taxonID	type	description	language	source
A5636478EE04FFB1FF3DBAE5FCEF76C1.taxon	distribution	Distribution Origin Nearctic Distribution Canada: MB, NB, ON; USA: not recorded New provincial Canada: Manitoba: Moose Lake, 2016 - IX- 19, 49.2018 ° N, 95.3423 ° W, 353 m, records mushroom, sifting, B. Godin & D. Horwood (BGC) 1 male, 5 females; same locality data (LFC) 2 females; Adam Lake, 2016 - IX- 11, 49.0508 ° N, 100.0666 ° W, 689 m, mushrooms, sifting, B. Godin & D. Horwood (LFC) 1 male, 3 females. References Webster et al. 2016, Klimaszewski et al. 2018 Collection and habitat data. Habitat (outside of the study area). In old red maple (Acer rubrum L.) forest, mixed forests, a wet alder (Alnus sp.) swamp, a mature red spruce (Picea rubens Sarg.) and red maple forest, a rich Appalachian hardwood forest, in a Carex marsh, small sedge marsh, marsh with scattered alders, in old-growth eastern white cedar (Thuja occidentalis L.) swamps and forests, and in 8.5 - year-old regenerating mixed forests. Adults in moss and litter near brooks, in moss and litter at the base of cedar, in moss and litter in red spruce and cedar forests, in leaf litter under alders near a stream, in leaf litter and grass on hummocks in a wet alder swamp, in grass litter and sphagnum in marshes, moist leaves on the margin of a vernal pool, in sphagnum and leaf litter at the bottom of old tire depressions, and one specimen was collected from a gilled mushroom. Collecting period. III – IX. Collecting method. Sifting various kinds of litter.	en	Klimaszewski, Jan, Godin, Benoit, Davies, Anthony, Bourdon, Caroline, Horwood, Denise (2018): Forty new records of aleocharine beetles, and two new species in the genera Acrotona Thomson and Atheta Thomson, for the province of Manitoba, Canada (Coleoptera: Staphylinidae: Aleocharinae). Insecta Mundi 2018 (641): 1-33, DOI: 10.5281/zenodo.5173990
A5636478EE05FFB0FF3DBED0FBF07460.taxon	description	Fig. 1 – 8	en	Klimaszewski, Jan, Godin, Benoit, Davies, Anthony, Bourdon, Caroline, Horwood, Denise (2018): Forty new records of aleocharine beetles, and two new species in the genera Acrotona Thomson and Atheta Thomson, for the province of Manitoba, Canada (Coleoptera: Staphylinidae: Aleocharinae). Insecta Mundi 2018 (641): 1-33, DOI: 10.5281/zenodo.5173990
A5636478EE05FFB0FF3DBED0FBF07460.taxon	materials_examined	Holotype. Canada: Manitoba, Max Lake, 2016 - IX- 10, 49.0714 ° N, 100.1384 ° W, 686 m, mushrooms, sifting, coll. B. Godin & D. Horwood (CNC) 1 male. Paratype. Canada: Manitoba, Katherine Lake, 2016 - IX- 14, 50.6603 ° N, 99.8947 ° W, 652 m, litter, sifting, coll. B. Godin & D. Horwood (LFC) 1 female.	en	Klimaszewski, Jan, Godin, Benoit, Davies, Anthony, Bourdon, Caroline, Horwood, Denise (2018): Forty new records of aleocharine beetles, and two new species in the genera Acrotona Thomson and Atheta Thomson, for the province of Manitoba, Canada (Coleoptera: Staphylinidae: Aleocharinae). Insecta Mundi 2018 (641): 1-33, DOI: 10.5281/zenodo.5173990
A5636478EE05FFB0FF3DBED0FBF07460.taxon	etymology	Etymology. Named after the Canadian province of Manitoba, where the type specimens were found.	en	Klimaszewski, Jan, Godin, Benoit, Davies, Anthony, Bourdon, Caroline, Horwood, Denise (2018): Forty new records of aleocharine beetles, and two new species in the genera Acrotona Thomson and Atheta Thomson, for the province of Manitoba, Canada (Coleoptera: Staphylinidae: Aleocharinae). Insecta Mundi 2018 (641): 1-33, DOI: 10.5281/zenodo.5173990
A5636478EE05FFB0FF3DBED0FBF07460.taxon	description	Description. Body narrow and subparallel (Fig. 1), moderately convex; length 2.8 - 2.9 mm, width at middle of pronotum 0.9 mm, width at base of elytra 0.6 mm; colour dark brown with elytra and apical segments of abdomen slightly paler rufobrunneous, appendages brunneotestaceous; punctation on forebody fine, moderately dense and slightly asperate, particularly on pronotum (Fig. 1); head distinctly narrower than pronotum, ratio of maximum width of head to maximum width of pronotum 0.7; antennomeres VII – X subquadrate; pronotum broad and shield-shaped, sides evenly arcuate, maximum width of pronotum greater than width of elytra at base, ratio of maximum width to length 1.4; elytra at suture shorter than pronotum; abdomen slightly tapering posteriorly. Male. Tergite VIII moderately transverse, apical margin arcuate (Fig. 4); sternite VIII elongate, broadly parabolically rounded from base to apex, base slightly sinuate (Fig. 5); tubus of median lobe of aedeagus in lateral view short, with ventral margin curved obliquely ventrad in basal half and almost straight in apical half, with apex narrow and rounded, internal sac with narrow, sclerotized, curved subapical structure (Fig. 2); in dorsal view median lobe with tubus narrow, subparallel, broadly rounded apically, bulbus very wide, subovoid, internal sac with two narrow arcuate structures (Fig. 3). Female. Tergite VIII similar to that of male (Fig. 6); sternite VIII elongate, apex rounded, base evenly arcuate (Fig. 7); spermatheca with capsule kidney-shaped, stem narrow, with single broad coil posteriorly (Fig. 8).	en	Klimaszewski, Jan, Godin, Benoit, Davies, Anthony, Bourdon, Caroline, Horwood, Denise (2018): Forty new records of aleocharine beetles, and two new species in the genera Acrotona Thomson and Atheta Thomson, for the province of Manitoba, Canada (Coleoptera: Staphylinidae: Aleocharinae). Insecta Mundi 2018 (641): 1-33, DOI: 10.5281/zenodo.5173990
A5636478EE05FFB0FF3DBED0FBF07460.taxon	distribution	Distribution. Known only from the type locality in Manitoba. Collection and habitat data. Captured in September by sifting organic litter in deciduous forest and mushrooms.	en	Klimaszewski, Jan, Godin, Benoit, Davies, Anthony, Bourdon, Caroline, Horwood, Denise (2018): Forty new records of aleocharine beetles, and two new species in the genera Acrotona Thomson and Atheta Thomson, for the province of Manitoba, Canada (Coleoptera: Staphylinidae: Aleocharinae). Insecta Mundi 2018 (641): 1-33, DOI: 10.5281/zenodo.5173990
A5636478EE05FFB0FF3DBED0FBF07460.taxon	discussion	Comments. This species differs from all other Nearctic Acrotona species known to us in combination of the following characters: broad and shield-shaped pronotum that is wider than base of elytra, elytra at suture shorter than pronotum, unique shape of median lobe of aedeagus with large bulbus and two elongate narrow structures of internal sac (Fig. 2, 3), distinct shape of tergite and sternite VIII of male and female (Fig. 4 – 7), and the distinct shape of spermatheca (Fig. 8).	en	Klimaszewski, Jan, Godin, Benoit, Davies, Anthony, Bourdon, Caroline, Horwood, Denise (2018): Forty new records of aleocharine beetles, and two new species in the genera Acrotona Thomson and Atheta Thomson, for the province of Manitoba, Canada (Coleoptera: Staphylinidae: Aleocharinae). Insecta Mundi 2018 (641): 1-33, DOI: 10.5281/zenodo.5173990
A5636478EE05FFB0FF3DBB5BFC4376C1.taxon	distribution	Distribution Origin Nearctic Distribution Canada: MB, NB, ON, QC; USA: NY New provincial record Canada: Manitoba: Birds Hill [Prov. Park] - bur oak [trail], 2016 - IX- 16, 50.0083 ° N, 96.9216 ° W, 264 m, mushrooms, sifting, B. Godin & D. Horwood (BGC) 1 male, 3 females; same locality data (LFC) 1 male, 3 females. References Casey 1910, Klimaszewski et al. 2018 Collection and habitat data. Habitat (outside of the study area). In ON, a common species in various kinds of deciduous forests, mixed forests, and a semi-open habitat (e. g. oak savannah); adults sifted from grass pile and leaves and captured in Malaise pans and Lindgren funnel traps. In NB, adults collected from decaying sea wrack under alders (Alnus sp.) and gravel and sand on the margin of a sea beach where it was common. Collecting period. V – IX. Collecting method. Pitfall traps, Lindgren funnel traps, flight intercept traps, Malaise pan traps, sifting litter, car net.	en	Klimaszewski, Jan, Godin, Benoit, Davies, Anthony, Bourdon, Caroline, Horwood, Denise (2018): Forty new records of aleocharine beetles, and two new species in the genera Acrotona Thomson and Atheta Thomson, for the province of Manitoba, Canada (Coleoptera: Staphylinidae: Aleocharinae). Insecta Mundi 2018 (641): 1-33, DOI: 10.5281/zenodo.5173990
A5636478EE02FFB7FF3DBED1FCE573BE.taxon	distribution	Distribution Origin Palearctic, adventive in North America Distribution Canada: MB, LB, NB, NF, NS, ON, PE, QC, SK. USA: CA, CO, IN, MA, ME, MI, NH, NY, PA, VT, WA New provincial Canada: Manitoba: Souris River, 2016 - IX- 9, 49.5022 ° N, 99.9915 ° W, 402 m, river record bank, oak litter, sifting, B. Godin & D. Horwood (BGC) 1 female; same locality data (LFC) 2 females. References Klimaszewski et al. 2018 Collection and habitat data. Habitat (outside of the study area). In many open and forested habitats; agricultural fields, pasture / meadow (commonly collected by sweeping grasses), urban fields, in grass litter in garden and lawn, coastal sand dunes amidst vegetation, river margins in flood debris and drift material, old-growth eastern white cedar swamp in moist sphagnum, black spruce bog in sphagnum, and red spruce forest in pitfall traps. Collecting period. IV – IX. Collecting method. Sifting various kinds of litter and moss, pitfall traps, sweeping grasses.	en	Klimaszewski, Jan, Godin, Benoit, Davies, Anthony, Bourdon, Caroline, Horwood, Denise (2018): Forty new records of aleocharine beetles, and two new species in the genera Acrotona Thomson and Atheta Thomson, for the province of Manitoba, Canada (Coleoptera: Staphylinidae: Aleocharinae). Insecta Mundi 2018 (641): 1-33, DOI: 10.5281/zenodo.5173990
