taxonID	type	description	language	source
AA067B1DFFF7FF97FF67FA8CAB8A3DEB.taxon	diagnosis	Diagnosis. Macrophyes species resemble those of Mesilla by having elongated and thin body; long and thin legs; median apophysis on anterior side of the tegulum and a large retromarginal tooth on distal side of chelicerae, but can been distinguished by the laminar ventral tegular projection; a short embolus basally inserted on the tegulum (Brescovit 1992: 105, figs 1 − 3, 6 − 11; 1993: 116, figs 1, 2; Figs 1 C−E). Females are characterized by their thin copulatory ducts, anteriorly curved and with small seminal receptacles on medial region; laminar and convergent lateral borders, forming a median, longer than wide atrium, and copulatory openings in anterior position (Brescovit 1992: 105, figs 4, 5; 1993: 116, figs 3 − 6; Figs 1 F, G).	en	Martínez, Leonel, Brescovit, Antonio Domingos, Oliveira, Luiz Fernando M. (2020): Two new species of the ghost spider genus Macrophyes O. Pickard-Cambridge 1893 from Colombia and description of the first male of Mesilla Simon, 1903 (Araneae: Anyphaenidae: Anyphaeninae). Zootaxa 4853 (4): 581-590, DOI: 10.11646/zootaxa.4853.4.7
AA067B1DFFF7FF95FF67F891AA293EA0.taxon	description	Figures 1 A−G, 4.	en	Martínez, Leonel, Brescovit, Antonio Domingos, Oliveira, Luiz Fernando M. (2020): Two new species of the ghost spider genus Macrophyes O. Pickard-Cambridge 1893 from Colombia and description of the first male of Mesilla Simon, 1903 (Araneae: Anyphaenidae: Anyphaeninae). Zootaxa 4853 (4): 581-590, DOI: 10.11646/zootaxa.4853.4.7
AA067B1DFFF7FF95FF67F891AA293EA0.taxon	materials_examined	Type material. Holotype: male from Jardin Botánico del Darién (8 ° 37 ′ 53.95 ″ N; 77 ° 21 ′ 23.43 ″ W), 90 m, Acandí, Capurgana, Chocó, Colombia, 28. III. 2009, M. Vargas & C. Zornasa leg. (MPUJ _ ENT 0061936). Paratypes: one male and female same data as holotype (MPUJ _ ENT 0061923 [m # 1]; MPUJ _ ENT 0061910 [f # 1]); one male and female same locality data, 250 m, 7. X. 2009 (MPUJ _ ENT 0061884 [m # 1]; MPUJ _ ENT 0061897 [f # 1]). One male same locality data, 3. XI. 2008, J. Aristizabal leg. (IBSP 244556).	en	Martínez, Leonel, Brescovit, Antonio Domingos, Oliveira, Luiz Fernando M. (2020): Two new species of the ghost spider genus Macrophyes O. Pickard-Cambridge 1893 from Colombia and description of the first male of Mesilla Simon, 1903 (Araneae: Anyphaenidae: Anyphaeninae). Zootaxa 4853 (4): 581-590, DOI: 10.11646/zootaxa.4853.4.7
AA067B1DFFF7FF95FF67F891AA293EA0.taxon	etymology	Etymology. The specific epithet is a patronymic given in honor of the Spanish singer Alejandro Sanz, who has greatly contributed to Spanish music.	en	Martínez, Leonel, Brescovit, Antonio Domingos, Oliveira, Luiz Fernando M. (2020): Two new species of the ghost spider genus Macrophyes O. Pickard-Cambridge 1893 from Colombia and description of the first male of Mesilla Simon, 1903 (Araneae: Anyphaenidae: Anyphaeninae). Zootaxa 4853 (4): 581-590, DOI: 10.11646/zootaxa.4853.4.7
