taxonID	type	description	language	source
9E45D034FFE49237FF35FDB87E52B459.taxon	diagnosis	Diagnosis. The shape and position of the labial palpi in the male are most similar to those of F. phycitipalpia Horak, they arise vertically and are appressed to the frons. However, the labial palpi of F. t r a t e n s i s are much shorter and broader and the inner surface of the second segment is only sparsely covered with scales; in contrast, the labial palpi of F. phycitipalpia are longer and more slender. The long, slender, modified scales on distal end of the second segment of the labial palpi and lower frons in the male of F. t r a te ns i s are unique within Olethreutinae and easily distinguished this species from all congeners. The phallus of F. t r a t e n s i s is most similar to those of F. neaera (Meyrick) and F. geniculata Pinkaew and Zhang. However, F. t r a t e n s i s differs from F. n e a e r a and F. geniculata in forewing pattern and in the presence of three large flattened spiniform setae from the outer surface of the valva; in the latter species there are many more large flattened spiniform setae.	en	Jaikla, Soraya, Pinkaew, Nantasak, Vitheepradit, Akekawat, Klangsap, Nathawat (2013): Two new species of Fibuloides (Lepidoptera: Tortricidae) from eastern Thailand. Zootaxa 3664 (1): 85-91, DOI: 10.11646/zootaxa.3664.1.7
9E45D034FFE49237FF35FDB87E52B459.taxon	description	Description. Head: Vertex light brown to brown mixed with dark brown laterally; upper frons light brown mixed with brown, lower frons light brown, male with vertical ridge medially, covered with short, appressed scales mixed with dense, long, slender yellowish white scales, pointing outward (Fig. 2 C); labial palpus in male appressed to the frons and rising upwardly (natural position) (Fig. 2 A), with first segment dark brown, second segment light brown with three, dark brown, transverse bands (at base, middle and apex), with inner surface bearing shallow excavation at 2 / 3 length and covered with dense, appressed, light brown scales, tip with group of moderately dense, long, narrow, light brown scales, apical segment narrow, orange white (Fig. 2 D); female with labial palpus long and porrect, first segment dark brown, second segment triangular, distally greatly widened, light brown with three, dark brown, transverse bands (at base, middle and apex), apical segment short, dark brown (Fig. 2 B); male antenna with notch at the first segment of flagellum. Thorax: Pronotal collar brown mixed with dark brown; mesothorax with tegula light brown mixed with brown; mesonotum light brown to brown with irregular, transverse band medially, with orange white scales on posterior 1 / 4. Forewing (Figs. 1 A – B) length 3.8 – 4.2 mm in males (n = 8), 4.3 – 4.5 mm in females (n = 2); male costal fold absent; termen slightly concave below apex, between R 5 and M 1; ground color light brown; costal strigula 1 indistinct, strigulae 2 – 4 and 7 – 9 paired, strigulae 5 and 6 single, pale orange white, paired strigulae separated by dark brown spots, termen with strigula 10 forming pale yellow spot between R 5 and M 1; basal fascia indistinct, brown mixed with dark brown; subbasal fascia present as small subtriangular spot on costa and extending as irregular transverse narrow band to inner margin, brown mixed with dark brown; median fascia fused with postmedian fascia, forming distinct subtriangular patch on costa, dark brown, extending from costa to midwing; preterminal fascia present as short and oblique band, brown, extending outward from costa to R 5 near outer margin, wing apex brown mixed with dark brown, strigulae 5, 6 and 7 with oblique silvery striae, confluent at R 3 and extending from R 3 to dorsum near tornus, strigula 8 with oblique silvery stria extending to R 4, strigula 9 with short, transverse silvery stria extending to R 5, a narrow, oblique brown band between strigulae 7 and 8 