A5636478EE02FFB7FF3DBD36FCC974C0.taxon	distribution	Distribution Origin Nearctic Distribution Canada: MB, NB, NS, ON, QC, YT. USA: not recorded. New provincial record Canada: Manitoba: Hecla, 2016 - IX- 17, 51.7849 ° N, 96.6152 ° W, 218 m, mushroom, sifting, B. Godin & D. Horwood (LFC) 1 female. References Klimaszewski et al. 2018 Collection and habitat data. Habitat (outside of the study area). In NB, from various forest types; upland black spruce forest, red spruce and yellow birch forest, mature red spruce and eastern white cedar forest, balsam fir stand, red pine forest, jack pine forest, various mixed forests, red oak forest, and various hardwood forests. Adults in various kinds of mushrooms (mostly gilled mushroom species) in various stages of decay (fresh to well rotten) on forest floor; also found in forest litter (moldy conifer duff, leaf litter), some from pitfall traps in regenerating red spruce forest and yellow birch forest. Others from decaying (moldy) corncobs and cornhusks near mixed forest. Collecting period. V – IX. Collecting method. Sifting fungi and forest litter, pitfall traps.	en	Klimaszewski, Jan, Godin, Benoit, Davies, Anthony, Bourdon, Caroline, Horwood, Denise (2018): Forty new records of aleocharine beetles, and two new species in the genera Acrotona Thomson and Atheta Thomson, for the province of Manitoba, Canada (Coleoptera: Staphylinidae: Aleocharinae). Insecta Mundi 2018 (641): 1-33, DOI: 10.5281/zenodo.5173990
A5636478EE02FFB6FF3DBBBBFA687546.taxon	description	Fig. 1 – 16	en	Klimaszewski, Jan, Godin, Benoit, Davies, Anthony, Bourdon, Caroline, Horwood, Denise (2018): Forty new records of aleocharine beetles, and two new species in the genera Acrotona Thomson and Atheta Thomson, for the province of Manitoba, Canada (Coleoptera: Staphylinidae: Aleocharinae). Insecta Mundi 2018 (641): 1-33, DOI: 10.5281/zenodo.5173990
A5636478EE02FFB6FF3DBBBBFA687546.taxon	materials_examined	Holotype. Canada: Manitoba, Moose Lake, 2016 - IX- 19, 49.2018 ° N, 95.3423 ° W, 353 m, Birch litter, sifting, B. Godin & D. Horwood (CNC) 1 female. Paratype. Canada: Manitoba, Moose Lake, 2016 - IX- 19, 49.2018 ° N, 95.3423 ° W, 353 m, Birch litter, sifting, B. Godin & D. Horwood (LFC) 1 male.	en	Klimaszewski, Jan, Godin, Benoit, Davies, Anthony, Bourdon, Caroline, Horwood, Denise (2018): Forty new records of aleocharine beetles, and two new species in the genera Acrotona Thomson and Atheta Thomson, for the province of Manitoba, Canada (Coleoptera: Staphylinidae: Aleocharinae). Insecta Mundi 2018 (641): 1-33, DOI: 10.5281/zenodo.5173990
A5636478EE02FFB6FF3DBBBBFA687546.taxon	etymology	Etymology. Named after the Canadian province of Manitoba, where the type specimens were found.	en	Klimaszewski, Jan, Godin, Benoit, Davies, Anthony, Bourdon, Caroline, Horwood, Denise (2018): Forty new records of aleocharine beetles, and two new species in the genera Acrotona Thomson and Atheta Thomson, for the province of Manitoba, Canada (Coleoptera: Staphylinidae: Aleocharinae). Insecta Mundi 2018 (641): 1-33, DOI: 10.5281/zenodo.5173990
A5636478EE02FFB6FF3DBBBBFA687546.taxon	description	Description. Body narrow, subparallel, length 2.7 – 2.8 mm; head, antennae, pronotum, and abdomen (except for its apex), dark brown, elytra and legs light brown or light reddish-brown (Fig. 9); antennomeres V-VI elongate, and VII-X subquadrate; pronotum transverse (maximum width 0.6 mm), evenly arcuate laterally, slightly narrower than maximum width of elytra, arcuate basally (Fig. 9); elytra strongly transverse (maximum width 0.7 mm), distinctly shorter than pronotum (Fig. 9); abdomen subparallel, with very coarse macrosetae apically; legs very long (Fig. 9). Male. Tergite VIII transverse, arcuate apically (Fig. 12); sternite VIII broadly parabolic, slightly truncate apically, with numerous strong macrosetae in apical part of disc except medially, antecostal suture almost straight and narrowly separated from basal margin medially (Fig. 13); median lobe of aedeagus in dorsal view with bulbus narrowly oval, tubus long and broad, slightly tapering to triangular apex (Fig. 11), in lateral view ventral margin of tubus strongly curved ventrad in basal half, almost straight to narrowly elongate subapical part, apex narrowly rounded (Fig. 10). Female. Tergite VIII, transverse, truncate medially at apex (Fig. 14); sternite VIII broadly parabolic from base to apex, with numerous strong macrosetae in apical part of disc, antecostal suture sinuate, moderately separated from basal margin (Fig. 15); spermatheca with capsule small, pear-shaped with deep apical invagination, stem long, thin and arcuate medially, narrowly recurved posteriorly (Fig. 16).	en	Klimaszewski, Jan, Godin, Benoit, Davies, Anthony, Bourdon, Caroline, Horwood, Denise (2018): Forty new records of aleocharine beetles, and two new species in the genera Acrotona Thomson and Atheta Thomson, for the province of Manitoba, Canada (Coleoptera: Staphylinidae: Aleocharinae). Insecta Mundi 2018 (641): 1-33, DOI: 10.5281/zenodo.5173990
A5636478EE02FFB6FF3DBBBBFA687546.taxon	distribution	Distribution. Known only from the type locality in Manitoba. Collection and habitat data. Habitat. associated with birch forest; adults found in litter. Collecting period. IX. Collecting method. sifting forest litter.	en	Klimaszewski, Jan, Godin, Benoit, Davies, Anthony, Bourdon, Caroline, Horwood, Denise (2018): Forty new records of aleocharine beetles, and two new species in the genera Acrotona Thomson and Atheta Thomson, for the province of Manitoba, Canada (Coleoptera: Staphylinidae: Aleocharinae). Insecta Mundi 2018 (641): 1-33, DOI: 10.5281/zenodo.5173990
A5636478EE02FFB6FF3DBBBBFA687546.taxon	discussion	Comments. This species differs from other Atheta species in having a broad and shield-shaped pronotum, evenly arcuate laterally, slightly wider than the maximum width of the elytra; elytra shorter than the pronotum; macrosetae of the apical part of the abdomen strong; antennomeres VII – X subquadrate; legs very long (middle and hind legs each as long as length of pronotum and elytra combined); and the shape of the median lobe of the aedeagus and spermatheca are distinctive (Fig. 10, 11). The shape of median lobe of the aedeagus is similar to some species of the subgenus Dimetrota of Atheta, but all species of this subgenus known to us have the apical margin of male tergite VIII modified, with lateral teeth and a median emargination, which is lacking in this species. In Benick and Lohse (1974) this species would key to “ Mischgruppe II ” [= mixed group] of Atheta. A female was designated as a holotype for this species because of the unique shape of the spermatheca.	en	Klimaszewski, Jan, Godin, Benoit, Davies, Anthony, Bourdon, Caroline, Horwood, Denise (2018): Forty new records of aleocharine beetles, and two new species in the genera Acrotona Thomson and Atheta Thomson, for the province of Manitoba, Canada (Coleoptera: Staphylinidae: Aleocharinae). Insecta Mundi 2018 (641): 1-33, DOI: 10.5281/zenodo.5173990
A5636478EE03FFB6FF3DBA3DFE8774B9.taxon	distribution	Distribution Origin Nearctic Distribution Canada: LB, MB, NB, NF, ON, QC, YT. USA: not recorded. New provincial record Canada: Manitoba: Bead Lake, 2016 - IX- 13, 50.7868 ° N, 99.9928 ° W, 620 m, mushrooms, sifting, B. Godin & D. Horwood (BGC) 1 male. References Klimaszewski et al. 2005, Klimaszewski et al. 2018 Collection and habitat data. Habitat (outside of the study area). In red spruce, balsam fir, jack pine, mixed and riparian type forests. Most specimens were from pitfall traps and flight intercept traps; some from forest litter. This species feeds on several species of yeasts and fungal spores based on contents in the posterior gut. Collecting period. VI – VIII. Collecting method. Sifting litter, pitfall traps, flight intercept traps.	en	Klimaszewski, Jan, Godin, Benoit, Davies, Anthony, Bourdon, Caroline, Horwood, Denise (2018): Forty new records of aleocharine beetles, and two new species in the genera Acrotona Thomson and Atheta Thomson, for the province of Manitoba, Canada (Coleoptera: Staphylinidae: Aleocharinae). Insecta Mundi 2018 (641): 1-33, DOI: 10.5281/zenodo.5173990
A5636478EE03FFB5FF3DB832FE877096.taxon	distribution	Distribution Origin Nearctic Distribution Canada: LB, MB, NB, NF, ON, QC, SK. USA: ME New provincial record Canada: Manitoba: Katherine Lake, 2016 - IX- 14, 50.6603 ° N, 99.8947 ° W, 652 m, mushrooms, sifting, B. Godin & D. Horwood (BGC) 1 male. References Bernhauer 1907, Klimaszewski et al. 2018 Collection and habitat data. Habitat (outside of the study area). In NB, found in various forested and open habitats, often in decaying organic materials; in compost near mixed forest, in dung in entrance to fox den in a pasture, in decaying sea wrack on sea beach, under dung on margin of lake, in litter of roof of an abandoned beaver lodge occupied by muskrats, in decaying moist litter in Carex marsh, and from unbaited pitfall traps in a red spruce forest. In SK, in compost, wrack on lakeshore; in NF, captured in carrion-baited pitfall traps and flight intercept traps in conifer-dominated and mixed forests and on coastal barrens in LB. Collecting period. IV – IX. Collecting method. Sifting litter, pitfall traps, flight intercept traps.	en	Klimaszewski, Jan, Godin, Benoit, Davies, Anthony, Bourdon, Caroline, Horwood, Denise (2018): Forty new records of aleocharine beetles, and two new species in the genera Acrotona Thomson and Atheta Thomson, for the province of Manitoba, Canada (Coleoptera: Staphylinidae: Aleocharinae). Insecta Mundi 2018 (641): 1-33, DOI: 10.5281/zenodo.5173990