AA067B1DFFF7FF95FF67F891AA293EA0.taxon	diagnosis	Diagnosis. Males of Macrophyes sanzi n. sp., resemble those of M. elongata Chickering, 1937 by the conical ventral tegular process and a retrolateral tibial apophysis with a small posterior projection (see Brescovit 1992, figs 6, 7), but can be distinguished by the quadrangular, straight retrolateral tibial apophysis and wider ventral tegular process (Figs 1 C−E). Females are similar to those of M. elongata by having an oval atrium, differing mainly by their rounded and large spermathecae. Also, resembles to M. pacoti Brescovit et al., 2019 by the large and rounded spermathecae, separating by the shape of the atrium (Figs 1 F, G).	en	Martínez, Leonel, Brescovit, Antonio Domingos, Oliveira, Luiz Fernando M. (2020): Two new species of the ghost spider genus Macrophyes O. Pickard-Cambridge 1893 from Colombia and description of the first male of Mesilla Simon, 1903 (Araneae: Anyphaenidae: Anyphaeninae). Zootaxa 4853 (4): 581-590, DOI: 10.11646/zootaxa.4853.4.7
AA067B1DFFF7FF95FF67F891AA293EA0.taxon	description	Description. Male. (Holotype, MPUJ _ ENT 0061936). Carapace yellowish dark in the cephalic region. Chelicerae brown, labium and endites brown. Sternum yellowish-brown. Legs yellow. Abdomen dorsally pale yellow, with two yellow lateral stripes; ventrally pale yellow. Spinnerets pale yellow (Fig. 1 A). Total length 7.12, carapace length 2.49, width 1.99, high 0.68. Clypeus height 0.15. Eye diameters and interdistances: AME 0.10, ALE 0.18, PME 0.2, PLE 0.21; AME – AME 0.23, AME – ALE 0.35, PME – PME 0.48, PME – PLE 0.38, ALE – PLE 0.43. Chelicerae 1.72 long, distal large retromarginal teeth, six promarginal teeth, six retromarginal denticle Leg I — femur 6.13 / patella 0.86 / tibia 7.08 / metatarsus 6.82 / tarsus 1.87 / total 22.76; II — 5.77 / 0.81 / 6.68 / 6.59 / 1.85 / 21.7; III — 3.3 / 0.81 / 2.63 / 3.54 / 1.17 / 11.45; IV — 4.42 / 0.73 / 4.77 / 6.31 / 1.55 / 17.78. Leg spination: III — tibia v 2 - 0 - 2 - 0, p 0 - 1 - 0 - 1, r 1 - 0 - 0 - 1, metatarsus p 1 - 1 - 0 - 0, r = I; IV — tibia v 2 - 0 - 2 - 2 - 0, p 0 - 1 - 0 - 1 - 0, r 1 - 0 - 0 - 0 - 1, metatarsus v 0 - 1 - 0 - 0 - 1, p 1 - 0 - 1 - 1 - 0, r 1 - 0 - 1 - 0 - 0. Abdomen, twice the length of carapace, length 4.61, epigastric furrow 0.68 from tracheal spiracle, spiracle 2.93 from base of spinnerets. Palp: retrolateral tibial apophysis small and apically straight, with a small projection on posterior side; cymbium shorter than tibia; subtegulum longer than wide; tegulum longer than wide, with ventral tegular projection wide, laminar; median apophysis long, curved, apically inserted on the tegulum; embolus short, apically filiform, with narrow base and proximally inserted on the tegulum (Figs 1 C – E). Female (Paratype, MPUJ _ ENT 0061923). Carapace yellowish. Chelicerae, labium and endites yellow. Sternum yellowish-brown. Legs pale yellow. Abdomen and spinnerets pale yellow (Fig. 1 B). Total length 7.02, carapace length 2.06, width 1.90, high 0.77. Clypeus height 0.09. Eye diameters and interdistances: AME 0.07, ALE 0.18, PME 0.15, PLE 0.17; AME – AME 0.22, AME – ALE 0.30, PME – PME 0.44, PME – PLE 0.42, ALE – PLE 0.37. Chelicerae 1.41 long, six promarginal teeth, six retromarginal teeth, sixth largest. Leg measurements: leg I — femur 6.84 / patella 1.12 / tibia 7.86 / metatarsus 6.98 / tarsus 2.01 / total 24.81; II — 5.88 / 1.02 / 5.56 / 5.94 / 1.81 / 20.21; III — 3.99 / 0.91 / 3.49 / 4.38 / 1.22 / 13.99; IV — 5.86 / 1.06 / 5.8 / 7.07 / 1.81 / 21.6. Leg spination: III — tibia v 2 - 2 - 2 - 2, p 1 - 