taxonID	type	description	language	source
A95287C7FFDAB54CFE79D390FB5EB4BC.taxon	type_taxon	Type species: Aulacophilinus rennellensis, by original designation and monotypy.	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFDAB54CFE79D390FB5EB4BC.taxon	discussion	TAXONOMIC HISTORY. — Lomholdt (1980) established the genus Aulacophilinus, a member of Trypoxylini, for his new species A. rennellensis from the Solomon Islands. He pointed out that the new genus has gastral segment I as long as in the Neotropical genus Aulacophilus and shares with it the presence of only two submarginal cells and the absence of a carina at the top of the propodeal side, but differs from it by the flagellum filiform (rather than claviform), the occipital carina continuous (rather than interrupted ventrally), and the mesopleuron non-ridged (rather than horizontally ridged). There are also three additional differences between these genera: in Aulacophilus the mandible is acuminate apically (truncate in Aulacophilinus, see below for details), the metapleuron is parallel-sided for most of its length (in Aulacophilinus the metapleuron is markedly wider above the upper metapleural pit than below it), and tergum I has a pair of longitudinal, admedian carinae (no such carinae are present in Aulacophilinus). Antropov (1999) questioned the close relationships of these two genera, claimed that Aulacophilinus has not a single autapomorphy within Trypoxylini and that its gaster is just an extreme case of the situation in Pison, in which several species have the gaster elongate to a various degree, with the length exceeding the maximum width (P. difficile Turner, P. icarioides Turner, P. obliteratum F. Smith, P. pistillum Menke, and P. woji Menke; in P. icarioides segment I is bulging apically, in P. woji the basal portion is approaching the condition of Aulacophilus). He concluded that Aulacophilinus should be treated as a specialized member of Pison and as its junior synonym. Neither Lomholdt nor Antropov, however, paid attention to the specialized, unique mandible and the specialized clypeus of Aulacophilinus. Naumann (1990 b), on the other hand, recognized the importance of the broadly truncate mandible and revised the four Australian species with this character, calling them the caliginosum group of Pison. He did not recognize the close relationship of the group to Aulacophilinus rennellensis. Menke (2016), however, demonstrated that the caliginosum species group should be included into Aulacophilinus, resurrected Aulacophilinus from synonymy, provided its definition, described one new species (A. amblygnathus), provided a key to determination of the species known to him, and commented on the types of A. rennellensis and A. mandibulatus. DEFINITION OF GENUS. — Aulacophilinus is characterized by a mandible that is unique within Trypoxylini: both the outer and inner surfaces are punctate and setose throughout (except narrowly impunctate and asetose adjacent to the apical margin), the acetabular and condylar grooves are absent, and the inner portion is broadly expanded preapically, thus forming an apical truncation (the apical margin of the truncation being slightly concave, e. g., Fig. 2). Unlike Pison, the clypeus is punctate throughout, without a shiny, medioventral lamella (see below for exception). I agree with Menke (2016) that these striking features justify recognition of Aulacophilinus as an independent genus. An undescribed Pison from Australia, however, is somewhat intermediate between Aulacophilinus and Pison: the clypeus without a lamella and the broadened preapically inner mandibular portion are as in Aulacophilinus. Unlike Aulacophilinus, however, the inner mandibular surface is impunctate and glabrous (as in Pison), and the following are unique: two large, preapical teeth on the inner mandibular margin, a broad, shallow condylar groove, and a sharp conspicuous acetabular carina. Also, the setae of tergum I are erect (appressed in Aulacophilinus). The unique mandible of Aulacophilinus, clearly an outstanding autapomorphy, may be related to a particular way of nest building. Unfortunately, nothing is known about the nesting habits of the included species.	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFDAB54CFE79D390FB5EB4BC.taxon	description	DESCRIPTION. — In addition to the specialized clypeus and mandible, the species share the following characters: eye asetose, ommatidia about equal in size; inner eye margin convergent above; frons without middle supraantennal carina except carina present in large females of A. caliginosus; distance between antennal socket and orbit larger than socket diameter (about equal to socket diameter in female of A. rennellensis); clypeus transverse; flagellum cylindrical, not thickened toward apex; mandibular posterior margin straight, neither emarginate nor step-like; labrum not emarginate or minimally emarginate apically; occipital carina continuous ventrally, not joining hypostomal carina, mostly not expanded but expanded in A. weiri, in many A. caliginosus, and slightly so ventrally in A. amblygnathus; hypostomal carina not expanded; gena narrow in dorsal view (e. g., Fig. 31), somewhat thicker in A. solitarius; scutum without notauli; scutellum not foveate along anterior margin except slightly foveate in A. carinatus and A. solitarius; postspiracular carina present, about as long as midocellar diameter (about twice as long as midocellar diameter in A. solitarius); propleuron and forecoxal venter closely punctate; metapleuron not ridged longitudinally between dorsal and ventral metapleural pits except ridged in A. weiri and most A. caliginosus; metapleural flange narrowly lamelliform; propodeal dorsum with median longitudinal carina; forewing with two or three submarginal cells; first recurrent vein received by submarginal cell I or interstitial with first intersubmarginal vein; marginal cell acuminate; midcoxae separated, hindcoxae contiguous; punctures of tergum I well defined; sterna without graduli, punctate throughout; setae silvery, appressed to suberect on tergum I, at most slightly shorter than midocellar diameter; female tergum VI without pygidial plate; male flagellomeres cylindrical, not emarginate nor expanded, and without tyli except tyli present in A. rennellensis; posterior margin of male sternum VIII emarginate.	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFDAB54CFE79D390FB5EB4BC.taxon	distribution	GEOGRAPHIC DISTRIBUTION. — Four species of Aulacophilinus inhabit Australia (A. caliginosus, A. mandibulatus, A. pyrrhicus, and A. weiri), four other occur in New Guinea (A. amblygnathus, A. carinatus, A. solitarius, and A. tegularis), and A. rennellensis is found on the Rennell Island in the Solomon Archipelago. Of the four Australian species, A. caliginosus is known from all parts of the continent including Tasmania, A. mandibulatum occurs in the South Australia and Western Australia, and A. pyrrhicum and A. weiri are found in the Northern Territory and Queensland.	