A5636478EE00FFB5FF3DBC0DFC2F75A9.taxon	distribution	Distribution Origin Holarctic, recorded from Europe, China, India, Japan and South Korea Distribution Canada: AB, BC, LB, MB, NB, NF, NS, ON, PE, QC, SK, YT. USA: AK, NY, PA, RI New provincial Canada: Manitoba: Birds Hill [Prov. Park] - bur oak [trail], 2016 - IX- 16, records 50.0083 ° N, 96.9216 ° W, 264 m, mushrooms, sifting, B. Godin & D. Horwood (BGC) 1 male, 1 female; Georges Creek, 2016 - IX- 15, 50.6729 ° N, 99.6212 ° W, 696 m, mushrooms, sifting, B. Godin & D. Horwood (BGC) 1 male, 1 female; same locality data (LFC) 1 male, 1 female; Adam Lake, 2016 - IX- 11, 49.0508 ° N, 100.0666 ° W, 689 m, mushrooms, sifting, B. Godin & D. Horwood (LFC) 1 male, 1 female. References Thomson 1867, Klimaszewski et al. 2018 Collection and habitat data. Habitat (outside of the study area). In various mixed, coniferous, and hardwood forest types, usually in decaying materials such as decaying fungi and carrion; from Lindgren funnel trap in red spruce forest with red maple and balsam fir, unbaited pitfall traps in red spruce forest, spruce forest in decaying fleshy fungi, mixed forests on Pleurotus sp. on Populus log, red oak and red maple forest in gilled mushrooms, alder swamp on remains of dead deer, hardwood forests in gilled fungi, and rich Appalachian hardwood forest in fleshy fungi in various stages of decay. Also in compost, poplar / spruce litter, carrion, bison and caribou dung, and under the bark of decaying spruce logs. Collecting period. V – IX. Collecting method. Sifting various kinds of organic materials, flight intercept traps, Lindgren funnel traps, unbaited and carrion-baited pitfall traps.	en	Klimaszewski, Jan, Godin, Benoit, Davies, Anthony, Bourdon, Caroline, Horwood, Denise (2018): Forty new records of aleocharine beetles, and two new species in the genera Acrotona Thomson and Atheta Thomson, for the province of Manitoba, Canada (Coleoptera: Staphylinidae: Aleocharinae). Insecta Mundi 2018 (641): 1-33, DOI: 10.5281/zenodo.5173990
A5636478EE00FFB4FF3DBB22FADB706A.taxon	distribution	Distribution Origin Nearctic Distribution Canada: AB, BC, LB, MB, NB, NF, NS, ON, QC, SK. USA: not recorded New provincial Canada: Manitoba: Adam Lake, 2016 - IX- 11, 49.0508 ° N, - 100.0666 ° W, 689 m, records mushrooms, sifting, B. Godin & D. Horwood (BGC) 1 male, 2 females; Birds Hill [Prov. Park] - bur oak [trail], 2016 - IX- 16, 50.0083 ° N, 96.9216 ° W, 264 m, mush- rooms, sifting, B. Godin & D. Horwood (LFC) 3 males, 3 females; Katherine Lake, 2016 - IX- 14, 50.6603 ° N, 99.8947 ° W, 652 m, mushrooms, sifting, B. Godin & D. Horwood (BGC) 1 male, 1 female. References Casey 1911, Klimaszewski et al. 2018 Collection and habitat data. Habitat (outside of the study area). In NB, in various forest types usually in dung or decaying mushrooms; eastern white cedar swamp in moose dung, mature red spruce and eastern white cedar forest in deer dung, mixed forest in bear dung, in decaying (moldy) corncobs and cornhusks near mixed forest, regenerating mixed forest in decaying mushrooms, near lakeshore in decaying fungi, old jack pine forest in rotten Boletus mushrooms, red maple swamp in leaf litter near stream, in unbaited pitfall traps in red spruce forest, and a few collected at u. v. light. Elsewhere, captured in flight intercept traps and carrion-baited pitfall traps in coniferous and mixed forests (NL) and on coastal barrens (LB); also from spruce litter, and dry and decaying mushrooms. Collecting period. VI – IX. Collecting method. Sifting litter, dung, and fungi, pitfall traps, flight intercept traps.	en	Klimaszewski, Jan, Godin, Benoit, Davies, Anthony, Bourdon, Caroline, Horwood, Denise (2018): Forty new records of aleocharine beetles, and two new species in the genera Acrotona Thomson and Atheta Thomson, for the province of Manitoba, Canada (Coleoptera: Staphylinidae: Aleocharinae). Insecta Mundi 2018 (641): 1-33, DOI: 10.5281/zenodo.5173990
A5636478EE01FFB4FF3DBF61FB93742D.taxon	distribution	Distribution Origin Nearctic Distribution Canada: BC, LB, MB, NB, NS, ON, QC, SK. USA: MA, NC, NH, NY, PA, RI, VT, WV New provincial Canada: Manitoba: Betula Lake, 2016 - IX- 20, 50.0667 ° N, 95.5887 ° W, 306 m, records mushrooms, sifting, B. Godin & D. Horwood (BGC) 4 males, 2 females; Katherine Lake, 2016 - IX- 14, 50.6603 ° N, 99.8947 ° W, 652 m, mushrooms, sifting, B. Godin & D. Horwood (BGC) 5 males, 2 females; Max Lake, 2016 - IX- 10, 49.0714 ° N, 100.1384 ° W, 686 m, mushrooms, sifting, B. Godin & D. Horwood (BGC) 1 male, 1 female; Birds Hill [Prov. Park] - bur oak [trail], 2016 - IX- 16, 50.0083 ° N, 96.9216 ° W, 264 m, mushrooms, sifting, B. Godin & D. Horwood (BGC) 4 males, 2 females; Hecla, 2016 - IX- 17, 51.7849 ° N, 96.6152 ° W, 218 m, mushroom, sifting, B. Godin & D. Horwood (LFC) 3 males, 3 females; Adam Lake, 2016 - IX- 11, 49.0508 ° N, 100.0666 W ˚, 689 m, mushrooms, sifting, B. Godin & D. Horwood (LFC) 7 males, 5 females; West Lake, 2016 - IX- 20, 49.7468 ° N, 95.2098 ° W, 341 m, mushrooms, sifting, B. Godin & D. Horwood (LFC) 1 male, 1 female; Moose Lake, 2016 - IX- 19, 49.2018 ° N, 95.3423 ° W, 353 m, mushroom, sifting, B. Godin & D. Horwood (LFC) 1 male, 1 female. References Bernhauer 1909, Klimaszewski et al. 2018 Collection and habitat data. Habitat (outside of the study area). In NB, in various coniferous, mixed, and hardwood forests in decaying fungi where it is often common; white pine and spruce forest in decaying mushrooms, old-growth eastern hemlock forest in decaying mushrooms, red spruce and eastern white cedar forest in decaying mushrooms, red pine forest in decaying mushrooms, jack pine forest in decaying mushrooms, mixed forests in decaying mushrooms, red oak forest in gilled mushrooms, rich Appalachian hardwood forest in decaying / rotten fleshy fungi; one from pitfall trap in red spruce forest and some in decaying (moldy) corncobs and cornhusks near a mixed forest. Elsewhere, from pitfall traps in open spruce forest with sandy soil and Cladina lichen cover (LB), birch-dominated forests (QC), in polypore fungus in coniferous forest, and on forest floor in red oak and deciduous forests (NS). Collecting period. VII – X. Collecting method. Sifting fungi and forest litter, pitfall traps.	en	Klimaszewski, Jan, Godin, Benoit, Davies, Anthony, Bourdon, Caroline, Horwood, Denise (2018): Forty new records of aleocharine beetles, and two new species in the genera Acrotona Thomson and Atheta Thomson, for the province of Manitoba, Canada (Coleoptera: Staphylinidae: Aleocharinae). Insecta Mundi 2018 (641): 1-33, DOI: 10.5281/zenodo.5173990
A5636478EE01FFABFF3DBBA6FC39736A.taxon	distribution	Distribution Origin Nearctic Distribution Canada: AB, BC, LB, MB, NB, NF, NS, ON, QC. USA: AK, CA, NC, NH, NY, OR, PA, RI, WA New provincial Canada: Manitoba: Adam Lake, 2016 - IX- 11, 49.0508 ° N, 100.0666 ° W, 689 m, records mushrooms, sifting, B. Godin & D. Horwood (BGC) 3 males; Hecla, 2016 - IX- 17, 51.7849 ° N, 96.6152 ° W, 218 m, mushroom, sifting, B. Godin & D. Horwood (BGC) 1 female; Georges Creek, 2016 - IX- 15, 50.6729 ° N, 99.6212 ° W, 696 m, mushrooms, sift- ing, B. Godin & D. Horwood (BGC) 1 male; Katherine Lake, 2016 - IX- 14, 50.6603 ° N, 99.8947 ° W, 652 m, mushrooms, sifting, B. Godin & D. Horwood (BGC) 1 male, 1 female; Birds Hill [Prov. Park] - bur oak [trail], 2016 - IX- 16, 50.0083 ° N, 96.9216 ° W, 264 m, mushrooms, sifting, B. Godin & D. Horwood (LFC) 4 males, 3 females. References Bernhauer 1909, Klimaszewski et al. 2018 Collection and habitat data. Habitat (outside of the study area). It occurs in various coniferous, mixed, and hardwood forests, usually in decaying organic materials such as rotting fungi, dung, and carrion; in NB, specimens from black spruce forest on gilled mushrooms, jack pine forest in decaying mushrooms, from pitfall traps in red spruce forest, old-growth hemlock forest in decaying bracket fungi (polypore), various mixed forests in gilled mushrooms and decaying fleshy and gilled mushrooms in various stages of decay, in decaying corncobs and cornhusks near mixed forest, alder swamp on remains of dead deer, rich Appalachian hardwood forest in fleshy fungi of various stages of decay, some from leaf litter and scat under great horned owl nest in hardwood forest; in NS, in compost, carrion, and mushrooms in coniferous and deciduous forests and open habitats. In NL, captured in carrion traps and flight intercept traps in mixed and coniferous forests. Collecting period. V-IX. Collecting method. sifting fungi and litter, unbaited and carrion-baited pitfall traps, flight intercept traps.	en	Klimaszewski, Jan, Godin, Benoit, Davies, Anthony, Bourdon, Caroline, Horwood, Denise (2018): Forty new records of aleocharine beetles, and two new species in the genera Acrotona Thomson and Atheta Thomson, for the province of Manitoba, Canada (Coleoptera: Staphylinidae: Aleocharinae). Insecta Mundi 2018 (641): 1-33, DOI: 10.5281/zenodo.5173990
