taxonID	type	description	language	source
A15787C8FF80034153DB6B8976D09838.taxon	description	Figs 1 A – C Male. Colour. Head brown, face somewhat paler. Antenna brown, including scape and pedicel. Clypeus and mouthparts pale yellowish. Thorax light brown to brown. Legs yellowish to light brown, apical half of hind femur, and apical and basal thirds of mid femur infuscated. Wing with light brownish tinge because of microtrichia; halter yellowish with blackish knob. Abdomen brown, tergites laterally and sternites lighter. All vestiture pale, yellowish or brownish, thicker setae and trichia seeming darker than finer ones. Head. Antennal flagellomere 4 ca. 1.7 times as long as wide. Palpomere 3 of maxillary palpus with apicomesial thumb-like extension, with 4 apically curved sensilla; palpomere 4 with parasegment. Nine strong postocular setae. Thorax. Anepisternum with 68 setae; anterior basalare and laterotergite non-setose; preepisternum 2 with 25 setae; metepisternum with 17 setae. Legs. Mid and hind tibial organs absent. Wing. R 1 meeting C within basal half of costal margin; sclerotized part of M 2 not extending to level of tip of R 1; wing length 2.3 mm. Hypopygium (Figs 1 A – C). Sternite 9 narrow, about two times as long as wide, posterior ¾ laterally free from gonocoxa, anterior ¼ fused with gonocoxa, posterior margin with deep V-shaped incision medially, extending over middle between bases of gonocoxa and gonostylus, anterior margin with a deep V-shaped incision medially, anterior half non-setose, setae on posterior half similar to those on ventral side of gonocoxa. Ventral medial margin of gonocoxa angular. Parastylar lobe large, plate-like, medially drawn out to a finger like process, which has two strong setae apically. No paraapodemal lobe identifiable. Dorsal medial margin of gonocoxa simple, slightly bulging medially. Dorsal posterior margin simple, forming together with medial margin an acute angle with a dense aggregation of fine setae on medial side. On ventral side of dorsal posterior margin of gonocoxa a finger-like lobe bearing two strong apical setae. Two juxtagonostylar setae present, a normal seta and apically curved simple megaseta, both arising from a common basal body, which is about as long as the megaseta. Gonostylus subcircular, setose, setae similar to those on gonocoxa except for conspicuously long setae along apical margin; apicomedially with a small lobe bearing two dorsally turned megasetae at apex. Aedeagus subtriangular, with lateral shoulders, apex curved ventrad. Hypoproct extending posteriorly over basal part of gonostylus, the ventral setae strong and forming a mesial longitudinal stripe of 7 – 8 setae on each half. Cerci medially separated; setae delimited on the apical half of cercus. Female. Unknown.	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FF80034153DB6B8976D09838.taxon	materials_examined	Etymology. The species is named after its occurrence in the state of Burundi by adding the Latin suffix – ensis [denoting place]. Comments. Manota burundiensis sp. n. belongs to a group of six Afrotropical species as defined under M. triseta. Among them, M. burundiensis is most similar to M. relicina (see Hippa & Kurina 2012: fig. 20) as both species have the cercal setae short and delimited on the apical half of cercus, and the gonostylus with an apical or subapical medial lobe bearing two simple megasetae. It differs in having (1) one juxtagonostylar seta as a normal seta (both juxtagonostylar setae are megasetae in M. relicina), (2) the common basal body of the juxtagonostylar setae is about as long as setae (the basal body is considerably longer than juxtagonostylar megasetae in M. relicina), (3) parastylar lobe with two unusually strong setae at medial corner (with normal setae in M. relicina), (4) gonostylus apically rounded (apically transverse in M. relicina), and (5) dorsal posterior margin of the gonocoxa forming together with medial margin an acute angle with a dense aggregation of fine setae on medial side (with a row of closely placed setae in M. relicina). Types. Holotype. Male, BURUNDI, Kayanza Prov., Parc National de la Kibira, Rwecura sector, Indigenous Afromontane forest, 02 ° 55.320 ’ S 29 ° 30.067 ’ E, Malaise trap, 21 – 26. xi. 2010, A. H. Kirk-Spriggs leg. (slide mounted, BMSA).	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FF8E034153DB68F872E09E74.taxon	description	Figs 2 A – C Male. Colour. Head brown, face somewhat paler. Antenna light brown, including scape and pedicel. Clypeus and mouthparts pale yellowish. Thorax light brown. Legs yellowish to light brown, apical fifth of hind femur infuscated. Wing with light brownish tinge because of microtrichia; halter entirely brownish. Abdomen brown, tergites laterally and sternites lighter. All vestiture pale, yellowish or brownish, thicker setae and trichia seeming darker than finer ones. Head. Antennal flagellomere 4 ca. as long as wide. Palpomere 3 of maxillary palpus with apicomesial thumb-like extension, with 4 apically curved sensilla; palpomere 4 with parasegment; palpomere 5 1.4 – 1.5 times longer than palpomere 4. 10 - 11 strong postocular setae. Thorax. Anepisternum with 52 setae; anterior basalare and laterotergite non-setose; preepisternum 2 with 12 – 15 setae; metepisternum with 14 – 19 setae. Legs. Mid and hind tibial organs absent. Wing. R 1 meeting C within basal half of costal margin; sclerotized part of M 2 almost extending to level of tip of R 1; wing length 1.9 mm. Hypopygium (Figs 2 A – C). Sternite 9 laterally free from gonocoxa, posterior margin membranous, slightly concave, extending over middle between bases of gonocoxa and gonostylus, anterior margin with deep and wide V-shaped incision medially, anterior 1 / 3 and posterior membranous part nonsetose, setae on posterior 2 / 3 slightly shorter than those on ventral side of gonocoxa, postero-laterally with fingerlike sclerotizations on dorsal surface of both sides, which bear one short and curved apical seta deviating from other setosity on sternite 9. Ventral medial margin simple, membranous. Parastylar lobe plate-like, subrectangular, medially constricted, apically with 3 short and curved setae. Dorsal medial margin of gonocoxa simple, posteriorly forming well delimited setose bill-like small lobe with dorsal posterior margin of gonocoxa. Three juxtagonostylar setae present, all megaseta-like, the dorsal one apically slightly curved, with an apically separate basal body, the two dorsal ones more prominent, apically hook-like curved, with a common basal body which is more than half of the length of megasetae. Gonostylus dorso-ventrally wide, lateroapically, between dorsal and ventral portions concave; dorsal portion narrow, tapering with basal heel-like lobe, subbasal setose hump and 2 – 3 more prominent setae dorsoapically; ventral portion wide, with two blunt, medially directed megasetae apically and two more prominent setae subapically. Aedeagus elongate subtriangular, without lateral shoulders, apex curved ventrad. Hypoproct posteriorly not extending to the basal part of gonostylus, ventral setae (sternite 10) forming a medial longitudinal stripe of ca. 8 setae on each half. Cerci (scarcely discernible on slides) medially separated, largely membranous, with unusually narrow apical parts, bearing 4 – 5 setae. Female. Unknown.	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FF8E034153DB68F872E09E74.taxon	materials_examined	Etymology. The name is Latin, cornuta, ‘ horned’, referring to the horn-like setigerous lobe at posterolateral corners of sternite 9. Comments. Manota cornuta sp. n. is similar to M. afra and the two key out in the same couplet in the species key. Both species have three juxtagonostylar megasetae that is unique among the Afrotropical species. The unique finger-like posterolateral lobes on sternite 9 and the unusually narrow apical part of cercus, distinguishes M. cornuta from all known Manota. Types. Holotype. Male, GHANA, Volta region, Wli Falls, Afegame (St. 10 A), 7 – 10.03.1993, NUFU-ZMBN, leg. J. S. Amakye & J. Kjaerandsen (slide mounted, NHMO). Paratypes. Male, GHANA, Volta region, Wli Falls, Afegame (St. 8 A), 11 – 20.11.1993, NUFU-ZMBN, leg. J. Kjaerandsen (slide mounted, NHMO).	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FF8E034253DB6E3274AE9D20.taxon	description	Figs 3 A – D Male. Colour. Head brown, face somewhat paler. Antenna light brown, including scape and pedicel. Clypeus and mouthparts pale yellowish. Thorax yellowish to light brown. Legs yellowish to light brown. Wing with light brownish tinge because of microtrichia; halter entirely brownish. Abdomen brown, tergites laterally and sternites lighter. All vestiture pale, yellowish or brownish, thicker setae and trichia seeming darker than finer ones. Head. Antennal flagellomere 4 ca. 1.6 times as long as wide. Palpomere 3 of maxillary palpus with apicomesial thumb-like extension, with 4 apically curved sensilla; palpomere 4 with parasegment; palpomere 5 ca. 1.5 times longer than palpomere 4. Ten strong postocular setae. Thorax. Anepisternum with 56 setae; anterior basalare and laterotergite non-setose; preepisternum 2 with 20 setae; metepisternum with 11 setae. Legs. Mid and hind tibial organs absent.	