taxonID	type	description	language	source
A676E712FFDBFF86050EF9AF167289B0.taxon	type_taxon	Type species Jacaena distincta Thorell, 1897	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2013): Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae). Journal of Natural History 47 (23 - 24): 1539-1567, DOI: 10.1080/00222933.2012.763059, URL: http://dx.doi.org/10.1080/00222933.2012.763059
A676E712FFDBFF86050EF9AF167289B0.taxon	diagnosis	Diagnosis Representatives of Jacaena can be recognized by a combined presence of characters: the integument of carapace is strongly granulate or punctate, forming radiating striae (Figures 1 – 2); the fovea is deep and circular (Figure 3 B); the dorsal opisthosomal scutum occupying approximately half the length of the opisthosoma or much less in males (Figure 1), completely absent in females (Figure 2); the dorsal pattern of the opisthosoma consists of paired, oval patches, followed by three to five pale chevrons (Figures 1 – 2) on a dark greenish background; the male palp is provided with an apically excavated proximal RTA and a truncated distal RTA (Figure 12 A); the pyriform tegulum bears a distal conductor and a hyaline tegular apophysis (Figure 9 D); the embolic base originates disto-prolaterally, its filiform embolus wrapping behind the tegulum (Figure 4 C, D); the elongated female internal ducts are coiled, moving peripherally before leading to poorly defined or elliptical spermathecae, which are posteriorly situated.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2013): Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae). Journal of Natural History 47 (23 - 24): 1539-1567, DOI: 10.1080/00222933.2012.763059, URL: http://dx.doi.org/10.1080/00222933.2012.763059
A676E712FFDBFF86050EF9AF167289B0.taxon	description	Description Carapace (Figures 1, 2) ovoid in dorsal view, attenuated in front, narrowed opposite palpal insertion, widest between coxae II and III, in profile highest between PME and fovea; pars thoracica rounded, marked with deep, circular fovea (Figure 3 B); ocular area and clypeus with few stiff setae; pars thoracica with fine recumbent setae; integument of carapace usually with numerous granules (Figures 1 A – B, D – H, 2 A – E, G – H) or punctures (Figures 1 C, 2 F) forming radiating striae; colour from dark chestnutbrown to brownish orange. Both eye rows straight in dorsal view (Figures 1, 2, 3 D); in frontal view, AER very slightly procurved, PER slightly procurved (Figure 3 E); AME circular, dark; PME oval, pearly white; ALE and PLE subcircular, light; anterior eyes separated by their diameter or less; posterior eyes evenly spaced, separated by 1.5 – 2 their diameter (except in J. mihun comb. nov. and J. thakek comb. nov. whose PME are distinctly enlarged); ALE and PLE separated by 1 – 1.5 times their diameter. MOQ longer than wide, slightly wider in front than behind (wider behind than in front in J. mihun comb. nov. and J. thakek comb. nov. because of enlarged PME). Clypeal height 1.5 times diameter of AME. Chelicerae convex anteriorly, with conspicuous condyles, fang furrow with three promarginal teeth and one to three retromarginal denticles. Chilum heavily sclerotised, trapezoidal (Figure 3 E). Mouthparts and sternum brownish orange, darker along sternal margin. Labium (Figure 3 A) subtriangular, longer than wide, invaginated at posterolateral corners, with few strong setae on anterior surface. Gnathocoxae (Figure 3 A) rectangular, with very faint oblique depressions, each with anterolateral serrula and anteromedian scopula arising from pale apical area. Sternum (Figure 3 A) scutiform, strongly rebordered, posterior margin truncated, not protruded between coxae IV; anterior margin medially excavated to accommodate labium; lateral margin with triangular extensions fitting to and between coxae. Leg formula 4123; legs brownish orange; femora I always with two elongated disto-prolateral spines; patellae unmodified; anterior tibiae ventrally not flattened, with five to nine pairs of ventral spines and single prolateral spine; anterior metatarsi with four to six pairs of ventral spines; posterior metatarsi with brush of distoventral hairs, anterior without such brush; anterior tarsi devoid of spines; posterior tarsi with few spines; two pectinate claws carrying two to three denticles; claw tufts indistinct; trichobothria long, in two dorsal rows on tarsi and metatarsi, one row on tibiae increasing in length distally. Opisthosoma (Figures 1, 2) ovoid in dorsal view, dark greenish dorsally, dorsal pattern consisting of two pale oval patches situated medially, followed by three to five chevrons. Dorsal scutum with poorly diffused edge, covering less than half the length of the opisthosoma in male, completely absent in female. Two pairs of muscle apodemes, their surface covered by minute, circular, heavily sclerotised spots; first pair of spots locating just behind dorsal scutum (Figures 1, 2), second pair always located at centre of second pale oval patches. Epigastric region entirely covered with scutum (Figure 3 C), slightly extending anteriorly to form short collar. Ventral scutum absent. Postgenital scuta (Figure 3 C) triangular or semi-circular, situated posterior laterally, surrounding book lung openings but clearly separated from epigastric scutum. Colour of venter uniform in all species (Figure 3 C), dark greenish, marked with dark trident on pale rectangular area, with numerous tiny sclerotised spots arranged into four longitudinal lines running between epigastric furrow and transverse tracheal spiracle. Spinnerets two-segmented; ALS conical, contiguous, distal segment short; PMS small, slender in males, flattened and elongated in females; PLS with short distal segment. Male palp. Palpal femur without modification. Retrolateral tibial apophysis consisting of basal and distal parts (Figure 12 A); basal RTA triangular or digitiform, apical surface usually with deep excavation; distal RTA short and membranous, digitiform. Cymbium dorsally with mid-longitudinal patch of chemosensitive hairs (Figure 4 B, E, H). Tegulum elongate-ovoid, with distinct apico-prolateral depression accommodating base of embolus. Embolus