taxonID	type	description	language	source
A86D8544B073FFAB77ADFB124AA844A7.taxon	materials_examined	Material examined: Males (n = 112), females (n = 131), and juveniles (n = 74) collected at the type locality of Torreya State Park in the Florida panhandle (IZ- 133791 – 2, IZ- 133796 – 8, IZ- 134557 – 8), nearby localities east of the Apalachicola River (IZ- 133793 – 5, IZ- 134492 – 3), and at Florida Caverns State Park (IZ- 133808 – 13). Two specimens from collection IZ- 133812 were dissected for genitalia (SPM 007418 and 7420), the latter of which was disarticulated for appendage measurements and SEMs. A total of 124 specimens covering all localities were sequenced for molecular analysis. Diagnostic molecular characters: In our COI amino acid fragment (described above) no. 53 is isoleucine (not valine), no. 115 is methionine (rarely threonine; not leucine), and no. 248 is valine or leucine (not isoleucine). See Table 3. M. americanus M. savannahensis M. sassafrasensis sp. nov. sp. nov. male male male male male male female male male female holotype holotype paratype paratype paratype holotype paratype paratype M. americanus M. savannahensis M. sassafrasensis sp. nov. sp. nov. male male male male - holotype paratype paratype 7420 (from Davis, 7185 7176 range size 1933) (lengths) Chelicer I (whole) 0.6 / 2.9 0.7 / 3.3 0.7 / 3.3 0.1 I (from crest) 0.4 / 1.9 0.4 / 1.9 0.4 / 2.0 0 II 0.8 / 5.0 0.8 / 5.0 0.9 / 5.6 0.1 III 0.2 / 5.7 0.2 / 4.9 0.2 / 4.1 0 Total length 1.6 1.4 1.7 1.8 0.4 Palp Trochanter 0.3 / 3.6 0.2 / 2.9 0.3 / 3.4 0.1 II 0.4 / 5.0 0.4 / 5.0 0.4 / 5.7 0 III 0.2 / 3.4 0.2 / 3.0 0.3 / 3.2 0.1 IV 0.3 / 4.0 0.3 / 4.0 0.3 / 3.9 0 ...... continued on the next page	en	Clouse, Ronald M., Wheeler, Ward C. (2014): Descriptions of two new, cryptic species of Metasiro (Arachnida: Opiliones: Cyphophthalmi: Neogoveidae) from South Carolina, USA, including a discussion of mitochondrial mutation rates. Zootaxa 3814 (2): 177-201, DOI: 10.11646/zootaxa.3814.2.2
A86D8544B07EFFA977ADF9D24ECC40DF.taxon	materials_examined	Material examined. Males (n = 62), females (n = 72), and juveniles (n = 51), with the following collection details: Jasper County, South Carolina, USA, at Kingfisher Pond, Savannah National Wildlife Refuge, 300 m south of parking lot (lat. 32.18923, long. - 81.08008, elev. 3 m), leg. P. Sharma and R. Clouse, 16 March 2010. Holotype, male, specimen IZ- 133799 (105645) - 7184. Paratypes, from collections IZ- 133799 – 807, 6 males (SPM 007215, 7230, 7240, 7266, 7295, 7315) and 7 females (SPM 007191, 7222, 7234, 7252, 7275, 7301, 7322); two males dissected for genitalia (SPM 007185 and 7335), SPM 007185 disarticulated for appendage measurements and SEMs, and SPM 7214 and 7221 – 3 mounted for dorsal and ventral SEMs. A total of 85 specimens were sequenced for molecular analysis. All specimens are deposited in MCZ.	en	Clouse, Ronald M., Wheeler, Ward C. (2014): Descriptions of two new, cryptic species of Metasiro (Arachnida: Opiliones: Cyphophthalmi: Neogoveidae) from South Carolina, USA, including a discussion of mitochondrial mutation rates. Zootaxa 3814 (2): 177-201, DOI: 10.11646/zootaxa.3814.2.2