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FF8E034253DB6E3274AE9D20.taxon	materials_examined	Wing. R 1 meeting C within basal half of costal margin; sclerotized part of M 2 almost extending to level of tip of R 1; wing length 1.6 mm. Hypopygium (Figs 3 A – D). Sternite 9 laterally free from gonocoxa, posterior margin straight, membranous, extending over middle between bases of gonocoxa and gonostylus, anterior margin with deep Vshaped incision medially, anterior 1 / 3 non-setose, setae on posterior 2 / 3 similar to those on ventral side of gonocoxa. Ventral medial margin simple, membranous. Parastylar lobe plate-like, rectangular, apical part with 3 setae. Dorsal medial margin of gonocoxa simple, posteriorly forming well delimited setose lobe with dorsal posterior margin of gonocoxa. Ventrally from dorsal posterior margin of gonocoxa a finger-like lobe bearing a strong apical seta. Two juxtagonostylar setae present, one is a rather unmodified seta, the other is a flattened apically pointed megaseta and twice as long as the former, both arising from a common basal body which is about as long as the unmodified seta. Gonostylus elongate, rectangular, apical third slightly bent medially, lateral margin folded dorsally at basal half, setae similar to these on gonocoxa except 2 – 3 apical setae and 2 setae on lateral margin which are stronger, deviating from others. Aedeagus subtriangular, without lateral shoulders, apex curved ventrad. Hypoproct extending posteriorly to base of gonostyli, ventral setae distributed only on apical fourth, number of ventral setae (sternite 10) ca. 10 on each half. Cerci medially separated, subapically constricted, with a row of apical setae only. Female. Unknown. Etymology. The name is Latin, fuscinula, ‘ small trident’, referring to the three strong setae apically on the gonostylus; noun in apposition. Comments. Manota fuscinula sp. n. belongs to a group of eight Afrotropical species as defined under M. limai. For further comments, see the latter species. Types. Holotype. Male, GHANA, G. Accra region, Legon, Botanical Garden, 1 – 3.12.1993, NUFU-ZMBN / leg. J. Kjaerandsen (slide mounted, NHMO).	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FF8D034453DB6D9074139AB0.taxon	description	Figs 4 A – D Male. Colour. Head brown, face somewhat paler. Antenna light brown, including scape and pedicel. Clypeus and mouthparts pale yellowish. Thorax light brown to brown. Legs yellowish, apical fourth of hind femur slightly infuscated. Wing with light brownish tinge because of microtrichia; halter yellowish with dark brown knob. Abdomen brown, tergites laterally and sternites lighter. All vestiture pale, yellowish or brownish, thicker setae and trichia seeming darker than finer ones. Head. Antennal flagellomere 4 ca. 1.7 times as long as wide. Palpomere 3 of maxillary palpus with apicomesial thumb-like extension, with 4 apically curved sensilla; palpomere 4 with parasegment; palpomere 5 not measurable in holotype. Ten strong postocular setae. Thorax. Anepisternum with 43 setae; anterior basalare with 5 setae, laterotergite with 21 setae, preepisternum 2 non-setose; metepisternum with 15 setae. Legs. Mid and hind tibial organs absent. Wing. R 1 meeting C within basal half of costal margin; sclerotized part of M 2 extending to level of tip of R 1; wing length 1.9 mm. Hypopygium (Figs 4 A – D). Sternite 9 laterally fused to gonocoxa, posterior margin semicircular, tongue-like, extending to base gonostylus, anterior margin with a shallow concavity, anterior 1 / 3 non-setose, posterior 2 / 3 setose, setae similar to those on ventral side of gonocoxa but arranged more densely. Ventral medial margin of gonocoxa oblique. Parastylar lobe plate-like, medially drawn out to a finger-like lobe that has an apical seta. No paraapodemal lobe identifiable. Dorsal medial margin of gonocoxa complex, posteriorly with a row of five flattened and apically pointed megasetae at medial margin and a single equal megaseta at posterolateral corner. At the dorsal medial margin of gonocoxa, on more ventral level a plate-lake submembranous lobe with two medially directed simple and blunt megasetae at anteromedial corner, and a posteriorly directed geniculate megasetae arising from a finger-like basal body. Gonocoxal apodeme large, polygonal. One juxtagonostylar seta present as a flattened and apically expanded megaseta arising from a curved basal body which is about five times longer than the megaseta. Gonostylus slightly less than half of the ventral length of gonocoxa, medially geniculate, basal half twice as wide as apical half, basal half ventrally setose, dorsally non-setose except for two strong setae, apical half with two strong setae at basal portion of medial margin and 4 – 5 apical setae, 2 – 3 of them stronger, deviating from others. Aedeagus elongated subtriangular, without lateral shoulders, apex curved ventrad. In holotype, aedeagal apodemes turned upward, pressed against the basal part of aedeagus. Hypoproct large, extending posteriorly near to apex of gonostylus, number of ventral setae (sternite 10) ca. 50 on each half, the setae indistinctly divided into an anterior and posterior groups with ca. 45 and 5 setae, respectively. Cerci medially separated, setae scattered over the surface, apical 4 – 5 setae about twice as long as the rest of cercal setae. Female. Unknown.	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FF8D034453DB6D9074139AB0.taxon	materials_examined	Etymology. The name is Latin, geniculata, ‘ like the bent knee’, referring to the geniculate gonostylus. Comments. Manota geniculata sp. n. belongs to a group of six Afrotropical species that have (1) setose laterotergite and anterior basalare, (2) the ventral setae of the hypoproct scattered over the whole ventral surface, (3) nonsetose preepisternum 2, and (4) the dorsal medial margin of gonocoxa posteriorly with a row of scale- or leaf-like megasetae. The other species being M. bracteata, M. foliolata, M. peltigera, M. pinnata and M. pinnulata. Manota geniculata, M. peltigera and M. pinnulata all have 3 flattened megasetae at the dorsal medial margin of gonocoxa posteriorly and a tongue-like posterior part of sternite 9. The latter character is shared also with M. foliolata, but that species has only two flattened megasetae posteriorly at the dorsal medial margin of gonocoxa. Of these three species, M. geniculata is most similar to M. pinnulata (cf. Hippa & Kurina 2012: fig. 18) but differs in having the gonostylus geniculate (subtriangular, widening from base to apex in M. pinnulata), and in having the megasetae posteriorly at the dorsal medial margin of gonocoxa comma-like, apically pointed, and becoming medially wider towards posterior end of the row (megasetae scale-like, apically widening and enlarging in size towards posterior end of the row in M. pinnulata). Types. Holotype. Male, GABON, Makoku M’Passa Bale affl., 200 m, J. Legrand rec., 28.05 – 5.06. 1979, Piège de Malaise (slide mounted, MNHN).	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FF8B034653DB6D0076D09B60.taxon	description	Figs 5 A – D Male. Colour. Head light brown, face somewhat paler. Antenna light brown, including scape and pedicel. Clypeus and mouthparts pale yellowish. Thorax yellowish. Legs yellowish, basal and apical fourths of hind femur infuscated. Wing with light brownish tinge because of microtrichia; halter yellowish with blackish knob. Abdomen brown, tergites laterally and sternites lighter. All vestiture pale, yellowish or brownish, thicker setae and trichia seeming darker than finer ones. Head. Antennal flagellomere 4 ca. 1.4 times as long as wide. Palpomere 3 of maxillary palpus with apicomesial thumb-like extension, with 4 apically curved sensilla; palpomere 4 with parasegment; palpomere 5 1.7 times as long as palpomere 4. Nine strong postocular setae. Thorax. Anepisternum with 58 setae; anterior basalare with 8 setae, laterotergite with 30 – 34 setae, preepisternum 2 non-setose; metepisternum with 14 setae. Legs. Mid and hind tibial organs absent. Wing. R 1 meeting C within basal half of costal margin; sclerotized part of M 2 extending to level of tip of R 1; wing length 1.9 – 2.2 mm. Hypopygium (Figs 5 A – D). Sternite 9 laterally fused to gonocoxa, posterior margin broad, convex, extending to base of gonostylus, anterior margin with a shallow and wide V-shaped concavity medially, anterior 1 / 3 non-setose, posterior 2 / 3 setose, setae similar to those on ventral side of gonocoxa. Ventral medial margin of gonocoxa short, simple. Parastylar lobe finger-like, tapering, with a stout and medially directed apical seta. No paraapodemal lobe identifiable. Dorsal medial margin of gonocoxa membranous, with a transverse conical lobe, bearing 5 – 6 pointed megasetae in marginal row posteriorly (lobe bend inwards in paratype). Dorsal posterior margin simple, forming together with medial margin an acute rounded angle. Setae on dorsal side of gonocoxa similar to those on ventral side. One juxtagonostylar seta present, a simple and pointed megaseta with finger-like basal body that is shorter than the megaseta. Gonostylus elongated, about 5 times as long as wide medially, with basal 1 / 3 slightly widened, setose, 2 – 3 apical setae somewhat longer and stronger, otherwise setae similar to those on ventral side of gonocoxa. Aedeagus elongate subtriangular, without lateral shoulders, apex curved ventrad. Hypoproct extending posteriorly to the middle of gonostylus, number of ventral setae (sternite 10) ca. 10 on each half. Cerci medially separated, setae scattered over the surface, apical setae somewhat longer, deviating from others. Female. Unknown.	