taxonID	type	description	language	source
AD4387F33104FF9E10D02488FC76FD94.taxon	materials_examined	Type species. Idanthyrsus macropaleus Schmarda, 1861 (after Kirtley 1994). Type locality. Valparaiso Harbour, Chile.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F33104FF9E10D02488FC76FD94.taxon	diagnosis	Diagnosis. Outer paleae straight (no angle between shaft and blade), flat and with pointed denticles along lateral margins and distal tips. Three thoracic segments and, if confirmed in all species of the genus, only fine lanceolate chaetae on neuropodia of parathoracic segments.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F33104FF9E10D02488FC76FD94.taxon	discussion	Remarks. Before the present study there were 19 accepted species in the genus Idanthyrsus (Kirtley 1994; Nishi and Kirtley 1999). We have synonymised I. glaessneri with I. australiensis and described an additional two new species increasing the total number of valid species to 20 (see Table 1).	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F33104FF9710D02693FB4AF9DF.taxon	description	Figures 1 A, B, 2 – 5, 6 A, Table 1	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F33104FF9710D02693FB4AF9DF.taxon	materials_examined	Material examined. TYPE?: Sabellaria (Hermella) australiensis Haswell, 1883, BMNH 1882.2.22.71 *, 47 mm in length excluding cauda, 6 mm in width (anteriorly), 47 chaetigers (in 2 parts, anterior fragment with 30 chaetigers and posterior with 17 chaetigers). Queensland: Thursday Island, 10 ° 35 ' S 142 ° 13 ' E, 7.8 – 12.3 m, collected by HMS Alert. Additional material examined. Queensland: Cape Yorke Peninsula, south of Fly Point, Putta Putta Beach, 10 ° 45 ' S 142 ° 36 ' E, 11. vii. 1976, 1, AM W 26926, on semi-exposed boulder bank; Thursday Island, 10 ° 35 ' S 142 ° 13 ' E, 4, AM W 7087 *, 1, AM W 26683 (mounted for SEM), 1, AM W 26684 (mounted for SEM), AM W 26851; Gladstone, Calliope River, 23 ° 55 ' S 151 ° 10 ' E, 1, AM W 199314 *; Gladstone Harbour, 23 ° 51 ' S 151 ° 16 ' E, 06. xi. 1975, 1, QM G 10564; Gladstone, Calliope River, 23 ° 55 ' S 151 ° 10 ' E, 5; Gladstone Harbour, 23 ° 51 ' S 151 ° 16 ' E, 06. xi. 1975, 3, QM G 10565 *; Mary River, 26 ° 23 ' S 152 ° 45 ' E, 31. xii. 1971, low tide, 1, AM W 26928; Hervey Bay, Point Vernon, 25 ° 15 ' S 152 ° 49 ' E, iv. 1972, 1, AM W 201700 *; Maroochydore, Alexandra Headland, 26 ° 39 ' S 153 ° 06 ' E, 01. iv. 1972, 1, AM W 26929; Caloundra, 26 ° 48 ' S 153 ° 08 ' E, viii. 1945, 4, QM G 4043, from south beach rocks; 2, QM G 4044; 1938, 1, QM G 212812; Moreton Bay, Peel Island, Horseshoe Bay, 27 ° 30 ' S 153 ° 21 ' E, 18. xii. 1977, 1, QM G 212813 *, from under rocks submerged at low tide. New South Wales: Lennox Heads, 28 ° 48 ' S 153 ° 36 ' E, 27. iii. 1972, 1, AM W 26971; south of Clarence River mouth, Minnie Water, 29 ° 47 ' S 153 ° 18 ' E, 2 – 3. xi. 1963, 3, AM W 26895; 25. ii. 1971, 1, AM W 26901, from shore reef, in crevices, covered in coralline algae, 1, AM W 26900, from shore reef, underneath boulders in rockpools; Wilson Headland, 29 ° 50 ' S 153 ° 17 ' E, 12. v. 1984, 6, AM W 26857; off Coffs Harbour, Arrawarra Head, 30 ° 04 ' S 153 ° 12 ' E, 14. x. 1974, 11, AM W 9196 *, cemented together forming small colony; Coffs Harbour, SW face of South Solitary Island, 30 ° 12 ' S 153 ° 16 ' E, 19. v. 1972, 7.6 – 10.7 m, 5, AM W 26917; 17. v. 1973, 15.2 m, 3, AM W 26916, from small boulders covered in ascidians; Coffs Harbour, west side of Solitary Island, 30 ° 12 ' S 153 ° 16 ' E, 18. v. 1972, 1, AM W 26923; Toowoon Bay, near Tuggerah Lake, 33 ° 22 ' S 151 ° 30 ' E, 13. iv. 1986, 0.6 m, 3, AM W 26858, from rocks loosely piled, but cemented together, covered in Homosira banksii; off Avalon, 33 ° 38 ' S, 151 ° 20 ' E, 31. i. 1973, 36.6 m, 1, AM W 26849, from reef flat; Long Reef, 33 ° 45 ' S 151 ° 19 ' E, 16. xi. 1970, 3, AM W 4524 * (mounted for SEM), from on underside of rocks, in sandy, firm tubes; Long Reef, north side of reef, 33 ° 45 ' S 151 ° 19 ' E, 23. ii. 1971, 1, AM W 26902; v. 1967, 2, MV F 78871 *; Port Jackson, 33 ° 50 ' S 151 ° 16 ' E, 10. i. 1983, intertidal, 1, AM W 198478 *; North Head, 33 ° 50 ' S 151 ° 18 ' E, 26. v. 1972, 28.9 m, 1, AM W 6963 *; 13. xii. 1972, 28.9 m, 1, AM W 26925; Maroubra, 33 ° 57 ' 06 " S 151 ° 17 ' 37 " E, 01. xii. 1954, 1, AM W 1575 *; Kurnell, Potter Point, 34 ° 03 ' S 151 ° 13 ' E, 27. ix. 1989, low tide, 4, AM W 26863, from on and under rocks; Port Hacking, near Jibbon Head, 34 ° 05 ' S 151 ° 10 ' E, 23 m, 1, AM W 16351 *, from reef; north of Wollongong, Bellambi Reef, 34 ° 26 ' S 150 ° 53 ' E, 14. xi. 1971, 1, AM W 26968, from rock pool, in holdfast of seaweed; Shellharbour, 34 ° 35 ' S 150 ° 52 ' E, between tide marks, 5, AM W 26903; Shellharbour, near bluestone quarry, 34 ° 35 ' S 150 ° 52 ' E, 05. ix. 1971, 7.6 – 9.1 m, 3, AM W 26974; Gerroa, Blackhead, 34 ° 46 ' S 150 ° 49 ' E, 25. ix. 1982, mid tide, 1, AM W 198479 *; north of Broulee, 35 ° 51 ' S 150 ° 11 ' E, i. 1971, 1, MV F 78866 *; near Jervis Bay, Wreck Bay, Cemetry Point, 35 ° 11 ' S 150 ° 38 ' E, 27. ii. 1976, intertidal, 1, AM W 26927, on coralline algae; Twofold Bay, Murrumbulga Point, 37 ° 05 ' S 149 ° 54 ' E, 09. x. 1984, intertidal, 1, AM W 199879 *, from gravel and algal washings on rock platform; 27. iii. 1985, intertidal, 1, AM W 199889 *, cryptic fauna and rock platform; 25. vi. 1985, subtidal, 2, AM W 199880 *, crevice fauna and amongst Diopatra tubes on rock platform; Twofold Bay, Red Point, 37 ° 05 ' S 149 ° 54 ' E, 20. v. 1995, 1, AM W 26907; Munganno Point, 37 ° 05 ' S 149 ° 54 ' E, 10. x. 1984, 6 m, 10, AM W 199892 * from erect bryozoan colony on wharf pile. Victoria: South tip of Mallacoota, Bastion Point, 37 ° 34 ' S 149 ° 46 ' E, 1, AM W 26525 (mounted for SEM); Western Port, Merricks, 38 ° 24 ' S 145 ° 07 ' E, 1969, 1, MV F 78856 *; Bass Strait, 5 km south of Point Reginald, 38 ° 48 ' S 143 ° 17 ' E, xi. 1981, 47 m, 1, MV F 78883 *; Western Port, Somers, 38 ° 24 ' S 145 ° 10 ' E, 2, MV F 78853 *; Flinders Ocean Beach, 38 ° 29 ' S 145 ° 02 ' E, 17. xii. 1969, 1, MV F 78872 *; Venus Bay, west of Eagles Nest, 38 ° 40 ' S 145 ° 41 ' E, 01. ii. 1966, 1, MV F 78868 *; Western Port, near Shoreham, Honeysuckle Point, 38 ° 26 ' S 145 ° 04 ' E, 14. xi. 1970, 1, MV F 78864 *; Flinders, south reef, 38 ° 29 ' S 145 ° 02 ' E, 13. xi. 1965, 1, MV F 78863 *; Airey's Inlet, 38 ° 28 ' S 144 ° 06 ' E, 31. i. 1967, 1, MV F 78869. South Australia: Sellicks Beach, 35 ° 20 ' S 138 ° 27 ' E, 16. iii. 1979, 3, AM W 198439 *, York Peninsula, Gleeson's Landing, 34 ° 35 ' S 137 ° 43 ' E, 27. xi. 1985, intertidal, 1, SAM E 3093, from under rocks; 3, SAM E 3094, from under rocks; Aldinga Bay, 35 ° 21 ' S 138 ° 25 ' E, 12. x. 1977, low tide, 1, AM W 14055 *, under rocks. Western Australia: King George Sound, Mistaken Island, 35 ° 04 ' S 117 ° 56 ' E, 1. i. 1988, 1, AM W 26894, crevice fauna; Cape Naturaliste, Bunker Bay, 33 ° 32 ' S 115 ° 02 ' E, 30. i. 1972, low tide, 6, WAM V 156, from amongst metamorphic rocks; Bunker Bay, 33 ° 31 ' 5.48 " S 115 ° 03 25.56 ' E, 30. i. 72, low tide, many, WAM V 284; 2 km off North Point *, 32 ° 01 ' S 115 ° 30 ' E, 11. i. 1991, 30 m, 1, AM W 26908, from reef and coral bommie surrounded by algal beds; Rottnest Island, Strickland Bay, 32 ° 00 ' S 115 ° 30 ' E, 20. i. 1991, 1, AM W 26904, from reef edge, rock and algae; Sandy Cape, Point Peron, 32 ° 01 ' S 115 ° 30 ' E, 1946, 2, AM W 26856, 1, AM W 26855; 2 km south of Cape Peron, reef west of Groyne, 32 ° 01 ' S 115 ° 30 ' E, 26. xii. 1983, 3, AM W 26862, from sponges and gorgonaceans from cove on reef; Point Clone, 32 ° 01 ' S 115 ° 30 ' E, 29. xi. 1945, 1, AM W 26854; Warnborough Sound, 32 ° 20 ' S 115 ° 43 ' E, 21. iii. 1993, 5 m, 1, AM W 26909, from Posidonia sp. beds; Fremantle, South Mole, 32 ° 03 ' S 115 ° 44 ' E, 1, AM W 26861; Cockburn Sound, 32 ° 01 ' S 115 ° 45 ' E, 15. ii. 1966, 1, WAM V 3814; Cottesloe Reef, near Groyne, 31 ° 59 ' S 115 ° 45 ' E, 29. iii. 1973, 2, WAM V 311, from under stones and in dead shells; Cottesloe Beach, 31 ° 59 ' S 115 ° 45 ' E, 08. i. 1988, intertidal, 2, AM W 26864; 29. iii. 1973, 2, WAM 72 – 74, from under stones and in dead shells; Kendrew Island, Dampier Archipelago, 20 ° 29 ' S 116 ° 32 ' E, 03. iv. 1987, 10 m, 1, AM W 26911, crevice fauna; Finucane Island, 20 ° 18 ' S 118 ° 33 ' E, 20. vi. 1970, AM W 26914; Roly Rocks, 20 ° 33 ' S 116 ° 32 ' E, 03. iv. 1987, 10 m, 1, AM W 26910; Port Hedland, Finucane Island, 20 ° 18 ' S 118 ° 33 ' E, 21. vi. 1970, 1, AM W 26912; Broome, Gantheame Point, 17 ° 59 ' S 122 ° 11 ' E, 04. vii. 1970, ELWM spring, 2, AM W 26859; 15. iii. 1987, 12 – 18 m, 2, NTM W 4916 *, from rocky reef; 15. iii. 1987, LWS, 1, NTM W 4548 *, from rock slabs in silt; Broome, Cable Beach, 17 ° 57 ' S 122 ° 12 ' E, 29. ix. 1984, LWS, 1, NTM W 2280 *, from under stones; 02. xii. 1981, 3 m, 2, MV F 78852; Riddell Point, Broome, 17 ° 56.30 ' S 122 ° 12.33 ' E, 19. viii. 1982, 1 WAM 1890. Northern Territory: Port Essington, Sandy Island, 11 ° 06 ' S 132 ° 18 ' E, 02. v. 1982, 14 m, 1, NTM W 1122 *, from muddy bottom; 1, NTM W 1124 *, from muddy bottom; Port Essington, Burford, 11 ° 29 ' S 131 ° 57 ' E, 13. x. 1981, 1, NTM W 1120 *, from reef flat; Cobourg Peninsula, Burford Island, 11 ° 29 ' S 131 ° 57 ' E, 13. x. 1981, 1, NTM W 25 *; Charles Point, Beagle Gulf, 12 ˚ 18.96 ' S 130 ˚ 40.74 ' E, 23 m, 13. x. 1993, NTM W 23480. Other species examined for comparison. HOLOTYPE of Idanthyrsus glaessneri Kirtley, 1994, LACM AHF N 6277, sand flat at low tide, Lakes Entrance, Victoria, 37.8 ° S 147.9 ° E, coll. B. Dew, 1.3 cm in length, 3.42 mm in width (maximum width of thorax without chaetae).	