taxonID	type	description	language	source
BA7D4A9F85273E795210F834AA6190F3.taxon	diagnosis	Diagnosis. Nemastygnus was transferred from Cranaidae to Gonyleptidae, Metasarcinae by Kury (2003). It is closely related to Avima Roewer, 1949, based on dorsal scutum shape (rectangular) and scutal area III (unarmed). It is impossible to distinguish both genera mainly because Avima, the largest genus of Leiosteninae with 33 species, is a heterogeneous genus based on penial characters. The type-species of Avima, Avima leucobunus Roewer, 1949, is known only from the original description (and only external morphology). However, we will not propose synonymy in this paper because Nemastygnus is older and 33 new combinations should be proposed. A review of Avima plus Nemastygnus species is needed.	en	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
BA7D4A9F85273E795210F834AA6190F3.taxon	description	Description. Ocularium unarmed, saddle-shaped. Areas and posterior margin of dorsal scutum and free tergites unarmed (Fig. 1 A). Scutal area I undivided. Lateral margin of dorsal scutum coriaceus. Chelicera dimorphic. Pedipalp with slender articles; femur without dorsoapical spine, with row of three ventral large setae and two ventroapical large setae; patella with prolateral large setae; tibia-tarsus with ectal-mesal large setae. Leg I filiform, three-segmented. Penis: basal setae of truncus and basal setae of ventral plate very long and bifid; stylus with a large, longitudinal keel.	en	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
9C262AEE532EB1D56E8C6DA826F87240.taxon	description	Description. Penis (Fig. 1 B-C; holotype): truncus with three pairs of bifid setae (basal, subdistal lateral and distal ventral). Ventral plate with rounded lobe on corners, two dorso-basal pairs of small single branched setae, a ventral distal pair of small single branched setae and a ventral median bifid setae. Glans dorsally projected, slender, with a small ventral crest and a long dorsal crest. Stylus smooth.	en	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
5D6F1EB143BCDAD269631F67BE1C64C8.taxon	diagnosis	Diagnosis. Taulisa (Fig. 2) differs from the other 10 genera of Leiosteninae by the following combination of characters: ocularium unarmed, saddle-shaped; areas of dorsal scutum with two large tubercles each, posterior margin of dorsal scutum and free tergites, each one with a central large tubercle; area I undivided; lateral margin of dorsal scutum coriaceus; pedipalp with slender articles, its femur with dorsoapical spine, with row of four ventral large setae; pedipalpal patella with prolateral large setae; pedipalpal tibia-tarsus with ectal-mesal large setae; leg I filiform, three-segmented. Male unknown.	en	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
E334DFD1D805B996F1DA3EE5205C551A.taxon	diagnosis	Diagnosis. Napostygnus was originally placed in Gonyleptidae, Prostygninae and transferred to the Gonyleptidae, Metasarcinae by Kury (2003). It differs from the other 75 genera of Cranaidae by the following combination of characters: ocularium, scutal areas I-IV, posterior margin of dorsal scutum and free tergite I unarmed; free tergites II-III with one spine; posterior margin of dorsal scutum concave; legs thin and unarmed; penis with ventral plate wider at the middle, three pairs of distal setae, two pairs of basal setae, glans with dorsal process, stylus with apex enlarged.	en	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
77324B80A96E80D91D08AE8037767ADC.taxon	diagnosis	Diagnosis. As for genus.	en	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
0D9602A1EB4016A1A68A22D96B2CBA02.taxon	description	Description. Male (MZSP 36132). Dorsum (Fig. 3 A). Measurements: dorsal scutum length 4.1; dorsal scutum maximum width 3.5; carapace length 2.1; carapace maximum width 2.9; femur IV length 11.2. Body outline nearly subrectangular. Anterior margin of dorsal scutum with a median frontal hump small-granulate. Ocularium near middle of carapace, saddle shaped, with small granules near the eyes, unarmed. Carapace higher than the rest of dorsal scutum, with 4 tubercles behind ocularium. Scutal areas I-III with 2 small median tubercles on each area; IV with 4 tubercles. Lateral margin of dorsal scutum with a low density of small granules. Posterior margin of dorsal scutum and free tergite I with a row of small granules, unarmed. Free tergites II-III each with a median spine, small granulate. Chelicera: segment I unarmed. Segment II swollen, finger II