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FF8B034653DB6D0076D09B60.taxon	materials_examined	Etymology. The species is named after Dr. Ashley Kirk-Spriggs (London, UK) in honour and recognition of his contribution to the study of Afrotropical Diptera. He is the collector of a number of specimens studied in this paper including both type specimens of M. kirkspriggsi sp. n. Comments. Manota kirkspriggsi sp. n. belongs to a group of 14 Afrotropical species as defined under M. polylobata. In having 4 – 6 pointed megasetae of subequal length at dorsal medial margin of the gonocoxa in marginal row on a transverse conical lobe, M. kirkspriggsi differs remarkably from all other species in this group. Types. Holotype. Male, MADAGASCAR, Province Fianarantsoa, Ranomafana National Park at: 21 ° 15.613 ’ S 47 ° 25.117 ’ E, 15 – 17. i. 2007, Malaise trap, A. H. Kirk-Spriggs leg. (slide mounted, AMGS). Paratype. Male, MAD- AGASCAR: Fianarantsoa Ranomafana National Park, Talatakely, 800 m SW entrance, 21 ° 15.48 ’ S 47 ° 25.27 ’ E; 16 – 19. x. 2014, 610 m, A. H. Kirk-Spriggs & R. Harin’Hala leg., Malaise trap secondary rainforest (slide mounted, BMSA).	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FF89035853DB6BD0735C98F0.taxon	description	Figs 6 A – C Male. Colour. Head brown, face somewhat paler. Antenna light brown, including scape and pedicel. Clypeus and mouthparts pale yellowish. Thorax light brown. Legs yellowish, apical third of hind femur and basal fourth of mid femur infuscated. Wing with brownish tinge because of microtrichia; halter entirely brownish. Abdomen brown, tergites laterally and sternites lighter. All vestiture pale, yellowish or brownish, thicker setae and trichia seeming darker than finer ones. Head. Antennal flagellomere 4 ca. 1.7 times as long as wide. Palpomere 3 of maxillary palpus with apicomesial thumb-like extension, with 4 apically curved sensilla; palpomere 4 subapically constricted, without parasegment; palpomere 5 ca. 1.6 – 1.7 times longer than palpomere 4. 8 – 9 strong postocular setae. Thorax. Anepisternum with 25 – 26 setae; anterior basalare with 5 – 6 setae; laterotergite with 19 setae; preepisternum 2 non-setose; metepisternum with 20 – 23 setae. Legs. Mid and hind tibial organs absent. Wing. R 1 meeting C within basal half of costal margin; sclerotized part of M 2 not extending to level of tip of R 1; wing length 1.6 – 1.7 mm. Hypopygium (Figs 6 A – C). Sternite 9 laterally fused to gonocoxa, posterior margin convex, extending over base of gonostylus, anterior margin with a narrow deep V-shaped incision medially, anterior half non-setose, setae on posterior half similar to those on ventral side of gonocoxa. Ventral medial margin of gonocoxa short, simple. Parastylar lobe finger-like with one strong apical seta. No paraapodemal lobe identifiable. Dorsal medial margin of gonocoxa medially simple, posteriorly drawn out into an apically rounded large lobe, covering basal third of gonostylus and bearing two subapical, medially directed megasetae: the posterior one short and pointed, the anterior one longer, with a whip-like apical part. Ventrally from dorsal medial margin of gonocoxa a large medially bulging plate-like lobe bearing a row of 6 simple and strong megasetae, 3 anteriors apically slightly tapering, 3 posteriors blunt, slightly widening apically. Two juxtagonostylar setae present, both are simple, apically pointed megasetae, arising from small separate basal bodies which are similar to normal sockets of setae. Gonostylus elongated, basal third heel-like bulging medially, with an extra-long apical seta and 8 – 9 longer setae along apical 2 / 3 of medial margin. Aedeagus elongate subtriangular, apical 2 / 3 almost parallel-sided, without lateral shoulders, apex curved ventrad. Hypoproct extending posteriorly to the middle of gonostylus, number of ventral setae (sternite 10) ca. 30 on each half. Cerci medially separated, setae scattered over the surface, apical setae deviating from others, longer than the subapical width of cercus. Female. Unknown.	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FF89035853DB6BD0735C98F0.taxon	materials_examined	Etymology. The species is named after Dr. Jostein Kjaerandsen (University of Tromsø, Norway), an active student of fungus gnats’ systematics and the collector of a number of the specimens from Ghana studied in this paper. Comments. Manota kjaerandseni sp. n. belongs to a group of 14 Afrotropical species that have (1) setose laterotergite and anterior basalare, (2) the ventral setae of the hypoproct scattered over the whole ventral surface, (3) non-setose preepisternum 2, and (4) the dorsal medial margin of gonocoxa posteriorly without a row of scale- or leaf-like megasetae. Among them, the new species is unique in having a row of 6 slender and blunt megasetae marginally on a bulging plate-like lobe at the dorsal medial margin of gonocoxa and in having the posterodorsal part of gonocoxa elongated and posteriorly rounded and with two megasetae on the medial margin. Types. Holotype. Male, GHANA, Volta region, Wli Falls, Afegame (St. 6 A), 9 – 12.03.1993, NUFU-ZMBN, leg. J. S. Amakye & J. Kjaerandsen (slide mounted, NHMO). Paratypes. Male, GHANA, Western region, Ankasa game prod. Reserve (St. 2), 06 – 12.12.1993, NUFU-ZMBN, leg. J. Kjaerandsen (slide mounted, NHMO); male, GHANA, Central Region, Kakum, Entwikrom, 8 – 15.10.1994, NUFU-project-ZMBN, leg. J. Kjaerandsen & T. Anderson (slide mounted, IZBE); male, same data except 8 – 15.11.1994 (slide mounted, IZBE); 2 males, CÔTE D’IVOIRE, rég. de Tai / 1 – 15. ii. 1985 / G. Gouturier & V. Van Zeijst Réc. // BIO (24) / Friche 6 ans / Piège malaise // ORSTOM-Paris / Mission UNESCO (slide mounted, MNHN). Additional material. 2 males, GHANA, Wli Falls, 4 – 13.03.1993, leg. J. Kjaerandsen (in alcohol, NHMO).	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FF97035853DB6B4077479E2C.taxon	description	Figs 7 A – D Male. Colour. Head brown, face somewhat paler. Antenna light brown to brown, including scape and pedicel. Clypeus and mouthparts pale yellowish. Thorax yellowish. Legs yellowish, basal third of hind femur infuscated. Wing with light brownish tinge because of microtrichia; halter yellowish with blackish knob. Abdomen brown to dark brown, tergites laterally and sternites lighter. All vestiture pale, yellowish or brownish, thicker setae and trichia seeming darker than finer ones. Head. Antennal flagellomere 4 ca. 1.8 times as long as wide. Palpomere 3 of maxillary palpus with apicomesial thumb-like extension, with 4 apically curved sensilla; palpomere 4 with parasegment; palpomere 5 ca. 1.5 times longer than palpomere 4. Ten strong postocular setae. Thorax. Anepisternum with 69 setae; anterior basalare with 13 setae, laterotergite with 32 setae, preepisternum 2 non-setose, metepisternum with 12 setae. Legs. Mid and hind tibial organs absent. Wing. R 1 meeting C within basal half of costal margin; sclerotized part of M 2 not extending to level of tip of R 1; wing length 2.2 mm. Hypopygium (Figs 7 A – D). Sternite 9 laterally fused to gonocoxa, posterior margin broad, convex, extending to base of gonostylus, anterior margin with a shallow and wide V-shaped concavity medially, anterior half non-setose, posterior half setose, setae similar to those on ventral side of gonocoxa. Ventral medial margin of gonocoxa short, simple. Parastylar lobe finger-like, tapering, with a stout and medially curved apical seta. No paraapodemal lobe identifiable. Dorsal medial margin of gonocoxa medially bulging, forming an obtuse lobe with dorsal posterior margin, dorsal surface of gonocoxa anteriorly and laterally setose, lateral setae similar to those on ventral side, anterior setae shorter and placed more densely, medial and posterior parts non-setose except two marginal setae posteromedially. Medioventrally from the dorsal medial margin a thumb-like lobe with a stout megaseta apically (in holotype that megaseta is broken in both sides). One juxtagonostylar seta present, a simple, apically whip-like megaseta arising from a finger-like basal body that is longer than the megaseta. Gonostylus bilobed: subrectangular lateral lobe about half of the gonocoxal height, with 7 – 8 strong setae along lateral and apical margins; subequal, medially directed and towards apex tapering lobe, with one basal and 2 apical setae along posterior margin and 2 more basal setae on dorsal surface. Aedeagus elongate subtriangular, without lateral shoulders, apex curved ventrad. Hypoproct extending posteriorly over gonostyli, number of ventral setae (sternite 10) ca. 30 on each half. Cerci medially separated, setae delimited on the apical part and medial margin, apical 4 setae somewhat longer, deviating from others. Female. Unknown.	