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFDAB54CFE79D390FB5EB4BC.taxon	description	CLADISTIC ANALYSIS The following is the list of characters used in the analysis below: 1. Clypeal lamella: 0. present, 1. absent. 2. Mandible surface: 0. partly impunctate, 1. punctate throughout on both inner and outer surface. 3. Mandible: acetabular and condylar groves: 1. grooves present, 2. grooves absent. 4. Mandible: apex: 0. acuminate, 1. broadly truncate. 5. Mandible: color: 0. black, 1. ferruginous 6. Width of labrum: 0. more than one midocellar diameter, 1. less than one midocellar diameter 7. Sculpture of frons, vertex, scutum, and mesopleuron: 0. punctate, 1. areolate. 8. Crenulate sulcus on gena: 0. absent, 1. present. 9. Tegula: 0. impunctate posteriorly, 1. punctate throughout. 10. Omalus: 0. absent, 1. present. 11. Number of submarginal cells: 0. three, 1. two. 12. Gaster: shape: 1. sessile, 2. pedunculate. 13. Gaster: color: 0. black, 1. ferruginous 14. Apical emargination of male sternum VIII: 0. shallow or moderately deep, 1. prominently deep. Pison atrum Spinola, the type species of Pison, was used as the outgroup, and the following data matrix was constructed: Pison atrum 00000 00000 0000 amblygnathus 11110 00000 0000 caliginosus 11110 10010 1001 carinatus 11110 00000 0000 mandibulatus 11111 10010 0000 pyrrhicus 11110 00100 0010 rennellensis 11110 10000 1100 solitarius 11110 00000 000? tegularis 11110 00010 000? weiri 11110 11111 1001 A cladistic analysis was kindly performed by Fernando Alvarez Padilla of the Universidad Nacional Autónoma de México through implicit enumeration, using T. N. T. program by Pablo Goloboff, Steve Farris, and Kevin Nixon (2003). One tree of 16 steps was found (Fig. 1), with the consistency index of 87 and the retention index of 80 (several nodes on the cladogram are without support because there are no characters that can be optimized unambiguously at those nodes).	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFDEB54EFFBFD536FD63B581.taxon	description	Figures 2 – 9.	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFDEB54EFFBFD536FD63B581.taxon	materials_examined	Aulacophilinus amblygnathus Menke, 2016: 335, ♀, ♂. Holotype: ♂, New Guinea: Morobe Province: Wau (AEI), examined.	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFDEB54EFFBFD536FD63B581.taxon	diagnosis	RECOGNITION. — Aulacophilinus amblygnathus is an all black endemic of New Guinea with three submarginal cells. It differs from all its congener in having the frontal punctures 2 – 3 diameters apart (rather than less than one to slightly more than one diameter apart), and also in having the frontal setae about as long as 1.5 – 2.0 midocellar diameters (while no longer than one midocellar diameter in the other species). The absence of silvery, setal fasciae on terga is a subsidiary recognition feature.	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFDEB54EFFBFD536FD63B581.taxon	description	DESCRIPTION. — Frons dull, shallowly punctate, punctures averaging 2 – 3 diameter apart (Figs. 3, 5). Free margin of clypeal lobe obtusely angulate to rounded (Figs. 2, 4). Occipital carina slightly expanded ventrally. Width of labrum equal to 1.1 × midocellar diameter. Anteromedian pronotal pit transversely elongate, about as long as midocellar diameter. Scutum not foveate along flange, with short longitudinal ridges adjacent to posterior margin; scutal punctures well defined, varying from about one diameter apart on average on disk to less than one diameter apart; interspaces aciculate. Tegula impunctate in posterior half. Mesopleural punctures well defined, less than one diameter apart except about one diameter apart near center. Propodeum in most specimens without longitudinal carina separating side from dorsum and posterior surface, but evanescent carina present in some specimens or replaced by vague linear series of short, transverse rugae; dorsum, side and posterior surface with well-defined punctures; dorsum with punctures slightly more than one diameter apart near midline, less than one diameter apart laterally, with well defined, oblique ridges basally; side with punctures less than one diameter apart except about one diameter apart posterodorsally, with fine ridges visible from certain angles; posterior surface with punctures less than one diameter apart near midline, about one diameter apart laterally, with transverse ridges in ventral half ridges near base Forewing with three submarginal cells. Posteroventral forefemoral surface with fine punctures several diameters apart. Hindcoxal dorsum with outer margin obtusely carinate. Outer surface of hindtibia with several fine spines. Punctures of tergum I averaging about two diameters apart anterior of apical depression. Setae silvery, on upper frons erect, sinuous, varying from about 1.5 × to 2.0 × midocellar diameters; on postocellar area erect, straight, shorter than midocellar diameter; on scutum erect, straight, about 0.5 × to 0.8 × midocellar width; on tergum I suberect, slightly shorter than midocellar width; on lower gena suberect, curved apically, slightly longer than midocellar diameter; not concealing integument on clypeus. Apical depressions of terga without silvery, setal fasciae. Body all black. ♀. – Upper interocular distance equal to 0.58 – 0.62 × lower interocular distance; ocellocular distance equal to 0.6 – 0.7 × hindocellar diameter, distance between hindocelli equal to 1.0 – 1.3 × hindocellar diameter; eye height equal to 0.94 – 1.00 × distance between eye notches. Dorsal length of flagellomere I 1.9 – 2.1 × apical width. Length 7.2 – 7. 3 mm; head width 2.1 mm. ♂. – Upper interocular distance equal to 0.7 × lower interocular distance; ocellocular distance equal to 0.7 × hindocellar diameter, distance between hindocelli equal to 1.4 × hindocellar diameter; eye height equal to 1.06 × distance between eye notches. Dorsal length of flagellomere I 1.8 × apical width. Apical margin of sternum VIII emarginate (Fig. 6). Genitalia (Figs. 7, 8): Length 7.0 mm; head width 2.0 – 2. 1 mm.	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFDEB54EFFBFD536FD63B581.taxon	distribution	GEOGRAPHIC DISTRIBUTION (Fig. 9). — Higher elevations of New Guinea.	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFDEB54EFFBFD536FD63B581.taxon	materials_examined	RECORDS. — INDONESIA: Western Papua: Paniai Lakes (as Wisselmeren): Enarotadi, elevation 1,850 m (1 ♀, BISH), Top Camp [of 1939 Dutch-American New Guinea Expedition = approximately 122 km SW Jaiapura] (1 ♂, RMNH). PAPUA NEW GUINEA: Madang Province: Pandambai 6 air km W Bundi at 5 º 38 ′ S 145 º 11 ′ E, elevation 2,330 m (2 ♀, CAS), Teptep at 5 ° 55 ′ S 146 ° 30 ′ E, elevation 1,900 m (1 ♂, CAS). Morobe Province: Wau [elevation 1,134 m] (1 ♀, 1 ♂, AEI, holotype and paratype of A. amblygnathus; 1 ♀, BISH).	