1 - 0 - 0 - 1, r 1 - 1 - 0 - 0 - 1, metatarsus v 2 - 1 - 1 - 2; IV — tibia v 1 - 2 - 1 - 1 - 1, p 0 - 1 - 1 - 0 - 0, r 1 - 1 - 0 - 0 - 1, metatarsus v 2 - 0 - 1 - 1 - 0, p 1 - 0 - 1 - 0 - 1, r 1 - 0 - 1 - 0 - 0 - 1 - 1. Abdomen: length 4.86, epigastric furrow 0.83 from tracheal spiracle, spiracle 3.07 from base of spinnerets. Epigynum: atrium longer than wide, oval-shaped lateral borders wide, sclerotized and laminar; internally with long, thin, curved copulatory ducts; seminal receptacles small, rounded, medially situated on the copulatory ducts; spermathecae large, rounded and posteriorly positioned; fertilization ducts as long as the spermathecae width (Figs 1 F, G). Variation. Males (n = 3): total length: 7.03 – 8.20; carapace length: 2.05 – 2.71; femur I length: 6.13 – 6.66. Females (n = 6): total length: 1.02 – 1.17; carapace length: 0.41 – 0.51. Habitat. The specimens were caught manually and beating foliage of shrubs and low vegetation in a fragment of rain forest in Chocó department, Colombia.	en	Martínez, Leonel, Brescovit, Antonio Domingos, Oliveira, Luiz Fernando M. (2020): Two new species of the ghost spider genus Macrophyes O. Pickard-Cambridge 1893 from Colombia and description of the first male of Mesilla Simon, 1903 (Araneae: Anyphaenidae: Anyphaeninae). Zootaxa 4853 (4): 581-590, DOI: 10.11646/zootaxa.4853.4.7
AA067B1DFFF7FF95FF67F891AA293EA0.taxon	distribution	Distribution. Only known from the type locality (Fig. 4).	en	Martínez, Leonel, Brescovit, Antonio Domingos, Oliveira, Luiz Fernando M. (2020): Two new species of the ghost spider genus Macrophyes O. Pickard-Cambridge 1893 from Colombia and description of the first male of Mesilla Simon, 1903 (Araneae: Anyphaenidae: Anyphaeninae). Zootaxa 4853 (4): 581-590, DOI: 10.11646/zootaxa.4853.4.7
AA067B1DFFF5FF93FF67FA58AB213A20.taxon	description	Figure 2 A−C, 4.	en	Martínez, Leonel, Brescovit, Antonio Domingos, Oliveira, Luiz Fernando M. (2020): Two new species of the ghost spider genus Macrophyes O. Pickard-Cambridge 1893 from Colombia and description of the first male of Mesilla Simon, 1903 (Araneae: Anyphaenidae: Anyphaeninae). Zootaxa 4853 (4): 581-590, DOI: 10.11646/zootaxa.4853.4.7
AA067B1DFFF5FF93FF67FA58AB213A20.taxon	materials_examined	Type material. Holotype: female from Finca El Llanito (6 ° 50 ′ 29.69 ″ N; 73 ° 23 ′ 13.49 ″ W), 1313 m, Parque Nacional Natural Serranía de los Yariguies San Vicente de Chucurí Vereda Centro, Santander, Colombia, 13 – 15. X. 2015, J. A. Moreno-González leg. (ICN-Ar 10644).	en	Martínez, Leonel, Brescovit, Antonio Domingos, Oliveira, Luiz Fernando M. (2020): Two new species of the ghost spider genus Macrophyes O. Pickard-Cambridge 1893 from Colombia and description of the first male of Mesilla Simon, 1903 (Araneae: Anyphaenidae: Anyphaeninae). Zootaxa 4853 (4): 581-590, DOI: 10.11646/zootaxa.4853.4.7
AA067B1DFFF5FF93FF67FA58AB213A20.taxon	etymology	Etymology. The specific epithet is a patronymic given in honor of the deceased Argentine singer Gustavo Cerati, who has greatly contributed to Spanish Rock music.	en	Martínez, Leonel, Brescovit, Antonio Domingos, Oliveira, Luiz Fernando M. (2020): Two new species of the ghost spider genus Macrophyes O. Pickard-Cambridge 1893 from Colombia and description of the first male of Mesilla Simon, 1903 (Araneae: Anyphaenidae: Anyphaeninae). Zootaxa 4853 (4): 581-590, DOI: 10.11646/zootaxa.4853.4.7