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F33104FF9710D02693FB4AF9DF.taxon	description	Description. Type flesh coloured, with dark brown blotches on sides of operculum and anterior end and darkly pigmented cauda. Body compact and robust (Fig. 1 A). Operculum with completely separate lobes, longer than wide (Figs 1 A, 3 B), with distal end sloped posteriorly (oblique to longitudinal axis). Fourteen pairs of paleae in outer row, with cylindrical shaft (Fig. 3 A, C), and flat, straight blade, ornamented with alternate, sharp-tipped, similar sized and slightly curved denticles on margins. Inner row with nine pairs of cylindrical and smooth paleae, tapering evenly to acute tip (Figs 1 A, B, 3 A, D). Opercular papillae, 14 pairs, peripheral to outer row of paleae on each lobe (Fig. 3 A). Three pairs of nuchal spines with bent tips (hooks) on each side, concave margins smooth, limbation absent (Figs 1 B, 3 A). Tentacular filaments, arranged in nine rows (Fig. 3 B). Median ridge and median organ present, extending to dorsal edge of junction of opercular lobes, raised and pigmented (illustrated in non type material, Fig. 2 B, C). Numerous eyespots present on both sides of median ridge. Palps about half length of operculum. Segment 1 (chaetiger 1), with neuropodial lobe on either side of U-shaped buccal organ (illustrated in non type material Fig. 2 B); capillary neurochaetae present, flattened and with finely denticulated margins (Fig. 3 E). Segment 2 (chaetiger 2) with three pairs of triangular lateral lobes, connecting dorsal branchiae to neuropodia; with similar neurochaetae as those on chaetiger 1. Neuropodia of chaetiger 1 and 2 not vertically aligned, those of chaetiger 2 situated more laterally (Fig. 1 B). Fourty six pairs of dorsal branchiae present from chaetiger 2, continuing along entire length of body but diminishing in size in posterior segments, some represented only by scars (Fig. 1 A). Branchiae conical, with transverse rows of cilia and blunt tips (Fig. 1 A, B). Segments 3 – 5 (parathoracic) with two types of chaetae arranged transversely, eight, straight, flattened, lanceolate, some with frayed twisted tips, majority represented only by stumps and eight fine, smooth capillaries, inserted between lanceolate chaetae (Fig. 3 F). Segments 3 – 5 with lancelolate neurochaetae arranged in two tiers of different lengths (longer about twice the length of shorter, as illustrated in non type material Fig. 4 I). Notopodial parathoracic chaetae larger and more robust than neuropodial. Abdominal region with 42 chaetigers. Notopodia as transverse tori with uncini, decreasing in number posteriorly. Each uncinus with two vertical rows of teeth, each with about eight teeth (Fig. 3 I). Neuropodia with capillaries becoming longer posteriorly, with thin and flattened blades ornamented with thecal laminar extensions distally forming oblique rows and splayed tips (Fig. 3 G, H). Cauda smooth, about a quarter of abdominal length (Fig. 1 A). Variation. As the type material and material from the type locality is in relatively poor condition we provide here some supplementary information based on the material from Arrawarra, northern NSW (Fig. 2 A – H). The median ridge and median organ (Fig. 2 C) have a line of eye spots on both sides. The lobes of chaetiger 1 have fine capillary neurochaetae laterally displaced in relationship to those of chaetiger 2 (Fig. 2 E). Chaetiger 2 with three pairs of triangular lateral lobes which connect the branchiae to the neuropodia (Fig. 2 D). The triangular shaped branchiae (Fig. 2 F) continue all along the abdomen. Parathoracic neuropodia were examined under SEM on material from Long Reef, NSW showing two tiers of lanceolate neurochaetae present (Fig. 4 I). A large number of specimens were examined from several localities, including sexually mature individuals, that vary in length (4 – 60 mm), width (1 – 6 mm), number of chaetigers (25 – 52), number of outer opercular paleae (8 – 28 pairs), inner opercular paleae (6 – 17 pairs), opercular papillae (5 – 16 pairs), nuchal hooks (2 – 3 pairs), number of rows of tentacular filaments (8 – 11), number of branchiae (14 – 46 pairs), number of teeth on abdominal uncini (8 – 10) and number of abdominal chaetigers (15 – 49). These variations can be mainly attributed to size of the specimens, with larger animals tending to have more paleae, papillae, uncinial teeth and abdominal chaetigers. As minor differences in structure of paleae and nuchal hooks have been used previously to differentiate species, we illustrate these structures from material collected from Long Reef, Sydney, NSW (Fig. 4 A – M) and from Mallacoota, Victoria (Fig. 5 A – G) the type locality of Idanthyrsus glaessneri Kirtley, 1994, in addition to material from type locality (Fig. 3 A – I). The outer paleae of specimens examined from each of these three localities vary slightly in terms of length of the denticles on margins, and we suggest that this represents a gradual gradation rather than separate species and represent a difficult character to measure (Figs 3 C, 4 C, 5 C). Lanceolate parathoracic notochaetae appear to be contractile within parapodia (compare Fig. 4 H and Fig. 5 E), and we suggest that these parapodia are highly mobile. Neurochaetae of the first two segments are missing from one side on holotype, but on non type material from type locality (eg. AM W 26683, AM W 26684) they are present on both sides. Abdominal neurochaetae are similar in these three distant localities (Figs 3 G, H, 4 J, K, 5 F). Divergence of the opercular lobes varies and we suggest this may be a function of size and fixation.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F33104FF9710D02693FB4AF9DF.taxon	discussion	Remarks. The diagnostic characters of I. australiensis are the combination of the absence of a limbation on the nuchal spines, the presence of three lateral lobes on segment 2, and the presence of branchiae on posterior segments. Kirtley (1994) primarily used the shape and thecal development of the outer and inner paleae to distinguish between species, but in most cases intraspecific variation was not considered. We have tried to group this variation observed in some groups regarding the arrangement, shape, relative length and relative numbers of denticles on both sides of outer paleae (Table 1). The structure of the notopodial chaetae of the parathoracic segments also differs between I. australiensis (Figs 3 F, 4 H) and I. nesos n. sp. (see this paper), in the former the capillaries imbetween the lanceolate chaetae are thin whereas in the new species the capillaries have a wider distal end, resembling lanceolate chaetae (Fig. 8 F, G). However, for virtually all described species these parathoracic notochaetae are poorly if at all described and comparison cannot be made at this stage, and this is also true for the parathoracic neurochaetae and therefore these characters are not given in Table 1, but we suggest they may provide useful characters to distinguish between species. Idanthyrsus glaessneri Kirtley, 1994 from Lakes Entrance, Victoria, Australia, was briefly described and only the outer and inner opercular paleae illustrated. In his key to species of Idanthyrsus, Kirtley (1994) separated this species from I. australiensis due to the length of denticles (without sharp tipped denticles in I. glaesneri and sharp in I. australiensis), ornamentation of the thecae (irregular wavy lines across blade in I. glaesneri) and overall shape of paleae (strongly bent in I. glaesneri and slightly sigmoidal in I. australiensis). After examination of the holotype of I. glaesneri, which has been partially dried at some stage, is posteriorly incomplete, and has lost the majority of the outer row of paleae, we would describe the outer opercular paleae as with alternate denticles, slightly curved, with a ratio of 1: 1. This together with other diagnostic features (presence of three lateral lobes on segment 2 and the presence of branchiae on posterior segments) appear to fall within the range observed for I. australiensis and we have therefore synonymised the two species. Examining a large amount of material of I. australiensis from many localities (Fig. 6 A) has allowed us to assess the variability of characters such as the ornamentation of the paleae which Kirtley (1994) had largely relied upon to distinguish species and we have concluded that additional characters such as numbers and distribution of denticles on the outer paleae, number and shape of nuchal spines, numbers of lateral lobes on segment 2 and absence or presence of branchiae in posterior abdominal segments are useful to distinguish between species (see Table 1). In summary, we suggest that I. australiensis exhibits some variation in a suite of characters but no discrete geographical patterns can be detected along the distribution range of this species (Fig. 6 A).	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F33104FF9710D02693FB4AF9DF.taxon	distribution	Distribution. Australia wide from NE Queensland to NW Western Australia, except Tasmania (Fig. 6 A). Habitat. Found from low intertidal to depths of 50 m attached to solid substrate, sometimes occurring as crevice fauna. Specimens typically form small colonies of 6 – 10 individuals, although specimens from Bunker Bay, in Western Australia, were found in larger colonies, consisting of several hundreds of specimens.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F3310DFF8A10D022D8FE25FC09.taxon	description	Figures 1 C – D, 6 A, 7, 8, Table 1	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F3310DFF8A10D022D8FE25FC09.taxon	materials_examined	Material examined. HOLOTYPE: 1, WAM V 7850 *, Dolly Beach, Christmas Island, Indian Ocean, 10 ° 31 ' S 105 ° 41 ' E, 27 – 28 xi. 1969, intertidal reef and pools, 18.5 mm in length, 1 mm in width, 33 chaetigers. PARATYPES: 12, WAM V 7851 *, same locality as holotype, 8 – 20 mm in length without cauda, 1 mm in width, 20 – 29 chaetigers; same locality as holotype, AM W 37778 (mounted for SEM on two pins).	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F3310DFF8A10D022D8FE25FC09.taxon	description	Description. Holotype thin (Fig. 1 C), body with slightly pigmented cauda, abdominal segments dorsally ridged, pigmented and glandular, ventrum with scattered pigmentation. Operculum with elongate, partially fused lobes; with distal end sloped posteriorly (oblique to longitudinal axis). Outer paleae, 24 pairs (Figs 1 D, 7 A, 8 A, B), with cylindrical shaft and ornamented thecae as compact rings with irregular edges, restricted to proximal third of length of paleae (Fig. 8 C) and flattened straight blades with alternate, mostly straight sharp-tipped denticles, increasing in length along shaft towards tip (Fig. 8 C). Inner row with 15 pairs of cylindrical paleae tapering evenly to acute tip (Fig. 8 D); ornamented distally with compact narrow ring thecae (Fig. 8 D). Outer paleae longer than inner paleae (Fig. 8 B). Opercular papillae, five pairs, reduced, stout, inserted external to outer row of paleae on each lobe (Fig. 8 B). One pair of stout and flattened nuchal spines present, strongly recurved tips (Figs 1 D, 7 C, 8 B), and concave margins with wide limbation. Tentacular filaments compound, arranged in 13 horizontal rows (Fig. 7 A, B). Median ridge and median organ present at dorsal junction of opercular lobes, with eyespots present on their margins. Palps about one fifth of length of opercular lobes. Segment 1 (chaetiger 1) with rounded lobe-shaped neuropodia (Fig. 7 A) on either side of U-shaped buccal organ, with fine capillary neurochaetae, flattened, with finely denticulated margins, widest at base and tapering towards fine tip (Fig. 8 E). Segment 2 (chaetiger 2) with two pairs of poorly developed triangular lobes with blunt tips, dorsal one larger than ventral, connecting branchiae to neuropodia (Fig. 1 D), and with similar chaetae as those of chaetiger 1. Neuropodia of chaetiger 1 and 2 not vertically aligned, those of chaetiger 2 situated more laterally (Fig. 1 D). Twenty one pairs of dorsal branchiae present from chaetiger 2, absent from posterior abdominal segments. Branchiae