A5636478EE1EFFABFF3DBC61FC39774D.taxon	distribution	Distribution Origin Nearctic Distribution Canada: AB, MB, NB, NF, QC. USA: ME, PA [all other previously published records (Klimaszewski et al. 2007 a, Dollin et al. 2008, Majka and Klimaszewski 2008 b, 2008 d, 2010, Bishop et al. 2009, Klimaszewski et al. 2011, Majka et al. 2011, Klimaszewski et al. 2012, 2013, 2015 c) of this species need to be reassessed] New provincial Canada: Manitoba: Adam Lake, 2016 - IX- 11, 49.0508 ° N, - 100.0666 ° W, 689 m, records mushrooms, sifting, B. Godin & D. Horwood (BGC) 2 males; same locality data (LFC) 1 female; Hecla, 2016 - IX- 17, 51.7849 ° N, 96.6152 ° W, 218 m, mushroom, sifting, B. Godin & D. Horwood (BGC) 6 males, 5 females; Birds Hill [Prov. Park] - bur oak [trail], 2016 - IX- 16, 50.0083 ° N, 96.9216 ° W, 264 m, mushrooms, sift- ing, B. Godin & D. Horwood (BGC) 1 male; Max Lake, 2016 - IX- 10, 49.0714 ° N, 100.1384 ° W, 686 m, mushrooms, sifting, B. Godin & D. Horwood (LFC) 5 males, 1 female; Bead Lake, 2016 - IX- 13, 50.7868 ° N, 99.9928 ° W, 620 m, mushrooms, sift- ing, B. Godin & D. Horwood (LFC) 2 males, 2 females; same locality data (BGC) 1 female; Katherine Lake, 2016 - IX- 14, 50.6603 ° N, 99.8947 ° W, 652 m, mushrooms, sifting, B. Godin & D. Horwood (LFC) 1 female. References Bernhauer 1909, Klimaszewski et al. 2018 Collection and habitat data. Habitat (outside of the study area). In NB, this species occurred in various forest types; hemlock forests, eastern white cedar swamps, red spruce and yellow birch forests, various mixed forests, a red oak forest, red pine forest, hardwood forests, and northern hardwood forests. Adults were found in various fungi, including fleshy polypore fungi, inside birch polypore (Piptoporus betulinus (Bull.) Karst.), gilled mushrooms, and decaying coral fungi. Some were found in compost (moldy corncobs and cornhusks, decaying vegetables) and inside a well rotted fungus-covered log. In NL, specimens from pitfall traps in boreal conifer forests. Collecting period. IV – X. Collecting method. Sifting fungi and litter, unbaited and carrion-baited pitfall traps, flight intercept traps.	en	Klimaszewski, Jan, Godin, Benoit, Davies, Anthony, Bourdon, Caroline, Horwood, Denise (2018): Forty new records of aleocharine beetles, and two new species in the genera Acrotona Thomson and Atheta Thomson, for the province of Manitoba, Canada (Coleoptera: Staphylinidae: Aleocharinae). Insecta Mundi 2018 (641): 1-33, DOI: 10.5281/zenodo.5173990
A5636478EE1FFFAAFF3DBEDDFAE475D9.taxon	distribution	Distribution Origin Nearctic Distribution Canada: AB, MB, NB, NS, ON, QC. USA: CT, DC, ME, MI, NY, PA, RI, VT, WV New provincial Canada: Manitoba: Betula Lake, 2016 - IX- 20, 50.0667 ° N, 95.5887 ° W, 306 m, records mushrooms, sifting, B. Godin & D. Horwood (BGC) 2 males; Georges Creek, 2016 - IX- 15, 50.6729 ° N, 99.6212 ° W, 696 m, mushrooms, sifting, B. Godin & D. Horwood (BGC) 1 male, 1 female; Katherine Lake, 2016 - IX- 14, 50.6603 ° N, 99.8947 ° W, 652 m, mushrooms, sifting, B. Godin & D. Horwood (BGC) 8 males, 11 females; Hecla, 2016 - IX- 17, 51.7849 ° N, 96.6152 ° W, 218 m, mushroom, sifting, B. Godin & D. Horwood (BGC) 4 females; same locality data (LFC) 3 males; Moose Lake, 2016 - IX- 19, 49.2018 ° N, 95.3423 ° W, 353 m, mushroom, sifting, B. Godin & D. Horwood (LFC) 1 male, 1 female; Max Lake, 2016 - IX- 10, 49.0714 ° N, 100.1384 ° W, 686 m, mushrooms, sifting, B. Godin & D. Horwood (LFC) 2 males, 1 female; Adam Lake, 2016 - IX- 11, 49.0508 ° N, 100.0666 ° W, 689 m, mushrooms, sifting, B. Godin & D. Horwood (LFC) 3 males, 4 females; Birds Hill [Prov. Park] - bur oak [trail], 2016 - IX- 16, 50.0083 ° N, 96.9216 ° W, 264 m, mushrooms, sifting, B. Godin & D. Horwood (LFC) 8 males, 7 females. References Melsheimer 1844, Klimaszewski et al. 2018 Collection and habitat data. Habitat (outside of the study area). In NB, found in various coniferous and hardwood forest types, usually in decaying fungi on forest floor; red oak forest in decaying gilled mushrooms, rich Appalachian hardwood forest in gilled mushrooms, and in entrance to mammal burrow in dung, various mixed forests in decaying gilled mushrooms and fleshy fungi, balsam fir forest in decaying Boletus mushrooms and gilled mushrooms, black spruce forest in Russula sp. mushrooms, mature red spruce and eastern white cedar forest in decaying mushrooms, and from unbaited pitfall traps in a red spruce forest (abundant); also from pitfall and flight intercept traps in yellow birch / balsam fir forest, mixed forest (Acer, Betula, Quercus, Ulmus, Tsuga, Pinus sp.), and a maple forest. Collecting period. V – IX. Collecting method. Sifting fungi and litter, pitfall traps, flight intercept traps.	en	Klimaszewski, Jan, Godin, Benoit, Davies, Anthony, Bourdon, Caroline, Horwood, Denise (2018): Forty new records of aleocharine beetles, and two new species in the genera Acrotona Thomson and Atheta Thomson, for the province of Manitoba, Canada (Coleoptera: Staphylinidae: Aleocharinae). Insecta Mundi 2018 (641): 1-33, DOI: 10.5281/zenodo.5173990
A5636478EE1FFFAAFF3DBAD2FA5A761C.taxon	distribution	Distribution Origin Nearctic Distribution Canada: LB, MB, NB, NF, NS, ON, QC. USA: IN, MA, MN, NY, PA, RI, VA, VT New provincial Canada: Manitoba: Birds Hill [Prov. Park] - bur oak [trail], 2016 - IX- 16, records 50.0083 ° N, 96.9216 ° W, 264 m, mushrooms, sifting, B. Godin & D. Horwood (BGC) 2 males, 1 female; same locality data (LFC) 2 males, 2 females; Hecla, 2016 - IX- 17, 51.7849 ° N, 96.6152 ° W, 218 m, mushroom, sifting, B. Godin & D. Horwood (BGC) 1 female. References Bernhauer, 1907, Klimaszewski et al. 2018 Collection and habitat data. Habitat (outside of the study area). This species is found in various conifer, mixed and hardwood / deciduous forests such as red spruce dominated forests, black spruce forests, balsam fir-spruce forests, jack pine forests, red pine forests, old-growth hemlock forests, riparian forests, red oak and maple forests, red oak forests, various mixed forests, northern hardwood forests, rich Appalachian hardwood forest, Populus stands, and yellow birch forests. Adults are often abundant in various kinds of decaying and fresh mushrooms (gilled, polypore; Polyporus varius on side of log, stalked polypores) on forest floor and also occur in forest litter, especially in the spring; often captured in abundance in pitfall traps. Collecting period. IV – X. Collecting method. Sifting forest litter, pitfall traps.	en	Klimaszewski, Jan, Godin, Benoit, Davies, Anthony, Bourdon, Caroline, Horwood, Denise (2018): Forty new records of aleocharine beetles, and two new species in the genera Acrotona Thomson and Atheta Thomson, for the province of Manitoba, Canada (Coleoptera: Staphylinidae: Aleocharinae). Insecta Mundi 2018 (641): 1-33, DOI: 10.5281/zenodo.5173990
A5636478EE1CFFA9FF3DBA46FA7D77A0.taxon	distribution	Distribution Origin Nearctic Distribution Canada: AB, BC, LB, MB, NB, NF, NS, ON, QC, SK, YT. USA: AK, NY New provincial Canada: Manitoba: Birds Hill [Prov. Park] - bur oak [trail], 2016 - IX- 16, records 50.0083 ° N, 96.9216 ° W, 264 m, mushrooms, sifting, B. Godin & D. Horwood (BGC) 1 male; Georges Creek, 2016 - IX- 15, 50.6729 ° N, 99.6212 ° W, 696 m, mushrooms, sifting, B. Godin & D. Horwood (BGC) 2 males, 1 female; Betula Lake, 2016 - IX- 20, 50.0667 ° N, 95.5887 ° W, 306 m, mushrooms, sifting, B. Godin & D. Horwood (BGC) 2 females; Adam Lake, 2016 - IX- 11, 49.0508 ° N, 100.0666 ° W, 689 m, mushrooms, sifting, B. Godin & D. Horwood (LFC) 1 male; Katherine Lake, 2016 - IX- 14, 50.6603 ° N, 99.8947 ° W, 652 m, mushrooms, sifting, B. Godin & D. Horwood (LFC) 2 males, 3 females. References Casey 1910, Klimaszewski et al. 2018 Collection and habitat data. Habitat (outside of the study area). In various forest types, usually in mushrooms; yellow birch / balsam fir forest, red spruce forest, old-growth hemlock forest, mixed forests, deciduous forests, and a red oak forest; most individuals in gilled mushrooms and fleshy fungi in various stages of decay. Also from leaf litter and moss in Carex marsh and forest litter. Collecting period. V – X. Collecting method. Sifting litter and fungi, pitfall traps, pitfall Luminoc traps, flight intercept traps.	en	Klimaszewski, Jan, Godin, Benoit, Davies, Anthony, Bourdon, Caroline, Horwood, Denise (2018): Forty new records of aleocharine beetles, and two new species in the genera Acrotona Thomson and Atheta Thomson, for the province of Manitoba, Canada (Coleoptera: Staphylinidae: Aleocharinae). Insecta Mundi 2018 (641): 1-33, DOI: 10.5281/zenodo.5173990
A5636478EE1CFFA9FF3DBED1FD61754E.taxon	distribution	Distribution Origin Nearctic Distribution Canada: AB, QC, MB, NB, NF, SK, YT. USA: [all published records of A. klagesi need to be re-examined because they may contain mixed series with A. pseudokla- gesi] New provincial Canada: Manitoba: Hecla, 2016 - IX- 17, 51.7849 ° N, 96.6152 ° W, 218 m, mushroom, records sifting, B. Godin & D. Horwood (BGC) 1 male; Georges Creek, 2016 - IX- 15, 50.6729 ° N, 99.6212 ° W, 696 m, mushrooms, sifting, B. Godin & D. Horwood (BGC) 1 male; Katherine Lake, 2016 - IX- 14, 50.6603 ° N, 99.8947 ° W, 652 m, mushrooms, sifting, B. Godin & D. Horwood (BGC) 1 male; Bead Lake, 2016 - IX- 13, 50.7868 ° N, 99.9928 ° W, 620 m, mushrooms, sifting, B. Godin & D. Horwood (LFC) 1 male, 1 female; Max Lake, 2016 - IX- 10, 49.0714 ° N, 100.1384 ° W, 686 m, mushrooms, sifting, B. Godin & D. Horwood (LFC) 1 male, 1 female. References Webster et al. 2016, Klimaszewski et al. 2018 Collection and habitat data. Habitat (outside of the study area). In NB, in mature mixed forest, old-growth and old white spruce and balsam fir forests, a mature red spruce forest, and in a wet alder swamp; adults in coral fungi on a Populus log, fleshy polypore fungi at base of a dead standing Populus, in decaying gilled mushrooms, in gilled mushrooms, and under bark of red spruce and dead aspen. In NF, from pitfall traps in boreal forests. In QC, in yellow birch / balsam fir forest, using pitfall traps, Luminoc pit / light traps, and intercept traps. In SK, in aspen woodland, in bracket / gilled fungi, under bark of dead aspen, and on fungus on dead lodgepole pine stump. In YT, in mushrooms. Collecting period. V – X. Collecting method. Sifting mushrooms.	en	Klimaszewski, Jan, Godin, Benoit, Davies, Anthony, Bourdon, Caroline, Horwood, Denise (2018): Forty new records of aleocharine beetles, and two new species in the genera Acrotona Thomson and Atheta Thomson, for the province of Manitoba, Canada (Coleoptera: Staphylinidae: Aleocharinae). Insecta Mundi 2018 (641): 1-33, DOI: 10.5281/zenodo.5173990
