taxonID	type	description	language	source
BED65A578088522DA2DA4CBDFDED8643.taxon	description	Figs 1, 2, 3, 4, 5	en	Nguyen, Anh D., Eguchi, Katsuyuki (2022): Mountainous millipedes in Vietnam. II. A conspicuous Tylopus species from Northern Vietnam (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 1091: 1-13, DOI: http://dx.doi.org/10.3897/zookeys.1091.80094, URL: http://dx.doi.org/10.3897/zookeys.1091.80094
BED65A578088522DA2DA4CBDFDED8643.taxon	materials_examined	Material examined. Holotype. Vietnam • 1 male; Vinh Phuc Province, Tam Dao National Park, on the way to Tam Dao 2; 1,100 m a. s. l.; 25 Feb. 2017; Anh D. Nguyen leg.; natural forests; IEBR-Myr 603 H. Paratypes. Vietnam • 2 females; same data as for holotype; IEBR-Myr 603 P • 1 male; Vinh Phuc Province, Tam Dao National Park, on way to TV tower; 21.46065 ° N, 105.64863 ° E; 1,081 m a. s. l.; 10 Dec. 2019; Hoang Quang Duy leg.; natural forests; IEBR-Myr 876. Non-type. Vietnam • 1 male; Ha Giang Province, Bac Me Natural Reserve, Minh Ngoc commune, Lung Can village; 22.71814 ° N, 105.17997 ° E; 361 m a. s. l.; 12 - 13 Dec. 2018; Anh D. Nguyen leg.; bushes (IEBR-Myr 809).	en	Nguyen, Anh D., Eguchi, Katsuyuki (2022): Mountainous millipedes in Vietnam. II. A conspicuous Tylopus species from Northern Vietnam (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 1091: 1-13, DOI: http://dx.doi.org/10.3897/zookeys.1091.80094, URL: http://dx.doi.org/10.3897/zookeys.1091.80094
BED65A578088522DA2DA4CBDFDED8643.taxon	diagnosis	Diagnosis. The new species distinctly differs from its congeners in having a three-times spiraled solenophore of the gonopods and postfemoral lamella l present while spine z and process h totally absent. The new species is slightly similar to its congener T. strongylomoides (Korsors & Golovatch, 1989), from the same locality (Tam Dao National Park), in having a twisted solenophore and solenomere. However, the new species obviously differs from T. strongylosomoides in having a more strongly coiled solenophore and solenomere (3 x vs 1.5 x). Regarding the gonopod conformation, the new species is somewhat similar to species of the genus Helicorthomorpha in the twist of both the solenophore and solenomere. However, the postfemoral region of Helicorthomorpha members is more elaborate, twisted, and carries no additional processes, whereas that of the new species is spiraled or coiled rather than twisted, and has a postfemoral lamella.	en	Nguyen, Anh D., Eguchi, Katsuyuki (2022): Mountainous millipedes in Vietnam. II. A conspicuous Tylopus species from Northern Vietnam (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 1091: 1-13, DOI: http://dx.doi.org/10.3897/zookeys.1091.80094, URL: http://dx.doi.org/10.3897/zookeys.1091.80094
BED65A578088522DA2DA4CBDFDED8643.taxon	etymology	Etymology. The species epithet, helicorthomorphoides, is used to emphasize the similarity of the gonopod solenophore between the new species and those of the genus Helicorthomorpha.	en	Nguyen, Anh D., Eguchi, Katsuyuki (2022): Mountainous millipedes in Vietnam. II. A conspicuous Tylopus species from Northern Vietnam (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 1091: 1-13, DOI: http://dx.doi.org/10.3897/zookeys.1091.80094, URL: http://dx.doi.org/10.3897/zookeys.1091.80094
BED65A578088522DA2DA4CBDFDED8643.taxon	description	Description. Body length ~ 14.4 mm (male) and 14.9 mm (female). Width of midbody pro- and metazona ~ 1.2 mm (male), 1.6 mm (female) and 1.6 mm (male), 1.9 mm (female), respectively. Body generally brownish yellow or darkish yellow except antennomere 7 and metaterga with a darker, median, V-shaped region (or median triangular-shaped area on metaterga) (Figs 2 - 4). Head (Fig. 2 B, C) slightly smaller than collum, labroclypeous region densely setose. Epicranial suture clearly distinct, dividing frons into two equal parts, with 2 + 2 setae along suture. Frons convex. Antenna (Fig. 2 A-C) short, reaching to approximately tergum 2 laterally. Antennomere 1 = 7 <2 = 3 = 4 = 5 <6; antennomere 6 largest, obviously clavate. Collum (Fig. 2 B) oval-shaped, surface smooth, without granulates or tubercles, but with 2 + 2 setae near anterior margin and 2 + 2 setae in intermediate area. Paratergum well developed, subtriangular, with broad corner. Body segments 3 <4 <2 = 5 - 17, thereafter gradually tapering towards telson. Prozonae and metazonae clearly divided by deep, striated waists. Prozonae smooth, shining, yellowish brown, with a median, broad, longitudinal yellow stria (Figs 2 B, 3 A). Metazonae (Figs 2 B, 3 A) smooth, shining, with 2 + 2 setae near anterior margin, two lateral spotted areas, and 2 + 2 or 3 + 3 oblong knobs near posterior margin. These knobs more obvious on posterior segments. Metatergal sulcus starting on segment 4 but clearly present from segment 5, nearly reaching base of paraterga (Fig. 2 B). Axial line distinct. Paraterga (Figs 2 A, B, D, 3 A-C) well developed, wing-shaped, with at least one setiferous incision near anterior, broadly round corner; caudal corner acute, more pointed from mid-body segment onwards, but never reaching following segment. Caudal corner of paraterga 17 - 19 very pointed, spine-like (Fig. 3 C, D). Ozopores located inside round hollows at the caudolateral sides of paraterga 5, 7, 9, 10, 12, 13, and 15 - 19 (Figs 2 A, D, 3 B). Pleurites (Figs 2 A, D, 3 B) smooth, not granulated. Pleurosternal carinae (Fig. 2 A) present until segment 8, with a small tubercle-like projection, then gradually reduced or missing on subsequent segments. Telson (Figs 3 C, D, 4 A). Epiproct strongly concave, forming two long lateral tubercles. Hypoproct sub-trapeziform, with two separated distolateral setiferous knobs. Sterna. Cross impression distinct. Sternum 5 with a large rectangular lamina between coxae 4 (Fig. 4 B). Leg short, ~ 1.5 x as long as midbody height. Prefemora swollen dorsally. Tarsal brushes present till legs 21, then gradually thinner and absent on subsequent legs. Adenostyles present on postfemora and tibiae, each with a small knob / tubercle (Fig. 4 C). Gonopod suberect, simple (Figs 4 D, 5, 6). Coxite short, with sparsely setose distoventral part. Telopodite long and erect. Prefemorite densely setose, separated from femorite by an oblique sulcus laterally. Femorite somewhat enlarged distally and slightly twisted, without modifications or additional processes. Postfemoral region consisting of a solenomere and a solenophore, both completely coiled 3 x; lamina l present, spine z and process h missing. Seminal groove running entirely on mesal side of femorite, then entering flagelliform solenomere completely sheathed by solenophore. Tip of gonopod strongly bifid.	en	Nguyen, Anh D., Eguchi, Katsuyuki (2022): Mountainous millipedes in Vietnam. II. A conspicuous Tylopus species from Northern Vietnam (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 1091: 1-13, DOI: http://dx.doi.org/10.3897/zookeys.1091.80094, URL: http://dx.doi.org/10.3897/zookeys.1091.80094
