Vancouver Island, spring run beside South Shore Road, 2.3 kilometers north of Lake Cowichan, 6 Jun 1979, by IM Smith, IMS 790007 * California, USA: 1 ♀ and 2 ♂ from Del Norte County, Six Rivers National Forest, Middle Fork Smith River (41 ° 51 ' 20 " N, 123 ° 53 ' 10 " W), 15 Aug 2013, by JR Fisher, JRF 13 - 0815 - 002 * 1 ♀ and 2 ♂ from Mendocino County, Cottaneva Creek, beside Route 1, 21.8 kilometers southwest of Route 101, 5 Aug 1987, by IM Smith, IMS 870129 A * 4 ♀ and 4 ♂ from Mendocino County, small stream at beach access road, off Route 1, 2.6 kilometers south of Westport, 5 Aug 1987, by IM Smith, IMS 870128 A * 1 ♀ and 1 ♂ from Mono County, Humboldt-Toiyabe National Forest, Leavitt Creek (38 ° 18 ' 40 " N, 119 ° 34 ' 49 " W), 31 Aug 2013, by JR Fisher, JRF 13 - 0831 - 004 * 1 ♀ and 1 ♂ from Mono County, Humboldt-Toiyabe National Forest, West Walker River (38 ° 21 ' 59 " N, 119 ° 28 ' 55 " W), 31 Aug 2013, by JR Fisher, JRF 13 - 0831 - 003 * 1 ♀ from Monterey County, Los Padres National Forest, Salmon Creek (35 ° 48 ' 57 " N, 121 ° 21 ' 29 " W), 6 Sep 2013, by JR Fisher, JRF 13 - 0906 - 003 * 1 ♀ and 2 ♂ from Monterey County, Nacimiento River, beside Nacimiento-Ferguson Road at Nacimiento campground, 30 Jul 1987, by IM Smith, IMS 870120 A * 2 ♀ from Monterey County, Pfeiffer State Park, Big Sur River (36 ° 14 ' 42 " N, 121 ° 46 ' 43 " W), 4 Sep 2013, by JR Fisher, JRF 13 - 0904 - 004 * 2 ♀ and 2 ♂ from Monterey County, Salmon Creek, beside Route 1, south of Gorda, 28 Jul 1987, by IM Smith, IMS 870114 A * 1 ♀ and 1 ♂ from Nevada County, beside Route 89, north of Hobart Mills, 13 Jun 1976, by IM Smith, IMS 760109 * 1 ♀ and 1 ♂ from Nevada County, Tahoe National Forest, Sagehen Creek (39 ° 26 ' 2 " N, 120 ° 12 ' 17 " W), 26 Aug 2013, by JR Fisher, JRF 13 - 0826 - 006 * 1 ♀ from Plumas County, beside Route 89, north of Greenville, 14 Jun 1976, by IM Smith, IMS 760113 * 1 ♀ from Siskiyou County, Klamath National Forest, Dead Cow Creek (41 ° 57 ' 16 " N, 122 ° 52 ' 23 " W), 16 Aug 2013, by JR Fisher, JRF 13 - 0816 - 002 * 1 ♀ and 1 ♂ from Tulare County, Stony Creek at Stony Creek Picnic Area, east of Sequoia National Park, 1 Aug 1987, by IM Smith, IMS 870124 A * 1 ♀ from Ventura County, Ojai, North Fork of Ventura River, beside Route 33, just above Wheeler Gorge, 25 - 26 Jul 1987, by IM Smith, IMS 870109 A * Idaho, USA: 1 ♀ from Blaine County, Salmon River, beside Route 75 between Obsidian & Galena Summit, 3 Jul 1985, by IM Smith, IMS 850067 * 1 ♂ from Blaine County, Sawtooth National Forest, Deer River (43 ° 32 ' 49 " N, 114 ° 26 ' 31 " W), 29 Jul 2012, by JR Fisher, WA Nelson, & JC O'Neill, ROW 12 - 0729 - 001 * 1 ♀ and 1 ♂ from Custer County, Basin Creek campground, Basin Creek, beside Route 75 between Sunbeam & Stanley, 2 Jul 1985, by IM Smith, IMS 850066 * 1 ♀ from Custer County, Salmon River at picnic area, beside Route 93 north of Morgan Creek, 2 Jul 1985, by IM Smith, IMS 850064 * 1 ♀ and 1 ♂ from Lemhi County, North Fork of Salmon River, beside Route 93, 15 kilometers north of North Fork, 1 Jul 1985, by IM Smith, IMS 850062 * 1 ♀ from Lemhi County, Salmon National Forest, Indian Creek (45 ° 24 ' 12 " N, 114 ° 10 ' 10 " W), 2 Aug 2012, by JR Fisher, WA Nelson, & JC O'Neill, ROW 12 - 0802 - 001 * Nevada, USA: 2 ♀ and 2 ♂ from Elko County, Lamoille, Lamoille Creek near car park, at end of Lamoille Canyon Road, 12 Aug 1987, by IM Smith, IMS 870143 A * 2 ♀ and 3 ♂ from Humboldt County, Paradise Valley, Dutch John Creek beside road, 8.3 kilometers north of Hinkey Summit, 11 Aug 1987, by IM Smith, IMS 870142 A * New Mexico, USA: 2 ♀ and 2 ♂ from Catron County, Whitewater Creek, Glenwood Whitewater Picnic Area, 5 May 2012, by IM Smith, IMS 120005 * 1 ♀ and 1 ♂ from Catron County, Glenwood Whitewater Creek at Whitewater Creek Picnic Area, 12 Jul 1987, by IM Smith, IMS 870084 * 1 ♀ and 1 ♂ from Lincoln County, Eagle Creek, beside Route 532, just above Route 37, 8 Jul 1987, by IM Smith, IMS 870077 * 1 ♀ and 1 ♂ from San Miguel County, Pecos, Pecos River, beside Route 63 at Dalton campground, 6 Jul 1987, by IM Smith, IMS 870071 * 1 ♀ and 1 ♂ from San Miguel County, stream near Route 63 in Dalton Canyon, 6 Jul 1987, by IM Smith, IMS 870073 * 1 ♀ from Sante Fe County, Sante Fe, Tesuque Creek, beside Route 22, south of Tesuque, 5 Jul 1987, by IM Smith, IMS 870070 * Montana, USA: 1 ♀ from Lake County, stream beside Route 83, 39.5 kilometers north of Condon, 30 Jun 1985, by IM Smith, IMS 850059 A * 1 ♀ from Ravalli County, Beaverhead National Forest, East Fork Bitterroot River (45 ° 51 ' 40 " N, 114 ° 1 ' 46 " W), 3 Aug 2012, by JR Fisher, WA Nelson, & JC O'Neill, ROW 12 - 0803 - 004 * 1 ♂ from Ravalli County, Bitterroot National Forest, Piquette Creek (45 ° 51 ' 24 " N, 114 ° 11 ' 37 " W), 6 Aug 2012, by JR Fisher, WA Nelson, & JC O'Neill, ROW 12 - 0806 - 002 * 1 ♀ and 2 ♂ from Ravalli County, Medicine Springs, Spring Gulch campground, East Fork of Bitterroot River, beside Route 93, 1 Jul 1985, by IM Smith, IMS 850060 * Oregon, USA: 1 ♀ from Clackamas County, Rhododendron Pioneer Tollgate campground, Zigzag River, 27 Jun 1976, by IM Smith, IMS 760164 * 1 ♀ and 2 ♂ from Coos County, Siskiyou National Forest, Coal Creek, Road 33 between Powers & Agness, 2 Jul 1983, by IM Smith, IMS 830015 * 2 ♀ and 2 ♂ from Coos County, Siskiyou National Forest, Road 33 between Powers & Agness, Daphne Grove campground, 2 Jul 1983, by IM Smith, IMS 830017 * 1 ♀ and 1 ♂ from Curry County, Port Orford, Butler Bar campground, Elk River, 25 Jun 1976, by IM Smith, IMS 760162 * 1 ♀ and 6 ♂ from Curry County, Port Orford, Humbug Mountain State Park Picnic Area, Brush Creek, beside Route 1, 3 Jul 1983, by IM Smith, IMS 830020 A & IMS 830020 B * 2 ♀ and 1 ♂ from Curry County, Port Orford, Humbug Mountain State Park Picnic Area, beside Route 1, Brush Creek, 1 Jul 1983, by IM Smith, IMS 830012 * 1 ♀ from Curry County, Rogue River National Forest, Elk River (42 ° 42 ' 46 " N, 124 ° 18 ' 41 " W), 13 Aug 2013, by JR Fisher, JRF 13 - 0813 - 003 * 1 ♀ and 1 ♂ from Curry County, Siskiyou National Forest, road 33 between Powers and Agness, North Fork of Foster Creek, 2 Jul 1983, by IM Smith, IMS 830019 * 2 ♀ and 1 ♂ from Curry County, Sixes, Sixes River, beside road at mouth of Edson Creek, 4 Jul 1983, by IM Smith, IMS 830021 A * 1 ♀ and 1 ♂ from Multnomah County, Columbia River Scenic Highway, Horsetail Falls, 27 Jun 1983, by IM Smith, IMS 830005 * Utah, USA: 2 ♀ and 1 ♂ from Summit County, North Fork of Provo River, beside Route 150, west of Provo River Overlook, 14 Aug 1987, by IM Smith, IMS 870148 * 1 ♂ from Utah County, Uinta National Forest, Hobble Creek Road, upstream on Right Fork (40 ° 10 ' 9 " N, 111 ° 28 ' 36 " W), 22 Jul 2012, by JR Fisher, WA Nelson, & JC O'Neill, ROW 12 - 0722 - 001 * 1 ♀ and 1 ♂ from Wasatch County, Provo River, beside Route 150 at Upper Provo River Bridge Picnic Area, 14 Aug 1987, by IM Smith, IMS 870149 * 2 ♀ and 1 ♂ from Wasatch County, Wasatch-Cache National Forest, Holden Fork (40 ° 47 ' 19 " N, 110 ° 53 ' 2 " W), 23 Jul 2012, by JR Fisher, WA Nelson, & JC O'Neill, ROW 12 - 0723 - 003 * Washington, USA: 2 ♀ from Jefferson County, Rocky Brook (47 ° 43 ' 11 " N, 122 ° 56 ' 32 " W), 22 Jul 2013, by JC O'Neill & WA Nelson, JNOW 13 - 0722 - 002 * 1 ♀ and 1 ♂ from Lewis County, Gifford Pinchot National Forest, Snake Creek (46 ° 38 ' 52 " N, 121 ° 43 ' 8 " W), 23 Jul 2013, by JC O'Neill, & WA Nelson, JNOW 13 - 0723 - 006 * Wyoming, USA: 2 ♀ and 1 ♂ from Carbon County, Medicine Bow Mountains, Medicine Bow River, west of Arlington, 30 Jul 2012, by IM Smith, IMS 120045 * 1 ♀ and 1 ♂ from Fremont County, Wind River Mountains, Sinks Canyon, Popo Agie River, 1 Aug 2012, by IM Smith, IMS 120049.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
FB959FB0C31D58B9BF53CE481BAC972E.taxon	diagnosis	Diagnosis. Torrenticola multiforma are similar to other members of the Ellipsoidalis Group (T. ellipsoidalis, T. occidentalis, and T. leviathan), in being among the largest Torrenticola in the west (dorsum length ♀ = 700 - 885; ♂ = 665 - 850), although T. sierrensis are also large (dorsum length ♀ = 700 - 880; ♂ = 590 - 735) but can easily be distinguished from the Ellipsoidalis Group by being circular instead of ellipsoid or rectangular (dorsum length / width = 1.17 - 1.28 in T. sierrensis, 1.30 - 1.67 in Ellipsoidalis Group). T. multiforma are best differentiated from other members of the Ellipsoidalis Group by having more elongate subcapitular rostra (length / width = 2.5 - 2.8 in T. multiforma, 1.84 - 2.27 in other Ellipsoidalis Group).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
FB959FB0C31D58B9BF53CE481BAC972E.taxon	description	Re-description. Female (Figure 159) (n = 6) with characters of the genus with following specifications. Dorsum - (700 - 850 (730) long; 500 - 615 (550) wide) ellipsoid with highly variable coloration, colorless to orange to purple without distinct pattern. Anterio-medial platelets (132.5 - 150 (145) long; 65 - 77.5 (65) wide). Anterio-lateral platelets (195 - 230 (195) long; 77.5 - 92.5 (82.5) wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 380 - 465 (400 )). Dorsal plate proportions: dorsum length / width 1.31 - 1.41 (1.33); dorsal width / distance between Dgl- 4 1.26 - 1.38 (1.38); anterio-medial platelet length / width 1.94 - 2.27 (2.23); anterio-lateral platelet length / width 2.34 - 2.87 (2.36); anterio-lateral / anterio-medial length 1.34 - 1.57 (1.34). Gnathosoma - Subcapitulum (340 - 385 long (ventral); 255 - 284 long (dorsal); 145 - 172.5 tall) colorless. Rostrum (132.5 - 152.5 long; 50 - 55 wide). Chelicerae (336 - 395 long) with curved fangs (62 - 79 long). Subcapitular proportions: ventral length / height 2.19 - 2.46; rostrum length / width 2.5 - 2.8. Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (37.5 - 45 long); femur (115 - 127.5 long); genu (75 - 90 long); tibia (87.5 - 95 long; 27.5 - 32.5 wide); tarsus (17.5 - 20 long). Palpomere proportions: femur / genu 1.42 - 1.53; tibia / femur 0.71 - 0.78; tibia length / width 2.77 - 3.18. Venter - (840 - 1010 long; 588 - 679 wide) colorless. Gnathosomal bay (165 - 217.5 long; 82.5 - 100 wide). Cxgl- 4 subapical. Medial suture (17.5 - 32.5 long). Genital plates (210 - 242.5 long; 197.5 - 222.5 wide). Additional measurements: Cx- 1 (295 - 380 long (total); 123 - 170 long (medial )); Cx- 3 (359 - 441 wide); anterior venter (175 - 212.5 long). Ventral proportions: gnathosomal bay length / width 1.89 - 2.44; anterior venter / genital field length 0.79 - 0.92; anterior venter length / genital field width 0.86 - 1.05; anterior venter / medial suture 5.83 - 11.14. Male (Figure 160) (n = 6) with characters of the genus with following specifications. Dorsum - (665 - 790 long; 470 - 580 wide) ellipsoid with highly variable coloration, colorless to orange to purple without distinct pattern. Anterio-medial platelets (125 - 165 long; 60 - 77.5 wide). Anterio-lateral platelets (185 - 222.5 long; 75 - 87.5 wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 365 - 450). Dorsal plate proportions: dorsum length / width 1.30 - 1.41; dorsal width / distance between Dgl- 4 1.27 - 1.38; anterio-medial platelet length / width 1.97 - 2.33; anterio-lateral platelet length / width 2.46 - 2.67; anterio-lateral / anterio-medial length 1.35 - 1.48. Gnathosoma - Subcapitulum (310 - 380 long (ventral); 236 - 284) long (dorsal); 137.5 - 170 tall) colorless. Rostrum (120 - 150 long; 45 - 60 wide). Chelicerae (309 - 382.5 long) with curved fangs (63 - 75 long). Subcapitular proportions: ventral length / height 2.16 - 2.34; rostrum length / width 2.50 - 2.79. Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (40 - 45 long); femur (102.5 - 122.5 long); genu (70 - 87.5 long); tibia (80 - 95 long; 26.25 - 32.5 wide); tarsus (17.5 - 20 long). Palpomere proportions: femur / genu 1.40 - 1.50; tibia / femur 0.73 - 0.79; tibia length / width 2.91 - 3.33. Venter - (805 - 940 long; 479 - 653 wide) colorless. Gnathosomal bay (172.5 - 197.5 long; 75 - 100 wide). Cxgl- 4 subapical. Medial suture (62.5 - 82.5 long). Genital plates (175 - 190 long; 140 - 155 wide). Additional measurements: Cx- 1 (308 - 375 long (total); 127 - 180 long (medial )); Cx- 3 (351 - 421 wide); anterior venter (225 - 270 long). Ventral proportions: gnathosomal bay length / width 1.92 - 2.30; anterior venter / genital field length 1.29 - 1.46; anterior venter length / genital field width 1.57 - 1.81; anterior venter / medial suture 3.27 - 4.00. Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
FB959FB0C31D58B9BF53CE481BAC972E.taxon	etymology	Etymology. Habeeb (1974) did not specify an etymology for the specific epithet (multiforma). However, he notes that the males " vary a lot in size. " Thus, we suspect the specific epithet refers to the variable body size males of this species (multus, L. many; forma, L. form).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
FB959FB0C31D58B9BF53CE481BAC972E.taxon	distribution	Distribution. Western (Figure 158). T. multiforma was previously known only from Prairie Creek, California (Habeeb 1974), but we expand its range to include most of western North America.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
C0774CE07C49A82863D9C6F6621DFA2D.taxon	materials_examined	Material examined. HOLOTYPE (♂): from Canada, New Brunswick, Victoria County, Salmon River, 21 Jun 1953, by H Habeeb, HH 530075. PARATYPES (1 ♀; 0 ♂): New Brunswick, Canada: 1 ♀ from Victoria County, Salmon River, 21 Jun 1953, by H Habeeb, HH 530075. OTHER MATERIAL (27 ♀; 26 ♂; 1 nymph): Arkansas, USA: 1 nymph from Montgomery County, Caddo Gap, access track off Manfred Road, 0.3 km west of Route 8, 29 Jul 2011, by IM Smith, IMS 110037 * 1 ♀ from Newton County, Buffalo National River, Mill Creek (36 ° 3 ' 42.12 " N, 93 ° 8 ' 7.62 " W), 30 May 2012, by TD Edwards, TDE 12 - 0530 - 004 * Kentucky, USA: 1 ♂ from McCreary County, White Oak Junction, Rock Creek, beside Forest Route 556, 2.3 kilometers south of Route 1363, 8 Jul 1990, by IM Smith, IMS 900082 A * Maine, USA: 1 ♀ and 1 ♂ from Aroostook County, Ashland, beside Route 11 at bridge, Aroostook River (46 ° 38 ' N, 68 ° 24 ' W), 4 Jul 1989, by IM Smith, IMS 890067 * 2 ♀ and 1 ♂ from Piscataquis County, Baxter State Park, Trout Brook, beside road, 25 Jul 1980, by IM Smith, IMS 800125 A * 1 ♂ from Washington County, Old Stream, off Route 9, 5.5 km west of Route 192 at Wesley, 6 Jun 2012, by IM Smith, IMS 120012 * Missouri, USA: 1 ♀ from Crawford County, Huzzah Creek, Red Bluff campground, east of Davisville, 23 Jul 2011, by IM Smith, IMS 110029 * 1 ♂ from McDonald County, Tiff City, beside Route 43, Buffalo Creek (36 ° 40 ' 17 " N, 94 ° 36 ' 17 " W), 2 May 1996, by IM Smith, IMS 960004 * New Brunswick, Canada: 2 ♀ and 6 ♂ from Charlotte County, Rollingham, Digdeguash River, beside Highway 770 at covered bridge, 30 Jun 1989, by IM Smith, IMS 890053 * 3 ♀ from Charlotte County, Rollingham, Digdegaush River, beside Highway 770, 3 Oct 2011, by IM Smith, IMS 110118 * 1 ♀ and 2 ♂ from York County, Magaguadavic River, beside Highway 3 just east of Thomaston Corners, 1 Jul 1989, by IM Smith, IMS 890055 A * 1 ♀ and 1 ♂ from York County, Nashwaak River, beside Highway 8, 1.7 kilometers north of Durham Bridge Road, 2 Jul 1989, by IM Smith, IMS 890058 * New York, USA: 1 ♀ and 2 ♂ from Essex County, Minerva Boreas River, beside Route 28 N, 13.8 kilometers northwest of Morse Memorial Parkway, 21 Jun 1990, by IM Smith, IMS 900050 A * 3 ♀ and 2 ♂ from Schuyler County, beside Town Line Road off Route 228, 0.6 kilometers south of Perry City, 21 July 1990, by IM Smith, IMS 900112 A * Nova Scotia, Canada: 1 ♀ and 1 ♂ from Antigonish County, Antigonish, West River, 1 Jul 1981, IMS 810050 * 1 ♂ from Guysborough County, Sherbrooke, St. Mary's River, 17 Sep 2011, by IM Smith, IMS 110087 * 1 ♀ and 1 ♂ from Victoria County, Cape Brenton Island, Baddeck River, beside road to Baddeck Forks, 18 Jul 1981, by IM Smith, IMS 810082 * Ontario, Canada: 3 ♀ and 3 ♂ from Grey County, Durham, Saugeen River, beside County Road 27 near Durham Conservation Area, 9 Jun 1989, by IM Smith, IMS 890028 A * Pennsylvania, USA: 1 ♀ from Fayette County, Ohiopyle State Park, Laurel Run (39 ° 50 ' 58 " N, 79 ° 30 ' 51 " W), 10 Aug 2014, by MJ Skvarla, MS 14 - 0810 - 005 * Tennessee, USA: 1 ♀ from Sevier County, Great Smokey Mountians National Park, Little River (35 ° 40 ' 56 " N, 83 ° 39 ' 2 " W), 8 Sep 2009, by IM Smith, IMS 090103 * 2 ♀ from Monroe County, Tellico River, beside Forest Route 210, 1.8 kilometers east of bridge at Bald River Falls, 5 Jul 1990, by IM Smith, IMS 900079 * Texas, USA: 1 ♂ from Shackelford County, Albany, beside Route 180, 4.6 kilometers east of Route 283 (32 ° 44 ' 29 " N, 99 ° 14 ' 17 " W), 3 May 1996, by IM Smith, IMS 960007 * Virginia, USA: 1 ♀ from Alleghany County, Longdale Furnace, Simpson Creek, beside Forest Road 108, 1.7 kilometers west of Route 850, 14 Jul 1990, by IM Smith, IMS 900094 * West Virginia, USA: 1 ♀ and 1 ♂ from Pendleton County, North Fork South Branch Potomac River, beside Route 28 / 55, 20.8 kilometers southwest of Route 42, 17 Jul 1990, by IM Smith, IMS 900104.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
C0774CE07C49A82863D9C6F6621DFA2D.taxon	diagnosis	Diagnosis. Torrenticola neoanomala are similar to species with similar dorsal patterning, such as the Rusetria " 4 - Plate " group (T. dunni, T. glomerabilis, T. kittatinniana, T. pollani, T. rufoalba, and T. shubini), Elongata Group (T. elongata, T. gorti, and T. reduncarostra) and T. interiorensis, T. bondi, T. erectirostra, T. robisoni, T. irapalpa, T. racupalpa, T. skvarlai, and T. arktonyx. They can be differentiated from Rusetria 4 - Plates and T. skvarlai by having distinct hind coxal margins. T. neoanomala can be differentiated from T. erectirostra and T. robisoni by having a straight, anteriorly-directed rostrum (upturned in T. erectirostra and T. robisoni). T. neoanomala can be differentiated from T. arktonyx by having an unmodified dorsal plate (T. arktonyx has distinctive longitudinal dark markings on the anterior portion of the dorsal plate that fade posteriorly). T. neoanomala can be differentiated from T. racupalpa and T. irapalpa by having more elongate anterio-lateral platelets (length / width = 2.79 - 3.23 in T. neoanomala, 2.17 - 2.67 in others) and Dgl- 4 closer to the dorsal edge (dorsal width / distance between Dgl- 4 = 1.37 - 1.5 in T. neoanomala, 1.58 - 2.77 in others). T. neoanomala can be differentiated from Elongata Group by being slightly more ovoid (dorsum length / width ♀ = 1.35 - 1.43 in T. neoanomala, 1.45 - 2.08 in Elongata Group; ♂ = 1.43 - 1.50 in T. neoanomala, 1.51 - 1.7 in Elongata Group) and having a stockier rostrum (length / width = 2.59 - 2.90 in T. neoanomala, 3.24 - 4.00 in Elongata Group). T. neoanomala can be differentiated from T. bondi by having a longer medial suture (♀ = 22 - 40 in T. neoanomala, 10 - 15 in T. bondi; ♂ = 95 - 108 in T. neoanomala, 55 - 70 in T. bondi) and by anterior venter / genital field width (♀ = 1.39 - 1.45 in T. neoanomala, 1.15 - 1.25 in T. bondi; ♂ = 2.42 - 2.66 in T. neoanomala, 1.95 - 2.05 in T. bondi). Female T. neoanomala can be differentiated from female T. interiorensis by having more elongate anterio-lateral platelets (length / width = 2.86 - 3.09 in T. neoanomala, 2.62 - 2.67 in T. interiorensis). Male T. neoanomala can be differentiated from male T. interiorensis by having a longer anterior venter (267.5 - 290 in T. neoanomala, 220 - 240 in T. interiorensis) and a longer genital field (145 - 160 in T. neoanomala, 132 - 138 in T. interiorensis).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
C0774CE07C49A82863D9C6F6621DFA2D.taxon	description	Re-description. Female (Figure 162) (n = 6) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (590 - 680 (650) long; 420 - 480 (480) wide) ovoid with bluish-purple to purple coloration separated into anterior and posterior portions with faint orange medially. Anterio-medial platelets (135 - 155 (140) long; 52.5 - 65 (60) wide). Anterio-lateral platelets (177.5 - 205 (205) long; 57.5 - 70 (67.5) wide) free from dorsal plate. Dgl- 4 closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 280 - 345 (345 )). Dorsal plate proportions: dorsum length / width 1.35 - 1.43 (1.35); dorsal width / distance between Dgl- 4 1.39 - 1.50 (1.39); anterio-medial platelet length / width 2.23 - 2.58 (2.33); anterio-lateral platelet length / width 2.86 - 3.09 (3.04); anterio-lateral / anterio-medial length 1.29 - 1.46 (1.46). Gnathosoma - Subcapitulum (320 - 350 long (ventral); 235 - 259 long (dorsal); 130 - 152.5 tall) colorless. Rostrum (120 - 142.5 long; 45 - 55 wide). Chelicerae (315 - 358 long) with curved fangs (56 - 61 long). Subcapitular proportions: ventral length / height 2.30 - 2.46; rostrum length / width 2.59 - 2.89. Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (40 - 47.5 (47.5) long); femur (110 - 128.75 (123.75) long); genu (67.5 - 75 (70) long); tibia (77.5 - 90 (90) long; 21.25 - 25 (25) wide); tarsus (16.25 - 20 (20) long). Palpomere proportions: femur / genu 1.63 - 1.81 (1.77); tibia / femur 0.67 - 0.74 (0.73); tibia length / width 3.58 - 3.68 (3.60). Venter - (680 - 820 (770) long; 480 - 570 (570) wide) mostly colorless with faint bluish-purple or purple in areas surrounding coxae. Gnathosomal bay (147.5 - 177.5 (160) long; 87.5 - 107.5 (107.5) wide). Cxgl- 4 subapical. Medial suture (22.5 - 40 (40) long). Genital plates (157.5 - 187.5 (187.5) long; 140 - 155 (145) wide). Additional measurements: Cx- 1 (300 - 334 (325) long (total); 144 - 169 (162.5) long (medial )); Cx- 3 (336 - 412.5 (412.5) wide); anterior venter (195 - 225 (207.5) long). Ventral proportions: gnathosomal bay length / width 1.49 - 1.78 (1.49); anterior venter / genital field length 1.11 - 1.27 (1.11); anterior venter length / genital field width 1.39 - 1.45 (1.43); anterior venter / medial suture 5.19 - 9.44 (5.19). Male (Figure 163) (n = 5) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (545 - 635 (555) long; 370 - 430 (370) wide) ovoid with bluish-purple to purple coloration separated into anterior and posterior portions with faint orange medially. Anterio-medial platelets (120 - 130 (120) long; 48.75 - 62.5 (48.75) wide). Anterio-lateral platelets (167.5 - 197.5 (167.5) long; 55 - 62.5 (60) wide) free from dorsal plate. Dgl- 4 closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 255 - 315 (265 )). Dorsal plate proportions: dorsum length / width 1.43 - 1.50 (1.50); dorsal width / distance between Dgl- 4 1.37 - 1.49 (1.40); anterio-medial platelet length / width 2.08 - 2.46 (2.46); anterio-lateral platelet length / width 2.79 - 3.23 (2.79); anterio-lateral / anterio-medial length 1.40 - 1.52 (1.40). Gnathosoma - Subcapitulum (272.5 - 315 (272.5) long (ventral); 207.5 - 235 (207.5) long (dorsal); 97.5 - 122.5 (97.5) tall) colorless. Rostrum (112.5 - 127.5 (112.5) long; 38.75 - 45 (38.75) wide). Chelicerae (262 - 307 (267.5) long) with curved fangs (48 - 58 (55) long). Subcapitular proportions: ventral length / height 2.57 - 2.79 (2.79); rostrum length / width 2.67 - 2.90 (2.90). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (40 - 42.5 (40) long); femur (100 - 115 (100) long); genu (60 - 70 (60) long); tibia (78.75 - 87.5 (78.75) long; 20 - 26.25 (20) wide); tarsus (15 - 20 (15) long). Palpomere proportions: femur / genu 1.60 - 1.69 (1.67); tibia / femur 0.74 - 0.80 (0.79); tibia length / width 3.33 - 3.94 (3.94). Venter - (675 - 770 (675) long; 420 - 515 (470) wide) mostly colorless with faint bluish-purple or purple in areas surrounding coxae. Gnathosomal bay (127.5 - 152.5 (127.5) long; 68.75 - 90 (73.75) wide). Cxgl- 4 subapical. Medial suture (95 - 107.5 (102.5) long). Genital plates (145 - 160 (150) long; 102.5 - 120 (102.5) wide). Additional measurements: Cx- 1 (270 - 323 (270) long (total); 142 - 169 (152.5) long (medial )); Cx- 3 (322 - 384 (340) wide); anterior venter (267.5 - 290 (272.5) long). Ventral proportions: gnathosomal bay length / width 1.69 - 2.15 (1.73); anterior venter / genital field length 1.79 - 1.90 (1.82); anterior venter length / genital field width 2.42 - 2.66 (2.66); anterior venter / medial suture 2.60 - 2.83 (2.66). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
C0774CE07C49A82863D9C6F6621DFA2D.taxon	etymology	Etymology. Habeeb (1957) did not specify an etymology for the specific epithet (neoanomala). In fact, Habeeb (1955) initially identified the specimens as the Palaearctic T. anomala, but later (1957) writes, " The mite I reported as Torrenticola anomala (Koch) is now seen to be distinct from this European form. " So, this name surely refers to the similarity of this species to the T. anomala (Koch, 1837) (neos, G. new).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
C0774CE07C49A82863D9C6F6621DFA2D.taxon	distribution	Distribution. T. neoanomala was previously known only from Albert County, New Brunswick, but we extend its range throughout eastern North America (Figure 161). Records from Shackelford County, Texas, make T. neoanomala one of the few eastern Torrenticola that are found in the Great Plains.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
4D8AB9D466E67C3DD142A824028A0B57.taxon	materials_examined	Material examined. HOLOTYPE (♀): from Canada, New Brunswick, Victoria County, Salmon River, 21 Jun 1953, by H Habeeb, HH 530075. PARATYPES (0 ♀; 1 ♂): New Brunswick, Canada: 1 ♂ (ALLOTYPE) from Victoria County, Salmon River, 21 Jun 1953, by H Habeeb, HH 530075. OTHER MATERIAL (83 ♀; 80 ♂): Alabama, USA: 2 ♀ and 1 ♂ from Clay County, beside Forest Route 649, 0.8 kilometers northeast of road from Forest Route 600 to Campbell Springs, Talladega Creek, 2 Jul 1990, by IM Smith, IMS 900075 A * 1 ♂ from Cleburne County, beside Route 431, 3.3 kilometers southeast of Calhoun, Jackson Creek (33 ° 36 ' N, 85 ° 42 ' W), 2 Jul 1990, by IM Smith, IMS 900074 * 1 ♀ and 1 ♂ from DeKalb County, Desoto State Park, beside Trail Y, West Fork of Little River (34 ° 29 ' N, 85 ° 32 ' W), by IM Smith, IMS 920053 A * Georgia, USA: 1 ♂ from Chattooga County, Cloudland, beside Route 48, East Fork Little River (34 ° 31 ' 25 " N, 85 ° 30 ' 23 " W), 29 Sep 2005, by IM Smith, IMS 050110 A * 1 ♂ from Floyd County, beside road from Everett Springs to Villanow, 1.8 kilometers south of The Pocket campground, Johns Creek, 4 Jul 1990, by IM Smith, IMS 900076 * Kentucky, USA: 2 ♀ and 2 ♂ from Bell County, Middlesboro, Sugar Run (36 ° 38 ' N, 83 ° 39 ' W), 9 Jul 1990, by IM Smith, IMS 900084 * Maine, USA: 1 ♀ and 1 ♂ from Aroostook County, Ashland, beside Route 11 at bridge, Aroostook River (46 ° 38 ' N, 68 ° 24 ' W), 4 Jul 1989, by IM Smith, IMS 890067 * 1 ♂ from Aroostook County, beside Route 2 A, 7 miles northeast of Macwahoc, Wytopitlock Stream, 5 Sep 1968, by DR Cook, DRC 680063 * 1 ♀ and 1 ♂ from Aroostook County, Monticello, North Branch Meduxnekeag River, 28 Aug 1964, by DR Cook, DRC 640042 * 1 ♀ and 1 ♂ from Franklin County, smalls Falls Picnic Area, beside Route 4, Sandy River (44 ° 52 ' N, 70 ° 31 ' W), 5 Jul 1989, by IM Smith, IMS 890069 * 1 ♀ and 1 ♂ from Piscataquis County, Baxter State Park, Trout Brook, 25 Jul 1980, by IM Smith, IMS 800125 A * 4 ♀ from Washington County, Old Stream, off Route 9, 5.5 km west of Route 192, 6 Jun 2012, by IM Smith, IMS 120012 * Michigan, USA: 1 ♀ and 1 ♂ from Cheboygan County, Pigeon River, 23 Jun 1952, by DR Cook, DRC 520023 * New Brunswick, Canada: 1 ♀ and 2 ♂ from Charlotte County, Rollingham, Digdegaush River, beside Highway 770, 30 Jun 1989, by IM Smith, IMS 890053 * 8 ♀ and 2 ♂ from Charlotte County, Rollingham, Digdegaush River, beside Highway 770, 3 Oct 2011, by IM Smith, IMS 110118 * 2 ♀ and 2 ♂ from Northumberland County, beside Highway 108, Renous River, 18 Jul 1980, by IM Smith, IMS 800111 * 1 ♀ and 1 ♂ from Restigouche County, Mount Carleton Provincial Park, Nictau River, 16 Jul 1980, by IM Smith, IMS 800109 * 2 ♂ from Victoria County, near Limestone, small stream, 26 Aug 1964, by DR Cook, DRC 640036 * 2 ♀ and 3 ♂ from York County, beside Highway 3, just east of Thomaston Corners, Magaguadavic River, 1 Jul 1989, by IM Smith, IMS 890055 A * 1 ♀ and 1 ♂ from York County, beside Highway 8, 1.7 kilometers north of Durham Bridge Road, Nashwaak River, 2 Jul 1989, by IM Smith, IMS 890058 * 1 ♀ and 1 ♂ from York County, beside Highway 107, 27.3 kilometers west of Road J- 19, North Branch of Southwest Miramichi River, 22 Jul 1981, by IM Smith, IMS 810092 B * 1 ♀ and 1 ♂ from York County, beside Road J- 19, 6.3 kilometers north of Nashwaak Experimental Watershed headquarters, Napadogan Brook, 23 Jul 1981, by IM Smith, IMS 810095 A * 1 ♀ and 1 ♂ from York County, Nashwaak Experimental Watershed, Narrow Mountain Brook, 21 Jul 1980, by IM Smith, IMS 800120 A * New Hampshire, USA: 1 ♀ and 1 ♂ from Coos County, White Mountain National Forest, Zealand Picnic Area, Ammonoosuc River, 29 Jul 1980, by IM Smith, IMS 800130 * New Jersey, USA: 1 ♀ and 1 ♂ from Sussex County, beside Flatbrook Road, 2.6 kilometers north of Route 206 at Tuttles Corner, Big Flat Brook, 23 Jun 1990, by IM Smith, IMS 900053 * New York, USA: 3 ♂ from Essex County, beside Route 28 N, 13.8 kilometers northwest of Morse Memorial Parkway at Minerva, Boreas River, 21 Jun 1990, by IM Smith, IMS 900050 A * North Carolina, USA: 3 ♀ and 1 ♂ from Haywood County, Great Smokey Mountains National Park, Big Creek (35 ° 45 ' 3.92 " N, 83 ° 6 ' 31.67 " W), 15 Sep 2009, by AJ Radwell, AJR 090008 A * 2 ♂ from Swain County, Oconaluftee River (35 ° 32 ' 34.76 " N, 83 ° 18 ' 14.13 " W), 6 Aug 2008, by AJ Radwell, AJR 080019 A * 1 ♂ from Yancey County, Lost Cove Picnic Area on Forest Route 472, 2.8 kilometers south of Route 80, South Toe River, 28 Jun 1990, by IM Smith, IMS 900065 A * Nova Scotia, Canada: 1 ♀ and 1 ♂ from Antigonish County, Antigonish, West River, 1 Jul 1981, by IM Smith, IMS 810050 * 2 ♀ and 2 ♂ from Cape Brenton Highlands National Park, beside Cabot Trail, south of Neils Harbour, Halfway Brook, 4 Jul 1981, by IM Smith, IMS 810058 * 5 ♀ and 3 ♂ from Cape Brenton Highlands National Park, picnic area above Mary Ann Falls, Mar Ann Brook, 5 Jul 1981, by IM Smith, IMS 810060 * 1 ♀ and 1 ♂ from Cape Brenton Island, Victoria County, Ingonish, beside Cabot Trail, Dundas Brook, 11 Jul 1981, by IM Smith, IMS 810074 * 2 ♀ from Inverness, Cheticamp River, 10 Sep 2011, by IM Smith, IMS 110071 * 1 ♂ from Luneburg County, New Germany, LaHave River, beside Highway 10, 23 Sep 2011, by IM Smith, IMS 110098 * 1 ♀ from Colchester County, Wentworth, picnic area beside Highway 104, tributary of Wallace River, 19 Jul 1981, by IM Smith, IMS 810085 * Ontario, Canada: 1 ♂ from Algoma District, Lake Superior Provincial Park, Agawa River, 28 Aug, 1965, by DR Cook, DRC 650001 * 2 ♀ and 2 ♂ from Grey County, Durham, beside County Road 27, near Durnham Conservation Area, Saugeen River, 9 Jun 1989, by IM Smith, IMS 890028 A * 2 ♀ and 1 ♂ from Hastings County, Madawaska, beside Highway 60, Opeongo River, 29 Aug 1981, by IM Smith, IMS 810033 A * 1 ♀ and 1 ♂ from Hastings County, Maple Leaf, Papineau Creek, east of Davis Road before Highway 62, 18 Aug 2011, by IM Smith, IMS 110053 * 2 ♀ and 1 ♂ from Lanark County, beside Lanark Road # 12, between Lanark and Fallbrook, Mississippi River, 6 October 1983, by IM Smith & CJ Hill, IMS 830094 A & IMS 830094 B * 1 ♀ and 1 ♂ from Muskoka District, Huntsville, East River Xing road to Dyer Memorial, 26 Aug 1981, by IM Smith, IMS 810032 A * 2 ♀ and 1 ♂ from Nipissing District, Algonquin Provincial Park, at Highway 60, near Lake of Two Rivers, Madawaska River, 15 May 1980, by IM Smith & CJ Hill, IMS 800004 B & IMS 800004 C * 1 ♀ and 1 ♂ from Peterborough County, crossing Highway 28 at picnic area just north of Woodview, Eels Creek, 13 Jun 1981, by IM Smith, IMS 810018 A * 2 ♀ and 2 ♂ from Thunder Bay District, Rossport, Rossport Provincial Campground, off Highway 17, McLeans Creek, 3 Aug 1988, by IM Smith, IMS 880065 * Pennsylvania, USA: 1 ♀ and 1 ♂ from Fayette County, Dunbar Creek (39 ° 56 ' 16 " N, 79 ° 35 ' 3.70 " W), 10 Aug 2014, by MJ Skvarla, MS 14 - 0810 - 002 * 1 ♀ from Fayette County, off Meadow Run Road, Laurel Run (39 ° 50 ' 58 " N, 79 ° 30 ' 51 " W), 10 Aug 2014, by MJ Skvarla, MS 14 - 0810 - 005 * Quebec, Canada: 1 ♂ from Stanstead County, Tompkin Stream, Tomifobia River (45 ° 0 ' 31 " N, 72 ° 7 ' 6 " W), 20 Aug 1996, by IM Smith & M MacKenzie, IMS 960056 * Tennessee, USA: 2 ♀ from Blount County, Great Smokey Mountains National Park, Cades Cove, Abrams River (35 ° 35 ' 31 " N, 83 ° 51 ' 21 " W), 17 Sep 2010, IMS 100141 * 2 ♀ and 6 ♂ from Monroe County, Turkey Creek at confluence with Tellico River, 12 Sep 2009, by AJ Radwell, AJR 090004 * 2 ♂ from Sevier County, Great Smokey Mountains National Park, middle prong Little Pigeon River (35 ° 43 ' 32 " N, 83 ° 24 ' 2 " W), 2 Sep 2009, by IM Smith, IMS 090093 * Vermont, USA: 2 ♀ from Addison County, beside road from Lincoln to Ripton, Middlebury River (44 ° 0 ' N, 73 ° 1 ' W), 6 Jul 1989, by IM Smith, IMS 890075 * 1 ♀ and 1 ♂ from Addison County, Lincoln, beside US Forest Service Road # 54, Haven River (44 ° 6 ' N, 72 ° 59 ' W), 6 Jul 1989, by IM Smith, IMS 890074 * Virginia, USA: 1 ♀ and 1 ♂ from Alleghany County, Covington, beside Route 18, 0.5 kilometers north of Route 657, Potts Creek, 13 Jul 1990, by IM Smith, IMS 900091 A * 1 ♀ from Amherst County, Upper Otter Creek Overlook beside Blue Ridge, Otter Creek (37 ° 36 ' 57 " N, 79 ° 19 ' 27 " W), 7 Sep 2007, by IM Smith, IMS 070056 A * 2 ♀ and 2 ♂ from Bath County, beside Route 287, 2.4 kilometers south of Bacova, Cowardin Run, 15 Jul 1990, by IM Smith, IMS 900097 * 1 ♀ and 1 ♂ from Bath County, beside Forest Route 1843, 3.5 kilometers south of Route 220, Jackson River, 16 Jul 1990, IMS 900100 * 1 ♀ from Giles County, Mechanicsburg, beside Dismal Creek Road, Standrock Brook (37 ° 11 ' 38 " N, 80 ° 53 ' 26 " W), 9 Sep 2005, by IM Smith, IMS 050066 * 3 ♂ from Highland County, 4.8 kilometers northwest of McDowell, Crab Run, 23 Jul 1964, by DR Cook, DRC 640013 * 1 ♀ from Montgomery County, Blacksburg, beside Route 321 at Caldwell, Craig Creek (37 ° 20 ' N, 80 ° 20 ' W), 12 Jul 1990, by IM Smith, IMS 900089 A * 2 ♀ and 2 ♂ from Page County, beside Route 730, 0.2 kilometers west of Route 675, Passage Creek, 25 Jun 1990, by IM Smith, IMS 900059 * 2 ♀ and 2 ♂ from Washington County, Damascus, beside Route 58 near boundary of Mount Rogers National Recreation Area, Laurel River, 10 Jul 1990, by IM Smith, IMS 900085 A * West Virginia, USA: 1 ♀ and 1 ♂ from Randolph County, Laurel Fork Campground, off Forest Route 14, south of Wymer, Laurel Fork of Cheat River, 17 Jul 1990, by IM Smith, IMS 900102.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
4D8AB9D466E67C3DD142A824028A0B57.taxon	diagnosis	Diagnosis. Torrenticola nigroalba are similar to other members of the Nigroalba Group (T. flangipalpa, T. solisorta, T. dentirostra) in being small, slightly elongate, having purple dorsal coloration restricted posteriorly, and having distinct yet poorly-defined hind coxal margins. T. nigroalba can be differentiated from T. flangipalpa in having tuberculate pedipalp femoral extension (flange-like and anteriorly-directed in T. flangipalpa); a shorter anterior venter (200 - 223 in T. nigroalba, 235 - 265 in T. flangipalpa); and more elongate pedipalpal tibiae (length / width ♀ = 5.38 - 5.83 in T. nigroalba, 4.79 - 5.00 in T. flangipalpa; ♂ = 5.08 - 5.33 in T. nigroalba, 4.40 - 4.86 in T. flangipalpa). T. nigroalba can be differentiated from T. dentirostra by having a smooth rostrum (T. dentirostra has a dentate bump midway on the dorsal edge of the rostrum) and by having more elongate pedipalpal tibiae (length / width ♀ = 5.3 - 5.9 in T. nigroalba, 4.5 - 5.0 in T. dentirostra; ♂ = 5.0 - 5.4 in T. nigroalba, 4.5 - 4.7 in T. dentirostra). T. nigroalba can be differentiated from T. solisorta by lacking orangish coloration immediately anterior to the purple dorsal coloration, although rarely specimens have been found with this coloration. Female T. nigroalba are also slightly larger (500 - 530 in T. nigroalba, 475 - 500 in T. solisorta); have a thinner gnathosomal bay (length / width = 1.25 - 1.55 in T. nigroalba, 1.3 - 1.5 in T. solisorta); and have a slightly thicker subcapitulum (3.00 - 3.14 in T. nigroalba, 3.14 - 3.30 in T. solisorta). Male T. nigroalba also have a longer medial suture with respect to the anterior venter (anterior venter / medial suture = 2.54 - 2.77 in T. nigroalba, 2.87 - 3.26 in T. solisorta) and a thinner dorsum (290 - 300 in T. nigroalba, 305 - 320 in T. solisorta). Other Torrenticola with purple dorsal coloration restricted posteriorly can be confused with T. nigroalba, such as T. tahoei and T. oregonensis. Both of these species are larger (dorsum length ♀ = 500 - 530 in T. nigroalba, 600 - 810 in others; ♂ = 440 - 455 in T. nigroalba, 560 - 820 in others) and distributed in the west (T. nigroalba is eastern).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
4D8AB9D466E67C3DD142A824028A0B57.taxon	description	Re-description. Female (Figure 165) (n = 6) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (500 - 530 (510) long; 330 - 365 (350) wide) ovoid with bluish-purple to purple coloration restricted posteriorly (rarely with faint orange at the anterior edge of the purple, especially medially). Anterio-medial platelets (105 - 112.5 (105) long; 40 - 47.5 (40) wide). Anterio-lateral platelets (145 - 160 (157.5) long; 46.25 - 52.5 (50) wide) free from dorsal plate. Dgl- 4 closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 230 - 260 (245 )). Dorsal plate proportions: dorsum length / width 1.42 - 1.53 (1.46); dorsal width / distance between Dgl- 4 1.33 - 1.46 (1.43); anterio-medial platelet length / width 2.32 - 2.65 (2.63); anterio-lateral platelet length / width 2.90 - 3.26 (3.15); anterio-lateral / anterio-medial length 1.29 - 1.51 (1.50). Gnathosoma - Subcapitulum (280 - 298 long (ventral); 197.5 - 220 long (dorsal); 92.5 - 97.5 tall) elongate and colorless. Rostrum (102.5 - 112.5 long; 35 - 40 wide). Chelicerae (273 - 287 long) with curved fangs (40 - 53 long). Subcapitular proportions: ventral length / height 3.03 - 3.14; rostrum length / width 2.81 - 3.10. Pedipalps elongate (especially tibiae) with tuberculate ventral extensions on femora and genua ending broadly and dentate. Palpomeres: trochanter (30 - 35 (32.5) long); femur (90 - 95 (95) long); genu (55 - 57.5 (57.5) long); tibia (85 - 92.5 (87.5) long; 15 - 16.25 (13.75) wide); tarsus (12.5 - 15 (12.5) long). Palpomere proportions: femur / genu 1.64 - 1.73 (1.65); tibia / femur 0.92 - 0.97 (0.92); tibia length / width 5.38 - 5.83 (5.38). Venter - (620 - 660 (640) long; 340 - 430 (430) wide) with faint bluish purple or purple coloration. Gnathosomal bay (110 - 120 (110) long; 72.5 - 82.5 (82.5) wide). Cxgl- 4 far from apex. Medial suture (50 - 55 (55) long). Genital plates (135 - 142.5 (137.5) long; 122.5 - 132.5 (123.75) wide). Additional measurements: Cx- 1 (243 - 250 (250) long (total); 98 - 140 (140) long (medial )); Cx- 3 (239 - 285 (285) wide); anterior venter (200 - 212.5 (203.75) long). Ventral proportions: gnathosomal bay length / width 1.33 - 1.55 (1.33); anterior venter / genital field length 1.42 - 1.53 (1.48); anterior venter length / genital field width 1.51 - 1.68 (1.65); anterior venter / medial suture 3.70 - 4.25 (3.70). Male (Figure 166) (n = 6) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (440 - 455 (455) long; 290 - 300 (295) wide) ovoid with bluish-purple to purple coloration restricted posteriorly. Anterio-medial platelets (92.5 - 100 (97.5) long; 35 - 42.5 (35) wide). Anterio-lateral platelets (137.5 - 150 (142.5) long; 40 - 47.5 (47.5) wide) free from dorsal plate. Dgl- 4 closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 205 - 225 (225 )). Dorsal plate proportions: dorsum length / width 1.50 - 1.55 (1.54); dorsal width / distance between Dgl- 4 1.31 - 1.41 (1.31); anterio-medial platelet length / width 2.26 - 2.86 (2.79); anterio-lateral platelet length / width 3.00 - 3.63 (3.00); anterio-lateral / anterio-medial length 1.38 - 1.58 (1.46). Gnathosoma - Subcapitulum (247.5 - 260 (250) long (ventral); 181 - 188.75 (188.75) long (dorsal); 77.5 - 82.5 (82.5) tall) elongate and colorless. Rostrum (92.5 - 97.5 (97.5) long; 32.5 - 35 (32.5) wide). Chelicerae (225 - 237.5 (237.5) long) with curved fangs (35 - 40 (40) long). Subcapitular proportions: ventral length / height 3.03 - 3.23 (3.03); rostrum length / width 2.79 - 3.00 (3.00). Pedipalps elongate (especially tibiae) with tuberculate ventral extensions on femora and genua ending broadly and dentate. Palpomeres: trochanter (25 - 30 (25) long); femur (77.5 - 82.5 (78.75) long); genu (50 - 55 (50) long); tibia (76.25 - 83.75 (80) long; 15 - 16.25 (15) wide); tarsus (15 - 17.5 (17.5) long). Palpomere proportions: femur / genu 1.50 - 1.65 (1.58); tibia / femur 0.95 - 1.05 (1.02); tibia length / width 5.08 - 5.33 (5.33). Venter - (550 - 570 (550) long; 313 - 380 (370) wide) with faint bluish purple or purple coloration. Gnathosomal bay (100 - 107.5 (102.5) long; 60 - 70 (67.5) wide). Cxgl- 4 far from apex. Medial suture (77.5 - 87.5 (80) long). Genital plates (105 - 118.75 (118.75) long; 85 - 92.5 (90) wide). Additional measurements: Cx- 1 (216 - 237 (235) long (total); 97 - 135 (127.5) long (medial )); Cx- 3 (230 - 263 (255) wide); anterior venter (212.5 - 222.5 (215) long). Ventral proportions: gnathosomal bay length / width 1.43 - 1.75 (1.52); anterior venter / genital field length 1.81 - 2.12 (1.81); anterior venter length / genital field width 2.32 - 2.51 (2.39); anterior venter / medial suture 2.54 - 2.77 (2.69). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
4D8AB9D466E67C3DD142A824028A0B57.taxon	etymology	Etymology. Habeeb (1955) did not specify an etymology for the specific epithet (nigroalba). However, surely this name refers to the dorsal coloration being half pigmented (purple) and half unpigmented (yellow), although Habeeb (1955) identified these colors as black and white (niger, L. black; albus, L. white).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
4D8AB9D466E67C3DD142A824028A0B57.taxon	distribution	Distribution. Northeastern and southward throughout the Appalachians (Figure 164). T. nigroalba was previously known only from Albert County, New Brunswick; we extend its distribution throughout the Appalachians and into Ottawa.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
EDEA8A9FEDF3BE55FE1C97F4B82C98C1.taxon	materials_examined	Material examined. HOLOTYPE (♀): from USA, California, Trinity County, Shasta-Trinity National Forest, Wilson Creek (40 ° 25 ' 17 " N, 123 ° 3 ' 5 " W), 20 Aug 2013, by JR Fisher, JRF 13 - 0820 - 003, DNA 3000. PARATYPES (9 ♀; 5 ♂): California, USA: 1 ♂ (ALLOTYPE) from USA, California, Trinity County, Shasta-Trinity National Forest, Wilson Creek (40 ° 25 ' 17 " N, 123 ° 3 ' 5 " W), 20 Aug 2013, by JR Fisher, JRF 13 - 0820 - 003, DNA 2996 * 5 ♀ and 2 ♂ from Plumas County, Plumas National Forest, Silver Creek (39 ° 56 ' 60 " N, 121 ° 2 ' 17 " W), 24 Aug 2013, by JR Fisher, JRF 13 - 0824 - 005 * 3 ♀ and 1 ♂ from Trinity County, Shasta-Trinity National Forest, Wilson Creek (40 ° 25 ' 17 " N, 123 ° 3 ' 5 " W), 20 Aug 2013, by JR Fisher, JRF 13 - 0820 - 003 * 1 ♀ and 1 ♂ from Tulare County, Sequoia National Forest, Brush Creek (35 ° 57 ' 57 " N, 118 ° 28 ' 43 " W), 3 Sep 2013, by JR Fisher, JRF 13 - 0903 - 002.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
EDEA8A9FEDF3BE55FE1C97F4B82C98C1.taxon	diagnosis	Diagnosis. Torrenticola nortoni are similar to other members of the Rusetria " Western 2 - Plates " group (T. mulleni, T. walteri, and T. welbourni) in having anterio-lateral platelets fused to the dorsal plate, having faint dorsal coloration (some are colorless), and being distributed in the west. Female T. nortoni can be differentiated from T. welbourni (female only known) by having stockier pedipalp tibiae (3.13 - 3.33 in A 33, 3.73 in A 30) and shorter pedipalp femora (112.5 - 122.5 in A 33, 137.5 in A 30). Female T. nortoni can be differentiated from female T. mulleni by having a shorter medial suture (10 - 12.5 in A 33; 20 - 22.5 in A 31) and a shorter genital field (177.5 - 192.5 in A 33, 195 - 205 in A 31). Male T. nortoni can be differentiated from male T. mulleni by having shorter pedipalp femora (85 - 93 in T. nortoni, 97 - 103 in T. mulleni). Additionally, T. nortoni can be differentiated from T. mulleni by being distributed in California, instead of in the Rocky Mountains (Idaho, Montana, Utah and Wyoming). Female T. nortoni can be differentiated from female T. walteri by having slightly longer pedipalp femora with respect to genua (1.69 - 1.82 in T. nortoni, 1.52 - 1.64 in T. walteri) and slightly more elongate anterio-medial platelets (2.74 - 3.06 in T. nortoni, 2.58 - 2.72 in T. walteri). Male T. nortoni can be differentiated from male T. walteri by having longer pedipalp femora (85 - 92.5 in T. nortoni, 95 - 100 in T. walteri) and slightly more elongate pedipalp tibiae (2.73 - 3.0 in T. nortoni, 3.05 - 3.10 in T. walteri).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
