identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
BA7D4A9F85273E795210F834AA6190F3.text	BA7D4A9F85273E795210F834AA6190F3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nemastygnus Roewer 1929	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Nemastygnus Roewer, 1929 new family assignment</p>
            <p> Nemastygnus Roewer, 1929: 277; Kury 2003: 145; (type species:  Nemastygnus ovalis Roewer, 1929, by monotypy). </p>
            <p>Diagnosis.</p>
            <p> Nemastygnus was transferred from  Cranaidae to  Gonyleptidae ,  Metasarcinae by Kury (2003). It is closely related to  Avima Roewer, 1949, based on dorsal scutum shape (rectangular) and scutal area III (unarmed). It is impossible to distinguish both genera mainly because  Avima , the largest genus of  Leiosteninae with 33 species, is a heterogeneous genus based on penial characters. The type-species of  Avima ,  Avima leucobunus Roewer, 1949, is known only from the original description (and only external morphology). However, we will not propose synonymy in this paper because  Nemastygnus is older and 33 new combinations should be proposed. A review of  Avima plus  Nemastygnus species is needed. </p>
            <p>Description.</p>
            <p> Ocularium unarmed, saddle-shaped. Areas and posterior margin of dorsal scutum and free tergites unarmed (Fig. 1A). Scutal area I undivided. Lateral margin of dorsal scutum coriaceus. Chelicera dimorphic. Pedipalp with slender articles; femur without dorsoapical spine, with row of three ventral large setae and two ventroapical large setae; patella with prolateral large setae;  tibia–tarsus with ectal-mesal large setae. Leg I filiform, three-segmented. Penis: basal setae of truncus and basal setae of ventral plate very long and bifid; stylus with a large, longitudinal keel. </p>
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	https://treatment.plazi.org/id/BA7D4A9F85273E795210F834AA6190F3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pinto-da-Rocha, Ricardo;Benedetti, Alipio Rezende;de Vasconcelos, Eduardo Gomes;Hara, Marcos Ryotaro	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
9C262AEE532EB1D56E8C6DA826F87240.text	9C262AEE532EB1D56E8C6DA826F87240.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nemastygnus ovalis Roewer 1929	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Nemastygnus ovalis Roewer, 1929 Fig. 1 </p>
            <p> Nemastygnus ovalis Roewer, 1929: 277, fig. 44 (♂); Kury 2003: 145; (male holotype, Colombia, Cundinamarca,  Bogotá , SMF RI 1005/4, examined). </p>
            <p>Description.</p>
            <p> Penis (Fig. 1  B–C ; holotype): truncus with three pairs of bifid setae (basal, subdistal lateral and distal ventral). Ventral plate with rounded lobe on corners, two dorso-basal pairs of small single branched setae, a ventral distal pair of small single branched setae and a ventral median bifid setae. Glans dorsally projected, slender, with a small ventral crest and a long dorsal crest. Stylus smooth. </p>
            <p>Taxonomical note.</p>
            <p> Nemastygnus was originally placed in  Gonyleptidae ,  Prostygninae . Kury (1994b) transferred  Prostygninae to  Cranaidae . Later, in his catalogue, Kury (2003) transferred  Nemastygnus to  Gonyleptidae ,  Metasarcinae without examining the type material. We herein propose the transfer of  Nemastygnus to  Agoristenidae Leiosteninae , based on characteristics of body and male genitalia, viz., filiform leg I, saddle-shaped ocularium, pedipalpus with well developed setae in ventral row of femur (basalmost longest, size including socket about length of pedipalpal femur) and prolateral/retrolateral of tibia and tarsus, penis with typical ventral plate and bifid setae (Kury 1993; Pinto-da-Rocha 1996; Kury 1997; Pinto-da-Rocha and Hara 2009). </p>
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	https://treatment.plazi.org/id/9C262AEE532EB1D56E8C6DA826F87240	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pinto-da-Rocha, Ricardo;Benedetti, Alipio Rezende;de Vasconcelos, Eduardo Gomes;Hara, Marcos Ryotaro	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
5D6F1EB143BCDAD269631F67BE1C64C8.text	5D6F1EB143BCDAD269631F67BE1C64C8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Taulisa Roewer 1956	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Taulisa Roewer, 1956 new family assignment</p>
            <p> Taulisa Roewer, 1956: 433; Kury 2003: 145; (type species:  Taulisa koepckei Roewer, 1956, by original designation). </p>
            <p>Diagnosis.</p>
            <p> Taulisa (Fig. 2) differs from the other 10 genera of  Leiosteninae by the following combination of characters: ocularium unarmed, saddle-shaped; areas of dorsal scutum with two large tubercles each, posterior margin of dorsal scutum and free tergites, each one with a central large tubercle; area I undivided; lateral margin of dorsal scutum coriaceus; pedipalp with slender articles, its femur with dorsoapical spine, with row of four ventral large setae; pedipalpal patella with prolateral large setae; pedipalpal  tibia–tarsus with ectal-mesal large setae; leg I filiform, three-segmented. Male unknown. </p>
            <p>Taxonomical note.</p>
            <p> Taulisa was originally placed in  Phalangodidae ,  Tricommatinae . Kury (1992a) elevated  Tricommatinae to family level. In 2003, Kury transferred  Taulisa from  Tricommatinae to  Gonyleptidae ,  Metasarcinae without further remarks. We propose to transfer  Taulisa to  Agoristenidae ,  Leiosteninae based on characteristics of the body, viz., filiform leg I, saddle-shaped ocularium, pedipalpus with well developed and long setae in ventral row of femur and on prolateral/retrolateral of tibiae/tarsi (Kury 1993; Pinto-da-Rocha 1996; Kury 1997; Pinto-da-Rocha and Hara 2009).  Information regarding male genitalia was unavailable, since this species is only known from the female holotype. </p>
            <p>Composition.</p>
            <p>Monotypic.</p>
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	https://treatment.plazi.org/id/5D6F1EB143BCDAD269631F67BE1C64C8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pinto-da-Rocha, Ricardo;Benedetti, Alipio Rezende;de Vasconcelos, Eduardo Gomes;Hara, Marcos Ryotaro	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
E334DFD1D805B996F1DA3EE5205C551A.text	E334DFD1D805B996F1DA3EE5205C551A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Napostygnus Roewer 1929	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Napostygnus Roewer, 1929 new family assignment</p>
            <p> Napostygnus Roewer, 1929: 274, 275; Kury 2003: 145; (type species:  Napostygnus bispinosus Roewer, 1929, by monotypy). </p>
            <p>Diagnosis.</p>
            <p> Napostygnus was originally placed in  Gonyleptidae ,  Prostygninae and transferred to the  Gonyleptidae ,  Metasarcinae by Kury (2003). It differs from the other 75 genera of  Cranaidae by the following combination of characters: ocularium, scutal areas  I–IV , posterior margin of dorsal scutum and free tergite I unarmed; free tergites  II–III with one spine; posterior margin of dorsal scutum concave; legs thin and unarmed; penis with ventral plate wider at the middle, three pairs of distal setae, two pairs of basal setae, glans with dorsal process, stylus with apex enlarged. </p>
            <p>Composition.</p>
            <p>Monotypic.</p>
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	https://treatment.plazi.org/id/E334DFD1D805B996F1DA3EE5205C551A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pinto-da-Rocha, Ricardo;Benedetti, Alipio Rezende;de Vasconcelos, Eduardo Gomes;Hara, Marcos Ryotaro	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
77324B80A96E80D91D08AE8037767ADC.text	77324B80A96E80D91D08AE8037767ADC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Taulisa koepckei Roewer 1956	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Taulisa koepckei Roewer, 1956 Fig. 2 </p>
            <p> Taulisa koepckei Roewer, 1956: 433, fig. 3-4 (♀); Kury 2003: 145; (female holotype, Peru, Lambayeque, Hacienda Taulis, 6°50'S, 79°10'W, 1700 m, SMF 9697, examined). </p>
            <p>Diagnosis.</p>
            <p>As for genus.</p>
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	https://treatment.plazi.org/id/77324B80A96E80D91D08AE8037767ADC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pinto-da-Rocha, Ricardo;Benedetti, Alipio Rezende;de Vasconcelos, Eduardo Gomes;Hara, Marcos Ryotaro	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
0D9602A1EB4016A1A68A22D96B2CBA02.text	0D9602A1EB4016A1A68A22D96B2CBA02.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Napostygnus bispinosus Roewer 1929	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Napostygnus bispinosus Roewer, 1929 Fig. 3 </p>
            <p> Napostygnus bispinosus Roewer, 1929: 275, fig. 42 (♀); Kury 2003: 145; (female holotype, Ecuador, Napo, Valley of Rio Napo, SMF RI 1004/3, examined). </p>
            <p>Material examined.</p>
            <p> ECUADOR. Napo: Valley of Rio Napo, 1 female holotype (SMF RI 1004/3);  Cantón Quijos, Parroquira Cozanga, Yanayacu Research Station (0°35'S, 78°57'W, 2128 m), 1 ♂ &amp; 2 ♀ (MZSP 36132); ditto, 1 ♀ (IBSP 10550)  .</p>
            <p>Description.</p>
            <p> Male (MZSP 36132). Dorsum (Fig. 3A). Measurements: dorsal scutum length 4.1; dorsal scutum maximum width 3.5; carapace length 2.1; carapace maximum width 2.9; femur IV length 11.2. Body outline nearly subrectangular. Anterior margin of dorsal scutum with a median frontal hump small-granulate. Ocularium near middle of carapace, saddle shaped, with small granules near the eyes, unarmed. Carapace higher than the rest of dorsal scutum, with 4 tubercles behind ocularium. Scutal areas  I–III with 2 small median tubercles on each area; IV with 4 tubercles. Lateral margin of dorsal scutum with a low density of small granules. Posterior margin of dorsal scutum and free tergite I with a row of small granules, unarmed. Free tergites  II–III each with a median spine, small granulate. </p>
            <p>Chelicera: segment I unarmed. Segment II swollen, finger II with 3 teeth, III with 4 teeth.</p>
            <p>Pedipalpus: trochanter with 2 ventral tubercles, femur and patella smooth. Tibial setation: retrolateral iiIii; prolateral iiiIiii. Tarsal setation: retrolateral IiIi; prolateral IiiiIi.</p>
            <p> Legs: legs  I–IV unarmed and without granules, except for trochanters, which are small granulate. Basitarsus I slightly inflated. Tarsal process present. Tarsal segmentation: 6(3); 16(3); 6; 7. </p>
            <p> Penis (Fig. 3  B–C ): ventral plate with almost straight distal margin, thick median lobe and folded ventrally to the distal setae, 3 pairs of distal setae and 2 pairs of basal setae. Glans with thumb-like dorsal process. Stylus with dorsal apical projection and ventral apical small trichomes. </p>
