taxonID	type	description	language	source
BE3D8799FFDEFFBAFF136546B165FE9F.taxon	type_taxon	Type species. Mercedula fuscomarginata Ponomarenko & M. Omelko, sp. nov. by present designation.	en	Ponomarenko, Margarita G., Omelko, Мichail M. (2025): New genus and two new species of gelechiid moths (Lepidoptera, Gelechiidae) from Malaysia and Indonesia. Zootaxa 5715 (1): 300-307, DOI: 10.11646/zootaxa.5715.1.28, URL: https://doi.org/10.11646/zootaxa.5715.1.28
BE3D8799FFDEFFBAFF136546B165FE9F.taxon	diagnosis	Diagnosis. The new genus can be distinguished from the genera of tribe Gelechiini by the male genitalia with long narrow uncus; longitudinally elongated tegumen, anterior part of which with complex apodeme; an absent gnathos; valva deeply divided into narrow dorsal part and ventral part broadened distally; shifted caudally phallic complex consisting of vinculum, juxta and aedeagus. Also, it differs by female genitalia with 8 th segment, consisting of two lateral sclerotisations and membranous areas between them, and two signa in corpus bursae.	en	Ponomarenko, Margarita G., Omelko, Мichail M. (2025): New genus and two new species of gelechiid moths (Lepidoptera, Gelechiidae) from Malaysia and Indonesia. Zootaxa 5715 (1): 300-307, DOI: 10.11646/zootaxa.5715.1.28, URL: https://doi.org/10.11646/zootaxa.5715.1.28
BE3D8799FFDEFFBAFF136546B165FE9F.taxon	description	Description. Adult. Head whitish with appressed large scales. Basal antennal segment (scapus) slightly flattened dorso-ventrally, brownish on upper side and light beige on under side, flagellum with light beige segments alternating with ones brownish on upper side and with black ones formed rings. Labial palpi with very short first segment, second segment 1.3 – 1.8 times longer than third segment, greyish brown on outer side and beige on inner side; third segment beige with two dark brown rings near the base and before apex (Figs 2, 7). Thorax, patagia and tegulae whitish or light beige. Forewings length 3.1 – 3.3 mm; light beige, with yellowish-brown stripe along costal margin from base to 3 / 5 or 4 / 5 of wing length, which widened distally, distinct small black spot at cell end formed by raised scales and large greyish brown spot posteriad the latter (Figs 1, 5, 6); fringe beige on the costal and outer edges of wing, and grey on posterior margin, with 7 sparse black dots around wing apex. Hindwings and fringe grey. Forelegs and middle legs light beige with brownish darkening on outer side of femur and tibia, and dark brown darkening on tarsi; hind legs light beige with brush of hair-like scales of same colour on upper and under sides of tibia and with light beige spurs, for except the outer spur of second pair, which dark brown. Second abdominal sternite with well-developed relatively long apodemes curved medially, long and sinuous venulae and medial fold arising from U-shaped narrow sclerotisation in proximal part of sternite (Figs 17, 24). Male genitalia (Figs 12, 13) (based on the male genitalia of M. fuscomarginata sp. nov.). Uncus narrow and long. Gnathos absent. Tegumen elongated caudally and widened at basal 2 / 5, with well-developed apodeme in anterior part bearing two pairs of longitudinal folds and rounded dorsal notch. Valva deeply divided into two separate parts joined with tegumen and vinculum through of short transversal basal sclerite. Valval dorsal part sclerotised, narrow and with pointed apex, broadened basally and parallel-sided beyond basal 2 / 5. Valval ventral part enlarged at about 3 / 5 of length and with rounded apex. Sternal