taxonID	type	description	language	source
0231086ADB3D509A9F8A15F37A8C8F83.taxon	diagnosis	Diagnosis (Figs 4 - 7). A species of Pristimantis characterized by the following combination of morphological characters: (1) dorsal skin shagreen with scattered larger tubercles; dorsolateral folds absent; discoidal fold visible; skin on venter areolate. (2) Tympanic membrane and tympanic annulus present, its dorsoposterior border converges with supratympanic fold; its diameters are 35.6 - 56.0 % of the eye diameter; small, barely visible, subconical postrictal tubercles. (3) Snout short, broadly rounded in dorsal view and rounded in lateral view; canthus rostralis sharp and concave. (4) Upper eyelid bearing one to three subconical tubercles, narrower than IOD. (5) Choanae small, subovoid; dentigerous processes of vomers prominent, oblique, and widely separated from each other, bearing 8 or 9 teeth. (6) Males with vocal slits; subgular vocal sac observable; nuptial pads not evident. (7) Finger I shorter than II, with discs expanded and rounded; bifid palmar tubercle. (8) Fingers bearing narrow lateral fringes. (9) Ulnar tubercles absent. (10) Tarsal tubercles present, subconical; heel tubercles present but nearly inconspicuous and conical. (11) Two metatarsal tubercles, with inner tubercle elongate, three times the length of the rounded and prominent outer tubercle; supernumerary plantar tubercles numerous, enlarged, and rounded. (12) Toes with lateral fringes and broad discs; toe V much longer than toe III (disc on toe III extends to the proximal edge of the medial subarticular tubercle on toe IV, disc on toe V extends beyond the distal edge of the penultimate subarticular tubercle on toe IV); webbing absent. (13) Dorsal surface pattern variable, with homogeneous color brown (with or without paravertebral line) or inverted V-shaped markings with dark brown blotches edged with pale cream; iris gray, medially reddish, with black reticulations; ventral surfaces cream-colored to light brown, finely peppered with irregular, diffuse, dark-brown reticulations or blotches; posterior surface of thighs brown; dark-brown labial bars present or absent; edge of the chin with irregular blotches of dark brown (Fig. 5). (14) apparently sexually dimorphic in size (Table 4), with an SVL in adult males 19.6 - 23.7 mm and 19.0 - 24.9 mm in adult females.	en	Acosta-Galvis, Andres R., Saldarriaga-Gomez, Ana M., Ramirez, Beatriz, Vargas-Ramirez, Mario (2020): A new Terrarana frog of genus Pristimantis from an unexplored cloud forest from the eastern Andes, Colombia. ZooKeys 961: 129-156, DOI: http://dx.doi.org/10.3897/zookeys.961.51971, URL: http://dx.doi.org/10.3897/zookeys.961.51971
0231086ADB3D509A9F8A15F37A8C8F83.taxon	description	Description of the holotype. An adult female (Figs 4, 5) with a snout-vent length (SVL) of 23.8 mm; the skin of cephalic region, dorsum, eyelids, lateral surfaces, and dorsal thighs shagreen with scattered larger tubercles; dorsolateral folds absent and discoidal folds visible; skin on venter areolate. Head length (HL), diagonally from the corner of mouth to tip of snout 10.4 mm; head width (HW) 9.8 mm, approximately equal to width of the body and 41.1 % of the SVL. Snout broadly rounded in dorsal view (type F, sensu Duellman and Lehr 2009; Fig. 4) and rounded in lateral view (type A, sensu Duellman and Lehr 2009; Fig. 4); internarial distance (between center of naris) 2.9 mm; nostril moderately protuberant, directed dorsolaterally; canthus rostralis well defined; loreal region slightly concave; lips not prominent. Eye diameter (ED) from its posterior to anterior corner 3.3 mm; its length 73.3 % of the ETS (distance between the anterior edge of the eye to the tip of snout); interorbital region wider than upper eyelid; the upper eyelid width (UEW) 55.2 % of interorbital distance (IOD); upper eyelid bearing three smaller subconical tubercles (Figs 4, 5); no cranial crests. Supratympanic fold low and short. Tympanic membrane and tympanic annulus present, small, and rounded (Figs 4, 5), its dorsoposterior border converges with supratympanic fold; its diameter 1.1 mm and equivalent to 33 % of eye diameter (ED). Choanae subovoid, not concealed by the palatal shelf of the maxillary arch; dentigerous processes of vomers prominent, nine teeth positioned posterior to level of choanae and widely separated from each other. Tongue rounded, its posterior border notched for half of its extension is adherent to the floor of mouth; teeth present on the maxillary arch. Forelimbs of moderate size, forearm length 6.4 mm; ulnar tubercles absent. Hand length (HnL) 7.4 mm its length 31.0 % of SVL. Palmar tubercle bifid, about two-thirds the length of oval thenar tubercle (Fig. 7). Supernumerary palmar tubercles