with 3 teeth, III with 4 teeth. Pedipalpus: trochanter with 2 ventral tubercles, femur and patella smooth. Tibial setation: retrolateral iiIii; prolateral iiiIiii. Tarsal setation: retrolateral IiIi; prolateral IiiiIi. Legs: legs I-IV unarmed and without granules, except for trochanters, which are small granulate. Basitarsus I slightly inflated. Tarsal process present. Tarsal segmentation: 6 (3); 16 (3); 6; 7. Penis (Fig. 3 B-C): ventral plate with almost straight distal margin, thick median lobe and folded ventrally to the distal setae, 3 pairs of distal setae and 2 pairs of basal setae. Glans with thumb-like dorsal process. Stylus with dorsal apical projection and ventral apical small trichomes. Coloration (in ethanol) (Fig. 3 A): body background yellow with brown spots mainly on carapace, scutal areas, lateral and posterior margins of dorsal scutum and free tergites. Mesotergum with one longitudinal yellow stripe surrounded by blackish pigment at grooves I-V. Pedipalpus and chelicera yellowish brown with a brown reticulate pattern. Legs yellowish brown.	en	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
8F100F3A3D4C034E369EF464C87CB5D3.taxon	diagnosis	Diagnosis. Neopachyloides was a hitherto monotypic genus that resembles Ampycinae genera with a paired armature on ocularium and free tergite III with a median, long spiniform apophysis asin Ampycella Roewer, 1929, Ampycus Simon, 1879, Hutamaia Soares & Soares, 1977 and Sibollus Roewer, 1929. Neopachyloides can be distinguished from these genera by the following combination of characteristics: scutal area III with a paramedian pair of enlarged, pointed tubercles, scutal area IV undivided and free tergite II unarmed. Neopachyloides peruvianus can be distinguished from Neopachyloides spinipes Roewer, 1913 by the dorsal scutum covered by granules and scutal areas I-II unarmed (Fig. 4 A).	en	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
8F100F3A3D4C034E369EF464C87CB5D3.taxon	description	Description. Penis (Fig. 4 B-C; holotype): ventral plate with sub hexagonal shape, deep cleft on distal margin, 4 pairs of distal setae (distalmost curved and basalmost small), 3 pairs of basal setae straight. Glans very long (⅔ of ventral plate length), stylus smooth and curved dorsally, without dorsal and ventral processes.	en	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
E315E5F58F07063706EFE43F502FBAC6.taxon	diagnosis	Diagnosis. Pirunipygus is a monotypic genus and resembles Ampycinae genera with a paramedian pair of enlarged tubercles on scutal areas I-II as Ampycus Simon, 1879, Hexabunus Roewer, 1913, Neopachyloides Roewer, 1913 and Parahernandria Go odnight & Goodnight, 1947. Pirunipygus paradoxus can be distinguished from these genera by the following combination of characters: ocularium with a median spiniform apophysis (Fig. 5 A); dorsal scutum with secondary tubercles (sensu Maury 1992); scutal area III with a paramedian pair of spiniform apophyses; free tergite II with a median spiniform apophysis; free tergite III with three spiniform apophyses, the middle one largest and bifid.	en	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
5EB53246070FE5B439C049CE6B6B3E65.taxon	diagnosis	Diagnosis. As in Kury (2003: 137).	en	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
CC5133B5F0FE1F303C44DF2BFF5FB97C.taxon	diagnosis	Diagnosis. Acanthogonyleptes singularis and Acanthogonyleptes variolosus can be distinguished from the remaining species of the genus by the presence of a median single elevation on scutal area III. Males of Acanthogonyleptes singularis can be distinguished from those of Acanthogonyleptes variolosus by the presence of a robust curved retrolateral apophysis on trochanter IV.	en	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
1084F131AFA4A2CD376292E57EEAF2B4.taxon	description	Description. Penis (Fig. 5 B-C; holotype): ventral plate sub hexagonal, deep U-cleft on distal margin, 3 - 4 distal pairs of setae (2 - 3 distalmost curved), 2 pairs of dorsal median setae of moderate size, 2 pairs of ventral median small setae. Stylus wide on basal 3 / 4, apex twisted, with a patch of ventral subapical trichomes and some scattered apically. Glans without dorsal and ventral processes.	en	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
F67722BD0705688E10A9D0F9615A933F.taxon	diagnosis	Diagnosis. Gonyleptes is a genus composed of generally large species (dorsal scutum length longer than 8 mm), which males present femur IV with conspicuous armature and scutal areas I-III convex in lateral view. The penis presents ventral plate very convex and apex of ventral process serrate and semicircular. See taxonomical note below.	en	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
