taxonID	type	description	language	source
A7576C4EFFD9FFED818095916640F897.taxon	diagnosis	Diagnosis Body slender, elongate; frons flat or slightly bulbous around base of antenna, often covered with elongate setae; wing strongly banded; wing cell m 3 closed; relatively elongate hind legs; epandrium arched, lateral margins extending ventrally; medial atrium present posteriorly between gonocoxites; gonocoxal apodemes approximately equal to length of gonocoxite; hypandrium with patch of strong, posteriorly directed setae; three spermathecae; spermathecal sac small rounded. Redescription Male and female. Body length: 6.0 – 8.0 mm. Head. Shape longer than high; frons flat, slightly raised around base of antennae, overlain with pubescence but sometimes with glossy patch above antennae, usually admixed with elongate setae, male frons width at narrowest point either narrower than or equal width to ocellar tubercle, female frons much wider than ocellar tubercle; lower frons and face with silver or grey pubescence; ocellar tubercle black, overlain with grey pubescence admixed with long, dark setae; male occiput concave or slightly convex; usually single row of postocular setae in male (2 – 3 poorly defined rows in male of A. olivacea sp. nov.); female occiput convex; 2 – 3 poorly defined rows of postocular setae; antennae positioned low on frons; length equal to or slightly shorter than head length; setae on antennae often enlarged; scape slightly wider than pedicel; flagellum gradually tapered (A. olivacea sp. nov.) to turbinate (A. pulchella), length approximately equal to combined length of scape and pedicel.	en	Winterton, Shaun L. (2007): Revision of the Australian stiletto fly genus Acatopygia Kröber (Diptera: Therevidae: Agapophytinae). Zootaxa 1405: 51-62, DOI: 10.5281/zenodo.175484
A7576C4EFFD9FFED818095916640F897.taxon	description	Thorax. Scutum dark, overlain with brown, green or grey pubescence, admixed with scattered dark setae, setae longer in male; macrosetae dark; scutellum relatively small, dark, sometimes overlain with matte, black pubescence anteriorly; pleuron glossy dark brown or black, partially overlain with dense, grey pubescence and long pale setae; coxae and legs dark or variously patterned with yellow; fore and hind femora with elongate velutum patch on ventral and posteroventral surfaces respectively; hind femur much longer than other femora; anteroventral subapical seta on hind femur absent (a row of dark, ventral setae present on all femora in A. pulchella); wing hyaline with dark banding, third medial cell (m 3) closed; scutal chaetotaxy: notopleural (np), 2 – 4; supraalar (sa), 1; postalar (pa), 1 – 2; dorsocentral (dc), 3 – 6; scutellar (sc), 1. Abdomen. Elongate, glossy black or dark brown, extensive velutum dorsally on male abdomen (absent in A. olivacea sp. nov.); setae on abdomen of uniform size and distribution, not concentrated posteromedially on tergite 2. Male genitalia. Epandrium width equal to length medially, posterolateral corners rounded or pointed, sometimes extending anterolaterally and ventrally to partially conceal gonocoxites; cercus elongate or short; tergite 8 emarginate medially; gonocoxite with large medial atrium with velutum­covered membrane; hypandrium large, quadrangular to sub­triangular, partially fused to gonocoxites laterally, numerous dark setae located medially along posterior margin; ventral lobe elongate, extending medially and dorsally, velutum present apically; gonocoxal apodeme narrow, shorter than or equal to gonocoxite length; gonocoxal process present, narrow, shorter than or equal in length to inner gonocoxal process; inner gonocoxal process spatulate apically; gonostylus curved medially along length; distiphallus straight or curved; dorsal apodeme of parameral sheath present, ‘ T’ ­ shaped, fused laterally to gonocoxal apodeme in A. olivacea sp. nov.; ventral apodeme of parameral sheath often elongate, length equal to ejaculatory apodeme, rarely shorter; lateral ejaculatory apodeme present but not enlarged; ejaculatory apodeme narrow, elongate. Female genitalia. Anterior process of tergite 8 narrow; three spherical spermathecae of equal size; spermathecal sac simple, spherical, without lobes; spermathecal ducts joining to common spermathecal sac duct; acanthophorite A 1 setae well developed.	en	Winterton, Shaun L. (2007): Revision of the Australian stiletto fly genus Acatopygia Kröber (Diptera: Therevidae: Agapophytinae). Zootaxa 1405: 51-62, DOI: 10.5281/zenodo.175484