present, rounded to elongated, and slightly elevated; subarticular tubercles large, round, and conical; fingers without lateral fringes; disks on all fingers rounded apically and extensively expanded (Fig. 7); disk of finger III equal in diameter to the tympanic annulus; disks bearing ventral pads; finger I shorter than II when appressed (Fig. 7). Relative lengths of appressed fingers III> IV> II> I. Subarticular tubercles 1 - 1 - 2 - 2. Hindlimbs slender; foot length (FtL) 12.1 mm, 50.8 % of SVL. Toe webbing and toe fringes absent. Relative lengths of appressed toes IV> V> III> II> I. Discs of the toes expanded; width of adjacent phalange 53.7 % of disc of toe IV; disc of toe III does not reach penultimate subarticular tubercle of toe IV; toe V beyond that of the level of penultimate subarticular tubercle of toe IV. Femur length (FL) 13.6 mm, tibia length (TL) 14.5 mm, its length is equivalent to 60.9 % of SVL. Subarticular tubercles 1 - 1 - 2 - 3 - 2; supernumerary plantar tubercles numerous, suboval, and low; inner metatarsal tubercle oval; outer metatarsal tubercle rounded, prominent, and smaller than inner metatarsal tubercle. Diameter outer metatarsal tubercle 52.8 % of inner metatarsal tubercle; outer tarsal fold absent; inner tarsal fold short. Numerous supernumerary plantar tubercles rounded and barely visible; subarticular tubercles large, round, and conical; toes without lateral fringes; no webbing. Cloacal sheath absent; subcloacal tubercles absent. Color of holotype in preservative (Fig. 4). Dorsum and flanks dark brown; hands in dorsal view, with fingers I and II cream-colored, while fingers III and IV brown with cream-colored bars; dorsal surfaces of the thigh with diffuse dark-brown transversal bars; hidden surfaces of thighs pale brown; venter light brown with a dark-brown suffusion and mottled brown; ventral surfaces of hindlimbs and forelimbs dark brown with a cream-colored suffusion; edge of chin with irregular blotches of dark brown; hands, in ventral view, with palmar tubercle cream-colored and palmar region dark brown. Color of holotype in life (Fig. 5). Dorsal surfaces of body and limbs pink-orange; flanks salmon and sides of the head pink-orange; venter reddish cream-colored on chest and belly, cream-colored on throat; axillary region, groin, and anterior thigh pale orange; ventral surfaces of thighs light brown; iris gray, medially reddish, with black reticulations.	en	Acosta-Galvis, Andres R., Saldarriaga-Gomez, Ana M., Ramirez, Beatriz, Vargas-Ramirez, Mario (2020): A new Terrarana frog of genus Pristimantis from an unexplored cloud forest from the eastern Andes, Colombia. ZooKeys 961: 129-156, DOI: http://dx.doi.org/10.3897/zookeys.961.51971, URL: http://dx.doi.org/10.3897/zookeys.961.51971
0231086ADB3D509A9F8A15F37A8C8F83.taxon	distribution	Distribution and natural history. This species is only known from the type locality at an altitude between 2125 - 2160 m a. s. l. in an Andean and relictual cloud forest in the Casanare region on the eastern flank of the Cordillera Oriental of Colombia (Fig. 1). The locality is within the Cordillera Oriental montane forest ecoregion (sensu Dinerstein et al. 1995) in the Andean region (Middle Orobiome). The forest (Fig. 2) is unaffected by human activities and is typified by a canopt of medium-height (up to 20 m). The annual precipitation is between 4600 and 5600 mm with bimodal seasonality. Specimens were found active during the second annual rainy season (August to November) at a temperature of 14 ° C resting on mosses and lower leaves of shrubs and ferns in the undergrowth. Pristimantis chamezensis is syntopic with an undescribed species of genus Pristimantis.	en	Acosta-Galvis, Andres R., Saldarriaga-Gomez, Ana M., Ramirez, Beatriz, Vargas-Ramirez, Mario (2020): A new Terrarana frog of genus Pristimantis from an unexplored cloud forest from the eastern Andes, Colombia. ZooKeys 961: 129-156, DOI: http://dx.doi.org/10.3897/zookeys.961.51971, URL: http://dx.doi.org/10.3897/zookeys.961.51971
0231086ADB3D509A9F8A15F37A8C8F83.taxon	etymology	Etymology. The specific epithet is derived from the Municipality of Chameza, a geopolitical area where the type locality is located. We decided on this name using a citizen science approach, where expert scientists and local people met and discussed a list of possible names and their corresponding meanings. There was consensus on P. chamezensis as the preferred name.	en	Acosta-Galvis, Andres R., Saldarriaga-Gomez, Ana M., Ramirez, Beatriz, Vargas-Ramirez, Mario (2020): A new Terrarana frog of genus Pristimantis from an unexplored cloud forest from the eastern Andes, Colombia. ZooKeys 961: 129-156, DOI: http://dx.doi.org/10.3897/zookeys.961.51971, URL: http://dx.doi.org/10.3897/zookeys.961.51971