with narrow base and weakly ridged, tapering gently to rounded tip, decreasing in size posteriorly. Segments 3 – 5 (parathoracic) with two types of notochaetae arranged transversely, five, large, flattened, lanceolate with frayed tips, with upturned, curved margins (Fig. 8 F, G); and thinner chaetae with straight tips, theca with expanded margins, interspersed with lanceolate (Fig. 8 F, G). Notochaetae appear to be highly retractile (Fig. 8 F). Segments 3 – 5 (parathoracic) with two tiers of lanceolate neurochaetae; seven, thick and seven thinner ones with finely denticulated margins. Neurochaetae less robust than notochaetae. Abdominal region with 27 chaetigers. Notopodia as transverse tori, with number of uncini per torus decreasing posteriorly. Notopodial tori becoming increasingly erect and decreasing in width posteriorly. Each uncinus with two vertical rows of teeth, each row with seven to eight teeth (Fig. 8 I). Neuropodia with capillaries becoming longer posteriorly, with thin and flattened blades ornamented with thecal laminar extensions distally forming oblique rows with elongate frayed tips (Fig. 8 H), similar on all abdominal chaetigers. Cauda, smooth, equivalent in length to 15 chaetigers (Fig. 7 D). Variations. The paratypes, including sexually mature female specimens, vary from 8 – 20 mm in length without cauda, no variation in width, with 20 – 29 segments, 12 – 24 pairs of outer paleae, 10 – 15 pairs of inner paleae; 4 – 9 pairs of opercular papillae; 9 – 19 pairs of dorsal branchiae; and 15 – 24 abdominal chaetigers. These variations can be attributed to size of specimens, with larger animals tending to have more paleae, papillae, uncinial teeth and abdominal chaetigers than smaller ones.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F3310DFF8A10D022D8FE25FC09.taxon	discussion	Remarks. Idanthyrsus nesos n. sp. is the only species described in the genus with the following combination of characters (see Table 1): presence of one pair of limbated nuchal hooks (shared with I. bihamatus (Caullery, 1944), I. cretus Chamberlin, 1919, and presumably I. bicornis (Schmarda, 1861), I. kornickeri Kirtley, 1994, I. luciae (Rochebrune, 1882), I. manningi Kirtley, 1994, and I. valentenei Kirtley, 1994), presence of two lateral lobes on segment 2 (shared with I. bihamatus) and presence of fine denticles (20 – 25) along the edges of the outer paleae, increasing in length towards the tip. Idanthyrsus nesos n. sp., was found at Christmas Island, an isolated location on the Indian Ocean and can be distinguished from I. australiensis, the most common species on the mainland, by the number and shape of nuchal spines (one pair of flattened hooks with large limbation in I. nesos n. sp., and 2 – 3 pairs of cylindrical hooks lacking limbations in I. australiensis) and the number of pairs of lateral lobes on segment 2 (two in I. nesos n. sp., and three in I. australiensis) and by the distribution of branchiae (restricted to anterior abdominal segments in I. nesos n. sp., and present on all abdominal segments in I. australiensis). As summarised in Table 1, other species reported from the Indo-Pacific differ from I. nesos n. sp., in one or more of the diagnostic features, although it should be noted that information is missing for many species. For example, I. albigens (Ehlers, 1908), described from deepwater off Diego Garcia, has two pairs of nuchal hooks whereas I. nesos n. sp., has one pair of these spines. Idanthyrsus bicornis (Schmarda, 1861), described from intertidal areas of Sri Lanka, has outer paleae with more than 30 denticles with an asymmetrical arrangement on both margins and with maximum length at mid paleae, (see Fig. 6.4. 1 in Kirtley 1994), in contrast to the 20 – 25 alternate denticles increasing in length towards the tip present in I. nesos n. sp. Idanthyrsus bihamatus (Caullery, 1944), known only from deep water in Indonesia, has outer paleae with denticles similar in size or slightly longer at mid length of paleae while in I. nesos n. sp., denticles increasing in length towards the tip (see Table 1 for comparison of other features). Idanthyrsus bicornis (Schmarda, 1861), described from intertidal areas of Sri Lanka, has outer paleae with more than 30 denticles which increase in size towards the tip and arising at 45 ˚ (see Fig. 6.4. 1 in Kirtley 1994, not illustrated by Schmarda and also no other diagnostic characters are given) which differs from the new species and I. bihamatus (Caullery, 1944) known only from deep water in Indonesia which has outer paleae with smooth shafts and numerous denticles of uniform size along shaft and arising at 45 ˚ whereas I. nesos n. sp., has straight denticles increasing in length along the strongly denticulate shaft, although they share many other characters (see Table 1). Idanthyrsus nesos n. sp., differs from I. okinawaensis Nishi & Kirtley, 1999, described from shallow waters in Japan, in the number of pairs of opercular papillae, 30 in I. okinawaensis and less than ten in the new species, and the number and shape of nuchal hooks, two pairs and without a limbation in I. okinawaensis and one pair of limbated hooks in I. nesos n. sp.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F3310DFF8A10D022D8FE25FC09.taxon	distribution	Distribution. Christmas Island, Indian Ocean. Habitat. Intertidal, found in small colonies.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F3310DFF8A10D022D8FE25FC09.taxon	etymology	Etymology. The specific name nesos is a Greek word for island and refers to the isolated type locality of this species, Christmas Island.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F33110FF8F10D02069FA03FB21.taxon	description	Figures 1 E – F, 6 A, 9, 10, Table 1	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F33110FF8F10D02069FA03FB21.taxon	materials_examined	Material examined. HOLOTYPE: 1, WAM V 7852 *, West Australia: Quondong, 17 ° 34 ' S 122 ° 09 ' E, 01. vii. 1975, collected beneath intertidal rock slabs, 45 mm in length, 5 mm in width, 48 chaetigers. PARATYPES: 1, WAM V 7853 *, 32 mm in length, 5 mm in width, 47 chaetigers, 1, AM W 37779 (mounted for SEM on two pins), both paratypes from same sample as holotype. Additional material examined. Western Australia: Dampier Archipelago Kendrew Isl. 20 ° 28 ' 30 " S 116 ° 325 ' 12 " E 7. v. 1973, 8 m, 1, WAM V 3812; transect N, Kendrew Isl. 20 ° 28 ' 28 " S 116 ° 25 ' 12 " E 11. ii. 1973, 1 – 2 m, 1 WAM V 899; Lady Nova Flat, NE Rosemary Isl. 2. xi. 1971, intertidal on oyster, 3, WAM V 677; northeast side of Cape Naturaliste, 33 ° 32 ' S 115 ° 00 " E, 26. xii. 1958, low tide, 2, WAM V 855, under stones; Port Denison, 32 ° 10 ' 51 " S 115 ° 45 ' 12 " E, 27. x. 1973, 9.7 – 10.6 m, 1, WAM V 626, on pieces of reef; Near Little Isl. off Sorento, 31 ° 34 ' 36 " S 115 ° 44.22 " E, vi. 1974, 3 m on bivalve shells, 1, WAM V 3820; Cottesloe Reef, 32 ° 35 ' 39 " S 115 ° 58.29 " E, 14. vi. 1973, intertidal on Ninella mollusc, 3, WAM V 313. Northern Territory: Trepang Bay, 11 ° 13 ' S 131 ° 57 ' E, 15. x. 1981, 5 m, 1, NTM W 26 *, from edge of reef.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F33110FF8F10D02069FA03FB21.taxon	description	Description. Holotype robust, compact (Fig. 1 E), with dark pigment patches throughout ventrum, on abdominal tori and caudas. Operculum partially fused lobes (Figs 9 A, B, 10 F), with an outer row of 17 pairs of paleae, with cylindrical and smooth shafts, flat and curved blades, with sharp-tipped denticles present on anterior half of shaft (Fig. 10 C), length of which declines towards to tip, more splayed basally (Figs 1 F, 9 A, 10 C). Inner row with 13 pairs of cylindrical paleae, slender, smooth, tapering gently with slightly curved tips (Fig. 10 C, D). Opercular papillae, 12 pairs, inserted externally to outer paleae on each lobe (Fig. 1 E). Four pairs of nuchal spines (hooks) on each side (Figs 1 F, 9 A), long shafted, and cylindrical, with subacute tips, lacking limbation; longest ones with more rounded tips, weakly recurved (Figs 1 E, F, 9 D, 10 B). Tentacular filaments compound, arranged in 20 transverse, long rows (Fig. 9 A, B). Small median organ present at dorsal juncture of opercular lobes (Figs 1 F, 9 A, 10 A). Eyespots along sides of median ridge. Palps tapered, about one third length of opercular lobes. Segment 1 (chaetiger 1) with lobe-shaped neuropodia (Fig. 9 C), with thin and flattened capillary neurochaetae, with serrations on their margins, decreasing in size towards fine tips (Fig. 10 E). Segment 2 (chaetiger 2) with four pairs of large triangular-shaped lobes, connecting branchiae to neuropodia (Figs 1 F, 9 B, C). Neuropodia of chaetiger 1 and 2 not vertically aligned, those of chaetiger 2 situated more laterally (Fig. 9 C), neurochaetae similar in structure. Fourty-five pairs of dorsal branchiae present from chaetiger 2, wide based (Fig. 9 C), not meeting mid dorsally, continuing along entire length of body, but diminishing in size posteriorly so by posterior segments very small (Figs 1 E, 9 E). Segments 3 – 5 (parathoracic) with two types of notochaetae arranged in two rows, 11 lanceolate slightly curved forming a shallow scoop with slightly twisted frayed tips and blades and 11 short, thinner lanceolate chaetae inserted between each of larger ones (Fig. 10 F). Segments 3 – 5, with lanceolate neurochaetae arranged in two tiers and significantly different in width, largest ones with smooth blades and smaller ones with textured blades. Parathoracic notochaetae more robust than parathoracic neurochaetae. Abdominal region with 43 chaetigers. Notopodia as transverse tori, with uncini, numbers per torus decreasing posteriorly. Each uncinus with two rows of vertical teeth, each with eight teeth (Fig. 10 H). Neuropodia with capillaries becoming longer posteriorly, with thin, flattened blades ornamented with thecal laminar extensions, distally forming oblique rows with elongate thin tips (Fig. 10 G), similar in structure on all abdominal chaetigers. Cauda smooth about one third the length of abdomen (Figs 1 E, 9 E). Variation. Material examined, including sexually mature females, varies from 10 – 35 mm in length without cauda, 2 – 6 mm in width, with 36 – 48 chaetigers, 14 – 18 pairs of outer paleae, 8 – 15 pairs of inner paleae, 8 – 17 pairs of opercular papillae, 1 – 4 pairs of nuchal hooks, 12 – 20 rows of compound tentacular filaments and 29 – 45 pairs of dorsal branchiae. These variations can be mainly attributed to size of the specimens, with larger animals tending to have more paleae, papillae, tentacular filaments and dorsal branchiae than smaller ones.