            <p> Coloration (in ethanol) (Fig. 3A): body background yellow with brown spots mainly on carapace, scutal areas, lateral and posterior margins of dorsal scutum and free tergites. Mesotergum with one longitudinal yellow stripe surrounded by blackish pigment at grooves  I–V . Pedipalpus and chelicera yellowish brown with a brown reticulate pattern. Legs yellowish brown. </p>
            <p>Taxonomical note.</p>
            <p> Napostygnus was originally placed in  Gonyleptidae ,  Prostygninae . Kury (1994b) transferred  Prostygninae to  Cranaidae and later transferred  Napostygnus to  Gonyleptidae ,  Metasarcinae (Kury 2003). We herein propose the removal of  Napostygnus from  Metasarcinae based on male genitalia, which does not present the diagnostic character for the subfamily (Kury and Maury 1998; Kury and Pinto-da-Rocha 2007b), viz., a pair of spiny laterobasal sacs on the ventral plate. Another remarkable difference from  Metasarcinae is the unarmed ventral pedipalpus femur. The combination of characteristics of penis and body morphology does not allow placing  Napostygnus bispinosus in any other family of Laniatores which bear tarsal process on legs  III–IV . The slightly swollen male basitarsus I is not equal to those of  Manaosbiidae , and furthermore the genitalia is distinct, since it possesses a dorsal process. The ocularium resembles those of  Gonyleptidae ,  Bourguyiinae , but the penis is not similar, because there is only a dorsal process and no ventral process. The aspect of the penis resembles those of  Cranaidae (presence of dorsal process, ventral plate and setae shape) and therefore, we propose its transfer to this family. It is noteworthy to mention that the morphology of the body of  Napostygnus bispinosus is somewhat distinct from the typical  Cranaidae . We will not assign it to any subfamily, following the opinion of Orrico and Kury (2009) on the meaninglessness of current subfamily classifications in  Cranaidae . </p>
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	https://treatment.plazi.org/id/0D9602A1EB4016A1A68A22D96B2CBA02	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pinto-da-Rocha, Ricardo;Benedetti, Alipio Rezende;de Vasconcelos, Eduardo Gomes;Hara, Marcos Ryotaro	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
8F100F3A3D4C034E369EF464C87CB5D3.text	8F100F3A3D4C034E369EF464C87CB5D3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neopachyloides peruvianus (Roewer 1956) Roewer 1956	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Neopachyloides peruvianus (Roewer, 1956) new comb., new subfamily assignment Fig. 4 </p>
            <p> Ceropachylinus peruvianus Roewer, 1956: 440, fig. 19 (♂); Kury 2003: 158; (male holotype, Peru, near San Luis de Shuaro, 700-750 m, Koepcke leg, 17.III.1955, SMF 9791/1, examined). </p>
            <p>Diagnosis.</p>
            <p> Neopachyloides was a hitherto monotypic genus that resembles  Ampycinae genera with a paired armature on ocularium and free tergite III with a median, long spiniform apophysis asin  Ampycella Roewer, 1929,  Ampycus Simon, 1879,  Hutamaia Soares &amp; Soares, 1977 and  Sibollus Roewer, 1929.  Neopachyloides can be distinguished from these genera by the following combination of characteristics: scutal area III with a paramedian pair of enlarged, pointed tubercles, scutal area IV undivided and free tergite II unarmed.  Neopachyloides peruvianus can be distinguished from  Neopachyloides spinipes Roewer, 1913 by the dorsal scutum covered by granules and scutal areas  I–II unarmed (Fig. 4A). </p>
            <p>Description.</p>
            <p> Penis (Fig. 4  B–C ; holotype): ventral plate with sub hexagonal shape, deep cleft on distal margin, 4 pairs of distal setae (distalmost curved and basalmost small), 3 pairs of basal setae straight. Glans very long (⅔ of ventral plate length), stylus smooth and curved dorsally, without dorsal and ventral processes. </p>
            <p>Taxonomical note.</p>
            <p> The assignment of  Neopachyloides peruvianus to  Neopachyloides is based on overall similarity and should be considered tentative.  Neopachyloides peruvianus is the only  Ampycinae which presents just one scutal area armed with a paramedian enlarged pair of tubercles (most genera present 3 scutal areas armed with a paramedian pair of enlarged tubercles, as  Ampycus ,  Hexabunus and  Pirunipygus or all of them unarmed). We preferred to place  Neopachyloides peruvianus under  Neopachyloides instead of proposing a new genus because monophyly of  Ampycinae genera (most of them monotypic) are doubtful. See taxonomical note of  Pirunipygus paradoxus for the reasons of the new placement of  Neopachyloides peruvianus . </p>
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	https://treatment.plazi.org/id/8F100F3A3D4C034E369EF464C87CB5D3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pinto-da-Rocha, Ricardo;Benedetti, Alipio Rezende;de Vasconcelos, Eduardo Gomes;Hara, Marcos Ryotaro	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
E315E5F58F07063706EFE43F502FBAC6.text	E315E5F58F07063706EFE43F502FBAC6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pirunipygus Roewer 1936	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Pirunipygus Roewer, 1936 new subfamily assignment</p>
            <p> Pirunipygus Roewer, 1936: 341; Kury 2003: 187; (type species:  Pirunipygus paradoxus Roewer, 1936, by monotypy). </p>
            <p>Diagnosis.</p>
            <p> Pirunipygus is a monotypic genus and resembles  Ampycinae genera with a paramedian pair of enlarged tubercles on scutal areas  I–II as  Ampycus Simon, 1879,  Hexabunus Roewer, 1913,  Neopachyloides Roewer, 1913 and  Parahernandria Go  odnight &amp; Goodnight, 1947.  Pirunipygus paradoxus can be distinguished from these genera by the following combination of characters: ocularium with a median spiniform apophysis (Fig. 5A); dorsal scutum with secondary tubercles (sensu Maury 1992); scutal area III with a paramedian pair of spiniform apophyses; free tergite II with a median spiniform apophysis; free tergite III with three spiniform apophyses, the middle one largest and bifid. </p>
            <p>Composition.</p>
            <p>Monotypic.</p>
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	https://treatment.plazi.org/id/E315E5F58F07063706EFE43F502FBAC6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pinto-da-Rocha, Ricardo;Benedetti, Alipio Rezende;de Vasconcelos, Eduardo Gomes;Hara, Marcos Ryotaro	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
5EB53246070FE5B439C049CE6B6B3E65.text	5EB53246070FE5B439C049CE6B6B3E65.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acanthogonyleptes Mello-Leitao 1922	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> 
Acanthogonyleptes 
Mello-Leitao
, 1922
</p>
            <p> Acanthogonyleptes Mello-Leitão , 1922: 336; Kury 2003:137; DaSilva and Pinto-da-Rocha 2010: 626; (type species:  Acanthogonyleptes pulcher Mello-Leitão , 1922, by original designation). </p>
            <p> Centroleptes Roewer, 1943: 45; Kury 2003: 122; (type species:  Centroleptes flavus Roewer, 1943, by monotypy). Syn. n. </p>
            <p>Diagnosis.</p>
            <p>As in Kury (2003: 137).</p>
            <p>Taxonomical note.</p>
            <p> The monotypic genus  Centroleptes is a subjective synonym of  Acanthogonyleptes , since the type examined material of  Centroleptes flavus and  Acanthogonyleptes singularis are identical and synonymized here. </p>
            <p>Composition.</p>
            <p> Acanthogonyleptes alticola (  Mello-Leitão , 1922);  Acanthogonyleptes editus (Roewer, 1943);  Acanthogonyleptes fallax (  Mello-Leitão , 1932);  Acanthogonyleptes fulvigranulatus (  Mello-Leitão , 1922);  Acanthogonyleptes marmoratus (  Mello-Leitão , 1940);  Acanthogonyleptes pictus (Piza, 1942);  Acanthogonyleptes singularis (  Mello-Leitão , 1935);  Acanthogonyleptes soaresi (  Mello-Leitão , 1944);  Acanthogonyleptes variolosus (  Mello-Leitão , 1940). </p>
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	https://treatment.plazi.org/id/5EB53246070FE5B439C049CE6B6B3E65	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pinto-da-Rocha, Ricardo;Benedetti, Alipio Rezende;de Vasconcelos, Eduardo Gomes;Hara, Marcos Ryotaro	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
CC5133B5F0FE1F303C44DF2BFF5FB97C.text	CC5133B5F0FE1F303C44DF2BFF5FB97C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acanthogonyleptes singularis (Mello-Leitao 1935) Mello-Leitao 1935	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> 
Acanthogonyleptes singularis (
Mello-Leitao
, 1935)
</p>
            <p> Adelphobunus singularis Mello-Leitão , 1935: 392, fig. 19 (♂); Kury 2003: 137; (male holotype; Brazil,  São Paulo,  Ribeirão Pires, IBSP 17, examined). </p>
            <p> Sphaerobunus singularis : Kury 2003: 139. </p>
            <p> Acanthogonyleptes singularis : DaSilva and Pinto-da-Rocha 2010: 626. </p>
            <p> Bocaina marmorata Piza, 1943: 46, fig. 5 (♂); Kury 2003: 139; (male holotype, Brazil,  São Paulo, Serra da  Bocaina , Fazenda  Águas de Santa Rosa, MZSP 810, examined). Synonymy established by Soares and Soares (1987). </p>
            <p> Centroleptes flavus Roewer, 1943: 45, fig. 50 (♀); Kury 2003: 122; (female holotype, Brazil, Santa  Catarina , Seara, Nova Teutonia, SMF 6430/63, examined). Syn. n. </p>
            <p>Material examined.</p>
            <p> BRAZIL.  São Paulo:  São José do Barreiro (Serra da  Bocaina , Fazendas  Águas de Santa Rosa), female holotype of  Bocaina marmorata Piza (MZSP 810);  Ribeirão Pires, male holotype of  Adelphobunus singularis Mello-Leitão (IBSP 17);  São José do Barreiro, 1 ♂ (HEMS 761); Santo  André , 1 ♂ &amp; 1 ♀ (MZSP 21257). Santa  Catarina : Seara (Nova  Teutônia ), ♀ holotype of  Centroleptes flavus Roewer (SMF 6430/63). </p>
            <p>Diagnosis.</p>
            <p> Acanthogonyleptes singularis and  Acanthogonyleptes variolosus can be distinguished from the remaining species of the genus by the presence of a median single elevation on scutal area III. Males of  Acanthogonyleptes singularis can be distinguished from those of  Acanthogonyleptes variolosus by the presence of a robust curved retrolateral apophysis on trochanter IV. </p>
            <p>Taxonomical note.</p>
            <p> The genus  Acanthogonyleptes is a morphologically homogeneous group, probably representing a monophyletic group (see  Sphaerobunus in Kury 2003).  Acanthogonyleptes singularis and  Centroleptes flavus bear a median single elevation on scutal area III. In addition, the morphology of female leg IV of both  Acanthogonyleptes singularis and  Centroleptes flavus are identical. The geographical records of  Acanthogonyleptes singularis include localities in the Brazilian states of  São Paulo and Rio de Janeiro. The only known specimen identified as  Centroleptes flavus is the holotype, supposedly collected in Seara (Nova  Teutônia ), and therefore the locality cited by Roewer (1943) is considered wrong. </p>