A5636478EE1DFFA8FF3DBD36FC3F7470.taxon	distribution	Distribution Origin Nearctic Distribution Canada: BC, LB, MB, NB, NF, ON, QC, SK, YT. USA: AK, NY New provincial record Canada: Manitoba: Katherine Lake, 2016 - IX- 14, 50.6603 ° N, 99.8947 ° W, 652 m, mushrooms, sifting, B. Godin & D. Horwood (BGC) 1 female; same locality data (LFC) 1 female. References Casey 1910, Klimaszewski et al. 2018 Collection and habitat data. Habitat (outside of the study area). It occurs in various forest types such as red spruce, mature red spruce and eastern white cedar, yellow birch and balsam fir, mixed, red oak, rich Appalachian hardwood, northern hardwood, and other deciduous forests. Adults usually found in decaying mushrooms (often common) on forest floor but also in leaf litter and possibly carrion. Collecting period. VI – X (most common in IX in NB). Collecting method. Sifting mushrooms and litter, carrion-baited and unbaited pitfall traps, flight intercept traps.	en	Klimaszewski, Jan, Godin, Benoit, Davies, Anthony, Bourdon, Caroline, Horwood, Denise (2018): Forty new records of aleocharine beetles, and two new species in the genera Acrotona Thomson and Atheta Thomson, for the province of Manitoba, Canada (Coleoptera: Staphylinidae: Aleocharinae). Insecta Mundi 2018 (641): 1-33, DOI: 10.5281/zenodo.5173990
A5636478EE1DFFAFFF3DBB6BFD18708A.taxon	distribution	Distribution Origin Nearctic Distribution Canada: LB, MB, NB, NF, ON, QC, SK, YT. USA: not recorded New provincial Canada: Manitoba: Hecla, 2016 - IX- 17, 51.7849 ° N, 96.6152 ° W, 218 m, mushroom, records sifting, B. Godin & D. Horwood (BGC) 3 males; same locality data (LFC) 4 males, 1 female; Bead Lake, 2016 - IX- 13, 50.7868 ° N, 99.9928 ° W, 620 m, mushrooms, sifting, B. Godin & D. Horwood (BGC) 1 male; same locality data (LFC) 1 male, 1 female. References Klimaszewski et al. 2011, 2018 Collection and habitat data. Habitat (outside of the study area). It occurs in many coniferous, mixed and hardwood / deciduous forest types in Canada. Adults commonly found in decaying mushrooms (polypore and gilled) and sometimes in fresh mushrooms. In NB, in an oak forest, rich Appalachian hardwood forest, northern hardwood forest, various mixed forests, red spruce forest, mature red spruce forest with red maple or yellow birch, black spruce forest, and eastern white cedar forests and swamps; specimens in fresh and decaying gilled mushrooms of various species, rotting lobster mushrooms, and a coral mushroom on spruce log. Also captured in unbaited and carrion-baited pitfall traps and flight intercept traps in various coniferous (spruce-feathermoss, spruce-lichen forests), mixed (spruce / aspen, riparian), and deciduous forests in NF and LB including some from decaying mushrooms; also from red maple, pine / willow, balsam fir / white birch forests, elsewhere. Collecting period. VI – X (most common in VIII and IX). Collecting method. Sifting litter and mushrooms, carrion-baited and unbaited pitfall traps, flight intercept traps, Lindgren funnel traps.	en	Klimaszewski, Jan, Godin, Benoit, Davies, Anthony, Bourdon, Caroline, Horwood, Denise (2018): Forty new records of aleocharine beetles, and two new species in the genera Acrotona Thomson and Atheta Thomson, for the province of Manitoba, Canada (Coleoptera: Staphylinidae: Aleocharinae). Insecta Mundi 2018 (641): 1-33, DOI: 10.5281/zenodo.5173990
A5636478EE1DFFA8FF3DBED1FD8773BD.taxon	distribution	Distribution Origin Nearctic Distribution Canada: MB, SK, YT. USA: not recorded New provincial Canada: Manitoba: Bead Lake, 2016 - IX- 13, 50.7868 ° N, 99.9928 ° W, 620 m, records mushrooms, sifting, B. Godin & D. Horwood (LFC) 2 males; Hecla, 2016 - IX- 17, 51.7849 ° N, 96.6152 ° W, 218 m, mushroom, sifting, B. Godin & D. Horwood (LFC) 1 male; Max Lake, 2016 - IX- 10, 49.0714 ° N, 100.1384 ° W, 686 m, mushrooms, sifting, B. Godin & D. Horwood (BGC) 2 males. References Klimaszewski et al. 2012, 2016 Collection and habitat data. Habitat (outside of the study area). In various forest types, usually in mushrooms; red spruce forest, old-growth hemlock forest, mixed forests, deciduous forests, and a red oak forest; most individuals were found in gilled mushrooms and fleshy fungi in various stages of decay. Also from leaf litter and moss in Carex marsh and forest litter. Collecting period. VI – IX. Collecting method. Sifting litter and fungi, pitfall traps.	en	Klimaszewski, Jan, Godin, Benoit, Davies, Anthony, Bourdon, Caroline, Horwood, Denise (2018): Forty new records of aleocharine beetles, and two new species in the genera Acrotona Thomson and Atheta Thomson, for the province of Manitoba, Canada (Coleoptera: Staphylinidae: Aleocharinae). Insecta Mundi 2018 (641): 1-33, DOI: 10.5281/zenodo.5173990
A5636478EE1AFFAFFF3DBC00FCA775CE.taxon	distribution	Distribution Origin Nearctic Distribution Canada: AB, BC, LB, MB, NB, NF, NS, ON, QC, SK, YT. USA: AK, CA, NV, OR New provincial Canada: Manitoba: Cyprus Creek, 2016 - IX- 15, 49.5774 ° N, 101.3645 ° W, 510 m, records Willow litter, sifting, B. Godin & D. Horwood (BGC) 1 male, 2 females; Souris River, 2016 - IX- 9, 49.5022 ° N, 99.9915 ° W, 402 m, river bank, oak litter, sifting, B. Godin & D. Horwood (BGC) 2 males, 1 female; Souris River, 2016 - IX- 9, 49.5022 ° N, 99.9915 ° W, 402 m, river bank, oak litter, sifting, B. Godin & D. Horwood (LFC) 3 males, 3 females. References Mäklin 1852, Klimaszewski et al. 2011, 2018 Collection and habitat data. Habitat (outside of the study area). In NF, frequently captured in pitfall traps in various forest types such as birch, spruce-lichen, spruce-poplar, fir; also in vegetation on coastal dunes, in disturbed fields among grass and weeds, and on shrubby limestone barrens. In QC, from pitfall traps in yellow birch / balsam fir forests, and sifted from litter in an oak savanna / alvar in ON. One individual from sphagnum and litter in an eastern white cedar swamp and another captured in Lindgren funnel trap in an old red pine forest in NB. Collecting period. IV – IX. Collecting method. Pitfall traps, Lindgren funnel trap, sifting forest litter.	en	Klimaszewski, Jan, Godin, Benoit, Davies, Anthony, Bourdon, Caroline, Horwood, Denise (2018): Forty new records of aleocharine beetles, and two new species in the genera Acrotona Thomson and Atheta Thomson, for the province of Manitoba, Canada (Coleoptera: Staphylinidae: Aleocharinae). Insecta Mundi 2018 (641): 1-33, DOI: 10.5281/zenodo.5173990
A5636478EE1AFFAFFF3DBAC5FA867641.taxon	distribution	Distribution Origin Nearctic Distribution Canada: MB, ON, QC, SK. USA: IA, MN New provincial Canada: Manitoba: Souris River, 2016 - IX- 9, 49.5022 ° N, 99.9915 ° W, 402 m, river records bank, oak litter, sifting, B. Godin & D. Horwood (BGC) 1 female; Cyprus Creek, 2016 - IX- 15, 49.5774 ° N, 101.3645 ° W, 510 m, willow litter, sifting, B. Godin & D. Horwood (BGC) 1 male; Birds Hill [Prov. Park] - bur oak [trail], 2016 - IX- 16, 50.0083 ° N, 96.9216 ° W, 264 m, mushrooms, sifting, B. Godin & D. Horwood (BGC) 5 males, 6 females; same locality data (LFC) 5 males, 4 females; Moose Lake, 2016 - IX- 19, 49.2018 ° N, 95.3423 ° W, 353 m, mushroom, sifting, B. Godin & D. Horwood (LFC) 2 males, 2 females. References Casey 1893, Klimaszewski et al. 2015 b, 2018 Collection and habitat data. Habitat (outside of the study area). Associated with hardwood / deciduous forests; adults in forest litter, deciduous leaf mold, in maple / oak forest litter and other unspecified deciduous forest litter. Collecting period. IV – X. Collecting method. Pitfall traps, sifting forest litter.	en	Klimaszewski, Jan, Godin, Benoit, Davies, Anthony, Bourdon, Caroline, Horwood, Denise (2018): Forty new records of aleocharine beetles, and two new species in the genera Acrotona Thomson and Atheta Thomson, for the province of Manitoba, Canada (Coleoptera: Staphylinidae: Aleocharinae). Insecta Mundi 2018 (641): 1-33, DOI: 10.5281/zenodo.5173990
A5636478EE1BFFAEFF3DBED0FC5A72FE.taxon	distribution	Distribution Origin Palearctic, adventive in North America Distribution Canada: AB, BC, LB, MB, NB, NF, NS, NU, ON, PE, QC, SK, YT. USA: AK, CA, ME, MA, MN, NH, NY, OR, RI, WA New provincial Canada: Manitoba: Birds Hill [Prov. Park] - bur oak [trail], 2016 - IX- 16, records 50.0083 ° N, 96.9216 ° W, 264 m, mushrooms, sifting, B. Godin & D. Horwood (BGC) 1 female; Cyprus Creek, 2016 - IX- 15, 49.5774 ° N, 101.3645 ° W, 510 m, willow litter, sifting, B. Godin & D. Horwood (BGC) 3 females; Souris River, 2016 - IX- 9, 49.5022 ° N, 99.9915 ° W, 402 m, river bank, oak litter, sifting, B. Godin & D. Horwood (LFC) 5 females; Hecla, 2016 - IX- 17, 51.7849 ° N, 96.6152 ° W, 218 m, mushroom, sifting, B. Godin & D. Horwood (LFC) 1 female. References Gravenhorst 1806, Klimaszewski et al. 2015 b, 2018 Collection and habitat data. Habitat (outside of the study area). Found in various forest types and wetlands, usually in litter. In NB, in silver maple swamps in leaf litter, in leaf litter in eastern white cedar swamps and forests, under alders in leaf litter and moss in a marsh, treading vegetation in boggy marsh and a cattail marsh, river margin on moist clay soil among tall grasses, a sea beach in decaying sea wrack, in pitfall traps in a red spruce forest. In NF, from pitfall traps in cut and burned balsam fir, birch, spruce-poplar and riparian forests, in agricultural fields and among vegetation on coastal sand dunes. Elsewhere, in mixed forests and yellow birch forests. Collecting period. V – IX. Collecting method. Pitfall traps, sifting forest litter, treading marsh vegetation.	en	Klimaszewski, Jan, Godin, Benoit, Davies, Anthony, Bourdon, Caroline, Horwood, Denise (2018): Forty new records of aleocharine beetles, and two new species in the genera Acrotona Thomson and Atheta Thomson, for the province of Manitoba, Canada (Coleoptera: Staphylinidae: Aleocharinae). Insecta Mundi 2018 (641): 1-33, DOI: 10.5281/zenodo.5173990