filiform, elongated, curving behind tegulum (Figure 4 A, B). Tegular apophysis hyaline, gradually tapered, originating near base of conductor (Figures 6 A, 9 D). Conductor originating on disto-prolateral surface of tegulum; its shape from simple, triangular projection (Figure 10 A, D) to elongated cylinder with median groove (Figures 5 D, 13 A) or large crescent-shaped lamina (Figures 6 A, 9 D). Female copulatory organ. Epigynal region (Figure 7) heavily sclerotised, with two copulatory orifices usually filled with dark secretory substance (Figure 7 A, B). Internal duct system consisting of semi-transparent proximal duct (Figures 6 F, 14 C) or enlarged chamber (Figures 6 D, 13 E) connected to loosely coiled insemination ducts (Figure 12 E: ID) and spermathecae. Spermathecae slightly enlarged, poorly defined (Figure 8 A – D, G) or with elliptical and spherical enlargement (Figure 8 E, F); surface of spermathecae usually provided with glandular pores (Figures 8 G, 12 E). Small, perforated ampulla (Figures 6 D, F, 12 E, 13 E, 14 B, E: AM) connected to proximal portion of insemination ducts or chambers (Figures 6 D, 13 E), usually with hardened secretion filling proximal portion of ducts (Figure 7 A, B).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2013): Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae). Journal of Natural History 47 (23 - 24): 1539-1567, DOI: 10.1080/00222933.2012.763059, URL: http://dx.doi.org/10.1080/00222933.2012.763059
A676E712FFDBFF86050EF9AF167289B0.taxon	biology_ecology	Natural history Most Jacaena species were collected by sifting decomposing organic litter in moist, primary evergreen hill forests at relatively high elevation. These forests are generally covered with a thick layer of humus on the forest floor and the forest canopy is often dense, allowing limited sunlight to reach the ground. A species from Thailand (J. mihun) inhabits deciduous forests and open plantations. Jacaena thakek was sifted from relatively dry litter in a narrow belt of trees, bamboo or Rattan (disturbed secondary habitat); the type locality of this species is almost of the same altitude of the Mekong basin (159 m alt.)	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2013): Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae). Journal of Natural History 47 (23 - 24): 1539-1567, DOI: 10.1080/00222933.2012.763059, URL: http://dx.doi.org/10.1080/00222933.2012.763059
A676E712FFDBFF86050EF9AF167289B0.taxon	discussion	Included species Jacaena angoonae sp. nov.; J. distincta Thorell, 1897; J. erawan (Deeleman-Reinhold, 2001) comb. nov.; J. lunulata sp. nov.; J. mihun Deeleman-Reinhold, 2001; J. punctata sp. nov.; J. peculiaris sp. nov.; J. schwendingeri (Deeleman-Reinhold, 2001) comb. nov.; J. thakek (Jäger, 2007) comb. nov.; J. zhui (Zhang and Fu, 2011) comb. nov. Two further species were also collected from Vietnam (male and female) and China (female); they have not yet been described.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2013): Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae). Journal of Natural History 47 (23 - 24): 1539-1567, DOI: 10.1080/00222933.2012.763059, URL: http://dx.doi.org/10.1080/00222933.2012.763059
A676E712FFDBFF86050EF9AF167289B0.taxon	distribution	Distribution Predominantly Indo-Burmese sub-region of the Oriental Region (Myanmar, Thailand, Laos, Vietnam), also in evergreen forests of southern China (Yunnan Province).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2013): Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae). Journal of Natural History 47 (23 - 24): 1539-1567, DOI: 10.1080/00222933.2012.763059, URL: http://dx.doi.org/10.1080/00222933.2012.763059
A676E712FFD1FF870519F9F511DE8A30.taxon	description	Figure 15	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2013): Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae). Journal of Natural History 47 (23 - 24): 1539-1567, DOI: 10.1080/00222933.2012.763059, URL: http://dx.doi.org/10.1080/00222933.2012.763059
A676E712FFD1FF870519F9F511DE8A30.taxon	materials_examined	Type material Holotype: ♀, Myanmar, Tanintharyi Region, Tenasserim Division, Mt Mooleyit, 1885 – 1889, L. Fea leg. (MSNG, examined). Original label: Cadius distinctus Thor., Birmania: Tenasserim, Mooleyit, Fea. For comments on species name and recent redescription of this species see Deeleman-Reinhold (2001: 467).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2013): Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae). Journal of Natural History 47 (23 - 24): 1539-1567, DOI: 10.1080/00222933.2012.763059, URL: http://dx.doi.org/10.1080/00222933.2012.763059
A676E712FFD1FF870519F9F511DE8A30.taxon	discussion	Remarks The type species was described based on a single female that closely resembles the female of J. peculiaris sp. nov. (Figure 2 H) in general appearance.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2013): Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae). Journal of Natural History 47 (23 - 24): 1539-1567, DOI: 10.1080/00222933.2012.763059, URL: http://dx.doi.org/10.1080/00222933.2012.763059
A676E712FFD1FF870519F9F511DE8A30.taxon	diagnosis	Diagnosis Jacaena distincta can be easily distinguished from its congeners by the widely separated copulatory orifices that are medially situated on the epigynal region, and the poorly defined spermathecae that are not distinctly enlarged (Figure 15 E).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2013): Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae). Journal of Natural History 47 (23 - 24): 1539-1567, DOI: 10.1080/00222933.2012.763059, URL: http://dx.doi.org/10.1080/00222933.2012.763059
A676E712FFD1FF870519F9F511DE8A30.taxon	distribution	Distribution Known only from the type locality (Mt Mooleyit, south-eastern Myanmar).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2013): Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae). Journal of Natural History 47 (23 - 24): 1539-1567, DOI: 10.1080/00222933.2012.763059, URL: http://dx.doi.org/10.1080/00222933.2012.763059