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F33110FF8F10D02069FA03FB21.taxon	discussion	Remarks. Idanthyrsus willora is described as a new species because of the presence of four pairs of lateral lobes on segment two, which have irregular margins. The number of lateral lobes is a feature only shared with I. saxicavus (Baird, 1863) although in the latter species these lobes are long and digitiform (Kirtley 1994: Fig. 6.15.1). These two species are also distinguished by the shape of outer paleae increasing in size slightly towards tips in the new species, while in I. saxicavus they are similar in size all along the paleae. Idanthyrsus willora n. sp., can be distinguished from the other Australian species by several features. Idanthyrsus willora has straight denticles increasing in length towards the tip on the outer paleae while there are curved denticles similar in length along the blade in I. australiensis. The number and shape of lateral lobes on segment 2 also varies, (four large triangular ones in I. willora n. sp., two poorly developed in I. nesos n. sp., and three triangular ones in I. australiensis). Idanthyrsus willora n. sp., also differs from I. nesos n. sp., in having branchiae continuing to the far posterior segments. Idanthyrsus willora n. sp., can be separated from other species recorded from the Indo-Pacific by the absence of branchiae on posterior segments, whereas they are present in L. boninensis Nishi & Kirtley, 1999, described from Japan. Another Japanese species, I. okudai Kirtley, 1994, can be separated from I. willora n. sp., based on the number of pairs of lobes on segment 2 (four in I. willora n. sp., and only two in I. okudai). Two other species from from Sri Lanka, I. kornickeri Kirtley, 1994 and I. bicornis (Schmarda 1861), both poorly described (see Table 1), are distinguished from I. willora n. sp., in the shape and arrangement of the denticles on the outer opercular paleae which are curved and similar in length along the blade in I. kornickeri, in contrast to those from I. willora n. sp., which are straight and increase in length towards the tip. Idanthyrsus bicornis has these denticles arranged opposite to each other along the shaft, and in I. willora they are arranged in an alternate pattern on both margins. More over, the nuchal hooks in I. bicornis are limbate whereas they are not in I. willora n. sp.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F33110FF8F10D02069FA03FB21.taxon	distribution	Distribution. Western Australia and Northern Territory. Habitat. Specimens found in the intertidal but there is no information available as to whether this species is gregarious or occurs as single individuals, since specimens examined had all been removed from their tubes. Some of the records indicate they were attached to mollusc shells.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F33110FF8F10D02069FA03FB21.taxon	etymology	Etymology. The specific name willora is the Aboriginal word for the type locality Quondong (Endacott 1973).	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F33115FF8F10D12141FB49F92B.taxon	materials_examined	Type species. Lygdamis indicus Kinberg, 1867 by monotypy. Type locality. Bangka Straits, Java, Indonesia.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F33115FF8F10D12141FB49F92B.taxon	diagnosis	Diagnosis. Operculum with separated lobes and distal end sloped posteriorly. Opercular paleae straight and smooth, the outer ones flattened and the inner ones cylindrical, resembling spines.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F33115FF8F10D12141FB49F92B.taxon	discussion	Remarks. The monophyly of Lygdamis was not assessed in the phylogenetic analyses undertaken by Capa et al. (2012) but all species currently assigned to the genus conform to the above diagnosis. The genus is known from 19 species (Kirtley 1994; Lechapt & Kirtley 1998; Nishi & Kirtley 1999) including the new species described here. Kirtley (1994) suggested that the shape and colouration of the median organ are useful characters especially in live material, but after fixation and preservation they become so distorted as to loose their usefulness.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F33115FF8210D02374FDC6FE62.taxon	description	Figures 6 B, 11 A – D, 12 – 13, Table 2	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F33115FF8210D02374FDC6FE62.taxon	materials_examined	Material examined. HOLOTYPE: As Eupallasia giardi Augener, 1927 (identified by Kirtley, 1994) off Disaster Bay, New South Wales: Australia: 37 ° 05 ' S 150 ° 05 ' E; 01. x. 1914; 53 – 73 m; ZMH V- 9569; Th. Mortensen's Endeavour, 20 mm in length (including damaged posterior end), 2 mm in width. PARATYPES: 3, from same sample as holotype. Additional material examined. New South Wales: (Tasman Sea) off The Entrance, 33 ° 14 ' 12.6 " S 156 ° 10 ' 40.8 " E, 02. v. 1989, 133 m, 1, AM W 27564 *, 16 mm long, 1 mm wide, 24 segments; AM W 27565 (mounted for SEM).	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F33115FF8210D02374FDC6FE62.taxon	description	Description. Holotype flesh coloured, no pigmentation present. Body elongate and slender (Fig. 11 A). Operculum with completely separated lobes, with distal end sloping; longer than wide (Fig. 11 A, B). Length of operculum more than one third the length of the body (Fig. 11 B, C). Paleae displayed in two rows, outer row with 42 pairs of paleae, with flattened shafts, smooth margins with tips curved outwards (Fig. 13 C); inner row with 30 pairs of paleae, cylindrical and tapering slowly to blunted tips (Fig. 13 C). Opercular papillae, 12 pairs, peripheral to outer row of paleae on each lobe, elongate, increasing in length towards nuchal spines (Figs 12 C, 13 B), pair adjacent to nuchal spines about one quarter length of outer paleae. One pair of nuchal spines, strongly recurved, margins smooth, with pointed tips (Figs 11 A, 12 D, 13 B). Tentacular filaments compound arranged in 19 horizontal rows. Eye spots present along sides of median ridge. Pair of elongate grooved palps with crinkled margins on either side of median ridge, about one quarter length of opercular lobes. Segment 1 with lobe-shaped neuropodia (Fig. 12 B – E), capillaries absent. Segment 2 with three triangular lateral lobes on each side of body (Fig. 12 E), with fine capillaries arranged in compact fascicle, inserted in neuropodia, posterior to U-shaped buccal organ (Fig. 12 B). Eight pairs of dorsal branchiae present from segment 2, with narrow base and tapering gently to a fine tip, slightly ridged, size decreasing posteriorly, not meeting mid dorsally (Fig. 11 A). Segments 3 – 6 (parathoracic) with notochaetae arranged in two transverse rows, six long lanceolate, with tapering frayed tips, blades slightly concave and textured, and six capillaries inserted imbetween (Fig. 13 D). Segments 3 – 6 with two types of neurochaetae, six lanceolate, with frayed twisted tips, and compact thecae and fine capillaries with thin flattened blades with short narrow thecae, alternating with lanceolate (Fig. 13 E). Parathoracic notochaetae more robust than neurochaetae. Abdominal region with 19 chaetigers. Notopodia as transverse tori, with number of uncini decreasing posteriorly. Each uncinus with two vertical rows of eight teeth (Fig. 13 F). Neuropodia with capillaries arranged in discrete fascicle, with thin, flattened blades with short, narrow thecal rings. Cauda smooth, about length of last five chaetigers (Fig. 11 C). Variation. Material examined, including sexually mature specimens, varies from 15 – 18 mm in length without cauda, 2 – 3 mm in width, with 19 – 23 chaetigers, 42 – 44 pairs of outer paleae, 30 – 36 pairs of inner paleae, 6 – 8 teeth on each row of uncini and 15 – 19 pairs of tentacular filaments. Pigmentation present along the rows of some tentacular filaments. We suggest that these variations are to some extent size dependent and pigmentation may be related to methods of preservation and subsequent storage.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F33115FF8210D02374FDC6FE62.taxon	discussion	Remarks. Kirtley (1994) described this species based on material examined by Augener (1927) who had identified it as L. giardi McIntosh, 1885. The two species differ not only in the range of the number of pairs of outer paleae, 42 – 44 in L. augeneri and 16 – 20 in L. giardi, but also in the shape of the outer paleae. I. augeneri has outer paleae with curved flattened tips (Fig. 13 C) whereas those of L. giardi have asymmetrical flattened pointed tips (Fig. 15 C).	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F33115FF8210D02374FDC6FE62.taxon	distribution	Distribution. South-eastern Australia. Habitat. Shelf depths of 53 – 133 m.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F33118FF8710D0258AFB2AFDBA.taxon	description	Figures 6 B, 11 D, E, 14, 15, Table 2	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F33118FF8710D0258AFB2AFDBA.taxon	materials_examined	Material examined. HOLOTYPE: as Sabellaria (Pallasia) giardi McIntosh 1885; off Port Jackson, New South Wales: 33 ° 51 ' S 151 ° 22 ' E; 03. vi. 1874; 64 m, BMNH 1885.12. 1.6. Specimen described as 7 mm in length, 2 mm in width and damaged, but currently in worse condition and only anterior end and posterior chaetigers are in vial. Additional material examined. New South Wales: Sydney, Malabar, 33 ° 49 ' 30 " S 151 ° 21 ' 36 " E, 27. ii. 1973, 66 m, 8, AM W 6191 *; 22. v. 1972, 28 m, 6, AM W 6185 *; North Head, 33 ° 49 ' 25 " S 151 ° 20 ' 43 " E, 27. ii. 1973, 63 m, 81, AM W 26682 (mounted for SEM); 7, AM W 6192 *, 2 km east of Long Bay, 33 ° 58 ' S 151 ° 15 ' E, 66 m, 2, AM W 6188 *, 5, AM W 6189 *; off Maroubra, 33 ° 57 ' 06 " S 151 ° 17 ' 37 " E, 20. vii. 1972, 58 m, 1, AM W 6190 *; Tasman Sea, off Sydney, 33 ° 53 ' 06 " S 151 ° 21 ' 54 " E, 30. iii. 1972, 76.8 m, 2, AM W 27553 *, from gritty calcareous mud; off Eden, 36 ° 39 ' 30 " S 150 ° 37 ' 48 " E, 01. iv. 1972, 86 m, 1, AM W 27554, from muddy quartz sand; off Broken Bay, 33 ° 37 ' – 39 ' S 152 ° 04 – 02 ' E, 10. xii. 1980, 1, AM W 27555 *; Sydney, off North Head, 33 ° 48 ' 46 " S 151 ° 20 ' 59 " E, 60 m, 13. iv. 1989, 1, AM W 20677 *, from sandy substratum; 15. ii. 1973, 66 m, 2, AM W 27556 *; Malabar, 33 ° 58 ' S 151 ° 19 ' E, 1973, 1, AM W 27557 *; 2, AM W 27558 *; 45 m, 1, AM W 27559; 17. v. 1972, 69 m, 3, AM W 27560 *; 22. v. 1972, 75 m, 1, AM W 27561; 02. i. 1973, 54.9 – 60.3 m, 4, AM W 27562 *; off Tuggerah Lakes, 33 ° 20 ' S 151 ° 39 ' 24 " E, 11. iii. 1972, 60 m, 5, AM W 27563. Tasmania, Bass Strait: King Island, 35 km northeast of Cape Wickham, 39 ° 16 ' 00 " S 144 ° 05 ' 24 " E, 23. xi. 1981, 82 m, 1, MV F 78887 *, from sandy shell. Northern Territory: 11 km, NE of Cape Ford, Anson Bay, Beagle Gulf, 13 ˚ 25 ' 02 " S 129 ˚ 55 '. 98 " E, 16 m, 01. x. 1993, NTM 23481.