A86D8544B07EFFA977ADF9D24ECC40DF.taxon	description	Description. Morphology as in the original description of M. americanus (Davis, 1933), with the following recap of characters and additional details. Body and appendage article measurements and proportions available in Tables 5 – 7. Tuberculate granulations, irregular in shape and spacing, covering entire body (Figs. 3 D – F; 7 B) and legs with the following exceptions: on dorsal prosoma just anterior to ozophores and above anterior tip of coxa I; on irregular medial strip of anal plate of males (being glossy) (Figs. 6 B, C; 7 B); on inner distal femur and patella of leg II (being glossy); on inner trochanter of leg IV (being glossy), and on all tarsi (being generally smooth). Anal plate in males with smooth, raised, medial strip, tapering posteriorly to width of anal gland pore on tergite IX, not reaching anterior edge of anal plate (Figs. 6 B, C; 7 B). Tergite IX distinctly bilobed in males and females (Figs. 6 B, C; 8 B). Gonostome in males oval, with straight posterior edge and a toothlike projection on either side of anterior edge, edges otherwise smooth (Fig. 7 A); gonostome more rounded in females, without large toothlike lateral projections, anterior edge with fringe of small, sharp projections (Fig. 8 A). Setation pattern even over most of body and appendages; especially concentrated on tarsi (Fig. 4 B, E, H, K), these bearing different kinds of setae, including weakly defined solae on tarsus I (Fig. 4 B), and solenidia (thick, curved setae; Willemart & Giribet 2010) on dorsal surface of tarsi I and II (Fig. 4 B, E); hairs nearly absent from chelicerae, especially from second article. Microtrichial formula of spermatopositor: 3 – 4, 6 – 8, 4 + 4 + 4 (Figs. 10 C, D, H; 11 C, D; 12 A; 13 B); ventral microtrichia at nearly same level (middle slightly more distal) across middle of ventral surface (Fig. 10 C, D, H); apical microtrichia in some cases separated by deep medial cleft (Fig. 11 C). Molecular diagnostic characters. In our COI amino acid fragment (described above) no. 43 is isoleucine (not methionine), no. 80 is threonine (not serine), no. 135 is methionine (not isoleucine), no. 145 is asparagine (not serine or glycine), no. 163 is valine (not isoleucine), and no. 231 is glutamine (not histadine). See Table 3.	en	Clouse, Ronald M., Wheeler, Ward C. (2014): Descriptions of two new, cryptic species of Metasiro (Arachnida: Opiliones: Cyphophthalmi: Neogoveidae) from South Carolina, USA, including a discussion of mitochondrial mutation rates. Zootaxa 3814 (2): 177-201, DOI: 10.11646/zootaxa.3814.2.2
A86D8544B07EFFA977ADF9D24ECC40DF.taxon	distribution	Distribution and etymology. Known only from the type locality, the Savannah River delta in South Carolina, for which the species is named.	en	Clouse, Ronald M., Wheeler, Ward C. (2014): Descriptions of two new, cryptic species of Metasiro (Arachnida: Opiliones: Cyphophthalmi: Neogoveidae) from South Carolina, USA, including a discussion of mitochondrial mutation rates. Zootaxa 3814 (2): 177-201, DOI: 10.11646/zootaxa.3814.2.2
