identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
B4A368D346145DE8B232C68F9B452625.text	B4A368D346145DE8B232C68F9B452625.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Isannaia fortunata Jeratthitikul & Sutcharit & Prasankok 2024	<div><p>Isannaia fortunata sp. nov.</p><p>Figs 1, 3, 4 A, B</p><p>Type material.</p><p>Holotype. Thailand • Nakhon Phanom Province, Tha Uthen District, Tha Uthen Subdistrict, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.609&amp;materialsCitation.latitude=17.5621" title="Search Plazi for locations around (long 104.609/lat 17.5621)">Thuai River</a>; 17.5621 ° N, 104.6090 ° E; E. Jeratthitikul, P. Prasankok, and K. Wisittikoson leg.; MUMNH -UNI 1950; shell length 54.13 mm, shell height 29.96 mm, and shell width 14.39 mm . Paratypes. Thailand • 3 shells; same collection data as for holotype; MUMNH -UNI 1951 to UNI 1953 .</p><p>Description.</p><p>Shell medium-sized (shell length 43.3–61.4 mm, shell height 24.5–35.1 mm, shell width 13.4–18.4 mm; Table 2), rhomboidal, elongate (H / L ratio = 0.55–0.57), rather thin, very inequilateral, moderately inflated. Anterior margin rounded, rather constricted, shouldered; posterior margin truncated, slightly elongate ventrally; ventral margin slightly curved. Dorsal margin slightly curved; anterior low, posterior end slightly higher. Umbo slightly elevated, heavily eroded even in young specimens. Posterior ridge wide and obtuse, not prominent; posterior slope with two faint lines running from umbo to posterior margin. Periostracum thin, brownish-black, eroded part white to coppery-brown. Shell surface with visible growth lines. Ligament long, very narrow, dark brown. One very shallow hill-like or triangulate pseudocardinal tooth on right valve; pseudocardinal tooth on left valve less developed, tubercule-like, or almost absent. Posterior end of hinge structure with small V-shaped fossette. Anterior adductor muscle scar shallow, ovate, contiguous with pedal retractor and anterior protractor muscle scars; posterior adductor muscle scar round, very shallow, almost invisible. Pallial line faint, continuous. Umbo cavity moderately deep, with 1 to 2 tiny muscle scars in the cavity. Nacre blue-whitish with yellowish tint around the umbo cavity.</p><p>Exhalant aperture almost smooth, with a row of tiny tubercle-like papillae on edge of aperture, shorter than inhalant. Inhalant aperture with a row of conical papillae, varying in length. Small epithelial fold divides exhalant and inhalant apertures. Gills elongated and slightly ribbed; inner gills slightly wider, and anterior margin of inner gills slightly longer than that of outer gills. Labial palps elongate, slightly curved, pointed at tip. Glochidia unknown.</p><p>Etymology.</p><p>The species name “ fortunata ” is derived from the Latin adjective “ fortunatus ” meaning lucky. This name refers to the fortuitous discovery of this rare freshwater species.</p><p>Distribution.</p><p>This new species is known only from the type locality.</p><p>Remarks.</p><p>This new species is rare and restricted to the Thuai River. Specimens were collected from hard clay along the river bank. It was found sympatrically with several unionid species endemic to Songkhram Basin and nearby tributaries of the Mekong Basin, including Lens maenamensis Pfeiffer et al., 2021, Physunio pellucidus Pfeiffer et al., 2021, Ensidens jaculus (Rochebrune, 1882), Scabiellus songkramensis (Kongim et. al., 2015), Pseudodon cambodjensis (Petit de la Saussaye, 1865) and Scabies scobinatus (Lea, 1856) .</p></div>	https://treatment.plazi.org/id/B4A368D346145DE8B232C68F9B452625	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Jeratthitikul, Ekgachai;Sutcharit, Chirasak;Prasankok, Pongpun	Jeratthitikul, Ekgachai, Sutcharit, Chirasak, Prasankok, Pongpun (2024): Two new genera and three new species of exceptionally rare and endemic freshwater mussels (Bivalvia, Unionidae) from the Mekong Basin. Zoosystematics and Evolution 100 (4): 1333-1345, DOI: 10.3897/zse.100.130929