extending to R 5, ocellar region with irregular, transverse narrow line, dark brown, extending from R 5 to tornus and silvery stria beyond extending from M 1 to between CuA 1 and CuA 2, outer margin with narrow line, brown, extending from between R 5 and M 1 to CuA 1. Underside greyish brown, strigulae on costa and termen yellowish grey. Hindwing brown dorsally, greyish brown ventrally, except basal 2 / 3 of both sides covered with dark brown, narrow scales. Abdomen: Male genitalia (Fig. 3 A) with tegumen widest in middle, dorsally with pronounced shoulders. Uncus distally bipartite, dilated medially, apices round, bent, spatulate. Socii subtriangular, moderately large, sparsely setose. Gnathos arising from mid-length of tegumen, moderately sclerotized, with two parallel, dorsally rising bands. Valva long and slender with moderately large basal excavation, small subtriangular lobe on ventral margin near middle of valva, with a cluster of spiniform setae and three flattened, widened spiniform setae arising from outer surface, neck very slender, inner surface with patch of dense setae at base of cucullus; cucullus small, transversely extended, peanut-shaped, dorsal portion small and narrow, apex rounded, moderately setose, ventral portion widened, tip round, with dense spiniform setae; juxta triangular; anellus closely surrounding base of phallus, cuplike, dorsally with plate extending to mid-length of phallus; phallus long, bent upward at right angle in middle, with seven, non-deciduous cornuti. Female genitalia (Fig. 4 A) with sternum VII with large sclerotized plate, with deep, U-shaped medial emargination in posterior margin; densely scaled except on posterior 1 / 4, medially with curved patch of dense, small sclerotized ridges and microtrichia. Tergum VIII with dense microtrichia on anterior half, lateral triangular extensions sparsely setose, anterior 1 / 3 with dense microtrichia, scales absent. Papillae anales with dense setae; apophyses short and stout, anteriores slightly longer than posteriores; ostium bursae opening into a wide shallow cup fused with emargination of S 7; lamella postvaginalis forming a transverse band with a well-sclerotized posterior margin in middle, spinulose, more densely scaled lateroposteriorly, anteromedially without scales and microtrichia, with patch of dense microtrichia extending on membrane laterally beyond lamella postvaginalis; ductus bursae rather short, widened towards corpus bursae; colliculum moderately large, sclerotized, not encircling ductus bursae, followed by a short membranous portion with large sclerotized, bi-pronged plate narrowly encircling ductus bursae near middle and the two pointed prongs extending into bursae; ductus seminalis arising from anterior 1 / 3 of ductus bursae; corpus bursae spinulose, except around the two slender blade-like signa.	en	Jaikla, Soraya, Pinkaew, Nantasak, Vitheepradit, Akekawat, Klangsap, Nathawat (2013): Two new species of Fibuloides (Lepidoptera: Tortricidae) from eastern Thailand. Zootaxa 3664 (1): 85-91, DOI: 10.11646/zootaxa.3664.1.7
9E45D034FFE49237FF35FDB87E52B459.taxon	materials_examined	Holotype. 3. Thailand: Trat Prov.: Trat Agroforestry R. St., 12 ˚ 23 ' 43 " N 102 ˚ 40 ' 32 " E, ca. 30 m, 18 – 19 Aug 2012, N. Pinkaew; np 5334; KKIC slide no. NP 1545. Deposited in BMNH. Paratypes. Thailand: Trat Prov.: Trat Agroforestry R. St., 12 ˚ 23 ' 43 " N 102 ˚ 40 ' 32 " E, ca. 30 m, 19 – 20 Oct 2011 (1 Ƥ) (np 4849; KKIC genitalia slide NP 1549); 19 – 20 Oct 2011 (13), (np 4832; KKIC genitalia slide NP 1541), deposited in TNHM; 24 – 25 Dec 2011 (13) (np 5154; KKIC genitalia slide NP 1542); 21 – 23 Apr 2012 (1 Ƥ) (np 5017; KKIC genitalia slide NP 1546); 17 – 18 Feb 2012 (33) (np 4959; KKIC genitalia slide NP 1538, np 4975 KKIC genitalia slide NP 1539, np 4970; KKIC genitalia slide NP 1543); 16 – 18 Jun 2012 (23) (np 5118; KKIC genitalia slide NP 1540, np 5075; KKIC genitalia slide NP 1537), all N. Pinkaew. Deposited in KKIC.	en	Jaikla, Soraya, Pinkaew, Nantasak, Vitheepradit, Akekawat, Klangsap, Nathawat (2013): Two new species of Fibuloides (Lepidoptera: Tortricidae) from eastern Thailand. Zootaxa 3664 (1): 85-91, DOI: 10.11646/zootaxa.3664.1.7