phallic complex, consisting of vinculum, juxta and aedeagus, shifted caudally and joined with rest genital structures by lateral arms of vinculum, which long and narrow. Vinculum of complex morphology: two lateral arms widened distally in triangular lateral plates located on sides of aedeagus and fused with juxta, its ventral part scletotised and with narrow band anteriorly; without saccus. Juxta integrated with vinculum, relatively large, with wedge-shaped ventral notch and two ventro-lateral lobes bearing beak-like apices directed ventrally, it wraps aedeagus from ventral and lateral sides. Aedeagus fixed in sternal phallic complex, it curved dorsally at basal 2 / 5, with inflated basal part and tube-like distal one, without caecum and with truncated apex. Vesica with minute needle-like cornuti (Fig. 13). Female genitalia (Figs 14 – 16, 18 – 23). Papillae anales scletotised. Ovipositor short, membrane between segments 9 and 8 shorter than papillae anales, 8 th segment consisting of two lateral sclerotisations with rounded posterior angles and membranous areas between them, of which dorsal one narrower. Ostium and antrum shifted anteriorly, ostium almost at level of posterior edge of 7 th segment. Antrum sclerotised, funnel-shaped or cup-shaped, joined with posterior edge of 7 th segment (Figs 18, 20), ductus bursae membranous and tubular. Corpus bursae oval, membranous and with two signa each of which on underlying sclerotisation.	en	Ponomarenko, Margarita G., Omelko, Мichail M. (2025): New genus and two new species of gelechiid moths (Lepidoptera, Gelechiidae) from Malaysia and Indonesia. Zootaxa 5715 (1): 300-307, DOI: 10.11646/zootaxa.5715.1.28, URL: https://doi.org/10.11646/zootaxa.5715.1.28
BE3D8799FFDEFFBAFF136546B165FE9F.taxon	distribution	Distribution. Malaysia (Sabah) and Indonesia (Sumatra).	en	Ponomarenko, Margarita G., Omelko, Мichail M. (2025): New genus and two new species of gelechiid moths (Lepidoptera, Gelechiidae) from Malaysia and Indonesia. Zootaxa 5715 (1): 300-307, DOI: 10.11646/zootaxa.5715.1.28, URL: https://doi.org/10.11646/zootaxa.5715.1.28
BE3D8799FFDEFFBAFF136546B165FE9F.taxon	etymology	Etymology. The generic name is derived from Latin word “ mercedula ” meaning a small reward, which corresponds to the satisfaction from discovery of the new taxa as a result of tropical material study; generic name is feminine.	en	Ponomarenko, Margarita G., Omelko, Мichail M. (2025): New genus and two new species of gelechiid moths (Lepidoptera, Gelechiidae) from Malaysia and Indonesia. Zootaxa 5715 (1): 300-307, DOI: 10.11646/zootaxa.5715.1.28, URL: https://doi.org/10.11646/zootaxa.5715.1.28
BE3D8799FFDDFFBAFF13669AB355F827.taxon	description	(Figs 1 – 5, 10, 12 – 21)	en	Ponomarenko, Margarita G., Omelko, Мichail M. (2025): New genus and two new species of gelechiid moths (Lepidoptera, Gelechiidae) from Malaysia and Indonesia. Zootaxa 5715 (1): 300-307, DOI: 10.11646/zootaxa.5715.1.28, URL: https://doi.org/10.11646/zootaxa.5715.1.28
BE3D8799FFDDFFBAFF13669AB355F827.taxon	materials_examined	Type material. Holotype: ♂, Borneo, East Malaysia, state of Sabah, 9 km NNW of Keningau, 23. iii 2018 (leg. M. Omelko), GS 355 MP, FSCB. Paratypes: 1 ♂, 1 ♀, same locality, date and collector, 354 (♂), 359 (♀) MP; 1 ♂, 24 km north of Tawau, 19. viii 2019 (leg. M. Omelko), GS 353 MP, FSCB. Additional material. 1 ♀, Indonesia, North Sumatra, Bukit Lawang village, 19. ii 2023 (leg. M. Omelko), GS 356 MP; 1 ♀, same locality and collector, 18. ii 2023, GS 358 MP; FSCB.	en	Ponomarenko, Margarita G., Omelko, Мichail M. (2025): New genus and two new species of gelechiid moths (Lepidoptera, Gelechiidae) from Malaysia and Indonesia. Zootaxa 5715 (1): 300-307, DOI: 10.11646/zootaxa.5715.1.28, URL: https://doi.org/10.11646/zootaxa.5715.1.28