C0910C0350AE55BFB0CF0786B4F04852.text	C0910C0350AE55BFB0CF0786B4F04852.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Isannaia Jeratthitikul & Sutcharit & Prasankok 2024	<div><p>Genus Isannaia gen. nov.</p><p>Type species.</p><p>Isannaia fortunata gen. et sp. nov., by present designation.</p><p>Species included.</p><p>The new genus is currently composed of two species, the type species and I. occultata sp. nov.</p><p>Diagnosis.</p><p>The new genus is distinguished from other genera in the Pseudodontini by its thin and moderately inflated shell, and with rhomboidal to ovate outline (Fig. 4). The overall shape is somewhat elongated but relatively shorter when compared to those of Bineurus, Pilsbryoconcha or Namkongnaia . The anterior margin is rounded, shouldered, and the posterior margin is truncated to rounded. Its pseudocardinal teeth are rather small, one on each valve, hill-like or triangulate shape, and less developed on the left valve. Its umbo is slightly elevated and the umbo cavity moderately deep. It is also represented as a distinct clade in multi-locus phylogenetic analyses (Fig. 1).</p><p>Description.</p><p>Shell medium-sized, rather thin, rhomboidal to ovate outline, elongate, very inequilateral, moderately inflated; anterior margin rounded, shouldered; posterior margin truncated to rounded. Umbo slightly elevated and usually eroded. Ligament very narrow. One shallow hill-like or triangulate pseudocardinal tooth on right valve; left valve with one less developed hill-like to tubercule-like pseudocardinal tooth. End of hinge structure with V-shaped fossette. Anterior adductor muscle scar shallow to well-developed, ovate, contiguous with pedal retractor and anterior protractor muscle scars; posterior adductor muscle scar round, very shallow. Pallial line faint to well-marked, continuous. Umbo cavity moderately deep.</p><p>Etymology.</p><p>The generic name “ Isannaia ” is from the word “ Isan ”, the name of the region of northeastern Thailand, and Greek word “ naiad ” meaning freshwater mussel. The name of this genus thus means “ freshwater mussels from northeastern Thailand ”. The gender is feminine.</p><p>Distribution.</p><p>Endemic to tributaries of the Mekong River in northeastern Thailand (Fig. 3).</p><p>Remarks.</p><p>Phylogenetic analyses have failed to recover a robustly supported position of Isannaia gen. nov. among other Pseudodontina genera. However, in terms of genetic distance, it is nearest to Namkongnaia, with a separation of 11.77 % p-distance of the COI gene (Table 1). Isannaia gen. nov. can be easily distinguished from Namkongnaia by having a much shorter shell (Jeratthitikul et al. 2021 b).</p><p>In general, Isannaia gen. nov. bears conchological resemblance to certain species within the Sundadontina, such as S. brandti Bolotov et al., 2020 or S. plugpomenica Konopleva et al., 2023 (Bolotov et al. 2020; Konopleva et al. 2023). Nevertheless, its thin shell with less developed pseudocardinal teeth set it apart from the latter genus, which typically features a thicker and sturdier shell with stout and tubercular-like pseudocardinal teeth (Bolotov et al. 2020).</p></div>	https://treatment.plazi.org/id/C0910C0350AE55BFB0CF0786B4F04852	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Jeratthitikul, Ekgachai;Sutcharit, Chirasak;Prasankok, Pongpun	Jeratthitikul, Ekgachai, Sutcharit, Chirasak, Prasankok, Pongpun (2024): Two new genera and three new species of exceptionally rare and endemic freshwater mussels (Bivalvia, Unionidae) from the Mekong Basin. Zoosystematics and Evolution 100 (4): 1333-1345, DOI: 10.3897/zse.100.130929
ACBEF56237745895AE444C49C4EE638E.text	ACBEF56237745895AE444C49C4EE638E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Isannaia occultata Jeratthitikul & Sutcharit & Prasankok 2024	<div><p>Isannaia occultata sp. nov.</p><p>Figs 1, 3, 4 C, D</p><p>Type material.</p><p>Holotype. Thailand • Udon Thani Province, Wang Sam Mo District, Nong Ya Sai Subdistrict, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.3638&amp;materialsCitation.latitude=16.9886" title="Search Plazi for locations around (long 103.3638/lat 16.9886)">Yang Stream</a> (a tributary of Lam Pao River); 16.9886 ° N, 103.3638 ° E; K. Wisittikoson leg.; MUMNH -UNI 2895; shell length 53.84 mm, shell height 31.68 mm, and shell width 20.41 mm . Paratypes. Thailand • 2 shells; same collection data as for holotype; MUMNH -UNI 2896, UNI 2897 .</p><p>Diagnosis.