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FF97035853DB6B4077479E2C.taxon	materials_examined	Etymology. The name is Latinized from the Greek λεπτóς (leptos) ‘ thin / slender / narrow’ and χαίτη (chaítç) ‘ bristle / seta’, referring to the considerably narrow juxtagonostylar megaseta. Comments. Manota leptochaeta sp. n. belongs to a group of 12 Afrotropical species as defined under M. platychaeta. Among these species M. leptochaeta is unique in having one megaseta on a lobe medially at dorsal medial margin of gonocoxa and gonostylus with a prominent medially directed lobe (all other species have at least two megasetae at the dorsal medial margin of gonocoxa or are without megasetae (M. edentula), and gonostylus without a prominent medial lobe). Types. Holotype. Male, MADAGASCAR: Tamatave Torotorofotsy, Andasibe (Perinet), 22 km NW, 18 ° 46,25 ’ S 48 ° 25,93 ’ E, 23 – 25. x. 2014, 960 m, leg. A. H. Kirk-Spriggs & R. Harin’Hala, Malaise trap primary rainforest (slide mounted, BMSA)	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FF95035A53DB69DB75CC9F2C.taxon	description	Figs 8 A – E Male. Colour. Head brown, face somewhat paler. Antenna light brown to brown, including scape and pedicel. Clypeus and mouthparts pale yellowish. Thorax light brown to brown. Legs yellowish, basal and apical fourths of hind femur and basal third of midfemur slightly infuscated. Wing with light brownish tinge because of microtrichia; halter yellowish with blackish knob. Abdomen brown to dark brown, tergites laterally and sternites lighter. All vestiture pale, yellowish or brownish, thicker setae and trichia seeming darker than finer ones. Head. Antennal flagellomere 4 1.1 – 1.2 times as long as wide. Palpomere 3 of maxillary palpus with apicomesial thumb-like extension, with 5 apically curved sensilla; palpomere 4 with parasegment; palpomere 5 1.5 – 1.6 times longer than palpomere 4. Nine strong postocular setae. Thorax. Anepisternum with 54 – 56 setae; anterior basalare and laterotergite non-setose, preepisternum 2 with 18 – 19 setae; metepisternum with 10 – 15 setae. Legs. Mid and hind tibial organs absent. Wing. R 1 meeting C within basal half of costal margin; sclerotized part of M 2 not extending to level of tip of R 1; wing length 2.2 – 2.3 mm. Hypopygium (Figs 8 A – E). Sternite 9 laterally free from gonocoxa, anterior margin with deep Ushaped incision medially, posterior margin transverse with V-shaped membranous area, extending over the middle between bases of gonocoxa and gonostylus, anterior 1 / 3 non-setose, setae on posterior 2 / 3 similar to those on ventral side of gonocoxa. Ventral medial margin of gonocoxa simple, oblique. Parastylar lobe elongate, subquadrangular, covered with conspicuous microtrichia and with 3 – 4 strong setae at anteromedial corner. Paraapodemal lobe large, apically rounded. Dorsal medial margin of gonocoxa simple. Dorsal posterior margin simple, forming together with medial margin a rounded acute angle with a dense aggregation of fine setae on medial side. Setae on dorsal side of gonocoxa similar to those on ventral side except for a few more prominent ones at posterolateral margin. On the ventral side of dorsal posterior margin (between this and the juxtagonostylar setae), a finger-like lobe bearing a strong apically curved seta at apex. Two juxtagonostylar setae present, one is a rather unmodified long pointed megaseta with a strongly hooked apex, the other an unmodified apically slightly curved megaseta of subequal length, both arising from a common basal body, which is shorter than the megasetae. Gonostylus subrectangular, with posteromedial part drawn out into a prominent rectangular lobe bearing two or three (both the holotype and the paratype are asymmetrical by having 3 megasetae on the left side and 2 on the right) simple, pointed and medially directed megasetae at apical margin, ventral surface of gonostylus setose, dorsal surface with three prominent setae medially, the microtrichia long. Aedeagus subtriangular, without lateral shoulders, apex curved ventrad. Hypoproct extending posteriorly to middle of gonostylus, the ventral setae (sternite 10) strong and forming a mesial longitudinal stripe of 4 setae on each half. Cerci medially separated, subapically constricted, setae confined to apex. Female. Unknown.	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FF95035A53DB69DB75CC9F2C.taxon	materials_examined	Etymology. The species is named after Dr. Ricardo Faustino de Lima (University of Lisbon, Portugal) in recognition of his contribution in organizing the Malaise trap sampling in São Tomé. Comments. The non-setose laterotergite, setose preepisternum 2 and a finger like lobe with one apical seta posterodorsally on gonocoxa group Manota limai sp. n. with eight other Afrotropical species, viz. M. issongo, M. joerni, M. mabokeensis, M. dissidens, M. freerki, M. toomasi, M. redunca and M. fuscinula. In having elongated rectangular gonostylus with strong setae or megasetae apically, M. limai resembles M. fuscinula, but can be distinguished in having one of juxtagonostylar megasetae apically hooked (straight or gently curved in M. fuscinula). Types. Holotype. Male, SÃO TOMÉ and PRINCIPE, São Tomé, Lagoa Amélia near Bom Sucesso, 00 ° 06 ° 35 ’ 27 ’’ E, 29. viii – 15. ix. 2016, Malaise trap, R. de Lima & L. Viegas leg. (slide mounted, IZBE). Paratype. Male, same data as holotype, except 15 – 31. xii. 2016 (slide mounted, IZBE).	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FF95035F53DB6FFB736E9D04.taxon	description	Figs 9 A – C Male. Colour. Head light brown, face somewhat paler. Antenna light brown, including scape and pedicel. Clypeus and mouthparts pale yellowish. Thorax yellowish to light brown. Legs yellowish to light brown, apical third of hind femur infuscated. Wing with light brownish tinge because of microtrichia; halter yellowish with blackish knob. Abdomen brown, tergites laterally and sternites lighter. All vestiture pale, yellowish or brownish, thicker setae and trichia seeming darker than finer ones. Head. Antennal flagellomere 4 ca. 1.3 times as long as wide. Palpomere 3 of maxillary palpus with apicomesial thumb-like extension, with 4 apically curved sensilla; palpomere 4 with parasegment. Nine strong postocular setae. Thorax. Anepisternum with 65 setae; anterior basalare and laterotergite nonsetose; preepisternum 2 with 13 setae; metepisternum with 9 setae. Legs. Mid and hind tibial organs absent. Wing. R 1 meeting C within basal half of costal margin; sclerotized part of M 2 not extending to level of tip of R 1; wing length 1.9 mm. Hypopygium (Figs 9 A – C). Sternite 9 laterally free from gonocoxa, posterior and anterior margins both with deep V-shaped incision medially, extending about middle between bases of gonocoxa and gonostylus, anterior half non-setose, setae on posterior half somewhat shorter than those on ventral side of gonocoxa. Ventral medial margin of gonocoxa angular. Parastylar lobe large, plate-like, triangular, with two setae at medial angle. No paraapodemal lobe identifiable. Dorsal medial margin of gonocoxa simple, convex. Dorsal posterior margin simple, forming together with medial margin a right angled lobe with a dense aggregation of fine setae on medial side. On ventral side of dorsal posterior margin of gonocoxa an elongated, subrectangular lobe, about twice as long as wide, with aggregation of fine setae at apex. One juxtagonostylar setae present, a simple, apically vasculiform megaseta, arising from a basal body, which is longer than the megaseta. Gonostylus subrectangular, apically transverse, with a very long seta deviating from other setosity at apicolateral corner, and with a short finger-like lobe subapically on medial side bearing two weak and simple apical megasetae. Aedeagus subtriangular, with lateral shoulders, apex curved ventrad. Hypoproct extending posteriorly over basal part of gonostylus, the ventral setae (sternite 10) strong and forming a mesial longitudinal stripe of 5 – 6 setae on each half. Cerci medially separated, setae delimited on the apical and medial margins of cercus. Female. Unknown.	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FF95035F53DB6FFB736E9D04.taxon	materials_examined	Etymology. The species is named after the mythical hero Oronna who defended the town Ilaro when it got under the threat of invasion by the Dahomean Army in the 17 th century. Comments. Because of the non-setose laterotergite, setose preepisternum 2 and gonocoxa posterodorsally with an apical setae bearing lobe, the new species groups together with 16 other Afrotropical species (see couplets 36 to 52 in the Key of the Afrotropical species above). Among them, M. oronnai sp. n. is unique as the lobe posterodorsally from the gonocoxa is subrectangular and with an aggregation of fine setae apically while all other species have this lobe finger-like and with 1 – 3 strong setae apically. In addition, the new species has a single apically vasculiform juxtagonostylar megaseta, while all other species have two juxtagonostylar setae, which are apically pointed. Types. Holotype. Male, W. NIGERIA, Ilaro Forest, 11. xi. 1973, M. A. Cornes (slide mounted, MNHN).	