A676E712FFD3FF8506D3FBB510028FF2.taxon	description	(Figures 1 D, 2 G, 3 D, 4 G – I, 6 F, 7 D, 8 A – D, 10 A – C)	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2013): Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae). Journal of Natural History 47 (23 - 24): 1539-1567, DOI: 10.1080/00222933.2012.763059, URL: http://dx.doi.org/10.1080/00222933.2012.763059
A676E712FFD3FF8506D3FBB510028FF2.taxon	materials_examined	Type material Holotype: ♂, Thailand: Nakhon Ratchasima Province: Khao Yai NP, 1000 m, secondary forest, sifting leaf litter, 25 October 1985, P. Deeleman leg. (RMNH, examined, Figures 4 G – I, 10 A – C). Paratype: One ♀, same data as holotype (RMNH, examined). Other material examined The following specimens were studied by Deeleman-Reinhold (2001) and included in the original description, but they were not designated as types: Thailand: Chiang Mai Province, Chiang Mai City, Mae Hia, deciduous forest and teak plantation, pitfall trap, 17 – 24 January 1988, 1 ♂ (MHNG, examined). From locality as previous, 7 – 13 February 1988, 1 ♀ (MHNG, examined). P. J. Schwendinger leg. New material Thailand: Sakon Nakhon Province: Phu Phan NP, 400 m, litter sample, 18 November 2007, P. Dankittipakul leg., 1 ♀ (TNHM). Loei Province: Phu Ruea NP, park headquarters, 850 – 8710 m, pan traps, 15 – 16 July 2007, P. Dankittipakul leg., 1 ♂ (TNHM). Khon Kaen Province: Phu Pha Man NP, in front of Tham (cave) Phiang Din, near the road Chumphae – Phu Kradung (at km 122) c. 300 m, 24 July 2000, sieving in a mixed deciduous forest, det. C. Deeleman, P. J. Schwendinger leg., 1 ♂ (MHNG, TH – 00 / 03). Sugarcane plantation near Phu Pha Man NP, 300 m, 15 June 2007, P. Dankittipakul leg., 1 ♂ (TNHM). Chiang Mai Province, Chiang Mai City, Mae Hia, 7 – 24 January 1988, det. C. Deeleman, P. J. Schwendinger leg., 1 ♂ (MHNG, examined).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2013): Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae). Journal of Natural History 47 (23 - 24): 1539-1567, DOI: 10.1080/00222933.2012.763059, URL: http://dx.doi.org/10.1080/00222933.2012.763059
A676E712FFD3FF8506D3FBB510028FF2.taxon	diagnosis	Diagnosis Males of J. mihun are very similar to those of J. thakek comb. nov. in having the strongly excavated apico-prolateral side of the tegulum (Figures 4 G cf. 5 G), and the short conductor (Figures 4 H cf. 5 H), but can be distinguished by the presence of a small triangular denticle on the dorsal surface of the conductor (Figure 10 A). Females can be recognised by the tremendously lengthened insemination ducts strongly convoluted; spermathecae are indistinct (Figure 8 A – D).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2013): Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae). Journal of Natural History 47 (23 - 24): 1539-1567, DOI: 10.1080/00222933.2012.763059, URL: http://dx.doi.org/10.1080/00222933.2012.763059
A676E712FFD3FF8506D3FBB510028FF2.taxon	distribution	Distribution Northern and north-eastern Thailand.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2013): Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae). Journal of Natural History 47 (23 - 24): 1539-1567, DOI: 10.1080/00222933.2012.763059, URL: http://dx.doi.org/10.1080/00222933.2012.763059
A676E712FFD2FF9A0625FC36107B8DB6.taxon	description	(Figures 1 E, 5 A – C, 11)	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2013): Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae). Journal of Natural History 47 (23 - 24): 1539-1567, DOI: 10.1080/00222933.2012.763059, URL: http://dx.doi.org/10.1080/00222933.2012.763059
A676E712FFD2FF9A0625FC36107B8DB6.taxon	materials_examined	Type material Holotype: ♂, Thailand, Kanchanaburi Province, Erawan Waterfalls NP, evergreen forest along river, leaf litter, 15 – 16 March 1986, P. P. and C. Deeleman leg. (RMNH, examined, Figures 5 A – C, 11). Other material examined The following specimen was studied by Deeleman-Reinhold (2001) and included in the original description, but it was not designated as a type: Thailand: Lamphang Province, Doi Khuntan NP, Doi Khuntan, disturbed evergreen forest, 1200 – 1300 m, 16 February 1992, P. J. Schwendinger leg., 1 ♂ (MHNG). New material Thailand: Kanchanaburi Province, Erawan NP and waterfalls, 100 m, 10 November 2000, P. J. Schwendinger leg., 1 ♂ (MHNG, TH – 00 / 05).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2013): Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae). Journal of Natural History 47 (23 - 24): 1539-1567, DOI: 10.1080/00222933.2012.763059, URL: http://dx.doi.org/10.1080/00222933.2012.763059
A676E712FFD2FF9A0625FC36107B8DB6.taxon	diagnosis	Diagnosis Jacaena erawan comb. nov. can be easily distinguished by the truncated dRTA of the male palp when viewed from the ventral side (Figure 5 A), and by the base of the conductor that is rather broad and gradually tapered towards its apex (Figures 5 A, 11 A – C).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2013): Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae). Journal of Natural History 47 (23 - 24): 1539-1567, DOI: 10.1080/00222933.2012.763059, URL: http://dx.doi.org/10.1080/00222933.2012.763059
A676E712FFD2FF9A0625FC36107B8DB6.taxon	distribution	Distribution Northern and Western Thailand.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2013): Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae). Journal of Natural History 47 (23 - 24): 1539-1567, DOI: 10.1080/00222933.2012.763059, URL: http://dx.doi.org/10.1080/00222933.2012.763059
A676E712FFCDFF9A06CCFDF3119089B0.taxon	description	(Figures 1 F, 5 G – I, 10 D – F)	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2013): Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae). Journal of Natural History 47 (23 - 24): 1539-1567, DOI: 10.1080/00222933.2012.763059, URL: http://dx.doi.org/10.1080/00222933.2012.763059
A676E712FFCDFF9A06CCFDF3119089B0.taxon	materials_examined	Type material Holotype: ♂, Laos, Khammouan Province, 9.5 km northeast of Thakek, 159 m, 17 ◦ 26.936 ′ N, 104 ◦ 52.499 ′ E, sieving leaf litter in front of cave, 30 October 2004, P. Jäger and V. Vedel leg. (SMF – 56635, examined, Figures 1 F, 5 G – I, 10 D – F).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2013): Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae). Journal of Natural History 47 (23 - 24): 1539-1567, DOI: 10.1080/00222933.2012.763059, URL: http://dx.doi.org/10.1080/00222933.2012.763059