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	https://treatment.plazi.org/id/CC5133B5F0FE1F303C44DF2BFF5FB97C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pinto-da-Rocha, Ricardo;Benedetti, Alipio Rezende;de Vasconcelos, Eduardo Gomes;Hara, Marcos Ryotaro	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
1084F131AFA4A2CD376292E57EEAF2B4.text	1084F131AFA4A2CD376292E57EEAF2B4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pirunipygus paradoxus Roewer 1936	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Pirunipygus paradoxus Roewer, 1936 Fig. 5 </p>
            <p> Pirunipygus paradoxus Roewer, 1936: 341, fig. 4 (♂); Kury 2003: 106; (male holotype, Peru, Junin, Tarma, SMF 6180/85, examined). </p>
            <p>Description.</p>
            <p> Penis (Fig. 5  B–C ; holotype): ventral plate sub hexagonal, deep U-cleft on distal margin, 3-4 distal pairs of setae (2-3 distalmost curved), 2 pairs of dorsal median setae of moderate size, 2 pairs of ventral median small setae. Stylus wide on basal  ¾ , apex twisted, with a patch of ventral subapical trichomes and some scattered apically. Glans without dorsal and ventral processes. </p>
            <p>Taxonomical note.</p>
            <p> Both  Neopachyloides peruvianus and  Pirunipygus paradoxus were formerly placed in  Gonyleptidae ,  Pachylinae . These two species show the following diagnostic characteristics of  Ampycinae (Kury 2003), viz., body coloration black, integument with huge rounded tubercles, stout apophysis on free tergite III, ventral plate of penis with deep U-shape cleft on distal margin, absence of dorsal and ventral processes of glans. </p>
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	https://treatment.plazi.org/id/1084F131AFA4A2CD376292E57EEAF2B4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pinto-da-Rocha, Ricardo;Benedetti, Alipio Rezende;de Vasconcelos, Eduardo Gomes;Hara, Marcos Ryotaro	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
92D62780ADB1EC34EE94ECDF69607E38.text	92D62780ADB1EC34EE94ECDF69607E38.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Geraeocormobius sylvarum Holmberg 1887	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Geraeocormobius sylvarum Holmberg, 1887</p>
            <p> Geraecormobius sylvarum Holmberg, 1887: 211, fig. unnumbered (♂); Kury 2003: 125; (1 male &amp; 1 female syntypes,  Argentina , Misiones, Santa Ana, near river  Piraí Mirín , type depository unknown)  .</p>
            <p> Discocyrtus serrifemur Roewer, 1943: 21, fig. 10 (♂); (1 male &amp; 8 female syntypes, Brazil, Santa  Catarina , Nova  Teutônia , SMF 6193/104, examined). Syn. n. </p>
            <p> Geraeocormobius sylvarum Acosta et al. 2007a: 303, figs 1-3 (map, ♂♀). </p>
            <p>Other material examined.</p>
            <p> Brazil.  Paraná : Cachoeirinha, 2 ♂ &amp; 5 ♀ (MNRJ 26912);  Paranavaí , 6 ♂ &amp; 7 ♀ (MNRJ 4732). </p>
            <p>Taxonomical note.</p>
            <p> Roewer (1943) placed  Discocyrtus serrifemur in the  Pachylinae based on the observation of pigmentation on posterior part of dorsal scutum, which he assumed erroneously to be the scutal area IV. Both sexes of type series of  Discocyrtus serrifemur are identical to  Geraeocormobius sylvarum . In general, females of  Gonyleptinae do not show conspicuous diagnostic characters, which are mainly seen in male tegumentary ornamentation. Both male and female specimens of  Geraeocormobius sylvarum preserved in 70% alcohol present a general black coloration with olive or yellowish lateral marks on the dorsal scutum, with white pedipalps.  Geraeocormobius sylvarum could be confused with  Geraeocormobius rohri (  Mello-Leitão ) and  Geraeocormobius salebrosus (Roewer) because of its relatively larger body size, presence of large rounded tubercles on the scutal area III, general black coloration and geographical occurrence. However,  Geraeocormobius sylvarum can be distinguished from them by the white pedipalps. </p>
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	https://treatment.plazi.org/id/92D62780ADB1EC34EE94ECDF69607E38	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pinto-da-Rocha, Ricardo;Benedetti, Alipio Rezende;de Vasconcelos, Eduardo Gomes;Hara, Marcos Ryotaro	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
BC9A736C5A776F4FA43BB981351D6CA6.text	BC9A736C5A776F4FA43BB981351D6CA6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gonyleptellus bimaculatus (Sorensen 1884) Sorensen 1884	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> 
Gonyleptellus bimaculatus (
Sorensen
, 1884)
 comb. n.</p>
            <p> Gonyleptes bimaculatus Sørensen , 1884: 605; (female holotype, Brazil, ZMUC, examined from digital photos). </p>
            <p> Paragonyleptes bimaculatus : Roewer 1913: 243. </p>
            <p> Gonyleptes cancellatus Roewer, 1917: 127, fig. 26 (♂); (male holotype, Brazil,  São Paulo, Santos, SMF 1320, examined). Syn. n. </p>
            <p> Gonyleptellus cancellatus : Kury 2003: 126. </p>
            <p> "Paragonyleptes" bimaculatus : Kury 2003: 121. </p>
            <p>Material examined.</p>
            <p> BRAZIL. Locality not specified further, female holotype of  Gonyleptes bimaculatus , examined by digital photos (ZMUC).  São Paulo: Santos, ♂ holotype of  Gonyleptes cancellatus (SMF 1320); Bananal (  Estação Ecológica do Bananal), A. Monteiro leg., IV.2004, 10 ♂ &amp; 10 ♀ (MZSP 27804). Rio de Janeiro: Itatiaia (Parque Nacional do Itatiaia), Equipe Biota leg., 8-15.VI.2001, 3 ♂ &amp; 1 ♀ (MZSP 21746);  Terezópolis (Parque Nacional da Serra dos  Órgãos , sede), R. Pinto-da-Rocha, S. Outeda-Jorge, C. Mattoni leg., 10.II.2007, 3 ♂ &amp; 5 ♀ (MZSP 28223); Nova Friburgo (Mury, Debossan, 950 m), R.S.  Bérnils &amp; P. Labiak leg., 29.VII.1996, 6 ♂ &amp; 3 ♀ (MZSP 15120). </p>
            <p>Taxonomical note.</p>
            <p> Sørensen (1884) described  Gonyleptes bimaculatus from Brazil without any illustration. Roewer (1913) transferred it to  Paragonyleptes Roewer,  1913 . Kury (2003) synonymized the type species (  Gonyleptes bicuspidatus Koch, 1839) of  Paragonyleptes with  Collonychium bicuspidatum Bertkau, 1880 and considered some species allocated in  Paragonyleptes as  Gonyleptinae incertae sedis, such as  Gonyleptellus bimaculatus . The examination of  Gonyleptellus bimaculatus from detailed photos of the holotype, which is reasonably well preserved, allowed us to recognize it as  Gonyleptellus cancellatus , a well-known species from the Brazilian states of Rio de Janeiro and  São Paulo (Serra da Bocaina mountain range). Therefore,  Gonyleptellus cancellatus is a junior synonym of  Gonyleptellus bimaculatus . The other species of  Gonyleptellus is  Gonyleptellus bufoninus (  Mello-Leitão , 1949), known only by its original description, which lacks illustration and its type depository is unknown (Kury 2003). </p>
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	https://treatment.plazi.org/id/BC9A736C5A776F4FA43BB981351D6CA6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pinto-da-Rocha, Ricardo;Benedetti, Alipio Rezende;de Vasconcelos, Eduardo Gomes;Hara, Marcos Ryotaro	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
F67722BD0705688E10A9D0F9615A933F.text	F67722BD0705688E10A9D0F9615A933F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gonyleptes Kirby 1819	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Gonyleptes Kirby, 1819</p>
            <p> Gonyleptes Kirby, 1819: 450; Kury 2003: 126; (type species:  Gonyleptes horridus Kirby, 1819, by subsequent designation, Roewer 1913). </p>
            <p> Collonychium Bertkau, 1880: 108; Kury 2003: 122; (type species:  Collonychium bicuspidatum Bertkau, 1880, by monotypy). Syn. n. </p>
            <p>Diagnosis.</p>
            <p> Gonyleptes is a genus composed of generally large species (dorsal scutum length longer than 8 mm), which males present femur IV with conspicuous armature and scutal areas  I–III convex in lateral view. The penis presents ventral plate very convex and apex of ventral process serrate and semicircular. See taxonomical note below. </p>
            <p>Taxonomical note.</p>
            <p> Gonyleptes possesses 23 species and its habitus resembles other genera placed in  Gonyleptinae . As Kury (2003) already suggested, the generic boundaries are not so clear cut.  Collonychium is an objective synonym of  Gonyleptes since both type species are synonymous. It can be only distinguished from related taxa, such as  Geraeocormobius Holmberg, 1887, by a combination of unsatisfactory Roewerian characteristics. The composition of the genus below includes the changes proposed in this article. </p>
            <p> Composition.  Gonyleptes acanthopus (Quoy &amp; Gaimard, 1824);  Gonyleptes armatus Perty, 1833;  Gonyleptes atrus Mello-Leitão , 1923, revalidated;  Gonyleptes barbiellinii Mello-Leitão , 1932;  Gonyleptes calcaripes (Roewer, 1917);  Gonyleptes curticornis (  Mello-Leitão , 1940);  Gonyleptes curvicornis (Roewer, 1932), revalidated, comb. n.;  Gonyleptes fragilis Mello-Leitão , 1923;  Gonyleptes gertschi Soares &amp; Soares, 1948;  Gonyleptes gonyleptoides (Soares &amp; Soares, 1945);  Gonyleptes granulatus (Piza, 1940);  Gonyleptes horridus Kirby, 1819;  Gonyleptes parcigranulatus Soares &amp; Soares, 1949;  Gonyleptes pectinatus Koch, 1845;  Gonyleptes pectinipes Roewer, 1917;  Gonyleptes perlatus (  Mello-Leitão , 1935), comb. n.;  Gonyleptes pseudogranulatus Soares &amp; Soares, 1946;  Gonyleptes pseudoguttatus Giltay, 1928;  Gonyleptes pustulatus Sørensen , 1884;  Gonyleptes recentissimus Mello-Leitão , 1932;  Gonyleptes saprophilus Mello-Leitão , 1922;  Gonyleptes vatius Bertkau, 1880;  Gonyleptes viridisagittatus Soares &amp; Soares, 1945. </p>
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	https://treatment.plazi.org/id/F67722BD0705688E10A9D0F9615A933F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pinto-da-Rocha, Ricardo;Benedetti, Alipio Rezende;de Vasconcelos, Eduardo Gomes;Hara, Marcos Ryotaro	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
7DC0E0AE964DD95B57CE8361E7CDBA12.text	7DC0E0AE964DD95B57CE8361E7CDBA12.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gonyleptes horridus Kirby 1819	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Gonyleptes horridus Kirby, 1819 Figs 6  A–D 7 </p>
            <p> Gonyleptes horridus Kirby, 1819: 452, pl. 22, fig. 16 (♂); Kury 2003: 128; (male holotype, Brazil, NHM 1863.41, examined from detailed photos). </p>
            <p> Gonyleptes bicuspidatus Koch, 1839: 13; (2 females syntypes, Brazil, NHMW, not examined; 2 males &amp; 2 females syntypes, Brazil, ZMB 905, not examined). Syn. n. </p>
            <p> Collonychium bicuspidatum Bertkau, 1880: 108, pl. 2, fig. 39 (♀); (female nymph holotype, Brazil, Rio de Janeiro, Rio de Janeiro [Copacabana], ISNB, not examined). Syn. n. </p>
            <p> Metagonyleptes hamatus Roewer, 1913: 213, fig. 89 (♂); (male holotype, Brazil,  São Paulo, SMF 891, examined). Syn. n. </p>
            <p> Paragonyleptes simoni Roewer, 1930: 379, fig. 12 (♂); (male holotype, Brazil, Santa Catarina, Serra Azul, SMF RII 1333, examined). Syn. n. </p>