A7576C4EFFD9FFED818095916640F897.taxon	discussion	Comments The shape of the gonocoxites and aedeagus indicates a close relationship between Acatopygia and Patanothrix. The banded wings and shape of the head and male genitalia are diagnostic for the genus. Acatopygia species are distributed throughout coastal south­eastern Australia. Acatopygia pulchella is common in southeastern Queensland while A. paradoxa is more common in southern New South Wales. Acatopygia olivacea sp. nov. is less common than the other two species in collections and is known from southern Queensland to Northern New South Wales. Included species	en	Winterton, Shaun L. (2007): Revision of the Australian stiletto fly genus Acatopygia Kröber (Diptera: Therevidae: Agapophytinae). Zootaxa 1405: 51-62, DOI: 10.5281/zenodo.175484
A7576C4EFFDBFFEA818090A96084FCC8.taxon	materials_examined	Holotype female, AUSTRALIA: QUEENSLAND: Mt. Coot [­ t] ha, 20. iii. 1948 (MEI 25318) (ANIC). Condition: Good. Paratypes, AUSTRALIA: QUEENSLAND: male, Pistol Gap, nr. Byfield, 23 10 ’ S, 150 40 ’ E, 10. i. 1970, mv lamp, G. A. Holloway (MEI 25319) (AMSA); female, 5 km N Leyburn, 27 58 ’ S, 151 38 ’ E, 3. iv. 1993, G. & A. Daniels, C. J. Burwell (MEI 33661) (GDCB / AMSA); male, Brisbane Forest Park: Scrub Road, 10. x. 1997, 27 25 ’ 05 ” S, 152 50 ’ 13 ” E, open eucalypt forest, S. Winterton, N. Power, D. White, Malaise trap 3 (ANIC 29 21105) (QDPI). NEW SOUTH WALES: male, Mosman, 20. ii. [19] 24, Mackerras [J. Mann Coll.] (MEI 25317) (QMBA); female, Warrumbungle National Park, Brownes Creek nr. Wombelong Creek, M. E. Irwin, 12 – 16. xii. 1995, nr. cliff overhang at water hole, 31 16 ’ 24 ” S, 148 57 ’ 38 ” E (MEIC / CASC).	en	Winterton, Shaun L. (2007): Revision of the Australian stiletto fly genus Acatopygia Kröber (Diptera: Therevidae: Agapophytinae). Zootaxa 1405: 51-62, DOI: 10.5281/zenodo.175484
A7576C4EFFDBFFEA818090A96084FCC8.taxon	diagnosis	Diagnosis Scutum covered with olive­green pubescence; male frons wider than ocellar tubercle, occiput with multiple, poorly defined rows of postocular setae; legs brown; male abdomen without silver velutum; male sternites 4 – 7 with black, scale­like setae.	en	Winterton, Shaun L. (2007): Revision of the Australian stiletto fly genus Acatopygia Kröber (Diptera: Therevidae: Agapophytinae). Zootaxa 1405: 51-62, DOI: 10.5281/zenodo.175484
A7576C4EFFDBFFEA818090A96084FCC8.taxon	description	Description Male: Body length: 7.5 – 8.0 mm. Head. Frons flat, width equal to ocellar tubercle at narrowest point; frons and face glossy brown; frons overlain with sparse, brown pubescence dorsally, silver pubescence laterally between eye and base of antenna, admixed with short, dark setae; eye facets uniform size throughout eye, transverse furrow absent; ocellar tubercle flattened, black, overlain with grey pubescence admixed with several dark setae; occiput convex, dark, overlain with dense green­grey pubescence dorsally, changing to silver on lower portion and on gena; 2 – 3 poorly­defined rows of relatively long, black, postocular setae; gena overlain with silver­white pubescence admixed with white setae; palp and labellum brown, palp covered with dark setae; antennal length shorter than head; antennna orange­brown, overlain with brown­grey pubescence, admixed with scattered, dark setae on scape, pedicel, and base of flagellum; flagellum tapered gradually with two apical segments; style pale. Thorax. Glossy dark brown; scutum overlain with olive­green pubescence, densely admixed with short setae; scutal macrosetae black; scutellum overlain with matte, black pubescence anteriorly, grey pubescence posteriorly; pleuron with