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F33118FF8710D0258AFB2AFDBA.taxon	description	Description. As the holotype is incomplete and cut into portions, this description is largely based on material collected from off the NSW coast in the region of the type locality. Body robust, preserved material pale cream, paleae golden with dark brown pigmentation on opercular lobes and tentacular filaments in some specimens. Operculum completely divided into two free long lobes with distal ends oblique (Figs 11 E, 15 B). Operculum with two rows of paleae, outer row with 20 pairs of paleae, cylindrical at base and flattened, with nearly symmetrical pointed tips, with smooth margins (Fig. 15 C); inner row with ten pairs of paleae, smooth cylindrical shafts with blunt tips (Fig. 15 D). Opercular papillae, 11 pairs, peripheral to outer paleae of each lobe (Figs 11 D, E, 14 E, F). One pair of nuchal spines on each side (Figs 11 D, 14 F), stout, strongly recurved without sharp limbations on concave margin. Tentacular filaments compound, arranged in 15 – 19 rows (Fig. 14 C). Median organ present, with eyespots on either side. Pair of ringed palps, less than quarter length of opercular lobes (Fig. 14 B). Segment 1 with lobe shaped neuropodia (Fig. 14 A), chaetae absent. Segment 2 with three pairs of triangular shaped lateral lobes, connecting branchiae to neuropodia (Figs 11 E, 14 D). Fourteen pairs of branchiae present from segment 2 (Fig. 14 D) not extending to posterior segments. Branchiae, taper gently and evenly to subacute tip (Fig. 14 D). Segments 3 – 6 (parathoracic), with two types of notochaetae arranged transversely, 7 – 8 lanceolate, with frayed twisted tips, interspersed with 7 – 8 capillaries (Fig. 15 E). Segments 3 – 6 with two types of neurochaetae arranged transversely, five lanceolate and five thinner capillaries with compact thecae alternating with lanceolate (Fig. 15 F). Abdominal notopodia with uncini decreasing in number posteriorly, each uncinus with two vertical rows each with about 7 – 9 teeth (Fig. 15 H). Neuropodia with capillaries, ornamented with irregular thecal laminar extensions (Fig. 15 G), similar in structure on all abdominal chaetigers. Cauda smooth about the length of five posterior abdomen segments (Figs 11 E, 14 H). Variations. Material examined, including sexually mature specimens, varies from 12 – 43 mm in length (without cauda), 2 – 5 mm in width, with 17 – 32 chaetigers, 16 – 20 pairs of outer paleae, 12 – 14 pairs of inner paleae, 8 – 12 pairs of opercular papillae, 17 – 19 pairs of tentacular filaments and 9 – 10 pairs of dorsal branchiae.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F33118FF8710D0258AFB2AFDBA.taxon	discussion	Remarks. McIntosh (1885) dissected the holotype and cut up the body into sections and not all are still present in the vial, hence we have also illustrated a complete specimen from the same location (AM W 6191). The diagnostic characters of L. giardi are the presence of 16 – 20 pairs of outer paleae which are straight, nearly symmetrical with distally smooth margins and pointed tips, 15 – 19 rows of tentacular filaments and three pairs of lateral lobes on segment 2. Lygdamis giardi can be separated from L. augeneri by the number of outer paleae, 16 – 20 and 42 – 44 respectively and by the structure of their tips L. giardi having flattened nearly symmetrical tips whereas L. augeneri has sharply bent outer paleae with broad, blunt flattened tips (see Fig. 15 C and 13 C).	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F33118FF8710D0258AFB2AFDBA.taxon	distribution	Distribution. Lygdamis giardi has a discontinuous distribution around Australia, and this may represent a true record or reflect the lack of collecting on the shelf around Australia; currently reported from New South Wales, Tasmania and Northern Territory. This species has also been recorded from the Moluccas Islands, Indonesia (Caullery, 1913) although this material has not been checked. Habitat. Inshore at depths of 16 – 86 m, in sediments ranging from coarse sand to muddy sand. Most records of solitary individuals but one sample of 81 individuals suggesting they can form small colonies.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F3311DFF8510D026E2FB3BFBD1.taxon	description	Figures 6 B, 11 F, G, 16, 17, Table 2.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F3311DFF8510D026E2FB3BFBD1.taxon	materials_examined	Material examined. HOLOTYPE: QM G 10505 *, 26 + mm in length (incomplete), 3 mm in width, 26 + segments (incomplete). PARATYPES: QM G 10338 *, 14 + mm in length (incomplete), 2 mm in width, 24 + segments (incomplete), plus one specimen mounted for SEM. All material from Queensland: Moreton Bay, Peel Island, 0.8 km southeast of South West Rocks, 27 ° 30 ' S, 153 ° 21 ' E, from shell, grit and sand, 7.62 m, collected vi. 1970, 7 m, and iii. 1970. Additional material examined.? Northern Territory: New Year Island, 10 ° 55 ' S 133 ° 02 ' E, 14. x. 1982, LWM, 1, NTM W 1121 *, from coarse sandy bottom.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F3311DFF8510D026E2FB3BFBD1.taxon	description	Description. Holotype, stout, colourless with golden yellow paleae and dark brown nuchal hooks. Operculum completely divided into two free divergent lobes, longer than wide (Figs 11 F, H, 16 A, B) with distal end sloping with two rows of paleae (Fig. 17 B). Outer row with 29 paleae, blades finely textured, colourless, margins straight, symmetrical, tapering to acute tip with small distal denticles (Fig. 17 B, C), inner row with 12 pairs of spines, yellow, smooth and acute tipped (Fig. 17 D); outer paleae longer than inner (Fig. 17 B) but thinner. Opercular papillae, 18 pairs peripheral to outer row of paleae on each lobe, about one third length of outer paleae (Figs 11 G, 16 C, 17 B). One pair of nuchal hooks present, strongly recurved, margins smooth, convave with pointed tips (Figs 11 F, 16 B, C). Tentacular filaments compound, arranged in about 30 horizontal rows (Fig. 16 A, B). Medial organ present at dorsal base of opercular lobes (Figs 11 F, 16 A, 17 A), palps shorter than a third of opercular lobes. Eyespots absent. Segment 1 with lobe shaped neuropodia (Fig. 16 A), chaetae absent. Segment 2 with three pairs of triangular lateral lobes (Figs 11 F, G, 16 B), with fine capillaries with plumose margins and extended fine tips, in compact fascicle, inserted posteriorly to U-shaped buccal organ. Twelve pairs of branchiae present from segment 2. Branchiae with broad base, tapering gently to rounded tip (Figs 11 G, 16 B), from chaetiger 7 onwards, triangular, larger in size in anterior abdominal segments, absent from posterior segments of incomplete type material. Segment 3 – 6 (parathoracic) with chaetae inserted in two transverse rows with eight lanceolate chaetae with frayed twisted tips (Fig. 17 E), and fine capillaries with compact thecae interspersed between them (Fig. 17 E), we suspect that the lanceolate chaetae can retract but not the capillaries. Segments 3 – 6 with two types of neurochaetae arranged in oblique transverse row, five lanceolate with fine capillaries. Abdominal region with 20 chaetigers, but holotype is incomplete lacking cauda. Notopodia as transverse tori, with numbers of uncini per row decreasing posteriorly, uncini with two vertical rows of teeth, each row with eight teeth. Neuropodia with capillaries arranged in discrete fascicles with compact theca, distal margins becoming more elongate towards tip (Fig. 17 F). Variation. Paratypes exhibit the following variation, number of rows of tentacular filaments (15 – 30), outer paleae (27 – 29 pairs), inner paleae (12 – 16 pairs). A specimen from Northern Territory is 25 mm in length, 4 mm in width and has 32 pairs of outer paleae, 20 pairs of inner paleae, one pair of hooks, 26 pairs of opercular papillae and parathoracic segments with 6 – 7 notopodial lanceolate chaetae. This specimen has considerably more inner paleae than the holotype which has 12 pairs and the paratype mounted for SEM has 13. At this stage we are referring the Northern Territory material to the species, Lygdamis wambiri n. sp., as it certainly does not represent either of the other two species of Lygadamis recorded from Australia, but additional material from this locality should be examined to determine if this is correct or whether material from the Northern Territory represents another undescribed species of the genus.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F3311DFF8510D026E2FB3BFBD1.taxon	discussion	Remarks. Lygdamis wambiri n. sp., is distinguished from other species in the genus by a combination of characters: 27 – 29 pairs of outer paleae, 12 – 16 pairs of inner paleae, three pairs of lateral lobes on segment 2 and having straight markedly asymmetrical outer paleae with distally denticulate margins (Table 2). Lygdamis wambiri n. sp., can be distinguished from the other species of the genus recorded from Australia by the shape of the distal margins of the outer paleae, in L. wambiri n. sp., they are finely denticulate whereas they are smooth in the other Australian species. Lygdamis wambiri n. sp., has short inflated lanceolate notochaetae on parathoracic segments, whereas the two Australian species have longer less inflated lanceolate notochaetae compared to those of the other two species. Lygdamis wambiri n. sp., has branchiae present on posterior segments whereas L. augeneri and L. giardi lack them posteriorly. The only other species which appears to have distal margins of the outer paleae denticulate is L. robinsi Jeldes and Lefevre, 1959 described from off Angola, but this species is poorly known with regard to other characters. Lygdamis wambiri n. sp., can be separated from other species described from the Indo-Pacific by the number of opercular papillae: L. bhaudi Kirtley, 1994, described from Madagascar, has 3 – 4 pairs whereas L. wambiri has 12 – 17. Lygdamis curvatus (Johansson, 1922) and L. japonicas Nishi & Kirtley, 1944, both described from Japan, have smooth distal tips of outer paleae in contrast to the denticulate tips of L. wambiri n. sp. Lygdamis ehlersi Caullery, 1913, described from Indonesia has only five pairs of outer paleae in contrast to the 27 – 29 pairs present in the new species.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F3311DFF8510D026E2FB3BFBD1.taxon	distribution	Distribution. Moreton Bay, Queensland, with a possible record from Northern Territory. Habitat. Inshore in sheltered shallow waters (around 7 m).	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F3311DFF8510D026E2FB3BFBD1.taxon	etymology	Etymology. The specific name wambiri is an Aboriginal word for the coast (Endacott 1973) and refers to the extensive coast line of Moreton Bay a shallow protected bay, the type locality of the species.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F3311FFF8510D020D1FA09F9FF.taxon	materials_examined	Type species. Phalacrostemma cidariophilum Marenzeller, 1895, by monotypy. Type locality. Off Pelagossa Island, Adriatic Sea.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F3311FFF8510D020D1FA09F9FF.taxon	diagnosis	Diagnosis. Buccal flaps present, arrangement of outer paleae in a spiral on each lobe, in most species.