            <p> Gonyleptes borgmeyeri Mello-Leitão , 1932: 305, fig. 167 (♂); (male holotype, MNRJ, lost, Brazil, Rio de Janeiro,  Petrópolis ). Syn. n. </p>
            <p> Gonyleptes curvicornis Mello-Leitão , 1932: 305, fig. 166 (♂); (male holotype, MNRJ, lost, Brazil, Rio de Janeiro, Itatiaia). Syn. n. </p>
            <p> Gonyleptes melloleitaoi Kury &amp; Alonso-Zarazaga, 2011: 54 (replacement name for  Gonyleptes curvicornis Mello-Leitão , secondary homonym of  Weyhia curvicornis Roewer). </p>
            <p>Material examined.</p>
            <p> BRAZIL. Locality not specified further, male holotype of  Gonyleptes horridus , examined by digital photos (NHM 1863.41).  São Paulo:  São Paulo, male holotype of  Metagonyleptes hamatus (SMF 891). Rio de Janeiro: Rio de Janeiro (  Gávea ), 1 male (MZSP 1370). Santa Catarina: (Serra Azul), male holotype of  Paragonyleptes simoni (SMF RII 1333). </p>
            <p>Diagnosis.</p>
            <p> Gonyleptes horridus can be recognized by: the relatively larger size; anterior border of carapace smooth (without tubercles or spines in the corners); frontal hump on anterior border of carapace and ocularium with white small tubercles; prolateral apical apophysis of coxa IV long, with abrupt curvature backwards.  Gonyleptes horridus ,  Gonyleptes perlatus and  Gonyleptes pustulatus can be distinguished from all other species of  Gonyleptes by the robust and bifid (two distal tips) retrolateral apical apophysis of coxa IV. Females of  Gonyleptes horridus and  Gonyleptes perlatus present dimorphic sulfur yellow spines on the free tergites, not present in females of  Gonyleptes pustulatus .  Gonyleptes horridus can be distinguished from  Gonyleptes perlatus by the armature pattern of femur IV of the males (see note on  Gonyleptes perlatus below).  Gonyleptes horridus has relatively smaller tubercles on posterior part of the lateral margins of abdominal scutum than in  Gonyleptes perlatus . </p>
            <p>Description.</p>
            <p> Dorsal scutum length 7-9 mm. Anterior margin of carapace without tubercles or spines. Median frontal hump on anterior margin of carapace and ocularium with a pair of tubercles on each one (Fig. 6A). Four scutal grooves delimiting three scutal areas. Scutal areas I, II with low white tubercles in the median region; III with high paramedian tubercles. Lateral margin of dorsal scutum with flattened white  tubercles . Pedipalps approximately same length as dorsal scutum. Prolateral apophyses of coxa IV elongated, strongly curved backwards, dorsal branch longest. Retrolateral apophysis of coxa IV bifid. Femur IV straight, with dorsobasal apophyses and longitudinal rows of spines (Fig. 6  B–C ). Penis with typical  Gonyleptinae -like pattern (Kury 1992b). Tarsal segmentation: 6(3); 10-12(3); 7-8; 8-9. Females with dimorphic cones (one yellowish huge median spine) on free tergites II and III. General coloration brown with white tubercles (Fig. 6D). Penis (Fig. 7  A–C ; MZSP 15496): ventral plate with a deep U-cleft on anterior margin (its distal tips convergent), 3 distal, 1 median and 4 basal pairs of setae, ventrally with a subdistal and a median pair of setae. Stylus sigmoid, with ventral subapical trichomes. Ventral process robust, apex as a flabellum with several small projections on distal margin. </p>
            <p>Taxonomical note.</p>
            <p> All synonyms given in Kury (2003) for  Gonyleptes horridus refer, in fact, to  Gonyleptes curvicornis (Roewer, 1913). The type material of  Gonyleptes curvicornis Mello-Leitão , 1932 was lost, but the original descriptions and illustrations are sufficient to propose the present synonymy. The types of  Metagonyleptes hamatus Roewer and  Paragonyleptes simoni Roewer are identical to that of  Gonyleptes horridus Kirby. The record of  Gonyleptes horridus in Santa Catarina (type locality of  Paragonyleptes simoni) is dubious. The synonymy of  Gonyleptes bicuspidatus with  Gonyleptes horridus is here proposed based on the reasoning of Kury (2003), who considered this nominal species as synonym of  Collonychium bicuspidatum . In turn,  Collonychium bicuspidatum is clearly a synonym of  Gonyleptes horridus . </p>
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	https://treatment.plazi.org/id/7DC0E0AE964DD95B57CE8361E7CDBA12	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pinto-da-Rocha, Ricardo;Benedetti, Alipio Rezende;de Vasconcelos, Eduardo Gomes;Hara, Marcos Ryotaro	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
92484EAA302D83651B3C9287FF39815C.text	92484EAA302D83651B3C9287FF39815C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gonyleptes curvicornis (Roewer 1913) Roewer 1913	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Gonyleptes curvicornis (Roewer, 1913) revalidated, comb. n. Figs 6  E–H 8 </p>
            <p> Weyhia curvicornis Roewer, 1913: 193, fig. 80 (♂); Kury 2003: 128; (holotype male, Brazil,  São Paulo, SMF 979, examined). </p>
            <p> Gonyleptes lacrimosus Mello-Leitão , 1932: 294, fig. 148 (♂); (male holotype, Brazil, Rio de Janeiro (Floresta da Tijuca), MNRJ 11789, not examined)  .</p>
            <p> Mendesius albipunctatus Roewer, 1943: 41, pl. 5, fig. 45 (♀); (female holotype, Brazil, Mendes, SMF 5389/47). </p>
            <p>Material examined.</p>
            <p> BRAZIL.  São Paulo: Locality not specified further, male holotype (SMF 979);  São Paulo (  Chácara Dr. J. L. Lane), male (MZSP 0871). Rio de Janeiro: Rio de Janeiro (  Grajaú ), 1 male (MZSP 1117). </p>
            <p>Diagnosis.</p>
            <p> Gonyleptes curvicornis resembles  Gonyleptes horridus (see diagnosis above) but can be distinguished by the relatively shorter prolateral apical apophysis of coxa IV, lacking the abrupt backwards curvature (Fig. 6E). The retrolateral apical apophysis of coxa IV is reduced and not bifid as in  Gonyleptes horridus . The male femur IV has a different pattern of armature (Fig. 6  F–G ) compared to  Gonyleptes horridus . Carapace with a pair of tubercles on the posterior area (posterior to ocularium). Females of  Gonyleptes curvicornis do not present dimorphic sulfur yellow spines on the free tergites. </p>
            <p>Description.</p>
            <p> Penis (Fig. 8  A–B ; MZSP 1117): ventral plate sub hexagonal, with a deep U-cleft on distal margin (its distal tips convergent), 3 distal, 1 median, 4 basal pairs of setae (distal ones curved, median ones shortest). Stylus sigmoid, with ventral subapical trichomes. Ventral process robust, apex flabelliform with several projections of varying sizes on distal margin. </p>
            <p>Taxonomical note.</p>
            <p> Roewer’s (1913; 1923) redescriptions of  Gonyleptes horridus and  Weyhia curvicornis are the same. Following  Gonyleptes horridus sensu Roewer, Soares and Soares (1988) considered  Weyhia curvicornis as its junior synonym. However, as seen above, such redescription of  Gonyleptes horridus was mistaken and, therefore,  Weyhia curvicornis can no longer be considered as a junior synonym.  Weyhia was synonymized with  Geraeocormobius by Soares and Soares (1949). The  Gonyleptinae genera definition is still unsatisfactory, but the placement of  Weyhia curvicornis in  Gonyleptes is more reasonable than its placement in  Geraeocormobius , due to morphological similarities shared with  Gonyleptes horridus , such as white tubercles on frontal hump on anterior margin of dorsal scutum, ocularium (Fig. 6H) and lateral borders of abdominal scutum convex and abdominal scutum itself convex. </p>
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	https://treatment.plazi.org/id/92484EAA302D83651B3C9287FF39815C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pinto-da-Rocha, Ricardo;Benedetti, Alipio Rezende;de Vasconcelos, Eduardo Gomes;Hara, Marcos Ryotaro	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
E67535A551E03FB4CB82811E01881AE7.text	E67535A551E03FB4CB82811E01881AE7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gonyleptes atrus Mello-Leitao 1923	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> 
Gonyleptes atrus 
Mello-Leitao
, 1923
 revalidated Figs 9  A–B 10 </p>
            <p> Gonyleptes atrus Mello-Leitão , 1923: 140; Kury 2003: 129; (male syntype, Brazil, Itatiaia, state of Rio de Janeiro or Minas Gerais, MNRJ 1462, examined; 3 males &amp; 3 females syntypes, MZSP 42). </p>
            <p> Weyhia brieni Giltay, 1928: 83; (male holotype; Brazil, Itatiaia; ISNB; not examined). Syn. n. </p>
            <p> Geraeocormobius jimi Soares &amp; Soares, 1974: 601, figs 8-25 (♂♀); (male holotype and 1 female paratype; Brazil, Rio de Janeiro; Itatiaia; J. Jim et al. leg, 28.XI.1977; HSPC 447; 2 males paratypes; same locality; A. Peracchi &amp; E. Izecksohn leg, 9.IV.1966; HSPC 463; examined). Syn. n. </p>
            <p>Material examined.</p>
            <p> BRAZIL. Minas Gerais: Lambari (Parque Estadual Nova Baden), (MZSP 32513); Delfim Moreira (MZSP 29393); Itamonte (MZSP 21251);  Poços de Caldas (MZSP 29366). Rio de Janeiro: Itatiaia, syntypes of  Weyhia bisignata (MNRJ 27321).  São Paulo: Campos do  Jordão (MZSP 21251). </p>
            <p>Diagnosis.</p>
            <p> Dorsal scutum length 8-10 mm. Corners of anterior margin of carapace smooth. Median frontal hump on anterior margin of carapace and ocularium with one pair of tubercles each one. Posterior region of carapace with 2 median tubercles. Mesotergum densely covered by tubercles. Scutal areas  I–III each with a pair of median enlarged tubercles (III with largest and round ones). Prolateral apical apophysis of male coxa IV slightly directed backwards, retrolateral apophysis of male coxa IV absent or very reduced (Fig. 9  A–B ). Dorsobasal apophysis of male femur IV robust, curved and retrolaterally oriented. Ornamentation of male femur IV variable, normally with two prolateral spines. </p>
            <p>Description.</p>
            <p> Penis (Fig. 10  A–C , MZSP 29393): ventral plate with 3 distal and 3 basal pairs of large setae, 1 median pair of short setae. Stylus with ventral trichomes. Ventral process with a triangular flabellum, with digitiform ventral median projection. </p>
            <p>Taxonomical note.</p>
            <p> Gonyleptes atrus was synonymized, without further remarks, with  Gonyleptes saprophilus by Kury (2003) in his catalogue of Laniatores of the New World. However, the study of type material of both species revealed no overlap of some diagnostic characteristics.  Gonyleptes atrus is relatively larger than  Gonyleptes saprophilus . Both species present different patterns of ornamentation on male femur IV.  Gonyleptes atrus shows a sigmoid femur IV with robust dorsal subasal apophysis and two remarkable retrolateral apophyses (Fig. 9  A–B ), while  Gonyleptes saprophilus shows a straight femur, reduced dorsal subasal apophysis, one remarkable retrolateral apophysis and three to five dorsal spines on anterior half of  the femur IV (Fig. 9  C–D ). Specimens of  Geraeocormobius jimi and  Gonyleptes atrus are identical. The description of  Weyhia brieni is sufficiently clear to consider this species synonymy of  Gonyleptes atrus , even without the examination of the type material. Only  Gonyleptes itatiayae Mello-Leitão ,  Weyhia bisignata Mello-Leitão and  Gyndesops pretiosus Mello-Leitão , and all its combination, remain in the synonymic list of  Gonyleptes saprophilus .  Gonyleptes atrus occurs in sympatry with  Gonyleptes saprophilus in almost all of its distribution range. </p>