A676E712FFCDFF9A06CCFDF3119089B0.taxon	diagnosis	Diagnosis This species seems to be closest to J. mihun, because both species have very distinctly enlarged PME; their palps can be distinguished from other members of Jacaena by the pear-shaped bulb anteriorly narrowed, and the apex of the short, triangular conductor is pointing apically (Figures 4 G – I cf. 5 G – I). The male of this species can be separated from that of J. mihun by the absence of a small, triangular denticle on the dorsal surface of the conductor (Figure 10 A cf. 10 D). However, we have a private notion that the boundary between these two species is vague and J. thakek comb. nov. is possibly a junior synonym of J. mihun. Much larger samples would be needed to corroborate these slight differences as species-specific rather than intraspecific variation because more material recently obtained suggests that J. mihun has relatively broad distribution covering northern and north-eastern parts of Thailand (and most likely extending eastwards to Laos where J. thakek comb. nov. was collected).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2013): Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae). Journal of Natural History 47 (23 - 24): 1539-1567, DOI: 10.1080/00222933.2012.763059, URL: http://dx.doi.org/10.1080/00222933.2012.763059
A676E712FFCDFF9A06CCFDF3119089B0.taxon	distribution	Distribution Known only from the type locality, Khammouan Province, Laos.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2013): Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae). Journal of Natural History 47 (23 - 24): 1539-1567, DOI: 10.1080/00222933.2012.763059, URL: http://dx.doi.org/10.1080/00222933.2012.763059
A676E712FFCDFF980647F9F510558AF0.taxon	description	(Figures 1 A, 2 A, 4 A – C, 7 A, 8 F, 9 A – C)	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2013): Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae). Journal of Natural History 47 (23 - 24): 1539-1567, DOI: 10.1080/00222933.2012.763059, URL: http://dx.doi.org/10.1080/00222933.2012.763059
A676E712FFCDFF980647F9F510558AF0.taxon	materials_examined	Type material Holotype: ♂, Thailand, Chiang Mai Province, Chiang Dao District, Doi Chiang Dao Wildlife Sanctuary, 510 m, pitfall trap, 22 September – 25 October 1990, P. J. Schwendinger leg. (MHNG, examined, Figures 1 A, 4 A – C, 9 A – C). Paratype: From type locality, 450 m, 7 March 1987, P. J. Schwendinger leg., 1 ♀ (MHNG, examined, Figures 2 A, 7 A, 8 F).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2013): Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae). Journal of Natural History 47 (23 - 24): 1539-1567, DOI: 10.1080/00222933.2012.763059, URL: http://dx.doi.org/10.1080/00222933.2012.763059
A676E712FFCDFF980647F9F510558AF0.taxon	diagnosis	Diagnosis Males of J. schwendingeri comb. nov. can be recognised by the large and curved conductor that is divided into a heavily sclerotised outer margin and a semi-transparent inner membrane (Figures 4 A, 9 A – C); the apex of conductor is gradually tapered and abruptly bent anteriorly (Figures 4 C, 9 B). Females can be distinguished by the large copulatory orifices that are situated at the centre of the epigynal region (Figure 7 A), and by the middle portion of the internal ducts, which are greatly enlarged, whereas the anterior part of the spermathecae is spherical (Figure 8 F).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2013): Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae). Journal of Natural History 47 (23 - 24): 1539-1567, DOI: 10.1080/00222933.2012.763059, URL: http://dx.doi.org/10.1080/00222933.2012.763059
A676E712FFCDFF980647F9F510558AF0.taxon	distribution	Distribution Known only from the type locality in northern Thailand. Other specimens from different localities previously known (Deeleman-Reinhold 2001: 461) were misplaced (see J. lunulata sp. nov. for more details).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2013): Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae). Journal of Natural History 47 (23 - 24): 1539-1567, DOI: 10.1080/00222933.2012.763059, URL: http://dx.doi.org/10.1080/00222933.2012.763059
A676E712FFCEFF990672FBFB10288A20.taxon	diagnosis	Diagnosis Females of J. zhui comb. nov. can be distinguished from those of J. schwendingeri comb. nov. by their smaller copulatory orifices that are located more anteriorly, internal ducts that are simple, and the spermathecae that are robust. In J. schwendingeri comb. nov. the copulatory orifices are distinctly larger and medially situated on the epigynal plate (Figure 7 A), the internal ducts are partially enlarged, and the anterior portion of the spermathecae is spherical (Figure 8 F). Males differ by the apex of the conductor, which is blunt, whereas in J. schwendingeri comb. nov. the conductor is sharply pointed and more elongate (Figures 4 A, 9 A).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2013): Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae). Journal of Natural History 47 (23 - 24): 1539-1567, DOI: 10.1080/00222933.2012.763059, URL: http://dx.doi.org/10.1080/00222933.2012.763059
A676E712FFCEFF990672FBFB10288A20.taxon	distribution	Distribution Yunnan Province, southern China.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2013): Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae). Journal of Natural History 47 (23 - 24): 1539-1567, DOI: 10.1080/00222933.2012.763059, URL: http://dx.doi.org/10.1080/00222933.2012.763059
A676E712FFCEFF9C06FAF9841095891A.taxon	description	(Figures 1 B, H, 2 B, C, 3 A – C, E, 4 D – F, 6 A – C, 7 C, 8 G, 9 D, E, 12)	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2013): Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae). Journal of Natural History 47 (23 - 24): 1539-1567, DOI: 10.1080/00222933.2012.763059, URL: http://dx.doi.org/10.1080/00222933.2012.763059