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FF90035153DB6DEC74319A64.taxon	description	Figs 10 A – D Male. Colour. Head brown, face somewhat paler. Antenna light brown, scape and pedicel somewhat paler. Clypeus and mouthparts pale yellowish. Thorax yellowish to light brown. Legs yellowish, basal third of hind femur infuscat- ed. Wing with light brownish tinge because of microtrichia; halter yellowish with blackish knob. Abdomen brown to dark brown, tergites laterally and sternites lighter. All vestiture pale, yellowish or brownish, thicker setae and trichia seeming darker than finer ones. Head. Antennal flagellomere 4 ca. 1.5 times as long as wide. Palpomere 3 of maxillary palpus with apicomesial thumb-like extension, with 3 apically curved sensilla; palpomere 4 subapically constricted, without parasegment; palpomere 5 ca. 1.4 times longer than palpomere 4. Eleven strong postocular setae. Thorax. Anepisternum with 63 setae; anterior basalare with 11 setae, laterotergite with 33 setae, preepisternum 2 non-setose, metepisternum with 14 setae. Legs. Mid and hind tibial organs absent. Wing. R 1 meeting C within basal half of costal margin; sclerotized part of M 2 extending to level of tip of R 1; wing length 2.2 mm. Hypopygium (Figs 10 A – D). Sternite 9 laterally entirely fused to gonocoxa, posterior margin broad, medially concave, lateral parts posteriorly bulging, extending to base of gonostylus, anterior margin with a V-shaped incision medially, anterior 1 / 3 non-setose, posterior 2 / 3 setose, setae similar to those on ventral side of gonocoxa but arranged more densely. Ventral medial margin of gonocoxa short, simple. Parastylar lobe finger-like, with one apical seta. No paraapodemal lobe identifiable. Dorsal medial margin of gonocoxa medially bulging, ventrally from the dorsal medial margin, on more ventral level a finger-like geniculate lobe with three slender, apically pointed megasetae apically. Dorsal posterior margin simple, forming together with medial margin a posteriorly projecting finger-like lobe, with 3 – 4 apical setae. Setae on dorsal side of gonocoxa similar to those on ventral side. One juxtagonostylar seta present, a complex spathulate megaseta with apical membranous leaf-like lobe dorsally, arising from a prominent basal body that is longer than the seta. Gonostylus elongated subquadrangular, about 5 times as long as wide medially, setose, at medial margin a submedial hump-like lobe with 2 – 3 apical setae, 3 apical and 1 apicolateral setae considerably stronger and longer, deviating from other setae on gonostylus. Aedeagus elongated, subtriangular, without lateral shoulders, apex curved ventrad. Hypoproct extending posteriorly to middle of gonostylus, number of ventral setae (sternite 10) ca. 30 on each half. Cerci medially separated, setae scattered over the surface, apical setae somewhat longer, deviating from others. Female. Unknown.	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FF90035153DB6DEC74319A64.taxon	materials_examined	Etymology. The name is Latinized from the Greek πλατύς (platys) ‘ wide / broad’ and χαίτη (chaítç) ‘ bristle / seta’, referring to the broad juxtagonostylar megaseta. Comments. Manota platychaeta sp. n. belongs to a group of 14 Afrotropical species that have setose laterotergite and anterior basalare, non-setose preepisternum 2, setae on hypoproct arranged over the whole ventral surface, and the dorsal medial margin of gonocoxa without a row of flattened megasetae posteriorly. Among these species, M. platychaeta is similar to M. styloides because the megasetae on the lobe at dorsal medial margin of gonocoxa are placed together in a group, not in a row. Manota platychaeta is distinguished e. g. by having the dorsal medial margin of gonocoxa posteriorly with a group of 3 slender megasetae (5 stout megasetae in M. styloides) and the juxtagonostylar megaseta is spathulate with an apical membranous leaf-like lobe (apically whip-like in M. styloides). Types. Holotype. Male, MADAGASCAR: Fianarantsoa, Ranomafana National Park, Talatakely, 800 m SW entrance, 21 ° 15,48 ’ S 47 ° 25,27 ’ E; 16 – 19. x. 2014, 610 m, leg. A. H. Kirk-Spriggs & R. Harin’Hala, Malaise trap secondary rainforest (slide mounted, BMSA).	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FF9E035353DB6ACC73B49B60.taxon	description	Figs 11 A – D Male. Colour. Head light brown, face somewhat paler. Antenna: scape and pedicel light brown, flagella absent in holotype. Clypeus and mouthparts pale yellowish. Thorax light brown to brown. Legs yellowish, apical fourth of hind femur slightly infuscated. Wing with light brownish tinge because of microtrichia; halter yellowish with blackish knob. Abdomen brown, tergites laterally and sternites lighter. All vestiture pale, yellowish or brownish, thicker setae and trichia seeming darker than finer ones. Head. Palpomere 3 of maxillary palpus with apicomesial thumb-like extension, with 4 apically curved sensilla; palpomere 4 with parasegment; palpomere 5 not measurable in holotype. Nine strong postocular setae. Thorax. Anepisternum with 40 setae; anterior basalare with 10 setae, laterotergite with 34 setae, preepisternum 2 non-setose; metepisternum with 21 setae. Legs. Mid and hind tibial organs absent. Wing. R 1 meeting C within basal half of costal margin; sclerotized part of M 2 extending to level of tip of R 1; wing length 2.2 mm. Hypopygium (Figs 11 A – D). Sternite 9 laterally fused to gonocoxa, posterior margin broad, convex, extending over middle between bases of gonocoxa and gonostylus, anterior margin with a deep and narrow V-shaped incision medially, anterior 1 / 3 non-setose, posterior 2 / 3 setose, setae similar to those on ventral side of gonocoxa. Ventral medial margin of gonocoxa oblique, laterally from gonostylus drawn into a suboval lobe about half of gonostylus length, bearing conspicuous microtrichia on ventral surface and two subapical twisted and apically whip-like megasetae. Parastylar lobe bifurcate, anterior branch short, smaller than posterior that has an apical seta. No paraapodemal lobes identifiable. Dorsal medial margin of gonocoxa simple with submembranous lobe on more ventral level, just anteriorly from juxtagonostylar setae, bearing row of three megasetae gradually diminishing in size posteriad: anterior megaseta flat and apically dilated, medial megaseta similar but apically somewhat narrower, posterior megaseta pointed. Gonocoxal apodeme unusual, posteriorly strongly widened. Setae on dorsal side of gonocoxa similar to those on ventral side. Two juxtagonostylar seta present, both short, flattened and expanded megasetae, arising from a common basal body which is shorter than megasetae. Gonostylus complex, with prominent medial lobe and two small lobes laterally; the medial prominent lobe longitudinally drawn out, about half as long as gonocoxa, apically conical with ca. 8 long marginal setae, medial margin with an aggregation of ca. 10 setae on ventral surface, dorsally with an additional medial lobe bearing a row of 6 – 7 setae; the lateral lobe subquadrangular with 6 – 8 small setae apically; the medial lobe finger-like with 4 apical setae. Aedeagus subtriangular, without lateral shoulders, apex curved ventrad. Hypoproct extending posteriorly near to apex of gonostylus, apical third tapering, number of ventral setae (sternite 10) ca. 40 on each half. Cerci medially separated, setae scattered over the surface, apical 4 – 5 setae about twice as long as rest of cercal setae. Female. Unknown.	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FF9E035353DB6ACC73B49B60.taxon	materials_examined	Etymology. The name is combined from Greek πολύ (polú) ‘ many’ and Latin lobata ‘ lobed, with lobes’, referring to the four-lobed gonostylus. Comments. Manota polylobata sp. n. belongs to a group of 14 Afrotropical species that have (1) setose laterotergite and anterior basalare, (2) the ventral setae of the hypoproct scattered over the whole ventral surface, (3) nonsetose preepisternum 2, and (4) the dorsal medial margin of gonocoxa posteriorly without a row of scale- or leaf-like megasetae (see couplets 21 to 33 in the Key of the Afrotropical species above). Among them, M. polylobata is most similar to M. tridactyla in having a complex gonostylus, two leaf-like juxtagonostylar megasetae, and megasetae at dorsal medial margin of the gonocoxa close to posterior end of the margin (should not to be confused with a distinct row of leaf- or scale like megasetae posteriorly on dorsal medial margin of gonocoxa as defined under M. geniculata). Manota polylobata differs in having 3 megasetae at dorsal medial margin of gonocoxa with the anteriormost megaseta apically dilated (4 megasetae with the anteriormost hook-like in M. tridactyla). Types. Holotype. Male, NIGERIA, SE State, Obudu CR, 27. ix. 1973, J. T. Medler coll. (slide mounted, MNHN).	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FF9C035553DB6BC6741598A8.taxon	description	Figs 12 A – C Male. Colour. Head brown, face somewhat paler. Antenna light brown, including scape and pedicel. Clypeus and mouthparts pale yellowish. Thorax yellowish. Legs yellowish to light brown, apical fourth to third of hind femur infuscated. Wing with light brownish tinge because of microtrichia; halter yellowish with brown knob. Abdomen brown, tergites laterally and sternites lighter. All vestiture pale, yellowish or brownish, thicker setae and trichia seeming darker than finer ones. Head. Antennal flagellomere 4 ca. 1.3 times as long as wide. Palpomere 3 of maxillary palpus with apicomesial thumb-like extension, with 4 apically curved sensilla; palpomere 4 with parasegment; palpomere 5 ca. 1.4 times as long as palpomere 4. 9 - 11 strong postocular setae. Thorax. Anepisternum with 52 – 57 setae; anterior basalare and laterotergite non-setose; preepisternum 2 with 13 – 17 setae; metepisternum with 15 – 16 setae. Legs. Mid and hind tibial organs absent. Wing. R 1 meeting C within basal half of costal margin; sclerotized part of M 2 not extending to level of tip of R 1; wing length 1.7 – 1.9 mm. Hypopygium (Figs 12 A – C). Sternite 9 laterally free from gonocoxa at posterior 2 / 3, anterior 1 / 3 fused with gonocoxa, posterior margin membranous, shallowly concave, extending about the middle between bases of gonocoxa and gonostylus, anterior margin with deep and wide V-shaped incision medially, anterior half non-setose, setae on posterior half similar to those on ventral side of gonocoxa. Ventral medial margin simple. Parastylar lobe large, plate-like, two setae anteriorly on ventral surface. Dorsal medial margin of gonocoxa simple, slightly convex, with tight aggregation of fine setae posteriorly on ventral surface. On the ventral side of the posteromedial angle a finger-like lobe bearing two apical setae. Two juxtagonostylar setae present, a normal seta and apically curved simple megaseta, both arising from a common basal body, which is about of the length of megaseta. Gonostylus elongated, about 5 times as long as wide, slightly curved, with a medial small hump-like lobe bearing 2 normal setae, and with 3 very long and medially directed setae apically, otherwise the gonostylar setosity is similar to that of gonocoxa. Aedeagus short, subtriangular, with prominent lateral shoulders, apex curved ventrad. Hypoproct posteriorly not reaching the base of gonostylus, the ventral setae strong and forming a mesial longitudinal stripe of ca. 5 setae on each half. Cerci medially separate, simple, the setae confined to apical part, the longest ones maximally two times longer than the apical width of cercus. Female. Unknown.	