</p><p>This species resembles I. fortunata sp. nov., but it can be distinguished by the overall shell shape being more rounded (vs rhomboidal), slightly higher shell (H / L ratio 0.58–0.60 vs 0.55–0.57), wider anterior margin (vs rather constricted), more truncated and rounded posterior margin (vs slightly ventrally elongate), less curved ventral margin (vs curved ventral margin). It also differs in characteristics inside the shell by having higher and more triangulate pseudocardinal tooth on the right valve (vs very shallow and hill-like pseudocardinal tooth), well-developed muscle scars (vs very shallow and sometimes almost invisible), 3 to 5 well-developed muscle scars in the umbo cavity (vs 1 to 2 tiny muscle scars in the umbo cavity), and well-marked pallial line (vs faint pallial line). It can also be distinguished from its congeners by fixed nucleotide substitution positions in the COI gene fragment of base A on the 54 th, 114 th, 363 rd, and 414 th; base T on the 90 th, 141 st, 591 st, and 609 th; base C on the 57 th, and 216 th; and base G on 150 th, 117 th, 192 nd, 558 th, 480 th, 552 nd, 627 th, 639 th, 645 th, and 657 th.</p><p>Description.</p><p>Shell medium-sized (shell length 43.8–53.8 mm, shell height 26.7–31.7 mm, shell width 13.8–20.4 mm; Table 2), ovate to rectangular, elongate (H / L ratio = 0.58–0.60), rather thin, very inequilateral, moderately inflated. Anterior margin rounded, shouldered; posterior margin truncated to rounded; ventral margin straight to slightly curved. Dorsal margin slightly curved; anterior low, posterior end slightly higher. Umbo slightly elevated, heavily eroded even in young specimens. Posterior ridge wide and obtuse, not prominent; posterior slope with two faint lines running from umbo to posterior margin, invisible in old specimens. Periostracum thin, brown to brownish-black, eroded part white to coppery-brown. Shell surface with visible growth lines. Ligament long, very narrow, dark brown. One hill-like or triangulate pseudocardinal tooth on right valve; pseudocardinal tooth on left valve less developed, hill-like. Posterior end of hinge structure with small V-shaped fossette. Anterior adductor muscle scar well-developed, ovate, contiguous with anterior pedal retractor and anterior protractor muscle scars; posterior adductor muscle scar round, shallow, contiguous with posterior retractor muscle scars; posterior retractor muscle scars triangular. Pallial line well-marked, continuous. Umbo cavity moderately deep, with 3 to 5 muscle scars in cavity. Nacre blue-whitish to yellowish, more yellowish toward the umbo cavity.</p><p>Exhalant aperture almost smooth, with a row of tiny tubercle-like papillae on the edge of aperture, similar length with inhalant. Inhalant aperture with a row of very short conical papillae. Small epithelial fold divides exhalant and inhalant apertures. Gills elongated and slightly ribbed; inner gills slightly wider, and anterior margin of inner gills slightly longer than that of outer gills. Labial palps elongate, slightly curved, pointed at tip. Glochidia unknown.</p><p>Etymology.</p><p>The species name “ occultata ” is derived from the Latin adjective “ occultatus ” meaning concealed or hidden, a reference to this new freshwater mussel being hidden in a small stream far from the main river.</p><p>Distribution.</p><p>This new species is known only from the type locality, Yang Stream in Udon Thani Province. Yang Stream is a headwater tributary of the Lam Pao River in the Chi River Basin. The type locality is above the Lam Pao Dam.</p><p>Remarks.</p><p>This new species is rare and difficult to find despite repeated surveys. They were collected from a network of dense tree roots on a clay wall of a small stream. Nearby, there was a community of unionid mussels living in muddy or sandy substrate of the stream bottom, including Scabies phaselus (Lea, 1856), Physunio modelli Brandt, 1974, Namkongnaia inkhavilayi Jeratthitikul et al., 2021, P. exilis, S. brandti, and Thaiconcha callifera (Martens, 1860) .</p></div>	https://treatment.plazi.org/id/ACBEF56237745895AE444C49C4EE638E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Jeratthitikul, Ekgachai;Sutcharit, Chirasak;Prasankok, Pongpun	Jeratthitikul, Ekgachai, Sutcharit, Chirasak, Prasankok, Pongpun (2024): Two new genera and three new species of exceptionally rare and endemic freshwater mussels (Bivalvia, Unionidae) from the Mekong Basin. Zoosystematics and Evolution 100 (4): 1333-1345, DOI: 10.3897/zse.100.130929