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFDCB542FFBFD03BFD57B39A.taxon	description	Figures 10 – 17.	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFDCB542FFBFD03BFD57B39A.taxon	diagnosis	RECOGNITION. — Aulacophilinus caliginosus shares with rennellensis and weiri the presence of only two submarginal cells. Unlike rennellensis, its gastral segment I is sessile rather than conspicuously pedunculate; the entirely punctate tegula is a subsidiary recognition feature. Unlike weiri, the frons, scutum, and mesopleuron of caliginosus are punctate rather than coarsely areolate (although the mesopleuron may be rugose), the gena has no crenulate sulcus along the posterior eye margin, there is no omaulus (omaulus present in weiri), and the pronotal collar has no crenulate furrow (crenulate furrow present in weiri).	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFDCB542FFBFD03BFD57B39A.taxon	materials_examined	SPECIES STATUS. — The material I have examined shows significantly more variation than described by Naumann (1990 b). Most variants of different characters, however, do not correlate with each other, indicating that one species, and not a number of closely related species, is involved.	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFDCB542FFBFD03BFD57B39A.taxon	description	DESCRIPTION. — Frons with well defined punctures less than one diameter apart in most specimens (compressed against each other in some), but punctures ill defined, shallow, up to about one diameter apart in females from Taree area, New South Wales; interspaces varying from unsculptured, shiny, to conspicuously microsculptured, dull; middle supraantennal line mostly absent, present in larger females. Free margin of clypeal lobe obtusely triangular (Fig. 10). Occipital carina expanded in many specimens. Width of labrum equal to 0.5 × midocellar diameter. Anteromedian pronotal pit slightly transversely elongate, about as long as midocellar diameter. Pronotal collar with ill-defined transverse furrow next to apical margin (furrow not foveolate). Scutum foveate along flange (only inconspicuously so in smallest specimen), with or without short longitudinal ridges adjacent to posterior margin; scutal punctures less than one diameter apart in most specimens, but some punctures more than one diameter apart in some individuals; interspaces varying from unsculptured and shiny to markedly microsculptured, dull. Tegula punctate throughout. Mesopleuron mostly punctate, but rugose in some specimens from Western Australia; punctures conspicuous, less than one diameter apart, but not conspicuous and more than one diameter apart ventrally in females from Taree area. Metapleuron longitudinally ridged between dorsal and ventral metapleural pits in most specimens, but minutely, sparsely punctate in females from Blundells Creek and from Taree area. Propodeum in most specimens without longitudinal carina separating side from dorsum and posterior surface and extending from gastral socket area toward spiracle, but such carina present (rudimentary to well defined) in some specimens; dorsum obliquely ridged, punctate between ridges (punctures fine in most specimens, but coarse in some males), punctate only in one male from Pilbara Region, Western Australia; side ridged, punctate between ridges in larger specimens, punctate only in small ones; posterior surface densely, coarsely punctured, transversely ridged ventrally in some specimens. Forewing with only two submarginal cells (Fig. 11); in one female from 50 km NW Taree, minuscule additional submarginal cell appears in lower right corner of second cell of left wing, making the wing appear to have three submarginal cells (Figs. 12, 13); length of posterior margin of second cell 1.2 – 2.4 × its height, second cell triangular in some males from Pilbara Region. Hindcoxal dorsum with outer margin sharply carinate, but carina obsolete basally in many specimens. Outer surface of hindtibia with fine but well-defined spines. Punctures of tergum I averaging about one diameter apart. Setae nearly appressed on upper frons and scutum, appressed on tergum I, completely concealing integument on clypeus or nearly so; erect (curved apically) on lower gena, about as long as 0.5 × midocellar diameter. Apical depressions of terga with silvery, setal fasciae. Body all black or mandibular apex ferruginous. ♀. – Upper interocular distance equal to 0.68 – 0.70 × lower interocular distance; ocellocular distance equal to 0.1 – 0.5 × hindocellar diameter, distance between hindocelli equal to 1.0 – 1.2 × hindocellar diameter; eye height equal to 0.98 – 1.04 × distance between eye notches. Dorsal length of flagellomere I 1.8 – 2.0 × apical width. Length 5.9 – 6.5 mm; head width 1.8 – 1.9 mm. ♂. – Upper interocular distance equal to 0.77 – 0.86 × lower interocular distance; ocellocular distance equal to 0.5 – 0.6 × hindocellar diameter, distance between hindocelli equal to 1.2 × hindocellar diameter; eye height equal to 0.92 – 1.04 × distance between eye notches. Dorsal length of flagellomere I 1.4 – 1.9 × apical width. Sternum VIII deeply emarginate (Fig. 14). Genitalia: Figs. 15 and 16. Length 4.7 – 6.8 mm; head width 1.5 – 1.7 mm.	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFDCB542FFBFD03BFD57B39A.taxon	distribution	GEOGRAPHIC DISTRIBUTION (Fig. 17). — Whole Australia including Tasmania, also Norfolk Island.	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFDCB542FFBFD03BFD57B39A.taxon	materials_examined	RECORDS (N = Naumann, 1990 b). — AUSTRALIA: Australian Capital Territory: Black Mountain at 35 ° 16 ′ S 149 ° 06 ′ E (3 ♀, 2 ♂, ANIC; 1 ♂, BMNH; 1 ♀, 2 ♂, CAS; 1 ♂, UCD), Blundells Creek at 35 ° 22 ′ S 148 ° 50 ′ E (7 ♀, ANIC), Canberra (1 ♀, ANIC), Hume (1 ♀, ANIC). New South Wales: 6 km NE Bilpin near Kurrajong (2 ♀, AMS), 100 km SE Broken Hill at 32 ° 51 ′ S 141 ° 37 ′ E (1 ♀, ANIC), Colo Heights (N), Congo Point (1 ♀, BMNH), Cullerin (1 ♂, UCD), Gilgandra (1 ♀, 1 ♂, AMS), Jinki Creek in Blue Mountains (1 ♀, AMS), 0.5 km SE Lansdowne near Taree (1 ♀, AMS), Lindfield at 33 ° 46 ′ S 151 ° 11 ′ E (1 ♀, CAS), Lorien Wildlife Refuge 3 km N and ca 1 km NNW Lansdowne near Taree (2 ♀, AMS), Mount Tomah (1 ♀, 1 ♂, AMS), 15 km SE Nimmitabel (N), Springs Creek 68 km SW Wilcannia at 31 ° 44 ′ S 142 ° 41 ′ E (N), 50 km NW Taree at 31 ° 31 ′ S 152 ° 14 ′ E (2 ♀, 2 ♂, AMS), Whiskers 7 km WNW Hoskinstown at 35 ° 24 ′ S 149 ° 23 ′ E (1 ♂, ANIC), Woodford (N). Norfolk Island: Mount Bates (1 ♀, BISH), Rocky Point Reserve at 29 ° 03 ′ S 167 ° 55 ′ E (N), Selwyn Pine Road at 29 ° 01 ′ S 167 ° 57 ′ E (N). Northern Territory: Areyonga (N), Roe Creek 12 km WSW Alice Springs (N), Gregory National Park at 16 ° 03 ′ 01 ″ S 130 ° 04 ′ 07 ″ E (1 ♂, ANIC), Keep River National Park at 15 ° 44 ′ 17 ″ S 129 ° 06 ′ 55 ″ E (2 ♂, CAS), at 15 ° 45 ′ 42 ″ S 129 ° 06 ′ 45 ″ E (1 ♂, USU), at 15 ° 54 ′ 55 ″ S 129 ° 04 ′ 11 ″ E (1 ♂, USU), at 15 ° 55 ′ 22 ″ S 129 ° 03 ′ 25 ″ E, at 15 ° 57 ′ 55 ″ S 129 ° 01 ′ 52 ″ E (1 ♀, ANIC), and at 16 ° 03 ′ 01 ″ S 130 ° 24 ′ 07 ’’ E (1 ♂, ANIC), Tennant Creek (1 ♂, WAM), 131 km N Tennant Creek at 18 ° 28 ′ S 133 ° 52.1 ′ E (1 ♂, CAS), Victoria Highway 109 km WSW Timber Creek at 15 ° 56 ′ 11 ″ S 129 ° 35 ′ 22 ″ E (1 ♂, USU), Waterhouse Range 19 km SSW Alice Springs (N), Yuendumu (N). Queensland: 5 km S Batavia Downs at 12 ° 41 ′ S 142 ° 41 ′ E (3 ♀, ANIC), 3 km W Batavia Downs at 12 ° 40 ′ S 142 ° 39 ′ E (1 ♀, ANIC), Beerwah (1 ♀, QMB), Bundaberg (N), 48 km S Cunnamulla (N), Davies Creek National Park at 17 ° 00.6 ′ S 145 ° 34.9 ′ E (1 ♂, CAS), Hann River at 15 ° 11 ′ S 143 ° 52 ′ E (2 ♀, 1 ♂, ANIC), Heathlands at 11 ° 45 ′ S 142 ° 35 ′ S (1 ♀, ANIC), 2 km S Horseshoe Lookout in Blackdown Tableland (N), Isaacs River 100 km NE Clermont (1 ♂, QMB), Lawn Hill National Park at 18 ° 35 ′ 15 ″ S 138 ° 04 ′ 28 ″ E (1 ♂, QMB), 18 ° 38 ′ 13 ″ S 138 ° 12 ′ 29 ″ E (1 ♀, QMB), and 18 ° 40 ’ 15 ″ S 138 ° 22 ’ 15 ’’ (1 ♀, QMB), Moonie (1 ♀, QMB), Mount Cook National Park (N), Mount Nebo (N), 29 km NW Mundubbera (1 ♀, QMB), Musselbrook Camp at 18 ° 36 ′ S 138 ° 08 ′ E (2 ♂, ANIC), North Stradbroke Island (N), Split Rock 14 km SE Laura at 15 ° 39 ′ S 144 ° 31 ′ E (2 ♀, ANIC), 6 km N Taroom at 25 ° 36 ′ S 149 ° 46 ′ E (1 ♂, QMB), 13 km SE Weipa at 12 ° 40 ′ S 143 ° 00 ′ E (4 ♀, 1 ♂, ANIC). South Australia: Mount Davies area in Tomkinson Range (N), 79 km NNW Renmark at 33 ° 31 ′ S 140 ° 24 ′ E (2 ♀, ANIC; 3 ♀, CAS). Tasmania: 14 km S Bronte Park at 42 ° 15 ′ S 146 ° 29 ′ E (1 ♀, ANIC), 1 km SSE Gladstone (N), Great Pine Tier 13 km NNW Bronte Peak (1 ♂, BMNH), Mount Field National Park (N). Western Australia: 48 km NW Carnarvon at 24 ° 35.2 ′ S 113 ° 31.2 ′ E (1 ♀, CAS), Charnley River 2 km SW Rolly Hill (1 ♀, ANIC), Drysdale River at 15 ° 02 ′ S 125 ° 55 ′ E (N), Great Northern Highway at 22 ° 41 ′ 36 ″ S 118 ° 42 ′ 19 ″ E (2 ♀, 4 ♂, AMS), 23 ° 02.6 ′ S 118 ° 50.2 ′ E (3 ♀, CAS; 1 ♂, USU), and 23 ° 07.3 ′ S 119 ° 05.5 ′ E (1 ♂, ANIC; 1 ♂, USU), Juna Downs road to Packsaddle Bore at 22 ° 51 ’ 30 ″ S 118 ° 40 ’ 14 ″ E (4 ♀, 4 ♂, AMS) and 22 ° 52 ′ 31 ″ S 118 ° 31 ′ 49 ″ E (1 ♂, AMS), 14 km SE Kalumburu Mission at 14 ° 25 ′ S 124 ° 50 ′ E (1 ♀, ANIC), Karijini National Park at 22 ° 26.3 ′ S 118 ° 22.9 ′ E (2 ♀, 2 ♂, USU), at 22 ° 28.4 ′ S 118 ° 32.6 ′ E (1 ♂, ANIC), and at 22 ° 30.1 ′ S 118 ° 24.4 ′ E (1 ♂, USU), Kennedy Range National Park at 24 ° 38.7 ′ S 115 ° 10.7 ′ E (1 ♀, USU), 11 km E Marble Bar at 21 ° 09.0 ′ S 119 ° 51.7 ′ E (2 ♂, ANIC; 1 ♂, CAS), 25 km N Marble Bar at 20 ° 56.2 ′ S 118 ° 51.0 ′ E (1 ♂, USU), 30 km E Marble Bar at 21 ° 11.0 ′ S 120 ° 01.7 ′ E (1 ♀, ANIC; 2 ♂, CAS), 133 km SW Marble Bar at 21 ° 41.6 ′ S 119 ° 04.8 ′ E (4 ♀, USU), 4 km SW Mining Camp in Mitchell Plateau at 14 ° 52 ′ S 125 ° 50 ′ E (4 ♀, ANIC), Mount Augustus National Park at 24 ° 21.9 ′ S 116 ° 52.2 ′ E (1 ♂, CAS), Murdoch, southern suburb of Perth (1 ♀, WAM), 65 km E Nanutarra Road House at 22 ° 27.8 ′ S 116 ° 02.6 ′ E (1 ♂, USU), Nanutarra-Wittenoom road at 22 ° 21 ′ 21 ″ S 117 ° 54 ’ 16 ’’ (1 ♂, AMS), 22 ° 26 ′ 08 ″ S 117 ° 49 ′ 56 ″ E (2 ♀, 2 ♂, SAM), and 22 ° 26 ’ 36 ″ S 117 ° 48 ′ 23 ″ E (1 ♀, 2 ♂, AMS), 47 km S Pardoo Roadhouse on Shay Gap road at 20 ° 22.7 ′ S 120 ° 01.3 ′ E (2 ♀, 4 ♂, ANIC), Perth (1 ♂, BMNH), Perth: Darling Range (1 ♀, BMNH), Perth: Darlington (1 ♂, WAM), Thomas River 23 km NNW Mount Aridat 33 ° 55 ′ S 123 ° 00 ′ E (N).	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFD0B544FFBFD38DFCD2B27F.taxon	description	Figures 18 – 23.	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFD0B544FFBFD38DFCD2B27F.taxon	etymology	NAME DERIVATION. — Carinatus, Latin for carinate; with reference to the carinate propodeum of his species.	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFD0B544FFBFD38DFCD2B27F.taxon	diagnosis	RECOGNITION. — Aulacophilinus carinatus is an all black inhabitant of New Guinea with three submarginal cells. It differs from its three other New Guinean congeners with these characters, A. amblygnathus, solitarius, and tegularis, in having the propodeum with a longitudinal carina that separates the side from the dorsum and the posterior surface, and with s series of transverse ridges meeting the carina on its dorsal side (Fig. 19). In the three other species, the longitudinal carina is absent or evanescent, and the transverse ridges on the sides of the propodeal dorsum are lacking or are evanescent. Unlike A. amblygnathus, the punctures of the frons are no more than one diameter apart (rather than 2 – 3 diameters apart), and unlike A. tegularis the tegula is impunctate posteriorly (rather than uniformly punctate throughout).	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFD0B544FFBFD38DFCD2B27F.taxon	description	DESCRIPTION. — Frons dull, punctate, punctures no more than one diameter apart. Free margin of clypeal lobe obtusely angulate to rounded (Fig. 18). Width of labrum equal to 1.3 × midocellar diameter. Scutum not foveate along flange, with short, inconspicuous, longitudinal ridges adjacent to posterior margin; scutal punctures well defined, almost contiguous. Scutellum slightly foveate along anterior margin. Tegula impunctate (only aciculate) in posterior half. Mesopleural punctures well defined, less than one diameter in female, some punctures about one diameter apart in male Propodeum with longitudinal carina separating side from dorsum and posterior surface and extending from gastral socket area toward spiracle (Fig. 19); dorsum obliquely ridged, punctate between ridges (ridges becoming more conspicuous as they meet longitudinal carina); side punctate and somewhat irregularly ridged; posterior surface with well defined punctures, transversely ridged in ventral half. Forewing with three submarginal cells. Posteroventral forefemoral surface minutely punctate, punctures more than one diameter apart. Hindcoxal dorsum with outer margin obtusely carinate. Outer surface of hindtibia with fine but well defined spines. Punctures of tergum I fine in female, well defined in male, several diameters apart anterior of apical depression. Setae silvery, suberect on frons, about 0.5 × midocellar diameter long in female, up to one diameter in male; appressed on postocellar area, subappressed on scutum and about 0.3 × as long as midocellar diameter in female, as 0.5 × diameter in male; suberect on tergum I and up to about 0.5 × midocellar diameter long; on lower gena erect and up to one midocellar diameter long; not concealing integument on clypeus in female, concealing in male. Apical depressions of terga with silvery, setal fasciae (fasciae ill defined in female). Body all black. ♀. – Upper interocular distance equal to 0.62 × lower interocular distance; ocellocular distance equal to 0.3 × hindocellar diameter, distance between hindocelli equal to 1.0 × hindocellar diameter; eye height equal to 0.98 × distance between eye notches. Dorsal length of flagellomere I 2.1 × apical width. Length 6.2 mm; head width 1.8 mm. ♂. – Upper interocular distance equal to 0.60 × lower interocular distance; ocellocular distance equal to 0.5 × hindocellar diameter, distance between hindocelli equal to 0.8 × hindocellar diameter; eye height equal to 1.06 × distance between eye notches. Dorsal length of flagellomere I 2.4 × apical width. Sternum VIII: (Fig. 20). Genitalia: (Figs. 21, 22). Length 6.3 mm; head width 2.0 mm.	