EDEA8A9FEDF3BE55FE1C97F4B82C98C1.taxon	description	Description. Female (Figure 168) (n = 5) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (570 - 645 (580) long; 420 - 480 (435) wide) ovoid with faint orange coloration or colorless. Anterio-medial platelets (122.5 - 131.25 (125) long; 40 - 47.5 (42.5) wide). Anterio-lateral platelets (155 - 180 (162.5) long; 50 - 57.5 (57.5) wide) fused to dorsal plate. Dgl- 4 closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 295 - 320 (310 )). Dorsal plate proportions: dorsum length / width 1.31 - 1.36 (1.33); dorsal width / distance between Dgl- 4 1.40 - 1.50 (1.40); anterio-medial platelet length / width 2.74 - 3.06 (2.94); anterio-lateral platelet length / width 2.83 - 3.20 (2.83); anterio-lateral / anterio-medial length 1.22 - 1.38 (1.30). Gnathosoma - Subcapitulum (310 - 330 (325) long (ventral); 232.5 - 243.25 (240) long (dorsal); 132.5 - 140 (132.5) tall) colorless. Rostrum (125 - 130 (130) long; 43.75 - 50 (45) wide). Chelicerae (315 - 325 (325) long) with curved fangs (60 - 66 (60) long). Subcapitular proportions: ventral length / height 2.34 - 2.45 (2.45); rostrum length / width 2.60 - 2.89 (2.89). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (45 - 50 (45) long); femur (112.5 - 122.5 (120) long); genu (62.5 - 72.5 (70) long); tibia (80 - 90 87.5) long; 25 - 28.75 (26.25) wide); tarsus (17.5 - 20 (17.5) long). Palpomere proportions: femur / genu 1.69 - 1.82 (1.71); tibia / femur 0.71 - 0.73 (0.73); tibia length / width 3.13 - 3.33 (3.33). Venter - (710 - 760 (730) long; 470 - 560 (490) wide) colorless. Gnathosomal bay (152.5 - 167.5 (167.5) long; 81.25 - 95 (85) wide). Cxgl- 4 subapical. Medial suture (10 - 12.5 (12.5) long). Genital plates (177.5 - 192.5 (182.5) long; 160 - 182.5 (160) wide). Additional measurements: Cx- 1 (270 - 281 (280) long (total); 110 - 124 (110) long (medial )); Cx- 3 (300 - 365 (320) wide); anterior venter (157.5 - 177.5 (157.5) long). Ventral proportions: gnathosomal bay length / width 1.61 - 1.97 (1.97); anterior venter / genital field length 0.86 - 0.92 (0.86); anterior venter length / genital field width 0.97 - 0.98 (0.98); anterior venter / medial suture 12.6 - 17.75 (12.6). Male (Figure 169) (n = 5) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (450 - 500 (455) long; 310 - 380 (320) wide) ovoid with faint orange coloration or colorless. Anterio-medial platelets (97.5 - 105 (102.5) long; 35 - 37.5 (37.5) wide). Anterio-lateral platelets (127.5 - 145 (137.5) long; 45 - 53.75 (45) wide) fused to dorsal plate. Dgl- 4 closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 240 - 285 (245 )). Dorsal plate proportions: dorsum length / width 1.32 - 1.45 (1.42); dorsal width / distance between Dgl- 4 1.29 - 1.37 (1.31); anterio-medial platelet length / width 2.67 - 2.80 (2.73); anterio-lateral platelet length / width 2.60 - 3.06 (3.06); anterio-lateral / anterio-medial length 1.28 - 1.44 (1.34). Gnathosoma - Subcapitulum (250 - 267.5 (252.5) long (ventral); 180 - 195 (192.5) long (dorsal); 95 - 107.5 (97.5) tall) colorless. Rostrum (97.5 - 105 (105) long; 35 - 37.5 (37.5) wide). Chelicerae (246 - 260 (250) long) with curved fangs (45 - 52 (50) long). Subcapitular proportions: ventral length / height 2.49 - 2.63 (2.59); rostrum length / width 2.67 - 2.80 (2.80). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (35 - 42.5 (37.5) long); femur (85 - 92.5 (92.5) long); genu (52.5 - 60 (56.25) long); tibia (66.25 - 75 (67.5) long; 22.5 - 27.5 (22.5) wide); tarsus (15 - 17.5 (17.5) long). Palpomere proportions: femur / genu 1.54 - 1.64 (1.64); tibia / femur 0.73 - 0.81 (0.73); tibia length / width 2.73 - 3.00 (3.00). Venter - (560 - 620 (590) long; 350 - 431 (370) wide) colorless. Gnathosomal bay (106 - 128 (122.5) long; 65 - 67.5 (65) wide). Cxgl- 4 subapical. Medial suture (82.5 - 110 (102.5) long). Genital plates (112.5 - 125 (115) long; 90 - 102.5 (90) wide). Additional measurements: Cx- 1 (211 - 230 (230) long (total); 100 - 110 (110) long (medial )); Cx- 3 (260 - 292 (270) wide); anterior venter (232.5 - 250 (240) long). Ventral proportions: gnathosomal bay length / width 1.60 - 1.89 (1.88); anterior venter / genital field length 1.96 - 2.13 (2.09); anterior venter length / genital field width 2.39 - 2.67 (2.67); anterior venter / medial suture 2.18 - 2.82 (2.34). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
EDEA8A9FEDF3BE55FE1C97F4B82C98C1.taxon	etymology	Etymology. Specific epithet (nortoni) named in honor of acarologist Roy Norton who taught JRF oribatids during the Soil Mite Course at The Ohio State University Acarology Summer Program; specifically for his talent in weaving biological stories into his lessons.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
EDEA8A9FEDF3BE55FE1C97F4B82C98C1.taxon	distribution	Distribution. California (Figure 167).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
90CFC3B959CDBF8B02D91BDED21C225E.taxon	materials_examined	Material examined. HOLOTYPE (♀): from USA, Wyoming, Medicine Bow National Forest, 1928, by JW Scott, RM 280072.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
90CFC3B959CDBF8B02D91BDED21C225E.taxon	diagnosis	Diagnosis. Torrenticola occidentalis are similar to other members of the Ellipsoidalis Group (T. ellipsoidalis, T. multiforma, and T. leviathan), and in being among the largest Torrenticola in the west (dorsum length ♀ = 700 - 885; ♂ = 665 - 850), although T. sierrensis are also large (dorsum length ♀ = 700 - 880; ♂ = 590 - 735) but can easily be distinguished from the Ellipsoidalis Group by being circular instead of ellipsoid or rectangular (dorsum length / width = 1.17 - 1.28 in T. sierrensis, 1.30 - 1.67 in Ellipsoidalis Group). T. occidentalis can be differentiated from T. ellipsoidalis by having a shorter medial suture (20 in T. occidentalis, 40 - 57.5 in T. ellipsoidalis) and by having more elongate anterio-lateral platelets (length / width = 2.54 in T. occidentalis, 2.00 - 2.39 in T. ellipsoidalis). T. occidentalis can be differentiated from T. multiforma by having stockier subcapitular rostra (length / width = 2.15 in T. occidentalis, 2.5 - 2.8 in T. multiforma). T. occidentalis can be differentiated from T. leviathan by having less elongate pedipalpal tibiae (length / width = 3.33 in T. occidentalis, 3.43 - 4.20 in T. leviathan) and a shorter dorsum (length ♀ = 770 in T. occidentalis, 845 - 870 in T. leviathan).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
90CFC3B959CDBF8B02D91BDED21C225E.taxon	description	Re-description. Female (Figure 171) (n = 1) (holotype only) with characters of the genus with following specifications. Dorsum- (770 long; 590 wide) ellipsoid with pink coloration without a distinct pattern. Anterio-medial platelets (145 long; 76.25 wide). Anterio-lateral platelets (235 long; 92.5 wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 455). Dorsal plate proportions: dorsum length / width 1.31; dorsal width / distance between Dgl- 4 1.30; anterio-medial platelet length / width 1.90; anterio-lateral platelet length / width 2.54; anterio-lateral / anterio-medial length 1.62. Gnathosoma - Subcapitulum (370 long (ventral); 282.5 long (dorsal); 170 tall) colorless. Rostrum (145 long; 67.5 wide) short. Chelicerae (370 long) with curved fangs (75 long). Subcapitular proportions: ventral length / height 2.18; rostrum length / width 2.15. Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (50 long); femur (127.5 long); genu (87.5 long); tibia (100 long; 30 wide); tarsus (15 long). Palpomere proportions: femur / genu 1.46; tibia / femur 0.78; tibia length / width 3.33. Venter - (980 long; 660 wide) colorless. Gnathosomal bay (202.5 long; 97.5 wide). Cxgl- 4 subapical. Medial suture (20 long). Genital plates (235 long; 210 wide). Additional measurements: Cx- 1 (385 long (total); 180 long (medial )); Cx- 3 (430 wide); anterior venter (217.5 long). Ventral proportions: gnathosomal bay length / width 2.08; anterior venter / genital field length 0.93; anterior venter length / genital field width 1.04; anterior venter / medial suture 10.88. Male unknown. Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
90CFC3B959CDBF8B02D91BDED21C225E.taxon	etymology	Etymology. Marshall (1933) did not specify an etymology for the specific epithet (occidentalis). However, surely this name refers to the type locality of this species in western United States (occidens, L. direction of the setting sun, west).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
90CFC3B959CDBF8B02D91BDED21C225E.taxon	distribution	Distribution. Known only from South-central Wyoming (Medicine Bow National Forest) (Figure 170), from a single female collected from trout stomach contents.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
B38B6DBAA45529D75BA5DF8188265BC6.taxon	materials_examined	Material examined. HOLOTYPE (♀): from USA, Oregon, Curry County, Siskiyou National Forest; beside Road # 33 between Powers and Agness, (42 ° 39 ' 39 " N, 124 ° 4 ' 4 " W), 2 July 1983, by IM Smith, IMS 830019. PARATYPES (4 ♀; 4 ♂): California, United States: 1 ♀ and 1 ♂ from Mendocino County, beside Rt. 1, 3.6 km southwest of Rt. 101, (39 ° 51 ' 51 " N, 123 ° 44 ' 44 " W), 5 August 1987, by IM Smith, IMS 870130 * 1 ♀ and 1 ♂ from Trinity County, beside Rt. 36, 5.2 km west of Forest Glen Station, (40 ° 23 ' 23 " N, 123 ° 21 ' 21 " W), 6 August 1987, by IM Smith & JD Smith, IMS 870132 * Oregon, United States: 1 ♂ (ALLOTYPE) from Curry County, Siskiyou National Forest; beside Road # 33 between Powers and Agness, (42 ° 39 ' 39 " N, 124 ° 4 ' 4 " W), 2 July 1983, by IM Smith, IMS 830019 * 1 ♀ from Curry County, Port Orford; Humbug Mountain State Park Picnic Area off Rt. 101, (42 ° 41 ' 41 " N, 124 ° 26 ' 26 " W), 1 July 1983, by IM Smith, IMS 830013 * 1 ♂ from Curry County, Port Orford; Humbug Mountain State Park Picnic Area off Rt. 101, (42 ° 41 ' 41 " N, 124 ° 25 ' 25 " W), 1 July 1983, by IM Smith, IMS 830012 * Washington, United States: 1 ♀ from Pierce County, Mt. Rainier National Park; Sunshine Point Campground, (46 ° 44 ' 44 " N, 121 ° 54 ' 54 " W), 30 June 1976, by IM Smith, IMS 760177.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
B38B6DBAA45529D75BA5DF8188265BC6.taxon	diagnosis	Diagnosis. Torrenticola oliveri are similar to members of the Miniforma group (T. copipalpa, T. manni, T. miniforma, T. pacificensis, T. rockyensis, and T. pinocchio) in having similar pedipalpal extensions (unique to members of this group). T. oliveri can be differentiated from all other members of Miniforma Complex by having a longer anterior venter (250 - 310 in T. oliveri, 173 - 235 in others) and more elongate pedipalpal tibiae (length / width = 3.68 - 4.13 in T. oliveri, 2.47 - 3.50 in others).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
B38B6DBAA45529D75BA5DF8188265BC6.taxon	description	Description. Female (Figure 173) (n = 5) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (615 - 710 (645) long; 480 - 545 (490) wide) ovoid with faint pink coloration without a distinct pattern. Anterio-medial platelets (125 - 157.5 (130) long; 65 - 75 (65) wide). Anterio-lateral platelets (210 - 245 (210) long; 75 - 87.5 (75) wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 370 - 430 (395 )). Dorsal plate proportions: dorsum length / width 1.27 - 1.34 (1.32); dorsal width / distance between Dgl- 4 1.24 - 1.30 (1.24); anterio-medial platelet length / width 1.92 - 2.10 (2.00); anterio-lateral platelet length / width 2.63 - 2.97 (2.80); anterio-lateral / anterio-medial length 1.56 - 1.68 (1.62). Gnathosoma - Subcapitulum (370 - 397.5 (397.5) long (ventral); 285 - 297.5 (297.5) long (dorsal); 112.5 - 122.5 (117.5) tall) colorless. Rostrum (150 - 160 (160) long; 47.5 - 50 (50) wide). Chelicerae (380 - 415 (415) long) with curved fangs (55 - 65 (60) long). Subcapitular proportions: ventral length / height 3.08 - 3.38 (3.38); rostrum length / width 3.00 - 3.20 (3.20). Pedipalps with tuberculate ventral extensions on femora and dentate, tuberculate ventral extensions on genua. Palpomeres: trochanter (33.75 - 37.5 (37.5) long); femur (110 - 118.75 (118.75) long); genu (75 - 83.75 (83.75) long); tibia (80 - 87.5 (85) long; 20 - 23.75 (22.5) wide); tarsus (12.5 - 15 (15) long). Palpomere proportions: femur / genu 1.38 - 1.53 (1.42); tibia / femur 0.72 - 0.75 (0.72); tibia length / width 3.68 - 4.00 (3.78). Venter - (770 - 835 (805) long; 550 - 650 (580) wide) with faint pink coloration. Gnathosomal bay (155 - 167.5 (167.5) long; 70 - 90 (72.5) wide). Cxgl- 4 subapical. Medial suture (80 - 95 (80) long). Genital plates (190 - 205 (205) long; 170 - 180 (180) wide). Additional measurements: Cx- 1 (320 - 340 (320) long (total); 140 - 185 (155) long (medial )); Cx- 3 (340 - 400 (360) wide); anterior venter (250 - 290 (250) long). Ventral proportions: gnathosomal bay length / width 1.75 - 2.31 (2.31); anterior venter / genital field length 1.22 - 1.45 (1.22); anterior venter length / genital field width 1.39 - 1.63 (1.39); anterior venter / medial suture 2.86 - 3.38 (3.13). Male (Figure 174) (n = 5) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (575 - 640 (590) long; 445 - 470 (470) wide ovoid with faint pink coloration without a distinct pattern. Anterio-medial platelets (125 - 140 (125) long; 60 - 75 (60) wide). Anterio-lateral platelets (205 - 220 (220) long; 65 - 90 (80) wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 355 - 375 (375 )). Dorsal plate proportions: dorsum length / width 1.26 - 1.39 (1.26); dorsal width / distance between Dgl- 4 1.25 - 1.28 (1.25); anterio-medial platelet length / width 1.87 - 2.17 (2.08); anterio-lateral platelet length / width 2.39 - 3.15 (2.75); anterio-lateral / anterio-medial length 1.54 - 1.76 (1.76). Gnathosoma - Subcapitulum (325 - 367.5 (355) long (ventral); 260 - 285 (262.5) long (dorsal); 105 - 110 (110) tall) colorless. Rostrum (135 - 145 (135) long; 45 - 47.5 (45) wide). Chelicerae (350 - 410 (365) long) with curved fangs (55 - 55 (55) long). Subcapitular proportions: ventral length / height 3.10 - 3.34 (3.23); rostrum length / width 3.00 - 3.11 (3.00). Pedipalps with tuberculate ventral extensions on femora and dentate, tuberculate ventral extensions on genua. Palpomeres: trochanter (32.5 - 35 (35) long); femur (105 - 107.5 (107.5) long); genu (75 - 80 (80) long); tibia (82.5 - 90 (87.5) long; 20 - 22.5 (21.25) wide); tarsus (12.5 - 15 (15) long). Palpomere proportions: femur / genu 1.34 - 1.40 (1.34); tibia / femur 0.79 - 0.86 (0.81); tibia length / width 4.00 - 4.13 (4.12). Venter - (725 - 795 (725) long; 515 - 540 (540) wide) with faint pink coloration. Gnathosomal bay (142.5 - 150 (150) long; 60 - 70 (70) wide). Cxgl- 4 subapical. Medial suture (105 - 130 (105) long). Genital plates (155 - 165 (160) long; 120 - 130 (130) wide). Additional measurements: Cx- 1 (290 - 310 (295) long (total); 145 - 165 (150) long (medial )); Cx- 3 (330 - 350 (350) wide); anterior venter (270 - 310 (270) long). Ventral proportions: gnathosomal bay length / width 2.14 - 2.40 (2.14); anterior venter / genital field length 1.69 - 1.88 (1.69); anterior venter length / genital field width 2.08 - 2.58 (2.08); anterior venter / medial suture 2.27 - 2.57 (2.57). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
B38B6DBAA45529D75BA5DF8188265BC6.taxon	etymology	Etymology. Specific epithet (oliveri) named in honor of comedian John Oliver and his team, whose commentary on Last Week Tonight breaches the realm of comedy and enters journalism. News and politics, like science, require both challenging misinformation and effectively communicating those challenges-tasks Oliver has humbly mastered.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
B38B6DBAA45529D75BA5DF8188265BC6.taxon	distribution	Distribution. Pacific Northwest (Figure 172).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
FDF673E9842E0784C17FA81DBE4FA3BF.taxon	materials_examined	Material examined. HOLOTYPE (♀): from USA, Oregon, Douglas County, Umpqua National Forest, Umpqua River (43 ° 17 ' 28 " N, 122 ° 37 ' 12 " W), 12 Aug 2013, by JC O'Neill, & WA Nelson, JNOW 13 - 0812 - 006. PARATYPES (9 ♀; 12 ♂): British Columbia, Canada: 1 ♀ and 1 ♂ from Vancouver Island, Malahat, Goldstream Provincial Park, Goldstream River, 26 Jun 1979, by IM Smith, IMS 790028 A * California, USA: 1 ♀ and 1 ♂ from Humboldt County, Honeydew, Mattole River, beside road to Bull Creek on east side of bridge, 8 Aug 1987, by IM Smith, IMS 870135 A * 1 ♀ and 1 ♂ from Trinity County, Trinity River, beside Route 299, 8.7 kilometers northwest of Del Loma, 9 Aug 1987, by IM Smith, IMS 870137 A * 1 ♀ from Trinity County, Weaver Creek, beside Route 299, 4.3 kilometers north of Route 3 West, 9 Aug 1987, by IM Smith, IMS 870138 A & IMS 870138 B * Oregon, USA: 1 ♂ from Coos County, Siskiyou National Forest, Road 33 between Powers & Agness, Daphne Grove campground, Coquille River, 2 Jul 1983, by IM Smith, IMS 830016 * 3 ♀ and 6 ♂ from Curry County, Port Orford, Butler Bar campground, Elk River, 25 - 26 Jun 1976, by IM Smith, IMS 760163 * 1 ♀ from Curry County, Port Orford, small spring run beside road from Humbug Mountain State Park to McGribble campground, 25 Jun 1976, by IM Smith, IMS 760161 * 1 ♂ (ALLOTYPE) from Douglas County, Umpqua National Forest, Umpqua River (43 ° 17 ' 28 " N, 122 ° 37 ' 12 " W), 12 Aug 2013, by JC O'Neill, & WA Nelson, JNOW 13 - 0812 - 006 * 1 ♀ and 1 ♂ from Douglas County, Umpqua National Forest, Umpqua River (43 ° 17 ' 28 " N, 122 ° 37 ' 12 " W), 12 Aug 2013, by JC O'Neill, & WA Nelson, JNOW 13 - 0812 - 006.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
FDF673E9842E0784C17FA81DBE4FA3BF.taxon	diagnosis	Diagnosis. Torrenticola olliei are similar to other members of the Tricolor Complex (T. bittikoferae, T. hoosieri, T. larvata, T. pearsoni, T. sierrensis, T. tricolor, T. trimaculata, T. unimaculata, T. cardia, T. kringi, T. dimorpha, and T. mohawk) in having a short, conical rostrum. T. olliei can be differentiated from most Torrenticola, including most other members of the Tricolor Complex, by having a very short rostrum (length / width = 1.56 - 1.81 in T. olliei, 1.80 - 3.13 in others), except T. bittikoferae (1.81 - 1.9) and T. dimorpha (1.3 - 1.6). Additionally, males have a shorter anterior venter than all other Tricolor Complex (♂ = 205 - 225 in T. olliei, 230 - 335 in others), except T. kringi (220 - 255), and both males and females have a wider genital field than most other Tricolor Complex (♀ = 190 - 202.5 in T. olliei, 145 - 180 in others; ♂ = 130 - 137.5 in T. olliei, 92.5 - 120 in others) except for T. sierrensis (♀ = 180 - 212.5; ♂ = 135 - 175) and female T. leviathan (180 - 195). Finally, T. olliei can be differentiated from most Tricolor Complex, except for T. sierrensis, by being distributed in the west (all others are eastern).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
FDF673E9842E0784C17FA81DBE4FA3BF.taxon	description	Description. Female (Figure 176) (n = 5) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (690 - 900 (690) long; 500 - 670 (500) wide) ovoid with orange coloration separated into posterior and anterior portions with faint orange medially. Anterio-medial platelets (130 - 172.5 (130) long; 60 - 80 (60) wide). Anterio-lateral platelets (190 - 235 (190) long; 70 - 92.5 (70) wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 360 - 470 (360 )). Dorsal plate proportions: dorsum length / width 1.34 - 1.42 (1.38); dorsal width / distance between Dgl- 4 1.31 - 1.43 (1.39); anterio-medial platelet length / width 1.95 - 2.23 (2.17); anterio-lateral platelet length / width 2.34 - 2.71 (2.71); anterio-lateral / anterio-medial length 1.36 - 1.46 (1.46). Gnathosoma - Subcapitulum (245 - 287.5 (245) long (ventral); 155 - 225 (155) long (dorsal); 137.5 - 152.5 (137.5) tall) colorless and tall. Rostrum (85 - 112.5 (85) long; 50 - 67.5 (50) wide) short and conical. Chelicerae (225 - 305 (225) long) with curved fangs (55 - 65 (55) long). Subcapitular proportions: ventral length / height 1.78 - 1.92 (1.78); rostrum length / width 1.62 - 1.76 (1.70). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (37.5 - 50 long); femur (87.5 - 122.5 (87.55) long); genu (58.75 - 82.5 (60) long); tibia (75 - 102.5 (75) long; 25 - 33.75 (25) wide); tarsus (25 - 30 (26.25) long). Palpomere proportions: femur / genu 1.46 - 1.51 (1.46); tibia / femur 0.84 - 0.90 (0.86); tibia length / width 3.00 - 3.08 (3.00). Venter - (790 - 1050 (810) long; 550 - 765 (550) wide) colorless. Gnathosomal bay (120 - 170 (150) long; 75 - 97.5 (75) wide). Cxgl- 4 subapical. Medial suture (12.5 - 50 (12.5) long). Genital plates (202.5 - 230 (212.5) long; 190 - 202.5 (190) wide). Additional measurements: Cx- 1 (260 - 375 (280) long (total); 130 - 195 (130) long (medial )); Cx- 3 (350 - 465 (350) wide); anterior venter (162.5 - 255 (162.5) long). Ventral proportions: gnathosomal bay length / width 1.37 - 2.00 (2.00); anterior venter / genital field length 0.76 - 1.11 (0.76); anterior venter length / genital field width 0.85 - 1.28 (0.86); anterior venter / medial suture 5.10 - 13.00 (13.00). Male (Figure 177) (n = 5) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (560 - 640 (570) long; 410 - 460 (410) wide) ovoid with orange coloration separated into posterior and anterior portions with faint orange medially. Anterio-medial platelets (102.5 - 125 (107.5) long; 50 - 70 (56.25) wide). Anterio-lateral platelets (170 - 187.5 (170) long; 62.5 - 80 (62.5) wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 320 - 375 (320 )). Dorsal plate proportions: dorsum length / width 1.33 - 1.44 (1.39); dorsal width / distance between Dgl- 4 1.19 - 1.33 (1.28); anterio-medial platelet length / width 1.71 - 2.25 (1.91); anterio-lateral platelet length / width 2.34 - 2.72 (2.72); anterio-lateral / anterio-medial length 1.50 - 1.68 (1.58). Gnathosoma - Subcapitulum (197.5 - 220 (212.5) long (ventral); 147.5 - 160 (159) long (dorsal); 115 - 125 (115) tall) colorless and tall. Rostrum (70 - 82.5 (72.5) long; 40 - 50 (40) wide) short and conical. Chelicerae (195 - 220 long) with curved fangs (47.5 - 52.5 long). Subcapitular proportions: ventral length / height 1.65 - 1.85 (1.85); rostrum length / width 1.56 - 1.81 (1.81). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (32.5 - 35 (32.5) long); femur (75 - 77.5 (76.25) long); genu (52.5 - 55 (52.5) long); tibia (67.5 - 75 (67.5) long; 22.5 - 25 (22.5) wide); tarsus (20 - 25 (20) long). Palpomere proportions: femur / genu 1.41 - 1.48 (1.45); tibia / femur 0.89 - 0.97 (0.89); tibia length / width 2.70 - 3.00 (3.00). Venter - (650 - 710 (650) long; 444 - 530 (444) wide) colorless. Gnathosomal bay (122.5 - 137.5 (127.5) long; 62.5 - 75 (62.5) wide). Cxgl- 4 subapical. Medial suture (70 - 80 (75) long). Genital plates (165 - 172.5 (165) long; 130 - 137.5 (130) wide). Additional measurements: Cx- 1 (234 - 270 (235) long (total); 102 - 135 (102) long (medial )); Cx- 3 (311 - 355 (312) wide); anterior venter (200 - 225 (207.5) long). Ventral proportions: gnathosomal bay length / width 1.70 - 2.04 (2.04); anterior venter / genital field length 1.19 - 1.30 (1.26); anterior venter length / genital field width 1.54 - 1.67 (1.60); anterior venter / medial suture 2.67 - 2.93 (2.77). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
FDF673E9842E0784C17FA81DBE4FA3BF.taxon	etymology	Etymology. Specific epithet (olliei) named in honor of Ollie-pet Boston Terrier of JRF and DMF-whose brachycephaly resembles the short rostrum of this species, the shortest of all Torrenticola.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
FDF673E9842E0784C17FA81DBE4FA3BF.taxon	distribution	Distribution. Pacific coastal mountains (Figure 175).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
E9614F351BAA169025C05FAA36084BFF.taxon	materials_examined	Material examined. HOLOTYPE (♀): from USA, Oregon, Grant County, Prairie City; Strawberry Forest Camp, (44 ° 19 ' 19 " N, 118 ° 40 ' 40 " W), 17 June 1976, by IM Smith, IMS 760126. PARATYPES (3 ♀; 3 ♂): Oregon, USA: 1 ♂ (ALLOTYPE) from Grant County, Prairie City; Strawberry Forest Camp, (44 ° 19 ' 19 " N, 118 ° 40 ' 40 " W), 17 June 1976, by IM Smith, IMS 760126 * 1 ♀ from Benton County, Philomath; Marys Peak Campground, (44 ° 30 ' 30 " N, 123 ° 33 ' 33 " W), 23 June 1976, by IM Smith, IMS 760147 * 2 ♂ from Benton County, Philomath; Marys Peak Campground, (44 ° 30 ' 30 " N, 123 ° 33 ' 33 " W), 27 June 1983, by IM Smith & AB Smith, IMS 830006 * 1 ♀ from Benton County, Philomath; Marys Peak Campground, (44 ° 30 ' 30 " N, 123 ° 33 ' 33 " W), 28 June 1983, by IM Smith, IMS 830008 * 1 ♀ from Grant County, Prairie City; beside Rt. 26 just east of Dixie Pass, (44 ° 32 ' 32 " N, 118 ° 32 ' 32 " W), 17 June 1976, by IM Smith, IMS 760125.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
E9614F351BAA169025C05FAA36084BFF.taxon	diagnosis	Diagnosis. Torrenticola oregonensis are similar to the other member of Tahoei group, T. tahoei by subcapitulum shape, having purple coloration restricted posteriorly and being distributed in the west and to members of the Nigroalba Group (T. flangipalpa, T. nigroalba, T. solisorta, and T. dentirostra) in having purple dorsal coloration restricted posteriorly. T. oregonensis can be differentiated from the Nigroalba Group by being larger (dorsum length ♀ = 760 - 840 in T. oregonensis, 475 - 565 in Nigroalba Group; ♂ = 690 - 820 in T. oregonensis, 425 - 510 in Nigroalba Group) and distributed in Oregon (Nigroalba Group are eastern). T. oregonensis can be differentiated from T. tahoei by being larger (dorsum length ♀ = 760 - 840 in T. oregonensis, 600 - 720 in T. tahoei; ♂ = 690 - 820 in T. oregonensis, 560 - 650 in T. tahoei; dorsum width ♀ = 560 - 640 in T. oregonensis, 430 - 515 in T. tahoei; ♂ = 520 - 605 in T. oregonensis, 400 - 460 in T. tahoei) and a less elongate subcapitulum (ventral length / width = 2.63 - 2.74 in T. oregonensis, 3.25 - 4.11 in T. tahoei).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
E9614F351BAA169025C05FAA36084BFF.taxon	description	Description. Female (Figure 179) (n = 4) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (760 - 840 (760) long; 560 - 640 (560) wide) ovoid with faint pink coloration medially, occasionally continuing posteriorly. Anterio-medial platelets (145 - 163.75 (145) long; 72.5 - 87.5 (72.5) wide). Anterio-lateral platelets (222.5 - 247.5 (222.5) long; 97.5 - 102.5 (100) wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 435 - 495 (435 )). Dorsal plate proportions: dorsum length / width 1.30 - 1.36 (1.36); dorsal width / distance between Dgl- 4 1.27 - 1.29 (1.29); anterio-medial platelet length / width 1.85 - 2.21 (2.00); anterio-lateral platelet length / width 2.23 - 2.45 (2.23); anterio-lateral / anterio-medial length 1.48 - 1.56 (1.53). Gnathosoma - Subcapitulum (380 - 415 (380) long (ventral); 280 - 307.5 (280) long (dorsal); 140 - 157.5 (140) tall) colorless. Rostrum (152.5 - 170 (152.5) long; 60 - 65 (60) wide). Chelicerae (380 - 425 (380) long) with curved fangs (70 - 75 (70) long). Subcapitular proportions: ventral length / height 2.63 - 2.71 (2.71); rostrum length / width 2.48 - 2.62 (2.54). Pedipalps with tuberculate ventral extensions on femora and tuberculate extensions on genua that are slightly dentate. Palpomeres: trochanter (42.5 - 46.25 (42.5) long); femur (115 - 133.75 (115) long); genu (83.75 - 95 (83.75) long); tibia (100 - 110 (100) long; 25 - 27.5 (25) wide); tarsus (16.25 - 17.5 (17.5) long). Palpomere proportions: femur / genu 1.37 - 1.43 (1.37); tibia / femur 0.78 - 0.87 (0.87); tibia length / width 3.81 - 4.00 (4.00). Venter - (900 - 990 (900) long; 660 - 750 (660) wide) colorless. Gnathosomal bay (170 - 185 (170) long; 75 - 100 (75) wide). Cxgl- 4 subapical. Medial suture (60 - 80 (60) long). Genital plates (210 - 230 (210) long; 185 - 205 (185) wide). Additional measurements: Cx- 1 (330 - 370 (330) long (total); 162.5 - 190 (170) long (medial )); Cx- 3 (410 - 470 (410) wide); anterior venter (230 - 270 (230) long). Ventral proportions: gnathosomal bay length / width 1.80 - 2.27 (2.27); anterior venter / genital field length 1.04 - 1.22 (1.10); anterior venter length / genital field width 1.24 - 1.32 (1.24); anterior venter / medial suture 3.00 - 3.83 (3.83). Male (Figure 180) (n = 3) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (690 - 820 (820) long; 520 - 605 (605) wide) ovoid with faint pink coloration medially, occasionally continuing posteriorly. Anterio-medial platelets (140 - 155 (150) long; 67.5 - 82.5 (82.5) wide). Anterio-lateral platelets (215 - 247.5 (247.5) long; 87.5 - 102.5 (102.5) wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 420 - 480 (480 )). Dorsal plate proportions: dorsum length / width 1.33 - 1.36 (1.36); dorsal width / distance between Dgl- 4 1.24 - 1.26 (1.26); anterio-medial platelet length / width 1.82 - 2.30 (1.82); anterio-lateral platelet length / width 2.25 - 2.46 (2.41); anterio-lateral / anterio-medial length 1.45 - 1.65 (1.65). Gnathosoma - Subcapitulum (370 - 380 (380) long (ventral); 275 - 290 (290) long (dorsal); 135 - 140 (140) tall) colorless. Rostrum (145 - 150 (150) long; 52.5 - 60 (60) wide). Chelicerae (375 - 395 (395) long) with curved fangs (65 - 70 (70) long). Subcapitular proportions: ventral length / height 2.64 - 2.74 (2.71); rostrum length / width 2.50 - 2.76 (2.50). Pedipalps with tuberculate ventral extensions on femora and tuberculate extensions on genua that are slightly dentate. Palpomeres: trochanter (40 - 43.75 (43.75) long); femur (115 - 127.5 (127.5) long); genu (82.5 - 87.5 (87.5) long); tibia (97.5 - 102.5 (102.5) long; 25 - 26.25 (26.25) wide); tarsus (15 - 16.25 (15) long). Palpomere proportions: femur / genu 1.31 - 1.46 (1.46); tibia / femur 0.80 - 0.85 (0.80); tibia length / width 3.90 - 3.90 (3.90). Venter - (840 - 930 (930) long; 610 - 710 (710) wide) colorless. Gnathosomal bay (157.5 - 172.5 (157.5) long; 70 - 100 (100) wide). Cxgl- 4 subapical. Medial suture (100 - 120 (120) long). Genital plates (165 - 175 (175) long; 130 - 140 (140) wide). Additional measurements: Cx- 1 (330 - 340 (340) long (total); 160 - 180 (180) long (medial )); Cx- 3 (390 - 455 (455) wide); anterior venter (275 - 325 (325) long). Ventral proportions: gnathosomal bay length / width 1.58 - 2.46 (1.58); anterior venter / genital field length 1.65 - 1.86 (1.86); anterior venter length / genital field width 2.11 - 2.32 (2.32); anterior venter / medial suture 2.71 - 2.75 (2.71). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
E9614F351BAA169025C05FAA36084BFF.taxon	etymology	Etymology. Specific epithet (oregonensis) refers to the distribution of this species, which was paradoxically found in several ecoregions across Oregon, but not in surrounding areas.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
E9614F351BAA169025C05FAA36084BFF.taxon	distribution	Distribution. Oregon (Figure 178).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
06B3F8E2AF9BFE1D2CAF842BAA8B417B.taxon	materials_examined	Material examined. HOLOTYPE (♀): from USA, Washington, Clallam County, Green Creek (48 ° 10 ' 45 " N, 124 ° 12 ' 21 " W), 24 Jul 2013, by JC O'Neill, & WA Nelson, JNOW 13 - 0724 - 005. PARATYPES (34 ♀; 31 ♂): British Columbia, Canada: 2 ♀ and 1 ♂ from Vancouver Island, Caycuse, Nixon Creek, 8 Jul 1976, by IM Smith, IMS 760197 & IMS 760198 * 1 ♀ and 1 ♂ from Vancouver Island, beside Highway 4, 35.6 kilometers east of Pacific Rim Road, 9 Jul 1976, by IM Smith, IMS 760206 * 1 ♂ from Vancouver Island, beside Highway 4, 1.3 kilometers east of Pacific Rim Road, Lost Shoe Creek, 9 Jul 1976, by IM Smith, IMS 760202 * 2 ♀ and 2 ♂ from Vancouver Island, Lake Cowichan, Skutz Falls, Skutz Creek, near Cowichan River, 9 Jul 1979, by IM Smith, IMS 790036 A & IMS 790036 B * 1 ♀ from Vancouver Island, Lake Cowichan, Cowichan River, above Skutz Falls, 9 Jul 1979, by IM Smith, IMS 790035 * 1 ♀ and 1 ♂ from Vancouver Island, Lake Cowichan, South Shore Road, north of Mesachie Lake, Robertson River, 4 Jul 1976, by IM Smith, IMS 760183 A * 1 ♀ and 1 ♂ from Vancouver Island, Lake Cowichan, South Shore Road, north of Mesachie Lake, tributary of Robertson River, 10 Jul 1976, by IM Smith, IMS 760183 B * 1 ♀ and 1 ♂ from Vancouver Island, Lake Cowichan, South Shore Road, north of Mesachie Lake, tributary of Robertson River, 4 - 10 Jul 1976, by IM Smith, IMS 760182 * 1 ♀ and 1 ♂ from Vancouver Island, Malahat, Goldstream Provincial Park, Goldstream River, 26 Jun 1979, by IM Smith, IMS 790028 A * 1 ♀ and 1 ♂ from Vancouver Island, Malahat, Goldstream Provincial Park, near Mt. Finlayson Trail, 26 Jun 1979, by IM Smith, IMS 790027 A * 1 ♂ from Vancouver Island, beside South Shore Road, 2.3 kilometers south of Caycuse, 26 Jul 1979, by IM Smith, IMS 790052 * 1 ♂ from Vancouver Island, beside North Shore Road, 1.7 kilometers north of Lake Cowichan, 7 Jun 1979, by IM Smith, IMS 790008 C * 1 ♀ and 1 ♂ from Vancouver Island, North Shore Road, 3.2 kilometers south of Youbou, 4 Jul 1976, by IM Smith, IMS 760190 * 1 ♂ from Vancouver Island, Youbou, North Shore Road, Shaw Creek, 4.3 kilometers south of north end of Cowichan Lake, by IM Smith, IMS 760196 * Oregon, USA: 1 ♀ and 1 ♂ from Benton County, Marys Peak near Philomath, Parker Creek, 27 - 28 Jun 1983, by IM Smith & AB Smith, IMS 830006 * 1 ♀ from Benton County, Marys Peak near Philomath, Parker Creek, 28 Jun 1983, by IM Smith, IMS 830007 * 1 ♂ from Benton County, Philomath, Marys Peak campground, Park Creek, 23 Jun 1976, by IM Smith, IMS 760146 * 1 ♀ from Coos County, Siskiyou National Forest, road 33 between Powers and Agness, Coal Creek, 2 Jul 1983, by IM Smith, IMS 830015 * 1 ♀ from Coos County, Siskiyou National Forest, road 33 between Powers and Agness, 0.5 kilometers east of Daphne Grove campground, 2 Jul 1983, by IM Smith, IMS 830017 * 3 ♀ and 2 ♂ from Curry County, Port Orford, small spring run beside road from Humbug Mountain State Park to McGribble campground, 25 Jun 1976, IMS 760161 * 1 ♂ from Curry County, Port Orford, Humbug Mountain State Park Picnic Area, Brush Creek, beside Route 1, 3 Jul 1983, by IM Smith, IMS 830020 A * 1 ♀ and 1 ♂ from Curry County, Siskiyou National Forest, road 33 between Powers and Agness, North Fork of Foster Creek, 2 Jul 1983, by IM Smith, IMS 830019 * 1 ♀ from Lane County, Gate Creek (44 ° 8 ' 48 " N, 122 ° 34 ' 20 " W), 11 Aug 2013, by JC O'Neill, & WA Nelson, JNOW 13 - 0811 - 001 * 1 ♀ and 1 ♂ from Lincoln County, Blackberry campground near Tidewater, Alsea River, 28 Jun 1983, by IM Smith & AB Smith, IMS 830009 * 4 ♀ and 2 ♂ from Marion County, Marion Forks Riverside campground, North Fork of Santiam River, 22 Jun 1976, by IM Smith, IMS 760145 * 2 ♀ and 2 ♂ from Multnomah County, Columbia River Scenic Highway, stream below Horsetail Falls, 27 Jun 1983, by IM Smith, IMS 830005 * 1 ♀ from Tillamook County, Siuslaw National Forest, Alder Creek (45 ° 9 ' 27 " N, 123 ° 47 ' 60 " W), 6 Aug 2013, by JC O'Neill, & WA Nelson, JNOW 13 - 0806 - 002 * Washington, USA: 1 ♂ (ALLOTYPE) from Clallam County, Green Creek (48 ° 10 ' 45 " N, 124 ° 12 ' 21 " W), 24 Jul 2013, by JC O'Neill, & WA Nelson, JNOW 13 - 0724 - 005 * 2 ♀ and 1 ♂ from Clallam County, Green Creek (48 ° 10 ' 45 " N, 124 ° 12 ' 21 " W), 24 Jul 2013, by JC O'Neill, & WA Nelson, JNOW 13 - 0724 - 005 * 2 ♀ and 1 ♂ from Lewis County, Gifford Pinchot National Forest, Snake Creek (46 ° 38 ' 52 " N, 121 ° 43 ' 8 " W), 23 Jul 2013, by JC O'Neill, & WA Nelson, JNOW 13 - 0723 - 006 * 1 ♀ and 3 ♂ from Snohomish County, Mount Baker National Forest, Marten River (48 ° 4 ' 19 " N, 121 ° 36 ' 24 " W), 28 Jul 2013, by JC O'Neill, & WA Nelson, JNOW 13 - 0728 - 002.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
06B3F8E2AF9BFE1D2CAF842BAA8B417B.taxon	diagnosis	Diagnosis. Torrenticola pacificensis are similar to members of the Miniforma group (T. copipalpa, T. manni, T. miniforma, T. rockyensis, T. oliveri, and T. pinocchio) in having short, stocky pedipalps (except T. oliveri and T. pinocchio); similar pedipalpal extensions (unique to members of this group); and being among the smallest Torrenticola in the west (dorsum 500 - 625 long) (except T. oliveri). T. pacificensis are best differentiated from T. rockyensis by females having a stockier rostrum (length / width ♀ = 2.59 - 2.68 in T. pacificensis, 2.72 - 2.91 in T. rockyensis); and by being distributed in the Pacific Coast Ranges of Washington and Oregon instead of the Rocky Mountains (Idaho & Montana). T. pacificensis are best differentiated from T. copipalpa by having tuberculate pedipalp femoral extensions (broad and flat in T. copipalpa) and by being distributed in the Pacific Coast Ranges of Washington and Oregon, whereas although T. pacificensis overlap, they are more southern (southwest Oregon & California). T. pacificensis are best differentiated from T. manni by having stockier tibiae (length / width = 2.67 - 3.00 in pacificensis, 3.13 - 3.38 in T. manni); having a stockier rostrum (length / width ♀ = 2.59 - 2.68 in T. pacificensis, 3.00 - 3.13 in T. manni; ♂ = 2.76 - 3.07 in T. pacificensis, 3.13 - 3.20 in T. manni); and by being distributed in the Pacific Coast Ranges of Washington and Oregon, whereas T. manni is only known from Catron County, New Mexico. T. pacificensis are best differentiated from T. miniforma by having stockier anterio-medial platelets (length / width = 2.00 - 2.16 in T. pacificensis, 2.44 - 2.65 in T. miniforma); stockier subcapitular rostra (length / width = 2.59 - 3.07 in T. pacificensis; 3.13 - 3.19 in T. miniforma); and by being distributed in the Pacific Coast Ranges of Washington and Oregon, whereas T. miniforma is only known from Humboldt County, California. T. pacificensis can be differentiated from T. oliveri by having a shorter anterior venter (177 - 235 in T. pacificensis, 250 - 310 in T. oliveri) and less elongate pedipalpal tibiae (length / width = 2.6 - 3.0 in T. pacificensis, 3.6 - 4.2 in T. oliveri). T. pacificensis can be differentiated from T. pinocchio by having a less elongate rostrum (length / width = 2.5 - 3.1 in T. pacificensis, 4.5 - 4.9 in T. pinocchio) and less elongate pedipalpal tibiae (length / width = 2.67 - 3.00 in T. pacificensis, 3.13 - 3.50 in T. pinocchio).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
06B3F8E2AF9BFE1D2CAF842BAA8B417B.taxon	description	Description. Female (Figure 182) (n = 6) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (530 - 625 (580) long; 335 - 430 (400) wide) ovoid with purple coloration restricted posteriorly. Anterio-medial platelets (100 - 117.5 (110) long; 47.5 - 55 (52.5) wide). Anterio-lateral platelets (145 - 170 (162.5) long; 52.5 - 63.75 (57.5) wide) free from dorsal plate. Dgl- 4 much closer to the edge of dorsum than to the muscle scars (distance between Dgl- 4 270 - 345 (300 )). Dorsal plate proportions: dorsum length / width 1.42 - 1.58 (1.45); dorsal width / distance between Dgl- 4 1.24 - 1.36 (1.33); anterio-medial platelet length / width 2.05 - 2.14 (2.10); anterio-lateral platelet length / width 2.63 - 2.95 (2.83); anterio-lateral / anterio-medial length 1.43 - 1.48 (1.48). Gnathosoma - Subcapitulum (287.5 - 325 (300) long (ventral); 210 - 236 (225) long (dorsal); 110 - 125 (115) tall) colorless. Rostrum (110 - 127.5 (120) long; 41.25 - 47.5 (45) wide). Chelicerae (265 - 331 (305) long) with curved fangs (50 - 62 (52.5) long). Subcapitular proportions: ventral length / height 2.52 - 2.65 (2.61); rostrum length / width 2.59 - 2.68 (2.67). Pedipalps short and stocky (especially tibiae) with nearly tuberculate, dentate ventral extensions on femora and dentate, flanged ventral extensions on genua. Palpomeres: trochanter (30 - 35 (32.5) long); femur (85 - 95 (95) long); genu (57.5 - 67.5 (65) long); tibia (55 - 67.5 (62.5) long; 20 - 23.75 (22.5) wide); tarsus (12.5 - 17.5 (15) long). Palpomere proportions: femur / genu 1.40 - 1.48 (1.46); tibia / femur 0.65 - 0.71 (0.66); tibia length / width 2.71 - 3.00 (2.78). Venter - (650 - 770 (720) long; 434 - 521 (440) wide) colorless or with faint purple coloration. Gnathosomal bay (122.5 - 142.5 (140) long; 66.25 - 87.5 (67.5) wide). Cxgl- 4 subapical. Medial suture (47.5 - 55 (50) long). Genital plates (162.5 - 185 (180) long; 143.75 - 172.5 (160) wide). Additional measurements: Cx- 1 (230 - 289 (250) long (total); 110 - 142 (120) long (medial )); Cx- 3 (280 - 331 (280) wide); anterior venter (177.5 - 205 (182.5) long). Ventral proportions: gnathosomal bay length / width 1.49 - 2.07 (2.07); anterior venter / genital field length 1.01 - 1.11 (1.01); anterior venter length / genital field width 1.07 - 1.23 (1.14); anterior venter / medial suture 3.38 - 3.89 (3.65). Male (Figure 183) (n = 5) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (525 - 590 (550) long; 355 - 375 (370) wide) ovoid with purple coloration restricted posteriorly. Anterio-medial platelets (97.5 - 115 (102.5) long; 47.5 - 55 (47.5) wide). Anterio-lateral platelets (147.5 - 162.5 (147.5) long; 52.5 - 57.5 52.5) wide) free from dorsal plate. Dgl- 4 much closer to the edge of dorsum than to the muscle scars (distance between Dgl- 4 280 - 290 (290 )). Dorsal plate proportions: dorsum length / width 1.48 - 1.57 (1.49); dorsal width / distance between Dgl- 4 1.28 - 1.30 (1.28); anterio-medial platelet length / width 2.00 - 2.16 (2.16); anterio-lateral platelet length / width 2.73 - 2.83 (2.81); anterio-lateral / anterio-medial length 1.41 - 1.54 (1.44). Gnathosoma - Subcapitulum (275 - 287.5 (285) long (ventral); 195 - 220 (212) long (dorsal); 103.75 - 115 (107.5) tall) colorless. Rostrum (107.5 - 117.5 (113.75) long; 35 - 42.5 (38.75) wide). Chelicerae (251 - 281 (273) long) with curved fangs (46 - 57 (47) long). Subcapitular proportions: ventral length / height 2.50 - 2.65 (2.65); rostrum length / width 2.76 - 3.07 (2.94). Pedipalps short and stocky (especially tibiae) with nearly tuberculate, dentate ventral extensions on femora and dentate, flanged ventral extensions on genua. Palpomeres: trochanter (30 - 32.5 (32.5) long); femur (82.5 - 90 (85) long); genu (57.5 - 65 (62.5) long); tibia (55 - 60 (57.5) long; 20 - 22.5 (21.25) wide); tarsus (15 - 16.25 (16.25) long). Palpomere proportions: femur / genu 1.36 - 1.50 (1.36); tibia / femur 0.65 - 0.68 (0.68); tibia length / width 2.67 - 2.94 (2.71). Venter - (645 - 740 (740) long; 388 - 460 (431) wide) colorless or with faint purple coloration. Gnathosomal bay (120 - 140 (135) long; 60 - 75 (60) wide). Cxgl- 4 subapical. Medial suture (82.5 - 87.5 (82.5) long). Genital plates (140 - 147.5 (140) long; 110 - 120 (113.75) wide). Additional measurements: Cx- 1 (243 - 277 (277) long (total); 120 - 146 (125) long (medial )); Cx- 3 (245 - 310 (300) wide); anterior venter (216.25 - 235 (220) long). Ventral proportions: gnathosomal bay length / width 1.60 - 2.25 (2.25); anterior venter / genital field length 1.50 - 1.59 (1.57); anterior venter length / genital field width 1.89 - 1.98 (1.93); anterior venter / medial suture 2.49 - 2.69 (2.67). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
06B3F8E2AF9BFE1D2CAF842BAA8B417B.taxon	etymology	Etymology. Specific epithet (pacificensis) refers to the distribution of this species in the Pacific Ranges of the Pacific Northwest. This location-based name reflects that locality is the best way to differentiate this species from others in the Miniforma Group, particularly T. rockyensis.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
06B3F8E2AF9BFE1D2CAF842BAA8B417B.taxon	distribution	Distribution. Western Oregon and western Washington (Figure 181). It seems reasonable to suspect T. pacificensis also occurs in coastal British Columbia and northwestern California.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
7C5962D67BEEE4C829749B99BAE0F827.taxon	materials_examined	Material examined. HOLOTYPE (♀): from USA, Arkansas, Montgomery County, Ouachita National Forest, South Fork Ouachita River, 29 Jul 2011, by AJ Radwell, & B Crump, AJR 110302. PARATYPES (7 ♀; 10 ♂): Arkansas, USA: 2 ♀ and 3 ♂ from Montgomery County, Ouachita National Forest, South Fork Ouachita River, 29 Jul 2011, by AJ Radwell, & B Crump, AJR 110302 * 1 ♀ from Newton County, Buffalo National River, Mill Creek (36 ° 3 ' 42.12 " N, 93 ° 8 ' 7.62 " W), 30 May 2012, by TD Edwards, TDE 12 - 0530 - 004 * 1 ♀ from Newton County, Buffalo National River, Mill Creek (36 ° 3 ' 42.12 " N, 93 ° 8 ' 7.62 " W), 20 Jun 2012, by TD Edwards, TDE 12 - 0620 - 004 * 1 ♀ from Newton County, Ozark-St Francis National Forest, Little Buffalo River, 11 Jul 2012, by TD Edwards, TDE 12 - 0711 - 004 * Missouri, USA: 1 ♂ (ALLOTYPE) from Crawford County, Huzzah Creek, Red Bluff campground, east of Davisville, 23 Jul 2011, by IM Smith, IMS 110029 * 1 ♂ from Crawford County, Huzzah Creek, Red Bluff campground, east of Davisville, 23 Jul 2011, by IM Smith, IMS 110029 * 2 ♂ from McDonald County, Tiff City, beside Route 43, Buffalo Creek (36 ° 40 ' 17 " N, 94 ° 36 ' 17 " W), 2 May 1996, by IM Smith, IMS 960004 * 1 ♀ and 1 ♂ from Oregon County, beside Route 19, north of Greer, Eleven Point (36 ° 48 ' N, 91 ° 20 ' W), 28 Jun 1987, by IM Smith, IMS 870056 * Oklahoma, USA: 1 ♀ and 2 ♂ from Pushmataha County, beside Route 271, south of Albion, Walnut Creek (34 ° 39 ' N, 95 ° 7 ' W), 1 Jul 1987, by IM Smith, IMS 870063 A.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
7C5962D67BEEE4C829749B99BAE0F827.taxon	diagnosis	Diagnosis. Torrenticola pearsoni are similar to other members of the Tricolor Complex (T. bittikoferae, T. hoosieri, T. larvata, T. olliei, T. sierrensis, T. tricolor, T. trimaculata, T. unimaculata, T. cardia, T. kringi, T. dimorpha, and T. mohawk) in having a short, conical rostrum. T. pearsoni can be differentiated from most other Tricolor Complex (except T. bittikoferae, T. hoosieri and T. dimorpha) by having diffuse pink dorsal coloration, whereas most other members have bold patterning. T. pearsoni can be differentiated from T. hoosieri by having ventral extensions on the pedipalp femora and genua (lacking in T. hoosieri) and having stockier pedipalp tibiae (length / width = 3.0 - 3.3 in T. pearsoni, 3.6 - 4.4 in T. hoosieri). Male T. pearsoni can be differentiated from T. bittikoferae (males only) by having Dgl- 4 closer to the dorsal edge (dorsal width / distance between Dgl- 4 = 1.2 - 1.3 in T. pearsoni, 1.6 - 1.7 in T. bittikoferae); more elongate pedipalp tibiae (length / width = 3.1 - 3.3 in T. pearsoni, 2.7 - 2.8 in T. bittikoferae); and a more elongate rostrum (length / width = 2.1 - 2.4 in T. pearsoni, 1.8 - 1.9 in T. bittikoferae). T. pearsoni can be differentiated from T. dimorpha by having an unmodified dorsal plate (T. dimorpha has a dorsal plate medial extension covering nearly half the length of the anterio-medial platelets) and by males having unmodified pedipalps (male T. dimorpha have large, highly modified pedipalps which are expanded vertically and laterally).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