A676E712FFCEFF9C06FAF9841095891A.taxon	materials_examined	Type material Holotype: ♂, Thailand: Chiang Mai Province, Chomthong District, Doi Inthanon NP, Doi Inthanon, evergreen hill forest, 1600 – 1680 m, extraction of soil sample, 11 January 2006, P. Dankittipakul leg. (MHNG, PDC 5746). Paratypes: From type locality, pitfall trap, 25 March – 23 April 2000, P. Dankittipakul leg., 1 ♀ (MHNG, PDC 5747); litter sample, 10 September 2005, P. Dankittipakul leg., 2 ♂, 1 ♀, (TNHM); evergreen hill forest, 1650 m, litter sample, 9 October 1999, P. Dankittipakul leg., 1 ♂ (THNM, PDC 5748). Doi Inthanon, evergreen hill forest, 1750 m, litter sample, 15 January 2000, P. Dankittipakul leg., 1 ♂ (THNM, PDC 5749). Doi Inthanon, evergreen hill forest, 1500 m, litter sample, 9 October 1999, P. Dankittipakul leg., 1 ♂, 2 ♀ (THNM, PDC 5750). Doi Inthanon, evergreen hill forest, 1250 m, 6 November 1985, P. J. Schwendinger leg., 1 ♂ (MHNG, misidentified as S. erawan). Chiang Mai Province and District, Doi Suthep-Pui NP, Doi Suthep, 1200 m, 18 June 1987, P. J. Schwendinger leg., 1 ♀ (MHNG, misidentified as S. schwendingeri). Doi Suthep, 960 m, pitfall trap, 31 November 1986 – 2 January 1987, P. J. Schwendinger leg., 1 ♂ (RMNH, misidentified as S. schwendingeri).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2013): Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae). Journal of Natural History 47 (23 - 24): 1539-1567, DOI: 10.1080/00222933.2012.763059, URL: http://dx.doi.org/10.1080/00222933.2012.763059
A676E712FFCEFF9C06FAF9841095891A.taxon	diagnosis	Diagnosis Males of J. lunulata sp. nov. are similar to those of J. schwendingeri comb. nov. but differ by having a crescent-shaped conductor that is distinctly broader and strongly curved backward (Figures 6 A, 9 D, E), a tegulum that is greatly expanded, and the distal RTA of the male palp that is digitiform (Figure 6 C). In comparison, in J. schwendingeri comb. nov. the conductor is narrower and its sharply pointed apex is abruptly bent (Figure 4 A, C), the tegulum and the distal RTA are significantly smaller. Females can be distinguished by the rather simple internal genitalia, the ID of which consists of circular proximal ducts, the more convoluted ID, and the poorly defined spermathecae (Figure 8 G cf. 8 F).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2013): Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae). Journal of Natural History 47 (23 - 24): 1539-1567, DOI: 10.1080/00222933.2012.763059, URL: http://dx.doi.org/10.1080/00222933.2012.763059
A676E712FFCEFF9C06FAF9841095891A.taxon	etymology	Etymology The specific epithet is derived from Latin adjective (lunulatus = crescent-shaped), referring to the lamina-shaped conductor on the male palp.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2013): Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae). Journal of Natural History 47 (23 - 24): 1539-1567, DOI: 10.1080/00222933.2012.763059, URL: http://dx.doi.org/10.1080/00222933.2012.763059
A676E712FFCEFF9C06FAF9841095891A.taxon	description	Description Male (holotype). Total length 6.6. Carapace 3.2 long, 2.5 wide. Opisthosoma 3.4 long, 2.0 wide. Eye sizes and interdistances: AME 0.15; ALE 0.13; PME 0.10; PLE 0.13; AME – AME 0.06; AME – ALE 0.05; PME – PME 0.15; PME – PLE 0.16. MOQ 0.33 long; front width 0.33; back width 0.35. Leg formula 4123. Spination. Leg I: femur 1 – 1 pl; tibia 8 pl 7 rl; metatarsus 4 pl 4 rl. Leg II: tibia 7 pl 6 rl; metatarsus 4 pl 4 rl. Leg III: tibia 1 – 1 v 1 pl; metatarsus 1 – 1 v 1 pl. Leg IV: tibia 1 – 2 – 2 v 1 – 1 rl; metatarsus 1 – 1 – 1 – 1 v 1 pl 1 rl. Leg measurements: Leg I 8.1 (2.3, 0.9, 2.2, 1.8, 0.9); II 7.5 (2.1, 0.9, 1.9, 1.7, 0.9); III 6.7 (1.9, 0.9, 1.4, 1.6, 0.9); IV 9.4 (2.6, 1.0, 2.1, 2.5, 1.2). Colouration and pattern (Figure 1 B): Carapace dark chestnut-brown; integument coarsely granulated. Chelicerae, sternum, labium and gnathocoxae dark reddishbrown. Coxae, trochanters and femora dark brown; patellae and tibiae brown, distal part of tibiae orange; metatarsi I – II brown, III – IV orange; tarsi orange. Opisthosoma ovoid; dorsum of opisthosoma dark grey, with pattern consisting of one pair of pale patches, followed by four transverse chevrons and pale area above spinnerets. Dorsal scutum occupying approximately one-third of opisthosoma length, posterior margin rounded, with oval extensions fusing with dorsal muscle apodemes. Palp (Figures 6 A – C, 9 D, E, 12 A – C): Proximal RTA triangular in ventral view (Figure 12 A); apical surface partially membranous, slightly excavated, forming shallow concavity (Figure 9 D, E). Distal RTA digitiform, its apex blunt (Figures 6 C, 12 A, C). Conductor (Figures 6 A, 9 D, E, 12 A – C) crescent-shaped, enlarged and broad, semi-transparent, with heavily pigmented outer margin, sharply pointed apex curving backwards. Tegular apophysis hyaline, lanceolate, originating behind base of conductor, aligning with tip of embolus (Figures 6 A, 9 D). Female (paratype, NHML, PDC 5747). Total length 8.1. Carapace 3.3 long, 2.5 wide. Opisthosoma 4.5 long, 2.7 wide. Eye sizes and interdistances: AME 0.11; ALE 0.13; PME 0.08; PLE 0.13; AME – AME 0.08; AME – ALE 0.05; PME – PME 0.15; PME – PLE 0.16. MOQ 0.38 long; front width 0.31; back width 0.35. Leg formula 1423. Spination. Leg I: femur 1 – 1 pl; tibia 8 pl 7 rl; metatarsus 4 pl 4 rl. Leg II: tibia 7 pl 6 rl; metatarsus 4 pl 4 rl. Leg III: tibia 2 – 1 v 1 pl; metatarsus 1 – 1 v 1 pl. Leg IV: tibia 1 – 1 – 2 v 1 rl; metatarsus 1 – 1 – 1 – 1 v 1 pl 1 rl. Leg measurements: Leg I 8.6 (2.3, 1.0, 2.3, 2.0, 1.0); II 7.9 (2.2, 1.0, 1.9, 1.8, 1.0); III 6.6 (1.8, 2.8, 1.5, 1.5, 1.0); IV 8.3 (2.3, 0.8, 2.0, 2.2, 1.0). Colouration and pattern (Figure 2 B): Carapace dark chestnut-brown; integument coarsely granulated. Chelicerae, sternum, labium and gnathocoxae dark reddishbrown. Legs I – II brown, femora slightly darker than other segments, tibiae orangebrown distally; legs III – IV orange-brown except femora dark brown. Opisthosoma elongate-ovoid; dorsum grey, with pair of pale patches connected anteriorly, followed by four chevrons and pale area above spinnerets (first pair very faint and narrowed, disconnected medially). Copulatory organ (Figures 7 C, 8 F, 12 D, E): Copulatory orifices situated medially on heavily sclerotized epigynal region (Figures 7 C, 12 D). Proximal half of insemination ducts forming two loops encircling copulatory ducts (Figures 8 G, 12 E); inner loop distinctly enlarged, filled with dark substance (Figure 8 G); distal half of ducts ascending anteriorly then traversing and descending to connect with poorly defined spermathecae (Figures 8 G, 12 E); glandular pores situated at anterior margin of spermathecae (Figure 12 E, GF).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2013): Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae). Journal of Natural History 47 (23 - 24): 1539-1567, DOI: 10.1080/00222933.2012.763059, URL: http://dx.doi.org/10.1080/00222933.2012.763059
