taxonID	type	description	language	source
B5A0191EDAC458AFB3B9529391532757.taxon	etymology	Etymology. Mercurio and Andreone (2007) dedicated G. azzurrae to F. Andreone's second daughter, Kintana Azzurra Andreone. Since this name turned out to be a junior synonym of G. corvus, F. Andreone and the other authors of this paper wish to dedicate the new species to honour her with the new name. The Malagasy word " kintana " means " star " and is used as a noun in apposition.	en	Cocca, Walter, Andreone, Franco, Belluardo, Francesco, Rosa, Goncalo M., Randrianirina, Jasmin E., Glaw, Frank, Crottini, Angelica (2020): Resolving a taxonomic and nomenclatural puzzle in mantellid frogs: synonymization of Gephyromantis azzurrae with G. corvus, and description of Gephyromantis kintana sp. nov. from the Isalo Massif, western Madagascar. ZooKeys 951: 133-157, DOI: http://dx.doi.org/10.3897/zookeys.951.51129, URL: http://dx.doi.org/10.3897/zookeys.951.51129
B5A0191EDAC458AFB3B9529391532757.taxon	diagnosis	Diagnosis. A species assigned to the genus Gephyromantis (sensu Glaw and Vences 2006), subgenus Phylacomantis, based on genetic and morphological similarities to the other known species (G. atsingy, G. corvus, and G. pseudoasper), and recognisable by the presence of the following morphological characters and natural history traits: (1) medium size (adult male SVL 36 - 44 mm), (2) webbing between toes present, (3) lateral metatarsalia partly connected, (4) inner and outer metatarsal tubercles present, (5) presence of femoral glands of " Type 2 " (sensu Glaw et al. 2000), (6) presence of a paired subgular vocal sac, (7) tongue bifid, (8) enlarged triangular finger tips; (9) dirty white throat, belly and thighs, (10) males with white vocal sacs; (11) brownish to olive-grey dorsal colouration with multiple and irregular brown-olive patches, (12) occurrence in young (shallow) canyons with limited (to almost no) vegetation, (13) mostly crepuscular / nocturnal activity, (14) advertisement call (see Mercurio and Andreone 2007 for the description of the advertisement call of specimen MRSN A 5313 (ZSM 0047 / 2011), now genetically assigned to G. kintana sp. nov.). The new species differs from the three other species of Phylacomantis by high genetic differentiation (pairwise 16 S distance ranging from 9.9 % to 13.4 %), as well as from a combination of morphological and natural history traits. Gephyromantis kintana sp. nov. is overall similar to the other three species of the subgenus Phylacomantis. Distinguished from G. pseudoasper by: (a) dirty white throat (vs. darker colouration); (b) ventrally dirty white thighs (vs. orange colouration); (c) presence of white vocal sacs (vs. blackish vocal sacs); (d) less granular dorsal skin; (e) larger size (maximum SVL in males 43.6 vs. 37.4 mm), (f) higher maximum number of granules in the femoral glands (96 vs. 43), (g) occurrence in young (shallow) canyons with limited vegetation (vs. mostly rainforest), (h) advertisement call (15 - 21 vs. 3 notes per call and lower dominant frequency, 3,000 - 3,200 Hz vs. 3,400 - 5,000 Hz). Distinguished from the sympatric G. corvus by: (a) brownish to olive grey dorsal colouration with multiple and irregular brown-olive patches (vs. darker brown dorsal colouration, often with a broad vertebral stripe), (b) dirty white throat (vs. dark brown throat), (c) dirty white belly (vs. brown belly), (d) dirty white thighs (vs. brown thighs); (e) presence of white vocal sacs (vs. brown-blackish vocal sacs), (f) higher maximum number of granules in the femoral glands (96 vs. 58), (g) occurrence in young (shallow) canyons with limited vegetation (vs. dry deciduous gallery forest in deep canyons), (g) advertisement call (15 - 21 vs. 10 - 14 notes per call and higher dominant frequency, 3,000 - 3,200 Hz vs. 2,400 - 2,700 Hz). Distinguished from G. atsingy by: (a) brownish to olive grey dorsal colouration with multiple and irregular brown-olive patches (vs. light brown-beige with a greenish shading), (b) less granular dorsal skin; (c) larger size (maximum SVL in males 43.6 vs. 36.6 mm), (d) higher maximum number of granules in the femoral glands (96 vs. 70), e) occurrence in young (shallow) canyons with limited / missing vegetation (vs. " tsingy " geological formations).	en	Cocca, Walter, Andreone, Franco, Belluardo, Francesco, Rosa, Goncalo M., Randrianirina, Jasmin E., Glaw, Frank, Crottini, Angelica (2020): Resolving a taxonomic and nomenclatural puzzle in mantellid frogs: synonymization of Gephyromantis azzurrae with G. corvus, and description of Gephyromantis kintana sp. nov. from the Isalo Massif, western Madagascar. ZooKeys 951: 133-157, DOI: http://dx.doi.org/10.3897/zookeys.951.51129, URL: http://dx.doi.org/10.3897/zookeys.951.51129