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFD0B544FFBFD38DFCD2B27F.taxon	distribution	GEOGRAPHIC DISTRIBUTION (Fig. 23). — Indonesian part of the island of New Guinea.	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFD0B544FFBFD38DFCD2B27F.taxon	materials_examined	RECORDS. — HOLOTYPE: ♀, INDONESIA: Western Papua: Waris S of Jayapura (as Hollandia), 1 – 2 Aug 1959, T. C. Maa (BISH). PARATYPE: INDONESIA: Western Papua: no specific locality, 10 Nov 1944, T. Aaron (1 ♂, CAS, labeled “ Neth. New Guinea ”).	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFD5B547FFBFD6E1FBFEB08C.taxon	description	Figures 24 – 29.	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFD5B547FFBFD6E1FBFEB08C.taxon	materials_examined	♀, ♂. Lectotype: ♀, Australia: Western Australia: Yallingup (BMNH), designated by Naumann, 1990 b: 242, examined.	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFD5B547FFBFD6E1FBFEB08C.taxon	diagnosis	RECOGNITION. — Aulacophilinus mandibulatus is the only species of the genus with the entire mandible ferruginous (the remaining body being black). Subsidiary recognition features are: forewing with three submarginal cells, tegula punctate throughout.	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFD5B547FFBFD6E1FBFEB08C.taxon	description	DESCRIPTION. — Frons dull, markedly microsculptured, punctate (punctures less than one diameter apart). Free margin of clypeal lobe obtusely angulate mesally (Fig. 24). Width of labrum equal to 0.7 × midocellar diameter. Anteromedian pronotal pit transversely elongate, smaller than midocellar diameter. Scutum not foveate along flange, without longitudinal ridges adjacent to posterior margin; scutal punctures well defined, interspaces somewhat microsculptured, linear in female, averaging less than one diameter apart in male. Tegula punctate throughout. Mesopleural punctures well defined, compressed against each other; interspaces merging into small ridges. Propodeum with well-defined, irregular carina separating side from dorsum and posterior surface and extending from gastral socket area toward spiracle; dorsum finely, obliquely ridged, punctate between ridges, with middle sulcus triangularly enlarging basally (Fig. 25); side punctate, interspaces merging into fine ridges; posterior surface in female conspicuously punctate (punctures compressed against each other), with well defined transverse ridges in male (punctate between ridges). Forewing with three submarginal cells. Posteroventral forefemoral surface microscopically, closely punctate. Hindcoxal dorsum with outer margin sharply carinate except anteriorly. Outer surface of hindtibia with fine but well-defined spines. Tarsomeres with plantulae. Punctures of tergum I on horizontal part averaging slightly more than one diameter apart mesally. Setae silvery, nearly appressed on upper frons, straight, suberect on lower gena (setal length about 0.6 × midocellar diameter), suberect but markedly shorter than midocellar diameter on scutum, appressed on tergum I, not concealing integument on clypeus. Apical depressions of terga without silvery, setal fasciae. Head, thorax, propodeum, legs, and gaster black, mandible ferruginous, female clypeus ferruginous next to lobe free margin. ♀. – Upper interocular distance equal to 0.72 × lower interocular distance; ocellocular distance equal to 0.9 × hindocellar diameter, distance between hindocelli equal to 1.3 × hindocellar diameter; eye height equal to 0.92 × distance between eye notches. Dorsal length of flagellomere I 2.1 × apical width. Mandible: trimmal carina with minimal preapical incision. Length 9.2 mm; head width 2.5 mm. ♂. – Upper interocular distance equal to 0.78 × lower interocular distance; ocellocular distance equal to 1.3 × hindocellar diameter, distance between hindocelli equal to 1.5 × hindocellar diameter; eye height equal to 0.96 × distance between eye notches. Dorsal length of flagellomere I 1.9 × apical width. Sternum VIII broadly emarginate apically (Fig. 26). Genitalia: Figs. 27, 28. Length 5.5 mm; head width 1.7 mm.	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFD5B547FFBFD6E1FBFEB08C.taxon	distribution	GEOGRAPHIC DISTRIBUTION (Fig. 29). — South Australia, Western Australia.	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFD5B547FFBFD6E1FBFEB08C.taxon	materials_examined	RECORDS. — AUSTRALIA: South Australia: 44 km NW Lock at 33 ° 31 ′ S 135 ° 16 ′ E (1 ♂, ANIC). Western Australia: Yallingup (1 ♀, BMNH, lectotype of Pison mandibulatum).	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFD4B55BFFBFD393FD51B443.taxon	description	Figures 30 – 37.	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFD4B55BFFBFD393FD51B443.taxon	materials_examined	♀, ♂. Holotype: ♀, Australia: Queensland: Kookaburra Cave in Carnarvon National Park (QMB), examined.	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFD4B55BFFBFD393FD51B443.taxon	diagnosis	RECOGNITION. — Aulacophilinus pyrrhicum is the only member of the genus with a ferruginous gaster and with the posterior propodeal surface with several ridges radiating up from the gastropropodeal articulation. It shares with A. weiri the gena with a crenulate sulcus along the posterior eye margin, the sulcus delimited posteriorly by a well-defined carina. The presence of three submarginal cells is a subsidiary recognition feature.	