A86D8544B078FFAF77ADFB3F4A364556.taxon	materials_examined	Material examined. Males (n = 8), females (n = 7), and juveniles (n = 1), with the following collection details: Pickens County, South Carolina, USA, at Jocosse Gorge, Sassafras Mt., off F. van Clayton Highway (lat. 35.06228, long. - 82.79500, elev. 760 m), over ridge in old forest, around rotting log, leg. P. Sharma and R. Clouse, 15 March 2010. Holotype, male, specimen IZ- 134535 (105644 - 7171). Paratypes, from collections IZ- 134534 – 5, 7 males (SPM 007168 and 7172 – 7), 7 females (SPM 007169 – 70 and 7178 – 82); male specimens SPM 007176 and 7177 dissected for genitalia, and SPM 007176 disarticulated for appendage measurements. Twelve specimens were sequenced for molecular analysis. All specimens are deposited in MCZ.	en	Clouse, Ronald M., Wheeler, Ward C. (2014): Descriptions of two new, cryptic species of Metasiro (Arachnida: Opiliones: Cyphophthalmi: Neogoveidae) from South Carolina, USA, including a discussion of mitochondrial mutation rates. Zootaxa 3814 (2): 177-201, DOI: 10.11646/zootaxa.3814.2.2
A86D8544B078FFAF77ADFB3F4A364556.taxon	description	Description. Morphology as in the original description of M. americanus and of M. savannahensis sp. nov. (see above; leg characters of M. sassafrasensis sp. nov. illustrated in Fig. 5; chelicer, palp, claw, and solenidia of M. sassafrasensis sp. nov. illustrated in Fig. 9). Microtrichial formula of spermatopositor: 3, 6, 4 + 4 + 4 (Figs. 10 A, B, G; 11 A, B; 13 A). Body and appendage article measurements and proportions available in Tables 5 – 7. Molecular diagnostic characters. In our COI amino acid fragment (described above) no. 87 is valine (not leucine). See Table 3.	en	Clouse, Ronald M., Wheeler, Ward C. (2014): Descriptions of two new, cryptic species of Metasiro (Arachnida: Opiliones: Cyphophthalmi: Neogoveidae) from South Carolina, USA, including a discussion of mitochondrial mutation rates. Zootaxa 3814 (2): 177-201, DOI: 10.11646/zootaxa.3814.2.2
A86D8544B078FFAF77ADFB3F4A364556.taxon	distribution	Distribution and etymology. Known only from the type locality, Sassafras Mountain, South Carolina, for which the species is named. M. americanus M. savannahensis M. sassafrasensis sp. nov. sp. nov. male male male male holotype paratype paratype 7420 (from Davis, 1933) 7185 7176 range size (all) Chelicer Article I 0.7 0.8 0.8 0.8 0.1 (whole) Article III 0.3 0.3 0.3 0.2 0.1 Palp Patella 0.7 0.7 0.6 0.7 0.1 Tibia 0.8 0.9 0.8 0.8 0.1 Tarsus 0.8 0.7 0.8 0.8 0.1 Leg I Patella 0.6 0.6 0.5 0.5 0.1 Tibia 0.7 0.6 0.6 0.7 0.1 Metatarsus 0.6 0.6 0.5 0.5 0.1 Tarsus 0.7 0.7 0.7 0.7 0.0 Leg II Patella 0.6 0.5 0.5 0.5 0.1 Tibia 0.6 0.6 0.6 0.7 0.1 Metatarsus 0.5 0.5 0.5 0.5 0 Tarsus 0.8 0.6 0.7 0.7 0.2 Leg III Patella 0.6 0.6 0.7 0.5 0.2 Tibia 0.7 0.8 0.9 0.7 0.2 Metatarsus 0.7 0.8 0.7 0.6 0.2 Tarsus 0.9 0.8 1.0 0.8 0.2 Leg IV Patella 0.6 0.6 0.6 0.5 0.1 Tibia 0.7 0.7 0.7 0.7 0 Metatarsus 0.6 0.7 0.6 0.5 0.2 Tarsus 0.7 0.7 0.8 0.7 0.1	en	Clouse, Ronald M., Wheeler, Ward C. (2014): Descriptions of two new, cryptic species of Metasiro (Arachnida: Opiliones: Cyphophthalmi: Neogoveidae) from South Carolina, USA, including a discussion of mitochondrial mutation rates. Zootaxa 3814 (2): 177-201, DOI: 10.11646/zootaxa.3814.2.2