7DC0E0AE964DD95B57CE8361E7CDBA12.taxon	diagnosis	Diagnosis. Gonyleptes horridus can be recognized by: the relatively larger size; anterior border of carapace smooth (without tubercles or spines in the corners); frontal hump on anterior border of carapace and ocularium with white small tubercles; prolateral apical apophysis of coxa IV long, with abrupt curvature backwards. Gonyleptes horridus, Gonyleptes perlatus and Gonyleptes pustulatus can be distinguished from all other species of Gonyleptes by the robust and bifid (two distal tips) retrolateral apical apophysis of coxa IV. Females of Gonyleptes horridus and Gonyleptes perlatus present dimorphic sulfur yellow spines on the free tergites, not present in females of Gonyleptes pustulatus. Gonyleptes horridus can be distinguished from Gonyleptes perlatus by the armature pattern of femur IV of the males (see note on Gonyleptes perlatus below). Gonyleptes horridus has relatively smaller tubercles on posterior part of the lateral margins of abdominal scutum than in Gonyleptes perlatus.	en	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
7DC0E0AE964DD95B57CE8361E7CDBA12.taxon	description	Description. Dorsal scutum length 7 - 9 mm. Anterior margin of carapace without tubercles or spines. Median frontal hump on anterior margin of carapace and ocularium with a pair of tubercles on each one (Fig. 6 A). Four scutal grooves delimiting three scutal areas. Scutal areas I, II with low white tubercles in the median region; III with high paramedian tubercles. Lateral margin of dorsal scutum with flattened white tubercles. Pedipalps approximately same length as dorsal scutum. Prolateral apophyses of coxa IV elongated, strongly curved backwards, dorsal branch longest. Retrolateral apophysis of coxa IV bifid. Femur IV straight, with dorsobasal apophyses and longitudinal rows of spines (Fig. 6 B-C). Penis with typical Gonyleptinae - like pattern (Kury 1992 b). Tarsal segmentation: 6 (3); 10 - 12 (3); 7 - 8; 8 - 9. Females with dimorphic cones (one yellowish huge median spine) on free tergites II and III. General coloration brown with white tubercles (Fig. 6 D). Penis (Fig. 7 A-C; MZSP 15496): ventral plate with a deep U-cleft on anterior margin (its distal tips convergent), 3 distal, 1 median and 4 basal pairs of setae, ventrally with a subdistal and a median pair of setae. Stylus sigmoid, with ventral subapical trichomes. Ventral process robust, apex as a flabellum with several small projections on distal margin.	en	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
92484EAA302D83651B3C9287FF39815C.taxon	diagnosis	Diagnosis. Gonyleptes curvicornis resembles Gonyleptes horridus (see diagnosis above) but can be distinguished by the relatively shorter prolateral apical apophysis of coxa IV, lacking the abrupt backwards curvature (Fig. 6 E). The retrolateral apical apophysis of coxa IV is reduced and not bifid as in Gonyleptes horridus. The male femur IV has a different pattern of armature (Fig. 6 F-G) compared to Gonyleptes horridus. Carapace with a pair of tubercles on the posterior area (posterior to ocularium). Females of Gonyleptes curvicornis do not present dimorphic sulfur yellow spines on the free tergites.	en	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
92484EAA302D83651B3C9287FF39815C.taxon	description	Description. Penis (Fig. 8 A-B; MZSP 1117): ventral plate sub hexagonal, with a deep U-cleft on distal margin (its distal tips convergent), 3 distal, 1 median, 4 basal pairs of setae (distal ones curved, median ones shortest). Stylus sigmoid, with ventral subapical trichomes. Ventral process robust, apex flabelliform with several projections of varying sizes on distal margin.	en	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
E67535A551E03FB4CB82811E01881AE7.taxon	diagnosis	Diagnosis. Dorsal scutum length 8 - 10 mm. Corners of anterior margin of carapace smooth. Median frontal hump on anterior margin of carapace and ocularium with one pair of tubercles each one. Posterior region of carapace with 2 median tubercles. Mesotergum densely covered by tubercles. Scutal areas I-III each with a pair of median enlarged tubercles (III with largest and round ones). Prolateral apical apophysis of male coxa IV slightly directed backwards, retrolateral apophysis of male coxa IV absent or very reduced (Fig. 9 A-B). Dorsobasal apophysis of male femur IV robust, curved and retrolaterally oriented. Ornamentation of male femur IV variable, normally with two prolateral spines.	en	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