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	https://treatment.plazi.org/id/E67535A551E03FB4CB82811E01881AE7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pinto-da-Rocha, Ricardo;Benedetti, Alipio Rezende;de Vasconcelos, Eduardo Gomes;Hara, Marcos Ryotaro	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
3C56F9A55B29EF714C40B1B7F0B9B6FB.text	3C56F9A55B29EF714C40B1B7F0B9B6FB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gonyleptes fragilis Mello-Leitao 1923	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> 
Gonyleptes fragilis 
Mello-Leitao
, 1923
 Fig. 11 </p>
            <p> Gonyleptes fragilis Mello-Leitão , 1923: 127; Kury 2003: 127; (female holotype, Brazil,  São Paulo, Santo  André , Alto da Serra, MZSP 56, examined). </p>
            <p> Mendesius guttatus Roewer, 1952: 57; (female holotype, Brazil, SMF RII 10311/72, examined). </p>
            <p> Gonyleptes banana Kury, 2003: 127 (replacement name for  Mendesius guttatus Roewer, 1952, preoccupied by  Gonyleptes guttatus Roewer, 1917). Syn. n. </p>
            <p>Diagnosis.</p>
            <p> Gonyleptes fragilis resembles  Gonyleptes horridus and  Gonyleptes curvicornis because of the overall similarities of the features of the dorsal scutum.  Gonyleptes fragilis can be distinguished from the other two species by the ocularium with a pair of spines (tubercles in  Gonyleptes horridus and  Gonyleptes curvicornis ) and the pale yellowish spots around the tubercles on abdominal scutum. </p>
            <p>Description.</p>
            <p> Penis (Fig. 11  A–C ): ventral plate inflated in the middle region, with a deep U-cleft on anterior margin (its distal tips convergent), 3 distal pairs of long, curved setae, 1 median pair of short setae, 4 basal pairs of spatulate setae. Glans sac slightly projected distally, with similar projection to a dorsal process. Stylus sigmoid, smooth. Ventral process robust, apex flabelliform with several small projections on distal margin. </p>
            <p>Taxonomical note.</p>
            <p> Soares and Soares (1988) proposed the new combination  Gonyleptes guttatus (Roewer, 1952) for  Melloleitaniella guttatus , making it a homonym of  Gonyleptes guttatus Roewer, 1917 [=  Melloleitaniella guttata (Roewer, 1917)]. Kury (2003) renamed the species as  Gonyleptes banana , based on the  Roewer’s citation about the peculiar collection site of the type specimen, a banana shipment for export. The female holotype of  Mendesius guttatus matches exactly the female holotype of  Gonyleptes fragilis . Both males and females of  Gonyleptes fragilis present a brown general coloration with some pale yellowish spots around the tubercles of the mesotergum, a diagnostic characteristic for this species. In addition, the occurrence area of  Gonyleptes fragilis includes the Vale do Ribeira and southern coast of  São Paulo (Kury 2003), an important area known for producing banana for exportation in the 1940's and 1950's. </p>
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	https://treatment.plazi.org/id/3C56F9A55B29EF714C40B1B7F0B9B6FB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pinto-da-Rocha, Ricardo;Benedetti, Alipio Rezende;de Vasconcelos, Eduardo Gomes;Hara, Marcos Ryotaro	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
3862CD1142AA3F98A5E664B4EC09C2BB.text	3862CD1142AA3F98A5E664B4EC09C2BB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gonyleptes perlatus (Mello-Leitao 1935) Mello-Leitao 1935	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> 
Gonyleptes perlatus (
Mello-Leitao
, 1935)
 comb. n.</p>
            <p> Moojenia perlata Mello-Leitão , 1935: 384, fig. 13 (♂); (2 males syntypes, Brazil, Minas Gerais,  Viçosa , MNRJ 1577, examined) </p>
            <p> Collonychium perlatum : Kury 2003: 123. </p>
            <p>Diagnosis.</p>
            <p> Gonyleptes perlatus is very similar to  Gonyleptes horridus (according to Kury 2003) but can be distinguished by the relatively larger tubercles on lateral margin of the do  rsal scutum and the absence of a robust prolateral row of apophysis on male femur IV (present in  Gonyleptes horridus ), although this article might present variable armature. </p>
            <p>Taxonomical note.</p>
            <p> We propose the allocation of this species in  Gonyleptes because it is very similar to  Gonyleptes horridus (see comments in Kury 2003), including the typical sexual dimorphism. According Kury (2003), both species are allopatric. </p>
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	https://treatment.plazi.org/id/3862CD1142AA3F98A5E664B4EC09C2BB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pinto-da-Rocha, Ricardo;Benedetti, Alipio Rezende;de Vasconcelos, Eduardo Gomes;Hara, Marcos Ryotaro	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
EC30BA5D5DBB468147F7AFDC68512D52.text	EC30BA5D5DBB468147F7AFDC68512D52.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gonyleptes pustulatus Sorensen 1884	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> 
Gonyleptes pustulatus 
Sorensen
, 1884
 Fig. 12 </p>
            <p> Gonyleptes pustulatus Sørensen , 1884: 603, Kury 2003: 128; (male holotype, Brazil, ZMUC, examined from detailed photo). </p>
            <p> Gonyleptes guttatus Roewer, 1917: 125, fig. 25 (♂); Kury 2003: 128; (male holotype, Brazil,  São Paulo, Santos, SMF 1321, examined). Syn. n. </p>
            <p>Material examined.</p>
            <p> BRAZIL. Locality not specified further, male holotype of  Gonyleptes pustulatus , examined from photo (ZMUC).  São Paulo: Santos, male holotype of  Gonyleptes guttatus (SMF 1321). Rio de Janeiro: Casimiro de Abreu, 5 ♂ &amp; 6 ♀ (MNRJ 17456). </p>
            <p>Diagnosis.</p>
            <p> Gonyleptes pustulatus (Fig. 12  A–B ) superficially resembles  Gonyleptes bimaculatus because of the conspicuous white patches on mesotergum. Males of  Gonyleptes pustulatus can be distinguished from  Gonyleptes bimaculatus by: femur IV armed (unarmed in  Gonyleptes bimaculatus ) and presence of a bifid retrolateral apical apophysis on coxa IV. </p>
            <p>Description.</p>
            <p> Penis (Fig. 12  C–D ; MNRJ 17456): ventral plate with slightly inflated middle region, a deep distal U-cleft on anterior margin (its distal tips are divergent), 3 distal pairs of long, curved setae, 1 median pair of short setae, 4 basal pairs of spatulate setae. Stylus slightly straight, with apex slightly swollen. Ventral process robust, apex flabelliform with serrated and rounded distal margin. </p>
            <p>Taxonomical note.</p>
            <p> Gonyleptes pustulatus is a remarkable species by body color pattern. It was described without any illustration. Probably this was the reason that kept this species virtually unknown and led Roewer to describe  Gonyleptes guttatus . </p>
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	https://treatment.plazi.org/id/EC30BA5D5DBB468147F7AFDC68512D52	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pinto-da-Rocha, Ricardo;Benedetti, Alipio Rezende;de Vasconcelos, Eduardo Gomes;Hara, Marcos Ryotaro	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
062F75CF17D59338F303B3899A7DC5D4.text	062F75CF17D59338F303B3899A7DC5D4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Liogonyleptoides minensis (Piza 1946) Piza 1946	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Liogonyleptoides minensis (Piza, 1946)</p>
            <p> Chaquesia minensis Piza, 1946: 365, fig. 1 (♂); (2 males &amp; 3 females syntypes; Brazil, Minas Gerais, Cachoeira do  Pajeú ; MZLQ A0053; examined). </p>
            <p> Liogonyleptoides minensis : Kury 2003: 130. </p>
            <p> Currala bahiensis Soares, 1972: 56, figs 1-4, 14-15 (♂♀); Kury 2003: 123; (male holotype and 1 female paratype; Brazil, Bahia,  Maracás ; MNRJ 5268; examined). Syn. n. </p>
            <p>Taxonomical note.</p>
            <p> Comparisons between the type material of  Currala bahiensis and  Liogonyleptoides minensis allowed us to conclude that they are synonymous.  Liogonyleptoides minensis is a species that can be easily diagnosed among gonyleptines by the relatively larger size, body dorsally rounded, without tubercles on corners, presence of scutal area IV; femur III curved (instead of sigmoid); male femur IV with a long, robust dorsobasal apophysis and a prolateral one that is curved dorsally on distal third. The genus  Currala is now composed only by its type species  Currala spinifrons Roewer, 1927. </p>
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	https://treatment.plazi.org/id/062F75CF17D59338F303B3899A7DC5D4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pinto-da-Rocha, Ricardo;Benedetti, Alipio Rezende;de Vasconcelos, Eduardo Gomes;Hara, Marcos Ryotaro	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
C477EC0827EAA848715B1C99ACA5AB5B.text	C477EC0827EAA848715B1C99ACA5AB5B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mischonyx Bertkau 1880	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Mischonyx Bertkau, 1880</p>
            <p> Mischonyx Bertkau, 1880: 106; Kury 2003: 132; (type species:  Mischonyx squalidus Bertkau, 1880, by monotypy). </p>
            <p> Eugonyleptes Roewer, 1913: 219; Kury 2003: 123; (type species:  Gonyleptes scaber Kirby, 1819, by monotypy). Syn. n. </p>
            <p> Gonazula Roewer, 1930: 417; Kury 2003: 126; (type species:  Gonazula gibbosa Roewer, 1930, by monotypy). Syn. n. </p>
            <p>Diagnosis.</p>
            <p> Mischonyx resembles  Currala Roewer, 1927 and  Schenkelibunus Strand, 1932, which are gonyleptine of relatively smaller size (about 5 mm of dorsal scutum length).  Mischonyx can be distinguished from those genera by the combination of the following characters: anterior border of the carapace with robust spines; frontal hump on anterior margin of carapace and ocularium with a pair of spines; scutal areas  I–III with a paramedian pair of tubercles, those on  I–II elliptical and widest pair on scutal area III; lateral margin of the dorsal scutum with large white tubercles (in specimens preserved in 70% ethanol)  .</p>
            <p>Taxonomical note.</p>
            <p> According DaSilva and Pinto-da-Rocha (2010),  Mischonyx is phylogenetically close to  Hernandariinae . It is probable that due to the Roewerian classification, some species which should be in  Mischonyx is placed elsewhere in  Gonyleptinae . Therefore, a comprehensive revision of the subfamily is needed. </p>
            <p>Composition.</p>