AA067B1DFFF5FF93FF67FA58AB213A20.taxon	diagnosis	Diagnosis. Females of Macrophyes ceratii n. sp., can be distinguished of the remain species of the genus by their long copulatory ducts, curved toward the medial region of the epigynum; long atrium, widening posteriorly to a circular shape and small, oval-shaped spermathecae (Figs 2 B, C).	en	Martínez, Leonel, Brescovit, Antonio Domingos, Oliveira, Luiz Fernando M. (2020): Two new species of the ghost spider genus Macrophyes O. Pickard-Cambridge 1893 from Colombia and description of the first male of Mesilla Simon, 1903 (Araneae: Anyphaenidae: Anyphaeninae). Zootaxa 4853 (4): 581-590, DOI: 10.11646/zootaxa.4853.4.7
AA067B1DFFF5FF93FF67FA58AB213A20.taxon	description	Description. Female (Holotype, ICN – Ar 10644). Coloration uniformly pale yellow (Fig. 2 A). Total length 6.32, carapace length 1.81, width 1.28, high 0.77. Clypeus height 0.04. Eye diameters and interdistances: AME 0.06, ALE 0.11, PME 0.10, PLE 0.09; AME – AME 0.14, AME – ALE 0.23, PME – PME 0.29, PME – PLE 0.27, ALE – PLE 0.21. Chelicerae 0.74 long, eight promarginal teeth, nine retromarginal teeth, being the distal longer than other. Leg measurements: leg I — femur 5.93 / patella 0.81 / tibia 6.71 / metatarsus 6.41 / tarsus 2.01 / total 24.87; II — 4.89 / 0.81 / 5.07 / 4.91 / 1.47 / 17.15; III — 3.27 / 0.63 / 2.78 / 3.39 / 0.99 / 11.06; IV — 5.44 / 0.79 / 4.61 / 5.94 / 1.49 / 18.27. Leg spination: I — metatarsus p 1 - 0 - 0 - 0 - 0; II — tibia d 0 - 0 - 0 - 0 - 1, p 0; II = I; III — tibia d 0 - 0 - 0 - 0 - 1, v 0 - 2 - 0 - 2 - 0, p 0 - 0 - 1 - 0 - 1, r 0 - 0 - 1 - 0 - 1, metatarsus v 2 - 0 - 0 - 2 - 0, r 1 - 0 - 1 - 0 - 0, p 1 - 0 - 1 - 0 - 0; IV — tibia d 1 - 0 - 0 - 0 - 0, v 0 - 1 - 0 - 0 - 1, p 0 - 0 - 1 - 1 - 1, r 0 - 0 - 1 - 1 - 1, metatarsus v 0 - 2 - 0 - 1 - 0, p 1 - 0 - 0 - 1 - 1, r 1 - 0 - 1 - 1 - 1. Abdomen: length 4.52, epigastric furrow 0.83 from tracheal spiracle, spiracle 3.07 from base of spinnerets. Epigynum: atrium longer than wide, cylindric-shaped, rounded posteriorly; lateral borders thin, sclerotized, laminar; internally with very long, thin copulatory ducts, curved at medial region; seminal receptacles small and rounded, sub-medially situated on the copulatory ducts; spermathecae large, ovalshaped, posteriorly positioned; fertilization ducts almost as long as the spermathecae (Figs 2 B, C). Male. Unknown.	en	Martínez, Leonel, Brescovit, Antonio Domingos, Oliveira, Luiz Fernando M. (2020): Two new species of the ghost spider genus Macrophyes O. Pickard-Cambridge 1893 from Colombia and description of the first male of Mesilla Simon, 1903 (Araneae: Anyphaenidae: Anyphaeninae). Zootaxa 4853 (4): 581-590, DOI: 10.11646/zootaxa.4853.4.7
AA067B1DFFF5FF93FF67FA58AB213A20.taxon	distribution	Distribution. Only known from the type locality (Fig. 4). Habitat. The specimens were collected beating low shrubs, on foliage, in a conserved high mountain wet forest ecosystem, at 1313 – 1702 meters high.	en	Martínez, Leonel, Brescovit, Antonio Domingos, Oliveira, Luiz Fernando M. (2020): Two new species of the ghost spider genus Macrophyes O. Pickard-Cambridge 1893 from Colombia and description of the first male of Mesilla Simon, 1903 (Araneae: Anyphaenidae: Anyphaeninae). Zootaxa 4853 (4): 581-590, DOI: 10.11646/zootaxa.4853.4.7
