taxonID	type	description	language	source
66EB7300C73C53C7BFA5AE266FBFD4F1.taxon	description	Figs 1, 2, 3, 4 Grumpy dwarfgoby	en	Nunes Peinemann, Viktor, Pombo-Ayora, Lucía, Tornabene, Luke, Berumen, Michael L. (2024): The Grumpy dwarfgoby, a new species of Sueviota (Teleostei, Gobiidae) from the Red Sea. ZooKeys 1212: 17-28, DOI: 10.3897/zookeys.1212.121135
66EB7300C73C53C7BFA5AE266FBFD4F1.taxon	diagnosis	Diagnosis. This is a species of Sueviota characterized by the following combination of characters: no cephalic sensory-canal pores; dorsal fin VI-I, 8, or I, 9, without filamentous spines; anal fin I, 7 or I, 8; pelvic fin I, 5, rays 1 – 4 branched, fifth ray unbranched or with two branches, elongate (77 – 88 % of fourth) and flattened towards the tips if unbranched, fourth ray longest; well-developed pelvic fin membrane fully joining fifth pelvic fin rays, frenum absent; 14 or 15 pectoral fin rays, some branched; body robust and deep, anterior slope of snout nearly vertical giving the head a blunt profile, terminal mouth inclined vertically forming a 72 ° angle to horizontal body axis.	en	Nunes Peinemann, Viktor, Pombo-Ayora, Lucía, Tornabene, Luke, Berumen, Michael L. (2024): The Grumpy dwarfgoby, a new species of Sueviota (Teleostei, Gobiidae) from the Red Sea. ZooKeys 1212: 17-28, DOI: 10.3897/zookeys.1212.121135
66EB7300C73C53C7BFA5AE266FBFD4F1.taxon	description	Description. Dorsal fin elements VI-I, 9 (I, 8), first dorsal fin rounded to square shaped, second and third spines slightly longer than the first spine; no elongate filaments on first dorsal fin; some or all soft rays of second dorsal fin branched, final ray branched to base; anal fin I, 8 (one paratype I, 7); pectoral fin rays 14 (14 – 15), 6 – 7 lower rays branched; pelvic fin I, 5; fifth ray 77 – 88 % of fourth ray; fourth pelvic fin ray with 4 branches, fifth pelvic fin ray unbranched or with 2 branches; 3 - 2 - 2 segments between consecutive branches of fourth pelvic fin ray; membrane connecting pelvic 5 th fin rays well developed extending out towards the tip, no frenum (Fig. 2 B); 17 (15 – 17) segmented and 13 (12 – 13) branched caudal fin rays; lateral scale rows 25 (24 – 25); transverse scale rows 7 (6 – 7); ctenoid scales on body, no scales on head and breast; anterior extent of scales does not reach the base of the pectoral fin, no scales on the base of the first dorsal fin, but present on the base of the second dorsal fin; scales on the trunk extend ventrally onto the abdomen to beneath the pelvic fin rays with a small naked section on the ventral midline of the abdomen; front of the head distinctively blunt; mouth inclined vertically forming an angle of 72 ° to horizontal body axis, lower jaw projecting; upper jaw extending posteriorly to a vertical reaching the middle of the pupil; anterior tubular nares about 50 % of the pupil diameter, posterior nares enlarged with an elevated rim adjacent to the eye; prominent canines; gill opening extending forward reaching the anterior edge of the operculum; lacking entire cephalic sensory canals and corresponding pores, but papillae (free neuromasts) are present where canal pores would be located (Fig. 2 A); female urogenital papilla short and bulbous (Fig. 3 B); male urogenital papilla elongated, smooth and bulbous with weakly fimbriate margin resembling type e (Fig. 3 A); vertebral count 10 + 15 = 25 (Fig. 4 C). Measurements (percentage of SL; based on holotype and nine paratypes, 9.2 – 16.7): head length 27.1 (22.8 – 27.1); origin of first dorsal fin 33.8 (33.5 – 37.9), slightly behind pectoral fin base and pelvic fin origin; origin of second dorsal fin 55.6 (53.2 – 59.8); origin of anal fin 58.6 (56.2 – 60.9); caudal peduncle length 20.3 (19.5 – 23.8); caudal peduncle depth 15.0 (14.9 – 17.5); body depth at origin of first dorsal fin 22.6 (19.5 – 25.2), body relatively slender; pectoral fin length 16.5 (17.3 – 19.9); pelvic fin length 19.3 (15.0 – 21.3). As a percentage of HL: eye diameter 25.8 (22.2 – 30.8); snout length 15 (15.8 – 21.3); upper-jaw length 52.8 (46.9 – 59.3). Teeth: Sueviota aethon has two irregular rows of conical teeth in both its upper and lower jaws. As described by Winterbottom and Hoese (1988) in the initial description of the Sueviota genus, S. aethon has enlarged conical canines in both the upper and lower jaws. A closer examination of the head osteology through a micro-CT scan (Fig. 4) shows this detail with one enlarged curved canine situated on each side of the premaxilla, as well as one enlarged curved canine on each side of the innermost row of teeth on the dentary.	en	Nunes Peinemann, Viktor, Pombo-Ayora, Lucía, Tornabene, Luke, Berumen, Michael L. (2024): The Grumpy dwarfgoby, a new species of Sueviota (Teleostei, Gobiidae) from the Red Sea. ZooKeys 1212: 17-28, DOI: 10.3897/zookeys.1212.121135
66EB7300C73C53C7BFA5AE266FBFD4F1.taxon	etymology	Etymology. The specific epithet stems from the ancient Greek Aethon, one of the four horses of the sun god Helios. The most similar species to S. aethon, Sueviota pyrios Greenfield & Randall, 2017, is named after a different horse of Helios. The specific name is a noun in apposition. The common name, Grumpy dwarfgoby, refers to the fish’s apparent grumpy and rather unhappy appearance, primarily due to the extremely upturned mouth position.	en	Nunes Peinemann, Viktor, Pombo-Ayora, Lucía, Tornabene, Luke, Berumen, Michael L. (2024): The Grumpy dwarfgoby, a new species of Sueviota (Teleostei, Gobiidae) from the Red Sea. ZooKeys 1212: 17-28, DOI: 10.3897/zookeys.1212.121135
66EB7300C73C53C7BFA5AE266FBFD4F1.taxon	distribution	Distribution and habitat. Sueviota aethon is a rare species, with only ten specimens found during extensive rotenone and clove oil collections along the Saudi Arabian Red Sea coast. These specimens were collected at depths between 10 and 30 meters. A sample from another expedition (not presented here) was confirmed from 53 meters depth. The species may be more common at similar mesophotic depths, but further collections are needed to confirm this. The specimens collected with clove oil were found on CCA-covered roofs of small caves. Six specimens were collected from a single mass sampling at a CCA-covered wall with small crevices and holes. All but one of the specimens analyzed in this study were collected from exposed offshore reefs. Sueviota aethon is presumably a Red Sea endemic. Our records range from Al Qunfudhah (18.9922 ° N, 40.6145 ° E) to Thuwal (22.4283 ° N, 38.9932 ° E), but it is likely that the species is more widely distributed throughout the main basin of the Red Sea (Fig. 5).	en	Nunes Peinemann, Viktor, Pombo-Ayora, Lucía, Tornabene, Luke, Berumen, Michael L. (2024): The Grumpy dwarfgoby, a new species of Sueviota (Teleostei, Gobiidae) from the Red Sea. ZooKeys 1212: 17-28, DOI: 10.3897/zookeys.1212.121135