A676E712FFCEFF9C06FAF9841095891A.taxon	biology_ecology	Natural history Types of this new species were collected in evergreen hill forests of Doi Suthep and Doi Inthanon between 1250 and 1750 m.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2013): Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae). Journal of Natural History 47 (23 - 24): 1539-1567, DOI: 10.1080/00222933.2012.763059, URL: http://dx.doi.org/10.1080/00222933.2012.763059
A676E712FFCEFF9C06FAF9841095891A.taxon	distribution	Distribution Chiang Mai Province, northern Thailand.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2013): Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae). Journal of Natural History 47 (23 - 24): 1539-1567, DOI: 10.1080/00222933.2012.763059, URL: http://dx.doi.org/10.1080/00222933.2012.763059
A676E712FFCBFF930539FA6F114F8F40.taxon	description	(Figures 1 C, 2 E, F, 5 D – F, 6 E, 7 B, 13)	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2013): Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae). Journal of Natural History 47 (23 - 24): 1539-1567, DOI: 10.1080/00222933.2012.763059, URL: http://dx.doi.org/10.1080/00222933.2012.763059
A676E712FFCBFF930539FA6F114F8F40.taxon	materials_examined	Type material Holotype: ♂, Thailand: Nan Province: Tha Wang Pha District, Nanthaburi NP, Doi Wao, Doi Wao, evergreen hill forest near the summit, 1300 – 1550 m, extraction of leaf litter samples, 7 December 2005, P. Dankittipakul leg. (MHNG, PDC 5130).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2013): Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae). Journal of Natural History 47 (23 - 24): 1539-1567, DOI: 10.1080/00222933.2012.763059, URL: http://dx.doi.org/10.1080/00222933.2012.763059
A676E712FFCBFF930539FA6F114F8F40.taxon	materials_examined	Paratypes: Same data as holotype, 2 ♀ (TNHM). From type locality, 17 December 2002, P. J. Schwendinger leg., 1 ♂, 1 ♀ (MHNG, TH – 02 / 21). Doi Wao, evergreen forest surrounding park bungalow, 1300 m, 9 December 2005, P. Dankittipakul leg., 1 ♀ (THNM, PDC 4658). Nan Province: Phu Ka District, Doi Phu Ka NP, Doi Phu Ka, evergreen forest along nature trail, 1300 m, 9 December 2005, P. Dankittipakul leg., 1 ♀ (THNM, PDC 4655).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2013): Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae). Journal of Natural History 47 (23 - 24): 1539-1567, DOI: 10.1080/00222933.2012.763059, URL: http://dx.doi.org/10.1080/00222933.2012.763059
A676E712FFCBFF930539FA6F114F8F40.taxon	diagnosis	Diagnosis Jacaena punctata sp. nov. can be easily distinguished from its congeners by the integument of the carapace, which is punctate (Figures 1 C, 2 F) instead of granulate. Males are easily recognised by a tuft of bristles (Figures 5 D, 13 A) on the retrolateral side of the cymbium – a character that is absent in other species. Females are recognised by the copulatory orifice connected to an enlarged chamber of the internal genitalia (Figure 13 E), and by the elliptical spermathecae (Figure 13 E).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2013): Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae). Journal of Natural History 47 (23 - 24): 1539-1567, DOI: 10.1080/00222933.2012.763059, URL: http://dx.doi.org/10.1080/00222933.2012.763059
A676E712FFCBFF930539FA6F114F8F40.taxon	etymology	Etymology The specific epithet is a derived from Latin adjective (punctatus = with punctures), referring to the punctated carapace of this new species.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2013): Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae). Journal of Natural History 47 (23 - 24): 1539-1567, DOI: 10.1080/00222933.2012.763059, URL: http://dx.doi.org/10.1080/00222933.2012.763059
A676E712FFCBFF930539FA6F114F8F40.taxon	description	Description Male (holotype). Total 7.4. Carapace 3.8 long, 2.8 wide. Opisthosoma 3.4 long, 2.0 wide. Eye sizes and interdistances: AME 0.15; ALE 0.15; PME 0.11; PLE 0.15; AME – AME 0.08; AME – ALE 0.06; PME – PME 0.16; PME – PLE 0.23. MOQ 0.45 long; front width 0.43; back width 0.43. Leg formula 4123. Spination. Leg I: femur 1 – 1 pl; tibia 8 pl 7 rl; metatarsus 5 pl 5 rl. Leg II: tibia 7 pl 6 rl; metatarsus 4 pl 4 rl. Leg III: tibia 2 – 1 – 1 – 2 v; metatarsus 1 – 1 – 1 v 1 pl 1 rl. Leg IV: tibia 1 – 1 – 1 – 2 v 1 rl; metatarsus 1 – 1 – 1 – 1 v 1 pl 1 rl. Leg measurements: Leg I 10.7 (3.1, 1.2, 2.8, 2.3, 1.3); II 9.0 (2.7, 1.0, 2.2, 1.9, 1.2); III 7.4 (2.2, 0.8, 1.5, 1.8, 1.1); IV 10.9 (3.0, 1.2, 2.5, 2.8, 1.4). Colouration and pattern (Figure 1 C): Carapace dark chestnut-brown; integument punctated, covered with punctures forming radiating striae. Carapace, chelicerae and sternum dark reddish brown. Coxae, trochanters and femora dark brown (except femora I – II brown, distally yellow); patellae, tibiae, metatarsi and tarsi orange. Opisthosoma ovoid; dorsum of opisthosoma with pattern consisting of paired, medially connected patches, followed by four chevrons and white area above spinnerets on dark grey background. Dorsal scutum orange-brown, wider in front than behind, posterior margin straight, occupying approximately half of opisthosoma length. Palp (Figures 5 D – F, 13 A – C): Proximal RTA triangular, apex sharply pointed, apical surface with shallow excavation. Distal RTA subtriangular in lateral view, apex blunt, ventral surface membranous. Cymbium with group of elongated bristles situated on meso-retrolateral margin (Figures 5 D, 13 A). Tegulum pyriform, gradually tapered towards apex, prolateral margin slightly compressed. Conductor cylindrical, gradually tapered towards bluntly pointed apex, with shallow groove running along dorsal