taxonID	type	description	language	source
AA52B01C0807FFB3D5B83DAA08F09A7B.taxon	description	Fig. 4 a-g	en	SCHLAGINTWEIT, FELIX, YAZDI-MOGHADAM, MOHSEN (2023): Pseudocyclammina Sarvakensis Sp. Nov. And Pseudotextulariella Brevicamerata Sp. Nov.: Further Evidence For The Cenomanian Megadiversity Of Larger Benthic Foraminifera From The Sarvak Formation Of Sw Iran. Acta Palaeontologica Romaniae 19 (2): 3-13, DOI: 10.35463/j.apr.2023.02.01, URL: http://dx.doi.org/10.35463/j.apr.2023.02.01
AA52B01C0807FFB3D5B83DAA08F09A7B.taxon	etymology	Derivatio nominis: The species name refers to the Sarvak Formation cropping out in southwestern Iran (Zagros Zone).	en	SCHLAGINTWEIT, FELIX, YAZDI-MOGHADAM, MOHSEN (2023): Pseudocyclammina Sarvakensis Sp. Nov. And Pseudotextulariella Brevicamerata Sp. Nov.: Further Evidence For The Cenomanian Megadiversity Of Larger Benthic Foraminifera From The Sarvak Formation Of Sw Iran. Acta Palaeontologica Romaniae 19 (2): 3-13, DOI: 10.35463/j.apr.2023.02.01, URL: http://dx.doi.org/10.35463/j.apr.2023.02.01
AA52B01C0807FFB3D5B83DAA08F09A7B.taxon	materials_examined	Holotype: Specimen illustrated in Figure 4 b, equatorial section, sample RAP 181. Type-locality: Tang-e Darbast section, Interior Fars, Zagros Mountains (Figs. 2 - 3).	en	SCHLAGINTWEIT, FELIX, YAZDI-MOGHADAM, MOHSEN (2023): Pseudocyclammina Sarvakensis Sp. Nov. And Pseudotextulariella Brevicamerata Sp. Nov.: Further Evidence For The Cenomanian Megadiversity Of Larger Benthic Foraminifera From The Sarvak Formation Of Sw Iran. Acta Palaeontologica Romaniae 19 (2): 3-13, DOI: 10.35463/j.apr.2023.02.01, URL: http://dx.doi.org/10.35463/j.apr.2023.02.01
AA52B01C0807FFB3D5B83DAA08F09A7B.taxon	description	Type-horizon: Lower part of the middle Cenomanian of the Sarvak Formation. Description: Medium-sized (max. diameter 1.35 mm) species of Pseudocylammina with a planispiral, involute test displaying a rounded periphery. The spiral stage is well developed, and a tendency to uncoil in the final stage has not been observed. In subaxial sections, the test is slightly compressed (nautiloid morphology). Test surface as seen in equatorial section is smooth with much reduced to almost no sutural depressions. The chambers following the subspherical proloculus are arranged in 2.5 to 3.5 whorls displaying 14 to 16 chambers in the final whorl. The dimensions for the four measured proloculi diameters (0.110 mm, 0.125 mm, 0.140 mm and 0.300 mm) suggest a test dimorphism that needs further clarification. The early postembryonic chambers are subrhombohedric later with increasing width become recurved. The chamber lumen is markedly higher as the thickness of the septa and also of the wall. Multiple and tiny foramina (diameter 20 - 35 µm), with parallel arrangement in axial sections and septal spaces about twice the width of the former (Fig. 4 f). Wall agglutinated, alveolar.	en	SCHLAGINTWEIT, FELIX, YAZDI-MOGHADAM, MOHSEN (2023): Pseudocyclammina Sarvakensis Sp. Nov. And Pseudotextulariella Brevicamerata Sp. Nov.: Further Evidence For The Cenomanian Megadiversity Of Larger Benthic Foraminifera From The Sarvak Formation Of Sw Iran. Acta Palaeontologica Romaniae 19 (2): 3-13, DOI: 10.35463/j.apr.2023.02.01, URL: http://dx.doi.org/10.35463/j.apr.2023.02.01
AA52B01C0807FFB3D5B83DAA08F09A7B.taxon	diagnosis	Comparisons: As can be deduced from the synonymy, the new species has been confounded with Pseudocyclammina rugosa (d’Orbigny, 1850). These assignments however must have been made on the basis simplicity as P. rugosa represented the only species of the genus described (until now) from Cenomanian strata. However, with respect to the traditional specific criteria, P. sarvakensis nov. sp. and P. rugosa are by no means comparable. According to Maync (1952), P. rugosa attains 3.1 to 3.8 mm millimeters in length, displaying five to six large inflated chambers in the last whorl (Fig. 4 i). It is worth mentioning that already Henson (1948, p. 10 and 15) mentioned that the wall structure of ‘ Lituola ’ rugosa d’Orbigny and Pseudocyclammina “ are essentially similar in construction ” and that “ separation of the two genera is somewhat uncertain ”. Another upper Cretaceous representative of Pseudocyclammina is P. massiliensis described by Maync (1959) from the Santonian of southeastern