7C5962D67BEEE4C829749B99BAE0F827.taxon	description	Description. Female (Figure 185) (n = 5) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (620 - 730 (620) long; 455 - 500 (455) wide) ellipsoid with faint pink coloration without a distinct pattern. Anterio-medial platelets (123.75 - 142.5 (123.75) long; 62.5 - 70 (62.5) wide). Anterio-lateral platelets (167.5 - 195 (167.5) long; 70 - 80 (72.5) wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 365 - 395 (365 )). Dorsal plate proportions: dorsum length / width 1.36 - 1.47 (1.36); dorsal width / distance between Dgl- 4 1.21 - 1.30 (1.25); anterio-medial platelet length / width 1.93 - 2.04 (1.98); anterio-lateral platelet length / width 2.31 - 2.64 (2.31); anterio-lateral / anterio-medial length 1.34 - 1.42 (1.35). Gnathosoma - Subcapitulum (247.5 - 290 (247.5) long (ventral); 170 - 210 (171) long (dorsal); 120 - 142.5 (120) tall) colorless. Rostrum (92.5 - 112.5 (92.5) long; 45 - 51.25 (45) wide) short and conical. Chelicerae (220 - 274 (221) long) with curved fangs (53 - 75 (64) long). Subcapitular proportions: ventral length / height 2.04 - 2.12 (2.06); rostrum length / width 2.06 - 2.25 (2.06). Pedipalps with tuberculate ventral extensions on femora and flat, flanged extensions on genua. Palpomeres: trochanter (37.5 - 50 (37.5) long); femur (96.25 - 120 (96.25) long); genu (57.5 - 75 (57.5) long); tibia (75 - 90 (75) long; 23.75 - 27.5 (23.75) wide); tarsus (20 - 25 (20) long). Palpomere proportions: femur / genu 1.60 - 1.69 (1.67); tibia / femur 0.70 - 0.80 (0.78); tibia length / width 3.00 - 3.27 (3.16). Venter - (742 - 802.75 (743) long; 505 - 610 (506) wide) colorless. Gnathosomal bay (117.5 - 145 (132.5) long; 80 - 100 (80) wide). Cxgl- 4 subapical. Medial suture (27.5 - 42.5 (30) long). Genital plates (180 - 187.5 (182.5) long; 147.5 - 155 (147.5) wide). Additional measurements: Cx- 1 (235 - 282 (260) long (total); 130 - 138 (137) long (medial )); Cx- 3 (335 - 420 (336) wide); anterior venter (180 - 200 (187.5) long). Ventral proportions: gnathosomal bay length / width 1.26 - 1.69 (1.66); anterior venter / genital field length 1.00 - 1.07 (1.03); anterior venter length / genital field width 1.18 - 1.33 (1.27); anterior venter / medial suture 4.53 - 6.55 (6.25). Male (Figure 186) (n = 5) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (620 - 675 (620) long; 430 - 495 (430) wide) ellipsoid with faint pink coloration without a distinct pattern. Anterio-medial platelets (117.5 - 127.5 (117.5) long; 62.5 - 75 (62.5) wide). Anterio-lateral platelets (172.5 - 195 (188.75) long; 67.5 - 82.5 (70) wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 355 - 410 (360 )). Dorsal plate proportions: dorsum length / width 1.34 - 1.44 (1.44); dorsal width / distance between Dgl- 4 1.19 - 1.24 (1.19); anterio-medial platelet length / width 1.70 - 2.04 (1.88); anterio-lateral platelet length / width 2.36 - 2.70 (2.70); anterio-lateral / anterio-medial length 1.35 - 1.61 (1.61). Gnathosoma - Subcapitulum (250 - 260 (260) long (ventral); 170 - 187 (186) long (dorsal); 112.5 - 117.5 (112.5) tall) colorless. Rostrum (92.5 - 102.5 (102.5) long; 42.5 - 46.25 (42.5) wide) short and conical. Chelicerae (225 - 244 (239) long) with curved fangs (55 - 63 (63) long). Subcapitular proportions: ventral length / height 2.20 - 2.31 (2.31); rostrum length / width 2.11 - 2.41 (2.41). Pedipalps with tuberculate ventral extensions on femora and flat, flanged extensions on genua. Palpomeres: trochanter (40 - 42.5 (42.5) long); femur (100 - 106.25 (106.25) long); genu (60 - 67.5 (67.5) long); tibia (75 - 85 (75) long; 23.75 - 26.25 (23.75) wide); tarsus (22.5 - 22.5 (22.5) long). Palpomere proportions: femur / genu 1.54 - 1.67 (1.57); tibia / femur 0.71 - 0.83 (0.71); tibia length / width 3.10 - 3.24 (3.16). Venter - (720 - 800 (720) long; 461 - 588 (461) wide) colorless. Gnathosomal bay (110 - 130 (125) long; 70 - 87.5 (70) wide). Cxgl- 4 subapical. Medial suture (107.5 - 135 (107.5) long). Genital plates (140 - 155 (152.5) long; 95 - 105 (95) wide). Additional measurements: Cx- 1 (276 - 302 (283) long (total); 140 - 167 (140) long (medial )); Cx- 3 (334 - 414 (334) wide); anterior venter (285 - 335 (285) long). Ventral proportions: gnathosomal bay length / width 1.26 - 1.79 (1.79); anterior venter / genital field length 1.87 - 2.18 (1.87); anterior venter length / genital field width 2.93 - 3.19 (3.00); anterior venter / medial suture 2.30 - 2.85 (2.65). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
7C5962D67BEEE4C829749B99BAE0F827.taxon	etymology	Etymology. Specific epithet (pearsoni) named in honor of Pearson Dowling, son of APGD, for his unquenchable curiosity for all things big and small.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
7C5962D67BEEE4C829749B99BAE0F827.taxon	distribution	Distribution. Interior Highlands of Missouri and Arkansas (Figure 184).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
941EF75F7EDC7A112B853401AB21E38E.taxon	materials_examined	Material examined. HOLOTYPE (♀): from USA, Maine, Washington County, Old Stream, off Route 9, 5.5 km west of Route 192 at Wesley, 6 June 2012, by IM Smith, IMS 120012. PARATYPES (1 ♀; 1 ♂): Maine, USA: 1 ♂ (ALLOTYPE) from Washington County, Old Stream, off Route 9, 5.5 km west of Route 192 at Wesley, 6 June 2012, by IM Smith, IMS 120012 * 1 ♀ from Washington County, Old Stream, off Route 9, 5.5 km west of Route 192 at Wesley, 6 June 2012, by IM Smith, IMS 120012.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
941EF75F7EDC7A112B853401AB21E38E.taxon	diagnosis	Diagnosis. Torrenticola pendula are similar to other members of the Rusetria " Eastern 2 - Plates " group (T. biscutella, T. caerulea, T. delicatexa, T. indistincta, T. malarkeyorum, T. sellersorum, T. tysoni, T. ululata, T. whitneyae, T. microbiscutella, T. feminellai) in having anterio-lateral platelets fused to the dorsal plate, having dorsal coloration separated into anterior and posterior portions (except T. ululata and T. indistincta), and being distributed in the east. It is one of only four Eastern 2 - Plates that have dark, bold, bluish-purple coloration (also T. tysoni, T. biscutella, and T. sellersorum). T. pendula can be differentiated from other Eastern 2 - Plates (except T. whitneyae and T. feminellai) by having a more elongate gnathosomal bay (2.42 - 2.9 in T. pendula, 1.4 - 2.24 in others) and often by having a dorsal pattern of dark bluish-purple separated into anterior and posterior portions connected medially. In the Rusetria Complex only T. whitneyae and T. magnexa (rarely) has a similar pattern. T. pendula can be differentiated from T. magnexa by having anterio-medial platelets fully fused to the dorsal plate (anterio-lateral platelets partially fused to the dorsal plate in T. magnexa). T. pendula can be differentiated from T. whitneyae by having a more elongate rostrum (length / width = 2.87 - 3.06 in T. pendula, 2.41 - 2.69 in T. whitneyae). T. pendula can be differentiated from T. feminellai by having less elongate pedipalpal tibiae (length / width = 2.78 - 3.05 in T. pendula; 3.63 - 4.11 in T. feminellai).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
941EF75F7EDC7A112B853401AB21E38E.taxon	description	Description. Female (Figure 188) (n = 2) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (630 - 650 (630) long; 490 - 490 (490) wide) ovoid with bold purple coloration both anteriorly and posteriorly connected medially. Anterio-medial platelets (145 - 145 (145) long; 45 - 47.5 (47.5) wide). Anterio-lateral platelets (170 - 177.5 (170) long; 62.5 - 75 (62.5) wide) fused to dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 350 - 350 (350 )). Dorsal plate proportions: dorsum length / width 1.29 - 1.33 (1.29); dorsal width / distance between Dgl- 4 1.40 - 1.40 (1.40); anterio-medial platelet length / width 3.05 - 3.22 (3.05); anterio-lateral platelet length / width 2.37 - 2.72 (2.72); anterio-lateral / anterio-medial length 1.17 - 1.22 (1.17). Gnathosoma - Subcapitulum (320 - 335 (335) long (ventral); 242 - 252 (252) long (dorsal); 147.5 - 147.5 (147.5) tall) colorless. Rostrum (127.5 - 137.5 (137.5) long; 42.5 - 45 (45) wide). Chelicerae (336 - 347 (347) long) with curved fangs (53 - 62 (62) long). Subcapitular proportions: ventral length / height 2.17 - 2.27 (2.27); rostrum length / width 3.00 - 3.06 (3.06). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (45 - 47.5 (47.5) long); femur (102.5 - 107.5 (102.5) long); genu (67.5 - 71.25 (67.5) long); tibia (72.5 - 75 (75) long; 23.75 - 25 (25) wide); tarsus (17.5 - 17.5 (17.5) long). Palpomere proportions: femur / genu 1.51 - 1.52 (1.52); tibia / femur 0.67 - 0.73 (0.73); tibia length / width 3.00 - 3.05 (3.00). Venter - (770 - 800 (770) long; 532 - 557 (557) wide) with bold purple coloration. Gnathosomal bay (187.5 - 195 (195) long; 75 - 77.5 (75) wide). Cxgl- 4 subapical. Medial suture (10 - 12.5 (12.5) long). Genital plates (180 - 186.25 (186.25) long; 168.75 - 172.5 (168.75) wide). Additional measurements: Cx- 1 (290 - 301 (290) long (total); 110 - 115 (115) long (medial )); Cx- 3 (345 - 350 (346) wide); anterior venter (145 - 157.5 (145) long). Ventral proportions: gnathosomal bay length / width 2.42 - 2.60 (2.60); anterior venter / genital field length 0.78 - 0.88 (0.78); anterior venter length / genital field width 0.86 - 0.91 (0.86); anterior venter / medial suture 11.60 - 15.75 (11.60). Male (Figure 189) (n = 1) (allotype only) with characters of the genus with following specifications. Dorsum - (500 long; 380 wide) ovoid with bold purple coloration separated into anterior and posterior portions. Anterio-medial platelets (130 long; 41.25 wide). Anterio-lateral platelets (155 long; 62.5 wide) fused to dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 275). Dorsal plate proportions: dorsum length / width 1.32; dorsal width / distance between Dgl- 4 1.38; anterio-medial platelet length / width 3.15; anterio-lateral platelet length / width 2.48; anterio-lateral / anterio-medial length 1.19. Gnathosoma - Subcapitulum (272.5 long (ventral); 197.5 long (dorsal); 110 tall) colorless. Rostrum (107.5 long; 37.5 wide). Chelicerae (280 long) with curved fangs 50 long). Subcapitular proportions: ventral length / height 2.48; rostrum length / width 2.87. Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (38.75 long); femur (85 long); genu (58.75 long); tibia (62.5 long; 22.5 wide); tarsus (15 long). Palpomere proportions: femur / genu 1.45; tibia / femur 0.74; tibia length / width 2.78. Venter - (620 long; 430 wide) with faint purple coloration. Gnathosomal bay (152.5 long; 52.5 wide). Medial suture (72.5 long). Cxgl- 4 subapical. Genital plates (126.25 long; 120 wide). Additional measurements: Cx- 1 (250 long (total); 100 long (medial )); Cx- 3 (300 wide); anterior venter (207.5 long). Ventral proportions: gnathosomal bay length / width 2.90; anterior venter / genital field length 1.64; anterior venter length / genital field width 1.73; anterior venter / medial suture 2.86. Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
941EF75F7EDC7A112B853401AB21E38E.taxon	etymology	Etymology. Specific epithet (pendula) named for the swinging torture axe described in " The Pit and the Pendulum " by Edgar Allan Poe, most artistic depictions of which resemble the dorsal patterning on this species.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
941EF75F7EDC7A112B853401AB21E38E.taxon	distribution	Distribution. Known only from Washington County, Maine (Figure 187).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
9A1C01BC0830104E3913D2D9D6D09574.taxon	materials_examined	Material examined. HOLOTYPE (♀): from USA, California, Mendocino County, beside Rt. 128, 7.3 km south of Boonville, (38 ° 57 ' 57 " N, 123 ° 19 ' 19 " W), 4 August 1987, by IM Smith, IMS 870126 A PARATYPES (3 ♀; 2 ♂): California, USA: 1 ♂ (ALLOTYPE) from Mendocino County, beside Rt. 128, 7.3 km south of Boonville, (38 ° 57 ' 57 " N, 123 ° 19 ' 19 " W), 4 August 1987, by IM Smith, IMS 870126 A * 1 ♀ from Trinity County, beside Rt. 36 at Forest Glen Station Campground, (40 ° 23 ' 23 " N, 123 ° 20 ' 20 " W), 6 August 1987, by IM Smith & JD Smith, IMS 870131 * Oregon, USA: 1 ♀ and 1 ♂ from Curry County, Port Orford; Butler Bar Campground off Elk River Road, (42 ° 43 ' 43 " N, 124 ° 16 ' 16 " W), 25 June 1976, by IM Smith, IMS 760162 * 1 ♀ from Curry County, Sixes; Edson Creek Campground beside Sixes River Road, (42 ° 48 ' 48 " N, 124 ° 24 ' 24 " W), 4 July 1983, by IM Smith, IMS 830021 A	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
9A1C01BC0830104E3913D2D9D6D09574.taxon	diagnosis	Diagnosis. Torrenticola pinocchio are similar to members of the Miniforma group (T. copipalpa, T. manni, T. miniforma, T. pacificensis, T. rockyensis, and T. oliveri) in having similar pedipalpal extensions (unique to members of this group); and being among the smallest Torrenticola in the west (dorsum 500 - 625 long) (except T. oliveri). T. pinocchio can be differentiated from all other Torrenticola by having a more elongate rostrum (length / width = 4.58 - 3.91 in T. pinocchio, 1.29 - 4.40 in others).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
9A1C01BC0830104E3913D2D9D6D09574.taxon	description	Description. Female (Figure 191) (n = 4) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (580 - 610 (580) long; 360 - 400 (370) wide) ellipsoid and colorless. Anterio-medial platelets (95 - 115 (102.5) long; 47.5 - 50 (47.5) wide). Anterio-lateral platelets (150 - 165 (150) long; 55 - 65 (60) wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 280 - 310 (295 )). Dorsal plate proportions: dorsum length / width 1.53 - 1.64 (1.57); dorsal width / distance between Dgl- 4 1.25 - 1.30 (1.25); anterio-medial platelet length / width 1.90 - 2.30 (2.16); anterio-lateral platelet length / width 2.50 - 2.82 (2.50); anterio-lateral / anterio-medial length 1.43 - 1.63 (1.46). Gnathosoma - Subcapitulum (300 - 310 (305) long (ventral); 230 - 240 (230) long (dorsal); 100 - 107.5 (107.5) tall) colorless. Rostrum (137.5 - 145 (140) long; 30 - 30 (30) wide). Chelicerae (290 - 305 (300) long) with curved fangs (40 - 45 (45) long). Subcapitular proportions: ventral length / height 2.84 - 3.05 (2.84); rostrum length / width 4.58 - 4.83 (4.67). Pedipalps with broad, dentate, and anteriorly-directed ventral extensions on femora and dentate, flanged ventral extensions on genua. Palpomeres: trochanter (30 - 30 (30) long); femur (97.5 - 102.5 (97.5) long); genu (67.5 - 71.25 (67.5) long); tibia (65 - 70 (70) long; 20 - 20 (20) wide); tarsus (15 - 17.5 (17.5) long). Palpomere proportions: femur / genu 1.39 - 1.45 (1.44); tibia / femur 0.65 - 0.72 (0.72); tibia length / width 3.25 - 3.50 (3.50). Venter - Subcapitulum (720 - 755 (730) long; 400 - 450 (410) wide) colorless. Gnathosomal bay (140 - 145 (145) long; 55 - 60 (60) wide). Cxgl- 4 subapical. Medial suture (52.5 - 55 (55) long). Genital plates (155 - 167.5 (160) long; 135 - 150 (140) wide). Additional measurements: Cx- 1 (260 - 270 (270) long (total); 120 - 130 (130) long (medial )); Cx- 3 (52.5 - 55 (55) wide); anterior venter (180 - 195 (192.5) long). Ventral proportions: gnathosomal bay length / width 2.42 - 2.59 (2.42); anterior venter / genital field length 1.16 - 1.20 (1.20); anterior venter length / genital field width 1.30 - 1.41 (1.38); anterior venter / medial suture 3.43 - 3.71 (3.50). Male (Figure 192) (n = 2) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (520 - 540 (520) long; 340 - 350 (340) wide) ellipsoid and colorless. Anterio-medial platelets (95 - 102.5 (95) long; 45 - 47.5 (45) wide). Anterio-lateral platelets (145 - 155 (145) long; 50 - 57.5 (50) wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 260 - 270 (260 )). Dorsal plate proportions: dorsum length / width 1.53 - 1.54 (1.53); dorsal width / distance between Dgl- 4 1.30 - 1.31 (1.31); anterio-medial platelet length / width 2.11 - 2.16 (2.11); anterio-lateral platelet length / width 2.70 - 2.90 (2.90); anterio-lateral / anterio-medial length 1.51 - 1.53 (1.53). Gnathosoma - Subcapitulum (275 - 290 (275) long (ventral); 215 - 225 (215) long (dorsal); 90 - 95 (95) tall) colorless. Rostrum (130 - 135 (130) long; 27.5 - 27.5 (27.5) wide). Chelicerae (( 265) long) with curved fangs (( 45) long). Subcapitular proportions: ventral length / height 2.89 - 3.22 (2.89); rostrum length / width 4.73 - 4.91 (4.73). Pedipalps with broad, dentate, and anteriorly-directed ventral extensions on femora and dentate, flanged ventral extensions on genua. Palpomeres: trochanter (30 - 32.5 (30) long); femur (90 - 96.25 (90) long); genu (67.5 - 67.5 (67.5) long); tibia (62.5 - 65 (65) long; 18.75 - 20 (18.75) wide); tarsus (15 - 15 (15) long). Palpomere proportions: femur / genu 1.33 - 1.43 (1.33); tibia / femur 0.65 - 0.72 (0.72); tibia length / width 3.13 - 3.47 (3.47). Venter - (650 - 670 (650) long; 385 - 410 (385) wide) colorless. Gnathosomal bay (120 - 125 (110) long; 50 - 52.5 (50) wide). Cxgl- 4 subapical. Medial suture (80 - 85 (80) long). Genital plates (117.5 - 125 (117.5) long; 97.5 - 100 (97.5) wide). Additional measurements: Cx- 1 (240 - 240 (240) long (total); 110 - 125 (110) long (medial )); Cx- 3 (260 - 280 (260) wide); anterior venter (210 - 225 (210) long). Ventral proportions: gnathosomal bay length / width 2.29 - 2.50 (2.50); anterior venter / genital field length 1.79 - 1.80 (1.79); anterior venter length / genital field width 2.15 - 2.25 (2.15); anterior venter / medial suture 2.63 - 2.65 (2.63). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
9A1C01BC0830104E3913D2D9D6D09574.taxon	etymology	Etymology. Specific epithet (pinocchio) refers to the elongate rostrum of this species, which resembles the ever-growing nose of Pinocchio (noun in apposition), a fictional character from the popular Italian children's novel, The Adventures of Pinocchio by Carlo Collodi (1883). Pinocchio is well-known for his nose, which grows every time he tells a lie.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
9A1C01BC0830104E3913D2D9D6D09574.taxon	distribution	Distribution. California and Oregon (Figure 190).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
E7A3FB77D53D20D4A9BEABB0E8412FD9.taxon	materials_examined	Material examined. HOLOTYPE (♀): from USA, Alabama, Lauderdale County, off Natchez Parkway, 7 km south of Tennessee state line (34 ° 56 ' 31 " N, 87 ° 49 ' 41 " W), 27 Sep 2010, by IM Smith, IMS 100162, DNA 1288. PARATYPES (4 ♀; 11 ♂): Alabama, USA: 2 ♂ from Clay County, beside Forest Route 649, 0.8 kilometers northeast of road from Forest Route 600 to Campbell Springs, Talladega Creek, 2 Jul 1990, by IM Smith, IMS 900075 A * 1 ♂ from Cleburne County, beside Route 431, 3.3 kilometers southeast of Calhoun, Jackson Creek (33 ° 36 ' N, 85 ° 42 ' W), 2 Jul 1990, by IM Smith, IMS 900074 * 1 ♂ (ALLOTYPE) from Lauderdale County, off Natchez Parkway, 7 km south of Tennessee state line (34 ° 56 ' 31 " N, 87 ° 49 ' 41 " W), 27 Sep 2010, by IM Smith, IMS 100162 * 1 ♀ and 1 ♂ from Lauderdale County, off Natchez Parkway, 7 km south of Tennessee state line (34 ° 56 ' 31 " N, 87 ° 49 ' 41 " W), 24 Sep 2009, by IM Smith, IMS 090121 * 3 ♀ and 2 ♂ from Lauderdale County, off Natchez Parkway, 7 km south of Tennessee state line (34 ° 56 ' 31 " N, 87 ° 49 ' 41 " W), 27 Sep 2010, by IM Smith, IMS 100162 * Georgia, USA: 1 ♂ from Floyd County, beside road from Everett Springs to Villanow, 1.8 kilometers south of The Pocket Campground, Johns Creek, 4 Jul 1990, by IM Smith, IMS 900076 * 1 ♂ from Floyd County, beside road from Everett Springs to Villanow, 1.4 kilometers south of The Pocket Campground, tributary of Johns Creek, 4 Jul 1990, by IM Smith, IMS 900076 * 2 ♂ from Floyd County, The Pocket Campground, between Everett Springs and Villanow, tributary of Johns Creek, 4 Jul 1990, by IM Smith, IMS 900073 A * 1 ♂ from White County, Helen, beside Road to Anna Ruby Falls, Smith Creek (34 ° 44 ' N, 83 ° 43 ' W), 24 Sep 1992, by IM Smith, IMS 920051.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
E7A3FB77D53D20D4A9BEABB0E8412FD9.taxon	diagnosis	Diagnosis. Torrenticola pollani are similar to other members of the Rusetria " 4 - Plates " group (T. dunni, T. glomerabilis, T. kittatinniana, T. rufoalba, and T. shubini) and T. skvarlai in having anterio-lateral platelets free from the dorsal plate, dorsal coloration separated into anterior and posterior portions, and indistinct hind coxal margins. T. pollani can be differentiated from T. dunni by having a smaller dorsum (length ♀ = 535 - 560 in T. pollani, 605 - 680 in T. dunni; ♂ = 440 - 490 in T. pollani, 500 - 540 in T. dunni; width ♀, 410 - 420 in T. pollani, 440 - 490 in T. dunni; ♂ = 350 - 370 in A 38, 310 - 340 in T. pollani); and a more elongate rostrum (length / width = 3.3 - 3.8 in T. pollani, 2.8 - 3.1 in T. dunni). T. pollani can be differentiated from T. shubini by having more elongate tibiae (length / width ♀ = 4.00 - 4.18 in T. pollani, 3.35 - 3.60 in T. shubini; ♂ = 3.44 - 3.75 in T. pollani, 3.11 - 3.22 in T. shubini) and a more elongate rostrum (length / width = 3.27 - 3.82 in T. pollani, 2.24 - 2.85 in T. shubini). T. pollani can be differentiated from T. glomerabilis by having more elongate anterio-medial platelets (length / width ♀ = 2.5 - 3.0 in T. pollani, 1.9 - 2.3 in T. glomerabilis; ♂ = 2.3 - 2.5 in T. pollani, 1.9 - 2.2 in T. glomerabilis) and thinner dorsum (♀ = 410 - 420 in T. pollani, 460 - 490 in T. glomerabilis; ♂ = 310 - 340 in T. pollani, 395 - 430 in T. glomerabilis). T. pollani can be differentiated from T. kittatinniana by having a more elongate rostrum (length / width = 3.27 - 3.82 in T. pollani, 2.71 - 3.16 in T. kittatinniana) and more elongate tibiae (length / width ♀ = 3.8 - 4.2 in T. pollani, 3.3 in T. kittatinniana; ♂ = 3.44 - 3.75 in T. pollani, 2.80 in T. kittatinniana). T. pollani can be differentiated from T. rufoalba by having a more elongate rostrum (length / width = 3.27 - 3.82 in T. pollani, 2.96 - 3.06 in T. rufoalba). Female T. pollani can be differentiated from female T. rufoalba by having more elongate tibiae (length / width = 3.8 - 4.2 in T. pollani, 3.5 in T. rufoalba). Male T. pollani can be differentiated from male T. rufoalba by having a longer anterior venter (235 - 250 in T. pollani, 195 in T. rufoalba). T. pollani can be differentiated from T. skvarlai by having a conical pedipalpal femoral tubercle, whereas T. skvarlai has a broad and flat pedipalpal femoral tubercle, and by having a longer anterior venter (♀ = 155 - 163 in T. pollani, 140 - 153 in T. skvarlai; ♂ = 235 - 250 in T. pollani, 177.5 - 205 in T. skvarlai).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
E7A3FB77D53D20D4A9BEABB0E8412FD9.taxon	description	Description. Female (Figure 194) (n = 5) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (535 - 560 (550) long; 410 - 420 (415) wide) ovoid with purple to bluish-purple coloration separated into anterior and posterior portions, and occasionally with faint strip of orange medially. Anterio-medial platelets (105 - 125 (105) long; 40 - 47.5 (40) wide). Anterio-lateral platelets (152.5 - 170 (152.5) long; 52.5 - 62.5 (52.5) wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 270 - 290 (270 )). Dorsal plate proportions: dorsum length / width 1.29 - 1.37 (1.33); dorsal width / distance between Dgl- 4 1.45 - 1.54 (1.54); anterio-medial platelet length / width 2.50 - 3.03 (2.63); anterio-lateral platelet length / width 2.64 - 2.93 (2.90); anterio-lateral / anterio-medial length 1.34 - 1.47 (1.45). Gnathosoma - Subcapitulum (310 - 332.5 (310) long (ventral); 236 - 257.5 (237) long (dorsal); 122.5 - 132.5 (122.5) tall) colorless. Rostrum (133.75 - 150 (133.75) long; 35 - 41.25 (35) wide). Chelicerae (310 - 340 (315) long) with curved fangs (53 - 60 (56) long). Subcapitular proportions: ventral length / height 2.43 - 2.61 (2.53); rostrum length / width 3.27 - 3.82 (3.82). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (37.5 - 42.5 (40) long); femur (115 - 127.5 (117.5) long); genu (65 - 70 (65) long); tibia (80 - 92.5 (80) long; 20 - 22.5 (20) wide); tarsus (17.5 - 20 (17.5) long). Palpomere proportions: femur / genu 1.70 - 1.92 (1.81); tibia / femur 0.68 - 0.78 (0.68); tibia length / width 3.89 - 4.18 (4.00). Venter - (610 - 675 (675) long; 461 - 490 (489) wide) colorless. Gnathosomal bay (142.5 - 170 (142.5) long; 77.5 - 97.5 (87.5) wide). Cxgl- 4 subapical. Medial suture (12.5 - 20 (17.5) long). Genital plates (157.5 - 175 (157.5) long; 137.5 - 152.5 (137.5) wide). Additional measurements: Cx- 1 (258 - 290 (259) long (total); 89 - 121 (96) long (medial )); Cx- 3 (304 - 364 (310) wide); anterior venter (155 - 162.5 (157.5) long). Ventral proportions: gnathosomal bay length / width 1.63 - 2.13 (1.63); anterior venter / genital field length 0.89 - 1.02 (1.00); anterior venter length / genital field width 1.02 - 1.15 (1.15); anterior venter / medial suture 7.75 - 12.70 (9.00). Male (Figure 195) (n = 5) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (440 - 490 (450) long; 310 - 340 (315) wide) ovoid with purple to bluish-purple coloration separated into anterior and posterior portions, and occasionally with faint strip of orange medially. Anterio-medial platelets (92.5 - 102.5 (100) long; 40 - 42.5 (42.5) wide). Anterio-lateral platelets (140 - 155 (142.5) long; 42.5 - 50 (45) wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 210 - 250 (215 )). Dorsal plate proportions: dorsum length / width 1.33 - 1.44 (1.43); dorsal width / distance between Dgl- 4 1.32 - 1.48 (1.47); anterio-medial platelet length / width 2.31 - 2.44 (2.35); anterio-lateral platelet length / width 3.10 - 3.29 (3.17); anterio-lateral / anterio-medial length 1.43 - 1.59 (1.43). Gnathosoma - Subcapitulum (265 - 285 (265) long (ventral); 202 - 208 (203) long (dorsal); 87.5 - 100 (97.5) tall) colorless. Rostrum (111.25 - 122.5 (111.25) long; 32.5 - 35 (32.5) wide). Chelicerae (257 - 280 (263) long) with curved fangs (45 - 50 (47) long). Subcapitular proportions: ventral length / height 2.72 - 3.06 (2.72); rostrum length / width 3.41 - 3.54 (3.42). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (35 - 40 (35) long); femur (100 - 103.75 (101.25) long); genu (60 - 62.5 (60) long); tibia (72.5 - 80 (75) long; 20 - 22.5 (20) wide); tarsus (15 - 17.5 (17.5) long). Palpomere proportions: femur / genu 1.64 - 1.73 (1.69); tibia / femur 0.70 - 0.78 (0.74); tibia length / width 3.41 - 3.75 (3.75). Venter - (540 - 600 (555) long; 358 - 408 (359) wide) colorless. Gnathosomal bay (95 - 127.5 (116.25) long; 65 - 77.5 (70) wide). Cxgl- 4 subapical. Medial suture (92.5 - 110 (93.75) long). Genital plates (105 - 120 (110) long; 80 - 90 (83.75) wide). Additional measurements: Cx- 1 (210 - 250 (246) long (total); 84 - 125 (111) long (medial )); Cx- 3 (266 - 300 (266) wide); anterior venter (235 - 250 (237.5) long). Ventral proportions: gnathosomal bay length / width 1.46 - 1.89 (1.66); anterior venter / genital field length 2.08 - 2.29 (2.16); anterior venter length / genital field width 2.78 - 3.03 (2.84); anterior venter / medial suture 2.27 - 2.54 (2.53). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
E7A3FB77D53D20D4A9BEABB0E8412FD9.taxon	etymology	Etymology. Specific epithet (pollani) named in honor of author Michael Pollan, whose influential books breach mere accounts on food culture and enter insightful discussions of human ecology.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
E7A3FB77D53D20D4A9BEABB0E8412FD9.taxon	distribution	Distribution. Southeastern (northern Alabama and Georgia) (Figure 193).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
E07A4763224520C247B312753E8EE6E5.taxon	materials_examined	Material examined. HOLOTYPE (♀): from USA, Texas, Tyler County, Spurger; beside Farm Road 1013, 8.2 km west of Rt. 92, (30 ° 41 ' 41 " N, 94 ° 15 ' 15 " W), 30 September 1994, by IM Smith, IMS 940027 A PARATYPES (5 ♀; 5 ♂): New Hampshire, USA: 4 ♀ and 1 ♂ from Woodstock County, beside Rt. 118, 3.2 km south of Rt. 112, (44 ° 0 ' 0 " N, 71 ° 45 ' 45 " W), 11 September 1992, by IM Smith, IMS 920036 * Texas, USA: 1 ♂ (ALLOTYPE) from Tyler County, Spurger; beside Farm Road 1013, 8.2 km west of Rt. 92, (30 ° 41 ' 41 " N, 94 ° 15 ' 15 " W), 30 September 1994, by IM Smith, IMS 940027 A * 1 ♀ and 3 ♂ from Tyler County, Spurger; beside Farm Road 1013, 8.2 km west of Rt. 92, (30 ° 41 ' 41 " N, 94 ° 15 ' 15 " W), 30 September 1994, by IM Smith, IMS 940027 A	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
E07A4763224520C247B312753E8EE6E5.taxon	diagnosis	Diagnosis. Torrenticola priapus are similar to other members of the Partial 2 - Plate Group (T. folkertsae, T. magnexa, and T. pulchra) in having anterio-lateral platelets partially fused to the dorsal plate and being distributed in the east. T. priapus can be differentiated from T. magnexa and T. pulchra by dorsal coloration and pattern. T. priapus can be further differentiated from T. magnexa by having a more elongate rostrum (length / width = 3.17 - 3.39 in T. priapus, 2.25 - 3.00 in T. magnexa). T. priapus can be further differentiated from T. pulchra by having a round dorsum (length / width = 1.18 - 1.22 in T. priapus, 1.4 - 1.61 in T. pulchra) and more elongate pedipalpal tibiae (length / width ♀ = 3.9 - 4.22 in T. priapus, 3.3 - 3.70 in T. pulchra, ♂ = 3.5 - 3.78 in T. priapus, 3.00 - 3.35 in T. pulchra). T. priapus can be differentiated from T. folkertsae by having less elongate pedipalpal tibiae (length / width ♀ = 3.9 - 4.22 in T. priapus, 4.5 - 4.83 in T. folkertsae, ♂ = 3.5 - 3.78 in T. priapus, 4.05 - 4.33 in T. folkertsae) and a more elongate rostrum (length / width = 3.17 - 3.39 in T. priapus, 2.55 - 3.00 in T. folkertsae).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
E07A4763224520C247B312753E8EE6E5.taxon	description	Description. Female (Figure 197) (n = 5) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (610 - 690 (650) long; 510 - 580 (540) wide) circular with purple coloration posteriorly and anteriorly connected medially but not extending to platelets. Anterio-medial platelets (150 - 170 (162.5) long; 62.5 - 70 (65) wide). Anterio-lateral platelets (175 - 190 (185) long; 75 - 95 (95) wide) partially fused to dorsal plate. Dgl- 4 nearly halfway between the edge of the dorsum and the muscle scars (distance between Dgl- 4 305 - 370 (345 )). Dorsal plate proportions: dorsum length / width 1.19 - 1.21 (1.20); dorsal width / distance between Dgl- 4 1.56 - 1.67 (1.57); anterio-medial platelet length / width 2.37 - 2.50 (2.50); anterio-lateral platelet length / width 1.95 - 2.33 (1.95); anterio-lateral / anterio-medial length 1.12 - 1.23 (1.14). Gnathosoma - Subcapitulum (325 - 355 (355) long (ventral); 245 - 270 (270) long (dorsal); 137.5 - 155 (150) tall) colorless. Rostrum (142.5 - 152.5 (152.5) long; 45 - 45 (45) wide). Chelicerae (330 - 370 (360) long) with curved fangs (55 - 65 (65) long). Subcapitular proportions: ventral length / height 2.28 - 2.37 (2.37); rostrum length / width 3.17 - 3.39 (3.39). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (42.5 - 50 (48.75) long); femur (118.75 - 135 (135) long); genu (70 - 80 (76.25) long); tibia (92.5 - 105 (97.5) long; 22.5 - 25 (25) wide); tarsus (17.5 - 22.5 (22.5) long). Palpomere proportions: femur / genu 1.69 - 1.77 (1.77); tibia / femur 0.72 - 0.84 (0.72); tibia length / width 3.90 - 4.22 (3.90). Venter - (720 - 850 (770) long; 590 - 645 (605) wide) colorless. Gnathosomal bay (160 - 192.5 (192.5) long; 77.5 - 90 (80) wide). Cxgl- 4 subapical. Medial suture (15 - 15 (15) long). Genital plates (180 - 190 (190) long; 167.5 - 185 (185) wide). Additional measurements: Cx- 1 (280 - 320 (315) long (total); 115 - 135 (130) long (medial )); Cx- 3 (355 - 390 (390) wide); anterior venter (150 - 165 (160) long). Ventral proportions: gnathosomal bay length / width 2.00 - 2.41 (2.41); anterior venter / genital field length 0.81 - 0.89 (0.84); anterior venter length / genital field width 0.86 - 0.97 (0.86); anterior venter / medial suture 10.00 - 11.00 (10.67). Male (Figure 198) (n = 4) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (530 - 585 (530) long; 435 - 495 (435) wide) circular with purple coloration posteriorly and anteriorly connected medially but not extending to platelets. Anterio-medial platelets (131.25 - 145 (131.25) long; 52.5 - 62.5 (55) wide). Anterio-lateral platelets (165 - 182.5 (170) long; 70 - 77.5 (70) wide) partially fused to dorsal plate. Dgl- 4 nearly halfway between the edge of the dorsum and the muscle scars (distance between Dgl- 4 265 - 330 (265 )). Dorsal plate proportions: dorsum length / width 1.18 - 1.22 (1.22); dorsal width / distance between Dgl- 4 1.50 - 1.70 (1.64); anterio-medial platelet length / width 2.32 - 2.76 (2.39); anterio-lateral platelet length / width 2.13 - 2.43 (2.43); anterio-lateral / anterio-medial length 1.14 - 1.30 (1.30). Gnathosoma - Subcapitulum (305 - 320 (305) long (ventral); 230 - 245 (235) long (dorsal); 125 - 130 (125) tall) colorless. Rostrum (127.5 - 140 (127.5) long; 40 - 42.5 (40) wide). Chelicerae (305 - 320 (305) long) with curved fangs (57.5 - 60 (60) long). Subcapitular proportions: ventral length / height 2.44 - 2.48 (2.44); rostrum length / width 3.19 - 3.29 (3.19). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (42.5 - 45 (42.5) long); femur (110 - 117.5 (110) long); genu (67.5 - 72.5 (67.5) long); tibia (85 - 93.75 (85) long; 22.5 - 25 (22.5) wide); tarsus (20 - 22.5 (20) long). Palpomere proportions: femur / genu 1.62 - 1.70 (1.63); tibia / femur 0.76 - 0.80 (0.77); tibia length / width 3.50 - 3.78 (3.78). Venter - (625 - 695 (625) long; 510 - 560 (540) wide) colorless. Gnathosomal bay (145 - 157.5 (145) long; 70 - 77.5 (70) wide). Cxgl- 4 subapical. Medial suture (75 - 80 (75) long). Genital plates (165 - 185 (165) long; 145 - 160 (150) wide). Additional measurements: Cx- 1 (270 - 305 (285) long (total); 120 - 140 (140) long (medial )); Cx- 3 (340 - 360 (350) wide); anterior venter (220 - 250 (230) long). Ventral proportions: gnathosomal bay length / width 2.00 - 2.25 (2.07); anterior venter / genital field length 1.33 - 1.42 (1.39); anterior venter length / genital field width 1.47 - 1.61 (1.53); anterior venter / medial suture 2.93 - 3.13 (3.07). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
E07A4763224520C247B312753E8EE6E5.taxon	etymology	Etymology. Specific epithet (priapus) is named for Priapus, the Greek fertility god who was marked by his oversized, permanent erection, which refers to the genital opening in the male-the largest of all North American Torrenticola.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
E07A4763224520C247B312753E8EE6E5.taxon	distribution	Distribution. Eastern (Figure 196). Distribution seemingly disjunct (Texas and New Hampshire).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
4708AA4A2B50799100A3E0E19606DF33.taxon	materials_examined	Material examined. LECTOTYPE (1 ♀): from USA, New York, Cayuga County, Moravia, brook, 22 May 1960, by H Habeeb, HH 600011. PARALECTOTYPE (1 ♂): from USA, New York, Cayuga County, Moravia, brook, 22 May 1960, by H Habeeb, HH 600011. OTHER MATERIAL (25 ♀; 27 ♂): Alabama, USA: 4 ♀ and 4 ♂ from Clay County, Talladega Creek, beside Forest Route 649, 0.8 kilometers northeast of road from Forest Route 600 to Campbell Springs, 2 Jul 1990, by IM Smith, IMS 900075 A * 1 ♂ from Cleburne County, beside Route 431, 3.3 kilometers southeast of Calhoun, Jackson Creek (33 ° 36 ' N, 85 ° 42 ' W), 2 Jul 1990, by IM Smith, IMS 900074 * 2 ♀ and 1 ♂ from Lauderdale County, off Natchez Trace Parkway (34 ° 56 ' 31 " N, 87 ° 49 ' 41 " W), 24 Sep 2009, by IM Smith, IMS 090121 * 1 ♂ from Lauderdale County, off Natchez Trace Parkway (34 ° 56 ' 31 " N, 87 ° 49 ' 41 " W), 27 Sep 2010, by IM Smith, IMS 100162 * Georgia, USA: 1 ♂ from Chattooga County, beside road from Everett Springs to Villanow, 1.4 kilometers south of Pocket Recreation Area, 4 Jul 1990, by IM Smith, IMS 900077 * 1 ♀ and 2 ♂ from Floyd County, tributary of Johns Creek, beside road from Everett Springs to Villanow, 1.4 kilometers south of The Pocket Campground, 4 Jul 1990, by IM Smith, IMS 900077 * New York, USA: 3 ♀ and 2 ♂ from Cayuga County, Dutch Hollow Brook, beside Route 38 A at Niles, 22 Jul 1990, by IM Smith, IMS 900113 A * 2 ♀ and 1 ♂ from Schuyler County, beside Town Line Road off Route 228, 0.6 kilometers south of Perry City, 21 Jul 1990, by IM Smith, IMS 900112 A * North Carolina, USA: 3 ♀ and 5 ♂ from Yancey County, South Toe River, Lost Cove Picnic Area on Forest Route 472, 2.8 kilometers south of Route 80, 28 Jun 1990, by IM Smith, IMS 900065 A * 2 ♂ from Yancey County, South Toe River (35 ° 45 ' 10 " N, 82 ° 12 ' 43 " W), 28 Jun 1990, by IM Smith, IMS 900065 A * Pennsylvania, USA: 1 ♀ from Fayette County, Dunbar Creek (39 ° 57 ' 50 " N, 79835 ' 3.70 " W), 10 Aug 2014, by MJ Skvarla, MS 14 - 0810 - 001 * 1 ♂ from Fayette County, Dunbar Creek (39 ° 56 ' 16.10 " N, 79 ° 35 ' 3.70 " W), 10 Aug 2014, by MJ Skvarla, MS 14 - 0810 - 002 * Tennessee, USA: 2 ♂ from Monroe County, Turkey Creek, beside Forest Route 35, 2.0 kilometers northeast of road from Route 165 to Miller Chapel Church, 5 Jul 1990, by IM Smith, IMS 900078 * 2 ♀ from Sevier County, Great Smoky Mountains National Park, Middle Prong Pigeon River (35 ° 43 ' 32 " N, 83 ° 24 ' 2 " W), 2 Sep 2009, by IM Smith, IMS 090093 * 3 ♀ and 1 ♂ from Sevier County, Great Smoky Mountains National Park, Sugarlands Nature Trail (35 ° 40 ' 47 " N, 83 ° 31 ' 51 " W), 10 Sep 2010, by IM Smith, IMS 100125 * Virginia, USA: 1 ♂ from Amherst County, Upper Otter Creek Overlook beside Blue Ridge, Otter Creek (37 ° 36 ' 57 " N, 79 ° 19 ' 27 " W), 7 Sep 2007, by IM Smith, IMS 070056 A * 4 ♀ and 2 ♂ from Washington County, Damascus, Laurel River, beside Route 58 near boundary of Mount Rogers National Recreation Area, 10 Jul 1990, by IM Smith, IMS 900085 A.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
4708AA4A2B50799100A3E0E19606DF33.taxon	diagnosis	Diagnosis. Torrenticola projector are unlike nearly all other Torrenticola in having such elongate bodies and subcapitula. T. elongata are also elongate, but have different dorsal coloration and do not have an elongate subcapitulum. T. tahoei also have elongate subcapitula, but have rounder bodies with different dorsal coloration. T. anoplopalpa also have elongate subcapitula, but have rounder bodies and incomplete hind coxae.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
4708AA4A2B50799100A3E0E19606DF33.taxon	description	Re-description. Female (Figure 200) (n = 5) with characters of the genus with following specifications. Dorsum - (625 - 745 long; 400 - 470 wide) ovoid and elongate with pink coloration without distinct pattern. Anterio-medial platelets (95 - 115 long; 55 - 62.5 wide). Anterio-lateral platelets (185 - 217.5 long; 50 - 57.5 wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 320 - 370). Dorsal plate proportions: dorsum length / width 1.52 - 1.68; dorsal width / distance between Dgl- 4 1.24 - 1.28; anterio-medial platelet length / width 1.73 - 2.05; anterio-lateral platelet length / width 3.36 - 3.90; anterio-lateral / anterio-medial length 1.76 - 1.95. Gnathosoma - Subcapitulum (382.5 - 445 long (ventral); 300 - 333 long (dorsal); 87.5 - 97.5 tall) colorless and elongate. Rostrum (142.5 - 172.5 long; 37.5 - 42.5 wide) elongate. Chelicerae (360 - 428 long) with curved fangs (51 - 62 long). Subcapitular proportions: ventral length / height 4.03 - 4.78; rostrum length / width 3.80 - 4.27. Pedipalps short and stocky (especially tibiae) without extensions on femora and genua. Palpomeres: trochanter (47.5 - 55 long); femur (108.75 - 125 long); genu (60 - 70 long); tibia (21.25 - 25 long; 21.25 - 25 wide); tarsus (20 - 22.5 long). Palpomere proportions: femur / genu 1.74 - 1.81; tibia / femur 0.55 - 0.60; tibia length / width 2.82 - 3.11. Venter - (830 - 931 long; 480 - 544 wide) colorless. Gnathosomal bay (92.5 - 110 long; 67.5 - 80 wide). Cxgl- 4 far from apex. Medial suture (50 - 77.5 long). Genital plates 183.75 - 188.75 long; 147.5 - 160 wide). Additional measurements: Cx- 1 (296.75 - 357 long (total); 215 - 250 long (medial )); Cx- 3 (310 - 374 wide); anterior venter (285 - 340 long). Ventral proportions: gnathosomal bay length / width 1.22 - 1.63; anterior venter / genital field length 1.55 - 1.84; anterior venter length / genital field width 1.78 - 2.16; anterior venter / medial suture 4.39 - 6.05. Male (Figure 201) (n = 6) with characters of the genus with following specifications. Dorsum - (540 - 630 (600) long; 335 - 400 wide) ovoid and elongate with pink coloration without distinct pattern. Anterio-medial platelets (90 - 112.5 long; 47.5 - 57.5 wide). Anterio-lateral platelets (170 - 205 long; 47.5 - 57.5 wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 265 - 335). Dorsal plate proportions: dorsum length / width 1.54 - 1.61); dorsal width / distance between Dgl- 4 1.16 - 1.26; anterio-medial platelet length / width 1.70 - 2.20; anterio-lateral platelet length / width 3.23 - 3.68; anterio-lateral / anterio-medial length 1.61 - 2.00. Gnathosoma - Subcapitulum (325 - 420 long (ventral); 245 - 334 long (dorsal); 78.75 - 90 tall) colorless and elongate. Rostrum (127.5 - 158.75 long; 31.25 - 35 wide) elongate. Chelicerae (297 - 382 long) with curved fangs (41 - 61 long). Subcapitular proportions: ventral length / height 3.94 - 4.94; rostrum length / width 3.64 - 4.54. Pedipalps short and stocky (especially tibiae) without extensions on femora and genua. Palpomeres: trochanter (45 - 50 long); femur (100 - 113.75 long); genu (62.5 - 67.5 long); tibia (62.5 - 67.5 long; 22.5 - 22.5 wide); tarsus (17.5 - 20 long). Palpomere proportions: femur / genu 1.58 - 1.75; tibia / femur 0.59 - 0.64; tibia length / width 2.78 - 3.00. Venter - (680 - 860 long; 405 - 492 wide) colorless. Gnathosomal bay (82.5 - 132.5 long; 60 - 75 wide). Cxgl- 4 far from apex. Medial suture (127.5 - 142.5 long). Genital plates (137.5 - 157.5 long; 100 - 111.25 wide). Additional measurements: Cx- 1 (252 - 392 long (total); 175 - 267 long (medial )); Cx- 3 (282 - 355 wide); anterior venter (320 - 410 long). Ventral proportions: gnathosomal bay length / width 1.38 - 2.04; anterior venter / genital field length 2.27 - 2.64; anterior venter length / genital field width 3.20 - 3.71; anterior venter / medial suture 2.39 - 3.15. Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
4708AA4A2B50799100A3E0E19606DF33.taxon	etymology	Etymology. Habeeb (1961) did not specify an etymology for the specific epithet (projector). However, surely this name reflects the elongated, projectable gnathosoma (proiectus, L. stick out)	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
4708AA4A2B50799100A3E0E19606DF33.taxon	distribution	Distribution. Appalachians (Figure 199). T. projector was previously known only from New York; we extend its range throughout the Appalachians.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
6C3B2296181797A75883B2FF3178DB21.taxon	materials_examined	Material examined. HOLOTYPE (♀): from, USA, Illinois, Pope County, Eddyville; Bell Smith Springs Recreation Area; just below low water bridge at Hunting Branch Picnic Area, (37 ° 31 ' 31 " N, 88 ° 39 ' 39 " W), 1 September 2006, by IM Smith, S Yi, IMS 060057. PARATYPES (4 ♀; 5 ♂): Illinois, USA: 1 ♂ (ALLOTYPE) from Pope County, Eddyville; Bell Smith Springs Recreation Area; just below low water bridge at Hunting Branch Picnic Area, (37 ° 31 ' 31 " N, 88 ° 39 ' 39 " W), 1 September 2006, by IM Smith, S Yi, IMS 060057 * 3 ♀ and 3 ♂ from Pope County, Eddyville; Bell Smith Springs Recreation Area; just below low water bridge at Hunting Branch Picnic Area, (37 ° 31 ' 31 " N, 88 ° 39 ' 39 " W), 1 September 2006, by IM Smith, S Yi, IMS 060057 * 1 ♀ and 1 ♂ from Pope County, Eddyville; Bell Smith Springs Recreation Area; Hunting Branch Picnic Area, (37 ° 31 ' 31 " N, 88 ° 40 ' 40 " W), 9 September 1994, by IM Smith, IMS 940004	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
6C3B2296181797A75883B2FF3178DB21.taxon	diagnosis	Diagnosis. Torrenticola pulchra are similar to other members of the Partial 2 - Plate Group (T. folkertsae, T. magnexa, and T. priapus) in having anterio-lateral platelets partially fused to the dorsal plate and being distributed in the east. T. pulchra can be differentiated from other Partial 2 - Plate Group by dorsal coloration and pattern. T. pulchra can be further differentiated from T. priapus and T. folkertsae by having a more ovoid dorsum (length / width = 1.4 - 1.61 in T. pulchra, 1.11 - 1.29 in others) and less elongate pedipalpal tibiae (length / width ♀ = 3.3 - 3.70 in T. pulchra, 3.9 - 4.83 in others, ♂ = 3.00 - 3.35 in T. pulchra, 3.5 - 4.33 in others). Male T. pulchra can be further differentiated from male T. magnexa by having a smaller genital field (length ♂ = 110 - 123 in T. pulchra, 125 - 148 in T. magnexa; width ♂ = 87 - 95 in T. pulchra, 115 - 125 in T. magnexa,) and less elongate pedipalpal tibiae (length / width ♂ = 3.00 - 3.35 in T. pulchra, 3.78 - 4.00 in T. magnexa,). Female T. pulchra can be differentiated from female T. magnexa by having a thinner genital field (♀ 147 - 160 in T. pulchra, 170 - 188 in T. magnexa) and shorter pedipalpal tibiae (♀ 82 - 93 in T. pulchra, 102 - 113 in T. magnexa).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