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F3311FFF8510D020D1FA09F9FF.taxon	discussion	Remarks. Kirtley (1994) recognised 10 species within the genus and four species have been described since then (Lechapt & Kirtley 1998 and present paper). This represents the first record of the genus from Australia and as this genus currently is only known from deep water, this is not surprising given the limited offshore sampling carried out in Australia. As can be seen from Table 3, many species of the genus are extremely poorly known and have been separated based solely on paleal features. A complete revision of the genus is therefore urgently needed.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F3311FFFB810D12238FE0FFC22.taxon	description	Figures 6 A, 18, 19, 20, Table 3	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F3311FFFB810D12238FE0FFC22.taxon	materials_examined	Material examined. HOLOTYPE: AM W 27566 *, New South Wales: Tasman Sea, off Newcastle, 33 ° 30 S 152 ° 06 ' E, iv. 1971, 548.6 m, in 2 pieces 10 + - 5 + mm in length, 2 mm in width, 15 - 4 chaetigers, attached to mollusc shell. PARATYPES: Tasman Sea, off Newcastle, 32 ° 30 ' S 152 ° 06 ' E, iv. 1971, 548.6 m, 1, AM W 27567; 457.2 – 502.9 m, 6, AM W 27568 * attached to mollusc shell; 33 ° 40 ' S 152 ° 45 ' E, 15 vi. 1971, 6, AM W 27569; 33 ° 30 ' S 152 ° 06 ' E, iv. 1971, 1, AM W 26681 (mounted for SEM). Additional material examined. MV F 78859, Stn S 05 / 84 29 from Tasman Sea, eastern slope 70 km S of Gabo Island, Victoria (1 specimen).	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F3311FFFB810D12238FE0FFC22.taxon	description	Description. Holotype with body stout, tapering to elongated abdomen, reddish brown with darker areas along noto- and neuropodia. Operculum completely divided into two free short lobes with distal end perpendicular to longitudinal axis (Figs 18 A, B, 19 A – D). Operculum with outer row of 12 pairs of paleae, arranged spirally (Fig. 18 C); paleae yellow as simple spines with acute tips and compact thecae with margins straight and not expanding (Figs 19 E, 20 B, C); inner row with two pairs of short spines, yellow and tapering tip (Fig. 18 C). Ten pairs of robust and tapering opercular papillae, peripheral to outer paleae (Figs 18 C, 19 B, C), almost as long as outer paleae. Five pairs of flattened nuchal hooks, with limbation present on concave side (Figs 18 B, C, 19 D, 20 B). Medial organ present at dorsal junction of opercular lobes (Fig. 18 C, see Capa et al., 2012). Eyes not observed. Three pairs of simple tentacular filaments (Fig. 19 B). A pair of buccal flaps below tentacular filaments (Fig. 19 A). Palps shorter than length of operculum (Fig. 19 B). Segment 1 (chaetiger 1) with long and tapering neuropodial lobe (Fig. 19 A) and capillary neurochaetae (Fig. 19 B, C). Segment 2 (chaetiger 2) with two triangular lateral lobes connecting branchiae to neuropodia (Figs 18 B, 19 C). Eight pairs of dorsal tapering branchiae present from chaetiger 2, not meeting mid-dorsally (Fig. 18 B), reducing in size posteriorly but present on posterior segments. Segments 3 – 6 (parathoracic) with two types of notochaetae inserted in transverse rows with two types of notochaetae; eight lanceolate and fine capillaries interspersed (Fig. 20 D, E). Segments 3 – 6 with two types of neurochaetae arranged transversely, four lanceolate and fine short capillaries interspersed (Fig. 20 F). Notopodial lanceolate much stouter than those in neuropodia (Fig. 20 D – F). Abdominal notopodia as erect expanded tori, with uncini decreasing in numbers posteriorly. Each uncinus with three vertical rows of teeth imbricated (Fig. 20 H). Neuropodia with long fine capillaries with compact thecae arranged in discrete fascicles in two tiers (Fig. 20 G). Cauda smooth and about one third length of abdomen (Fig. 18 A). Variations. The paratypes including sexually mature females, vary from 7 – 18 mm in length (without cauda) none of which is complete, 2 mm in width with 12 – 22 chaetigers, 12 – 16 pairs of outer paleae, 2 – 3 pairs of inner paleae, 7 – 13 pairs of opercular papillae, and with 3 – 9 pairs of dorsal branchiae. All these variations could be size dependent. None of the material is well preserved and fragile.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F3311FFFB810D12238FE0FFC22.taxon	discussion	Remarks. Phalacrostemma maloga n. sp., is characterised by a unique combination of: 12 – 16 pairs of outer paleae with straight edges and without expanded thecae, five pairs of nuchal spines, three pairs of tentacular filaments, buccal flaps present and with some abdominal uncini provided with three vertical rows of teeth. Other species without expanded thecae on the outer paleae (see Table 3) include P. setosa Treadwell, 1906 described from the Caribbean which has irregular edges and P. cidariophilum Marezeller, 1895 described from the Northern Adriatic Sea, which has straight edges but lacks tentacular filaments. The closest species geographically is P. tenue Lechapt & Kirtley, 1998, described from deep water off New Caledonia but this has only two pairs of nuchal spines whereas P. maloga n. sp., has five pairs. Most species of this genus, which typically occur in deep waters (see Table 3), are poorly known. We suggest that the number of pairs of lateral lobes on segment 2, the presence of buccal flaps and the number of pairs of tentacular filaments are useful characters to distinguish between species. The dentition of the abdominal uncini, with three rows of teeth instead of the two rows typical of other sabellariids, may be a synapomorphy for the species. But it could have also been overlooked in other species of the genus and therefore needs further study.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F3311FFFB810D12238FE0FFC22.taxon	distribution	Distribution. South-eastern Australia, type known only from type locality and from a single record from south of Gabo Island, off the coast of Victoria .. Habitat. Continental shelf in depths of 457 – 548 m and attached to mollusc shells in soft sediment substrates forming small colonies.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F3311FFFB810D12238FE0FFC22.taxon	etymology	Etymology. The specific name maloga is an Aboriginal word for deep water (Endacott 1973) and refers to the habitat of this species.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F33122FFB810D0204AFD06F987.taxon	materials_examined	Type species. Sabella alveolata Linnaeus, 1767, by monotypy. Type locality. Europe, Hartman (1959).	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F33122FFB810D0204AFD06F987.taxon	diagnosis	Diagnosis. Operculum completely divided in two symmetrical lobes, inner paleae arranged in two rows (with mid and inner paleae) and three parathoracic segments.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F33122FFB810D0204AFD06F987.taxon	discussion	Remarks. The monophyly of Sabellaria has not being assessed in the phylogenetic analysis undertaken by Capa et al. (2012) and the above diagnosis includes characters that are shared by all species in the genus. The total number of species of Sabellaria described to date (including those in Nishi et al. 2010, Santos et al. 2011 and herein) are 39. Features such as the shape and ornamentation of paleae and nuchal spines (as in Nishi et al. 2010, Santos et al. 2011) together with some others have been used for the comparison of the species herein (Table 4). According to Kirtley (1994), all species within the genus are gregarious and capable of forming large colonies and reefs along shores where suitable hydrodynamic and sedimentary conditions occur but the new species described below are known from only a few solitary individuals. First record of the genus in Australian waters.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F33122FFB010D022E0FD9BFDD9.taxon	description	Figures 6 B, 21, 22, 23, Table 4	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F33122FFB010D022E0FD9BFDD9.taxon	materials_examined	Material examined. HOLOTYPE: 1, AM W 27333 *, 18 mm long, 2 mm wide, 22 chaetigers. PARATYPE: AM W 27195 *, 19 mm long, 3 mm wide, 22 chaetigers, from Albany Passage, Queensland, Australia, 10 ° 45 ' S 142 ° 37 ' E, 10 m; paratype AM W 37820.001 (mounted for SEM) Shell Island, East Arm, Darwin Harbour, Northern Territory, St. RH 88 / 37, 12 ˚ 29.4 '' S 130 ˚ 53.2 '' E, rocky intertidal, 18 iii. Mar 1988. Additional material examined. Northern Territory: West of Elizabeth Reef, Cape Hotham, Beagle Gulf, 12 ˚ 4.02 ' S 131 ˚ 20.04 ' E, 21 m 29. x. 1993, 1, NTM W 23479, 09. x. 93, 1, NTM W 23477; 2 nm E of Quail Isl., Bynoe Harbour, Beagle Gulf, 12 ˚ 31.02 ' S 130 ˚ 28.98 ' E, 28 m, 6. x. 1993, 1, NTM W 23478; East Arm, Shell Island, Darwin Harbour, 12 ° 29 ' S 130 ° 53 ' E, 18. iii. 1988, low water, rocky intertidal, 1, AM W 37820 *, (mounted for SEM); Darwin Harbour, 12 ˚ 32.72 ' S 130 ˚ 51.32 ' E, DW 105 A, 7 m, coll. 24. iii. 1994, 1, NTM W 15033; 12 ˚ 32.72 ' S 130 ˚ 51.32 ' E, DW 1054, 7 m, coll. 24. iii. 1994, 1, NTM W 15031; 12 ˚ 36.17 ' S 130 ˚ 46.87 ' E, DW 76 A, 4 m, coll. 17. iii. 1994, 1, NTM W 15023; 12 ° 32.9 ' S 130 ° 50.02 E, 1, D 103 A, 8 m, coll. 15. vi. 1993, 1, NTM W 10042; 12 ° 31.98 ' S 130 ° 50.09 ' E, DH 100 A, 24. iii. 1994, 15 m, coll. 24. iii. 94, 1, NTM W 15030; 12 ˚ 29.27 ' S 130 ˚ 49.48 ' E, DW 204 A, 14 m, coll. 25. iii. 1994, fine & coarse sand, 1, NTM W 15026; 12 ˚ 30.07 ' S 130 ˚ 55.88 ' E, DW 187 A, 4 m, coll. 22. iii. 1994, 2, NTM W 15027; 12 ˚ 26.93 ' S 130 ˚ 46.5 ' E, DW 544, 14 m, coll. 16. iii. 1994, 1, NTM W 15032; 12 ˚ 34 ' S 130 ˚ 52.22 ' E, DW 116 A, 12 m, coll. 18. iii. 1994, 1, NTM W 15024; 12 ˚ 34 ' S 130 ˚ 52.22 ' E, DW 116 A, 18. iii. 1994, 12 m, 1, NTM W 15035, 12 ˚ 31.68 ' S 130 ˚ 51.3 ' E, DW 101 A, 8 m, coll. 24. iii. 1994, 1, NTM W 15028; 12 ˚ 31.7 ' S 130 ˚ 51.28 ' E, 8 m, 14. vii. 1993, 1, NTM W 23482; 12 ˚ 31 ' 54 '' S 130 ˚ 50 ' 25 '' E, 24. iii. 1994, 1, NTM W 15029; 12 ˚ 27 ' 57 '' S 130 ˚ 46 ' 30 '' E, 08. vi. 1994, 1, NTM W 10046; 12 ˚ 28 ' 57 '' S 130 ˚ 50 ' 25 '' E, 17. iii. 1994, 1, NTM W 15034; Melville Bay, Gove, 12 ˚ 11.7 ' S 136 ˚ 41.3 ' E, 8. vii. 1991,7.5 m, 1, NTM W 16880, 1, NTM W 16887, 1, NTM W 16890; Melville Bay, Gove, 12 ˚ 12 ' 31 '' S 136 ˚ 40 ' 30 '' E, 9. vi. 1991, 1, NTM W 16882, 1, NTM W 6881; 12 ˚ 10 ' 31 '' S 136 ˚ 39 ' E, Nov 1991 – March 1992, 1, NTM W 08265; NW of Granite Islet, Melville Bay, Gove, 12 ˚ 14 ' S 136 ˚ 40 ' E, 22. iii. 1992, 12.9 m, 1, NTM W 16891; Bing Bong, McArthur River, Gulf of Carpentaria, 15 ˚ 37 ' S 136 ˚ 23 ' E, ix. 1992 – iii. 1993, 8 m, 1, NTM W 007737, 1, NTM W 16899.