E67535A551E03FB4CB82811E01881AE7.taxon	description	Description. Penis (Fig. 10 A-C, MZSP 29393): ventral plate with 3 distal and 3 basal pairs of large setae, 1 median pair of short setae. Stylus with ventral trichomes. Ventral process with a triangular flabellum, with digitiform ventral median projection.	en	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
3C56F9A55B29EF714C40B1B7F0B9B6FB.taxon	diagnosis	Diagnosis. Gonyleptes fragilis resembles Gonyleptes horridus and Gonyleptes curvicornis because of the overall similarities of the features of the dorsal scutum. Gonyleptes fragilis can be distinguished from the other two species by the ocularium with a pair of spines (tubercles in Gonyleptes horridus and Gonyleptes curvicornis) and the pale yellowish spots around the tubercles on abdominal scutum.	en	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
3C56F9A55B29EF714C40B1B7F0B9B6FB.taxon	description	Description. Penis (Fig. 11 A-C): ventral plate inflated in the middle region, with a deep U-cleft on anterior margin (its distal tips convergent), 3 distal pairs of long, curved setae, 1 median pair of short setae, 4 basal pairs of spatulate setae. Glans sac slightly projected distally, with similar projection to a dorsal process. Stylus sigmoid, smooth. Ventral process robust, apex flabelliform with several small projections on distal margin.	en	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
3862CD1142AA3F98A5E664B4EC09C2BB.taxon	diagnosis	Diagnosis. Gonyleptes perlatus is very similar to Gonyleptes horridus (according to Kury 2003) but can be distinguished by the relatively larger tubercles on lateral margin of the do rsal scutum and the absence of a robust prolateral row of apophysis on male femur IV (present in Gonyleptes horridus), although this article might present variable armature.	en	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
EC30BA5D5DBB468147F7AFDC68512D52.taxon	diagnosis	Diagnosis. Gonyleptes pustulatus (Fig. 12 A-B) superficially resembles Gonyleptes bimaculatus because of the conspicuous white patches on mesotergum. Males of Gonyleptes pustulatus can be distinguished from Gonyleptes bimaculatus by: femur IV armed (unarmed in Gonyleptes bimaculatus) and presence of a bifid retrolateral apical apophysis on coxa IV.	en	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
EC30BA5D5DBB468147F7AFDC68512D52.taxon	description	Description. Penis (Fig. 12 C-D; MNRJ 17456): ventral plate with slightly inflated middle region, a deep distal U-cleft on anterior margin (its distal tips are divergent), 3 distal pairs of long, curved setae, 1 median pair of short setae, 4 basal pairs of spatulate setae. Stylus slightly straight, with apex slightly swollen. Ventral process robust, apex flabelliform with serrated and rounded distal margin.	en	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
C477EC0827EAA848715B1C99ACA5AB5B.taxon	diagnosis	Diagnosis. Mischonyx resembles Currala Roewer, 1927 and Schenkelibunus Strand, 1932, which are gonyleptine of relatively smaller size (about 5 mm of dorsal scutum length). Mischonyx can be distinguished from those genera by the combination of the following characters: anterior border of the carapace with robust spines; frontal hump on anterior margin of carapace and ocularium with a pair of spines; scutal areas I-III with a paramedian pair of tubercles, those on I-II elliptical and widest pair on scutal area III; lateral margin of the dorsal scutum with large white tubercles (in specimens preserved in 70 % ethanol).	en	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
DE6965D44E10B9B79055DA2350A09D97.taxon	diagnosis	Diagnosis. Mischonyx cuspidatus resembles Mischonyx poeta and Mischonyx scaber (see diagnosis of Mischonyx scaber). Mischonyx cuspidatus can be distinguished from them by the median high, acuminated tubercle on the free tergites I-III.	en	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