6C3B2296181797A75883B2FF3178DB21.taxon	description	Description. Female (Figure 203) (n = 5) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (640 - 710 (710) long; 450 - 490 (480) wide) ovoid with bold blue coloration separated into anterior and posterior portions, posterior portion not meeting posterior limit of dorsal plate, and bold red medially. Anterio-medial platelets (130 - 145 (135) long; 50 - 55 (55) wide). Anterio-lateral platelets (177.5 - 200 (190) long; 60 - 70 (60) wide) nearly fused to dorsal plate posteriorly. Dgl- 4 closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 320 - 375 (355 )). Dorsal plate proportions: dorsum length / width 1.40 - 1.48 (1.48); dorsal width / distance between Dgl- 4 1.30 - 1.45 (1.35); anterio-medial platelet length / width 2.45 - 2.90 (2.45); anterio-lateral platelet length / width 2.77 - 3.17 (3.17); anterio-lateral / anterio-medial length 1.33 - 1.41 (1.41). Gnathosoma - Subcapitulum (295 - 345 (345) long (ventral); 215 - 252.5 (252.5) long (dorsal); 130 - 155 (155) tall) faint blue coloration. Rostrum (117.5 - 135 (135) long; 40 - 45 (45) wide). Chelicerae (295 - 350 (350) long) with curved fangs (60 - 70 (70) long). Subcapitular proportions: ventral length / height 2.03 - 2.33 (2.23); rostrum length / width 2.94 - 3.06 (3.00). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (47.5 - 52.5 (52.5) long); femur (118.75 - 132.5 (130) long); genu (67.5 - 75 (72.5) long); tibia (82.5 - 92.5 (90) long; 25 - 26.25 (25) wide); tarsus (17.5 - 20 (20) long). Palpomere proportions: femur / genu 1.73 - 1.80 (1.79); tibia / femur 0.69 - 0.72 (0.69); tibia length / width 3.30 - 3.70 (3.60). Venter - (740 - 840 (840) long; 530 - 560 (560) wide) with blue coloration. Gnathosomal bay (155 - 190 (190) long; 75 - 90 (75) wide). Cxgl- 4 subapical. Medial suture (10 - 20 (10) long). Genital plates (180 - 200 (192.5) long; 147.5 - 160 (155) wide). Additional measurements: Cx- 1 (280 - 350 (350) long (total); 130 - 162.5 (162.5) long (medial )); Cx- 3 (325 - 365 (360) wide); anterior venter (160 - 175 (175) long). Ventral proportions: gnathosomal bay length / width 1.94 - 2.53 (2.53); anterior venter / genital field length 0.80 - 0.96 (0.91); anterior venter length / genital field width 1.00 - 1.13 (1.13); anterior venter / medial suture 8.00 - 17.50 (17.50). Male (Figure 204) (n = 5) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (490 - 520 (500) long; 310 - 330 (315) wide ovoid with bold blue coloration separated into anterior and posterior portions, posterior portion not meeting posterior limit of dorsal plate, and bold red medially. Anterio-medial platelets (95 - 110 (100) long; 40 - 47.5 (42.5) wide). Anterio-lateral platelets (140 - 160 (145) long; 45 - 55 (50) wide) nearly fused to dorsal plate posteriorly. Dgl- 4 closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 235 - 265 (245 )). Dorsal plate proportions: dorsum length / width 1.53 - 1.61 (1.59); dorsal width / distance between Dgl- 4 1.23 - 1.36 (1.29); anterio-medial platelet length / width 2.24 - 2.50 (2.35); anterio-lateral platelet length / width 2.80 - 3.56 (2.90); anterio-lateral / anterio-medial length 1.45 - 1.55 (1.45). Gnathosoma - Subcapitulum (255 - 262.5 (255) long (ventral); 182.5 - 195 (182.5) long (dorsal); 90 - 95 (90) tall) faint blue coloration. Rostrum (97.5 - 105 (97.5) long; 30 - 32.5 (30) wide). Chelicerae (240 - 255 (245) long) with curved fangs (45 - 50 (45) long). Subcapitular proportions: ventral length / height 2.68 - 2.83 (2.83); rostrum length / width 3.23 - 3.50 (3.25). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (37.5 - 42.5 (37.5) long); femur (97.5 - 100 (100) long); genu (55 - 57.5 (57.5) long); tibia (67.5 - 72.5 (70) long; 21.25 - 22.5 (22.5) wide); tarsus (15 - 17.5 (15) long). Palpomere proportions: femur / genu 1.70 - 1.82 (1.74); tibia / femur 0.69 - 0.73 (0.70); tibia length / width 3.00 - 3.35 (3.11). Venter - (600 - 625 (600) long; 355 - 390 (355) wide) with blue coloration. Gnathosomal bay (120 - 130 (120) long; 60 - 67.5 (67.5) wide). Cxgl- 4 subapical. Medial suture (105 - 130 (125) long). Genital plates (110 - 122.5 (115) long; 87.5 - 95 (92.5) wide). Additional measurements: Cx- 1 (240 - 255 (240) long (total); 122.5 - 130 (125) long (medial )); Cx- 3 (275 - 295 (280) wide); anterior venter (235 - 270 (255) long). Ventral proportions: gnathosomal bay length / width 1.78 - 2.08 (1.78); anterior venter / genital field length 2.00 - 2.32 (2.22); anterior venter length / genital field width 2.61 - 2.91 (2.76); anterior venter / medial suture 2.04 - 2.24 (2.04). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
6C3B2296181797A75883B2FF3178DB21.taxon	etymology	Etymology. Specific epithet (pulchra) refers to the bright and distinctive coloration of this species (pulchra, L. beautiful).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
6C3B2296181797A75883B2FF3178DB21.taxon	distribution	Distribution. Southern Illinois (Figure 202).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
69615CB3F40CD6A66422478143FE8335.taxon	materials_examined	Material examined. HOLOTYPE (♀): from USA, Tennessee, Wayne County, Glenrock Branch Creek (35 ° 15 ' 50 " N, 87 ° 37 ' 34 " W), 24 Sep 2009, by IM Smith, IMS 090124, DNA 1867. PARATYPES (4 ♀; 2 ♂): Virginia, USA: 4 ♀ and 2 ♂ (ALLOTYPE) from Smyth County, Mount Rogers National Recreation Area, Little Laurel Creek, beside Route 600, 2.2 kilometers north of Route 603, 10 Jul 1990, by IM Smith, IMS 900086.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
69615CB3F40CD6A66422478143FE8335.taxon	diagnosis	Diagnosis. Torrenticola racupalpa are similar to other members of the Raptor Group (T. gnoma, T. irapalpa, T. longitibia, T. mjolniri, T. elusiva, T. raptor, T. danielleae, T. daemon, and T. ivyae) in having round bodies; Dgl- 4 close to muscles scars; long, thin subcapitular rostra; and long, thin pedipalp tibiae. Female T. racupalpa can be differentiated from T. elusiva by having a more elongate subcapitulum (ventral length / height = 2.48 - 2.73 in T. racupalpa, 2.39 in T. elusiva); and by dorsal pattern. T. racupalpa can be differentiated from T. irapalpa and T. daemon by having Dgl- 4 closer to the muscle scars (dorsal width / distance between Dgl- 4 = 2.19 - 2.77 in T. racupalpa, 1.45 - 2.09 in others) and a more elongate rostrum (length / width = 3.56 - 3.88 in T. racupalpa, 2.66 - 3.39 in others). T. racupalpa can be differentiated from T. gnoma by having a more elongate rostrum (length / width = 3.56 - 3.88 in T. racupalpa, 2.56 - 3.13 in T. gnoma) and dorsal coloration and pattern. T. racupalpa can be differentiated from T. mjolniri by having a shorter anterior venter (♀ = 152.5 - 165 in T. racupalpa, 180 - 195 in T. mjolniri; ♂ = 200 - 205 in T. racupalpa, 230 - 255 in T. mjolniri); and by dorsal pattern. Male T. racupalpa can be differentiated from T. longitibia (males only) by having a shorter femur with respect to the genu (femur / genu = 1.92 - 1.92 in T. racupalpa, 2.10 - 2.17 in T. longitibia); a stockier rostrum (length / width = 3.88 - 3.88 in T. racupalpa, 4.15 - 4.23 in T. longitibia); and dorsal pattern. T. racupalpa can be differentiated from T. raptor by having Dgl- 4 closer to the muscle scars (dorsal width / distance between Dgl- 4 = 2.19 - 2.77 in T. racupalpa, 1.68 - 2.02 in T. raptor); and shorter anterior venter (♀ = 152.5 - 165 in T. racupalpa, 205 - 240 in T. raptor, ♂ = 200 - 205 in T. racupalpa, 245 - 305 in T. raptor). Female T. racupalpa can be differentiated from female T. raptor by having stockier pedipalp tibiae (length / width = 4.44 - 5.00 in T. racupalpa, 6 - 7.54 in T. raptor). T. racupalpa can be differentiated from T. danielleae by having Dgl- 4 closer to the muscle scars (dorsal width / distance between Dgl- 4 = 2.19 - 2.77 in T. racupalpa, 1.42 - 1.70 in T. danielleae) and by dorsal coloration and pattern. Female T. racupalpa can be differentiated from female T. ivyae by having a less elongate rostrum (length / width = 3.56 - 3.82 in T. racupalpa, 4.00 - 4.15 in T. ivyae) and less elongate pedipalpal tibiae (length / width = 4.44 - 5.00 in T. racupalpa, 5.07 - 5.64 in T. ivyae). Male T. racupalpa can be differentiated from male T. ivyae by having a shorter anterior venter (♂ 200 - 205 in T. racupalpa, 220 - 230 in T. ivyae) and a longer genital field (♂ 160 - 165 in T. racupalpa, 142 - 148 in T. ivyae).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
69615CB3F40CD6A66422478143FE8335.taxon	description	Description. Female (Figure 206) (n = 3) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (570 - 630 (570) long; 450 - 540 (450) wide) ovoid with bluish-purple coloration separated into anterior and posterior portions with bold orange medially. Anterio-medial platelets (130 - 145 (130) long; 52.5 - 65 (52.5) wide). Anterio-lateral platelets (157.5 - 200 (157.5) long; 71.25 - 75 (71.25) wide) free from dorsal plate. Dgl- 4 much closer to the muscle scars than to dorsum edge (distance between Dgl- 4 170 - 205 (170 )). Dorsal plate proportions: dorsum length / width 1.17 - 1.27 (1.27); dorsal width / distance between Dgl- 4 2.59 - 2.77 (2.65); anterio-medial platelet length / width 2.19 - 2.48 (2.48); anterio-lateral platelet length / width 2.21 - 2.67 (2.21); anterio-lateral / anterio-medial length 1.21 - 1.40 (1.21). Gnathosoma - Subcapitulum (330 - 375 (330) long (ventral); 251 - 297.5 (252) long (dorsal); 130 - 145 (130) tall) faint bluish-purple coloration. Rostrum (142.5 - 170 (142.5) long; 40 - 45 (40) wide) elongate. Chelicerae (319 - 375 (320) long) with curved fangs (50 - 60 (50) long). Subcapitular proportions: ventral length / height 2.48 - 2.73 (2.54); rostrum length / width 3.56 - 3.82 (3.56). Pedipalps elongate (especially tibia) with long tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (45 - 45 (45) long); femur (125 - 145 (125) long); genu (67.5 - 75 (67.5) long); tibia (100 - 125 (100) long; 22.5 - 25 (22.5) wide); tarsus (17.5 - 20 (17.5) long). Palpomere proportions: femur / genu 1.85 - 1.93 (1.85); tibia / femur 0.80 - 0.86 (0.80); tibia length / width 4.44 - 5.00 (4.44). Venter - (700 - 800 (700) long; 490 - 610 (491) wide) with bold bluish-purple coloration. Gnathosomal bay (152.5 - 185 (152.5) long; 80 - 87.5 (87.5) wide). Cxgl- 4 subapical. Medial suture (17.5 - 20 (17.5) long). Genital plates (152.5 - 185 (152.5) long; 140 - 170 (140) wide). Additional measurements: Cx- 1 (285 - 320 (286) long (total); 114 - 140 (115) long (medial )); Cx- 3 (332 - 380 (332) wide); anterior venter (152.5 - 165 (152.5) long). Ventral proportions: gnathosomal bay length / width 1.74 - 2.31 (1.74); anterior venter / genital field length 0.89 - 1.00 (1.00); anterior venter length / genital field width 0.97 - 1.09 (1.09); anterior venter / medial suture 8.00 - 8.71 (8.71). Male (Figure 207) (n = 2) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (555 - 570 (570) long; 460 - 475 (475) wide) ovoid with bluish-purple coloration separated into anterior and posterior portions with bold orange medially. Anterio-medial platelets (130 - 135 (135) long; 57.5 - 65 (65) wide). Anterio-lateral platelets (182.5 - 190 (190) long; 70 - 80 (80) wide) free from dorsal plate. Dgl- 4 much closer to the muscle scars than to dorsum edge (distance between Dgl- 4 200 - 210 (200 )). Dorsal plate proportions: dorsum length / width 1.20 - 1.21 (1.20); dorsal width / distance between Dgl- 4 2.19 - 2.38 (2.38); anterio-medial platelet length / width 2.08 - 2.26 (2.08); anterio-lateral platelet length / width 2.38 - 2.61 (2.38); anterio-lateral / anterio-medial length 1.40 - 1.41 (1.41). Gnathosoma - Subcapitulum (315 - 317.5 (317.5) long (ventral); 250 - 250 (250) long (dorsal); 110 - 115 (115) tall) faint bluish-purple coloration. Rostrum (155 - 155 (155) long; 40 - 40 (40) wide) elongate. Chelicerae (295 - 310 (310) long) with curved fangs (50 - 55 (50) long). Subcapitular proportions: ventral length / height 2.76 - 2.86 (2.76); rostrum length / width 3.88 - 3.88 (3.88). Pedipalps elongate (especially tibia) with long tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (40 - 40 (40) long); femur (125 - 125 (125) long); genu (65 - 65 (65) long); tibia (107.5 - 110 (107.5) long; 20 - 20 (20) wide); tarsus (15 - 15 (15) long). Palpomere proportions: femur / genu 1.92 - 1.92 (1.92); tibia / femur 0.86 - 0.88 (0.86); tibia length / width 5.38 - 5.50 (5.38). Venter - (695 - 710 (710) long; 500 - 505 (500) wide) with bold bluish-purple coloration. Gnathosomal bay (157.5 - 160 (157.5) long; 70 - 70 (70) wide). Cxgl- 4 subapical. Medial suture (45 - 45 (45) long). Genital plates (160 - 165 (165) long; 125 - 125 (125) wide). Additional measurements: Cx- 1 (300 - 305 (300) long (total); 145 - 145 (145) long (medial )); Cx- 3 (340 - 345 (340) wide); anterior venter (200 - 205 (205) long). Ventral proportions: gnathosomal bay length / width 2.25 - 2.29 (2.25); anterior venter / genital field length 1.24 - 1.25 (1.24); anterior venter length / genital field width 1.60 - 1.64 (1.64); anterior venter / medial suture 4.44 - 4.56 (4.56). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
69615CB3F40CD6A66422478143FE8335.taxon	etymology	Etymology. Specific epithet (racupalpa) refers to the spined, rake-like pedipalps which have long, thin tibiae and elongate tubercles (racu, Old English feminine, rake; palpus, L. hand, feeler).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
69615CB3F40CD6A66422478143FE8335.taxon	distribution	Distribution. Southeastern, Tennessee and Virginia (Figure 205).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
DE61DF71E7EB0BFEBF0B5584332321ED.taxon	materials_examined	Material examined (11 ♀; 15 ♂). Arizona, USA: 3 ♂ from Coconino County, Oak Creek Canyon, Oak Creek, beside Route 89 A, between Banjo Bill & Bootlegger campgrounds, 21 Jul 1987, by IM Smith, IMS 870100 A * 1 ♂ and 1 ♂ from Cochise County, Chiricahua Mountains, Cave Creek Recreation Area, Cave Creek at John Hand Picnic Area, 15 Jul 1987, by IM Smith, IMS 870092 B * 1 ♀ from Cochise County, Chiricahua Mountains, Cave Creek Recreation Area, Cave Creek at Stewart campground, 16 Jul 1987, by IM Smith, IMS 870094 * 1 ♀ from Cochise County, Chiricahua Mountains, Cave Creek, Herb Martyr Campground, Forest Road 42 A 10 May 2012, by IM Smith, IMS 120009 * 3 ♀ and 1 ♂ from Gila County, East Verde River, beside Route 87, north of Payson, 19 Jul 1987, by IM Smith, IMS 870097 * 2 ♂ from Yavapai County, West Clear Creek at Clear Creek campground, off Forest Road 9, east of Camp Verde, 19 - 22 Jul 1987, by IM Smith, IMS 870098 * New Mexico, USA: * 1 ♂ from Catron County, Glenwood Whitewater Creek at Whitewater Creek Picnic Area, 12 Jul 1987, by IM Smith, IMS 870084 * 2 ♀ and 4 ♂ from Catron County, Little Creek, beside Route 15, 65 kilometers north of Route 180 in Silver City, 10 Jul 1987, by IM Smith, IMS 870081 A * Texas, USA: 1 ♀ and 1 ♂ from Bandera County, Lost Maples State Natural Area, Sabinal River (29 ° 49 ' N, 99 ° 34 ' W), 27 Sep 1995, by IM Smith, IMS 950052 * 1 ♀ from Bandera County, Vanderpool, beside Route 187, Sabinal River (29 ° 48 ' 10 " N, 99 ° 34 ' 30 " W), 2 May 2009, by IM Smith, IMS 090007 * 1 ♀ and 1 ♂ from Val Verde County, Bakers Crossing Campground, off Route 163, Devils River (29 ° 58 ' N, 101 ° 9 ' W), 5 Oct 1999, by IM Smith, IMS 990061 A * 1 ♀ from Uvalde County, Garner State Park, Frio River (29 ° 35 ' 22 " N, 99 ° 44 ' 12 " W), 28 May 1998, by IM Smith, IMS 980027 A.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
DE61DF71E7EB0BFEBF0B5584332321ED.taxon	diagnosis	Diagnosis. Torrenticola rala are similar to other members of the Rala Group (T. boettgeri, T. keesdavidsi, T. kurtvietsi, T. lamellipalpis, T. dolichodactyla, and T. anoplopalpa) by being colorless, having incomplete hind coxal margins and being distributed in the southwest. T. rala can be differentiated from all other Rala Group by having a stockier rostrum (length / width = 1.63 - 1.83 in T. rala, 2.19 - 3.57 in others). Additionally, T. rala can be differentiated from all other Rala Group by having a shorter anterior venter (♀ = 170 - 185 in T. rala 192 - 260 in others ♂ = 200 - 225 in T. rala, 235 - 308 in others), except male T. keesdavidsi (♂ 225 - 260).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
DE61DF71E7EB0BFEBF0B5584332321ED.taxon	description	Re-description. Female (Figure 209) (n = 5) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (630 - 700 long; 465 - 505 wide) ellipsoid and colorless. Anterio-medial platelets (150 - 162.5 long; 62.5 - 72.5 wide). Anterio-lateral platelets (172.5 - 195 long; 70 - 80 wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 370 - 420). Dorsal plate proportions: dorsum length / width 1.30 - 1.46; dorsal width / distance between Dgl- 4 1.17 - 1.26; anterio-medial platelet length / width 2.14 - 2.41; anterio-lateral platelet length / width 2.38 - 2.79; anterio-lateral / anterio-medial length 1.15 - 1.26. Gnathosoma - Subcapitulum (247.5 - 260 long (ventral); 160 - 170 long (dorsal); 115 - 120 tall) tall and colorless. Rostrum (79 - 89 long; 45 - 50 wide) short and conical. Chelicerae (217.5 - 247.5 long) with curved fangs (52.5 - 60 long). Subcapitular proportions: ventral length / height 2.06 - 2.17; rostrum length / width 1.70 - 1.83. Pedipalps short and stocky (especially tibiae) with tuberculate ventral extensions on femora and ventral extensions on genua absent. Palpomeres: trochanter (35 - 37.5 long); femur (70 - 77.5 long); genu (50 - 57.5 long); tibia (55 - 62.5 long; 20 - 22.5 wide); tarsus (15 - 17.5 long). Palpomere proportions: femur / genu 1.35 - 1.43; tibia / femur 0.79 - 0.81; tibia length / width 2.75 - 2.88. Venter - (760 - 840 long; 527 - 585 wide) colorless. Gnathosomal bay (150 - 162.5 long; 55 - 62.5 wide). Cxgl- 4 apical. Medial suture (25 - 35 long). Genital plates (175 - 190 long; 155 - 170 wide). Additional measurements: Cx- 1 (280 - 300 long (total); 130 - 145 long (medial )); Cx- 3 (345 - 380 wide); anterior venter (170 - 185 long). Ventral proportions: gnathosomal bay length / width 2.58 - 2.73; anterior venter / genital field length 0.92 - 1.01; anterior venter length / genital field width 1.00 - 1.19; anterior venter / medial suture 5.00 - 6.80. Male (Figure 210) (n = 5) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (600 - 655 long; 435 - 480 wide) ellipsoid and colorless. Anterio-medial platelets (135 - 165 long; 55 - 67.5 wide). Anterio-lateral platelets (177.5 - 190 long; 64 - 82.5 wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 375 - 400). Dorsal plate proportions: dorsum length / width 1.35 - 1.42; dorsal width / distance between Dgl- 4 1.16 - 1.20; anterio-medial platelet length / width 2.40 - 2.48; anterio-lateral platelet length / width 2.30 - 2.78; anterio-lateral / anterio-medial length 1.15 - 1.31. Gnathosoma - Subcapitulum (230 - 249 long (ventral); 152.5 - 162.5 long (dorsal); 107.5 - 112.5 tall) tall and colorless. Rostrum (77.5 - 81 long; 45 - 47.5 wide) short and conical. Chelicerae (215 - 232.5 long) with curved fangs (42.5 - 50 long). Subcapitular proportions: ventral length / height 2.14 - 2.21; rostrum length / width 1.63 - 1.72. Pedipalps short and stocky (especially tibiae) with tuberculate ventral extensions on femora and ventral extensions on genua absent. Palpomeres: trochanter (32.5 - 37.5 long); femur (65 - 72.5 long); genu (49 - 52.5 long); tibia (55 - 62.5 long; 20 - 21 wide); tarsus (15 - 17.5 long). Palpomere proportions: femur / genu 1.33 - 1.45; tibia / femur 0.79 - 0.89; tibia length / width 2.71 - 2.94. Venter - (710 - 765 long; 500 - 540 wide) colorless. Gnathosomal bay (135 - 145 long; 55 - 62.5 wide). Cxgl- 4 apical. Medial suture (65 - 80 long). Genital plates (160 - 185 long; 127.5 - 135 wide). Additional measurements: Cx- 1 (270 - 295 long (total); 125 - 145 long (medial )); Cx- 3 (337.5 - 375 wide); anterior venter (200 - 225 long). Ventral proportions: gnathosomal bay length / width 2.20 - 2.45; anterior venter / genital field length 1.14 - 1.33; anterior venter length / genital field width 1.57 - 1.67; anterior venter / medial suture 2.66 - 3.31. Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
DE61DF71E7EB0BFEBF0B5584332321ED.taxon	etymology	Etymology. Cook (1980) did not specify an etymology for this specific epithet (rala) and we are unable to offer helpful speculation.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
DE61DF71E7EB0BFEBF0B5584332321ED.taxon	distribution	Distribution. Southwestern U. S. (Arizona, New Mexico, Texas) and southward through Central America (Figure 208).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
46F7DB2589049FAAC0D2A5DC660FD28D.taxon	materials_examined	Material examined. HOLOTYPE (♀): from USA, Ontario, Hastings County, Maple Leaf, Papineau Creek, beside Highway 62, 18 Aug 2011, by IM Smith, IMS 110054, DNA 2864. PARATYPES (58 ♀; 40 ♂): Alabama, USA: 3 ♀ and 3 ♂ from Clay County, Talladega Creek, beside Forest Route 649, 0.8 kilometers northeast of road from Forest Route 600 to Campbell Springs, 2 Jul 1990, by IM Smith, IMS 900075 A * Maine, USA: 1 ♀ from Aroostook County, Ashland, beside Route 11 at bridge, Aroostook River (46 ° 38 ' N, 68 ° 24 ' W), 4 Jul 1989, by IM Smith, IMS 890067 * 1 ♀ from Franklin County, Smalls Falls Picnic Area, beside Route 4, Sandy River (44 ° 52 ' N, 70 ° 31 ' W), 5 Jul 1989, by IM Smith, IMS 890069 * 2 ♀ from Washington County, Old Stream, off Route 9, 5.5 km west of Route 192, 6 Jun 2012, by IM Smith, IMS 120012 * New Brunswick, Canada: 2 ♀ and 1 ♂ from Charlotte County, Rollingham, Digdeguash River, beside Highway 770 at covered bridge, 30 Jun 1989, by IM Smith, IMS 890053 * 2 ♀ and 1 ♂ from Charlotte County, Rollingham, Digdegaush River, beside Highway 770, 3 Oct 2011, by IM Smith, IMS 110118 * 1 ♂ from Northumberland County, Renous River, beside Highway 108, 18 Jul 1980, by IM Smith, IMS 800111 * 1 ♀ and 1 ♂ from Restigouche County, Mt. Carleton Provincial Park, Nictau River, 16 Jul 1980, by IM Smith, IMS 800109 * 1 ♀ and 1 ♂ from York County, Magaguadavic River, beside Highway 3, just east of Thomaston Corners, 1 Jul 1989, by IM Smith, IMS 890055 A * 1 ♂ from York County, Napadogan Brook, beside Road J- 19, 6.3 kilometers north of Nashwaak Exper. Watershed headquarters, 23 Jul 1981, by IM Smith, IMS 810095 B * 1 ♀ from York County, Nashwaak Exper. Watershed, Nashwaak River, at trunk road, 20 Jul 1980, by IM Smith, IMS 800118 A * New Hampshire, USA: 1 ♀ from Coos County, picnic area beside Route 110, Ammonoosuc River (44 ° 36 ' N, 71 ° 24 ' W), 5 Jul 1989, by IM Smith, IMS 890071 * New Jersey, USA: 1 ♀ and 1 ♂ from Sussex County, Big Flat Brook beside Flatbrook Road, 2.6 kilometers north of Route 206 at Tuttles Corner, 23 Jun 1990, by IM Smith, IMS 900053 * New York, USA: 2 ♀ from Essex County, Minerva, Boreas River, beside Route 28 N, 13.8 kilometers northwest of Morse Memorial Parkway, 21 Jun 1990, by IM Smith, IMS 900050 A * 1 ♀ and 1 ♂ from Greene County, Schoharie Creek, beside Route 23 A, 9.6 kilometers west of Route 296 in Hunter, 22 Jun 1990, by IM Smith, IMS 900052 * 1 ♀ from Ulster County, beside Route 28, 1.6 kilometers south of Mt. Tremper, 17 Aug 1964, by DR Cook, DRC 640023 * 2 ♀ and 1 ♂ from Warren County, East Branch of Sacandaga River, beside Route 8, 14.5 kilometers east of Route 30, 21 Jun 1990, by IM Smith, IMS 900051 A * North Carolina, USA: 2 ♀ and 2 ♂ from Yancey County, Lost Cove Picnic Area, South Toe River, on Forest Route 472, 2.8 kilometers south of Route 80, 28 Jun 1990, by IM Smith, IMS 900065 A * Nova Scotia, Canada: 1 ♀ from Inervess County, Cape Brenton Highlands National Park, Cheticamp entrance, pond near Salmon Pools Trailhead, 10 Sep 2011, by IM Smith, IMS 110072 * 1 ♀ from Inervess County, Inervess, Cheticamp River, 10 Sep 2011, by IM Smith, IMS 110071 * 1 ♀ and 1 ♂ from Luneburg County, New Germany, LaHave River, beside Highway 10, 23 Sep 2011, by IM Smith, IMS 110098 * 2 ♀ and 1 ♂ from Victoria County, Cape Brenton Island, Baddeck River, beside road to Baddeck Forks, 18 Jul 1981, by IM Smith, IMS 810082 * Ontario, Canada: 1 ♂ from Cochrane County, Hearst, Pitopiko River, beside Highway 11, 24 Jul 1975, by IM Smith & D Spaner, IMS 750120 * 3 ♀ and 3 ♂ from Hastings County, Madawaska, Opeongo River, beside Highway 60, 29 Aug 1981, by IM Smith, IMS 810033 A * 2 ♀ and 2 ♂ from Hastings County, Maple Leaf, Papineau Creek, 10 - 11 Jun 1981, by IM Smith, IMS 810014 A * 1 ♂ (ALLOTYPE) from Hastings County, Maple Leaf, Papineau Creek, east of Davis Road before Highway 62, 18 Aug 2011, by IM Smith, IMS 110053, DNA 1257 * 2 ♀ and 1 ♂ from Hastings County, Maple Leaf, Papineau Creek, east of Davis Road before Highway 62, 18 Aug 2011, by IM Smith, IMS 110053 * 1 ♀ from Hastings County, Maple Leaf, Papineau Creek, beside Highway 62, 18 Aug 2011, by IM Smith, IMS 110054 * 1 ♀ from Kenora County, Ignace, beside Highway 599, 4.2 kilometers north of Highway 17, 31 Jul 1975, by IM Smith, IMS 750216 * 2 ♀ and 2 ♂ from Kenora County, Revell River, beside Highway 17, 40.2 kilometers east of Highway 72, 3 Jun 1980, by IM Smith, IMS 800052 * 2 ♀ and 2 ♂ from Muskoka County, Huntsville, East River, Xing road to Dyer Memorial, 26 Aug 1981, by IM Smith, IMS 810032 A * 1 ♀ and 1 ♂ from Lanark County, Mississippi River, beside Lanark Road # 12, between Lanark & Fallbrook, 6 Oct 1983, by IM Smith & CJ Hill, IMS 830094 A * 1 ♂ from Nipissing County, Algonquin Provincial Park, Madawaska River, at Highway 60, near Lake of Two Rivers, 15 May 1980, by IM Smith & CJ Hill, IMS 800004 C * 1 ♀ and 1 ♂ from Nipissing County, Aumond Creek, beside Highway 17, east of Mattawa, 30 Aug 1983, by IM Smith & CJ Hill, IMS 830079 A * 1 ♀ and 1 ♂ from Nipissing County, Aumond Creek, beside Highway 17, east of Mattawa, 30 Aug 1983, by IM Smith & CJ Hill, IMS 830079 B * 1 ♀ from Nipissing County, Bastien Creek, at picnic area, beside Highway 17, east of Mattawa, 30 Aug 1983, by IM Smith & CJ Hill, IMS 830076 A * 1 ♀ and 1 ♂ from Peterborough County, Apsley, Eels Creek, crossing Highway 28, just south of Eels Lake, 13 Jun 1981, by IM Smith, IMS 810017 A * 1 ♀ and 1 ♂ from Peterborough County, Eels Creek, crossing Highway 28 at picnic area, just north of Woodview, 13 Jun 1981, by IM Smith, IMS 810018 A * 1 ♂ from Thunder Bay County, west of Geraldton, Creelman Creek, beside Highway 11, 26 Jul 1975, by IM Smith & D Spaner, IMS 750147 * Quebec, Canada: 1 ♀ from Gatineau County, Gatineau Park stream crossing, Gatineau Parkway at Meech Lake Road, 15 Sep 1981, by IM Smith & C Cramer, IMS 810034 A * 1 ♀ from Gatineau County, beside Gatineau Parkway, at Meech Lake Road, 13 Aug 1982, by IM Smith & CJ Hill, IMS 820001 A * 1 ♂ from Gatineau County, beside Gatineau Parkway, at Meech Lake Road, 13 Aug 1982, by IM Smith & CJ Hill, IMS 820001 B * 1 ♀ and 1 ♂ from Pontiac County, beside road at east end of Thorne Lake (45 ° 32 ' N, 76 ° 0 ' W), 1 May 1986, by IM Smith & CJ Hill, IMS 860002 A * South Carolina, USA: 1 ♀ from Greenville County, Matthews Creek, 24 Apr 2014, by D Eargle, JRF 14 - 0424 - 001 * Tennessee, USA: 1 ♀ and 1 ♂ from Monroe County, Tellico River, beside Forest Route 210, 1.8 kilometers east of bridge at Bald River Falls, 5 Jul 1990, by IM Smith, IMS 900079 * 1 ♀ from Monroe County, Tellico River (35 ° 20 ' 27 " N, 84 ° 11 ' 31 " W), 12 Sep 2009, by IM Smith, IMS 090111 * 1 ♀ from Sevier County, Great Smokey Mountians National Park, Little River (35 ° 40 ' 56 " N, 83 ° 39 ' 2 " W), 8 Sep 2009, by IM Smith, IMS 090103 * 2 ♀ from Sevier County, Great Smokey Mountianss National Park, Laurel Creek (35 ° 39 ' 7 " N, 83 ° 42 ' 32 " W), 17 Sep 2010, by IM Smith, IMS 100145 * West Virginia, USA: 1 ♀ and 1 ♂ from Pocahontas County, Island Campground, East Fork of Greenbrier River, beside Route 28, northeast of Thornwood, 16 Jul 1990, by IM Smith, IMS 900101 A * 1 ♀ and 1 ♂ from Randolph County, Laurel Fork Campground, Laurel Fork of Cheat River, off Forest Route 14, south of Wymer, 17 Jul 1990, by IM Smith, IMS 900102.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
46F7DB2589049FAAC0D2A5DC660FD28D.taxon	diagnosis	Diagnosis. Torrenticola raptor are similar to other members of the Raptor Group (T. gnoma, T. irapalpa, T. longitibia, T. mjolniri, T. elusiva, T. racupalpa, T. danielleae, T. daemon, and T. ivyae) in having round bodies; Dgl- 4 close to muscles scars; long, thin subcapitular rostra; and long, thin pedipalp tibiae. T. raptor can be differentiated from all other members of the Raptor Group by having a more elongate subcapitulum (ventral length / height ♀ = 2.98 - 3.18 in T. raptor, 2.26 - 2.90 in others; ♂ = 3.13 - 3.27 in T. raptor, 2.29 - 3.00 in others). T. raptor can be further differentiated from T. elusiva, T. irapalpa, T. gnoma, T. danielleae, T. daemon, and T. ivyae by having more elongate pedipalp tibiae (length / width ♀ = 6.00 - 7.54 in T. raptor, 4.09 - 5.67 in others; ♂ = 5.29 - 5.63 in T. raptor, 3.88 - 5.20 in others). T. raptor can be further differentiated from T. mjolniri, T. longitibia, T. gnoma, T. elusiva, T. racupalpa, and T. ivyae by having Dgl- 4 closer to the dorsal edge (dorsal width / distance between Dgl- 4 = 1.66 - 2.02 in T. raptor, 2.06 - 3.29 in others).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
46F7DB2589049FAAC0D2A5DC660FD28D.taxon	description	Description. Female (Figure 212) (n = 13) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (570 - 660 (625) long; 465 - 550 (505) wide) circular with coloration posteriorly extending in a strip anteriorly to the edge of the dorsal plate (rarely without anterior extension), coloration variable from navy blue to purple to reddish purple. Anterio-medial platelets (122.5 - 150 (135) long; 60 - 80 (75) wide). Anterio-lateral platelets (182.5 - 210 (195) long; 75 - 90 (80) wide) free from dorsal plate. Dgl- 4 closer to the muscle scars than to the dorsum edge (distance between Dgl- 4 230 - 295 (275 )). Dorsal plate proportions: dorsum length / width 1.19 - 1.31 (1.24); dorsal width / distance between Dgl- 4 1.80 - 2.02 (1.84); anterio-medial platelet length / width 1.75 - 2.07 (1.80); anterio-lateral platelet length / width 2.28 - 2.49 (2.44); anterio-lateral / anterio-medial length 1.39 - 1.57 (1.44). Gnathosoma - Subcapitulum (335 - 380 (365) long (ventral); 249 - 283 (270) long (dorsal); 107.5 - 125 (120) tall) colorless. Rostrum (150 - 167.5 (160) long; 37.5 - 45 (37.5) wide) elongate. Chelicerae (316 - 368 (355) long) with curved fangs (43 - 60 (60) long). Subcapitular proportions: ventral length / height 2.98 - 3.18 (3.04); rostrum length / width 3.44 - 4.40 (4.27). Pedipalps elongate (especially tibiae) with long tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (37.5 - 42.5 (42.5) long); femur (122.5 - 143.75 (137.5) long); genu (65 - 77.5 (70) long); tibia (112.5 - 142.5 (122.5) long; 16.25 - 20 (18.75) wide); tarsus (17.5 - 22.5 (22.5) long). Palpomere proportions: femur / genu 1.68 - 1.96 (1.96); tibia / femur 0.82 - 1.03 (0.89); tibia length / width 6.00 - 7.54 (6.53). Venter - (720 - 830 (800) long; 535 - 609 (550) wide) with faint navy blue to purple coloration or colorless. Gnathosomal bay (135 - 192.5 (172.5) long; 62.5 - 85 (75) wide). Cxgl- 4 far from apex. Medial suture (35 - 45 (45) long). Genital plates (157.5 - 182.5 (170) long; 135 - 153.75 (145) wide). Additional measurements: Cx- 1 (271 - 330 (330) long (total); 92 - 160 (160) long (medial )); Cx- 3 (336 - 373 (340) wide); anterior venter (205 - 240 (220) long). Ventral proportions: gnathosomal bay length / width 1.74 - 2.48 (2.30); anterior venter / genital field length 1.23 - 1.47 (1.29); anterior venter length / genital field width 1.41 - 1.65 (1.52); anterior venter / medial suture 4.56 - 6.29 (4.89). Male (Figure 213) (n = 5) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (540 - 610 (540) long; 415 - 450 (440) wide) circular with coloration posteriorly extending in a strip anteriorly to the edge of the dorsal plate (rarely without anterior extension), coloration variable from navy blue to purple to reddish purple. Anterio-medial platelets (126.25 - 132.5 (127.5) long; 57.5 - 65 (57.5) wide). Anterio-lateral platelets (177.5 - 190 (177.5) long; 68 - 80 (75) wide) free from dorsal plate. Dgl- 4 closer to the muscle scars than to the dorsum edge (distance between Dgl- 4 230 - 265 (230 )). Dorsal plate proportions: dorsum length / width 1.23 - 1.36 (1.23); dorsal width / distance between Dgl- 4 1.66 - 1.91 (1.91); anterio-medial platelet length / width 1.94 - 2.22 (2.22); anterio-lateral platelet length / width 2.34 - 2.58 (2.37); anterio-lateral / anterio-medial length 1.39 - 1.47 (1.39). Gnathosoma - Subcapitulum (297.5 - 327.5 (297.5) long (ventral); 220 - 247 (221) long (dorsal); 95 - 102.5 (95) tall) colorless. Rostrum (131.25 - 147.5 (131.25) long; 30 - 35 (30) wide) elongate. Chelicerae (274 - 306 (274) long) with curved fangs (44 - 54 (45) long). Subcapitular proportions: ventral length / height 3.13 - 3.27 (3.13); rostrum length / width 3.89 - 4.38 (4.38). Pedipalps elongate (especially tibiae) with long tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (37.5 - 40 (40) long); femur (111.25 - 122.5 (111.25) long); genu (62.5 - 65 (62.5) long); tibia (102.5 - 112.5 (102.5) long; 18.75 - 21.25 (18.75) wide); tarsus (17.5 - 20 (20) long). Palpomere proportions: femur / genu 1.73 - 1.88 (1.78); tibia / femur 0.89 - 0.99 (0.92); tibia length / width 5.29 - 5.63 (5.47). Venter - (680 - 790 (680) long; 457 - 523 (458) wide) with faint navy blue to purple coloration or colorless. Gnathosomal bay (132.5 - 145 (132.5) long; 52.5 - 77.5 (52.5) wide). Cxgl- 4 far from apex. Medial suture (80 - 107.5 (80) long). Genital plates (137.5 - 152.5 (141.25) long; 110 - 125 (120) wide). Additional measurements: Cx- 1 (268 - 322 (269) long (total); 112 - 160 (113) long (medial )); Cx- 3 (332 - 347 (335) wide); anterior venter (245 - 305 (245) long). Ventral proportions: gnathosomal bay length / width 1.81 - 2.52 (2.52); anterior venter / genital field length 1.73 - 2.11 (1.73); anterior venter length / genital field width 2.04 - 2.60 (2.04); anterior venter / medial suture 2.84 - 3.06 (3.06). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
46F7DB2589049FAAC0D2A5DC660FD28D.taxon	etymology	Etymology. Specific epithet (raptor) refers to the long, thin pedipalps (especially tibiae) of this species which, combined with pronounced tubercles, appear especially capable of grasping slippery prey (rapio L. to seize; known in apposition).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
46F7DB2589049FAAC0D2A5DC660FD28D.taxon	distribution	Distribution. Northeastern, but extending southward in the Appalachians (Figure 211).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
01174C58015DB3149D1BB69E6C655B21.taxon	materials_examined	Material examined. HOLOTYPE (♀): from USA, New Mexico, Catron County, Little Creek, Gila Hot Springs, Little Creek Recreation Area off Route 15, 6 May 2012, by IM Smith, IMS 120006, DNA 2895. PARATYPES (8 ♀; 8 ♂): Arizona, USA: 2 ♀ and 2 ♂ from Coconino County, Oak Creek Canyon, Oak Creek, beside Route 89 A, between Banjo Bill & Bootlegger campgrounds, 21 Jul 1987, by IM Smith, IMS 870100 B * 1 ♀ from Coconino County, Oak Creek Canyon, Oak Creek, beside Route 89 A, just north of Pine Flat campground, 21 Jul 1987, by IM Smith, IMS 870099 B * 1 ♀ and 1 ♂ from Yavapai County, West Clear Creek at Clear creek campground, off Forest Road 9, east of Camp Verde, 22 Jul 1987, by IM Smith, IMS 870102 * New Mexico, USA: 1 ♂ (ALLOTYPE) from Grant County, East Fork Gila River, Grapevine Recreation Area off Route 15, north of Silver City, 5 May 2012, by IM Smith, IMS 120007, DNA 2900 * 1 ♂ from Catron County, Gila River, beside Route 15, just below mouth of Little Creek, 11 Jul 1987, by IM Smith, IMS 870083 A * 2 ♀ and 1 ♂ from Catron County, Little Creek, Gila Hot Springs, Little Creek Recreation Area off Route 15, 6 May 2012, by IM Smith, IMS 120006 * 2 ♀ and 2 ♂ from Catron County, Little Creek, beside Route 15, 65 kilometers north of Route 180 in Silver City, 10 Jul 1987, by IM Smith, IMS 870081 A.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
01174C58015DB3149D1BB69E6C655B21.taxon	diagnosis	Diagnosis. Torrenticola raptoroides are unlike all other western species in having round bodies with dorsal coloration restricted posteriorly and long, thin pedipalp tibiae. Additionally, they are only known from Catron & Grant Counties, New Mexico.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
01174C58015DB3149D1BB69E6C655B21.taxon	description	Description. Female (Figure 215) (n = 5) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (610 - 690 (650) long; 485 - 530 (520) wide) ovoid with navy blue coloration separated into anterior and posterior portions with faint orange medially. Anterio-medial platelets (140 - 167.5 (157.5) long; 56 - 67.5 (61.25) wide). Anterio-lateral platelets (165 - 200 (195) long; 72.5 - 82.5 (75) wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 295 - 380 (380 )). Dorsal plate proportions: dorsum length / width 1.19 - 1.30 (1.25); dorsal width / distance between Dgl- 4 1.37 - 1.64 (1.37); anterio-medial platelet length / width 2.36 - 2.60 (2.57); anterio-lateral platelet length / width 2.27 - 2.60 (2.60); anterio-lateral / anterio-medial length 1.15 - 1.24 (1.24). Gnathosoma - Subcapitulum (315 - 335 (330) long (ventral); 240 - 250 (249) long (dorsal); 135 - 140 (137.5) tall) colorless. Rostrum (135 - 142.5 (142.5) long; 40 - 45 (42.5) wide). Chelicerae (335 - 360 (345) long) with curved fangs (62 - 69 (63) long). Subcapitular proportions: ventral length / height 2.25 - 2.48 (2.40); rostrum length / width 3.17 - 3.38 (3.35). Pedipalps elongate (especially tibiae) with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (40 - 47.5 (45) long); femur (120 - 131.25 (131.25) long); genu (63.75 - 75 (72.5) long); tibia (100 - 112.5 (112.5) long; 21.25 - 22.5 (22.5) wide); tarsus (15 - 17.5 (17.5) long). Palpomere proportions: femur / genu 1.60 - 1.88 (1.81); tibia / femur 0.79 - 0.94 (0.86); tibia length / width 4.44 - 5.00 (5.00). Venter - (710 - 805 (805) long; 562 - 600 (562) wide) colorless. Gnathosomal bay (167.5 - 182.5 (180) long; 72.5 - 100 (82.5) wide). Cxgl- 4 far from apex. Medial suture (10 - 17.5 (12.5) long). Genital plates (172.5 - 182.5 (172.5) long; 167.5 - 177.5 (167.5) wide). Additional measurements: Cx- 1 (300 - 311 (311) long (total); 111 - 140 (112) long (medial )); Cx- 3 (350 - 410 (373) wide); anterior venter (150 - 187.5 (170) long). Ventral proportions: gnathosomal bay length / width 1.83 - 2.41 (2.18); anterior venter / genital field length 0.86 - 1.04 (0.99); anterior venter length / genital field width 0.88 - 1.07 (1.01); anterior venter / medial suture 10.71 - 16.75 (13.60). Male (Figure 216) (n = 5) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (530 - 605 (590) long; 415 - 500 (455) wide) ovoid with navy blue coloration separated into anterior and posterior portions with faint orange medially. Anterio-medial platelets (130 - 147.5 (140) long; 50 - 62.5 (62.5) wide). Anterio-lateral platelets (170 - 192.5 (177.5) long; 60 - 77.5 (75) wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 290 - 365 (320 )). Dorsal plate proportions: dorsum length / width 1.21 - 1.30 (1.30); dorsal width / distance between Dgl- 4 1.34 - 1.43 (1.42); anterio-medial platelet length / width 2.24 - 2.60 (2.24); anterio-lateral platelet length / width 2.33 - 2.83 (2.37); anterio-lateral / anterio-medial length 1.25 - 1.33 (1.27). Gnathosoma - Subcapitulum (260 - 300 (290) long (ventral); 197.5 - 225 (215) long (dorsal); 105 - 122.5 (112.5) tall) colorless. Rostrum (112.5 - 127.5 (122.5) long; 37.5 - 42.5 (40) wide). Chelicerae (255 - 295 (282) long) with curved fangs (45 - 64 (64) long). Subcapitular proportions: ventral length / height 2.40 - 2.58 (2.58); rostrum length / width 2.94 - 3.06 (3.06). Pedipalps elongate (especially tibiae) with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (35 - 42.5 (40) long); femur (85 - 113.75 (113.75) long); genu (55 - 67.5 (67.5) long); tibia (85 - 107.5 (102.5) long; 20 - 22.5 (22.5) wide); tarsus (15 - 17.5 (17.5) long). Palpomere proportions: femur / genu 1.55 - 1.69 (1.69); tibia / femur 0.90 - 1.00 (0.90); tibia length / width 4.25 - 4.78 (4.56). Venter - (630 - 710 (710) long; 480 - 580 (498) wide) colorless. Gnathosomal bay (135 - 162.5 (157.5) long; 60 - 72.5 (70) wide). Cxgl- 4 far from apex. Medial suture (60 - 70 (62.5) long). Genital plates (135 - 155 (147.5) long; 120 - 130 (127.5) wide). Additional measurements: Cx- 1 (260 - 300 (280) long (total); 130 - 155 (131) long (medial )); Cx- 3 (330 - 375 (340) wide); anterior venter (195 - 235 (227.5) long). Ventral proportions: gnathosomal bay length / width 1.90 - 2.32 (2.25); anterior venter / genital field length 1.37 - 1.66 (1.54); anterior venter length / genital field width 1.56 - 1.86 (1.78); anterior venter / medial suture 3.25 - 3.64 (3.64). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
01174C58015DB3149D1BB69E6C655B21.taxon	etymology	Etymology. Specific epithet (raptoroides) refers to the long, thin pedipalp tibia and long genual / femoral tubercles, which are similar to members of the Raptor Group (rapio, L. to seize; - oides, G. resembling).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
01174C58015DB3149D1BB69E6C655B21.taxon	distribution	Distribution. Southeastern, Arizona and New Mexico (Figure 214).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
E29A77F2CF569E00A09297FBE9E73EC2.taxon	materials_examined	Material examined. HOLOTYPE (♀): from USA, Virginia, Washington County, Damascus; beside Rt. 58 just inside boundary of Mount Rogers National Recreation Area, (36 ° 38 ' 38 " N, 81 ° 45 ' 45 " W), 10 July 1990, by IM Smith, IMS 900085 A. PARATYPES (8 ♀; 7 ♂): Alabama, USA: 1 ♂ from Clay County, beside Forest Route 649, 0.8 km northeast of road from Campbell Springs to Forest Route 600, (33 ° 22 ' 22 " N, 85 ° 52 ' 52 " W), 3 July 1990, by IM Smith, IMS 900075 A * Maine, USA: 1 ♀ and 1 ♂ from Aroostook County, Ashland; beside Rt. 11 at bridge over Aroostook River, (46 ° 38 ' 38 " N, 68 ° 24 ' 24 " W), 4 July 1989, by IM Smith, IMS 890067 * New Brunswick, Canada: 2 ♀ and 1 ♂ from York County, beside Hwy. 8, 1.7 km north of road to Durham Bridge, (46 ° 7 ' 7 " N, 66 ° 36 ' 36 " W), 2 July 1989, by IM Smith, IMS 890058 * New York, USA: 1 ♀ and 1 ♂ from Schuyler County, beside Town Line Road off Route 228, 0.6 km south of Perry City, (42 ° 29 ' 29 " N, 76 ° 42 ' 42 " W), 21 July 1990, by IM Smith, IMS 900112 A * Tennessee, USA: 1 ♀ from Monroe County, beside Forest Route 35, 2.0 km northeast of road from Rt. 165 to Miller Chapel Baptist Church, (35 ° 21 ' 21 " N, 84 ° 9 ' 9 " W), 5 July 1990, by IM Smith, IMS 900078 * 2 ♀ and 2 ♂ from Monroe County, beside Forest Route 210, 1.8 km east of bridge at Bald River Falls, (35 ° 19 ' 19 " N, 84 ° 10 ' 10 " W), 5 July 1990, by IM Smith, IMS 900079 * Virginia, USA: 1 ♂ (ALLOTYPE) from Washington County, Damascus; beside Rt. 58 just inside boundary of Mount Rogers National Recreation Area, (36 ° 38 ' 38 " N, 81 ° 45 ' 45 " W), 10 July 1990, by IM Smith, IMS 900085 A * 1 ♀ from Washington County, Damascus; beside Rt. 58 just inside boundary of Mount Rogers National Recreation Area, (36 ° 38 ' 38 " N, 81 ° 45 ' 45 " W), 10 July 1990, by IM Smith, IMS 900085 A	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
E29A77F2CF569E00A09297FBE9E73EC2.taxon	diagnosis	Diagnosis. Torrenticola reduncarostra are similar to species with similar dorsal patterning, such as the Rusetria " 4 - Plate " group (T. dunni, T. glomerabilis, T. kittatinniana, T. pollani, T. rufoalba and T. shubini), Neoanomala Group (T. interiorensis and T. neoanomala), and T. bondi, T. gorti, T. elongata, T. erectirostra, T. robisoni, T. irapalpa, T. racupalpa, T. skvarlai, and T. arktonyx. T. reduncarostra can be differentiated from all Torrenticola, except Erectirostra Group, by having an upturned rostrum. T. reduncarostra can be differentiated from T. erectirostra, T. karambita, and T. robisoni by lacking dentation on the rostrum (others have strong dentation on the lateral edge of the rostrum).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