surface to accommodate tip of embolus. Embolus filiform. Tegular apophysis indistinct. Female (paratype, THNM, PDC 4658). Total length 8.7. Carapace 3.6 long, 2.8 wide. Opisthosoma 4.7 long, 3.0 wide. Eye sizes and interdistances: AME 0.15; ALE 0.15; PME 0.10; PLE 0.16; AME – AME 0.08; AME – ALE 0.05; PME – PME 0.18; PME – PLE 0.16. MOQ 0.41 long; front width 0.36; back width 0.41. Leg formula 1423. Spination. Leg I: femur 1 – 1 pl; tibia 8 pl 8 rl; metatarsus 5 pl 5 rl. Leg II: tibia 7 pl 7 rl; metatarsus 4 pl 5 rl. Leg III: tibia 1 – 1 – 1 – 1 – 1 v; metatarsus 1 – 1 – 1 – 1 v 1 rl. Leg IV: tibia 1 – 1 – 1 – 1 pv 2 v 1 – 1 rl; metatarsus 1 – 1 – 1 – 1 – 1 v 1 pl 1 rl. Leg measurements: Leg I 10.6 (3.0, 1.2, 2.6, 2.5, 1.3); II 9.0 (2.6, 1.1, 2.3, 1.9, 1.1); III 7.5 (2.1, 1.0, 1.5, 1.8, 1.1); IV 10.8 (2.8, 1.1, 2.4, 3.0, 1.5). Colouration and pattern (Figure 2 F): Carapace dark chestnut-brown; integument punctate, with large pits forming striae. Fovea obsolete. Chelicerae, sternum, labium and gnathocoxae dark reddish-brown. Legs I – II brown, femora slightly darker than other segments, tibiae orange-brown distally; legs III – IV orange-brown except femora dark brown, distally pale yellow. Opisthosoma elongate-ovoid; dorsum grey, with paired, irregular-shaped patches, followed by four chevrons and white area above spinnerets. Copulatory organ (Figures 6 E, 7 B, 13 D, E): Copulatory orifices subcircular, situated anteriorly, separated by twice their diameter, plugged with black secretory substance (Figure 7 B). Internally with large, semi-transparent chamber connected to posterior tubular duct (Figure 13 E), abruptly ascends anteriorly then curves along anterior margin of copulatory orifices and connects to enlarged, elliptical spermathecae. Secretory ampullae spherical, heavily perforated (Figure 6 E), situated on proximal part of insemination ducts (Figures 6 E, 13 E). Natural history Jacaena punctata sp. nov. inhabits evergreen hill forests of Doi Wao and Doi Phu Kha (both localities are higher than 1300 m).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2013): Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae). Journal of Natural History 47 (23 - 24): 1539-1567, DOI: 10.1080/00222933.2012.763059, URL: http://dx.doi.org/10.1080/00222933.2012.763059
A676E712FFCBFF930539FA6F114F8F40.taxon	distribution	Distribution Known from two mountains in Nan Province, northern Thailand.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2013): Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae). Journal of Natural History 47 (23 - 24): 1539-1567, DOI: 10.1080/00222933.2012.763059, URL: http://dx.doi.org/10.1080/00222933.2012.763059
A676E712FFC4FF9606C1FCA4166F8DC4.taxon	description	(Figures 2 D, 7 E, 8 E, 14 A – C)	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2013): Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae). Journal of Natural History 47 (23 - 24): 1539-1567, DOI: 10.1080/00222933.2012.763059, URL: http://dx.doi.org/10.1080/00222933.2012.763059
A676E712FFC4FF9606C1FCA4166F8DC4.taxon	materials_examined	Type material Holotype: ♀, Thailand: Huay Nam Dang NP, joint border between Mae Taeng District of Chiang Mai Province and Pai District of Mae Hong Son Province, Doi Chang, evergreen hill forest, 1930 m, extraction of soil sample, 4 June 1983, P. J. Schwendinger leg. (MHNG, PDC 4650). Paratypes: Same data as holotype, 3 ♀ (NHML & THNM, PDC 4651).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2013): Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae). Journal of Natural History 47 (23 - 24): 1539-1567, DOI: 10.1080/00222933.2012.763059, URL: http://dx.doi.org/10.1080/00222933.2012.763059
A676E712FFC4FF9606C1FCA4166F8DC4.taxon	diagnosis	Diagnosis Females of this new species bear very close resemblance to those of J. schwendingeri comb. nov. in having large, medially situated copulatory orifices, and similar shape of spermathecae. However, in J. angoonae sp. nov. the copulatory orifices are teardrop-shaped (Figure 7 E) and the insemination ducts are tubular without any modification (Figure 8 E) whereas in J. schwendingeri comb. nov. the orifices are subcircular (Figure 7 A) and the median part of the ducts is greatly enlarged (Figure 8 F).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2013): Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae). Journal of Natural History 47 (23 - 24): 1539-1567, DOI: 10.1080/00222933.2012.763059, URL: http://dx.doi.org/10.1080/00222933.2012.763059
A676E712FFC4FF9606C1FCA4166F8DC4.taxon	etymology	Etymology The specific epithet is an eponymous patronym (genitive form of a noun) dedicated to Dr Angoon Lewvanich, a prolific Thai taxonomist.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2013): Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae). Journal of Natural History 47 (23 - 24): 1539-1567, DOI: 10.1080/00222933.2012.763059, URL: http://dx.doi.org/10.1080/00222933.2012.763059
A676E712FFC4FF9606C1FCA4166F8DC4.taxon	description	Description Female (holotype). Total length 7.5. Carapace 3.0 long, 2.4 wide. Opisthosoma 4.5 long, 2.6 wide. Eye sizes and interdistances: AME 0.10; ALE 0.13; PME 0.08; PLE 0.11; AME – AME 0.06; AME – ALE 0.05; PME – PME 0.13; PME – PLE 0.16. MOQ 0.36 long; front width 0.28; back width 0.33. Leg formula 4123. Spination. Leg I: femur 1 – 1 pl; tibia 8 pl 8 rl; metatarsus 4 pl 4 rl. Leg II: tibia 7 pl 6 rl; metatarsus 4 pl 4 rl. Leg III: tibia 1 – 1 – 1 v 1 pl; metatarsus 1 – 1 v 1 pl. Leg IV: tibia 1 – 1 – 1 – 1 – 2 v 1 – 1 rl; metatarsus 1 – 1 – 1 v 1 pl 1 rl. Leg measurements: Leg I 8.6 (2.5, 1.0, 2.3, 1.85, 1.0); II 7.7 (2.2, 0.9, 1.9, 1.7, 1.0); III 6.1 (1.7, 0.8, 1.4, 0.85); IV 9.2 (2.5, 1.0, 2.1, 2.4, 1.2). Colouration and pattern (Figure 2 D): Carapace dark chestnut-brown; integument coarsely granulated. Chelicerae, sternum, labium and gnathocoxae dark reddishbrown; coxae, trochanters and femora dark brown; patellae and tibiae brown, tibiae orange distally; metatarsi and tarsi I – II light brown, III – IV orange. Opisthosoma ovoid; dorsum with pattern consisting of paired, oblique patches, followed by three chevrons on dark grey background. Copulatory organ (Figures 7 E, 8 E, 14 A – C): Copulatory orifices teardrop-shaped (Figure 14 A). Insemination ducts elongated, loosely coiled (Figure 14 B); proximal duct connected to copulatory orifice semi-transparent (Figure 8 E), forming sigmoid arch; secretory ampullae situated on posterior surface of proximal ducts; distal half of duct ascending anteriorly then descending posteriorly to connect with spermathecae. Spermathecae elliptical, anterior part distinctly enlarged, more or less spherical with glandular pore (Figure 14 B). Male. Unknown. Natural history Jacaena angoonae sp. nov. inhabits evergreen hill forest of Doi Chang (1930 m). This new species marks the highest altitudinal record among members of the genus Jacaena.