BE4673223C875761292F24387C2A45B2.taxon	diagnosis	Diagnosis. Mischonyx scaber resembles Mischonyx cuspidatus and Mischonyx poeta because of armature of the male femur IV: presence of a dorsobasal apophysis relatively short and curved; one retrolateral apophysis on distal third; and a row of high, acuminated tubercles increasing in size from the base to distal third. Mischonyx scaber can be distinguished from Mischonyx cuspidatus by the presence of a short gap between the retrolateral row and the retrolateral apophysis on femur IV (the retrolateral row is continuous to the retrolateral apophysis on femur IV in Mischonyx cuspidatus). Mischonyx scaber can be distinguished from Mischonyx poeta by the absence of enlarged tubercles (present in Mischonyx poeta) on the posterior part of the lateral margin of the dorsal scutum.	en	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
29E7A90639329AA1642DBC274E2D4090.taxon	description	Description. Penis (Fig. 13 A-C; MZSP 16837): ventral plate with slightly convergent lateral sides deep U-cleft on distal margin, its apex folded ventrally, 3 distal pairs of large setae, slightly curved at apex, 1 median pair of short setae and, 4 basal pairs of spatulate setae, directed frontwards. Glans with dorsal process. Stylus thin, long, sigmoid, with subapical ventral trichomes. Ventral process shaft folded ventrally, apex flabelliform, with serrate distal margins and long distal tip.	en	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
BDC701706E61516999256D31A4330AE9.taxon	diagnosis	Diagnosis. Parampheres resembles Acanthogonyleptes, Adhynastes Roewer, 1930 and Deltaspidium Roewer, 1927 by the presence of a more elongated prolateral apical apophysis on male coxa IV (greater length / basal diameter ratio of the apophysis) than that in Gonyleptes and related Gonyleptinae genera. Parampheres can be distinguished from Adhynastes and Deltaspidium by the scutal area III with tubercles (instead of spines). Parampheres can be distinghished from Acanthogonyleptes by the absence of a clearly defined dorsobasal apophysis on the male femur IV. Parampheres species bear a row of apophyses on the dorsal basal area of the femur IV.	en	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
A9326F98A6FF24DC1201463930754490.taxon	description	Description. Penis (Fig. 14 A-C; MZSP 14571): ventral plate with the lateral sides slightly convergent, deep U-cleft on distal margin, its apex folded ventrally and convergent, 3 distal pairs of setae curved on apex, 1 pair of short median setae, 4 basal pairs of setae arranged in a row, 2 pairs of ventral distal short setae. Glans with small dorsal process. Stylus slender, sigmoid, smooth. Ventral process apex flabelliform with serrate distal margin.	en	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
CD7076E26DBDCC91273BD7479D7C9F6C.taxon	description	Description. Penis (Fig. 15 A-C; MZSP 29075): ventral plate with the sides subparallel, with deep cleft on distal margin (U-cleft in ventral view and V-cleft in dorsal view; its apex convergent), 3 pairs of long helycoidal setae, 2 pairs of ventral distal short setae, 1 pair of median short setae, 4 pairs of spatulate, large setae arranged in an oblique row in lateral view, basal lobe short and projected dorsad. Stylus sigmoid, smooth. Ventral process shaft short and wide, apex flabelliform with slightly serrated distal margin.	en	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
1CDE6E195D5233C7F606C1C65E48E519.taxon	diagnosis	Diagnosis. See Mendes (2011: 463).	en	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
D6ABA4CEA1BC6568F5BA682E46A07366.taxon	diagnosis	Diagnosis. See Mendes (2011: 453). Composition. Pseudopucrolia discrepans (Roewer, 1943); Pseudopucrolia incerta (Mello-Leitao, 1928); Pseudopucrolia mutica (Perty, 1833) and Pseudopucrolia rugosa (Roewer, 1930).	en	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
43F1E56775DE7A8252700D512CAB0078.taxon	diagnosis	Diagnosis. See Acosta (2001: 59).	en	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
467B6CBC2F12ABD97E1E6C6CFE6D175C.taxon	diagnosis	Diagnosis. Acrographinotus ornatus can be easily distinguished from other species of the genus by the blister-like, enlarged, brown tubercles on whitish scutal areas I-IV; posterior margin of dorsal scutum and free tergites I-II with a row of enlarged tubercles and white spots on lateral and posterior margin of dorsal scutum and coxa IV.	en	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
467B6CBC2F12ABD97E1E6C6CFE6D175C.taxon	description	Description. Penis (Fig. 16 A-B; SMF 2881): ventral plate with 3 distal pairs of long setae, 0 - 2 median pairs of short setae, 4 - 5 basal pairs of setae. Stylus almost straight (apex slightly bent), smooth. Distal ventral process shaped as an ibis head.	en	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