E29A77F2CF569E00A09297FBE9E73EC2.taxon	description	Description. Female (Figure 218) (n = 5) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (610 - 665 (625) long; 420 - 450 (420) wide) ovoid with faint purple coloration separated into anterior and posterior portions, occasionally colorless. Anterio-medial platelets (120 - 130 (120) long; 55 - 57.5 (55) wide). Anterio-lateral platelets (175 - 205 (185) long; 57.5 - 67.5 (60) wide) free from dorsal plate. Dgl- 4 approximately half-way between the edge of the dorsum and the muscle scars (distance between Dgl- 4 280 - 315 (290 )). Dorsal plate proportions: dorsum length / width 1.45 - 1.5 (1.49); dorsal width / distance between Dgl- 4 1.40 - 1.50 (1.45); anterio-medial platelet length / width 2.18 - 2.27 (2.18); anterio-lateral platelet length / width 2.59 - 3.42 (3.08); anterio-lateral / anterio-medial length 1.46 - 1.58 (1.54). Gnathosoma - Subcapitulum (330 - 340 (330) long (ventral); 240 - 255 (240) long (dorsal); 110 - 120 (120) tall) colorless. Rostrum (130 - 140 (132.5) long; 35 - 37.5 (35) wide) elongate and upturned. Chelicerae (345 - 360 (350) long) with curved fangs (55 - 60 (55) long). Subcapitular proportions: ventral length / height 2.75 - 3.09 (2.75); rostrum length / width 3.71 - 4.00 (3.79). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (45 - 50 (47.5) long); femur (115 - 120 (115) long); genu (62.5 - 65 (62.5) long); tibia (67.5 - 72.5 (70) long; 23.75 - 26.25 (26.25) wide); tarsus (16.25 - 20 (17.5) long). Palpomere proportions: femur / genu 1.81 - 1.85 (1.84); tibia / femur 0.58 - 0.62 (0.61); tibia length / width 2.67 - 2.85 (2.67). Venter - (740 - 800 (740) long; 470 - 515 (500) wide) colorless. Gnathosomal bay (145 - 165 (155) long; 65 - 70 (67.5) wide). Cxgl- 4 subapical. Medial suture (25 - 35 (25) long). Genital plates (170 - 192.5 (170) long; 150 - 157.5 (155) wide). Additional measurements: Cx- 1 (290 - 320 (305) long (total); 150 - 165 (155) long (medial )); Cx- 3 (325 - 350 (330) wide); anterior venter (190 - 210 (190) long). Ventral proportions: gnathosomal bay length / width 2.07 - 2.46 (2.30); anterior venter / genital field length 1.07 - 1.24 (1.12); anterior venter length / genital field width 1.23 - 1.40 (1.23); anterior venter / medial suture 6.00 - 8.10 (7.60). Male (Figure 219) (n = 5) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (480 - 580 (525) long; 295 - 360 (335) wide) ovoid with faint purple coloration separated into anterior and posterior portions, occasionally colorless. Anterio-medial platelets (95 - 115 (115) long; 42.5 - 52.5 (50) wide). Anterio-lateral platelets (165 - 195 (170) long; 52.5 - 55 (52.5) wide) free from dorsal plate. Dgl- 4 approximately halfway between the edge of the dorsum and the muscle scars (distance between Dgl- 4 215 - 255 (220 )). Dorsal plate proportions: dorsum length / width 1.51 - 1.68 (1.57); dorsal width / distance between Dgl- 4 1.37 - 1.52 (1.52); anterio-medial platelet length / width 2.19 - 2.30 (2.30); anterio-lateral platelet length / width 3.14 - 3.71 (3.24); anterio-lateral / anterio-medial length 1.48 - 1.79 (1.48). Gnathosoma - Subcapitulum (255 - 300 (290) long (ventral); 195 - 220 (207.5) long (dorsal); 87.5 - 95 (95) tall) colorless. Rostrum (102.5 - 120 (110) long; 27.5 - 32.5 (30) wide) elongate and upturned. Chelicerae (250 - 300 (300) long) with curved fangs (45 - 55 (55) long). Subcapitular proportions: ventral length / height 2.91 - 3.16 (3.05); rostrum length / width 3.54 - 3.73 (3.67). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (40 - 42.5 (42.5) long); femur (90 - 105 (102.5) long); genu (52.5 - 60 (58.75) long); tibia (62.5 - 72.5 (67.5) long; 22.5 - 25 (25) wide); tarsus (15 - 17.5 (17.5) long). Palpomere proportions: femur / genu 1.69 - 1.83 (1.74); tibia / femur 0.66 - 0.72 (0.66); tibia length / width 2.70 - 3.11 (2.70). Venter - (595 - 710 (650) long; 345 - 410 (370) wide) colorless. Gnathosomal bay (122.5 - 145 (137.5) long; 57.5 - 67.5 (57.5) wide). Cxgl- 4 subapical. Medial suture (75 - 105 (85) long). Genital plates (135 - 150 (140) long; 100 - 105 (102.5) wide). Additional measurements: Cx- 1 (250 - 280 (270) long (total); 130 - 140 (135) long (medial )); Cx- 3 (270 - 300 (295) wide); anterior venter (222.5 - 257.5 (245) long). Ventral proportions: gnathosomal bay length / width 1.96 - 2.39 (2.39); anterior venter / genital field length 1.53 - 1.81 (1.75); anterior venter length / genital field width 2.23 - 2.45 (2.39); anterior venter / medial suture 2.45 - 3.19 (2.88). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
E29A77F2CF569E00A09297FBE9E73EC2.taxon	etymology	Etymology. Specific epithet (reduncarostra) refers to the rostrum, which is curved upwards anteriorly, a rare condition outside of the Erectirostra Group (reduncus, L. bent backward; rostrum, L. snout).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
E29A77F2CF569E00A09297FBE9E73EC2.taxon	distribution	Distribution. Appalachians (Figure 217).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
BA11A5AF95017F05CBFEE5228313CA44.taxon	materials_examined	Material examined. HOLOTYPE (♂): from USA, Oregon, Curry County, Port Orford, beside Elk River Road, 9 km east of Elk River Fish Hatchery (42 ° 42 ' 22 " N, 124 ° 20 ' 28 " W), 18 Jun 2010, by IM Smith, IMS 100075, DNA 1442. PARATYPES (2 ♀; 6 ♂): Oregon, USA: 1 ♀ (ALLOTYPE) from Curry County, Port Orford; beside Elk River Road 9.0 km east of Elk River Fish Hatchery, (42 ° 42 ' 42 " N, 124 ° 20 ' 20 " W), 18 June 2010, by IM Smith, IMS 100075 * 1 ♀ and 6 ♂ from Curry County, Port Orford; beside Elk River Road 9.0 km east of Elk River Fish Hatchery, (42 ° 42 ' 42 " N, 124 ° 20 ' 20 " W), 18 June 2010, by IM Smith, IMS 100075	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
BA11A5AF95017F05CBFEE5228313CA44.taxon	diagnosis	Diagnosis. Torrenticola regalis can be differentiated from all other western Torrenticola by having purple coloration covering the entire dorsal plate and by being the only member of the Miniforma Complex with indistinct or partial hind coxal margins. Occasionally its coloration can look similar to T. tahoei, but T. regalis can be differentiated from T. tahoei by having a shorter anterior venter (200 - 265 in T. regalis, 285 - 325 in T. tahoei), a stockier subcapitulum (ventral length / height = 2.41 - 2.68 in T. regalis, 3.25 - 4.11 in T. tahoei), and by having indistinct hind coxal margins (distinct in T. tahoei).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
BA11A5AF95017F05CBFEE5228313CA44.taxon	description	Description. Female (Figure 221) (n = 2) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (680 - 695 (695) long; 540 - 540 (540) wide) circular with purple coloration, excepting platelets. Anterio-medial platelets (152.5 - 160 (160) long; 65 - 65 (65) wide). Anterio-lateral platelets (207.5 - 222.5 (222.5) long; 80 - 83.75 (83.75) wide) free from dorsal plate. Dgl- 4 much closer to the edge of dorsum than to the muscle scars (distance between Dgl- 4 405 - 420 (420 )). Dorsal plate proportions: dorsum length / width 1.26 - 1.29 (1.29); dorsal width / distance between Dgl- 4 1.29 - 1.33 (1.29); anterio-medial platelet length / width 2.35 - 2.46 (2.46); anterio-lateral platelet length / width 2.59 - 2.66 (2.66); anterio-lateral / anterio-medial length 1.36 - 1.39 (1.39). Gnathosoma - Subcapitulum (400 - 410 (410) long (ventral); 295 - 307.5 (307.5) long (dorsal); 160 - 170 (170) tall) colorless. Rostrum (155 - 165 (165) long; 52.5 - 55 (55) wide). Chelicerae (410 - 420 (420) long) with curved fangs (70 - 75 (70) long). Subcapitular proportions: ventral length / height 2.41 - 2.50 (2.41); rostrum length / width 2.95 - 3.00 (3.00). Pedipalps with tuberculate ventral extensions on femora and tuberculate ventral extensions with dentate tip on genua. Palpomeres: trochanter (45 - 47.5 (45) long); femur (132.5 - 132.5 (132.5) long); genu (80 - 85 (85) long); tibia (90 - 93.75 (93.75) long; 30 - 30 (30) wide); tarsus (17.5 - 17.5 (17.5) long). Palpomere proportions: femur / genu 1.56 - 1.66 (1.56); tibia / femur 0.68 - 0.71 (0.71); tibia length / width 3.00 - 3.13 (3.13). Venter - (860 - 860 (860) long; 630 - 630 (630) wide) colorless. Gnathosomal bay (197.5 - 205 (205) long; 75 - 80 (75) wide). Cxgl- 4 subapical. Medial suture (30 - 40 (30) long). Genital plates (225 - 232.5 (232.5) long; 205 - 205 (205) wide). Additional measurements: Cx- 1 (340 - 350 (350) long (total); 140 - 145 (145) long (medial )); Cx- 3 (395 - 405 (405) wide); anterior venter 200 - 200 (200) long). Ventral proportions: gnathosomal bay length / width 2.47 - 2.73 (2.73); anterior venter / genital field length 0.86 - 0.89 (0.86); anterior venter length / genital field width 0.98 - 0.98 (0.98); anterior venter / medial suture 5.00 - 6.67 (6.67). Male (Figure 222) (n = 5) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (580 - 640 (630) long; 465 - 520 (490) wide) circular with purple coloration, excepting platelets. Anterio-medial platelets (127.5 - 145 (140) long; 55 - 67.5 (67.5) wide). Anterio-lateral platelets (177.5 - 217.5 (203.75) long; 72.5 - 85 (82.5) wide) free from dorsal plate. Dgl- 4 much closer to the edge of dorsum than to the muscle scars (distance between Dgl- 4 375 - 420 (400 )). Dorsal plate proportions: dorsum length / width 1.23 - 1.29 (1.29); dorsal width / distance between Dgl- 4 1.23 - 1.27 (1.23); anterio-medial platelet length / width 2.07 - 2.32 (2.07); anterio-lateral platelet length / width 2.44 - 2.64 (2.47); anterio-lateral / anterio-medial length 1.39 - 1.50 (1.46). Gnathosoma - Subcapitulum (355 - 375 (365) long (ventral); 255 - 280 (270) long (dorsal); 132.5 - 150 (145) tall) colorless. Rostrum (140 - 150 (145) long; 45 - 55 (55) wide). Chelicerae (350 - 375 (365) long) with curved fangs (65 - 70 (67.5) long). Subcapitular proportions: ventral length / height 2.50 - 2.68 (2.52); rostrum length / width 2.64 - 3.16 (2.64). Pedipalps with tuberculate ventral extensions on femora and tuberculate ventral extensions with dentate tip on genua. Palpomeres: trochanter (41.25 - 50 (45) long); femur (113.75 - 125 (125) long); genu (72.5 - 82.5 (80) long); tibia (82.5 - 92.5 (92.5) long; 23.75 - 27.5 (27.5) wide); tarsus (17.5 - 17.5 (17.5) long). Palpomere proportions: femur / genu 1.48 - 1.57 (1.56); tibia / femur 0.71 - 0.74 (0.74); tibia length / width 3.36 - 3.50 (3.36). Venter - (730 - 780 (770) long; 535 - 640 (640) wide) colorless. Gnathosomal bay (160 - 185 (160) long; 72.5 - 92.5 (92.5) wide). Cxgl- 4 subapical. Medial suture (75 - 90 (90) long). Genital plates (185 - 190 (185) long; 145 - 155 (152.5) wide). Additional measurements: Cx- 1 (305 - 340 (320) long (total); 135 - 160 (160) long (medial )); Cx- 3 (350 - 410 (410) wide); anterior venter 237.5 - 265 (265) long). Ventral proportions: gnathosomal bay length / width 1.73 - 2.55 (1.73); anterior venter / genital field length 1.28 - 1.43 (1.43); anterior venter length / genital field width 1.62 - 1.74 (1.74); anterior venter / medial suture 2.64 - 3.47 (2.94). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
BA11A5AF95017F05CBFEE5228313CA44.taxon	etymology	Etymology. Specific epithet (regalis) refers to the coloration of this species, which is nearly colorless except the purple dorsum, giving the appearance of a mite wearing a purple cape, such as those often depicted as worn by royalty (rēgalis, L. royal).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
BA11A5AF95017F05CBFEE5228313CA44.taxon	distribution	Distribution. Known only from Curry County, Oregon (Figure 220).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
EB2822DBA6B378E9A4ADEA7C71B60E54.taxon	materials_examined	Material examined. HOLOTYPE (♀): from USA, Arkansas, Polk County, East Saline Creek, 30 Jul 2011, by IM Smith, IMS 110041 PARATYPES (1 ♀; 0 ♂): Oklahoma, USA: Pushmataha County, beside Route 271, Walnut Creek (34 ° 39 ' N 95 ° 7 ' W), 1 Jul 1987, by IM Smith, IMS 870063 A	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
EB2822DBA6B378E9A4ADEA7C71B60E54.taxon	diagnosis	Diagnosis. Torrenticola robisoni are similar to other members of the Erectirostra Group (T. karambita and T. erectirostra). They are similar to other species with similar dorsal patterning, but can be differentiated from these by having a dentate, upturned rostrum that is wide when viewed ventrally. T. robisoni can be differentiated from T. erectirostra by having more elongate anterio-lateral platelets (length / width ♀ = 2.96 - 3.00 in T. robisoni, 2.52 - 2.69 in T. erectirostra). T. robisoni can be differentiated from T. karambita by having dorsal coloration (T. karambita is colorless) and having a more elongate rostrum (length / width ♀ = 2.09 - 2.09 in T. robisoni, 1.57 - 1.62 in T. karambita). T. robisoni can be further differentiated from T. erectirostra and T. karambita by being distributed in the Interior Highlands, while the others are in the Appalachians.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
EB2822DBA6B378E9A4ADEA7C71B60E54.taxon	description	Description. Female (Figure 224) (n = 2) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum- (650 - 670 (650) long; 455 - 500 (455) wide) ovoid with reddish purple coloration separated into anterior and posterior portions with orange medially. Anterio-medial platelets (147.5 - 150 (150) long; 60 - 62.5 (60) wide). Anterio-lateral platelets (200 - 210 (200) long; 67.5 - 70 (67.5) wide) free from dorsal plate. Dgl- 4 closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 305 - 340 (305 )). Dorsal plate proportions: dorsum length / width 1.34 - 1.43 (1.34); dorsal width / distance between Dgl- 4 1.47 - 1.49 (1.49); anterio-medial platelet length / width 2.36 - 2.50 (2.50); anterio-lateral platelet length / width 2.96 - 3.00 (2.96); anterio-lateral / anterio-medial length 1.33 - 1.42 (1.33). Gnathosoma - Subcapitulum (325 - 335 (325) long (ventral); 218 - 250 (218) long (dorsal); 125 - 132.5 (132.5) tall) colorless. Rostrum (115 - 120 (115) long; 55 - 57.5 (55) wide) wide and upturned with dentation. Chelicerae (315 - 330 (315) long) with curved fangs (46 - 50 (46) long). Subcapitular proportions: ventral length / height 2.45 - 2.68 (2.45); rostrum length / width 2.09 - 2.09 (2.09). Pedipalps short and stocky (especially tibiae) with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (52.5 - 52.5 (52.5) long); femur (95 - 107.5 (95) long); genu (60 - 62.5 (60) long); tibia (47.5 - 58.75 (47.5) long; 25 - 27.5 (25) wide); tarsus (15 - 17.5 (15) long). Palpomere proportions: femur / genu 1.58 - 1.72 (1.58); tibia / femur 0.50 - 0.55 (0.50); tibia length / width 1.90 - 2.14 (1.90). Venter - (779 - 825 (779) long; 543 - 550 (543) wide) colorless. Gnathosomal bay (165 - 190 (165) long; 115 - 120 (115) wide). Cxgl- 4 far from apex. Medial suture (12.5 - 20 (12.5) long). Genital plates (168.75 - 202.5 (202.5) long; 157.5 - 180 (180) wide). Additional measurements: Cx- 1 (308 - 350 (308) long (total); 142 - 155 (142) long (medial )); Cx- 3 (367 - 375 (367) wide); anterior venter (182.5 - 200 (182.5) long). Ventral proportions: gnathosomal bay length / width 1.43 - 1.58 (1.43); anterior venter / genital field length 1.05 - 1.08 (1.08); anterior venter length / genital field width 1.16 - 1.25 (1.16); anterior venter / medial suture 10.00 - 14.60 (14.60). Male unknown. Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
EB2822DBA6B378E9A4ADEA7C71B60E54.taxon	etymology	Etymology. Specific epithet (robisoni) named in honor of Henry W. Robison for his efforts in communicating the importance of the Interior Highlands (Ozarks and Ouachitas) - the type locality-which have a high proportion of endemic species, yet are understudied with respect to other areas of increased endemism (e. g., California floristic province, coastal plains, southern Appalachians, Pacific Northwest). His contributions instill passion for the region to even the casual reader, and have inspired many, including JRF, to pursue biodiversity research in the area.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
EB2822DBA6B378E9A4ADEA7C71B60E54.taxon	distribution	Distribution. Known only from the Ouachita Mountains, possibly endemic (Figure 223).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
4317571272FA31EC2467B43B878D9711.taxon	materials_examined	Material examined. HOLOTYPE (♀): from USA, Idaho, Blaine County, Sawtooth National Forest, Salmon River (43 ° 53 ' 7 " N, 114 ° 46 ' 15 " W), 29 Jul 2012, by JR Fisher, WA Nelson, & JC O'Neill, ROW 12 - 0729 - 002, DNA 2623. PARATYPES (4 ♀; 6 ♂): Idaho, USA: 2 ♀ and 2 ♂ from Blaine County, Salmon River, beside Route 75 between Obsidian & Galena Summit, 3 Jul 1985, by IM Smith, IMS 850067 * 1 ♂ from Blaine County, Sawtooth National Forest, Salmon River (43 ° 53 ' 7 " N, 114 ° 46 ' 15 " W), 29 Jul 2012, by JR Fisher, WA Nelson, & JC O'Neill, ROW 12 - 0729 - 002 * 2 ♀ and 3 ♂ from Custer County, Basin Creek campground, beside Route 75 between Sunbeam & Stanley Basin Creek, 2 Jul 1985, by IM Smith, IMS 850066 * 1 ♀ and 1 ♂ from Custer County, Challis National Forest, Stanley Creek (44 ° 15 ' 12 " N, 115 ° 0 ' 19 " W), 30 Jul 2012, by JR Fisher, WA Nelson, & JC O'Neill, ROW 12 - 0730 - 005 * 1 ♀ and 1 ♂ from Custer County, Salmon River (44 ° 12 ' 31 " N, 114 ° 55 ' 51 " W), 29 Jul 2012, by JR Fisher, WA Nelson, & JC O'Neill, ROW 12 - 0729 - 003 * 1 ♂ from Custer County, Stanley, Little Redfish Lake, 3 Jul 1985, by IM Smith, IMS 850069 * 2 ♀ from Lemhi County, North Fork of Salmon River, beside Route 93, 15 kilometers north of North Fork, 1 Jul 1985, by IM Smith, IMS 850062 * 1 ♂ (ALLOTYPE) from Lemhi County, Salmon National Forest, Niapas Creek (45 ° 8 ' 15 " N, 114 ° 13 ' 4 " W), 2 Aug 2012, by JR Fisher, WA Nelson, & JC O'Neill, ROW 12 - 0802 - 003, DNA 2626 * 1 ♀ from Lemhi County, Salmon National Forest, Niapas Creek (45 ° 8 ' 15 " N, 114 ° 13 ' 4 " W), 2 Aug 2012, by JR Fisher, WA Nelson, & JC O'Neill, ROW 12 - 0802 - 003 * Montana, USA 1 ♂ from Missoula County, Lolo National Forest, Lolo Creek (46 ° 41 ' 51 " N, 114 ° 32 ' 34 " W), 7 Aug 2012, by JR Fisher, WA Nelson & JC O'Neill, ROW 12 - 0807 - 002 * 1 ♀ from Missoula County, Lolo National Forest, Lolo Creek (46 ° 46 ' 7 " N, 114 ° 27 ' 53 " W), 7 Aug 2012, by JR Fisher, WA Nelson & JC O'Neill, ROW 12 - 0807 - 003 * 1 ♂ from Ravalli County, Bitterroot National Forest, Soda Spring Creek (45 ° 47 ' 12 " N, 114 ° 21 ' 2 " W), 6 Aug 2012, by JR Fisher, WA Nelson & JC O'Neill, ROW 12 - 0806 - 001 * 1 ♀ and 1 ♂ from Ravalli County, Medicine Springs, Spring Gulch campground, East Fork of Bitterroot River, beside Route 93, 1 Jul 1985, by IM Smith, IMS 850060.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
4317571272FA31EC2467B43B878D9711.taxon	diagnosis	Diagnosis. Torrenticola rockyensis are similar to members of the Miniforma group (T. copipalpa, T. manni, T. miniforma, T. pacificensis, T. oliveri, and T. pinocchio) in having short, stocky pedipalps (except T. oliveri and T. pinocchio); similar pedipalpal extensions (unique to members of this group); and being among the smallest Torrenticola in the west (dorsum 500 - 625 long) (except T. oliveri). T. rockyensis can be differentiated from all other Miniforma group by being distributed in the Rocky Mountains. T. rockyensis are best differentiated from T. copipalpa by having tuberculate pedipalp femoral extensions (broad and flat in T. copipalpa). T. rockyensis are best differentiated from T. pacificensis by females having more elongate subcapitular rostra (length / width ♀ = 2.72 - 2.91 in T. rockyensis, 2.59 - 2.68 in T. pacificensis). T. rockyensis are best differentiated from T. manni by having stockier pedipalp tibiae (length / width = 2.47 - 3.11 in T. rockyensis, 3.13 - 3.38 in T. manni). T. rockyensis are best differentiated from T. miniforma by being larger (dorsum length ♀ = 570 - 620 in T. rockyensis, 545 in T. miniforma; ♂ = 525 - 545 in T. rockyensis, 485 in T. miniforma) and having stockier subcapitular rostra (length / width ♀ = 2.72 - 2.91 in T. rockyensis, 3.13 in T. miniforma; ♂ = 2.83 - 3.00 in T. rockyensis, 3.19 in T. miniforma). T. rockyensis can be differentiated by having a shorter anterior venter (175 - 223 in T. rockyensis, 250 - 310 in T. oliveri) and less elongate pedipalpal tibiae (length / width = 2.4 - 3.1 in T. rockyensis, 3.6 - 4.2 in T. oliveri). T. rockyensis can be differentiated from T. pinocchio by having a less elongate rostrum (length / width = 2.7 - 3.0 in T. rockyensis, 4.5 - 4.9 in T. pinocchio).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
4317571272FA31EC2467B43B878D9711.taxon	description	Description. Female (Figure 226) (n = 5) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (570 - 620 (620) long; 400 - 420 (400) wide) ovoid with purple coloration often restricted posteriorly, occasionally encroaching anteriorly nearly to the platelets (one specimen with purple on the platelets). Anterio-medial platelets (110 - 120 (120) long; 43.75 - 52.5 (52.5) wide). Anterio-lateral platelets (160 - 172.5 (172.5) long; 52.5 - 62.5 (62.5) wide) free from dorsal plate. Dgl- 4 much closer to the edge of dorsum than to the muscle scars (distance between Dgl- 4 290 - 310 (300 )). Dorsal plate proportions: dorsum length / width 1.43 - 1.55 (1.55); dorsal width / distance between Dgl- 4 1.33 - 1.38 (1.33); anterio-medial platelet length / width 2.19 - 2.51 (2.29); anterio-lateral platelet length / width 2.76 - 3.05 (2.76); anterio-lateral / anterio-medial length 1.43 - 1.55 (1.44). Gnathosoma - Subcapitulum (300 - 315 (302.5) long (ventral); 222.5 - 235 (231) long (dorsal); 115 - 122.5 (117.5) tall) colorless. Rostrum (117.5 - 130 (123.75) long; 42.5 - 45 (42.5) wide). Chelicerae (283 - 310 (283) long) with curved fangs (52.5 - 68 (67) long). Subcapitular proportions: ventral length / height 2.49 - 2.68 (2.57); rostrum length / width 2.72 - 2.91 (2.91). Pedipalps short and stocky (especially tibiae) with tuberculate (occasionally broadly tuberculate), dentate ventral extensions on femora and dentate, flanged ventral extensions on genua. Palpomeres: trochanter 35 - 42.5 (42.5) long); femur (90 - 95 (95) long); genu (63.75 - 67.5 (67.5) long); tibia (62.5 - 70 (70) long; 21.25 - 23.75 (22.5) wide); tarsus (12.5 - 17.5 (15) long). Palpomere proportions: femur / genu 1.38 - 1.46 (1.41); tibia / femur 0.68 - 0.74 (1.74); tibia length / width 2.78 - 3.11 (3.11). Venter - (615 - 770 (760) long; 339 - 470 (455) wide) colorless. Gnathosomal bay (120 - 147.5 (145) long; 65 - 80 (67.5) wide). Cxgl- 4 subapical. Medial suture (42.5 - 60 (52.5) long). Genital plates (172.5 - 190 (185) long; 157.5 - 173.75 (170) wide). Additional measurements: Cx- 1 (239 - 279 (279) long (total); 112 - 133 (121) long (medial )); Cx- 3 (247 - 310 (296) wide); anterior venter (175 - 202.5 (190) long). Ventral proportions: gnathosomal bay length / width 1.71 - 2.27 (2.15); anterior venter / genital field length 0.99 - 1.07 (1.03); anterior venter length / genital field width 1.08 - 1.19 (1.12); anterior venter / medial suture 3.13 - 4.29 (3.62). Male (Figure 227) (n = 5) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (525 - 545 (535) long; 335 - 350 (350) wide) ovoid with purple coloration often restricted posteriorly, occasionally encroaching anteriorly nearly to the platelets. Anterio-medial platelets (102.5 - 106.25 (105) long; 45 - 47.5 (45) wide). Anterio-lateral platelets (145 - 155 (152.5) long; 52.5 - 55 (52.5) wide) free from dorsal plate. Dgl- 4 much closer to the edge of dorsum than to the muscle scars (distance between Dgl- 4 255 - 285 (280 )). Dorsal plate proportions: dorsum length / width 1.53 - 1.57 (1.53); dorsal width / distance between Dgl- 4 1.23 - 1.31 (1.25); anterio-medial platelet length / width 2.21 - 2.36 (2.33); anterio-lateral platelet length / width 2.68 - 2.90 (2.90); anterio-lateral / anterio-medial length 1.36 - 1.51 (1.45). Gnathosoma - Subcapitulum (270 - 285 (285) long (ventral); 197.5 - 208 (207) long (dorsal); 102.5 - 107.5 (107.5) tall) colorless. Rostrum (106.25 - 112.5 (112.5) long; 36.25 - 38.75 (37.5) wide). Chelicerae (269 - 284 (282) long) with curved fangs (41 - 52 (46) long). Subcapitular proportions: ventral length / height 2.62 - 2.67 (2.65); rostrum length / width 2.83 - 3.00 (3.00). Pedipalps short and stocky (especially tibiae) with tuberculate (occasionally broadly tuberculate), dentate ventral extensions on femora and dentate, flanged ventral extensions on genua. Palpomeres: trochanter (30 - 40 (31.25) long); femur (81.25 - 86.25 (86.25) long); genu (57.5 - 60 (60) long); tibia (52.5 - 62.5 (62.5) long; 21.25 - 22.5 (22.5) wide); tarsus (12.5 - 15 (15) long). Palpomere proportions: femur / genu 1.41 - 1.44 (1.44); tibia / femur 0.62 - 0.74 (0.72); tibia length / width 2.47 - 2.82 (2.78). Venter - (580 - 690 (690) long; 352 - 420 (385) wide) colorless. Gnathosomal bay (130 - 140 (140) long; 57.5 - 62.5 (62.5) wide). Cxgl- 4 subapical. Medial suture (75 - 92.5 (92.5) long). Genital plates (137.5 - 147.5 (143.75) long; 105 - 113.75 (110) wide). Additional measurements: Cx- 1 (234 - 266 (266) long (total); 98.25 - 137 (136) long (medial )); Cx- 3 (245 - 270 (260) wide); anterior venter (197.5 - 222.5 (215) long). Ventral proportions: gnathosomal bay length / width 2.12 - 2.26 (2.24); anterior venter / genital field length 1.42 - 1.51 (1.50); anterior venter length / genital field width 1.82 - 1.98 (1.95); anterior venter / medial suture 2.32 - 2.73 (2.32). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
4317571272FA31EC2467B43B878D9711.taxon	etymology	Etymology. Specific epithet (rockyensis) refers to the distribution of this species in the Rocky Mountains. This location-based naming reflects that locality is the easiest way to differentiate this species from others in the Miniforma Group, particularly T. pacificensis.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
4317571272FA31EC2467B43B878D9711.taxon	distribution	Distribution. Rocky Mountains of Idaho and Montana (Figure 225).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
B7509B1B316247B8D21629B0B62F7246.taxon	materials_examined	Material examined. HOLOTYPE (♂): from USA, New Jersey, Morris County, Brook, Brookside, 20 May 1953, by H Habeeb. PARATYPES (1 ♀ and 0 ♂): New Jersey, USA: 1 ♀ (ALLOTYPE) from Sussex County, Little Flatbrook, north of Bevans, 12 Oct 1953, by H Habeeb.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
B7509B1B316247B8D21629B0B62F7246.taxon	diagnosis	Diagnosis. Torrenticola rufoalba are similar to other members of the Rusetria " 4 - Plates " group (T. dunni, T. glomerabilis, T. kittatinniana, T. pollani, and T. shubini) and T. skvarlai in having anterio-lateral platelets free from the dorsal plate, dorsal coloration separated into anterior and posterior portions, and indistinct hind coxal margins. Male T. rufoalba can be differentiated from other male Rusetria four plates by having a shorter anterior venter (195 in T. rufoalba, 215 - 285 in others). T. rufoalba can be differentiated from T. dunni by having a shorter dorsum (♀ = 550 in T. rufoalba, 605 - 680 in T. dunni; ♂ = 440 in T. rufoalba, 500 - 540 in T. dunni) and a thinner dorsum (♀ = 400 in T. rufoalba, 440 - 490 in T. dunni; ♂ = 320 in T. rufoalba, 350 - 370 in T. dunni). T. rufoalba can be differentiated from T. pollani by having a stockier rostrum (length / width = 2.96 - 3.06 in T. rufoalba, 3.27 - 3.82 in T. pollani). Female T. rufoalba can be differentiated from female T. pollani by having stockier tibiae (length / width = 3.5 in T. rufoalba, 3.8 - 4.2 in T. pollani). T. rufoalba can be differentiated from T. shubini by having a more elongate rostrum (length / width = 2.96 - 3.06 in T. rufoalba, 2.24 - 2.92 in T. shubini). Female T. rufoalba can be differentiated from female T. shubini by having a wider genital field (157.5 in T. rufoalba, 137 - 145 in T. shubini). T. rufoalba can be differentiated from T. glomerabilis and by having more elongate anterio-medial platelets (length / width = 2.45 - 2.61 in T. rufoalba, 1.9 - 2.3 in T. glomerabilis) and thinner dorsum (♀ = 400 in T. rufoalba, 460 - 490 in T. glomerabilis; ♂ = 320 in T. rufoalba, 395 - 430 in T. glomerabilis). T. rufoalba can be differentiated from T. kittatinniana by having a shorter dorsum (♀ = 550 in T. rufoalba, 640 in T. kittatinniana; ♂ = 440 in T. rufoalba, 500 in T. kittatinniana) and stockier anterio-medial platelets (length / width = 2.45 - 2.61 in T. rufoalba, 2.83 - 2.88 in T. kittatinniana). T. rufoalba can be differentiated from T. skvarlai by having a conical pedipalpal femoral tubercle, whereas T. skvarlai has a broad and flat pedipalpal femoral tubercle.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
B7509B1B316247B8D21629B0B62F7246.taxon	description	Re-description. Female (Figure 229) (n = 1) (allotype only) with characters of the genus with following specifications. Dorsum - (550 long; 400 wide) ovoid with reddish-purple coloration separated into anterior and posterior portions with a strip of orange medially. Anterio-medial platelets (107.5 long; 41.25 wide). Anterio-lateral platelets (168.75 long; 55 wide) free from dorsal plate. Dgl- 4 closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 255). Dorsal plate proportions: dorsum length / width 1.38; dorsal width / distance between Dgl- 4 1.57; anterio-medial platelet length / width 2.61; anterio-lateral platelet length / width 3.07; anterio-lateral / anterio-medial length 1.57. Gnathosoma - Subcapitulum (310 long (ventral); 235 long (dorsal); 127.5 tall) colorless. Rostrum (130 long; 42.5 wide). Chelicerae (315 long) with curved fangs (62.5 long). Subcapitular proportions: ventral length / height 2.43; rostrum length / width 3.06. Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (42.5 long); femur (115 long); genu (65 long); tibia (87.5 long; 25 wide); tarsus (17.5 long). Palpomere proportions: femur / genu 1.77; tibia / femur 0.76; tibia length / width 3.50. Venter - (640 long; 450 wide) mostly colorless with reddish-purple coloration in areas surrounding coxae. Gnathosomal bay (142.5 long; 92.5 wide). Cxgl- 4 subapical. Medial suture (17.5 long). Genital plates (167.5 long; 155 wide). Additional measurements: Cx- 1 (125 long (total); 125 long (medial )); Cx- 3 (335 wide); anterior venter (155 long). Ventral proportions: gnathosomal bay length / width 1.54; anterior venter / genital field length 0.93; anterior venter length / genital field width 1.00; anterior venter / medial suture 8.86. Male (Figure 230) (n = 1) (holotype only) with characters of the genus with following specifications. Dorsum - (440 long; 320 wide) ovoid with reddish-purple coloration separated into anterior and posterior portions with a strip of orange medially. Anterio-medial platelets (95 long; 38.75 wide). Anterio-lateral platelets (142.5 long; 47.5 wide) free from dorsal plate. Dgl- 4 closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 237.5). Dorsal plate proportions: dorsum length / width 1.38; dorsal width / distance between Dgl- 4 1.35; anterio-medial platelet length / width 2.45; anterio-lateral platelet length / width 3.00; anterio-lateral / anterio-medial length 1.50. Gnathosoma - Subcapitulum (257.5 long (ventral); 190 long (dorsal); 95 tall) colorless. Rostrum (103.75 long; 35 wide). Chelicerae (237.5 long) with curved fangs (50 long). Subcapitular proportions: ventral length / height 2.71; rostrum length / width 2.96. Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (35 long); femur (92.5 long); genu (55 long); tibia (76.25 long; 21.25 wide); tarsus (17.5 long). Palpomere proportions: femur / genu 1.68; tibia / femur 0.82; tibia length / width 3.59. Venter - (530 long; 400 wide) mostly colorless with reddish-purple coloration in areas surrounding coxae. Gnathosomal bay (120 long; 67.5 wide). Cxgl- 4 subapical. Medial suture (72.5 long). Genital plates (120 long; 97.5 wide). Additional measurements: Cx- 1 (235 long (total); 112.5 long (medial )); Cx- 3 (280 wide); anterior venter (195 long). Ventral proportions: gnathosomal bay length / width 1.78; anterior venter / genital field length 1.63; anterior venter length / genital field width 2.00; anterior venter / medial suture 2.69. Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
B7509B1B316247B8D21629B0B62F7246.taxon	etymology	Etymology. Habeeb (1955) did not specify an etymology for this specific epithet (rufoalba). However, we speculate that it refers to the dorsal coloration as Habeeb (1955) notes, " color red and white " (rufus, L. red; albus, L. white).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
B7509B1B316247B8D21629B0B62F7246.taxon	distribution	Distribution. Northern New Jersey (Figure 228).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
18D6CE7482E32F84B1F9C7BBF054A698.taxon	materials_examined	Material examined. HOLOTYPE (♀): from USA, Pennsylvania, Fayette County, Ohiopyle State Park, Laurel Run (39 ° 50 ' 58 " N, 79 ° 30 ' 51 " W), 10 Aug 2014, by MJ Skvarla, MS 14 - 0810 - 005, DNA 2831. PARATYPES (18 ♀; 8 ♂): Manitoba, Canada: 1 ♀ and 1 ♂ from Eating Point Creek, 0.3 kilometers east of Highway 6 (53 ° 15 ' 7 " N, 99 ° 18 ' 54 " W), 4 Jul 2009, by IM Smith, IMS 090030 A * 1 ♀ and 1 ♂ from Ochre River, beside Highway 5 (51 ° 3 ' N, 99 ° 46 ' W), 6 Sep 1970, by DW Barr & H Frania, ROM 700638 * New Mexico, USA: 2 ♀ from Catron County, Whitewater Creek, Glenwood Whitewater Picnic Area, 5 May 2012, by IM Smith, IMS 120005 * 1 ♀ from Grant County, east fork of Gila River, Grapevine Recreation Area off Route 15 north of Silver City, 5 May 2012, by IM Smith, IMS 120007 * Ohio, USA: 2 ♀ and 1 ♂ from Hocking County, beside road near Ash Cave, East Branch of Queer Creek (39 ° 24 ' N, 82 ° 33 ' W), 5 May 1993, by IM Smith & DR Cook, IMS 930001 A * Pennsylvania, USA: 1 ♂ (ALLOTYPE) from Fayette County, Ohiopyle State Park, Laurel Run (39 ° 50 ' 58 " N, 79 ° 30 ' 51 " W), 10 Aug 2014, by MJ Skvarla, MS 14 - 0810 - 005, DNA 2835 * 1 ♂ from Fayette County, Ohiopyle State Park, Laurel Run (39 ° 50 ' 58 " N, 79 ° 30 ' 51 " W), 10 Aug 2014, by MJ Skvarla, MS 14 - 0810 - 005 * 2 ♀ from Westmoreland County, Irwin Park (40 ° 19 ' 38 " N, 79 ° 42 ' 30 " W), 4 Aug 2014, by MJ Skvarla, MS 14 - 0804 - 004 * 4 ♀ from Somerset County, Laurel Hill State Park Laurel Hill Creek (40 ° 1 ' 6 " N, 79 ° 14 ' 4 " W), 8 Aug 2014, by MJ Skvarla, MS 14 - 0808 - 001 * Saskatchewan, Canada: 2 ♀ and 1 ♂ from Smeaton Torch River, beside Highway 106, 26.2 km north of Highway 55, 20 Aug 2012, by IM Smith, IMS 120079 * South Dakota, USA: 2 ♀ and 2 ♂ from Pennington County, Willow Creek Trail Head off Route 244, Willow Creek (43 ° 54 ' N, 103 ° 32 ' W), 10 Sep 1999, by IM Smith, IMS 990033.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
18D6CE7482E32F84B1F9C7BBF054A698.taxon	diagnosis	Diagnosis. Torrenticola sellersorum are similar to other members of the Rusetria " Eastern 2 - Plates " group (T. biscutella, T. caerulea, T. delicatexa, T. indistincta, T. malarkeyorum, T. pendula, T. tysoni, T. ululata, T. whitneyae, T. microbiscutella, and T. feminellai) in having anterio-lateral platelets fused to the dorsal plate, having dorsal coloration separated into anterior and posterior portions (except T. ululata and T. indistincta), and being distributed in the east. It is one of only four Eastern 2 - Plates that have dark, bold, bluish-purple coloration (also T. tysoni, T. biscutella, and T. pendula). T. sellersorum can be differentiated from T. ululata, T. indistincta, and T. feminellai by dorsal coloration and pattern. T. sellersorum can be differentiated from T. tysoni by having a stockier rostrum (length / width ♀ = 2.44 - 2.68 in T. sellersorum, 3.06 - 3.31 in T. tysoni; ♂ = 2.71 - 3.05 in T. sellersorum, 3.14 - 3.50 in T. tysoni). Female T. sellersorum can be differentiated from female T. biscutella by anterior venter / genital field length (0.69 - 0.77 in T. sellersorum, 0.82 - 0.88 in T. biscutella). Male T. sellersorum can be differentiated from male T. biscutella by having slightly more elongate anterio-lateral platelets (length / width = 2.76 - 3.00 in T. sellersorum, 2.58 - 2.74 in T. biscutella). T. sellersorum can be differentiated from T. pendula by having a stockier gnathosomal bay (length / width = 1.56 - 2.08 in T. sellersorum, 2.42 - 2.90 in T. pendula) and more elongate tibiae (length / width = 3.13 - 3.8 in T. sellersorum, 2.78 - 3.05 in T. pendula). T. sellersorum can be differentiated from T. whitneyae by having more elongate pedipalpal tibiae (length / width = 3.13 - 3.80 in T. sellersorum, 2.42 - 2.95 in T. whitneyae). T. sellersorum can be differentiated from T. microbiscutella by having a less elongate dorsum (length / width = 1.23 - 1.54 in T. sellersorum, 1.63 - 1.75 in T. microbiscutella). Female T. sellersorum can be differentiated from female T. malarkeyorum by anterior venter / genital field length (0.69 - 0.77 in T. sellersorum, 0.85 - 0.89 in T. malarkeyorum). Female T. sellersorum can be differentiated from female T. caerulea by having a wider genital field (150 - 182 in T. sellersorum, 120 - 145 in T. caerulea). Female T. sellersorum can be differentiated from female T. delicatexa by having a rounder dorsum (length / width = 1.23 - 1.37 in T. sellersorum, 1.38 - 1.44 in T. delicatexa). Male T. sellersorum do not have any measurement differences with male T. malarkeyorum, T. caerulea, and T. delicatexa; however, they can be differentiated by dorsal coloration.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
18D6CE7482E32F84B1F9C7BBF054A698.taxon	description	Description. Female (Figure 232) (n = 5) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (540 - 650 (540) long; 400 - 520 (400) wide) ovoid with bold (occasionally faint) bluish-purple coloration separated into anterior and posterior portions with a thin or thick strip of red medially. Anterio-medial platelets (112.5 - 135 (117.5) long; 35 - 50 (35) wide). Anterio-lateral platelets (147.5 - 172.5 (147.5) long; 47.5 - 75 (47.5) wide) fused to dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 290 - 370 (295 )). Dorsal plate proportions: dorsum length / width 1.23 - 1.37 (1.35); dorsal width / distance between Dgl- 4 1.35 - 1.42 (1.36); anterio-medial platelet length / width 2.65 - 3.36 (3.36); anterio-lateral platelet length / width 2.23 - 3.11 (3.11); anterio-lateral / anterio-medial length 1.26 - 1.36 (1.26). Gnathosoma - Subcapitulum (282.5 - 330 (285) long (ventral); 210 - 245 (219) long (dorsal); 130 - 157.5 (135) tall) colorless. Rostrum (110 - 125 (113.75) long; 42.5 - 47.5 (42.5) wide). Chelicerae 280 - 325 (280) long) with curved fangs (51 - 66 (62) long). Subcapitular proportions: ventral length / height 2.05 - 2.17 (2.11); rostrum length / width 2.44 - 2.68 (2.68). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (40 - 50 (42.5) long); femur (107.5 - 122.5 (110) long); genu (62.5 - 72.5 (65) long); tibia (82.5 - 95 (85) long; 22.5 - 27.5 (22.5) wide); tarsus (17.5 - 20 (20) long). Palpomere proportions: femur / genu 1.69 - 1.85 (1.69); tibia / femur 0.75 - 0.79 (0.77); tibia length / width 3.14 - 3.80 (3.78). Venter - (640 - 800 (645) long; 449 - 529 (470) wide) colorless. Gnathosomal bay (132.5 - 187.5 (157.5) long; 82.5 - 97.5 (82.5) wide). Cxgl- 4 subapical. Medial suture (0 - 10 (0) long) often absent. Genital plates (160 - 195 (165) long; 150 - 181.25 (157.5) wide). Additional measurements: Cx- 1 (230 - 322 (270) long (total); 83 - 126 (106) long (medial )); Cx- 3 (310 - 365 (319) wide); anterior venter (120 - 142.5 (120) long). Ventral proportions: gnathosomal bay length / width 1.56 - 2.08 (1.91); anterior venter / genital field length 0.69 - 0.77 (0.73); anterior venter length / genital field width 0.74 - 0.80 (0.76); anterior venter / medial suture (proportion cannot be calculated for specimens without a medial suture) 12.25 - 17.00. Male (Figure 233) (n = 5) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (390 - 470 (400) long; 255 - 330 (280) wide) ovoid with bold (occasionally faint) bluish-purple coloration separated into anterior and posterior portions with a thin or thick strip of red medially. Anterio-medial platelets (82.5 - 97.5 (82.5) long; 30 - 40 (30) wide). Anterio-lateral platelets (112.5 - 145 (120) long; 37.5 - 52.5 (42.5) wide) fused to dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 190 - 250 (220 )). Dorsal plate proportions: dorsum length / width 1.42 - 1.54 (1.43); dorsal width / distance between Dgl- 4 1.27 - 1.34 (1.27); anterio-medial platelet length / width 2.44 - 2.83 (2.75); anterio-lateral platelet length / width 2.76 - 3.00 (2.82); anterio-lateral / anterio-medial length 1.32 - 1.49 (1.45). Gnathosoma - Subcapitulum (205 - 245 (215) long (ventral); 151 - 179 (166) long (dorsal); 80 - 107.5 (91.25) tall) colorless. Rostrum 80 - 95 (85) long; 27.5 - 35 (30) wide). Chelicerae (195 - 235 (208) long) with curved fangs (40 - 50 (46) long). Subcapitular proportions: ventral length / height 2.24 - 2.59 (2.36); rostrum length / width 2.71 - 3.05 (2.83). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (30 - 38.75 (30) long); femur (75 - 87.5 (80) long); genu (46.25 - 52.5 (50) long); tibia (62.5 - 70 (67.5) long; 18.75 - 21.25 (20) wide); tarsus (15 - 17.5 (15) long). Palpomere proportions: femur / genu 1.60 - 1.67 (1.60); tibia / femur 0.79 - 0.87 (0.84); tibia length / width 3.13 - 3.47 (3.38). Venter - (465 - 570 (490) long; 290 - 367 (325) wide) colorless. Gnathosomal bay (105 - 135 (117.5) long; 57.5 - 66.25 (65) wide). Cxgl- 4 subapical. Medial suture (57.5 - 72.5 (57.5) long). Genital plates (102.5 - 122.5 (107.5) long; 91.25 - 115 (96.25) wide). Additional measurements: Cx- 1 (189 - 237 (199) long (total); 79 - 96 (85) long (medial )); Cx- 3 (230 - 290 (232) wide); anterior venter (150 - 185 (150) long). Ventral proportions: gnathosomal bay length / width 1.75 - 2.08 (1.81); anterior venter / genital field length 1.39 - 1.59 (1.40); anterior venter length / genital field width 1.55 - 1.78 (1.56); anterior venter / medial suture 2.32 - 2.61 (2.61). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
18D6CE7482E32F84B1F9C7BBF054A698.taxon	etymology	Etymology. Specific epithet (sellersorum) named in honor of the grandparents of DMF, who always supported and encouraged her love of nature and discovery.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
18D6CE7482E32F84B1F9C7BBF054A698.taxon	distribution	Distribution. Widespread (Figure 231). T. sellersorum are unlike all other Torrenticola in our collections in that they span either side of the Great Plains, which act as a barrier for most other species. T. sellersorum are likely absent from cool, highland streams of the Appalachians, Interior Highlands, and Rocky Mountains, but are likely widely distributed throughout the Interior. Explaining the peculiar distribution of T. sellersorum will depend on knowledge of dispersal capabilities, including host preference by the larvae and the dispersal capabilities of the host.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
B3129E05902983A06FF481551A47BDD1.taxon	materials_examined	Material examined. HOLOTYPE (♀): from USA, New Mexico, Catron County, Gila Hot Springs, Little Creek Recreational Area, Little Creek, off Route 15, 6 May 2012, by IM Smith, IMS 120006, DNA 2891. PARATYPES (13 ♀; 11 ♂): Arizona, USA: 2 ♀ and 2 ♂ from Coconino County, Oak Creek Canyon, Oak Creek, beside Route 89 A, between Banjo Bill & Bootlegger campgrounds, 21 Jul 1987, by IM Smith, IMS 870100 A & IMS 870100 B * 1 ♀ and 1 ♂ from Yavapai County, West Clear Creek at Clear Creek campground, off Forest Road 9, east of Camp Verde, 19 - 22 Jul 1987, by IM Smith, IMS 870098 & IMS 870102 * New Mexico, USA: 1 ♀ and 1 ♂ from Catron County, Gila Cliff Dwellers National Monument, West Fork of Gila River, 10 Jul 1987, by IM Smith, IMS 870080 A * 1 ♂ (ALLOTYPE) from Catron County, Gila Hot Springs, Little Creek, Little Creek Recreational Area off Route 15, 6 May 2012, by IM Smith, IMS 120006, DNA 2892 * 1 ♀ and 1 ♂ from Catron County, Gila River, beside Route 15, just below mouth of Little Creek, 11 Jul 1987, by IM Smith, IMS 870083 A * 2 ♀ and 1 ♂ from Catron County, Gila Hot Springs, Little Creek, Little Creek Recreational Area off Route 15, 6 May 2012, by IM Smith, IMS 120006 * 2 ♀ and 3 ♂ from Catron County, Little Creek, beside Route 15, 65 kilometers north of Route 180 in Silver City, 10 Jul 1987, by IM Smith, IMS 870081 A * 3 ♀ and 2 ♂ from Grant County, Gila River Recreation Area, East Fork of Gila River, beside Route 15, 11 Jul 1987, by IM Smith, IMS 870082 A * 1 ♀ and 1 ♂ from Grant County, Grapevine Recreational Area, East Fork of Gila River, off Route 15 north of Silver City, 5 May 2012, by IM Smith, IMS 120007.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
B3129E05902983A06FF481551A47BDD1.taxon	diagnosis	Diagnosis. Torrenticola sharkeyi is unlike all other western species by having the following combination of characters: dorsal coloration, although faint, restricted posteriorly; indistinct hind coxal margins; and ellipsoid body. It is only known from Catron & Grant Counties, New Mexico. It is most similar to members of the Rala Group (T. rala, T. keesdavidsi, T. kurtvietsi, T. boettgeri, T. lamellipalpis, T. dolichodactyla, and T. anoplopalpa); however, members of this group are colorless. T. sharkeyi can be differentiated from T. boettgeri, T. kurtvietsi, T. lamellipalpis, T. keesdavidsi, and T. anoplopalpa by having tuberculate ventral extensions on the pedipalpal femora (others are lacking extensions, have dentate flanged extensions or wide lamellate extensions). T. sharkeyi can be differentiated from T. rala, T. kurtvietsi, T. boettgeri, T. dolichodactyla and T. anoplopalpa by having a more elongate pedipalpal tibiae (length / width = 3.81 - 4.10 in T. sharkeyi, 1.75 - 3.38 in others). T. sharkeyi can be differentiated from T. kurtvietsi, T. keesdavidsi, T. lamellipalpis, and T. anoplopalpa by having a stockier subcapitulum (ventral length / width = 2.03 - 2.40 in T. sharkeyi, 2.51 - 4.16 in others).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