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F33122FFB010D022E0FD9BFDD9.taxon	description	Description. Holotype with speckled faint brown pigment on ridged cilated dorsum (Fig. 21 C), rectangular pigment patches across ventrum of chaetiger 5 (3 rd parathoracic). Operculum similar in width to length, with lobes completely separated, and three rows of golden paleae arranged in concentric semicircles (Figs 21 B, 23 A). Outer row with 12 geniculate paleae with flattened blades, smooth lateral margins and serrated distal margins with a middle tapering denticle occupying half of distal end which has marked surface ornamentation (Figs 21 B, 22 G, 23 D, E, 22 D, E). Twelve pairs of middle geniculate paleae, with smooth concave shafts and tapering blades pointing towards to outer edge of operculum (Figs 21 B, 22 H, 23 C, F). Fourteen pairs of inner paleae, strongly geniculated, concave blades, similar to middle paleae but wider, shorter and pointing inwards to center of operculum (Fig. 21 B). Outer paleae longer than those of inner rows. Opercular papillae, 19 pairs, short, peripheral to outer paleae on each lobe (Figs. 21 B, C, 22 D, 23 B). Three pairs of nuchal spines present (four in paratype), stout, flattened, slightly curved, with smooth blunt tips (Figs 21 B, 22 I, 23 G). Tentacular filaments, compound, arranged in 14 horizontal rows (Fig. 22 A). Median ridge and median organ present. Eyespots present. Palps slightly longer than opercular lobes. Segment 1 (chaetiger 1) with lobe-shaped neuropodia, with capillary neurochaetae, on either side of U-shaped buccal organ (Fig. 22 B). Segment 2 (chaetiger 2) with one elongate triangular shaped lateral lobe, connecting branchiae to neuropodia, with fine capillary neurochaetae (Figs 21 A, C, 22 E, F). Neuropodia of chaetigers 1 and 2 not vertically aligned, those of chaetiger 2 situated more laterally (Figs 21 B, 23 H). Sixteen pairs of dorsal branchiae present from segment 2 (Figs 22 F, 23 H), continuing along entire length of body, with broad base and strongly ciliated ridges and slightly pigmented, all detached except one, which tapers distally with filiform tip, more remain on paratype (Figs 21 A, 23 H). Segments 3 – 5 (parathoracic) with two types of notochaetae arranged transversely, seven (many represented only by stumps) lanceolate with frayed tips and short, flattened smooth shafts with strongly frayed tips, and fine short capillaries with expanded theca inserted between lanceolate (Fig. 23 I). Segments 3 – 5 with two types of neurochaetae arranged in compact fascicle, with seven lanceolate and seven fine capillaries (Fig. 23 H). Parathoracic notopodia larger than neuropodia (Fig. 22 J). Abdominal region with 16 chaetigers. Notopodia as elongate, erect tori, with long-handled uncini, numbers per torus decreasing posteriorly, each uncinus with two vertical rows, each with seven teeth. Neuropodia becoming considerably elongated posteriorly (Fig. 22 I) with ventral bundle of capillaries (Fig. 23 J). Cauda smooth and more than half length of abdomen (Figs 22 I, K, 23 L). Paratype a gravid female with eggs released from between posterior abdominal notopodia. Variations. Material examined varies from 4 – 12 mm in length, 1 – 2 mm in width with 12 – 22 chaetigers, but much of the material is posteriorly incomplete, 12 – 16 pairs of outer paleae, 10 – 12 pairs of middle paleae, 16 – 19 pairs of opercular papillae and 3 – 4 pairs of nuchal spines.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F33122FFB010D022E0FD9BFDD9.taxon	discussion	Remarks. Sabellaria kooraltha n. sp., is characterised by the following combination of characters: 12 – 16 pairs of outer paleae with a smooth median distal spike which has marked thecal bands, and two smaller, smooth basal, lateral spines (Fig. 23 D, E) and 11 – 12 pairs of middle paleae which are geniculate and with a broad based triangular shaped central tooth (Fig. 23 F), one pair of lateral lobes on chaetiger 2 and branchiae absent from posterior abdominal segments. This species can easily be separated from the other species of Sabellaria recorded from Australia by the structure of the outer paleae. Sabellaria kooraltha n. sp., has a smooth midline tooth or spike although the thecae are very conspicuous on the outer paleae (Fig. 23 D, E) whereas Sabellaria lungalla n. sp., (this paper) has a finely serrated distal plume on the outer paleae (Figs 24 G, 25 C). With regard to other Indo-Pacific species which have one kind of middle paleae, both S. gilchristi (McIntosh, 1925) described from South Africa, and S. javanica Augener, 1934, described from Indonesia, have outer paleae with midline plume whereas S. kooraltha n. sp., has a smooth distal spike. The value of SEM is highlighted for the characterisation of this species although the strongly ornamented midline plume of the outer paleae can be seen under high magnification using a compound microscope. Details of the other described species of Sabellari a are given in Table 4.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F33122FFB010D022E0FD9BFDD9.taxon	distribution	Distribution. North-eastern Australia, from Albany Passage, just south of Cape York in North Queensland and Darwin Harbour, Northern Territory. Habitat. No information on habitat is available for the type locality, but co-ordinates suggest a depth of 10 m and collected by dredging. Non type material collected from muddy substrates from low water mark to 15 m.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F33122FFB010D022E0FD9BFDD9.taxon	etymology	Etymology. The specific name kooraltha is an Aboriginal word for spotted (Endacott 1973) and refers to the pigmented nature of the dorsum.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F3312AFFB610D026D9FD10FDBA.taxon	description	Figs 6 B, 24, 25, Table 4	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F3312AFFB610D026D9FD10FDBA.taxon	materials_examined	Material examined. HOLOTYPE: NTM W 16893, Northern Territory: Bing Bong, McAthur River, 15 ° 23 ' S 136 ° 30.35 ' E, 17. iii. 1993, complete except for cauda, 3 mm in length, 1 mm in width, 12 chaetigers. PARATYPES: 2, NTM W 16896), Drimmie Arm, Melville Bay, Gove, 12 ° 14 ' S 136 ° 42 ' E, 4 mm in length, 1.5 mm in width for 14 chaetigers, 3.45 mm in length 1.00 mm in width for 14 chaetigers; 2, NTM W 10043, Darwin Harbour, D 92 A, 12 ° 29.52 ' S 130 ° 50.18 ' E, 7 m, 14. vii. 1993, 7 m, 3 mm in length, 1 mm in width, 18 chaetigers, 5 mm in length, 1 mm in width, 16 segments. Additional material examined. Northern Territory: Bing Bong, McArthur River, Gulf of Carpentaria, 15 ° 36.72 ' S 136 ° 23.45 ' E, 18. iii. 1993, 1, AM W 37780 (mounted for SEM); Darwin Harbour, 12 ° 29 ' S 130 ° 50.18 E, 14. vi. 1993, 1, NTM W 10043; 12 ° 32.72 ' S 130 ° 51.32 ' E, 24. iii. 1994, 1, NTM W 24725; 12 ° 32.9 ' S 130 ° 50.02 ' E, 15. vi. 1993, 1, NTM W 10042; 1, NTM W 16889. Majority of this material is posteriorly incomplete. Western Australia: Mermaid Sound, Dampier Archipelago, 20 ° 52 ' S 116 ° 44 ' E, WAM V 3816, 1981.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F3312AFFB610D026D9FD10FDBA.taxon	description	Description. Holotype with short compact body, colourless except for scattered pigment around base of opercular lobes (Fig. 24 A). Operculum short, similar in width to length, with lobes completely separated along its length. Paleae arranged in three semicircular concentric rows (Fig. 25 A, B, D). Outer row with 22 pairs of geniculate golden paleae with flattened blades, smooth lateral margins and distal tip with midline plume and 5 – 7 smaller denticles on each side (Figs 24 F, 25 A – C). Middle row with 10 – 11 pairs of paleae, short, geniculate, with broad and concave blades directed outwards and tapering tips and inner row with 11 pairs of similar in shape paleae, slightly smaller arranged in between the middle paleae directed inwards, broadly flattened, geniculate, yellow-brown, smooth, pointed tips (Fig. 25 A, D). Outer paleae 5 – 6 times longer than inner and middle rows, middle paleae not covering the inner opercular paleae (Fig. 25 A). Sixteen pairs of short opercular papillae (Figs 24 C, D, 25 B) peripheral to outer paleae. Three pairs of nuchal spines present, slightly curved inwards with blunt tips (Figs 24 F, 25 E). Tentacular filaments compound (branching) arranged in eight rows (Fig. 24 B, C). Palps slightly longer than length of operculum. Median organ present at dorsal junction of lobes of operculum, with eye spots on lateral margins. Segment 1 (chaetiger 1) with one pair of neuropodial cirri with small tuft of fine capillary chaetae on either side of U-shaped buccal organ (Fig. 24 B). Segment 2 (chaetiger 2) with one pair of elongated lateral lobes, connecting branchiae to neuropodia (Fig. 24 C, D). Neuropodia of segments 1 and 2 not vertically aligned, those of segment 2 situated more laterally, neurochaetae similar in structure on both segments. Five to six pairs of branchiae present (difficult to count as holotype slightly damaged). Branchiae narrow based, strongly ciliated, long and tapering to fine tip. Segment 3 – 5 (parathoracic) with two types of notochaetae arranged transversely (Fig. 25 H); six lanceolate chaetae, tapering to an elongate frayed tip (Fig. 25 H) and fine capillaries with frayed margins inserted between lanceolate ones (Fig. 25 H). Segments 3 – 5 with two types of neurochaetae arranged in compact lateral fascicle, with about six to eight lanceolate chaetae plus fine capillaries. Parathoracic notopodia more robust that neuropodia (Fig. 24 A). Abdominal region with 12 chaetigers. Notopodia as transverse tori, with long handled uncini, numbers per torus decreasing posteriorly, each uncinus with two vertical rows, each row with six to eight major teeth (Fig. 25 J), similar throughout abdomen. Neuropodia with ventral bundle of fine notochaetal capillaries with extended fine tips, shafts composed of stacks of thecae with margins extending to fine filamentous tip (Fig. 25 I), similar throughout abdomen. Cauda smooth and about half of length of abdomen on paratype. Variation. Material examined varies from 3 – 5 mm in length, without cauda, 1 – 2 in width with 13 – 18 chaetigers, 22 – 23 pairs of outer paleae, 10 – 11 pairs of inner paleae, 11 – 12 pairs of middle paleae and 16 – 23 pairs of opercular papillae.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F3312AFFB610D026D9FD10FDBA.taxon	discussion	Remarks. Sabellaria lungalla n. sp., is characterised by the distal margin of the outer paleae being serrated with a midline plume, one kind of middle paleae which are geniculate with smooth pointed tips and inner paleae which are asymmetrical with short blunt tips, three pairs of nuchal spines and one pair of lateral lobes on segment 2, which no other species share (Table 4). Sabellaria lungalla n. sp., can be separated from S. kooraltha n. sp., by the shape of the distal margin of the outer paleae: S. lungalla n. sp., has an elongate finely serrated distal plume (Fig. 25 C) whereas S. kooraltha n. sp., has a large central distal tooth which is smooth with two lateral teeth on either side (Fig. 23 D, E). The other Australian species S. pyramis n. sp., differs from S. lungalla n. sp., by the number and shape of the middle paleae (Fig. 26 E, F). Sabellaria lungalla n. sp., belongs to a group of Sabellaria with only one kind of middle paleae (Table 4), and of those occurring in the Indo-Pacific they can be separated from S. lungalla n. sp., on the basis on the dentition of the single kind of mid-paleae. Sabellaria gilchristi (McIntosh, 1925) has only two pairs of teeth on the distal margin of the outer paleae Sabellaria javanica Augener, 1934 described from Indonesia, has no teeth on the lateral margins and S. tottoriensis Nishi et al., 2004, described from Japan, has only three teeth whereas S. lungalla n. sp., has 7 – 8 pairs of teeth.