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FF9C035553DB6BC6741598A8.taxon	materials_examined	Etymology. The name is Latin, triseta, ‘ three setae’, referring to the three very long medially directed apical setae of gonostylus; noun in apposition. Comments. The non-setose laterotergite, setose preepisternum 2, and the gonocoxa posterodorsally with a fingerlike lobe with two apical setae group M. triseta sp. n. together to M. comata, M. pedicellata, M. relicina, M. petiolata and M. burundiensis. Among them, M. triseta is unique in having gonostylus apically or subapically without small lobe that bears two rather short megasetae. Otherwise, M. triseta is similar to M. issongo because of very long, medially directed setae apically on the gonostylus but distinguished in having (1) the finger-like lobe posterodorsally on the gonocoxa with two setae at apex (one seta in M. issongo), (2) the three very long setae apically on the gonostylus (two setae in M. issongo), (3) the two setae medially on the gonostylus arise from a distinct lobe (no distinct lobe discernible in M. issongo), (4) the two setae on the parastylar lobe separated by a distinct gap (two setae on the parastylar lobe close to each other in M. issongo), and (5) the ventral setae of hypoproct are in one row on each side, flanking the apex of aedeagus (setae on the hypoproct arranged as a patch in M. issongo). Types. Holotype. Male, GHANA, Volta region, Wli Falls, Agumatsa (St. 6 A), 12 – 21.11.1993, NUFU-ZMBN, leg. J. Kjaerandsen (slide mounted, NHMO). Paratypes. 4 males, as holotype (slide mounted, IZBE and NHMO); 3 males, as holotype except 11 – 20.11.1993 (slide mounted, IZBE and NHMO); 2 males, GHANA, Volta region, Wli Falls, Afegame (St. 9 A), 7 – 10.03.1993, NUFU-ZMBN / leg. J. Kjaerandsen (slide mounted, NHMO); male, GUINEA, Monts Nimba / DL-HP-JYR // Source 1200 M / Piége Malaise / 27 - iv – 7 - v- 1993 (slide mounted, MNHN); male, GUINEA, Mt. Nimba, Ft. Gal. du Zougué, 750 m, 18 – 29. VI. 1991, Girard et Legrand col., Piège de Malaise (slide mounted, MNHN). Additional material. 23 males, GHANA, Volta region; Wli Falls, Agumatsa; St: 11, 11 - 20.11.1993; NUFUproject ZMBN; leg. J. Kjaerandsen (1 slide mounted, 22 in alcohol, IZBE and NHMO); male same data except 13 – 20.11.1993 (in alcohol, NHMO).	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FF9A035553DB6B3C747B9A37.taxon	materials_examined	Material studied. GHANA: male, Central Region, Kakum, Entwikrom, 31.10 – 8.11.1994, NUFU-project-ZMBN, leg. J. Kjaerandsen & T. Anderson (slide mounted, NHMO); 2 males, same data, except 8 – 15.11.1994 (slide mount- ed, NHMO, IZBE). Remarks. The species has so far been known from Kakum National Park in Ghana and from Kibale National Park in Uganda (Hippa & Kurina 2012).	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FF9A035553DB6AEC73A09CE0.taxon	materials_examined	Material studied. GHANA: 2 males, Volta region, Wli Falls, Afegame (St. 6 A), 12 – 15.03.1993, NUFU-ZMBN, leg. J. Kjaerandsen (slide mounted, NHMO); male, Volta region, Wli Falls, Afegame (St. 8 B), 10 – 13.03.1993, NUFU-ZMBN, leg. J. S. Amakye & J. Kjaerandsen (slide mounted, IZBE); male, Volta region, Wli Falls, Afegame (St. 10 A), 7 – 10.03.1993, NUFU-ZMBN, leg. J. S. Amakye & J. Kjaerandsen (slide mounted, NHMO); male, Volta region, Wli Falls, Agumatsa (St. 10), 11 – 20.11.1993, NUFU-ZMBN, leg. J. Kjaerandsen (slide mounted, IZBE); male, same data except St: 9 – 11, 4 – 14.03.1993 (slide mounted, NHMO); 2 males, Volta region, Wli Falls, 4 – 13.03.1993, leg. J. Kjaerandsen (slide mounted, IZBE and NHMO); 2 males, Boti Falls, 28.10 – 4.11.1994 (slide mounted, IZBE); 3 males, Central Region, Kakum, Entwikrom, 31.10 – 8.11.1994, NUFU-project-ZMBN, leg. J. Kjaerandsen & T. Anderson (in alcohol, NHMO); 4 males, same data except 8 – 15.11.1994 (in alcohol, IZBE and NHMO). D. R. CONGO: male, Oriental Prov. Likombo forest, 2 km SW Bomane, 1.28349, 23.72358, 20 – 22. v. 2010, leg. A. H. Kirk-Spriggs, Malaise traps lowland evergreen primary forest (disturbed), BMSA (D) 58310 (slide mount- ed, BMSA). Remarks. The species was described from Uganda (Hippa & Kurina 2012) and has not been recorded since.	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FF9A035553DB6F45736F9F94.taxon	materials_examined	Material. MADAGASCAR: male, Tamatave Torotorofotsy, Andasibe (Perinet), 22 km NW, 18 ° 46,25 ’ S 48 ° 25,93 ’ E; 23 – 25. x. 2014, 960 m, leg. A. H. Kirk-Spriggs & R. Harin’Hala, Malaise trap primary rainforest (slide mounted, BMSA); male, Perinet, 26. iv – 4. v. 1983, M. C. Day & J. S. Noyes leg. (slide mounted, NHM). Remarks. The species has so far been known only from the type locality in Madagascar (Hippa 2008).	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FF9A035453DB6E5177349B79.taxon	materials_examined	Material. CÔTE D’IVOIRE: male, rég. de Tai, 1 – 15. ii. 1985, G. Gouturier & V. Van Zeijst Réc., BIO (24), Friche de 6 ans, Piége malaise, ORSTOM-Paris, Mission UNESCO (slide mounted, MNHN). Remarks. The species was described from Uganda (Hippa & Kurina 2012) and has not been recorded since. Manota comata Hippa & Kurina, 2012 Material. GHANA: 9 males, Volta region, Wli Falls, Agumatsa (St. 10), 11 – 20.11.1993, NUFU-ZMBN, leg. J. Kjaerandsen (slide mounted, IZBE and NHMO); 6 males, Central Region, Kakum, Entwikrom, 31.10 – 8.11.1994, NUFU-project-ZMBN, leg. J. Kjaerandsen & T. Anderson (3 slide mounted, IZBE, 3 in alcohol, NHMO); 5 males, same data except 8 – 15.11.1994 (2 slide mounted NHMO, 3 in alcohol, NHMO); male, Western region, Ankasa game prod. Reserve, 6 – 12. XII. 1993 (St: 1 – 8, 13), NUFU-project ZMBN, leg. J. Kjaerandsen & T. Andersen (in alcohol, NHMO). GUINEA: male, Mt. Nimba, Ft. Gal. du Zougué, 750 m, 18 – 29. vi. 1991, Piège de Malaise, Girard et Legrand col. (slide mounted, MNHN). D. R. CONGO: male, Oriental Prov., Likombo forest, 2 km SW, Bomane, 1.28349, 23.72358, 20 – 22. v. 2010, leg. A. H. Kirk-Spriggs, Malaise traps lowland evergreen primary forest (disturbed), Boyekoli Ebale Congo Expedition 2010, BECE 00466 (slide mounted, BMSA). UGANDA: male, Western region, Budongo Forest; N 01 ° 43 ’ 18,9 ’’ E 031 ° 31 ’ 46,7 ’’ (Site 9) 4 – 8 July 2010 G. Kvifte leg., Malaise trap (slide mounted, NHMO). Remarks. The species has so far been known from the type locality in Ghana and from Uganda (Hippa & Kurina 2012).	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FF9B035453DB6B3876C79A19.taxon	materials_examined	Material. MADAGASCAR: 2 males, Mahajanga, Boeny, Ankarafantsika NP, 16.30268 S 46.81041 E, 58 m, Malaise trap, # MAD 09 - 38, 9. xii. 2009 – 5. i. 2010, Bergsten, J. et. al. leg. (slide mounted, NHRS-BYWS 000000732). Remarks. The species has so far been known only by type material from two localities in Madagascar (Hippa 2008).	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FF9B035453DB6AD8746F9DA9.taxon	materials_examined	Material. GHANA: 10 males, Central Region, Kakum, Entwikrom, 31.10 – 8.11.1994, NUFU-project-ZMBN, leg. J. Kjaerandsen & T. Anderson (2 slide mounted, IZBE; 8 in alcohol, NHMO); 27 males, same data except 8 – 15.10.1994, leg. J. Kjaerandsen & T. Anderson (2 slide mounted, IZBE; 25 in alcohol, NHMO). CÔTE D’IVOIRE: 2 males, rég. de Tai, 16. i. - 1. ii. 1985, G. Gouturier & V. Van Zeijst Réc., BIO (21), (Gilbert), Friche 17 ans, Piége malaise, ORSTOM-Paris, Mission UNESCO (slide mounted, MNHN); male, same data except 1 – 15. ii. 1985 (slide mounted, MNHN); male, same data except 7 – 11. ii. 1985, BIO (25) Forêt noire station (slide mounted, MNHN). Remarks. The species has so far been known only from the type locality in Ghana: Central region, Kakum National Park (Hippa & Kurina 2012).	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FF9B035453DB6C0873A09F49.taxon	materials_examined	Material. CAMEROON: male, Northwest Reg. Mezam, Bafut village at: 06 ° 05,026 ’ N 10 ° 07,442 ’ E, 17 – 19. viii. 2013, 1060 m, leg. A. H. Kirk-Spriggs, Malaise traps cultivated plots & degraded forest, BMSA (D) 58256 (slide mounted, BMSA). Remarks. The species was described from Uganda (Hippa & Kurina 2012) and has not been recorded since.	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FF9B035453DB6F2874E29E05.taxon	materials_examined	Material. GHANA: 5 males, Central Region, Kakum, Entwikrom, 31.10 – 8.11.1994, NUFU-project-ZMBN, leg. J. Kjaerandsen & T. Anderson (1 slide mounted, IZBE; 4 in alcohol, NHMO); 4 males, same data, except 8 – 15.11.1994 (3 slide mounted, IZBE; 1 in alcohol, NHMO).	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FF9B035453DB6F2874E29E05.taxon	discussion	Remarks. The species has so far been known only from the holotype collected from exactly the same locality as our studied specimens (Hippa & Kurina 2012).	