            <p> Mischonyx anomalus (  Mello-Leitão , 1936);  Mischonyx antiquus (  Mello-Leitão , 1934);  Mischonyx cuspidatus (Roewer, 1913);  Mischonyx fidelis (  Mello-Leitão , 1931);  Mischonyx insulanus (Soares, 1972);  Mischonyx intermedius (  Mello-Leitão , 1935);  Mischonyx kaisara Vasconcelos, 2004;  Mischonyx poeta Vasconcelos, 2005;  Mischonyx processigerus (Soares &amp; Soares, 1970);  Mischonyx squalidus Bertkau, 1880 and  Mischonyx sulinus (Soares &amp; Soares, 1947). </p>
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	https://treatment.plazi.org/id/C477EC0827EAA848715B1C99ACA5AB5B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pinto-da-Rocha, Ricardo;Benedetti, Alipio Rezende;de Vasconcelos, Eduardo Gomes;Hara, Marcos Ryotaro	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
DE6965D44E10B9B79055DA2350A09D97.text	DE6965D44E10B9B79055DA2350A09D97.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mischonyx cuspidatus (Roewer 1913) Roewer 1913	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Mischonyx cuspidatus (Roewer, 1913)</p>
            <p> Ilhaia cuspidata Roewer, 1913: 221; (male holotype; Brazil, Rio de Janeiro, Ilha Grande, SMF 900; examined). </p>
            <p> Mischonyx cuspidatus : Kury 2003: 133. </p>
            <p> Gonazula gibbosa Roewer, 1930: 418, fig. 32 (♂); Kury 2003: 126; (male holotype; Brazil, Santa Catarina, Serra Azul; SMF 1328; examined). Syn. n. </p>
            <p>Material examined.</p>
            <p> BRAZIL. Rio de Janeiro: Ilha Grande, male holotype (SMF 900);  Macaé , 16 ♂ &amp; 6 ♀ (MNRJ 4601). Santa Catarina: Serra Azul, male holotype (SMF 1328); Gaspar, 1 ♂ (MNRJ 11584). </p>
            <p>Diagnosis.</p>
            <p> Mischonyx cuspidatus resembles  Mischonyx poeta and  Mischonyx scaber (see diagnosis of  Mischonyx scaber ).  Mischonyx cuspidatus can be distinguished from them by the median high, acuminated tubercle on the free tergites  I–III . </p>
            <p>Taxonomical note.</p>
            <p> The type locality of  Gonazula gibbosa , Serra Azul (in the state of Santa Catarina), is an unknown toponym from where seven species are recorded (see Kury 2003). As some of these species were rediscovered in Serra dos  Órgãos (in the state of Rio de Janeiro), we believe that the opilionids from Serra Azul were mislabeled, although  Mischonyx cuspidatus is a synanthropic gonyleptid (Mestre and Pinto-da-Rocha 2004) with one of the largest geographical distributions known among Neotropical harvestmen (see records in Kury 2003). </p>
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	https://treatment.plazi.org/id/DE6965D44E10B9B79055DA2350A09D97	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pinto-da-Rocha, Ricardo;Benedetti, Alipio Rezende;de Vasconcelos, Eduardo Gomes;Hara, Marcos Ryotaro	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
BE4673223C875761292F24387C2A45B2.text	BE4673223C875761292F24387C2A45B2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mischonyx scaber (Kirby 1819) Kirby 1819	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Mischonyx scaber (Kirby, 1819) comb. n.</p>
            <p> Gonyleptes scaber Kirby, 1819: 453; (3 males &amp; 1 female syntypes; Brazil; NHM 1863.41; examined from detailed photo). </p>
            <p> Eugonyleptes scaber : Roewer, 1913: 219; Kury 2003: 123. </p>
            <p> Xundarava holacantha Mello-Leitão , 1927: 20; (female holotype; Brazil, Rio de Janeiro, Niteroi; MNRJ 1469; examined).Syn. n. </p>
            <p> Mischonyx holacanthus : Kury 2003: 133. </p>
            <p>Material examined.</p>
            <p> BRAZIL. Locality not specified further, 3 males &amp; 1 female syntypes (NHM 1863.41). Rio de Janeiro:  Niterói , female holotype of  Xundarava holacantha (MNRJ 1469); ditto, male holotype of  Weyhia absconsa (MNRJ 1483). </p>
            <p>Diagnosis.</p>
            <p> Mischonyx scaber resembles  Mischonyx cuspidatus and  Mischonyx poeta because of armature of the male femur IV: presence of a dorsobasal apophysis relatively short and curved; one retrolateral apophysis on distal third; and a row of high, acuminated tubercles increasing in size from the base to distal third.  Mischonyx scaber can be distinguished from  Mischonyx cuspidatus by the presence of a short gap between the retrolateral row and the retrolateral apophysis on femur IV (the retrolateral row is continuous to the retrolateral apophysis on femur IV in  Mischonyx cuspidatus ).  Mischonyx scaber can be distinguished from  Mischonyx poeta by the absence of enlarged tubercles (present in  Mischonyx poeta ) on the posterior part of the lateral margin of the dorsal scutum. </p>
            <p>Taxonomical note.</p>
            <p> Mischonyx scaber was described from Brazil, however, many authors, including Kury (2003), cited other countries as its type locality. It is clearly written  “Brazil” on the label of the  Gonyleptes scaber holotype. The only other known specimens of this species are the types of  Xundarava holacantha and  Weyhia absconsa (both in poor condition). The precise distribution records are Rio de Janeiro (Ilha do Governador) and  Niterói . </p>
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	https://treatment.plazi.org/id/BE4673223C875761292F24387C2A45B2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pinto-da-Rocha, Ricardo;Benedetti, Alipio Rezende;de Vasconcelos, Eduardo Gomes;Hara, Marcos Ryotaro	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
29E7A90639329AA1642DBC274E2D4090.text	29E7A90639329AA1642DBC274E2D4090.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Megapachylus grandis Roewer 1913	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Megapachylus grandis Roewer, 1913 Fig. 13 </p>
            <p> Megapachylus grandis Roewer, 1913: 124, fig. 56 (♂); Kury 2003: 131. </p>
            <p> Metapachyloides almeidai Soares &amp; Soares, 1946: 317, fig. 2 (♂); Kury 2003: 176; (male holotype; Brazil, Batea,  São Paulo, Lane &amp; Soares leg., V.1943; MZSP 883; examined) Syn. n. </p>
            <p>Description.</p>
            <p> Penis (Fig. 13  A–C ; MZSP16837): ventral plate with slightly convergent lateral sides deep U-cleft on distal margin, its apex folded ventrally, 3 distal pairs of large setae, slightly curved at apex, 1 median pair of short setae and, 4 basal pairs of spatulate setae, directed frontwards. Glans with dorsal process. Stylus thin, long, sigmoid, with subapical ventral trichomes. Ventral process shaft folded ventrally, apex flabelliform, with serrate distal margins and long distal tip. </p>
            <p>Taxonomical note.</p>
            <p> The male of the species described by Soares &amp; Soares shows a moderate development of armature of leg IV but the number and arrangement of tubercles are perfectly coincident with those of  Megapachylus grandis . It could be a beta male, although an extensive population study should be conducted to confirm this. </p>
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	https://treatment.plazi.org/id/29E7A90639329AA1642DBC274E2D4090	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pinto-da-Rocha, Ricardo;Benedetti, Alipio Rezende;de Vasconcelos, Eduardo Gomes;Hara, Marcos Ryotaro	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
BDC701706E61516999256D31A4330AE9.text	BDC701706E61516999256D31A4330AE9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parampheres Roewer 1913	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Parampheres Roewer, 1913</p>
            <p> Parampheres Roewer, 1913: 345; (type species:  Parampheres pectinatus Roewer, 1913, by monotypy). </p>
            <p> Metapachyloides Roewer, 1917: 120; (type species:  Metapachyloides rugosus Roewer, 1917, by monotypy) Syn. n. </p>
            <p>Diagnosis.</p>
            <p> Parampheres resembles  Acanthogonyleptes ,  Adhynastes Roewer, 1930 and  Deltaspidium Roewer, 1927 by the presence of a more elongated prolateral apical apophysis on male coxa IV (greater length/basal diameter ratio of the apophysis) than that in  Gonyleptes and related  Gonyleptinae genera.  Parampheres can be distinguished from  Adhynastes and  Deltaspidium by the scutal area III with tubercles (instead of spines).  Parampheres can be distinghished from Acanthogonyleptes by the absence of a clearly defined dorsobasal apophysis on the male femur IV.  Parampheres species bear a row of apophyses on the dorsal basal area of the femur IV. </p>
            <p>Taxonomical note.</p>
            <p> Metapachyloides is a monotypic genus and its type species is junior synonym of  Parampheres tibialis Roewer, 1917. </p>
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	https://treatment.plazi.org/id/BDC701706E61516999256D31A4330AE9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pinto-da-Rocha, Ricardo;Benedetti, Alipio Rezende;de Vasconcelos, Eduardo Gomes;Hara, Marcos Ryotaro	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
A9E7A3D446B03DD7E4D2BAD982F1B93B.text	A9E7A3D446B03DD7E4D2BAD982F1B93B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parampheres tibialis Roewer 1917	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Parampheres tibialis Roewer, 1917</p>
            <p> Parampheres tibialis Roewer, 1917: 144, fig. 37 (♂); (male holotype; Brazil,  São Paulo, Santos; SMF 1330; examined). </p>
            <p> Metapachyloides rugosus Roewer, 1917: 121, fig. 22 (♀); (female holotype; Brazil,  São Paulo, Santos; SMF 1322; examined). Syn. n. </p>
            <p>Taxonomical note.</p>
            <p> The monotypic genus  Metapachyloides was formerly placed in  Gonyleptidae ,  Pachylinae . Instead of a male, as stated in the original description, the  holotype of  Metapachyloides rugosus is a female, and its general aspect is similar to that of  Parampheres Roewer, 1913. Surprisingly, in the same year, Roewer described  Parampheres tibialis in  Gonyleptinae , based on a male collected in the same locality (Santos), the most important harbor in Brazil. Roewer described many harvestmen species indicating Santos as their type locality, but those were never collected again there or in the area of endemism to which Santos belongs (Serra do Mar of  São Paulo, see Pinto-da-Rocha et al. 2005 for more details). We conclude that  Parampheres tibialis , as well as many other species supposedly collected in Santos were probably mislabeled. </p>
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	https://treatment.plazi.org/id/A9E7A3D446B03DD7E4D2BAD982F1B93B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pinto-da-Rocha, Ricardo;Benedetti, Alipio Rezende;de Vasconcelos, Eduardo Gomes;Hara, Marcos Ryotaro	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
A9326F98A6FF24DC1201463930754490.text	A9326F98A6FF24DC1201463930754490.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parapachyloides uncinatus (Sorensen 1879) Sorensen 1879	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> 
Parapachyloides uncinatus (
Sorensen
, 1879)
 Fig. 14 </p>
            <p> Gonyleptes uncinatus Sørensen , 1879: 214, figs 1-5, 9-10, 14, 18, 22-26, 28-29 (♂♀); (syntypes males &amp; females, ZMUC, type locality not specified, see Kury 2003; examined from detailed photos). </p>