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2013): Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae). Journal of Natural History 47 (23 - 24): 1539-1567, DOI: 10.1080/00222933.2012.763059, URL: http://dx.doi.org/10.1080/00222933.2012.763059
A676E712FFC4FF9606C1FCA4166F8DC4.taxon	distribution	Distribution Known only from the type locality, Mae Hongson Province, northern Thailand.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2013): Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae). Journal of Natural History 47 (23 - 24): 1539-1567, DOI: 10.1080/00222933.2012.763059, URL: http://dx.doi.org/10.1080/00222933.2012.763059
A676E712FFC1FF97053DFE2111A38F1F.taxon	description	(Figures 2 H, 6 D, 7 F, 14 D – E)	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2013): Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae). Journal of Natural History 47 (23 - 24): 1539-1567, DOI: 10.1080/00222933.2012.763059, URL: http://dx.doi.org/10.1080/00222933.2012.763059
A676E712FFC1FF97053DFE2111A38F1F.taxon	materials_examined	Type material Holotype: ♀, Thailand: Nan Province: Tha Wang Pha District, Nanthaburi NP, Doi Wao, evergreen hill forest near the summit, 1300 – 1550 m, extraction of leaf litter samples, 7 December 2005, P. Dankittipakul leg. (MHNG, PDC 4640). Paratypes: Same data as holotype, 2 ♀ (THNM, PDC 4654).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2013): Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae). Journal of Natural History 47 (23 - 24): 1539-1567, DOI: 10.1080/00222933.2012.763059, URL: http://dx.doi.org/10.1080/00222933.2012.763059
A676E712FFC1FF97053DFE2111A38F1F.taxon	diagnosis	Diagnosis Unlike most females of Jacaena, J. peculiaris sp. nov. can be readily distinguished by the very enlarged spermathecae (Figures 7 F, 14 E), a character that is shared among the females of Teutamus species. However, females of Jacaena can be separated from those of Teutamus in the lack of the dorsal scutum and their epigastric scutum is not protruding and forming an elongated collar; the spination of legs and other somatic characters are also different from Teutamus.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2013): Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae). Journal of Natural History 47 (23 - 24): 1539-1567, DOI: 10.1080/00222933.2012.763059, URL: http://dx.doi.org/10.1080/00222933.2012.763059
A676E712FFC1FF97053DFE2111A38F1F.taxon	etymology	Etymology The specific epithet refers to the peculiar shape of the spermathecae, which are the largest among Jacaena females (Latin, pecularis = belonging exclusively to one person).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2013): Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae). Journal of Natural History 47 (23 - 24): 1539-1567, DOI: 10.1080/00222933.2012.763059, URL: http://dx.doi.org/10.1080/00222933.2012.763059
A676E712FFC1FF97053DFE2111A38F1F.taxon	description	Description Female (holotype). Total length 6.1. Carapace 2.8 long, 2.2 wide. Opisthosoma 3.3 long, 1.7 wide. Eye sizes and interdistances: AME 0.13; ALE 0.13; PME 0.11; PLE 0.15; AME – AME 0.08; AME – ALE 0.05; PME – PME 0.11; PME – PLE 0.15. MOQ 0.33 long; front width 0.31; back width 0.33. Leg formula 4123. Spination. Leg I: femur 1 – 1 pl; tibia 8 pl 7 rl; metatarsus 5 pl 5 rl. Leg II: tibia 7 pl 6 rl; metatarsus 4 pl 4 rl. Leg III: tibia 1 – 1 – 1 v 1 pl 1 rl; metatarsus 1 – 1 v 1 rl. Leg IV: tibia 1 – 1 – 1 – 1 – 2 v 1 – 1 rl; metatarsus 1 – 1 – 1 – 1 – 1 v 1 pl 1 rl. Leg measurements: Leg I 7.5 (2.2, 0.8, 2.0, 1.6, 0.9); II 6.8 (2.0, 0.9, 1.5, 1.5, 0.9); III 5.6 (1.6, 0.7, 1.2, 1.3, 2.8); IV 8.8 (2.3, 0.9, 2.0, 2.5, 1.1). Colouration and pattern (Figure 2 H): Carapace dark reddish-brown; integument finely granulated. Fovea obsolete. Chelicerae, sternum, labium and gnathocoxae dark reddish-brown. Legs I – II dark brown, femora darkest, tibiae yellow distally; legs III – IV orange-brown, femora brown. Opisthosoma elongate-ovoid; dorsum grey, with paired, enlarged pale patches, followed by broad pale band and pale area above spinnerets. Copulatory organ (Figures 6 D, 7 F, 14 D – E): Copulatory orifices (Figure 14 D) circular, anterior margin poorly defined, posterior margin represented by elevated ridge, connected to bell-shaped, semi-transparent chamber (Figure 6 D). Insemination ducts tubular, elongated, slender, opening to mesolateral surface of enlarged spermathecae (Figure 14 E). Spermathecae more or less reniform, with large glandular pore situated anteriorly (Figures 6 D, 14 E). Secretory ampullae attached to apex of semi-transparent chamber (Figure 6 D). Male. Unknown. Natural history Types of J. peculiaris sp. nov. were collected from evergreen hill forests near the summit of Doi Wao where the humidity is constantly high (> 85 % relatively humidity), even during summer.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2013): Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae). Journal of Natural History 47 (23 - 24): 1539-1567, DOI: 10.1080/00222933.2012.763059, URL: http://dx.doi.org/10.1080/00222933.2012.763059
A676E712FFC1FF97053DFE2111A38F1F.taxon	distribution	Distribution Known only from the type locality, Nan Province, northern Thailand.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2013): Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae). Journal of Natural History 47 (23 - 24): 1539-1567, DOI: 10.1080/00222933.2012.763059, URL: http://dx.doi.org/10.1080/00222933.2012.763059