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FF98035753DB6936779498D3.taxon	materials_examined	Material. D. R. CONGO: male, Oriental Prov, Eyolo forest, ca 2 km E, Lieki, 0.67642, 24.24186, 25 – 29. v. 2010, leg. A. H. Kirk-Spriggs, Malaise traps lowland evergreen swamp forest, Boyekoli Ebale Congo Expedition 2010, BECE 02270 (slide mounted, BMSA).	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FF98035753DB6936779498D3.taxon	discussion	Remarks. The species has so far been known only from the holotype collected from Yangambi NP in D. R. Congo, North from Congo River (Kurina & Hippa 2014). The studied specimen was collected from same area but South from Congo River.	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FF98035753DB689E77349A3F.taxon	materials_examined	Material. GHANA: male, Volta region, Wli Falls, Afegame (St. 2 A), 11 – 14.03.1993, NUFU-ZMBN, leg. J. Kjaerandsen (slide mounted, NHMO); male, Volta region, Wli Falls, Agumatsa (St. 6), 11 – 20.11.1993, NUFU- ZMBN leg. J. Kjaerandsen. (slide mounted, NHMO); male, Central Region, Kakum, Entwikrom, 31.10 – 8.11.1994, NUFU-project-ZMBN, leg. J. Kjaerandsen & T. Anderson (slide mounted, IZBE). UGANDA: male, Western region, Budongo Forest / N 01 ° 43 ’ 04,2 ’’ E 031 ° 32 ’ 46,2 ’’ (Site 16) / 9 – 14 July 2010 G. Kvifte leg. (Malaise trap) (slide mounted, NHMO). GUINEA: male, Mt. Nimba, Girard et Legrand col., Ft. Gal. du Gouan, 500 m, 19 – 26. V. 1991, Piège de Malaise (slide mounted, MNHN). Remarks. The species has so far been known from the type locality in Uganda and from Ghana (Hippa & Kurina 2012).	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FF98035753DB6AF275E39DDF.taxon	materials_examined	Material. D. R. CONGO: male, Gal. for. De la Djoumouna, Yakayaka piège adh. Lum., xii. 1977, J. Trouillet leg. (slide mounted, MNHN). Remarks. The species was described from the Côte d’Ivoire (Matile 1972) and subsequently recorded from Uganda and D. R. Congo (Hippa & Kurina 2012, Kurina & Hippa 2014).	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FF98035753DB6D9272C19F53.taxon	materials_examined	Material. D. R. CONGO: male, Oriental Prov. Likombo forest, 2 km SW Bomane, 1.28349, 23.72358; 20 – 22. v. 2010, A. H. Kirk-Spriggs, Malaise traps lowland evergreen primary forest (disturbed), BMSA (D) 58310 (slide mounted, BMSA). UGANDA: male, Semuliki NP, 0 ° 50 ’ 8 ’’ N 30 ° 9 ’ 46 ’’ E, 670 m, Malaise trap, 9 – 15. ii. 2013, F. Molleman et al. leg. (slide mounted, IZBE); male, same data except 15 – 22. ii. 2013 (slide mounted, IZBE); male, Bwindi NP, 01 ° 03 ’ N 29 ° 43 ’ E, Malaise trap (No 1), 10 – 25. iii. 2014, F. Molleman et al. leg. (slide mounted, IZBE). Remarks. The species was described from the Central African Republic (Matile 1972) and subsequently recorded from Uganda and D. R. Congo (Hippa 2008, Hippa & Kurina 2012, Kurina & Hippa 2014).	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FF98035753DB6F1E72A29EC7.taxon	materials_examined	Material. MADAGASCAR: male, Fianarantsoa, Ranomafana National Park, Talatakely, 800 m SW entrance, 21 ° 15,48 ’ S 47 ° 25,27 ’ E, 16 – 19. x. 2014, 610 m, leg. A. H. Kirk-Spriggs & R. Harin’Hala, Malaise trap secondary rainforest (slide mounted, BMSA); 4 males, Tamatave Torotorofotsy, Andasibe (Perinet), 22 km NW, 18 ° 46,25 ’ S 48 ° 25,93 ’ E; 23 – 25. x. 2014, 960 m, leg. A. H. Kirk-Spriggs & R. Harin’Hala, Malaise trap primary rainforest (slide mounted, BMSA and IZBE). Remarks. The species has so far been known only from the holotype collected from Province Fianarantsoa in Madagascar (Hippa 2008). The current records indicate a wider distribution on the Island.	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FF99035653DB69DB77D39AF8.taxon	description	Figs 13 A – C	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FF99035653DB69DB77D39AF8.taxon	materials_examined	Material. GUINEA: 3 males, Monts Nimba, DL-HP-JYR, Source 1200 m, Piége Malaise, 27 - iv – 7 - v- 1993 (slide mounted, MNHN); male, Mont Nimba, 27. iii. 1993, 1220 m, D. Lachaise leg. (slide mounted, MNHN); 2 males, Ft. Gal. du Zougué, 750 m, 18 – 29. vi. 1991, Piège de Malaise, Girard et Legrand col. (slide mounted, MNHN); male, same dada except 3 – 17. vi. 1991 (slide mounted, MNHN). GHANA: 10 males, Central Region, Kakum, Entwikrom, 31.10 – 8.11.1994, NUFU-project-ZMBN, leg. J. Kjaerandsen & T. Anderson (3 slide mounted, IZBE; 7 in alcohol, NHMO); 4 males, same data except 8 - 15.11.1994 (3 slide mounted, NHMO; 1 in alcohol, IZBE). UGANDA: male, Western region, Budongo Forest, N 01 ° 43 ’ 04,2 ’’ E 031 ° 32 ’ 46,2 ’’ (Site 16), 9 – 14 July 2010, leg. G. Kvifte (Malaise trap) (slide mounted, NHMO). Remarks. In several studied specimens from Guinea and Ghana the preepisternum 2 has 2 – 4 setae, while other specimens have none (most common) or 1 seta. There are minute dissimilarities in the male terminalia, here illustrated by a Guinean specimen (Figs 13 A – C), but we regard them to be within infraspecific variation, and may even be caused by slightly different positions of the hypopygia in the slide mounts. Variability in male terminalia of M. mazumbaiensis is depicted also by Hippa and Kurina (2012: fig 28). Manota mazumbaiensis is one of the most widely distributed Manota species in Africa. Having been described from Usambara Mountains in Tanzania (Søli 1993), the species has subsequently been recorded from Uganda, D. R. Congo and Ghana (Hippa & Kurina 2012, Kurina & Hippa 2014), and now also from Nimba Mountains in Guinea. However, such an extensive distribution, and variation in male terminalia may eventually point to a complex of species impossible to distinguish on the basis of the morphology only. Additional studies, including that of DNA sequences from freshly collected material are needed for further conclusions.	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FF99035653DB6ABB77569FBC.taxon	materials_examined	Material. REPUBLIC OF SOUTH AFRICA: male, Western Cape, Bloukrantz, Platbos forest at: 33 ° 56.558 ’ S 23 ° 37.566 ’ E, 26 – 28. iii. 2009, 300 m, A. H. Kirk-Spriggs & S. Otto, Malaise traps (2), indigenous forest, BMSA (D) 08010 (slide mounted, BMSA); male, same data, except BMSA (D) 07989 (slide mounted, BMSA); 3 males, Western Cape, Keurbos forest, 500 m, 33 ° 54.435 ’ S 23 ° 43.714 ’ E, 28 – 30. iii. 2009, A. H. Kirk-Spriggs & S. Otto, Malaise trap indigenous montane forest, BMSA (D) 08307 (slide mounted, BMSA); male, KNZ, Royal Natal N. P., Thendele, 1600 m, 28 ° 42.378 ’ S 23 ° 56.083 ’ E, 15 – 17. ii. 2010, A. H. Kirk-Spriggs, Malaise trap Protea caffra wetland gully, BMSA (D) 20745 (slide mounted, BMSA); 2 males, E. Cape, Grahamstown (plot 5280), Three Chimneys farm, 33 ° 18.542 ’ S 26 ° 29.846 ’ E, 2 – 13. iii. 2008, A. H. Kirk-Spriggs, Malaise trap (slide mounted, AMGS); male, E. Cape, Grahamstown, Albany Museum grounds, 33 ° 18.822 ’ S 26 ° 31.315 ’ E, 9 – 15. x. 2007, A. H. Kirk- Spriggs, Malaise trap (slide mounted, AMGS); male, Eastern Cape, Plaatbos forest at: 33 ° 57.863 ’ S 23 ° 54.484 ’ E, 31. iii – 1. iv. 2009, 300 m, A. H. Kirk-Spriggs & S. Otto, Malaise traps (2) / indigenous forest, BMSA (D) 06881 (slide mounted, BMSA); male, Eastern Cape Prov., Lower Elandsbos River at: 33 ° 58.007 ’ S 23 ° 46.492 ’ E, 3 – 5. iv. 2008, A. H. Kirk-Spriggs, Malaise trap, indigenous forest (slide mounted, AMGS); 2 males, Tsitsikamma N. P., Plaatbos Nat. Reserve at: 33 ° 59.137 ’ S 23 ° 54.895 ’ E, 20 – 22. i. 2009, A. H. Kirk-Spriggs & S. Otto, Malaise traps (2), indigenous forest, BMSA (D) 02641 and BMSA (D) 02622 (slide mounted, BMSA); male, Cape province, Tsitsikamma Nat. Park., Malaise trap, 19. x – 13. xi. 1994, Allen, J. leg. (slide mounted, NHRS-BYWS 000000734); 6 males, same data except 21. ii – 20 iii. 1995 (2 slide mounted, 4 in alcohol, NHRS-BYWS 000000736).	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FF99035653DB6ABB77569FBC.taxon	discussion	Remarks. The species is previously known from several localities in South Africa (Jaschhof & Mostovski 2006, Hippa 2008).	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FFA7036853DB693F74769813.taxon	materials_examined	Material. GHANA: male, Central Region, Kakum, Entwikrom, 31.10 – 8.11.1994, NUFU-project-ZMBN, leg. J. Kjaerandsen & T. Anderson (slide mounted, NHMO).	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FFA7036853DB693F74769813.taxon	discussion	Remarks. The species has so far been known only from the holotype collected from Yangambi National Park in D. R. Congo (Kurina & Hippa 2014).	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FFA7036853DB68DF75C99BEB.taxon	materials_examined	Material. GHANA: male, Volta region, Wli Falls, Afegame (St. 10 A), 4 – 7.03.1993, NUFU-ZMBN, leg. J. S. Amakye & J. Kjaerandsen (slide mounted, NHMO); male, Volta region, Wli Falls, Agumatsa (St. 12), 12 – 21.11.1993, NUFU-ZMBN, leg. J. Kjaerandsen (slide mounted, IZBE). UGANDA: male, Western region, Budongo Forest, N 01 ° 43 ’ 04,2 ’’ E 031 ° 32 ’ 46,2 ’’ (Site 6), 1 – 4 July 2010, G. Kvifte leg. (Malaise trap) (slide mounted, NHMO). Remarks. The species was recently described from the material collected from Garamba and Yangamby National Parks in D. R. Congo (Kurina & Hippa 2014), and not recorded since.	