BE3D8799FFDDFFBAFF13669AB355F827.taxon	diagnosis	Diagnosis. New species is very similar in appearance to M. storozhenkoi sp. nov. described below. It can be distinguished by the relatively smaller size of moth, head with two tufts of elongated scales, each of which is placed on sides of frons, and forewing with costal stripe not reaching the level of cell end (Fig. 10). The new species also differs by female genitalia with large funnel-shaped antrum, antevaginal plate setaceous and with lateral drop-like lobes, small band-like postvaginal plate without honeycomb-like structure on surface and relatively small thorn-shaped signa (Figs 15, 16).	en	Ponomarenko, Margarita G., Omelko, Мichail M. (2025): New genus and two new species of gelechiid moths (Lepidoptera, Gelechiidae) from Malaysia and Indonesia. Zootaxa 5715 (1): 300-307, DOI: 10.11646/zootaxa.5715.1.28, URL: https://doi.org/10.11646/zootaxa.5715.1.28
BE3D8799FFDDFFBAFF13669AB355F827.taxon	description	Description. Adult (Figs 1 – 5, 10). Head with two dense tufts of elongated scales, each of which extends from behind antennal socket, covers scapus basally and eye anteriorly (Fig. 3). Flagellum in proximal part with one wide ring formed by four blackish segments before middle, in distal part with four black rings, two first of them equal in width and located just beyond the middle (Fig. 4). Forewing length 3.1 – 3.2 mm. Forewing light beige, interspersed with brownish scales, located more densely along dorsal margin; with yellowish-brown stripe along costal margin from base to 3 / 5 of wing length, which widened distally and not reaching level of cell end (Figs 1, 5, 10); distinct small black spot of raised scales at cell end; large greyish brown spot posteriad the latter; and with indistinct brownish band along distal third of costal edge; fringe light beige on costal and outer edges and dark grey on posterior margin. Hindwings and fringe dark grey. Male genitalia (Figs 12, 13). See the generic description above. Female genitalia (Figs 14 – 16). Ovipositor short, membrane between 9 th and 8 th segments about two times shorter than papillae anales. Glandular sac located at the base of apophyses posteriores on dorsal side. Apophyses posteriores and apophyses anteriores equal in length. 8 th segment consists of two lateral sclerotisations and two membranous areas between them, of which ventral wider. Ostium placed almost at level of posterior margin of 7 th segment, wide, about half of 8 th segment width, covered by rectangular antevaginal plate bearing small setae on inner surface and two drop-like lobes at each side (Fig. 15). Postvaginal plate as trapezoid transverse band. Antrum funnel-shaped, sclerotised, relatively long, almost equal to 8 th segment in length, immerged under 7 th segment and joined with its posterior margin, which with rounded sclerotised pockets on medial side at each side of antrum; anterior part of antrum weakly sclerotised and flattened in place where passing into ductus bursae. Ductus bursae membranous, smoothly widened towards corpus bursae. Corpus bursae oval with two small thorn-shaped signa, each placed on oval sclerotisation (Fig. 16).	en	Ponomarenko, Margarita G., Omelko, Мichail M. (2025): New genus and two new species of gelechiid moths (Lepidoptera, Gelechiidae) from Malaysia and Indonesia. Zootaxa 5715 (1): 300-307, DOI: 10.11646/zootaxa.5715.1.28, URL: https://doi.org/10.11646/zootaxa.5715.1.28