France (Fig. 4 h). This is also a comparably large species commonly displaying uncoiled elongated tests (up to 2.3 mm) with five to seven chambers in the final whorl. So far no specimen of P. sarvakensis displaying uncoiled test has been observed. The number of chambers in the final whorl might differentiate P. sarvakensis from all other Upper Jurassic – Lower Cretaceous Pseudocyclammina species (Maync, 1959, tab. 1; Banner & Whittaker, 1991). For example, the Upper Jurassic-Lower Cretaceous type-species Pseudocyclammina lituus (Yokoyama, 1890) typically has 8 to 12 chambers in the last whorl (Banner & Whittaker, 1991). The Lower Cretaceous (Aptian-Albian) P. hedbergi Maync, 1953 and P. vasconica, 1959 have 4 - 5 and 7 - 8 chambers in the final whorl. Pseudocyclammina sphaeroidea Gendrot, 1968 (= Mayncina sphaeroidea acc. to De Castro, 1991) described from the upper Santonian of southeastern France is distinctly smaller than P. sarvakensis with test length of 0.5 to 0.6 mm, often uncoiling (Fig. 4 j-m). The youngest record of the genus Pseudocyclammina seems to be from the upper Campanian (Parente, 1994, pl. 1, fig. 5, Pseudocyclammina sp., in our opinion referable to P. massiliensis Maync).	en	SCHLAGINTWEIT, FELIX, YAZDI-MOGHADAM, MOHSEN (2023): Pseudocyclammina Sarvakensis Sp. Nov. And Pseudotextulariella Brevicamerata Sp. Nov.: Further Evidence For The Cenomanian Megadiversity Of Larger Benthic Foraminifera From The Sarvak Formation Of Sw Iran. Acta Palaeontologica Romaniae 19 (2): 3-13, DOI: 10.35463/j.apr.2023.02.01, URL: http://dx.doi.org/10.35463/j.apr.2023.02.01
AA52B01C0807FFB0D61F390D0E699B57.taxon	description	Fig. 5 a-i, k	en	SCHLAGINTWEIT, FELIX, YAZDI-MOGHADAM, MOHSEN (2023): Pseudocyclammina Sarvakensis Sp. Nov. And Pseudotextulariella Brevicamerata Sp. Nov.: Further Evidence For The Cenomanian Megadiversity Of Larger Benthic Foraminifera From The Sarvak Formation Of Sw Iran. Acta Palaeontologica Romaniae 19 (2): 3-13, DOI: 10.35463/j.apr.2023.02.01, URL: http://dx.doi.org/10.35463/j.apr.2023.02.01
AA52B01C0807FFB0D61F390D0E699B57.taxon	etymology	Derivatio nominis: Prefix brevis (latin) = short combined with camerata = chamber.	en	SCHLAGINTWEIT, FELIX, YAZDI-MOGHADAM, MOHSEN (2023): Pseudocyclammina Sarvakensis Sp. Nov. And Pseudotextulariella Brevicamerata Sp. Nov.: Further Evidence For The Cenomanian Megadiversity Of Larger Benthic Foraminifera From The Sarvak Formation Of Sw Iran. Acta Palaeontologica Romaniae 19 (2): 3-13, DOI: 10.35463/j.apr.2023.02.01, URL: http://dx.doi.org/10.35463/j.apr.2023.02.01
AA52B01C0807FFB0D61F390D0E699B57.taxon	materials_examined	Holotype: Specimen illustrated in Figure 5 e, oblique axial section, sample RAP 188. Type-locality: Tang-e Darbast section, Interior Fars, Zagros Mountains (Figs. 2 - 3).	en	SCHLAGINTWEIT, FELIX, YAZDI-MOGHADAM, MOHSEN (2023): Pseudocyclammina Sarvakensis Sp. Nov. And Pseudotextulariella Brevicamerata Sp. Nov.: Further Evidence For The Cenomanian Megadiversity Of Larger Benthic Foraminifera From The Sarvak Formation Of Sw Iran. Acta Palaeontologica Romaniae 19 (2): 3-13, DOI: 10.35463/j.apr.2023.02.01, URL: http://dx.doi.org/10.35463/j.apr.2023.02.01
AA52B01C0807FFB0D61F390D0E699B57.taxon	description	Type-horizon: Lower part of the middle Cenomanian of the Sarvak Formation. Description: Conical to wedge-shaped test, slightly compressed, higher than wide and with a central depression at the base. The test exhibits a well developed biserial part with chambers of distinctly reduced height compared for example with the test diameter. The short initial triserial stage as typical for the genus (Barnard, 1963) has not been observed due to inadequate sections. The biseriate chambers are subdivided by both horizontal and vertical (= radial) partitions. It appears that the horizontal partitions (rafters) are not present in the early part but appear later during ontogeny. In contrast hereto, the radial vertical partitions (up to three orders) seem to be present already in the early stage. The main vertical partitions (beams) stretch far into the test interior (length up to 0.2 mm in the middle part of the test), while the other orders (intercalary beams) are distinctly shorter. The rather short third order partitions (see ib in Fig. 5 f) reach up to 0.05 mm in length and are confined to the marginal part of the chamber.	en	SCHLAGINTWEIT, FELIX, YAZDI-MOGHADAM, MOHSEN (2023): Pseudocyclammina Sarvakensis Sp. Nov. And Pseudotextulariella Brevicamerata Sp. Nov.: Further Evidence For The Cenomanian Megadiversity Of Larger Benthic Foraminifera From The Sarvak Formation Of Sw Iran. Acta Palaeontologica Romaniae 19 (2): 3-13, DOI: 10.35463/j.apr.2023.02.01, URL: http://dx.doi.org/10.35463/j.apr.2023.02.01