B3129E05902983A06FF481551A47BDD1.taxon	description	Description. Female (Figure 235) (n = 5) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (650 - 750 (715) long; 450 - 530 (490) wide) ovoid with faint bluish-purple coloration restricted posteriorly or colorless. Anterio-medial platelets (130 - 155 (137.5) long; 50 - 57.5 (52.5) wide). Anterio-lateral platelets (187.5 - 225 (202.5) long; 62.5 - 75 (67.5) wide) free from dorsal plate. Dgl- 4 closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 305 - 360 (335 )). Dorsal plate proportions: dorsum length / width 1.41 - 1.46 (1.46); dorsal width / distance between Dgl- 4 1.38 - 1.52 (1.46); anterio-medial platelet length / width 2.60 - 2.77 (2.62); anterio-lateral platelet length / width 2.91 - 3.08 (3.00); anterio-lateral / anterio-medial length 1.39 - 1.47 (1.47). Gnathosoma - Subcapitulum (300 - 335 (310) long (ventral); 215 - 245 (227) long (dorsal); 135 - 157.5 (152.5) tall) colorless. Rostrum (117.5 - 137.5 (125) long; 45 - 55 (46.25) wide). Chelicerae (290 - 340 (314) long) with curved fangs (62.5 - 70 (68) long). Subcapitular proportions: ventral length / height 2.03 - 2.25 (2.03); rostrum length / width 2.45 - 2.89 (2.70). Pedipalps with short tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (42.5 - 50 (50) long); femur (112.5 - 132.5 (125) long); genu (66.25 - 75 (68.75) long); tibia (100 - 107.5 (105) long; 25 - 27.5 (26.25) wide); tarsus (23.75 - 27.5 (27.5) long). Palpomere proportions: femur / genu 1.67 - 1.83 (1.82); tibia / femur 0.81 - 0.91 (0.84); tibia length / width 3.91 - 4.10 (4.00). Venter - (730 - 880 (870) long; 513 - 610 (543) wide) colorless. Gnathosomal bay (145 - 186.25 (181.25) long; 92.5 - 105 (105) wide). Cxgl- 4 apical. Medial suture (20 - 25 (20) long). Genital plates (177.5 - 200 (200) long; 157.5 - 180 (167.5) wide). Additional measurements: Cx- 1 (280 - 350 (323) long (total); 125 - 160 (140) long (medial )); Cx- 3 (365 - 405 (365) wide); anterior venter (167.5 - 186.25 (181.25) long). Ventral proportions: gnathosomal bay length / width 1.57 - 1.84 (1.73); anterior venter / genital field length 0.91 - 0.97 (0.91); anterior venter length / genital field width 1.03 - 1.08 (1.08); anterior venter / medial suture 7.30 - 9.25 (9.06). Male (Figure 236) (n = 5) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (580 - 650 (650) long; 415 - 470 (470) wide) ovoid with faint bluish-purple coloration restricted posteriorly or colorless. Anterio-medial platelets (125 - 137.5 (137.5) long; 40 - 60 (60) wide). Anterio-lateral platelets (180 - 197.5 (197.5) long; 62.5 - 82.5 (82.5) wide) free from dorsal plate. Dgl- 4 closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 280 - 320 (320 )). Dorsal plate proportions: dorsum length / width 1.36 - 1.44 (1.38); dorsal width / distance between Dgl- 4 1.46 - 1.49 (1.47); anterio-medial platelet length / width 2.29 - 3.13 (2.29); anterio-lateral platelet length / width 2.39 - 2.96 (2.39); anterio-lateral / anterio-medial length 1.37 - 1.51 (1.44). Gnathosoma - Subcapitulum (250 - 300 (300) long (ventral); 195 - 216 (215.75) long (dorsal); 105 - 127.5 (127.5) tall) colorless. Rostrum (100 - 119 (118.75) long; 37.5 - 50 (42.5) wide). Chelicerae (270 - 303 (303) long) with curved fangs (46 - 50 (47) long). Subcapitular proportions: ventral length / height 2.31 - 2.40 (2.35); rostrum length / width 2.30 - 2.91 (2.79). Pedipalps with short tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (45 - 47.5 (47.5) long); femur (102.5 - 115 (110) long); genu (57.5 - 67.5 (67.5) long); tibia (88 - 104 (103.75) long; 22.5 - 27 (26.25) wide); tarsus (22.5 - 27.5 (22.5) long). Palpomere proportions: femur / genu 1.63 - 1.80 (1.63); tibia / femur 0.83 - 0.94 (0.94); tibia length / width 3.80 - 3.95 (3.95). Venter - (670 - 771 (770) long; 460 - 520 (496) wide) colorless. Gnathosomal bay (127.5 - 170 (170) long; 80 - 90 (90) wide). Cxgl- 4 apical. Medial suture (85 - 105 (100) long). Genital plates (177.5 - 202 (201.25) long; 135 - 147.5 (147.5) wide). Additional measurements: Cx- 1 (270 - 315 (305) long (total); 140 - 157.5 (157.5) long (medial )); Cx- 3 (325 - 370 (360) wide); anterior venter (237.5 - 270 (270) long). Ventral proportions: gnathosomal bay length / width 1.46 - 1.94 (1.89); anterior venter / genital field length 1.28 - 1.41 (1.34); anterior venter length / genital field width 1.70 - 1.93 (1.83); anterior venter / medial suture 2.43 - 2.79 (2.70). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
B3129E05902983A06FF481551A47BDD1.taxon	etymology	Etymology. Specific epithet (sharkeyi) named in honor of braconid systematist, Michael Sharkey, who advised JRF during his master's degree and solidified his life-long passion for taxonomy.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
B3129E05902983A06FF481551A47BDD1.taxon	distribution	Distribution. Southwestern, New Mexico and Arizona (Figure 234).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
918A2FF5C6F8B6EEBDF48A0673969472.taxon	materials_examined	Material examined. HOLOTYPE (♀): USA, Tennessee, Sevier County, Great Smokey Mountains National Park, middle prong Little Pigeon River (35 ° 44 ' 12 " N, 83 ° 24 ' 51 " W), 12 Sep 2010, by IM Smith, IMS 100132. PARATYPES (5 ♀; 5 ♂): Pennsylvania, USA: 1 ♂ (ALLOTYPE) from Somerset County, Laurel Hill State Park, Laurel Hill Creek (40 ° 1 ' 6 " N, 79 ° 14 ' 4 " W), 8 Aug 2014, by MJ Skvarla, MS 14 - 0808 - 001, DNA 2845 * Tennessee, USA: 1 ♀ from Blount County, Great Smokey Mountains National Park, Abrams River (35 ° 35 ' 31 " N, 83 ° 51 ' 21 " W), 17 Sep 2010, by IM Smith, IMS 100141 * 2 ♀ from Monroe County, Turkey Creek (35 ° 21 ' 47 " N, 84 ° 9 ' 47 " W), 12 Sep 2009, by IM Smith, IMS 090112 * 1 ♂ from Sevier County, Great Smokey Mountains National Park, middle prong Little Pigeon River (35 ° 43 ' 33 " N, 83 ° 24 ' 1 " W), 12 Sep 2010, by IM Smith, IMS 100131 * 1 ♂ from Sevier County, Great Smokey Mountains National Park, middle prong Little Pigeon River (35 ° 44 ' 12 " N, 83 ° 24 ' 51 " W), 12 Sep 2010, by IM Smith, IMS 100132 * Vermont, USA: 2 ♀ from Addison County, Lincoln, beside Forest Service Road # 54, New Haven River (44 ° 6 ' N, 72 ° 59 ' W), 6 Jul 1989, by IM Smith, IMS 890074 * Virginia, USA: 2 ♂ from Scott County, beside Route 58 / 421, 0.9 kilometers east of Route 709, North Fork of Holston River (36 ° 39 ' N, 82 ° 28 ' W), by IM Smith, IMS 900080.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
918A2FF5C6F8B6EEBDF48A0673969472.taxon	diagnosis	Diagnosis. Torrenticola shubini are similar to other members of the Rusetria " 4 - Plates " group (T. dunni, T. glomerabilis, T. kittatinniana, T. pollani, and T. rufoalba) and T. skvarlai in having anterio-lateral platelets free from the dorsal plate, dorsal coloration separated into anterior and posterior portions, and indistinct hind coxal margins. Female T. shubini can be differentiated from female T. dunni by having a thinner rostrum (length / width = 2.5 - 2.7 in T. shubini, 2.8 - 3.0 in T. dunni). Male T. dunni can be differentiated from male T. shubini by having a longer anterior venter (215 - 238 in T. shubini, 277 - 285 in T. dunni). T. shubini can be differentiated from T. pollani by having stockier tibiae (length / width ♀ = 3.30 - 3.60 in T. shubini, 3.89 - 4.18 in T. pollani; ♂ = 3.11 - 3.22 in T. shubini, 3.41 - 3.75 in T. pollani) and a stockier rostrum (length / width = 2.24 - 2.92 in T. shubini, 3.27 - 3.82 in T. pollani). T. shubini can be differentiated from T. glomerabilis by having the Dgl- 4 closer to the dorsal edge (dorsal width / length between Dgl- 4 = 1.2 - 1.4 in T. shubini; 1.5 - 1.7 in T. glomerabilis); and stockier tibiae (length / width ♀ = 3.3 - 3.6 in T. shubini, 4.1 - 4.5 in T. glomerabilis; ♂ = 3.1 - 3.2 in T. shubini, 3.5 - 4.4 in T. glomerabilis). Female T. shubini can be differentiated from female T. kittatinniana by having a stockier rostrum (length / width = 2.5 - 2.7 in T. shubini, 3.16 in T. kittatinniana) and a taller subcapitulum (140 - 145 in T. shubini, 125 in T. kittatinniana). Male T. shubini can be differentiated from male T. kittatinniana by having a shorter dorsum (400 - 465 in T. shubini, 500 in T. kittatinniana) and a shorter genital field (90 - 108 in T. shubini, 115 in T. kittatinniana). T. shubini can be differentiated from T. rufoalba by having a stockier rostrum (length / width = 2.24 - 2.92 in T. shubini, 2.96 - 3.06 in T. rufoalba). Female T. shubini can be differentiated from female T. rufoalba by having a thinner genital field (137 - 145 in T. shubini, 157.5 in T. rufoalba). Male T. shubini can be differentiated from male T. rufoalba by having a longer anterior venter (215 - 238 in T. pollani, 195 in T. rufoalba). T. shubini can be differentiated from T. skvarlai by having a conical pedipalpal femoral tubercle, whereas T. skvarlai has a broad and flat pedipalpal femoral tubercle, and by having a longer anterior venter (♀ = 156 - 170 in T. shubini, 140 - 153 in T. skvarlai; ♂ = 215 - 238 in T. shubini, 177 - 205 in T. skvarlai).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
918A2FF5C6F8B6EEBDF48A0673969472.taxon	description	Description. Female (Figure 238) (n = 5) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (550 - 640 (590) long; 415 - 455 (430) wide) ovoid with faint bluish-purple coloration separated into anterior and posterior portions, and with faint orange medially. Anterio-medial platelets (115 - 125 (122.5) long; 40 - 47.5 (46.25) wide). Anterio-lateral platelets (160 - 170 (165) long; 55 - 60 (56.25) wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 310 - 335 (325 )). Dorsal plate proportions: dorsum length / width 1.33 - 1.45 (1.37); dorsal width / distance between Dgl- 4 1.32 - 1.42 (1.32); anterio-medial platelet length / width 2.63 - 2.88 (2.65); anterio-lateral platelet length / width 2.75 - 2.96 (2.93); anterio-lateral / anterio-medial length 1.28 - 1.43 (1.35). Gnathosoma - Subcapitulum (300 - 320 (310) long (ventral); 223 - 255 (232) long (dorsal); 140 - 145 (140) tall) colorless. Rostrum (117.5 - 122.5 (120) long; 45 - 48.75 (45) wide). Chelicerae (288 - 320 (316) long) with curved fangs (54 - 63 (57) long). Subcapitular proportions: ventral length / height 2.14 - 2.21 (2.21); rostrum length / width 2.51 - 2.67 (2.67). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (42.5 - 47.5 (47.5) long); femur (112.5 - 121.25 (117.5) long); genu (63.75 - 70 (66.25) long); tibia (82.5 - 90 (83.75) long; 23.75 - 25 (25) wide); tarsus (17.5 - 17.5 (17.5) long). Palpomere proportions: femur / genu 1.71 - 1.83 (1.77); tibia / femur 0.68 - 0.75 (0.71); tibia length / width 3.30 - 3.60 (3.35). Venter - (640 - 780 (740) long; 446 - 521 (447) wide) colorless. Gnathosomal bay (127.5 - 165 (165) long; 85 - 96.25 (96.25) wide). Cxgl- 4 subapical. Medial suture (7.5 - 22.5 (7.5) long). Genital plates (156.25 - 175 (167.5) long; 137.5 - 145 wide). Additional measurements: Cx- 1 (238 - 308 (291) long (total); 98 - 138 (138) long (medial )); Cx- 3 (307 - 356 (307) wide); anterior venter (157.5 - 170 (157.5) long). Ventral proportions: gnathosomal bay length / width 1.50 - 1.94 (1.71); anterior venter / genital field length 0.94 - 1.05 (0.94); anterior venter length / genital field width 1.14 - 1.24; anterior venter / medial suture 7.50 - 21.00 (7.50). Male (Figure 239) (n = 5) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (400 - 465 (460) long; 260 - 305 (300) wide) ovoid with faint bluish-purple coloration separated into anterior and posterior portions, and with faint orange medially. Anterio-medial platelets (82.5 - 100 (92.5) long; 30 - 40 (35) wide). Anterio-lateral platelets (115 - 135 (127.5) long; 35 - 47.5 (42.5) wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 210 - 250 (250 )). Dorsal plate proportions: dorsum length / width 1.52 - 1.57 (1.53); dorsal width / distance between Dgl- 4 1.20 - 1.27 (1.20); anterio-medial platelet length / width 2.31 - 2.92 (2.64); anterio-lateral platelet length / width 2.84 - 3.29 (3.00); anterio-lateral / anterio-medial length 1.29 - 1.41 (1.38). Gnathosoma - Subcapitulum (210 - 245 (240) long (ventral); 162.5 - 180 (171) long (dorsal); 78.75 - 87.5 (85) tall) colorless. Rostrum (87.5 - 95 (92.5) long; 30 - 42.5 (32.5) wide). Chelicerae (210 - 234 (229) long) with curved fangs (38 - 42.5 (40) long). Subcapitular proportions: ventral length / height 2.67 - 2.82 (2.82); rostrum length / width 2.24 - 2.92 (2.85). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (32.5 - 37.5 (37.5) long); femur (82.5 - 91.25 (87.5) long); genu (50 - 55 (52.5) long); tibia (62.5 - 72.5 (70) long; 20 - 22.5 (22.5) wide); tarsus (15 - 16.25 (15) long). Palpomere proportions: femur / genu 1.62 - 1.75 (1.67); tibia / femur 0.76 - 0.83 (0.80); tibia length / width 3.11 - 3.22 (3.11). Venter - (490 - 571 (571) long; 310 - 390 (342) wide) colorless. Gnathosomal bay (92.5 - 115 (115) long; 55 - 65 (65) wide). Cxgl- 4 subapical. Medial suture (95 - 105 (95) long). Genital plates (90 - 107.5 (107.5) long; 75 - 80 (80) wide). Additional measurements: Cx- 1 (200 - 235 (235) long (total); 96 - 121 (98) long (medial )); Cx- 3 (235 - 293 (260) wide); anterior venter (215 - 237.5 (232.5) long). Ventral proportions: gnathosomal bay length / width 1.54 - 1.82 (1.77); anterior venter / genital field length 2.16 - 2.39 (2.16); anterior venter length / genital field width 2.84 - 3.07 (2.91); anterior venter / medial suture 2.05 - 2.45 (2.45). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
918A2FF5C6F8B6EEBDF48A0673969472.taxon	etymology	Etymology. Specific epithet (shubini) named in honor of author and palaeontologist Neil Shubin for his efforts to popularize stories of human evolution with his book (2009) and TV series (2014), Your Inner Fish. As with many of the species that Shubin studies (e. g., Tiktaalik roseae Daeschler, Shubin & Jenkins, 2006), Torrenticola shubini may represent a key evolutionary transition in the Rusetria Complex, between species that do and do not have their lateral platelets fused to the dorsal shield.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
918A2FF5C6F8B6EEBDF48A0673969472.taxon	distribution	Distribution. Appalachians (Figure 237).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
93BB2111CE6F022ED76A3F804C344D57.taxon	materials_examined	Material examined. LECTOTYPE (1 ♀): USA, California, Santa Cruz County, Waddell Creek, 25 - 26 Aug 1932, by PR Needham, RM 320007 PARALECTOTYPE (1 ♂): California, Santa Cruz County, Waddell Creek, 1 Dec 1932, by PR Needham, RM 320008. TOPOTYPES (1 ♀; 1 ♂): from USA, California, Santa Cruz County, Waddell Creek, 30 - 31 Aug 1933, by PR Needham, RM 330016. OTHER MATERIAL (54 ♀; 34 ♂): British Columbia, Canada: 3 ♀ and 1 ♂ from Ryan Rest Area off Highway 3, east of Yahk Moyie River, 15 Aug 2012, by IM Smith, IMS 120071 * 1 ♀ from Vancouver Island, Lake Cowichan, Cowichan River, above Skutz Falls, 9 Jul 1979, by IM Smith, IMS 790035 * California, USA: 1 ♀ from Del Norte County, Six Rivers National Forest, Middle Fork Smith River (41 ° 51 ' 20 " N, 123 ° 53 ' 10 " W), 15 Aug 2013, by JR Fisher, JRF 13 - 0815 - 002 * 1 ♀ from El Dorado County, Upper Truckee River (38 ° 50 ' 56 " N, 120 ° 1 ' 39 " W), 29 Aug 2013, by JR Fisher, JRF 13 - 0829 - 003 * 1 ♀ and 1 ♂ from El Dorado County, Upper Truckee River (38 ° 50 ' 56 " N, 120 ° 1 ' 39 " W), 29 Aug 2013, by JR Fisher, JRF 13 - 0829 - 004 * 2 ♀ and 2 ♂ from Mariposa County, El Portal, Indian Flat campground, Merced River, 9 - 10 Jun 1976, by IM Smith, IMS 760087 * 2 ♀ from Mariposa County, Yosemite Valley, East Fork of Merced River, 22 Aug 1933, by PR Needham, RM 330012 * 1 ♀ from Mendocino County, Jackson Demonstration State Park, North Fork of Big River (39 ° 20 ' 46 " N, 123 ° 30 ' 35 " W), 22 Aug 2013, by JR Fisher, JRF 13 - 0822 - 002 * 1 ♀ from Mendocino County, Paul M. Dimmick Recreation Area, North Fork of Navarro River, beside Route 128, 4 Aug 1987, by IM Smith, IMS 870127 A * 1 ♂ from Mendocino County, Rancheria Creek, beside Route 128, 7.3 kilometers south of Boonville, 4 Aug 1987, by IM Smith, IMS 870126 A * 1 ♀ and 1 ♂ from Monterey County, Big Sur River, beside Route 1 near Pfeiffer-Big Sur State Park, 28 - 29 Jul 1987, by IM Smith, IMS 870116 A * 3 ♀ from Monterey County, Big Sur, Pfeiffer-Big Sure State Park, Big Sur Creek, 7 Jun 1976, by IM Smith, IMS 760074 * 1 ♀ from Monterey County, Los Padres National Forest, Salmon Creek (35 ° 48 ' 57 " N, 121 ° 21 ' 29 " W), 6 Sep 2013, by JR Fisher, JRF 13 - 0906 - 003 * 1 ♀ from Monterey County, Pfeiffer State Park, Big Sur River (36 ° 14 ' 42 " N, 121 ° 46 ' 43 " W), 4 Sep 2013, by JR Fisher, JRF 13 - 0904 - 004 * 1 ♀ and 1 ♂ from Monterey County, Salmon Creek, beside Route 1, south of Gorda, 28 Jul 1987, by IM Smith, IMS 870114 A * 1 ♀ from Nevada County, beside Route 89, north of Hobart Mills, 13 Jun 1976, by IM Smith, IMS 760109 * 1 ♀ from Nevada County, Tahoe National Forest, Sagehen Creek (39 ° 26 ' 2 " N, 120 ° 12 ' 17 " W), 26 Aug 2013, by JR Fisher, JRF 13 - 0826 - 006 * 2 ♀ from Trinity County, Shasta-Trinity National Forest, Wilson Creek (40 ° 25 ' 17 " N, 123 ° 3 ' 5 " W), 20 Aug 2013, by JR Fisher, JRF 13 - 0820 - 003 * 1 ♂ from Shasta County, Battle Creek, beside Route 44, 5.6 kilometers west of Viola, 10 Aug 1987, by IM Smith, IMS 870139 A * 1 ♀ from Trinity County, South Fork of Trinity River, beside Route 36 at Forest Glen campground, 6 Aug 1987, by IM Smith, IMS 870131 * 1 ♀ and 1 ♂ from Tulare County, Brush Creek, Brush Creek Flat, beside SM 99 between Roads End Station & Johnsondale, 31 Jul 1987, by IM Smith, IMS 870122 * 1 ♀ and 3 ♂ from Tulare County, Kern River, Brush Creek Flat, beside SM 99 between Roads End Station & Johnsondale, 31 Jul 1987, by IM Smith, IMS 870121 * 1 ♂ from Tulare County, Stony Creek, Stony Creek Picnic Area, east of Sequoia National Park, 1 Aug 1987, by IM Smith, IMS 870124 A * Idaho, USA: 1 ♀ and 1 ♂ from Custer County, Challis National Forest, Stanley Creek (44 ° 15 ' 12 " N, 115 ° 0 ' 19 " W), 30 Jul 2012, by JR Fisher, WA Nelson, & JC O'Neill, ROW 12 - 0730 - 005 * 1 ♀ and 1 ♂ from Custer County, Salmon River at picnic area, beside Route 93, north of Morgan Creek, 2 Jul 1985, by IM Smith, IMS 850064 * 1 ♀ from Fremont County, Targhee National Forest, Rock Creek (44 ° 6 ' 44 " N, 111 ° 15 ' 4 " W), by JR Fisher, WA Nelson, & JC O'Neill, ROW 12 - 0725 - 001 * Montana, USA: 1 ♀ from Ravalli County, Bitterroot National Forest, West Fork Bitterroot River (45 ° 54 ' 38 " N, 114 ° 9 ' 43 " W), 6 Aug 2012, by JR Fisher, WA Nelson, & JC O'Neill, ROW 12 - 0806 - 003 * 1 ♀ from Ravalli County, Medicine Springs, Spring Gulch campground, East Fork of Bitterroot River, beside Route 93, 1 Jul 1985, by IM Smith, IMS 850060 * Nevada, USA: 1 ♀ and 1 ♂ from Humboldt County, Paradise Valley, Cabin Creek beside road, 10.7 kilometers north of Hinkey Summit, 11 Aug 1987, by IM Smith, IMS 870141 A * 1 ♀ and 2 ♂ from Humboldt County, Paradise Valley, Dutch John Creek beside road, 8.3 kilometers north of Hinkey Summit, 11 Aug 1987, by IM Smith, IMS 870142 A * Oregon, USA: 1 ♀ and 2 ♂ from Coos County, Siskiyou National Forest, Road 33 between Powers & Agness, 2 Jul 1983, by IM Smith, IMS 830015 * 2 ♀ from Curry County, Port Orford, Butler Bar campground, Elk River, 25 Jun 1976, by IM Smith, IMS 760162 * 1 ♀ and 1 ♂ from Curry County, Port Orford, Humbug Mountain State Park Picnic Area, beside Route 1, Brush Creek, 1 Jul 1983, by IM Smith, IMS 830012 * 1 ♀ and 2 ♂ from Curry County, Port Orford, Humbug Mountain State Park Picnic Area, Brush Creek, beside Route 1, 3 Jul 1983, by IM Smith, IMS 830020 B * 1 ♀ from Curry County, Quosatana Creek (42 ° 29 ' 21 " N, 124 ° 14 ' 2 " W), 14 Aug 2013, by JR Fisher, JRF 13 - 0814 - 003 * 1 ♀ from Curry County, Rogue River National Forest, Elk River (42 ° 42 ' 46 " N, 124 ° 18 ' 41 " W), 13 Aug 2013, by JR Fisher, JRF 13 - 0813 - 003 * 3 ♀ from Lane County, Gate Creek (44 ° 8 ' 48 " N, 122 ° 34 ' 20 " W), 11 Aug 2013, by JC O'Neill, & WA Nelson, JNOW 13 - 0811 - 001 * 1 ♀ and 1 ♂ from Curry County, Sixes, Sixes River, beside road at mouth of Edson Creek, 4 Jul 1983, by IM Smith, IMS 830021 B * 1 ♀ and 1 ♂ from Lincoln County, Blackberry campground near Tidewater, Alsea River, 28 Jun 1983, by IM Smith, IMS 830009 * 2 ♀ and 2 ♂ from Lincoln County, Blackberry campground near Tidewater, Alsea River, 29 Jun 1983, by IM Smith, IMS 830010 * 2 ♀ and 2 ♂ from Lincoln County, Siuslaw National Forest, Five Rivers Creek (44 ° 19 ' 53 " N, 123 ° 50 ' 59 " W), 8 Aug 2013, by JC O'Neill, & WA Nelson, JNOW 13 - 0808 - 001 * Washington, USA: 2 ♀ and 2 ♂ from Cowlitz County (46 ° 22 ' 24 " N, 122 ° 34 ' 45 " W), 16 Jul 2013, by JC O'Neill, & WA Nelson, JNOW 13 - 0716 - 001 * 1 ♀ and 1 ♂ from Grays Harbor County, Stewarts Creek (47 ° 15 ' 49 " N, 123 ° 55 ' 12 " W), 25 Jul 2013, by JC O'Neill, & WA Nelson, JNOW 13 - 0725 - 001 * Wyoming, USA: 1 ♀ and 2 ♂ from Sublette County, Boulder Creek, beside Route 191, just north of Boulder, 15 Aug 1987, by IM Smith, IMS 870151 A & IMS 870151 B.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
93BB2111CE6F022ED76A3F804C344D57.taxon	diagnosis	Diagnosis. Torrenticola sierrensis are similar to other members of the Tricolor Complex (T. bittikoferae, T. hoosieri, T. larvata, T. pearsoni, T. olliei, T. tricolor, T. trimaculata, T. unimaculata, T. cardia, T. kringi, T. dimorpha, and T. mohawk) in having a short, conical rostrum. T. sierrensis can be differentiated from most Tricolor Complex by being distributed in the west and having a wider genital field (♀ = 180 - 213 in T. sierrensis, 145 - 180 in others; ♂ = 135 - 175 in T. sierrensis, 92 - 120 in others), except for T. olliei (♀ = 190 - 203, ♂ = 130 - 138). T. sierrensis can be differentiated from T. olliei by having a more elongate rostrum (length / width = 1.95 - 2.14 in T. sierrensis, 1.56 - 1.81 in T. olliei).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
93BB2111CE6F022ED76A3F804C344D57.taxon	description	Re-description. Female (Figure 241) (n = 9) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (700 - 880 (865) long; 550 - 740 (710) wide) circular with orange or reddish purple coloration separated into anterior and posterior portions with faint orange medially (occasionally colorless). Anterio-medial platelets (162.5 - 197.5 (187.5) long; 65 - 82.5 (81.25) wide). Anterio-lateral platelets (192.5 - 250 (225) long; 82.5 - 100 (92.5) wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 435 - 520 (500 )). Dorsal plate proportions: dorsum length / width 1.19 - 1.27 (1.22); dorsal width / distance between Dgl- 4 1.22 - 1.43 (1.42); anterio-medial platelet length / width 2.10 - 2.50 (2.31); anterio-lateral platelet length / width 2.33 - 2.50 (2.43); anterio-lateral / anterio-medial length 1.18 - 1.27 (1.20). Gnathosoma - Subcapitulum 320 - 365 (365) long (ventral); 227 - 265 (260) long (dorsal); 140 - 165 (152.5) tall) colorless. Rostrum (120 - 140 (140) long; 60 - 70 (70) wide) short and conical. Chelicerae (320 - 355 long) with curved fangs (61 - 77.5 long). Subcapitular proportions: ventral length / height 2.19 - 2.39 (2.39); rostrum length / width 1.96 - 2.08 (2.00). Pedipalps with tuberculate ventral extensions with dentate tips on femora and genua. Palpomeres: trochanter (45 - 55 (55) long); femur (115 - 135 (135) long); genu (80 - 92.5 (92.5) long); tibia (107.5 - 117.5 (117.5) long; 32.5 - 35 (35) wide); tarsus (21.25 - 35 32.5) long). Palpomere proportions: femur / genu 1.35 - 1.55 (1.46); tibia / femur 0.86 - 0.95 (0.87); tibia length / width 3.07 - 3.48 (3.36). Venter - (775 - 1040 (1015) long; 619 - 820 (770) wide) colorless. Gnathosomal bay (155 - 205 (195) long; 95 - 112.5 (112.5) wide). Cxgl- 4 subapical. Medial suture (17.5 - 37.5 (22.5) long). Genital plates (210 - 235 (222.5) long; 180 - 212.5 (212.5) wide). Additional measurements: Cx- 1 (192.5 - 370 (370) long (total); 136 - 185 (172.5) long (medial )); Cx- 3 (250 - 475 (475) wide); anterior venter (192.5 - 237.5 (212.5) long). Ventral proportions: gnathosomal bay length / width 1.50 - 1.91 (1.73); anterior venter / genital field length 0.90 - 1.06 (0.96); anterior venter length / genital field width 1.00 - 1.13 (1.00); anterior venter / medial suture 6.33 - 12.14 (9.44). Male (Figure 242) (n = 7) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (590 - 735 (715) long; 460 - 590 (580) wide) circular with orange or reddish purple coloration separated into anterior and posterior portions with faint orange medially (occasionally colorless). Anterio-medial platelets (137.5 - 172.5 (172.5) long; 60 - 75 (75) wide). Anterio-lateral platelets (185 - 222.5 (215) long; 72.5 - 95 (92.5) wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 355 - 460 (460 )). Dorsal plate proportions: dorsum length / width 1.17 - 1.28 (1.23); dorsal width / distance between Dgl- 4 1.26 - 1.33 (1.26); anterio-medial platelet length / width 2.24 - 2.48 (2.30); anterio-lateral platelet length / width 2.12 - 2.55 (2.32); anterio-lateral / anterio-medial length 1.22 - 1.38 (1.25). Gnathosoma - Subcapitulum (270 - 330 (325) long (ventral); 200 - 235 (235) long (dorsal); 118.75 - 140 (135) tall) colorless. Rostrum (100 - 122.5 (122.5) long; 50 - 60 (60) wide) short and conical. Chelicerae (262 - 317.5 (315) long) with curved fangs (50 - 67.5 (60) long). Subcapitular proportions: ventral length / height 2.23 - 2.46 (2.41); rostrum length / width 1.95 - 2.14 (2.04). Pedipalps with tuberculate ventral extensions with dentate tips on femora and genua. Palpomeres: trochanter (40 - 50 (50) long); femur (92.5 - 120 (120) long); genu (65 - 80 (78.75) long); tibia (82.5 - 107.5 (102.5) long; 27.5 - 32.5 (32.5) wide); tarsus (25 - 32.5 (32.5) long). Palpomere proportions: femur / genu 1.42 - 1.52 (1.52); tibia / femur 0.85 - 0.95 (0.85); tibia length / width 3.00 - 3.31 (3.15). Venter - (690 - 895 (875) long; 538 - 667 (665) wide) colorless. Gnathosomal bay (112.5 - 180 (175) long; 80 - 100 (92.5) wide). Cxgl- 4 subapical. Medial suture (72.5 - 92.5 (90) long). Genital plates (172.5 - 220 (217.5) long; 135 - 175 (175) wide). Additional measurements: Cx- 1 (269 - 360 (360) long (total); 137 - 182.5 (182.5) long (medial )); Cx- 3 (364 - 455 (455) wide); anterior venter (255 - 295 (290) long). Ventral proportions: gnathosomal bay length / width 1.41 - 1.89 (1.89); anterior venter / genital field length 1.32 - 1.51 (1.33); anterior venter length / genital field width 1.66 - 1.89 (1.66); anterior venter / medial suture 3.03 - 3.59 (3.22). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
93BB2111CE6F022ED76A3F804C344D57.taxon	etymology	Etymology. Although Marshall (1943) did not specify an etymology for the specific epithet (sierrensis), surely it refers to the type locality of this species within the Sierra Nevada in California.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
93BB2111CE6F022ED76A3F804C344D57.taxon	distribution	Distribution. Western (Figure 240), but likely not found in the southwest. T. sierrensis was previously known only from a few localities in California; we extend the range in western North America, but not in the southwest.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
5EF74D187F35EDFCAA74AC2477D0B833.taxon	materials_examined	Material examined. HOLOTYPE (♀): from USA, Pennsylvania, Somerset County, Laurel Hill State Park, Laurel Hill Creek (40 ° 1 ' 6 " N, 79 ° 14 ' 4 " W), 8 Aug 2014, by MJ Skvarla, MS 14 - 0808 - 001. PARATYPES (17 ♀; 24 ♂): New Hampshire, USA: 1 ♀ and 1 ♂ from Coos County, Randolph, Moose River, beside Route 2, 30 Jul 1981, by IM Smith, IMS 810105 * 2 ♀ and 2 ♂ from Coos County, White Mountain National Forest, Peabody River, Dolly Copp Campground, south of Gorham, 27 Jul 1981, IMS 810099 A & IMS 810099 B * New York, USA: 1 ♀ and 1 ♂ from Essex County, Minerva, Boreas River, beside Route 28 N, 13.8 kilometers northwest of Morse Memorial Parkway, 21 Jun 1990, by IM Smith, IMS 900050 A * 2 ♀ and 1 ♂ from Greene County, Schoharie Creek, beside Route 23 A, 9.6 kilometers west of Route 296 in Hunter, 22 Jun 1990, by IM Smith, IMS 900052 * North Carolina, USA: 1 ♀ and 1 ♂ from Macon County, Cullasaja River, Highlands beside Route 64 / 28, 4.1 kilometers northwest of Cliffside Lake Road, 30 Jun 1990, by IM Smith, IMS 900070 A * 2 ♂ from Macon County, Rainbow Springs, Nantahala River, beside Forest Route 67, south of Standing Indian Campground, 1 Jul 1990, IMS 900072 * 2 ♀ and 2 ♂ from Yancey County, South Toe River, Lost Cove Picnic Area on Forest Route 472, 2.8 kilometers south of Route 80, 28 Jun 1990, by IM Smith, IMS 900065 A * Pennsylvania, USA: 1 ♂ from Bedford County, Chaneysville, Sweet Root Picnic Area beside Route 326, 18 Jul 1990, by IM Smith, IMS 900105 * 1 ♀ and 3 ♂ from Fayette County, Dunbar Creek (39 ° 57 ' 50 " N, 79 ° 35 ' 8.70 " W), 10 Aug 2014, by MJ Skvarla, MS 14 - 0810 - 001 * 2 ♀ and 1 ♂ from Huntingdon County, Alan Seeger Natural Area, Stone Creek, beside road from McAlevys Fort to Route 322, 19 Jul 1990, by IM Smith, IMS 900107 * 1 ♂ (ALLOTYPE) from Somerset County, Laurel Hill State Park, Laurel Hill Creek (40 ° 1 ' 6 " N, 79 ° 14 ' 4 " W), 8 Aug 2014, by MJ Skvarla, MS 14 - 0808 - 001 * 2 ♀ from Somerset County, Laurel Hill State Park, Laurel Hill Creek (40 ° 1 ' 6 " N, 79 ° 14 ' 4 " W), 8 Aug 2014, by MJ Skvarla, MS 14 - 0808 - 001 * Tennessee, USA: 1 ♀ and 1 ♂ from Monroe County, Tellico River, beside Forest Route 210, 1.8 kilometers east of bridge at Bald River Falls, 5 Jul 1990, by IM Smith, IMS 900079 * 1 ♂ from Monroe County, Turkey Creek, beside Forest Route 35, 2.0 kilometers northeast of road from Route 165 to Miller Chapel Church, 5 Jul 1990, by IM Smith, IMS 900078 * Virginia, USA: 1 ♀ and 4 ♂ from Washington County, Damascus, Laurel River, beside Route 58 near boundary of Mount Rogers National Recreation Area, 10 Jul 1990, by IM Smith, IMS 900085 A * West Virginia, USA: 1 ♂ from Pendleton County, Spruce Knob, beside Forest Route 112, 10.2 kilometers west of Route 33, 17 Jul 1990, by IM Smith, IMS 900103 * 1 ♀ and 1 ♂ from Randolph County, Laurel Fork of Cheat River, Laurel Fork Campground off Forest Route 14, south of Wymer, 17 Jul 1990, by IM Smith, IMS 900102.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
5EF74D187F35EDFCAA74AC2477D0B833.taxon	diagnosis	Diagnosis. Torrenticola skvarlai are similar to species with similar dorsal patterning, such as the Rusetria " 4 - Plate " group (T. dunni, T. glomerabilis, T. kittatinniana, T. pollani, T. rufoalba, and T. shubini), Elongata Group (T. elongata, T. gorti, and T. reduncarostra), Neoanomala Group (T. interiorensis and T. neoanomala), T. bondi, T. racupalpa, T. irapalpa, T. erectirostra, and T. arktonyx. They can be differentiated from all of these except Rusetria 4 - Plates by having indistinct hind coxal margins. T. skvarlai can be differentiated from all Rusetria 4 - Plates by having broadly tuberculate, dentate pedipalp femoral extensions (all Rusetria Complex have conical tuberculate pedipalp femoral extensions, usually without dentation) and by having a shorter anterior venter (♀ = 140 - 152.5 in T. skvarlai, 155 - 213 in Rusetria 4 - Plates; ♂ = 177 - 205 in T. skvarlai, 215 - 285 in Rusetria 4 - Plates).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
5EF74D187F35EDFCAA74AC2477D0B833.taxon	description	Description. Female (Figure 244) (n = 5) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (495 - 560 (495) long; 370 - 420 (370) wide) ovoid with reddish-purple or bluish-purple coloration separated into anterior and posterior portions, occasionally with orange medially. Anterio-medial platelets (107.5 - 117.5 (107.5) long; 42.5 - 47.5 (42.5) wide). Anterio-lateral platelets (155 - 167.5 (155) long; 52.5 - 62.5 (52.5) wide) free from dorsal plate. Dgl- 4 closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 245 - 285 (255 )). Dorsal plate proportions: dorsum length / width 1.33 - 1.39 (1.34); dorsal width / distance between Dgl- 4 1.45 - 1.58 (1.45); anterio-medial platelet length / width 2.37 - 2.53 (2.53); anterio-lateral platelet length / width 2.68 - 2.95 (2.95); anterio-lateral / anterio-medial length 1.36 - 1.44 (1.44). Gnathosoma - Subcapitulum (290 - 315 (290) long (ventral); 218 - 240 (219) long (dorsal); 120 - 135 (120) tall) colorless. Rostrum (115 - 130 (115) long; 40 - 47.5 (40) wide). Chelicerae (286.75 - 321 (299) long) with curved fangs (49 - 65 (54) long). Subcapitular proportions: ventral length / height 2.33 - 2.48 (2.42); rostrum length / width 2.74 - 2.94 (2.88). Pedipalps with broadly tuberculate, dentate ventral extensions on femora and tuberculate ventral extensions on genua. Palpomeres: trochanter (37.5 - 45 (40) long); femur (105 - 119 (105) long); genu (62.5 - 67.5 (62.5) long); tibia (82.5 - 90 (82.5) long; 22.5 - 25 (22.5) wide); tarsus (17.5 - 20 (17.5) long). Palpomere proportions: femur / genu 1.67 - 1.76 (1.68); tibia / femur 0.76 - 0.79 (0.79); tibia length / width 3.50 - 3.78 (3.67). Venter - (600 - 690 (600) long; 435 - 480 (435) wide) with faint reddish-purple or bluish-purple coloration. Gnathosomal bay (125 - 165 (135) long; 85 - 105 (85) wide). Cxgl- 4 subapical. Medial suture (12.5 - 15 (12.5) long). Genital plates (165 - 172.5 (166.25) long; 150 - 152.5 (150) wide). Additional measurements: Cx- 1 (243 - 285 (257) long (total); 109 - 120 (109) long (medial )); Cx- 3 (295 - 345 (300) wide); anterior venter (140 - 152.5 (145) long). Ventral proportions: gnathosomal bay length / width 1.47 - 1.71 (1.59); anterior venter / genital field length 0.84 - 0.92 (0.87); anterior venter length / genital field width 0.93 - 1.02 (0.97); anterior venter / medial suture 10.17 - 12.00 (11.60). Male (Figure 245) (n = 5) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (425 - 510 (450) long; 310 - 360 (325) wide) ovoid with reddish-purple or bluish-purple coloration separated into anterior and posterior portions, occasionally with orange medially. Anterio-medial platelets (95 - 102.5 (100) long; 38.75 - 47.5 (40) wide). Anterio-lateral platelets (142.5 - 177.5 (147.5) long; 45 - 57.5 (47.5) wide) free from dorsal plate. Dgl- 4 closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 220 - 240 (240 )). Dorsal plate proportions: dorsum length / width 1.32 - 1.42 (1.38); dorsal width / distance between Dgl- 4 1.35 - 1.50 (1.35); anterio-medial platelet length / width 2.16 - 2.50 (2.50); anterio-lateral platelet length / width 3.09 - 3.28 (3.11); anterio-lateral / anterio-medial length 1.48 - 1.73 (1.48). Gnathosoma - Subcapitulum (245 - 290 (255) long (ventral); 177.5 - 220 (193) long (dorsal); 92.5 - 105 (97.5) tall) colorless. Rostrum (97.5 - 125 (102.5) long; 32.5 - 45 (35) wide). Chelicerae (235 - 280 (245) long) with curved fangs (47.5 - 55 (54) long). Subcapitular proportions: ventral length / height 2.62 - 2.76 (2.62); rostrum length / width 2.78 - 3.00 (2.93). Pedipalps with broadly tuberculate, dentate ventral extensions on femora and tuberculate ventral extensions on genua. Palpomeres: trochanter (30 - 40 (36.25) long); femur (87.5 - 105 (87.5) long); genu (50 - 60 (52.5) long); tibia (75 - 82.5 (78.75) long; 20 - 22.5 (21.25) wide); tarsus (15 - 18.75 (17.5) long). Palpomere proportions: femur / genu 1.67 - 1.80 (1.67); tibia / femur 0.79 - 0.90 (0.90); tibia length / width 3.65 - 3.88 (3.71). Venter - (530 - 620 (550) long; 358 - 415 (359) wide) with faint reddish-purple or bluish-purple coloration. Gnathosomal bay (130 - 145 (130) long; 62.5 - 75 (73.75) wide). Cxgl- 4 subapical. Medial suture (55 - 75 (75) long). Genital plates (120 - 135 (120) long; 100 - 110 (100) wide). Additional measurements: Cx- 1 (225 - 280 (242) long (total); 100 - 135 (102) long (medial )); Cx- 3 (270 - 320 (272) wide); anterior venter (177.5 - 205 (192.5) long). Ventral proportions: gnathosomal bay length / width 1.76 - 2.16 (1.76); anterior venter / genital field length 1.48 - 1.63 (1.60); anterior venter length / genital field width 1.78 - 1.95 (1.93); anterior venter / medial suture 2.57 - 3.23 (2.57). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
5EF74D187F35EDFCAA74AC2477D0B833.taxon	etymology	Etymology. Specific epithet (skvarlai) named in honor of Michael Skvarla, who collected specimens of the species and is a dear friend and colleague of JRF.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
5EF74D187F35EDFCAA74AC2477D0B833.taxon	distribution	Distribution. Appalachians (Figure 243).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
02900DCF4969DD929E148EA2E89B4BF7.taxon	materials_examined	Material examined. HOLOTYPE (♀): from USA, Arkansas, Polk County, East Saline Creek, beside Forest Road 38, north of Shady Lake Recreation Area, 30 Jul 2011, by IM Smith, IMS 110041, DNA 1300. PARATYPES (4 ♀; 6 ♂): Arkansas, USA: 2 ♂ from Montgomery County, Gaston, South Fork Ouachita River, 29 Jul 2011, by IM Smith, IMS 110040 * 1 ♂ (ALLOTYPE) from Montgomery County, Ouachita National Forest, South Fork Ouachita River, 29 Jul 2011, by AJ Radwell, & B Crump, AJR 110302 * 4 ♀ and 3 ♂ from Polk County, East Saline Creek, beside Forest Road 38, north of Shady Lake Recreation Area, 30 Jul 2011, by IM Smith, IMS 110041.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
02900DCF4969DD929E148EA2E89B4BF7.taxon	diagnosis	Diagnosis. Torrenticola solisorta are similar to other members of the Nigroalba Group (T. flangipalpa, T. nigroalba, and T. dentirostra) in being small, slightly elongate, having purple dorsal coloration restricted posteriorly, and having distinct yet poorly-defined hind coxal margins. T. solisorta can be differentiated from T. flangipalpa by having tuberculate pedipalp femoral extension (flange-like and anteriorly-directed in T. flangipalpa); a shorter anterior venter (192 - 225 in T. solisorta, 235 - 265 in T. flangipalpa); and more elongate pedipalp tibia (length / width ♀ = 5.67 - 5.82 in T. solisorta, 4.79 - 5.00 in T. flangipalpa; ♂ = 5.08 - 5.17 in T. solisorta, 4.40 - 4.86 in T. flangipalpa). T. solisorta can be differentiated from T. dentirostra by having a smooth rostrum (T. dentirostra has a dentate bump midway on the dorsal edge of the rostrum) and more elongate pedipalpal tibiae (length / width ♀ = 5.6 - 5.9 in T. solisorta, 4.5 - 5.0 in T. dentirostra; ♂ = 5.0 - 5.2 in T. solisorta, 4.5 - 4.7 in T. dentirostra). T. solisorta can be differentiated from T. nigroalba by having orangish coloration immediately anterior to the purple dorsal coloration. Additionally, female T. solisorta can be differentiated from female T. nigroalba by having a slightly shorter dorsum (475 - 500 in T. solisorta, 500 - 530 in T. nigroalba); a thicker gnathosomal bay (length / width = 1.3 - 1.5 in T. solisorta, 1.25 - 1.55 in T. nigroalba); and a slightly thinner subcapitulum (3.14 - 3.30 in T. solisorta, 3.00 - 3.14 in T. nigroalba). Male T. solisorta also can be differentiated from T. nigroalba by anterior venter / medial suture (2.87 - 3.26 in T. solisorta, 2.54 - 2.77 in T. nigroalba) and having a wider dorsum (305 - 320 in T. solisorta, 290 - 300 in T. nigroalba). Other Torrenticola with purple dorsal coloration restricted posteriorly can be confused with T. solisorta, such as T. tahoei and T. oregonensis. Both of these species are larger (dorsum length ♀ = 475 - 500 in T. solisorta, 600 - 840 in others; ♂ = 425 - 460 in T. solisorta, 560 - 820 in others) and distributed in the west (T. solisorta is known only from the Ouachita Mountains in Arkansas).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
02900DCF4969DD929E148EA2E89B4BF7.taxon	description	Description. Female (Figure 247) (n = 5) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (475 - 500 (500) long; 325 - 360 (350) wide) ovoid with bluish-purple to purple coloration restricted posteriorly with orange immediately anterior and fading anteriorly. Anterio-medial platelets (102.5 - 112.5 (112.5) long; 42.5 - 46.25 (45) wide). Anterio-lateral platelets (138.75 - 150 (147.5) long; 47.5 - 50 (47.5) wide) free from dorsal plate. Dgl- 4 closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 220 - 240 (240 )). Dorsal plate proportions: dorsum length / width 1.38 - 1.46 (1.43); dorsal width / distance between Dgl- 4 1.45 - 1.50 (1.46); anterio-medial platelet length / width 2.22 - 2.53 (2.50); anterio-lateral platelet length / width 2.78 - 3.16 (3.11); anterio-lateral / anterio-medial length 1.31 - 1.43 (1.31). Gnathosoma - Subcapitulum (280 - 300 (300) long (ventral); 199 - 217 (217) long (dorsal); 85 - 92.5 (92.5) tall) elongate and colorless. Rostrum (105 - 110 (110) long; 35 - 37.5 (36.25) wide). Chelicerae (256 - 281 (278) long) with curved fangs (34 - 44 (43) long). Subcapitular proportions: ventral length / height 3.14 - 3.29 (3.24); rostrum length / width 2.87 - 3.03 (3.03). Pedipalps elongate (especially tibiae) with tuberculate ventral extensions on femora and genua ending broadly and dentate. Palpomeres: trochanter (27.5 - 30 (30) long); femur (86.25 - 93.75 (93.75) long); genu (52.5 - 55 (55) long); tibia (80 - 85 (85) long; 13.75 - 15 (15) wide); tarsus (12.5 - 15 (12.5) long). Palpomere proportions: femur / genu 1.64 - 1.70 (1.70); tibia / femur 0.91 - 0.99 (0.91); tibia length / width 5.67 - 5.82 (5.67). Venter - (580 - 640 (630) long; 356 - 439 (380) wide) with bluish-purple or purple coloration. Gnathosomal bay (97.5 - 110 (110) long; 70 - 77.5 (77.5) wide). Cxgl- 4 far from apex. Medial suture (47.5 - 50 (47.5) long). Genital plates (132.5 - 140 (140) long; 117.5 - 122.5 (122.5) wide). Additional measurements: Cx- 1 (228 - 249 (249) long (total); 88 - 121 (121) long (medial )); Cx- 3 (236.5 - 259.75 (252) wide); anterior venter (192.5 - 207.5 (205) long). Ventral proportions: gnathosomal bay length / width 1.30 - 1.50 (1.42); anterior venter / genital field length 1.45 - 1.54 (1.46); anterior venter length / genital field width 1.60 - 1.77 (1.67); anterior venter / medial suture 3.85 - 4.37 (4.32). Male (Figure 248) (n = 6) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (425 - 460 (435) long; 305 - 320 (305) wide) ovoid with bluish-purple to purple coloration restricted posteriorly with orange immediately anterior and fading anteriorly. Anterio-medial platelets (97.5 - 105 (100) long; 40 - 45 (42.5) wide). Anterio-lateral platelets (130 - 145 (145) long; 45 - 50 (45) wide) free from dorsal plate. Dgl- 4 closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 205 - 220 (215 )). Dorsal plate proportions: dorsum length / width 1.37 - 1.48 (1.43); dorsal width / distance between Dgl- 4 1.42 - 1.51 (1.42); anterio-medial platelet length / width 2.33 - 2.44 (2.35); anterio-lateral platelet length / width 2.85 - 3.22 (3.22); anterio-lateral / anterio-medial length 1.32 - 1.45 (1.45). Gnathosoma - Subcapitulum (252.5 - 265 (255) long (ventral); 181 - 196 (190) long (dorsal); 80 - 87.5 (80) tall) elongate and colorless. Rostrum (92.5 - 100 (100) long; 32.5 - 35 (35) wide). Chelicerae (240 - 246 (240) long) with curved fangs (33 - 45 (45) long). Subcapitular proportions: ventral length / height 2.97 - 3.19 (3.19); rostrum length / width 2.81 - 2.92 (2.86). Pedipalps elongate (especially tibiae) with tuberculate ventral extensions on femora and genua ending broadly and dentate. Palpomeres: trochanter (27.5 - 30 (27.5) long); femur (80 - 82.5 (80) long); genu (48.75 - 51.25 (48.75) long); tibia (77.5 - 82.5 (77.5) long; 15 - 16.25 (15) wide); tarsus (12.5 - 15 (12.5) long). Palpomere proportions: femur / genu 1.59 - 1.65 (1.64); tibia / femur 0.94 - 1.00 (0.97); tibia length / width 5.08 - 5.17 (5.17). Venter - (520 - 570 (545) long; 335 - 382 (335) wide) with bluish-purple or purple coloration. Gnathosomal bay (85 - 107.5 (102.5) long; 60 - 72.5 (67.5) wide). Cxgl- 4 far from apex. Medial suture (65 - 77.5 (75) long). Genital plates (105 - 115 (105) long; 90 - 95 (92.5) wide). Additional measurements: Cx- 1 (208 - 231 (230) long (total); 91 - 130 (130) long (medial )); Cx- 3 (240 - 281 (240) wide); anterior venter (210 - 225 (215) long). Ventral proportions: gnathosomal bay length / width 1.17 - 1.52 (1.52); anterior venter / genital field length 1.87 - 2.10 (2.05); anterior venter length / genital field width 2.27 - 2.50 (2.32); anterior venter / medial suture 2.87 - 3.26 (2.87). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
02900DCF4969DD929E148EA2E89B4BF7.taxon	etymology	Etymology. Specific epithet (solisorta) refers to the dorsal coloration of this species, which is the easiest way to differentiate it from its sister species (T. nigroalba). If one imagines the posterior purple color as the landscape and horizon, then the orangish coloration resembles the sun beginning to rise (solis ortus, L. sunrise).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
02900DCF4969DD929E148EA2E89B4BF7.taxon	distribution	Distribution. Ouachita Mountains (Figure 246).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