9E45D034FFE49237FF35FDB87E52B459.taxon	distribution	Distribution. Thailand (Trat).	en	Jaikla, Soraya, Pinkaew, Nantasak, Vitheepradit, Akekawat, Klangsap, Nathawat (2013): Two new species of Fibuloides (Lepidoptera: Tortricidae) from eastern Thailand. Zootaxa 3664 (1): 85-91, DOI: 10.11646/zootaxa.3664.1.7
9E45D034FFE49237FF35FDB87E52B459.taxon	etymology	Etymology. The species name refers to the type locality of Trat province.	en	Jaikla, Soraya, Pinkaew, Nantasak, Vitheepradit, Akekawat, Klangsap, Nathawat (2013): Two new species of Fibuloides (Lepidoptera: Tortricidae) from eastern Thailand. Zootaxa 3664 (1): 85-91, DOI: 10.11646/zootaxa.3664.1.7
9E45D034FFE19235FF35FA8B7FD5B22A.taxon	diagnosis	Diagnosis. This species is most similar to F. t r a t e n s i s in forewing pattern, but the two species differ the male and female genitalia. The frons and labial palpi of the male of F. bulla lack modified scales characteristic of F. tratensis. F. b u l l a is distinguished by a dense cluster of long, parallel spiniform setae on the ventral part of the cucullus, which is unique and easily separates it from all other congeners.	en	Jaikla, Soraya, Pinkaew, Nantasak, Vitheepradit, Akekawat, Klangsap, Nathawat (2013): Two new species of Fibuloides (Lepidoptera: Tortricidae) from eastern Thailand. Zootaxa 3664 (1): 85-91, DOI: 10.11646/zootaxa.3664.1.7
9E45D034FFE19235FF35FA8B7FD5B22A.taxon	description	Description. Head: Upper frons yellowish white with lateral areas narrowly dark brown, lower frons yellowish white; vertex pale grey mixed with brown to dark brown laterally; labial palpus of male and female long, porrect, first segment light grey, second segment triangular, distally greatly widened, yellowish white, with brownish grey spots at base, dorsomedially, ventromedially and apex, apical segment yellowish white; male antenna with notch at of basal segment of flagellum. Thorax: Pronotal collar brown mixed with dark brown; mesothorax with tegula light brown mixed with dark brown; mesonotum light brown with diffuse spots, brown to dark brown. Forewing (Figs. 1 C – D) length 3.5 – 3.8 mm in males (n = 3), 5.0 mm in female (n = 1); male costal fold absent; termen slightly concave between R 5 and M 1; ground color light brown with diffused transverse, narrow, darker streaks; costal strigula 1 indistinct, strigulae 2 – 4 and 7 – 8 paired, strigulae 5, 6 and 9 single, light brown, paired strigulae separated by dark brown spots, termen with strigula 10 forming yellowish white spot between R 5 and M 1; basal fascia present as irregular patch extending from costa to midwing; subbasal fascia a dark brown, subrectangular patch extending from strigula 2 to 3 and from costa to R; median fascia a distinct, dark brown, oblique patch extending from costa to middle of discal cell, separated from small dark brown spot at base of M 2; postmedian fascia present as short, oblique brown band, extending outward from costa to R 5, wing apices brown mixed with dark brown, strigula 6 with irregular silvery stria extending from R 2 to inner margin, strigula 7 with silvery stria extending obliquely to R 5, ocellar region with irregular transverse band, brown mixed with dark brown, extending from R 5 to tornus, near outer margin with short, transverse silvery stripe between M 2 and CuA 1 near outer margin, outer margin with narrow line of dark brown scales, extending from M 1 to CuA 1. Hindwing brown. Underside of forewing greyish yellow, strigulae on costa yellowish grey, strigula on termen yellowish white; underside of hindwing light brown. Abdomen: Male genitalia (Fig. 3 B) with tegumen densely setose