BE3D8799FFDDFFBAFF13669AB355F827.taxon	distribution	Distribution. East Malaysia (Sabah) and Indonesia (Sumatra).	en	Ponomarenko, Margarita G., Omelko, Мichail M. (2025): New genus and two new species of gelechiid moths (Lepidoptera, Gelechiidae) from Malaysia and Indonesia. Zootaxa 5715 (1): 300-307, DOI: 10.11646/zootaxa.5715.1.28, URL: https://doi.org/10.11646/zootaxa.5715.1.28
BE3D8799FFDDFFBAFF13669AB355F827.taxon	etymology	Etymology. The name of the new species is derived from the Latin roots “ fusc - ” and “ margin - ”, meaning “ dark ”, and “ border ”, referring to the pattern of the moth’s forewing with a dark stripe along the costal edge.	en	Ponomarenko, Margarita G., Omelko, Мichail M. (2025): New genus and two new species of gelechiid moths (Lepidoptera, Gelechiidae) from Malaysia and Indonesia. Zootaxa 5715 (1): 300-307, DOI: 10.11646/zootaxa.5715.1.28, URL: https://doi.org/10.11646/zootaxa.5715.1.28
BE3D8799FFDDFFBAFF13669AB355F827.taxon	discussion	Remarks. Two specimens, both females, collected in Sumatra are not included in type series of M. fuscomarginata sp. nov. since they differ from Malaysian specimens from Sabah by brownish costal stripe (Fig. 5) and some structures of the female genitalia. One of them has relatively narrower and longer antrum and closely placed drop-like lobes of antevaginal plate (Fig. 18), the other female has larger signa and underlying sclerotisation, whereas size of antrum corresponds to that in the paratype (Figs 19 – 21).	en	Ponomarenko, Margarita G., Omelko, Мichail M. (2025): New genus and two new species of gelechiid moths (Lepidoptera, Gelechiidae) from Malaysia and Indonesia. Zootaxa 5715 (1): 300-307, DOI: 10.11646/zootaxa.5715.1.28, URL: https://doi.org/10.11646/zootaxa.5715.1.28
BE3D8799FFDCFFBDFF1361B8B453FC17.taxon	description	(Figs 6 – 9, 11, 22 – 24)	en	Ponomarenko, Margarita G., Omelko, Мichail M. (2025): New genus and two new species of gelechiid moths (Lepidoptera, Gelechiidae) from Malaysia and Indonesia. Zootaxa 5715 (1): 300-307, DOI: 10.11646/zootaxa.5715.1.28, URL: https://doi.org/10.11646/zootaxa.5715.1.28
BE3D8799FFDCFFBDFF1361B8B453FC17.taxon	materials_examined	Type material. Holotype: ♀, Borneo, East Malaysia, state of Sabah, 24 km north of Tawau, 22. viii 2019 (leg. M. Omelko), GS 357 MP, FSCB.	en	Ponomarenko, Margarita G., Omelko, Мichail M. (2025): New genus and two new species of gelechiid moths (Lepidoptera, Gelechiidae) from Malaysia and Indonesia. Zootaxa 5715 (1): 300-307, DOI: 10.11646/zootaxa.5715.1.28, URL: https://doi.org/10.11646/zootaxa.5715.1.28
BE3D8799FFDCFFBDFF1361B8B453FC17.taxon	diagnosis	Diagnosis. New species differs from M. fuscomarginata sp. nov. by the relatively larger size of moth, head without tuft of elongated scales on each side of frons and forewing with costal stripe exceeding the level of cell end (Fig. 11). It can be also distinguished by female genitalia with small ostium about ¼ of 8 th segment width, smaller cup-shaped antrum, postvaginal plate with honeycomb-like sculpture on surface and relatively large thorn-shaped signa in corpus bursae.	en	Ponomarenko, Margarita G., Omelko, Мichail M. (2025): New genus and two new species of gelechiid moths (Lepidoptera, Gelechiidae) from Malaysia and Indonesia. Zootaxa 5715 (1): 300-307, DOI: 10.11646/zootaxa.5715.1.28, URL: https://doi.org/10.11646/zootaxa.5715.1.28