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFD4B55BFFBFD393FD51B443.taxon	description	DESCRIPTION. — Frons coarsely punctate or punctatorugose, interspaces shiny, linear. Free margin of clypeal lobe arcuate mesally, with minute apical point (Fig. 30). Occipital carina expanded ventrally. Width of labrum equal to 1.3 × midocellar diameter. Anteromedian pronotal pit transversely elongate, about three times as long as midocellar diameter. Scutum not foveate along flange, without longitudinal ridges adjacent to posterior margin; scutum and mesopleuron coarsely punctate or punctatorugose, interspaces linear, unsculptured (Fig. 32). Tegula posterolaterally impunctate or with microscopic, scattered punctures. Propodeum with or without longitudinal carina separating side from dorsum and posterior surface and extending from gastral socket area toward spiracle; dorsum obliquely ridged, rugose laterally; side coarsely punctate, interspaces merging into fine ridges; posterior surface irregularly transversely ridged, coarsely punctate between ridges, with several ridges radiating up from gastropropodeal articulation. Forewing with three submarginal cells. Posteroventral forefemoral surface minutely punctate, punctures averaging about two diameters apart. Hindcoxal dorsum with outer margin sharply carinate. Outer surface of hindtibia with fine but well-defined spines. Punctures of tergum I conspicuous, no more than one diameter apart on horizontal part in female and some males, slightly more in other males. Setae silvery, erect on frons and scutum, up to about as long as midocellar diameter; on lower gena subappressed to suberect, curved or sinuous, about as long as midocellar diameter; appressed on tergum I; not concealing integument on clypeus. Apical depressions of terga without setal fasciae. Apical depressions of terga without silvery, setal fasciae. Head, thorax, and propodeum black; flagellum ferruginous (apical flagellomere partly dark). Fore- and midfemora black, ferruginous apically, hindfemur ferruginous in apical third to all ferruginous; tibiae and tarsi ferruginous. Gaster ferruginous. ♀. – Upper interocular distance equal to 0.60 – 0.62 × lower interocular distance; ocellocular distance equal to 0.5 – 0.6 × hindocellar diameter, distance between hindocelli equal to 1.1 × hindocellar diameter (Fig. 31); eye height equal to 0.94 – 0.96 × distance between eye notches. Dorsal length of flagellomere I 2.2 × apical width. Length 6.5 – 7. 5 mm; head width 2.1 – 2. 3 mm. ♂. – Upper interocular distance equal to 0.68 – 0.76 × lower interocular distance; ocellocular distance equal to 0.7 – 0.8 × hindocellar diameter, distance between hindocelli equal to 1.1 × hindocellar diameter; eye height equal to 0.92 – 0.96 × distance between eye notches. Dorsal length of flagellomere I 2.0 – 2.3 × apical width. Sternum VIII shallowly, broadly emarginate apically (Fig. 33), with long, erect setae apically (Fig. 34. Genitalia: Figs. 35 and 36. Length 5.5 – 6.0 mm; head width 1.9 – 2.1 mm.	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFD4B55BFFBFD393FD51B443.taxon	distribution	GEOGRAPHIC DISTRIBUTION (Fig. 37). — Northern Territory, Queensland.	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFD4B55BFFBFD393FD51B443.taxon	materials_examined	RECORDS. — AUSTRALIA: Northern Territory: Kakadu National Park (1 ♂, CAS), Nourlangie Creek and Obiri Rock in Kakadu National Park (Naumann, 1990). Queensland: Arcadia on Magnetic Island (1 ♀, ANIC), Calamvale, a southern suburb of Brisbane (1 ♀, USNM), Fletcher Creek 43 km NW Charters Towers at 19 ° 48.9 ′ S 146 ° 03.3 ′ E (3 ♀, CAS), Kookaburra Cave in Carnarvon National Park (Naumann, 1990), Pendland at 20 ° 31.0 ′ S 145 ° 24.2 ′ E (3 ♀, 1 ♂, CAS), 2 km N Rokeby at 13 ° 39 ′ S 142 ° 40 ′ E (1 ♀, ANIC).	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFC9B55CFFBED2A5FDFBB722.taxon	description	Figures 38 – 46.	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFC9B55CFFBED2A5FDFBB722.taxon	materials_examined	♀, ♂. Holotype: ♀, Solomon Islands: Rennell Island: Hutuna (ZMUC), examined.	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFC9B55CFFBED2A5FDFBB722.taxon	diagnosis	RECOGNITION. — Aulacophilinus rennellensis differs from all its congeners in having a conspicuously pedunculate gastral segment I (its length is about 3.0 × maximum width in the female and 3.5 × in the male rather than about equal to width) and also conspicuously swollen apically (Figs. 41, 42). The presence of only two submarginal cells is a subsidiary recognition feature.	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFC9B55CFFBED2A5FDFBB722.taxon	description	DESCRIPTION. — Frons dull, microareolate, finely punctate, punctures averaging more than one diameter apart. Distance between antennal socket and orbit about equal to socket diameter in female, slightly greater in male. Free margin of clypeal lobe roundly arcuate (Figs. 38, 39). Occipital carina not expanded. Width of labrum equal to 0.6 × midocellar diameter. Anteromedian pronotal pit transversely elongate, about as long as midocellar diameter. Scutum foveate along flange, with short longitudinal ridges adjacent to posterior margin; scutal punctures minute, several diameters apart. Tegula impunctate in posterior half. Mesopleural punctures of medium size, several diameters apart. Propodeum without longitudinal carina separating side from dorsum and posterior surface; dorsum obliquely ridged; side punctate, finely, transversely ridged anteriorly in male; posterior surface punctate. Forewing with two submarginal cells, length of posterior margin of second cell equals 1.8 – 1.9 × its height. Posteroventral forefemoral surface minutely punctate, punctures several diameters apart. Hindcoxal dorsum with outer margin obtusely carinate. Outer surface of hindtibia without spines. Gastral segment I conspicuously pedunculate (its length about 3.0 × maximum width in female and 3.5 × in male), conspicuously swollen apically (Figs. 42). Apical swelling of tergum I with minute punctures several diameters apart. Setae silvery, subappressed on upper frons, on postocellar area appressed in female, erect in male, on scutum subappressed in female, suberect in male, appressed on tergum I, on lower gena suberect, straight, up to one midocellar diameter long, partly concealing integument on clypeus. Terga without setal, silvery, fasciae. Body all black, mandible with ferruginous tint. ♀. – Upper interocular distance equal to 0.4 × lower interocular distance; ocellocular distance equal to 0.2 × hindocellar diameter, distance between hindocelli equal to 0.5 × hindocellar diameter (Fig. 40); eye height equal to 1.6 × distance between eye notches. Dorsal length of flagellomere I 2.5 × apical width. Length 9.0 mm; head width 2.0 mm. ♂. – Upper interocular distance equal to 0.48 × lower interocular distance; ocellocular distance equal to 0.4 × hindocellar diameter, distance between hindocelli equal to 0.8 × hindocellar diameter; eye height equal to 1.12 × distance between eye notches. Dorsal length of flagellomere I 2.4 × apical width; flagellomeres I-IV with polished, elevated tyli. Sternum VIII and male genitalia missing from the specimens examined, but figured by Lomholdt (1980) and here reproduced as Figs. 43 – 45); sternum VIII emarginate apically (Fig. 43). Genitalia: (Figs. 44, 45). Length 8.5 mm; head width 1.7 mm.	