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FFA7036853DB6A47741F9AA7.taxon	materials_examined	Material. GHANA: male, Central Region, Kakum, Entwikrom, 31.10 – 8.11.1994, NUFU-project-ZMBN, leg. J. Kjaerandsen & T. Anderson (in alcohol, NHMO); 3 males, same data except 8 – 15.11.1994 (slide mounted, IZBE and NHMO). Remarks. The species has so far been known from the type locality in Uganda and from D. R. Congo (Hippa & Kurina 2012, Kurina & Hippa 2014).	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FFA7036853DB6D0B72299CF7.taxon	materials_examined	Material. MADACASCAR: male, Limite forêt / cultures, 3.03.1988, M. Donskoff, Piège de Malaise (slide mount- ed, MNHN); male, Manjakatompo, xii. 1951, N. S. H. Krauss (slide mounted, MNHN); 3 males, Fianarantsoa, Ranomafana National Park, Talatakely, 800 m SW entrance; 21 ° 15,48 ’ S 47 ° 25,27 ’ E; 16 – 19. x. 2014, 610 m, leg. A. H. Kirk-Spriggs & R. Harin’Hala; Malaise trap secondary rainforest (slide mounted, BMSA and IZBE); 4 males, Tamatave Torotorofotsy, Andasibe (Perinet), 22 km NW; 18 ° 46,25 ’ S 48 ° 25,93 ’ E, 23 – 25. x. 2014, 960 m, leg. A. H. Kirk-Spriggs & R. Harin’Hala, Malaise trap primary rainforest (slide mounted, BMSA and IZBE). Remarks. The species has so far been known only from the holotype collected from northern Madagascar (Hippa 2008). The current records indicate to a wider distribution in the Island.	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FFA7036B53DB6CBB741F9819.taxon	materials_examined	Material. GHANA: male, Gr. Accra Region, Accra, Airpost res. Area. Inst. Aq. Biology, 30.10 – 6.11.1992, NUFU- ZMBN, leg. J. S. Amakye & J. Kjaerandsen (slide mounted, NHMO); male, same data except 06 – 13.11.1992 (slide mounted, IZBE); male, Volta region, Wli Falls, Afegame (St. 11 A), 7 – 10.03.1993, NUFU-ZMBN, leg. J. S. Amakye & J. Kjaerandsen (slide mounted, NHMO); male, same data except 10 – 13.03.1993 (slide mounted, IZBE); 5 males, Volta region, Wli Falls, Agumatsa (St. 10), 11 – 20.11.1993, NUFU-ZMBN, leg. J. Kjaerandsen (slide mounted, IZBE and NHMO); male, same data, except St: 9 – 8, 4 – 14.03.1993 (slide mounted, IZBE); male, Volta region, Wli Falls, 4 – 13.03.1993, leg. J. Kjaerandsen (slide mounted, NHMO); male, Northern Region, River Pru, Asubende, 17 – 19.06.1993, J. Amakye leg. (slide mounted, NHMO); male, Eastern Region, small stream near Akropong / Mamfe, 31.10.1993, J. Kjaerandsen leg. (slide mounted, NHMO); 16 males, Central Region, Kakum, Entwikrom, 31.10 – 8.11.1994, NUFU-project-ZMBN, leg. J. Kjaerandsen & T. Anderson (3 slide mounted, IZBE; 13 in alcohol NHMO); 30 males, same data except 8 – 15.11.1994 (4 slide mounted, IZBE; 26 in alcohol, NHMO). GUINEA: male, Mt. Nimba, Girard et Legrand col., Ft. Gal. du Zougué, 750 m, 3 – 17. VI. 1991, Piège de Malaise (slide mounted, MNHN). CÔTE D’IVOIRE: male, rég. de Tai, 16. i. - 1. ii. 1985, G. Gouturier & V. Van Zeijst Réc., BIO (21), (Gilbert), Friche 17 ans, Piege malaise, ORSTOM-Paris, Mission UNESCO (slide mounted, MNHN); male, same data except 1 – 15. ii. 1985 (slide mounted, MNHN). NIGERIA: male, Lagos, 3454, Koyi, 3. ii. 1974, M. A. Cornes (slide mounted, MNHN).	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FFA7036B53DB6CBB741F9819.taxon	discussion	Remarks. The species has so far been known from the type locality in Uganda and from D. R. Congo (Hippa & Kurina 2012, Kurina & Hippa 2014).	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FFA4036B53DB68D873AE9B39.taxon	materials_examined	Material. D. R. CONGO: male, Oriental Prov, Eyolo forest, ca 2 km E, Lieki, 0.67642, 24.24186, 25 – 29. v. 2010, leg. A. H. Kirk-Spriggs, Malaise traps lowland evergreen swamp forest, Boyekoli Ebale Congo Expedition 2010, BECE 02304 (slide mounted, BMSA). Remarks. The species has so far been known only from the type locality in Uganda (Hippa & Kurina 2012).	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FFA4036B53DB6BF873AE9A61.taxon	materials_examined	Material. GHANA: male, Boti Falls, 28.10 – 4.11.1994 (slide mounted, NHMO). Remarks. The species has so far been known only from the type locality in Uganda (Hippa & Kurina 2012).	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FFA4036B53DB6AD072DB9FC1.taxon	description	Figs 14 A – D	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FFA4036B53DB6AD072DB9FC1.taxon	materials_examined	Material. GHANA: 4 males, Volta region, Wli Falls, Afegame (St. 10 A), 3 – 7.03.1993, NUFU-ZMBN, leg. J. S. Amakye & J. Kjaerandsen (slide mounted, IZBE and NHMO); male, same data except 4 – 7.03.1993 (slide mounted, NHMO); male, same data except 6 – 9.03.1993 (in alcohol, NHMO); 2 males, same data except 9 – 12.03.1993 (in alcohol, NHMO); 2 males, same data except 7 – 10.03.1993 (1 slide mounted, IZBE; 1 in alcohol, NHMO); 4 males, same data except 10 – 13.03.1993 (3 slide mounted, NHMO; 1 in alcohol, IZBE); 2 males, same data except 12 – 15.03.1993 (slide mounted, NHMO); male, same data except 11.03.1993, light trap (in alcohol, NHMO); 2 males, Volta region, Wli Falls, Agumatsa (St. 10), 11 – 20.11.1993, NUFU-ZMBN, leg. J. Kjaerandsen (slide mounted, NHMO); male, Central Region, Kakum, Entwikrom, 31.10 – 8.11.1994, NUFU-project-ZMBN, leg. J. Kjaerandsen & T. Anderson (slide mounted, IZBE); 3 males, same data except 8 – 15.10.1994, leg. J. Kjaerandsen & T. Anderson (2 slide mounted, IZB; 1 in alcohol, NHMO); 2 males, Volta region, Wli Falls, 4 – 13.03.1993, leg. J. Kjaerandsen (1 slide mounted, NHMO; 1 in alcohol, IZBE). CÔTE D’IVOIRE: male, rég. de Tai, 1 – 15. ii. 1985, G. Gouturier & V. Van Zeijst Réc., BIO (24), Friche 6 ans, Piège malaise // ORSTOM - Paris, Mission UNESCO (slide mounted, MNHN); male, same data except 7 – 11. ii. 1985, BIO (25) Forêt noire station (slide mounted, MNHN). GUINEA: 3 males, Ft. Gal. du Zougué, 750 m, 18 – 29. vi. 1991, Piège de Malaise, Girard et Legrand col. (2 slide mounted, IZBE and MNHN; 1 pinned, MNHN); 2 males, same data, except 3 – 17. vi. 1991 (pinned, MNHN). Remarks. The species is described from the male holotype collected from Kona on Itimbti river in D. R. Congo and not recorded since (Kurina & Hippa 2014). The study of more extensive material disclosed a small variation in male terminalia (Figs 14 A – D), especially in details of the gonostylus and the setigerous lobe dorsally from juxtagonostylar megasetae, which are well exposed in the slide mounts of the newly studied specimens.	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FFA4036B53DB6FB073AE9EC5.taxon	materials_examined	Material. UGANDA: mal e, Bwindi NP, 01 ° 03 ’ N 29 ° 43 ’ E, Malaise trap (No 2), 10 – 25. iii. 2014, F. Molleman et al. leg. (slide mounted, IZBE).	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FFA4036B53DB6FB073AE9EC5.taxon	discussion	Remarks. The species has so far been known only from the type locality in Uganda (Hippa & Kurina 2012).	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FFA2036D53DB693F741F9B5F.taxon	materials_examined	Material. GHANA: male, Central Region, Kakum, Entwikrom, 8 – 15.10.1994, NUFU-project-ZMBN, leg. J. Kjaerandsen & T. Anderson (slide mounted, NHMO). GABON: male, Makoku M’Passa, Bale Affl. 200 m, J. Legrand rec., Piège de Malaise, 28.05 – 5.06.1979 (slide mounted, MNHN); male, same data except 18 – 25.06.1979 (slide mounted, MNHN). GUINEA: 2 males, Mt. Nimba, Girard et Legrand col., Ft. Gal. du Zougué, 750 m, 3 – 17. VI. 1991, Piège de Malaise (slide mounted, MNHN); 1 male, same data except 18 – 24. VI. 1991 (slide mount- ed, MNHN). CENTRAL AFRICAN REPUBLIC: male, La Maboke, 29. VIII. 1970, L. Matile rec. (slide mounted, MNHN).	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FFA2036D53DB693F741F9B5F.taxon	discussion	Remarks. The species has so far been known from the type locality in Uganda and from D. R. Congo (Hippa & Kurina 2012, Kurina & Hippa 2014).	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FFA2036D53DB6B1372299AD3.taxon	materials_examined	Material. MADAGASCAR: male, Tamatave Torotorofotsy, Andasibe (Perinet), 22 km NW, 18 ° 46,25 ’ S 48 ° 25,93 ’ E, 23 – 25. x. 2014, 960 m, leg. A. H. Kirk-Spriggs & R. Harin’Hala, Malaise trap primary rainforest, BMSA (D) 68190 (slide mounted, BMSA); male, Fianarantsoa, Ranomafana National Park, Talatakely, 800 m SW entrance; 21 ° 15,48 ’ S 47 ° 25,27 ’ E, 16 – 19. x. 2014, 610 m, leg. A. H. Kirk-Spriggs & R. Harin’Hala, Malaise trap secondary rainforest, BMSA (D) 60354 (slide mounted, BMSA). Remarks. The species has so far been known only from the holotype collected from southern Madagascar (Hippa 2008). The current records indicate to a wider distribution in the Island.	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
A15787C8FFA2036D53DB6A9F73379C63.taxon	materials_examined	Material. GHANA: 3 males, Central region, Kakum, Entwikrom 31.10 – 8.11.1994, NUFU-project ZMBN, leg. J. Kjaerandsen & T. Anderson (slide mounted, IZBE and NHMO). CAMEROON: male, North Reg. Vina, Tchabal village, 07 ° 35,195 ’ N 13 ° 33,566 ’ E; 13. viii. 2013, 1362 m, A. H. Kirk-Spriggs; Malaise traps degraded savanna hill forest, BMSA (D) 54969 (slide mounted, BMSA). Remarks. The species was described from KwaZulu-Natal, South Africa, but was recorded also from Côte d’Ivoire (Hippa 2008). The studied specimens are more similar to the specimen figured from Côte d’Ivoire (Hippa 2008: fig. 12 B) because the dorsal medial margin of the gonocoxa is only slightly lobe-like produced posteriorly, while it is a more prominent lobe in South African specimens (cf. Jaschhof & Mostovski 2006: fig. 5).	en	Hippa, Heikki, Søli, Geir, Kurina, Olavi (2019): New data on the genus Manota Williston (Diptera: Mycetophilidae) from Africa, with an updated key to the species. Zootaxa 4652 (3): 401-441, DOI: 10.11646/zootaxa.4652.3.1