sparse, grey pubescence anteriorly on postpronotum and proepisternum, also on meron, metanepisternum and posterior margin of katepisternum; coxae glossy dark­brown, overlain with grey pubescence, admixed with setae on anterolateral surfaces of mid and fore coxae; setae dark on fore coxa and posterolateral surface of hind coxa; setae pale on mid and hind coxae; legs glossy brown with sparse, pale setae, longer on fore femur; wing hyaline with brown or black banding, two bands across basal medial (bm) and discal (d) cells respectively, third band across the subcostal and radial fields, basal bands connected approximately midway; venation brown; haltere stem brown, knob white; scutal chaetotaxy (pairs): np, 3; sa, 1; pa, 1; dc, 3 – 4; sc, 1. Abdomen. Glossy brown with scattered pale setae, setae on posterior segments a mixture of pale and dark setae, setae on terminalia pale; tergite 1 with grey pubescence laterally, brown pubescence medially on tergites 1 – 5; intersegmental membrane of segments 2 – 3 white; sternites 4 – 7 with covering of black, scale­like setae; terminalia dark. Male genitalia. Epandrium quadrangular, length much shorter along midline than greatest width, posterolateral corners rounded (Fig. 1 A); cercus rounded in shape; tergite 8 broad, slightly emarginate medially (Fig. 1 B); sternite 8 ovate, anterior margin straight, posterior margin rounded with medial patch of elongate setae (Fig. 1 E); gonocoxite (Figs 1 C – D) slightly rounded in profile; gonocoxal process narrow, length equal to inner gonocoxal process, elongate setae on posterior surface; medial atrium present, relatively small; hypandrium large, fused laterally to gonocoxites, large patch of posteriorly projecting setae medially; ventral lobe broad, projecting dorsally, relatively short, velutum on apically posterior surface; gonostylus broad basally, curved medially along length, erect setae on medial surface; inner gonocoxal process spatulate, equal in length to gonostylus; gonocoxal apodeme approximately equal to length of gonocoxite; distiphallus curved ventrally (Figs 1 F – G), ventral apodeme of parameral sheath relatively broad, much shorter than ejaculatory apodeme; dorsal apodeme recurved dorsally, ‘ Y’ ­ shaped, fused to ejaculatory apodeme laterally; lateral ejaculatory apodeme small and tapered; ejaculatory apodeme acuminate anteriorly. Female: Body length: 7.0 – 9.0 mm. Similar to male except: Frons slightly wider than ocellar tubercle, covered with greenish­grey pubescence dorsally; shorter setae on head, thorax and abdomen; scutum with short dark setae; abdominal sternites without black scale­like setae. Female genitalia. Spermathecal sac present, spherical in shape; spermathecal sac duct thickened at common junction with spermathecal ducts; furca quadrangular in shape; spermathecae spherical.	en	Winterton, Shaun L. (2007): Revision of the Australian stiletto fly genus Acatopygia Kröber (Diptera: Therevidae: Agapophytinae). Zootaxa 1405: 51-62, DOI: 10.5281/zenodo.175484
A7576C4EFFDBFFEA818090A96084FCC8.taxon	discussion	Comments Acatopygia olivacea sp. nov. is easily recognised from other species of Acatopygia based on the scutal colouration and male abdominal characters such as the presence of black, scale­like setae on the sternites and the lack of silver velutum on the tergites. This species is known from a disparate series of specimens from northern New South Wales and Queensland.	en	Winterton, Shaun L. (2007): Revision of the Australian stiletto fly genus Acatopygia Kröber (Diptera: Therevidae: Agapophytinae). Zootaxa 1405: 51-62, DOI: 10.5281/zenodo.175484
A7576C4EFFDDFFE8818092B26760F947.taxon	materials_examined	Other material examined. AUSTRALIA: NEW SOUTH WALES: near Braidwood, Monga, 35.582 S, 149.918 E, 14. x. 1954, S. J. Paramonov, 19 males, 2 females, (MEI 109489 ­ 507) (ANIC).	en	Winterton, Shaun L. (2007): Revision of the Australian stiletto fly genus Acatopygia Kröber (Diptera: Therevidae: Agapophytinae). Zootaxa 1405: 51-62, DOI: 10.5281/zenodo.175484