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFC9B55CFFBED2A5FDFBB722.taxon	distribution	GEOGRAPHIC DISTRIBUTION (Fig. 46). — Rennell Island (Solomon Islands archipelago).	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFC9B55CFFBED2A5FDFBB722.taxon	materials_examined	RECORDS. — SOLOMON ISLANDS: Rennell Island: Hutuna (1 ♀, 1 ♂, ZMUC, holotype and paratype of Aulacophilinus rennellensis).	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFCEB55FFFBFD5BEFCB8B583.taxon	description	Figures 47 – 48.	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFCEB55FFFBFD5BEFCB8B583.taxon	etymology	NAME DERIVATION. — Solitarius is a Latin adjective meaning solitary, lonely; with reference to the fact that only one specimen of this species is known.	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFCEB55FFFBFD5BEFCB8B583.taxon	diagnosis	RECOGNITION. — Aulacophilinus solitarius is an all black endemic of New Guinea with three submarginal cells. It differs as follows from this island’s congeners: unlike A. amblygnathus, the frons punctures are no more than one diameter apart (rather than 2 – 3 diameters apart); unlike A. carinatum, it lacks the longitudinal carina separating the propodeal side from the dorsum and posterior surface and the ridges at the side of the propodeal dorsum are evanescent (carina present in carinatum, with well-defined ridges meeting its dorsal side); and unlike A. tegularis, the tegula is impunctate posteriorly (rather than punctate throughout).	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFCEB55FFFBFD5BEFCB8B583.taxon	description	DESCRIPTION. — Frons dull, markedly microsculptured, with well-defined punctures that average about one diameter apart. Free margin of clypeal lobe obtusely angulate (Fig. 47). Width of labrum equal to 1.5 × midocellar diameter. Gena somewhat thicker in dorsal view than in other Aulacophilinus. Anteromedian pronotal pit transversely elongate, about twice as long as midocellar diameter. Scutum foveate along flange, with minute, inconspicuous longitudinal ridges adjacent to posterior margin; scutal punctures well defined, less than one diameter apart. Tegula impunctate in posterior half. Mesopleural punctures well defined, less than one diameter apart except about one diameter apart ventrally. Postspiracular carina about twice as long as midocellar diameter. Propodeum without longitudinal carina separating side from dorsum and posterior surface; dorsum regularly, obliquely ridged (ridges evanescent laterally), with middle carina that is visible only from certain angles; side punctate, interspaces merging into fine, irregular ridges; posterior surface punctate, transversely ridged in ventral half. Forewing with three submarginal cells. Posteroventral forefemoral surface minutely punctate, punctures averaging about 2 – 3 diameters apart. Hindcoxal dorsum with outer margin obtusely carinate. Outer surface of hindtibia with fine spines. Punctures of tergum I fine, more than one diameter apart (except on apical depression). Setae silvery, suberect on frons and about as long as midocellar diameter, appressed on postocellar area, suberect on scutum and tergum I and up to about 0.5 × as long as midocellar diameter; on lower gena curved, about as long as midocellar diameter; not concealing integument on clypeus. Apical depressions of terga with silvery, setal fasciae. Body all black. ♀. – Upper interocular distance equal to 0.56 × lower interocular distance; ocellocular distance equal to 0.8 × hindocellar diameter, distance between hindocelli equal to 1.2 × hindocellar diameter; eye height equal to 0.96 × distance between eye notches. Both flagella missing. Length 9.0 mm; head width 2.5 mm. ♂. – Unknown.	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFCEB55FFFBFD5BEFCB8B583.taxon	distribution	GEOGRAPHIC DISTRIBUTION (Fig. 48). — Known from one locality in the Indonesian part of New Guinea.	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFCEB55FFFBFD5BEFCB8B583.taxon	materials_examined	RECORDS. — HOLOTYPE: ♀, INDONESIA: Western Papua: Paniai Lakes (as Wisselmeren): Enarotadi, elevation 1,800 – 1,900 m, 22 Aug 1962, J. Sedlacek (BISH).	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFCDB551FFBFD3E4FF31B588.taxon	description	Figures 49 – 51.	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFCDB551FFBFD3E4FF31B588.taxon	etymology	NAME DERIVATION. — Tegularis is an adjective derived from tegula, Latin for tile; with reference to the uniformly punctate tegula of this species.	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFCDB551FFBFD3E4FF31B588.taxon	diagnosis	RECOGNITION. — Aulacophilinus tegularis is an all black endemic of New Guinea with three submarginal cells. It differs from the other three New Guinean species with these characters, A. amblygnathus, A. solitarius, and A. carinatus, in having the tegula uniformly punctate throughout (Fig. 50) rather than impunctate or with evanescent, microscopic punctures posteriorly. Also, unlike A. amblygnathus, it has the frontal and scutal punctures less than one diameter apart (rather than more than one diameter apart), and the frontal setae about as long as 0.5 × midocellar diameter (rather than about one diameter long), It differs from A. carinatum in lacking the longitudinal carina separating the propodeal side from the dorsum and posterior surface and in lacking ridges on the side of the propodeal dorsum (longitudinal carina and transverse carinae present in A. carinatum).	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFCDB551FFBFD3E4FF31B588.taxon	description	DESCRIPTION. — Frons dull, markedly microsculptured, with well-defined punctures that are less than one diameter apart. Free margin of clypeal lobe obtusely angulate (Fig. 49). Width of labrum equal to 1.4 × midocellar diameter. Anteromedian pronotal pit transversely elongate, almost as long as midocellar diameter. Scutum minutely foveae along flange, with rudimentary longitudinal ridges adjacent to posterior margin; scutal punctures well defined, less than one diameter apart. Tegula uniformly punctate throughout (Fig. 50). Mesopleural punctures well defined, less than one diameter apart, interspaces merging posteriorly into small ridges. Propodeum without longitudinal carina separating side from dorsum and posterior surface; dorsum obliquely ridged, punctate between ridges; side irregularly ridged, punctate between ridges; posterior surface punctate, also transversely ridged in ventral half. Forewing with three submarginal cells. Posteroventral forefemoral surface microscopically, closely punctate. Hindcoxal dorsum with outer margin sharply carinate. Outer surface of hindtibia with fine but well-defined spines. Tarsomeres with plantulae. Punctures of tergum I, on horizontal part, averaging slightly more than one diameter apart. Setae silvery, suberect on frons and about 0.5 × as long as midocellar diameter, appressed on postocellar area, on scutum and tergum I suberect and about 0.3 × as long as midocellar diameter; on lower gena partly straight, partly curved, up to one midocellar diameter long; not concealing integument on clypeus. Apical depressions of terga with silvery, setal fasciae. Body all black, mandible narrowly ferrugineus apicoventrally in paratype. ♀. – Upper interocular distance equal to 0.56 – 0.58 × lower interocular distance; ocellocular distance equal to 0.5 – 0.7 × hindocellar diameter, distance between hindocelli equal to 0.5 – 0.8 × hindocellar diameter; eye height equal to 1.06 – 12.10 × distance between eye notches. Dorsal length of flagellomere I 2.3 – 2.5 × apical width. Length 9.4 – 9. 