A5636478EE1BFFAEFF3DBDF5FBE17730.taxon	distribution	Distribution Origin Nearctic Distribution Canada: MB, NB, NF, ON, QC. USA: DC, IA, IN, MA, MI, MN, MO, MS, NC, NY, PA, RI, WI New provincial Canada: Manitoba: Cyprus Creek, 2016 - IX- 15, 49.5774 ° N, 101.3645 ° W, 510 records m, willow litter, sifting, B. Godin & D. Horwood (BGC) 1 female; same locality data (LFC) 1 female; Bead Lake, 2016 - IX- 13, 50.7868 ° N, 99.9928 ° W, 620 m, mushrooms, sifting, B. Godin & D. Horwood (LFC) 1 male. References Erichson 1839, Klimaszewski et al. 2015 b, 2018 Collection and habitat data. Habitat (outside of the study area). Most specimens from QC were captured in pitfall traps in yellow birch and balsam fir dominated forests. In NB, under decaying seaweed on sea beach, under drift material on a riverbank, in grass, moss, and leaf litter in alder swamps, eastern white cedar swamps, and Carex marshes; in sphagnum moss and leaf litter in a young regenerating mixed forest, and in decaying (moldy) corncobs and cornhusks. Elsewhere, adults in litter around raspberry bushes near a bog, in a Typha marsh in a nest of Microtus pennsylvanicus (ON), on a lake shore and in a Microtus pennsylvanicus nest (MN), and sifting vegetable debris in a dump (IN). Collecting period. III – X. Collecting method. Sifting forest and marsh litter, pitfall traps.	en	Klimaszewski, Jan, Godin, Benoit, Davies, Anthony, Bourdon, Caroline, Horwood, Denise (2018): Forty new records of aleocharine beetles, and two new species in the genera Acrotona Thomson and Atheta Thomson, for the province of Manitoba, Canada (Coleoptera: Staphylinidae: Aleocharinae). Insecta Mundi 2018 (641): 1-33, DOI: 10.5281/zenodo.5173990
A5636478EE18FFADFF3DBED1FADB727D.taxon	distribution	Distribution Origin Nearctic Distribution Canada: MB, NB, NF, ON, SK, YT. USA: not recorded New provincial Canada: Manitoba: Adam Lake, 2016 - IX- 11, 49.0508 ° N, 100.0666 ° W, 689 m, records mushrooms, sifting, B. Godin & D. Horwood (LFC) 1 male. References Klimaszewski et al. 2011, 2018 Collection and habitat data. Habitat (outside of the study area). It occurs in various riparian and wetland habitats; adults sifted from moist leaves on vernal pond margins in mixed forests, hardwood forests, and a silver maple swamp, sifted from sedges and leaves on pond margin, treaded from Carex and grasses on lake margin, treaded from vegetation in seasonally flooded marsh, sifted from moss and leaves in seepage area in hardwood forest and debris on muddy soil near brook, from drift material on river margin, and sifted from leaf litter and moss under alders near brook. Also hand collected from cobblestones on partially shaded cobblestone bar along medium-sized stream and from moss near splash zone of waterfalls by splashing. In ON, one specimen from pitfall in hedgerow and flight intercept traps in a mixed forest in NF. Collecting period. V – VII, IX – X. Collecting method. Sifting mushrooms (new data), sifting litter, treading vegetation, splashing moss, flight intercept traps, pitfall traps.	en	Klimaszewski, Jan, Godin, Benoit, Davies, Anthony, Bourdon, Caroline, Horwood, Denise (2018): Forty new records of aleocharine beetles, and two new species in the genera Acrotona Thomson and Atheta Thomson, for the province of Manitoba, Canada (Coleoptera: Staphylinidae: Aleocharinae). Insecta Mundi 2018 (641): 1-33, DOI: 10.5281/zenodo.5173990
A5636478EE18FFADFF3DBD76FE2E7710.taxon	distribution	Distribution Origin Nearctic Distribution Canada: AB, BC, MB, NB, NF, NS, ON, QC, SK. USA: AZ, CO, ID, MI, MN, MT, NH, NM, SD, WI. MEXICO. GUATEMALA. HONDURAS New provincial Canada: Manitoba: Hecla, 2016 - IX- 17, 51.7849 ° N, 96.6152 ° W, 218 m, mushroom, records sifting, B. Godin & D. Horwood (BGC) 1 female; Bead Lake, 2016 - IX- 13, 50.7868 ° N, 99.9928 ° W, 620 m, mushrooms, sifting, B. Godin & D. Horwood (BGC) 2 males, 2 females; Katherine Lake, 2016 - IX- 14, 50.6603 ° N, 99.8947 ° W, 652 m, mushrooms, sifting, B. Godin & D. Horwood (LFC) 2 males, 2 females; Max Lake, 2016 - IX- 10, 49.0714 ° N, 100.1384 ° W, 686 m, mushrooms, sifting, B. Godin & D. Horwood (LFC) 1 male. References Bernhauer 1907, Gusarov 2003, Ashe 1986 [as S. bispinosa], Klimaszewski et al. 2011, 2018 Collection and habitat data. Habitat (outside of the study area). It was recorded from old-growth northern hardwood forest, old-growth white spruce and balsam fir forest, fir and riparian forests, mountain forests, oak forest, yellow birch dominated forest, balsam fir forest, and meadow; specimens from moose dung, soil and moss under logs, leaf litter near a body of water, river debris, humus under maple, Populus litter at the edge of spruce-cedar forest, pine stump buttress, litter near the sea. Collecting period. III – X. Collecting method. Sifting mushrooms (new data), sifting forest litter, pitfall traps, Lindgren funnel traps.	en	Klimaszewski, Jan, Godin, Benoit, Davies, Anthony, Bourdon, Caroline, Horwood, Denise (2018): Forty new records of aleocharine beetles, and two new species in the genera Acrotona Thomson and Atheta Thomson, for the province of Manitoba, Canada (Coleoptera: Staphylinidae: Aleocharinae). Insecta Mundi 2018 (641): 1-33, DOI: 10.5281/zenodo.5173990
A5636478EE19FFACFF3DBF2DFA837389.taxon	distribution	Distribution Origin Nearctic Distribution Canada: MB, NB, ON, QC. USA: MI, NC, NY, PA New provincial Canada: Manitoba: Bead Lake, 2016 - IX- 13, 50.7868 ° N, 99.9928 ° W, 620 m, record mushrooms, sifting, B. Godin & D. Horwood (BGC) 4 males; same locality data (LFC) 3 males. References Seevers 1951, Webster et al 2012, Brunke et al. 2012, Klimaszewski et al. 2018 Collection and habitat data. Habitat (outside of the study area). Found in hardwood forest, a mixed forest, black spruce, balsam fir, and eastern white cedar forests. Adults in fresh, gilled mushrooms and Pleurotus sp. on the side of a log. Collecting period. VIII-IX. Collecting method. Sifting mushrooms.	en	Klimaszewski, Jan, Godin, Benoit, Davies, Anthony, Bourdon, Caroline, Horwood, Denise (2018): Forty new records of aleocharine beetles, and two new species in the genera Acrotona Thomson and Atheta Thomson, for the province of Manitoba, Canada (Coleoptera: Staphylinidae: Aleocharinae). Insecta Mundi 2018 (641): 1-33, DOI: 10.5281/zenodo.5173990
A5636478EE19FFACFF3DBD02FE4A746C.taxon	distribution	Distribution Origin Nearctic Distribution Canada: MB, NB, NS, ON, QC. USA: CT, DC, IL, IN, KY, MA, MD, ME, MI, MO, NC, NH, NJ, NY, PA, RI, TN, VA, VT, WI, WV New provincial Canada: Manitoba: Moose Lake, 2016 - IX- 19, 49.2018 ° N, 95.3423 ° W, 353 m, record mushroom, sifting, B. Godin & D. Horwood (BGC) 1 male. References Melsheimer 1844, Seevers 1951, Klimaszewski et al. 2018 Collection and habitat data. Habitat (outside of the study area). In mixed forest, hardwood forest, rich Appalachian hardwood forest, red oak forest, red oak and red maple forest, 8.5 - year-old regenerating mixed forest, mature red spruce and red maple forest. Adults on / in gilled mushrooms on forest floor, in gilled mushroom on stump, on Pleurotus species growing on a log, on polypore (bracket) fungus on side of log, in Porodaedalea piceina (Peck) Niemalä on dead standing beech tree, on Russula delica, in decaying mushrooms, and on white fungus on white birch. Collecting period. VI – IX. Collecting method. Sifting mushrooms.	en	Klimaszewski, Jan, Godin, Benoit, Davies, Anthony, Bourdon, Caroline, Horwood, Denise (2018): Forty new records of aleocharine beetles, and two new species in the genera Acrotona Thomson and Atheta Thomson, for the province of Manitoba, Canada (Coleoptera: Staphylinidae: Aleocharinae). Insecta Mundi 2018 (641): 1-33, DOI: 10.5281/zenodo.5173990
A5636478EE19FFACFF3DBB67FEAC7667.taxon	distribution	Distribution Origin Nearctic Distribution Canada: AB, MB, ON, PE. USA: IL, IN, MA, MI, PA, WI New provincial Canada: Manitoba: Adam Lake, 2016 - IX- 11, 49.0508 ° N, 100.0666 ° W, 689 m, records mushrooms, sifting, B. Godin & D. Horwood (BGC) 1 male; Hecla, 2016 - IX- 17, 51.7849 ° N, 96.6152 ° W, 218 m, mushroom, sifting, B. Godin & D. Horwood (BGC) 1 male; Katherine Lake, 2016 - IX- 14, 50.6603 ° N, 99.8947 ° W, 652 m, mushrooms, sifting, B. Godin & D. Horwood (LFC) 3 males. References Casey 1906, Seevers 1951, Webster et al. 2012, Klimaszewski et al. 2018 Collection and habitat data. Habitat. Mushrooms. Collecting period. V-IX. Collecting method. Sifting mushrooms.	en	Klimaszewski, Jan, Godin, Benoit, Davies, Anthony, Bourdon, Caroline, Horwood, Denise (2018): Forty new records of aleocharine beetles, and two new species in the genera Acrotona Thomson and Atheta Thomson, for the province of Manitoba, Canada (Coleoptera: Staphylinidae: Aleocharinae). Insecta Mundi 2018 (641): 1-33, DOI: 10.5281/zenodo.5173990
A5636478EE16FFA3FF3DBED1FC1D733E.taxon	distribution	Distribution Origin Nearctic Distribution Canada: MB, NB, ON. USA: DC, IL, IN, KS, MI, NC, NY, PA, VA, WV New provincial Canada: Manitoba: Moose Lake, 2016 - IX- 19, 49.2018 ° N, 95.3423 ° W, 353 m, record mushroom, sifting, B. Godin & D. Horwood (BGC) 1 male. References Casey 1906, Seevers 1951, Klimaszewski et al. 2018 Collection and habitat data. Habitat (outside of the study area). Mixed forest, hardwood forest, on ridge with red oak surrounded by a silver maple forest, red oak forest, oak and red maple forest, and spruce forest. Adults on / in fresh gilled mushrooms on forest floor, a few in decaying fleshy mushrooms. Collecting period. VIII – IX. Collecting method. Sifting mushrooms.	en	Klimaszewski, Jan, Godin, Benoit, Davies, Anthony, Bourdon, Caroline, Horwood, Denise (2018): Forty new records of aleocharine beetles, and two new species in the genera Acrotona Thomson and Atheta Thomson, for the province of Manitoba, Canada (Coleoptera: Staphylinidae: Aleocharinae). Insecta Mundi 2018 (641): 1-33, DOI: 10.5281/zenodo.5173990