39F9F39A3CD2D93E4F4849EC03CF93E9.taxon	diagnosis	Diagnosis. Gyndesops resembles the largest Pachylinae genus, Discocyrtus Holmberg, 1878, which presents an ocularium and scutal area III with paired armature, four scutal areas, scutal area II-IV and free tergites I-III unarmed. This combination of characters also occurs in genera such as Gyndoides Mello-Leitao, 1927, Lacronia Strand, 1942, Paraluederwaldtia Mello-Leitao, 1927 and Parapucrolia Roewer, 1917. Gyndesops can be distinguished from Lacronia and Parapucrolia by the coloration, which is uniformly auburn (Lacronia has green / yellow spots and / or stripes on dorsal scutum and Parapucrolia has white patches on dorsal scutum). It is very difficult to distinguish Gyndesops from the remaining genera, because no revision was ever made including those groups, and their monophyly is doubtful. It is possible that Gyndesops (as well as Gyndoides and Paraluederwaldtia) is a junior synonym of Discocyrtus, but we refrained ourselves from such a rash nomenclatural act.	en	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
5B85277898C1C02F927856B717A3C04D.taxon	description	Description. Penis (Fig. 17 A-B, holotype): ventral plate subrectangular, basal half of lateral margin projected laterad, distal margin slightly concave, 3 distal pairs of cyli ndrical setae, 1 pair of median setae (long left seta and moderate sized right seta), 3 basal pairs of cylindrical setae (median largest). Glans sac almost reaching distal margin of ventral plate. Stylus with swollen apex, smooth. Ventral process with large shaft, apex flabelliform with serrated lateral margins.	en	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
6F7FB1330433B939B9798750F52C8F74.taxon	diagnosis	Diagnosis. Haversia resembles Pachylinae with three scutal areas, such as Corralia Roewer, 1913, Diconospelta Canals, 1934, Graphinotus Koch, 1839, Huassampilia Roewer, 1913, Neogonyleptes Roewer, 1913, Oxapampeus Roewer, 1936, Spinivunus Roewer, 1943, Tumbesia Loman, 1899, Ubatubesia B. Soares, 1945, and two species of Sadocus Sorensen, 1886 (Sadocus conspicillatus and Sadocus polyacanthus). Haversia can be distinguished from those genera by the unarmed and smooth (i. e. without tubercles) scutal areas and the complex apophyses of male trochanter IV: three prolateral apophyses (anterior median, posterior median and an apical apophysis).	en	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
24ABA0A2DBD00DC7C296A64F0A91CC58.taxon	description	Description. Penis (Fig. 18 A-B; SMF 1320 / 8): ventral plate with lateral margins slightly concave, distal margin straight, 3 distal pairs of cylindrical, straight, long setae, 1 pair of median short setae, 2 pairs of basal cylindrical, straight, long setae. Glans sac projected dorsally, ventral process and stylus arising ventrally. Stylus long, slightly curved, with apical ventral and lateral trichomes. Ventral process swollen apically, with nailhead-like apex.	en	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
19B4FDD0EAB9EC825E32C8307E8180A2.taxon	diagnosis	Diagnosis. Based on external morphology, Oxapampeus resembles Pachylinae with three scutal areas (subfamily classical diagnosis is the presence of four areas) and scutal area III with a paramedian pair of spines, such as Corralia Roewer, 1913, Diconospelta Canals, 1934, Haversia Roewer, 1913, Huassampilia Roewer, 1913, Neogonyleptes Roewer, 1913, Spinivunus Roewer, 1943, and two species of Sadocus Sorensen, 1886 (Sadocus conspicillatus and Sadocus polyacanthus). However, Oxapampeus can be distinguished from those genera by the combination of the following characters: ocularium with a paramedian pair of pointed tubercles; frontal hump on anterior border of dorsal scutum moderately developed; scutal areas I-II each with a paramedian pair of enlarged tubercles; posterior margin of dorsal scutum straight; and free tergites I-III unarmed.	en	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
BA19CCEE6E345D0F1FC9E3453149BDBE.taxon	description	Description. Penis (Fig. 19 A-B; holotype): ventral plate with lateral and distal margins slightly concave, with 3 - 4 distal pairs of cylindrical, straight, long setae, 0 - 1 median seta (visible in lateral view), 5 basal pairs of setae. Glans sac projected dorsally, ventral process and stylus arising ventrally. Stylus short and straight, with ventral subapical trichomes. Ventral process swollen apically, more elongated and thinner than stylus.	en	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