0A854173AC93A310E2A8A486DF7BEA12.taxon	materials_examined	Material examined. HOLOTYPE (♀): from USA, California, Santa Cruz County, Waddell Creek, 28 April 1933, PR Needham, RM 330007. PARATYPES (1 ♀; 0 ♂): from USA, California, El Dorado County, South Lake Tahoe, Taylor Creek, 1 Sep 1932, by PR Needham, RM 320005. OTHER MATERIAL (49 ♀; 59 ♂): British Columbia, Canada: 2 ♀ and 1 ♂ from Ryan Rest Area off Highway 3, east of Yahk Moyie River, 15 Aug 2012, by IM Smith, IMS 120071 * 1 ♀ and 1 ♂ from Vancouver Island, beside Highway 4, 35.6 kilometers east of Pacific Rim Road, 9 Jul 1976, by IM Smith, IMS 760206 * 2 ♀ and 1 ♂ from Vancouver Island, Caycuse, Nixon Creek, 8 Jul 1976, by IM Smith, IMS 760197 & IMS 760198 * 1 ♀ from Vancouver Island, Lake Cowichan, Cowichan River, above Skutz Falls, 9 Jul 1979, by IM Smith, IMS 790035 * 1 ♀ and 2 ♂ from Vancouver Island, Lake Cowichan, Robertson River, South Shore Road, north of Mesachie Lake, 10 Jul 1976, by IM Smith, IMS 760183 A * 1 ♂ from Vancouver Island, Lake Cowichan, tributary of Robertson River, South Shore Road, north of Mesachie Lake, 10 Jul 1976, by IM Smith, IMS 760183 B * 2 ♀ and 2 ♂ from Vancouver Island, Lake Cowichan, Skutz Falls, Skutz Creek, near Cowichan River, 9 Jul 1979, by IM Smith, IMS 790036 A & IMS 790036 B * 1 ♀ and 1 ♂ from Vancouver Island, Lost Shoe Creek, beside Highway 4, 1.3 kilometers east of Pacific Rim Road, 9 Jul 1976, by IM Smith, IMS 760203 * 1 ♀ from Vancouver Island, spring run beside South Shore Road, 2.3 kilometers north of Lake Cowichan, 6 Jun 1979, by IM Smith, IMS 790007 * 1 ♀ and 1 ♂ from Vancouver Island, Ucluelet, beside Highway 4, 16.6 kilometers east of Pacific Rim Road, 18 - 19 Jul 1979, by IM Smith, IMS 790047 * 1 ♀ from Vancouver Island, Youbou, Shaw Creek, North Shore Road, 4.3 kilometers south of north end of Cowichan Lake, 8 Jul 1976, by IM Smith, IMS 760196 * California, USA: 1 ♂ from Calaveras County, Stanislaus National Forest, North Fork Stanislaus River (38 ° 25 ' 20 " N, 120 ° 2 ' 47 " W), 30 Aug 2013, by JR Fisher, JRF 13 - 0830 - 005 * 1 ♀ and 1 ♂ from El Dorado County, El Dorado National Forest, Taylor Creek (38 ° 55 ' 59 " N, 120 ° 3 ' 21 " W), 27 Aug 2013, by JR Fisher, JRF 13 - 0827 - 003 * 1 ♀ from Mendocino County, Jackson Demonstration State Park, North Fork of Big River (39 ° 20 ' 46 " N, 123 ° 30 ' 35 " W), 22 Aug 2013, by JR Fisher, JRF 13 - 0822 - 002 * 1 ♀ and 1 ♂ from Mendocino County, Cottaneva Creek, beside Route 1, 21.8 kilometers southwest of Route 101, 5 Aug 1987, by IM Smith, IMS 870129 A * 1 ♀ and 1 ♂ from Mendocino County, small stream at beach access road, off Route 1, 2.6 kilometers south of Westport, 5 Aug 1987, by IM Smith, IMS 870128 A * 2 ♀ from Monterey County, Nacimiento River, beside Nacimiento-Ferguson Road at Nacimiento campground, 30 Jul 1987, by IM Smith, IMS 870120 A * 1 ♀ and 1 ♂ from Monterey County, Salmon Creek, beside Route 1, south of Gorda, 28 Jul 1987, by IM Smith, IMS 870114 A * 1 ♀ and 1 ♂ from Monterey County, spring run on north side of Salmon Creek, beside Route 1, south Gorda, 29 Jul 1987, by IM Smith, IMS 870117 * 1 ♂ from Nevada County, Tahoe National Forest, Sagehen Creek (39 ° 26 ' 2 " N, 120 ° 12 ' 17 " W), 26 Aug 2013, by JR Fisher, JRF 13 - 0826 - 006 * 1 ♀ and 1 ♂ from Plumas County, beside Route 89, north of Greenville, 14 Jun 1976, by IM Smith, IMS 760113 * 3 ♀ and 3 ♂ from Shasta County, Battle Creek, beside Route 44, 5.6 kilometers west of Viola, 10 Aug 1987, by IM Smith, IMS 870139 A * 1 ♂ from Trinity County, small cascading trickle beside Route 36, 5.2 kilometers west of Forest Glen Station, 6 Aug 1987, by IM Smith, IMS 870132 * 1 ♀ and 1 ♂ from Tulare County, Johnsondale, Double Bunk Creek, beside SM 50, 2.2 kilometers east of Double Bunk Meadow, 31 Jul 1987, by IM Smith, IMS 870123 * 1 ♂ from Tulare County, Kern River, Brush Creek Flat, beside SM 99 between Roads End Station & Johnsondale, 31 Jul 1987, by IM Smith, IMS 870121 * 1 ♂ from Tulare County, Stony Creek, Stony Creek Picnic Area, east of Sequoia National Park, 1 Aug 1987, by IM Smith, IMS 870124 A * 1 ♀ and 2 ♂ from Ventura County, Ojai, North Fork of Ventura River, beside Route 33, just above Wheeler Gorge, 25 - 26 Jul 1987, by IM Smith, IMS 870109 A & IMS 870109 B * 1 ♀ and 1 ♂ from Ventura County, Ojai, North Fork of Ventura River just below Wheeler Gorge campground, beside Route 33, 27 Jul 1987, by IM Smith, IMS 870112 * Idaho, USA: 1 ♀ from Custer County, Challis National Forest, Squaw Creek (44 ° 19 ' 35 " N, 114 ° 28 ' 15 " W), 30 Jul 2012, by JR Fisher, WA Nelson, & JC O'Neill, ROW 12 - 0730 - 002 * 1 ♂ from Lemhi County, Salmon National Forest, Niapas Creek (45 ° 8 ' 15 " N, 114 ° 13 ' 4 " W), 2 Aug 2012, by JR Fisher, WA Nelson, & JC O'Neill, ROW 12 - 0802 - 003 * Montana, USA: 1 ♀ and 1 ♂ from Missoula County, Lolo National Forest, Lolo Creek (46 ° 46 ' 7 " N, 114 ° 27 ' 53 " W), 7 Aug 2012, by JR Fisher, WA Nelson, & JC O'Neill, ROW 12 - 0807 - 003 * 1 ♀ and 1 ♂ from Ravalli County, Bitterroot National Forest, West Fork Bitterroot River (45 ° 54 ' 38 " N, 114 ° 9 ' 43 " W), 6 Aug 2012, by JR Fisher, WA Nelson, & JC O'Neill, ROW 12 - 0806 - 003 * 1 ♀ from Ravalli County, Medicine Springs, Spring Gulch campground, East Fork of Bitterroot River, beside Route 93, 1 Jul 1985, by IM Smith, IMS 850060 * Oregon, USA: 2 ♀ and 3 ♂ from Coos County, Siskiyou National Forest, Road 33 between Powers & Agness, Coal Creek, 2 Jul 1983, by IM Smith, IMS 830015 * 1 ♂ from Curry County, Port Orford, Butler Bar campground, Elk River, 25 Jun 1976, by IM Smith, IMS 760162 * 1 ♂ from Curry County, Port Orford, Butler Bar campground, Elk River, 25 - 26 Jun 1976, by IM Smith, IMS 760163 * 1 ♀ and 2 ♂ from Curry County, Port Orford, Humbug Mountain State Park Picnic Area, beside Route 1, Brush Creek, 1 Jul 1983, by IM Smith, IMS 830012 * 2 ♀ and 6 ♂ from Curry County, Port Orford, Humbug Mountain State Park Picnic Area, Brush Creek, beside Route 1, 3 Jul 1983, by IM Smith, IMS 830020 A & IMS 830020 B * 1 ♀ from Curry County, Port Orford, Humbug Mountain State Park Picnic Area, beside Route 1, 1 Jul 1983, by IM Smith, IMS 830013 * 3 ♀ and 5 ♂ from Curry County, Sixes, Sixes River, beside road at mouth of Edson Creek, 4 Jul 1983, by IM Smith, IMS 830021 A & IMS 830021 B * 2 ♂ from Lane County, Gate Creek (44 ° 8 ' 48 " N, 122 ° 34 ' 20 " W), 11 Aug 2013, by JC O'Neill, & WA Nelson, JNOW 13 - 0811 - 001 * 2 ♀ and 1 ♂ from Lincoln County, Blackberry campground, near Tidewater, Alsea River, 28 Jun 1983, by IM Smith & AB Smith, IMS 830009 * 2 ♀ and 3 ♂ from Multnomah County, Columbia River Scenic Highway, Horsetail Falls, 27 Jun 1983, by IM Smith, IMS 830005 * Washington, USA: 2 ♀ from Clallam County, Green Creek (48 ° 10 ' 45 " N, 124 ° 12 ' 21 " W), 24 Jul 2013, by JC O'Neill, & WA Nelson, JNOW 13 - 0724 - 005 * 1 ♂ from Clallam County, Olympic National Forest, Jimmy Come Lately Creek (47 ° 59 ' 5 " N, 123 ° 0 ' 5 " W), 23 Jul 2013, by JC O'Neill, & WA Nelson, JNOW 13 - 0724 - 001 * 1 ♀ from Clallam County, Whiskey Creek (48 ° 8 ' 23 " N, 123 ° 47 ' 7 " W), 24 Jul 2013, by JC O'Neill, & WA Nelson, JNOW 13 - 0724 - 004 * 1 ♀ and 1 ♂ from Lewis County, Gifford Pinchot National Forest, Snake Creek (46 ° 38 ' 52 " N, 121 ° 43 ' 8 " W), 23 Jul 2013, by JC O'Neill, & WA Nelson, JNOW 13 - 0723 - 006 * 2 ♂ from Snohomish County, Mount Baker National Forest, Marten River (48 ° 4 ' 19 " N, 121 ° 36 ' 24 " W), 28 Jul 2013, by JC O'Neill, & WA Nelson, JNOW 13 - 0728 - 002.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
0A854173AC93A310E2A8A486DF7BEA12.taxon	diagnosis	Diagnosis. Torrenticola tahoei are similar to the other member of Tahoei group, T. oregonensis by subcapitulum shape, having purple coloration restricted posteriorly and being distributed in the west and to members of the Nigroalba Group (T. flangipalpa, T. nigroalba, T. solisorta, and T. dentirostra) in having purple dorsal coloration restricted posteriorly. T. tahoei can be differentiated from the Nigroalba Group by being larger (dorsum length ♀ = 600 - 720 in T. tahoei, 475 - 565 in Nigroalba Group; ♂ = 560 - 650 in T. tahoei, 425 - 510 in Nigroalba Group) and distributed in the west (Nigroalba Group are eastern). T. tahoei can be differentiated from T. oregonensis by being smaller (dorsum length ♀ = 600 - 720 in T. tahoei, 760 - 840 in T. oregonensis; ♂ = 560 - 650 in T. tahoei, 690 - 820 in T. oregonensis; dorsum width ♀ = 430 - 515 in T. tahoei, 560 - 640 in T. oregonensis; ♂ = 400 - 460 in T. tahoei, 520 - 605 in T. oregonensis) and a more elongate subcapitulum (ventral length / width = 3.25 - 4.11 in T. tahoei, 2.63 - 2.74 in T. oregonensis).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
0A854173AC93A310E2A8A486DF7BEA12.taxon	description	Re-description. Female (Figure 250) (n = 7) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (600 - 720 (640) long; 430 - 515 (515) wide) ovoid with purple coloration restricted posteriorly. Anterio-medial platelets (115 - 140 (132.5) long; 62.5 - 77.5 (72.5) wide). Anterio-lateral platelets (177.5 - 202.5 (192.5) long; 65 - 85 (75) wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 335 - 400 (380 )). Dorsal plate proportions: dorsum length / width 1.24 - 1.44 (1.24); dorsal width / distance between Dgl- 4 1.25 - 1.43 (1.36); anterio-medial platelet length / width 1.68 - 1.96 (1.83); anterio-lateral platelet length / width 2.31 - 2.76 (2.57); anterio-lateral / anterio-medial length 1.40 - 1.57 (1.45). Gnathosoma - Subcapitulum (345 - 390 (390) long (ventral); 266 - 315 (315) long (dorsal); 95 - 110 (95) tall) elongate and colorless. Rostrum (142.5 - 157.5 (157.5) long; 40 - 47.5 (45) wide) elongate. Chelicerae (358 - 395 (395) long) with curved fangs (50 - 56 (52.5) long). Subcapitular proportions: ventral length / height 3.41 - 4.11 (4.11); rostrum length / width 3.11 - 3.81 (3.50). Pedipalps elongate (especially tibiae) with broad, flat, dentate, and anteriorly-directed ventral extensions on femora and broadly tuberculate, dentate ventral extensions on genua. Palpomeres: trochanter (32.5 - 40 long); femur (102.5 - 117.5 long); genu (72.5 - 85 long); tibia (87.5 - 97.5 long; 21.25 - 25 wide); tarsus (15 - 17.5 long). Palpomere proportions: femur / genu 1.38 - 1.52; tibia / femur 0.82 - 0.86; tibia length / width 3.89 - 4.22. Venter - (770 - 850 (810) long; 466 - 565 (565) wide) mostly colorless with areas of purple coloration. Gnathosomal bay (95 - 112.5 (100) long; 65 - 80 (70) wide). Cxgl- 4 far from apex. Medial suture (95 - 130 (122.5) long). Genital plates (160 - 172.5 (166.25) long; 145 - 160 (160) wide). Additional measurements: Cx- 1 (266 - 301 (290) long (total); 159 - 195 (190) long (medial )); Cx- 3 (297 - 340 (335) wide); anterior venter (285 - 322.5 (312.5) long) elongate. Ventral proportions: gnathosomal bay length / width 1.19 - 1.59 (1.43); anterior venter / genital field length 1.70 - 1.88 (1.88); anterior venter length / genital field width 1.84 - 2.08 (1.95); anterior venter / medial suture 2.37 - 3.11 (2.55). Male (Figure 251) (n = 5) with characters of the genus with following specifications. Dorsum - (560 - 650 long; 400 - 460 wide) ovoid with purple coloration restricted posteriorly. Anterio-medial platelets (112.5 - 125 long; 57.5 - 67.5 wide). Anterio-lateral platelets (163.75 - 200 long; 65 - 80 wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 300 - 370). Dorsal plate proportions: dorsum length / width 1.30 - 1.41; dorsal width / distance between Dgl- 4 1.24 - 1.33; anterio-medial platelet length / width 1.70 - 1.96; anterio-lateral platelet length / width 2.38 - 2.71; anterio-lateral / anterio-medial length 1.44 - 1.65. Gnathosoma - Subcapitulum (325 - 347.5 long (ventral); 244 - 268 long (dorsal); 92.5 - 106.25 tall) elongate and colorless. Rostrum (130 - 140 long; 40 - 45 wide) elongate. Chelicerae (320 - 353 long) with curved fangs (41 - 56 long). Subcapitular proportions: ventral length / height 3.25 - 3.57; rostrum length / width 3.06 - 3.31. Pedipalps elongate (especially tibia) with broad, flat, dentate, and anteriorly-directed ventral extensions on femora and broadly tuberculate, dentate ventral extensions on genua. Palpomeres: trochanter (35 - 37.5 long); femur (100 - 107.5 long); genu (67.5 - 75 long); tibia (82.5 - 92.5 long; 20 - 23.75 wide); tarsus (12.5 - 17.5 long). Palpomere proportions: femur / genu 1.43 - 1.48; tibia / femur 0.80 - 0.88; tibia length / width 3.67 - 4.13. Venter - (720 - 790 long; 450 - 517 wide) mostly colorless with areas of purple coloration. Gnathosomal bay (95 - 105 long; 65 - 77.5 wide). Cxgl- 4 far from apex. Medial suture (120 - 147.5 long). Genital plates (132.5 - 142.5 long; 110 - 122.5 wide). Additional measurements: Cx- 1 (232 - 264 long (total); 138 - 183 long (medial )); Cx- 3 (285 - 315 wide); anterior venter (305 - 325 long) elongate. Ventral proportions: gnathosomal bay length / width 1.27 - 1.54; anterior venter / genital field length 2.21 - 2.34; anterior venter length / genital field width 2.57 - 2.82; anterior venter / medial suture 2.18 - 2.54. Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
0A854173AC93A310E2A8A486DF7BEA12.taxon	etymology	Etymology. Marshall (1943) did not specify an etymology for this specific epithet (tahoei). It surely refers to the collection locality of the paratype, Taylor's Creek near Lake Tahoe, California, although the holotype was collected from Waddell Creek near the coast.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
0A854173AC93A310E2A8A486DF7BEA12.taxon	distribution	Distribution. Western, but not known from the southwest (Figure 249). T. tahoei was originally reported only from a few localities in California; we expand its known range into Oregon and Washington, and the Rocky Mountains.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
64D9C6D4DDE6A3C99C38384FD4C395C3.taxon	materials_examined	Material examined. HOLOTYPE (♂): from USA, New Jersey, Sussex County, Little Flatbrook, north of Bevans, 12 Oct 1953, by H Habeeb, HH 530110. PARATYPES (1 ♀; 0 ♂): New Jersey, USA: 1 ♀ (ALLOTYPE) from Sussex County, Little Flatbrook, north of Bevans, 12 Oct 1953, by H Habeeb, HH 530110. OTHER MATERIAL (15 ♀; 8 ♂): Georgia, USA: 1 ♀ and 2 ♂ from Chattooga County, East Fork of Little River, Cloudland (34 ° 31 ' 25 " N, 85 ° 30 ' 23 " W), 28 Sep 1992, by IM Smith, IMS 920056 A * New Hampshire, USA: 1 ♀ from Coos County, picnic area beside Route 110, Upper Ammonoosuc River (44 ° 36 ' N, 71 ° 24 ' W), 5 Jul 1989, by IM Smith, IMS 890071 * New Jersey, USA: 1 ♀ and 1 ♂ from Warren County, Pequest River, 16 kilometers west of Hackettstown, 15 Aug 1964, by DR Cook, DRC 640020 * New York, USA: 1 ♀ from Hamilton County, beside Route 8, 4.8 kilometers from Warren County line, 19 Aug 1964, by DR Cook, DRC 640026 * Nova Scotia, Canada: 2 ♀ and 1 ♂ from Guysborough County, Sherbrooke, Sherbrooke Picnic Park beside Highway 7, 17 Sep 2011, by IM Smith, IMS 110087 * Ontario, Canada: 1 ♀ from Hastings County, Moira River, Vanderwater Conservation Area, off Highway 37 south of Tweed, 20 Sep 1983, by IM Smith & CJ Hill, IMS 830086 C * 1 ♀ from Muskoka District, Baysville, shallow bay on south side of Echo Lake, 18 Aug 1981, by IM Smith, IMS 810023 * South Carolina, USA: 1 ♀ from Greenville County, Matthews Creek, 24 Apr 2014, by D Eargle, JRF 14 - 0424 - 001 * Tennessee, USA: 1 ♀ from Blount County, Great Smoky National Park, Abrams River (35 ° 35 ' 30 " N, 83 ° 51 ' 20 " W), 17 Sep 2010, by IM Smith, IMS 100142 * 1 ♀ from Monroe County, Tellico River (35 ° 19 ' N, 84 ° 10 ' W), 5 Jul 1990, by IM Smith, IMS 900079 * 1 ♂ from Monroe County, Tellico River (35 ° 20 ' 27 " N, 84 ° 11 ' 31 " W), 12 Sep 2009, by IM Smith, IMS 090111 * 3 ♀ and 1 ♂ from Sevier County, Great Smoky Mountains National Park, Middle Prong Little Pigeon River (35 ° 43 ' 33 " N, 83 ° 24 ' 1 " W), 12 Sep 2010, by IM Smith, IMS 100131 * 1 ♂ from Sevier County, Great Smoky Mountains National Park, Middle Prong Little Pigeon River (35 ° 44 ' 12 " N, 83 ° 24 ' 51 " W), 12 Sep 2010, by IM Smith, IMS 100132 * Virginia, USA: 1 ♂ from Alleghany County, Covington, Potts Creek, beside Route 18, 0.5 kilometers north of Route 657, 13 Jul 1990, by IM Smith, IMS 900091 B * 1 ♀ from Rappahannock County, Sperrysville, North Fork of Thornton River, beside Route 612, 0.5 miles from Route 211, 8 Sep 1968, by DR Cook, DRC 680066.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
64D9C6D4DDE6A3C99C38384FD4C395C3.taxon	diagnosis	Diagnosis. Torrenticola tricolor are similar to other members of the Tricolor Complex (T. bittikoferae, T. hoosieri, T. larvata, T. pearsoni, T. olliei, T. sierrensis, T. trimaculata, T. unimaculata, T. cardia, T. kringi, T. dimorpha, and T. mohawk) in having a short, conical rostrum. T. tricolor can be differentiated from all Torrenticola, including other members of the Tricolor Complex, by having a distinct dorsal pattern. Although the spots always have this basic construction, there is great variability between specimens. T. tricolor are most similar to other members of the Tricolor Complex that have bold patterning (T. larvata, T. unimaculata, T. trimaculata, cardia, kringi, and T. mohawk). T. tricolor can be further differentiated from T. larvata by being rounder (dorsum length / width = 1.2 - 1.38 in T. tricolor, 1.41 - 1.57 in T. larvata) and stockier pedipalp tibiae (length / width ♀ = 3.0 - 3.11 in T. tricolor, 3.25 - 3.5 in T. larvata; ♂ = 2.7 - 2.8 in T. tricolor, 3.1 - 3.2 in T. larvata). T. tricolor can be further differentiated from T. trimaculata by having longer genital field (♀ = 187.5 - 210 in T. tricolor, 157.5 - 185 in T. trimaculata; ♂ = 145 - 170 in T. tricolor, 120 - 140 in T. trimaculata) and a slightly more elongate rostrum (length / width ♀ = 2.14 - 2.39 in T. tricolor, 1.91 - 2.1 in T. trimaculata; ♂ = 2.37 - 2.5 in T. tricolor, 2.05 - 2.22 in T. trimaculata). T. tricolor can be further differentiated from T. unimaculata by having stockier pedipalp tibiae (♀ = 3.0 - 3.11 in T. tricolor, 3.2 - 3.4 in T. unimaculata; ♂ = 2.69 - 2.8 in T. tricolor, 2.9 - 3.11 in T. unimaculata) and shorter medial suture in females (15 - 22.5 in T. tricolor, 40 - 47.5 in T. unimaculata). T. tricolor can be further differentiated from T. cardia by having a rounder dorsum (length / width = 1.2 - 1.38 in T. tricolor, 1.39 - 1.54 in T. cardia) and longer pedipalpal genua (75 - 83 in T. tricolor, 60 - 70 in T. cardia). T. tricolor can be further differentiated from T. kringi by having a stockier rostrum (length / width = 2.14 - 2.50 in T. tricolor, 2.67 - 3.13 in T. kringi). T. tricolor can be differentiated from T. mohawk by having a more elongate rostrum (length / width ♀ = 2.14 - 2.39 in T. tricolor, 1.80 - 2.00 in T. mohawk; ♂ = 2.37 - 2.50 in T. tricolor, 2.00 - 2.13 in T. mohawk).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
64D9C6D4DDE6A3C99C38384FD4C395C3.taxon	description	Re-description. Male (Figure 253) (n = 6) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (580 - 660 (640) long; 435 - 520 (520) wide) circular to ellipsoid with reddish-purple or bluish-purple to navy blue coloration in two large posterior spots and a smaller anterior spot all merged together, often with orange between the posterior spots. Anterio-medial platelets (120 - 137.5 (137.5) long; 62.5 - 70 (70) wide). Anterio-lateral platelets (175 - 207.5 (192.5) long; 72.5 - 95 (95) wide) free from dorsal plate. Dgl- 4 usually closer to the edge of the dorsum than to the muscle scars, occasionally halfway between the dorsum edge and muscle scars (distance between Dgl- 4 295 - 375 (325 )). Dorsal plate proportions: dorsum length / width 1.23 - 1.38 (1.23); dorsal width / distance between Dgl- 4 1.33 - 1.60 (1.60); anterio-medial platelet length / width 1.92 - 2.00 (1.96); anterio-lateral platelet length / width 2.03 - 2.55 (2.03); anterio-lateral / anterio-medial length 1.40 - 1.58 (1.40). Gnathosoma - Subcapitulum (247.5 - 295 (247.5) long (ventral); 197.5 - 226 (197.5) long (dorsal); 105 - 122.5 (112.5) tall) colorless. Rostrum (107.5 - 125 (107.5) long; 45 - 52.5 (45) wide) short and conical. Chelicerae (254 - 290 (260) long) with curved fangs (42 - 56 (52.5) long). Subcapitular proportions: ventral length / height 2.20 - 2.62 (2.20); rostrum length / width 2.37 - 2.50 (2.39). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (37.5 - 45 (37.5) long); femur (95 - 110 (95) long); genu (70 - 80 (75) long); tibia (76.25 - 90 (76.25) long; 28.75 - 32.5 (28.75) wide); tarsus (22.5 - 27.5 (22.5) long). Palpomere proportions: femur / genu 1.27 - 1.45 (1.27); tibia / femur 0.80 - 0.86 (0.80); tibia length / width 2.65 - 2.80 (2.65). Venter - (720 - 804 (790) long; 448 - 660 (660) wide) colorless. Gnathosomal bay (110 - 145 (126.25) long; 75 - 97.5 (85) wide). Cxgl- 4 subapical. Medial suture (105 - 137.5 (105) long). Genital plates (145 - 170 (157.5) long; 107.5 - 115 (107.5) wide). Additional measurements: Cx- 1 (271 - 290.5 (280) long (total); 139 - 163 (150) long (medial )); Cx- 3 (316 - 401 (382.5) wide); anterior venter (255 - 297.5 (262.5) long). Ventral proportions: gnathosomal bay length / width 1.33 - 1.61 (1.49); anterior venter / genital field length 1.67 - 1.92 (1.67); anterior venter length / genital field width 2.37 - 2.77 (2.44); anterior venter / medial suture 2.16 - 2.50 (2.50). Female (Figure 254) (n = 6) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (600 - 755 (610) long; 470 - 575 (510) wide) circular to ellipsoid with reddish-purple or bluish-purple to navy blue coloration in two large posterior spots and a smaller anterior spot all merged together, often with orange between the posterior spots. Anterio-medial platelets (135 - 148.75 (137.5) long; 66.25 - 75 (70) wide). Anterio-lateral platelets (195 - 207.5 (195) long; 82.5 - 100 (100) wide) free from dorsal plate. Dgl- 4 usually closer to the edge of the dorsum than to the muscle scars, occasionally halfway between the dorsum edge and muscle scars (distance between Dgl- 4 320 - 410 (320 )). Dorsal plate proportions: dorsum length / width 1.20 - 1.35 (1.20); dorsal width / distance between Dgl- 4 1.36 - 1.59 (1.59); anterio-medial platelet length / width 1.93 - 2.20 (1.96); anterio-lateral platelet length / width 1.95 - 2.52 (1.95); anterio-lateral / anterio-medial length 1.33 - 1.46 (1.42). Gnathosoma - Subcapitulum (285 - 330 (285) long (ventral); 210 - 251 (210) long (dorsal); 132.5 - 142.5 (132.5) tall) colorless. Rostrum (112.5 - 137.5 (112.5) long; 52.5 - 57.5 (52.5) wide) short and conical. Chelicerae (300 - 332 (300) long) with curved fangs (49 - 63 (52.5) long). Subcapitular proportions: ventral length / height 2.15 - 2.42 (2.15); rostrum length / width 2.14 - 2.39 (2.14). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (42.5 - 47.5 (42.5) long); femur (107.5 - 120 (107.5) long); genu (75 - 82.5 (75) long); tibia (90 - 105 (90) long; 30 - 33.75 (30) wide); tarsus (22.5 - 27.5 (22.5) long). Palpomere proportions: femur / genu 1.41 - 1.48 (1.43); tibia / femur 0.84 - 0.89 (0.84); tibia length / width 3.00 - 3.11 (3.00). Venter - (690 - 890 (740) long; 561 - 665 (665) wide) colorless. Gnathosomal bay (145 - 165 (145) long; 91.25 - 102.5 (91.25) wide). Cxgl- 4 subapical. Medial suture (15 - 22.5 (20) long). Genital plates (187.5 - 210 (187.5) long; 155 - 171.25 (167.5) wide). Additional measurements: Cx- 1 (265 - 311 (265) long (total); 125 - 151.75 (125) long (medial )); Cx- 3 (375 - 433 (375) wide); anterior venter (152.5 - 192.5 (152.5) long). Ventral proportions: gnathosomal bay length / width 1.53 - 1.70 (1.59); anterior venter / genital field length 0.81 - 0.95 (0.81); anterior venter length / genital field width 0.91 - 1.18 (0.91); anterior venter / medial suture 7.63 - 11.83 (7.63). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
64D9C6D4DDE6A3C99C38384FD4C395C3.taxon	etymology	Etymology. Although Habeeb (1957) did not specify an etymology for the specific epithet (tricolor), it surely refers to his comment on the dorsal coloration, " color black with some red and white " (tres, L. three; color, L. color).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
64D9C6D4DDE6A3C99C38384FD4C395C3.taxon	distribution	Distribution. Northeastern and southward throughout the Appalachians (Figure 252). T. tricolor was previously known only from northern New Jersey.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
1E6A2697BD3ACBE651EDE5DCD05E03AF.taxon	materials_examined	Material examined. HOLOTYPE (♀): USA, Arkansas, Madison County, Withrow Springs State Park, War Eagle Creek (36 ° 8 ' 59.3 " N, 93 ° 44 ' 26.94 " W), 27 Jul 2011, by IM Smith, IMS 110034. PARATYPES (49 ♀; 37 ♂): Arkansas, USA: 1 ♂ (ALLOTYPE) from Madison County, Withrow Springs State Park, War Eagle Creek (36 ° 8 ' 59.3 " N, 93 ° 44 ' 26.94 " W), 27 Jul 2011, by IM Smith, IMS 110034 * 2 ♀ and 3 ♂ from Madison County, Withrow Springs State Park, War Eagle Creek (36 ° 8 ' 59.3 " N, 93 ° 44 ' 26.94 " W), 27 Jul 2011, by IM Smith, IMS 110034 * 1 ♀ from Marion County, Crooked Creek ex. Northern hogsucker (Hypentelium nigricans) (36 ° 15 ' 9.9 " N, 94 ° 26 ' 25.8 " W), 22 Jul 2014, by CT McAllister * 3 ♀ and 2 ♂ from Montgomery County, Ouachita National Forest, Ouachita River (34 ° 34 ' 53.20 " N, 93 ° 53 ' 0.16 " W), 5 Oct 2007, by AJ Radwell, & HW Robison, AJR 070300 A * 8 ♀ and 5 ♂ from Montgomery County, Ouachita National Forest, South Fork of Ouachita River, 29 Jul 2011, by AJ Radwell, & B Crump, AJR 110302 * 2 ♀ and 1 ♂ from Montgomery County, Ouachita National Forest, Ouachita River, 27 Aug 2011, by AJ Radwell, AJR 110307 * 4 ♀ and 4 ♂ from Montgomery County, Ouachita National Forest, South Fork of Ouachita River, 29 Jul 2011, by IM Smith, IMS 110040 * 1 ♀ from Montgomery County, Caddo River, 29 Jul 2011, by IM Smith, IMS 110037 * 1 ♂ from Newton County, Ozark National Forest, Mill Creek (36 ° 3 ' 42.12 " N, 93 ° 8 ' 7.62 " W), 20 Jun 2012, by TD Edwards, TDE 12 - 0620 - 010 * 2 ♀ and 2 ♂ from Newton County, Ozark National Forest, Little Buffalo River, 2 Sep 2012, by TD Edwards, TDE 12 - 0902 - 003 * 1 ♂ from Newton County, Buffalo National River, Whiteley Creek (35 ° 59 ' 28.14 " N, 93 ° 23 ' 57.24 " W), 23 May 2012, by TD Edwards, TDE 12 - 0523 - 002 * Illinois, USA: 2 ♀ and 1 ♂ from Union County, Clear Creek (37 ° 33 ' N, 89 ° 23 ' W), 13 Sep 1991, by IM Smith, IMS 910036 A * Indiana, USA: 1 ♀ from Wayne County (39 ° 51 ' 13 " N, 85 ° 8 ' 4 " W), 24 Jul 2014, by MJ Skvarla, MS 14 - 0731 - 001 * Georgia, USA: 1 ♀ from Chattooga County, Johns Creek (34 ° 34 ' N, 80 ° 5 ' W), 4 Jul 1990, by IM Smith, IMS 900076 * Kentucky, USA: 1 ♀ and 2 ♂ from McCreary County, Rock Creek (36 ° 42 ' N, 84 ° 36 ' W), 8 Jul 1990, by IM Smith, IMS 900082 B * Michigan, USA: 2 ♀ and 2 ♂ from Barry County, Thornapple River (42 ° 39 ' N, 85 ° 17 ' W), 29 Jul 1959, by DR Cook, DRC 590034 * Missouri, USA: 2 ♀ and 1 ♂ from Crawford County, Huzzah Creek, 23 Jul 2011, by IM Smith, IMS 110029 * New York, USA: 3 ♀ and 1 ♂ from St. Lawrence County, Canton (44 ° 35 ' N, 75 ° 10 ' W), 15 May 1986, by BP Smith, BPS 860508 * 1 ♀ from USA, New York, Delaware Co., Roscoe (41 ° 55 ' N, 74 ° 54 ' W), 11 June 1988, by PW Schefter and R MacCulloch, IMS 880110 * Nova Scotia, Canada: 1 ♀ from Victoria County, Baddeck River (44 ° 52 ' N, 61 ° 5 ' W), 18 Jul 1981, by IM Smith, IMS 810082 * Ontario, Canada: 4 ♀ and 2 ♂ from Grey County, Saugeen River (44 ° 10 ' N, 80 ° 49 ' W), 9 Jun 1989, by IM Smith, IMS 890028 A * 1 ♀ from Madoc (44 ° 30 ' N, 77 ° 28 ' W), 4 May 1980, by IM Smith, IMS 800003 A * 1 ♂ from Renfrew County, Madawaska River (45 ° 21 ' N, 76 ° 40 ' W), 25 May 1980, by IM Smith, IMS 800012 * 1 ♀ and 1 ♂ from Lanark County, Mississippi River (45 ° 3 ' N, 76 ° 23 ' W), 6 Oct 1983, by IM Smith and CJ Hill, IMS 830093 A * Texas, USA: 1 ♀ from Gillespie County, Fredericksburg, Pedernales River (30 ° 14 ' 42 " N, 98 ° 54 ' 50 " W), 30 May 1998, by IM Smith, IMS 980029 * Virginia, USA: 1 ♀ and 1 ♂ from Scott County, North Fork of Holston River (36 ° 39 ' N, 82 ° 28 ' W), 7 Jul 1990, by IM Smith, IMS 0900080 * 2 ♀ and 4 ♂ from Alleghany County, Potts Creek (37 ° 44 ' N, 80 ° 2 ' W), 13 Jul 1990, by IM Smith, IMS 900091 B * 1 ♀ and 1 ♂ from Bath County, Jackson River (38 ° 8 ' N, 79 ° 46 ' W), 16 Jul 1990, by IM Smith, IMS 900100 * West Virginia, USA: 2 ♀ from Pendleton County, North Fork of South Branch of Potomac River (39 ° 0 ' N, 79 ° 22 ' W), 17 Jul 1990, by IM Smith, IMS 900104.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
1E6A2697BD3ACBE651EDE5DCD05E03AF.taxon	diagnosis	Diagnosis. Torrenticola trimaculata are similar to other members of the Tricolor Complex (T. bittikoferae, T. hoosieri, T. larvata, T. pearsoni, T. olliei, T. sierrensis, T. tricolor, T. unimaculata, T. cardia, T. kringi, T. dimorpha, and T. mohawk) in having a short, conical rostrum. T. trimaculata can be differentiated from all Torrenticola, including other members of the Tricolor Complex, by having a distinct dorsal pattern. Individuals are reported in two distinct morphs primarily based upon ventral coloration. T. trimaculata are most similar to other members of the Tricolor Complex that have bold patterning (T. larvata, T. unimaculata, T. tricolor, T. cardia, T. kringi, and T. mohawk). T. trimaculata can be further differentiated from T. larvata by being rounder (dorsum length / width = 1.2 - 1.37 in T. trimaculata, 1.41 - 1.57 in T. larvata); Dgl- 4 closer to the muscle scars (dorsal width / distance between Dgl- 4 = 1.49 - 1.69 in T. trimaculata, 1.18 - 1.35 in T. larvata); and a stockier rostrum (length / width = 1.91 - 2.22 in T. trimaculata, 2.32 - 2.53 in T. larvata). T. trimaculata can be further differentiated from T tricolor by having shorter genital field (♀ = 157.5 - 185 in T. trimaculata, 187.5 - 210 in T. tricolor; ♂ = 120 - 140 in T. trimaculata, 145 - 170 in T. tricolor) and a stockier rostrum (length / width ♀ = 1.91 - 2.1 in T. trimaculata, 2.14 - 2.39 in T. tricolor; ♂ = 2.05 - 2.22 in T. trimaculata, 2.37 - 2.5 in T. tricolor). T. trimaculata can be further differentiated from T. unimaculata by Dgl- 4 closer to the muscles scars (dorsal width / distance between Dgl- 4 = 1.49 - 1.69 in T. trimaculata, 1.23 - 1.41 in T. unimaculata) and females with shorter medial suture (♀ = 17.5 - 27.5 in T. trimaculata, 40 - 47.5 in T. unimaculata). T. trimaculata can be further differentiated from T. cardia by having Dgl- 4 further from the edge of the dorsum (dorsum width / distance between Dgl- 4 = 1.5 - 1.7 in T. trimaculata, 1.15 - 1.4 in T. cardia) and having a stockier rostrum (length / width ♀ = 1.191 - 2.10 in T. trimaculata, 2.24 - 2.50 in T. cardia; ♂ = 2.05 - 2.22 in T. trimaculata, 2.27 - 2.47 in T. cardia). T. trimaculata can be differentiated from T. kringi by having a stockier rostrum (length / width = 1.9 - 2.3 in T. trimaculata, 2.6 - 3.2 in T. kringi). T. trimaculata can be differentiated from T. mohawk by having a shorter anterior venter (♀ = 157 - 180 in T. trimaculata, 187 - 200 in T. mohawk, ♂ = 230 - 260 in T. trimaculata, 290 - 308 in T. mohawk). Additionally, female T. trimaculata can be differentiated from female T. mohawk by having longer pedipalpal tibiae (♀ = 90 - 97 in T. trimaculata, 77 - 88 in T. mohawk).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
1E6A2697BD3ACBE651EDE5DCD05E03AF.taxon	description	Re-description (amended from Fisher et al. 2015). Female (Figure 256) (n = 6) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (570 - 725 (725) long; 455 - 550 (550) wide) circular to ellipsoid with bluish-purple to navy blue coloration in three distinct spots, one anteriorly and two posteriorly, and orange medially. Anterio-medial platelets (125 - 145 (145) long; 60 - 72.5 (67.5) wide). Anterio-lateral platelets (162.5 - 180 (180) long; 80 - 85 (82.5) wide) free from dorsal plate. Dgl- 4 approaching midway between muscle scars and the dorsum edge (distance between Dgl- 4 295 - 335 (325 )). Dorsal plate proportions: dorsum length / width 1.20 - 1.32 (1.32); dorsal width / distance between Dgl- 4 1.49 - 1.69 1.69); anterio-medial platelet length / width 1.93 - 2.19 (2.15); anterio-lateral platelet length / width 2.00 - 2.19 (2.18); anterio-lateral / anterio-medial length 1.23 - 1.30 (1.24). Gnathosoma - Subcapitulum (250 - 280 (280) long (ventral); 202 - 252 (210) long (dorsal); 115 - 135 (130) tall) colorless or with bluish-purple to navy blue coloration. Rostrum (97.5 - 110 (107.5) long; 47.5 - 55 (52.5) wide) short and conical. Chelicerae (261 - 326 (270) long) with curved fangs (48 - 62 (55) long). Subcapitular proportions: ventral length / height 2.04 - 2.17 (2.15); rostrum length / width 1.91 - 2.10 (2.05). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (37.5 - 42.5 (42.5) long); femur (97.5 - 107.5 (103.75) long); genu (65 - 77.5 (70) long); tibia (90 - 96.25 (92.5) long; 27.5 - 30 (27.5) wide); tarsus (22.5 - 25 (22.5) long). Palpomere proportions: femur / genu 1.39 - 1.50 (1.48); tibia / femur 0.88 - 0.92 (0.89); tibia length / width 3.17 - 3.36 (3.36). Venter - (615 - 840 (840) long; 533 - 700 (700) wide) colorless or with variable amount of bluish-purple to navy blue coloration. Gnathosomal bay (130 - 145 (130) long; 80 - 90 (80) wide). Cxgl- 4 subapical. Medial suture (17.5 - 27.5 (20) long). Genital plates (157.5 - 185 (180) long; 152.5 - 185 (180) wide). Additional measurements: Cx- 1 (258 - 306.75 (260) long (total); 109 - 154 (145) long (medial )); Cx- 3 (328 - 390 (390) wide); anterior venter (157.5 - 180 (180) long). Ventral proportions: gnathosomal bay length / width 1.49 - 1.75 (1.63); anterior venter / genital field length 0.90 - 1.03 (1.00); anterior venter length / genital field width 1.00 - 1.14 (1.14); anterior venter / medial suture 6.09 - 9.29 (9.00). Male (Figure 257) (n = 5) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (530 - 595 (595) long; 390 - 440 (435) wide) circular to ellipsoid with bluish-purple to navy blue coloration in three distinct spots, one anteriorly and two posteriorly, and orange medially. Anterio-medial platelets (107.5 - 130 (120) long; 55 - 67.5 (62.5) wide). Anterio-lateral platelets (70 - 87.5 (75) long; 70 - 87.5 (75) wide) free from dorsal plate. Dgl- 4 approaching midway between muscle scars and the dorsum edge (distance between Dgl- 4 250 - 285 (270 )). Dorsal plate proportions: dorsum length / width 1.30 - 1.37 (1.37); dorsal width / distance between Dgl- 4 1.51 - 1.61 (1.61); anterio-medial platelet length / width 1.92 - 2.00 (1.92); anterio-lateral platelet length / width 1.91 - 2.23 (2.23); anterio-lateral / anterio-medial length 1.29 - 1.46 (1.40). Gnathosoma - Subcapitulum (207.5 - 242.5 (230) long (ventral); 175 - 215 (194.75) long (dorsal); 92.5 - 112.5 (102.5) tall) colorless or with bluish-purple to navy blue coloration. Rostrum (87.5 - 100 (93.75) long; 40 - 47.5 (42.5) wide) short and conical. Chelicerae (232 - 286 (251) long) with curved fangs (40 - 53 (49) long). Subcapitular proportions: ventral length / height 2.16 - 2.43 (2.24); rostrum length / width 2.05 - 2.22 (2.21). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (32.5 - 40 (32.5) long); femur (85 - 95 (93.75) long); genu (60 - 70 (65) long); tibia (75 - 82.5 (75) long; 25 - 27.5 (27.5) wide); tarsus (20 - 25 (22.5) long). Palpomere proportions: femur / genu 1.34 - 1.44 (1.44); tibia / femur 0.80 - 0.88 (0.80); tibia length / width 2.73 - 3.00 (2.73). Venter - (588 - 719 (715) long; 431 - 571 (568) wide) colorless or with variable amount of bluish-purple to navy blue coloration. Gnathosomal bay (82.5 - 112.5 (102.5) long; 62.5 - 77.5 (77.5) wide). Cxgl- 4 subapical. Medial suture (87.5 - 105 (105) long). Genital plates (120 - 140 (127.5) long; 92.5 - 105 (95) wide). Additional measurements: Cx- 1 (216 - 297 (261) long (total); 130 - 168 (152) long (medial )); Cx- 3 (287 - 372 (349) wide); anterior venter (230 - 260 (255) long). Ventral proportions: gnathosomal bay length / width 1.32 - 1.55 (1.32); anterior venter / genital field length 1.79 - 2.04 (2.00); anterior venter length / genital field width 92.5 - 105 (95); anterior venter / medial suture 2.43 - 2.63 (2.43). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
1E6A2697BD3ACBE651EDE5DCD05E03AF.taxon	etymology	Etymology. Fisher et al. (2015) named the specific epithet (trimaculata) in reference to the dorsal coloration of three dark spots (tres, L. three; macula, L. spot).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
1E6A2697BD3ACBE651EDE5DCD05E03AF.taxon	distribution	Distribution. Eastern North America (Figure 255).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
6D98F30878C979E4CBA327D09CC84276.taxon	materials_examined	Material examined. HOLOTYPE (♀): from USA, Alabama, Lauderdale County, off Natchez Trace Parkway, 7 km south of Tennessee state line (34 ° 56 ' 31 " N, 87 ° 49 ' 41 " W), 24 Sep 2009, by IM Smith, IMS 090121, DNA 2871. PARATYPES (6 ♀; 5 ♂): Alabama, USA: 1 ♂ (ALLOTYPE) from Lauderdale County, off Natchez Trace Parkway, 7 km south of Tennessee state line (34 ° 56 ' 31 " N, 87 ° 49 ' 41 " W), 24 Sep 2009, by IM Smith, IMS 090121, DNA 2870 * 1 ♀ from Lauderdale County, off Natchez Trace Parkway, 7 km south of Tennessee state line (34 ° 56 ' 31 " N, 87 ° 49 ' 41 " W), 24 Sep 2009, by IM Smith, IMS 090121 * 2 ♀ and 2 ♂ from Lauderdale County, off Natchez Trace Parkway, 7 km south of Tennessee state line (34 ° 56 ' 32 " N, 87 ° 49 ' 43 " W), 27 Sep 2010, by IM Smith, IMS 100163 * 1 ♂ from Lauderdale County, off Natchez Trace Parkway, 7 km south of Tennessee state line (34 ° 56 ' 31 " N, 87 ° 49 ' 41 " W), 27 Sep 2010, by IM Smith, IMS 100162 * Tennessee, USA: 3 ♀ and 1 ♂ from Wayne County, Glenrock Branch Creek (35 ° 15 ' 50 " N, 87 ° 37 ' 34 " W), 24 Sep 2009, by IM Smith, IMS 090124.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
6D98F30878C979E4CBA327D09CC84276.taxon	diagnosis	Diagnosis. Torrenticola tysoni are similar to other members of the Rusetria " Eastern 2 - Plates " group (T. biscutella, T. caerulea, T. delicatexa, T. indistincta, T. malarkeyorum, T. pendula, T. sellersorum, T. ululata, T. whitneyae, T. microbiscutella, and T. feminellai) in having anterio-lateral platelets fused to the dorsal plate, having dorsal coloration separated into anterior and posterior portions (except T. ululata and T. indistincta), and being distributed in the east. It is one of only four Eastern 2 - Plates that have dark, bold, bluish-purple coloration (also T. biscutella, T. sellersorum, and T. pendula). T. tysoni can be further differentiated from other Eastern 2 - Plates by having a more elongate rostrum (length / width = ♀ = 3.06 - 3.31 in T. tysoni, 2.33 - 3.0 in others; ♂ = 3.14 - 3.50 in T. tysoni, 2.50 - 3.05 in others), except T. feminellai (3.05 - 3.38) and female T. pendula (3.0 - 3.06). T. tysoni can be differentiated from T. feminellai and T. pendula by dorsal coloration and pattern.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
6D98F30878C979E4CBA327D09CC84276.taxon	description	Description. Female (Figure 259) (n = 5) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (610 - 670 (670) long; 450 - 475 (475) wide) ovoid with bold bluish-purple coloration separated into anterior and posterior portions, and with faint orange medially. Anterio-medial platelets (125 - 137.5 (135) long; 42.5 - 52.5 (52.5) wide). Anterio-lateral platelets (162.5 - 175 (172.5) long; 55 - 67.5 (67.5) wide) fused to dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 320 - 330 (330 )). Dorsal plate proportions: dorsum length / width 1.35 - 1.41 (1.41); dorsal width / distance between Dgl- 4 1.38 - 1.44 (1.44); anterio-medial platelet length / width 2.50 - 3.18 (2.57); anterio-lateral platelet length / width 2.56 - 3.18 (2.56); anterio-lateral / anterio-medial length 1.18 - 1.40 (1.28). Gnathosoma - Subcapitulum (310 - 327.5 (327.5) long (ventral); 230 - 245 (245) long (dorsal); 130 - 135 (135) tall) mostly colorless. Rostrum (125 - 132.5 (130) long; 40 - 42.5 (42.5) wide). Chelicerae (315 - 380 (350) long) with curved fangs (57.5 - 60 (60) long). Subcapitular proportions: ventral length / height 2.30 - 2.46 (2.43); rostrum length / width 3.06 - 3.31 (3.06). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (37.5 - 47.5 (45) long); femur (112.5 - 120 (120) long); genu (62.5 - 70 (70) long); tibia (80 - 90 (87.5) long; 22.5 - 25 (25) wide); tarsus (17.5 - 22.5 (20) long). Palpomere proportions: femur / genu 1.61 - 1.80 (1.71); tibia / femur 0.70 - 0.80 (0.73); tibia length / width 3.50 - 3.67 (3.50). Venter - (715 - 790 (790) long; 505 - 560 (540) wide) with bold bluish-purple coloration. Gnathosomal bay (162.5 - 172.5 (172.5) long; 72.5 - 100 (85) wide). Cxgl- 4 subapical. Medial suture (13.75 - 22.5 (22.5) long). Genital plates (175 - 185 (180) long; 152.5 - 155 (155) wide). Additional measurements: Cx- 1 (280 - 315 (310) long (total); 115 - 145 (140) long (medial )); Cx- 3 (310 - 350 (335) wide); anterior venter (150 - 172.5 (172.5) long). Ventral proportions: gnathosomal bay length / width 1.65 - 2.24 (2.03); anterior venter / genital field length 0.81 - 0.99 (0.96); anterior venter length / genital field width 0.97 - 1.11 (1.11); anterior venter / medial suture 7.67 - 11.27 (7.67). Male (Figure 260) (n = 5) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (430 - 485 (460) long; 310 - 340 (320) wide) ovoid with bold bluish-purple coloration separated into anterior and posterior portions, and with faint orange medially. Anterio-medial platelets (97.5 - 127.5 (127.5) long; 35 - 40 (40) wide). Anterio-lateral platelets (127.5 - 137.5 (127.5) long; 42.5 - 50 (47.5) wide) fused to dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 235 - 245 (240 )). Dorsal plate proportions: dorsum length / width 1.39 - 1.48 (1.44); dorsal width / distance between Dgl- 4 1.32 - 1.39 (1.33); anterio-medial platelet length / width 2.60 - 3.19 (3.19); anterio-lateral platelet length / width 2.65 - 3.00 (2.68); anterio-lateral / anterio-medial length 1.00 - 1.31 (1.00). Gnathosoma - Subcapitulum (250 - 265 (250) long (ventral); 192.5 - 200 (195) long (dorsal); 77.5 - 95 (90) tall) mostly colorless. Rostrum (100 - 110 (100) long; 30 - 35 (30) wide). Chelicerae (237.5 - 260 (250) long) with curved fangs (45 - 47.5 (45) long). Subcapitular proportions: ventral length / height 2.76 - 3.23 (2.78); rostrum length / width 3.14 - 3.50 (3.33). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (33.75 - 48.75 (37.5) long); femur (87.5 - 97.5 (92.5) long); genu (52.5 - 62.5 (57.5) long); tibia (67.5 - 75 (75) long; 20 - 21.25 (21.25) wide); tarsus (15 - 17.5 (15) long). Palpomere proportions: femur / genu 1.56 - 1.68 (1.61); tibia / femur 0.72 - 0.81 (0.81); tibia length / width 3.18 - 3.63 (3.53). Venter- (540 - 585 (560) long; 350 - 410 (360) wide) with bold bluish-purple coloration. Gnathosomal bay (110 - 122.5 (112.5) long; 62.5 - 67.5 (62.5) wide). Cxgl- 4 subapical. Medial suture (85 - 115 (85) long). Genital plates (110 - 117.5 (110) long; 87.5 - 92.5 (92.5) wide). Additional measurements: Cx- 1 (240 - 250 (240) long (total); 115 - 127.5 (122.5) long (medial )); Cx- 3 (250 - 290 (265) wide); anterior venter (227.5 - 250 (232.5) long). Ventral proportions: gnathosomal bay length / width 1.69 - 1.85 (1.80); anterior venter / genital field length 1.98 - 2.13 (2.11); anterior venter length / genital field width 2.51 - 2.86 (2.51); anterior venter / medial suture 2.17 - 2.74 (2.74). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
6D98F30878C979E4CBA327D09CC84276.taxon	etymology	Etymology. Specific epithet (tysoni) named in honor of Neil Degrasse Tyson for his efforts in popularizing cosmology and science in general with Cosmos: A Spacetime Odyssey (2014), which was a worthy update to Carl Sagan's Cosmos: A Personal Voyage (1980).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
6D98F30878C979E4CBA327D09CC84276.taxon	distribution	Distribution. Known only from several localities along Natchez Trace Parkway (Figure 258).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
9AD2717BE6A0510B9D37D2D78E2FF3FA.taxon	materials_examined	Material examined. HOLOTYPE (♀): from USA, Mississippi, Attala County, Hurricane Creek (33 ° 4 ' N, 89 ° 32 ' W), 13 Oct 1999, by IM Smith, IMS 990071. PARATYPES (2 ♀; 4 ♂): Alabama, USA: 1 ♀ from Lauderdale County, off Natchez Trace parkway, 7 km south of Tennessee state line (34 ° 56 ' 32 " N, 87 ° 49 ' 43 " W), 24 Sep 2009, by IM Smith, IMS 090122 * 2 ♂ from Lauderdale County, off Natchez Trace parkway, 7 km south of Tennessee state line (34 ° 56 ' 32 " N, 87 ° 49 ' 43 " W), 27 Sep 2010, by IM Smith, IMS 100163 * Mississippi, USA: 1 ♂ (ALLOTYPE) from Attala County, Hurricane Creek (33 ° 4 ' 58 " N, 89 ° 31 ' 31 " W), 30 Sep 2010, by IM Smith, IMS 100168 * 1 ♀ and 1 ♂ from Attala County, Hurricane Creek (33 ° 4 ' 58 " N, 89 ° 31 ' 31 " W), 30 Sep 2010, by IM Smith, IMS 100168.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
9AD2717BE6A0510B9D37D2D78E2FF3FA.taxon	diagnosis	Diagnosis. Torrenticola ululata are similar to other members of the Rusetria " Eastern 2 - Plates " group (T. biscutella, T. caerulea, T. delicatexa, T. indistincta, T. malarkeyorum, T. pendula, T. sellersorum, T. tysoni, T. whitneyae, T. microbiscutella, and T. feminellai) in having anterio-lateral platelets fused to the dorsal plate, and being distributed in the east. T. ululata can be differentiated from most Torrenticola, including other Eastern 2 - Plates, by having a distinct dorsal pattern with a single dark spot posteriorly and an orange spot posterior to the dark spot. The only other species with this pattern is T. unimaculata and T. kringi, which both have anterio-lateral platelets free from the dorsal plate.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
9AD2717BE6A0510B9D37D2D78E2FF3FA.taxon	description	Description. Female (Figure 262) (n = 3) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (540 - 580 (570) long; 400 - 450 (450) wide) circular or occasionally ovoid with coloration restricted to a single dark spot anteriorly, often with an orange spot posterior to the dark spot. Anterio-medial platelets (130 - 137.5 (130) long; 50 - 55 (55) wide). Anterio-lateral platelets (150 - 180 (150) long; 72.5 - 77.5 (72.5) wide) fused to dorsal plate. Dgl- 4 approaching midway between muscle scars and dorsum edge (distance between Dgl- 4 265 - 295 (265 )). Dorsal plate proportions: dorsum length / width 1.27 - 1.41 (1.27); dorsal width / distance between Dgl- 4 1.36 - 1.70 (1.70); anterio-medial platelet length / width 2.36 - 2.70 (2.36); anterio-lateral platelet length / width 2.07 - 2.32 (2.07); anterio-lateral / anterio-medial length 1.15 - 1.31 (1.15). Gnathosoma - Subcapitulum (290 - 305 (290) long (ventral); 216 - 235 (216) long (dorsal); 130 - 140 (130) tall) colorless. Rostrum (125 - 125 (125) long; 42.5 - 45 (42.5) wide). Chelicerae (289 - 317 (289) long) with curved fangs (59 - 64 (64) long). Subcapitular proportions: ventral length / height 2.18 - 2.23 (2.23); rostrum length / width 2.78 - 2.94 (2.94). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (42.5 - 45 (45) long); femur (117.5 - 122.5 (117.5) long); genu (67.5 - 75 (67.5) long); tibia (97.5 - 105 (97.5) long; 21.25 - 25 (21.25) wide); tarsus (17.5 - 20 (17.5) long). Palpomere proportions: femur / genu 1.63 - 1.74 (1.74); tibia / femur 0.83 - 0.86 (0.83); tibia length / width 4.20 - 4.59 (4.59). Venter - (595 - 670 (670) long; 517 - 540 (518) wide) colorless. Gnathosomal bay (152.5 - 160 (152.5) long; 85 - 102.5 (85) wide). Cxgl- 4 subapical. Medial suture (15 - 15 (15) long). Genital plates (155 - 170 (155) long; 147.5 - 157.5 (147.5) wide). Additional measurements: Cx- 1 (276 - 282 (281) long (total); 115 - 126 (120) long (medial )); Cx- 3 (357 - 382 (358) wide); anterior venter 155 - 160 (160) long). Ventral proportions: gnathosomal bay length / width 1.56 - 1.79 (1.79); anterior venter / genital field length 0.93 - 1.03 (1.03); anterior venter length / genital field width 0.98 - 1.08 (1.08); anterior venter / medial suture 10.33 - 10.67 (10.67). Male (Figure 263) (n = 4) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (460 - 510 (500) long; 355 - 380 (380) wide) circular or occasionally ovoid with coloration restricted to a single dark spot anteriorly, often with an orange spot posterior to the dark spot. Anterio-medial platelets (107.5 - 117.5 (110) long; 42.5 - 50 (42.5) wide). Anterio-lateral platelets (145 - 155 (155) long; 55 - 60 (60) wide) fused to dorsal plate. Dgl- 4 approaching midway between muscle scars and dorsum edge (distance between Dgl- 4 235 - 250 (250 )). Dorsal plate proportions: dorsum length / width 1.30 - 1.38 (1.32); dorsal width / distance between Dgl- 4 1.45 - 1.57 (1.52); anterio-medial platelet length / width 2.26 - 2.59 (2.59); anterio-lateral platelet length / width 2.52 - 2.77 (2.58); anterio-lateral / anterio-medial length 1.32 - 1.41 (1.41). Gnathosoma - Subcapitulum (227.5 - 255 (247.5) long (ventral); 165 - 190 (176) long (dorsal); 90 - 100 (100) tall) colorless. Rostrum (90 - 102.5 (100) long; 32.5 - 38.75 (37.5) wide). Chelicerae (225 - 254 (225) long) with curved fangs (34 - 49 (46) long). Subcapitular proportions: ventral length / height 2.48 - 2.55 (2.48); rostrum length / width 2.61 - 2.77 (2.67). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (37.5 - 40 long); femur (92.5 - 100 (100) long); genu (57.5 - 65 (60) long); tibia (77.5 - 92.5 (85) long; 20 - 22.5 (20) wide); tarsus (15 - 17.5 (15) long). Palpomere proportions: femur / genu 1.54 - 1.67 (1.67); tibia / femur 0.84 - 0.93 (0.85); tibia length / width 3.88 - 4.25 (4.25). Venter - (580 - 620 (580) long; 425 - 487 (426) wide) colorless. Gnathosomal bay (110 - 125 (110) long; 67.5 - 75 (67.5) wide). Cxgl- 4 subapical. Medial suture (80 - 100 (100) long). Genital plates (117.5 - 122.5 (120) long; 102.5 - 110 (107.5) wide). Additional measurements: Cx- 1 (246 - 257 (257) long (total); 124 - 129 (125) long (medial )); Cx- 3 (283 - 326 (290) wide); anterior venter (220 - 257.5 (252.5) long). Ventral proportions: gnathosomal bay length / width 1.63 - 1.67 (1.63); anterior venter / genital field length 1.87 - 2.17 (2.10); anterior venter length / genital field width 2.05 - 2.49 (2.35); anterior venter / medial suture 2.53 - 3.19 (2.53). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