37E406E32C2254DF9F22C25E8BA1B68F.text	37E406E32C2254DF9F22C25E8BA1B68F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lannanaia Jeratthitikul & Sutcharit & Prasankok 2024	<div><p>Genus Lannanaia gen. nov.</p><p>Type species.</p><p>Lannanaia kokensis gen. et sp. nov., by present designation.</p><p>Species included.</p><p>The new genus currently contains only one species, L. kokensis sp. nov.</p><p>Diagnosis.</p><p>This new genus is distinguished from other Pseudodontini genera by its compressed, rather short and high shell, with sub-trigonal outline, and distinct posterior wing. The hinge plate is without dentition, or with very rudimentary broad pseudocardinal tooth in each valve, and posterior end of the hinge structure with rather wide V-shaped fossette (Fig. 2). It is also represented as a distinct clade in multi-locus phylogenetic analyses (Fig. 1).</p><p>Description.</p><p>Shell medium-sized, thin, rather short and high, winged, sub-trigonal outline, inequilateral, rather compressed. Anterior margin rounded; posterior margin angulate, somewhat pointed. Umbo tiny, not prominent, eroded. Ligament long, very narrow. Hinge without dentition, or with very rudimentary broad pseudocardinal tooth in each valve; posterior end of the hinge structure with rather wide V-shaped fossette. Anterior adductor muscle scar very shallow, ovate, contiguous with anterior pedal retractor muscle scar, but separated from anterior protractor scar; posterior adductor muscle scar round, very shallow, almost invisible. Pallial line very faint, continuous. Umbo cavity shallow.</p><p>Etymology.</p><p>The generic name “ Lannanaia ” is from the word “ Lanna ”, a name of the kingdom located in present-day northern Thailand during the 13 th to 18 th centuries, and the Greek word “ naiad ” meaning freshwater mussel. The name of this genus thus means “ freshwater mussels from Lanna ” or “ freshwater mussels from northern Thailand ”. The gender is feminine.</p><p>Distribution.</p><p>Endemic to Kok River, a tributary of the Mekong River in northern Thailand (Fig. 3).</p><p>Remarks.</p><p>The phylogenetic tree generated in this study places Lannanaia gen. nov. as a sister group to Songkhlanaia, with strong support from both analyses (BS = 100 %, bpp = 0.99; Fig. 1), and with a 11.42 % p-distance based on the COI gene (Table 1). Both genera share general characteristics of a short shell outline and rather compressed shell. However, Lannanaia gen. nov. can be distinguished from Songkhlanaia by having a sub-trigonal shell outline and a curved ventral margin (vs a rectangular shell outline and an almost straight ventral margin in Songkhlanaia), distinct posterior wing (vs absent posterior wing), and not prominent umbo (vs slightly elevated and easily visible umbo) (Konopleva et al. 2023). In addition, the biogeography of the two genera is also distinct. The monotypic genus Songkhlanaia was described from the Songkhla Lake Basin in southern Thailand (Konopleva et al. 2023), while Lannanaia gen. nov. is restricted to the Kok River, a tributary of the Mekong River in northern Thailand.</p></div>	https://treatment.plazi.org/id/37E406E32C2254DF9F22C25E8BA1B68F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Jeratthitikul, Ekgachai;Sutcharit, Chirasak;Prasankok, Pongpun	Jeratthitikul, Ekgachai, Sutcharit, Chirasak, Prasankok, Pongpun (2024): Two new genera and three new species of exceptionally rare and endemic freshwater mussels (Bivalvia, Unionidae) from the Mekong Basin. Zoosystematics and Evolution 100 (4): 1333-1345, DOI: 10.3897/zse.100.130929
6A0BC990AA2155A9A787499DACD59067.text	6A0BC990AA2155A9A787499DACD59067.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lannanaia kokensis Jeratthitikul & Sutcharit & Prasankok 2024	<div><p>Lannanaia kokensis sp. nov.</p><p>Figs 1, 2, 3</p><p>Type material.</p><p>Holotype. Thailand • Chiang Rai Province, Mueang Chiang Rai District, Rop Wiang Subdistrict, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.8697&amp;materialsCitation.latitude=19.9131" title="Search Plazi for locations around (long 99.8697/lat 19.9131)">Kok River</a>; 19.9131 ° N, 99.8697 ° E; E. Jeratthitikul and K. Wisittikoson leg.; MUMNH -UNI 0825; shell length 57.2 mm, shell height 40.8 mm, and shell width 14.4 mm . Paratype. Thailand • 1 shell; same collection data as for holotype; MUMNH -UNI 0824 .