A7576C4EFFDDFFE8818092B26760F947.taxon	diagnosis	Diagnosis Antenna and foreleg bright yellow; eye without transverse furrow; dark, enlarged setae on scape and pedicel; scutum overlain with dark­grey pubescence; male abdomen with dense, silver velutum; ventral lobe of male genitalia acuminate. Redescription Male: Body length: 7.0 – 8.0 mm. Head. Frons flat, slightly bulbous above antennal base, narrower than ocellar tubercle at narrowest point; frons and face glossy brown; frons overlain with sparse, brown pubescence dorsally, admixed with sparsely distributed, elongate, dark setae; round, glabrous patch immediately above antenna; silver pubescence around base of antenna and on face; eye with facets larger in upper portion, transverse furrow absent; ocellar tubercle raised, black, overlain with grey pubescence admixed with dark setae; occiput concave, dark, overlain with light grey pubescence; single row of relatively long, black, postocular setae; gena overlain with silver­white pubescence along eye margin, admixed with white setae; palp bright yellow, labellum brown, covered with dark setae; antennal length approximately equal to length of head; antenna yellow, overlain with yellow pubescence, admixed with erect, elongate, dark setae on scape and pedicel; flagellum tapered; style pale.	en	Winterton, Shaun L. (2007): Revision of the Australian stiletto fly genus Acatopygia Kröber (Diptera: Therevidae: Agapophytinae). Zootaxa 1405: 51-62, DOI: 10.5281/zenodo.175484
A7576C4EFFDDFFE8818092B26760F947.taxon	description	Thorax. Glossy black; scutum overlain with dark­grey pubescence, densely admixed with long, setae; scutal macrosetae black; scutellum overlain with matte, black pubescence anteriorly, glabrous posteriorly; pleuron with dense, silver pubescence except on anepisternum and anterior part of anepimeron; long pale setae on postpronotum, proepisternum anepisternum and katatergite; fore coxa yellow, mostly glabrous; mid and hind coxae black, overlain with dense silver pubescence; fore leg yellow, end of basitarsus and rest of tarsomeres brown; mid and hind legs brown­black with apical 1 / 5 of femora, tibia and basitarsi dark yellow; wing hyaline with brown or black banding, two bands across cells bm and d respectively, a third band across subcostal and radial fields less distinct, dark spot on bifurcation of veins R 4 and R 5; venation brown; haltere stem brown, knob white; scutal chaetotaxy (pairs): np, 3; sa, 1; pa, 1; dc, 2 – 3; sc, 1. Abdomen. Glossy black with scattered pale setae, setae on anterior segments longer laterally; setae on terminalia both pale and dark; dense, silver, velutinous pubescence on all tergites; intersegmental membrane of segments 2 – 4 white; terminalia dark. Male genitalia. Epandrium quadrangular, length much shorter along midline than greatest width, posterolateral corners rounded (Fig. 2 A); cercus elongate; tergite 8 narrow, band­like, deeply emarginate (Fig. 2 B); sternite 8 narrow (Fig. 2 E); gonocoxite (Figs 2 C – D) slightly rounded in profile; gonocoxal process narrow, 1 / 2 length of inner gonocoxal process, elongate setae on posterior surface; medial atrium present, membranous area covered with velutum; hypandrium large, fused laterally to gonocoxites, broadly emarginate medially with large, posteriorly projecting setae (concentrated medially); ventral lobe narrow, apically acuminate with velutum present at apex; gonostylus curved medially along length, erect setae on medial surface; inner gonocoxal process spatulate, equal in length to gonostylus; gonocoxal apodeme approximately equal to length of gonocoxite; distiphallus straight (Figs 2 F – G), ventral apodeme of parameral sheath elongate, narrow, approximately equal length to ejaculatory apodeme; dorsal apodeme recurved dorsally, narrowly ‘ Y’ ­ shaped, poorly sclerotised; lateral ejaculatory apodeme small and tapered; ejaculatory apodeme narrow anteriorly. Female: Body length: 7.0 – 9.0 mm. Similar to male except: Frons slightly wider than ocellar tubercle, covered with brown pubescence dorsally; shorter setae on head, thorax and abdomen; abdominal tergites without dense velutum covering. Female genitalia. Furca well sclerotised; spermathecal sac present, spherical in shape; spermathecae spherical (Fig. 2 H).	en	Winterton, Shaun L. (2007): Revision of the Australian stiletto fly genus Acatopygia Kröber (Diptera: Therevidae: Agapophytinae). Zootaxa 1405: 51-62, DOI: 10.5281/zenodo.175484