laterally, elongate subrectangular, shoulders with dense microtrichia. Uncus short, small, bifid, subtriangular, pointing outward. Socii moderately large, teardrop-shaped, pendent, from base of uncus, with dense spiniform setae mixed with less dense smaller ones. Gnathos arising near mid-length of tegumen, with two rising bands, moderately sclerotized. Valva straight and rather wide subbasally, with large basal opening, with dense microtrichia from near base of sacculus to neck; sacculus sparsely setose basally, posterior margin to neck with patch of dense setae, dorsal and ventral margin sparsely setose, ventral margin with row of large, flattened spiniform setae with multidentate apices from outer surface of valva; neck sparsely setose; cucullus simple, short, rounded, densely setose, base of inner surface with group of dense, narrow spines, pointing basally, ventral margin with dense cluster of moderately long, flattened spiniform setae; juxta subtrapezoid, caulis moderately long; anellus closely surrounding basal 1 / 5 of phallus; phallus moderately long, sinuate, with five, deciduous cornuti. Female genitalia (Fig. 4 B) with sternum VII moderately sclerotized, moderately densely scaled, scales denser near posterior margin, except sparsely setose near sterigma, posterior margin with small U-shaped emargination medially. Tergum VIII with two wide, shallow pouches, lateral triangular extensions with dense scales, moderately dense setae and microtrichia. Papillae anales densely setose; apophyses anteriores long and slender (same length as apophyses posteriores); sterigma simple, ostium bursae rather small, fused with emargination of posterior margin of S 7, antrum forming narrow sclerotized cup, 1 / 4 length of ductus bursae; ductus bursae moderately long, widened toward corpus bursae, colliculum small, moderately sclerotized, not encircling ductus bursae, with large sclerotized bi-pronged plate narrowly encircling ductus bursae near middle and with two band-like projections with rounded tips extending into posterior portion of corpus bursae; corpus bursae spinulose except around two strong, blade-like signa with acute tips, left one larger than right one.	en	Jaikla, Soraya, Pinkaew, Nantasak, Vitheepradit, Akekawat, Klangsap, Nathawat (2013): Two new species of Fibuloides (Lepidoptera: Tortricidae) from eastern Thailand. Zootaxa 3664 (1): 85-91, DOI: 10.11646/zootaxa.3664.1.7
9E45D034FFE19235FF35FA8B7FD5B22A.taxon	materials_examined	Holotype. 3. Thailand: Trat Prov.: Trat Agroforestry R. St., 12 ˚ 23 ' 43 " N 102 ˚ 40 ' 32 " E, ca. 30 m, 12 – 14 Oct 2012, N. Pinkaew; np 5526; KKIC slide no. NP 1547. Deposited in BMNH. Paratype. Thailand: Trat Prov.: Trat Agroforestry R. St., 12 ˚ 23 ' 43 " N 102 ˚ 40 ' 32 " E, ca. 30 m, 12 – 14 Oct 2011 (1 Ƥ) (np 5527; KKIC genitalia slide NP 1548); 18 – 19 Aug 2012 (23) (np 5371; KKIC genitalia slide NP 1544), 12 – 14 Oct 2011 (np 5528; KKIC genitalia slide NP 1550), all N. Pinkaew et al. Deposited in THNM and KKIC.	en	Jaikla, Soraya, Pinkaew, Nantasak, Vitheepradit, Akekawat, Klangsap, Nathawat (2013): Two new species of Fibuloides (Lepidoptera: Tortricidae) from eastern Thailand. Zootaxa 3664 (1): 85-91, DOI: 10.11646/zootaxa.3664.1.7
9E45D034FFE19235FF35FA8B7FD5B22A.taxon	distribution	Distribution. Thailand (Trat).	en	Jaikla, Soraya, Pinkaew, Nantasak, Vitheepradit, Akekawat, Klangsap, Nathawat (2013): Two new species of Fibuloides (Lepidoptera: Tortricidae) from eastern Thailand. Zootaxa 3664 (1): 85-91, DOI: 10.11646/zootaxa.3664.1.7
9E45D034FFE19235FF35FA8B7FD5B22A.taxon	etymology	Etymology. The specific name is derived from Latin bulla (= bubble), referring to the shape of the cucullus.	en	Jaikla, Soraya, Pinkaew, Nantasak, Vitheepradit, Akekawat, Klangsap, Nathawat (2013): Two new species of Fibuloides (Lepidoptera: Tortricidae) from eastern Thailand. Zootaxa 3664 (1): 85-91, DOI: 10.11646/zootaxa.3664.1.7