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F3312AFFB610D026D9FD10FDBA.taxon	distribution	Distribution. Gulf of Carpentaria, Northern Australia, and Dampier Archipelago, Western Australia. Habitat. Intertidal in sheltered bays from intertidal to 15 m.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F3312AFFB610D026D9FD10FDBA.taxon	etymology	Etymology. The specific name lungalla refers to an Aboriginal word for a lagoon in the Northern Territory and this species is found in inshore shallow bays.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F3312FFFB410D024C1FB40FBD1.taxon	description	Fig. 6 A, 26, Table 4	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F3312FFFB410D024C1FB40FBD1.taxon	materials_examined	Material examined. Holotype: NTM W 10045, 6 mm long, 1 mm wide, posteriorly incomplete with 17 chaetigers, paratypes: 1 NTM W 023787, 10 mm long, 1 mm wide, complete with 28 chaetigers, plus cauda, operculum incomplete; 1, NTM W 10044, 8 mm long, 1 mm wide, incomplete. All type material from Northern Territory: Darwin Harbour, Holotype and NTM W 023787 from St. D 24 A, 12 ° 25.05 ' S 130 ° 48.58 ' E, 8 m, 7. i. 1993 and NTM W 10044, from St D 98 A, 12 ° 31.9 ' S 130 ° 50.43 ' E, 18 m, coll 14. vii. 1993.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F3312FFFB410D024C1FB40FBD1.taxon	description	Description. Holotype pale yellow with scattered brown pigmentation on dorsum of last parathoracic segments and at base of tentacular filaments. Operculum longer than wide, with completely separated lobes. Paleae arranged in three semicircular concentric rows; outer row with 18 pairs, clear and golden paleae with broad flattened smooth blade, distally serrated with fine extended midline plume (majority broken). Middle row with one kind of paleae, five pairs, stout, robust, shaft forming a four sided pyramid with blunt tips (Fig. 26 D). Inner row with nine, geniculate with symmetrical blunt tips directed inwards paleae. Middle paleae longest, outer paleae cover bases of middle, inner shortest. Opercular papillae 12 pairs, shorter than wide, cylindrical (Fig. 26 B). Nuchal spines not observed. Tentacular filaments compound, long, arranged in five transverse rows (Fig. 26 A). Median organ and median ridge present, with eye spots on both margins. Segment 1 (chaetiger 1) with one pair of elongate neuropodial cirri, with long capillaries (Fig. 26 G). Segment 2 (chaetiger 2) with one pair of lateral lobes connecting branchiae to neuropodia (Fig. 26 H). Neurochaetae similar on both segments. Six pairs of branchiae, elongate, strongly ridged, narrow based, tapering to fine tip, not meeting mid dorsally. Segments 3 – 5 (parathoracic) with two types of notochaetae arranged transversely, eight lanceolate chaetae with fine capillaries imbetween. Segments 3 – 5 with two types of neurochaetae, 3 – 4 lanceolate and capillaries imbetween. Abdomen of 23 segments with short neuropodia with capillary chaetae and expanded rectangular notopodia with terminal transverse torus with long handled uncini with 6 – 8 teeth arranged vertically, similar throughout abdomen. Cauda smooth and equal in length to last ten segments. Variations. The two paratypes vary in the number of pairs of middle paleae (4 – 5), inner paleae (9 – 10) and branchiae (5 – 6) present, and both are incomplete posteriorly and the opercular lobes are slightly damaged.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F3312FFFB410D024C1FB40FBD1.taxon	discussion	Remarks. Sabellaria pyramis n. sp., differs from other species of the genus in the particular pyramidal shape of the middle paleae, unique in the genus. All other described species of Sabellaria have geniculate middle paleae. Even though the genus Sabellaria was not recovered as mophyletic in Capa et al. (2012) or the present study (Fig. 28 C) we have positioned this species within this genus since it shares the combination of features of the rest of the species (operculum completely divided in two symmetrical lobes, inner paleae arranged in two rows, with mid and inner paleae, and three parathoracic segments) together with the shape of outer paleae provided with a distal plume and denticles on both sides.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F3312FFFB410D024C1FB40FBD1.taxon	distribution	Distribution. Darwin Harbour, Northern Territory, known only from type locality. Habitat. Muddy sediments in depths of 8 – 18 m.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F3312FFFB410D024C1FB40FBD1.taxon	etymology	Etymology. The specific name of pyramis refers to the pyramidal shape of the middle paleae.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F3312EFFB410D020D1FC05FA13.taxon	materials_examined	Type species. Hermella varians Treadwell, 1901, designation by Kirtley, 1994. Type locality: Mayagüez Harbour, Puerto Rico.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F3312EFFB410D020D1FC05FA13.taxon	diagnosis	Diagnosis. Characterised by the arrangement of the inner paleae in a short ventral line on each inner margin of the opercular lobes. Nuchal hooks are large and with broadened shafts, unique among sabellariids.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F3312EFFB410D020D1FC05FA13.taxon	discussion	Remarks. This genus currently contains ten species plus the one described in this paper, with most known from deep water (188 – 4825 m). Traditional diagnostic characters of these species, based almost entirely on paleal and nuchal hook characters (as in Kirtley, 1994) are listed in Table 5.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F3312EFFA910D0221CFBF9FD61.taxon	description	Figs 6 B, 27, Table 5	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F3312EFFA910D0221CFBF9FD61.taxon	materials_examined	Material examined. Holotype: NTM W 4994 *, FRV Soela, cruise 06 / 85 st. 5: 17. xi. 1985, S of Saumarez Reef, Marian Plateau, 22 ˚ 40.0 ' S, 154 ˚ 05.5 ' E to 22 ˚ 42.5 ' S, 154 ˚ 05.9 ' E, 416 – 419 m, collected with a lobster trawl. Specimen posteriorly incomplete with 15 chaetigers, 35 mm in length, 6 mm in width.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F3312EFFA910D0221CFBF9FD61.taxon	description	Description. Holotype robust, laterally flattened body, preserved animal pale yellow colourless except for dark brown nuchal hooks. Operculum with two partially fused lobes, as long as wide with deep indentation on ventral margin, laterally compressed (Fig. 27 A). Outer paleae, 40 pairs, arranged in semicircles, stout, golden, flattened, blades with three transverse thecal bands and asymmetrical pointed tips (Fig. 27 A). Inner row of paleae with three pairs of paleae arranged as a short ventral line on each inner margin of the opercular lobes, about one quarter length of outer paleae. One pair of flat and strongly curved nuchal spines without limbation (Fig. 25 C). Opercular papillae, 14 pairs, triangular, almost as long as outer paleae (Fig. 27 A, B). Dorsal papilla, digitiform, located between nuchal spines (Fig. 27 C). Palps, thick, rounded, with crenulated margins about one third longer than operculum. Median and median ridge present. Eye spots not observed, as difficult to separate lobes. Tentacular filaments, simple, nine pairs. Segment 1 (chaetiger 1) with neuropodial cirri on either side of buccal organ, elongate and tapering and a fascicle of capillaries. Segment 2 (chaetiger 2) with well fascicle of neuropodial capillaries, three elongate narrow lateral lobes present. Branchiae from segment 2, extending to chaetiger 13, last two chaetigers lacking branchiae, long thin flattened, meeting mid dorsally. Segments 3 – 6 (parathoracic) with two types of notochaetae inserted transversely, eight stout lanceolate with frayed tips and much shorter finer capillaries inserted in between. First parathoracic notopodia very small about one quarter size of subsequent ones which increase in size posteriorly. Notochaetae consist of two rows of five large lanceolate chaetae and short fine capillaries respectively. Segments 3 – 6 (parathoracic segments) with only one kind of neurochaetae, six capillaries. Abdomen incomplete. Abdominal neuropodia with compact fascicle of long capillaries arranged in two tiers with expanded fine tips, thecae expanded and margins appear denticulate. Notopodia erect expanded tori with transverse row of uncini with two rows of teeth, each row with 7 – 8 teeth. Cauda missing in holotype.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F3312EFFA910D0221CFBF9FD61.taxon	discussion	Remarks. Tetreres terribilis n. sp., is characterised by the presence of outer paleae with flattened blades symmetrical tips and transverse thecal bands a feature only shared with T. perryi (Kirtley, 1994), however it should be noted that the latter species is poorly described with only an illustration of the outer paleae and a comment that the nuchal looks are large with sharply recurvedd tips (Table 5). These two species are distinguished by the shape of the tip of these outer paleae with a pointed tip in the new species and a long spike in T. perryi, and the presence of an internal fusiform outline in T. terribilis n. sp., absent in T. perryi (Table 5). The only species described from nearby localities to Marian Plateau is T. robustus Lechapt & Kirtley, 1998 at depths of 440 – 450 m off New Caledonia. This species was described as having a larger number of outer paleae (70 – 80 pairs) but with a smooth surface, lacking thecal bands but they can be separated by the number of lateral lobes on segment 2, being three in T. terribilis n. sp. and four in T. robustus.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F3312EFFA910D0221CFBF9FD61.taxon	distribution	Distribution. Known only from type locality, Marion Plateau in depths of 416 – 419 m. Habitat. Substrate unknown, appears to be a solitary species.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
AD4387F3312EFFA910D0221CFBF9FD61.taxon	etymology	Etymology. The name of this species terribilis refers to the large size of the species and which is much larger than any other material examined and if complete would represent a very scary animal.	en	Hutchings, Pat, Capa, María, Peart, Rachael (2012): 3306. Zootaxa 3306: 1-60