A7576C4EFFDDFFE8818092B26760F947.taxon	discussion	Comments The holotype of A. paradoxa was apparently destroyed in 1956 during the fire in the Hungarian National Museum of Natural History (Winterton et al. 2001) (pers. com., M. Foldvari, HNMNH) and no paratypes or other specimens are known from the type locality. Kröber (1913: fig. 23) presents a colour image of the head of A. paradoxa. A neotype is designated to clarify and stabilise the taxonomic status of Spatulipalpa paradoxa Kröber. The neotype specimen designated herein is a male from the Australian Capital Territory, a region relatively close to the original type locality. Female specimens from the series mentioned above match the original description by Kröber, but a male is designated here as most of the diagnostic characters for the species are found in the male and not the female. There is no doubt that the neotype designated here represents the male counterpart of the female described originally by Kröber (1912 b). Acatopygia paradoxa is readily differentiated from other members of Acatopygia by the distinctive, bright­yellow antennae and foreleg colouration.	en	Winterton, Shaun L. (2007): Revision of the Australian stiletto fly genus Acatopygia Kröber (Diptera: Therevidae: Agapophytinae). Zootaxa 1405: 51-62, DOI: 10.5281/zenodo.175484
A7576C4EFFDFFFE58180960461DAFA51.taxon	materials_examined	Holotype male, AUSTRALIA: NEW SOUTH WALES: “ Australien ”, A. L. Schrader leg. ded., 25. ix. 1896 (MEI 90753) (DEIC). Condition: poor; specimen damaged. Other material examined. AUSTRALIA, NEW SOUTH WALES: female, Minnamurra Falls, [­ 34.632, 150.717], 18. xii. 1968, mv light, C. E. Chadwick (MEI 025309) (NSWA). QUEENSLAND: 3 males, Brisbane, [­ 27.465, 153.017], 3. ix. 1927, J. Mann (MEI 109628 ­ 30) (ANIC); female, Brisbane, [­ 27.465, 153.017], 20. viii. 1927, J. Mann (MEI 109636) (ANIC); male, Bribie Island, [­ 27.05, 153.15], 4. ix. 1980, M. G. Jefferies (MEI 109802) (MGJC / ANIC).	en	Winterton, Shaun L. (2007): Revision of the Australian stiletto fly genus Acatopygia Kröber (Diptera: Therevidae: Agapophytinae). Zootaxa 1405: 51-62, DOI: 10.5281/zenodo.175484
A7576C4EFFDFFFE58180960461DAFA51.taxon	diagnosis	Diagnosis Antenna and foreleg brown; eye with transverse furrow; scutum overlain with brown pubescence, distinct white­grey dorsocentral stripes; femora with dark ventral setae; abdomen overlain with silver velutum; epandrium large, cerci elongate. Redescription Male: Body length: 6.0 – 7.5 mm. Head. Frons flat, bulbous around antennal base (Figs 3 A – B), narrower than ocellar tubercle at narrowest point; frons and face glossy black; frons overlain brown pubescence tending towards silver in lower portion, admixed with elongate, dark setae; bulbous area above antennal base glabrous; silver pubescence around base of antenna and on face, admixed with several elongate dark setae; eye with facets slightly larger in upper portion, transverse furrow present; ocellar tubercle raised, black, overlain with brown pubescence admixed with elongate, dark setae; occiput concave, overlain with light­grey pubescence, single row of relatively long, black, postocular setae dorsally; gena overlain with silver­white pubescence, admixed with white setae; mouthparts light orange­brown, covered with dark setae; antennal length approximately equal to length of head, antenna orange­brown, overlain with brown pubescence, admixed with erect, elongate, dark setae on scape and pedicel; flagellum turbinate, slightly shorter than combined length of scape and pedicel.	en	Winterton, Shaun L. (2007): Revision of the Australian stiletto fly genus Acatopygia Kröber (Diptera: Therevidae: Agapophytinae). Zootaxa 1405: 51-62, DOI: 10.5281/zenodo.175484