AA52B01C0807FFB0D61F390D0E699B57.taxon	diagnosis	Comparisons: Pseudotextulariella brevicamerata sp. nov. may exhibit the same number of biserial chambers, but within a test height half the size of the late Albian-Cenomanian P. cretosa (Cushman, 1932). This means, that the chamber height (= lumen) is distinctly reduced in the former. While the height of the chamber lumen often equals the thickness of the septa in P. brevicamerata sp. nov., it may be up to three times higher in P. cretosa. Also, the overall test dimensions (height, diameter) are larger in P. cretosa (Table 1). With its several orders of rafters (at least in the adult part), P. cretosa is structurally more complex than P. brevicamerata. P. courtionensis Brönnimann, 1967 from the Valanginian of Switzerland exhibits an uncompressed test yielding rounded transverse sections (Brönnimann, 1967, pl. 1, fig. 3, pl. 2, fig. 1). For all species of Pseudotextulariella however it appears that transverse sections through the juvenile part of the test are always subcircular and that a compression in the plane of biseriality appears throughout ontogeny. Like P. brevicamerata, the Swiss form also has only one order of rafters equally appearing in the adult part of the test. Note that in oblique sections, vertical partitions can be misidentified as rafters resulting in a (false) higher number of the latter (Fig. 5 l, left lower part). Exhibiting equivalent dimensions (height, diameter) compared with P. courtionensis, P. brevicamerata has more chambers due to reduced thickness of septa and chamber height (lumen). Last but not least, the different stratigraphy (late Berriasian-Valanginian vs. middle Cenomanian) as well as the different palaeogeographic locations (northern margin vs. southern margin, here Arabian Plate, of Neotethys) merits the differentiation of the new species described herein.	en	SCHLAGINTWEIT, FELIX, YAZDI-MOGHADAM, MOHSEN (2023): Pseudocyclammina Sarvakensis Sp. Nov. And Pseudotextulariella Brevicamerata Sp. Nov.: Further Evidence For The Cenomanian Megadiversity Of Larger Benthic Foraminifera From The Sarvak Formation Of Sw Iran. Acta Palaeontologica Romaniae 19 (2): 3-13, DOI: 10.35463/j.apr.2023.02.01, URL: http://dx.doi.org/10.35463/j.apr.2023.02.01
AA52B01C0807FFB0D61F390D0E699B57.taxon	discussion	Remarks: The late Albian-Cenomanian Pseudotextulariella cretosa (Cushman) represents a typical taxon of the boreal realm (type-locality England; Germany, Frieg, 1989: Poland: Gawor-Biedowa, 1972) and the western part of the northern margin of Neotethys (Romania: Bucur & Baltres, 2002). Usually, it occurs in marly shelf lithologies (‘ chalk facies’) delivering free, isolated specimens, but it may also be found in mixed siliciclastic-carbonate orbitolinid-bearing facies (Bucur & Baltres, 2002). Pseudotextulariella brevicamerata sp. nov. instead occurs in foraminiferal wackestones-packstones of typical inner platform facies. The palaeogeographic setting of the Iranian Sarvak Formation refers its occurrence to the former southern margin of the Neotethys, i. e. the Arabian Plate (e. g., Ziegler, 2001). Both, Pseudocyclammina sarvakensis and Pseudotextulariella brevicamerata are further elements of the so-called ‘ Nezzazata - alveolinid’ assemblage zone sensu Wynd (1965) characterizing the late early to upper Cenomanian inner platform carbonates of the Sarvak Formation. For both taxa, the complete stratigraphic range could be modified by future new finds.	en	SCHLAGINTWEIT, FELIX, YAZDI-MOGHADAM, MOHSEN (2023): Pseudocyclammina Sarvakensis Sp. Nov. And Pseudotextulariella Brevicamerata Sp. Nov.: Further Evidence For The Cenomanian Megadiversity Of Larger Benthic Foraminifera From The Sarvak Formation Of Sw Iran. Acta Palaeontologica Romaniae 19 (2): 3-13, DOI: 10.35463/j.apr.2023.02.01, URL: http://dx.doi.org/10.35463/j.apr.2023.02.01