8 mm; head width 2.5 mm. ♂. – Unknown.	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFCDB551FFBFD3E4FF31B588.taxon	distribution	GEOGRAPHIC DISTRIBUTION (Fig. 51). — Known from two localities in the Madang Province of Papua New Guinea.	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFCDB551FFBFD3E4FF31B588.taxon	materials_examined	RECORDS. — HOLOTYPE: ♀, PAPUA NEW GUINEA: Madang Province: Bundi at 5 º 45 ′ S 145 º 15 ′ E, 20 May 1988, W. J. Pulawski (CAS). PARATYPE: PAPUA NEW GUINEA: Madang Province: Sapi Forest Reserve 30 km W Madang at 5 º 12 ′ S 145 º 30 ′ E, 10 Feb 1987, W. J. Pulawski (1 ♀, CAS).	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFC3B552FFBFD5FEFD68B5D0.taxon	description	Figures 52 – 62.	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFC3B552FFBFD5FEFD68B5D0.taxon	materials_examined	♀, ♂. Holotype: ♀, Australia: Northern Territory: Island of Rimbija (ANIC), examined.	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFC3B552FFBFD5FEFD68B5D0.taxon	diagnosis	RECOGNITION. — Aulacophilinus weiri shares with A. caliginosus and A. rennellensis the presence of only two submarginal cells. It is unique among its congeners in having the frons (Fig. 54), scutum (Fig. 58), and mesopleuron coarsely areolate (rather than punctate), the pronotal collar with a crenulate furrow posteriorly (no such furrow elsewhere or furrow ill defined), and in having an omalus (no omalus in the other Aulacophilinus). It shares with A. pyrrhicum the gena with a crenulate sulcus along the posterior eye margin, the sulcus delimited posteriorly by a well-defined carina (Fig. 56). Unlike A. rennellensis, its gastral segment I is sessile rather than conspicuously pedunculate. Unlike these two species, the tegula of A. weiri is minutely punctate throughout (rather than impunctate posteriorly).	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFC3B552FFBFD5FEFD68B5D0.taxon	description	DESCRIPTION. — Frons shiny, coarsely areolate (Fig. 54). Free margin of clypeal lobe widely rounded (Figs. 52, 53). Occipital carina expanded (Fig. 57), as high ventrally as midocellar diameter (less than that in some specimens). Width of labrum equal to 0.9 × midocellar diameter. Gena with crenulate sulcus along posterior eye margin (Fig. 56), sulcus delimited posteriorly by well-defined carina. Anteromedian pronotal pit rounded elongate, smaller than midocellar diameter. Scutum foveate along flange, with short longitudinal ridges adjacent to posterior margin, conspicuously areolate. Tegula microscopically punctate throughout. Mesopleuron conspicuously areolate, with omaulus. Metapleuron longitudinally ridged between dorsal and ventral metapleural pits. Propodeum without longitudinal carina separating side from dorsum and posterior surface and extending from gastral socket area toward spiracle; dorsum rugose, with short, transverse carinae emerging from middle carina; side with well-defined punctures (interspaces in some specimens merging into fine longitudinal ridges); posterior surface areolate, with tendency to form transverse ridges in ventral half. Forewing with two submarginal cells; length of posterior margin of second cell equals 1.3 – 1.4 × its height. Posteroventral forefemoral surface minutely, closely punctate. Hindcoxal dorsum with outer margin sharply carinate. Outer surface of hindtibia with evanescent spines. Punctures of tergum I well defined, about one diameter apart on horizontal portion. Setae silvery, appressed on frons, scutum, and tergum I; almost completely concealing integument on clypeus; genal setae suberect, straight, curved apically, longest setae about equal to midocellar diameter. Apical depressions of terga with ill-defined, silvery, setal fasciae. Body all black except mandibular apex ferruginous. ♀. – Upper interocular distance equal to 0.70 – 0.72 × lower interocular distance; ocellocular distance equal to 0.3 × hindocellar diameter, distance between hindocelli equal to 1.1 – 1.3 × hindocellar diameter (Fig. 55); eye height equal to 0.90 – 0.92 × distance between eye notches. Dorsal length of flagellomere I 1.4 – 1.8 × apical width. Length 4.8 – 6.0 mm; head width 1.4 – 1.6 mm. ♂. – Upper interocular distance equal to 0.80 × lower interocular distance; ocellocular distance equal to 0.4 × hindocellar diameter, distance between hindocelli equal to 1.2 × hindocellar diameter; eye height equal to 0.92 × distance between eye notches. Dorsal length of flagellomere I 1.3 × apical width. Sternum VIII conspicuously emarginate apically (Fig. 59). Genitalia: Figs. 60 and 61. Length 4.2 mm; head width 1.3 – 1.4 mm.	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFC3B552FFBFD5FEFD68B5D0.taxon	distribution	GEOGRAPHIC DISTRIBUTION (Fig. 62). — Northern part of Northern Territory, northern Queensland.	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
A95287C7FFC3B552FFBFD5FEFD68B5D0.taxon	materials_examined	RECORDS. — AUSTRALIA: Northern Territory: Cattle Creek 54 km S Borroloola (Naumann, 1990 b), Gregory National Park at 15 ° 58.3 ′ S 130 ° 29.3 ′ E (1 ♀, 1 ♂, ANIC), at 15 ° 58 ′ 17 ″ S 130 ° 29 ′ 17 ″ E (1 ♂, ANIC), at 16 ° 03.7 ′ S 130 ° 27.1 ′ E (1 ♂, USU), and 16 ° 12 ′ 47 ″ S 130 ° 25 ′ 11 ″ E (1 ♀, CAS; 1 ♂, USU), Gregory National Park near Timber Creek on Victoria River bank at 15 ° 37.8 ′ S 130 ° 28.6 ′ E (1 ♀, CAS), Island of Rimbija (1 ♀, 2 ♂, ANIC, holotype and paratypes of weiri), Keep River National Park at 15 ° 45 ′ 30 ″ S 129 ° 06 ′ 28 ″ E (1 ♂, CAS). Queensland: 4 km NE Batavia at 12 ° 39 ′ S 142 ° 42 ′ E (2 ♀, ANIC; 1 ♀, CAS), 4 km SW Casuarina Hill (Naumann, 1990 b), 13 km SE Weipa at 12 ° 40 ′ S 143 ° 00 ′ E (2 ♀, ANIC).	en	Pulawski, Wojciech J. (2017): A Revision of the Wasp Genus Aulacophilinus Lomholdt, 1980 with Descriptions of Three New Species (Hymenoptera: Crabronidae). Proceedings of the California Academy of Sciences 64 (1): 1-29, DOI: 10.5281/zenodo.11066844