A7576C4EFFDFFFE58180960461DAFA51.taxon	description	Thorax. Glossy black; scutum overlain with brown pubescence, anterolateral corner of scutum and dorsocentral stripes of white­grey pubescence, admixed with long, erect setae; postalar callus glabrous; scutal macrosetae black; scutellum overlain with matte, black pubescence anteriorly, glabrous posteriorly; pleuron and coxae with dense, silver pubescence except on anepisternum, anepimeron and katatergite, admixed with long, pale setae on postpronotum, proepisternum, anepisternum, katatergite and coxae; legs dark brown, bases of tarsomeres and anterior surfaces of mid and hind tibia dark yellow; single row of dark macrosetae on ventral surfaces of mid and hind femora and on anteroventral surface of fore femur; wing hyaline with brown or black banding, bands across cells bm and d respectively, third band across subcostal and radial fields; venation brown in infuscate areas, dark yellow in hyaline areas; haltere crème­white colour; macrosetae difficult to discern from regular setation of scutum; scutal chaetotaxy (pairs): np, 3; sa, 1; pa, 1 – 2; dc, 2 – 4; sc, 1. Abdomen. Glossy brown­black with scattered, erect, pale setae; setae longer laterally; setae on terminalia dark; dense, silver, velutinous pubescence tergite 1 – 5 and epandrium; intersegmental membrane of segments 2 – 4 white; terminalia dark. Male genitalia. Epandrium large, quadrangular, length slightly shorter along midline than greatest width, posterolateral corners rounded (Fig. 4 A), extending laterally and ventrally to partially conceal gonocoxites; cercus greatly elongate; tergite 8 narrow, band­like with round lateral lobes (Fig. 4 B); sternite 8 broad with few setae along posterior margin (Fig. 4 E); gonocoxite (Figs 4 C – D) wedge­like in profile; gonocoxal process narrow, much shorter than inner gonocoxal process, elongate setae on posterior surface; medial atrium present with membranous area covered with velutum; hypandrium large, fused laterally to gonocoxites, large, posteriorly projecting setae along posterior margin; ventral lobe elongate, projecting dorsally, broadly club shape apically with velutum present at apex; gonostylus curved medially towards apex, erect setae on medial surface; inner gonocoxal process equal in length to gonostylus, curved medially towards apex, short setae on ventral surface of apex; gonocoxal apodeme approximately equal to length of gonocoxite; distiphallus straight (Figs 4 F – G), ventral apodeme of parameral sheath elongate, narrow, approximately equal in length to ejaculatory apodeme; dorsal apodeme of parameral sheath reduced; lateral ejaculatory apodeme small and tapered; ejaculatory apodeme narrowed anteriorly, projecting dorsally at apex. Female: Body length: 6.5 – 8.0 mm. Similar to male except: Frons wider than ocellar tubercle at narrowest point, extensively covered with brown pubescence dorsally; shorter setae on head, thorax and abdomen; abdominal tergites without velutum covering, silver­grey pubescence extensively on tergite 1. Female genitalia. Spermathecal sac present, spherical in shape; spermathecal sac duct thickened at common junction with spermathecal ducts; spermathecae spherical.	en	Winterton, Shaun L. (2007): Revision of the Australian stiletto fly genus Acatopygia Kröber (Diptera: Therevidae: Agapophytinae). Zootaxa 1405: 51-62, DOI: 10.5281/zenodo.175484
A7576C4EFFDFFFE58180960461DAFA51.taxon	discussion	Comments Acatopygia pulchella is the type species for the genus. It is readily differentiated from other species of Acatopygia based on furrowed eyes, row of macrosetae on the femora, brown legs and antennae, scutum colouration, and abdominal velutum covering and enlarged epandrium in the male. This species is known from central, eastern Australia and has been associated with coastal heath habitats. Mann (1929) noted that males of this species are found hovering in swarms in sandy habitats.	en	Winterton, Shaun L. (2007): Revision of the Australian stiletto fly genus Acatopygia Kröber (Diptera: Therevidae: Agapophytinae). Zootaxa 1405: 51-62, DOI: 10.5281/zenodo.175484