AA067B1DFFF3FF93FF67FED8A8F239D1.taxon	diagnosis	Diagnosis. Mesilla species are similar to those of Macrophyes species by having elongated and thin body; long and thin legs; median apophysis on anterior side of the tegulum and by having a large retrolateral, marginal tooth on distal side of chelicerae, but can been distinguished by the long and laminar median apophysis; ventral tegular projection absent and long embolus basally inserted on the tegulum (Figs 3 C−E). Females are characterized by their short copulatory ducts, restricted to the posterior portion of the epigynum and with small, tubular seminal receptacles on medial region; long, wide, divergent lateral borders, with rounded apex (Brescovit 1992: 12, 13; 1997: figs 150 − 151; Figs 3 G, H).	en	Martínez, Leonel, Brescovit, Antonio Domingos, Oliveira, Luiz Fernando M. (2020): Two new species of the ghost spider genus Macrophyes O. Pickard-Cambridge 1893 from Colombia and description of the first male of Mesilla Simon, 1903 (Araneae: Anyphaenidae: Anyphaeninae). Zootaxa 4853 (4): 581-590, DOI: 10.11646/zootaxa.4853.4.7
AA067B1DFFF3FF90FF67FC8BAD193BC4.taxon	description	Figures 3 A – H, 4.	en	Martínez, Leonel, Brescovit, Antonio Domingos, Oliveira, Luiz Fernando M. (2020): Two new species of the ghost spider genus Macrophyes O. Pickard-Cambridge 1893 from Colombia and description of the first male of Mesilla Simon, 1903 (Araneae: Anyphaenidae: Anyphaeninae). Zootaxa 4853 (4): 581-590, DOI: 10.11646/zootaxa.4853.4.7
AA067B1DFFF3FF90FF67FC8BAD193BC4.taxon	diagnosis	Diagnosis. Males of Mesilla vittiventris Simon, 1903 can be distinguished by their very long, thin cymbium; a long median apophysis distally erected and single, extremely short retrolateral tibial apophysis (Figs 3 C – E). Females are characterized by the epigynum with long, wide lateral borders, curved inward and with a septum; a long atrium and small seminal receptacles situated on the anterior third of the copulatory ducts (Figs 3 G, H). Note. Brescovit (1997) illustrated the internal genitalia of the Mesilla vittiventris type specimen. That epigynum, being detached from the abdomen and suffering from dissection, did not present the copulatory ducts, only fertilization ducts (Brescovit 1997, fig. 151). Here, new illustrations of the well preserved epigynal plate of the Colombian specimen belonging to this species are included.	en	Martínez, Leonel, Brescovit, Antonio Domingos, Oliveira, Luiz Fernando M. (2020): Two new species of the ghost spider genus Macrophyes O. Pickard-Cambridge 1893 from Colombia and description of the first male of Mesilla Simon, 1903 (Araneae: Anyphaenidae: Anyphaeninae). Zootaxa 4853 (4): 581-590, DOI: 10.11646/zootaxa.4853.4.7