            <p> Parapachyloides uncinatus : Kury 2003: 186. </p>
            <p> Goyazella armata Mello-Leitão , 1931: 120, fig. 1 (♀); (female holotype; MNRJ 11373; Brazil;  Goiás ; Chapada dos Veadeiros; Blases leg., examined). syn. n. </p>
            <p> Parapachyloides armatus : B. Soares 1944b: 164; Kury 2003: 186; Kury et al. 2010: 566. </p>
            <p>Material examined.</p>
            <p> BRAZIL.  São Paulo: Porto Cabral, 4 ♂ (MZSP 787); Usina  Hidrelétrica de Rosana, 2 males &amp; 2 females (MZSP 14571). Mato Grosso do Sul:  Taquaruçu , 2 ♂ &amp; 1 ♀ (MZSP 19113); ditto, (Canal do Rio  Baía ), 1 ♂ (MZSP 22084). </p>
            <p>Description.</p>
            <p> Penis (Fig. 14  A–C ; MZSP 14571): ventral plate with the lateral sides slightly convergent, deep U-cleft on distal margin, its apex folded ventrally and convergent, 3 distal pairs of setae curved on apex, 1 pair of short median setae, 4 basal pairs of setae arranged in a row, 2 pairs of ventral distal short setae. Glans with small dorsal process. Stylus slender, sigmoid, smooth. Ventral process apex flabelliform with serrate distal margin. </p>
            <p> Taxonomical note. </p>
            <p> Males of this species show polymorphism in the posterior tubercle on the lateral margin of dorsal scutum (from low, small and blunt to high, enlarged, curved and pointed apex), posterior margin of dorsal scutum (from low tubercles to two high tubercles) and armature of free tergite I (with one large median tubercle without enlarged lateral tubercles, or with 2-4 high lateral tubercles); retrolateral basal tubercle on male femur IV of same width as femur to 1.5 times wider, short (1.3 times femur width) to long (3.2  × ). </p>
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	https://treatment.plazi.org/id/A9326F98A6FF24DC1201463930754490	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pinto-da-Rocha, Ricardo;Benedetti, Alipio Rezende;de Vasconcelos, Eduardo Gomes;Hara, Marcos Ryotaro	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
CD7076E26DBDCC91273BD7479D7C9F6C.text	CD7076E26DBDCC91273BD7479D7C9F6C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Schubartesia singularis B. Soares 1944	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Schubartesia singularis B. Soares, 1944a Fig. 15 </p>
            <p> Schubartesia singularis B. Soares, 1944a: 34, fig. 1 (♂); Kury 2003: 192; Kury et al. 2010: 566; (male holotype, Brazil, Bahia, Os Gerais, Rio Branco Valley, MZSP, examined). </p>
            <p>Material examined.</p>
            <p> BRAZIL. Bahia: (Os Gerais, Rio Branco Valley), male holotype of  Schubartesia singularis (MZSP). Minas Gerais:  Januária (Parque Nacional Cavernas do  Peruaçu ), 1 ♂ &amp; 1 ♀ (MZSP 29075); ditto, 2 ♂ (MZSP 29839). </p>
            <p>Description.</p>
            <p> Penis (Fig. 15  A–C ; MZSP 29075): ventral plate with the sides subparallel, with deep cleft on distal margin (U-cleft in ventral view and V-cleft in dorsal view; its apex convergent), 3 pairs of long helycoidal setae, 2 pairs of ventral distal short setae, 1 pair of median short setae, 4 pairs of spatulate, large setae arranged in an oblique row in lateral view, basal lobe short and projected dorsad. Stylus sigmoid, smooth. Ventral process shaft short and wide, apex flabelliform with slightly serrated distal margin. </p>
            <p>Taxonomical note.</p>
            <p> Schubartesia singularis and  Parapachyloides armatus were both formerly placed in  Pachylinae and transferred recently to  Gonyleptinae by Kury et al. (2010). In that occasion, they justified the transfer arguing that penial features are typical of those of  Gonyleptinae , but they did not formally described or characterized them. Despite the typical "  Pachylinae " general aspect (four scutal areas on dorsal  scutum , robust pedipalps and conspicuous trochanter armature), the penis morphology, such as the deep cleft on distal margin of the penis ventral plate, indicates that it belongs to a more derivative gonyleptine lineage. </p>
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	https://treatment.plazi.org/id/CD7076E26DBDCC91273BD7479D7C9F6C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pinto-da-Rocha, Ricardo;Benedetti, Alipio Rezende;de Vasconcelos, Eduardo Gomes;Hara, Marcos Ryotaro	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
1CDE6E195D5233C7F606C1C65E48E519.text	1CDE6E195D5233C7F606C1C65E48E519.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudopucrolia mutica (Perty 1833) Perty 1833	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Pseudopucrolia mutica (Perty, 1833)</p>
            <p> Eusarcus muticus Perty, 1833: 203. </p>
            <p> Pseudopucrolia mutica : Kury 2003: 143; Mendes 2011: 462. </p>
            <p> Meteusarcus armatus Roewer, 1913: 74, fig. 33 (♀); Kury 2003: 177; (female holotype; Brazil; SMF 801; examined). Syn. n. </p>
            <p>Diagnosis.</p>
            <p>See Mendes (2011: 463).</p>
            <p>Taxonomical note.</p>
            <p>See genus "Taxonomical note".</p>
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	https://treatment.plazi.org/id/1CDE6E195D5233C7F606C1C65E48E519	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pinto-da-Rocha, Ricardo;Benedetti, Alipio Rezende;de Vasconcelos, Eduardo Gomes;Hara, Marcos Ryotaro	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
D6ABA4CEA1BC6568F5BA682E46A07366.text	D6ABA4CEA1BC6568F5BA682E46A07366.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudopucrolia Roewer 1912	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Pseudopucrolia Roewer,1912</p>
            <p> Pseudopucrolia Roewer, 1912: 167; Mendes 2011: 451; (type species:  Pseudopucrolia spinosa Roewer, 1912, by monotypy). </p>
            <p> Meteusarcus Roewer, 1913: 74; Kury 2003: 177; (type species:  Meteusarcus armatus Roewer, 1913, by monotypy). Syn. n. </p>
            <p>Diagnosis.</p>
            <p>See Mendes (2011: 453).</p>
            <p> Composition.  Pseudopucrolia discrepans (Roewer, 1943);  Pseudopucrolia incerta (  Mello-Leitão , 1928);  Pseudopucrolia mutica (Perty, 1833) and  Pseudopucrolia rugosa (Roewer, 1930). </p>
            <p>Taxonomical note.</p>
            <p> Meteusarcus armatus was formerly placed in  Gonyleptidae ,  Pachylinae . The examination of the female holotype revealed it was a female of the heteropachyline  Pseudopucrolia mutica . </p>
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	https://treatment.plazi.org/id/D6ABA4CEA1BC6568F5BA682E46A07366	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pinto-da-Rocha, Ricardo;Benedetti, Alipio Rezende;de Vasconcelos, Eduardo Gomes;Hara, Marcos Ryotaro	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
43F1E56775DE7A8252700D512CAB0078.text	43F1E56775DE7A8252700D512CAB0078.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acrographinotus Holmgren 1916	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Acrographinotus Holmgren, 1916</p>
            <p> Acrographinotus Holmgren, 1916: 89; Roewer 1929: 240; Kury 2003: 155; (type species:  Acrographinotus erectispina Roewer, 1929, by subsequent designation of Roewer (1929)). </p>
            <p> Unduavius Roewer, 1929: 244; Kury 2003: 195; (type species:  Unduavius ornatus Roewer, 1929, by monotypy). Syn. n. </p>
            <p>Diagnosis.</p>
            <p>See Acosta (2001: 59).</p>
            <p>Composition.</p>
            <p> Acrographinotus ceratopygus (Soares &amp; Bauab-Vianna, 1972);  Acrographinotus curvispina Roewer, 1929;  Acrographinotus erectispina Roewer, 1929;  Acrographinotus mitmaj Acosta, 2002;  Acrographinotus niawpaq Acosta, 2001;  Acrographinotus ortizi (Roewer, 1957);  Acrographinotus ornatus (Roewer, 1929), comb. n. </p>
            <p>Taxonomical note.</p>
            <p> The genus  Unduavius is monotypic, but the remarkable ventral process of the penis, with its "ibis head" shape, fits perfectly in the generic concept presented by Acosta (2001), who rediagnosed the genus  Acrographinotus . </p>
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	https://treatment.plazi.org/id/43F1E56775DE7A8252700D512CAB0078	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pinto-da-Rocha, Ricardo;Benedetti, Alipio Rezende;de Vasconcelos, Eduardo Gomes;Hara, Marcos Ryotaro	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
467B6CBC2F12ABD97E1E6C6CFE6D175C.text	467B6CBC2F12ABD97E1E6C6CFE6D175C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acrographinotus ornatus (Roewer 1929) Roewer 1929	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Acrographinotus ornatus (Roewer, 1929) comb. n. Fig. 16 </p>
            <p> Unduavius ornatus Roewer, 1929: 244, fig. 28 (♂); Kury 2003: 195; (syntypes from NHMW, not examined, probably lost). </p>
            <p>Material examined.</p>
            <p>BOLIVIA. La Paz: Nor Yungas (Unduavi), 1 male &amp; 6 females (SMF 2881/79; syntypes according to Acosta (1996), which was previously referred to as belonging to Naturhistorisches Museum, Wien. We acknowledge them as syntypes together with those in SMF RII 995/52).</p>
            <p>Diagnosis.</p>
            <p> Acrographinotus ornatus can be easily distinguished from other species of the genus by the blister-like, enlarged, brown tubercles on whitish scutal areas  I–IV ; posterior margin of dorsal scutum and free tergites  I–II with a row of enlarged tubercles and white spots on lateral and posterior margin of dorsal scutum and coxa IV. </p>
            <p>Description.</p>
            <p> Penis (Fig. 16  A–B ; SMF 2881): ventral plate with 3 distal pairs of long setae, 0-2 median pairs of short setae, 4-5 basal pairs of setae. Stylus almost straight (apex slightly bent), smooth. Distal ventral process shaped as an ibis head. </p>
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	https://treatment.plazi.org/id/467B6CBC2F12ABD97E1E6C6CFE6D175C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pinto-da-Rocha, Ricardo;Benedetti, Alipio Rezende;de Vasconcelos, Eduardo Gomes;Hara, Marcos Ryotaro	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
39F9F39A3CD2D93E4F4849EC03CF93E9.text	39F9F39A3CD2D93E4F4849EC03CF93E9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gyndesops Roewer 1943	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Gyndesops Roewer, 1943 new subfamilial assignment</p>
            <p> Gyndesops Roewer, 1943: 29; Kury 2003: 129; (type species:  Gyndesops denisi Roewer, 1943). </p>
            <p>Diagnosis.</p>
            <p> Gyndesops resembles the largest  Pachylinae genus,  Discocyrtus Holmberg, 1878, which presents an ocularium and scutal area III with paired armature, four scutal areas, scutal area  II–IV and free tergites  I–III unarmed. This combination of characters also occurs in genera such as  Gyndoides Mello-Leitão , 1927,  Lacronia Strand, 1942,  Paraluederwaldtia Mello-Leitão , 1927 and  Parapucrolia Roewer, 1917.  Gyndesops can be distinguished  from Lacronia and  Parapucrolia by the coloration, which is uniformly auburn (  Lacronia has green/yellow spots and/or stripes on dorsal scutum and  Parapucrolia has white patches on dorsal scutum). It is very difficult to distinguish  Gyndesops from the remaining genera, because no revision was ever made including those groups, and their monophyly is doubtful. It is possible that  Gyndesops (as well as  Gyndoides and  Paraluederwaldtia ) is a junior synonym of  Discocyrtus , but we refrained ourselves from such a rash nomenclatural act. </p>