BE3D8799FFDCFFBDFF1361B8B453FC17.taxon	description	Description. Adult, female (Figs 6 – 9). Head without tufts of elongated scales. Flagellum in proximal part with one wide ring formed by three blackish segments before middle, in distal part with four black rings, first of which wider, than in M. fuscomarginata sp. nov. (Figs 8, 9). Forewing length 3.3 mm. Forewing light beige, interspersed with brownish scales located more densely on distal ¼ of wing before apex; with yellowish-brown stripe along costal margin from base to almost 4 / 5 of wing length, which exceeds level of cell end (Fig. 11); small black spot of raised scales at cell end and greyish brown spot posteriad the latter; fringe light beige on costal and outer edges and dark grey on posterior margin. Hindwings and fringe dark grey. Female genitalia (Figs 22, 23). Papillae anales oblong and scletotised. Ovipositor short, membrane between 9 th and 8 th segments slightly shorter than papillae anales. Apophyses posteriores and apophyses anteriores equal in length. 8 th segment with wide ventral membranous area and narrower dorsal one. Ostium small, about 1 / 5 of 8 th segment width. Postvaginal plate more or less trapezoidal, with flat posterior edge and deep rounded notch on anterior half, in which ostium and posterior edge of antrum placed. Antrum cup-shaped, sclerotised and relatively short, about 1 / 4 of 8 th segment length. Ductus bursae membranous, narrow and tubular, slightly shorter than corpus bursae. Corpus bursae oval with two thorn-shaped signa in posterior half, each signum with oval underlying sclerotisation. Male unknown.	en	Ponomarenko, Margarita G., Omelko, Мichail M. (2025): New genus and two new species of gelechiid moths (Lepidoptera, Gelechiidae) from Malaysia and Indonesia. Zootaxa 5715 (1): 300-307, DOI: 10.11646/zootaxa.5715.1.28, URL: https://doi.org/10.11646/zootaxa.5715.1.28
BE3D8799FFDCFFBDFF1361B8B453FC17.taxon	distribution	Distribution. East Malaysia (Sabah).	en	Ponomarenko, Margarita G., Omelko, Мichail M. (2025): New genus and two new species of gelechiid moths (Lepidoptera, Gelechiidae) from Malaysia and Indonesia. Zootaxa 5715 (1): 300-307, DOI: 10.11646/zootaxa.5715.1.28, URL: https://doi.org/10.11646/zootaxa.5715.1.28
BE3D8799FFDCFFBDFF1361B8B453FC17.taxon	etymology	Etymology. The species is named after Prof. Sergey Yu. Storozhenko, a famous expert with the orthopteroid insects and who made a great contribution to the study of morphology, fauna, taxonomy, and evolution of these insects. A noun in the genitive case.	en	Ponomarenko, Margarita G., Omelko, Мichail M. (2025): New genus and two new species of gelechiid moths (Lepidoptera, Gelechiidae) from Malaysia and Indonesia. Zootaxa 5715 (1): 300-307, DOI: 10.11646/zootaxa.5715.1.28, URL: https://doi.org/10.11646/zootaxa.5715.1.28
BE3D8799FFDCFFBDFF1361B8B453FC17.taxon	discussion	Remarks. This species is described within genus Mercedula gen. nov. based on the similarity in appearance and some morphostructures of the female genitalia with those in the type species M. fuscomarginata sp. nov. However, M. storozhenkoi sp. nov. has significant differences in the ostial area, namely trapezoidal postvaginal plate with honey-like structure on surface and smaller cup-shaped antrum. Therefore, this species is tentatively placed in this genus until the male morphology will be studied to clarify its taxonomic position.	en	Ponomarenko, Margarita G., Omelko, Мichail M. (2025): New genus and two new species of gelechiid moths (Lepidoptera, Gelechiidae) from Malaysia and Indonesia. Zootaxa 5715 (1): 300-307, DOI: 10.11646/zootaxa.5715.1.28, URL: https://doi.org/10.11646/zootaxa.5715.1.28