A5636478EE16FFA3FF3DBCB6FB7D75B0.taxon	distribution	Distribution Origin Nearctic Distribution Canada: AB, BC, LB, MB, NB, NF, NS, ON, QC, YT. USA: FL, MA, ME, MT, NH, NY, OR, TN, WA, WY New provincial Canada: Manitoba: Bead Lake, 2016 - IX- 13, 50.7868 ° N, 99.9928 ° W, 620 m, record mushrooms, sifting, B. Godin & D. Horwood (LFC) 1 male; same locality data (BGC) 1 male. References Casey 1911, Seevers 1951, Klimaszewski et al. 2018 Collection and habitat data. Habitat (outside of the study area). Mature mixed forest, 8.5 - year-old regenerating mixed forest, red oak forest, eastern white cedar (Thuja occidentalis L.) swamps, red spruce (Picea rubens Sarg.) and red maple (Acer rubrum L.) forest (80 – 120 years old), black spruce Populus forests, recently burned coniferous forest, and an eastern hemlock forest [Tsuga canadensis (L.)] 120 + years old. Adults in in gilled mushrooms, in gilled fungi on rotten log, in gilled mushrooms on stump, in moss near brook, on polypore fungi on dead standing Populus sp. Collecting period. VI – IX. Collecting method. S ifting mushrooms, sifting moss, flight intercept trap, and pitfall trap.	en	Klimaszewski, Jan, Godin, Benoit, Davies, Anthony, Bourdon, Caroline, Horwood, Denise (2018): Forty new records of aleocharine beetles, and two new species in the genera Acrotona Thomson and Atheta Thomson, for the province of Manitoba, Canada (Coleoptera: Staphylinidae: Aleocharinae). Insecta Mundi 2018 (641): 1-33, DOI: 10.5281/zenodo.5173990
A5636478EE16FFA3FF3DBB2BFB957674.taxon	distribution	Distribution Origin Nearctic Distribution Canada: MB, NB, NF. USA: DC, IL, IN, KY, MA, NY, PA, TN, VA, WI New provincial Canada: Manitoba: Birds Hill [Prov. Park] bur oak [trail], 2016 - IX- 16, 50.0083 ° N, record 96.9216 ° W, 264 m, mushrooms, sifting, B. Godin & D. Horwood (BGC) 1 male; Max Lake, 2016 - IX- 10, 49.0714 ° N, 100.1384 ° W, 686 m, mushrooms, sifting, B. Godin & D. Horwood (LFC) 1 male. References Casey 1906, Seevers 1951, Klimaszewski et al. 2018 Collection and habitat data. Habitat (outside of the study area). Mixed forests (mature to old), regenerating mixed forest, red oak forest, red spruce and yellow birch forest, mature red spruce and red maple forest, old jack pine forest, and forested black spruce bog with red maple. Adults on / in fresh gilled mushrooms, on Pleurotus sp. on dead standing Populus tremuloides Michx, and in fresh Boletus mushrooms. Collecting period. VI – IX. Collecting method. S ifting mushrooms.	en	Klimaszewski, Jan, Godin, Benoit, Davies, Anthony, Bourdon, Caroline, Horwood, Denise (2018): Forty new records of aleocharine beetles, and two new species in the genera Acrotona Thomson and Atheta Thomson, for the province of Manitoba, Canada (Coleoptera: Staphylinidae: Aleocharinae). Insecta Mundi 2018 (641): 1-33, DOI: 10.5281/zenodo.5173990
A5636478EE17FFA2FF3DBEDDFC007559.taxon	distribution	Distribution Origin Nearctic Distribution Canada: AB, MB, NB, NF, NS, ON. USA: IL, IN, MI, MN, NH, NY, WI New provincial Canada: Manitoba: Hecla, 2016 - IX- 17, 51.7849 ° N, 96.6152 ° W, 218 m, mushroom, records sifting, B. Godin & D. Horwood (LFC) 2 males; same locality data (BGC) 2 males; Adam Lake, 2016 - IX- 11, 49.0508 ° N, 100.0666 ° W, 689 m, mushrooms, sifting, B. Godin & D. Horwood (LFC) 1 male; Bead Lake, 2016 - IX- 13, 50.7868 ° N, 99.9928 ° W, 620 m, mushrooms, sifting, B. Godin & D. Horwood (BGC) 1 male; Katherine Lake, 2016 - IX- 14, 50.6603 ° N, 99.8947 ° W, 652 m, mushrooms, sifting, B. Godin & D. Horwood (LFC) 4 males; same locality data (BGC) 3 males; Birds Hill [Prov. Park] - bur oak [trail], 2016 - IX- 16, 50.0083 ° N, 96.9216 ° W, 264 m, mushrooms, sifting, B. Godin & D. Horwood (BGC) 1 female; same locality data (LFC) 1 female; West Lake, 2016 - IX- 20, 49.7468 ° N, 95.2098 ° W, 341 m, mushrooms, sifting, B. Godin & D. Horwood (BGC) 1 female; Max Lake, 2016 - IX- 10, 49.0714 ° N, 100.1384 ° W, 686 m, mushrooms, sifting, B. Godin & D. Horwood (BGC) 1 male. References Casey 1906, Seevers 1951, Klimaszewski et al. 2018 Collection and habitat data. Habitat (outside of the study area). Mixed forests, hardwood forests, old-growth northern hardwood forest, red spruce forest, red spruce and yellow birch forest, mature red spruce and red maple forest, mature red spruce and eastern white cedar forest, red oak and red maple forest. Adults on / in fresh gilled mushrooms on forest floor, in decaying gilled mushroom (one occurrence), and on Pleurotus sp. on log. This species has also been collected in association with grasses. Collecting period. VIII – X. Collecting method. Sifting mushrooms, pitfall traps.	en	Klimaszewski, Jan, Godin, Benoit, Davies, Anthony, Bourdon, Caroline, Horwood, Denise (2018): Forty new records of aleocharine beetles, and two new species in the genera Acrotona Thomson and Atheta Thomson, for the province of Manitoba, Canada (Coleoptera: Staphylinidae: Aleocharinae). Insecta Mundi 2018 (641): 1-33, DOI: 10.5281/zenodo.5173990
A5636478EE17FFA2FF3DBA52FC2877FC.taxon	distribution	Distribution Origin Nearctic Distribution Canada: MB, NB, NF, ON, QC. USA: DC, IA, IN, KS, MA, NC, TN, VA, WI New provincial Canada: Manitoba: Bead Lake, 2016 - IX- 13, 50.7868 ° N, 99.9928 ° W, 620 m, records mushrooms, sifting, B. Godin & D. Horwood (BGC) 1 male; Birds Hill [Prov. Park] - bur oak [trail], 2016 - IX- 16, 50.0083 ° N, 96.9216 ° W, 264 m, mushrooms, sifting, B. Godin & D. Horwood (LFC) 1 male. References Seevers 1951, Klimaszewski et al. 2011, Webster et al. 2012, Klimaszewski et al. 2018 Collection and habitat data. Habitat (outside of the study area). In various deciduous and coniferous forest types, including hardwood forests with sugar maple, American beech, and white ash (Fraxinus americana L.), an old red oak forest, a mixed forest with eastern hemlock, mixed forests, red spruce forests, a black spruce forest, and a black spruce, balsam fir and eastern white cedar forest. Most adults in NB were collected from fresh (not decaying) gilled mushrooms, including a Russula sp. Some specimens were collected from a stalked polypore fungus on forest floor, a Boletus sp., and a Pleurotus sp. on a log. Collecting period. VII – X. Collecting method. S ifting mushrooms.	en	Klimaszewski, Jan, Godin, Benoit, Davies, Anthony, Bourdon, Caroline, Horwood, Denise (2018): Forty new records of aleocharine beetles, and two new species in the genera Acrotona Thomson and Atheta Thomson, for the province of Manitoba, Canada (Coleoptera: Staphylinidae: Aleocharinae). Insecta Mundi 2018 (641): 1-33, DOI: 10.5281/zenodo.5173990
A5636478EE14FFA1FF3DBED1FB0F73FE.taxon	distribution	Distribution Origin Nearctic Distribution Canada: MB, NB, ON, PE, QC. USA: AL, IL, IN, KY, ME, MI, NC, NH, NY, PA, TN, WV, WI New provincial Canada: Manitoba: Hecla, 2016 - IX- 17, 51.7849 ° N, 96.6152 ° W, 218 m, mushroom, record sifting, B. Godin & D. Horwood (BGC) 1 male; same locality data (LFC) 1 male. References Casey 1906, Seevers 1951, Klimaszewski et al. 2018 Collection and habitat data. Habitat (outside of the study area). In mixed forest, hardwood forest, red oak forest, rich Appalachian hardwood forest, flood plain forest, eastern white cedar swamps, black spruce forest with Populus sp., and mature red spruce and red maple forest. Adults on / in fresh, gilled mushrooms on forest floor, on small gilled mushrooms on side of log, on bracket fungi, and on Trametes hirsuta on a poplar log. Collecting period. VI – IX. Collecting method. Sifting mushrooms.	en	Klimaszewski, Jan, Godin, Benoit, Davies, Anthony, Bourdon, Caroline, Horwood, Denise (2018): Forty new records of aleocharine beetles, and two new species in the genera Acrotona Thomson and Atheta Thomson, for the province of Manitoba, Canada (Coleoptera: Staphylinidae: Aleocharinae). Insecta Mundi 2018 (641): 1-33, DOI: 10.5281/zenodo.5173990
A5636478EE14FFA1FF3DBCF6FC5F7431.taxon	distribution	Distribution Origin Nearctic Distribution Canada: MB, AB, BC, LB, NB, NF, NS, ON, PE, QC, SK, YT. USA: AK, CA, MN New provincial Canada: Manitoba: Katherine Lake, 2016 - IX- 14, 50.6603 ° N, 99.8947 ° W, 652 m, record mushrooms, sifting, B. Godin & D. Horwood (LFC) 1 female. References Bernhauer 1905, Klimaszewski et al. 2018 Collection and habitat data. Habitat (outside of the study area). In many hardwood and conifer forest types; unspecified hardwood forest types, rich Appalachian hardwood forest, red oak forest, old mixed forest with red oak, mixed forests, northern hardwood forest, coastal red spruce and birch forest, mature red spruce forest with eastern white cedar, old red spruce forest, spruce forest, white spruce forest, yellow birch / balsam fir forest, trembling aspen forest with a small amount of eastern balsam poplar, white birch forest, white spruce, and willow forests. In NB, adults common in decaying mushrooms in late summer; in fleshy fungi, in decaying gilled mushrooms, baited with pile of decaying mushrooms, fleshy fungi in various stages of decay, in gilled mushrooms, in rotten fungi. Elsewhere, in wet moss on forest floor, marten dung on moss, and in mushrooms. Collecting period. VII – X. Collecting method. Sifting mushrooms, baiting with mushrooms, pitfall traps, Luminoc pit-light traps, malaise traps, flight intercept traps, carrion baited traps, and sifting forest litter.	en	Klimaszewski, Jan, Godin, Benoit, Davies, Anthony, Bourdon, Caroline, Horwood, Denise (2018): Forty new records of aleocharine beetles, and two new species in the genera Acrotona Thomson and Atheta Thomson, for the province of Manitoba, Canada (Coleoptera: Staphylinidae: Aleocharinae). Insecta Mundi 2018 (641): 1-33, DOI: 10.5281/zenodo.5173990