</p><p>Description.</p><p>Shell medium-sized (shell length 57.1–57.2 mm, shell height 40.8–43.8 mm, shell width 14.4–14.8 mm; Table 2), thin, rather short and high (H / L ratio = 0.71–0.77), winged, sub-trigonal outline, inequilateral, rather compressed. Anterior margin rounded; posterior margin angulate, somewhat pointed; ventral margin curved. Dorsal margin straight; anterior low and short, gradually elevated to posterior end; posterior end high and winged. Umbo tiny, not prominent, eroded. Posterior ridge wide and obtuse, not prominent; posterior slope with two fine lines running as curved line from umbo to posterior margin. Periostracum thin, greenish to dark brown, the eroded part white to coppery-brown. Shell surface with very fine growth lines. Ligament long, very narrow, dark brown. Hinge without dentition, or with very rudimentary broad pseudocardinal tooth in each valve; posterior end of hinge structure with rather broad V-shaped fossette. Anterior adductor muscle scar very shallow, ovate, contiguous with anterior pedal retractor muscle scar, but separated from anterior protractor scar; posterior adductor muscle scar round, very shallow, almost invisible. Pallial line very faint, continuous. Umbo cavity shallow. Nacre blue-whitish.</p><p>Exhalant aperture smooth, shorter than inhalant. Inhalant aperture with a row of conical papillae, varying in length. Small epithelial fold divides exhalant and inhalant apertures. Gills elongated and slightly ribbed; outer gills much narrower (about half) than inner gills; anterior margin of inner gills slightly longer than that of outer gills. Labial palps elongate, round at tip. Glochidia unknown.</p><p>Etymology.</p><p>The species name “ kokensis ” refers to the type locality, the Kok River in Chiang Rai Province, northern Thailand.</p><p>Distribution.</p><p>This new species is known only from the type locality.</p><p>Remarks.</p><p>Two living specimens were collected from a small stream that is a tributary to the Kok River, with a muddy bottom. We searched multiple times and locations for this species but failed to find additional specimens. In contrast, Lens contradens (Lea, 1838) and Pilsbryoconcha exilis (Lea, 1838) were found nearby in high abundance, both living and as empty shells. We assume this new species has a low population density and is possibly endemic to the Kok River Basin.</p></div>	https://treatment.plazi.org/id/6A0BC990AA2155A9A787499DACD59067	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Jeratthitikul, Ekgachai;Sutcharit, Chirasak;Prasankok, Pongpun	Jeratthitikul, Ekgachai, Sutcharit, Chirasak, Prasankok, Pongpun (2024): Two new genera and three new species of exceptionally rare and endemic freshwater mussels (Bivalvia, Unionidae) from the Mekong Basin. Zoosystematics and Evolution 100 (4): 1333-1345, DOI: 10.3897/zse.100.130929
B0BBEAF274EB5BB4BBCE92A5B6DD59B2.text	B0BBEAF274EB5BB4BBCE92A5B6DD59B2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudodontina Frierson 1927	<div><p>Subtribe Pseudodontina Frierson, 1927</p><p>Remarks.</p><p>Currently the Pseudodontina is composed of 45 species in ten genera, including the new taxa described herein (Bolotov et al. 2023). These genera are Bineurus Simpson, 1900 (5 species), Isannaia gen. nov. (2 species), Lannanaia gen. nov. (1 species) Namkongnaia Jeratthitikul et al., 2021 (2 species), Nyeinchanconcha Bolotov et al., 2020 (1 species), Pilsbryoconcha Simpson, 1900 (9 species), Pseudodon Gould, 1844 (9 species), Songkhlanaia Konopleva et al., 2023 (1 species), Sundadontina Bolotov et al., 2020 (12 species), and Thaiconcha Bolotov et al., 2020 (3 species). Pseudodon mekongi (Bolotov et al., 2020) was nested within a clade of Pseudodon vondembuschianus (Lea, 1840) in the phylogenetic analysis in this study. Therefore, we recognize it as a subspecies of P. vondembuschianus .</p></div>	https://treatment.plazi.org/id/B0BBEAF274EB5BB4BBCE92A5B6DD59B2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Jeratthitikul, Ekgachai;Sutcharit, Chirasak;Prasankok, Pongpun	Jeratthitikul, Ekgachai, Sutcharit, Chirasak, Prasankok, Pongpun (2024): Two new genera and three new species of exceptionally rare and endemic freshwater mussels (Bivalvia, Unionidae) from the Mekong Basin. Zoosystematics and Evolution 100 (4): 1333-1345, DOI: 10.3897/zse.100.130929