            <p>Composition.</p>
            <p>Monotypic.</p>
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	https://treatment.plazi.org/id/39F9F39A3CD2D93E4F4849EC03CF93E9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pinto-da-Rocha, Ricardo;Benedetti, Alipio Rezende;de Vasconcelos, Eduardo Gomes;Hara, Marcos Ryotaro	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
5B85277898C1C02F927856B717A3C04D.text	5B85277898C1C02F927856B717A3C04D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gyndesops denisi Roewer 1943	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Gyndesops denisi Roewer, 1943 Fig. 17 </p>
            <p> Gyndesops denisi Roewer, 1943: 29, fig. 23 (♂); Kury 2003: 129; (male holotype, Brazil [Brasilien: Nova Teutonia], SMF 8833/117, examined). </p>
            <p>Description.</p>
            <p> Penis (Fig. 17  A–B , holotype): ventral plate subrectangular, basal half of lateral margin projected laterad, distal margin slightly concave, 3 distal pairs of cyli  ndrical setae, 1 pair of median setae (long left seta and moderate sized right seta), 3 basal pairs of cylindrical setae (median largest). Glans sac almost reaching distal margin of ventral plate. Stylus with swollen apex, smooth. Ventral process with large shaft, apex flabelliform with serrated lateral margins. </p>
            <p>Taxonomical note.</p>
            <p> Originally,  Gyndesops denisi was allocated in  Pachylinae , and later transferred to  Gonyleptinae by Kury (2003), without further explanation. It is known that  Gonyleptinae and  Pachylinae are possibly not monophyletic. A phylogenetic analysis of  Gonyleptidae is currently being carried out and indicates, however, that most  Gonyleptinae have a deep cleft in the distal margin of the penis ventral plate, as well as some other features, absent in this species. Since  Gyndesops denisi (i) does not possess such cleft in the penis ventral plate, and (ii) the exterior morphology might be misleading regarding  Gonyleptinae and  Pachylinae , we opted to allocate it back in  Pachylinae . </p>
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	https://treatment.plazi.org/id/5B85277898C1C02F927856B717A3C04D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pinto-da-Rocha, Ricardo;Benedetti, Alipio Rezende;de Vasconcelos, Eduardo Gomes;Hara, Marcos Ryotaro	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
6F7FB1330433B939B9798750F52C8F74.text	6F7FB1330433B939B9798750F52C8F74.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Haversia Roewer 1913	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Haversia Roewer, 1913 new subfamilial assignment</p>
            <p> Haversia Roewer, 1913: 170; Kury 2003: 129; (type species:  Gonyleptes defensus Butler, 1876). </p>
            <p> Hoggellula Roewer, 1930: 397; Kury 2003: 129; (type species:  Sadocus vallentini Hogg, 1913, by monotypy). Syn. n. </p>
            <p>Diagnosis.</p>
            <p> Haversia resembles  Pachylinae with three scutal areas, such as  Corralia Roewer, 1913,  Diconospelta Canals, 1934,  Graphinotus Koch, 1839,  Huassampilia Roewer, 1913,  Neogonyleptes Roewer, 1913,  Oxapampeus Roewer, 1936,  Spinivunus Roewer, 1943,  Tumbesia Loman, 1899,  Ubatubesia B. Soares, 1945, and two species of  Sadocus Sørensen , 1886 (  Sadocus conspicillatus and  Sadocus polyacanthus ).  Haversia can be distinguished from those genera by the unarmed and smooth (i.e. without tubercles) scutal areas and the complex apophyses of male trochanter IV: three prolateral apophyses (anterior median, posterior median and an apical apophysis). </p>
            <p>Taxonomical note.</p>
            <p> The two type species (  Gonyleptes defensus and  Sadocus vallentini )are synonymous, resulting in a generic synonymy, and the new assignment proposed here is based on penis morphology (see note on  Gonyleptes denisi ). The penis of  Haversia follows the basic pattern of those gonyleptids placed in the large (and non-monophyletic) subfamily  Pachylinae (ventral plate with straight anterior margin; glans without dorsal process; distal, median and basal setae of ventral plate short [seta length inferior to ventral plate width]; stylus and ventral process with variable shape). However,  Haversia presents two features that stand out: (i) the penis glans projected dorsally and (ii) the detailed shape of apex of penis ventral process. Those features are typical of some Chilean genera, such as  Fonckia ,  Neogonyleptes and  Sadocus (unpublished data), and might be the evidence that those four genera are closely related. </p>
            <p>Composition.</p>
            <p>Monotypic.</p>
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	https://treatment.plazi.org/id/6F7FB1330433B939B9798750F52C8F74	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pinto-da-Rocha, Ricardo;Benedetti, Alipio Rezende;de Vasconcelos, Eduardo Gomes;Hara, Marcos Ryotaro	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
24ABA0A2DBD00DC7C296A64F0A91CC58.text	24ABA0A2DBD00DC7C296A64F0A91CC58.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Haversia defensa (Butler 1876) Butler 1876	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Haversia defensa (Butler, 1876) Fig. 18 </p>
            <p> Gonyleptes defensus Butler, 1876: 152, fig. 4 (♂); (4 males &amp; 1 female syntypes; Argentina, Falklands Island; NHM; examined by detailed photos). </p>
            <p> Haversia defensa : Roewer, 1913: 171, fig. 72 (♂); Kury 2003: 129. </p>
            <p> Sadocus vallentini Hogg, 1913: 48, fig. 7 (♂); (syntypes males and females; Falklands Island; NHM 1299.1304; not examined). Syn. n. </p>
            <p> Hoggellula vallentini : Roewer 1930: 397, fig. 22 (♂); Kury 2003: 129. </p>
            <p>Material examined.</p>
            <p>ARGENTINA. Falklands (Islas Malvinas), 1 ♂ (SMF 1320/8).</p>
            <p>Description.</p>
            <p> Penis (Fig. 18  A–B ; SMF 1320/8): ventral plate with lateral margins slightly concave, distal margin straight, 3 distal pairs of cylindrical, straight, long setae, 1 pair of median short setae, 2 pairs of basal cylindrical, straight, long setae. Glans sac projected dorsally, ventral process and stylus arising ventrally. Stylus long, slightly curved, with apical ventral and lateral trichomes. Ventral process swollen apically, with nailhead-like apex. </p>
            <p> Taxonomical note. </p>
            <p> The examination of detailed photos of  Gonyleptes defensus type material and subsequent comparison of those with the description of  Sadocus vallentini allowed recognizing them as conspecific. </p>
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	https://treatment.plazi.org/id/24ABA0A2DBD00DC7C296A64F0A91CC58	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pinto-da-Rocha, Ricardo;Benedetti, Alipio Rezende;de Vasconcelos, Eduardo Gomes;Hara, Marcos Ryotaro	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
19B4FDD0EAB9EC825E32C8307E8180A2.text	19B4FDD0EAB9EC825E32C8307E8180A2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Oxapampeus Roewer 1963	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Oxapampeus Roewer,1963 new subfamilial assignment</p>
            <p> Oxapampeus Roewer, 1963: 62; Kury 2003: 135; (type species:  Oxapampeus weyrauchi Roewer, 1963). </p>
            <p>Diagnosis.</p>
            <p> Based on external morphology,  Oxapampeus resembles Pachylinae with three scutal areas (subfamily classical diagnosis is the presence of four areas) and scutal area III with a paramedian pair of spines, such as  Corralia Roewer, 1913,  Diconospelta Canals, 1934,  Haversia Roewer, 1913,  Huassampilia Roewer, 1913,  Neogonyleptes Roewer, 1913,  Spinivunus Roewer, 1943, and two species of  Sadocus Sørensen , 1886 (  Sadocus conspicillatus and  Sadocus polyacanthus ). However,  Oxapampeus can be distinguished from those genera by the combination of the following characters: ocularium with a paramedian pair of pointed tubercles; frontal hump on anterior border of dorsal scutum moderately developed; scutal areas  I–II each with a paramedian pair of enlarged tubercles; posterior margin of dorsal scutum straight; and free tergites  I–III unarmed. </p>
            <p>Taxonomical note.</p>
            <p> Oxapampeus has been included in  Gonyleptinae due to the presence of three scutal areas on the dorsal scutum and absence of autapomorphies of other gonyleptidean subfamilies. However, its penis does not present the typical gonyleptine pattern. In addition, its ventral plate, with slightly concave lateral sides, as well as the absence of dorsally projected basal lobes, makes it resemble several Andean  Pachylinae male genitalia. </p>
            <p>Composition.</p>
            <p>Monotypic.</p>
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	https://treatment.plazi.org/id/19B4FDD0EAB9EC825E32C8307E8180A2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pinto-da-Rocha, Ricardo;Benedetti, Alipio Rezende;de Vasconcelos, Eduardo Gomes;Hara, Marcos Ryotaro	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
BA19CCEE6E345D0F1FC9E3453149BDBE.text	BA19CCEE6E345D0F1FC9E3453149BDBE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Oxapampeus weyrauchi Roewer 1963	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Oxapampeus weyrauchi Roewer, 1963 Fig. 19 </p>
            <p> Oxapampeus weyrauchi Roewer, 1963: 62, fig. 29 (♂); Kury 2003: 135; (male holotype, 2 males &amp; 5 females paratypes, Peru, Pasco, Western slope of Eastern Andes, Villarica near Oxapampa, 1500 m, in Ucayali drainage, SMF RII 13964/74, examined). </p>
            <p>Description.</p>
            <p> Penis (Fig. 19  A–B ; holotype): ventral plate with lateral and distal margins slightly concave, with 3-4 distal pairs of cylindrical, straight, long setae, 0-1 median seta (visible in lateral view), 5 basal pairs of setae. Glans sac projected dorsally, ventral process and stylus arising ventrally. Stylus short and straight, with ventral subapical trichomes. Ventral process swollen apically, more elongated and thinner than stylus. </p>
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	https://treatment.plazi.org/id/BA19CCEE6E345D0F1FC9E3453149BDBE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Pinto-da-Rocha, Ricardo;Benedetti, Alipio Rezende;de Vasconcelos, Eduardo Gomes;Hara, Marcos Ryotaro	Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes, Hara, Marcos Ryotaro (2012): New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores). ZooKeys 198: 25-68, DOI: http://dx.doi.org/10.3897/zookeys.198.2337, URL: http://dx.doi.org/10.3897/zookeys.198.2337