A5636478EE14FFA0FF3DBBFBFCD070B6.taxon	distribution	Distribution Origin Nearctic Distribution Canada: AB, LB, MB, NB, NF, NS, ON, QC, SK. USA: AL, FL, IA, IL, MA, ME, NH, NJ, TN, WV. MEXICO New provincial Canada: Manitoba: Adam Lake, 2016 - IX- 11, 49.0508 ° N, 100.0666 ° W, 689 m, records mushrooms, sifting, B. Godin & D. Horwood (BGC) 1 male, 1 female; same locality data (LFC) 1 female; Max Lake, 2016 - IX- 10, 49.0714 ° N, 100.1384 ° W, 686 m, mushrooms, sifting, B. Godin & D. Horwood (LFC) 1 female. References Casey 1911, Klimaszewski et al. 2018 Collection and habitat data. Habitat (outside of the study area). Lives in debris in various riparian and wetland habitats; in leaf litter in areas with Carex on brook margin, river margin among cobblestones near outflow of brook, margin of large brook under alders among gravel on gravel bar, in leaf litter near seepage and brook in hardwood forest, in moist leaves on vernal pond margin in mixed forest, lake margins in wet leaf litter, inland margin of salt marsh in leaf litter, in Carex hummocks in marsh near small stream, in sphagnum hummocks on margin of tamarack bog, one from sifted from dead grass on pond margin, one from beaver lodge, some collected near streams, and in beaver lodge debris. Collecting period. II – XI (mainly V – VIII). Collecting method. sifting leaf litter, mushrooms (new data), turning gravel and cobblestones and aspirating specimens.	en	Klimaszewski, Jan, Godin, Benoit, Davies, Anthony, Bourdon, Caroline, Horwood, Denise (2018): Forty new records of aleocharine beetles, and two new species in the genera Acrotona Thomson and Atheta Thomson, for the province of Manitoba, Canada (Coleoptera: Staphylinidae: Aleocharinae). Insecta Mundi 2018 (641): 1-33, DOI: 10.5281/zenodo.5173990
A5636478EE15FFA0FF3DBC7DFD9075F9.taxon	distribution	Distribution Origin Nearctic Distribution Canada: AB, LB, MB, NB, NF, NS, ON, QC. USA: IA, MO, NY New provincial Canada: Manitoba: Moose Lake, 2016 - IX- 19, 49.2018 ° N, 95.3423 ° W, 353 m, record mushroom, sifting, B. Godin & D. Horwood (BGC) 1 female. References Casey 1893, Klimaszewski et al. 2011, 2018 Collection and habitat data. Habitat (outside of the study area). In various forest types including white spruce, mature black spruce, red spruce, mature 50 - year-old balsam fir, oak, maple, yellow birch / balsam fir white pine forests, red oak forests, mixed forests, and old eastern white cedar forest; often very common in decaying gilled mushrooms in the above forest habitats, especially those dominated hardwoods. Also in riparian habitats, and a meadow with Salix shrubs. Several collected from decaying corncobs and cornhusks. Collecting period. V – XI (most common in August and September). Collecting method. Unbaited and carrion-baited pitfall traps, processing forest litter through Berlese funnels, Luminoc pitfall-light traps, Lindgren traps, intercept traps, and treading leaf litter at the edge of a pond, sifting decaying mushrooms.	en	Klimaszewski, Jan, Godin, Benoit, Davies, Anthony, Bourdon, Caroline, Horwood, Denise (2018): Forty new records of aleocharine beetles, and two new species in the genera Acrotona Thomson and Atheta Thomson, for the province of Manitoba, Canada (Coleoptera: Staphylinidae: Aleocharinae). Insecta Mundi 2018 (641): 1-33, DOI: 10.5281/zenodo.5173990
A5636478EE15FFA0FF3DBAF2FC8C773C.taxon	distribution	Distribution Origin Nearctic Distribution Canada: MB, NB, ON, QC. USA: RI New provincial Canada: Manitoba: Katherine Lake, 2016 - IX- 14, 50.6603 ° N, 99.8947 ° W, 652 m, record mushrooms, sifting, B. Godin & D. Horwood (LFC) 1 male. References Casey 1911, Klimaszewski et al. 2018 Collection and habitat data. Habitat (outside of the study area). Mixed forest among moist leaves along the margin of a large (35 m × 15 m) vernal pond; also found along vernal pools in Ontario. Collecting period. V – X. Collecting method. Sifting leaf litter.	en	Klimaszewski, Jan, Godin, Benoit, Davies, Anthony, Bourdon, Caroline, Horwood, Denise (2018): Forty new records of aleocharine beetles, and two new species in the genera Acrotona Thomson and Atheta Thomson, for the province of Manitoba, Canada (Coleoptera: Staphylinidae: Aleocharinae). Insecta Mundi 2018 (641): 1-33, DOI: 10.5281/zenodo.5173990
A5636478EE12FFA7FF3DBED1FDAA729D.taxon	distribution	Distribution Origin Nearctic Distribution Canada: AB, LB, MB, NB, NF, NS, NT, ON, QC, YT. USA: AK, NH New provincial Canada: Manitoba: Moose Lake, 2016 - IX- 19, 49.2018 ° N, 95.3423 ° W, 353 m, records mushroom, sifting, B. Godin & D. Horwood (BGC) 1 male, 4 females; same locality data (LFC) 1 male, 3 females; Souris River, 2016 - IX- 9, 49.5022 ° N, 99.9915 ° W, 402 m, river bank, oak litter, sifting, B. Godin & D. Horwood (BGC) 1 male, 1 female; same locality data (LFC) 1 male, 1 female; Katherine Lake, 2016 - IX- 14, 50.6603 ° N, 99.8947 ° W, 652 m, mushrooms, sifting, B. Godin & D. Horwood (BGC) 1 male; Adam Lake, 2016 - IX- 11, 49.0508 ° N, 100.0666 ° W, 689 m, mushrooms, sifting, B. Godin & D. Horwood (LFC) 1 female. References Casey 1911, Klimaszewski et al. 2018 Collection and habitat data. Habitat (outside of the study area). In hardwood (silver maple, sugar maple, red oak, birch, alder) and mixed forests with some in fir, pine forests, mature spruce and eastern white cedar forest, and a tamarack bog margin. Adults in leaf litter, moss, litter and Sphagnum near small body of water, in moist leaves on vernal pond margins, in litter at base of tree in Carex marsh, in lining of deserted beaver lodge, in moist grass litter on abandoned beaver pond margin, Microtus nests, marmot (Marmota sp.) burrows, Sphagnum moss, litter on barren, litter around raspberry, in frass in tree holes of birch and maple, and in leaf litter on river margin. One individual in lichen on tree trunk. Collecting period. III-X. Collecting method. Sifting various kinds of litter, pitfall traps, Luminoc ® pitfall-light traps, Malaise traps.	en	Klimaszewski, Jan, Godin, Benoit, Davies, Anthony, Bourdon, Caroline, Horwood, Denise (2018): Forty new records of aleocharine beetles, and two new species in the genera Acrotona Thomson and Atheta Thomson, for the province of Manitoba, Canada (Coleoptera: Staphylinidae: Aleocharinae). Insecta Mundi 2018 (641): 1-33, DOI: 10.5281/zenodo.5173990
A5636478EE12FFA7FF3DBA16FC5377D0.taxon	distribution	Distribution Origin Nearctic Distribution Canada: AB, MB, NB, NF, NS, ON, QC, SK. USA: NH New provincial Canada: Manitoba: Katherine Lake, 2016 - IX- 14, 50.6603 ° N, 99.8947 ° W, 652 m, records mushrooms, sifting, B. Godin & D. Horwood (BGC) 1 male, 2 females; Hecla, 2016 - IX- 17, 51.7849 ° N, 96.6152 ° W, 218 m, mushroom, sifting, B. Godin & D. Horwood (LFC) 1 male, 1 female. References Klimaszewski et al. 2006, 2018 Collection and habitat data. Habitat (outside of the study area). In many kinds of forests and open habitats such as mixed forests, mature hardwood forests, birch forests, red maple forests and swamps, eastern white cedar swamps, fir forests (old-growth and 50 - year-old plantations), forested black spruce bogs, agricultural fields, meadow with Salix shrubs, and lake and river margins. Adults in Sphagnum moss, moss and Sphagnum along creek in forest, moss on tree trunks, moss and litter near vernal ponds in forests, Sphagnum in mist zone of waterfall, leaf litter, birch litter, litter on barren, litter in alpine and subalpine habitats, drift material along lake and river margins, and from lining of deserted beaver lodge. Collecting period. IV – XII, IX. Collecting method. Sifting mushrooms (new data), sifting moss, various kinds of forest litter, and drift material, pitfall traps.	en	Klimaszewski, Jan, Godin, Benoit, Davies, Anthony, Bourdon, Caroline, Horwood, Denise (2018): Forty new records of aleocharine beetles, and two new species in the genera Acrotona Thomson and Atheta Thomson, for the province of Manitoba, Canada (Coleoptera: Staphylinidae: Aleocharinae). Insecta Mundi 2018 (641): 1-33, DOI: 10.5281/zenodo.5173990
A5636478EE13FFA6FF3DBED1FB9173A6.taxon	distribution	Distribution Origin Nearctic Distribution Canada: AB, BC, LB, MB, NB, NF, NS, ON, QC, SK, YT. USA: AK, ME, NH New provincial Canada: Manitoba: Hecla, 2016 - IX- 17, 51.7849 ° N, 96.6152 ° W, 218 m, mushroom, record sifting, B. Godin & D. Horwood (LFC) 1 female. References Casey 1906, Klimaszewski et al. 2006 [as O. grandipennis], 2018 Collection and habitat data. Habitat (outside of the study area). In many forest types, including hardwood, red oak, mixed, mature red spruce, red spruce and birch, old red pine, spruce and balsam fir forests, and old-growth eastern white cedar forests. Adults common in decaying and rotten mushrooms in many of the above forests types. Some, in group of Pholiota mushrooms of trunk of dead Populus; another under coyote dung on forest road. Collecting period. V – X (most common in September). Collecting method. Sifting mushrooms, forest litter, dung, and moss; some of above material also processed through Berlese funnels, Luminoc ® pitfall-light traps, unbaited pitfall traps.	en	Klimaszewski, Jan, Godin, Benoit, Davies, Anthony, Bourdon, Caroline, Horwood, Denise (2018): Forty new records of aleocharine beetles, and two new species in the genera Acrotona Thomson and Atheta Thomson, for the province of Manitoba, Canada (Coleoptera: Staphylinidae: Aleocharinae). Insecta Mundi 2018 (641): 1-33, DOI: 10.5281/zenodo.5173990