B5A0191EDAC458AFB3B9529391532757.taxon	description	Description of the holotype (Figs 3 D, 4 D). Adult male in good state of preservation, distal phalanx of the 5 th toe of the left foot removed as tissue sample and part of the ventral surface of thighs cut and opened to count the number of the granules of the femoral gland. SVL 38.2 mm; for other measurements see Table 2. Body slender; head slightly wider than long; snout slightly pointed in dorsal view, rather rounded in lateral view; nostrils directed laterally much nearer to tip of snout than to eye; canthus rostralis moderately defined; tympanum distinct, rounded, its horizontal diameter 0.8 of eye diameter; supratympanic fold well distinct, regularly curved; tongue distinctly bifid posteriorly. Arms slender; subarticular tubercles single; outer metacarpal tubercle poorly developed, inner metacarpal tubercle relatively well developed; fingers without webbing; finger disks triangular and distinctly enlarged; nuptial pads absent. Hind limbs slender; tibiotarsal articulation reaching beyond the snout tip when hindlimbs are depressed along body; lateral metatarsalia partly connected; inner metatarsal tubercle distinct, outer metatarsal tubercle small but recognisable; webbing of foot 1 (1), 2 i (1.75), 2 e (0.75), 3 i (2), 3 e (1), 4 i (2), 4 e (2), 5 (0.5). Skin slightly granular on dorsum and belly, ventral skin smooth on throat and chest. Femoral glands are distinctly recognisable from external view and are arranged in a typical glandular cluster (" Type 2 ", according to Glaw et al. 2000), including 71 single whitish granular glands of ca. 0.3 mm diameter. The vocal sacs in the male holotype are white and distinct.	en	Cocca, Walter, Andreone, Franco, Belluardo, Francesco, Rosa, Goncalo M., Randrianirina, Jasmin E., Glaw, Frank, Crottini, Angelica (2020): Resolving a taxonomic and nomenclatural puzzle in mantellid frogs: synonymization of Gephyromantis azzurrae with G. corvus, and description of Gephyromantis kintana sp. nov. from the Isalo Massif, western Madagascar. ZooKeys 951: 133-157, DOI: http://dx.doi.org/10.3897/zookeys.951.51129, URL: http://dx.doi.org/10.3897/zookeys.951.51129
B5A0191EDAC458AFB3B9529391532757.taxon	distribution	Distribution. Gephyromantis kintana sp. nov. is currently known from localities inside (Piscine Naturelle, Zahavola, Sakamalio, Malaso, Tsiombivositra, and Ambovo) and outside (Andranomena, Andranombilahy, Andohasahenina, and possibly Ilakaka) the borders of Isalo National Park (Fig. 1). However, this known distributional area remains restricted to the southern and western portion of the Isalo Massif. Now that we provided a straightforward way to distinguish this species from its sympatric sister species G. corvus more field surveys should be conducted to characterise its distribution in detail. The range encompasses elevations from 726 - 999 m a. s. l. The population densities are not known but it can be locally abundant, with several individuals grouping together close to remaining water bodies forming ponds in shallow (young) canyons (see Fig. 5 C).	en	Cocca, Walter, Andreone, Franco, Belluardo, Francesco, Rosa, Goncalo M., Randrianirina, Jasmin E., Glaw, Frank, Crottini, Angelica (2020): Resolving a taxonomic and nomenclatural puzzle in mantellid frogs: synonymization of Gephyromantis azzurrae with G. corvus, and description of Gephyromantis kintana sp. nov. from the Isalo Massif, western Madagascar. ZooKeys 951: 133-157, DOI: http://dx.doi.org/10.3897/zookeys.951.51129, URL: http://dx.doi.org/10.3897/zookeys.951.51129