9AD2717BE6A0510B9D37D2D78E2FF3FA.taxon	etymology	Etymology. Specific epithet (ululata) refers to the dorsal coloration resembling a wailing mouth, where the dark anterior spot is the oral cavity and the posterior red spot is the tongue (ululatus, L. shriek, wail).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
9AD2717BE6A0510B9D37D2D78E2FF3FA.taxon	distribution	Distribution. Southeastern, Mississippi and Alabama (Figure 261).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
168752C31C3173F94C72299940881BBD.taxon	materials_examined	Material examined. HOLOTYPE (♀): from Canada, New Brunswick, York County, SW Mirimachi River, beside Highway 107 between Napdogan and Juniper, 21 Jun 2012, by IM Smith, IMS 120036, DNA 3010. PARATYPES (7 ♀; 7 ♂): Arkansas, USA: 2 ♀ and 1 ♂ from Montgomery County, Caddo Gap, access track off Manfred Road, 0.3 km west of Route 8, 29 Jul 2011, by IM Smith, IMS 110037 * 4 ♀ from Montgomery County, Gaston, South Fork of Ouachita River, access off County Road 17 at Forest Road 903, 29 Jul 2011, by IM Smith, IMS 110040 * 2 ♂ from Montgomery County, Ouachita River, Pine Ridge, 5 Oct 2007, by AJ Radwell, & HW Robison, AJR 070300 A * 2 ♂ from Montgomery County, Ouachita National Forest, South Fork of Ouachita River, 29 Jul 2011, by AJ Radwell, & B Crump, AJR 110302 * New Brunswick, Canada: 1 ♂ (ALLOTYPE) from York County, SW Mirimachi River, beside Highway 107 between Napdogan and Juniper, 21 Jun 2012, by IM Smith, IMS 120036, DNA 3011 * 1 ♀ and 1 ♂ from York County, SW Mirimachi River, beside Highway 107 between Napdogan and Juniper, 21 Jun 2012, by IM Smith, IMS 120036.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
168752C31C3173F94C72299940881BBD.taxon	diagnosis	Diagnosis. Torrenticola unimaculata are similar to other members of the Tricolor Complex (T. bittikoferae, T. hoosieri, T. larvata, T. pearsoni, T. olliei, T. sierrensis, T. tricolor, T. trimaculata, T. cardia, T. kringi, T. dimorpha, and T. mohawk) in having a short, conical rostrum. T. unimaculata can be differentiated from most Torrenticola, including other members of the Tricolor Complex, by having a distinct dorsal pattern of a large anterior dorsal spot. The only other species with this pattern is T. ululata, which, like all Rusetria 2 - Plates, have anterio-lateral platelets fused to the dorsal plate and T. kringi, which has a more elongate rostrum (length / width = 1.9 - 2.2 in T. unimaculata, 2.6 - 3.2 in T. kringi). T. unimaculata are most similar to other members of the Tricolor Complex that have bold patterning (T. larvata, T. tricolor, T. trimaculata, T. cardia, T. kringi, and T. mohawk). Female T. unimaculata can be further differentiated from these members of the complex by having a longer medial suture (♀ = 40 - 47.5 in T. unimaculata, 15 - 35 in others), except T. cardia (25 - 50) and T. mohawk (30 - 45).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
168752C31C3173F94C72299940881BBD.taxon	description	Description. Female (Figure 265) (n = 6) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (650 - 730 (720) long; 490 - 600 (600) wide) ovoid with coloration restricted to a single dark spot anteriorly (occasionally extending medially), with an orange spot posterior to the dark spot. Anterio-medial platelets (127.5 - 145 (140) long; 62.5 - 70 (67.5) wide). Anterio-lateral platelets (172.5 - 200 (200) long; 72.5 - 80 (80) wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 350 - 425 (425 )). Dorsal plate proportions: dorsum length / width 1.20 - 1.39 (1.20); dorsal width / distance between Dgl- 4 1.34 - 1.41 (1.41); anterio-medial platelet length / width 1.89 - 2.08 (2.07); anterio-lateral platelet length / width 2.29 - 2.57 (2.50); anterio-lateral / anterio-medial length 1.35 - 1.44 (1.43). Gnathosoma - Subcapitulum (242.5 - 265 (265) long (ventral); 176.25 - 194 (190) long (dorsal); 110 - 125 (125) tall) colorless. Rostrum (85 - 100 (92.5) long; 42.5 - 47.5 (47.5) wide) short and conical. Chelicerae (236 - 252 (250) long) with curved fangs (51 - 61.5 (55) long). Subcapitular proportions: ventral length / height 2.12 - 2.26 (2.12); rostrum length / width 1.89 - 2.18 (1.95). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (35 - 42.5 (42.5) long); femur (91.25 - 100 (100) long); genu (62.5 - 67.5 (67.5) long); tibia (80 - 87.5 (87.5) long; 25 - 26.25 (26.25) wide); tarsus (20 - 25 (22.5) long). Palpomere proportions: femur / genu 1.46 - 1.52 (1.48); tibia / femur 0.81 - 0.93 (0.88); tibia length / width 3.20 - 3.40 (3.33). Venter - (700 - 860 (860) long; 539 - 630 (630) wide) colorless. Gnathosomal bay (110 - 140 (140) long; 75 - 87.5 (80) wide). Cxgl- 4 subapical. Medial suture (40 - 47.5 (40) long). Genital plates (180 - 210 (210) long; 152.5 - 170 (170) wide). Additional measurements: Cx- 1 (243 - 290 (290) long (total); 128 - 162 (145) long (medial )); Cx- 3 (358 - 426 (390) wide); anterior venter (182.5 - 205 (195) long). Ventral proportions: gnathosomal bay length / width 1.40 - 1.75 (1.75); anterior venter / genital field length 0.93 - 1.14 (0.93); anterior venter length / genital field width 1.15 - 1.30 (1.15); anterior venter / medial suture 4.06 - 4.88 (4.88). Male (Figure 266) (n = 6) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum- (525 - 690 (690) long; 400 - 520 (520) wide) ovoid with coloration restricted to a single dark spot anteriorly (occasionally extending medially), with an orange spot posterior to the dark spot. Anterio-medial platelets (112.5 - 137.5 (137.5) long; 57.5 - 72.5 (72.5) wide). Anterio-lateral platelets (160 - 205 (205) long; 65 - 82.5 (82.5) wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 320 - 405 (405 )). Dorsal plate proportions: dorsum length / width 1.30 - 1.43 (1.33); dorsal width / distance between Dgl- 4 1.23 - 1.28 (1.28); anterio-medial platelet length / width 1.90 - 2.09 (1.90); anterio-lateral platelet length / width 2.28 - 2.54 (2.48); anterio-lateral / anterio-medial length 1.35 - 1.43 (1.33). Gnathosoma - Subcapitulum (212.5 - 255 (255) long (ventral); 155 - 185 (185) long (dorsal); 90 - 110 (110) tall) colorless. Rostrum (80 - 95 (95) long; 37.5 - 48.75 (48.75) wide) short and conical. Chelicerae (195 - 235 (235) long) with curved fangs (42 - 55 (55) long). Subcapitular proportions: ventral length / height 2.32 - 2.42 (2.32); rostrum length / width 1.95 - 2.2 (1.95). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (32.5 - 40 (40) long); femur (80 - 97.5 (97.5) long); genu (55 - 67.5 (67.5) long); tibia (70 - 80 (80) long; 22.5 - 27.5 (27.5) wide); tarsus (20 - 22.5 (22.5) long). Palpomere proportions: femur / genu 1.39 - 1.48 (1.44); tibia / femur 0.82 - 0.91 (0.82); tibia length / width 2.90 - 3.11 (2.91). Venter - (640 - 800 (800) long; 448 - 570 (570) wide) colorless. Gnathosomal bay (97.5 - 132.5 (132.5) long; 62.5 - 77.5 (72.5) wide). Cxgl- 4 subapical. Medial suture (107.5 - 125 (125) long). Genital plates (130 - 161.25 (161.25) long; 92.5 - 112.5 (112.5) wide). Additional measurements: Cx- 1 (218 - 338 (290) long (total); 109 - 165 (165) long (medial )); Cx- 3 (218 - 338 (290) wide); anterior venter (265 - 300 (300) long). Ventral proportions: gnathosomal bay length / width 1.44 - 1.83 (1.83); anterior venter / genital field length 1.86 - 2.13 (1.86); anterior venter length / genital field width 2.67 - 2.92 (2.67); anterior venter / medial suture 2.30 - 2.49 (2.40). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
168752C31C3173F94C72299940881BBD.taxon	etymology	Etymology. Specific epithet (unimaculata) named for the single dark dorsal spot of this species (unus, L. one; macula, L. spot).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
168752C31C3173F94C72299940881BBD.taxon	distribution	Distribution. Known only from Arkansas and New Brunswick (Figure 264).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
30FC6CADDB0A532821ACECCC8922F976.taxon	materials_examined	Material examined. LECTOTYPE (1 ♀): from USA, California, Ventura County, Upper Ojai, Sisar Canyon, 4 Nov 1973, by H Habeeb, HH 730009. PARALECTOTYPES (1 ♀; 2 ♂): California, USA: 1 ♂ (ALLOTYPE) from Ventura County, Upper Ojai, Sisar Canyon, 4 Nov 1973, by H Habeeb, HH 730009 * 1 ♀ and 1 ♂ from Ventura County, Upper Ojai, Sisar Canyon, 4 Nov 1973, by H Habeeb, HH 730009. OTHER MATERIAL (30 ♀; 37 ♂): California, USA: 1 ♀ and 1 ♂ from Alpine County, Markleeville Creek (38 ° 41 ' 39 " N, 119 ° 46 ' 41 " W), 30 Aug 2013, by JR Fisher, JRF 13 - 0830 - 001 * 3 ♀ and 3 ♂ from Humboldt County, Honeydew, Mattole River, beside road to Bull Creek on east side of bridge, 8 Aug 1987, by IM Smith, IMS 870135 A * 1 ♀ and 1 ♂ from Los Angeles County, Azusa, San Gabriel Canyon, East Fork of San Gabriel River at East Fork Station, 24 Jul 1987, by IM Smith, IMS 870108 * 1 ♀ and 1 ♂ from Mariposa County, El Portal, Indian Flat campground, Merced River, 9 - 10 Jun 1976, by IM Smith, IMS 760087 * 2 ♀ and 3 ♂ from Mendocino County, Paul M. Dimmick Recreation Area, North Fork of Navarro River, beside Route 128, 4 Aug 1987, by IM Smith, IMS 870127 A * 3 ♀ and 3 ♂ from Mendocino County, Rancheria Creek, beside Route 128, 7.3 kilometers south of Boonville, 4 Aug 1987, by IM Smith, IMS 870126 A * 2 ♀ and 2 ♂ from Monterey County, Big Sur River, beside Route 1 near Pfeiffer-Big Sur State Park, 28 - 29 Jul 1987, by IM Smith, IMS 870116 A * 1 ♀ from Plumas County, Plumas National Forest, Silver Creek (39 ° 56 ' 60 " N, 121 ° 2 ' 17 " W), 24 Aug 2013, by JR Fisher, JRF 13 - 0824 - 005 * 2 ♀ and 4 ♂ from Trinity County, Shasta-Trinity National Forest, Wilson Creek (40 ° 25 ' 17 " N, 123 ° 3 ' 5 " W), 20 Aug 2013, by JR Fisher, JRF 13 - 0820 - 003 * 1 ♂ from Trinity County, small cascading trickle beside Route 36, 5.2 kilometers west of Forest Glen Station, 6 Aug 1987, by IM Smith, IMS 870132 * 2 ♀ and 3 ♂ from Trinity County, South Fork of Trinity River, beside Route 36 at Forest Glen campground, 6 Aug 1987, by IM Smith, IMS 870131 * 2 ♀ and 4 ♂ from Trinity County, Trinity River, beside Route 299, 8.7 kilometers northwest of Del Loma, 9 Aug 1987, by IM Smith, IMS 870137 A * 1 ♀ and 1 ♂ from Trinity County, Weaver Creek, beside Route 299, 4.3 kilometers north of Route 3 West, 9 Aug 1987, by IM Smith, IMS 870138 A & IMS 870138 B * 1 ♀ and 4 ♂ from Ventura County, Ojai, North Fork of Ventura River, beside Route 33 just above Wheeler Gorge, 25 - 26 Jul 1987, by IM Smith, IMS 870109 A & IMS 870109 B * 1 ♀ and 1 ♂ from Ventura County, Ojai, North Fork of Ventura River, beside Route 33, 9.8 kilometers north of Ojai, 25 Jul 1987, by IM Smith, IMS 870110 * 1 ♀ and 1 ♂ from Ventura County, Ojai, North Fork of Ventura River just below Wheeler Gorge campground, beside Route 33, 27 Jul 1987, by IM Smith, IMS 870112 * 1 ♀ and 1 ♂ from Ventura County, Sespe Creek at Middle Lion campground, off Road 6 N 31, 26 Jul 1987, by IM Smith, IMS 870111 A * 1 ♀ from Yuba County, Tahoe National Forest, Oregon Creek (39 ° 23 ' 50 " N, 121 ° 4 ' 54 " W), 25 Aug 2013, by JR Fisher, JRF 13 - 0825 - 006 * Oregon, USA: 1 ♀ and 1 ♂ from Coos County, Gaylord, Coquille Myrtle Grove State Park, Coquille River, 2 Jul 1983, by IM Smith, IMS 830014 * 1 ♀ from Curry County, Quosatana Creek (42 ° 29 ' 21 " N, 124 ° 14 ' 2 " W), 14 Aug 2013, JR Fisher, JRF 13 - 0814 - 003 * 2 ♀ and 2 ♂ from Curry County, Sixes, Sixes River, beside road at mouth of Edson Creek, 4 Jul 1983, by IM Smith, IMS 830021 A.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
30FC6CADDB0A532821ACECCC8922F976.taxon	diagnosis	Diagnosis. Torrenticola ventura is unlike all other western species by having the following combination of characters: anterio-lateral platelets free (fused to dorsal plate in Rusetria Complex); dorsal coloration separated into anterior and posterior portions (restricted posteriorly in T. tahoei, T. oregonensis, T. raptoroides, and T. sharkeyi, without pattern in T. regalis, colorless in T. occidentalis, Rala Group, T. wiedenmanni); ellipsoid body (circular in T. sierrensis and T. raptoroides, rectangular in T. ellipsoidalis); unmodified rostrum (short and conical in T. olliei, T. sierrensis, T. ellipsoidalis and T. leviathan); tuberculate ventral extensions on pedipalp genua (flanged in Miniforma group); and smaller body size than T. multiforma (dorsum length ♀ = 650 - 780 in T. ventura, 765 - 885 in T. multiforma; ♂ = 540 - 630 in T. ventura, 725 - 850 in T. multiforma). T. ventura are most similar to members of the Neoanomala Group, which are eastern.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
30FC6CADDB0A532821ACECCC8922F976.taxon	description	Re-description. Female (Figure 268) (n = 7) with characters of the genus with following specifications. Dorsum - (650 - 780 long; 470 - 550 wide) ovoid with bluish-purple coloration (often faint) separated into anterior and posterior portions with orange medially. Anterio-medial platelets (145 - 152.5 long; 57.5 - 62.5 wide). Anterio-lateral platelets (195 - 227.5 long; 67.5 - 80 wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 355 - 400). Dorsal plate proportions: dorsum length / width 1.34 - 1.46; dorsal width / distance between Dgl- 4 1.29 - 1.42; anterio-medial platelet length / width 2.32 - 2.65; anterio-lateral platelet length / width 2.70 - 3.03; anterio-lateral / anterio-medial length 1.28 - 1.57. Gnathosoma - Subcapitulum (350 - 397.5 long (ventral); 267 - 297 long (dorsal); 147.5 - 167.5 tall) colorless. Rostrum (137.5 - 155 long; 50 - 60 wide). Chelicerae (375 - 415 long) with curved fangs (63 - 80 long). Subcapitular proportions: ventral length / height 2.30 - 2.47; rostrum length / width 2.58 - 2.95. Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (47.5 - 60 long); femur (125 - 143.75 long); genu (75 - 82.5 long); tibia (92.5 - 102.5 long; 25 - 28.75 wide); tarsus (17.5 - 25 long). Palpomere proportions: femur / genu 1.67 - 1.80; tibia / femur 0.71 - 0.75; tibia length / width 3.36 - 3.73. Venter - (740 - 910 long; 550 - 667 wide) often with faint bluish-purple coloration. Gnathosomal bay (175 - 202.5 long; 87.5 - 122.5 wide). Cxgl- 4 subapical. Medial suture (22.5 - 30 long). Genital plates (170 - 200 long; 167.5 - 182.5 wide). Additional measurements: Cx- 1 (322 - 365 long (total); 137 - 173 long (medial )); Cx- 3 (22.5 - 30 wide); anterior venter 197.5 - 220 long). Ventral proportions: gnathosomal bay length / width 1.49 - 2.25; anterior venter / genital field length 1.05 - 1.16; anterior venter length / genital field width 1.08 - 1.22; anterior venter / medial suture 7.18 - 8.78. Male (Figure 269) (n = 7) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (540 - 630 long; 370 - 430 wide) ovoid with bluish-purple coloration (often faint) separated into anterior and posterior portions with orange medially. Anterio-medial platelets (105 - 135 long; 47.5 - 55 wide). Anterio-lateral platelets (172.5 - 197.5 long; 60 - 62.5 wide) free from dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 280 - 315). Dorsal plate proportions: dorsum length / width 1.43 - 1.53; dorsal width / distance between Dgl- 4 1.30 - 1.38; anterio-medial platelet length / width 2.21 - 2.63; anterio-lateral platelet length / width 2.76 - 3.16; anterio-lateral / anterio-medial length 1.40 - 1.64. Gnathosoma - Subcapitulum (300 - 332.5 long (ventral); 221.05 - 247.5 long (dorsal); 106.25 - 122.5 tall) colorless. Rostrum (120 - 135 long; 42.5 - 47.5 wide). Chelicerae (289 - 340 long) with curved fangs (56 - 65 long). Subcapitular proportions: ventral length / height 2.65 - 2.93; rostrum length / width 2.67 - 3.06. Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (40 - 45 long); femur (106.25 - 117.5 long); genu (65 - 72.5 long); tibia (81.25 - 91.25 long; 23.75 - 27.5 wide); tarsus (15 - 21.25 long). Palpomere proportions: femur / genu 1.62 - 1.70; tibia / femur 0.74 - 0.81; tibia length / width 3.32 - 3.42. Venter - (650 - 785 long; 436 - 500 wide) often with faint bluish-purple coloration. Gnathosomal bay (122.5 - 162.5 long; 78.75 - 85 wide). Cxgl- 4 subapical. Medial suture (97.5 - 125 long). Genital plates (122.5 - 142.5 long; 100 - 112.5 wide). Additional measurements: Cx- 1 (263.5 - 325 long (total); 155 - 164 long (medial )); Cx- 3 (319 - 357.5 wide); anterior venter (265 - 305 long). Ventral proportions: gnathosomal bay length / width 1.53 - 1.97; anterior venter / genital field length 2.02 - 2.25; anterior venter length / genital field width 2.55 - 2.81; anterior venter / medial suture 2.36 - 2.72. Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
30FC6CADDB0A532821ACECCC8922F976.taxon	etymology	Etymology. Although Habeeb (1973) did not specify an etymology for the specific epithet (ventura), it surely refers to the type locality-Ventura River in Ventura County, California.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
30FC6CADDB0A532821ACECCC8922F976.taxon	distribution	Distribution. California and southwest Oregon (Figure 267). T. ventura was previously known only from Ventura County in southwestern California; we extend the range northward.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
5940735C225768EBAA430E10F1F1140A.taxon	materials_examined	Material examined. HOLOTYPE (♀): from Canada, British Columbia, Ryan Rest Area off Hwy 3, East of Yahk Moyie River, 15 Aug 2012, by IM Smith, IMS 120071, DNA 2955. PARATYPES (12 ♀; 4 ♂): California, USA: 1 ♀ and 1 ♂ from El Dorado County, Upper Truckee River (38 ° 50 ' 56 " N, 120 ° 1 ' 39 " W), 29 Aug 2013, by JR Fisher, JRF 13 - 0829 - 003 * 2 ♀ and 2 ♂ El Dorado County, Upper Truckee River (38 ° 50 ' 56 " N, 120 ° 1 ' 39 " W), 29 Aug 2013, by JR Fisher, JRF 13 - 0829 - 004 * 2 ♀ from Mono County, Humboldt-Toiyabe National Forest, West Walker River (38 ° 21 ' 59 " N, 119 ° 28 ' 55 " W), 31 Aug 2013, by JR Fisher, JRF 13 - 0831 - 003 * 2 ♀ from Nevada County, Tahoe National Forest, Sagehen Creek (39 ° 26 ' 2 " N, 120 ° 12 ' 17 " W), 26 Aug 2013, by JR Fisher, JRF 13 - 0826 - 006 * Oregon, USA: 1 ♂ (ALLOTYPE) from Douglas County, Umpqua NF, Umpqua River (43 ° 17 ' 28 " N, 122 ° 37 ' 12 " W), 12 Aug 2013, by JC O'Neill, & WA Nelson, JNOW 13 - 0812 - 006 * 1 ♀ from Coos County, Middle Fork of Coquille River (43 ° 1 ' 56 " N, 124 ° 6 ' 1 " W), 12 Aug 2013, by JR Fisher, JRF 13 - 0812 - 001 * 2 ♀ from Douglas County, Umpqua NF, Umpqua River (43 ° 17 ' 28 " N, 122 ° 37 ' 12 " W), 12 Aug 2013, by JC O'Neill, & WA Nelson, JNOW 13 - 0812 - 006 * 2 ♀ from Lane County, Gate Creek (44 ° 8 ' 48 " N, 122 ° 34 ' 20 " W), 11 Aug 2013, by JC O'Neill, & WA Nelson, JNOW 13 - 0811 - 001.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
5940735C225768EBAA430E10F1F1140A.taxon	diagnosis	Diagnosis. Torrenticola walteri are similar to other members of the Rusetria " Western 2 - Plates " group (T. mulleni, T. nortoni, and T. welbourni) in having anterio-lateral platelets fused to the dorsal plate, having faint dorsal coloration, and being distributed in the west. Female T. walteri can be differentiated from T. welbourni (female only known) by having stockier pedipalp tibiae (3.09 - 3.23 in A 32, 3.73 in A 30); shorter pedipalp femora (112.5 - 125 in A 32, 137.5 in A 30); and a more elongate subcapitulum (ventral length / height: 2.21 - 2.34 in A 32, 2.47 in A 30). T. walteri can be differentiated from T. mulleni by having a slightly stockier gnathosomal bay (♀ = 1.57 - 1.84 in T. walteri, 1.89 - 2.16 in T. mulleni, ♂ = 1.55 - 1.73 in T. walteri, 1.77 - 1.93 in T. mulleni) and by being distributed in California, Oregon and British Columbia, instead of in the Rocky Mountains (Idaho, Montana, Utah and Wyoming). Additionally, male T. walteri can be differentiated from male T. mulleni by having a shorter genital field (115 - 117.5 in A 32, 130 - 140 in A 31), and female T. walteri can be differentiated from female T. mulleni by having a shorter medial suture (10 - 12.5 in T. walteri, 20 - 22.5 in T. mulleni). Female T. walteri can be differentiated from female T. nortoni by having slightly shorter pedipalp femora with respect to genua (1.52 - 1.64 in T. walteri, 1.69 - 1.82 in T. nortoni) and slightly stockier anterio-medial platelets (2.58 - 2.72 in T. walteri, 2.74 - 3.06 in T. nortoni). Male T. walteri can be differentiated from male T. nortoni by having longer pedipalp femora (95 - 100 in T. walteri, 85 - 92.5 in T. nortoni) and slightly more elongate pedipalp tibiae (3.05 - 3.10 in T. walteri, 2.73 - 3.0 in T. nortoni).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
5940735C225768EBAA430E10F1F1140A.taxon	description	Description. Female (Figure 271) (n = 5) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (580 - 640 (590) long; 420 - 450 (420) wide) ovoid with orange coloration separated into anterior and posterior portions, occasionally faint. Anterio-medial platelets (120 - 137.5 (120) long; 45 - 51.25 (45) wide). Anterio-lateral platelets (157.5 - 190 (157.5) long; 55 - 65 (55) wide) fused with dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 300 - 325 (300 )). Dorsal plate proportions: dorsum length / width 1.35 - 1.44 (1.40); dorsal width / distance between Dgl- 4 1.37 - 1.45 (1.40); anterio-medial platelet length / width 2.58 - 2.72 (2.67); anterio-lateral platelet length / width 2.63 - 3.27 (2.86); anterio-lateral / anterio-medial length 1.29 - 1.46 (1.31). Gnathosoma - Subcapitulum (305 - 335 (317.5) long (ventral); 223 - 243 (224) long (dorsal); 135 - 147.5 (137.5) tall) colorless. Rostrum (122.5 - 127.5 (123.75) long; 45 - 50 (47.5) wide). Chelicerae (304 - 328 (304) long) with curved fangs (55.75 - 64 (63) long). Subcapitular proportions: ventral length / height 2.21 - 2.34 (2.31); rostrum length / width 2.45 - 2.72 (2.61). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (47.5 - 52.5 (50) long); femur (112.5 - 125 (115) long); genu (70 - 82.5 (70) long); tibia (85 - 88.75 (88.75) long; 27.5 - 27.5 (27.5) wide); tarsus (17.5 - 20 (20) long). Palpomere proportions: femur / genu 1.52 - 1.64 (1.64); tibia / femur 0.70 - 0.77 (0.77); tibia length / width 3.09 - 3.23 (3.23). Venter - (690 - 730 (725) long; 467 - 520 (467) wide) colorless. Gnathosomal bay (141.25 - 175 (157.5) long; 86.25 - 95 (86.25) wide). Cxgl- 4 subapical. Medial suture (10 - 12.5 (10) long). Genital plates (185 - 197.5 (197.5) long; 158.75 - 181.25 (180) wide). Additional measurements: Cx- 1 (250 - 300 (285) long (total); 102 - 127 (119) long (medial )); Cx- 3 (313 - 362 (313) wide); anterior venter (140 - 158.75 (150) long). Ventral proportions: gnathosomal bay length / width 1.57 - 1.84 (1.83); anterior venter / genital field length 0.75 - 0.86 (0.76); anterior venter length / genital field width 0.83 - 0.96 (0.83); anterior venter / medial suture 11.20 - 15.88 (15.00). Male (Figure 272) (n = 4) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (490 - 510 (510) long; 340 - 350 (350) wide) ovoid with orange coloration separated into anterior and posterior portions, occasionally faint. Anterio-medial platelets (105 - 115 (115) long; 40 - 42.5 (42.5) wide). Anterio-lateral platelets (125 - 142.5 (142.5) long; 47.5 - 52.5 (52.5) wide) fused with dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 255 - 270 (270 )). Dorsal plate proportions: dorsum length / width 1.43 - 1.46 (1.46); dorsal width / distance between Dgl- 4 1.30 - 1.33 (1.30); anterio-medial platelet length / width 2.63 - 2.71 (2.71); anterio-lateral platelet length / width 2.63 - 2.71 (2.71); anterio-lateral / anterio-medial length 1.19 - 1.36 (1.24). Gnathosoma - Subcapitulum (265 - 270 (270) long (ventral); 179 - 202.5 (195) long (dorsal); 93.75 - 105 (95) tall) colorless. Rostrum (100 - 102.5 (102.5) long; 37.5 - 38.75 (37.5) wide). Chelicerae (240 - 265 (265) long) with curved fangs (38 - 50 (50) long). Subcapitular proportions: ventral length / height 2.52 - 2.85 (2.84); rostrum length / width 2.65 - 2.73 (2.73). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (35 - 40 (35) long); femur (95 - 100 (100) long); genu (57.5 - 60 (57.5) long); tibia (72.5 - 77.5 (72.5) long; 23.75 - 25 (23.75) wide); tarsus (17.5 - 20 (17.5) long). Palpomere proportions: femur / genu 1.65 - 1.74 (1.74); tibia / femur 0.73 - 0.82 (0.73); tibia length / width 3.05 - 3.10 (3.05). Venter - (590 - 625 (625) long; 387.5 - 410 (410) wide) colorless. Gnathosomal bay (112.5 - 127.5 (127.5) long; 72.5 - 78.75 (73.75) wide). Cxgl- 4 subapical. Medial suture (85 - 107.5 (107.5) long). Genital plates (115 - 117.5 (116.25) long; 95 - 98.75 (95) wide). Additional measurements: Cx- 1 (255 - 273 (270) long (total); 136 - 157 (150) long (medial )); Cx- 3 (253 - 316 (285) wide); anterior venter (245 - 262.5 (262.5) long). Ventral proportions: gnathosomal bay length / width 1.55 - 1.73 (1.73); anterior venter / genital field length 2.12 - 2.26 (2.26); anterior venter length / genital field width 2.48 - 2.76 (2.76); anterior venter / medial suture 2.44 - 2.88 (2.44). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
5940735C225768EBAA430E10F1F1140A.taxon	etymology	Etymology. Specific epithet (walteri) named in honor of acarologist Dave Walter for solidifying JRF's interest in mites with his popular book on mites (Mites: Ecology, Evolution & Behaviour - Life at a Microscale) and by teaching JRF the mesostigmatan mite section of the Acarology Summer Program at The Ohio State University in 2009.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
5940735C225768EBAA430E10F1F1140A.taxon	distribution	Distribution. Probably throughout the Pacific Coastal Ranges of California, Oregon, Washington, and southern British Columbia (Figure 270). We also collected T. walteri in the Rocky Mountains of British Columbia, indicating this species might occur in the northern Rockies of Canada. However, given our sampling effort in the Rockies, we doubt the occurrence of this species in most of the US Rockies.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
696CA3E68C0B6764D210754A81AC8C09.taxon	materials_examined	Material examined. HOLOTYPE (♀): from USA, California, Trinity County, Shasta-Trinity National Forest, Wilson Creek (40 ° 25 ' 17 " N, 123 ° 3 ' 5 " W), 20 Aug 2013, by JR Fisher, JRF 13 - 0820 - 003, DNA 1638.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
696CA3E68C0B6764D210754A81AC8C09.taxon	diagnosis	Diagnosis. Torrenticola welbourni are similar to other members of the Rusetria " Western 2 - Plates " group (T. mulleni, T. nortoni, and T. walteri) in having anterio-lateral platelets fused to the dorsal plate, having faint dorsal coloration, and being distributed in the west. T. welbourni (only female known) can be differentiated from all other Western 2 - plates by being larger (dorsal length: 690 in T. welbourni, 570 - 645 in others; dorsal width: 500 in T. welbourni, 415 - 480 in others), having more elongate pedipalp tibiae (length / width: 3.73 in T. welbourni, 3.0 - 3.33 in others) and longer pedipalp femora (137.5 in A 30, 112.5 - 125 in others).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
696CA3E68C0B6764D210754A81AC8C09.taxon	description	Description. Female (Figure 274) (n = 1) (holotype only) with characters of the genus with following specifications. Dorsum - (690 long; 500 wide) ovoid with faint orange coloration separated into anterior and posterior portions. Anterio-medial platelets (152.5 long; 57.5 wide). Anterio-lateral platelets (195 long; 72.5 wide) fused to dorsal plate. Dgl- 4 closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 / dorsal width 325). Dorsal plate proportions: dorsum length / width 1.38; dorsum width / distance between Dgl- 4 1.54; anterio-medial platelet length / width 2.65; anterio-lateral platelet length / width 2.69; anterio-lateral / anterio-medial length 1.28. Gnathosoma - Subcapitulum (357.5 long (ventral); 270 long (dorsal); 145 tall) colorless. Rostrum (145 long; 52.5 wide). Chelicerae (356 long) with curved fangs (70 long). Subcapitular proportions: ventral length / height 2.47; rostrum length / width 2.76. Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (50 long); femur (137.5 long); genu (75 long); tibia (102.5 long; 27.5 wide); tarsus (20 long). Palpomere proportions: femur / genu 1.83; tibia / femur 0.75; tibia length / width 3.73. Venter - (820 long; 580.25 wide) colorless. Gnathosomal bay (180 long; 100 wide). Cxgl- 4 subapical. Medial suture (12.5 long). Genital plates (180 long; 167.5 wide). Additional measurements: Cx- 1 (324 long (total); 122 long (medial )); Cx- 3 (371 wide); anterior venter (177.5 long). Ventral proportions: gnathosomal bay length / width 1.80; anterior venter / genital field length 0.99; anterior venter length / genital field width 1.06; anterior venter / medial suture 14.20. Male unknown. Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
696CA3E68C0B6764D210754A81AC8C09.taxon	etymology	Etymology. Specific epithet (welbourni) named in honor of acarologist Cal Welbourn, who has been instrumental in teaching terrestrial Parasitengona to JRF and for teaching JRF the prostigmatan mite section of the Acarology Summer Program at The Ohio State University in 2009.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
696CA3E68C0B6764D210754A81AC8C09.taxon	distribution	Distribution. Only known from Trinity County, California (Figure 273).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
EB1E1CB7A89BA2DDB9255A631ED5F87A.taxon	materials_examined	Material examined. HOLOTYPE (♀): from USA, North Carolina, Yancey County, Lost Cove Picnic Area beside Forest Road 472, 2.8 km west of Rt. 80, (35 ° 45 ' 45 " N, 82 ° 12 ' 12 " W), 12 September 2005, by IM Smith, IMS 050074 PARATYPES (4 ♀; 3 ♂): North Carolina, USA: 1 ♀ and 1 ♂ from Haywood County, Great Smoky Mountains National Park; Cataloochee; beside Mt. Sterling Rd. near bridge 1.7 km n. of road to Campground, (35 ° 38 ' 38 " N, 83 ° 4 ' 4 " W), 6 September 2009, by IM Smith, IMS 090099 * 2 ♀ from Swain County, Great Smokey Mountain National Park, Deep Creek upstream of picnic area, (35 ° 27 ' 27 " N, 83 ° 26 ' 26 " W), 14 September 2009, by AJ Radwell, AJR 090007 A * 1 ♂ (ALLOTYPE) from Yancey County, Lost Cove Picnic Area beside Forest Road 472, 2.8 km west of Rt. 80, (35 ° 45 ' 45 " N, 82 ° 12 ' 12 " W), 12 September 2005, by IM Smith, IMS 050074 * 1 ♀ and 1 ♂ from Yancey County, Lost Cove Picnic Area beside Forest Road 472, 2.8 km west of Rt. 80, (35 ° 45 ' 45 " N, 82 ° 12 ' 12 " W), 12 September 2005, by IM Smith, IMS 050074.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
EB1E1CB7A89BA2DDB9255A631ED5F87A.taxon	diagnosis	Diagnosis. Torrenticola whitneyae are similar to other members of the Rusetria " Eastern 2 - Plates " group (T. biscutella, T. caerulea, T. delicatexa, T. indistincta, T. malarkeyorum, T. pendula, T. sellersorum, T. tysoni, T. ululata, T. microbiscutella, and T. feminellai) in having anterio-lateral platelets fused to the dorsal plate, having dorsal coloration separated into anterior and posterior portions (except T. ululata and T. indistincta), and being distributed in the east. T. whitneyae is most similar to T. pendula, which also has purple coloration separated into anterior and posterior portions and often connected by a stripe medially. T. whitneyae is best differentiated from T. pendula by having a stockier rostrum (length / width = 2.41 - 2.69 in T. whitneyae, 2.87 - 3.06 in T. pendula). T. whitneyae can be differentiated from T. ululata, T. indistincta, and T. feminellai by dorsal coloration and pattern. T. whitneyae can be differentiated from all other Eastern 2 - Plates by having stockier pedipalpal tibiae (♀ = 2.42 - 2.95 in T. whitneyae, 3.00 - 4.59 in others; ♂ = 2.48 - 2.70 in T. whitneyae, 2.78 - 4.25 in others), except female T. delicatexa (♀ = 2.92 - 3.61). T. whitneyae can be differentiated from T. delicatexa by having a slightly rounder dorsum (length / width ♀ = 1.26 - 1.38 in T. whitneyae, 1.38 - 1.44 in T. delicatexa; ♂ = 1.35 - 1.37 in T. whitneyae, 1.44 - 1.56 in T. delicatexa) and by dorsal coloration.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
EB1E1CB7A89BA2DDB9255A631ED5F87A.taxon	description	Description. Female (Figure 276) (n = 5) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (550 - 690 (630) long; 400 - 520 (500) wide) ovoid with reddish-purple coloration both anteriorly and posteriorly connected medially. Anterio-medial platelets (120 - 163.75 (142.5) long; 41.25 - 52.5 (52.5) wide). Anterio-lateral platelets (170 - 220 (200 )) long; 65 - 85 (85) wide) partially fused, at least posteriorly, to dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 310 - 385 (370 )). Dorsal plate proportions: dorsum length / width 1.26 - 1.38 (1.26); dorsal width / distance between Dgl- 4 1.29 - 1.37 (1.35); anterio-medial platelet length / width 2.71 - 3.12 (2.71); anterio-lateral platelet length / width 2.35 - 2.84 (2.35); anterio-lateral / anterio-medial length 1.32 - 1.42 (1.40). Gnathosoma - Subcapitulum (310 - 385 (362.5) long (ventral); 237.5 - 285 (270) long (dorsal); 145 - 185 (177.5) tall) colorless. Rostrum (125 - 140 (135) long; 47.5 - 55 (55) wide). Chelicerae (310 - 415 (395) long) with curved fangs (65 - 80 (75) long). Subcapitular proportions: ventral length / height 1.95 - 2.14 (2.04); rostrum length / width 2.41 - 2.63 (2.45). Pedipalps stocky with short, tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (50 - 56.25 (55) long); femur (67.5 - 80 (75) long); genu (67.5 - 80 (75) long); tibia (72.5 - 80 (80) long; 26.25 - 30 (28.75) wide); tarsus (17.5 - 20 (17.5) long). Palpomere proportions: femur / genu 1.59 - 1.71 (1.63); tibia / femur 0.59 - 0.68 (0.65); tibia length / width 2.42 - 2.95 (2.78). Venter - (630 - 810 (780) long; 470 - 630 (590) wide) with faint reddish-purple coloration. Gnathosomal bay (175 - 222.5 (210) long; 80 - 105 (105) wide). Cxgl- 4 subapical. Medial suture absent. Genital plates (185 - 195 (187.5) long; 165 - 195 (180) wide). Additional measurements: Cx- 1 (280 - 360 (350) long (total); 110 - 140 (135) long (medial )); Cx- 3 (320 - 435 (405) wide); anterior venter (110 - 145 (140) long). Ventral proportions: gnathosomal bay length / width 2.00 - 2.59 (2.00); anterior venter / genital field length 0.59 - 0.75 (0.75); anterior venter length / genital field width 0.67 - 0.80 (0.78). Male (Figure 277) (n = 3) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (540 - 560 (540) long; 395 - 410 (395) wide) ovoid with reddish-purple coloration both anteriorly and posteriorly connected medially. Anterio-medial platelets (122.5 - 135 (135) long; 45 - 50 (50) wide). Anterio-lateral platelets (170 - 170 (170) long; 60 - 67.5 (60) wide) partially fused, at least posteriorly, to dorsal plate. Dgl- 4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl- 4 290 - 310 (295 )). Dorsal plate proportions: dorsum length / width 1.35 - 1.37 (1.37); dorsal width / distance between Dgl- 4 1.32 - 1.38 (1.34); anterio-medial platelet length / width 2.55 - 2.72 (2.70); anterio-lateral platelet length / width 2.52 - 2.83 (2.83); anterio-lateral / anterio-medial length 1.26 - 1.39 (1.26). Gnathosoma - Subcapitulum (290 - 305 (300) long (ventral); 220 - 225 (222.5) long (dorsal); 120 - 135 (135) tall) colorless. Rostrum (107.5 - 112.5 (112.5) long; 40 - 42.5 (42.5) wide). Chelicerae (310 - 310 (310) long) with curved fangs (60 - 60 (60) long). Subcapitular proportions: ventral length / height 2.22 - 2.42 (2.22); rostrum length / width 2.65 - 2.69 (2.65). Pedipalps stocky with short, tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (47.5 - 47.5 (47.5) long); femur (97.5 - 100 (97.5) long); genu (62.5 - 62.5 (62.5) long); tibia (65 - 67.5 (67.5) long; 25 - 26.25 (25) wide); tarsus (16.25 - 17.5 (16.25) long). Palpomere proportions: femur / genu 1.56 - 1.60 (1.56); tibia / femur 0.65 - 0.69 (0.69); tibia length / width 2.48 - 2.70 (2.70). Venter - (640 - 680 (640) long; 440 - 495 (495) wide) with faint reddish-purple coloration. Gnathosomal bay (170 - 175 (170) long; 70 - 75 (70) wide). Cxgl- 4 subapical. Medial suture (55 - 75 (55) long). Genital plates (140 - 150 (140) long; 130 - 140 (130) wide). Additional measurements: Cx- 1 (290 - 300 (300) long (total); 110 - 135 (135) long (medial )); Cx- 3 (325 - 360 (360) wide); anterior venter (195 - 210 (200) long). Ventral proportions: gnathosomal bay length / width 2.33 - 2.50 (2.43); anterior venter / genital field length 1.37 - 1.43 (1.43); anterior venter length / genital field width 1.50 - 1.54 (1.54); anterior venter / medial suture 2.60 - 3.64 (3.64). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
EB1E1CB7A89BA2DDB9255A631ED5F87A.taxon	etymology	Etymology. Specific epithet (whitneyae) named in honor of Whitney Nelson, one of two students (including JRF) of APGD studying water mite taxonomy for their doctoral studies.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
EB1E1CB7A89BA2DDB9255A631ED5F87A.taxon	distribution	Distribution. Southern Appalachians (Figure 275).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
B7BEA7DC1B7120458394F3524C970360.taxon	materials_examined	Material examined. HOLOTYPE (♀): from USA, California, Monterey County, beside Rt. 1 south of Gorda, south side of Salmon Creek, (35 ° 49 ' 49 " N, 121 ° 22 ' 22 " W), 28 July 1987, by IM Smith, IMS 870115. PARATYPES (5 ♀; 4 ♂): California, USA: 1 ♂ (ALLOTYPE) from Monterey County, beside Rt. 1 south of Gorda, south side of Salmon Creek, (35 ° 49 ' 49 " N, 121 ° 22 ' 22 " W), 28 July 1987, by IM Smith, IMS 870115 * 1 ♀ and 1 ♂ from San Bernardino County, Claremont; Mount Baldy; beside road 3.5 km east of Mount Baldy Village, (34 ° 15 ' 15 " N, 117 ° 39 ' 39 " W), 24 July 1987, by IM Smith, IMS 870107 * 2 ♀ and 2 ♂ from San Bernardino County, Lytle Creek Recreation Area off Rt. 15 west of Devore; Applewhite Picnic Grounds, (34 ° 16 ' 16 " N, 117 ° 30 ' 30 " W), 23 July 1987, by IM Smith, IMS 870103 * 1 ♀ from Tulare County, beside Rt. 180 at Stony Creek Picnic Area east of Sequoia National Park boundary, (36 ° 40 ' 40 " N, 118 ° 50 ' 50 " W), 1 August 1987, by IM Smith, IMS 870124 A * 1 ♀ from Ventura County, Ojai; beside Rt. 33 just above Wheeler Gorge Campground, (34 ° 31 ' 31 " N, 119 ° 16 ' 16 " W), 25 July 1987, by IM Smith, IMS 870109 B.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
B7BEA7DC1B7120458394F3524C970360.taxon	diagnosis	Diagnosis. Torrenticola wiedenmanni is unlike all other western species by having this combination of characters: anterio-lateral platelets free (fused to dorsal plate in Rusetria Complex); colorless (dorsum with distinct coloration in Tahoei group, T. ventura, and T. raptoroides); distinct and complete hind coxal margins (incomplete in Rala Group, and T. sharkeyi); round body (rectangular or ovoid in Ellipsoidalis Group); unmodified rostrum (short and conical in T. sierrensis, T. olliei, T. leviathan, and T. ellipsoidalis); and tuberculate ventral extensions on pedipalpal genua (dentate and flanged in Miniforma group).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
B7BEA7DC1B7120458394F3524C970360.taxon	description	Description. Female (Figure 279) (n = 5) (holotype measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (560 - 640 (610) long; 445 - 540 (530) wide) circular and colorless. Anterio-medial platelets (110 - 150 (135) long; 55 - 75 (67.5) wide). Anterio-lateral platelets (182.5 - 200 (190) long; 65 - 85 (80) wide) free from dorsal plate. Dgl- 4 approximately halfway between the edge of the dorsum and the muscle scars and occasionally anterior to the muscle scars (distance between Dgl- 4 295 - 350 (315 )). Dorsal plate proportions: dorsum length / width 1.15 - 1.26 (1.15); dorsal width / distance between Dgl- 4 1.33 - 1.73 (1.68); anterio-medial platelet length / width 2.00 - 2.11 (2.00); anterio-lateral platelet length / width 2.35 - 2.81 (2.38); anterio-lateral / anterio-medial length 1.31 - 1.66 (1.41). Gnathosoma - Subcapitulum (302.5 - 355 (352.5) long (ventral); 222.5 - 267.5 (260) long (dorsal); 110 - 130 (120) tall) colorless. Rostrum (120 - 142.5 (142.5) long; 37.5 - 42.5 (40) wide). Chelicerae (310 - 385 (385) long) with curved fangs (47.5 - 50 (50) long). Subcapitular proportions: ventral length / height 2.73 - 2.94 (2.94); rostrum length / width 3.20 - 3.56 (3.56). Pedipalps with tuberculate ventral extensions on femora and genua, with femora sharply taper proximally and tibial setae near tarsi. Palpomeres: trochanter (30 - 38.75 (36.25) long); femur (90 - 107.5 (102.5) long); genu (65 - 75 (72.5) long); tibia (72.5 - 87.5 (87.5) long; 17.5 - 21.25 (21.25) wide); tarsus (13.75 - 17.5 (16.25) long). Palpomere proportions: femur / genu 1.37 - 1.43 (1.41); tibia / femur 0.79 - 0.85 (0.85); tibia length / width 4.00 - 4.27 (4.12). Venter - (710 - 810 (780) long; 560 - 630 (615) wide) colorless. Gnathosomal bay (132.5 - 160 (160) long; 62.5 - 65 (62.5) wide). Cxgl- 4 subapical. Medial suture (40 - 60 (40) long). Genital plates (172.5 - 187.5 (180) long; 150 - 165 (155) wide). Additional measurements: Cx- 1 (270 - 320 (320) long (total); 140 - 162.5 (162.5) long (medial )); Cx- 3 (330 - 385 (380) wide); anterior venter (200 - 235 (220) long). Ventral proportions: gnathosomal bay length / width 2.42 - 2.56 (2.56); anterior venter / genital field length 1.16 - 1.25 (1.22); anterior venter length / genital field width 1.32 - 1.42 (1.42); anterior venter / medial suture 3.92 - 5.50 (5.50). Male (Figure 280) (n = 4) (allotypic measurements in parentheses when available) with characters of the genus with following specifications. Dorsum - (560 - 610 (575) long; 470 - 515 (480) wide) circular and colorless. Anterio-medial platelets (130 - 140 (135) long; 62.5 - 70 (65) wide). Anterio-lateral platelets (165 - 195 (180) long; 70 - 75 (75) wide) free from dorsal plate. Dgl- 4 approximately halfway between the edge of the dorsum and the muscle scars and occasionally anterior to the muscle scars (distance between Dgl- 4 285 - 315 (300 )). Dorsal plate proportions: dorsum length / width 1.18 - 1.20 (1.20); dorsal width / distance between Dgl- 4 1.54 - 1.67 (1.60); anterio-medial platelet length / width 2.00 - 2.08 (2.08); anterio-lateral platelet length / width 2.20 - 2.60 (2.40); anterio-lateral / anterio-medial length 1.27 - 1.39 (1.33). Gnathosoma - Subcapitulum (307.5 - 345 (325) long (ventral); 225 - 257.5 (240) long (dorsal); 107.5 - 117.5 (107.5) tall) colorless. Rostrum (125 - 140 (130) long; 35 - 40 (37.5) wide). Chelicerae (335 - 357.5 (345) long) with curved fangs (45 - 50 (45) long). Subcapitular proportions: ventral length / height 2.80 - 3.02 (3.02); rostrum length / width 3.33 - 3.57 (3.47). Pedipalps with tuberculate ventral extensions on femora and genua, femora sharply taper proximally and tibial setae near tarsi. Palpomeres: trochanter (32.5 - 35 (32.5) long); femur (90 - 100 (90) long); genu (65 - 72.5 (65) long); tibia (75 - 80 (75) long; 20 - 20 (20) wide); tarsus (13.75 - 15 (15) long). Palpomere proportions: femur / genu 1.36 - 1.38 (1.38); tibia / femur 0.80 - 0.89 (0.83); tibia length / width 3.75 - 4.00 (3.75). Venter - (690 - 755 (715) long; 465 - 590 (465) wide) colorless. Gnathosomal bay (140 - 152.5 (145) long; 55 - 60 (60) wide). Cxgl- 4 subapical. Medial suture (70 - 85 (80) long). Genital plates (145 - 155 (150) long; 115 - 125 (117.5) wide). Additional measurements: Cx- 1 (280 - 300 (300) long (total); 140 - 160 (160) long (medial )); Cx- 3 (335 - 370 (345) wide); anterior venter (230 - 262.5 (260) long). Ventral proportions: gnathosomal bay length / width 2.42 - 2.64 (2.42); anterior venter / genital field length 1.53 - 1.73 (1.73); anterior venter length / genital field width 1.92 - 2.21 (2.21); anterior venter / medial suture 3.09 - 3.29 (3.25). Immatures unknown.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
B7BEA7DC1B7120458394F3524C970360.taxon	etymology	Etymology. Specific epithet (wiedenmanni) named in honor of Rob Wiedenmann, Professor of Entomology at University of Arkansas, whose advice, guidance, and friendship to JRF was an inspiration in how to be a true mentor-lessons that JRF anticipates paying forward to his own students someday.	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
B7BEA7DC1B7120458394F3524C970360.taxon	distribution	Distribution. Southern California (Figure 278).	en	Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A., Dowling, Ashley P. G. (2017): Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. ZooKeys 701: 1-496, DOI: http://dx.doi.org/10.3897/zookeys.701.13261, URL: http://dx.doi.org/10.3897/zookeys.701.13261