AA067B1DFFF3FF90FF67FC8BAD193BC4.taxon	description	Description. Male. (ICN – Ar 9696). Carapace yellowish. Chelicerae brown, labium and endites brown. Sternum reddish-brown. Legs yellow. Abdomen dorsally grayish; ventrally yellow. Spinnerets yellow (Fig. 3 A). Total length 7.12, carapace length 2.49, width 1.99, high 0.68. Clypeus height 0.15. Eye diameters and interdistances: AME 0.10, ALE 0.18, PME 0.2, PLE 0.21; AME – AME 0.23, AME – ALE 0.35, PME – PME 0.48, PME – PLE 0.38, ALE – PLE 0.43. Chelicerae 1.72 long, distal large retromarginal teeth, six promarginal teeth, six retromarginal denticle Leg I — femur 6.13 / patella 0.86 / tibia 7.08 / metatarsus 6.82 / tarsus 1.87 / total 22.76; II — 5.77 / 0.81 / 6.68 / 6.59 / 1.85 / 21.7; III — 3.3 / 0.81 / 2.63 / 3.54 / 1.17 / 11.45; IV — 4.42 / 0.73 / 4.77 / 6.31 / 1.55 / 17.78. Leg spination (Fig. 3 F): I — II d 1 - 1 - 0 - 0 - 0, v 2 - 2 - 2 - 2 - 2, p 0, metatarsus p 1 - 0 - 0 - 0 - 0; III — tibia v 0 - 2 - 2 - 0 - 0, p 0 - 1 - 1 - 0 - 0, r 1 - 1 - 1 - 1 - 0, metatarsus v 2 - 0 - 0 - 2 - 0; IV — tibia v 1 - 1 - 1 - 2 - 1, metatarsus v 1 - 0 - 1 - 1 - 1, p 0 - 2 - 0 - 1 - 2. Abdomen, twice the length of carapace, length 4.61, epigastric furrow 0.68 from tracheal spiracle, spiracle 2.93 from base of spinnerets. Palp: retrolateral tibial apophysis very small, apically sharpened; cymbium shorter than tibia; subtegulum inconspicuous; tegulum longer than wide; median apophysis very long, sinuous and apically inserted; embolus long, basally inserted on the tegulum, apically filiform, with narrow base (Figs 3 C – E). Female. Described by Brescovit (1997), figs 150 - 151. Complementary description: Dorsal body as in Fig. 3 B. Epigynum: lateral borders sclerotized, parallel and wide; atrium long; internally with short, wide copulatory ducts; seminal receptacles small and medially situated on the copulatory ducts; spermathecae large, oval, posteriorly positioned; fertilization ducts as long as the spermathecae (Figs 3 G, H). Variation. Males (n = 3): total length: 7.03 – 8.20; carapace length: 2.05 – 2.71; femur I length: 6.13 – 6.66.	en	Martínez, Leonel, Brescovit, Antonio Domingos, Oliveira, Luiz Fernando M. (2020): Two new species of the ghost spider genus Macrophyes O. Pickard-Cambridge 1893 from Colombia and description of the first male of Mesilla Simon, 1903 (Araneae: Anyphaenidae: Anyphaeninae). Zootaxa 4853 (4): 581-590, DOI: 10.11646/zootaxa.4853.4.7
AA067B1DFFF3FF90FF67FC8BAD193BC4.taxon	distribution	Distribution. Known from Colombia and Ecuador (Fig. 4; Brescovit 1997: 65).	en	Martínez, Leonel, Brescovit, Antonio Domingos, Oliveira, Luiz Fernando M. (2020): Two new species of the ghost spider genus Macrophyes O. Pickard-Cambridge 1893 from Colombia and description of the first male of Mesilla Simon, 1903 (Araneae: Anyphaenidae: Anyphaeninae). Zootaxa 4853 (4): 581-590, DOI: 10.11646/zootaxa.4853.4.7
AA067B1DFFF3FF90FF67FC8BAD193BC4.taxon	materials_examined	Material examined. Three males from surroundings of the entrance of the Reserva Natural Río Ñambí, (1 ° 17 ′ 4.38 ″ N; 78 ° 5 ′ 18.24 ″ W), 1179 m, Vereda Altaquer, Barbacoas, Nariño, Colombia, 5. VII. 2017, W. Galvis leg. (ICN-Ar 9696 [m # 1], ICN-Ar 10644 [m # 2]). Female from Alto del Mirador (2 ° 58 ′ 0 ″ N, 78 ° 11 ′ 0 ″ W), 180 m, Parque Nacional Natural Gorgona, Cauca, Colombia, 30. XI. 2000, H. Torrez leg. (ICN-Ar 6271).	en	Martínez, Leonel, Brescovit, Antonio Domingos, Oliveira, Luiz Fernando M. (2020): Two new species of the ghost spider genus Macrophyes O. Pickard-Cambridge 1893 from Colombia and description of the first male of Mesilla Simon, 1903 (Araneae: Anyphaenidae: Anyphaeninae). Zootaxa 4853 (4): 581-590, DOI: 10.11646/zootaxa.4853.4.7
